Add updates up until 2025-01-02 13:12:51

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Key for species of the genus
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(mainly females)

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Key for the
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(mainly adult specimens)
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1. Pedipalp chelae very robust (
<figureCitation id="B9E82A60FFE8FF8CFE52FCA3FDBC5DD7" box="[456,522,875,899]" captionStart="Figures 27" captionStartId="4.[113,190,1595,1617]" captionTargetBox="[113,1474,234,1577]" captionTargetId="figure-249@4.[113,1474,234,1577]" captionTargetPageId="4" captionText="Figures 27: Diplocentrid characters. Figures 2, 4, 7. Heteronebo portoricensis, female in vivo habitus (2), coxosternal area (4); metasoma V, ventral aspect (7). Figures 3, 5. Oiclus sp., pedipalp patella, ventral aspect (3); leg IV tarsomere II (5). Figure 6. Cazierius garridoi, metasoma V, ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4648514" httpUri="https://zenodo.org/record/4648514/files/figure.png" pageId="7" pageNumber="6">Fig. 2</figureCitation>
). Ventral surface of the pedipalp patella with three trichobothria (
<figureCitation id="B9E82A60FFE8FF8CFDCBFC43FD255DF7" box="[593,659,907,931]" captionStart="Figures 27" captionStartId="4.[113,190,1595,1617]" captionTargetBox="[113,1474,234,1577]" captionTargetId="figure-249@4.[113,1474,234,1577]" captionTargetPageId="4" captionText="Figures 27: Diplocentrid characters. Figures 2, 4, 7. Heteronebo portoricensis, female in vivo habitus (2), coxosternal area (4); metasoma V, ventral aspect (7). Figures 3, 5. Oiclus sp., pedipalp patella, ventral aspect (3); leg IV tarsomere II (5). Figure 6. Cazierius garridoi, metasoma V, ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4648514" httpUri="https://zenodo.org/record/4648514/files/figure.png" pageId="7" pageNumber="6">Fig. 3</figureCitation>
). Sternum clearly pentagonal (
<figureCitation id="B9E82A60FFE8FF8CFEFFFC63FE115D97" box="[357,423,939,963]" captionStart="Figures 27" captionStartId="4.[113,190,1595,1617]" captionTargetBox="[113,1474,234,1577]" captionTargetId="figure-249@4.[113,1474,234,1577]" captionTargetPageId="4" captionText="Figures 27: Diplocentrid characters. Figures 2, 4, 7. Heteronebo portoricensis, female in vivo habitus (2), coxosternal area (4); metasoma V, ventral aspect (7). Figures 3, 5. Oiclus sp., pedipalp patella, ventral aspect (3); leg IV tarsomere II (5). Figure 6. Cazierius garridoi, metasoma V, ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4648514" httpUri="https://zenodo.org/record/4648514/files/figure.png" pageId="7" pageNumber="6">Fig. 4</figureCitation>
). Pectines with six to eight teeth (
<figureCitation id="B9E82A60FFE8FF8CFF03FC03FF565DB7" box="[153,224,971,995]" captionStart="Figures 27" captionStartId="4.[113,190,1595,1617]" captionTargetBox="[113,1474,234,1577]" captionTargetId="figure-249@4.[113,1474,234,1577]" captionTargetPageId="4" captionText="Figures 27: Diplocentrid characters. Figures 2, 4, 7. Heteronebo portoricensis, female in vivo habitus (2), coxosternal area (4); metasoma V, ventral aspect (7). Figures 3, 5. Oiclus sp., pedipalp patella, ventral aspect (3); leg IV tarsomere II (5). Figure 6. Cazierius garridoi, metasoma V, ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4648514" httpUri="https://zenodo.org/record/4648514/files/figure.png" pageId="7" pageNumber="6">Fig. 4</figureCitation>
). Leg tarsomere II (telotarsus) with two rows of spine-like setae (
<figureCitation id="B9E82A60FFE8FF8CFEDFFC23FE305A56" box="[325,390,1002,1027]" captionStart="Figures 27" captionStartId="4.[113,190,1595,1617]" captionTargetBox="[113,1474,234,1577]" captionTargetId="figure-249@4.[113,1474,234,1577]" captionTargetPageId="4" captionText="Figures 27: Diplocentrid characters. Figures 2, 4, 7. Heteronebo portoricensis, female in vivo habitus (2), coxosternal area (4); metasoma V, ventral aspect (7). Figures 3, 5. Oiclus sp., pedipalp patella, ventral aspect (3); leg IV tarsomere II (5). Figure 6. Cazierius garridoi, metasoma V, ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4648514" httpUri="https://zenodo.org/record/4648514/files/figure.png" pageId="7" pageNumber="6">Fig. 5</figureCitation>
). ........................(
<taxonomicName id="E6D34D66FFE8FF8CFDDFFC23FD515A57" authorityName="Karsch" authorityYear="1880" box="[581,743,1003,1027]" class="Arachnida" family="Diplocentridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="family">Diplocentridae</taxonomicName>
) 2
</keyLead>
</paragraph>
<paragraph id="216C36E5FFE8FF8CFFEBFBC3FCB45AF7" blockId="7.[113,771,875,1187]" pageId="7" pageNumber="6">
<keyLead id="9A2295D0FFE8FF8CFFEBFBC3FCB45AF7" pageId="7" pageNumber="6">
Pedipalp chelae feeble (
