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<mods:title id="F50D981C716E2BF870501FBB16E702E6">Papiri gen. nov., a new genus of huntsman spiders (Araneae: Sparassidae: Sparianthinae) from Brazil</mods:title>
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<treatment id="03988797FFE08C0DECECF89BA5EB77B4" LSID="urn:lsid:plazi:treatment:03988797FFE08C0DECECF89BA5EB77B4" httpUri="http://treatment.plazi.org/id/03988797FFE08C0DECECF89BA5EB77B4" lastPageId="5" lastPageNumber="531" pageId="1" pageNumber="527">
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<subSubSection id="C32B650AFFE08C09ECECF89BA63777AB" box="[151,343,1904,1930]" pageId="1" pageNumber="527" type="nomenclature">
<paragraph id="8B8E3681FFE08C09ECECF89BA63777AB" blockId="1.[151,343,1904,1930]" box="[151,343,1904,1930]" pageId="1" pageNumber="527">
<heading id="D0C681EDFFE08C09ECECF89BA63777AB" bold="true" box="[151,343,1904,1930]" fontSize="11" level="1" pageId="1" pageNumber="527" reason="1">
@ -64,7 +65,7 @@ species.
</emphasis>
</paragraph>
</subSubSection>
<caption id="DF4E6609FFE38C0AECECF87CA3F377D9" pageId="2" pageNumber="528" startId="2.[151,264,1943,1967]" targetBox="[166,1422,181,1919]" targetPageId="2" targetType="figure">
<caption id="DF4E6609FFE38C0AECECF87CA3F377D9" ID-DOI="http://doi.org/10.5281/zenodo.14804631" ID-Zenodo-Dep="14804631" httpUri="https://zenodo.org/record/14804631/files/figure.png" pageId="2" pageNumber="528" startId="2.[151,264,1943,1967]" targetBox="[166,1422,181,1919]" targetPageId="2" targetType="figure">
<paragraph id="8B8E3681FFE38C0AECECF87CA3F377D9" blockId="2.[151,1437,1943,2040]" pageId="2" pageNumber="528">
<emphasis id="B945EA93FFE38C0AECECF87CA65A778E" bold="true" box="[151,314,1943,1967]" pageId="2" pageNumber="528">FIGURES 1–6.</emphasis>
1, 3
@ -91,7 +92,7 @@ species.
(female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral.
</paragraph>
</caption>
<caption id="DF4E6609FFE28C0BECECF87CA30677D9" pageId="3" pageNumber="529" startId="3.[151,264,1943,1967]" targetBox="[167,1419,181,1919]" targetPageId="3" targetType="figure">
<caption id="DF4E6609FFE28C0BECECF87CA30677D9" ID-DOI="http://doi.org/10.5281/zenodo.14804635" ID-Zenodo-Dep="14804635" httpUri="https://zenodo.org/record/14804635/files/figure.png" pageId="3" pageNumber="529" startId="3.[151,264,1943,1967]" targetBox="[167,1419,181,1919]" targetPageId="3" targetType="figure">
<paragraph id="8B8E3681FFE28C0BECECF87CA30677D9" blockId="3.[151,1436,1943,2040]" pageId="3" pageNumber="529">
<emphasis id="B945EA93FFE28C0BECECF87CA627778E" bold="true" box="[151,327,1943,1967]" pageId="3" pageNumber="529">FIGURES 7–12.</emphasis>
7, 9
@ -118,7 +119,7 @@ species.
(10 male IBSP 275288, 11 female IBSP 275471). 7–10 Tarsus claws, left leg, prolateral (7 leg I, 8 leg II, 9 leg III, 10 leg IV); 11 Opisthosoma, epiandrium, epiandrous spigots, ventral; 12 Opisthosoma colulus, ventral.
</paragraph>
</caption>
<caption id="DF4E6609FFE58C0CECECFA85A41075EF" pageId="4" pageNumber="530" startId="4.[151,264,1390,1414]" targetBox="[151,1435,181,1366]" targetPageId="4" targetType="figure">
<caption id="DF4E6609FFE58C0CECECFA85A41075EF" ID-DOI="http://doi.org/10.5281/zenodo.14804637" ID-Zenodo-Dep="14804637" httpUri="https://zenodo.org/record/14804637/files/figure.png" pageId="4" pageNumber="530" startId="4.[151,264,1390,1414]" targetBox="[151,1435,181,1366]" targetPageId="4" targetType="figure">
<paragraph id="8B8E3681FFE58C0CECECFA85A41075EF" blockId="4.[151,1436,1390,1487]" pageId="4" pageNumber="530">
<emphasis id="B945EA93FFE58C0CECECFA85A63575A7" bold="true" box="[151,341,1390,1414]" pageId="4" pageNumber="530">FIGURES 13–16.</emphasis>
13–14
@ -191,13 +192,13 @@ and
<emphasis id="B945EA93FFE58C0CED69F95AA6FC76EA" box="[274,412,1713,1739]" italics="true" pageId="4" pageNumber="530">Thomasettia</emphasis>
</taxonomicName>
by the eyes arranged in two relatively straight rows or with posterior row slightly procurved (e.g.
