From c45d83c7b8504a71459fc406503b8d11ca190dec Mon Sep 17 00:00:00 2001 From: ggserver Date: Fri, 22 Nov 2024 19:13:34 +0000 Subject: [PATCH] Add updates up until 2024-11-22 19:08:24 --- .../87/03F087FEFFC7FF9DFC9CC6342D58FAFA.xml | 268 ++++++++-- .../87/03F087FEFFC9FF9DFF30C6742AF1FB5A.xml | 131 +++-- .../87/03F087FEFFC9FF9EFC42C5152D53F9F9.xml | 499 ++++++++++++++++++ .../87/03F087FEFFCAFF98FF5EC6D72C6CF9F8.xml | 150 ++++-- .../87/03F087FEFFCFFF84FC74C6752EF7FAB9.xml | 167 ++++-- .../87/03F087FEFFD0FF84FF03C7172E96F85F.xml | 113 ++++ .../87/03F087FEFFD0FF84FF54C5F52EC0F91F.xml | 102 ++++ .../87/03F087FEFFD0FF87FC91C0F12C6FFB7A.xml | 171 ++++++ .../87/03F087FEFFD3FF86FBC2C7342D64FA7A.xml | 178 +++++++ .../87/03F087FEFFD5FF80FEC1C2D32D46FEC5.xml | 249 +++++++++ 10 files changed, 1824 insertions(+), 204 deletions(-) create mode 100644 data/03/F0/87/03F087FEFFC9FF9EFC42C5152D53F9F9.xml create mode 100644 data/03/F0/87/03F087FEFFD0FF84FF03C7172E96F85F.xml create mode 100644 data/03/F0/87/03F087FEFFD0FF84FF54C5F52EC0F91F.xml create mode 100644 data/03/F0/87/03F087FEFFD0FF87FC91C0F12C6FFB7A.xml create mode 100644 data/03/F0/87/03F087FEFFD3FF86FBC2C7342D64FA7A.xml create mode 100644 data/03/F0/87/03F087FEFFD5FF80FEC1C2D32D46FEC5.xml diff --git a/data/03/F0/87/03F087FEFFC7FF9DFC9CC6342D58FAFA.xml b/data/03/F0/87/03F087FEFFC7FF9DFC9CC6342D58FAFA.xml index 068e213bcba..46ad762d71f 100644 --- a/data/03/F0/87/03F087FEFFC7FF9DFC9CC6342D58FAFA.xml +++ b/data/03/F0/87/03F087FEFFC7FF9DFC9CC6342D58FAFA.xml @@ -1,62 +1,62 @@ - - - -Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain) + + + +Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain) - - -Author + + +Author -Van Den Hoek Ostende, Lars W. -Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands) lars. vandenhoekostende @ naturalis. nl -lars.vandenhoekostende@naturalis.nl +Van Den Hoek Ostende, Lars W. +Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands) +lars.vandenhoekostende@naturalis.nl - - -Author + + +Author -Casanovas-Vilar, Isaac -Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) isaac. casanovas @ icp. cat -isaac.casanovas@icp.cat +Casanovas-Vilar, Isaac +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) +isaac.casanovas@icp.cat - - -Author + + +Author -Furió, Marc -Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain) marc. furio @ uab. cat marc. furio @ icp. cat (corresponding author) -marc.furio@uab.cat +Furió, Marc +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain) +marc.furio@uab.cat -text - - -Comptes Rendus Palevol +text + + +Comptes Rendus Palevol - -2020 - -2020-07-22 + +2020 + +2020-07-22 - -19 + +19 - -1 + +1 - -1 -25 + +1 +25 - -http://dx.doi.org/10.5852/cr-palevol2020v19a1 + +http://dx.doi.org/10.5852/cr-palevol2020v19a1 -journal article -10.5852/cr-palevol2020v19a1 -1777-571X -14203901 -urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3 +journal article +10.5852/cr-palevol2020v19a1 +1777-571X +14203901 +urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3 @@ -72,7 +72,7 @@ ( -Fig. 3 +Fig. 3 A-G) @@ -82,23 +82,79 @@ A-G) AND MEASUREMENTS (in cm). — + Spain -. El Canyet, IPS19361, mandible with m1-m4, partially encaged in sediment; El Palau 3B, +. +El Canyet +, +IPS19361 +, mandible with m1-m4, partially encaged in sediment + +; + +El Palau +3B, 1 m -2 sin., IPS86230, 2.17 × 1.17 × 1.26; -1 m -3 dext., IPS86231, 2.11 × 1.18 × 1.12; Can Martí Vell II, M3 dext., IPS102010, LvK 2.16 × WvK 2.36; LCr 2.12 × WCr 2.75; -1 m -1 sin., IPS86262, 2.00 × 1.00 × 1.12; -1 m -3 dext., IPS86260, 2.16 × 1.18 × 1.14; Can Martí Vell I, 1 M4 dext., IPS86243, 1.10 × 2.30; Les Cases de la Valenciana 1, 1 M3 sin., IPS86622, – × –, 2 fragmented upper molars, IPS86615, IPS86623; Turó de les Forques, 1 M2 dext., IPS86822, – × –; p2? dext., IPS86823, – × 0.75, m3 sin., IPS85484, 2.19 × 1.22 × 1.16; Sant Andreu de la Barca 1, dP3 dext., IPS86180, LvK 1.63 × WvK 1.36; LCr 1.37 × WCr 1.68. +2 sin., +IPS86230 +, 2.17 × 1.17 × 1.26; 1 m3 dext., +IPS86231 +, 2.11 × 1.18 × 1.12 + +; + +Can Martí +Vell II, M3 dext., +IPS102010 +, LvK 2.16 × WvK 2.36; LCr 2.12 × WCr 2.75; 1 m1 sin., +IPS86262 +, 2.00 × 1.00 × 1.12; 1 m3 dext., +IPS86260 +, 2.16 × 1.18 × 1.14 + +; + +Can Martí +Vell I, 1 M4 dext., +IPS86243 +, 1.10 × 2.30 + +; + +Les Cases de la Valenciana +1, 1 M3 sin., +IPS86622 +, – × –, 2 fragmented upper molars, +IPS86615 +, +IPS86623 + +; + +Turó de les Forques +, 1 M2 dext., +IPS86822 +, – × –; p2? dext., +IPS86823 +, – × 0.75, m3 sin., +IPS85484 +, 2.19 × 1.22 × 1.16 + +; + +Sant Andreu de la Barca +1, dP3 dext., +IPS86180 +, LvK 1.63 × WvK 1.36; LCr 1.37 × WCr 1.68 + +. - + -TABLE +TABLE 1. — Localities studied with their corresponding stage and zone and their relative position (when possible). Abbreviations: srm , stratigraphic refinement method used to determine the relative position; @@ -110,6 +166,108 @@ A-G) , biostratigraphy. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
+Localities + +Stage and zone + +Relative position + +Srm +
La Vinya VellaMN4very high in successionB+L
El Palau 3BMN4?–B
El CanyetMN4?–B
Les Escletxes del PapiolAragonian zone Csubzone Cb? younger than VilobíB+L
Sant MametAragonian zone Csubzone Cb? younger than VilobíB+L+M
Vilobí del PenedèsAragonian zone Csubzone Cb? younger than Els CasotsB+L
Els CasotsAragonian zone Csubzone Cb?B+L
Can JuliàAragonian zone C–B+L
Can Martí Vell 1-3Aragonian zone Csubzone Ca?B+L
Les Cases de la ValencianaAragonian zone Csubzone Ca?B+L
El Molí de Can CalopaRamblian zone Ayounger than CostablancaB+L
La Costablanca 1-2Ramblian zone A–B+L
Turó de les ForquesRamblian zone Abelow CostablancaB+L
Sant Andreu de la Barca 1-3Ramblian zone A–B
