Add updates up until 2024-09-09 14:39:38
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<mods:title id="28F1D7352B1C6DB8A311D5CB3BE7387F">The bees of Early Eocene Cambay amber (Hymenoptera: Apidae)</mods:title>
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<mods:namePart id="59890D85BB050C3C7CD0B7855B77A742">Engel, Michael S.</mods:namePart>
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<mods:affiliation id="B922CE2F36434127B78D89E12EFDA06C">Division of Entomology, Natural History Museum, and Department of Ecology & Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045, USA (msengel @ ku. edu; jaimeortega @ ku. edu). & Division of Invertebrate Zoology, American Museum of Natural History, Central Park West</mods:affiliation>
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<mods:namePart id="A050DFB255F0CB8E2A774AE5A3374CCE">Ortega-Blanco, Jaime</mods:namePart>
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<mods:affiliation id="B6E67088B63858508E8F680616727262">Division of Entomology, Natural History Museum, and Department of Ecology & Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045, USA (msengel @ ku. edu; jaimeortega @ ku. edu).</mods:affiliation>
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<mods:namePart id="E3F0AD0122B991730BC3C6EB74B87545">Nascimbene, Paul C.</mods:namePart>
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<mods:namePart id="3FE84227D2C8DA6977851FDDD4F64A3B">Singh, Hukam</mods:namePart>
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<mods:affiliation id="97FA4AD4FC2C357296CE5A22D67AF437">Birbal Sahni Institute of Palaeobotany, 53 University Road, Lucknow 226 007, India (hukams @ gmail. com).</mods:affiliation>
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<subSubSection id="C35765227578FFC5FE07FA9CFCE52279" box="[456,872,1313,1337]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8BF236A97578FFC5FE07FA9CFCE52279" blockId="2.[456,872,1249,1337]" box="[456,872,1313,1337]" pageId="2" pageNumber="3">
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<heading id="D0BA81C57578FFC5FE07FA9CFCE52279" box="[456,872,1313,1337]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="3" reason="2">
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<taxonomicName id="4C4D4D2A7578FFC5FE07FA9CFDDB2279" authority="Cockerell" authorityName="Cockerell" authorityYear="1908" box="[456,598,1313,1337]" class="Insecta" family="Apidae" genus="Protobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B939EABB7578FFC5FE07FA9CFDDB2279" box="[456,598,1313,1337]" italics="true" pageId="2" pageNumber="3">Protobombus</emphasis>
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</taxonomicName>
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? species indeterminate
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</heading>
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</paragraph>
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<paragraph id="8BF236A97578FFC5FF1FFADFFCA122D9" blockId="2.[160,1168,1377,1785]" pageId="2" pageNumber="3">
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MATERIAL:
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<collectionCode id="ED5CAE6C7578FFC5FE9BFADFFE23223A" box="[340,430,1378,1402]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="3">AMNH</collectionCode>
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<materialsCitation id="3B253CF47578FFC5FE77FADCFCA522D9" collectingDate="2009-01-07" collectingDateMax="2009-01-12" collectingDateMin="2009-01-07" collectionCode="AMNH" country="India" county="Formation" location="Gujurat" municipality="Tadkeshwar" pageId="2" pageNumber="3" specimenCount="1">
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Tad-41-A;
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<collectingCountry id="F35A76397578FFC5FDF9FADCFDFA2239" box="[566,631,1377,1401]" name="India" pageId="2" pageNumber="3">India</collectingCountry>
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:
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<location id="8E9260727578FFC5FD4AFADCFD6E2239" LSID="urn:lsid:plazi:treatment:03E487BF7578FFC4FE07FA9CFE7822F8:8E9260727578FFC5FD4AFADCFD6E2239" box="[645,739,1377,1401]" country="India" county="Formation" municipality="Tadkeshwar" name="Gujurat" pageId="2" pageNumber="3">Gujurat</location>
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,
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<collectingMunicipality id="6B96ACD37578FFC5FD3EFADCFC0C2239" box="[753,897,1377,1401]" pageId="2" pageNumber="3">Tadkeshwar</collectingMunicipality>
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lignite mine; Cambay
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<collectingCounty id="62934E257578FFC5FF6FFA3CFE9622D9" box="[160,283,1409,1433]" pageId="2" pageNumber="3">Formation</collectingCounty>
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(Paleo-Eocene),
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<date id="FFF310697578FFC5FE12FA3CFD3F22D9" box="[477,690,1409,1434]" pageId="2" pageNumber="3" value="2009-01-07" valueMax="2009-01-12" valueMin="2009-01-07">
