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<document id="957293D7C052363BDF1F4EDAAB15052E" ID-DOI="10.3897/zookeys.1215.129410" ID-ZooBank="7473119D-1697-4767-AE4D-691F5778C6EE" ID-publisher-id="129410" URI-arpha="57CE2FBC-0BA6-51E1-B1E4-09356F6E249D" URI-zoobank="http://zoobank.org/7473119D-1697-4767-AE4D-691F5778C6EE" article-type="research-article" checkinTime="1729177021934" checkinUser="pensoft" docAuthor="Doménech, Carles" docDate="2024" docId="C7A1D79B93A55F6CAEEF71D0D627DA9F" docLanguage="en" docName="ZooKeys 1215: 311-334" docOrigin="ZooKeys 1215" docSource="https://zookeys.pensoft.net/article/129410/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Scolopendra spinosissima sensu Kraepelin 1903" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="334" masterDocId="57CE2FBC0BA651E1B1E409356F6E249D" masterDocTitle="Type designation and redescription of Scolopendra spinosissima Kraepelin, 1903 (Scolopendromorpha, Scolopendridae), with remarks on related taxa" masterLastPageNumber="334" masterPageNumber="311" pageNumber="311" updateTime="1729177395330" updateUser="ExternalLinkService">
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<mods:title id="E94ABBBBB255F592BF42BFEF44D85B3F">Type designation and redescription of Scolopendra spinosissima Kraepelin, 1903 (Scolopendromorpha, Scolopendridae), with remarks on related taxa</mods:title>
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<mods:namePart id="A88C413F660A6FB5E034F0B9D2B519B1">Doménech, Carles</mods:namePart>
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<mods:affiliation id="30187DB81967EB39C3952A4E810FF9B1">Departament de Ciències Ambientals i Recursos Naturals, Universitat d’Alacant, Carretera de Sant Vicent del Raspeig s / n C. P. 03690, San Vicent del Raspeig, Alacant, Spain</mods:affiliation>
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<mods:date id="4A76351ECAF0127FEB9438A18F606F3D">2024</mods:date>
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<mods:number id="835024C509BCFA8389E5E935225B2496">2024-10-17</mods:number>
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<mods:identifier id="3E501050E21C87DF7CA4938E22FC335C" type="DOI">10.3897/zookeys.1215.129410</mods:identifier>
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<treatment id="C7A1D79B93A55F6CAEEF71D0D627DA9F" ID-DOI="http://doi.org/10.5281/zenodo.13946716" ID-Zenodo-Dep="13946716" ID-arpha="C7A1D79B-93A5-5F6C-AEEF-71D0D627DA9F" LSID="urn:lsid:plazi:treatment:C7A1D79B93A55F6CAEEF71D0D627DA9F" httpUri="http://treatment.plazi.org/id/C7A1D79B93A55F6CAEEF71D0D627DA9F">
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<taxonomicName id="7D1373AF4A72CD1156A6914B8FF83297" ID-CoL="4VSC6" authority="Kraepelin, 1903" authorityName="sensu Kraepelin" authorityYear="1903" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="spinosissima">
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<emphasis id="BF1C8AA1A4DD6064E0ACEBDDE181C5F0" italics="true">Scolopendra spinosissima</emphasis>
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<bibRefCitation id="3F16CE0025C42793D01F340B38FE3410" author="Kraepelin K" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pagination="1-276" refId="B17" refString="Kraepelin K (1903) Revision der Scolopendriden. Mitteilungen aus dem Naturhistorischen Museum in Hamburg 20: 1 - 276." title="Revision der Scolopendriden." volume="20" year="1903">Kraepelin, 1903</bibRefCitation>
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<subSubSection id="62CDF6B0F23B1539888A12CDE4239147" type="description">
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<paragraph id="CCE35DDA260A5FBEBAAFCDE5294CFDF5">
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<figureCitation id="DF3BA7DA193FD1FA186DA71B66B5C079" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scolopendra spinosissima Kraepelin, 1903 A, B habitus in vivo showing the only two known variations of colouration; specimens from Panay Is. (A) and Guimaras Is. (B) (photographs courtesy of E. Währen; individuals uncollected) C distribution, 20 th century records: red square = lectotype, Manila, Luzon Is.; red circle = paralectotypes 1–3, Camarines Peninsula, Luzon Is.; yellow square = non-types, Dolores, Quezon province (previously Tayabas), Luzon Is. (in the literature erroneously placed in Dolores, Camarines Peninsula, indicated by a red circle). 21 st century data: orange squares = (top) Mt. Isarog, Camarines Sur province, Luzon Is.; (middle) Cadiz, Negros Occidental province, Negros Is.; (bottom) Barili, Cebu Prov., Cebu Is. New data: blue squares: (top) Idiacacan, Pandan (Antique), Panay Is. (~ 11.683537; 122.114986); (bottom) Jordan, Guimaras Is. (~ 10.652871; 122.602054) (credits: E. Währen). Mindanao Is. indicates the origin of the specimen “ Mus. Paris. I. XI. 03 ” misidentified by Kraepelin as “ S. spinosissima ” (specimen currently excluded from the type series)." figureDoi="10.3897/zookeys.1215.129410.figure1" httpUri="https://binary.pensoft.net/fig/1155662">Figs 1</figureCitation>
|
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,
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||||
<figureCitation id="9241BFE8FEADCB94F76F04C106E55798" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A Scolopendra spinosissima Kraepelin, 1903; lectotype in original container (ZMH - A 0000633). Notice the two labels inside and the red label fixed outside the jar, indicating type status B labels of the original type series proposed by Kraepelin in 1903: lectotype from Manila (two labels on top), paralectotypes from Camarines (bottom left), and specimen “ Mus. Paris. I. XI. 03 ” from Mindanao (bottom right, specimen excluded from the type series) C the manuscript list in which Kraepelin pointed out the type status of these specimens, localities, and identification dates. Notice that these five specimens, making up the type series, are ascribed to the data “ S. 262–263 ”: S. is an abbrev. of “ Seiten ”, meaning pages in German, which is a reference to the page numbers of the 1903 publication containing the original description. Photograph courtesy of N. Dupérré." figureDoi="10.3897/zookeys.1215.129410.figure2" httpUri="https://binary.pensoft.net/fig/1155663">2</figureCitation>
|
||||
,
|
||||
<figureCitation id="C292A28BD05B338D7AA85767A3350060" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A cephalic plate and right antenna, dorsal view B forcipular segment, ventral view C tooth-plates, ventral view D left spiracle on segment 8. Scale bars: 0.5 mm (C, D); 2 mm (A, B)." figureDoi="10.3897/zookeys.1215.129410.figure3" httpUri="https://binary.pensoft.net/fig/1155664">3</figureCitation>
|
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,
|
||||
<figureCitation id="034FD7F574470C75FAED156723D3B1AC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A tergites 3–5 B sternites 10 and 11 C ultimate leg-bearing segment, ventral view D left coxopleural process, lateral view E ultimate leg, right lateral view. Scale bars: 1 mm (A, B, D); 2 mm (C, E)." figureDoi="10.3897/zookeys.1215.129410.figure4" httpUri="https://binary.pensoft.net/fig/1155665">4</figureCitation>
|
||||
,
|
||||
<figureCitation id="87A7DED7820349521692735DB7B0295A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) B, D, F Scolopendra paradoxa Doménech, 2018; holotype (CEUA 017 - Mr 0000) A, B right coxopleura and coxopleural processes, lateral views C, D ultimate leg prefemora, dorsal views E, F ultimate leg prefemora, ventral views. The coxopleuron and coxopleural processes are different shapes and lengths; with the different sizes, morphologies, numbers, and positions of the prefemoral spinous processes, these are the most remarkable characters differentiating these species. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1215.129410.figure5" httpUri="https://binary.pensoft.net/fig/1155666">5 A, C, E</figureCitation>
|
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,
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<geoCoordinate id="4CA1B1D03426487F58765E5BADEF5C9F" degrees="6, 8" direction="east" orientation="longitude" precision="5555" value="6.8">
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<figureCitation id="0873AB40DC3CAE5D1C11E64E10E090AE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopendra spinosissima Kraepelin, 1903; genital apparatus A paralectotype 2, female (ZMH - A 00016059) B paralectotype 3, male (ZMH - A 00016060). Abbreviations: AV = anal valve; LA = lamina adanalis; LS = lamina subanalis; SGS I = sternite of genital segment I; SGS II = sternite of genital segment II. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1215.129410.figure6" httpUri="https://binary.pensoft.net/fig/1155667">6</figureCitation>
|
||||
,
|
||||
<figureCitation id="D6EAB4C0BF26C7C617C8BC14417A45FB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A – D Scolopendra paradoxa Doménech, 2018; non-type (MNHN N ° 387), identified by Kraepelin (1904 b) as S. spinosissima (total length 176 mm) E S. spinosissima; non-type (CEUA 016 - Mr 0009, voucher) A habitus, dorsal view B coxopleuron and ultimate legs prefemur, right lateral view C ultimate leg-bearing segment and ultimate legs prefemur, ventral view D right ultimate leg prefemoral spinous processes (left ultimate leg regenerated), dorsolateral view E ultimate leg-bearing segment and ultimate legs prefemora, ventral view. Notice the smaller extra aberrant median VL spinous process in the left prefemur observed exclusively in this specimen. Scale bars: 1 mm (E); 20 mm (A)." figureDoi="10.3897/zookeys.1215.129410.figure8" httpUri="https://binary.pensoft.net/fig/1155669">8 E</figureCitation>
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,
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<tableCitation id="5F7CC541D4F0DCA7FF385D25547A625F" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparison between the type series of Scolopendra spinosissima Kraepelin, 1903, its voucher, and S. paradoxa Doménech, 2018 holotype. A = Absent; C = Complete; I = Incomplete; D = Distal; P = Proximal; PR = Partially retracted; R = Retracted. * = Appendix damaged. N / A = Not applicable." httpUri="http://table.plazi.org/id/BC7916AA261A079E149D4E809E484642" tableUuid="BC7916AA261A079E149D4E809E484642">Table 1</tableCitation>
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</paragraph>
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<treatmentCitation id="CCEC18517BAF4BBF27AB7131010232DF" author="Kraepelin K" page="263" year="1903">
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<taxonomicName id="95B81CD3C0112BC12FB76E19EBBE5305" ID-CoL="4VSC6" authority="Kraepelin, 1903: 262 - 263" authorityName="Kraepelin" authorityPageNumber="262 - 263" authorityYear="1903" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="variety" species="subspinipes" variety="spinosissima">
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||||
<emphasis id="7075653919A0E11B4D2CF00C8BCEBAEC" italics="true">Scolopendra subspinipes var. spinosissima</emphasis>
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<comment id="5080EA06553F49F039A5A963F2F4A9B9">
|
||||
<bibRefCitation id="8FE1232E8F494B7652AEA2D1000D0F41" author="Kraepelin K" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pagination="1-276" refId="B17" refString="Kraepelin K (1903) Revision der Scolopendriden. Mitteilungen aus dem Naturhistorischen Museum in Hamburg 20: 1 - 276." title="Revision der Scolopendriden." volume="20" year="1903">Kraepelin, 1903: 262–263</bibRefCitation>
|
||||
.