+ DESCRIPTION diff --git a/data/03/F0/87/03F087FEFFC9FF9DFF30C6742AF1FB5A.xml b/data/03/F0/87/03F087FEFFC9FF9DFF30C6742AF1FB5A.xml index 62bac8c5e26..67e85854a0a 100644 --- a/data/03/F0/87/03F087FEFFC9FF9DFF30C6742AF1FB5A.xml +++ b/data/03/F0/87/03F087FEFFC9FF9DFF30C6742AF1FB5A.xml @@ -1,62 +1,62 @@ - - - -Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain) + + + +Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain) - - -Author + + +Author -Van Den Hoek Ostende, Lars W. -Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands) lars. vandenhoekostende @ naturalis. nl -lars.vandenhoekostende@naturalis.nl +Van Den Hoek Ostende, Lars W. +Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands) +lars.vandenhoekostende@naturalis.nl - - -Author + + +Author -Casanovas-Vilar, Isaac -Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) isaac. casanovas @ icp. cat -isaac.casanovas@icp.cat +Casanovas-Vilar, Isaac +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) +isaac.casanovas@icp.cat - - -Author + + +Author -Furió, Marc -Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain) marc. furio @ uab. cat marc. furio @ icp. cat (corresponding author) -marc.furio@uab.cat +Furió, Marc +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain) +marc.furio@uab.cat -text - - -Comptes Rendus Palevol +text + + +Comptes Rendus Palevol - -2020 - -2020-07-22 + +2020 + +2020-07-22 - -19 + +19 - -1 + +1 - -1 -25 + +1 +25 - -http://dx.doi.org/10.5852/cr-palevol2020v19a1 + +http://dx.doi.org/10.5852/cr-palevol2020v19a1 -journal article -10.5852/cr-palevol2020v19a1 -1777-571X -14203901 -urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3 +journal article +10.5852/cr-palevol2020v19a1 +1777-571X +14203901 +urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3 @@ -72,7 +72,7 @@ Van den Hoek Ostende, 2003 ( -Fig. 3 +Fig. 3 H-J) @@ -82,13 +82,47 @@ H-J) AND MEASUREMENTS (in cm). — + Spain . -Turó de les Forques, 1 M2 dext. damaged, IPS85480, – × 3.06; 1 M3 sin., IPS85486, 1.36 × 2.24; +Turó de les Forques +, 1 M2 dext. damaged, + +IPS +85480 + +, – × 3.06; 1 M3 sin., + +IPS +85486 + +, 1.36 × 2.24; 1 m -2 dext., IPS85482, 2.98 × 2.06; Sant Andreu de la Barca 3, 1 P4 sin. damaged, IPS86202; Sant Andreu de la Barca 1, 1 p1, IPS86194, 1.45 × 0.69. +2 dext., + +IPS +85482 + +, 2.98 × 2.06 + +; + +Sant Andreu de la Barca +3, +1 P +4 sin. damaged, +IPS86202 +; + + +Sant Andreu de la Barca +1, 1 p1, +IPS86194 +, 1.45 × 0.69 + +. @@ -98,8 +132,7 @@ Turó de les Forques, 1 M2 dext. damaged, IPS85480, – × 3.06; 1 M3 sin., IPS8 P4 -The specimen from Sant Andreu de la Barca 3 is heavily damaged, lacking the posterolabial part of the premolar, having the protocone restored and being abraded around the parastyle. -The overall appearance of the tooth is quite sturdy. The conical protocone is about twice the height and size of the hypocone. The two lingual cusps stand well isolated from each other. +The specimen from Sant Andreu de la Barca 3 is heavily damaged, lacking the posterolabial part of the premolar, having the protocone restored and being abraded around the parastyle. The overall appearance of the tooth is quite sturdy. The conical protocone is about twice the height and size of the hypocone. The two lingual cusps stand well isolated from each other. M2 diff --git a/data/03/F0/87/03F087FEFFC9FF9EFC42C5152D53F9F9.xml b/data/03/F0/87/03F087FEFFC9FF9EFC42C5152D53F9F9.xml new file mode 100644 index 00000000000..4c7c86c0453 --- /dev/null +++ b/data/03/F0/87/03F087FEFFC9FF9EFC42C5152D53F9F9.xml @@ -0,0 +1,499 @@ + + + +Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain) + + + +Author + +Van Den Hoek Ostende, Lars W. +Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands) +lars.vandenhoekostende@naturalis.nl + + + +Author + +Casanovas-Vilar, Isaac +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) +isaac.casanovas@icp.cat + + + +Author + +Furió, Marc +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain) +marc.furio@uab.cat + +text + + +Comptes Rendus Palevol + + +2020 + +2020-07-22 + + +19 + + +1 + + +1 +25 + + + + +http://dx.doi.org/10.5852/cr-palevol2020v19a1 + +journal article +10.5852/cr-palevol2020v19a1 +1777-571X +14203901 +urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3 + + + + + + +Galerix symeonidisi +Doukas, 1986 + + + + + + +( +Fig. 3 +K-M) + + + + +MATERIAL +AND +MEASUREMENTS +(in cm). — + + +Spain +. + +El Canyet +, 1 mandible sin. with broken m2 and slightly damaged m3, +IPS20978 +, – × –; 1.67 × 1.19 + +; + +Can Julià +, 1 m3 dext., +IPS86238 +, 1.78 × 1.16; + + +Can Martí Vell +I: 1 P2 sin., +IPS96863 +, 1.67 × 0.86 + +; + +1 mandible dext. with m1 and m2, +IPS96861 +, 2.75 × 1.81; 2.32 × 1.72; 1 labial half of P4 dext., +IPS96862 +, 2.27 × – + +; + +Can Martí Vell +III, 1 M2 dext., +IPS90033 +, 1.75 × 2.29; 2 M2 sin., +IPS90034 +-35, 1.75 × 2.42; 1.79 × 2.48 + +; + +Les Cases de la Valenciana +1, 1 M2 sin., +IPS86824 +, 1.86 × 2.47; 1 M3 sin., +IPS86596 +, 1.11 × –; 1 p1 sin., +IPS86598 +, 1.11 × 0.60; 1 p2 dext., +IPS86597 +, 1.14 × 0.56; 1 m3 sin., +IPS86825 +, 1.90 × 1.21 + +; + +Vilobí de Penedès +, 1 P3 lingual fragment, +IPS86856 +; 2 damaged M1 dext., +IPS86858 +, 86860; 1 p4 sin., +IPS86852 +, 1.91 × 1.12 + +; + +La Vinya Vella +, 1 M1 dext., +IPS106596 +, 2.33 × 3.05 + +; + +Sant Mamet +1, 1 M1 sin., +IPS105162 +, +1.94 × +2.60; 1 p4 dext., +IPS105166 +, 1.70 × 0.97; 1 m2 sin., +IPS105164 +, 2.36 × 1.48; 1 m2 dext., +IPS105165 +, 2.41 × – + +. + + + + +DESCRIPTION + + +P2 + +The outline is elliptical and mesiodistally elongated. The two roots of this tooth are divergent and the main cusp is a bit anterior to the centre of the premolar. + +P4 + +Only the labial side is preserved in the single available specimen. The parastyle is somewhat protruding. The preserved part of the posterior margin is covered by a thin but well-defined cingulum. + +M1 + +The two available specimens from VdP are damaged. Only the specimen