<figureCitation id="130A2A04FFE58C0CECAFF93EA64D76CE" box="[212,301,1749,1775]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1949,1973]" captionTargetBox="[200,1384,429,1911]" captionTargetId="figure-125@7.[189,1397,417,1925]" captionTargetPageId="7" captionText="FIGURES 17–20. P. cabruca sp. nov., habitus. 17–18 Male, IBSP 47105 (17 dorsal, 18 ventral); 19–20 Female, IBSP 47125 (19 dorsal, 20 ventral). Scale lines: 2 mm." pageId="4" pageNumber="530">Figs 19</figureCitation>
<figureCitation id="130A2A04FFE58C0CECAFF93EA64D76CE" box="[212,301,1749,1775]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1949,1973]" captionTargetBox="[200,1384,429,1911]" captionTargetId="figure-125@7.[189,1397,417,1925]" captionTargetPageId="7" captionText="FIGURES 17–20. P. cabruca sp. nov., habitus. 17–18 Male, IBSP 47105 (17 dorsal, 18 ventral); 19–20 Female, IBSP 47125 (19 dorsal, 20 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804639" httpUri="https://zenodo.org/record/14804639/files/figure.png" pageId="4" pageNumber="530">Figs 19</figureCitation>
,
<figureCitation id="130A2A04FFE58C0CED47F93EA63A76CE" box="[316,346,1749,1775]" captionStart="FIGURES 32–35" captionStartId="10.[151,264,1928,1952]" captionTargetBox="[154,1424,395,1890]" captionTargetId="figure-95@10.[152,1434,381,1905]" captionTargetPageId="10" captionText="FIGURES 32–35. P. cacau sp. nov., habitus. 32–33 Male, IBSP 47114 (32 dorsal, 33 ventral); 34–35 Female, IBSP 47106 (34 dorsal, 35 ventral). Scale lines: 2 mm." pageId="4" pageNumber="530">32</figureCitation>
<figureCitation id="130A2A04FFE58C0CED47F93EA63A76CE" box="[316,346,1749,1775]" captionStart="FIGURES 32–35" captionStartId="10.[151,264,1928,1952]" captionTargetBox="[154,1424,395,1890]" captionTargetId="figure-95@10.[152,1434,381,1905]" captionTargetPageId="10" captionText="FIGURES 32–35. P. cacau sp. nov., habitus. 32–33 Male, IBSP 47114 (32 dorsal, 33 ventral); 34–35 Female, IBSP 47106 (34 dorsal, 35 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804647" httpUri="https://zenodo.org/record/14804647/files/figure.png" pageId="4" pageNumber="530">32</figureCitation>
,
<figureCitation id="130A2A04FFE58C0CED11F93EA6E976CE" box="[362,393,1749,1775]" captionStart="FIGURES 47–50" captionStartId="13.[151,264,1967,1991]" captionTargetBox="[218,1372,646,1931]" captionTargetId="figure-183@13.[203,1384,633,1944]" captionTargetPageId="13" captionText="FIGURES 47–50. P. caete sp. nov., habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm." pageId="4" pageNumber="530">49</figureCitation>
<figureCitation id="130A2A04FFE58C0CED11F93EA6E976CE" box="[362,393,1749,1775]" captionStart="FIGURES 47–50" captionStartId="13.[151,264,1967,1991]" captionTargetBox="[218,1372,646,1931]" captionTargetId="figure-183@13.[203,1384,633,1944]" captionTargetPageId="13" captionText="FIGURES 47–50. P. caete sp. nov., habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804657" httpUri="https://zenodo.org/record/14804657/files/figure.png" pageId="4" pageNumber="530">49</figureCitation>
,
<figureCitation id="130A2A04FFE58C0CEDE3F93EA6D576D1" box="[408,437,1749,1776]" captionStart="FIGURES 63–66" captionStartId="17.[151,264,1654,1678]" captionTargetBox="[166,1415,192,1616]" captionTargetId="figure-20@17.[165,1422,181,1629]" captionTargetPageId="17" captionText="FIGURES 63–66. P. tacomare sp. nov., habitus. 63–64 Male, IBSP 97195 (63 dorsal, 64 ventral); 65–66 Female, UFMG 9488 (65 dorsal, 66 ventral). Scale lines: 2 mm." pageId="4" pageNumber="530">65</figureCitation>
<figureCitation id="130A2A04FFE58C0CEDE3F93EA6D576D1" box="[408,437,1749,1776]" captionStart="FIGURES 63–66" captionStartId="17.[151,264,1654,1678]" captionTargetBox="[166,1415,192,1616]" captionTargetId="figure-20@17.[165,1422,181,1629]" captionTargetPageId="17" captionText="FIGURES 63–66. P. tacomare sp. nov., habitus. 63–64 Male, IBSP 97195 (63 dorsal, 64 ventral); 65–66 Female, UFMG 9488 (65 dorsal, 66 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804665" httpUri="https://zenodo.org/record/14804665/files/figure.png" pageId="4" pageNumber="530">65</figureCitation>
) (two recurved rows in the latter genera), from
<taxonomicName id="4C314D02FFE58C0CEF97F93EA3E176CE" authorityName="Simon" authorityYear="1898" box="[1004,1153,1749,1775]" class="Arachnida" family="Sparassidae" genus="Rhacocnemis" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="530" phylum="Arthropoda" rank="genus">
<emphasis id="B945EA93FFE58C0CEF97F93EA3E176CE" box="[1004,1153,1749,1775]" italics="true" pageId="4" pageNumber="530">Rhacocnemis</emphasis>
@ -207,29 +208,29 @@ and
<emphasis id="B945EA93FFE58C0CE8BBF93EA22976CE" box="[1216,1353,1749,1775]" italics="true" pageId="4" pageNumber="530">Thelcticopis</emphasis>
</taxonomicName>
by the presence of only two retromarginal teeth in the chelicerae (
<figureCitation id="130A2A04FFE58C0CEF46F912A41F7732" box="[829,895,1785,1811]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." pageId="4" pageNumber="530">Fig. 1</figureCitation>
<figureCitation id="130A2A04FFE58C0CEF46F912A41F7732" box="[829,895,1785,1811]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." figureDoi="http://doi.org/10.5281/zenodo.14804631" httpUri="https://zenodo.org/record/14804631/files/figure.png" pageId="4" pageNumber="530">Fig. 1</figureCitation>
) (more than four in the latter two genera), and from
<taxonomicName id="4C314D02FFE58C0CECADF8F6A6E97716" authorityName="Rheims" authorityYear="2019" box="[214,393,1821,1847]" class="Arachnida" family="Sparassidae" genus="Extraordinarius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="530" phylum="Arthropoda" rank="genus">
<emphasis id="B945EA93FFE58C0CECADF8F6A6E97716" box="[214,393,1821,1847]" italics="true" pageId="4" pageNumber="530">Extraordinarius</emphasis>
</taxonomicName>
by the male palps with a T-shaped, hood-like conductor with one end of the hood modified and curved (Hc) the other simple and rounded (e.g.
<figureCitation id="130A2A04FFE58C0CEE96F8AAA425777A" box="[749,837,1857,1883]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="4" pageNumber="530">Figs 27</figureCitation>
<figureCitation id="130A2A04FFE58C0CEE96F8AAA425777A" box="[749,837,1857,1883]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="4" pageNumber="530">Figs 27</figureCitation>
,
<figureCitation id="130A2A04FFE58C0CEF28F8AAA412777A" box="[851,882,1857,1883]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="4" pageNumber="530">42</figureCitation>
<figureCitation id="130A2A04FFE58C0CEF28F8AAA412777A" box="[851,882,1857,1883]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="4" pageNumber="530">42</figureCitation>
,
<figureCitation id="130A2A04FFE58C0CEF04F8AAA4FC777D" box="[895,924,1857,1884]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="4" pageNumber="530">57</figureCitation>
<figureCitation id="130A2A04FFE58C0CEF04F8AAA4FC777D" box="[895,924,1857,1884]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="4" pageNumber="530">57</figureCitation>
) (Z-shaped in
<taxonomicName id="4C314D02FFE58C0CE83CF8AAA39A777A" authorityName="Rheims" authorityYear="2019" box="[1095,1274,1857,1883]" class="Arachnida" family="Sparassidae" genus="Extraordinarius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="530" phylum="Arthropoda" rank="genus">
<emphasis id="B945EA93FFE58C0CE83CF8AAA39A777A" box="[1095,1274,1857,1883]" italics="true" pageId="4" pageNumber="530">Extraordinarius</emphasis>
</taxonomicName>
). Females are distinguished from those of the latter genera by the vulva with IDS divided into a proximal hyaline part (Dh) connecting to a distal sclerotized (Ds) part (e.g.