|
||||
</comment>
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</taxonomicName>
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</treatmentCitation>
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</paragraph>
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<paragraph id="8A0938E09A6943C7B940B64533DC9C62">
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<taxonomicName id="CC614ED8883096788D410983BA5252C0" ID-CoL="4VSC6" authorityName="sensu Kraepelin" authorityYear="1903" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="spinosissima">
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<emphasis id="C8AD7F6CAE8D660A9B26B755ECF0594F" italics="true">Scolopendra spinosissima</emphasis>
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</taxonomicName>
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||||
<comment id="4918EF81B6A4BAC644482F986238C96A">
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:
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<treatmentCitation id="5AC0F04C18473F16D4372140AA92793F" author="Attems C" page="31 - 32" year="1930">
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||||
<bibRefCitation id="EABA29B8EDC960F26A035DE047A3B73C" DOI="10.1515/9783112373002" author="Attems C" firstAuthor="Attems" page="31 - 32" refId="B2" refString="Attems C (1930) Myriapoda ”. 2. Scolopendromorpha. Das Tierreich 54: 308. https://doi.org/10.1515/9783112373002" year="1930">Attems 1930: 31–32</bibRefCitation>
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||||
</treatmentCitation>
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||||
, fig. 44 (transcription, illustration; specimen unidentified);
|
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<treatmentCitation id="CF582D31A80AA9D9A90494CC816E7D00" author="Lewis JGE" page="92" year="2010">
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<bibRefCitation id="FF89DABDF60CC2FCF9ABF700D4BBE7FF" DOI="10.1163/187525410X12578602960380" author="Lewis JGE" firstAuthor="Lewis" issue="3" journalOrPublisher="International Journal of Myriapodology" page="92" pagination="83-122" refId="B24" refString="Lewis JGE (2010) A key and annotated list of the Scolopendra species of the Old World with a reappraisal of Arthrorhabdus (Chilopoda: Scolopendromorpha: Scolopendridae). International Journal of Myriapodology 1 (3): 83–122. https://doi.org/10.1163/187525410X12578602960380" title="A key and annotated list of the Scolopendra species of the Old World with a reappraisal of Arthrorhabdus (Chilopoda: Scolopendromorpha: Scolopendridae)." volume="1" year="2010">Lewis 2010: 92</bibRefCitation>
|
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</treatmentCitation>
|
||||
, 110, fig. 18 (in keys);
|
||||
<treatmentCitation id="915EF6B32470968B07F6062BC7DB82AD" author="Kronmüller C & Lewis JGE" page="269 - 278" year="2015">
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<bibRefCitation id="595E0BB350F6DAD1289490A7EB27D280" DOI="10.3897/zookeys.510.8674" author="Kronmüller C & Lewis JGE" editor="Tuf IH & Tajovský K" firstAuthor="Kronmuller" journalOrPublisher="ZooKeys" page="269 - 278" pagination="269-278" refId="B21" refString="Kronmüller C, Lewis JGE (2015) On the function of the ultimate legs of some Scolopendridae (Chilopoda, Scolopendromorpha). In: Tuf IH, Tajovský K (Eds) Proceedings of the 16 th International Congress of Myriapodology, Olomouc, Czech Republic. ZooKeys 510: 269–278. https://doi.org/10.3897/zookeys.510.8674" title="On the function of the ultimate legs of some Scolopendridae (Chilopoda, Scolopendromorpha)." volume="510" volumeTitle="Proceedings of the 16 th International Congress of Myriapodology, Olomouc, Czech Republic." year="2015">Kronmüller and Lewis 2015: 269–278</bibRefCitation>
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</treatmentCitation>
|
||||
, fig. 2 A (not type);
|
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<treatmentCitation id="7BF34B43E2EF3F1EB0EE0C3C8BC48EAB" author="Doménech C & Barbera VM & Larriba E" page="401 - 427" year="2018">
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<bibRefCitation id="E693A30ABA41C5BC0908356461E7B18B" DOI="10.11646/zootaxa.4483.3.1" author="Doménech C & Barbera VM & Larriba E" etAl="et al." firstAuthor="Domenech" issue="3" journalOrPublisher="Zootaxa" page="401 - 427" pagination="401-427" refId="B10" refString="Doménech C, Barbera VM, Larriba E (2018) A phylogenetic approach to the Philippines endemic centipedes of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae), with the description of a new species. Zootaxa 4483 (3): 401–427. https://doi.org/10.11646/zootaxa.4483.3.1" title="A phylogenetic approach to the Philippines endemic centipedes of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae), with the description of a new species." volume="4483" year="2018">Doménech et al. 2018: 401–427</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
, table 4, figs 3 A, B (not types), 4 (current
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<typeStatus id="3B07ED0D841556C42E8CF27E9C2CB202" type="paralectotype">paralectotype</typeStatus>
|
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3), and 15 (voucher, not type);
|
||||
<treatmentCitation id="0CB4F9587425B5C28CE16FA0537BFCA4" author="Acuña DC & Isagani N & Pitogo KME" page="417 - 419" year="2021">
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||||
<bibRefCitation id="B21EBC11FA702F784D1294C4B5DB277D" DOI="10.17161/randa.v28i3.15782" author="Acuña DC & Isagani N & Pitogo KME" etAl="et al." firstAuthor="Acuna" issue="3" journalOrPublisher="Reptiles & Amphibians" page="417 - 419" pagination="417-419" refId="B1" refString="Acuña DC, Isagani N, Pitogo KME (2021) Predation on a McClung’s Philippine False Coralsnake, Hemibungarus mcclungi (Weigmann 1835), by a Giant Spiny Centipede, Scolopendra spinosissima Kraepelin 1903, on Luzon Island, the Philippines. Reptiles & Amphibians 28 (3): 417–419. https://doi.org/10.17161/randa.v28i3.15782" title="Predation on a McClung’s Philippine False Coralsnake, Hemibungarus mcclungi (Weigmann 1835), by a Giant Spiny Centipede, Scolopendra spinosissima Kraepelin 1903, on Luzon Island, the Philippines." volume="28" year="2021">Acuña et al. 2021: 417–419</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
, figs 1, 2 (not type).
|
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</comment>
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</paragraph>
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||||
</subSubSection>
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<subSubSection id="SECID0E24AE" type="diagnosis">
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<paragraph id="CE5D402B113E2F16B86D43C9A887BBD4">
|
||||
<heading id="764A6FD40B9D0B866A5C6ECCF61DE061" reason="title">Diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8DBC4AFC23C9114B7D22EE6406FC07D8">
|
||||
Colouration dark red to brownish. Antennae reaching posterior border of
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<collectionCode id="02FB05E967E13E48AB31872E2849C964" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
3, rarely
|
||||
<collectionCode id="A55FDC0A6D170D9759245FB472AAAA5E" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
4; with 19 antennal articles, basal four glabrous. Paramedian sutures on tergites highly variable in TT 1–7, in TT 8–20 complete. Paramedian sutures on sternite incomplete in
|
||||
<abbrev id="ABBRID0EB5AE" xlink_title="sternite/s">SS</abbrev>
|
||||
2–20. Free coxopleuron edge not extending beyond the
|
||||
<collectionCode id="E9AF6526B3D204939E2657DBC01987EF" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
21 posterior edge. Coxopleural process moderately long and not inflected with coxopleuron, forming together an angle of ~ 120 °. Coxopleural process with one
|
||||
<abbrev id="ABBRID0EF5AE" xlink_title="apical spine">AP</abbrev>
|
||||
and one smaller dorsal
|
||||
<abbrev id="ABBRID0EJ5AE" xlink_title="subapical spine">SAP</abbrev>
|
||||
, rarely with an extra ventral
|
||||
<abbrev id="ABBRID0EN5AE" xlink_title="subapical spine">SAP</abbrev>
|
||||
.
|
||||
<abbrev id="ABBRID0ER5AE" xlink_title="ultimate legs">
|
||||
<collectionCode id="0667CC0E48B5729AD9E4024607734FE4" country="USA" name="University of Louisville">UL</collectionCode>
|
||||
</abbrev>
|
||||
prefemur with single spine tipping long spinous processes disposed in
|
||||
<abbrev id="ABBRID0EV5AE" xlink_title="ventro-lateral process">VL</abbrev>
|
||||
: 1,
|
||||
<collectionCode id="776B63108C214AC2FCF20BF19CB9FCCC" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
|
||||
: 2,
|
||||
<abbrev id="ABBRID0EZ5AE" xlink_title="ventro-median process">VM</abbrev>
|
||||
: 2, M: 1,
|
||||
<abbrev id="ABBRID0E45AE" xlink_title="dorso-median process">
|
||||
<collectionCode id="CA4DADBF94FFFB61161EAD07D2577115" country="New Zealand" name="Dominion Museum">DM</collectionCode>
|
||||
</abbrev>
|
||||
: 2 and
|
||||
<abbrev id="ABBRID0EB6AE" xlink_title="spine on prefemoral corner process">SP</abbrev>
|
||||
: 1. Penis, gonopods, and secondary sexual characters in males absent.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EF6AE" type="Lectotype">
|
||||
<paragraph id="256091D66D2F13E829376AD979C5986A">
|
||||
<heading id="44C306D7BD4B48A311AE3603FCE3A622" reason="title">
|
||||
<typeStatus id="AF3F6CB8FD6A73E0C2ADC6A3196DDEE1" type="lectotype">Lectotype</typeStatus>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="4C1AC19F78921E5013C8CDAF1C468F27">
|
||||
<emphasis id="491AC3A639D6F6264E57064E37C829E1" bold="true">(new designation).</emphasis>
|
||||
<materialsCitation id="E1637C23746178CED10CEC2DA27908DB" collectionCode="VI, ZMH" collectorName="H. W. Brolemann & Paralectotypes." country="Philippines" location="Luzon Island" municipality="Luzon Island" specimenCode="A 0000633, A 00016058, A 00016060" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Manila" typeStatus="lectotype">
|
||||
<collectingCountry id="E5F330E4BBCC270541AE6B061A8A5110" name="Philippines">Philippines</collectingCountry>
|
||||
• 1 unsexed adult;
|
||||
<collectingMunicipality id="637A91B23705E8588F30AA5DD5504548">Luzon Island</collectingMunicipality>
|
||||
,
|
||||
<collectingRegion id="75C3021D21ED2FD33F76A3A13DFAFC77" country="Philippines" name="Manila">Manila</collectingRegion>
|
||||
; 1902;
|
||||
<collectorName id="A4990CCFD4BB4336E9D0BBB3F87D583F">H. W. Brölemann</collectorName>
|
||||
leg.; “
|
||||
<taxonomicName id="19D4746FAF8B56D53D2958DD209E7924" authority="Krpln. ex Typis. Mus. Paris." class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="variety" species="subspinipes" variety="spinosissima">
|
||||
<emphasis id="BF4451DACC4D8DC95147793BBD960F56" italics="true">Scolopendra subspinipes</emphasis>
|
||||
Leach.
|
||||
<emphasis id="5EB3881991B6C40387D2DA701437F373" italics="true">var. spinosissima</emphasis>
|
||||
Krpln. ex Typis. Mus. Paris.
|
||||
</taxonomicName>
|
||||
[unreadable].
|
||||
<collectionCode id="F49914BDD6D01290E16F39CB75C4E699" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
|
||||
. 03.
|
||||
<collectingRegion id="B5D5B16C77A8F141AE65593E509C9654" country="Philippines" name="Manila">Manila</collectingRegion>
|
||||
”, “
|
||||
<emphasis id="9698FD22B7D6B87534BB83A44786DEEF" italics="true">
|
||||
Scolop.
|
||||
<taxonomicName id="4361184094FA54CE541CAAEF854A9134" authority="Krpln. ex Typis. Mus. Paris. VI." class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="variety" species="subspinipes" variety="spinosissima">
|
||||
subspinipes Leach.
|
||||
<emphasis id="3D2603ACE0D1DF929EA215E2630907AA" italics="true">var. spinosissima</emphasis>
|
||||
Krpln. ex Typis. Mus. Paris. VI.
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
03.
|
||||
<collectingRegion id="79E4630E9E55B55B0FAFDC2453DB5882" country="Philippines" name="Manila">Manila</collectingRegion>
|
||||
”, “
|
||||
<taxonomicName id="8B06CB85F1AD9F241D80F21A1B2B5077" authority="Kraepelin" authorityName="Kraepelin" authorityYear="1903" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="subSpecies" species="subspinipes" subSpecies="spinosissima">
|
||||
<emphasis id="E713D1F5B1984551ED628D8CAA531071" italics="true">Scolopendra subspinipes spinosissima</emphasis>
|
||||
Kraepelin
|
||||
</taxonomicName>
|
||||
[red label fixed in jar] ”;
|
||||
<abbrev id="ABBRID0EFBAG" xlink_title="Zoological Museum of Hamburg">
|
||||
<collectionCode id="27AFAACC90C8A98918540B51E50A6D95" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" type="Museum">ZMH</collectionCode>
|
||||
</abbrev>
|
||||
-
|
||||
<specimenCode id="2F102ECA850C2CB58DA672EA3C7AB8D1" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" type="Museum">A 0000633</specimenCode>
|
||||
.
|
||||
<emphasis id="C10327EBA1ADEF22F9DFBED0F88AC999" bold="true">
|
||||
<collectorName id="CCF7D0266FB3DBD9A7FBBFFE2D30FE48">
|
||||
<emphasis id="183E2AEB3B48ABA16B3321C2C2C7E874" bold="true" italics="true">
|
||||
<typeStatus id="09CCE24015E9C3472EA89273BBE82DF9" type="paralectotype">Paralectotypes</typeStatus>
|
||||
</emphasis>
|
||||
.
|
||||
</collectorName>
|
||||
</emphasis>
|
||||
<collectingCountry id="C3ADDE3DCF4EBFD579C05953BEFD65F6" name="Philippines">Philippines</collectingCountry>
|
||||
•
|
||||
<specimenCount id="3FBE6BA696E59B8A8A99076D536D585D" count="1" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="F8D8D6F024D70851836909384DFE6123" count="1" type="female">1 ♀</specimenCount>
|
||||
, 1 unsexed;
|
||||
<location id="4F3686E211E2AC54674C173289A32BBA" LSID="urn:lsid:plazi:treatment:C7A1D79B93A55F6CAEEF71D0D627DA9F:4F3686E211E2AC54674C173289A32BBA" country="Philippines" municipality="Luzon Island" name="Luzon Island" stateProvince="Manila">Luzon Island</location>
|
||||
,
|
||||
<collectingRegion id="0C87A6663BE2A8B31DED004CCCFB7397" country="Philippines" name="Quezon">Quezón Province</collectingRegion>
|
||||
, Dolores; same collection data as for the lectotype; “
|
||||
<emphasis id="431CD11EA34C16D3C971EB4BF1D76E3B" italics="true">
|
||||
Scolop.
|
||||
<taxonomicName id="956D2AE157AEA0B88F8A4D80949A0F40" authority="Krpln. Mus. Paris." class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="variety" species="subspinipes" variety="spinosissima">
|
||||
subspinipes Leach.
|
||||
<emphasis id="01FEAB4A6DD5DEFB7C045DFFA88FF764" italics="true">var. spinosissima</emphasis>
|
||||
Krpln. Mus. Paris.
