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<mods:title>A New Species of Thomasomys (Rodentia: Muridae) from Eastern Ecuador, with Remarks on Mammalian Diversity and Biogeography in the Cordillera Oriental</mods:title>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4734860" ID-GBIF-Taxon="189547082" ID-Zenodo-Dep="4734860" LSID="urn:lsid:plazi:treatment:03FAB267FF97FFE9FCADFEACDB97DA47" httpUri="http://treatment.plazi.org/id/03FAB267FF97FFE9FCADFEACDB97DA47" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
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<paragraph blockId="33.[807,1119,318,341]" box="[807,1119,318,341]" pageId="33" pageNumber="34">
<heading box="[807,1119,318,341]" centered="true" fontSize="36" level="2" pageId="33" pageNumber="34" reason="8">
<taxonomicName authority="Thomas" authorityName="Thomas" authorityYear="1899" box="[807,1119,318,341]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="quichua">
<emphasis box="[807,1016,318,341]" italics="true" pageId="33" pageNumber="34">Coendou quichua</emphasis>
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<subSubSection id="C34950FAFF97FFEAFCADFEACDF1CDEFD" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8BEC0371FF97FFEAFCADFEACDF76DE20" blockId="33.[807,1119,318,341]" box="[807,1119,318,341]" pageId="33" pageNumber="34">
<heading id="D0A4B41DFF97FFEAFCADFEACDF76DE20" box="[807,1119,318,341]" centered="true" fontSize="36" level="2" pageId="33" pageNumber="34" reason="8">
<taxonomicName id="4C5378F2FF97FFEAFCADFEACDF76DE20" ID-CoL="C49RW" authority="Thomas" authorityName="Thomas" authorityYear="1899" box="[807,1119,318,341]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="quichua">
<emphasis id="B927DF63FF97FFEAFCADFEACD8D1DE20" box="[807,1016,318,341]" italics="true" pageId="33" pageNumber="34">Coendou quichua</emphasis>
Thomas
</taxonomicName>
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</paragraph>
<paragraph blockId="33.[699,1228,369,1741]" box="[728,1077,369,392]" pageId="33" pageNumber="34">SPECIMENS COLLECTED: None.</paragraph>
<paragraph id="8BEC0371FF97FFEAFD52FEE3DF1CDEFD" blockId="33.[699,1228,369,1741]" box="[728,1077,369,392]" pageId="33" pageNumber="34">SPECIMENS COLLECTED: None.</paragraph>
</subSubSection>
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<paragraph blockId="33.[699,1228,369,1741]" pageId="33" pageNumber="34">
<subSubSection id="C34950FAFF97FFE9FD52FE1CDB9DDE45" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="description">
<paragraph id="8BEC0371FF97FFEAFD52FE1CDFEEDD6E" blockId="33.[699,1228,369,1741]" pageId="33" pageNumber="34">
OTHER MATERIAL: I examined three NHRS specimens (A58/2822, A58/2962, A59/2962) collected by Ludovic Söderström in 1911 at ‘‘Tablon above Tumbaco’’ with recorded elevations of 9000–
<quantity box="[914,1022,516,539]" metricMagnitude="0" metricUnit="m" metricValue="3.3528" pageId="33" pageNumber="34" unit="ft" value="11.0">11,000 ft</quantity>
<quantity id="4CABAE94FF97FFEAFC18FD96D8D7DD6E" box="[914,1022,516,539]" metricMagnitude="0" metricUnit="m" metricValue="3.3528" pageId="33" pageNumber="34" unit="ft" value="11.0">11,000 ft</quantity>
[
<quantity box="[1038,1212,516,539]" metricMagnitude="3" metricUnit="m" metricValue="3.049" metricValueMax="3.354" metricValueMin="2.7439999999999998" pageId="33" pageNumber="34" unit="m" value="3049.0" valueMax="3354.0" valueMin="2744.0">2744–3354 m</quantity>
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].
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<paragraph id="8BEC0371FF97FFEAFD52FDB3DF07DC0E" blockId="33.[699,1228,369,1741]" pageId="33" pageNumber="34">
TAXONOMY:
<taxonomicName authorityName="Thomas" authorityYear="1899" box="[880,1094,545,568]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="quichua">
<emphasis box="[880,1094,545,568]" italics="true" pageId="33" pageNumber="34">Coendou quichua</emphasis>
<taxonomicName id="4C5378F2FF97FFEAFCFAFDB3DF6FDD4D" authorityName="Thomas" authorityYear="1899" box="[880,1094,545,568]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="quichua">
<emphasis id="B927DF63FF97FFEAFCFAFDB3DF6FDD4D" box="[880,1094,545,568]" italics="true" pageId="33" pageNumber="34">Coendou quichua</emphasis>
</taxonomicName>
is a morphologically distinctive porcupine whose diagnostic characters were accurately described by
<bibRefCitation author="Thomas, O." box="[736,919,633,656]" pageId="33" pageNumber="34" pagination="152 - 155" refId="ref25154" refString="Thomas, O. 1899. On new small mammals from South America. Annals and Magazine of Natural History 7 (3): 152 - 155." type="journal article" year="1899">Thomas (1899)</bibRefCitation>
<bibRefCitation id="EFC27E80FF97FFEAFD6AFDEBD8BEDDE5" author="Thomas, O." box="[736,919,633,656]" pageId="33" pageNumber="34" pagination="152 - 155" refId="ref25154" refString="Thomas, O. 1899. On new small mammals from South America. Annals and Magazine of Natural History 7 (3): 152 - 155." type="journal article" year="1899">Thomas (1899)</bibRefCitation>
.
<bibRefCitation author="Cabrera, A." box="[930,1110,633,656]" pageId="33" pageNumber="34" pagination="309 - 732" refId="ref21669" refString="Cabrera, A. 1961. Catalogo de los mamiferos de America del Sur. Revista del Museo Argentino de Ciencias Naturales '' Bernardino Rivadavia' ' (Ciencias Zoologicas) 4 (2): 309 - 732." type="journal article" year="1961">Cabrera (1961)</bibRefCitation>
<bibRefCitation id="EFC27E80FF97FFEAFC28FDEBDF7FDDE5" author="Cabrera, A." box="[930,1110,633,656]" pageId="33" pageNumber="34" pagination="309 - 732" refId="ref21669" refString="Cabrera, A. 1961. Catalogo de los mamiferos de America del Sur. Revista del Museo Argentino de Ciencias Naturales '' Bernardino Rivadavia' ' (Ciencias Zoologicas) 4 (2): 309 - 732." type="journal article" year="1961">Cabrera (1961)</bibRefCitation>
, however, treated
<taxonomicName authorityName="Thomas" authorityYear="1899" box="[787,881,663,686]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="quichua">
<emphasis box="[787,881,663,686]" italics="true" pageId="33" pageNumber="34">quichua</emphasis>
<taxonomicName id="4C5378F2FF97FFEAFC99FD05D858DDDB" authorityName="Thomas" authorityYear="1899" box="[787,881,663,686]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="quichua">
<emphasis id="B927DF63FF97FFEAFC99FD05D858DDDB" box="[787,881,663,686]" italics="true" pageId="33" pageNumber="34">quichua</emphasis>
</taxonomicName>
as a subspecies of
<taxonomicName authority="Tschudi" authorityName="Tschudi" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bicolor">
<emphasis box="[1108,1227,663,686]" italics="true" pageId="33" pageNumber="34">C. bicolor</emphasis>
<taxonomicName id="4C5378F2FF97FFEAFBDEFD05D83EDDBE" authority="Tschudi" authorityName="Tschudi" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B927DF63FF97FFEAFBDEFD05DFE2DDDB" box="[1108,1227,663,686]" italics="true" pageId="33" pageNumber="34">C. bicolor</emphasis>
Tschudi
</taxonomicName>
without providing any justification for doing so. Although
<bibRefCitation author="Emmons, L. H." box="[981,1175,721,744]" pageId="33" pageNumber="34" refId="ref22285" refString="Emmons, L. H. 1990. Neotropical rainforest mammals: a field guide. Chicago: University of Chicago Press." type="book" year="1990">Emmons (1990)</bibRefCitation>
<bibRefCitation id="EFC27E80FF97FFEAFC5FFD43DFBEDD9D" author="Emmons, L. H." box="[981,1175,721,744]" pageId="33" pageNumber="34" refId="ref22285" refString="Emmons, L. H. 1990. Neotropical rainforest mammals: a field guide. Chicago: University of Chicago Press." type="book" year="1990">Emmons (1990)</bibRefCitation>
,
<bibRefCitation author="Albuja" pageId="33" pageNumber="34" pagination="163 - 203" refId="ref21159" refString="Albuja V. L. 1991. Lista de vertebrados del Ecuador: mamiferos. Revista Politecnica 16: 163 - 203." type="journal article" year="1991">Albuja (1991)</bibRefCitation>
<bibRefCitation id="EFC27E80FF97FFEAFB2EFD43D863DC70" author="Albuja" pageId="33" pageNumber="34" pagination="163 - 203" refId="ref21159" refString="Albuja V. L. 1991. Lista de vertebrados del Ecuador: mamiferos. Revista Politecnica 16: 163 - 203." type="journal article" year="1991">Albuja (1991)</bibRefCitation>
,
<bibRefCitation author="Tirira" box="[860,1019,750,773]" pageId="33" pageNumber="34" refId="ref25510" refString="Tirira S., D. 1999. Mamiferos del Ecuador [vol.] II. Quito: Museo de Zoologia, Pontifica Universidad Catolica del Ecuador." type="book" year="1999">Tirira (1999)</bibRefCitation>
<bibRefCitation id="EFC27E80FF97FFEAFCD6FD7CD8D2DC70" author="Tirira" box="[860,1019,750,773]" pageId="33" pageNumber="34" refId="ref25510" refString="Tirira S., D. 1999. Mamiferos del Ecuador [vol.] II. Quito: Museo de Zoologia, Pontifica Universidad Catolica del Ecuador." type="book" year="1999">Tirira (1999)</bibRefCitation>
, and
<bibRefCitation author="Alberico, M. &amp; V. Rojas-Diaz &amp; J. G. Moreno" pageId="33" pageNumber="34" pagination="595 - 612" refId="ref21101" refString="Alberico, M., V. Rojas-Diaz, and J. G. Moreno. 1999. Aporte sobre la taxonomia y distribucion de los puercoespines (Rodentia: Erethizontidae) en Colombia. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas, y Naturales 23 (supplemento especial): 595 - 612." type="journal article" year="1999">Alberico et al. (1999)</bibRefCitation>
<bibRefCitation id="EFC27E80FF97FFEAFBC9FD7CD804DC56" author="Alberico, M. &amp; V. Rojas-Diaz &amp; J. G. Moreno" pageId="33" pageNumber="34" pagination="595 - 612" refId="ref21101" refString="Alberico, M., V. Rojas-Diaz, and J. G. Moreno. 1999. Aporte sobre la taxonomia y distribucion de los puercoespines (Rodentia: Erethizontidae) en Colombia. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas, y Naturales 23 (supplemento especial): 595 - 612." type="journal article" year="1999">Alberico et al. (1999)</bibRefCitation>
have subsequently recognized that
<taxonomicName authorityName="Thomas" authorityYear="1899" box="[699,793,809,832]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="quichua">
<emphasis box="[699,793,809,832]" italics="true" pageId="33" pageNumber="34">quichua</emphasis>
<taxonomicName id="4C5378F2FF97FFEAFD31FCBBD830DC35" authorityName="Thomas" authorityYear="1899" box="[699,793,809,832]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="quichua">
<emphasis id="B927DF63FF97FFEAFD31FCBBD830DC35" box="[699,793,809,832]" italics="true" pageId="33" pageNumber="34">quichua</emphasis>
</taxonomicName>
is a valid species, some checklists (e.g.,
<bibRefCitation author="Woods, C. A." box="[774,937,838,861]" pageId="33" pageNumber="34" pagination="771 - 806" refId="ref25944" refString="Woods, C. A. 1993. Suborder Hystricognathi. In D. E. Wilson and D. M. Reeder (editors), Mammal species of the world, 2 nd ed.: 771 - 806. Washington, DC: Smithsonian Institution Press." type="book chapter" year="1993">Woods, 1993</bibRefCitation>
<bibRefCitation id="EFC27E80FF97FFEAFC8CFCD4D880DC28" author="Woods, C. A." box="[774,937,838,861]" pageId="33" pageNumber="34" pagination="771 - 806" refId="ref25944" refString="Woods, C. A. 1993. Suborder Hystricognathi. In D. E. Wilson and D. M. Reeder (editors), Mammal species of the world, 2 nd ed.: 771 - 806. Washington, DC: Smithsonian Institution Press." type="book chapter" year="1993">Woods, 1993</bibRefCitation>
) continue to treat this name as a synonym of
<taxonomicName box="[979,1063,868,891]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bicolor">
<emphasis box="[979,1063,868,891]" italics="true" pageId="33" pageNumber="34">bicolor</emphasis>
<taxonomicName id="4C5378F2FF97FFEAFC59FCF6DF0EDC0E" box="[979,1063,868,891]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B927DF63FF97FFEAFC59FCF6DF0EDC0E" box="[979,1063,868,891]" italics="true" pageId="33" pageNumber="34">bicolor</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="33.[699,1228,369,1741]" lastBlockId="34.[124,653,193,480]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<paragraph id="8BEC0371FF97FFE9FD52FC13DB9DDE45" blockId="33.[699,1228,369,1741]" lastBlockId="34.[124,653,193,480]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
To date, no rationale has been provided for the zoogeographically incoherent and morphologically divergent collection of taxa that
<bibRefCitation author="Cabrera, A." box="[699,878,985,1008]" pageId="33" pageNumber="34" pagination="309 - 732" refId="ref21669" refString="Cabrera, A. 1961. Catalogo de los mamiferos de America del Sur. Revista del Museo Argentino de Ciencias Naturales '' Bernardino Rivadavia' ' (Ciencias Zoologicas) 4 (2): 309 - 732." type="journal article" year="1961">Cabrera (1961)</bibRefCitation>
<bibRefCitation id="EFC27E80FF97FFEAFD31FC4BD847DC85" author="Cabrera, A." box="[699,878,985,1008]" pageId="33" pageNumber="34" pagination="309 - 732" refId="ref21669" refString="Cabrera, A. 1961. Catalogo de los mamiferos de America del Sur. Revista del Museo Argentino de Ciencias Naturales '' Bernardino Rivadavia' ' (Ciencias Zoologicas) 4 (2): 309 - 732." type="journal article" year="1961">Cabrera (1961)</bibRefCitation>
lumped together as
<taxonomicName class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bicolor">
<emphasis italics="true" pageId="33" pageNumber="34">Coendou bicolor</emphasis>
<taxonomicName id="4C5378F2FF97FFEAFBE8FC4BD826DB7B" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B927DF63FF97FFEAFBE8FC4BD826DB7B" italics="true" pageId="33" pageNumber="34">Coendou bicolor</emphasis>
</taxonomicName>
. Although this name has been applied by authors to a wide range of morphologies, specimens collected in the vicinity of the Peruvian
<typeStatus box="[788,838,1102,1125]" pageId="33" pageNumber="34">type</typeStatus>
<typeStatus id="54E8BDD3FF97FFEAFC9EFBDCD86FDB10" box="[788,838,1102,1125]" pageId="33" pageNumber="34">type</typeStatus>
locality (e.g., AMNH 147500, FMNH 65799) are distinctively large porcupines (ca.