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<collectingDate id="EFB7E9817578FFC5FE12FA3CFD3F22D9" box="[477,690,1409,1434]" pageId="2" pageNumber="3" value="2009-01-07" valueMax="2009-01-12" valueMin="2009-01-07">7–12 January 2009</collectingDate>
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</date>
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(
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<collectionCode id="ED5CAE6C7578FFC5FD0EFA3CFCAD22D9" box="[705,800,1409,1433]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="3">AMNH</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<subSubSection id="C35765227578FFC4FF1FFA1CFE7822F8" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="discussion">
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<paragraph id="8BF236A97578FFC4FF1FFA1CFE7822F8" blockId="2.[160,1168,1377,1785]" lastBlockId="3.[160,1168,1376,1464]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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COMMENTS: A poorly preserved female worker (length approximately
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<quantity id="4CB59B4C7578FFC5FBE1FA1CFB0622F9" box="[1070,1163,1441,1465]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="3" unit="mm" value="7.5">7.5 mm</quantity>
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, forewing length approximately
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<quantity id="4CB59B4C7578FFC5FDEAFA7CFD0F229A" box="[549,642,1473,1498]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="3" unit="mm" value="6.7">6.7 mm</quantity>
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) putatively of the electrapine genus
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<taxonomicName id="4C4D4D2A7578FFC5FB83FA7FFEE122B9" authority="Cockerell" authorityName="Cockerell" authorityYear="1908" class="Insecta" family="Apidae" genus="Protobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B939EABB7578FFC5FB83FA7FFF7F22B9" italics="true" pageId="2" pageNumber="3">Protobombus</emphasis>
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Cockerell
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</taxonomicName>
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as indicated by the shape of the metabasitarsus. The specimen is poorly preserved with the wings outstretched from the body, the mesosoma of which is largely opened ventrally and cleared. The head is also only partially preserved, most of the right side and ventral portions being incomplete at the amber surface, with only the scape, pedicel, and basalmost flagellomeres preserved from the bee’s left side. The wings themselves are also somewhat partial with the anterior-apical portions of the bee’s right forewing missing and the left forewing largely crumpled and obscured. Given the diversity of these bees in the somewhat contemporaneous amber of the Baltic region (
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<bibRefCitation id="EFDC4B587578FFC5FE5AF95CFDA021B9" author="Engel, M. S." box="[405,557,1761,1785]" pageId="2" pageNumber="3" pagination="1 - 192" refId="ref3854" refString="Engel, M. S. 2001 a. A monograph of the Baltic amber bees and evolution of the Apoidea (Hymenoptera). Bulletin of the American Museum of Natural History 259: 1 - 192." type="journal article" year="2001">Engel, 2001a</bibRefCitation>
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,
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<bibRefCitation id="EFDC4B587578FFC5FDF4F95CFDFF21B9" author="Engel, M. S." box="[571,626,1761,1785]" pageId="2" pageNumber="3" pagination="9 - 33" refId="ref3941" refString="Engel, M. S. 2004. Geological history of the bees (Hymenoptera: Apoidea). Revista de Tecnologia e Ambiente 10 (2): 9 - 33." type="journal article" year="2004">2004</bibRefCitation>
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) as well as other deposits (
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<bibRefCitation id="EFDC4B587578FFC4FC76F95CFF542238" author="Wappler, T. & M. S. Engel" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" pagination="908 - 921" refId="ref5013" refString="Wappler, T., & M. S. Engel. 2003. The middle Eocene bee faunas of Eckfeld and Messel, Germany (Hymenoptera: Apoidea). Journal of Paleontology 77 (5): 908 - 921." type="journal article" year="2003">Wappler & Engel, 2003</bibRefCitation>
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), it is perhaps not surprising that such similar species should be discovered in
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<collectingCountry id="F35A76397579FFC4FF6FFA3DFE8E22D8" box="[160,259,1408,1432]" name="India" pageId="3" pageNumber="4">Cambay</collectingCountry>
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amber. It is greatly hoped that more complete and well-preserved specimens will eventually be recovered.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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|
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@ -1,67 +1,70 @@
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@ -113,9 +116,9 @@ DIAGNOSIS: This new group can be differentiated from