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
30.
|
||||
<collectionCode id="3675DFF59DF33921477ABDC4CFE19950" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
|
||||
. 03. Camarines. Philippinen ”;
|
||||
<abbrev id="ABBRID0EBCAG" xlink_title="Zoological Museum of Hamburg">
|
||||
<collectionCode id="7338F42759B0B8238185FEDF735647C8" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" type="Museum">ZMH</collectionCode>
|
||||
</abbrev>
|
||||
-
|
||||
<specimenCode id="52CC601C35F85F0688129EA091F1CA2C" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" type="Museum">A 00016058</specimenCode>
|
||||
to
|
||||
<specimenCode id="80C52BD8E36CC4AC8FEEE21EFBE3C93D" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" type="Museum">A 00016060</specimenCode>
|
||||
.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EFCAG" type="type locality">
|
||||
<paragraph id="BB97603E41C7DA732B9064D4D142A143">
|
||||
<heading id="7F6BC88DEC40C62785316EC3DF6D9994" reason="title">Type locality.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="BC260B9779B6A0D4A78E49175BF1C40B">
|
||||
Since a
|
||||
<typeStatus id="C4CA16E13FEB051153238CED3B1F3195" type="lectotype">lectotype</typeStatus>
|
||||
is now designated,
|
||||
<collectingRegion id="3B347F99B182E9E72069523A5871304E" country="Philippines" name="Manila">Manila</collectingRegion>
|
||||
, Luzon Island,
|
||||
<collectingCountry id="56E3120C041C94A1238F19B2B44BA1DE" name="Philippines">Philippines</collectingCountry>
|
||||
(and not just
|
||||
<collectingCountry id="273C49C20C0305A3AA840E3B43A073CD" name="Philippines">Philippines</collectingCountry>
|
||||
) is the current type locality (Figs
|
||||
<figureCitation id="E195442E84129F6D2FA4CEDA98B7101A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scolopendra spinosissima Kraepelin, 1903 A, B habitus in vivo showing the only two known variations of colouration; specimens from Panay Is. (A) and Guimaras Is. (B) (photographs courtesy of E. Währen; individuals uncollected) C distribution, 20 th century records: red square = lectotype, Manila, Luzon Is.; red circle = paralectotypes 1–3, Camarines Peninsula, Luzon Is.; yellow square = non-types, Dolores, Quezon province (previously Tayabas), Luzon Is. (in the literature erroneously placed in Dolores, Camarines Peninsula, indicated by a red circle). 21 st century data: orange squares = (top) Mt. Isarog, Camarines Sur province, Luzon Is.; (middle) Cadiz, Negros Occidental province, Negros Is.; (bottom) Barili, Cebu Prov., Cebu Is. New data: blue squares: (top) Idiacacan, Pandan (Antique), Panay Is. (~ 11.683537; 122.114986); (bottom) Jordan, Guimaras Is. (~ 10.652871; 122.602054) (credits: E. Währen). Mindanao Is. indicates the origin of the specimen “ Mus. Paris. I. XI. 03 ” misidentified by Kraepelin as “ S. spinosissima ” (specimen currently excluded from the type series)." figureDoi="10.3897/zookeys.1215.129410.figure1" httpUri="https://binary.pensoft.net/fig/1155662">1 C</figureCitation>
|
||||
,
|
||||
<figureCitation id="F229F465F3C0C84C74E55FDD289E76DB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A Scolopendra spinosissima Kraepelin, 1903; lectotype in original container (ZMH - A 0000633). Notice the two labels inside and the red label fixed outside the jar, indicating type status B labels of the original type series proposed by Kraepelin in 1903: lectotype from Manila (two labels on top), paralectotypes from Camarines (bottom left), and specimen “ Mus. Paris. I. XI. 03 ” from Mindanao (bottom right, specimen excluded from the type series) C the manuscript list in which Kraepelin pointed out the type status of these specimens, localities, and identification dates. Notice that these five specimens, making up the type series, are ascribed to the data “ S. 262–263 ”: S. is an abbrev. of “ Seiten ”, meaning pages in German, which is a reference to the page numbers of the 1903 publication containing the original description. Photograph courtesy of N. Dupérré." figureDoi="10.3897/zookeys.1215.129410.figure2" httpUri="https://binary.pensoft.net/fig/1155663">2 B, C</figureCitation>
|
||||
) (
|
||||
<bibRefCitation id="BE72F45E30A799DE7DF067263FACCBEC" journalOrPublisher="The International Trust for Zoological Nomenclature, London, UK" refId="B13" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK, 124 pp." title="International Code of Zoological Nomenclature. Fourth Edition." year="1999">ICZN 1999</bibRefCitation>
|
||||
: Art. 73.2. 3, 76.2;
|
||||
<bibRefCitation id="3E2DEE0A7A31121343020CFBC0479CF0" author="Kraepelin" firstAuthor="Kraepelin" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pagination="1-276" refId="B17" refString="Kraepelin K (1903) Revision der Scolopendriden. Mitteilungen aus dem Naturhistorischen Museum in Hamburg 20: 1–276." title="Revision der Scolopendriden." volume="20" year="1903">Kraepelin 1903</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E2CAG" type="Type depository">
|
||||
<paragraph id="DC407C9B860A9E48756F3C7D2220E191">
|
||||
<heading id="F8BA2844820A597658D7F48BCC797E5E" reason="title">Type depository</heading>
|
||||
</paragraph>
|
||||
<paragraph id="66B7799E782737038D8FB3B6EE499077">
|
||||
<emphasis id="DCEC5321F3D8187D5CC83E450DB4FEF2" bold="true">(new data).</emphasis>
|
||||
All
|
||||
<typeStatus id="BE5AC900F3328445DBD4AA27D241FDB6">type</typeStatus>
|
||||
material is deposited in the collection of
|
||||
<abbrev id="ABBRID0EDDAG" xlink_title="Zoological Museum of Hamburg">
|
||||
<collectionCode id="AA2A679A3E33E0B00B94498F4AA8C6CA" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" type="Museum">ZMH</collectionCode>
|
||||
</abbrev>
|
||||
,
|
||||
<collectingRegion id="8DD860C1AE3C3E965EB2BC2F0565D42B" country="Germany" name="Hamburg">Hamburg</collectingRegion>
|
||||
(
|
||||
<collectingCountry id="8CF001F0D4DDC792B6A4592C2AA7AB1B" name="Germany">Germany</collectingCountry>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EHDAG" type="Legitum">
|
||||
<paragraph id="FD35F61FF048ADC368EF65E413BC51A0">
|
||||
<heading id="7F13203F2102B2A057169ED4DD4436C4" reason="title">Legitum</heading>
|
||||
</paragraph>
|
||||
<paragraph id="7A9D0713914152CB60CED57727951402">
|
||||
<emphasis id="A2C24BF0762AC30EC5A88A9EA919762F" bold="true">(new data).</emphasis>
|
||||
H. W. Brölemann, 1902.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EPDAG" type="distribution">
|
||||
<paragraph id="71102B45579D3B8DB505BAB6F25D51F8">
|
||||
<heading id="B36F15389F2AC183BDA36343FE0DA6A3" reason="title">Distribution.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="DDDB784348D32E4D4C952676852D77D5">
|
||||
<collectingCountry id="16423927FDB8FD6CCD16AD8498171CD2" name="Philippines">Philippines</collectingCountry>
|
||||
, endemic. Known from the islands of Luzon,
|
||||
<collectingRegion id="0CA6B858CA5E884EF1965F51A3A3FF2B" country="Philippines" name="Cebu">Cebu</collectingRegion>
|
||||
, Negros,
|
||||
<collectingRegion id="05BD2E49554A9F4BADA8BAB4512271E7" country="Philippines" name="Guimaras">Guimaras</collectingRegion>
|
||||
, and Panay (Fig.
|
||||
<figureCitation id="2DD7BBF371744CCF642A62AB5B54F3D1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scolopendra spinosissima Kraepelin, 1903 A, B habitus in vivo showing the only two known variations of colouration; specimens from Panay Is. (A) and Guimaras Is. (B) (photographs courtesy of E. Währen; individuals uncollected) C distribution, 20 th century records: red square = lectotype, Manila, Luzon Is.; red circle = paralectotypes 1–3, Camarines Peninsula, Luzon Is.; yellow square = non-types, Dolores, Quezon province (previously Tayabas), Luzon Is. (in the literature erroneously placed in Dolores, Camarines Peninsula, indicated by a red circle). 21 st century data: orange squares = (top) Mt. Isarog, Camarines Sur province, Luzon Is.; (middle) Cadiz, Negros Occidental province, Negros Is.; (bottom) Barili, Cebu Prov., Cebu Is. New data: blue squares: (top) Idiacacan, Pandan (Antique), Panay Is. (~ 11.683537; 122.114986); (bottom) Jordan, Guimaras Is. (~ 10.652871; 122.602054) (credits: E. Währen). Mindanao Is. indicates the origin of the specimen “ Mus. Paris. I. XI. 03 ” misidentified by Kraepelin as “ S. spinosissima ” (specimen currently excluded from the type series)." figureDoi="10.3897/zookeys.1215.129410.figure1" httpUri="https://binary.pensoft.net/fig/1155662">1 C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EZDAG" type="Current rank and status">
|
||||
<paragraph id="4920F950CF508286F1DD26BEE0B757B2">
|
||||
<heading id="31D06632887FB66471C8C2D66D4A56F4" reason="title">Current rank and status.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="6DFDCF51D0E46CFEF385A4423840CF34">Accepted species.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E5DAG" type="description">
|
||||
<paragraph id="2DA563EEFA155763FA554F24985617AE">
|
||||
<heading id="07AB430EB685C0CD2A172E66CE3DB0BA" reason="title">
|
||||
<typeStatus id="4DCC24386FA93B874E4C9A7A4D789376" type="lectotype">Lectotype</typeStatus>
|
||||
redescription
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="138CEE8ACFBBCB7D718CA5815831FECB">
|
||||
(variation of
|
||||
<typeStatus id="04A6C8C15F2AF8592EB6E230B657E4DA" type="paralectotype">paralectotypes</typeStatus>
|
||||
given in parentheses). Body length reaching
|
||||
<quantity id="742D0B68DF0092B1F609A70C6BCE61BF" metricMagnitude="-1" metricUnit="m" metricValue="1.47" unit="mm" value="147.0">147 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="9AF0B9A176E13E626C601FD1BE87BCAD">
|
||||
Live specimens dark red to brownish with cephalic plate and TT 8–11 usually darker. Antennae and coxopleuron orange. Legs reddish to yellowish orange. Coxosternal surface and
|
||||
<abbrev id="ABBRID0EFEAG" xlink_title="sternite/s">SS</abbrev>
|
||||
pale yellow (Fig.
|
||||
<figureCitation id="2ED013C6B03581FA56CC263DF9E19323" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scolopendra spinosissima Kraepelin, 1903 A, B habitus in vivo showing the only two known variations of colouration; specimens from Panay Is. (A) and Guimaras Is. (B) (photographs courtesy of E. Währen; individuals uncollected) C distribution, 20 th century records: red square = lectotype, Manila, Luzon Is.; red circle = paralectotypes 1–3, Camarines Peninsula, Luzon Is.; yellow square = non-types, Dolores, Quezon province (previously Tayabas), Luzon Is. (in the literature erroneously placed in Dolores, Camarines Peninsula, indicated by a red circle). 21 st century data: orange squares = (top) Mt. Isarog, Camarines Sur province, Luzon Is.; (middle) Cadiz, Negros Occidental province, Negros Is.; (bottom) Barili, Cebu Prov., Cebu Is. New data: blue squares: (top) Idiacacan, Pandan (Antique), Panay Is. (~ 11.683537; 122.114986); (bottom) Jordan, Guimaras Is. (~ 10.652871; 122.602054) (credits: E. Währen). Mindanao Is. indicates the origin of the specimen “ Mus. Paris. I. XI. 03 ” misidentified by Kraepelin as “ S. spinosissima ” (specimen currently excluded from the type series)." figureDoi="10.3897/zookeys.1215.129410.figure1" httpUri="https://binary.pensoft.net/fig/1155662">1 A, B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="5D5613CD5F9F9AF121099DBB2AB97091">
|
||||
Antennae reaching posterior border on
|
||||
<collectionCode id="B208588CED9A2B750A306718A00AC434" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
3 (
|
||||
<collectionCode id="AD18AC9C30686E9A5E04BC6A432EE1E0" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
<quantity id="8B5DD57524B97609E4A3DFB61796829A" metricMagnitude="-1" metricUnit="m" metricValue="1.016" unit="in" value="4.0">4 in</quantity>
|
||||
<typeStatus id="527B4B1A7BC377BC5F04E3527576DFA3" type="paralectotype">paralectotype</typeStatus>
|
||||
1), with 19 articles (
|
||||
<quantity id="DA951F9F160B0607B9470E88867256E6" metricMagnitude="-1" metricUnit="m" metricValue="4.699" metricValueMax="5.08" metricValueMin="4.318" unit="in" value="18.5" valueMax="20.0" valueMin="17.0">17–20 in</quantity>
|
||||
<typeStatus id="28FFA67FFD831FC7E0C2D8750A12FF0D" type="paralectotype">paralectotypes</typeStatus>
|
||||
1 and 2), the basal four glabrous dorsally and ventrally (Fig.
