From c4b0c0fabdba41c0a65a5dcd6160e17cc2b6f2e2 Mon Sep 17 00:00:00 2001 From: ggserver Date: Tue, 10 Dec 2024 23:42:51 +0000 Subject: [PATCH] Add updates up until 2024-12-10 23:37:32 --- .../87/03E687A9993CFFB12B8F7DF77493FD86.xml | 597 ++++++++++++++++++ .../97/CE229727FF9BFF93FC003182FD08FA96.xml | 135 ++-- .../97/CE229727FF9BFF96FE893762FA8CF896.xml | 129 ++-- 3 files changed, 732 insertions(+), 129 deletions(-) create mode 100644 data/03/E6/87/03E687A9993CFFB12B8F7DF77493FD86.xml diff --git a/data/03/E6/87/03E687A9993CFFB12B8F7DF77493FD86.xml b/data/03/E6/87/03E687A9993CFFB12B8F7DF77493FD86.xml new file mode 100644 index 00000000000..d119bd6b3ac --- /dev/null +++ b/data/03/E6/87/03E687A9993CFFB12B8F7DF77493FD86.xml @@ -0,0 +1,597 @@ + + + +New species of Paraodontomma from mid-Cretaceous Burmese amber with muscle tissue preservation (Coleoptera: Archostemata: Ommatidae) + + + +Author + +Li, Yan-Da + + + +Author + +Yamamoto, Shûhei + + + +Author + +Huang, Di-Ying + + + +Author + +Cai, Chen-Yang + +text + + +Papéis Avulsos de Zoologia + + +2021 + +2021-06-04 + + +61 + + +1 +13 + + + + +http://dx.doi.org/10.11606/1807-0205/2021.61.53 + +journal article +10.11606/1807-0205/2021.61.53 +1807-0205 +5213117 +D9EBBBAC-B94A-4347-8E30-C9FA8E3D5B4D + + + + + + +Paraodontommaleptocristatum + +Li,Yamamoto & Cai sp.nov. ( +Figs. 1-8 +) + + + + + +Material: + +Holotype +, +NIGP174676 +, +adult +male +, covered with numerous bubbles + +. + + +Locality and horizon: +Amber mine located +near Noije Bum Village +, +Tanai Township +, +Myitkyina District +, +Kachin State +, +Myanmar +; unnamed horizon, mid-Cretaceous, +Upper Albian to Lower Cenomanian +. + + + + +Diagnosis: +Head ( +Figs. 2A, B +, +5 +) subquadrate (elongate in + +P. burmiticum + +and + +P. szwedoi + +), without prominent posterior protuberances (with prominent protuberances above eyes in + +P. szwedoi +, + +and also possibly in + +P. burmiticum +; + +Fig. 10A +). Pronotal disc without prominent ridges ( +Fig. 2C +) (with longitudinal ridges in + +P. szwedoi + +; +Fig. 10C +). Elytra with indistinct longitudinal and transverse ridges ( +Figs. 2F +, +6D +) (strongly developed in + +P. burmiticum + +and + +P. szwedoi +; + +Fig. 10I +). Teeth along outer elytral margins not forming a wave pattern ( +Figs. 2E +, +4D +) (wavily arranged in + +P. burmiticum + +and + +P.szwedoi +; + +Figs. 10F +, +11C +). + + + + +Figure 2. Details of + +Paraodontomma leptocristatum + +sp. nov. +, holotype, NIGP174676, under widefield fluorescence. (A) Head, anteroventral view. (B) Head, posterodorsal view. (C) Prothorax, dorsal view. (D) Prothorax, ventral view. (E) Mid leg and metathorax, ventral view. (F) Elytra, dorsal view. (G) Hind leg, ventral view. (H) Abdominal apex, ventral view. Abbreviations: an1-3 = antennomeres 1-3; ep = epipleuron; es = elytral suture; ey = compound eye; hd = head; msf = mesofemur;msts = mesotarsus;mtf = metafemur;mttb = metatibia; mtv = metaventrite;pc = procoxa;pf = profemur;pn = pronotum; pp = propleuron;ps = prosternum;ptb = protibia;pts = protarsus;v1-5 = ventrites1-5. Scale bars:500 μm. + + + + +Description: +Body length about +8.1 mm +; maximum width about +3.6 mm +between elytral margins in posterior +⅓ +of body. Head and prothorax slender compared to compact unit formed by pterothorax and abdomen; shoulders pronounced. Integument covered with rounded tubercles and setae. + + +Head ( +Figs. 2A, B +, +5 +, +6A +) prognathous, constricted posteriorly to form a neck (though not well visible on the specimen); maximum width about +1.4 mm +(including eyes); dorsal surface without prominent posterior protuberances. Compound eyes relatively large, protruding laterally. Antennal insertion area located anteriorly, anteromesad anterior margin of compound eyes, separated by more than one but less than two diameters of scapus. Antennal grooves absent on dorsal or ventral side. Postocular temples short. + + + +Figure 3. Details of + +Paraodontomma leptocristatum + +sp.nov. +, holotype,NIGP174676,under confocal microscopy.