<figureCitation id="B9E82A60FFE8FF8CFE23FBC3FDBE5A77" box="[441,520,1035,1059]" captionStart="Figures 811" captionStartId="5.[116,193,1031,1053]" captionTargetBox="[113,1474,234,1016]" captionTargetId="figure-490@5.[113,1474,234,1016]" captionTargetPageId="5" captionText="Figures 811. Buthid characters. Figure 8. Tityus obtusus, female in vivo habitus. Figure 9. Heteroctenus abudi, female, coxosternal region and sternite III. Figure 10. Heteroctenus abudi, pedipalp movable finger dentition, showing the median rows of denticles and the accessory denticles (10). Figure 11. Tityus sp., pedipalp movable finger dentition." figureDoi="http://doi.org/10.5281/zenodo.4648518" httpUri="https://zenodo.org/record/4648518/files/figure.png" pageId="7" pageNumber="6">Fig. 8</figureCitation>
). Ventral surface of the pedipalp patella without trichobothria. Sternum subtriangular to subpentagonal (
<figureCitation id="B9E82A60FFE8FF8CFD9EFB83FDF95A37" box="[516,591,1099,1123]" captionStart="Figures 811" captionStartId="5.[116,193,1031,1053]" captionTargetBox="[113,1474,234,1016]" captionTargetId="figure-490@5.[113,1474,234,1016]" captionTargetPageId="5" captionText="Figures 811. Buthid characters. Figure 8. Tityus obtusus, female in vivo habitus. Figure 9. Heteroctenus abudi, female, coxosternal region and sternite III. Figure 10. Heteroctenus abudi, pedipalp movable finger dentition, showing the median rows of denticles and the accessory denticles (10). Figure 11. Tityus sp., pedipalp movable finger dentition." figureDoi="http://doi.org/10.5281/zenodo.4648518" httpUri="https://zenodo.org/record/4648518/files/figure.png" pageId="7" pageNumber="6">Fig. 9</figureCitation>
). Pectines with more than eight teeth (
<figureCitation id="B9E82A60FFE8FF8CFE08FBA3FE575AD7" box="[402,481,1131,1155]" captionStart="Figures 811" captionStartId="5.[116,193,1031,1053]" captionTargetBox="[113,1474,234,1016]" captionTargetId="figure-490@5.[113,1474,234,1016]" captionTargetPageId="5" captionText="Figures 811. Buthid characters. Figure 8. Tityus obtusus, female in vivo habitus. Figure 9. Heteroctenus abudi, female, coxosternal region and sternite III. Figure 10. Heteroctenus abudi, pedipalp movable finger dentition, showing the median rows of denticles and the accessory denticles (10). Figure 11. Tityus sp., pedipalp movable finger dentition." figureDoi="http://doi.org/10.5281/zenodo.4648518" httpUri="https://zenodo.org/record/4648518/files/figure.png" pageId="7" pageNumber="6">Figs. 9</figureCitation>
,
<figureCitation id="B9E82A60FFE8FF8CFE6BFBA3FD855AD7" box="[497,563,1131,1155]" captionStart="Figures 1214" captionStartId="6.[116,193,1336,1358]" captionTargetBox="[113,1474,234,1325]" captionTargetId="figure-362@6.[113,1474,234,1325]" captionTargetPageId="6" captionText="Figures 1214: Microtityus spp., pectines. Figure 12. M. vieques, male holotype, basal middle lamella conspicuously enlarged, modified from Teruel et al. (2015: fig. 2b). Figure 13. M. santosi, female holotype, basal plate and the distal area of the left comb, showing two egg-like organisms firmly attached (outside square), modified from Teruel et al. (2014: fig. 5c). Figure 14. M. borincanus, female holotype, basal plate, modified from Teruel et al. (2014: fig. 2c)." figureDoi="http://doi.org/10.5281/zenodo.4648520" httpUri="https://zenodo.org/record/4648520/files/figure.png" pageId="7" pageNumber="6">12-14</figureCitation>
). Leg tarsomere II (telotarsus) without spine-like setae. .............. (
<taxonomicName id="E6D34D66FFE8FF8CFD19FB43FD515AF7" authorityName="C.L.Koch" authorityYear="1837" box="[643,743,1163,1187]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
) 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A272E40FFE8FF8CFFEBFB03FCB45BD7" pageId="7" pageNumber="6">
<paragraph id="216C36E5FFE8FF8CFFEBFB03FCB45B76" blockId="7.[113,771,1227,1411]" pageId="7" pageNumber="6">
<keyLead id="9A2295D0FFE8FF8CFFEBFB03FCB45B76" pageId="7" pageNumber="6">
2. Ventral surface of the metasomal segment V with a distal semicircular area, anteriorly delimited for well-defined transversal carina (
<figureCitation id="B9E82A60FFE8FF8CFEC1FAC3FE2B5B77" box="[347,413,1291,1315]" captionStart="Figures 27" captionStartId="4.[113,190,1595,1617]" captionTargetBox="[113,1474,234,1577]" captionTargetId="figure-249@4.[113,1474,234,1577]" captionTargetPageId="4" captionText="Figures 27: Diplocentrid characters. Figures 2, 4, 7. Heteronebo portoricensis, female in vivo habitus (2), coxosternal area (4); metasoma V, ventral aspect (7). Figures 3, 5. Oiclus sp., pedipalp patella, ventral aspect (3); leg IV tarsomere II (5). Figure 6. Cazierius garridoi, metasoma V, ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4648514" httpUri="https://zenodo.org/record/4648514/files/figure.png" pageId="7" pageNumber="6">Fig. 6</figureCitation>
). ...................................