from La Vinya Vella is completely preserved. The anterior width is quite similar to the lingual length. The posterolabial corner is elongated. The protocone-hypocone connection is stronger than the protocone-metaconule connection. The latter is even missing on one of the specimens. The posterior arm of the metaconule is short and it does not reach the posterior cingulum in any specimen. +M2 + +The protocone-hypocone connection is strong. Two out of three M2 from Can Martí Vell III lack the protocone-metaconule connection. The posterior cingulum is continuous in the +two specimens +in which this character can be ascertained. + + +M3 + +Even though the anterolabial part of the only available specimen is damaged, the last upper molar is triangular in occlusal view, in spite of the broken anterolabial part. The central basin is closed, except for a small notch near the paracone, thus partially interrupting the anterior crest. + +Mandible + +The only fragment from El Canyet is very damaged. It is a posterior fragment of the horizontal ramus with no significant characters other than a rather high mandibular corpus. The position of the mental foramen is unknown. + +p1 + +The outline of the occlusal surface is slightly ovoid. The tip of the unicuspid lies in the anterior part of the tooth; there is a small flattening behind it. The p1 is one-rooted. + +p2 + +In contrast to the p1, the p2 is double rooted. The outline is elliptical and the tip more centrally placed. There is no posterior flattening. + +p4 +The premolar consists of a trigonid with a very short posterior flattening. The metaconid is well developed. In our hardly worn specimen, the paraconid is formed like a sharp ridge. The paralophid is interrupted by a clear notch between the paraconid and the protoconid. The posterior flattening is bordered by a low posterior ridge. + + +m1 and m2 + +These elements show the invariable aspect typical of the genus, being the m1 larger than the m2 and having a more pointed paraconid, which is rounded in m2. Other than that, the only remarkable trait on these elements is the posterior cingulid, which rises obliquely the distal margin of the tooth, but it does not reach the postcristid. + +m3 + + +Specimens have been found in Les Cases de la Valenciana 1, El Canyet and Can Julià. The element from El Canyet is not complete, thus hampering comparisons. The element from Can Julià only differs from that of Les Cases de la Valenciana described in Jovells-Vaqué +et al. +(2018) in having a slightly smaller talonid. + + + + +REMARKS + + + +Galerix + +is a very common element in the Miocene faunas of the late and middle Miocene of Europe. The genus probably originated in +Anatolia +( + +Van +den Hoek Ostende 1992 + +). The earliest representatives in Europe are known from MN 3 onwards. In Spain, the oldest + +Galerix + +known hitherto is + +G. remmerti +Van den Hoek Ostende, 2003 a + +large-sized species which seems closely related to the central European + +G. aurelianensis +(Van den Hoek Ostende 2003) + +. In addition, +Van +den Hoek Ostende (2003) noted the presence of a very large + +Galerix + +in some Ramblian assemblages, known from a couple of elements only. At the start of the Aragonian, + +G. remmerti + +is replaced by the much smaller-sized + +G. symeonidisi + +in the Daroca-Calamocha basin ( + +Van +den Hoek Ostende & Doukas 2003 + +). A similar situation occurs in central Europe, but here + +G. symeonidisi + +gradually replaces + +G. aurelianensis + +. + +Galerix symeonidisi + +is in its turn replaced by + +G. exilis + +. Initially, this replacement was considered a matter of gradual evolution ( +Ziegler & Fahlbusch 1986 +) but according to the cladistic analysis of + +Borrani +et al. +(2018) + +, + +G. symeonidisi + +and + +G. exilis + +are not closely related species. Based on the large overall variation in some + +Galerix + +assemblage of the Daroca-Calamocha area, + +Van +den Hoek Ostende and Doukas (2003) + +showed that in fact the two species must have occurred together. This discussion clearly demonstrated the difficulties of interpreting + +Galerix + +assemblages if more than one species could be present. In the case of the material from the Vallès-Penedès, identifications are hampered by the scarce material and the absence of characteristic elements (P3, p4) in most assemblages. + + +In the P4 from SAB3, protocone and hypocone are separated, a condition also found in +one specimen +of + +G. remmerti + +from San Roque 4A ( +Van +den Hoek Ostende 2003). The M1 from Sant Mamet 1 and the M2 from Turó de les Forques are peculiar, as their metaconules do not have the crescent-shape that is considered typical for the Galericini ( + +Van +den Hoek Ostende 2001 + +). However, this morphotype is also known from Petersbuch 28 ( + +Klietmann +et al. +2014a + +) and from Zone B localities in the Daroca-Calamocha area (LHO, pers. obs.). The molar somewhat exceeds the largest specimen of + +Galerix remmerti + +from the inland localities, as do the m2 and M3 from the same locality. Also the p1 from Sant Andreu de la Barca 1 is remarkably large. The molars do fall in the size range of + +G. aurelianensis +Ziegler, 1990 + +, a central European species that mainly differs from + +G. remmerti + +in size and presumably was part of the same migration event at the beginning of the Ramblian. Actually, the distinction between + +G. aurelianensis + +and + +G. remmerti + +is not always easy without a large sample. For instance, + +Klietmann +et al. +(2014a) + +suggested that the form present in the French localities of Beaulieu and Bouzigues ( + +Aguilar +et al. +2003 + +; + +Sigé +et al. +1997 + +) was likely + +G. remmerti + +. Would it be the case, the presence of + +G. aurelianensis + +in the Vallès-Penedès basin would create a biogeographic problem of difficult interpretation. Even the possibility that these two forms were conspecific and the differences observed are only a matter of latitudinal variation cannot be discarded. In the absence of further elements to judge, a conservative taxonomic solution is herein preferred, noting that these questions must deserve attention in future works. + + + +FIG +. 