(A) Mouthparts,anterior view.(B)Procoxae,ventral view.(C) Mesocoxae,ventral view.(D) Epipleuron,ventral view,showing the nonwavily arranged teeth along the outer elytral margin(arrowheads).(E)Mesotarsus, ventral view.(F) Metatarsus,ventral view.(G) Abdominal apex,ventral view.(H) Anterior portion of head,posterodorsal view. (I) Elytra,dorsal view.Abbreviations: an1 = antennomere 1; ep = epipleuron; es = elytral suture; lb = labrum; lbp = labial palp; md = mandible; msc = mesocoxa; mst1-5 = mesotarsomeres 1-5; mtt1-5 = metatarsomeres 1-5; mtv = metaventrite; mxp = maxillary palp; pc = procoxa; ps = prosternum; ptb = protibia; v2,5 = ventrite 2,5; wp = window punctures.Scale bars:200 μm. + + + +Labrum transverse, with tuberculate dorsal surface and anterior margin densely set with setae ( +Fig. 3A +). Antenna ( +Figs. 2A +, +5 +) short, extending beyond anterior prothoracic margin when posteriorly directed, but not reaching posterior prothoracic margin, 11-segmented and submoniliform, with thin and short setae; antennomere 1 distinctly wider than other antennomeres, somewhat globular; antennomere 2 distinctly smaller, subquadrate, about as wide as long; antennomere 3 elongate,slightly widening distally;antennomeres 4-10 short, subequal in length; antennomere 11 spindle-shaped, tapering apically. Ratio of antennomere lengths: ∼ 3.4: 2.3: 3.4: 2.1: 2.1: 1.9: 1.8: 1.8: 1.9: 1.8: 2.9. Mandibles tridentate with vertically arranged teeth ( +Fig. 3A +). Apical maxillary and labial palpomeres slender, not expanded, with a small cavity near apex ( +Fig. 3A +). Prementum with deep posteromedian pit. Separate mentum present, transverse. Gular sutures not visible. + + + +Figure 4. X-ray microtomographic reconstruction of + +Paraodontomma leptocristatum + +sp. nov. +, holotype, NIGP174676.(A) Dorsal view. (B) Ventral view.(C) Lateral view.Scale bar:1.5 mm. + + + +Pronotal disc ( +Fig. 2C +) anteriorly distinctly narrower than head, posteriorly about as wide, roughly hexagonal, with rounded lateral edges; length about +1.3 mm +; maximum width about +1.6 mm +; lateral margin strongly serrated; anterolateral and posterolateral pronotal corners angles not projecting and rounded; surface without protuberances or furrows, evenly convex. Pronotal hypomeron narrow, separated from exposed propleuron by slightly curved suture ( +Fig. 1B +). Pleurosternal suture straight ( +Fig. 1B +). Prosternum ( +Fig. 2D +) quadrangular, about as wide as long, very slightly narrowing anteriorly, with slightly convex anterior margin; prosternal process short and narrow, not reaching middle region of procoxae ( +Fig. 3B +). Procoxal cavities contiguous posterior to prosternal process, posteriorly open ( +Fig. 3B +). Procoxae globular to cone-shaped. + + +Shoulder region pronounced. Mesoscutellar shield not visible (hidden by incrustations). Mesopleura and mesoventrite not clearly visible, partly concealed by legs and partly blurred. Mesocoxal cavities contiguous ( +Fig. 3C +). Mesocoxae similar to procoxae in size and shape. Metanepisternum large, broad anteriorly ( +Fig. 3E +); exposed part of metepimeron moderately sized, visible between elytral epipleura and concave posterolateral metanepisternal edge ( +Fig. 3E +). Metaventrite trapezoidal, strongly narrowing anteriorly, with distinct anteromedian projection; discrimen not visible, probably missing; metakatepisternal suture distinct and complete; metatrochantin distinctly