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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.138.38815" ID-PMC="PMC6969051" ID-Pensoft-Pub="1314-2003-138-225" ID-Pensoft-UUID="77ADF7E7928D55DEA2B11FA59D73D7DB" ID-PubMed="31988616" ModsDocID="1314-2003-138-225" checkinTime="1578758057223" checkinUser="pensoft" docAuthor="Peng, Yulan, Yang, Chenxuan &amp; Luo, Yan" docDate="2020" docId="6EF01D16146756EE920D572BF5A71C5F" docLanguage="en" docName="PhytoKeys 138: 225-232" docOrigin="PhytoKeys 138" docSource="http://dx.doi.org/10.3897/phytokeys.138.38815" docTitle="Blumea htamanthii Y. L. Peng, C. X. Yang &amp; Y. Luo 2020, sp. nov." docType="treatment" docVersion="5" id="77ADF7E7928D55DEA2B11FA59D73D7DB" lastPageNumber="225" masterDocId="77ADF7E7928D55DEA2B11FA59D73D7DB" masterDocTitle="Blumea htamanthii (Asteraceae), a new species from Myanmar" masterLastPageNumber="232" masterPageNumber="225" pageNumber="225" updateTime="1668139617434" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Blumea htamanthii (Asteraceae), a new species from Myanmar</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Peng, Yulan</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Chenxuan</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Luo, Yan</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>138</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>225</mods:start>
<mods:end>232</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.138.38815</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.138.38815</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-138-225</mods:identifier>
<mods:identifier type="Pensoft-UUID">77ADF7E7928D55DEA2B11FA59D73D7DB</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="161672051" LSID="urn:lsid:plazi:treatment:6EF01D16146756EE920D572BF5A71C5F" httpUri="http://treatment.plazi.org/id/6EF01D16146756EE920D572BF5A71C5F" lastPageNumber="225" pageId="0" pageNumber="225">
<subSubSection pageId="0" pageNumber="225" type="nomenclature">
<paragraph pageId="0" pageNumber="225">
<taxonomicName LSID="6EF01D16-1467-56EE-920D-572BF5A71C5F" authority="Y. L. Peng, C. X. Yang &amp; Y. Luo" authorityName="Y. L. Peng, C. X. Yang &amp; Y. Luo" authorityYear="2020" class="Magnoliopsida" family="Asteraceae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea htamanthii" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="htamanthii" status="sp. nov.">Blumea htamanthii Y.L.Peng, C.X.Yang &amp; Y.Luo</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="225">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Blumea htamanthii Y. L. Peng, C. X. Yang &amp; Y. Luo, sp. nov. A habit B the inner phyllary C the outer phyllary D bisexual floret E female floret F style of bisexual floret G magnified part of upper surface of the leaf. Drawings: Jian Gu based on the holotype." figureDoi="10.3897/phytokeys.138.38815.figure1" httpUri="https://binary.pensoft.net/fig/370311" pageId="0" pageNumber="225">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Blumea htamanthii Y. L. Peng, C. X. Yang &amp; Y. Luo, sp. nov. in the field and magnified leaves and receptacle A the whole plant B inflorescences C the abaxial surface of the leaf D receptacle E the upper surface of the leaf F magnified part of upper surface of the leaf G basal leaves. Photos: Y. L. Peng." figureDoi="10.3897/phytokeys.138.38815.figure2" httpUri="https://binary.pensoft.net/fig/370312" pageId="0" pageNumber="225">, 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="225" type="diagnosis">
<paragraph pageId="0" pageNumber="225">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="225">
This new species is the most similar to
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea bifolia" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="bifolia">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea bifolia</emphasis>
</taxonomicName>
(Linn.) DC. in its obovate-oblong leaves, reflexed linear phyllaries, flat, alveolate, glabrous receptacles. However, it is distinguished by its leaf blades with papillary hairs and sparse multicellular villous, abaxial purple, 1-4 capitula at the ends of the peduncles and its unribbed achenes.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="225" type="materials_examined">
<paragraph pageId="0" pageNumber="225">Type.</paragraph>
<paragraph pageId="0" pageNumber="225">
<emphasis bold="true" pageId="0" pageNumber="225">Myanmar</emphasis>
: Htamanthi Wildlife Sanctuary, Hkamti District of Sagaing Region, the cliff near the edges of the forest along the branch river of Chindwin River, elevation 127 m,
<geoCoordinate degrees="25.4948593" direction="north" orientation="latitude" precision="1" value="25.49486">25.4948593°N</geoCoordinate>
,
<geoCoordinate degrees="95.4319749" direction="east" orientation="longitude" precision="1" value="95.43198">95.4319749°E</geoCoordinate>
, May 23 2019,Y.L. Peng, C.X. Yang &amp; Y. Luo, SE02614 (Holotype CDBI!, Isotype HITBC!, RAF!).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="225" type="description">
<paragraph pageId="0" pageNumber="225">Description.</paragraph>
<paragraph pageId="0" pageNumber="225">
Annual herbs, herbaceous, 5-25 cm tall. Stems erect, occasionally procumbent, villous with multicellular hairs, leaves basal rosette or sub-basal rosette and a few cauline, petioles 0.2 to 0.3 cm long, at the base of petioles with white pilose hairs, lamina obovate or obovate-oblong, thinly papyraceous, 0.9-3.5
<normalizedToken originalValue="×">x</normalizedToken>
0.3-1.2 cm, acute at the apex, base abruptly constricted into winged petiole, margins distantly dentate, villous with multicellular hairs, both surfaces hairy, significantly discoloured, upper surface bright green, leaf blade with papillary hairs and sparse multicellular pilose, multicellular pilose on the veins are dense, lower surface purple, the base of margins serrate, apex acute; inflorescences loose panicles, 3-10 cm long, capitula terminal, rarely axillary, 1-4 heads at the ends of the peduncles, 4-6 mm in diameter, peduncles 5-25 mm long with white pilose hairs; phyllaries herbaceous, slightly longer than the florets, 10-20 mm long, phyllaries in 5 (-6)-seriate, reflexed, outer phyllaries linear, with colleters and pilose hairs, lower part of the inner phyllaries lanceolate, upper part abruptly reduced to a linear tip, the middle and upper part margin of the inner phyllaries lacerate, with sparse multicellular hairs, receptacle 0.5-1 mm in diameter, flat or slightly convex, alveolate, glabrous. Florets yellow, tubular, glabrous; those of the bisexual florets, corolla tube 3.5-4.5 mm long, with 5 ovate, papillate lobes, styles of the hermaphroditic flowers are wrapped in a slightly longer stamen tube; those of the female florets are filiform, up to 3.5 mm long, with 2 to 3 lobed, corolla tube 1-1.5 mm long. Cypselas pale brown, oblong, pubescent, not ribbed, 0.4-0.6 mm long, pilose; pappus carducous, white, 3-4 mm long.