<figureCitation id="130A2A04FFE58C0CEECAF862A4677785" box="[689,775,1929,1956]" captionStart="FIGURES 21–25" captionStartId="8.[151,264,1366,1390]" captionTargetBox="[180,1417,194,1329]" captionTargetId="figure-16@8.[166,1422,181,1342]" captionTargetPageId="8" captionText="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." pageId="4" pageNumber="530">Figs 25</figureCitation>
<figureCitation id="130A2A04FFE58C0CEECAF862A4677785" box="[689,775,1929,1956]" captionStart="FIGURES 21–25" captionStartId="8.[151,264,1366,1390]" captionTargetBox="[180,1417,194,1329]" captionTargetId="figure-16@8.[166,1422,181,1342]" captionTargetPageId="8" captionText="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." figureDoi="http://doi.org/10.5281/zenodo.14804643" httpUri="https://zenodo.org/record/14804643/files/figure.png" pageId="4" pageNumber="530">Figs 25</figureCitation>
,
<figureCitation id="130A2A04FFE58C0CEF68F862A4527782" box="[787,818,1929,1955]" captionStart="FIGURES 36–40" captionStartId="11.[151,264,1815,1839]" captionTargetBox="[195,1392,576,1779]" captionTargetId="figure-193@11.[180,1408,561,1792]" captionTargetPageId="11" captionText="FIGURES 36–40. P. cacau sp. nov. 36–38 Male (IBSP 47114), left palp (36 prolateral, 37 ventral, 38 retrolateral); 39–40 Female (IBSP 275186), genitalia (39 epigyne, ventral, 40 vulva, dorsal). Scale lines: 1mm." pageId="4" pageNumber="530">40</figureCitation>
<figureCitation id="130A2A04FFE58C0CEF68F862A4527782" box="[787,818,1929,1955]" captionStart="FIGURES 36–40" captionStartId="11.[151,264,1815,1839]" captionTargetBox="[195,1392,576,1779]" captionTargetId="figure-193@11.[180,1408,561,1792]" captionTargetPageId="11" captionText="FIGURES 36–40. P. cacau sp. nov. 36–38 Male (IBSP 47114), left palp (36 prolateral, 37 ventral, 38 retrolateral); 39–40 Female (IBSP 275186), genitalia (39 epigyne, ventral, 40 vulva, dorsal). Scale lines: 1mm." figureDoi="http://doi.org/10.5281/zenodo.14804651" httpUri="https://zenodo.org/record/14804651/files/figure.png" pageId="4" pageNumber="530">40</figureCitation>
,
<figureCitation id="130A2A04FFE58C0CEF46F862A43B7785" box="[829,859,1929,1956]" captionStart="FIGURES 51–55" captionStartId="14.[151,264,1345,1369]" captionTargetBox="[169,1413,195,1307]" captionTargetId="figure-16@14.[158,1428,181,1320]" captionTargetPageId="14" captionText="FIGURES 51–55. P. caete sp. nov. 51–53 Male (IBSP 135153), left palp (51 prolateral, 52 ventral, 53 retrolateral); 54–55 Female (IBSP 234826), genitalia (54 epigyne ventral, 55 vulva dorsal). Scale lines: 1 mm." pageId="4" pageNumber="530">55</figureCitation>
<figureCitation id="130A2A04FFE58C0CEF46F862A43B7785" box="[829,859,1929,1956]" captionStart="FIGURES 51–55" captionStartId="14.[151,264,1345,1369]" captionTargetBox="[169,1413,195,1307]" captionTargetId="figure-16@14.[158,1428,181,1320]" captionTargetPageId="14" captionText="FIGURES 51–55. P. caete sp. nov. 51–53 Male (IBSP 135153), left palp (51 prolateral, 52 ventral, 53 retrolateral); 54–55 Female (IBSP 234826), genitalia (54 epigyne ventral, 55 vulva dorsal). Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14804659" httpUri="https://zenodo.org/record/14804659/files/figure.png" pageId="4" pageNumber="530">55</figureCitation>
,
<figureCitation id="130A2A04FFE58C0CEF1CF862A4E67782" box="[871,902,1929,1955]" captionStart="FIGURES 67–71" captionStartId="18.[151,264,1441,1465]" captionTargetBox="[166,1421,219,1407]" captionTargetId="figure-16@18.[152,1435,201,1418]" captionTargetPageId="18" captionText="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." pageId="4" pageNumber="530">71</figureCitation>
<figureCitation id="130A2A04FFE58C0CEF1CF862A4E67782" box="[871,902,1929,1955]" captionStart="FIGURES 67–71" captionStartId="18.[151,264,1441,1465]" captionTargetBox="[166,1421,219,1407]" captionTargetId="figure-16@18.[152,1435,201,1418]" captionTargetPageId="18" captionText="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14804669" httpUri="https://zenodo.org/record/14804669/files/figure.png" pageId="4" pageNumber="530">71</figureCitation>
).
</paragraph>
</subSubSection>
@ -244,47 +245,47 @@ Coloration pattern: prosoma brown, slightly darker along thoracic striae and lat
<taxonomicNameLabel id="A27657E8FFE48C0DEE6CFECCA5107160" box="[535,624,295,321]" pageId="5" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, without U-shaped mark) (
<figureCitation id="130A2A04FFE48C0DEFE1FECCA37A7160" box="[922,1050,295,321]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1949,1973]" captionTargetBox="[200,1384,429,1911]" captionTargetId="figure-125@7.[189,1397,417,1925]" captionTargetPageId="7" captionText="FIGURES 17–20. P. cabruca sp. nov., habitus. 17–18 Male, IBSP 47105 (17 dorsal, 18 ventral); 19–20 Female, IBSP 47125 (19 dorsal, 20 ventral). Scale lines: 2 mm." pageId="5" pageNumber="531">Figs 17–18</figureCitation>
<figureCitation id="130A2A04FFE48C0DEFE1FECCA37A7160" box="[922,1050,295,321]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1949,1973]" captionTargetBox="[200,1384,429,1911]" captionTargetId="figure-125@7.[189,1397,417,1925]" captionTargetPageId="7" captionText="FIGURES 17–20. P. cabruca sp. nov., habitus. 17–18 Male, IBSP 47105 (17 dorsal, 18 ventral); 19–20 Female, IBSP 47125 (19 dorsal, 20 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804639" httpUri="https://zenodo.org/record/14804639/files/figure.png" pageId="5" pageNumber="531">Figs 17–18</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DE85FFECCA30D7160" box="[1060,1133,295,321]" captionStart="FIGURES 32–35" captionStartId="10.[151,264,1928,1952]" captionTargetBox="[154,1424,395,1890]" captionTargetId="figure-95@10.[152,1434,381,1905]" captionTargetPageId="10" captionText="FIGURES 32–35. P. cacau sp. nov., habitus. 32–33 Male, IBSP 47114 (32 dorsal, 33 ventral); 34–35 Female, IBSP 47106 (34 dorsal, 35 ventral). Scale lines: 2 mm." pageId="5" pageNumber="531">32–33</figureCitation>
<figureCitation id="130A2A04FFE48C0DE85FFECCA30D7160" box="[1060,1133,295,321]" captionStart="FIGURES 32–35" captionStartId="10.[151,264,1928,1952]" captionTargetBox="[154,1424,395,1890]" captionTargetId="figure-95@10.[152,1434,381,1905]" captionTargetPageId="10" captionText="FIGURES 32–35. P. cacau sp. nov., habitus. 32–33 Male, IBSP 47114 (32 dorsal, 33 ventral); 34–35 Female, IBSP 47106 (34 dorsal, 35 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804647" httpUri="https://zenodo.org/record/14804647/files/figure.png" pageId="5" pageNumber="531">32–33</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DE80CFECCA3A07160" box="[1143,1216,295,321]" captionStart="FIGURES 47–50" captionStartId="13.[151,264,1967,1991]" captionTargetBox="[218,1372,646,1931]" captionTargetId="figure-183@13.[203,1384,633,1944]" captionTargetPageId="13" captionText="FIGURES 47–50. P. caete sp. nov., habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm." pageId="5" pageNumber="531">47–48</figureCitation>