exposed. Metacoxae medially contiguous, transverse, distinctly reaching beyond lateral margin of metaventrite and reaching elytral epipleura laterally; metacoxal plates absent but excavation for metafemur at rest present. + + +Elytra about +5.5 mm +long, each +1.8 mm +wide in middle region, with setae rooted on tubercles along longitudinal ridges (elytral veins) and small rounded scales on surface ( +Figs. 3F +, +4I +). Elytral disc separated into 2 × 2 squares by very indistinct longitudinal and transverse ridges ( +Fig. 6D +). Epipleural rim moderately wide, about +⅓ +of elytral width anteriorly, evenly narrowing posteriorly, without window punctures ( +Figs. 2E +, +3D +). Teeth along outer elytral margins present, but not forming a wave pattern. + + +Legs ( +Figs. 2D, E, G +, +3F +) slender; all three pairs similar in size and shape. Femora robust, widest in middle region. Tibiae about as long as femur, slender. Tarsi 5-segmented, cylindrical; with simple tarsomeres, without distinct brushes of adhesive microtrichia. Length ratio of protarsomeres: ∼ 1.8: 1.3: 1.0: 1.0: 1.1. Length ratio of mesotarsomeres:∼ 5.3: 1.5: 1.4: 1.1: 3.4. Length ratio of metatarsomeres: ∼ 6.7: 2.2: 1.4: 1.2: 3.8. + + +Abdomen broad, with five coplanar ventrites, separated by distinct grooves ( +Fig. 2H +); ventrites 2-4 subequal in length; ventrites 5 about 1.3 times as long as ventrite 4, with broadly rounded apex ( +Fig. 3G +). Aedeagus possibly trilobate, as shown in +Fig. 6E +. + + + + +Etymology: +The specific name is derived from Greek +“leptos”, +thin, and Latin +“crista”, +crest, referring to the weak ridges on the elytra. + + + +Figure 5. X-ray microtomographic reconstruction of + +Paraodontomma leptocristatum + +sp. nov. +, holotype,NIGP174676. Note that some bubbles were not removed. (A) Head,dorsal view.(B) Head,ventral view.Scale bar:750 μm. + + + + +Figure 6. Virtual slices of + +Paraodontomma leptocristatum + +sp. nov. +, holotype, NIGP174676, based on X-ray microtomographic reconstruction. (A) Head, horizontal sections.(B)Metathorax,sagittal sections.(C) Metathorax,subhorizontal sections.(D) Elytron,horizontal sections.(E) Aedeagus,horizontal sections.Abbreviations: Mh11 = Musculus cranio-mandibularis internus;Mt84 = Musculus metanoto-sternalis;pe = penis.Scale bars:800 μm. + + + +The genus name + +Paraodontomma + +is an extended version of the genus name + +Omma +Newman + +, which is a neuter noun in Ancient Greek, meaning eye. Thus, according to Article 30.1.2 of ICZN (1999), the gender of + +Paraodontomma + +should also be neuter, rather than feminine as originally claimed by +Yamamoto (2017) +. + + + + + +DISCUSSION + + + +Yamamoto (2017) +placed + +Paraodontomma + +in the tribe +Brochocoleini +, and further suggested a close relationship between + +Paraodontomma + +and + +Odontomma +. + +Brochocoleini +defined by Tan +et al. +(2012) is characterized by more than one row of window punctures on the elytral epipleura ( +e.g., +fig. +73 in +Hong, 1982 +; fig. +1 in +Liu +et al., +2017). However, our new observation revealed that window punctures are absent on the epipleural rim of + +Paraodontomma + +( +Figs. 2E +, +3D +, +10F +, +11C +). Therefore, its tribal placement should be questioned and investigated in the future. +Yamamoto (2017) +ruled out a close relationship of the extinct genus to fossils with wide epipleural rims assigned to + +Tetraphalerus +, + +based on the lack of characteristic antennal grooves in + +Paraodontomma +. + +However, no antennal grooves have been validly observed in any of the Mesozoic + +“ +Tetraphalerus +” + +fossils (Li +et al., +2021), and many of them have been transferred into genus + +Allophalerus +Kirejtshuk + +just recently ( +Kirejtshuk, 2020 +). The overall shape of some + +Allophalerus + +is also similar to + +Paraodontomma +. + +Therefore, it is ambivalent whether + +Paraodontomma + +is closer to + +Odontomma + +or + +Allophalerus + +based on currently available evidence. + + + +Figure 7. Details of + +Paraodontomma leptocristatum + +sp.nov. +, holotype,NIGP174676,under confocal microscopy,with depth color-coding.