3. — Marsupial and galericines from the early Miocene of the Vallès-Penedès: + +Amphiperatherium frequens +von Meyer, 1846 + +( +A -G +). +A +, IPS86180. dP3 dext. (rev) SAB1. +B +, IPS102010. M3 dext. (rev) CMV-II. +C +, IPS86243. M4 dext. (rev). CMV-I. +D +, IPS86823. p2? dext. (rev). TFR. +E +, IPS86262. m1 sin. CMV-II. (E1-ocl; E2-lab). +F +, IPS86230. m2 sin. Palau 3B. (F1-ocl; F2-lab). +G +, IPS86260. m3 dext. CMV-II. + +Galerix remmerti +Van den Hoek Ostende, 2003 + +( +H -J +). +H +, IPS85480. M2 dext. (rev). TFR. +I +, IPS85486. M3 sin. TFR. +J +, IPS85482. m2 dext. (rev). TFR. (J1-ocl; J2-lab). + +Galerix symeonidisi +Doukas, 1986 + +( +K -M +). +K +, IPS106596. M1 dext. (rev). LVV. +L +, IPS96861. m1+m2 dext. (rev). CMV-I. (L1-ocl; L2-lab). +M +, IPS86238. m3 dext. (rev). CJ. (M1-ocl; M2-lab). Scale bar: 1 mm. + + + +The younger material of + +Galerix + +from the MN 4 localities in our study clearly belongs to a smaller species.Jovells-Vaqué +et al. +(2018) already identified the galericine from Cases de la Valenciana as + +Galerix +cf. +symeonidisi + +based on the size and the absence of a protocone-metaconule connections in two of the three upper molars. Their identification is now confirmed by a lingual fragment of a P3 from Vilobí del Penedès which clearly shows a hypocone, one of the main characteristics of + +G. symeonidisi + +. Of the two damaged M1 from this locality, one misses the protocone-metaconule connection. The complete M1 from La Vinya Vella (the only insectivore fossil found in this site) is tentatively ascribed to the same species due to the strong similarities with the material from Vilobí del Penedès. However, because of its somewhat larger size it could even belong to + +G. exilis + +, which is known to co-occur and ultimately replace + +G. symeonidisi + +in the Daroca-Calamocha area in Zone C ( + +Van +den Hoek Ostende & Doukas 2003 + +). This would be in line with La Vinya Vella being one of the youngest localities in this study. The material from LVV is, however, too limited and we thus refrain from ascribing it to a different species. + + +In short, the localities of the Vallès-Penedès seem to fit the pattern observed in the inland basins, in which the large-sized + +G. remmerti + +is replaced by + +G. symeonidisi + +around the Ramblian-Aragonian boundary. Moreover, the large size of the larger species fits with that of the central European + +G. aurelianensis + +, suggesting a near continuous distribution between the northern species and + +G. remmerti + +. As yet, the material is limited and some caution must be observed in these conclusions, but they fit in nicely with the current ideas about early Miocene galericine distribution patterns. + + + + \ No newline at end of file diff --git a/data/03/F0/87/03F087FEFFCAFF98FF5EC6D72C6CF9F8.xml b/data/03/F0/87/03F087FEFFCAFF98FF5EC6D72C6CF9F8.xml index c3ad8baad03..e4191840acc 100644 --- a/data/03/F0/87/03F087FEFFCAFF98FF5EC6D72C6CF9F8.xml +++ b/data/03/F0/87/03F087FEFFCAFF98FF5EC6D72C6CF9F8.xml @@ -1,62 +1,62 @@ - - - -Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain) + + + +Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain) - - -Author + + +Author -Van Den Hoek Ostende, Lars W. -Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands) lars. vandenhoekostende @ naturalis. nl -lars.vandenhoekostende@naturalis.nl +Van Den Hoek Ostende, Lars W. +Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands) +lars.vandenhoekostende@naturalis.nl - - -Author + + +Author -Casanovas-Vilar, Isaac -Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) isaac. casanovas @ icp. cat -isaac.casanovas@icp.cat +Casanovas-Vilar, Isaac +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) +isaac.casanovas@icp.cat - - -Author + + +Author -Furió, Marc -Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain) marc. furio @ uab. cat marc. furio @ icp. cat (corresponding author) -marc.furio@uab.cat +Furió, Marc +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain) +marc.furio@uab.cat -text - - -Comptes Rendus Palevol +text + + +Comptes Rendus Palevol - -2020 - -2020-07-22 + +2020 + +2020-07-22 - -19 + +19 - -1 + +1 - -1 -25 + +1 +25 - -http://dx.doi.org/10.5852/cr-palevol2020v19a1 + +http://dx.doi.org/10.5852/cr-palevol2020v19a1 -journal article -10.5852/cr-palevol2020v19a1 -1777-571X -14203901 -urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3 +journal article +10.5852/cr-palevol2020v19a1 +1777-571X +14203901 +urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3 @@ -72,7 +72,7 @@ ( -Fig. 4 +Fig. 4 A-F) @@ -82,15 +82,64 @@ A-F) AND MEASUREMENTS (in cm). — + Spain . -El Palau 3B, +El Palau +3B, 1 m -2 dext., IPS86232, – × –; Can Martí Vell III, +2 dext., +IPS86232 +, – × –; + + +Can Martí Vell +III, 1 m -3 sin., IPS86278, 1.24 × 0.78; Sant Andreu de la Barca 3, 1 M3 sin., IPS86203, 1.12 × 1.67; Sant Andreu de la Barca 1, 1 P? dext., IPS86195, 1.45 × 0.69; 2 P4 dext., IPS86192-93, 1.43 × 1.17; – × –; 2 M1 sin., IPS86183- 84, – × –; 1.62 × 1.50; 1 M2 dext., IPS86181, 1.77 × 2.25; Turó de les Forques, 1 P4 dext., IPS85483, 1.36 × 1.24; 1 damaged M1 dext., IPS85488, – × –; 1 M1 sin., IPS86816, 1.73 × 2.32; 1 M2 sin., IPS86817, 1.74 × 1.97; 1 M2 dext., IPS86818, 1.76 × 2.13; Costablanca, 1 M2 sin., IPS19372, 1.91 × 1.84. +3 sin., +IPS86278 +, 1.24 × 0.78 + +; + +Sant Andreu de la Barca +3, 1 M3 sin., +IPS86203 +, 1.12 × 1.67; Sant Andreu de la Barca 1, 1 