|
||||
<figureCitation id="1D1F3FFFF8998764F4FE6BD58764CD45" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A cephalic plate and right antenna, dorsal view B forcipular segment, ventral view C tooth-plates, ventral view D left spiracle on segment 8. Scale bars: 0.5 mm (C, D); 2 mm (A, B)." figureDoi="10.3897/zookeys.1215.129410.figure3" httpUri="https://binary.pensoft.net/fig/1155664">3 A</figureCitation>
|
||||
). Cephalic plate with four ocelli in each side. Surface covered by dispersed small puncta allocating a short sensillum each; median sulcus absent. Posterior part of cephalic plate without paramedian sulci, overlapping the anterior margin of
|
||||
<collectionCode id="3408AD5ECBBC395448D0BD279490AB7C" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
1 (Fig.
|
||||
<figureCitation id="FCD258D881F26A5C9758B078A82F7A45" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A cephalic plate and right antenna, dorsal view B forcipular segment, ventral view C tooth-plates, ventral view D left spiracle on segment 8. Scale bars: 0.5 mm (C, D); 2 mm (A, B)." figureDoi="10.3897/zookeys.1215.129410.figure3" httpUri="https://binary.pensoft.net/fig/1155664">3 A</figureCitation>
|
||||
). Coxosternite surface essentially smooth, counting with few isolated and less deep puncta; median suture absent (Fig.
|
||||
<figureCitation id="361EB3AA1B0A33E1C740227C029D3EE8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A cephalic plate and right antenna, dorsal view B forcipular segment, ventral view C tooth-plates, ventral view D left spiracle on segment 8. Scale bars: 0.5 mm (C, D); 2 mm (A, B)." figureDoi="10.3897/zookeys.1215.129410.figure3" httpUri="https://binary.pensoft.net/fig/1155664">3 B, C</figureCitation>
|
||||
). Article 2 of second maxillary telopodite with spur (Fig.
|
||||
<figureCitation id="D93D2198A03CB8B7B36C85444CD30B67" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A cephalic plate and right antenna, dorsal view B forcipular segment, ventral view C tooth-plates, ventral view D left spiracle on segment 8. Scale bars: 0.5 mm (C, D); 2 mm (A, B)." figureDoi="10.3897/zookeys.1215.129410.figure3" httpUri="https://binary.pensoft.net/fig/1155664">3 C</figureCitation>
|
||||
). Forcipula surface covered by dispersed small puncta. Left tarsungulum lost. Forcipular trochanteroprefemoral process with denticles in two groups, apically with two teeth on the right and three on the left, and proximally, one tooth on the right and two teeth on the left (a total of
|
||||
<quantity id="88F921F5D884A230186F5ACCCB68275A" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="12.7" metricValueMin="5.08" unit="in" value="3.5" valueMax="5.0" valueMin="2.0">2–5 in</quantity>
|
||||
<typeStatus id="C9395AF8ADB8829C3CBB51AAAA079A0B" type="paralectotype">paralectotypes</typeStatus>
|
||||
). Tooth-plates longer than wide, with small dispersed puncta and 7 + 8 teeth divided in two groups (5 +
|
||||
<quantity id="78458D3D8A4DAF31C731C6C419ED82DD" metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
|
||||
<typeStatus id="8D414662602542C1495791A30995330B" type="paralectotype">paralectotype</typeStatus>
|
||||
1). Tooth-plate with straight, transverse basal suture (Fig.
|
||||
<figureCitation id="F1440931D0AF0FAAF2BF480E2DE922EC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A cephalic plate and right antenna, dorsal view B forcipular segment, ventral view C tooth-plates, ventral view D left spiracle on segment 8. Scale bars: 0.5 mm (C, D); 2 mm (A, B)." figureDoi="10.3897/zookeys.1215.129410.figure3" httpUri="https://binary.pensoft.net/fig/1155664">3 C</figureCitation>
|
||||
; Table
|
||||
<tableCitation id="7F4BFD3E9944E144B7B34AE82FED93BE" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparison between the type series of Scolopendra spinosissima Kraepelin, 1903, its voucher, and S. paradoxa Doménech, 2018 holotype. A = Absent; C = Complete; I = Incomplete; D = Distal; P = Proximal; PR = Partially retracted; R = Retracted. * = Appendix damaged. N / A = Not applicable." httpUri="http://table.plazi.org/id/BC7916AA261A079E149D4E809E484642" tableUuid="BC7916AA261A079E149D4E809E484642">1</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="87B9F7DC4AE29A56CDAA1DBDDD015F3A" ID-DOI="10.3897/zookeys.1215.129410.figure3" ID-arpha="4923675E-4CA7-50B8-96AE-6AEB2267C270" httpUri="https://binary.pensoft.net/fig/1155664" startId="F3">
|
||||
<paragraph id="3CFEDDA40A364EE5657C75BB30BC38EF">
|
||||
<label id="C54A34E0815B4EF7FCB1F2CD6AA77E3A">Figure 3.</label>
|
||||
</paragraph>
|
||||
<paragraph id="654B8226E26A7CE88D6D18071D2201F8">
|
||||
<taxonomicName id="BCE0936549ED243C861FAD3E34A53192" authority="Kraepelin, 1903" authorityName="sensu Kraepelin" authorityYear="1903" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="spinosissima">
|
||||
<emphasis id="200EEDE16E2EAF6FE5F6DBF82FFBF690" italics="true">Scolopendra spinosissima</emphasis>
|
||||
<bibRefCitation id="AB50B7DD8159369C2BBE5461168B06DC" author="Kraepelin K" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pagination="1-276" refId="B17" refString="Kraepelin K (1903) Revision der Scolopendriden. Mitteilungen aus dem Naturhistorischen Museum in Hamburg 20: 1 - 276." title="Revision der Scolopendriden." volume="20" year="1903">Kraepelin, 1903</bibRefCitation>
|
||||
</taxonomicName>
|
||||
; lectotype (
|
||||
<abbrev id="ABBRID0E1FAG" xlink_title="Zoological Museum of Hamburg">ZMH</abbrev>
|
||||
- A 0000633)
|
||||
<emphasis id="2997B87C670DB59DBFBC2B6F6D62C954" bold="true">A</emphasis>
|
||||
cephalic plate and right antenna, dorsal view
|
||||
<emphasis id="D260B09BE22E8CAB4488C1F6570B8722" bold="true">B</emphasis>
|
||||
forcipular segment, ventral view
|
||||
<emphasis id="2545759A321AED823A02B2A6C096B249" bold="true">C</emphasis>
|
||||
tooth-plates, ventral view
|
||||
<emphasis id="AF108BEED2362CCCD76E1040770C3733" bold="true">D</emphasis>
|
||||
left spiracle on segment 8. Scale bars: 0.5 mm (
|
||||
<emphasis id="567E76F04E9DFDE042872C975B77AF43" bold="true">C, D</emphasis>
|
||||
); 2 mm (
|
||||
<emphasis id="52B3E7B6A20FF909144339D5D1A163C6" bold="true">A, B</emphasis>
|
||||
).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="BC7916AA261A079E149D4E809E484642" ID-Table-UUID="BC7916AA261A079E149D4E809E484642" httpUri="http://table.plazi.org/id/BC7916AA261A079E149D4E809E484642" startId="T1" targetIsTable="true">
|
||||
<paragraph id="4C0F5F4E6C4427AA43CF0AAD702FCB35">
|
||||
<label id="8804D483817FF0DA26B126633063DA9D">Table 1.</label>
|
||||
</paragraph>
|
||||
<paragraph id="C3211A5BE1E9D697D16F18C8C17EC1C9">
|
||||
Morphological comparison between the type series of
|
||||
<taxonomicName id="A84236AFFF2ECF3B1E457A0AE9EF1882" authority="Kraepelin, 1903" authorityName="sensu Kraepelin" authorityYear="1903" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="spinosissima">
|
||||
<emphasis id="237F2214CDC03038BE3137A272A89BC3" italics="true">Scolopendra spinosissima</emphasis>
|
||||
<bibRefCitation id="284B24DC0722456192B0C25471CD78E5" author="Kraepelin K" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pagination="1-276" refId="B17" refString="Kraepelin K (1903) Revision der Scolopendriden. Mitteilungen aus dem Naturhistorischen Museum in Hamburg 20: 1 - 276." title="Revision der Scolopendriden." volume="20" year="1903">Kraepelin, 1903</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, its voucher, and
|
||||
<taxonomicName id="F46E3914038BBD47EF080F4C68A4C183" authority="Domenech, 2018" authorityName="Domenech" authorityYear="2018" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="paradoxa">
|
||||
<emphasis id="2E60B6CB5B5932A9E9948CFF32D75C29" italics="true">S. paradoxa</emphasis>
|
||||
<bibRefCitation id="6CF63B339163682EC913F1A165C0C3CF" DOI="10.11646/zootaxa.4483.3.1" author="Doménech C & Barbera VM & Larriba E" issue="3" journalOrPublisher="Zootaxa" pagination="401-427" refId="B10" refString="Domenech C, Barbera VM, Larriba E (2018) A phylogenetic approach to the Philippines endemic centipedes of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae), with the description of a new species. Zootaxa 4483 (3): 401 - 427. https: // doi. org / 10.11646 / zootaxa. 4483.3. 1" title="A phylogenetic approach to the Philippines endemic centipedes of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae), with the description of a new species." volume="4483" year="2018">Doménech, 2018</bibRefCitation>
|
||||
</taxonomicName>
|
||||
holotype. A = Absent; C = Complete; I = Incomplete; D = Distal; P = Proximal; PR = Partially retracted; R = Retracted. * = Appendix damaged. N / A = Not applicable.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="7C414DCAAD743EF839EE8CE4060942D2">
|
||||
<table id="TID0EDSBG" rules="all">
|
||||
<tbody id="F123D35C36A4F7AA5F560C3C4598C033">
|
||||
<tr id="B4DCFA2ABC59E7CAA498BF27F49640E8">
|
||||
<th id="01EDC4F636B935D25C80B2EF373EA9CF" colspan="1" isEmpty="true" rowspan="1">-</th>
|
||||
<th id="2AF4C2D285D200B4841F45D184DEFE3C" colspan="1" rowspan="1">
|
||||
<taxonomicName id="98274155FDE166355D607C96D66A9339" authority="Domenech, 2018" authorityName="Domenech" authorityYear="2018" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="paradoxa">
|
||||
<emphasis id="5BD92159FC19CFBEDEC517ED85D88515" italics="true">Scolopendra paradoxa</emphasis>
|
||||
<bibRefCitation id="C2C72A96905888A70BF6C7DCA9410E78" DOI="10.11646/zootaxa.4483.3.1" author="Doménech C & Barbera VM & Larriba E" issue="3" journalOrPublisher="Zootaxa" pagination="401-427" refId="B10" refString="Domenech C, Barbera VM, Larriba E (2018) A phylogenetic approach to the Philippines endemic centipedes of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae), with the description of a new species. Zootaxa 4483 (3): 401 - 427. https: // doi. org / 10.11646 / zootaxa. 4483.3. 1" title="A phylogenetic approach to the Philippines endemic centipedes of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae), with the description of a new species." volume="4483" year="2018">Doménech, 2018</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</th>