|
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(
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<bibRefCitation id="EFDC4B587579FFC4FB8AF97DFF6921B8" author="Engel, M. S." pageId="3" pageNumber="4" pagination="1 - 192" refId="ref3854" refString="Engel, M. S. 2001 a. A monograph of the Baltic amber bees and evolution of the Apoidea (Hymenoptera). Bulletin of the American Museum of Natural History 259: 1 - 192." type="journal article" year="2001">Engel, 2001a</bibRefCitation>
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) by the presence of only two submarginal cells (
|
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<figureCitation id="13762A2C7579FFC4FCF5F95DFC0621B8" box="[826,907,1760,1784]" captionStart="Figures 1–2" captionStartId="3.[160,241,1254,1276]" captionTargetBox="[160,1168,224,1249]" captionTargetId="figure-123@3.[160,1168,224,1249]" captionTargetPageId="3" captionText="Figures 1–2. Photographs of holotype worker (AMNH Tad-41-A) of Melikertes (Paramelikertes) gujaratensis, new subgenus and species. 1. Entire specimen as preserved in dorsal oblique view. 2. Detail of forewing." pageId="3" pageNumber="4">Figs. 2</figureCitation>
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<figureCitation id="13762A2C7579FFC4FCF5F95DFC0621B8" box="[826,907,1760,1784]" captionStart="Figures 1–2" captionStartId="3.[160,241,1254,1276]" captionTargetBox="[160,1168,224,1249]" captionTargetId="figure-123@3.[160,1168,224,1249]" captionTargetPageId="3" captionText="Figures 1–2. Photographs of holotype worker (AMNH Tad-41-A) of Melikertes (Paramelikertes) gujaratensis, new subgenus and species. 1. Entire specimen as preserved in dorsal oblique view. 2. Detail of forewing." figureDoi="http://doi.org/10.5281/zenodo.13736875" httpUri="https://zenodo.org/record/13736875/files/figure.png" pageId="3" pageNumber="4">Figs. 2</figureCitation>
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,
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<figureCitation id="13762A2C7579FFC4FC55F95DFC2521B8" box="[922,936,1760,1784]" captionStart="Figure 3" captionStartId="4.[160,231,1267,1289]" captionTargetBox="[194,1164,242,1243]" captionTargetId="figure-161@4.[160,1168,225,1262]" captionTargetPageId="4" captionText="Figure 3. Holotype worker (AMNH Tad-41-A) of Melikertes (Paramelikertes) gujaratensis, new subgenus and species." pageId="3" pageNumber="4">3</figureCitation>
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<figureCitation id="13762A2C7579FFC4FC55F95DFC2521B8" box="[922,936,1760,1784]" captionStart="Figure 3" captionStartId="4.[160,231,1267,1289]" captionTargetBox="[194,1164,242,1243]" captionTargetId="figure-161@4.[160,1168,225,1262]" captionTargetPageId="4" captionText="Figure 3. Holotype worker (AMNH Tad-41-A) of Melikertes (Paramelikertes) gujaratensis, new subgenus and species." figureDoi="http://doi.org/10.5281/zenodo.13736877" httpUri="https://zenodo.org/record/13736877/files/figure.png" pageId="3" pageNumber="4">3</figureCitation>
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), the confluence of 2rs-m with 2m-cu, the more elongate and narrow marginal cell, the marginal cell apex acutely pointed on the wing margin, the more densely setose metasoma, and the more strongly developed and dense stout bristles of the mid- and hind legs. Like
|
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<taxonomicName id="4C4D4D2A757EFFC3FBD0FA11FB1D2284" authorityName="Engel" authorityYear="1998" box="[1055,1168,1452,1476]" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
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<emphasis id="B939EABB757EFFC3FBD0FA11FB1D2284" box="[1055,1168,1452,1476]" italics="true" pageId="4" pageNumber="5">Melikertes</emphasis>
|
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|
|
@ -128,7 +131,7 @@ Engel
|
|||
, the mesoscutellum apparently does not project over the metanotum, and the metasomal terga are not banded.
|
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</paragraph>
|
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</subSubSection>
|
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<emphasis id="B939EABB757EFFC3FF6FFB4EFF722248" bold="true" box="[160,255,1267,1289]" pageId="4" pageNumber="5">Figure 3.</emphasis>
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Holotype worker (AMNH Tad-41-A) of
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@ -1,67 +1,70 @@
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(
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<figureCitation id="13762A2C757FFFC2FDABFEFCFD462619" box="[612,715,321,345]" captionStart-0="Figures 1–2" captionStart-1="Figure 3" captionStartId-0="3.[160,241,1254,1276]" captionStartId-1="4.[160,231,1267,1289]" captionTargetBox-0="[160,1168,224,1249]" captionTargetBox-1="[194,1164,242,1243]" captionTargetId-0="figure-123@3.[160,1168,224,1249]" captionTargetId-1="figure-161@4.[160,1168,225,1262]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figures 1–2. Photographs of holotype worker (AMNH Tad-41-A) of Melikertes (Paramelikertes) gujaratensis, new subgenus and species. 1. Entire specimen as preserved in dorsal oblique view. 2. Detail of forewing." captionText-1="Figure 3. Holotype worker (AMNH Tad-41-A) of Melikertes (Paramelikertes) gujaratensis, new subgenus and species." pageId="5" pageNumber="6">Figs. 1–3</figureCitation>