<figureCitation id="130A2A04FFE48C0DE80CFECCA3A07160" box="[1143,1216,295,321]" captionStart="FIGURES 47–50" captionStartId="13.[151,264,1967,1991]" captionTargetBox="[218,1372,646,1931]" captionTargetId="figure-183@13.[203,1384,633,1944]" captionTargetPageId="13" captionText="FIGURES 47–50. P. caete sp. nov., habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804657" httpUri="https://zenodo.org/record/14804657/files/figure.png" pageId="5" pageNumber="531">47–48</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DE8B1FECCA2747160" box="[1226,1300,295,321]" captionStart="FIGURES 63–66" captionStartId="17.[151,264,1654,1678]" captionTargetBox="[166,1415,192,1616]" captionTargetId="figure-20@17.[165,1422,181,1629]" captionTargetPageId="17" captionText="FIGURES 63–66. P. tacomare sp. nov., habitus. 63–64 Male, IBSP 97195 (63 dorsal, 64 ventral); 65–66 Female, UFMG 9488 (65 dorsal, 66 ventral). Scale lines: 2 mm." pageId="5" pageNumber="531">63–64</figureCitation>
<figureCitation id="130A2A04FFE48C0DE8B1FECCA2747160" box="[1226,1300,295,321]" captionStart="FIGURES 63–66" captionStartId="17.[151,264,1654,1678]" captionTargetBox="[166,1415,192,1616]" captionTargetId="figure-20@17.[165,1422,181,1629]" captionTargetPageId="17" captionText="FIGURES 63–66. P. tacomare sp. nov., habitus. 63–64 Male, IBSP 97195 (63 dorsal, 64 ventral); 65–66 Female, UFMG 9488 (65 dorsal, 66 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804665" httpUri="https://zenodo.org/record/14804665/files/figure.png" pageId="5" pageNumber="531">63–64</figureCitation>
). Spinnerets yellowish brown, distally pale yellow. Coloration pattern in females as in males (
<figureCitation id="130A2A04FFE48C0DE86EFEA0A3F67144" box="[1045,1174,331,357]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1949,1973]" captionTargetBox="[200,1384,429,1911]" captionTargetId="figure-125@7.[189,1397,417,1925]" captionTargetPageId="7" captionText="FIGURES 17–20. P. cabruca sp. nov., habitus. 17–18 Male, IBSP 47105 (17 dorsal, 18 ventral); 19–20 Female, IBSP 47125 (19 dorsal, 20 ventral). Scale lines: 2 mm." pageId="5" pageNumber="531">Figs 19–20</figureCitation>
<figureCitation id="130A2A04FFE48C0DE86EFEA0A3F67144" box="[1045,1174,331,357]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1949,1973]" captionTargetBox="[200,1384,429,1911]" captionTargetId="figure-125@7.[189,1397,417,1925]" captionTargetPageId="7" captionText="FIGURES 17–20. P. cabruca sp. nov., habitus. 17–18 Male, IBSP 47105 (17 dorsal, 18 ventral); 19–20 Female, IBSP 47125 (19 dorsal, 20 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804639" httpUri="https://zenodo.org/record/14804639/files/figure.png" pageId="5" pageNumber="531">Figs 19–20</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DE8DBFEA1A3897144" box="[1184,1257,330,357]" captionStart="FIGURES 32–35" captionStartId="10.[151,264,1928,1952]" captionTargetBox="[154,1424,395,1890]" captionTargetId="figure-95@10.[152,1434,381,1905]" captionTargetPageId="10" captionText="FIGURES 32–35. P. cacau sp. nov., habitus. 32–33 Male, IBSP 47114 (32 dorsal, 33 ventral); 34–35 Female, IBSP 47106 (34 dorsal, 35 ventral). Scale lines: 2 mm." pageId="5" pageNumber="531">34–35</figureCitation>
<figureCitation id="130A2A04FFE48C0DE8DBFEA1A3897144" box="[1184,1257,330,357]" captionStart="FIGURES 32–35" captionStartId="10.[151,264,1928,1952]" captionTargetBox="[154,1424,395,1890]" captionTargetId="figure-95@10.[152,1434,381,1905]" captionTargetPageId="10" captionText="FIGURES 32–35. P. cacau sp. nov., habitus. 32–33 Male, IBSP 47114 (32 dorsal, 33 ventral); 34–35 Female, IBSP 47106 (34 dorsal, 35 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804647" httpUri="https://zenodo.org/record/14804647/files/figure.png" pageId="5" pageNumber="531">34–35</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DE888FEA1A25C7144" box="[1267,1340,330,357]" captionStart="FIGURES 47–50" captionStartId="13.[151,264,1967,1991]" captionTargetBox="[218,1372,646,1931]" captionTargetId="figure-183@13.[203,1384,633,1944]" captionTargetPageId="13" captionText="FIGURES 47–50. P. caete sp. nov., habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm." pageId="5" pageNumber="531">49–50</figureCitation>
<figureCitation id="130A2A04FFE48C0DE888FEA1A25C7144" box="[1267,1340,330,357]" captionStart="FIGURES 47–50" captionStartId="13.[151,264,1967,1991]" captionTargetBox="[218,1372,646,1931]" captionTargetId="figure-183@13.[203,1384,633,1944]" captionTargetPageId="13" captionText="FIGURES 47–50. P. caete sp. nov., habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804657" httpUri="https://zenodo.org/record/14804657/files/figure.png" pageId="5" pageNumber="531">49–50</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DE93DFEA1A2F17144" box="[1350,1425,330,357]" captionStart="FIGURES 63–66" captionStartId="17.[151,264,1654,1678]" captionTargetBox="[166,1415,192,1616]" captionTargetId="figure-20@17.[165,1422,181,1629]" captionTargetPageId="17" captionText="FIGURES 63–66. P. tacomare sp. nov., habitus. 63–64 Male, IBSP 97195 (63 dorsal, 64 ventral); 65–66 Female, UFMG 9488 (65 dorsal, 66 ventral). Scale lines: 2 mm." pageId="5" pageNumber="531">65–66</figureCitation>
<figureCitation id="130A2A04FFE48C0DE93DFEA1A2F17144" box="[1350,1425,330,357]" captionStart="FIGURES 63–66" captionStartId="17.[151,264,1654,1678]" captionTargetBox="[166,1415,192,1616]" captionTargetId="figure-20@17.[165,1422,181,1629]" captionTargetPageId="17" captionText="FIGURES 63–66. P. tacomare sp. nov., habitus. 63–64 Male, IBSP 97195 (63 dorsal, 64 ventral); 65–66 Female, UFMG 9488 (65 dorsal, 66 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804665" httpUri="https://zenodo.org/record/14804665/files/figure.png" pageId="5" pageNumber="531">65–66</figureCitation>
). Total length of males 7.1–13.2, of females 10.4–17.0. Dorsal shield of prosoma longer than wide. Cephalic region slightly higher than thoracic region, flattening posteriorly. Fovea conspicuous on posterior third of prosoma. Eyes arranged in two straight rows;AME larger than ALE, more separated from each other than from laterals; PME smaller than PLE, mostly equidistant (e.g.