(A) Mouthparts,anterior view.(B) Procoxae,ventral view.(C) Mesocoxae,ventral view.(D)Epipleuron,ventral view.(E) Mesotarsus,ventral view.(F) Metatarsus,ventral view.(G) Abdominal apex,ventral view.(H) Anterior portion of head,posterodorsal view.(I) Elytra,dorsal view.Scale bars:200 μm. + + + + +Figure 8. Serial horizontal virtual sections of + +Paraodontomma leptocristatum + +sp. nov. +, holotype, NIGP174676, based on X-ray microtomographic reconstruction. Scale bar:1.5 mm. + + + +An eye-catching character of + +Paraodontomma + +is the presence of transverse ridges on the elytra, which are rarely seen in other +Ommatidae +and in +Cupedidae +. The transverse ridges, together with longitudinal ridges (elytral veins), partition the elytra into 2 × 2 squares, each containing four window punctures. These transverse ridges are rather prominent in + +P. burmiticum + +and + +P. szwedoi + +(fig. +1 in +Yamamoto, 2017 +; figs. +1-2 in +Jarzembowski +et al., +2018). Although these ridges are quite indistinct in + +P. leptocristatum + + +sp. nov. + +, the 2 × 2 squares are still discernable through the micro-CT slices ( +Fig. 6D +). We thus suggest that the transverse ridges on elytra could potentially be an autapomorphy of genus + +Paraodontomma +. + +The body and elytra of some +Cupedidae +and +Ommatidae +are decorated with different scales. Here confocal microscopy revealed a new +type +of small and rounded scales on the elytra of + +Paraodontomma + +( +Figs. 3I +, +11F +).The outer elytral margins of + +P. burmiticum + +and + +P. szwedoi + +are armed with somewhat wavily arranged teeth (serration) ( +Figs. 10F +, +11C +; fig. 1A in +Yamamoto, 2017 +), while in + +P.leptocristatum + + +sp. nov. + +the arrangement of elytral teeth does not form a wave pattern ( +Figs. 2E +, +4D +). The head of + +P. burmiticum + +and + +P. szwedoi + +is relatively elongate, with the portion in front of eyes longer than the eye ( +Fig. 10A +; fig. 2A in +Yamamoto, 2017 +), while the head of + +P. leptocristatum + + +sp. nov. + +is subquadrate, with portion in front of eyes shorter than eye ( +Fig. 5A +). + + +Under micro-CT, the beetle from Burmese amber often appears as a subhomogenous highly-absorbing solid substrate or a void cavity ( +e.g., +Jałoszyński +et al., +2020). In contrast, micro-CT revealed fine internal softparts of our + +P. leptocristatum + +specimen ( +Figs. 6 +, +8 +). The striated musculature could be clearly seen in the metathorax of the beetle ( +Figs. 6B, C +), and correspond well to that of extant ommatid + +Tetraphalerus +Waterhouse (Friedrich +et al., +2009) + +. The muscles showed no apparent shrinkage (similar to Grimaldi +et al., +1994; but different from +Henwood, 1992b +), possibly suggesting a very rapid mummification. Unfortunately, due to the presence of taphonomic artefact and the limitation of resolution, we were only able to identify a few very large muscles. For example, +Fig. 6C +shows the largest muscle of metathorax, Musculus metanoto-sternalis, with one end attached on metaventrite. This muscle is present in +Archostemata +as a plesiomorphy but missing in many other groups. The minor muscles with more phylogenetic information were hard to discern in our specimen. + + + + \ No newline at end of file diff --git a/data/CE/22/97/CE229727FF9BFF93FC003182FD08FA96.xml b/data/CE/22/97/CE229727FF9BFF93FC003182FD08FA96.xml