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<figureCitation id="13762A2C757FFFC2FDABFEFCFD462619" box="[612,715,321,345]" captionStart-0="Figures 1–2" captionStart-1="Figure 3" captionStartId-0="3.[160,241,1254,1276]" captionStartId-1="4.[160,231,1267,1289]" captionTargetBox-0="[160,1168,224,1249]" captionTargetBox-1="[194,1164,242,1243]" captionTargetId-0="figure-123@3.[160,1168,224,1249]" captionTargetId-1="figure-161@4.[160,1168,225,1262]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figures 1–2. Photographs of holotype worker (AMNH Tad-41-A) of Melikertes (Paramelikertes) gujaratensis, new subgenus and species. 1. Entire specimen as preserved in dorsal oblique view. 2. Detail of forewing." captionText-1="Figure 3. Holotype worker (AMNH Tad-41-A) of Melikertes (Paramelikertes) gujaratensis, new subgenus and species." figureDoi-0="http://doi.org/10.5281/zenodo.13736875" figureDoi-1="http://doi.org/10.5281/zenodo.13736877" httpUri-0="https://zenodo.org/record/13736875/files/figure.png" httpUri-1="https://zenodo.org/record/13736877/files/figure.png" pageId="5" pageNumber="6">Figs. 1–3</figureCitation>
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)
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@ -121,9 +124,9 @@ DESCRIPTION:
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,
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<collectionCode id="ED5CAE6C757FFFC2FEA0FC1FFE4424FA" box="[367,457,930,954]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="5" pageNumber="6">AMNH</collectionCode>
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Tad-41-A (
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,
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);
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<collectingCountry id="F35A7639757FFFC2FD76FC1CFD7724F9" box="[697,762,929,953]" name="India" pageId="5" pageNumber="6">India</collectingCountry>
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:
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@ -138,7 +141,7 @@ lignite mine; Cambay
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|
||||
in- clude an aphelinid (Chalcidoidea), spider, and the aforementioned specimen of
|
||||
include an aphelinid (Chalcidoidea), spider, and the aforementioned specimen of
|
||||
<taxonomicName id="4C4D4D2A757FFFC2FB83FC5FFF7F2359" authorityName="Cockerell" authorityYear="1908" class="Insecta" family="Apidae" genus="Protobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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||||
<emphasis id="B939EABB757FFFC2FB83FC5FFF7F2359" italics="true" pageId="5" pageNumber="6">Protobombus</emphasis>
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</taxonomicName>
|
||||
|
|
@ -159,7 +162,7 @@ ETYMOLOGY: The specific epithet refers to the Indian State of
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<paragraph id="8BF236A9757FFFC2FF1FFBDCFEFB23B9" blockId="5.[160,1168,385,1273]" pageId="5" pageNumber="6">
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COMMENTS: Although the body of the
|
||||
<typeStatus id="54F6880B757FFFC2FD41FBDCFD7A2339" box="[654,759,1121,1145]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||||
is not ideally preserved, the struc- ture of the legs, mesoscutellum, metasomal terga, and absence of a clypeal protrusion (such protrusions are present in
|
||||
is not ideally preserved, the structure of the legs, mesoscutellum, metasomal terga, and absence of a clypeal protrusion (such protrusions are present in
|
||||
<taxonomicName id="4C4D4D2A757FFFC2FDD5FB1FFD0223F9" authorityName="Engel" authorityYear="2001" box="[538,655,1186,1209]" class="Insecta" family="Apidae" genus="Succinapis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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||||
<emphasis id="B939EABB757FFFC2FDD5FB1FFD0223F9" box="[538,655,1186,1209]" italics="true" pageId="5" pageNumber="6">Succinapis</emphasis>
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</taxonomicName>
|
||||
|
|
|
|||
|
|
@ -1,67 +1,70 @@
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|
||||
<figureCitation id="13762A2C757FFFC2FD91FA3CFD5F22D9" box="[606,722,1409,1433]" captionStart-0="Figures 4–6" captionStart-1="Figures 7–8" captionStart-2="Figures 9–11" captionStartId-0="6.[160,241,1409,1431]" captionStartId-1="7.[160,241,1412,1434]" captionStartId-2="8.[160,241,1214,1236]" captionTargetBox-0="[160,1168,224,1403]" captionTargetBox-1="[160,1168,224,1407]" captionTargetBox-2="[176,1144,259,1174]" captionTargetId-0="figure-144@6.[160,1168,224,1403]" captionTargetId-1="figure-103@7.[160,1168,224,1407]" captionTargetId-2="figure-148@8.[160,1168,224,1209]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="Figures 4–6. Photographs of holotype worker (AMNH Tad-272-A) of Melikertes (Melikertes) kamboja, new species. 4. Frontal view of specimen as preserved. 5. Dorsal oblique view. 6. Detail of forewing." captionText-1="Figures 7–8. Photographs of paratype worker (STB-092-T’10) of Melikertes (Melikertes) kamboja, new species. 7. Dorsal view of specimen as preserved. 8. Detail of forewing." captionText-2="Figures 9–11. Holotype worker (AMNH Tad-272-A) of Melikertes (Melikertes) kamboja, new spe- cies. 9. Sketch of specimen as preserved. 10. Detail of forewing. 11. Detail of metabasitarsal setae." pageId="5" pageNumber="6">Figs. 4–11</figureCitation>