<figureCitation id="130A2A04FFE48C0DEE5CFE30A51F71D4" box="[551,639,475,501]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1949,1973]" captionTargetBox="[200,1384,429,1911]" captionTargetId="figure-125@7.[189,1397,417,1925]" captionTargetPageId="7" captionText="FIGURES 17–20. P. cabruca sp. nov., habitus. 17–18 Male, IBSP 47105 (17 dorsal, 18 ventral); 19–20 Female, IBSP 47125 (19 dorsal, 20 ventral). Scale lines: 2 mm." pageId="5" pageNumber="531">Figs 17</figureCitation>
<figureCitation id="130A2A04FFE48C0DEE5CFE30A51F71D4" box="[551,639,475,501]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1949,1973]" captionTargetBox="[200,1384,429,1911]" captionTargetId="figure-125@7.[189,1397,417,1925]" captionTargetPageId="7" captionText="FIGURES 17–20. P. cabruca sp. nov., habitus. 17–18 Male, IBSP 47105 (17 dorsal, 18 ventral); 19–20 Female, IBSP 47125 (19 dorsal, 20 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804639" httpUri="https://zenodo.org/record/14804639/files/figure.png" pageId="5" pageNumber="531">Figs 17</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DEEF6FE30A5CB71D4" box="[653,683,475,501]" captionStart="FIGURES 32–35" captionStartId="10.[151,264,1928,1952]" captionTargetBox="[154,1424,395,1890]" captionTargetId="figure-95@10.[152,1434,381,1905]" captionTargetPageId="10" captionText="FIGURES 32–35. P. cacau sp. nov., habitus. 32–33 Male, IBSP 47114 (32 dorsal, 33 ventral); 34–35 Female, IBSP 47106 (34 dorsal, 35 ventral). Scale lines: 2 mm." pageId="5" pageNumber="531">34</figureCitation>
<figureCitation id="130A2A04FFE48C0DEEF6FE30A5CB71D4" box="[653,683,475,501]" captionStart="FIGURES 32–35" captionStartId="10.[151,264,1928,1952]" captionTargetBox="[154,1424,395,1890]" captionTargetId="figure-95@10.[152,1434,381,1905]" captionTargetPageId="10" captionText="FIGURES 32–35. P. cacau sp. nov., habitus. 32–33 Male, IBSP 47114 (32 dorsal, 33 ventral); 34–35 Female, IBSP 47106 (34 dorsal, 35 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804647" httpUri="https://zenodo.org/record/14804647/files/figure.png" pageId="5" pageNumber="531">34</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DEEC2FE30A5B771D4" box="[697,727,475,501]" captionStart="FIGURES 47–50" captionStartId="13.[151,264,1967,1991]" captionTargetBox="[218,1372,646,1931]" captionTargetId="figure-183@13.[203,1384,633,1944]" captionTargetPageId="13" captionText="FIGURES 47–50. P. caete sp. nov., habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm." pageId="5" pageNumber="531">47</figureCitation>
<figureCitation id="130A2A04FFE48C0DEEC2FE30A5B771D4" box="[697,727,475,501]" captionStart="FIGURES 47–50" captionStartId="13.[151,264,1967,1991]" captionTargetBox="[218,1372,646,1931]" captionTargetId="figure-183@13.[203,1384,633,1944]" captionTargetPageId="13" captionText="FIGURES 47–50. P. caete sp. nov., habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804657" httpUri="https://zenodo.org/record/14804657/files/figure.png" pageId="5" pageNumber="531">47</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DEE9EFE31A46571D4" box="[741,773,474,501]" captionStart="FIGURES 63–66" captionStartId="17.[151,264,1654,1678]" captionTargetBox="[166,1415,192,1616]" captionTargetId="figure-20@17.[165,1422,181,1629]" captionTargetPageId="17" captionText="FIGURES 63–66. P. tacomare sp. nov., habitus. 63–64 Male, IBSP 97195 (63 dorsal, 64 ventral); 65–66 Female, UFMG 9488 (65 dorsal, 66 ventral). Scale lines: 2 mm." pageId="5" pageNumber="531">65</figureCitation>
<figureCitation id="130A2A04FFE48C0DEE9EFE31A46571D4" box="[741,773,474,501]" captionStart="FIGURES 63–66" captionStartId="17.[151,264,1654,1678]" captionTargetBox="[166,1415,192,1616]" captionTargetId="figure-20@17.[165,1422,181,1629]" captionTargetPageId="17" captionText="FIGURES 63–66. P. tacomare sp. nov., habitus. 63–64 Male, IBSP 97195 (63 dorsal, 64 ventral); 65–66 Female, UFMG 9488 (65 dorsal, 66 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804665" httpUri="https://zenodo.org/record/14804665/files/figure.png" pageId="5" pageNumber="531">65</figureCitation>
). Clypeus low, less than ALE diameter. Chelicerae longer than wide with three promarginal teeth, the basal one smallest, and two small retromarginal teeth. Intermarginal teeth absent. Internal margin with one escort setae (
<figureCitation id="130A2A04FFE48C0DEE9EFDC8A448721C" box="[741,808,547,573]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." pageId="5" pageNumber="531">Fig. 1</figureCitation>
<figureCitation id="130A2A04FFE48C0DEE9EFDC8A448721C" box="[741,808,547,573]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." figureDoi="http://doi.org/10.5281/zenodo.14804631" httpUri="https://zenodo.org/record/14804631/files/figure.png" pageId="5" pageNumber="531">Fig. 1</figureCitation>
). Labium rebordered, as wide as long. Endites slightly convergent, longer than wide, with dense scopula on internal margin. Serrula with single row of denticles (
<figureCitation id="130A2A04FFE48C0DE932FDACA2EF7240" box="[1353,1423,583,609]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." pageId="5" pageNumber="531">Fig. 2</figureCitation>
<figureCitation id="130A2A04FFE48C0DE932FDACA2EF7240" box="[1353,1423,583,609]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." figureDoi="http://doi.org/10.5281/zenodo.14804631" httpUri="https://zenodo.org/record/14804631/files/figure.png" pageId="5" pageNumber="531">Fig. 2</figureCitation>
). Sternum longer than wide, slightly projected between coxae IV. Female palp with single pectinate claw with six short teeth (
<figureCitation id="130A2A04FFE48C0DED65FD64A6037288" box="[286,355,655,681]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." pageId="5" pageNumber="531">Fig. 6</figureCitation>
<figureCitation id="130A2A04FFE48C0DED65FD64A6037288" box="[286,355,655,681]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." figureDoi="http://doi.org/10.5281/zenodo.14804631" httpUri="https://zenodo.org/record/14804631/files/figure.png" pageId="5" pageNumber="531">Fig. 6</figureCitation>