<taxonomicName id="E6D34D66FFE8FF8CFD06FAC3FCB45B76" authorityName="Francke" authorityYear="1978" box="[668,770,1291,1314]" class="Arachnida" family="Diplocentridae" genus="Cazierius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="13A7EAF7FFE8FF8CFD06FAC3FCB45B76" box="[668,770,1291,1314]" italics="true" pageId="7" pageNumber="6">Cazierius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="216C36E5FFE8FF8CFFEBFAE3FCB45BD7" blockId="7.[113,771,1227,1411]" pageId="7" pageNumber="6">
<keyLead id="9A2295D0FFE8FF8CFFEBFAE3FCB45BD7" pageId="7" pageNumber="6">
Ventral surface of the metasomal segment V without a distal semicircular area, with the transversal carina poorly defined (
<figureCitation id="B9E82A60FFE8FF8CFF75FAA3FE865BD7" box="[239,304,1387,1411]" captionStart="Figures 27" captionStartId="4.[113,190,1595,1617]" captionTargetBox="[113,1474,234,1577]" captionTargetId="figure-249@4.[113,1474,234,1577]" captionTargetPageId="4" captionText="Figures 27: Diplocentrid characters. Figures 2, 4, 7. Heteronebo portoricensis, female in vivo habitus (2), coxosternal area (4); metasoma V, ventral aspect (7). Figures 3, 5. Oiclus sp., pedipalp patella, ventral aspect (3); leg IV tarsomere II (5). Figure 6. Cazierius garridoi, metasoma V, ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4648514" httpUri="https://zenodo.org/record/4648514/files/figure.png" pageId="7" pageNumber="6">Fig. 7</figureCitation>
). ...............................................
<taxonomicName id="E6D34D66FFE8FF8CFD1FFAA3FCB45BD7" authorityName="Pocock" authorityYear="1899" box="[645,770,1387,1411]" class="Arachnida" family="Diplocentridae" genus="Heteronebo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="13A7EAF7FFE8FF8CFD1FFAA3FCB45BD7" box="[645,770,1387,1411]" italics="true" pageId="7" pageNumber="6">Heteronebo</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A272E40FFE8FF8CFFEBFA63FCB45837" pageId="7" pageNumber="6">
<paragraph id="216C36E5FFE8FF8CFFEBFA63FCB45856" blockId="7.[113,771,1451,1635]" pageId="7" pageNumber="6">
<keyLead id="9A2295D0FFE8FF8CFFEBFA63FCB45856" pageId="7" pageNumber="6">
3. Pedipalp fixed finger with six rows of denticles, being the basal one very long (it occupies almost the basal one-half of the finger). ......................................................
<taxonomicName id="E6D34D66FFE8FF8CFD03FA23FCB45856" box="[665,770,1515,1538]" class="Arachnida" family="Buthidae" genus="Isometrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="13A7EAF7FFE8FF8CFD03FA23FCB45856" box="[665,770,1515,1538]" italics="true" pageId="7" pageNumber="6">Isometrus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="216C36E5FFE8FF8CFFEBF9C3FCB45837" blockId="7.[113,771,1451,1635]" pageId="7" pageNumber="6">
<keyLead id="9A2295D0FFE8FF8CFFEBF9C3FCB45837" pageId="7" pageNumber="6"> Pedipalp fixed finger with more than six rows of denticles, being the basal one very short (it occupies approximately one-eighth of the total length of the finger). ................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A272E40FFE8FF8CFFEBF943FCB45957" pageId="7" pageNumber="6">
<paragraph id="216C36E5FFE8FF8CFFEBF943FCB45896" blockId="7.[113,771,1675,1795]" pageId="7" pageNumber="6">
<keyLead id="9A2295D0FFE8FF8CFFEBF943FCB45896" pageId="7" pageNumber="6">
4. Pedipalp fingers with inner accessory denticles and accessory outer denticles (
<figureCitation id="B9E82A60FFE8FF8CFE31F963FE4C5897" box="[427,506,1707,1731]" captionStart="Figures 811" captionStartId="5.[116,193,1031,1053]" captionTargetBox="[113,1474,234,1016]" captionTargetId="figure-490@5.[113,1474,234,1016]" captionTargetPageId="5" captionText="Figures 811. Buthid characters. Figure 8. Tityus obtusus, female in vivo habitus. Figure 9. Heteroctenus abudi, female, coxosternal region and sternite III. Figure 10. Heteroctenus abudi, pedipalp movable finger dentition, showing the median rows of denticles and the accessory denticles (10). Figure 11. Tityus sp., pedipalp movable finger dentition." figureDoi="http://doi.org/10.5281/zenodo.4648518" httpUri="https://zenodo.org/record/4648518/files/figure.png" pageId="7" pageNumber="6">Fig. 10</figureCitation>