index ec7e5078b68..f018e8f7aad 100644 --- a/data/CE/22/97/CE229727FF9BFF93FC003182FD08FA96.xml +++ b/data/CE/22/97/CE229727FF9BFF93FC003182FD08FA96.xml @@ -1,53 +1,56 @@ - - - -Magnana, a new genus of Gyponini from Southeastern Brazil (Hemiptera: Cicadellidae: Iassinae) + + + +Magnana, a new genus of Gyponini from Southeastern Brazil (Hemiptera: Cicadellidae: Iassinae) - - -Author + + +Author -Domahovski, Alexandre Cruz -Universidade Federal do Paraná (UFPR), Setor de Ciências Biológicas, Departamento de Zoologia (DZOO). Curitiba, PR, Brasil. +Domahovski, Alexandre Cruz +Universidade Federal do Paraná (UFPR), Setor de Ciências Biológicas, Departamento de Zoologia (DZOO). Curitiba, PR, Brasil. - - -Author + + +Author -Cavichioli, Rodney Ramiro -Universidade Federal do Paraná (UFPR), Setor de Ciências Biológicas, Departamento de Zoologia (DZOO). Curitiba, PR, Brasil. +Cavichioli, Rodney Ramiro +Universidade Federal do Paraná (UFPR), Setor de Ciências Biológicas, Departamento de Zoologia (DZOO). Curitiba, PR, Brasil. -text - - -Papéis Avulsos de Zoologia +text + + +Papéis Avulsos de Zoologia - -2021 - -Pap. Avulsos Zool., S. Paulo + +2021 + +Pap. Avulsos Zool., S. Paulo - -2021-03-31 + +2021-03-31 - -61 + +61 - -1 -5 + +1 +5 - -http://dx.doi.org/10.11606/1807-0205/2021.61.40 + +http://dx.doi.org/10.11606/1807-0205/2021.61.40 -journal article -10.11606/1807-0205/2021.61.40 -1807-0205 -2A9E2AB5-CD8D-4C5C-BACF-28388282ACA3 +journal article +306322 +10.11606/1807-0205/2021.61.40 +707a46e7-a896-40fa-b2ce-65603cb807a8 +1807-0205 +14371050 +2A9E2AB5-CD8D-4C5C-BACF-28388282ACA3 - + @@ -63,7 +66,7 @@ - + Figs. 1-17 @@ -73,15 +76,15 @@ Diagnosis: Inner margin of pygofer ( -Fig. 9 +Fig. 9 ) serrated apically; style ( -Fig. 13 +Fig. 13 ) elongated and sigmoid; shaft of aedeagus ( -Figs. 14, 15 +Figs. 14, 15 ) bearing one short subapical process on anterior surface, curved ventrally, with pair of apical processes, slightly longer, branched at apex, branches directed ventrally. - + Figures 1-15. @@ -112,33 +115,33 @@ Total length: 18.7; head length: 1.4; transocular width: 3.6; humeral width of p Color: Mostly yellow ( -Figs. 16, 17 +Figs. 16, 17 ). Crown ( -Fig. 1 +Fig. 1 ) with black maculae on following areas: posterior margin, behind ocelli; inner margin of eyes, surrounding ocelli and extending to anterior margin; and apex of anterior margin. Pronotum ( -Fig. 1 +Fig. 1 ): several large round black maculae on disc and anterior portion; posterior margin with transverse irregular-shape black macula. Mesonotum and scutellum ( -Fig. 1 +Fig. 1 ) with transverse black macula. Face ( -Fig. 2 +Fig. 2 ): transition crown-face black; area surrounding antenal pits black; gena with large black spots medially; ventral portion of frons, lora and clypeus black. Proepimeron ( -Fig. 3 +Fig. 3 ) with ventral half black. Forewing ( -Fig. 4 +Fig. 4 ) with several large black maculae; costal margin orange. Profemur ( -Fig. 3 +Fig. 3 ) with base and apex black. Protibia ( -Fig. 3 +Fig. 3 ) and protarsus black. Mesofemur and mesotibia ( -Fig. 17 +Fig. 17 ) with base and apex black. Mesotarsus black. Metafemur and metatibia ( -Figs. 16, 17 +Figs. 16, 17 ) with base and apex black. Metatarsus ( -Fig. 5 +Fig. 5 ) with third tarsomere black. Abdomen with dorsal portion black; sternite III almost entire black; sternites IV-VII with pair of black maculae on anterolateral angles; sternite VIII ( -Fig. 6 +Fig. 6 ) almost entire black, anterior and lateral margins yellow. Pygofer ( -Fig. 8 +Fig. 8 ) with two black spots on anterior portion. @@ -148,28 +151,28 @@ External morphological characters as in generic description. Male terminalia: Sternite VIII ( -Fig. 5 +Fig. 5 ), in ventral view, 1.5× wider than long; lateral