</paragraph>
<caption doi="10.3897/phytokeys.138.38815.figure1" httpUri="https://binary.pensoft.net/fig/370311" pageId="0" pageNumber="225" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="225">
<emphasis bold="true" pageId="0" pageNumber="225">Figure 1.</emphasis>
<taxonomicName authorityName="Y. L. Peng, C. X. Yang &amp; Y. Luo" authorityYear="2020" class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea htamanthii" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="htamanthii">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea htamanthii</emphasis>
</taxonomicName>
Y.L.Peng, C.X.Yang &amp; Y.Luo, sp. nov.
<emphasis bold="true" pageId="0" pageNumber="225">A</emphasis>
habit
<emphasis bold="true" pageId="0" pageNumber="225">B</emphasis>
the inner phyllary
<emphasis bold="true" pageId="0" pageNumber="225">C</emphasis>
the outer phyllary
<emphasis bold="true" pageId="0" pageNumber="225">D</emphasis>
bisexual floret
<emphasis bold="true" pageId="0" pageNumber="225">E</emphasis>
female floret
<emphasis bold="true" pageId="0" pageNumber="225">F</emphasis>
style of bisexual floret
<emphasis bold="true" pageId="0" pageNumber="225">G</emphasis>
magnified part of upper surface of the leaf. Drawings: Jian Gu based on the holotype.
</paragraph>
</caption>
<caption doi="10.3897/phytokeys.138.38815.figure2" httpUri="https://binary.pensoft.net/fig/370312" pageId="0" pageNumber="225" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="225">
<emphasis bold="true" pageId="0" pageNumber="225">Figure 2.</emphasis>
<taxonomicName authorityName="Y. L. Peng, C. X. Yang &amp; Y. Luo" authorityYear="2020" class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea htamanthii" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="htamanthii">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea htamanthii</emphasis>
</taxonomicName>
Y.L.Peng, C.X.Yang &amp; Y.Luo, sp. nov. in the field and magnified leaves and receptacle
<emphasis bold="true" pageId="0" pageNumber="225">A</emphasis>
the whole plant
<emphasis bold="true" pageId="0" pageNumber="225">B</emphasis>
inflorescences
<emphasis bold="true" pageId="0" pageNumber="225">C</emphasis>
the abaxial surface of the leaf
<emphasis bold="true" pageId="0" pageNumber="225">D</emphasis>
receptacle
<emphasis bold="true" pageId="0" pageNumber="225">E</emphasis>
the upper surface of the leaf
<emphasis bold="true" pageId="0" pageNumber="225">F</emphasis>
magnified part of upper surface of the leaf
<emphasis bold="true" pageId="0" pageNumber="225">G</emphasis>
basal leaves. Photos: Y.L. Peng.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="225" type="etymology">
<paragraph pageId="0" pageNumber="225">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="225">
The new specific epithet
<normalizedToken originalValue="“htamanthii”">&quot;htamanthii&quot;</normalizedToken>
refers to the name of the town along the Chindwin River, Hkamti District of Sagaing, Myanmar, where the novel species was discovered.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="225" type="phenology">
<paragraph pageId="0" pageNumber="225">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="225">Flowering and fruiting April to June.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="225" type="distribution">
<paragraph pageId="0" pageNumber="225">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="225">
Myanmar. Sagaing, Htamanthi;
<taxonomicName authorityName="Y. L. Peng, C. X. Yang &amp; Y. Luo" authorityYear="2020" class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea htamanthii" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="htamanthii">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea htamanthii</emphasis>
</taxonomicName>
is only known from the type collection along the branch river of Chindwin River, growing on the steep rocks near the forest from 66-366 m altitude above mean sea level, 25.4948°-25.5152°N, 95.4319°-95.5268°E in the Htamanthii Nature Reserve.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="225" type="materials_examined">
<paragraph pageId="0" pageNumber="225">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="225">
<geoCoordinate degrees="25.4947931" direction="north" orientation="latitude" precision="1" value="25.494793">25.4947931°N</geoCoordinate>
,
<geoCoordinate degrees="95.4319147" direction="east" orientation="longitude" precision="1" value="95.431915">95.4319147°E</geoCoordinate>
, elevation 121-129 m, 23 May 2019,Y.L Peng, C.X. Yang &amp;Y. Luo SE02645, SE02694 (CDBI, HITBC, RAF);
<geoCoordinate degrees="25.5132139" direction="north" orientation="latitude" precision="1" value="25.513214">25.5132139°N</geoCoordinate>
,
<geoCoordinate degrees="95.5269449" direction="east" orientation="longitude" precision="1" value="95.52695">95.5269449°E</geoCoordinate>
, elevation 36-367 m, 26 May 2019, Y.L. Peng, C.X. Yang &amp; Y. Luo SE02730, SE02731, SE02736 (CDBI, HITBC, RAF);
<geoCoordinate degrees="25.5127053" direction="north" orientation="latitude" precision="1" value="25.512705">25.5127053°N</geoCoordinate>
,
<geoCoordinate degrees="95.5267582" direction="east" orientation="longitude" precision="1" value="95.52676">95.5267582°E</geoCoordinate>
, elevation 366 m, 27 May 2019, Y.L Peng, C.X. Yang &amp; Y. Luo, SE02769 (CDBI, HITBC, RAF);
<geoCoordinate degrees="25.5128305" direction="north" orientation="latitude" precision="1" value="25.51283">25.5128305°N</geoCoordinate>
,
<geoCoordinate degrees="95.5268144" direction="east" orientation="longitude" precision="1" value="95.52682">95.5268144°E</geoCoordinate>
, elevation 366 m, 27 May 2019, SE02770 (CDBI, HITBC, RAF),
<geoCoordinate degrees="25.5133152" direction="north" orientation="latitude" precision="1" value="25.513315">25.5133152°N</geoCoordinate>
,
<geoCoordinate degrees="95.5262927" direction="east" orientation="longitude" precision="1" value="95.52629">95.5262927°E</geoCoordinate>
, elevation 340 m, 27 May 2019, Y.L Peng, C.X. Yang &amp; Y. Luo SE02777 (CDBI, HITBC, RAF);
<geoCoordinate degrees="25.5128089" direction="north" orientation="latitude" precision="1" value="25.51281">25.5128089°N</geoCoordinate>
,
<geoCoordinate degrees="95.5266037" direction="east" orientation="longitude" precision="1" value="95.5266">95.5266037°E</geoCoordinate>
, elevation 160 m, 27 May 2019, Y.L Peng, C.X. Yang &amp; Y. Luo SE02806, SE02861(CDBI, HITBC, RAF).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="225" type="discussion">
<paragraph pageId="0" pageNumber="225">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="225">
<taxonomicName authorityName="Y. L. Peng, C. X. Yang &amp; Y. Luo" authorityYear="2020" class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea htamanthii" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="htamanthii">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea htamanthii</emphasis>
</taxonomicName>
resembles
<taxonomicName lsidName="B. bifoliata" pageId="0" pageNumber="225" rank="species" species="bifoliata">
<emphasis italics="true" pageId="0" pageNumber="225">B. bifoliata</emphasis>
</taxonomicName>
(Linn.) DC. and
<taxonomicName lsidName="B. diffusa" pageId="0" pageNumber="225" rank="species" species="diffusa">