P? dext., +IPS86195 +, 1.45 × 0.69; 2 P4 dext., +IPS86192-93 +, 1.43 × 1.17; – × –; 2 M1 sin., +IPS86183 +- 84, – × –; 1.62 × 1.50; 1 M2 dext., +IPS86181 +, 1.77 × 2.25 + +; + +Turó de les Forques +, 1 P4 dext., +IPS85483 +, 1.36 × 1.24; 1 damaged M1 dext., +IPS85488 +, – × –; 1 M1 sin., +IPS86816 +, 1.73 × 2.32; 1 M2 sin., +IPS86817 +, 1.74 × 1.97; 1 M2 dext., +IPS86818 +, 1.76 × 2.13 + +; + +Costablanca +, 1 M2 sin., +IPS19372 +, 1.91 × 1.84 + +. @@ -135,8 +184,8 @@ from Turó de les Forques, a slight thickening in the posterior arm reveals the M3 -It has an incompletely divided mesostyle, with the two cusplets clearly separated in the unworn specimen from Sant Andreu - +It has an incompletely divided mesostyle, with the two cusplets clearly separated in the unworn specimen from Sant Andreu de la Barca 3. The anterior arm of the protocone is only slightly longer than the posterior arm, and bends sharply at its end to form a protruding parastyle. There is a short stretch of cingulum between the parastyle and the anterior flank of the paracone. The hypocone is a small but distinct cusp, well isolated from the metacone. + FIG . 4. — Talpid and dimylids from the early Miocene of the Vallès-Penedès. @@ -185,7 +234,6 @@ sp. , IPS86185. mand. with c-m2 sin.SAB1.(K1-ocl; K2-lab). Scale bar:1 mm. -de la Barca 3. The anterior arm of the protocone is only slightly longer than the posterior arm, and bends sharply at its end to form a protruding parastyle. There is a short stretch of cingulum between the parastyle and the anterior flank of the paracone. The hypocone is a small but distinct cusp, well isolated from the metacone. m2 diff --git a/data/03/F0/87/03F087FEFFCFFF84FC74C6752EF7FAB9.xml b/data/03/F0/87/03F087FEFFCFFF84FC74C6752EF7FAB9.xml index 0a4ddeba55d..ae4ab2e991d 100644 --- a/data/03/F0/87/03F087FEFFCFFF84FC74C6752EF7FAB9.xml +++ b/data/03/F0/87/03F087FEFFCFFF84FC74C6752EF7FAB9.xml @@ -1,62 +1,62 @@ - - - -Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain) + + + +Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain) - - -Author + + +Author -Van Den Hoek Ostende, Lars W. -Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands) lars. vandenhoekostende @ naturalis. nl -lars.vandenhoekostende@naturalis.nl +Van Den Hoek Ostende, Lars W. +Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands) +lars.vandenhoekostende@naturalis.nl - - -Author + + +Author -Casanovas-Vilar, Isaac -Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) isaac. casanovas @ icp. cat -isaac.casanovas@icp.cat +Casanovas-Vilar, Isaac +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) +isaac.casanovas@icp.cat - - -Author + + +Author -Furió, Marc -Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain) marc. furio @ uab. cat marc. furio @ icp. cat (corresponding author) -marc.furio@uab.cat +Furió, Marc +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain) +marc.furio@uab.cat -text - - -Comptes Rendus Palevol +text + + +Comptes Rendus Palevol - -2020 - -2020-07-22 + +2020 + +2020-07-22 - -19 + +19 - -1 + +1 - -1 -25 + +1 +25 - -http://dx.doi.org/10.5852/cr-palevol2020v19a1 + +http://dx.doi.org/10.5852/cr-palevol2020v19a1 -journal article -10.5852/cr-palevol2020v19a1 -1777-571X -14203901 -urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3 +journal article +10.5852/cr-palevol2020v19a1 +1777-571X +14203901 +urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3 @@ -72,7 +72,7 @@ ( -Fig. 5 +Fig. 5 A-G) @@ -82,12 +82,81 @@ A-G) AND MEASUREMENTS (in cm) — + Spain -.Can Julià, 1 M1 sin., IPS21034, 1.65 × –; 1 M2 dext., IPS86235, 1.49 × –; 1 M2 sin., IPS86236, 1.48 × –; Can Martí Vell III, 1 i1 dext. fragmented, IPS90036; Can Martí Vell II, 1 P4 dext., IPS21023, 1.52 × 1.45; 1 M1 sin., IPS86257, 1.60 × 1.86; +. +Can Julià +, 1 M1 sin., +IPS21034 +, 1.65 × –; 1 M2 dext., +IPS86235 +, 1.49 × –; 1 M2 sin., +IPS86236 +, 1.48 × – + +; + +Can Martí Vell +III, 1 i1 dext. fragmented, +IPS90036 + +; + +Can Martí Vell +II, 1 P4 dext., +IPS21023 +, 1.52 × 1.45; 1 M1 sin., +IPS86257 +, 1.60 × 1.86; 1 m -1 dext., IPS86259, 1.95 × 1.29; Can Martí Vell I, 1 Ax sin., IPS86252, 1.10 × 0.89; 1 maxillary with M1 + M2 dext., IPS21025, 1.60 × 1.79, 1.38 × 1.71; 1 maxillary with M2 + M3 sin., IPS86241, 1.45 × 1.82, 0.83 × 1.27; 2 ax dext., IPS86249-50, 1.26 × 1.01; 1.19 × 0.92; 1 ax sin., IPS86251, 1.25 × 1.01; 1 i1 sin., IPS86254,> 4.75; 1 i1 dext (IPS86255,> 5.06), 2 mandibles with m1-m3 sin. (IPS86244-45, 2.08 × 1.30, 1.68 × 1.21, 1.30 × 0.98; 2.15 × 1.18, 1.74 × 1.15, 1.36 × 0.97; 1 mandible with m2, m3 dext., IPS86246, 1.63 × 1.28, 1.29 × 0.96; Cases de la Valenciana 1, 1 M1 dext., IPS86826, 1.67 × 1.93; ax, IPS86600, 1.22 × 1.00; Costablanca 2, 1 maxillary with M1 + M2 sin., IPS85666, 1.64 × 1.84; 1.54 × –; 1 M3 sin., IPS85667, 0.88 × –. +1 dext., +IPS86259 +, 1.95 × 1.29 + +; + +Can Martí Vell +I, 1 Ax sin., +IPS86252 +, 1.10 × 0.89; 1 maxillary with M1 + M2 dext., +IPS21025 +, 1.60 × 1.79, 1.38 × 1.71; 1 maxillary with M2 + M3 sin., +IPS86241 +, 1.45 × 1.82, 0.83 × 1.27; 2 ax dext., +IPS86249 +-50, 1.26 × 1.01; 1.19 × 0.92; 1 ax sin., +IPS86251 +, 1.25 × 1.01; 1 i1 sin., +IPS86254 +,> 4.75; 1 i1 dext ( +IPS86255 +,> 5.06), 2 mandibles with m1-m3 sin. ( +IPS86244 +-45, 2.08 × 1.30, 1.68 × 1.21, 1.30 × 0.98; 2.15 × 1.18, 1.74 × 1.15, 1.36 × 0.97; 1 mandible with m2, m3 dext., +IPS86246 +, 1.63 × 1.28, 1.29 × 0.96 + +; + +Cases de la Valenciana +1, 1 M1 dext., +IPS86826 +, 1.67 × 1.93; ax, +IPS86600 +, 1.22 × 1.00 + +; + +Costablanca +2, 1 maxillary with M1 + M2 sin., +IPS85666 +, 1.64 × 1.84; 1.54 × –; 1 M3 sin., +IPS85667 +, 0.88 × – + +. @@ -135,9 +204,9 @@ A-G) The trigonid is much longer and wider than the talonid. Contrary to m1 and m2, the metaconid does not bear a metacristid. Nor is there a well-developed entocristid. The talonid is quite reduced, although it still preserves an inner basin. In the talonid the hypoconid is the only discernible cuspid as such. The basal cingulid covers the labial side of the tooth from the paraconid to the base of the hypoconid. - + -FIG +FIG . 