). .................................. 5
</keyLead>
</paragraph>
<paragraph id="216C36E5FFE8FF8CFFEBF903FCB45957" blockId="7.[113,771,1675,1795]" pageId="7" pageNumber="6">
<keyLead id="9A2295D0FFE8FF8CFFEBF903FCB45957" pageId="7" pageNumber="6">
Pedipalp fingers without either inner accessory denticles or outer accessory denticles (
<figureCitation id="B9E82A60FFE8FF8CFE31F923FE4E5957" box="[427,504,1771,1795]" captionStart="Figures 811" captionStartId="5.[116,193,1031,1053]" captionTargetBox="[113,1474,234,1016]" captionTargetId="figure-490@5.[113,1474,234,1016]" captionTargetPageId="5" captionText="Figures 811. Buthid characters. Figure 8. Tityus obtusus, female in vivo habitus. Figure 9. Heteroctenus abudi, female, coxosternal region and sternite III. Figure 10. Heteroctenus abudi, pedipalp movable finger dentition, showing the median rows of denticles and the accessory denticles (10). Figure 11. Tityus sp., pedipalp movable finger dentition." figureDoi="http://doi.org/10.5281/zenodo.4648518" httpUri="https://zenodo.org/record/4648518/files/figure.png" pageId="7" pageNumber="6">Fig. 11</figureCitation>
). ................................. 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A272E40FFE8FF8CFFEBF8E2FCB55997" pageId="7" pageNumber="6">
<paragraph id="216C36E5FFE8FF8CFFEBF8E2FCB459D6" blockId="7.[113,771,1834,1987]" pageId="7" pageNumber="6">
<keyLead id="9A2295D0FFE8FF8CFFEBF8E2FCB459D6" pageId="7" pageNumber="6">
5. Sternite III (the first post-pectinal) with a well-developed pair of longitudinal furrows, which converge towards the anterior part of the plate (
<figureCitation id="B9E82A60FFE8FF8CFE38F8A3FE5259D7" box="[418,484,1899,1923]" captionStart="Figures 811" captionStartId="5.[116,193,1031,1053]" captionTargetBox="[113,1474,234,1016]" captionTargetId="figure-490@5.[113,1474,234,1016]" captionTargetPageId="5" captionText="Figures 811. Buthid characters. Figure 8. Tityus obtusus, female in vivo habitus. Figure 9. Heteroctenus abudi, female, coxosternal region and sternite III. Figure 10. Heteroctenus abudi, pedipalp movable finger dentition, showing the median rows of denticles and the accessory denticles (10). Figure 11. Tityus sp., pedipalp movable finger dentition." figureDoi="http://doi.org/10.5281/zenodo.4648518" httpUri="https://zenodo.org/record/4648518/files/figure.png" pageId="7" pageNumber="6">Fig. 9</figureCitation>
). ..................
<taxonomicName id="E6D34D66FFE8FF8CFDEFF8A3FCB459D6" authorityName="Pocock" authorityYear="1893" box="[629,770,1899,1922]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="13A7EAF7FFE8FF8CFDEFF8A3FCB459D6" box="[629,770,1899,1922]" italics="true" pageId="7" pageNumber="6">Heteroctenus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="216C36E5FFE8FF8CFFEBF843FCB55997" blockId="7.[113,771,1834,1987]" pageId="7" pageNumber="6">
<keyLead id="9A2295D0FFE8FF8CFFEBF843FCB55997" pageId="7" pageNumber="6">
Sternite III (the first post-pectinal) without longitudinal furrows or a feebly submedian pair only. .....
<taxonomicName id="E6D34D66FFE8FF8CFDEDF863FCB55997" authorityName="Marx" authorityYear="1890" box="[631,771,1963,1987]" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="13A7EAF7FFE8FF8CFDEDF863FCB55997" box="[631,771,1963,1987]" italics="true" pageId="7" pageNumber="6">Centruroides</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A272E40FFE8FF8CFCAAFF23FA745FF6" pageId="7" pageNumber="6">
<paragraph id="216C36E5FFE8FF8CFCAAFF23FA775F16" blockId="7.[816,1474,235,419]" pageId="7" pageNumber="6">
<keyLead id="9A2295D0FFE8FF8CFCAAFF23FA775F16" pageId="7" pageNumber="6">
6. Carapace subtriangular. Tergites with at least three welldeveloped longitudinal carinae. Pedipalp fingers with the median rows of denticles not imbricate. ..........