<emphasis italics="true" pageId="0" pageNumber="225">B. diffusa</emphasis>
</taxonomicName>
R. Br. ex Benth. in its reflexed linear phyllaries, flat, alveolate, glabrous receptacle and obovate leaves.
<taxonomicName authorityName="Y. L. Peng, C. X. Yang &amp; Y. Luo" authorityYear="2020" class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea htamanthii" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="htamanthii">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea htamanthii</emphasis>
</taxonomicName>
differs from
<taxonomicName lsidName="B. bifoliata" pageId="0" pageNumber="225" rank="species" species="bifoliata">
<emphasis italics="true" pageId="0" pageNumber="225">B. bifoliata</emphasis>
</taxonomicName>
by erect stem and basal bicolour rosette leaves, abaxially purple, with short petioles, leaf blade with papillary hairs and sparse multicellular villous and 1-4 capitula at the ends of the peduncles, achenes not ribbed (vs. leaves sessile, one colour, villous with multicellular hairs and stipitate glands, solitary capitula, achenes 6-10 ribbed) (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. A list of the morphological differences between Blumea tamanthii, B. bifoliata, B. diffusa and B. bicolor." httpUri="http://table.plazi.org/id/66D871D5F9695AFBF50228BAE754D1D1" pageId="0" pageNumber="225" tableUuid="66D871D5F9695AFBF50228BAE754D1D1">1</tableCitation>
).
<taxonomicName authorityName="Y. L. Peng, C. X. Yang &amp; Y. Luo" authorityYear="2020" class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea htamanthii" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="htamanthii">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea htamanthii</emphasis>
</taxonomicName>
differs from
<taxonomicName lsidName="B. diffusa" pageId="0" pageNumber="225" rank="species" species="diffusa">
<emphasis italics="true" pageId="0" pageNumber="225">B. diffusa</emphasis>
</taxonomicName>
in erect stems and leaves with short petioles, cauline leaf base not amplexicaulous, and 1-4 capitula at the ends of the peduncles (vs. stems procumbent, leaves sessile, one colour, cauline amplexicaulous, solitary capitula) (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. A list of the morphological differences between Blumea tamanthii, B. bifoliata, B. diffusa and B. bicolor." httpUri="http://table.plazi.org/id/66D871D5F9695AFBF50228BAE754D1D1" pageId="0" pageNumber="225" tableUuid="66D871D5F9695AFBF50228BAE754D1D1">1</tableCitation>
). In addition,
<taxonomicName lsidName="B. bicolor" pageId="0" pageNumber="225" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="225">B. bicolor</emphasis>
</taxonomicName>
is endemic in the Philippines with abaxially purple leaves (
<bibRefCitation author="Merrill, ED" journalOrPublisher="The Philippine Journal of Sciences (Section C, Botany)" pageId="0" pageNumber="225" pagination="259 - 357" refId="B6" refString="Merrill, ED, 1912. New or Noteworthy Philippine Plants, IX. The Philippine Journal of Sciences (Section C, Botany) 7: 259 - 357" title="New or Noteworthy Philippine Plants, IX." volume="7" year="1912">Merrill 1912</bibRefCitation>
,
<bibRefCitation author="Randeria, AJ" journalOrPublisher="Blumea" pageId="0" pageNumber="225" pagination="176 - 317" refId="B10" refString="Randeria, AJ, 1960. The Compositae genus Blumea: A taxonomic revision. Blumea 10: 176 - 317" title="The Compositae genus Blumea: A taxonomic revision." volume="10" year="1960">Randeria 1960</bibRefCitation>
). However, it is a tall erect herb with leaves aggregated in the middle portion of the stem, leave blade oblong elliptic, 5.5-23.5
<normalizedToken originalValue="×">x</normalizedToken>
1.3-8.6 cm and achenes ribbed. Its morphological traits are significantly different from those of
<taxonomicName lsidName="B. htamanthii" pageId="0" pageNumber="225" rank="species" species="htamanthii">
<emphasis italics="true" pageId="0" pageNumber="225">B. htamanthii</emphasis>
</taxonomicName>
(Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. A list of the morphological differences between Blumea tamanthii, B. bifoliata, B. diffusa and B. bicolor." httpUri="http://table.plazi.org/id/66D871D5F9695AFBF50228BAE754D1D1" pageId="0" pageNumber="225" tableUuid="66D871D5F9695AFBF50228BAE754D1D1">1</tableCitation>
).