|
||||
<figureCitation id="13762A2C757FFFC2FD91FA3CFD5F22D9" box="[606,722,1409,1433]" captionStart-0="Figures 4–6" captionStart-1="Figures 7–8" captionStart-2="Figures 9–11" captionStartId-0="6.[160,241,1409,1431]" captionStartId-1="7.[160,241,1412,1434]" captionStartId-2="8.[160,241,1214,1236]" captionTargetBox-0="[160,1168,224,1403]" captionTargetBox-1="[160,1168,224,1407]" captionTargetBox-2="[176,1144,259,1174]" captionTargetId-0="figure-144@6.[160,1168,224,1403]" captionTargetId-1="figure-103@7.[160,1168,224,1407]" captionTargetId-2="figure-148@8.[160,1168,224,1209]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="Figures 4–6. Photographs of holotype worker (AMNH Tad-272-A) of Melikertes (Melikertes) kamboja, new species. 4. Frontal view of specimen as preserved. 5. Dorsal oblique view. 6. Detail of forewing." captionText-1="Figures 7–8. Photographs of paratype worker (STB-092-T’10) of Melikertes (Melikertes) kamboja, new species. 7. Dorsal view of specimen as preserved. 8. Detail of forewing." captionText-2="Figures 9–11. Holotype worker (AMNH Tad-272-A) of Melikertes (Melikertes) kamboja, new spe- cies. 9. Sketch of specimen as preserved. 10. Detail of forewing. 11. Detail of metabasitarsal setae." figureDoi-0="http://doi.org/10.5281/zenodo.13736881" figureDoi-1="http://doi.org/10.5281/zenodo.13736883" figureDoi-2="http://doi.org/10.5281/zenodo.13736887" httpUri-0="https://zenodo.org/record/13736881/files/figure.png" httpUri-1="https://zenodo.org/record/13736883/files/figure.png" httpUri-2="https://zenodo.org/record/13736887/files/figure.png" pageId="5" pageNumber="6">Figs. 4–11</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -119,20 +122,20 @@ DESCRIPTION:
|
|||
<quantity id="4CB59B4C757FFFC2FC7DF99CFB942179" box="[946,1049,1569,1593]" metricMagnitude="-3" metricUnit="m" metricValue="3.64" pageId="5" pageNumber="6" unit="mm" value="3.64">3.64 mm</quantity>
|
||||
; forewing length
|
||||
<quantity id="4CB59B4C757FFFC2FF3BF9FCFED02119" box="[244,349,1601,1625]" metricMagnitude="-3" metricUnit="m" metricValue="2.77" pageId="5" pageNumber="6" unit="mm" value="2.77">2.77 mm</quantity>
|
||||
. Mandible elongate, curved, with at least two apical rounded teeth. Clypeus damaged but clearly without medial cleft or protuberances. Scape elongate and narrow; pedicel compressed and drawn out as preserved; flagellum distinctly wider than scape and pedicel. Mesosoma and legs covered by thin branched setae (particularly well visible on tibiae and basitarsi); mesoscutellum apparently not bulg- ing nor covering metanotum or propodeum. Metabasitarsus slightly widened with parallel sides; setae distinctly branched (
|
||||
<figureCitation id="13762A2C757CFFC1FD49FA40FD552156" box="[646,728,1533,1558]" captionStart="Figures 9–11" captionStartId="8.[160,241,1214,1236]" captionTargetBox="[176,1144,259,1174]" captionTargetId="figure-148@8.[160,1168,224,1209]" captionTargetPageId="8" captionText="Figures 9–11. Holotype worker (AMNH Tad-272-A) of Melikertes (Melikertes) kamboja, new spe- cies. 9. Sketch of specimen as preserved. 10. Detail of forewing. 11. Detail of metabasitarsal setae." pageId="6" pageNumber="7">Fig. 11</figureCitation>
|
||||
. Mandible elongate, curved, with at least two apical rounded teeth. Clypeus damaged but clearly without medial cleft or protuberances. Scape elongate and narrow; pedicel compressed and drawn out as preserved; flagellum distinctly wider than scape and pedicel. Mesosoma and legs covered by thin branched setae (particularly well visible on tibiae and basitarsi); mesoscutellum apparently not bulging nor covering metanotum or propodeum. Metabasitarsus slightly widened with parallel sides; setae distinctly branched (
|
||||
<figureCitation id="13762A2C757CFFC1FD49FA40FD552156" box="[646,728,1533,1558]" captionStart="Figures 9–11" captionStartId="8.[160,241,1214,1236]" captionTargetBox="[176,1144,259,1174]" captionTargetId="figure-148@8.[160,1168,224,1209]" captionTargetPageId="8" captionText="Figures 9–11. Holotype worker (AMNH Tad-272-A) of Melikertes (Melikertes) kamboja, new spe- cies. 9. Sketch of specimen as preserved. 10. Detail of forewing. 11. Detail of metabasitarsal setae." figureDoi="http://doi.org/10.5281/zenodo.13736887" httpUri="https://zenodo.org/record/13736887/files/figure.png" pageId="6" pageNumber="7">Fig. 11</figureCitation>
|
||||
). Forewing with basal vein straight, basad cu-a by about one-half length of cu-a; M angled posteriorly after separating from Rs; r-rs slightly more than three-quarters length second abscissa of Rs; second abscissa Rs straight; 1rs-m present (thus three submarginal cells) (
|
||||