margins slightly rounded and convergent posterad; posterior margin slightly undulated. Valve ( -Fig. 7 +Fig. 7 ), in ventral view, 2× wider than long; integument thickened on anterior margin only; posterior margin notched medially. Pygofer ( -Fig. 8 +Fig. 8 ), in lateral view, 1.8× longer than maximum height; ventral margin broadly round, bearing small setae; apex slightly tapered and round; macrosetae dispersed near apex and dorsal margin; in dorsal view ( -Fig. 9 +Fig. 9 ), apical portion of inner margin finely serrated, but not produced into process. Subgenital plate ( -Fig. 8 +Fig. 8 ), in lateral view, produced posteriorly almost as far as pygofer apex; in ventral view ( -Fig. 10 +Fig. 10 ), elongated, 4.5× longer than wide; lateral margins approximately straight; external margin and ventral surface bearing short setae; apex round. Connective ( -Fig. 11 +Fig. 11 ), in dorsal view, stalk present, wider than long; arms curved anterad, dorsal keel small. Style ( -Fig. 12 +Fig. 12 ), in dorsal view, with outer lobe well-developed, round; in lateral view ( -Fig. 13 +Fig. 13 ), blade elongated and sigmoid; external surface bearing small setae except on apical portion; apical portion with ventral margin serrated; apex slightly curved dorsally. Aedeagus ( -Figs. 14, 15 +Figs. 14, 15 ) with preatrium reduced; dorsal apodeme not expanded laterally; atrial processes with dorsal and ventral margins almost parallel; apex tapered and rounded, curved inwards; shaft tubular and elongated, curved dorsally near base and straight distally; apex with one short subapical process on anterior surface, curved ventrally, and pair of slightly longer apical processes, with branched apex, directed ventrally. - + Figures 16-17. diff --git a/data/CE/22/97/CE229727FF9BFF96FE893762FA8CF896.xml b/data/CE/22/97/CE229727FF9BFF96FE893762FA8CF896.xml index b5c3d8a38a9..16088117304 100644 --- a/data/CE/22/97/CE229727FF9BFF96FE893762FA8CF896.xml +++ b/data/CE/22/97/CE229727FF9BFF96FE893762FA8CF896.xml @@ -1,53 +1,56 @@ - - - -Magnana, a new genus of Gyponini from Southeastern Brazil (Hemiptera: Cicadellidae: Iassinae) + + + +Magnana, a new genus of Gyponini from Southeastern Brazil (Hemiptera: Cicadellidae: Iassinae) - - -Author + + +Author -Domahovski, Alexandre Cruz -Universidade Federal do Paraná (UFPR), Setor de Ciências Biológicas, Departamento de Zoologia (DZOO). Curitiba, PR, Brasil. +Domahovski, Alexandre Cruz +Universidade Federal do Paraná (UFPR), Setor de Ciências Biológicas, Departamento de Zoologia (DZOO). Curitiba, PR, Brasil. - - -Author + + +Author -Cavichioli, Rodney Ramiro -Universidade Federal do Paraná (UFPR), Setor de Ciências Biológicas, Departamento de Zoologia (DZOO). Curitiba, PR, Brasil. +Cavichioli, Rodney Ramiro +Universidade Federal do Paraná (UFPR), Setor de Ciências Biológicas, Departamento de Zoologia (DZOO). Curitiba, PR, Brasil. -text - - -Papéis Avulsos de Zoologia +text + + +Papéis Avulsos de Zoologia - -2021 - -Pap. Avulsos Zool., S. Paulo + +2021 + +Pap. Avulsos Zool., S. Paulo - -2021-03-31 + +2021-03-31 - -61 + +61 - -1 -5 + +1 +5 - -http://dx.doi.org/10.11606/1807-0205/2021.61.40 + +http://dx.doi.org/10.11606/1807-0205/2021.61.40 -journal article -10.11606/1807-0205/2021.61.40 -1807-0205 -2A9E2AB5-CD8D-4C5C-BACF-28388282ACA3 +journal article +306322 +10.11606/1807-0205/2021.61.40 +707a46e7-a896-40fa-b2ce-65603cb807a8 +1807-0205 +14371050 +2A9E2AB5-CD8D-4C5C-BACF-28388282ACA3 - + @@ -63,7 +66,7 @@ - + Figs. 1-17 @@ -85,27 +88,27 @@ species: Diagnosis: Large-sized and robust ( -Figs. 16, 17 +Figs. 16, 17 ); head, in dorsal view, distinctly narrower than humeral pronotal width, moderately produced anterad, crown surface strongly rugose ( -Figs. 1 +Figs. 1 , -16 +16 ); frons