5. — Heterosoricid and soricids from the early Miocene of the Vallès-Penedès: Heterosorex neumayrianus diff --git a/data/03/F0/87/03F087FEFFD0FF84FF03C7172E96F85F.xml b/data/03/F0/87/03F087FEFFD0FF84FF03C7172E96F85F.xml new file mode 100644 index 00000000000..4037b26e8b0 --- /dev/null +++ b/data/03/F0/87/03F087FEFFD0FF84FF03C7172E96F85F.xml @@ -0,0 +1,113 @@ + + + +Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain) + + + +Author + +Van Den Hoek Ostende, Lars W. +Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands) +lars.vandenhoekostende@naturalis.nl + + + +Author + +Casanovas-Vilar, Isaac +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) +isaac.casanovas@icp.cat + + + +Author + +Furió, Marc +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain) +marc.furio@uab.cat + +text + + +Comptes Rendus Palevol + + +2020 + +2020-07-22 + + +19 + + +1 + + +1 +25 + + + + +http://dx.doi.org/10.5852/cr-palevol2020v19a1 + +journal article +10.5852/cr-palevol2020v19a1 +1777-571X +14203901 +urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3 + + + + + +cf. + +Oligosorex thauensis +( +Crochet, 1975 +) + + + + + + +( +Fig. 5I +) + + + + +MATERIAL +AND +MEASUREMENTS +(in cm). — + + +Spain +. + +Can Martí Vell +III, +1 m +1 sin., +IPS90097 +, 1.11 × 0.67 × 0.69 + +; + +Vilobí de Penedès +, +1 m +2 sin., +IPS86866 +, 1.15 × 0.66 × 0.68 + +. + + + + \ No newline at end of file diff --git a/data/03/F0/87/03F087FEFFD0FF84FF54C5F52EC0F91F.xml b/data/03/F0/87/03F087FEFFD0FF84FF54C5F52EC0F91F.xml new file mode 100644 index 00000000000..2d2db89b000 --- /dev/null +++ b/data/03/F0/87/03F087FEFFD0FF84FF54C5F52EC0F91F.xml @@ -0,0 +1,102 @@ + + + +Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain) + + + +Author + +Van Den Hoek Ostende, Lars W. +Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands) +lars.vandenhoekostende@naturalis.nl + + + +Author + +Casanovas-Vilar, Isaac +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) +isaac.casanovas@icp.cat + + + +Author + +Furió, Marc +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain) +marc.furio@uab.cat + +text + + +Comptes Rendus Palevol + + +2020 + +2020-07-22 + + +19 + + +1 + + +1 +25 + + + + +http://dx.doi.org/10.5852/cr-palevol2020v19a1 + +journal article +10.5852/cr-palevol2020v19a1 +1777-571X +14203901 +urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3 + + + + + +cf. + +Clapasorex alvarezae +Van den Hoek Ostende, 2003 + + + + + + +( +Fig. 5H +) + + + + +MATERIAL +AND +MEASUREMENTS +(in cm). — + + +Spain +. + +Turó de les Forques +, 1 m1 dext. in ramus, +IPS86819 +, 1.17 × 0.69 × 0.70; 1 m1 dext., +IPS86820 +, 1.20 × 0.67 × 0.73 + +. + + + + \ No newline at end of file diff --git a/data/03/F0/87/03F087FEFFD0FF87FC91C0F12C6FFB7A.xml b/data/03/F0/87/03F087FEFFD0FF87FC91C0F12C6FFB7A.xml new file mode 100644 index 00000000000..c600bc0b3b0 --- /dev/null +++ b/data/03/F0/87/03F087FEFFD0FF87FC91C0F12C6FFB7A.xml @@ -0,0 +1,171 @@ + + + +Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain) + + + +Author + +Van Den Hoek Ostende, Lars W. +Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands) +lars.vandenhoekostende@naturalis.nl + + + +Author + +Casanovas-Vilar, Isaac +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) +isaac.casanovas@icp.cat + + + +Author + +Furió, Marc +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain) +marc.furio@uab.cat + +text + + +Comptes Rendus Palevol + + +2020 + +2020-07-22 + + +19 + + +1 + + +1 +25 + + + + +http://dx.doi.org/10.5852/cr-palevol2020v19a1 + +journal article +10.5852/cr-palevol2020v19a1 +1777-571X +14203901 +urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3 + + + + + +cf. + +Clapasorex alvarezae + +vel cf. + +Oligosorex thauensis + + + + + + +MATERIAL +EXAMINED +. — + + +Spain +. + +Sant Andreu de la Barca +1, 1 m2 sin. trigonid in ramus, +IPS86187 + +. + + + + +DESCRIPTION + + +m1 + +In the m1 from Turó de les Forques, the trigonid and talonid are of similar length; the trigonid is only slightly narrower.The protoconid is rather massive; the metaconid lies close to the protoconid. The oblique cristid ends just lingually to the base of the protoconid. The hypolophid curves at just its lingual end, behind the entoconid, with little or no space for a post-entoconid valley. The entocristid slopes down before levelling, ending against the posterior flank of the metaconid. There is a well-developed cingulid on the anterior and posterior sides; the cingulum on the labial and lingual sides is narrow but continuous. The m1 from Can Martí Vell III has more slender cusps, the protoconid and metaconid further apart and a more open trigonid basin. Between the hypolophid and the entoconid, there is a narrow post-entoconid valley. + +m2 + +The trigonid is just slightly narrower than the talonid and of similar length. The oblique cristid ends just a bit more lingually to the centre of the posterior wall of the trigonid.The hypolophid is relatively straight, with just a slight curve at the lingual end, rounding the entoconid and connecting to the lingual cingulum. The entocristid extends to the base of the metaconid.The cingulum is best developed on the anterior side, somewhat less on the labial and posterior sides and weak on the lingual side. + + + +REMARKS + + +Van +den Hoek Ostende (2003) commented on the difficulties in the taxonomy of the small shrews in the early Miocene and how these regularly led in literature to possible lumping of multiple species. In the material described in that paper, two of these small soricids were present as also testified by