<figureCitation id="13762A2C757CFFC1FC52F9E0FC672135" box="[925,1002,1629,1653]" captionStart="Figures 4–6" captionStartId="6.[160,241,1409,1431]" captionTargetBox="[160,1168,224,1403]" captionTargetId="figure-144@6.[160,1168,224,1403]" captionTargetPageId="6" captionText="Figures 4–6. Photographs of holotype worker (AMNH Tad-272-A) of Melikertes (Melikertes) kamboja, new species. 4. Frontal view of specimen as preserved. 5. Dorsal oblique view. 6. Detail of forewing." pageId="6" pageNumber="7">Figs. 6</figureCitation>
|
||||
<figureCitation id="13762A2C757CFFC1FC52F9E0FC672135" box="[925,1002,1629,1653]" captionStart="Figures 4–6" captionStartId="6.[160,241,1409,1431]" captionTargetBox="[160,1168,224,1403]" captionTargetId="figure-144@6.[160,1168,224,1403]" captionTargetPageId="6" captionText="Figures 4–6. Photographs of holotype worker (AMNH Tad-272-A) of Melikertes (Melikertes) kamboja, new species. 4. Frontal view of specimen as preserved. 5. Dorsal oblique view. 6. Detail of forewing." figureDoi="http://doi.org/10.5281/zenodo.13736881" httpUri="https://zenodo.org/record/13736881/files/figure.png" pageId="6" pageNumber="7">Figs. 6</figureCitation>
|
||||
,
|
||||
<figureCitation id="13762A2C757CFFC1FC3BF9E3FB8E2136" box="[1012,1027,1630,1654]" captionStart="Figures 7–8" captionStartId="7.[160,241,1412,1434]" captionTargetBox="[160,1168,224,1407]" captionTargetId="figure-103@7.[160,1168,224,1407]" captionTargetPageId="7" captionText="Figures 7–8. Photographs of paratype worker (STB-092-T’10) of Melikertes (Melikertes) kamboja, new species. 7. Dorsal view of specimen as preserved. 8. Detail of forewing." pageId="6" pageNumber="7">8</figureCitation>
|
||||
<figureCitation id="13762A2C757CFFC1FC3BF9E3FB8E2136" box="[1012,1027,1630,1654]" captionStart="Figures 7–8" captionStartId="7.[160,241,1412,1434]" captionTargetBox="[160,1168,224,1407]" captionTargetId="figure-103@7.[160,1168,224,1407]" captionTargetPageId="7" captionText="Figures 7–8. Photographs of paratype worker (STB-092-T’10) of Melikertes (Melikertes) kamboja, new species. 7. Dorsal view of specimen as preserved. 8. Detail of forewing." figureDoi="http://doi.org/10.5281/zenodo.13736883" httpUri="https://zenodo.org/record/13736883/files/figure.png" pageId="6" pageNumber="7">8</figureCitation>
|
||||
,
|
||||
<figureCitation id="13762A2C757CFFC1FBC1F9E3FBA62136" box="[1038,1067,1630,1654]" captionStart="Figures 9–11" captionStartId="8.[160,241,1214,1236]" captionTargetBox="[176,1144,259,1174]" captionTargetId="figure-148@8.[160,1168,224,1209]" captionTargetPageId="8" captionText="Figures 9–11. Holotype worker (AMNH Tad-272-A) of Melikertes (Melikertes) kamboja, new spe- cies. 9. Sketch of specimen as preserved. 10. Detail of forewing. 11. Detail of metabasitarsal setae." pageId="6" pageNumber="7">10</figureCitation>
|
||||
<figureCitation id="13762A2C757CFFC1FBC1F9E3FBA62136" box="[1038,1067,1630,1654]" captionStart="Figures 9–11" captionStartId="8.[160,241,1214,1236]" captionTargetBox="[176,1144,259,1174]" captionTargetId="figure-148@8.[160,1168,224,1209]" captionTargetPageId="8" captionText="Figures 9–11. Holotype worker (AMNH Tad-272-A) of Melikertes (Melikertes) kamboja, new spe- cies. 9. Sketch of specimen as preserved. 10. Detail of forewing. 11. Detail of metabasitarsal setae." figureDoi="http://doi.org/10.5281/zenodo.13736887" httpUri="https://zenodo.org/record/13736887/files/figure.png" pageId="6" pageNumber="7">10</figureCitation>
|
||||
); second submarginal cell narrow, narrowed anteriorly, anterior border along Rs shorter than r-rs; third submarginal cell larger than second, with anterior border along Rs about three times longer than anterior border of second submarginal cell; 1m-cu meeting second submarginal cell near midpoint; 2rs-m weakly arched apically in posterior half, confluent with 2m-cu; discal and subdiscal cells somewhat longer than high (around 1.6 times); marginal cell elongate, broadly tubular (
|
||||
<emphasis id="B939EABB757DFFC0FD34F9B7FC9E2161" box="[763,787,1546,1569]" italics="true" pageId="7" pageNumber="8">i.e</emphasis>
|
||||
., not tapering in width along its length), apex broadly rounded and truncate and appendiculate. Hind wing with six distinct hamuli on anterior margin. Metasoma with reddish aspect as preserved, apparently not banded and without fasciae; sting short and straight.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF326621757CFFC1FF6FFA3CFEAF2291" pageId="6" pageNumber="7" startId="6.[160,241,1409,1431]" targetBox="[160,1168,224,1403]" targetPageId="6" targetType="figure">
|
||||
<caption id="DF326621757CFFC1FF6FFA3CFEAF2291" ID-DOI="http://doi.org/10.5281/zenodo.13736881" ID-Zenodo-Dep="13736881" httpUri="https://zenodo.org/record/13736881/files/figure.png" pageId="6" pageNumber="7" startId="6.[160,241,1409,1431]" targetBox="[160,1168,224,1403]" targetPageId="6" targetType="figure">
|
||||
<paragraph id="8BF236A9757CFFC1FF6FFA3CFEAF2291" blockId="6.[160,1168,1409,1489]" pageId="6" pageNumber="7">
|
||||
<emphasis id="B939EABB757CFFC1FF6FFA3CFEAC22D6" bold="true" box="[160,289,1409,1431]" pageId="6" pageNumber="7">Figures 4–6.</emphasis>
|
||||
Photographs of holotype worker (AMNH Tad-272-A) of
|
||||
|
|
@ -154,7 +157,7 @@ Dorsal oblique view.