rugose ( -Fig. 2 +Fig. 2 ); crown-face transition distinct, slender but not foliaceous ( -Figs. 3 +Figs. 3 , -17 +17 ); appendix of forewing very narrow ( -Fig. 4 +Fig. 4 ); metatarsomere I with two double rows of setae, setae not cuculate ( -Fig. 5 +Fig. 5 ); pygofer lacking processes ( -Fig. 8 +Fig. 8 ); connective Y-shaped ( -Fig. 11 +Fig. 11 ); aedeagus bearing atrial processes ( -Figs. 14, 15 +Figs. 14, 15 ). @@ -113,52 +116,52 @@ Large-sized and robust ( Description: Large-sized and robust. Head, dorsal view ( -Figs. 1 +Figs. 1 , -16 +16 ): distinctly narrower than pronotum, transocular width seven-tenths of pronotal humeral width, moderately produced anterad, median length longer than half interocular width, anterior margin of crown subtriangular, surface strongly rugose, slightly concave near anterior margin;ocellus small-sized,approximately equidistant between median line and eye margin, and between anterior and posterior margins of crown; coronal suture indistinct. Head, ventral view ( -Fig. 2 +Fig. 2 ): face slightly wider than long; frontogenal sutures distant from eye margins by slightly less than maximum width of clypeus, surpassing antennal ledges, extended to anterior margin of crown; antennal ledge carinate, obliquely descending and slightly extended over frons; frons 1.6× longer than wide, texture rugose; epistomal suture indistinct; gena with deep striae, lateral margin excavated just below eyes, ventrolateral margin straight; maxillary plates reaching apex of clypeus; lora striate and rugose; clypeus 1.3× longer than its maximum width, lateral margins parallel, apex straight and carinated. Head, lateral view ( -Figs. 3 +Figs. 3 , -17 +17 ): crown-face transition distinct, slender but not foliaceous, irregular striae laterally, middle portion smooth; frons and clypeus not inflated. Pronotum, dorsal view ( -Fig. 1 +Fig. 1 ): deep transverse striae on disc and posterior half, anterior portion rugose; lateral margins long, 1.5× longer than eye length, carinate and convergent anterad; anterior margin broadly round; posterior margin excavated; lateral view ( -Fig. 3 +Fig. 3 ): moderately declivous, head and pronotum in continuous slope; proepimeron with transverse striae. Mesonotum, dorsal view ( -Fig. 1 +Fig. 1 ): longer than wide, rugose, lateral angles shagreen; scutellum flat in lateral view ( -Fig. 3 +Fig. 3 ). Forewing opaque, wide, 2.7× longer than maximum width ( -Fig. 4 +Fig. 4 ); venation distinct, extra cross-veins or veinlets absent; texture finely rugose; three anteapical cells and five apical, vein R1 present; appendix very narrow. Profemur: AD, AM and PD rows reduced and poorly defined, except for apical setae AD₁, AM₁ and PD₁, respectively; AV and PV including 3-4 and 3 setae respectively; IC row formed by slightly arched comb of fine setae,from distal third of femur to AM₁. Protibia: dorsal surface concave, round longitudinal carina adjacent to PD row; AV row formed by short setae, slightly longer and thicker toward apex, apical setae shorter than maximum diameter of protibia; AD and PD rows formed by undifferentiated setae, bearing few slightly longer and thicker intercalary setae; PV row with 7 longer setae intercalated by undifferentiated setae. Hind leg femoral setal formula 2:2:1. Metatibia: AD row lacking intercalary setae between macrosetae; PD, AD and AV rows bearing 20, 12 and 16 macrosetae, respectively. Metatarsomere I ( -Fig. 5 +Fig. 5 ): two double rows of setae, base not cucullate; apex with 6 platellae. Metatarsomere II ( -Fig. 5 +Fig. 5 ): apex with 4 platellae. Male terminalia: Pygofer ( -Fig. 8 +Fig. 8 ) lacking processes. Connective Y-shaped ( -Fig. 11 +Fig. 11 ). Aedeagus bearing atrial processes ( -Figs. 14, 15 +Figs. 14, 15 ).