<quantity box="[852,951,1161,1184]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="34" unit="mm" value="900.0">900 mm</quantity>
<quantity id="4CABAE94FF97FFEAFCDEFB1BD89EDBD5" box="[852,951,1161,1184]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="34" unit="mm" value="900.0">900 mm</quantity>
total length) with tails that are almost as long as the combined length of head­and­body; the visible dorsal pelage consists entirely of bicolored (blacktipped) quills, of which those over the forequarters are conspicuously longer than those over the lower back and rump. By contrast,
<taxonomicName authorityName="Thomas" authorityYear="1899" box="[699,830,1367,1390]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="quichua">
<emphasis box="[699,830,1367,1390]" italics="true" pageId="33" pageNumber="34">C. quichua</emphasis>
<taxonomicName id="4C5378F2FF97FFEAFD31FAC5D817DA1B" authorityName="Thomas" authorityYear="1899" box="[699,830,1367,1390]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="quichua">
<emphasis id="B927DF63FF97FFEAFD31FAC5D817DA1B" box="[699,830,1367,1390]" italics="true" pageId="33" pageNumber="34">C. quichua</emphasis>
</taxonomicName>
is much smaller (ca.
<quantity box="[1093,1192,1366,1389]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="33" pageNumber="34" unit="mm" value="600.0">600 mm</quantity>
<quantity id="4CABAE94FF97FFEAFBCFFAC4DF81DA18" box="[1093,1192,1366,1389]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="33" pageNumber="34" unit="mm" value="600.0">600 mm</quantity>
or less) with a proportionately much shorter tail (approximately half the length of head­andbody) and tricolored (pale­tipped) dorsal quills that are not conspicuously longer over the forequarters than on the lower back and rump. Cranially,
<taxonomicName authorityName="Thomas" authorityYear="1899" box="[909,1003,1543,1566]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="quichua">
<emphasis box="[909,1003,1543,1566]" italics="true" pageId="33" pageNumber="34">quichua</emphasis>
<taxonomicName id="4C5378F2FF97FFEAFC07F995D8C2D96B" authorityName="Thomas" authorityYear="1899" box="[909,1003,1543,1566]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="quichua">
<emphasis id="B927DF63FF97FFEAFC07F995D8C2D96B" box="[909,1003,1543,1566]" italics="true" pageId="33" pageNumber="34">quichua</emphasis>
</taxonomicName>
has a proportionately narrower rostrum than
<taxonomicName box="[1041,1125,1572,1595]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bicolor">
<emphasis box="[1041,1125,1572,1595]" italics="true" pageId="33" pageNumber="34">bicolor</emphasis>
<taxonomicName id="4C5378F2FF97FFEAFB9BF9B6DF4CD94E" box="[1041,1125,1572,1595]" class="Mammalia" family="Erethizontidae" genus="Coendou" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B927DF63FF97FFEAFB9BF9B6DF4CD94E" box="[1041,1125,1572,1595]" italics="true" pageId="33" pageNumber="34">bicolor</emphasis>
</taxonomicName>
, smaller orbits, less expanded jugals, and less inflated frontal sinuses. Other relevant morphological comparisons will be provided in an upcoming generic revision (Voss, in prep.), but the characters given here together with other traits mentioned by
<bibRefCitation author="Emmons, L. H." box="[363,553,193,216]" pageId="34" pageNumber="35" refId="ref22285" refString="Emmons, L. H. 1990. Neotropical rainforest mammals: a field guide. Chicago: University of Chicago Press." type="book" year="1990">Emmons (1990)</bibRefCitation>
<bibRefCitation id="EFC27E80FF94FFE9FEE1FF53D900DFAD" author="Emmons, L. H." box="[363,553,193,216]" pageId="34" pageNumber="35" refId="ref22285" refString="Emmons, L. H. 1990. Neotropical rainforest mammals: a field guide. Chicago: University of Chicago Press." type="book" year="1990">Emmons (1990)</bibRefCitation>
and
<bibRefCitation author="Alberico, M. &amp; V. Rojas-Diaz &amp; J. G. Moreno" pageId="34" pageNumber="35" pagination="595 - 612" refId="ref21101" refString="Alberico, M., V. Rojas-Diaz, and J. G. Moreno. 1999. Aporte sobre la taxonomia y distribucion de los puercoespines (Rodentia: Erethizontidae) en Colombia. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas, y Naturales 23 (supplemento especial): 595 - 612." type="journal article" year="1999">Alberico et al. (1999)</bibRefCitation>
<bibRefCitation id="EFC27E80FF94FFE9FDECFF53DA4CDF80" author="Alberico, M. &amp; V. Rojas-Diaz &amp; J. G. Moreno" pageId="34" pageNumber="35" pagination="595 - 612" refId="ref21101" refString="Alberico, M., V. Rojas-Diaz, and J. G. Moreno. 1999. Aporte sobre la taxonomia y distribucion de los puercoespines (Rodentia: Erethizontidae) en Colombia. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas, y Naturales 23 (supplemento especial): 595 - 612." type="journal article" year="1999">Alberico et al. (1999)</bibRefCitation>
are sufficient for unambiguous identifications of these dissimilar taxa.
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="discussion">
<paragraph blockId="34.[124,653,193,480]" pageId="34" pageNumber="35">
<subSubSection id="C34950FAFF94FFE9FF13FEA4DAE5DD4A" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8BEC0371FF94FFE9FF13FEA4D947DE95" blockId="34.[124,653,193,480]" pageId="34" pageNumber="35">
REMARKS: The original specimen tag of NHRS A58/2822 notes that the animal was ‘‘found in the underbrush’’.
<bibRefCitation author="Lonnberg, E." box="[455,653,369,392]" pageId="34" pageNumber="35" pagination="1 - 36" refId="ref23677" refString="Lonnberg, E. 1913. Mammals from Ecuador and related forms. Arkiv for Zoologi 8 (16): 1 - 36, pl. 1." type="journal article" year="1913">Lönnberg (1913)</bibRefCitation>
<bibRefCitation id="EFC27E80FF94FFE9FE4DFEE3D9A4DEFD" author="Lonnberg, E." box="[455,653,369,392]" pageId="34" pageNumber="35" pagination="1 - 36" refId="ref23677" refString="Lonnberg, E. 1913. Mammals from Ecuador and related forms. Arkiv for Zoologi 8 (16): 1 - 36, pl. 1." type="journal article" year="1913">Lönnberg (1913)</bibRefCitation>
originally reported this material as having been collected above Tumbaco, without mentioning the actual collecting site (Tablón).
</paragraph>
<paragraph blockId="34.[180,597,523,575]" pageId="34" pageNumber="35">
<taxonomicName authority="(Stolzmann)" baseAuthorityName="Stolzmann" baseAuthorityYear="1865" class="Mammalia" family="Cuniculidae" genus="Cuniculus" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="taczanowskii" subGenus="Stictomys">
<emphasis box="[180,298,523,546]" italics="true" pageId="34" pageNumber="35">Cuniculus</emphasis>
<paragraph id="8BEC0371FF94FFE9FF3EFD99DAE5DD4A" blockId="34.[180,597,523,575]" pageId="34" pageNumber="35">
<taxonomicName id="4C5378F2FF94FFE9FF3EFD99DAE5DD4A" authority="(Stolzmann)" baseAuthorityName="Stolzmann" baseAuthorityYear="1865" class="Mammalia" family="Cuniculidae" genus="Cuniculus" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="taczanowskii" subGenus="Stictomys">
<emphasis id="B927DF63FF94FFE9FF3EFD99DA03DD57" box="[180,298,523,546]" italics="true" pageId="34" pageNumber="35">Cuniculus</emphasis>
(
<emphasis box="[317,428,523,546]" italics="true" pageId="34" pageNumber="35">Stictomys</emphasis>
<emphasis id="B927DF63FF94FFE9FEB7FD99DA85DD57" box="[317,428,523,546]" italics="true" pageId="34" pageNumber="35">Stictomys</emphasis>
)
<emphasis box="[447,597,523,546]" italics="true" pageId="34" pageNumber="35">taczanowskii</emphasis>
<emphasis id="B927DF63FF94FFE9FE35FD99D97CDD57" box="[447,597,523,546]" italics="true" pageId="34" pageNumber="35">taczanowskii</emphasis>
(Stolzmann)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="description">
<paragraph blockId="34.[124,653,603,1330]" box="[153,502,603,626]" pageId="34" pageNumber="35">SPECIMENS COLLECTED: None.</paragraph>
<paragraph blockId="34.[124,653,603,1330]" pageId="34" pageNumber="35">OTHER MATERIAL: A single specimen that I have not examined (QCAZ 954) was collected in the páramo near Paso de Guamaní by G. Onore in 1993 (D. Tirira, personal commun.).</paragraph>
<paragraph blockId="34.[124,653,603,1330]" pageId="34" pageNumber="35">
<subSubSection id="C34950FAFF94FFE9FF13FDC9DB97DA47" pageId="34" pageNumber="35" type="description">
<paragraph id="8BEC0371FF94FFE9FF13FDC9DADFDD07" blockId="34.[124,653,603,1330]" box="[153,502,603,626]" pageId="34" pageNumber="35">SPECIMENS COLLECTED: None.</paragraph>
<paragraph id="8BEC0371FF94FFE9FF13FDEBDBD2DC70" blockId="34.[124,653,603,1330]" pageId="34" pageNumber="35">OTHER MATERIAL: A single specimen that I have not examined (QCAZ 954) was collected in the páramo near Paso de Guamaní by G. Onore in 1993 (D. Tirira, personal commun.).</paragraph>
<paragraph id="8BEC0371FF94FFE9FF13FC99DB97DA47" blockId="34.[124,653,603,1330]" pageId="34" pageNumber="35">
TAXONOMY: The mountain paca is morphologically distinctive (
<bibRefCitation author="Thomas, O." box="[420,592,809,832]" pageId="34" pageNumber="35" pagination="237 - 238" refId="ref25484" refString="Thomas, O. 1924. Some notes on pacas. Annals and Magazine of Natural History 9 (13): 237 - 238." type="journal article" year="1924">Thomas, 1924</bibRefCitation>
<bibRefCitation id="EFC27E80FF94FFE9FE2EFCBBD979DC35" author="Thomas, O." box="[420,592,809,832]" pageId="34" pageNumber="35" pagination="237 - 238" refId="ref25484" refString="Thomas, O. 1924. Some notes on pacas. Annals and Magazine of Natural History 9 (13): 237 - 238." type="journal article" year="1924">Thomas, 1924</bibRefCitation>
) and was formerly distinguished generically (as
<emphasis box="[124,235,868,891]" italics="true" pageId="34" pageNumber="35">Stictomys</emphasis>
<emphasis id="B927DF63FF94FFE9FFF6FCF6DBC2DC0E" box="[124,235,868,891]" italics="true" pageId="34" pageNumber="35">Stictomys</emphasis>
) from the lowland paca (
<taxonomicName class="Mammalia" family="Cuniculidae" genus="Cuniculus" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="paca">
<emphasis italics="true" pageId="34" pageNumber="35">Cuniculus paca</emphasis>
<taxonomicName id="4C5378F2FF94FFE9FD9DFCF6DB9DDCED" class="Mammalia" family="Cuniculidae" genus="Cuniculus" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="paca">
<emphasis id="B927DF63FF94FFE9FD9DFCF6DB9DDCED" italics="true" pageId="34" pageNumber="35">Cuniculus paca</emphasis>
</taxonomicName>
). Although only a single species of mountain paca is currently recognized, no critical analysis of morphological or molecular data is currently available to test the hypothesis that
<taxonomicName baseAuthorityName="Stolzmann" baseAuthorityYear="1865" box="[283,470,1014,1037]" class="Mammalia" family="Cuniculidae" genus="Cuniculus" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="taczanowskii">
<emphasis box="[283,470,1014,1037]" italics="true" pageId="34" pageNumber="35">C. taczanowskii</emphasis>
<taxonomicName id="4C5378F2FF94FFE9FE91FC64DAFFDB78" baseAuthorityName="Stolzmann" baseAuthorityYear="1865" box="[283,470,1014,1037]" class="Mammalia" family="Cuniculidae" genus="Cuniculus" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="taczanowskii">