|
||||
<taxonomicName id="FC1769B6BA2979F43D1DBD1C9FBE2996" authority="Kraepelin, 1903 Current" authorityName="sensu Kraepelin" authorityYear="1903" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="spinosissima">
|
||||
<th id="A125854D78F5490DB6E03BBBD388F543" colspan="5" rowspan="1">
|
||||
<emphasis id="B3C0D7998CBB998C7036320EB16A36F6" italics="true">Scolopendra spinosissima</emphasis>
|
||||
<bibRefCitation id="E8CA12418085B62B2472821D7020C5B9" author="Kraepelin K" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pagination="1-276" refId="B17" refString="Kraepelin K (1903) Revision der Scolopendriden. Mitteilungen aus dem Naturhistorischen Museum in Hamburg 20: 1 - 276." title="Revision der Scolopendriden." volume="20" year="1903">Kraepelin, 1903</bibRefCitation>
|
||||
</th>
|
||||
<tr id="D1D9E07BF9E6713DAE9F26FA356D9E38">
|
||||
<td id="D26BC4A9F5817CE31D46D1F4F321D80E" colspan="1" rowspan="1">Current type condition</td>
|
||||
<td id="3D190C540C2389E4B13F4AA2BD88CA0E" colspan="1" rowspan="1">Holotype – and voucher –</td>
|
||||
<td id="59FCC5FB5E3E8E36F5E9AAFCC74DE495" colspan="1" rowspan="1">Lectotype</td>
|
||||
<td id="D068B5B81E760E04A9273B1102848654" colspan="1" rowspan="1">Paralectotype 1</td>
|
||||
<td id="73713B3C383C86F312B8021FB1621C8B" colspan="1" rowspan="1">Paralectotype 2</td>
|
||||
<td id="5D51F4F8C554020A72178FDA27A660C5" colspan="1" rowspan="1">Paralectotype 3</td>
|
||||
<td id="33D05E0590470A320B8D4B9380A6850D" colspan="1" rowspan="1">Voucher (non type)</td>
|
||||
</tr>
|
||||
</taxonomicName>
|
||||
</tr>
|
||||
<tr id="011B877D4ED5E5C475E8AC3E772D7020">
|
||||
<td id="7ED4F3E8850D9B120768DF52E5FEE7B6" colspan="1" rowspan="1">Specimen number</td>
|
||||
<td id="F8C273FA23A46A084AD8B04839A26EDA" colspan="1" rowspan="1">CEUA 017 - Mr 0000</td>
|
||||
<td id="CEF9B8313C1BF16E6ABBB459DD4FCE3A" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EXJAG" xlink_title="Zoological Museum of Hamburg">ZMH</abbrev>
|
||||
- A 0000633
|
||||
</td>
|
||||
<td id="AAFE8E53F07B06CF5D246F16748EA1C4" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0E6JAG" xlink_title="Zoological Museum of Hamburg">ZMH</abbrev>
|
||||
- A 00016058
|
||||
</td>
|
||||
<td id="11D79DB470865A9E86A4FD65079E03C8" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EHKAG" xlink_title="Zoological Museum of Hamburg">ZMH</abbrev>
|
||||
- A 00016059
|
||||
</td>
|
||||
<td id="E75567CE6E895BA369D90194C50072B4" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EPKAG" xlink_title="Zoological Museum of Hamburg">ZMH</abbrev>
|
||||
- A 00016060
|
||||
</td>
|
||||
<td id="C15DF67E8F2F7B6746665E8C3A312299" colspan="1" rowspan="1">CEUA 016 - Mr 0009</td>
|
||||
</tr>
|
||||
<tr id="04500809D295D5F640D66E95B3190841">
|
||||
<td id="305489D754651F2A77D9F152AE57AF71" colspan="1" rowspan="1">Reference in previous labels and catalogue</td>
|
||||
<td id="3905CBC9E5D47AB76C2DF3A316777775" colspan="1" rowspan="1">N / A</td>
|
||||
<td id="D981758F29451D50923D6A9E00C1C583" colspan="1" rowspan="1">Mus. Paris c.; VI. 03; “ ex Typis ” (holotype)</td>
|
||||
<td id="FD3696FB80F5701691D72AC5A7902CCA" colspan="1" rowspan="1">Mus. Paris c.; 30. VI. 03; paratype</td>
|
||||
<td id="4932B4F6D7F6F7328B0336493846B937" colspan="1" rowspan="1">Mus. Paris c.; 30. VI. 03; paratype</td>
|
||||
<td id="D18D1ACA15D2C6D68A9E89EB5FAC402E" colspan="1" rowspan="1">Mus. Paris c.; 30. VI. 03; paratype</td>
|
||||
<td id="28D90F35282C56EB3E5D3E32B5390C48" colspan="1" rowspan="1">N / A</td>
|
||||
</tr>
|
||||
<tr id="473498201653FEBED128560CDB9E2DDD">
|
||||
<td id="976B9FDF739E91E44D4B741A7FC7615D" colspan="1" rowspan="1">Body length in mm</td>
|
||||
<td id="9E63FC0DEE66D85BF580DB290ECC5B34" colspan="1" rowspan="1">132</td>
|
||||
<td id="B0C561E382A0597D37DF36BC04163E3C" colspan="1" rowspan="1">147</td>
|
||||
<td id="17B376604006030F40B73026AE228BBE" colspan="1" rowspan="1">126</td>
|
||||
<td id="79917B133FD2A39A0A504A1F4956E017" colspan="1" rowspan="1">120</td>
|
||||
<td id="7DB1FB2D24E2D6A251874AB3D335DF1B" colspan="1" rowspan="1">99</td>
|
||||
<td id="9E81CCBC49838619E334C546F91F78E0" colspan="1" rowspan="1">87</td>
|
||||
</tr>
|
||||
<tr id="D7A0A2EE02247E8307B13992832C272A">
|
||||
<td id="B070DE1278033BDBDFD6B3B39761B1E3" colspan="1" rowspan="1">Sex</td>
|
||||
<td id="B8CEFF41EE2087A99C049378D0161B60" colspan="1" rowspan="1">R-Female</td>
|
||||
<td id="34D9B08962449172395EE507E2FDDC3B" colspan="1" rowspan="1">R</td>
|
||||
<td id="00F47B4F42AC70AA8B4FED79219B4B3F" colspan="1" rowspan="1">PR</td>
|
||||
<td id="F2959F5014D07A2B47E2F4619E040C2F" colspan="1" rowspan="1">Female</td>
|
||||
<td id="3438538BFD357FED22150363FF799FFC" colspan="1" rowspan="1">PR-Male</td>
|
||||
<td id="3BF036E256507BBDDB729AD30C3171D3" colspan="1" rowspan="1">* (probably female)</td>
|
||||
</tr>
|
||||
<tr id="459DF47350EFED14C986FA3ADDC3FAD3">
|
||||
<td id="649A42E9B09B2FE66C6E36DA230A65FD" colspan="1" rowspan="1">Antenna reaching to tergite</td>
|
||||
<td id="B6DE3BB62C5F56C9583C1EC8AA3DD15E" colspan="1" rowspan="1">T 5</td>
|
||||
<td id="2CD23A3C70B4D0056720471AC95B9CBF" colspan="1" rowspan="1">T 3</td>
|
||||
<td id="63CC3EA3385D071821A92F7B4BE26402" colspan="1" rowspan="1">T 4</td>
|
||||
<td id="076C89C1A02195388B2803D597A81EBF" colspan="1" rowspan="1">T 3</td>
|
||||
<td id="1BBCD92607540A384D221F634C81C4BC" colspan="1" rowspan="1">T 3</td>
|
||||
<td id="DBDA41838EAA504279F41C333E802A11" colspan="1" rowspan="1">T 4</td>
|
||||
</tr>
|
||||
<tr id="0DE5CF0DD029BEB7A0E1B010A077927C">
|
||||
<td id="67CD5CA908AC16E1DDAF9582F3921AFB" colspan="1" rowspan="1">Number of antennal articles</td>
|
||||
<td id="7CBA10172D2F4F5ABE3B8D306BF3A9DE" colspan="1" rowspan="1">19 / 19</td>
|
||||
<td id="A2AD6D12C1048E79E7A5E573CC1B25C8" colspan="1" rowspan="1">19 / 19</td>
|
||||
<td id="F474E41BD1F6F62047D90AEEC12061D0" colspan="1" rowspan="1">17 / 20</td>
|
||||
<td id="0D81CC30C77C31AD48177713AAAE1176" colspan="1" rowspan="1">18 / 19</td>
|
||||
<td id="A4F7822D84C7AAED970AA9AC0A3CD61D" colspan="1" rowspan="1">19 / 19</td>
|
||||
<td id="4A63347BA84185A29D074EB1F2672951" colspan="1" rowspan="1">19 / 19</td>
|
||||
</tr>
|
||||
<tr id="3AEE8807F117ECECC5A12A2D8E28A41F">
|
||||
<td id="A8E2DBAD4DFBD8CAA5BF6C4D4A6142DF" colspan="1" rowspan="1">Number of proximal glabrous articles</td>
|
||||
<td id="CAFD8D68DA72DC57E98E729401CBD8D8" colspan="1" rowspan="1">4 / 4 dorsally 5 ½ / 5 ½ ventrally</td>
|
||||
<td id="E3116C9B6B97084FB526BD64C7A6645F" colspan="1" rowspan="1">4 / 4 dorsally and ventrally</td>
|
||||
<td id="59A00D743C6917A963E974816C877FD4" colspan="1" rowspan="1">4 / 4 dorsally and ventrally</td>
|
||||
<td id="C56A7BA8818879AB322F44596C36306B" colspan="1" rowspan="1">4 / 4 dorsally and ventrally</td>
|
||||
<td id="1D1C055B09711649650C008922C7F976" colspan="1" rowspan="1">4 / 4 dorsally and ventrally</td>
|
||||
<td id="74A49E8F8805E93ECB2D423AEABEE90E" colspan="1" rowspan="1">4 / 4 dorsally and ventrally</td>
|
||||
</tr>
|
||||
<tr id="51C27A3F53B535F5A00419F5830732F8">
|
||||
<td id="DA1B861BE15EF9BB73ABB634C643B329" colspan="1" rowspan="1">Teeth on tooth-plate</td>
|
||||
<td id="185AE61B1868BBF68BAD8E291CBB34B1" colspan="1" rowspan="1">5 + 5</td>
|
||||
<td id="09E79DDCFE8FC6F9F4056CED735D88B6" colspan="1" rowspan="1">7 + 8</td>
|
||||
<td id="BF6E25FF21BBA4A0A34F3C3C41655F86" colspan="1" rowspan="1">5 + 5</td>
|
||||
<td id="C11E215F97E7FE76ECFCD52C30B34EDD" colspan="1" rowspan="1">7 + 7</td>
|
||||
<td id="F5B16E6292DA791402E006C3A5AA45D1" colspan="1" rowspan="1">7 + 7</td>
|
||||
<td id="3BA1D83FB862155F62F2EB5CB3AEC71E" colspan="1" rowspan="1">7 + 7</td>
|
||||
</tr>
|
||||
<tr id="84022CDA36EE3B5150627F03CA784EC4">
|
||||
<td id="083E30D11BF935C8E3F009CF08BE83B4" colspan="1" rowspan="1">Teeth on forcipular trochanteroprefemoral processes as total (upper group / lower group)</td>
|
||||
<td id="15BA02F91D8D3F958C42B9FECA511C0A" colspan="1" rowspan="1">2 (1 / 1) – 1 (1 / 0)</td>
|
||||
<td id="9B67A9C3DBB23D189D2CAB4F1732BB02" colspan="1" rowspan="1">3 (2 / 1) – 5 (3 / 2)</td>
|
||||
<td id="0A85DB7BB0A119F53173BA6A2D7ACAD5" colspan="1" rowspan="1">5 (3 / 2) – 3 (2 / 1)</td>
|
||||
<td id="A9695A4E8EA993497125FFF8954FB8D9" colspan="1" rowspan="1">2 (1 / 1) – 4 (3 / 1)</td>
|
||||
<td id="18FF375C294B54E5FC22E49A54338FE4" colspan="1" rowspan="1">4 (3 / 1) – 4 (3 / 1)</td>
|
||||
<td id="486F14301ACE3A054CA50357A11EE977" colspan="1" rowspan="1">4 (3 / 1) – 5 (3 / 2)</td>
|
||||
</tr>
|
||||
<tr id="BF0836F5AF4D047E2824524CA4060CEF">
|
||||
<td id="94B5D9979E58EC5373BCD743BB31A771" colspan="1" rowspan="1">Tergite paramedian sutures</td>
|
||||
<td id="8CECEFB6F568AB4A4DB9AF3CFB5F90D8" colspan="1" rowspan="1">TT 1–2 A; TT 3–4 IDP; T 5 C; T 6 C (right side ID); T 7 IDP; TT 9–20 C; T 21 A</td>
|
||||
<td id="8C716BD764C618533106C228548A3F60" colspan="1" rowspan="1">TT 1–2 A; T 3 IP; T 4 C; T 5 IP; T 6 C; T 7 IDP; TT 9–20 C; T 21 A</td>
|
||||
<td id="12E657BDAE87C113BC838F95CCC15FAE" colspan="1" rowspan="1">TT 1 A; TT 2 ID; TT 3–5 IDP; T 6 C; TT 7–8 IDP; TT 9–20 C; T 21 A</td>
|
||||
<td id="52ACE0BFC5F0B6DE3CBCE554D38EBA5A" colspan="1" rowspan="1">T 1 A; T 2 ID; T 3 IDP; T 4 C; T 5 IDP; TT 6–20 C; T 21 A</td>
|
||||
<td id="827812C436475E61CE9872F2E99C0803" colspan="1" rowspan="1">T 1 A; T 2 ID; T 3 IDP; T 4 C; T 5 IDP; T 6 C; TT 7–20 C; T 21 A</td>
|
||||
<td id="9B686F8EDDF3050D27EEBABFB8EFC53B" colspan="1" rowspan="1">TT 1–2 A; T 3 IP; T 4–20 C; T 21 A</td>
|
||||
</tr>
|
||||
<tr id="5B167D827EFD249037FFE003341EC713">
|
||||
<td id="DED8C5752E08ABCFC8A82781EE1FC5E1" colspan="1" rowspan="1">First tergite with complete margination</td>
|