</paragraph>
<caption ID-Table-UUID="66D871D5F9695AFBF50228BAE754D1D1" httpUri="http://table.plazi.org/id/66D871D5F9695AFBF50228BAE754D1D1" pageId="0" pageNumber="225" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="225">
<emphasis bold="true" pageId="0" pageNumber="225">Table 1.</emphasis>
A list of the morphological differences between
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea tamanthii" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="tamanthii">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea tamanthii</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="B. bifoliata" pageId="0" pageNumber="225" rank="species" species="bifoliata">
<emphasis italics="true" pageId="0" pageNumber="225">B. bifoliata</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="B. diffusa" pageId="0" pageNumber="225" rank="species" species="diffusa">
<emphasis italics="true" pageId="0" pageNumber="225">B. diffusa</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="B. bicolor" pageId="0" pageNumber="225" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="225">B. bicolor</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="225">
<table pageId="0" pageNumber="225">
<tr pageId="0" pageNumber="225">
<th colspan="1" pageId="0" pageNumber="225" rowspan="1">Characters</th>
<th colspan="1" pageId="0" pageNumber="225" rowspan="1">
<taxonomicName authorityName="Y. L. Peng, C. X. Yang &amp; Y. Luo" authorityYear="2020" class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea htamanthii" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="htamanthii">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea htamanthii</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="225" rowspan="1">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea bifoliata" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="bifoliata">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea bifoliata</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="225" rowspan="1">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea diffusa" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="diffusa">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea diffusa</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="225" rowspan="1">
<taxonomicName authorityName="Merr" authorityYear="1912" class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea bicolor" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea bicolor</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Leaf arrangement patterns</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Basal rosette or sub-basal rosette and a few cauline, the cauline ones are all alternate</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Mostly cauline, the uppermost pair are subopposite</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Basal rosette or sub-basal rosette and a few cauline; the cauline ones are all alternate and amplexicaulous</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">The lower part of the stem naked, leafless; the leaves mostly aggregated in the middle portion of the stem</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Leaf morphology</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
Petioles 0.2 to 0.3 cm long, lower surface purplish, apex acute, leaf blade obovate or obovate-oblong, villous with papillary hair and sparsely multicellular villous, 0.9-3.5
<normalizedToken originalValue="×">x</normalizedToken>
0.3-1.2 cm
</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
Sessile, both surfaces green, the apex acute or apiculate, leaf blade oblong or ovate, villous with multicellular hairs and stipitate glands, radical leaves 0.7-3
<normalizedToken originalValue="×">x</normalizedToken>
0.4-1.5 cm
</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
Sessile, both surfaces green, the apex acute to apiculate, obovate or rarely oblanceolate, pilose with colleters and multicellular hairs, 2-6 cm
<normalizedToken originalValue="×">x</normalizedToken>
1.0-2.5 cm wide
</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
Lower surface purplish, apex sharply acuminate, blade oblong elliptic, sparsely pilose with multicellular hairs, 5.5-23.5
<normalizedToken originalValue="×">x</normalizedToken>
1.3-8.6 cm
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Stem</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Erect, occasionally procumbent, usually unbranched, pilose with long, white hairs</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Erect, branched from the base, ascending or rarely procumbent</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Procumbent, stems branched from the base, pilose with long, white hairs</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Erect, generally unbranched, puberulous</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Capitula</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">1-4, colleters and pilose on the outer phyllaries, the middle and upper part margin of the inner phyllaries lacerate</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">1, glands on the phyllaries</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">1, pilose on the phyllaries</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Several formed a lax, terminal panicle, pubescent on the phyllaries</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Florets</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Glabrous</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Sparsely pubescent on the lobes</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Glabrous</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Bisexual florets pubescent, female florets glabrous</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Cypselas</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Pilose, not ribbed</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Pilose, 6-10-ribbed</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Sparsely pilose, 10-ribbed</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Ribbed, pubescent</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="225" type="key to blumea species in myanmar (including the closely related species b. bicolor in the philippines and b. diffusa in australia)">
<paragraph pageId="0" pageNumber="225">
Key to
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea</emphasis>
</taxonomicName>
species in Myanmar (including the closely related species
<taxonomicName lsidName="B. bicolor" pageId="0" pageNumber="225" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="225">B. bicolor</emphasis>
</taxonomicName>
in the Philippines and
<taxonomicName lsidName="B. diffusa" pageId="0" pageNumber="225" rank="species" species="diffusa">
<emphasis italics="true" pageId="0" pageNumber="225">B. diffusa</emphasis>
</taxonomicName>
in Australia)
</paragraph>
<paragraph pageId="0" pageNumber="225">
<table pageId="0" pageNumber="225">
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">1a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Plants densely white-woolly</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">2</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">2a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Outer phyllaries oblong-lanceolate, acute</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea hieraciifolia" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="hieraciifolia">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea hieraciifolia</emphasis>
</taxonomicName>
(Don) DC.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">2b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Outer phyllaries linear and tapering</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">3a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Capitula in large lax panicles; pappus red; corolla lobes of bisexual florets glabrous</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea densiflora" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="densiflora">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea densiflora</emphasis>
</taxonomicName>
DC.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">3b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Capitula in compact, spiciform panicles; pappus white; corolla lobes of bisexual florets hairy</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea lacera" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="lacera">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea lacera</emphasis>
</taxonomicName>
(Burm. f.) DC.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">1b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Plants glabrous or variously pubescent</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">4</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">4a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Phyllaries at least the outer phyllaries, oblong-ovate to oblong-lanceolate</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">5</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">5a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Climber; receptacle densely pubescent; corolla lobes of female florets with multicellular hairs</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea riparia" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="riparia">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea riparia</emphasis>
</taxonomicName>
(Blume) DC.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">5b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Erect, receptacle fimbrillate or rarely pilose; corolla lobes of female florets glabrous</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">6</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">6a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Receptacle fimbrillate or rarely pilose glabrous</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName authorityName="Druce" authorityYear="1917" baseAuthorityName="Roxb." class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea lanceolaria" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="lanceolaria">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea lanceolaria</emphasis>
</taxonomicName>
(Roxb.) Druce