<emphasis id="B927DF63FF94FFE9FE91FC64DAFFDB78" box="[283,470,1014,1037]" italics="true" pageId="34" pageNumber="35">C. taczanowskii</emphasis>
</taxonomicName>
(from
<collectingCountry box="[556,653,1014,1037]" name="Ecuador" pageId="34" pageNumber="35">Ecuador</collectingCountry>
<collectingCountry id="F34443E1FF94FFE9FDA6FC64D9A4DB78" box="[556,653,1014,1037]" name="Ecuador" pageId="34" pageNumber="35">Ecuador</collectingCountry>
and
<collectingCountry box="[177,232,1043,1066]" name="Peru" pageId="34" pageNumber="35">Peru</collectingCountry>
<collectingCountry id="F34443E1FF94FFE9FF3BFB81DBC1DB5F" box="[177,232,1043,1066]" name="Peru" pageId="34" pageNumber="35">Peru</collectingCountry>
) is actually conspecific with populations from
<collectingCountry box="[288,408,1073,1096]" name="Venezuela" pageId="34" pageNumber="35">Venezuela</collectingCountry>
<collectingCountry id="F34443E1FF94FFE9FEAAFBA3DAB1DB3D" box="[288,408,1073,1096]" name="Venezuela" pageId="34" pageNumber="35">Venezuela</collectingCountry>
and
<collectingCountry box="[480,595,1073,1096]" name="Colombia" pageId="34" pageNumber="35">Colombia</collectingCountry>
<collectingCountry id="F34443E1FF94FFE9FE6AFBA3D97ADB3D" box="[480,595,1073,1096]" name="Colombia" pageId="34" pageNumber="35">Colombia</collectingCountry>
that were formerly known as
<taxonomicName baseAuthorityName="Thomas" baseAuthorityYear="1905" box="[426,545,1102,1125]" class="Mammalia" family="Cuniculidae" genus="Cuniculus" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="sierrae">
<emphasis box="[426,545,1102,1125]" italics="true" pageId="34" pageNumber="35">C. sierrae</emphasis>
<taxonomicName id="4C5378F2FF94FFE9FE20FBDCD908DB10" baseAuthorityName="Thomas" baseAuthorityYear="1905" box="[426,545,1102,1125]" class="Mammalia" family="Cuniculidae" genus="Cuniculus" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="sierrae">
<emphasis id="B927DF63FF94FFE9FE20FBDCD908DB10" box="[426,545,1102,1125]" italics="true" pageId="34" pageNumber="35">C. sierrae</emphasis>
</taxonomicName>
(e.g., by
<bibRefCitation author="Thomas, O." box="[124,299,1131,1154]" pageId="34" pageNumber="35" pagination="584 - 591" refId="ref25254" refString="Thomas, O. 1905. New Neotropical Molossus, Conepatus, Nectomys, Proechimys, and Agouti, with a note on the genus Mesomys. Annals and Magazine of Natural History 7 (15): 584 - 591." type="journal article" year="1905">Thomas, 1905</bibRefCitation>
<bibRefCitation id="EFC27E80FF94FFE9FFF6FBF9DA02DBF7" author="Thomas, O." box="[124,299,1131,1154]" pageId="34" pageNumber="35" pagination="584 - 591" refId="ref25254" refString="Thomas, O. 1905. New Neotropical Molossus, Conepatus, Nectomys, Proechimys, and Agouti, with a note on the genus Mesomys. Annals and Magazine of Natural History 7 (15): 584 - 591." type="journal article" year="1905">Thomas, 1905</bibRefCitation>
;
<bibRefCitation author="Krumbiegel, V. I." box="[316,537,1131,1154]" pageId="34" pageNumber="35" pagination="223 - 238" refId="ref23428" refString="Krumbiegel, V. I. 1940. Die Saugetiere der Sudamerika-Expeditionen Prof. Dr. Kriegs. 7. Pakas. Zoologische Anzeiger 132: 223 - 238." type="journal article" year="1940">Krumbiegel, 1940</bibRefCitation>
<bibRefCitation id="EFC27E80FF94FFE9FEB6FBF9D930DBF7" author="Krumbiegel, V. I." box="[316,537,1131,1154]" pageId="34" pageNumber="35" pagination="223 - 238" refId="ref23428" refString="Krumbiegel, V. I. 1940. Die Saugetiere der Sudamerika-Expeditionen Prof. Dr. Kriegs. 7. Pakas. Zoologische Anzeiger 132: 223 - 238." type="journal article" year="1940">Krumbiegel, 1940</bibRefCitation>
). The International Commission on Zoological Nomenclature (ICZN, 1998) recently ruled that
<taxonomicName authority="Brisson, 1762" authorityName="Brisson" authorityYear="1762" box="[124,419,1219,1243]" class="Mammalia" family="Cuniculidae" genus="Cuniculus" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis box="[124,242,1220,1243]" italics="true" pageId="34" pageNumber="35">Cuniculus</emphasis>
<bibRefCitation author="M. J. Brisson" box="[252,419,1219,1242]" pageId="34" pageNumber="35" pagination="64 - 71" refId="ref23181" refString="ICZN. 1998. Opinion 1894 [of the International Commission on Zoological Nomenclature]. Regnum animale..., ed. 2 (M. J. Brisson, 1762): rejected for nomenclatural purposes, with the conservation of the mammalian generic names Philander (Marsupialia), Pteropus (Chiroptera), Glis, Cuniculus, and Hydrochoerus (Rodentia), Meles, Lutra and Hyaena (Carnivora), Tapirus (Perissodactyla), Tragulus and Giraffa (Artiodactyla). Bulletin of Zoological Nomenclature 55: 64 - 71." type="journal article" year="1762">Brisson, 1762</bibRefCitation>
<taxonomicName id="4C5378F2FF94FFE9FFF6FB56DA8ADBAF" authority="Brisson, 1762" authorityName="Brisson" authorityYear="1762" box="[124,419,1219,1243]" class="Mammalia" family="Cuniculidae" genus="Cuniculus" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="B927DF63FF94FFE9FFF6FB56DBDBDBAE" box="[124,242,1220,1243]" italics="true" pageId="34" pageNumber="35">Cuniculus</emphasis>
<bibRefCitation id="EFC27E80FF94FFE9FF76FB51DA8ADBAF" author="M. J. Brisson" box="[252,419,1219,1242]" pageId="34" pageNumber="35" pagination="64 - 71" refId="ref23181" refString="ICZN. 1998. Opinion 1894 [of the International Commission on Zoological Nomenclature]. Regnum animale..., ed. 2 (M. J. Brisson, 1762): rejected for nomenclatural purposes, with the conservation of the mammalian generic names Philander (Marsupialia), Pteropus (Chiroptera), Glis, Cuniculus, and Hydrochoerus (Rodentia), Meles, Lutra and Hyaena (Carnivora), Tapirus (Perissodactyla), Tragulus and Giraffa (Artiodactyla). Bulletin of Zoological Nomenclature 55: 64 - 71." type="journal article" year="1762">Brisson, 1762</bibRefCitation>
</taxonomicName>
, is the oldest available name for pacas, previously referred by most American authors to
<taxonomicName authority="Lacepede, 1799" authorityName="Lacepede" authorityYear="1799" class="Mammalia" family="Cuniculidae" genus="Agouti" higherTaxonomySource="CoL" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis box="[446,524,1278,1301]" italics="true" pageId="34" pageNumber="35">Agouti</emphasis>
<taxonomicName id="4C5378F2FF94FFE9FE34FB6CDB93DA47" authority="Lacepede, 1799" authorityName="Lacepede" authorityYear="1799" class="Mammalia" family="Cuniculidae" genus="Agouti" higherTaxonomySource="CoL" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="B927DF63FF94FFE9FE34FB6CD925DA60" box="[446,524,1278,1301]" italics="true" pageId="34" pageNumber="35">Agouti</emphasis>
Lacépède, 1799
</taxonomicName>
.

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<mods:title>A New Species of Thomasomys (Rodentia: Muridae) from Eastern Ecuador, with Remarks on Mammalian Diversity and Biogeography in the Cordillera Oriental</mods:title>
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<mods:title id="319EEC48D9C0DF4EBDB42C31CAE918B2">A New Species of Thomasomys (Rodentia: Muridae) from Eastern Ecuador, with Remarks on Mammalian Diversity and Biogeography in the Cordillera Oriental</mods:title>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5605917" ID-GBIF-Taxon="189547089" ID-Zenodo-Dep="5605917" LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D" httpUri="http://treatment.plazi.org/id/03FAB267FFA3FFDEFF70FC51D884DB3D" lastPageNumber="22" pageId="21" pageNumber="22">
<subSubSection pageId="21" pageNumber="22" type="nomenclature">
<paragraph blockId="21.[250,524,963,986]" box="[250,524,963,986]" pageId="21" pageNumber="22">
<heading box="[250,524,963,986]" centered="true" fontSize="36" level="2" pageId="21" pageNumber="22" reason="8">
<taxonomicName authority="Thomas" authorityName="Thomas" authorityYear="1894" box="[250,524,963,986]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis box="[250,420,963,986]" italics="true" pageId="21" pageNumber="22">Akodon mollis</emphasis>
<treatment id="03FAB267FFA3FFDEFF70FC51D884DB3D" ID-DOI="http://doi.org/10.5281/zenodo.5605917" ID-GBIF-Taxon="189547089" ID-Zenodo-Dep="5605917" LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D" httpUri="http://treatment.plazi.org/id/03FAB267FFA3FFDEFF70FC51D884DB3D" lastPageNumber="22" pageId="21" pageNumber="22">
<subSubSection id="C34950FAFFA3FFDEFF70FC51DBCADBAE" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BEC0371FFA3FFDEFF70FC51D925DCAF" blockId="21.[250,524,963,986]" box="[250,524,963,986]" pageId="21" pageNumber="22">
<heading id="D0A4B41DFFA3FFDEFF70FC51D925DCAF" box="[250,524,963,986]" centered="true" fontSize="36" level="2" pageId="21" pageNumber="22" reason="8">
<taxonomicName id="4C5378F2FFA3FFDEFF70FC51D925DCAF" authority="Thomas" authorityName="Thomas" authorityYear="1894" box="[250,524,963,986]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis id="B927DF63FFA3FFDEFF70FC51DA8DDCAF" box="[250,420,963,986]" italics="true" pageId="21" pageNumber="22">Akodon mollis</emphasis>
Thomas
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="21.[123,651,1014,1741]" pageId="21" pageNumber="22">
<paragraph id="8BEC0371FFA3FFDEFF12FC64DBCADBAE" blockId="21.[123,651,1014,1741]" pageId="21" pageNumber="22">
SPECIMENS COLLECTED: 12.5 km W (by road) Papallacta,
<quantity box="[326,432,1044,1067]" metricMagnitude="0" metricUnit="m" metricValue="4.120896" pageId="21" pageNumber="22" unit="ft" value="13.52">13,520 ft</quantity>
<quantity id="4CABAE94FFA3FFDEFECCFB86DA99DB5E" box="[326,432,1044,1067]" metricMagnitude="0" metricUnit="m" metricValue="4.120896" pageId="21" pageNumber="22" unit="ft" value="13.52">13,520 ft</quantity>
(UMMZ 155597); 10.6 km W (by road) Papallacta,
<quantity box="[541,651,1073,1096]" metricMagnitude="0" metricUnit="m" metricValue="3.84048" pageId="21" pageNumber="22" unit="ft" value="12.6">12,600 ft</quantity>
<quantity id="4CABAE94FFA3FFDEFD97FBA3D9A2DB3D" box="[541,651,1073,1096]" metricMagnitude="0" metricUnit="m" metricValue="3.84048" pageId="21" pageNumber="22" unit="ft" value="12.6">12,600 ft</quantity>
(UMMZ 155596, 155777); 7.5 km W (by road) Papallacta,
<quantity box="[327,433,1132,1155]" metricMagnitude="0" metricUnit="m" metricValue="3.6576" pageId="21" pageNumber="22" unit="ft" value="12.0">12,000 ft</quantity>
<quantity id="4CABAE94FFA3FFDEFECDFBFEDA98DBF6" box="[327,433,1132,1155]" metricMagnitude="0" metricUnit="m" metricValue="3.6576" pageId="21" pageNumber="22" unit="ft" value="12.0">12,000 ft</quantity>
(UMMZ 155591– 155595); 6.2 km W (by road) Papallacta,
<quantity box="[123,229,1190,1213]" metricMagnitude="0" metricUnit="m" metricValue="3.56616" pageId="21" pageNumber="22" unit="ft" value="11.7">11,700 ft</quantity>
<quantity id="4CABAE94FFA3FFDEFFF1FB34DBCCDBC8" box="[123,229,1190,1213]" metricMagnitude="0" metricUnit="m" metricValue="3.56616" pageId="21" pageNumber="22" unit="ft" value="11.7">11,700 ft</quantity>
(UMMZ 155586–155590, 155778– 155780).