); sensory setae long, distally curved, with barbules along the entire setae and with distal region bearing a large, rounded pore and a single filiform extension, scattered dorsally along tarsus. Legs laterigrade, mostly 4123 but can be 1423 or 4213. Spination pattern in males: femora I–III: p1-1-1, d0-1-1, r1-1-1; femur IV: p1-1-1, d0-11, r0-1-1; patellae I–IV: p0, r0; tibiae I–II: p1-0-1, d0-0-1; r1-0-1, v3–6 pairs; tibiae III–IV: p1-0-1; d0-0-1; r1-0-1; v2-2-2; metatarsi I–II: p1-1-1; r1-1-0; v2-0-0; metatarsus III: p1-1-1; r1-1-0; v2-2-0; metatarsus IV: p1-1-1; r1-1-1; v2-2-0. Palp: femur: p0-0-1; d0-1-2; r0; patella: p1, r1; tibia: p2-1-0; d1-0-0; r0; in females: Femora I–III: p1-1-1, d0-1-1, r1-1-1; femur IV: p1-1-1, d0-11, r0-1-1; patellae I–IV: p0, r0; tibiae I–II: v4–6 pairs; tibia III: v2-2-2; tibia IV: p1-0-1; r1-0-1; v2-2-2; metatarsi I–III: v2-0-0; metatarsus IV: p1-1-1; r1-1-1; v2-2-0. Palp: femur: p0-0-1; d0-1-2; r0-0-1; tibia: p2-1-0; r1-1-0; tarsus: p2-1-0; r1-1-0. Trochanter smooth. Metatarsi I–IV distally with dorsal trilobate membrane with median hook much smaller than lateral projections (
<figureCitation id="130A2A04FFE48C0DE832FC38A3EE73CC" box="[1097,1166,979,1005]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." pageId="5" pageNumber="531">Fig. 3</figureCitation>
<figureCitation id="130A2A04FFE48C0DE832FC38A3EE73CC" box="[1097,1166,979,1005]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." figureDoi="http://doi.org/10.5281/zenodo.14804631" httpUri="https://zenodo.org/record/14804631/files/figure.png" pageId="5" pageNumber="531">Fig. 3</figureCitation>
). Tarsi and anterior half of metatarsi scopulate. Trichobothria present on dorsal tibiae, metatarsi and tarsi, arranged in several rows on tarsi, converging to a single row on metatarsi. Dorsal plate with 6–7 transversal rows, projecting over a smooth proximal plate. Trichobothrial seta base thickened in a basal bulb (
<figureCitation id="130A2A04FFE48C0DEF49FBD4A41A7478" box="[818,890,1087,1113]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." pageId="5" pageNumber="531">Fig. 4</figureCitation>
<figureCitation id="130A2A04FFE48C0DEF49FBD4A41A7478" box="[818,890,1087,1113]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." figureDoi="http://doi.org/10.5281/zenodo.14804631" httpUri="https://zenodo.org/record/14804631/files/figure.png" pageId="5" pageNumber="531">Fig. 4</figureCitation>
). Tarsal organ capsulate, with tear-drop-shaped opening, located dorsally, at distal end of tarsi (
<figureCitation id="130A2A04FFE48C0DEEC7FB88A463745C" box="[700,771,1122,1149]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." pageId="5" pageNumber="531">Fig. 5</figureCitation>
<figureCitation id="130A2A04FFE48C0DEEC7FB88A463745C" box="[700,771,1122,1149]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." figureDoi="http://doi.org/10.5281/zenodo.14804631" httpUri="https://zenodo.org/record/14804631/files/figure.png" pageId="5" pageNumber="531">Fig. 5</figureCitation>
). Tarsi with pair of pectinate claws with 9–12 small teeth with claw tufts. The distal teeth inserted entally and the proximal ones inserted ectally (
<figureCitation id="130A2A04FFE48C0DE806FB6CA3947480" box="[1149,1268,1159,1185]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-20@3.[167,1419,181,1919]" captionTargetPageId="3" captionText="FIGURES 7–12. 7, 9 P. cabruca sp. nov. (male, IBSP 47126); 8, 12 P. caete sp. nov. (male, IBSP 249602); 10–11 P. cacau sp. nov. (10 male IBSP 275288, 11 female IBSP 275471). 7–10 Tarsus claws, left leg, prolateral (7 leg I, 8 leg II, 9 leg III, 10 leg IV); 11 Opisthosoma, epiandrium, epiandrous spigots, ventral; 12 Opisthosoma colulus, ventral." pageId="5" pageNumber="531">Figs 7–10</figureCitation>
<figureCitation id="130A2A04FFE48C0DE806FB6CA3947480" box="[1149,1268,1159,1185]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-20@3.[167,1419,181,1919]" captionTargetPageId="3" captionText="FIGURES 7–12. 7, 9 P. cabruca sp. nov. (male, IBSP 47126); 8, 12 P. caete sp. nov. (male, IBSP 249602); 10–11 P. cacau sp. nov. (10 male IBSP 275288, 11 female IBSP 275471). 7–10 Tarsus claws, left leg, prolateral (7 leg I, 8 leg II, 9 leg III, 10 leg IV); 11 Opisthosoma, epiandrium, epiandrous spigots, ventral; 12 Opisthosoma colulus, ventral." figureDoi="http://doi.org/10.5281/zenodo.14804635" httpUri="https://zenodo.org/record/14804635/files/figure.png" pageId="5" pageNumber="531">Figs 7–10</figureCitation>
). Opisthosoma oval, longer than wide. Male epiandrium with small groups of epiandrous spigots (
<figureCitation id="130A2A04FFE48C0DE81BFB40A3D574E4" box="[1120,1205,1195,1221]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-20@3.[167,1419,181,1919]" captionTargetPageId="3" captionText="FIGURES 7–12. 7, 9 P. cabruca sp. nov. (male, IBSP 47126); 8, 12 P. caete sp. nov. (male, IBSP 249602); 10–11 P. cacau sp. nov. (10 male IBSP 275288, 11 female IBSP 275471). 7–10 Tarsus claws, left leg, prolateral (7 leg I, 8 leg II, 9 leg III, 10 leg IV); 11 Opisthosoma, epiandrium, epiandrous spigots, ventral; 12 Opisthosoma colulus, ventral." pageId="5" pageNumber="531">Fig. 11</figureCitation>
<figureCitation id="130A2A04FFE48C0DE81BFB40A3D574E4" box="[1120,1205,1195,1221]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-20@3.[167,1419,181,1919]" captionTargetPageId="3" captionText="FIGURES 7–12. 7, 9 P. cabruca sp. nov. (male, IBSP 47126); 8, 12 P. caete sp. nov. (male, IBSP 249602); 10–11 P. cacau sp. nov. (10 male IBSP 275288, 11 female IBSP 275471). 7–10 Tarsus claws, left leg, prolateral (7 leg I, 8 leg II, 9 leg III, 10 leg IV); 11 Opisthosoma, epiandrium, epiandrous spigots, ventral; 12 Opisthosoma colulus, ventral." figureDoi="http://doi.org/10.5281/zenodo.14804635" httpUri="https://zenodo.org/record/14804635/files/figure.png" pageId="5" pageNumber="531">Fig. 11</figureCitation>
). Colulus triangular with sparse setae (
<figureCitation id="130A2A04FFE48C0DED1CFB24A6DB74C8" box="[359,443,1231,1257]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-20@3.[167,1419,181,1919]" captionTargetPageId="3" captionText="FIGURES 7–12. 7, 9 P. cabruca sp. nov. (male, IBSP 47126); 8, 12 P. caete sp. nov. (male, IBSP 249602); 10–11 P. cacau sp. nov. (10 male IBSP 275288, 11 female IBSP 275471). 7–10 Tarsus claws, left leg, prolateral (7 leg I, 8 leg II, 9 leg III, 10 leg IV); 11 Opisthosoma, epiandrium, epiandrous spigots, ventral; 12 Opisthosoma colulus, ventral." pageId="5" pageNumber="531">Fig. 12</figureCitation>
<figureCitation id="130A2A04FFE48C0DED1CFB24A6DB74C8" box="[359,443,1231,1257]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-20@3.[167,1419,181,1919]" captionTargetPageId="3" captionText="FIGURES 7–12. 7, 9 P. cabruca sp. nov. (male, IBSP 47126); 8, 12 P. caete sp. nov. (male, IBSP 249602); 10–11 P. cacau sp. nov. (10 male IBSP 275288, 11 female IBSP 275471). 7–10 Tarsus claws, left leg, prolateral (7 leg I, 8 leg II, 9 leg III, 10 leg IV); 11 Opisthosoma, epiandrium, epiandrous spigots, ventral; 12 Opisthosoma colulus, ventral." figureDoi="http://doi.org/10.5281/zenodo.14804635" httpUri="https://zenodo.org/record/14804635/files/figure.png" pageId="5" pageNumber="531">Fig. 12</figureCitation>