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">6b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Receptacle pilose</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">7</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">7a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Capitula in narrow panicles</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea hirsuta" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="hirsuta">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea hirsuta</emphasis>
</taxonomicName>
(Less.) M. R. Almeid
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">7b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Capitula in large, spreading panicles</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName authorityName="Hand - Mazz" authorityYear="1936" baseAuthorityName="Roxb." class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea repanda" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="repanda">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea repanda</emphasis>
</taxonomicName>
(Roxb.) Hand.-Mazz
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">4b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Phyllaries all linear or linear-lanceolate</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">8</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">8a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Receptacle fimibrillate</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea aromatica" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="aromatica">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea aromatica</emphasis>
</taxonomicName>
DC.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">8b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Receptacle glabrous or pilose.</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">9</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">9a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Leaves purplish on abaxial surface</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">10</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">10a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Height 20-100 cm, the lower part of the stem naked, leafless, mostly aggregated in the middle portion of the stem</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName authorityName="Merr" authorityYear="1912" class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea bicolor" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea bicolor</emphasis>
</taxonomicName>
Merr
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">10b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Height 5-25 cm, leaves basal rosette or sub-basal rosette and a few cauline</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName authorityName="Y. L. Peng, C. X. Yang &amp; Y. Luo" authorityYear="2020" class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea htamanthii" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="htamanthii">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea htamanthii</emphasis>
</taxonomicName>
Y.L.Peng, C.X.Yang &amp; Y.Luo, sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">9b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Leaves not purplish on abaxial surface</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">11</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">11a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Pappus reddish</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea balsamifera" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="balsamifera">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea balsamifera</emphasis>
</taxonomicName>
(L.) DC.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">11b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Pappus white</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">12</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">12a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Capitula solitary, axillary and terminal</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">13</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">13a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Diffuse herbs; leaves mostly radical, the cauline ones all alternate</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea diffusa" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="diffusa">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea diffusa</emphasis>
</taxonomicName>
R. Br. ex Benth
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">13b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Erect herbs; leaves mostly cauline, the uppermost pair subopposite</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea bifoliata" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="bifoliata">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea bifoliata</emphasis>
</taxonomicName>
(L.) DC.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">12b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Capitula interruptedly spiciform paniculate or loose or dense paniculate</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">14</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">14a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Inflorescence an interrupted spiciform panicle</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName authorityName="Kurz" authorityYear="1877" baseAuthorityName="Roxb." class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea fistulosa" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="fistulosa">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea fistulosa</emphasis>
</taxonomicName>
(Roxb.) Kurz
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">14b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Inflorescence a loose or dense paniculate</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">15</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">15a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Leaves spinous-toothed, stems procumbent</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea oxyodonta" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="oxyodonta">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea oxyodonta</emphasis>
</taxonomicName>
DC.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">15b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Leaves not spinous-toothed, stems erect</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">16</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">16a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Receptacle minutely pilose</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">17</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">17a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Leaves not lyrately lobed</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName authorityName="Franch" authorityYear="1896" class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea adenophora" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="adenophora">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea adenophora</emphasis>
</taxonomicName>
Franch
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">17b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Leaves lyrately lobed</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea sinulata" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="sinulata">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea sinulata</emphasis>
</taxonomicName>
(Lour.) Merr
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">16b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Receptacle glabrous</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">18</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">18a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Achenes ribbed</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">19</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">19a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Plants more or less glabrate</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea virens" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="virens">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea virens</emphasis>
</taxonomicName>
Wall. ex DC.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">19b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Plants pubescent or variously glandular</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea napifolia" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="napifolia">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea napifolia</emphasis>
</taxonomicName>
DC.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">18b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Achenes subangulate to terete</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">20</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">20a</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Leaves usually not lobed; corollas purple</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea axillaris" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="axillaris">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea axillaris</emphasis>
</taxonomicName>
(Lam.) DC.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="225">
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">20b</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">Leaves often lyrately lobed; corollas yellow</td>
<td colspan="1" pageId="0" pageNumber="225" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="225">
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Blumea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blumea lacera" order="Asterales" pageId="0" pageNumber="225" phylum="Tracheophyta" rank="species" species="lacera">
<emphasis italics="true" pageId="0" pageNumber="225">Blumea lacera</emphasis>
</taxonomicName>
(Burm. f.) DC.
</emphasis>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>

452
data/6E/F0/27/6EF0271060925773AB4626B8152E3F55.xml Normal file
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@ -0,0 +1,452 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1098.73223" ID-Pensoft-Pub="1313-2970-1098-1" ID-Pensoft-UUID="B22184E75B1A59C9B985C0DD9F092196" ID-ZooBank="B4F4033F296E43CCBE54B9413BC19268" LSID="urn:lsid:zoobank.org:act:C050A383-5A8A-4437-A01A-8AC5746E3F70" ModsDocID="1313-2970-1098-1" checkinTime="1651618226429" checkinUser="pensoft" docAuthor="Rakotonirina, Jean Claude &amp; Fisher, Brian L." docDate="2022" docId="6EF0271060925773AB4626B8152E3F55" docLanguage="en" docName="ZooKeys 1098: 1-180" docOrigin="ZooKeys 1098" docPubDate="2022-05-03" docSource="http://dx.doi.org/10.3897/zookeys.1098.73223" docTitle="Camponotus tapia Rakotonirina &amp; Fisher 2022, sp. nov." docType="treatment" docUuid="C050A383-5A8A-4437-A01A-8AC5746E3F70" docUuidSource="ZooBank" docVersion="1" id="B22184E75B1A59C9B985C0DD9F092196" lastPageNumber="1" masterDocId="B22184E75B1A59C9B985C0DD9F092196" masterDocTitle="Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology" masterLastPageNumber="180" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1651618226429" updateUser="pensoft">