<taxonomicName id="E6D34D66FFE8FF8CFADDFEE3FA775F16" authorityName="Kjellesvig-Waering" authorityYear="1966" box="[1351,1473,299,322]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="13A7EAF7FFE8FF8CFADDFEE3FA775F16" box="[1351,1473,299,322]" italics="true" pageId="7" pageNumber="6">Microtityus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="216C36E5FFE8FF8CFCAAFE83FA745FF6" blockId="7.[816,1474,235,419]" pageId="7" pageNumber="6">
<keyLead id="9A2295D0FFE8FF8CFCAAFE83FA745FF6" pageId="7" pageNumber="6">
Carapace subrectangular. Tergites with a single longitudinal carina. Pedipalp fingers with the median rows of denticles clearly imbricate (
<figureCitation id="B9E82A60FFE8FF8CFB89FE43FBD75FF7" box="[1043,1121,395,419]" captionStart="Figures 811" captionStartId="5.[116,193,1031,1053]" captionTargetBox="[113,1474,234,1016]" captionTargetId="figure-490@5.[113,1474,234,1016]" captionTargetPageId="5" captionText="Figures 811. Buthid characters. Figure 8. Tityus obtusus, female in vivo habitus. Figure 9. Heteroctenus abudi, female, coxosternal region and sternite III. Figure 10. Heteroctenus abudi, pedipalp movable finger dentition, showing the median rows of denticles and the accessory denticles (10). Figure 11. Tityus sp., pedipalp movable finger dentition." figureDoi="http://doi.org/10.5281/zenodo.4648518" httpUri="https://zenodo.org/record/4648518/files/figure.png" pageId="7" pageNumber="6">Fig. 11</figureCitation>
). ........................................
<taxonomicName id="E6D34D66FFE8FF8CFA18FE43FA745FF6" authorityName="C. L. Koch" authorityYear="1836" box="[1410,1474,395,418]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="13A7EAF7FFE8FF8CFA18FE43FA745FF6" box="[1410,1474,395,418]" italics="true" pageId="7" pageNumber="6">Tityus</emphasis>
</taxonomicName>
</keyLead>
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</keyStep>
</key>
</subSubSection>
</treatment>
</document>

View file

@ -1,42 +1,42 @@
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<taxonomicName id="E6D34D66FFECFF88FCAAFA34FAA05840" ID-CoL="SBTR" authority="Santiago-Blay, 2009" authorityName="Santiago-Blay" authorityYear="2009" box="[816,1302,1532,1556]" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="jorgeorum">
<emphasis id="13A7EAF7FFECFF88FCAAFA34FB855840" box="[816,1075,1532,1556]" italics="true" pageId="3" pageNumber="2">Centruroides jorgeorum</emphasis>
Santiago-Blay, 2009
</taxonomicName>
@ -45,7 +45,6 @@ Santiago-Blay, 2009
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DISTRIBUTION.
<collectingCounty id="C80D4E69FFECFF88FC4DF9D4FBBA5860" box="[983,1036,1564,1588]" pageId="3" pageNumber="2">Only</collectingCounty>
known from the
@ -63,9 +62,9 @@ locality:
</date>
, personal effort for rediscovering it in the
<typeStatus id="FE688847FFECFF88FAA1F954FADF58E0" box="[1339,1385,1692,1716]" pageId="3" pageNumber="2">type</typeStatus>
locality was fruitless. It remains as a species pendant of confirmation (Teruel et al., 2015: 12)
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.
locality was fruitless. It remains as a species pendant of confirmation (
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).
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@ -1,42 +1,42 @@
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<emphasis id="13A7EAF7FFECFF88FFEBF9B4FEE758C0" box="[113,337,1660,1684]" italics="true" pageId="3" pageNumber="2">Centruroides griseus</emphasis>
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@ -54,32 +54,36 @@ and
,
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, Caja de Muertos, Magueyes). The presence of
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<taxonomicName id="E6D34D66FFECFF88FD97F915FDCF58A0" baseAuthorityName="C. L. Koch" baseAuthorityYear="1844" box="[525,633,1757,1780]" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="13A7EAF7FFECFF88FD97F915FDCF58A0" box="[525,633,1757,1780]" italics="true" pageId="3" pageNumber="2">C. griseus</emphasis>
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in Desecheo Island (Torres-González, 1965, cited by Santiago-Blay, 2009: 112) was considered by Teruel et al. (2015: 13) as needed of verification and probably erroneous. The Puerto Rican population was described by
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in Desecheo Island (Torres-González, 1965, cited by Santiago-Blay, 2009: 112) was considered by
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as needed of verification and probably erroneous. The Puerto Rican population was described by