||||
<td id="E0A9DA251216B6ECEB48F13D48B9758C" colspan="1" rowspan="1">10</td>
|
||||
<td id="526A19738C99C48ACB54D5385270AB0D" colspan="1" rowspan="1">10</td>
|
||||
<td id="FA0A7427BBEE783816D7548154333AEB" colspan="1" rowspan="1">12</td>
|
||||
<td id="E560B4BDA47D18174F87E9F4ECF25776" colspan="1" rowspan="1">10</td>
|
||||
<td id="6D62C3504FA634A53AB5D49BE1379034" colspan="1" rowspan="1">10</td>
|
||||
<td id="691256C2EC5E457D4D1025EC13EA5C24" colspan="1" rowspan="1">12</td>
|
||||
</tr>
|
||||
<tr id="4AA934FAE0DFA3491A00B40FE6CC29B4">
|
||||
<td id="A13D0FEE13BB0CC91C05EECBA5F3FE32" colspan="1" rowspan="1">Paramedian sutures on sternites</td>
|
||||
<td id="89D146BF1A2D9B8BE2AB4D5B1973447C" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0E4RAG" xlink_title="sternite/s">SS</abbrev>
|
||||
1–2 A; SS 3–18 C; SS 19–21 A
|
||||
</td>
|
||||
<td id="5EF943E3D59035EB13EC5FDA2846DA4E" colspan="1" rowspan="1">S 1 A; SS 2–20 IDP; S 21 A</td>
|
||||
<td id="9108BE379406E672C343F8071A8A2243" colspan="1" rowspan="1">S 1 A; SS 2–20 IDP; S 21 A</td>
|
||||
<td id="FBC7519255E4AB0D32EEBF7828621F63" colspan="1" rowspan="1">S 1 A; SS 2–4 IP; SS 5–19 IPD; SS 20–21 A</td>
|
||||
<td id="0A913FF836EF236B06B7E69B4E87823A" colspan="1" rowspan="1">S 1 A; SS 2–5 C; SS 6–19 IPD; SS 20–21 A</td>
|
||||
<td id="C36F01E4ECC7359C154E3CECCF05A380" colspan="1" rowspan="1">S 1 A; SS 2–12 C; SS 13–19 IP; S 20–21 A</td>
|
||||
</tr>
|
||||
<tr id="4E35C996A100943AAA4DB8193A0462FC">
|
||||
<td id="4B4D22403932E612A77C56D7952ACFAE" colspan="1" rowspan="1">Spines in coxopleural process</td>
|
||||
<td id="B6E59F7FE6F4AA1FAF638E5217E534E6" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EYSAG" xlink_title="apical spine">AP</abbrev>
|
||||
: 1;
|
||||
<abbrev id="ABBRID0E3SAG" xlink_title="subapical spine">SAP</abbrev>
|
||||
: 1
|
||||
</td>
|
||||
<td id="C19A63DB796F3BF57555CED51D063B88" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EETAG" xlink_title="apical spine">AP</abbrev>
|
||||
: 1;
|
||||
<abbrev id="ABBRID0EITAG" xlink_title="subapical spine">SAP</abbrev>
|
||||
: 1; (left with 1 extra ventral
|
||||
<abbrev id="ABBRID0EMTAG" xlink_title="subapical spine">SAP</abbrev>
|
||||
spinula)
|
||||
</td>
|
||||
<td id="C96C2344237E3DDC3751EA6E3C8D1153" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EUTAG" xlink_title="apical spine">AP</abbrev>
|
||||
: 1;
|
||||
<abbrev id="ABBRID0EYTAG" xlink_title="subapical spine">SAP</abbrev>
|
||||
: 1
|
||||
</td>
|
||||
<td id="3EDBCE25AC73DE8CD744BA78D603E729" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EAUAG" xlink_title="apical spine">AP</abbrev>
|
||||
: 1;
|
||||
<abbrev id="ABBRID0EEUAG" xlink_title="subapical spine">SAP</abbrev>
|
||||
: 1
|
||||
</td>
|
||||
<td id="5AEDEFD2FCD83B7D47CB6E79FC983C30" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EMUAG" xlink_title="apical spine">AP</abbrev>
|
||||
: 1;
|
||||
<abbrev id="ABBRID0EQUAG" xlink_title="subapical spine">SAP</abbrev>
|
||||
: 1
|
||||
</td>
|
||||
<td id="A54551102C5C49AAFDD32C0A8E4336A0" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EYUAG" xlink_title="apical spine">AP</abbrev>
|
||||
: 1;
|
||||
<abbrev id="ABBRID0E3UAG" xlink_title="subapical spine">SAP</abbrev>
|
||||
: 1
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="094702109704D47C183653AD4BC660DA">
|
||||
<td id="798FDADD77DF6520FA12935FF64FD9AE" colspan="1" rowspan="1">Coxopleural process extending beyond T 21</td>
|
||||
<td id="54A553F3945591E0043EF5CC4C94ADFF" colspan="1" rowspan="1">Yes</td>
|
||||
<td id="02DF623C9D0893C14AC5FADC259470CC" colspan="1" rowspan="1">No</td>
|
||||
<td id="3476F931E5C038FB2FA96141D060B35E" colspan="1" rowspan="1">No</td>
|
||||
<td id="93B15B5190853561AFF463F5BFCA127D" colspan="1" rowspan="1">No</td>
|
||||
<td id="616945B7BB7F19E779E3EE73C1556975" colspan="1" rowspan="1">No</td>
|
||||
<td id="1DCCEE7A0F9B0CBFA01F8FC009BC22F1" colspan="1" rowspan="1">No</td>
|
||||
</tr>
|
||||
<tr id="BF065925A3E520063AE8AFB3303D5D91">
|
||||
<td id="118B59ABF28877895F5373BA7F94F2A0" colspan="1" rowspan="1">Spinous process formula on prefemora of ultimate legs</td>
|
||||
<td id="25EF63BF92A18361FA9D017C841CA3F4" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0E5VAG" xlink_title="ventro-lateral process">VL</abbrev>
|
||||
: 2 / 2; V: 0 / 0;
|
||||
<abbrev id="ABBRID0ECWAG" xlink_title="ventro-median process">VM</abbrev>
|
||||
: 1 / 1; M: 1 / 1;
|
||||
<abbrev id="ABBRID0EGWAG" xlink_title="dorso-median process">DM</abbrev>
|
||||
: 2 / 2;
|
||||
<abbrev id="ABBRID0EKWAG" xlink_title="spine on prefemoral corner process">SP</abbrev>
|
||||
: 1 / 1
|
||||
</td>
|
||||
<td id="3327630A2026F060F71F0FA9BD7AECD5" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0ESWAG" xlink_title="ventro-lateral process">VL</abbrev>
|
||||
: 1 / 1; V: 2 / 2;
|
||||
<abbrev id="ABBRID0EWWAG" xlink_title="ventro-median process">VM</abbrev>
|
||||
: 2 / 2; M: 1 / 1;
|
||||
<abbrev id="ABBRID0E1WAG" xlink_title="dorso-median process">DM</abbrev>
|
||||
: 2 / 2;
|
||||
<abbrev id="ABBRID0E5WAG" xlink_title="spine on prefemoral corner process">SP</abbrev>
|
||||
: 1 / 1
|
||||
</td>
|
||||
<td id="9468A762AA10AD707CB297A32109C5A0" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EGXAG" xlink_title="ventro-lateral process">VL</abbrev>
|
||||
: 1 / 1; V: 2 / 2;
|
||||
<abbrev id="ABBRID0EKXAG" xlink_title="ventro-median process">VM</abbrev>
|
||||
: 2 / 2; M: 1 / 1;
|
||||
<abbrev id="ABBRID0EOXAG" xlink_title="dorso-median process">DM</abbrev>
|
||||
: 2 / 2;
|
||||
<abbrev id="ABBRID0ESXAG" xlink_title="spine on prefemoral corner process">SP</abbrev>
|
||||
: 1 / 1; (left M and
|
||||
<abbrev id="ABBRID0EWXAG" xlink_title="ventro-median process">VM</abbrev>
|
||||
proximal spinous processes with spines *)
|
||||
</td>
|
||||
<td id="918FE341037AC5FEE627E5341293FC24" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0E5XAG" xlink_title="ventro-lateral process">VL</abbrev>
|
||||
: 1 / 1; V: 2 / 2;
|
||||
<abbrev id="ABBRID0ECYAG" xlink_title="ventro-median process">VM</abbrev>
|
||||
: 2 / 2; M: 1 / 1;
|
||||
<abbrev id="ABBRID0EGYAG" xlink_title="dorso-median process">DM</abbrev>
|
||||
: 2 / 2;
|
||||
<abbrev id="ABBRID0EKYAG" xlink_title="spine on prefemoral corner process">SP</abbrev>
|
||||
: 1 / 1
|
||||
</td>
|
||||
<td id="806F10C2D76EAD17806B110B83BE9DEC" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0ESYAG" xlink_title="ventro-lateral process">VL</abbrev>
|
||||
: 1 / 1; V: 2 / 2;
|
||||
<abbrev id="ABBRID0EWYAG" xlink_title="ventro-median process">VM</abbrev>
|
||||
: 2 / 2; M: 1 / 1;
|
||||
<abbrev id="ABBRID0E1YAG" xlink_title="dorso-median process">DM</abbrev>
|
||||
: 2 / 2;
|
||||
<abbrev id="ABBRID0E5YAG" xlink_title="spine on prefemoral corner process">SP</abbrev>
|
||||
: 1 / 1; (right
|
||||
<abbrev id="ABBRID0ECZAG" xlink_title="ventro-median process">VM</abbrev>
|
||||
proximal spinous process hardly noticeable, without spine); (
|
||||
<bibRefCitation id="1003D337475ECB8FCA937E9F2D637A38" DOI="10.11646/zootaxa.4483.3.1" author="Domenech" etAl="et al." firstAuthor="Domenech" issue="3" journalOrPublisher="Zootaxa" pagination="401-427" refId="B10" refString="Doménech C, Barbera VM, Larriba E (2018) A phylogenetic approach to the Philippines endemic centipedes of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae), with the description of a new species. Zootaxa 4483 (3): 401–427. https://doi.org/10.11646/zootaxa.4483.3.1" title="A phylogenetic approach to the Philippines endemic centipedes of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae), with the description of a new species." volume="4483" year="2018">Doménech et al. 2018</bibRefCitation>
|
||||
: fig. 4 A – C)
|
||||
</td>
|
||||
<td id="7E8A678005B81DFCFE1FE0527928070E" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EOZAG" xlink_title="ventro-lateral process">VL</abbrev>
|
||||
: 1 / 1; V: 2 / 2;
|
||||
<abbrev id="ABBRID0ESZAG" xlink_title="ventro-median process">VM</abbrev>
|
||||
: 2 / 2; M: 1 / 1;
|
||||
<abbrev id="ABBRID0EWZAG" xlink_title="dorso-median process">DM</abbrev>
|
||||
: 2 / 2;
|
||||
<abbrev id="ABBRID0E1ZAG" xlink_title="spine on prefemoral corner process">SP</abbrev>
|
||||
: 1 / 1; (left
|
||||
<abbrev id="ABBRID0E5ZAG" xlink_title="ventro-lateral process">VL</abbrev>
|
||||
with a small medial extra aberrant process)
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="114AC2E80351BBE1725F825C8FF00B41">
|
||||
<td id="4516B67EC279AFB4E08CEFD61348317D" colspan="1" rowspan="1">Legs with one tarsal spur</td>
|
||||
<td id="B15AC80E0EFCD62882D773533540120B" colspan="1" rowspan="1">1–18 (left leg 18 *)</td>
|
||||
<td id="F0BA1FF0B0A722DD7F5591471E331AB5" colspan="1" rowspan="1">1–20 (right leg 19)</td>
|
||||
<td id="65078BCA660815E13E80FF4C8AD9A440" colspan="1" rowspan="1">1–20</td>
|
||||
<td id="B9414A287B96F1A2411E55C00701D092" colspan="1" rowspan="1">1–20 (several mid body legs *)</td>
|
||||
<td id="5DC36966BB4AFD4CB6BC858977A95088" colspan="1" rowspan="1">1–20</td>
|
||||
<td id="AB897FBD866C0FA0E4B79B96EB8B544B" colspan="1" rowspan="1">1–20 (right leg 19)</td>
|
||||
</tr>
|
||||
</tbody>
|
||||
</table>
|
||||
</paragraph>
|
||||
<paragraph id="A850E0330F88E09710F0F1A5008B69F0">
|
||||
Spiracles positioned in segments 3, 5, 8, 10, 12, 14, 16, 18 and 20, triangular in form and tri-valved (Fig.
|
||||
<figureCitation id="2361F18BA7AD83397559C6057245DF2A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A cephalic plate and right antenna, dorsal view B forcipular segment, ventral view C tooth-plates, ventral view D left spiracle on segment 8. Scale bars: 0.5 mm (C, D); 2 mm (A, B)." figureDoi="10.3897/zookeys.1215.129410.figure3" httpUri="https://binary.pensoft.net/fig/1155664">3 D</figureCitation>
|
||||
). Tergite surface with shallow, small, and more dispersed puncta compared to the cephalic plate (Fig.