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<mods:title>Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rakotonirina, Jean Claude</mods:namePart>
<mods:affiliation>Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar &amp; Departement d'Entomologie-Culture, Elevage, Sante; Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar</mods:affiliation>
<mods:nameIdentifier type="email">jcrakoto25@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Brian L.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
<mods:affiliation>Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:date>2022</mods:date>
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<mods:number>2022-05-03</mods:number>
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<mods:number>1098</mods:number>
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<treatment LSID="urn:lsid:zoobank.org:act:C050A383-5A8A-4437-A01A-8AC5746E3F70" httpUri="http://treatment.plazi.org/id/6EF0271060925773AB4626B8152E3F55" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/C050A383-5A8A-4437-A01A-8AC5746E3F70" authority="Rakotonirina &amp; Fisher, 2022" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus tapia" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tapia" status="sp. nov.">Camponotus tapia</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Head in full-face view A C. mahafaly (CASENT 0115287) B C. tapia (CASENT 0493939) C C. quadrimaculatus (CASENT 0096044)." figureDoi="10.3897/zookeys.1098.73223.figure15" httpUri="https://binary.pensoft.net/fig/678074" pageId="0" pageNumber="1">Figs 15B</figureCitation>
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Head in full-face view A C. tapia (CASENT 0217310) B C. aro (CASENT 0489917)." figureDoi="10.3897/zookeys.1098.73223.figure25" httpUri="https://binary.pensoft.net/fig/678084" pageId="0" pageNumber="1">, 25A</figureCitation>
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Head in lateral view A C. strangulatus (CASENT 0136581) B C. tapia (CASENT 0493939)." figureDoi="10.3897/zookeys.1098.73223.figure29" httpUri="https://binary.pensoft.net/fig/678088" pageId="0" pageNumber="1">, 29B</figureCitation>
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Mesosoma in lateral view A C. atimo (CASENT 0454042) B C. tapia (CASENT 0217310)." figureDoi="10.3897/zookeys.1098.73223.figure30" httpUri="https://binary.pensoft.net/fig/678089" pageId="0" pageNumber="1">, 30B</figureCitation>
<figureCitation captionStart="Figure 75" captionStartId="F75" captionText="Figure 75. Camponotus tapia A lateral view B head in full-face view C dorsal view of minor worker CASENT 0217310 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure75" httpUri="https://binary.pensoft.net/fig/678134" pageId="0" pageNumber="1">, 75</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="holotype worker">
<paragraph pageId="0" pageNumber="1">Holotype worker.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Madagascar</emphasis>
: Province
<emphasis bold="true" pageId="0" pageNumber="1">Fianarantsoa</emphasis>
:
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="d’Atsirakambiaty">d'Atsirakambiaty</normalizedToken>
, 7.6 km 285° WNW Itremo,
<geoCoordinate degrees="20.59333" direction="south" orientation="latitude" precision="1" value="-20.59333">-20.59333</geoCoordinate>
,
<geoCoordinate degrees="46.56333" direction="east" orientation="longitude" precision="1" value="46.56333">46.56333</geoCoordinate>
, 1550 m, grassland, under stone, 22-26 Jan 2003 (Fisher, Griswold et al.) collection code: BLF07214, specimen code: CASENT0493939 (CAS).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="paratypes">
<paragraph pageId="0" pageNumber="1">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="1">5 minor workers and 1 major worker with same data as holotype but respectively with specimen codes: CASENT0826114, CASENT0826115, CASENT0493938, CASENT0837568, CASENT0837569, CASENT0493942 (NHMUK, MHNG, MSNG, PBZT, CAS).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Andohony IV Non Protected Area, A. Ravelomanana, Beapombo I Non Protected Area, Beapombo II Non Protected Area, Navoatra V Non Protected Area" country="Madagascar" county="Non Protected Area" elevation="1526" latitude="-18.97667" location="Non Protected Area" longLatPrecision="1" longitude="47.11889" municipality="Non Protected Area" specimenCount="1" stateProvince="Antananarivo">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
:
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
</emphasis>
:
<collectorName>
Andohony IV
<collectingCounty>Non Protected Area</collectingCounty>
</collectorName>
,
<locationDeviation location="Antsirabe">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.251" unit="km" value="22.51">22.51 km</quantity>
SW Antsirabe
</locationDeviation>
,
<geoCoordinate degrees="20.06718" direction="south" orientation="latitude" precision="1" value="-20.06718">-20.06718</geoCoordinate>
,
<geoCoordinate degrees="46.99274" direction="east" orientation="longitude" precision="1" value="46.99274">46.99274</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.526" unit="m" value="1526.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.526" unit="m" value="1526.0">1526 m</elevation>
</quantity>
,
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" lsidName="Uapaca" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Uapaca</emphasis>
</taxonomicName>
woodland (
<collectorName>A. Ravelomanana</collectorName>
) (CAS);
<collectorName>
Beapombo I
<collectingMunicipality>Non Protected Area</collectingMunicipality>
</collectorName>
,
<locationDeviation location="Antsirabe">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.251" unit="km" value="22.51">22.51 km</quantity>
SW Antsirabe
</locationDeviation>
,
<geoCoordinate degrees="20.06892" direction="south" orientation="latitude" precision="1" value="-20.06892">-20.06892</geoCoordinate>
,
<geoCoordinate degrees="47.00404" direction="east" orientation="longitude" precision="1" value="47.00404">47.00404</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.663" unit="m" value="1663.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.663" unit="m" value="1663.0">1663 m</elevation>
</quantity>
, Savannah grassland (
<collectorName>A. Ravelomanana</collectorName>
) (CAS);
<collectorName>
Beapombo II
<location LSID="urn:lsid:plazi:treatment:6EF0271060925773AB4626B8152E3F55:D6F5D45746862FB20057333F845C88FC" country="Madagascar" county="Non Protected Area" latitude="-18.97667" longLatPrecision="1" longitude="47.11889" municipality="Non Protected Area" name="Non Protected Area" stateProvince="Antananarivo">Non Protected Area</location>
</collectorName>
,
<locationDeviation location="Antsirabe">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.2649999999999997" unit="km" value="22.65">22.65 km</quantity>
SW Antsirabe
</locationDeviation>
,
<geoCoordinate degrees="20.07022" direction="south" orientation="latitude" precision="1" value="-20.07022">-20.07022</geoCoordinate>
,
<geoCoordinate degrees="47.00555" direction="east" orientation="longitude" precision="1" value="47.00555">47.00555</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.689" unit="m" value="1689.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.689" unit="m" value="1689.0">1689 m</elevation>
</quantity>
, Savannah grassland (
<collectorName>A. Ravelomanana</collectorName>
) (CAS); Navoatra III
<location LSID="urn:lsid:plazi:treatment:6EF0271060925773AB4626B8152E3F55:FB4A58703E24F29CBAFA5D881EDAFC6E" country="Madagascar" county="Non Protected Area" latitude="-18.97667" longLatPrecision="1" longitude="47.11889" municipality="Non Protected Area" name="Non Protected Area" stateProvince="Antananarivo">Non Protected Area</location>
,
<locationDeviation location="Arivonimamo">
<quantity metricMagnitude="3" metricUnit="m" metricValue="7.39" unit="km" value="7.39">7.39 km</quantity>
NW Arivonimamo
</locationDeviation>
,
<geoCoordinate degrees="18.98028" direction="south" orientation="latitude" precision="1" value="-18.98028">-18.98028</geoCoordinate>
,
<geoCoordinate degrees="47.12071" direction="east" orientation="longitude" precision="1" value="47.12071">47.12071</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.321" unit="m" value="1321.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.321" unit="m" value="1321.0">1321 m</elevation>
</quantity>
,
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" lsidName="Uapaca" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Uapaca</emphasis>
</taxonomicName>
woodland (
<collectorName>A. Ravelomanana</collectorName>
) (CAS);
<collectorName>
Navoatra V
<location LSID="urn:lsid:plazi:treatment:6EF0271060925773AB4626B8152E3F55:94408C355B174E7758B55A8FEA1F1120" country="Madagascar" county="Non Protected Area" latitude="-18.97667" longLatPrecision="1" longitude="47.11889" municipality="Non Protected Area" name="Non Protected Area" stateProvince="Antananarivo">Non Protected Area</location>
</collectorName>
,
<locationDeviation location="Arivonimamo">
<quantity metricMagnitude="3" metricUnit="m" metricValue="7.76" unit="km" value="7.76">7.76 km</quantity>
NW Arivonimamo
</locationDeviation>
,
<geoCoordinate degrees="18.97667" direction="south" orientation="latitude" precision="1" value="-18.97667">-18.97667</geoCoordinate>
,
<geoCoordinate degrees="47.11889" direction="east" orientation="longitude" precision="1" value="47.11889">47.11889</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.316" unit="m" value="1316.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.316" unit="m" value="1316.0">1316 m</elevation>
</quantity>
,
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" lsidName="Uapaca" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Uapaca</emphasis>
</taxonomicName>
woodland (
<collectorName>A. Ravelomanana</collectorName>
) (CAS)
</materialsCitation>
.