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="description">
<paragraph blockId="21.[123,651,1014,1741]" pageId="21" pageNumber="22">
<materialsCitation ID-GBIF-Occurrence="3395013323" collectionCode="OTHER, MATERIAL" collectorName="L. Soderstrom " county="Forty-two" elevation="3" location="Three" municipality="Mt. Antisana" pageId="21" pageNumber="22" specimenCode="AMNH 47156-47164, AMNH 66450 -66491" specimenCount="1">
<collectionCode box="[152,356,1249,1272]" pageId="21" pageNumber="22">OTHER MATERIAL</collectionCode>
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<paragraph id="8BEC0371FFA3FFDEFF12FB73D926D9C5" blockId="21.[123,651,1014,1741]" pageId="21" pageNumber="22">
<materialsCitation id="3B3B092CFFA3FFDEFF12FB73DAFEDA18" ID-GBIF-Occurrence="3395013323" collectionCode="OTHER, MATERIAL" collectorName="L. Soderstrom " county="Forty-two" elevation="3" location="Three" municipality="Mt. Antisana" pageId="21" pageNumber="22" specimenCode="AMNH 47156-47164, AMNH 66450 -66491" specimenCount="1">
<collectionCode id="ED429BB4FFA3FFDEFF12FB73DA4DDB82" box="[152,356,1249,1272]" pageId="21" pageNumber="22">OTHER MATERIAL</collectionCode>
:
<location LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFEF2FB73DA95DB8D" box="[376,444,1249,1272]" county="Forty-two" municipality="Mt. Antisana" name="Three" pageId="21" pageNumber="22">Three</location>
<location id="8E8C55AAFFA3FFDEFEF2FB73DA95DB8D" LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFEF2FB73DA95DB8D" box="[376,444,1249,1272]" county="Forty-two" municipality="Mt. Antisana" name="Three" pageId="21" pageNumber="22">Three</location>
additional specimens were collected at ‘‘
<location LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFE2AFB6CDAD0DA60" box="[416,505,1278,1301]" county="Forty-two" municipality="Mt. Antisana" name="Tablon" pageId="21" pageNumber="22">Tablon</location>
<location id="8E8C55AAFFA3FFDEFE2AFB6CDAD0DA60" LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFE2AFB6CDAD0DA60" box="[416,505,1278,1301]" county="Forty-two" municipality="Mt. Antisana" name="Tablon" pageId="21" pageNumber="22">Tablon</location>
, road to
<location LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFDEEFB6CDBC5DA46" county="Forty-two" municipality="Mt. Antisana" name="Papallacta'" pageId="21" pageNumber="22">Papallacta’</location>
’ with recorded elevations of
<quantity box="[123,325,1337,1360]" metricMagnitude="0" metricUnit="m" metricValue="3.3528" metricValueMax="3.5052" metricValueMin="3.2003999999999997" pageId="21" pageNumber="22" unit="ft" value="11.0" valueMax="11.5" valueMin="10.5">
<elevation box="[123,325,1337,1360]" metricMagnitude="0" metricUnit="m" metricValue="3.3528" metricValueMax="3.5052" metricValueMin="3.2003999999999997" pageId="21" pageNumber="22" unit="ft" value="11.0" valueMax="11.5" valueMin="10.5">10,500–11,500 ft</elevation>
</quantity>
[
<quantity box="[340,509,1337,1360]" metricMagnitude="3" metricUnit="m" metricValue="3.3525" metricValueMax="3.505" metricValueMin="3.2" pageId="21" pageNumber="22" unit="m" value="3352.5" valueMax="3505.0" valueMin="3200.0">
<elevation box="[340,509,1337,1360]" metricMagnitude="3" metricUnit="m" metricValue="3.3525" metricValueMax="3.505" metricValueMin="3.2" pageId="21" pageNumber="22" unit="m" value="3352.5" valueMax="3505.0" valueMin="3200.0">3200–3505 m</elevation>
<location id="8E8C55AAFFA3FFDEFDEEFB6CDBC5DA46" LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFDEEFB6CDBC5DA46" county="Forty-two" municipality="Mt. Antisana" name="Papallacta'" pageId="21" pageNumber="22">Papallacta’</location>
’ with recorded elevations of 10,500–11,500 ft [
<quantity id="4CABAE94FFA3FFDEFEDEFAABDAD4DA25" box="[340,509,1337,1360]" metricMagnitude="3" metricUnit="m" metricValue="3.3525" metricValueMax="3.505" metricValueMin="3.2" pageId="21" pageNumber="22" unit="m" value="3352.5" valueMax="3505.0" valueMin="3200.0">
<elevation id="007EE442FFA3FFDEFEDEFAABDAD4DA25" box="[340,509,1337,1360]" metricMagnitude="3" metricUnit="m" metricValue="3.3525" metricValueMax="3.505" metricValueMin="3.2" pageId="21" pageNumber="22" unit="m" value="3352.5" valueMax="3505.0" valueMin="3200.0">3200–3505 m</elevation>
</quantity>
] by
<collectorName pageId="21" pageNumber="22">L. Söderström</collectorName>
in 1903 and 1913.
<location LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFE62FAC4D96EDA18" box="[488,583,1366,1389]" county="Forty-two" municipality="Mt. Antisana" name="Another" pageId="21" pageNumber="22">Another</location>
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<collectorName id="26A666A7FFA3FFDEFDB9FAABDBFDDA18" pageId="21" pageNumber="22">L. Söderström</collectorName>
in 1903 and 1913.
</materialsCitation>
<materialsCitation id="3B3B092CFFA3FFDEFE62FAC4D920DAB0" ID-GBIF-Occurrence="3395013323" collectionCode="OTHER, MATERIAL" collectorName="L. Soderstrom " county="Forty-two" elevation="3" location="Three" municipality="Mt. Antisana" pageId="21" pageNumber="22" specimenCode="AMNH 47156-47164, AMNH 66450 -66491" specimenCount="1">
<location id="8E8C55AAFFA3FFDEFE62FAC4D96EDA18" LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFE62FAC4D96EDA18" box="[488,583,1366,1389]" county="Forty-two" municipality="Mt. Antisana" name="Another" pageId="21" pageNumber="22">Another</location>
<specimenCount id="9D55C8F8FFA3FFDEFDD3FAC4DBDDDAFE" pageId="21" pageNumber="22" type="generic">nine specimens</specimenCount>
(
<specimenCode box="[263,528,1396,1419]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH 47156–47164</specimenCode>
<specimenCode id="DBF5AB0AFFA3FFDEFE8DFAE6D939DAFE" box="[263,528,1396,1419]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH 47156–47164</specimenCode>
) taken by the same collector in 1914 are labeled ‘‘
<collectingMunicipality pageId="21" pageNumber="22">Mt. Antisana</collectingMunicipality>
<collectingMunicipality id="6B88990BFFA3FFDEFDD7FA03DBCBDAB0" pageId="21" pageNumber="22">Mt. Antisana</collectingMunicipality>
12000 feet’’ [
<quantity box="[407,510,1454,1477]" metricMagnitude="3" metricUnit="m" metricValue="3.658" pageId="21" pageNumber="22" unit="m" value="3658.0">
<elevation box="[407,510,1454,1477]" metricMagnitude="3" metricUnit="m" metricValue="3.658" pageId="21" pageNumber="22" unit="m" value="3658.0">3658 m</elevation>
<quantity id="4CABAE94FFA3FFDEFE1DFA3CDAD7DAB0" box="[407,510,1454,1477]" metricMagnitude="3" metricUnit="m" metricValue="3.658" pageId="21" pageNumber="22" unit="m" value="3658.0">
<elevation id="007EE442FFA3FFDEFE1DFA3CDAD7DAB0" box="[407,510,1454,1477]" metricMagnitude="3" metricUnit="m" metricValue="3.658" pageId="21" pageNumber="22" unit="m" value="3658.0">3658 m</elevation>
</quantity>
].
<collectingCounty box="[535,651,1454,1477]" pageId="21" pageNumber="22">Forty­two</collectingCounty>
].