|
|||
Detail of forewing.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF326621757DFFC0FF6FFA39FC4022F7" pageId="7" pageNumber="8" startId="7.[160,241,1412,1434]" targetBox="[160,1168,224,1407]" targetPageId="7" targetType="figure">
|
||||
<caption id="DF326621757DFFC0FF6FFA39FC4022F7" ID-DOI="http://doi.org/10.5281/zenodo.13736883" ID-Zenodo-Dep="13736883" httpUri="https://zenodo.org/record/13736883/files/figure.png" pageId="7" pageNumber="8" startId="7.[160,241,1412,1434]" targetBox="[160,1168,224,1407]" targetPageId="7" targetType="figure">
|
||||
<paragraph id="8BF236A9757DFFC0FF6FFA39FC4022F7" blockId="7.[160,1157,1412,1463]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B939EABB757DFFC0FF6FFA39FEAC22D9" bold="true" box="[160,289,1412,1434]" pageId="7" pageNumber="8">Figures 7–8.</emphasis>
|
||||
Photographs of paratype worker (STB-092-T’10) of
|
||||
|
|
@ -181,9 +184,9 @@ Detail of forewing.
|
|||
:
|
||||
<collectionCode id="ED5CAE6C757DFFC0FE9AF934FE2221E1" box="[341,431,1673,1697]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="7" pageNumber="8">AMNH</collectionCode>
|
||||
Tad-272-A (
|
||||
<figureCitation id="13762A2C757DFFC0FD88F934FD3421E1" box="[583,697,1673,1697]" captionStart="Figures 4–6" captionStartId="6.[160,241,1409,1431]" captionTargetBox="[160,1168,224,1403]" captionTargetId="figure-144@6.[160,1168,224,1403]" captionTargetPageId="6" captionText="Figures 4–6. Photographs of holotype worker (AMNH Tad-272-A) of Melikertes (Melikertes) kamboja, new species. 4. Frontal view of specimen as preserved. 5. Dorsal oblique view. 6. Detail of forewing." pageId="7" pageNumber="8">Figs. 4, 5</figureCitation>
|
||||
<figureCitation id="13762A2C757DFFC0FD88F934FD3421E1" box="[583,697,1673,1697]" captionStart="Figures 4–6" captionStartId="6.[160,241,1409,1431]" captionTargetBox="[160,1168,224,1403]" captionTargetId="figure-144@6.[160,1168,224,1403]" captionTargetPageId="6" captionText="Figures 4–6. Photographs of holotype worker (AMNH Tad-272-A) of Melikertes (Melikertes) kamboja, new species. 4. Frontal view of specimen as preserved. 5. Dorsal oblique view. 6. Detail of forewing." figureDoi="http://doi.org/10.5281/zenodo.13736881" httpUri="https://zenodo.org/record/13736881/files/figure.png" pageId="7" pageNumber="8">Figs. 4, 5</figureCitation>
|
||||
,
|
||||
<figureCitation id="13762A2C757DFFC0FD06F934FD5521E1" box="[713,728,1673,1697]" captionStart="Figures 9–11" captionStartId="8.[160,241,1214,1236]" captionTargetBox="[176,1144,259,1174]" captionTargetId="figure-148@8.[160,1168,224,1209]" captionTargetPageId="8" captionText="Figures 9–11. Holotype worker (AMNH Tad-272-A) of Melikertes (Melikertes) kamboja, new spe- cies. 9. Sketch of specimen as preserved. 10. Detail of forewing. 11. Detail of metabasitarsal setae." pageId="7" pageNumber="8">9</figureCitation>
|
||||
<figureCitation id="13762A2C757DFFC0FD06F934FD5521E1" box="[713,728,1673,1697]" captionStart="Figures 9–11" captionStartId="8.[160,241,1214,1236]" captionTargetBox="[176,1144,259,1174]" captionTargetId="figure-148@8.[160,1168,224,1209]" captionTargetPageId="8" captionText="Figures 9–11. Holotype worker (AMNH Tad-272-A) of Melikertes (Melikertes) kamboja, new spe- cies. 9. Sketch of specimen as preserved. 10. Detail of forewing. 11. Detail of metabasitarsal setae." figureDoi="http://doi.org/10.5281/zenodo.13736887" httpUri="https://zenodo.org/record/13736887/files/figure.png" pageId="7" pageNumber="8">9</figureCitation>
|
||||
);
|
||||
<collectingCountry id="F35A7639757DFFC0FD3FF934FCBC21E1" box="[752,817,1673,1697]" name="India" pageId="7" pageNumber="8">India</collectingCountry>
|
||||
:
|
||||
|
|
@ -202,7 +205,7 @@ lignite mine; Cambay Fm. (Paleo-Eocene),
|
|||
<paragraph id="8BF236A9757DFFC0FF1FF974FE542041" blockId="7.[160,1168,1513,1793]" pageId="7" pageNumber="8">