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="A. Ravelomanana, Antapia IV Non Protected Area" country="Madagascar" county="Foret d'Atsirakambiaty" elevation="1550" latitude="-20.73583" location="District of Ambositra" longLatPrecision="1" longitude="47.08399" municipality="Amoron'i Mania Region" specimenCount="1" stateProvince="Fianarantsoa">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="Madagascar" name="Fianarantsoa">Fianarantsoa</collectingRegion>
</emphasis>
:
<collectingCounty>
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="d’Atsirakambiaty">d'Atsirakambiaty</normalizedToken>
</collectingCounty>
,
<locationDeviation location="Itremo">
<quantity metricMagnitude="3" metricUnit="m" metricValue="7.6" unit="km" value="7.6">7.6 km</quantity>
285° WNW Itremo
</locationDeviation>
,
<geoCoordinate degrees="20.59333" direction="south" orientation="latitude" precision="1" value="-20.59333">-20.59333</geoCoordinate>
,
<geoCoordinate degrees="46.56333" direction="east" orientation="longitude" precision="1" value="46.56333">46.56333</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.55" unit="m" value="1550.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.55" unit="m" value="1550.0">1550 m</elevation>
</quantity>
, grassland (Fisher, Griswold et al.) (CAS);
<collectingMunicipality>
<normalizedToken originalValue="Amoron’i">Amoron'i</normalizedToken>
Mania Region
</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:6EF0271060925773AB4626B8152E3F55:8E94F790BAC71ED6EF433E45C219E7E5" country="Madagascar" county="Foret d'Atsirakambiaty" latitude="-20.73583" longLatPrecision="1" longitude="47.08399" municipality="Amoron'i Mania Region" name="District of Ambositra" stateProvince="Fianarantsoa">District of Ambositra</location>
, Italaviana
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" lsidName="Uapaca" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Uapaca</emphasis>
</taxonomicName>
forest,
<locationDeviation location="Antsirabe">
<quantity metricMagnitude="4" metricUnit="m" metricValue="3.5" unit="km" value="35.0">35 km</quantity>
SE of Antsirabe
</locationDeviation>
,
<geoCoordinate degrees="20.17333" direction="south" orientation="latitude" precision="1" value="-20.17333">-20.17333</geoCoordinate>
,
<geoCoordinate degrees="47.086" direction="east" orientation="longitude" precision="55" value="47.086">47.086</geoCoordinate>
, 1359 m,
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" lsidName="Uapaca" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Uapaca</emphasis>
</taxonomicName>
forest (
<normalizedToken originalValue="Rin’Ha">Rin'Ha</normalizedToken>
, Mike) (CAS); Ampangabe III
<location LSID="urn:lsid:plazi:treatment:6EF0271060925773AB4626B8152E3F55:13D4B11B1291C7138CA0617ED20E6766" country="Madagascar" county="Foret d'Atsirakambiaty" latitude="-20.73583" longLatPrecision="1" longitude="47.08399" municipality="Amoron'i Mania Region" name="Non Protected Area" stateProvince="Fianarantsoa">Non Protected Area</location>
,
<locationDeviation location="Itremo">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.126" unit="km" value="21.26">21.26 km</quantity>
W Itremo
</locationDeviation>
,
<geoCoordinate degrees="20.6125" direction="south" orientation="latitude" precision="5" value="-20.6125">-20.6125</geoCoordinate>
,
<geoCoordinate degrees="46.60883" direction="east" orientation="longitude" precision="1" value="46.60883">46.60883</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.412" unit="m" value="1412.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.412" unit="m" value="1412.0">1412 m</elevation>
</quantity>
, savannah woodland (
<collectorName>A. Ravelomanana</collectorName>
) (CAS); Ampangabe VII
<location LSID="urn:lsid:plazi:treatment:6EF0271060925773AB4626B8152E3F55:C5BA99D65017BA84BE3C8730F0AC2539" country="Madagascar" county="Foret d'Atsirakambiaty" latitude="-20.73583" longLatPrecision="1" longitude="47.08399" municipality="Amoron'i Mania Region" name="Non Protected Area" stateProvince="Fianarantsoa">Non Protected Area</location>
,
<locationDeviation location="Itremo">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.12" unit="km" value="21.2">21.2 km</quantity>
W Itremo
</locationDeviation>
,
<geoCoordinate degrees="20.61417" direction="south" orientation="latitude" precision="1" value="-20.61417">-20.61417</geoCoordinate>
,
<geoCoordinate degrees="46.60989" direction="east" orientation="longitude" precision="1" value="46.60989">46.60989</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.42" unit="m" value="1420.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.42" unit="m" value="1420.0">1420 m</elevation>
</quantity>
, Shrubland (
<collectorName>A. Ravelomanana</collectorName>
) (CAS);
<collectorName>
Antapia IV
<location LSID="urn:lsid:plazi:treatment:6EF0271060925773AB4626B8152E3F55:0099CC0E6187301240A0C8041758B3C3" country="Madagascar" county="Foret d'Atsirakambiaty" latitude="-20.73583" longLatPrecision="1" longitude="47.08399" municipality="Amoron'i Mania Region" name="Non Protected Area" stateProvince="Fianarantsoa">Non Protected Area</location>
</collectorName>
,
<locationDeviation location="Ambositra">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.642" unit="km" value="26.42">26.42 km</quantity>
SW Ambositra
</locationDeviation>
,
<geoCoordinate degrees="20.71917" direction="south" orientation="latitude" precision="1" value="-20.71917">-20.71917</geoCoordinate>
,
<geoCoordinate degrees="47.0868" direction="east" orientation="longitude" precision="5" value="47.0868">47.0868</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.494" unit="m" value="1494.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.494" unit="m" value="1494.0">1494 m</elevation>
</quantity>
,
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" lsidName="Uapaca" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Uapaca</emphasis>
</taxonomicName>
woodland (
<collectorName>A. Ravelomanana</collectorName>
) (CAS); Antohatsahomby III
<location LSID="urn:lsid:plazi:treatment:6EF0271060925773AB4626B8152E3F55:8B92EB9E2041DA1F7B5A7C740009A319" country="Madagascar" county="Foret d'Atsirakambiaty" latitude="-20.73583" longLatPrecision="1" longitude="47.08399" municipality="Amoron'i Mania Region" name="Non Protected Area" stateProvince="Fianarantsoa">Non Protected Area</location>
,
<locationDeviation location="Itremo">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.279" unit="km" value="22.79">22.79 km</quantity>
NW Itremo
</locationDeviation>
,
<geoCoordinate degrees="20.54806" direction="south" orientation="latitude" precision="1" value="-20.54806">-20.54806</geoCoordinate>
,
<geoCoordinate degrees="46.58599" direction="east" orientation="longitude" precision="1" value="46.58599">46.58599</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.499" unit="m" value="1499.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.499" unit="m" value="1499.0">1499 m</elevation>
</quantity>
,
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" lsidName="Uapaca" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Uapaca</emphasis>
</taxonomicName>
woodland (
<collectorName>A. Ravelomanana</collectorName>
) (CAS); Mampiarika III