</materialsCitation>
<materialsCitation id="3B3B092CFFA3FFDEFD9DFA3CDADDD968" ID-GBIF-Occurrence="3395013323" collectionCode="OTHER, MATERIAL" collectorName="L. Soderstrom " county="Forty-two" elevation="3" location="Three" municipality="Mt. Antisana" pageId="21" pageNumber="22" specimenCode="AMNH 47156-47164, AMNH 66450 -66491" specimenCount="1">
<collectingCounty id="628D7BFDFFA3FFDEFD9DFA3CD9A2DAB0" box="[535,651,1454,1477]" pageId="21" pageNumber="22">Forty­two</collectingCounty>
more specimens (
<specimenCode box="[356,642,1484,1507]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH 66450 –66491</specimenCode>
<specimenCode id="DBF5AB0AFFA3FFDEFEEEFA5ED9ABDA96" box="[356,642,1484,1507]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH 66450 –66491</specimenCode>
) were collected by
<collectorName box="[342,506,1513,1536]" pageId="21" pageNumber="22">H.E. Anthony</collectorName>
<collectorName id="26A666A7FFA3FFDEFEDCFA7BDAD3D975" box="[342,506,1513,1536]" pageId="21" pageNumber="22">H.E. Anthony</collectorName>
and
<collectorName pageId="21" pageNumber="22">G.H.H. Tate</collectorName>
<collectorName id="26A666A7FFA3FFDEFDBCFA7BDB85D968" pageId="21" pageNumber="22">G.H.H. Tate</collectorName>
between 4115 and
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<elevation box="[409,500,1542,1565]" metricMagnitude="3" metricUnit="m" metricValue="4.57" pageId="21" pageNumber="22" unit="m" value="4570.0">4570 m</elevation>
<quantity id="4CABAE94FFA3FFDEFE13F994DADDD968" box="[409,500,1542,1565]" metricMagnitude="3" metricUnit="m" metricValue="4.57" pageId="21" pageNumber="22" unit="m" value="4570.0">
<elevation id="007EE442FFA3FFDEFE13F994DADDD968" box="[409,500,1542,1565]" metricMagnitude="3" metricUnit="m" metricValue="4.57" pageId="21" pageNumber="22" unit="m" value="4570.0">4570 m</elevation>
</quantity>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3395013309" collectionCode="AMNH" pageId="21" pageNumber="22" specimenCode="AMNH 67330, 67332, 67334" specimenCount="1">
<materialsCitation id="3B3B092CFFA3FFDEFE77F994D9A2D92D" ID-GBIF-Occurrence="3395013309" collectionCode="AMNH" pageId="21" pageNumber="22" specimenCode="AMNH 67330, 67332, 67334" specimenCount="1">
on Antisana in 1923. Lastly,
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<specimenCount id="9D55C8F8FFA3FFDEFEDFF9B6D931D94E" box="[341,536,1572,1595]" pageId="21" pageNumber="22" type="generic">three specimens</specimenCount>
(
<specimenCode collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH 67330</specimenCode>
<specimenCode id="DBF5AB0AFFA3FFDEFDBAF9B6DBEED92D" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH 67330</specimenCode>
,
<specimenCode box="[215,291,1601,1624]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">67332</specimenCode>
<specimenCode id="DBF5AB0AFFA3FFDEFF5DF9D3DA0AD92D" box="[215,291,1601,1624]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">67332</specimenCode>
,
<specimenCode box="[306,381,1601,1624]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">67334</specimenCode>
<specimenCode id="DBF5AB0AFFA3FFDEFEB8F9D3DA54D92D" box="[306,381,1601,1624]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">67334</specimenCode>
) collected in 1924 by
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3395013324" collectionCode="R" county="Papallacta'" elevation="4" location="Olalla" municipality="Mt. Guamani'" pageId="21" pageNumber="22" specimenCount="1">
<collectionCode box="[123,146,1630,1653]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="22">R</collectionCode>
<materialsCitation id="3B3B092CFFA3FFDEFFF1F9CCD922D9C5" ID-GBIF-Occurrence="3395013324" collectionCode="R" county="Papallacta'" elevation="4" location="Olalla" municipality="Mt. Guamani'" pageId="21" pageNumber="22" specimenCount="1">
<collectionCode id="ED429BB4FFA3FFDEFFF1F9CCDBBBD900" box="[123,146,1630,1653]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="22">R</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFF15F9CCDBCFD900" box="[159,230,1630,1653]" county="Papallacta'" municipality="Mt. Guamani'" name="Olalla" pageId="21" pageNumber="22">Olalla</location>
<location id="8E8C55AAFFA3FFDEFF15F9CCDBCFD900" LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFF15F9CCDBCFD900" box="[159,230,1630,1653]" county="Papallacta'" municipality="Mt. Guamani'" name="Olalla" pageId="21" pageNumber="22">Olalla</location>
are labeled ‘‘
<collectingMunicipality box="[390,555,1630,1653]" pageId="21" pageNumber="22">Mt. Guamanı´</collectingMunicipality>
<collectingMunicipality id="6B88990BFFA3FFDEFE0CF9CCD902D900" box="[390,555,1630,1653]" pageId="21" pageNumber="22">Mt. Guamanı´</collectingMunicipality>
, road to
<collectingCounty box="[123,267,1660,1683]" pageId="21" pageNumber="22">Papallacta’</collectingCounty>
’ with recorded elevations of
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<elevation box="[123,327,1689,1712]" metricMagnitude="0" metricUnit="m" metricValue="3.81" metricValueMax="3.9624" metricValueMin="3.6576" pageId="21" pageNumber="22" unit="ft" value="12.5" valueMax="13.0" valueMin="12.0">12,000–13,000 ft</elevation>
</quantity>
[
<quantity box="[343,515,1689,1712]" metricMagnitude="3" metricUnit="m" metricValue="3.81" metricValueMax="3.9619999999999997" metricValueMin="3.658" pageId="21" pageNumber="22" unit="m" value="3810.0" valueMax="3962.0" valueMin="3658.0">
<elevation box="[343,515,1689,1712]" metricMagnitude="3" metricUnit="m" metricValue="3.81" metricValueMax="3.9619999999999997" metricValueMin="3.658" pageId="21" pageNumber="22" unit="m" value="3810.0" valueMax="3962.0" valueMin="3658.0">3658–3962 m</elevation>
<collectingCounty id="628D7BFDFFA3FFDEFFF1F9EEDA22D9E6" box="[123,267,1660,1683]" pageId="21" pageNumber="22">Papallacta’</collectingCounty>
’ with recorded elevations of 12,000–13,000 ft [
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<elevation id="007EE442FFA3FFDEFEDDF90BD92AD9C5" box="[343,515,1689,1712]" metricMagnitude="3" metricUnit="m" metricValue="3.81" metricValueMax="3.9619999999999997" metricValueMin="3.658" pageId="21" pageNumber="22" unit="m" value="3810.0" valueMax="3962.0" valueMin="3658.0">3658–3962 m</elevation>
</quantity>
]
</materialsCitation>
.
</paragraph>
<paragraph blockId="21.[123,651,1014,1741]" lastBlockId="21.[699,1228,193,1096]" pageId="21" pageNumber="22">
<paragraph id="8BEC0371FFA3FFDEFF12F924DF8EDD9D" blockId="21.[123,651,1014,1741]" lastBlockId="21.[699,1228,193,1096]" pageId="21" pageNumber="22">
TAXONOMY: The Papallacta material closely resembles the
<typeStatus box="[911,961,193,216]" pageId="21" pageNumber="22">type</typeStatus>
<typeStatus id="54E8BDD3FFA3FFDEFC05FF53D8E8DFAD" box="[911,961,193,216]" pageId="21" pageNumber="22">type</typeStatus>
of
<taxonomicName authorityName="Thomas" authorityYear="1913" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="subSpecies" species="mollis" subSpecies="altorum">
<emphasis italics="true" pageId="21" pageNumber="22">Akodon mollis altorum</emphasis>
<taxonomicName id="4C5378F2FFA3FFDEFC78FF53D829DF83" authorityName="Thomas" authorityYear="1913" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="subSpecies" species="mollis" subSpecies="altorum">
<emphasis id="B927DF63FFA3FFDEFC78FF53D829DF83" italics="true" pageId="21" pageNumber="22">Akodon mollis altorum</emphasis>
</taxonomicName>
as originally described by
<bibRefCitation author="Thomas, O." pageId="21" pageNumber="22" pagination="404 - 409" refId="ref25296" refString="Thomas, O. 1913. New forms of Akodon and Phyllotis, and a new genus for '' Akodon '' teguina. Annals and Magazine of Natural History 8 (11): 404 - 409." type="journal article" year="1913">Thomas (1913)</bibRefCitation>
<bibRefCitation id="EFC27E80FFA3FFDEFBE7FF4CD820DE66" author="Thomas, O." pageId="21" pageNumber="22" pagination="404 - 409" refId="ref25296" refString="Thomas, O. 1913. New forms of Akodon and Phyllotis, and a new genus for '' Akodon '' teguina. Annals and Magazine of Natural History 8 (11): 404 - 409." type="journal article" year="1913">Thomas (1913)</bibRefCitation>
based on a specimen collected at
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="21" pageNumber="22" unit="m" value="2600.0">2600 m</quantity>
<quantity id="4CABAE94FFA3FFDEFB1BFF6ED9FBDE45" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="21" pageNumber="22" unit="m" value="2600.0">2600 m</quantity>
in the Ecuadorean province of
<collectingRegion box="[1107,1178,281,304]" country="Ecuador" name="Canar" pageId="21" pageNumber="22">Cañar</collectingRegion>
<collectingRegion id="4997CD93FFA3FFDEFBD9FE8BDFB3DE45" box="[1107,1178,281,304]" country="Ecuador" name="Canar" pageId="21" pageNumber="22">Cañar</collectingRegion>
. No substantive analysis of character data, however, is available to support the current hypothesis (
<bibRefCitation author="Cabrera, A." box="[814,987,369,392]" pageId="21" pageNumber="22" pagination="309 - 732" refId="ref21669" refString="Cabrera, A. 1961. Catalogo de los mamiferos de America del Sur. Revista del Museo Argentino de Ciencias Naturales '' Bernardino Rivadavia' ' (Ciencias Zoologicas) 4 (2): 309 - 732." type="journal article" year="1961">Cabrera, 1961</bibRefCitation>
<bibRefCitation id="EFC27E80FFA3FFDEFCA4FEE3D8F2DEFD" author="Cabrera, A." box="[814,987,369,392]" pageId="21" pageNumber="22" pagination="309 - 732" refId="ref21669" refString="Cabrera, A. 1961. Catalogo de los mamiferos de America del Sur. Revista del Museo Argentino de Ciencias Naturales '' Bernardino Rivadavia' ' (Ciencias Zoologicas) 4 (2): 309 - 732." type="journal article" year="1961">Cabrera, 1961</bibRefCitation>
;
<bibRefCitation author="Musser, G. G. &amp; M. D. Carleton" pageId="21" pageNumber="22" pagination="501 - 755" refId="ref23955" refString="Musser, G. G., and M. D. Carleton. 1993. Family Muridae. In D. E. Wilson and D. M. Reeder (editors), Mammal species of the world, 2 nd ed.: 501 - 755. Washington, DC: Smithsonian Institution Press." type="book chapter" year="1993">Musser and Carleton, 1993</bibRefCitation>
<bibRefCitation id="EFC27E80FFA3FFDEFC66FEE3D805DED0" author="Musser, G. G. &amp; M. D. Carleton" pageId="21" pageNumber="22" pagination="501 - 755" refId="ref23955" refString="Musser, G. G., and M. D. Carleton. 1993. Family Muridae. In D. E. Wilson and D. M. Reeder (editors), Mammal species of the world, 2 nd ed.: 501 - 755. Washington, DC: Smithsonian Institution Press." type="book chapter" year="1993">Musser and Carleton, 1993</bibRefCitation>
) that
<emphasis box="[884,976,399,422]" italics="true" pageId="21" pageNumber="22">altorum</emphasis>
<emphasis id="B927DF63FFA3FFDEFCFEFE1DD8F9DED3" box="[884,976,399,422]" italics="true" pageId="21" pageNumber="22">altorum</emphasis>
, a highland taxon, is really conspecific with the geographically adjacent lowland forms
<taxonomicName authority="Thomas (1894)" authorityName="Thomas" authorityYear="1894" box="[963,1227,457,480]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis box="[963,1034,457,480]" italics="true" pageId="21" pageNumber="22">mollis</emphasis>
<bibRefCitation author="Thomas, O." box="[1045,1227,457,480]" pageId="21" pageNumber="22" pagination="346 - 366" refId="ref24959" refString="Thomas, O. 1894. Descriptions of some new Neotropical Muridae. Annals and Magazine of Natural History 6 (14): 346 - 366." type="journal article" year="1894">Thomas (1894)</bibRefCitation>
<taxonomicName id="4C5378F2FFA3FFDEFC49FE5BDFE2DE95" authority="Thomas (1894)" authorityName="Thomas" authorityYear="1894" box="[963,1227,457,480]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis id="B927DF63FFA3FFDEFC49FE5BDF23DE95" box="[963,1034,457,480]" italics="true" pageId="21" pageNumber="22">mollis</emphasis>
<bibRefCitation id="EFC27E80FFA3FFDEFB9FFE5BDFE2DE95" author="Thomas, O." box="[1045,1227,457,480]" pageId="21" pageNumber="22" pagination="346 - 366" refId="ref24959" refString="Thomas, O. 1894. Descriptions of some new Neotropical Muridae. Annals and Magazine of Natural History 6 (14): 346 - 366." type="journal article" year="1894">Thomas (1894)</bibRefCitation>
</taxonomicName>
and
<taxonomicName authority="Hershkovitz (1940)" authorityName="Hershkovitz" authorityYear="1940" box="[750,1110,486,510]" class="Mammalia" family="Muridae" genus="Oligoryzomys" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="fulvescens">
<emphasis box="[750,869,487,510]" italics="true" pageId="21" pageNumber="22">fulvescens</emphasis>
<bibRefCitation author="Hershkovitz, P." box="[878,1110,486,509]" pageId="21" pageNumber="22" pagination="1 - 3" refId="ref22833" refString="Hershkovitz, P. 1940. Notes on the distribution of the akodont rodent, Akodon mollis, in Ecuador with a description of a new race. Occasional Papers of the Museum of Zoology, University of Michigan 418: 1 - 3." type="journal article" year="1940">Hershkovitz (1940)</bibRefCitation>
<taxonomicName id="4C5378F2FFA3FFDEFD64FE75DF7FDE88" authority="Hershkovitz (1940)" authorityName="Hershkovitz" authorityYear="1940" box="[750,1110,486,510]" class="Mammalia" family="Muridae" genus="Oligoryzomys" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="fulvescens">
<emphasis id="B927DF63FFA3FFDEFD64FE75D84CDE8B" box="[750,869,487,510]" italics="true" pageId="21" pageNumber="22">fulvescens</emphasis>
<bibRefCitation id="EFC27E80FFA3FFDEFCE4FE74DF7FDE88" author="Hershkovitz, P." box="[878,1110,486,509]" pageId="21" pageNumber="22" pagination="1 - 3" refId="ref22833" refString="Hershkovitz, P. 1940. Notes on the distribution of the akodont rodent, Akodon mollis, in Ecuador with a description of a new race. Occasional Papers of the Museum of Zoology, University of Michigan 418: 1 - 3." type="journal article" year="1940">Hershkovitz (1940)</bibRefCitation>
</taxonomicName>
. As noted by
<bibRefCitation author="Myers, P. &amp; J. L. Patton" box="[739,1042,516,539]" pageId="21" pageNumber="22" pagination="1 - 35" refId="ref24077" refString="Myers, P., and J. L. Patton. 1989. Akodon of Peru and Bolivia - revision of the fumeus group (Rodentia: Sigmodontinae). Occasional Papers of the Museum of Zoology, University of Michigan 721: 1 - 35." type="journal article" year="1989">Myers and Patton (1989)</bibRefCitation>
<bibRefCitation id="EFC27E80FFA3FFDEFD69FD96DF3BDD6E" author="Myers, P. &amp; J. L. Patton" box="[739,1042,516,539]" pageId="21" pageNumber="22" pagination="1 - 35" refId="ref24077" refString="Myers, P., and J. L. Patton. 1989. Akodon of Peru and Bolivia - revision of the fumeus group (Rodentia: Sigmodontinae). Occasional Papers of the Museum of Zoology, University of Michigan 721: 1 - 35." type="journal article" year="1989">Myers and Patton (1989)</bibRefCitation>
,
<taxonomicName authorityName="Thomas" authorityYear="1894" box="[1057,1227,516,539]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis box="[1057,1227,516,539]" italics="true" pageId="21" pageNumber="22">Akodon mollis</emphasis>
<taxonomicName id="4C5378F2FFA3FFDEFBABFD96DFE2DD6E" authorityName="Thomas" authorityYear="1894" box="[1057,1227,516,539]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis id="B927DF63FFA3FFDEFBABFD96DFE2DD6E" box="[1057,1227,516,539]" italics="true" pageId="21" pageNumber="22">Akodon mollis</emphasis>
</taxonomicName>
as currently recognized has a very large geographic range and exhibits substantial geographic variation. Given their conclusion that
<emphasis box="[699,781,633,656]" italics="true" pageId="21" pageNumber="22">fumeus</emphasis>
<bibRefCitation author="Thomas, O." box="[794,974,633,656]" pageId="21" pageNumber="22" pagination="125 - 143" refId="ref25218" refString="Thomas, O. 1902. On mammals from Cochabamba, Bolivia, and the region north of that place. Annals and Magazine of Natural History 7 (9): 125 - 143." type="journal article" year="1902">Thomas (1902)</bibRefCitation>
<emphasis id="B927DF63FFA3FFDEFD31FDEBD824DDE5" box="[699,781,633,656]" italics="true" pageId="21" pageNumber="22">fumeus</emphasis>
<bibRefCitation id="EFC27E80FFA3FFDEFC90FDEBD8E7DDE5" author="Thomas, O." box="[794,974,633,656]" pageId="21" pageNumber="22" pagination="125 - 143" refId="ref25218" refString="Thomas, O. 1902. On mammals from Cochabamba, Bolivia, and the region north of that place. Annals and Magazine of Natural History 7 (9): 125 - 143." type="journal article" year="1902">Thomas (1902)</bibRefCitation>
—another taxon formerly ranked as a subspecies or synonym of
<taxonomicName authorityName="Thomas" authorityYear="1894" box="[699,770,692,715]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis box="[699,770,692,715]" italics="true" pageId="21" pageNumber="22">mollis</emphasis>
<taxonomicName id="4C5378F2FFA3FFDEFD31FD26D82BDDBE" authorityName="Thomas" authorityYear="1894" box="[699,770,692,715]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis id="B927DF63FFA3FFDEFD31FD26D82BDDBE" box="[699,770,692,715]" italics="true" pageId="21" pageNumber="22">mollis</emphasis>
</taxonomicName>
—is a valid species, a comprehensive revision of this complex is long overdue.