). Six spinnerets: anterior lateral spinnerets contiguous, conical and bi-segmented. Basal segment elongated and cylindrical, distal segment short and truncated. Posterior median spinnerets conical and short. Posterior lateral spinnerets conical and bi-segmented. Basal segment elongated and cylindrical, distal segment short and truncated. Male palp: tibia short, less than half cymbium length with three long prolateral spines (two in
<taxonomicName id="4C314D02FFE48C0DECECFAB4A67D7559" authorityName="Rheims" authorityYear="2025" box="[151,285,1375,1400]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="531" phylum="Arthropoda" rank="species" species="tacomare" status="sp. nov.">
<emphasis id="B945EA93FFE48C0DECECFAB4A67D7559" box="[151,285,1375,1400]" italics="true" pageId="5" pageNumber="531">P. tacomare</emphasis>
@ -300,69 +301,69 @@ Coloration pattern: prosoma brown, slightly darker along thoracic striae and lat
<taxonomicNameLabel id="A27657E8FFE48C0DE8E8FAB4A38F7558" box="[1171,1263,1375,1401]" pageId="5" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
); RTA inserted distally, single (
<figureCitation id="130A2A04FFE48C0DED37FA68A6C675BC" box="[332,422,1411,1437]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="5" pageNumber="531">Figs 28</figureCitation>
<figureCitation id="130A2A04FFE48C0DED37FA68A6C675BC" box="[332,422,1411,1437]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="5" pageNumber="531">Figs 28</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DEDCFFA68A6B375BC" box="[436,467,1411,1437]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="5" pageNumber="531">43</figureCitation>
<figureCitation id="130A2A04FFE48C0DEDCFFA68A6B375BC" box="[436,467,1411,1437]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="5" pageNumber="531">43</figureCitation>
), with two (
<figureCitation id="130A2A04FFE48C0DEE1AFA68A5D575BC" box="[609,693,1411,1437]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">Fig. 74</figureCitation>
<figureCitation id="130A2A04FFE48C0DEE1AFA68A5D575BC" box="[609,693,1411,1437]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804671" httpUri="https://zenodo.org/record/14804671/files/figure.png" pageId="5" pageNumber="531">Fig. 74</figureCitation>
) or three (
<figureCitation id="130A2A04FFE48C0DEF4AFA68A4E575BC" box="[817,901,1410,1437]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">Fig. 58</figureCitation>
<figureCitation id="130A2A04FFE48C0DEF4AFA68A4E575BC" box="[817,901,1410,1437]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="5" pageNumber="531">Fig. 58</figureCitation>
) branches; cymbium oval, with large RbP and small rounded scopulae dorsally on distal third; alveolus oval; ST cup-shaped; spermophore ring-like, narrowing at the base of the embolus; TBE hyaline, keel-like, distally extending towards retrolateral margin of cymbium; TBC laminar, partially sclerotized (
<figureCitation id="130A2A04FFE48C0DEE5DFA04A51E7628" box="[550,638,1519,1545]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="5" pageNumber="531">Figs 27</figureCitation>
<figureCitation id="130A2A04FFE48C0DEE5DFA04A51E7628" box="[550,638,1519,1545]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="5" pageNumber="531">Figs 27</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DEEF1FA04A5C77628" box="[650,679,1519,1545]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="5" pageNumber="531">42</figureCitation>
<figureCitation id="130A2A04FFE48C0DEEF1FA04A5C77628" box="[650,679,1519,1545]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="5" pageNumber="531">42</figureCitation>
) or reduced to a sclerotized area (
<figureCitation id="130A2A04FFE48C0DE853FA04A31A7628" box="[1064,1146,1519,1545]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">Fig. 73</figureCitation>
<figureCitation id="130A2A04FFE48C0DE853FA04A31A7628" box="[1064,1146,1519,1545]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804671" httpUri="https://zenodo.org/record/14804671/files/figure.png" pageId="5" pageNumber="531">Fig. 73</figureCitation>
) or elevation (
<figureCitation id="130A2A04FFE48C0DE95BFA04A2157628" box="[1312,1397,1518,1545]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">Fig. 57</figureCitation>
<figureCitation id="130A2A04FFE48C0DE95BFA04A2157628" box="[1312,1397,1518,1545]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="5" pageNumber="531">Fig. 57</figureCitation>
); C sclerotized, hood-shaped, with basal end curved, arising from the center of the tegulum; E arising from tegulum between 7–9 o’clock positions, filiform (
<figureCitation id="130A2A04FFE48C0DEE18F9DCA5DA7670" box="[611,698,1591,1617]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="5" pageNumber="531">Figs 27</figureCitation>
<figureCitation id="130A2A04FFE48C0DEE18F9DCA5DA7670" box="[611,698,1591,1617]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="5" pageNumber="531">Figs 27</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DEEBCF9DCA5867670" box="[711,742,1591,1617]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">73</figureCitation>
<figureCitation id="130A2A04FFE48C0DEEBCF9DCA5867670" box="[711,742,1591,1617]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804671" httpUri="https://zenodo.org/record/14804671/files/figure.png" pageId="5" pageNumber="531">73</figureCitation>
), with subdistal projection (
<figureCitation id="130A2A04FFE48C0DE85BF9DCA3127670" box="[1056,1138,1591,1617]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="5" pageNumber="531">Fig. 42</figureCitation>
<figureCitation id="130A2A04FFE48C0DE85BF9DCA3127670" box="[1056,1138,1591,1617]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="5" pageNumber="531">Fig. 42</figureCitation>
) or modified tip (
<figureCitation id="130A2A04FFE48C0DE941F9DCA2EF7670" box="[1338,1423,1590,1617]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">Fig. 59</figureCitation>
<figureCitation id="130A2A04FFE48C0DE941F9DCA2EF7670" box="[1338,1423,1590,1617]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="5" pageNumber="531">Fig. 59</figureCitation>
); MA arising from the center of the tegulum, cup-shaped (
<figureCitation id="130A2A04FFE48C0DEF71F9B0A4007654" box="[778,864,1627,1653]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="5" pageNumber="531">Figs 27</figureCitation>
<figureCitation id="130A2A04FFE48C0DEF71F9B0A4007654" box="[778,864,1627,1653]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="5" pageNumber="531">Figs 27</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DEF10F9B1A4E97654" box="[875,905,1626,1653]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">57</figureCitation>