<location LSID="urn:lsid:plazi:treatment:6EF0271060925773AB4626B8152E3F55:E21160A6D737A2D5964087D55021106A" country="Madagascar" county="Foret d'Atsirakambiaty" latitude="-20.73583" longLatPrecision="1" longitude="47.08399" municipality="Amoron'i Mania Region" name="Non Protected Area" stateProvince="Fianarantsoa">Non Protected Area</location>
,
<locationDeviation location="Ambositra">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.893" unit="km" value="28.93">28.93 km</quantity>
SW Ambositra
</locationDeviation>
,
<geoCoordinate degrees="20.73583" direction="south" orientation="latitude" precision="1" value="-20.73583">-20.73583</geoCoordinate>
,
<geoCoordinate degrees="47.08399" direction="east" orientation="longitude" precision="1" value="47.08399">47.08399</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.4869999999999999" unit="m" value="1487.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.4869999999999999" unit="m" value="1487.0">1487 m</elevation>
</quantity>
,
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" lsidName="Uapaca" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Uapaca</emphasis>
</taxonomicName>
woodland (
<collectorName>A. Ravelomanana</collectorName>
) (CAS)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">With head in full-face view, lateral margins of head anterior to eye level parallel, lacking erect hairs; clypeus with distinct anterolateral corner; in oblique profile, four or more pairs of erect hairs arranged successively from level of anterior ocular margin towards posterior cephalic margin; in profile, junction of propodeal dorsum to declivity surface broadly angulate; petiole nodelike and not anteroposteriorly compressed.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor worker.</emphasis>
In full-face view, head sides anterior to level of eye approximately parallel, strongly converging to form a short area posterior to level of eye; eye protruding and large (EL/CS: 0.29
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01; 0.28-0.31), breaking lateral cephalic margin, level of its posterior margin located at ca. posterior 1/4 of head (PoOc/CL: 0.24
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01; 0.23-0.25); frontal carinae close to each other, their distance equal to or smaller than their smallest distance to the eye (FR/CS: 0.25
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01; 0.23-0.27); clypeus with anterolateral angle, its anteromedian margin broadly triangular or convex; mandible with six teeth, its two apical teeth distantly spaced; antennal scape relatively long (SL/CS: 1.61
<normalizedToken originalValue="±">+/-</normalizedToken>
0.08; 1.48-1.71). Promesonotum weakly convex, mesopropodeum almost flat; mesonotum flat immediately anterior to metanotal groove; metanotal groove weakly visible; propodeal dorsum anteriorly slightly convex and posteriorly flat, joining declivity at a blunt angle; propodeal declivity 1/3 length of dorsum. Petiole nodiform, its dorsal margin rounding to anterior face, posterior face higher than anterior; femur of hind leg rounded axially, without twist basally.
</paragraph>
<paragraph pageId="0" pageNumber="1">First and second gastral tergites without a pair of white spots; erect hairs on lateral margin of head absent; posterior margin of head with two erect hairs; with head in profile, four pairs of erect hairs arranged from level of anterior margin of eye to posterior cephalic margin; antennal scape covered only with appressed hairs; pronotum with a pair of erect hairs; posterodorsal angle of propodeum with 2-4 erect hairs.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major worker.</emphasis>
Differing from minor worker in the following characters: enlarged head (CS: 3.20
<normalizedToken originalValue="±">+/-</normalizedToken>
0.21; 2.84-3.38; CWb/CL: 0.99
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03; 0.94-1.01) with noticeably concave posterior margin; apical 1/3 of antennal scape surpassing posterior cephalic margin; robust mesosoma, propodeal dorsum convex immediately posterior to metanotum, propodeal dorsum &lt;2
<normalizedToken originalValue="×">x</normalizedToken>
as long as height of declivity, rounded at junction; petiolar node tapering dorsally.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution and biology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The distribution of
<taxonomicName genus="C." lsidName="C. tapia" pageId="0" pageNumber="1" rank="species" species="tapia">
<emphasis italics="true" pageId="0" pageNumber="1">C. tapia</emphasis>
</taxonomicName>
is limited to the high central plateau of Madagascar (Fig.
<figureCitation captionStart="Figure 75" captionStartId="F75" captionText="Figure 75. Camponotus tapia A lateral view B head in full-face view C dorsal view of minor worker CASENT 0217310 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure75" httpUri="https://binary.pensoft.net/fig/678134" pageId="0" pageNumber="1">75D</figureCitation>
). The species inhabits grassland, savannah woodland, shrubland, and
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" lsidName="Uapaca" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Uapaca</emphasis>
</taxonomicName>
woodland in the region. Across these areas, this species nests most often under stones and in rotting tree stumps, and forages mainly on the ground.
</paragraph>
<caption doi="10.3897/zookeys.1098.73223.figure75" httpUri="https://binary.pensoft.net/fig/678134" pageId="0" pageNumber="1" start="Figure 75" startId="F75">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 75.</emphasis>
<taxonomicName authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus tapia" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tapia">
<emphasis italics="true" pageId="0" pageNumber="1">Camponotus tapia</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
lateral view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
head in full-face view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
dorsal view of minor worker CASENT0217310
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
distribution map.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="discussion">
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="1">
See discussion under
<taxonomicName genus="C." lsidName="C. atimo" pageId="0" pageNumber="1" rank="species" species="atimo">
<emphasis italics="true" pageId="0" pageNumber="1">C. atimo</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
In the present study, the definition of
<taxonomicName genus="C." lsidName="C. tapia" pageId="0" pageNumber="1" rank="species" species="tapia">
<emphasis italics="true" pageId="0" pageNumber="1">C. tapia</emphasis>
</taxonomicName>
is based on both qualitative morphological analysis and morphometrics, which are congruent. The grouping of the samples of this species obtained from the NC-clustering is confirmed by LDA with an identification success of 100%.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The species name
<emphasis italics="true" pageId="0" pageNumber="1">tapia</emphasis>
is non-Latin singular noun used in apposition and refers to the Malagasy name for
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" lsidName="Uapaca" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Uapaca</emphasis>
</taxonomicName>
in reference to the type of habitat where the ant was most frequently found.
</paragraph>
</subSubSection>
</treatment>
</document>