</paragraph>
<paragraph blockId="21.[699,1228,193,1096]" pageId="21" pageNumber="22">
<materialsCitation ID-GBIF-Occurrence="3395013329" collectionCode="FIELD, OBSERVATIONS" elevation="3880" pageId="21" pageNumber="22" specimenCount="16">
<collectionCode box="[728,968,750,773]" pageId="21" pageNumber="22">FIELD OBSERVATIONS</collectionCode>
<paragraph id="8BEC0371FFA3FFDEFD52FD7CD884DB3D" blockId="21.[699,1228,193,1096]" pageId="21" pageNumber="22">
<materialsCitation id="3B3B092CFFA3FFDEFD52FD7CD880DB3D" ID-GBIF-Occurrence="3395013329" collectionCode="FIELD, OBSERVATIONS" elevation="3880" pageId="21" pageNumber="22" specimenCount="16">
<collectionCode id="ED429BB4FFA3FFDEFD52FD7CD8E1DC70" box="[728,968,750,773]" pageId="21" pageNumber="22">FIELD OBSERVATIONS</collectionCode>
: The
<specimenCount box="[1037,1195,750,773]" pageId="21" pageNumber="22" type="generic">16 specimens</specimenCount>
<specimenCount id="9D55C8F8FFA3FFDEFB87FD7CDF82DC70" box="[1037,1195,750,773]" pageId="21" pageNumber="22" type="generic">16 specimens</specimenCount>
of
<taxonomicName authorityName="Thomas" authorityYear="1894" box="[699,868,780,803]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis box="[699,868,780,803]" italics="true" pageId="21" pageNumber="22">Akodon mollis</emphasis>
<taxonomicName id="4C5378F2FFA3FFDEFD31FC9ED84DDC56" authorityName="Thomas" authorityYear="1894" box="[699,868,780,803]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis id="B927DF63FFA3FFDEFD31FC9ED84DDC56" box="[699,868,780,803]" italics="true" pageId="21" pageNumber="22">Akodon mollis</emphasis>
</taxonomicName>
that I collected near Papallacta in 1980 were trapped at elevations ranging from
<quantity box="[764,954,838,861]" metricMagnitude="3" metricUnit="m" metricValue="3.88" metricValueMax="4.16" metricValueMin="3.6" pageId="21" pageNumber="22" unit="m" value="3880.0" valueMax="4160.0" valueMin="3600.0">
<elevation box="[764,954,838,861]" metricMagnitude="3" metricUnit="m" metricValue="3.88" metricValueMax="4.16" metricValueMin="3.6" pageId="21" pageNumber="22" unit="m" value="3880.0" valueMax="4160.0" valueMin="3600.0">3600 to 4160 m</elevation>
<quantity id="4CABAE94FFA3FFDEFD76FCD4D893DC28" box="[764,954,838,861]" metricMagnitude="3" metricUnit="m" metricValue="3.88" metricValueMax="4.16" metricValueMin="3.6" pageId="21" pageNumber="22" unit="m" value="3880.0" valueMax="4160.0" valueMin="3600.0">
<elevation id="007EE442FFA3FFDEFD76FCD4D893DC28" box="[764,954,838,861]" metricMagnitude="3" metricUnit="m" metricValue="3.88" metricValueMax="4.16" metricValueMin="3.6" pageId="21" pageNumber="22" unit="m" value="3880.0" valueMax="4160.0" valueMin="3600.0">3600 to 4160 m</elevation>
</quantity>
. Of these, 13 were taken in the shrubby páramo/forest ecotone, and
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and the other (
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69
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@ -56,7 +56,7 @@ cf.
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<collectionCode id="ED429BB4FFA4FFD9FF13FAABDA44DA3A" box="[153,365,1337,1360]" pageId="18" pageNumber="19">OTHER MATERIAL</collectionCode>
:
<materialsCitation id="3B3B092CFFA4FFD9FE02FAABDAFDD94E" ID-GBIF-Occurrence="3395013320" collectionCode="AMNH, QCAZ" collectorName="H. E. Anthony" county="Baeza" elevation="4145" location="Cerro Antisana" municipality="Cuyuja" pageId="18" pageNumber="19" specimenCode="AMNH 66719, QCAZ 0638, QCAZ 0726" specimenCount="1">
<materialsCitation id="3B3B092CFFA4FFD9FE02FAABDA34DADD" ID-GBIF-Occurrence="3395013320" collectionCode="AMNH, QCAZ" collectorName="H. E. Anthony" county="Baeza" elevation="4145" location="Cerro Antisana" municipality="Cuyuja" pageId="18" pageNumber="19" specimenCode="AMNH 66719, QCAZ 0638, QCAZ 0726" specimenCount="1">
I examined a single specimen (
<specimenCode id="DBF5AB0AFFA4FFD9FF71FAC4DA80DA18" box="[251,425,1366,1389]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="19">AMNH 66719</specimenCode>
) that was collected by
@ -67,7 +67,9 @@ at
</quantity>
on
<collectorName id="26A666A7FFA4FFD9FD89FAE6DB99DADD" pageId="18" pageNumber="19">Cerro Antisana</collectorName>
in 1923.
in 1923.
</materialsCitation>
<materialsCitation id="3B3B092CFFA4FFD9FEADFA03DA8FD975" ID-GBIF-Occurrence="3395013320" collectionCode="AMNH, QCAZ" collectorName="H. E. Anthony" county="Baeza" elevation="4145" location="Cerro Antisana" municipality="Cuyuja" pageId="18" pageNumber="19" specimenCode="AMNH 66719, QCAZ 0638, QCAZ 0726" specimenCount="1">
<collectorName id="26A666A7FFA4FFD9FEADFA03DA40DADD" box="[295,361,1425,1448]" pageId="18" pageNumber="19">Other</collectorName>
material that I have not seen (
<collectorName id="26A666A7FFA4FFD9FF4DFA3CDA4ADAB0" box="[199,355,1454,1477]" pageId="18" pageNumber="19">Diego Tirira</collectorName>
@ -81,7 +83,10 @@ on
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(
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), and another from
),
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<materialsCitation id="3B3B092CFFA4FFD9FE3FFA7BDAFDD94E" ID-GBIF-Occurrence="3395013320" collectionCode="AMNH, QCAZ" collectorName="H. E. Anthony" county="Baeza" elevation="4145" location="Cerro Antisana" municipality="Cuyuja" pageId="18" pageNumber="19" specimenCode="AMNH 66719, QCAZ 0638, QCAZ 0726" specimenCount="1">
and another from
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<elevation id="007EE442FFA4FFD9FFF6F994DBFED968" box="[124,215,1542,1565]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="18" pageNumber="19" unit="m" value="2800.0">2800 m</elevation>
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82
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(Desmarest)
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<paragraph id="8BEC0371FFA4FFD9FF13FD85DADFDD5B" blockId="18.[124,653,535,881]" box="[153,502,535,558]" pageId="18" pageNumber="19">SPECIMENS COLLECTED: None.</paragraph>
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OTHER MATERIAL: A single specimen (FMNH 43291) was collected by R. Olalla at
<quantity id="4CABAE94FFA4FFD9FF15FDFDDBD6DDF3" box="[159,255,623,646]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="19" unit="m" value="4000.0">4000 m</quantity>
elevation on Cerro Antisana in 1934.
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TAXONOMY: The pampas cat genus
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<emphasis id="B927DF63FFA4FFD9FDDDFD38DBDBDDAB" italics="true" pageId="18" pageNumber="19">Lynchailurus</emphasis>
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was revised by
<bibRefCitation id="EFC27E80FFA4FFD9FE60FD55DBE4DD8E" author="Garcia-Perea, R." pageId="18" pageNumber="19" pagination="1 - 35" refId="ref22340" refString="Garcia-Perea, R. 1994. The pampas cat group (genus Lynchailurus Severtzov, 1858) (Carnivora: Felidae), a systematic and biogeographic review. American Museum Novitates 3096: 1 - 35." type="journal article" year="1994">García­Perea (1994)</bibRefCitation>
, who examined FMNH 43291 and identified the local population as
<taxonomicName id="4C5378F2FFA4FFD9FD99FC90DA80DC43" authority="Lonnberg (1913)" authorityName="Lonnberg" authorityYear="1913" class="Mammalia" family="Felidae" genus="Lynchailurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="pajeros" subSpecies="thomasi">
<emphasis id="B927DF63FFA4FFD9FD99FC90DBF1DC43" italics="true" pageId="18" pageNumber="19">L. pajeros thomasi</emphasis>
<bibRefCitation id="EFC27E80FFA4FFD9FF55FC8DDA80DC43" author="Lonnberg, E." box="[223,425,799,822]" pageId="18" pageNumber="19" pagination="1 - 36" refId="ref23677" refString="Lonnberg, E. 1913. Mammals from Ecuador and related forms. Arkiv for Zoologi 8 (16): 1 - 36, pl. 1." type="journal article" year="1913">Lönnberg (1913)</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="EFC27E80FFA4FFD9FE3EFC8DD9A4DC43" author="Wozencraft, W. C." box="[436,653,799,822]" pageId="18" pageNumber="19" pagination="279 - 348" refId="ref25994" refString="Wozencraft, W. C. 1993. Order Carnivora. In D. E. Wilson and D. M. Reeder (editors), Mammal species of the world, 2 nd ed.: 279 - 348. Washington, DC: Smithsonian Institution Press." type="book chapter" year="1993">Wozencraft (1993)</bibRefCitation>
included this taxon in the synonymy of
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<emphasis id="B927DF63FFA4FFD9FDD5FCAFDA06DC04" italics="true" pageId="18" pageNumber="19">Oncifelis colocolo</emphasis>
(Molina)
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73
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@ -56,11 +56,14 @@
<subSubSection id="C34950FAFFA5FFDFFD52F90BD9AEDDA1" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8BEC0371FFA5FFD8FD52F90BDF1CD9C5" blockId="19.[728,1227,1689,1741]" box="[728,1077,1689,1712]" pageId="19" pageNumber="20">SPECIMENS COLLECTED: None.</paragraph>
<paragraph id="8BEC0371FFA5FFDFFD52F924DA5CDD4B" blockId="19.[728,1227,1689,1741]" lastBlockId="20.[124,653,193,963]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
OTHER MATERIAL: Two specimens of the
<materialsCitation id="3B3B092CFFA2FFDFFFF6FF53DA58DD4B" ID-GBIF-Occurrence="3395013340" collectionCode="AMNH" collectorName="C. Cordier" elevation="4" location="New York" pageId="20" pageNumber="21" specimenCode="AMNH 70521, AMNH 149370" specimenCount="1" stateProvince="New York">
woolly tapir are known from Papallacta. The first (
OTHER MATERIAL:
<materialsCitation id="3B3B092CFFA5FFDFFC36F924DAF6DE3A" ID-GBIF-Occurrence="3395013340" collectionCode="AMNH" collectorName="C. Cordier" elevation="4" lastPageId="20" lastPageNumber="21" location="New York" pageId="19" pageNumber="20" specimenCode="AMNH 70521, AMNH 149370" specimenCount="1" stateProvince="New York">
Two specimens of the woolly tapir are known from Papallacta. The first (
<specimenCode id="DBF5AB0AFFA2FFDFFF35FF4DDA5CDF83" box="[191,373,223,246]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="20" pageNumber="21">AMNH 70521</specimenCode>
), consisting of a skull collected by the Olallas (a family of professional collectors) in 1925, is unaccompanied by other geographic data. The second (
), consisting of a skull collected by the Olallas (a family of professional collectors) in 1925, is unaccompanied by other geographic data.