|
||||
<figureCitation id="AB91F65F438DBB689E25CAE73DB48747" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A tergites 3–5 B sternites 10 and 11 C ultimate leg-bearing segment, ventral view D left coxopleural process, lateral view E ultimate leg, right lateral view. Scale bars: 1 mm (A, B, D); 2 mm (C, E)." figureDoi="10.3897/zookeys.1215.129410.figure4" httpUri="https://binary.pensoft.net/fig/1155665">4 A</figureCitation>
|
||||
). Paramedian sutures of tergites faint and variable; in
|
||||
<typeStatus id="80B6DD039F1606AF86D018BA53D9F07D" type="paralectotype">paralectotype</typeStatus>
|
||||
sutures on
|
||||
<collectionCode id="F4FEA93961AF5EB6C95ED089E73054C6" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
1 and
|
||||
<collectionCode id="38F5C6DBE5DFBC0081C402C315783C0D" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
21 absent,
|
||||
<collectionCode id="A159913A3054F7B9672EA77945C9A020" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
2 incomplete posteriorly,
|
||||
<collectionCode id="E6E9BB818D5D1AAD9D7C42E144031F0E" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
3,
|
||||
<collectionCode id="B736D6CCD69352B8C096EECA0A3B9F4A" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
5, and
|
||||
<collectionCode id="130B89E93603E29B8B886959540579D4" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
7 incomplete posteriorly and anteriorly,
|
||||
<collectionCode id="29FC6A0FF53190E25775D3E7A1ABF27E" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
4,
|
||||
<collectionCode id="031E9FB38389D14B3B285F51D12A53F1" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
6, and TT 8–20 complete (see Table
|
||||
<tableCitation id="8E68979139FE11475BF284614FB6F39A" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparison between the type series of Scolopendra spinosissima Kraepelin, 1903, its voucher, and S. paradoxa Doménech, 2018 holotype. A = Absent; C = Complete; I = Incomplete; D = Distal; P = Proximal; PR = Partially retracted; R = Retracted. * = Appendix damaged. N / A = Not applicable." httpUri="http://table.plazi.org/id/BC7916AA261A079E149D4E809E484642" tableUuid="BC7916AA261A079E149D4E809E484642">1</tableCitation>
|
||||
for
|
||||
<typeStatus id="CB7AAC4D5F14F59BA2AEC5411C675323" type="paralectotype">paralectotypes</typeStatus>
|
||||
). Complete margination starting on
|
||||
<collectionCode id="6EBB07D34D73025A131EA5223B5FF03A" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
10 (on
|
||||
<collectionCode id="850EC84B584278990CA0510A802AC47C" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
<quantity id="B60043712ED6546BDEC3DCE6F0075593" metricMagnitude="-1" metricUnit="m" metricValue="3.048" unit="in" value="12.0">12 in</quantity>
|
||||
<typeStatus id="3B52D0CADB5E29DA985C92AACDB44A8F" type="paralectotype">paralectotype</typeStatus>
|
||||
1). Tergite of ultimate leg-bearing segment with disperse and non-deep puncta, without depression or sutures; posterior margin rounded. Ratio of width: length of tergite of ultimate leg-bearing segment 1.14: 1. Sternite surfaces essentially smooth, with dispersed, small, shallow puncta. Paramedian sutures in S 1 and S 21 absent; in SS 2–20 incomplete and confined to proximal 10–25 % and distal 5–10 % of sternite length (see Table
|
||||
<tableCitation id="28E2EDB2488421AFF51DC623CAD72B27" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparison between the type series of Scolopendra spinosissima Kraepelin, 1903, its voucher, and S. paradoxa Doménech, 2018 holotype. A = Absent; C = Complete; I = Incomplete; D = Distal; P = Proximal; PR = Partially retracted; R = Retracted. * = Appendix damaged. N / A = Not applicable." httpUri="http://table.plazi.org/id/BC7916AA261A079E149D4E809E484642" tableUuid="BC7916AA261A079E149D4E809E484642">1</tableCitation>
|
||||
for
|
||||
<typeStatus id="39277CFA38356E93D912874D4702AF01" type="paralectotype">paralectotypes</typeStatus>
|
||||
; Fig.
|
||||
<figureCitation id="BA10815672BDD5EF11DC9AF19957B9B1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A tergites 3–5 B sternites 10 and 11 C ultimate leg-bearing segment, ventral view D left coxopleural process, lateral view E ultimate leg, right lateral view. Scale bars: 1 mm (A, B, D); 2 mm (C, E)." figureDoi="10.3897/zookeys.1215.129410.figure4" httpUri="https://binary.pensoft.net/fig/1155665">4 B</figureCitation>
|
||||
). Space between sutures sometimes weakly depressed. Sternite of ultimate leg-bearing segment with sides converging posteriorly (Fig.
|
||||
<figureCitation id="129125B87FFA21C119C0623F7911A2D9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A tergites 3–5 B sternites 10 and 11 C ultimate leg-bearing segment, ventral view D left coxopleural process, lateral view E ultimate leg, right lateral view. Scale bars: 1 mm (A, B, D); 2 mm (C, E)." figureDoi="10.3897/zookeys.1215.129410.figure4" httpUri="https://binary.pensoft.net/fig/1155665">4 C</figureCitation>
|
||||
); surface without depressions or sutures.
|
||||
</paragraph>
|
||||
<caption id="CECD8B50E73640FE952A8B21223C2C00" ID-DOI="10.3897/zookeys.1215.129410.figure4" ID-arpha="292B0AA3-EC7F-5D81-8883-BD257054539A" httpUri="https://binary.pensoft.net/fig/1155665" startId="F4">
|
||||
<paragraph id="471A7C87D266F2A8EBE6ABFC92C72286">
|
||||
<label id="81DD6F3352CA93C5DB3552550E98F50E">Figure 4.</label>
|
||||
</paragraph>
|
||||
<paragraph id="41232D258E980EAFDE6BE608B0F2BA20">
|
||||
<taxonomicName id="EFDF203B37451A5490808A924861F355" authority="Kraepelin, 1903" authorityName="sensu Kraepelin" authorityYear="1903" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="spinosissima">
|
||||
<emphasis id="A7EA86EE5E46CBB6D4F0FC51F4654FB1" italics="true">Scolopendra spinosissima</emphasis>
|
||||
<bibRefCitation id="DFBF50E168A0174BA4676AFF14DECD01" author="Kraepelin K" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pagination="1-276" refId="B17" refString="Kraepelin K (1903) Revision der Scolopendriden. Mitteilungen aus dem Naturhistorischen Museum in Hamburg 20: 1 - 276." title="Revision der Scolopendriden." volume="20" year="1903">Kraepelin, 1903</bibRefCitation>
|
||||
</taxonomicName>
|
||||
; lectotype (
|
||||
<abbrev id="ABBRID0EF3AG" xlink_title="Zoological Museum of Hamburg">ZMH</abbrev>
|
||||
- A 0000633)
|
||||
<emphasis id="575679CD34CB754DAF32013BA54C094E" bold="true">A</emphasis>
|
||||
tergites 3–5
|
||||
<emphasis id="F71C6F558BB1EDFC651E28A7522D75DF" bold="true">B</emphasis>
|
||||
sternites 10 and 11
|
||||
<emphasis id="51F29B675D162AFB3D53D25DAC1FF770" bold="true">C</emphasis>
|
||||
ultimate leg-bearing segment, ventral view
|
||||
<emphasis id="4D677F365547538D5E2E720B980BB176" bold="true">D</emphasis>
|
||||
left coxopleural process, lateral view
|
||||
<emphasis id="AE8C034947A68BE65EB755CEC876072D" bold="true">E</emphasis>
|
||||
ultimate leg, right lateral view. Scale bars: 1 mm (
|
||||
<emphasis id="E592C24F911401F9C36BAC885D2B765B" bold="true">A, B, D</emphasis>
|
||||
); 2 mm (
|
||||
<emphasis id="FC3C61C60528D99BBD3C6406DC2E5230" bold="true">C, E</emphasis>
|
||||
).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="39AFFC7BB83B31FA11ADE57DAE7CBE1A">
|
||||
Coxopleuron with numerous coxal pores; reaching but excluding spines of the coxopleural process, not extending beyond
|
||||
<collectionCode id="E046A2D1DCA22F0AC9DD54ADB3E97462" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
21 posterior margin of
|
||||
<collectionCode id="D403969115F773E3035026FEDBF1F844" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
21. Free edge on coxopleuron moderately long, with straight dorsal and ventral margins. Posterodorsal margin of coxopleuron not inflected into dorsal margin of coxopleural process, forming both margins at ~ 120 ° angle (Figs
|
||||
<figureCitation id="A3A2A2E847226AB1848BB015818C8E7E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A tergites 3–5 B sternites 10 and 11 C ultimate leg-bearing segment, ventral view D left coxopleural process, lateral view E ultimate leg, right lateral view. Scale bars: 1 mm (A, B, D); 2 mm (C, E)." figureDoi="10.3897/zookeys.1215.129410.figure4" httpUri="https://binary.pensoft.net/fig/1155665">4 C, D</figureCitation>
|
||||
,
|
||||
<figureCitation id="ABF89ADE5D50041A4E5F9890CCF809AC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) B, D, F Scolopendra paradoxa Doménech, 2018; holotype (CEUA 017 - Mr 0000) A, B right coxopleura and coxopleural processes, lateral views C, D ultimate leg prefemora, dorsal views E, F ultimate leg prefemora, ventral views. The coxopleuron and coxopleural processes are different shapes and lengths; with the different sizes, morphologies, numbers, and positions of the prefemoral spinous processes, these are the most remarkable characters differentiating these species. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1215.129410.figure5" httpUri="https://binary.pensoft.net/fig/1155666">5 A</figureCitation>
|
||||
). Coxopleural process moderately long, with isolated small pores and with two or three distal spines, two on right (one each
|
||||
<abbrev id="ABBRID0EG4AG" xlink_title="apical spine">AP</abbrev>
|
||||
and smaller dorsal
|
||||
<abbrev id="ABBRID0EK4AG" xlink_title="subapical spine">SAP</abbrev>
|
||||
) and three on left (an additional minute spine in ventral
|
||||
<abbrev id="ABBRID0EO4AG" xlink_title="subapical spine">SAP</abbrev>
|
||||
; Fig.
|
||||
<figureCitation id="7827630602F72EF8FD97EEAB61E4C21B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A tergites 3–5 B sternites 10 and 11 C ultimate leg-bearing segment, ventral view D left coxopleural process, lateral view E ultimate leg, right lateral view. Scale bars: 1 mm (A, B, D); 2 mm (C, E)." figureDoi="10.3897/zookeys.1215.129410.figure4" httpUri="https://binary.pensoft.net/fig/1155665">4 D</figureCitation>
|
||||
). Lateral or dorsal spines absent. Pore-free area extending ventrally 30 % of length from proximal part of coxopleural process to margin of sternite of the ultimate leg-bearing segment (Figs
|
||||
<figureCitation id="24371E5A9F4A43479D4CD4CC55349734" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A tergites 3–5 B sternites 10 and 11 C ultimate leg-bearing segment, ventral view D left coxopleural process, lateral view E ultimate leg, right lateral view. Scale bars: 1 mm (A, B, D); 2 mm (C, E)." figureDoi="10.3897/zookeys.1215.129410.figure4" httpUri="https://binary.pensoft.net/fig/1155665">4 C</figureCitation>
|
||||
,
|
||||
<figureCitation id="001FD4FC7A703F1D46941611864CB964" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) B, D, F Scolopendra paradoxa Doménech, 2018; holotype (CEUA 017 - Mr 0000) A, B right coxopleura and coxopleural processes, lateral views C, D ultimate leg prefemora, dorsal views E, F ultimate leg prefemora, ventral views. The coxopleuron and coxopleural processes are different shapes and lengths; with the different sizes, morphologies, numbers, and positions of the prefemoral spinous processes, these are the most remarkable characters differentiating these species. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1215.129410.figure5" httpUri="https://binary.pensoft.net/fig/1155666">5 A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="F10E0DB5F9DFF25EFC815967A7312FD5" ID-DOI="10.3897/zookeys.1215.129410.figure5" ID-arpha="0DE148F6-DA79-532C-9B57-75D33B26EC23" httpUri="https://binary.pensoft.net/fig/1155666" startId="F5">
|
||||
<paragraph id="161201682A7E0B301B6BA7332B07CD64">
|
||||
<label id="41E23AD5E40CB2804161C7FA9B31CA3E">Figure 5.</label>
|
||||
</paragraph>
|
||||
<paragraph id="CA15F9697AA1AAC07E2B7DD13C11DE65">
|
||||
<emphasis id="93A992259A3FAD0A763AA489787F20A3" bold="true">A, C, E</emphasis>
|
||||
<taxonomicName id="83D5EB5139D27428C2F97916BB0B3131" authority="Kraepelin, 1903" authorityName="sensu Kraepelin" authorityYear="1903" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="spinosissima">
|
||||
<emphasis id="79E846FCD3226C87F727D9AFACC7530E" italics="true">Scolopendra spinosissima</emphasis>
|
||||
<bibRefCitation id="2FCA5A8DFB5F05CBF6DF15963A079A23" author="Kraepelin K" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pagination="1-276" refId="B17" refString="Kraepelin K (1903) Revision der Scolopendriden. Mitteilungen aus dem Naturhistorischen Museum in Hamburg 20: 1 - 276." title="Revision der Scolopendriden." volume="20" year="1903">Kraepelin, 1903</bibRefCitation>
|
||||
</taxonomicName>
|
||||
; lectotype (
|
||||
<abbrev id="ABBRID0ET5AG" xlink_title="Zoological Museum of Hamburg">ZMH</abbrev>
|
||||
- A 0000633)
|
||||
<emphasis id="3B53D4A9EC7F5054BE60E126E5D20AF3" bold="true">B, D, F</emphasis>
|
||||
<taxonomicName id="02E5990C7394C27119BFE251807F05D6" authority="Domenech, 2018" authorityName="Domenech" authorityYear="2018" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="paradoxa">
|
||||
<emphasis id="2FDDDC10C2EBDC2F1819AF8C4F192E93" italics="true">Scolopendra paradoxa</emphasis>
|
||||
<bibRefCitation id="FD205652CE6FFA1A3049A55DBFF80C3E" DOI="10.11646/zootaxa.4483.3.1" author="Doménech C & Barbera VM & Larriba E" issue="3" journalOrPublisher="Zootaxa" pagination="401-427" refId="B10" refString="Domenech C, Barbera VM, Larriba E (2018) A phylogenetic approach to the Philippines endemic centipedes of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae), with the description of a new species. Zootaxa 4483 (3): 401 - 427. https: // doi. org / 10.11646 / zootaxa. 4483.3. 1" title="A phylogenetic approach to the Philippines endemic centipedes of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae), with the description of a new species." volume="4483" year="2018">Doménech, 2018</bibRefCitation>
|
||||
</taxonomicName>
|
||||
; holotype (CEUA 017 - Mr 0000)
|
||||
<emphasis id="3BB3E895BA2644732C499244C831C615" bold="true">A, B</emphasis>
|
||||
right coxopleura and coxopleural processes, lateral views
|
||||
<emphasis id="BDBA772A069BDC17B884158AE815C0F4" bold="true">C, D</emphasis>
|
||||
ultimate leg prefemora, dorsal views
|
||||
<emphasis id="AFA164D32E382A150E89F2E2B50FF4A3" bold="true">E, F</emphasis>
|
||||
ultimate leg prefemora, ventral views. The coxopleuron and coxopleural processes are different shapes and lengths; with the different sizes, morphologies, numbers, and positions of the prefemoral spinous processes, these are the most remarkable characters differentiating these species. Scale bars: 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="F1741E0C4AF1CCE5F35AA5CC0E74D77B">
|
||||
All legs without tibial spurs. Surface with shallow, dispersed, small puncta allocating a short sensillum each. One tarsal spur on legs 1–19 or 20, right and left legs, respectively (all
|
||||
<typeStatus id="C62C9C536F212005CFD46ABAA5261D84" type="paralectotype">paralectotypes</typeStatus>
|
||||
with spur on legs 1–20).