<bibRefCitation id="45424B14FFECFF88FE2FF894FDFB5920" author="Armas" box="[437,589,1884,1908]" firstAuthor="Armas" pageId="3" pageNumber="2" pagination="1 - 25" refId="ref3503" refString="ARMAS, L. F. DE. 1982. Adiciones a las escorpiofaunas (Arachnida: Scorpiones) de Puerto Rico y de la Republica Dominicana. Poeyana, 237: 1 - 25." type="journal article" year="1982">Armas (1982)</bibRefCitation>
as
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<taxonomicName id="E6D34D66FFECFF88FDEDF894FEEA59C0" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="subSpecies" species="griseus" subSpecies="borinquensis">
<emphasis id="13A7EAF7FFECFF88FDEDF894FEEA59C0" italics="true" pageId="3" pageNumber="2">Centruroides griseus borinquensis</emphasis>
</taxonomicName>
, but later downgraded by him to subspecies (
<bibRefCitation id="45424B14FFECFF88FF6CF854FE0159E0" author="ARMAS" box="[246,439,1948,1972]" pageId="3" pageNumber="2" refId="ref3576" refString="ARMAS, L. F. DE. 1988. Sinopsis de los escorpiones antillanos. Editorial Cientifico-Tecnica, La Habana, 102 pp." type="book" year="1988">Armas, 1988: 50</bibRefCitation>
<bibRefCitation id="45424B14FFECFF88FF6CF854FE0159E0" author="Armas" box="[246,439,1948,1972]" firstAuthor="Armas" page="50" pageId="3" pageNumber="2" refId="ref3576" refString="ARMAS, L. F. DE. 1988. Sinopsis de los escorpiones antillanos. Editorial Cientifico-Tecnica, La Habana, 102 pp." type="book" year="1988">Armas, 1988: 50</bibRefCitation>
). Nevertheless, Santiago-Blay (2009: 112) mentioned it as a subspecies of
<taxonomicName id="E6D34D66FFECFF88FDD4F875FF4F59A0" authority="(C. L. Koch, 1844)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1844" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="griseus">
<taxonomicName id="E6D34D66FFECFF88FDD4F875FF4F59A0" authority="(C. L. Koch, 1844)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1844" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="13A7EAF7FFECFF88FDD4F875FD0D5980" box="[590,699,1981,2004]" italics="true" pageId="3" pageNumber="2">C. griseus</emphasis>
(C. L. Koch, 1844)
</taxonomicName>
, whereas Esposito &amp; Prendini (2019) removed it from the synonymy with
<taxonomicName id="E6D34D66FFECFF88FBA3FB14FB125AA0" baseAuthorityName="C. L. Koch" baseAuthorityYear="1844" box="[1081,1188,1244,1268]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="griseus">
, whereas
<bibRefCitation id="45424B14FFECFF88FEF9F814FD3F59A0" author="ESPOSITO, L. A. &amp; L. PRENDINI." box="[355,649,2012,2036]" journalOrPublisher="Scientific Reports" pageId="3" pageNumber="2" pagination="3500" part="9" refId="ref4016" refString="ESPOSITO, L. A. &amp; L. PRENDINI. 2019. Island ancestors and New World biogeography: A case study from the scorpions (Buthidae: Centruroidinae). Scientific Reports, (2019) 9: 3500 (https: // doi. org / 10.1038 / s 41598 - 018 - 33754 - 8)." title="Island ancestors and New World biogeography: A case study from the scorpions (Buthidae: Centruroidinae)" type="journal article" year="2019">Esposito &amp; Prendini (2019)</bibRefCitation>
removed it from the synonymy with
<taxonomicName id="E6D34D66FFECFF88FBA3FB14FB125AA0" baseAuthorityName="C. L. Koch" baseAuthorityYear="1844" box="[1081,1188,1244,1268]" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="13A7EAF7FFECFF88FBA3FB14FB125AA0" box="[1081,1188,1244,1268]" italics="true" pageId="3" pageNumber="2">C. griseus</emphasis>
</taxonomicName>
, and raised it to the species level, a nomenclatural action overlooked by Crew &amp; Esposito (2020: fig. 11), who again mentioned it as a subspecies of
<taxonomicName id="E6D34D66FFECFF88FA30FAD4FCCB5B00" baseAuthorityName="C. L. Koch" baseAuthorityYear="1844" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="griseus">
<taxonomicName id="E6D34D66FFECFF88FA30FAD4FCCB5B00" baseAuthorityName="C. L. Koch" baseAuthorityYear="1844" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="13A7EAF7FFECFF88FA30FAD4FCCB5B00" italics="true" pageId="3" pageNumber="2">C. griseus</emphasis>
</taxonomicName>
. If
<taxonomicName id="E6D34D66FFECFF88FC32FAF4FB1F5B00" box="[936,1193,1340,1364]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="subSpecies" species="griseus" subSpecies="borinquensis">
<taxonomicName id="E6D34D66FFECFF88FC32FAF4FB1F5B00" box="[936,1193,1340,1364]" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="subSpecies" species="griseus" subSpecies="borinquensis">
<emphasis id="13A7EAF7FFECFF88FC32FAF4FB1F5B00" box="[936,1193,1340,1364]" italics="true" pageId="3" pageNumber="2">C. griseus borinquensis</emphasis>
</taxonomicName>