|
||||
<typeStatus id="54F6880B757DFFC0FF1FF974FECE21A1" box="[208,323,1737,1761]" pageId="7" pageNumber="8" type="paratype">PARATYPE</typeStatus>
|
||||
: STB-092-T’10 (
|
||||
<figureCitation id="13762A2C757DFFC0FE3BF974FDBA21A1" box="[500,567,1737,1761]" captionStart="Figures 7–8" captionStartId="7.[160,241,1412,1434]" captionTargetBox="[160,1168,224,1407]" captionTargetId="figure-103@7.[160,1168,224,1407]" captionTargetPageId="7" captionText="Figures 7–8. Photographs of paratype worker (STB-092-T’10) of Melikertes (Melikertes) kamboja, new species. 7. Dorsal view of specimen as preserved. 8. Detail of forewing." pageId="7" pageNumber="8">Fig. 7</figureCitation>
|
||||
<figureCitation id="13762A2C757DFFC0FE3BF974FDBA21A1" box="[500,567,1737,1761]" captionStart="Figures 7–8" captionStartId="7.[160,241,1412,1434]" captionTargetBox="[160,1168,224,1407]" captionTargetId="figure-103@7.[160,1168,224,1407]" captionTargetPageId="7" captionText="Figures 7–8. Photographs of paratype worker (STB-092-T’10) of Melikertes (Melikertes) kamboja, new species. 7. Dorsal view of specimen as preserved. 8. Detail of forewing." figureDoi="http://doi.org/10.5281/zenodo.13736883" httpUri="https://zenodo.org/record/13736883/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
|
||||
);
|
||||
<materialsCitation id="3B253CF4757DFFC0FD84F974FBAF21A1" box="[587,1058,1737,1761]" country="India" location="Gujarat" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Gujarat" typeStatus="paratype">
|
||||
<collectingCountry id="F35A7639757DFFC0FD84F974FD0121A1" box="[587,652,1737,1761]" name="India" pageId="7" pageNumber="8">India</collectingCountry>
|
||||
|
|
@ -220,7 +223,7 @@ Fm. (Paleo-Eocene) (
|
|||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF3266217572FFCFFF6FFB03FF51224E" pageId="8" pageNumber="9" startId="8.[160,241,1214,1236]" targetBox="[176,1144,259,1174]" targetPageId="8" targetType="figure">
|
||||
<caption id="DF3266217572FFCFFF6FFB03FF51224E" ID-DOI="http://doi.org/10.5281/zenodo.13736887" ID-Zenodo-Dep="13736887" httpUri="https://zenodo.org/record/13736887/files/figure.png" pageId="8" pageNumber="9" startId="8.[160,241,1214,1236]" targetBox="[176,1144,259,1174]" targetPageId="8" targetType="figure">
|
||||
<paragraph id="8BF236A97572FFCFFF6FFB03FF51224E" blockId="8.[160,1165,1214,1294]" pageId="8" pageNumber="9">
|
||||
<emphasis id="B939EABB7572FFCFFF6FFB03FEA32393" bold="true" box="[160,302,1214,1236]" pageId="8" pageNumber="9">Figures 9–11.</emphasis>
|
||||
Holotype worker (AMNH Tad-272-A) of
|
||||
|
|
@ -248,9 +251,9 @@ ETYMOLOGY: The name Kamboja refers to the Indo-Iranian Kshatriya tribe who are r
|
|||
<superScript id="7C389BE17572FFCFFC7AFAE1FC45222A" attach="left" box="[949,968,1372,1386]" fontSize="6" pageId="8" pageNumber="9">nd</superScript>
|
||||
Century B.C. the Kambojas invaded northern
|
||||
<collectingCountry id="F35A76397572FFCFFE3CFAC0FDBC22D5" box="[499,561,1405,1429]" name="India" pageId="8" pageNumber="9">India</collectingCountry>
|
||||
and wrestled control of various Indo-Arayan terri- tories including
|
||||
and wrestled control of various Indo-Arayan territories including
|
||||
<collectingRegion id="4989F84B7572FFCFFEAEFA20FE3122F5" box="[353,444,1437,1461]" country="India" name="Gujarat" pageId="8" pageNumber="9">Gujarat</collectingRegion>
|
||||
, eventually settling and giving their name to the area (Kham- bat or
|
||||
, eventually settling and giving their name to the area (Khambat or
|
||||
<collectingCountry id="F35A76397572FFCFFF24FA00FEDF2295" box="[235,338,1469,1493]" name="India" pageId="8" pageNumber="9">Cambay</collectingCountry>
|
||||
). The name is treated as a noun in apposition.
|
||||
</paragraph>
|
||||
|
|
|
|||
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Reference in a new issue