|
||||
<abbrev id="ABBRID0ER6AG" xlink_title="ultimate legs">
|
||||
<collectionCode id="AA9E11EB0B138483B9327F235CBD4450" country="USA" name="University of Louisville">UL</collectionCode>
|
||||
</abbrev>
|
||||
long, slender, with length ratios prefemur and femur = 1.2: 1, femur and tibia = 1.07: 1, tibia and tarsus 2 = 2: 1; tarsus 1 and tarsus 2 = 1.45: 1 (Fig.
|
||||
<figureCitation id="67A1F084BCCC6E808762ECCDBF50E10E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A tergites 3–5 B sternites 10 and 11 C ultimate leg-bearing segment, ventral view D left coxopleural process, lateral view E ultimate leg, right lateral view. Scale bars: 1 mm (A, B, D); 2 mm (C, E)." figureDoi="10.3897/zookeys.1215.129410.figure4" httpUri="https://binary.pensoft.net/fig/1155665">4 E</figureCitation>
|
||||
). Prefemora flattened dorsally, with long wider base processes located backwards at 45 ° angle with respect to the prefemur. Spines of the spinous processes slightly curved backwards. Prefemoral spinous processes formula:
|
||||
<abbrev id="ABBRID0EZ6AG" xlink_title="ventro-lateral process">VL</abbrev>
|
||||
: 1,
|
||||
<collectionCode id="A1E30E87CA0ACD0AAF358AF8B1709EB6" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
|
||||
: 2,
|
||||
<abbrev id="ABBRID0E46AG" xlink_title="ventro-median process">VM</abbrev>
|
||||
: 2, M: 1,
|
||||
<abbrev id="ABBRID0EBABG" xlink_title="dorso-median process">
|
||||
<collectionCode id="C1189A3DD2B3517037E5D6BB24206ADC" country="New Zealand" name="Dominion Museum">DM</collectionCode>
|
||||
</abbrev>
|
||||
: 2 (Figs
|
||||
<figureCitation id="F1A8F747D3B275F5B2B62FE2D643BE93" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A tergites 3–5 B sternites 10 and 11 C ultimate leg-bearing segment, ventral view D left coxopleural process, lateral view E ultimate leg, right lateral view. Scale bars: 1 mm (A, B, D); 2 mm (C, E)." figureDoi="10.3897/zookeys.1215.129410.figure4" httpUri="https://binary.pensoft.net/fig/1155665">4 E</figureCitation>
|
||||
,
|
||||
<figureCitation id="09E0D200123DA61A9EF1F05D94ADA876" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) B, D, F Scolopendra paradoxa Doménech, 2018; holotype (CEUA 017 - Mr 0000) A, B right coxopleura and coxopleural processes, lateral views C, D ultimate leg prefemora, dorsal views E, F ultimate leg prefemora, ventral views. The coxopleuron and coxopleural processes are different shapes and lengths; with the different sizes, morphologies, numbers, and positions of the prefemoral spinous processes, these are the most remarkable characters differentiating these species. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1215.129410.figure5" httpUri="https://binary.pensoft.net/fig/1155666">5 C, E</figureCitation>
|
||||
), (in
|
||||
<typeStatus id="7DC37E888CE82384F664F9CAA52071F1" type="paralectotype">paralectotype</typeStatus>
|
||||
3, proximal spine in
|
||||
<abbrev id="ABBRID0ENABG" xlink_title="ventro-median process">VM</abbrev>
|
||||
position in right prefemur is absent (preserving the prefemoral process);
|
||||
<bibRefCitation id="BA2087980637B708AC9418EB372F623A" DOI="10.11646/zootaxa.4483.3.1" author="Domenech" etAl="et al." firstAuthor="Domenech" issue="3" journalOrPublisher="Zootaxa" pagination="401-427" refId="B10" refString="Doménech C, Barbera VM, Larriba E (2018) A phylogenetic approach to the Philippines endemic centipedes of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae), with the description of a new species. Zootaxa 4483 (3): 401–427. https://doi.org/10.11646/zootaxa.4483.3.1" title="A phylogenetic approach to the Philippines endemic centipedes of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae), with the description of a new species." volume="4483" year="2018">Doménech et al. 2018</bibRefCitation>
|
||||
: fig. 4). Prefemoral corner process slightly longer and with a narrow base in respect to other prefemoral processes, ending with a single non-curved spine (Figs
|
||||
<figureCitation id="A26D2EE8645F0990C5FCA95A958AB8DC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A tergites 3–5 B sternites 10 and 11 C ultimate leg-bearing segment, ventral view D left coxopleural process, lateral view E ultimate leg, right lateral view. Scale bars: 1 mm (A, B, D); 2 mm (C, E)." figureDoi="10.3897/zookeys.1215.129410.figure4" httpUri="https://binary.pensoft.net/fig/1155665">4 E</figureCitation>
|
||||
,
|
||||
<figureCitation id="585B7E2AFE5D6D526AA79567C68D373F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) B, D, F Scolopendra paradoxa Doménech, 2018; holotype (CEUA 017 - Mr 0000) A, B right coxopleura and coxopleural processes, lateral views C, D ultimate leg prefemora, dorsal views E, F ultimate leg prefemora, ventral views. The coxopleuron and coxopleural processes are different shapes and lengths; with the different sizes, morphologies, numbers, and positions of the prefemoral spinous processes, these are the most remarkable characters differentiating these species. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1215.129410.figure5" httpUri="https://binary.pensoft.net/fig/1155666">5 C, E</figureCitation>
|
||||
). Tarsus 1 partially lost in left
|
||||
<abbrev id="ABBRID0E4ABG" xlink_title="ultimate legs">
|
||||
<collectionCode id="0114AAE492A82D06E76809C315DD87C4" country="USA" name="University of Louisville">UL</collectionCode>
|
||||
</abbrev>
|
||||
(Fig.
|
||||
<figureCitation id="75E41428BAA87924D278DB01B9A1E730" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopendra spinosissima Kraepelin, 1903; lectotype (ZMH - A 0000633) A tergites 3–5 B sternites 10 and 11 C ultimate leg-bearing segment, ventral view D left coxopleural process, lateral view E ultimate leg, right lateral view. Scale bars: 1 mm (A, B, D); 2 mm (C, E)." figureDoi="10.3897/zookeys.1215.129410.figure4" httpUri="https://binary.pensoft.net/fig/1155665">4 E</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="EBDD6ACB190AA2A45D2DAFEC3F7168D2">
|
||||
Genitalia in the
|
||||
<typeStatus id="36E702020094AFC999C83D7B3A1AB4C2" type="lectotype">lectotype</typeStatus>
|
||||
and
|
||||
<typeStatus id="AE6ABB99FD8A308E6F9EBA7A4B18A8B3" type="paralectotype">paralectotype</typeStatus>
|
||||
1 retracted. In
|
||||
<typeStatus id="04AFC2088BA4743EDB0FFF90EA1F006E" type="paralectotype">paralectotypes</typeStatus>
|
||||
2 and 3 well-developed (Fig.
|
||||
<figureCitation id="486E4E4B416CB41BF0CF586CE0DFAB6C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopendra spinosissima Kraepelin, 1903; genital apparatus A paralectotype 2, female (ZMH - A 00016059) B paralectotype 3, male (ZMH - A 00016060). Abbreviations: AV = anal valve; LA = lamina adanalis; LS = lamina subanalis; SGS I = sternite of genital segment I; SGS II = sternite of genital segment II. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1215.129410.figure6" httpUri="https://binary.pensoft.net/fig/1155667">6 A, B</figureCitation>
|
||||
, respectively), partially retracted, reaching further than the distance between posterior margin of
|
||||
<abbrev id="ABBRID0ELBBG" xlink_title="ultimate leg-bearing segment">
|
||||
<collectionCode id="423FF9AA24FA758791C0F7634CACE828">ULBS</collectionCode>
|
||||
</abbrev>
|
||||
sternite and distal part of the coxopleural process. The genital segment sternite 1 rounded, convex posteriorly, with a median suture. Tergite of the genital segment without small setae. In male
|
||||
<typeStatus id="154ADC57376BF83E253119BAF38C0C49" type="paralectotype">paralectotype</typeStatus>
|
||||
3 (Fig.
|
||||
<figureCitation id="E915A09984E5C788FB73A3FB4591B533" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopendra spinosissima Kraepelin, 1903; genital apparatus A paralectotype 2, female (ZMH - A 00016059) B paralectotype 3, male (ZMH - A 00016060). Abbreviations: AV = anal valve; LA = lamina adanalis; LS = lamina subanalis; SGS I = sternite of genital segment I; SGS II = sternite of genital segment II. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1215.129410.figure6" httpUri="https://binary.pensoft.net/fig/1155667">6 B</figureCitation>
|
||||
) genital segment 2 is small, horseshoe-shaped, with small shallow puncta; penis, gonopods, and secondary sexual characters absent.
|
||||
</paragraph>
|
||||
<caption id="DA5B4B22B2034669BFAD424D2A2F444A" ID-DOI="10.3897/zookeys.1215.129410.figure6" ID-arpha="911695A8-B10A-5205-9C2F-1E782D18E99D" httpUri="https://binary.pensoft.net/fig/1155667" startId="F6">
|
||||
<paragraph id="071C162B79D4D999BFC78D572DB8697D">
|
||||
<label id="214661ED52C61AED1D008518C50FF1DA">Figure 6.</label>
|
||||
</paragraph>
|
||||
<paragraph id="4C54450EEC842BDA24A5DF1CD5C2DA6B">
|
||||
<taxonomicName id="4CFBF0AB09AE6CE16034EAE84D12A6CB" authority="Kraepelin, 1903" authorityName="sensu Kraepelin" authorityYear="1903" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="spinosissima">
|
||||
<emphasis id="42D6F4D7E33B0EA8B03D86C9740B97C4" italics="true">Scolopendra spinosissima</emphasis>
|
||||
<bibRefCitation id="5836CD9D979182E5F93FF68D67FF378F" author="Kraepelin K" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pagination="1-276" refId="B17" refString="Kraepelin K (1903) Revision der Scolopendriden. Mitteilungen aus dem Naturhistorischen Museum in Hamburg 20: 1 - 276." title="Revision der Scolopendriden." volume="20" year="1903">Kraepelin, 1903</bibRefCitation>
|
||||
</taxonomicName>
|
||||
; genital apparatus
|
||||
<emphasis id="5CC4DF5F840AA43AF5E574D038D305B1" bold="true">A</emphasis>
|
||||
paralectotype 2, female (
|
||||
<abbrev id="ABBRID0EICBG" xlink_title="Zoological Museum of Hamburg">ZMH</abbrev>
|
||||
- A 00016059)
|
||||
<emphasis id="E86314D98374A977D0D35AC6CD9A0B87" bold="true">B</emphasis>
|
||||
paralectotype 3, male (
|
||||
<abbrev id="ABBRID0EOCBG" xlink_title="Zoological Museum of Hamburg">ZMH</abbrev>
|
||||
- A 00016060). Abbreviations:
|
||||
<abbrev id="ABBRID0ESCBG" xlink_title="anal valve">AV</abbrev>
|
||||
= anal valve;
|
||||
<abbrev id="ABBRID0EWCBG" xlink_title="lamina adanalis">LA</abbrev>
|
||||
= lamina adanalis;
|
||||
<abbrev id="ABBRID0E1CBG" xlink_title="lateral spine">LS</abbrev>
|
||||
= lamina subanalis;
|
||||
<abbrev id="ABBRID0E5CBG" xlink_title="sternite of genital segment 1">SGS I</abbrev>
|
||||
= sternite of genital segment I;
|
||||
<abbrev id="ABBRID0ECDBG" xlink_title="sternite of genital segment 2">SGS II</abbrev>
|
||||
= sternite of genital segment II. Scale bars: 0.5 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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