</materialsCitation>
<materialsCitation id="3B3B092CFFA2FFDFFE79FEAADA58DD4B" ID-GBIF-Occurrence="3395013340" collectionCode="AMNH" collectorName="C. Cordier" elevation="4" location="New York" pageId="20" pageNumber="21" specimenCode="AMNH 70521, AMNH 149370" specimenCount="1" stateProvince="New York">
The second (
<specimenCode id="DBF5AB0AFFA2FFDFFF0EFEC4DA6BDE18" box="[132,322,342,365]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="20" pageNumber="21">AMNH 149370</specimenCode>
) is the skin and skull of an individual that was captured alive by
<collectorName id="26A666A7FFA2FFDFFDBCFEE6DB83DEDC" pageId="20" pageNumber="21">C. Cordier</collectorName>
@ -69,9 +72,7 @@ woolly tapir are known from Papallacta. The first (
Zoological Society in 1952; an index card in the
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collection archives indicates that this specimen was captured at
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<elevation id="007EE442FFA2FFDFFEF2FE7EDACADD76" box="[376,483,492,515]" metricMagnitude="0" metricUnit="m" metricValue="3.5052" pageId="20" pageNumber="21" unit="ft" value="11.5">11,500 ft</elevation>
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(
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<elevation id="007EE442FFA2FFDFFE79FE7ED978DD76" box="[499,593,492,515]" metricMagnitude="3" metricUnit="m" metricValue="3.505" pageId="20" pageNumber="21" unit="m" value="3505.0">3505 m</elevation>
@ -79,9 +80,7 @@ collection archives indicates that this specimen was captured at
), but
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reported the capture elevation as
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.
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90
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<mods:title id="300FF1779B0DC2E6F684D8B8463C2031">A New Species of Thomasomys (Rodentia: Muridae) from Eastern Ecuador, with Remarks on Mammalian Diversity and Biogeography in the Cordillera Oriental</mods:title>
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<emphasis id="B927DF63FFA7FFDAFC82FACCDF24DA00" box="[776,1037,1374,1397]" italics="true" pageId="17" pageNumber="18">Pseudalopex culpaeus</emphasis>
(Molina)
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<paragraph id="8BEC0371FFA7FFDAFD52FA03DF1CDADD" blockId="17.[699,1227,1425,1741]" box="[728,1077,1425,1448]" pageId="17" pageNumber="18">SPECIMENS COLLECTED: None.</paragraph>
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OTHER MATERIAL: I examined a single specimen (AMNH 66739) collected at
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elevation on Cerro Antisana by G.H.H. Tate in 1923. Another specimen (MNCN 3652), which I have not seen, was collected on Antisana by M. Jiménez de la Espada in 1865 (J. Barreiro, personal commun.).
</paragraph>
<paragraph id="8BEC0371FFA7FFD9FD52F9EEDBDCDE38" blockId="17.[699,1227,1425,1741]" lastBlockId="18.[124,654,193,451]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
TAXONOMY: There has been no comprehensive revision of the wolf­like canid taxa currently referred to this species. According to
<bibRefCitation id="EFC27E80FFA4FFD9FF14FF53DA41DFAD" author="Lonnberg, E." box="[158,360,193,216]" pageId="18" pageNumber="19" pagination="1 - 23" refId="ref23742" refString="Lonnberg, E. 1922. A third contribution to the mammalogy of Ecuador. Arkiv for Zoologi 14 (20): 1 - 23." type="journal article" year="1922">Lönnberg (1922)</bibRefCitation>
and
<bibRefCitation id="EFC27E80FFA4FFD9FE23FF53D974DFAD" author="Cabrera, A." box="[425,605,193,216]" pageId="18" pageNumber="19" pagination="1 - 307" refId="ref21625" refString="Cabrera, A. 1958 ['' 1957 '']. Catalogo de los mamiferos de America del Sur. Revista del Museo Argentino de Ciencias Naturales '' Bernardino Rivadavia' ' (Ciencias Zoologicas) 4 (1): 1 - 307." type="journal article" year="1958">Cabrera (1958)</bibRefCitation>
the local form is
<emphasis id="B927DF63FFA4FFD9FEB3FF4DDAAEDF83" box="[313,391,223,246]" italics="true" pageId="18" pageNumber="19">reissii</emphasis>
<bibRefCitation id="EFC27E80FFA4FFD9FE1CFF4CD9A1DF80" author="Hilzheimer, M." box="[406,648,222,245]" pageId="18" pageNumber="19" pagination="109 - 118" refId="ref23065" refString="Hilzheimer, M. 1906. Papio mundamensis, Felis deliensis, Canis reissii und andere neue Saugetiere. Zoologischer Anzeiger (Leipzig) 30: 109 - 118." type="journal article" year="1906">Hilzheimer (1906)</bibRefCitation>
, which was originally described from material collected on
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(
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,
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).
</paragraph>
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REMARKS: According to
<bibRefCitation id="EFC27E80FFA4FFD9FE68FEC6DA1DDEFD" author="Emerson, M. R. &amp; R. H. Johnson" pageId="18" pageNumber="19" refId="ref22243" refString="Emerson, M. R., and R. H. Johnson. 1960. An account of travels in Ecuador. Expedition to Ecuador 1960, general report: 6 - 25. Oxford: Oxford University Exploration Club." type="book" year="1960">Emerson and Johnson (1960)</bibRefCitation>
and
<bibRefCitation id="EFC27E80FFA4FFD9FEFCFEE3D93FDEFD" author="Black, J." box="[374,534,369,392]" pageId="18" pageNumber="19" pagination="25 - 52" refId="ref21519" refString="Black, J. 1982. Los paramos del Antisana. Revista Geografica 17: 25 - 52." type="journal article" year="1982">Black (1982)</bibRefCitation>
,
<taxonomicName id="4C5378F2FFA4FFD9FDAFFEE3DA02DED3" baseAuthorityName="Molina" baseAuthorityYear="1782" class="Mammalia" family="Canidae" genus="Pseudalopex" higherTaxonomySource="CoL" kingdom="Animalia" order="Carnivora" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="culpaeus">
<emphasis id="B927DF63FFA4FFD9FDAFFEE3DA02DED3" italics="true" pageId="18" pageNumber="19">Pseudalopex culpaeus</emphasis>
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is common in the páramo landscapes surrounding Cerro Antisana.
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@ -56,21 +56,29 @@ Anthony
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<paragraph id="8BEC0371FFABFFD6FD52FD43DFE2DCC0" blockId="29.[699,1228,721,1741]" pageId="29" pageNumber="30">
SPECIMENS COLLECTED: 6.2 km (by road) W Papallacta,
SPECIMENS COLLECTED:
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6.2 km (by road) W Papallacta,
<quantity id="4CABAE94FFABFFD6FCFBFD7CD8CBDC70" box="[881,994,750,773]" metricMagnitude="0" metricUnit="m" metricValue="3.56616" pageId="29" pageNumber="30" unit="ft" value="11.7">11,700 ft</quantity>
(UMMZ 155711); Río Papallacta valley [
(UMMZ 155711);
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Río Papallacta valley [
<quantity id="4CABAE94FFABFFD6FC4FFC9EDF09DC56" box="[965,1056,780,803]" metricMagnitude="3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="29" pageNumber="30" unit="km" value="4.0" valueMax="5.0" valueMin="3.0">3–5 km</quantity>
by trail NNW Papallacta],
<quantity id="4CABAE94FFABFFD6FCD4FCBBD8FFDC35" box="[862,982,809,832]" metricMagnitude="0" metricUnit="m" metricValue="3.38328" pageId="29" pageNumber="30" unit="ft" value="11.1">11,100 ft</quantity>
(UMMZ 155640– 155642, 155712, 155713); 1.4 km (by road) E Papallacta (AMNH 248283; UMMZ 155633–155639, 155709, 155710);
(UMMZ 155640– 155642, 155712, 155713);
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<materialsCitation id="3B3B092CFFABFFD6FC75FCD4DF7EDCED" pageId="29" pageNumber="30">1.4 km (by road) E Papallacta (AMNH 248283; UMMZ 155633–155639, 155709, 155710);</materialsCitation>
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<quantity id="4CABAE94FFABFFD6FBEAFC13DFB5DCED" box="[1120,1180,897,920]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="29" pageNumber="30" unit="km" value="9.0">9 km</quantity>
(by road) E Papallacta,
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(UMMZ 127133).
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<paragraph id="8BEC0371FFABFFD6FD52FC2ED8D2DCA6" blockId="29.[699,1228,721,1741]" box="[728,1019,956,979]" pageId="29" pageNumber="30">OTHER MATERIAL: None.</paragraph>
<paragraph id="8BEC0371FFABFFD6FD52FC4BD8ACDAB0" blockId="29.[699,1228,721,1741]" pageId="29" pageNumber="30">
<materialsCitation id="3B3B092CFFABFFD6FD52FC4BD8A8DAB0" ID-GBIF-Occurrence="3395013319" collectingDate="1924-06-10" collectionCode="AMNH" country="Ecuador" county="Papallacta" elevation="2591" latitude="-0.56666666" location="Olallas" longLatPrecision="1308" longitude="-77.63333" municipality="Sumaco" pageId="29" pageNumber="30" specimenCode="AMNH 68195" specimenCount="1" stateProvince="Napo">
TAXONOMY:
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<emphasis id="B927DF63FFABFFD6FCE4FC4BDF6DDC85" box="[878,1092,985,1008]" italics="true" pageId="29" pageNumber="30">Thomasomys erro</emphasis>
@ -125,11 +133,9 @@ of montane organisms on the upper slopes of
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are therefore likely to be ecologically disjunct from those in the vicinity of
<collectingCounty id="628D7BFDFFABFFD6FC8CFA3CD8A8DAB0" box="[774,897,1454,1477]" pageId="29" pageNumber="30">Papallacta</collectingCounty>
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.
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The
<location id="8E8C55AAFFABFFD6FC84FA5ED8ACDA96" LSID="urn:lsid:plazi:treatment:03FAB267FFABFFD4FC96FD12DAC6D9B8:8E8C55AAFFABFFD6FC84FA5ED8ACDA96" box="[782,901,1484,1507]" county="Papallacta" municipality="In" name="Papallacta" pageId="29" pageNumber="30">Papallacta</location>
specimens are the only additional material of
@ -152,9 +158,7 @@ all qualitative craniodental comparisons, the
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appears to be indistinguishable from the
<collectingCounty id="628D7BFDFFA8FFD5FF6DFAABDA77DA25" box="[231,350,1337,1360]" pageId="30" pageNumber="31">Papallacta</collectingCounty>
series and appears to represent the same taxon
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.
series and appears to represent the same taxon.
</paragraph>
<caption id="DF2C53F9FFA8FFD5FF1CFC2DDABADB51" ID-DOI="http://doi.org/10.5281/zenodo.4734956" ID-Zenodo-Dep="4734956" httpUri="https://zenodo.org/record/4734956/files/figure.png" pageId="30" pageNumber="31" startId="30.[150,188,959,980]" targetBox="[127,1230,190,939]" targetPageId="30">
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@ -219,6 +223,7 @@ from the Cordillera Oriental—are smaller than the nominotypical form but do no
</taxonomicName>
.