is a good species, then it would be most strongly demonstrated. Consequently, at this moment caution seems preferable.This is the most common scorpion in

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@ -1,42 +1,42 @@
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<subSubSection id="69C9656EFFECFF88FFEBFA97FD1F5B2C" box="[113,681,1375,1400]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="216C36E5FFECFF88FFEBFA97FD1F5B2C" blockId="3.[113,772,1343,1623]" box="[113,681,1375,1400]" pageId="3" pageNumber="2">
<taxonomicName id="E6D34D66FFECFF88FFEBFA97FD135B2C" authority="Armas &amp; Marcano Fondeur, 1987" authorityName="Armas &amp; Marcano Fondeur" authorityYear="1987" box="[113,677,1375,1400]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="bani">
<taxonomicName id="E6D34D66FFECFF88FFEBFA97FD135B2C" ID-CoL="SBSM" authority="Armas &amp; Marcano Fondeur, 1987" authorityName="Armas &amp; Marcano Fondeur" authorityYear="1987" box="[113,677,1375,1400]" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="bani">
<emphasis id="13A7EAF7FFECFF88FFEBFA97FE855B23" box="[113,307,1375,1399]" italics="true" pageId="3" pageNumber="2">Centruroides bani</emphasis>
Armas &amp; Marcano Fondeur, 1987
</taxonomicName>
@ -48,14 +48,16 @@ Armas &amp; Marcano Fondeur, 1987
DISTRIBUTION. Hispaniola and Mona Island (
<collectingCountry id="59C47675FFECFF88FDF7FA48FD4E5BC3" box="[621,760,1407,1432]" name="Puerto Rico" pageId="3" pageNumber="2">Puerto Rico</collectingCountry>
). This species was recorded from Mona Island by
<bibRefCitation id="45424B14FFECFF88FD11FA57FEB25B8C" author="ESPOSITO" pageId="3" pageNumber="2" pagination="1 - 134" refId="ref4077" refString="ESPOSITO, L. A., H. Y. YAMAGUTI, C. A. DE SOUZA, R. PINTO-DA-ROCHA &amp; L. PRENDINI. 2017. Systematic revision of the Neotropical club-tailed scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, revalidation of Heteroctenus, and descriptions of two new genera and three new species (Buthidae: Rhopalurusinae). Bulletin of the American Museum of Natural History, 415: 1 - 134." type="journal article" year="2017">Esposito et al. (2017: 27)</bibRefCitation>
, but without mentioning the material examined. Teruel et al. (2017) synonymizes
<taxonomicName id="E6D34D66FFECFF88FE69FA17FEFB584C" authority="Santiago-Blay, 2009" authorityName="Santiago-Blay" authorityYear="2009" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="mariaorum">
<bibRefCitation id="45424B14FFECFF88FD11FA57FEB25B8C" author="Esposito" etAl="et al." firstAuthor="Esposito" page="27" pageId="3" pageNumber="2" pagination="1 - 134" refId="ref4077" refString="ESPOSITO, L. A., H. Y. YAMAGUTI, C. A. DE SOUZA, R. PINTO-DA-ROCHA &amp; L. PRENDINI. 2017. Systematic revision of the Neotropical club-tailed scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, revalidation of Heteroctenus, and descriptions of two new genera and three new species (Buthidae: Rhopalurusinae). Bulletin of the American Museum of Natural History, 415: 1 - 134." type="journal article" year="2017">Esposito et al. (2017: 27)</bibRefCitation>
, but without mentioning the material examined.
<bibRefCitation id="45424B14FFECFF88FFEBFA17FEF85BAC" author="TERUEL" box="[113,334,1503,1528]" pageId="3" pageNumber="2" pagination="1 - 15" refId="ref5073" refString="TERUEL, R., M. J. RIVERA &amp; A. J. SANCHEZ. 2017. The scorpion fauna of Mona Island, Puerto Rico (Scorpiones: Buthidae, Scorpionidae). Euscorpius, 250: 1 - 15." type="journal article" year="2017">Teruel et al. (2017)</bibRefCitation>
synonymizes
<taxonomicName id="E6D34D66FFECFF88FE69FA17FEFB584C" authority="Santiago-Blay, 2009" authorityName="Santiago-Blay" authorityYear="2009" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="mariaorum">
<emphasis id="13A7EAF7FFECFF88FE69FA17FCB55BA3" box="[499,771,1503,1527]" italics="true" pageId="3" pageNumber="2">Centruroides mariaorum</emphasis>
Santiago-Blay, 2009
</taxonomicName>
, described from Mona Island, to
<taxonomicName id="E6D34D66FFECFF88FD34F9C8FD4A5843" authorityName="Armas &amp; Marcano Fondeur" authorityYear="1987" box="[686,764,1535,1559]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="bani">
<taxonomicName id="E6D34D66FFECFF88FD34F9C8FD4A5843" authorityName="Armas &amp; Marcano Fondeur" authorityYear="1987" box="[686,764,1535,1559]" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="bani">
<emphasis id="13A7EAF7FFECFF88FD34F9C8FD4A5843" box="[686,764,1535,1559]" italics="true" pageId="3" pageNumber="2">C. bani</emphasis>
</taxonomicName>
, and added new localities from this small insular territory, as well as notes on its habitat.