mandibles showing the alveoles of the antemolars. One of the species from the Ramblian +type +section was identified as + +Clapasorex alvarezae + +, the other as + +Oligosorex thauensis + +. Based on a change in the relative size of the m1 and m2, the younger assemblages of the latter were classified as + +O. +aff. +thauensis + +. Although the molar morphology of these two species, as indeed of most of the soricids of that period, is very close, +Van +den Hoek Ostende (2003) ventured to indicate some characters used to identify isolated molars. In the case of the lower molars, the cusps of + +Clapasorex + +had a more massive appearance, protoconid and metaconid of the m1 were situated close together and the hypolophid ran directly behind the entoconid without having a post-entoconid valley. + + +In the series from the early Miocene insectivore assemblages of the Vallès-Penedès, only a few isolated molars of small soricids have been found. Without the foreknowledge on the shrews from the Ramblian +type +section, these would almost certainly have been combined in one taxon.However, following the above-mentioned characteristics, both + +Clapasorex alvarezae + +and + +Oligosorex +aff. +thauensis + +appear to be represented in the Vallès-Penedès. Given the difficulties in identifying these isolated molars, we do take some caution in assigning the material to one or the other taxon, hence presenting them as cf. + +Clapasorex alvarezae + +and cf. + +Oligosorex thauensis + +, respectively. Despite these reservations, it is notable that cf. + +Clapasorex alvarezae + +was identified from the oldest assemblage in our series, Turó de les Forques. In the Ramblian +type +area, + +C. alvarezae + +has only been identified from older localities (zone Z). + + + + \ No newline at end of file diff --git a/data/03/F0/87/03F087FEFFD3FF86FBC2C7342D64FA7A.xml b/data/03/F0/87/03F087FEFFD3FF86FBC2C7342D64FA7A.xml new file mode 100644 index 00000000000..daa69d5399c --- /dev/null +++ b/data/03/F0/87/03F087FEFFD3FF86FBC2C7342D64FA7A.xml @@ -0,0 +1,178 @@ + + + +Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain) + + + +Author + +Van Den Hoek Ostende, Lars W. +Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands) +lars.vandenhoekostende@naturalis.nl + + + +Author + +Casanovas-Vilar, Isaac +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) +isaac.casanovas@icp.cat + + + +Author + +Furió, Marc +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain) +marc.furio@uab.cat + +text + + +Comptes Rendus Palevol + + +2020 + +2020-07-22 + + +19 + + +1 + + +1 +25 + + + + +http://dx.doi.org/10.5852/cr-palevol2020v19a1 + +journal article +10.5852/cr-palevol2020v19a1 +1777-571X +14203901 +urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3 + + + + + +cf. + +Florinia + + + + + + +( +Fig. 5 +N-O) + + + + +MATERIAL AND MEASUREMENTS (in cm). — + + +Spain +. + +Escletxes del Papiol +, 1 P4 sin., +IPS86423 +, 1.26 × 1.09 + +; + +Can Martí Vell +III, +1 m +1 dext., +IPS90052 +, 1.24 × 0.61 × 0.68 + +; + +Sant Andreu de la Barca +1, 1 mandible with m1, m2 dext., +IPS86186 +, 1.33 × 0.69 × 0.72, 1.21 × 0.73 × 0.73 + +; + +Costablanca +2, 1 P4 dext., +IPS85665 +, 1.18 × 1.22 + +. + + + + +DESCRIPTION + + +P4 + +The premolar is about as long as it is wide, and has a slight posterior emargination. The paracone looks rather gracile, with a relatively short posterocrista; its tip lies just in front of the middle of the molar. The parastylar region is damaged in both available specimens, but, based on the P4 from Escletxes del Papiol, parastyle and protocone appear to be of the same size and height. The two cusps are connected by a low ridge. A low ridge following the lingual outline of the premolar starts at the basis of the protocone. At the position of the hypocone, this ridge is somewhat thickened. + +m1 + +The outline of the occlusal surface is subtriangular. The trigonid is clearly narrower and much longer than the talonid. The paralophid is long and the metaconid is slightly lingualposteriorly to the protoconid; the trigonid basin is widely open. The oblique cristid ends just lingually to the base of the protoconid. The entoconid is close to the metaconid, with a short entocristid. The hypolophid bends behind the entoconid and it is separated from that cusp by a wide postentoconid valley. The cingulum is well developed on the anterior and posterior sides, more narrow on the labial side. + +m2 + +The m2 is far more rectangular than the m1. The trigonid is longer than the talonid; the trigonid basin is wide. In the other features it resembles the first lower molar. + + + +REMARKS + + +The genus + +Florinia + +, including the species + +F.stehlini + +, is a regular occurrence in MN 3-MN 5 faunas in central Europe ( +Ziegler 1989 +, +2006b +; +Ziegler & Mörs 2000 +; + +Klietmann +et al. +2014b + +). Here, the genus is for the first time described from +Spain +, although + +Van der Meulen +et al. +(2012) + +did list “ + +Florinia + +” (their quotation marks) for a number of localities in the Aragonian +type +area, ranging from the uppermost Ramblian (San Roque 4A, Zone A, latest MN 3) to the middle Aragonian (Las Umbrias 10, Zone Dd, MN 5). + + + + \ No newline at end of file diff --git a/data/03/F0/87/03F087FEFFD5FF80FEC1C2D32D46FEC5.xml b/data/03/F0/87/03F087FEFFD5FF80FEC1C2D32D46FEC5.xml new file mode 100644 index 00000000000..98a4c6feb60 --- /dev/null +++ b/data/03/F0/87/03F087FEFFD5FF80FEC1C2D32D46FEC5.xml @@ -0,0 +1,249 @@ + + + +Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain) + + + +Author + +Van Den Hoek Ostende, Lars W. +Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands) +lars.vandenhoekostende@naturalis.nl + + + +Author + +Casanovas-Vilar, Isaac +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) +isaac.casanovas@icp.cat + + + +Author + +Furió, Marc +Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain) +marc.furio@uab.cat + +text + + +Comptes Rendus Palevol + + +2020 + +2020-07-22 + + +19 + + +1 + + +1 +25 + + + + +http://dx.doi.org/10.5852/cr-palevol2020v19a1 + +journal article +10.5852/cr-palevol2020v19a1 +1777-571X +14203901 +urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3 + + + + + + +SORICINAE + +gen. et sp. indet. + + + + + +( +Fig. 5 +S-U) + + + + +MATERIAL +AND +MEASUREMENTS +(in cm). — + + +Spain +. + +El Canyet +, 1 I1 sin., +IPS85553 +, 1.81 × 075 × 1.30; 2 i1 dext., +IPS85552 +, –, +IPS85555 +, 3.20; 2 i1 sin., +IPS85554 +, 3.60, +IPS85556 +, –,> 2.94; 1 mandible with i1 and p4 sin., +IPS85557 +, i1 3.35, p4 1.27 × 0.90); 1 mandible with i1 and m1 dext., +IPS20979 +, i1 3.18, m1 1.55 × 0.88 × 0.98; 1 p4 sin., +IPS85550 +, 1.20 × 0.82; 1 mandible with a1- m1 sin., +IPS85551 +, 0.81 × 0.65, 1.10 × 0.79, 1.54 × 0.86 × 0.98 + +. + + + + +DESCRIPTION + + +I1 + +The upper incisor is not fissident, but there is a medial cuspule visible in dorsal view. The dorsal margin is regularly curved. The talon is not much developed. There is a narrow cingulum covering the base of the tooth. The root is only somewhat shorter than the crown. + +Mandible + +The mandible shows that the mental foramen is placed below the reentrant valley of the m1. It is also noteworthy that the i1 extends back labially as much as below the paraconid of the m1. + +i1 + +The lower incisor is bicuspulate. The anterior tip and the posterior cuspule have a similar height, whereas the intermediate one is a bit lower. + +a1 + +The antemolar has a heart-shaped occlusal outline. It is smaller in size than its posterior counterpart, the p4. There is no posterolingual basin, but it has a tetrahedral morphology. + +p4 + +The last antemolar has a heart-shaped occlusal outline and a ‘paralophid-like’ cristid, which defines a lingual valley. + +m1 + +The most significant characters are the presence of a high entoconid and its corresponding cristid, the oblique cristid ending much labially and a continuous cingulid covering all the base of the tooth, also by the lingual side. The rest of the characters (or their ‘absence’) are detailed in the discussion about the taxonomic ascription. + + + +REMARKS + + +Surprisingly, this species does not show the typical characters of the +Crocidosoricinae +, which were the dominant shrews during the early Miocene. Instead, the elements recovered have typical derived +Soricinae +traits. This is something unusual in localities of this age, as red-toothed shrews become frequent in Eurasia at the end of the Miocene. Some anterior teeth are isolated elements which have been tentatively ascribed to this form because they all come from the same locality, and their general aspect fits well with those of the limited dental series recovered. The +two specimens +with m1 show that this element is larger than in the rest. + + +The most significant derived features are the presence of only two lower antemolars (including the p4), a mental foramen placed under the reentrant valley of m1 and a p4 with a posterolingual basin (instead of a two-branched protoconid). If not a soricine shrew in itself, this form could be also ascribed to one of those species purportedly leading to the +Soricinae +clade (according to + +Furió +et al. +2007 + +), like + +Carposorex sylviae + +. Actually, the dimensions of the m1 match perfectly with that species, according to the data provided by +Crochet (1975) +. Moreover, the teeth have a massive aspect, the p4 has a labial branch more developed than the lingual one, and the m1 has a developed entoconid. However, the position of the mental foramen (under the reentrant valley of the m1; more advanced in crocidosoricines, also in + +Carposorex + +, in which it is placed under the p4), the absence of any connection between posterolabial crest of the protoconid and the labial cingulid, and the smooth enamel (instead of wrinkled as expected for the genus) discards such generic and specific allocation. In the other known species of the genus, + +Carposorex burkarti + +described by + +Hugueney +et al. +(2012) + +, the differences are even more, because of the higher number of lower antemolars (at least three in + +C. sylviae + +) and the smaller size. + + +The oldest representative of the subfamily +Soricinae +in Europe is + +Hemisorex robustus + +. This species has been described in the French sites of Vieux-Collonges, Sansan and La Grive L3, L7 and M ( +Baudelot 1967 +; +Engesser 1972 +, +2009 +; + +Hugueney +et al. +2012 + +). In fact, the measurements of the m +1 in +this form also fit quite well within the ranges provided by +Engesser (2009) +and + +Hugueney +et al. +(2012) + +, with a length of about +1.50 mm +and a width a bit less than 1.00 mm. The massive aspect, the bicuspulate i1, the p4 with a posterolingual basin and the position of the mental foramen would also support such ascription. However, the occlusal outline of the m1 (not as much rectangular as in + +Hemisorex + +) and the entoconid cristids (more developed than in + +Hemisorex + +), make a difference. + + + +Paenesorex bicuspis + +from Petersbuch 31 described byZiegler (2003) could be another possible candidate.However, it differs from the species found in El Canyet in the presence of a vestigial a2 between a1 and p4. Moreover, the overall size of teeth in + +Paenesorex + +is smaller than that of this undetermined soricid. + + +As there is hitherto no soricid species described in the early or middle Miocene matching all these characters, and in the absence of sufficient material we refrain from describing a new species, the most sensible option is leaving the identification of this form as +Soricinae +indet. + + + + \ No newline at end of file