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<footnote id="E8481F7FFFA8FFD5FD59F9BBD836D9B8" pageId="30" pageNumber="31">
<paragraph id="8BEC0371FFA8FFD5FD59F9BBD836D9B8" blockId="30.[700,1229,1577,1741]" pageId="30" pageNumber="31">
<materialsCitation id="3B3B092CFFA8FFD5FD59F9BBD832D9B8" ID-GBIF-Occurrence="3395013322" collectionCode="AMNH" country="Colombia" county="Popayan in Departamento" elevation="3152" location="Pacific" municipality="Cordillera Occidental" pageId="30" pageNumber="31" specimenCode="AMNH 32436, AMNH 32417, 32419, 32421-32425, 32427, 32428, 32430, 32433-32435, 32437-32440" specimenCount="17" stateProvince="Cauca" typeStatus="holotype">
<superScript id="7C26AE39FFA8FFD5FD59F9BBD9F3D941" attach="right" box="[723,730,1577,1588]" fontSize="5" pageId="30" pageNumber="31">7</superScript>
@ -264,6 +269,7 @@ west of
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.
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<paragraph id="8BEC0371FFA9FFD4FEDAFF50DACCDE78" blockId="31.[159,615,194,269]" pageId="31" pageNumber="32">
TABLE 6

97
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Among the material collected near Papallacta is a previously unknown murid rodent belonging to the sigmodontine genus
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Coues. Formerly
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construed broadly to include southeastern Brazilian taxa (e.g., by
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<bibRefCitation id="EFC27E80FFB1FFCCFD69FA36D8A7DACE" author="Osgood, W. H." box="[739,910,1444,1467]" pageId="7" pageNumber="8" pagination="11 - 14" refId="ref24248" refString="Osgood, W. H. 1933. Two new rodents from Ar- gentina. Zoological Series of the Field Museum of Natural History 20: 11 - 14." type="journal article" year="1933">Osgood, 1933</bibRefCitation>
; Ellerman, 1940;
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;
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),
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has subsequently been restricted (
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<bibRefCitation id="EFC27E80FFB1FFCCFC54FA73DF4EDA8D" author="Voss, R. S." box="[990,1127,1505,1528]" pageId="7" pageNumber="8" pagination="1 - 44" refId="ref25692" refString="Voss, R. S. 1993. A revision of the Brazilian muroid rodent genus Delomys with remarks on '' thomasomyine' ' characters. American Museum Novitates 3073: 1 - 44." type="journal article" year="1993">Voss, 1993</bibRefCitation>
;
<bibRefCitation author="Gonzalez, E. M." pageId="7" pageNumber="8" pagination="1 - 12" refId="ref22558" refString="Gonzalez, E. M. 2000. Un nuevo genero de roedor sigmodontino de Argentina y Brasil (Mammalia: Rodentia: Sigmodontinae). Comunicaciones Zoologicas del Museo de Historia Natural de Montevideo 196: 1 - 12." type="journal article" year="2000">González, 2000</bibRefCitation>
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) to a smaller but still speciose group that is endemic to tropical Andean cloud forests from
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to
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. Apparently, the center of diversity for the genus includes eastern
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, where the new species described below may occur sympatrically with at least seven other congeners.
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<mods:identifier id="60CAA2CD88C4A93D3FFFFD076718DF55" type="ISSN">0003-0082</mods:identifier>
<mods:identifier id="282FABF2E08524D3CDE9E29E4ABFE954" type="Zenodo-Dep">4734917</mods:identifier>
<mods:classification id="597AABFCD0F04BD8B6E2F0E2B53E9257">journal article</mods:classification>
<mods:identifier id="176A6F6D1964D1F1903391AA0DF861F1" type="CLB-Dataset">3793</mods:identifier>
<mods:identifier id="C597FFFFAD4583FBA4C0C9B0D0CA2C01" type="DOI">10.1206/0003-0082(2003)421&lt;0001:ANSOTR&gt;2.0.CO;2</mods:identifier>
<mods:identifier id="4C074FA2645FF1A694B36AD9C978C070" type="GBIF-Dataset">f4c2f80e-4917-4b5b-9415-0ee6377973ce</mods:identifier>
<mods:identifier id="457AD7F5DE753FD1BFB4165E6E5717B6" type="ISSN">0003-0082</mods:identifier>
<mods:identifier id="209CDAFBA587EAB7AFB64DE49619A009" type="Zenodo-Dep">4734917</mods:identifier>
</mods:mods>
<treatment id="03FAB267FFBFFFC5FF24FF53DA07DBE6" ID-DOI="http://doi.org/10.5281/zenodo.5818480" ID-GBIF-Taxon="189547085" ID-Zenodo-Dep="5818480" LSID="urn:lsid:plazi:treatment:03FAB267FFBFFFC5FF24FF53DA07DBE6" httpUri="http://treatment.plazi.org/id/03FAB267FFBFFFC5FF24FF53DA07DBE6" lastPageId="14" lastPageNumber="14" pageId="9" pageNumber="10">
<subSubSection id="C34950FAFFBFFFC2FF24FF53DAE8DF80" pageId="9" pageNumber="10" type="nomenclature">
@ -88,19 +88,11 @@ TYPE
<date id="FFED25B1FFBFFFC2FE2AFEF8D96EDEF4" box="[416,583,362,385]" pageId="9" pageNumber="10" value="1980-04-26">
<collectingDate id="EFA9DC59FFBFFFC2FE2AFEF8D96EDEF4" box="[416,583,362,385]" pageId="9" pageNumber="10" value="1980-04-26">26 April 1980</collectingDate>
</date>
at an elevation of
<quantity id="4CABAE94FFBFFFC2FE85FE15DA53DEEB" box="[271,378,391,414]" metricMagnitude="0" metricUnit="m" metricValue="3.38328" pageId="9" pageNumber="10" unit="ft" value="11.1">
<elevation id="007EE442FFBFFFC2FE85FE15DA53DEEB" box="[271,378,391,414]" metricMagnitude="0" metricUnit="m" metricValue="3.38328" pageId="9" pageNumber="10" unit="ft" value="11.1">11,100 ft</elevation>
</quantity>
(
<quantity id="4CABAE94FFBFFFC2FE01FE15DACEDEEB" box="[395,487,391,414]" metricMagnitude="3" metricUnit="m" metricValue="3.3839999999999995" pageId="9" pageNumber="10" unit="m" value="3384.0">
at an elevation of 11,100 ft (
<elevation id="007EE442FFBFFFC2FE01FE15DACEDEEB" box="[395,487,391,414]" metricMagnitude="3" metricUnit="m" metricValue="3.3839999999999995" pageId="9" pageNumber="10" unit="m" value="3384.0">3384 m</elevation>
</quantity>
) in the valley of the
<location id="8E8C55AAFFBFFFC2FF58FE36DAA9DECE" LSID="urn:lsid:plazi:treatment:03FAB267FFBFFFC5FF24FF53DA07DBE6:8E8C55AAFFBFFFC2FF58FE36DAA9DECE" box="[210,384,420,443]" country="Ecuador" name="Rio Papallacta" pageId="9" pageNumber="10" stateProvince="Napo">Río Papallacta</location>
(ca.
<quantity id="4CABAE94FFBFFFC2FE4CFE36D90DDECE" box="[454,548,420,443]" metricMagnitude="3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="9" pageNumber="10" unit="km" value="4.0" valueMax="5.0" valueMin="3.0">3–5 km</quantity>
by trail NNW Papallacta),
(ca. 3–5 km by trail NNW Papallacta),
<collectingRegion id="4997CD93FFBFFFC2FED6FE50D93EDEAC" box="[348,535,450,473]" country="Ecuador" name="Napo" pageId="9" pageNumber="10">Provincia Napo</collectingRegion>
,
<collectingCountry id="F34443E1FFBFFFC2FDAFFE50D9AEDEAC" box="[549,647,450,473]" name="Ecuador" pageId="9" pageNumber="10">Ecuador</collectingCountry>
@ -207,14 +199,12 @@ at Tablón, in Provincia
</subSubSection>
<subSubSection id="C34950FAFFBFFFC2FF12FC40DBE8DB15" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BEC0371FFBFFFC2FF12FC40DBE8DB15" blockId="9.[123,652,274,1741]" pageId="9" pageNumber="10">
<materialsCitation id="3B3B092CFFBFFFC2FF12FC40DB97DB15" ID-GBIF-Occurrence="3395013308" elevation="3700" location="Pichincha and Napo" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Pichincha and Napo">
DISTRIBUTION: Known only from the crest of the Cordillera Oriental (between ca. 3400 and
<quantity id="4CABAE94FFBFFFC2FF24FB9CDA23DB50" box="[174,266,1038,1061]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="10" unit="m" value="3700.0">
<elevation id="007EE442FFBFFFC2FF24FB9CDA23DB50" box="[174,266,1038,1061]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="10" unit="m" value="3700.0">3700 m</elevation>
</quantity>
) just south of the equator in the Ecuadorean provinces of
<collectingRegion id="4997CD93FFBFFFC2FE59FBB9DB97DB15" isEnumeration="true" pageId="9" pageNumber="10">Pichincha and Napo</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
@ -323,8 +313,8 @@ and
</taxonomicName>
differ in other points of comparison.
</paragraph>
<footnote id="E8481F7FFFBDFFC0FD58FA6BDF7AD9B8" pageId="11" pageNumber="12">
<paragraph id="8BEC0371FFBDFFC0FD58FA6BDF7AD9B8" blockId="11.[699,1228,1529,1741]" pageId="11" pageNumber="12">
<materialsCitation id="3B3B092CFFBDFFC0FD58FA6BD88FD9F0" ID-GBIF-Occurrence="3395013330" collectionCode="FMNH, AMNH" collectorName="de Tama" country="Venezuela" location="Paramo de Tama" pageId="11" pageNumber="12" specimenCode="FMNH 18583, AMNH 143667, 143668" specimenCount="3" stateProvince="Norte de Santander" typeStatus="holotype">
<specimenCount id="9D55C8F8FFBDFFC0FD58FA6BD86DD978" box="[722,836,1529,1549]" pageId="11" pageNumber="12" type="generic">
<superScript id="7C26AE39FFBDFFC0FD58FA6BD9F0D971" attach="right" box="[722,729,1529,1540]" fontSize="5" pageId="11" pageNumber="12">3</superScript>
Specimens
@ -352,7 +342,6 @@ on the border between
<specimenCode id="DBF5AB0AFFBDFFC0FD31F9E0D878D9F0" box="[699,849,1650,1669]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="11" pageNumber="12">AMNH 143667</specimenCode>
,
<specimenCode id="DBF5AB0AFFBDFFC0FCD7F9E0D88FD9F0" box="[861,934,1650,1669]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="11" pageNumber="12">143668</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B3B092CFFBDFFC0FC3BF9E0DFEED9E8" ID-GBIF-Occurrence="3395013314" collectionCode="FMNH" collectorName="de Tama" country="Venezuela" isTailEnumeration="true" location="Paramo de Tama" pageId="11" pageNumber="12" specimenCode="FMNH 18563, 18565, 18566, 18576, 18580, 18584, 18587, 18591, 18593, 18595" specimenCount="1" stateProvince="Norte de Santander" typeStatus="holotype">
<specimenCode id="DBF5AB0AFFBDFFC0FC3BF9E0DF11D9F0" box="[945,1080,1650,1669]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="12" type="Museum">FMNH 18563</specimenCode>
@ -391,6 +380,7 @@ on the border between
</materialsCitation>
.
</paragraph>
</footnote>
<caption id="DF2C53F9FFBAFFC7FF1CFD5ADA05DCED" ID-DOI="http://doi.org/10.5281/zenodo.4734936" ID-Zenodo-Dep="4734936" httpUri="https://zenodo.org/record/4734936/files/figure.png" pageId="12" pageNumber="13" startId="12.[150,188,712,733]" targetBox="[125,653,187,691]" targetPageId="12">
<paragraph id="8BEC0371FFBAFFC7FF1CFD5ADA05DCED" blockId="12.[124,653,712,920]" pageId="12" pageNumber="13">
Fig. 8. Lateral mandibular views of
@ -524,6 +514,7 @@ that is consistently present in
</taxonomicName>
.
</paragraph>
<footnote id="E8481F7FFFBAFFC7FD59FA6BD897D9B8" pageId="12" pageNumber="13">
<paragraph id="8BEC0371FFBAFFC7FD59FA6BD897D9B8" blockId="12.[700,1229,1529,1741]" pageId="12" pageNumber="13">
<superScript id="7C26AE39FFBAFFC7FD59FA6BD9F3D971" attach="right" box="[723,730,1529,1540]" fontSize="5" pageId="12" pageNumber="13">4</superScript>
This angle is equivalent to the incisive index of
@ -538,6 +529,7 @@ are orthodont, whereas those of
</taxonomicName>
are opisthodont.
</paragraph>
</footnote>
<caption id="DF2C53F9FFBBFFC6FF1FFC7BD8E3DB3B" ID-DOI="http://doi.org/10.5281/zenodo.4734940" ID-Zenodo-Dep="4734940" httpUri="https://zenodo.org/record/4734940/files/figure.png" pageId="13" pageNumber="14" startId="13.[149,187,1001,1022]" targetBox="[151,1200,187,981]" targetPageId="13">
<paragraph id="8BEC0371FFBBFFC6FF1FFC7BD8E3DB3B" blockId="13.[123,1227,1001,1102]" pageId="13" pageNumber="14">
Fig. 9. Left upper and right lower molars of