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@ -1,42 +1,42 @@
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<paragraph id="216C36E5FFECFF88FCAAF8D4FB505960" blockId="3.[816,1474,1820,2036]" box="[816,1254,1820,1844]" pageId="3" pageNumber="2">
<taxonomicName id="E6D34D66FFECFF88FCAAF8D4FB545960" authority="Santiago-Blay, 2009" authorityName="Santiago-Blay" authorityYear="2009" box="[816,1250,1820,1844]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="sasae">
<taxonomicName id="E6D34D66FFECFF88FCAAF8D4FB545960" ID-CoL="5XGZZ" authority="Santiago-Blay, 2009" authorityName="Santiago-Blay" authorityYear="2009" box="[816,1250,1820,1844]" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="sasae">
<emphasis id="13A7EAF7FFECFF88FCAAF8D4FC485960" box="[816,1022,1820,1844]" italics="true" pageId="3" pageNumber="2">Centruroides sasae</emphasis>
Santiago-Blay, 2009
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@ -48,13 +48,17 @@ Santiago-Blay, 2009
DISTRIBUTION. Only known from the
<typeStatus id="FE688847FFECFF88FB48F8F4FAB65900" box="[1234,1280,1852,1876]" pageId="3" pageNumber="2">type</typeStatus>
locality: Caja de Muertos Island. This species was described on the basis of a single adult male, which is the only known specimen. Its description is very inadequate and efforts for rediscovering it have been unproductive, because all the collected specimens were
<taxonomicName id="E6D34D66FFECFF88FCF6F814FC6C59A0" baseAuthorityName="C. L. Koch" baseAuthorityYear="1844" box="[876,986,2012,2036]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="griseus">
<taxonomicName id="E6D34D66FFECFF88FCF6F814FC6C59A0" baseAuthorityName="C. L. Koch" baseAuthorityYear="1844" box="[876,986,2012,2036]" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="13A7EAF7FFECFF88FCF6F814FC6C59A0" box="[876,986,2012,2036]" italics="true" pageId="3" pageNumber="2">C. griseus</emphasis>
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(
<bibRefCitation id="45424B14FFECFF88FC73F814FBCF59A0" author="ARMAS" box="[1001,1145,2012,2036]" pageId="3" pageNumber="2" pagination="55 - 64" refId="ref3858" refString="ARMAS, L. F. DE. 2010. Nuevos aracnidos de Puerto Rico (Arachnida: Amblypygi, Araneae, Opiliones, Parasitiformes, Schizomida, Scorpiones). Boletin de la Sociedad Entomologica Aragonesa, 47: 55 - 64." type="journal article" year="2010">Armas, 2010</bibRefCitation>
; Teruel et al., 2015). In a very recent paper on Caribbean arthropod biogeography, Crew &amp; Esposito (2020: fig. 11) included this dubious species in their analyses, but likely they were in error. Its taxonomical status is pendant of confirmation (Teruel et al., 2015: 12; but also see
<bibRefCitation id="45424B14FFEBFF8FFFEBF8EDFE915969" author="ARMAS" box="[113,295,1829,1853]" pageId="4" pageNumber="3" pagination="55 - 64" refId="ref3858" refString="ARMAS, L. F. DE. 2010. Nuevos aracnidos de Puerto Rico (Arachnida: Amblypygi, Araneae, Opiliones, Parasitiformes, Schizomida, Scorpiones). Boletin de la Sociedad Entomologica Aragonesa, 47: 55 - 64." type="journal article" year="2010">Armas, 2010: 62</bibRefCitation>
<bibRefCitation id="45424B14FFECFF88FC73F814FBCF59A0" author="Armas" box="[1001,1145,2012,2036]" firstAuthor="Armas" pageId="3" pageNumber="2" pagination="55 - 64" refId="ref3858" refString="ARMAS, L. F. DE. 2010. Nuevos aracnidos de Puerto Rico (Arachnida: Amblypygi, Araneae, Opiliones, Parasitiformes, Schizomida, Scorpiones). Boletin de la Sociedad Entomologica Aragonesa, 47: 55 - 64." type="journal article" year="2010">Armas, 2010</bibRefCitation>
;
<bibRefCitation id="45424B14FFECFF88FB1EF814FAF859A0" author="TERUEL" box="[1156,1358,2012,2036]" pageId="3" pageNumber="2" pagination="1 - 15" refId="ref5117" refString="TERUEL, R., M. J. RIVERA &amp; C. J. SANTOS. 2015. Two new scorpions from the Puerto Rican island of Vieques, Greater Antilles (Scorpiones: Buthidae). Euscorpius, 208: 1 - 15." type="journal article" year="2015">Teruel et al., 2015</bibRefCitation>
). In a very recent paper on Caribbean arthropod biogeography, Crew &amp; Esposito (2020: fig. 11) included this dubious species in their analyses, but likely they were in error. Its taxonomical status is pendant of confirmation (
<bibRefCitation id="45424B14FFEBFF8FFE0EF8CDFDC85949" author="TERUEL" box="[404,638,1797,1821]" pageId="4" pageNumber="3" pagination="1 - 15" refId="ref5117" refString="TERUEL, R., M. J. RIVERA &amp; C. J. SANTOS. 2015. Two new scorpions from the Puerto Rican island of Vieques, Greater Antilles (Scorpiones: Buthidae). Euscorpius, 208: 1 - 15." type="journal article" year="2015">Teruel et al., 2015: 12</bibRefCitation>
; but also see
<bibRefCitation id="45424B14FFEBFF8FFFEBF8EDFE915969" author="Armas" box="[113,295,1829,1853]" firstAuthor="Armas" page="62" pageId="4" pageNumber="3" pagination="55 - 64" refId="ref3858" refString="ARMAS, L. F. DE. 2010. Nuevos aracnidos de Puerto Rico (Arachnida: Amblypygi, Araneae, Opiliones, Parasitiformes, Schizomida, Scorpiones). Boletin de la Sociedad Entomologica Aragonesa, 47: 55 - 64." type="journal article" year="2010">Armas, 2010: 62</bibRefCitation>
).
</paragraph>
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