<figureCitation id="130A2A04FFE48C0DEF10F9B1A4E97654" box="[875,905,1626,1653]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="5" pageNumber="531">57</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DEFEEF9B0A4D27654" box="[917,946,1627,1653]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">73</figureCitation>
<figureCitation id="130A2A04FFE48C0DEFEEF9B0A4D27654" box="[917,946,1627,1653]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804671" httpUri="https://zenodo.org/record/14804671/files/figure.png" pageId="5" pageNumber="531">73</figureCitation>
) or knob-shaped (
<figureCitation id="130A2A04FFE48C0DE807F9B0A3B07654" box="[1148,1232,1627,1653]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="5" pageNumber="531">Fig. 42</figureCitation>
<figureCitation id="130A2A04FFE48C0DE807F9B0A3B07654" box="[1148,1232,1627,1653]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="5" pageNumber="531">Fig. 42</figureCitation>
). Female epigyne: EF as wide as long or longer than wide with anterior lateral MAB conspicuous or imbedded; LL smooth (
<figureCitation id="130A2A04FFE48C0DE93BF994A2F876B8" box="[1344,1432,1663,1689]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="5" pageNumber="531">Figs 44</figureCitation>
<figureCitation id="130A2A04FFE48C0DE93BF994A2F876B8" box="[1344,1432,1663,1689]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="5" pageNumber="531">Figs 44</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DECECF949A7D4769C" box="[151,180,1698,1725]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">75</figureCitation>
<figureCitation id="130A2A04FFE48C0DECECF949A7D4769C" box="[151,180,1698,1725]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804671" httpUri="https://zenodo.org/record/14804671/files/figure.png" pageId="5" pageNumber="531">75</figureCitation>
) or with posterior, lateral projections (
<figureCitation id="130A2A04FFE48C0DEE0DF948A5AF769C" box="[630,719,1699,1725]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="5" pageNumber="531">Figs 29</figureCitation>
<figureCitation id="130A2A04FFE48C0DEE0DF948A5AF769C" box="[630,719,1699,1725]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="5" pageNumber="531">Figs 29</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DEEA5F948A59D769C" box="[734,765,1699,1725]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">60</figureCitation>
<figureCitation id="130A2A04FFE48C0DEEA5F948A59D769C" box="[734,765,1699,1725]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="5" pageNumber="531">60</figureCitation>
), partially covering MS (
<figureCitation id="130A2A04FFE48C0DE858F948A31C769C" box="[1059,1148,1699,1725]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">Figs 60</figureCitation>
<figureCitation id="130A2A04FFE48C0DE858F948A31C769C" box="[1059,1148,1699,1725]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="5" pageNumber="531">Figs 60</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DE8F0F949A3C8769C" box="[1163,1192,1698,1725]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">75</figureCitation>
<figureCitation id="130A2A04FFE48C0DE8F0F949A3C8769C" box="[1163,1192,1698,1725]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804671" httpUri="https://zenodo.org/record/14804671/files/figure.png" pageId="5" pageNumber="531">75</figureCitation>
) or partially covered by MS (
<figureCitation id="130A2A04FFE48C0DEC8DF92CA62E76C0" box="[246,334,1735,1761]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="5" pageNumber="531">Figs 29</figureCitation>
<figureCitation id="130A2A04FFE48C0DEC8DF92CA62E76C0" box="[246,334,1735,1761]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="5" pageNumber="531">Figs 29</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DED27F92CA61C76C0" box="[348,380,1735,1761]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="5" pageNumber="531">44</figureCitation>
<figureCitation id="130A2A04FFE48C0DED27F92CA61C76C0" box="[348,380,1735,1761]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="5" pageNumber="531">44</figureCitation>
); anterior grove straight or slightly recurved (
<figureCitation id="130A2A04FFE48C0DEFF2F92CA48276C0" box="[905,994,1735,1761]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">Figs 60</figureCitation>
<figureCitation id="130A2A04FFE48C0DEFF2F92CA48276C0" box="[905,994,1735,1761]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="5" pageNumber="531">Figs 60</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DEF8BF92DA36D76C0" box="[1008,1037,1734,1761]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">75</figureCitation>
<figureCitation id="130A2A04FFE48C0DEF8BF92DA36D76C0" box="[1008,1037,1734,1761]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804671" httpUri="https://zenodo.org/record/14804671/files/figure.png" pageId="5" pageNumber="531">75</figureCitation>
) or projecting over MS, forming a scape-like structure (
<figureCitation id="130A2A04FFE48C0DEDF8F900A6BB7724" box="[387,475,1771,1797]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="5" pageNumber="531">Figs 29</figureCitation>
<figureCitation id="130A2A04FFE48C0DEDF8F900A6BB7724" box="[387,475,1771,1797]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="5" pageNumber="531">Figs 29</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DED92F900A5697724" box="[489,521,1771,1797]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="5" pageNumber="531">44</figureCitation>
<figureCitation id="130A2A04FFE48C0DED92F900A5697724" box="[489,521,1771,1797]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="5" pageNumber="531">44</figureCitation>
). Vulva: IDS divided into a proximal, expanded hyaline part (Dh), closer to CO, connecting to a distal sclerotized part (Ds) closer to FD; FD short and hook shaped (
<figureCitation id="130A2A04FFE48C0DE83CF8E4A3FD7708" box="[1095,1181,1807,1833]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="5" pageNumber="531">Figs 30</figureCitation>
<figureCitation id="130A2A04FFE48C0DE83CF8E4A3FD7708" box="[1095,1181,1807,1833]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="5" pageNumber="531">Figs 30</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DE8D2F8E5A3A77708" box="[1193,1223,1806,1833]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="5" pageNumber="531">45</figureCitation>
<figureCitation id="130A2A04FFE48C0DE8D2F8E5A3A77708" box="[1193,1223,1806,1833]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="5" pageNumber="531">45</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DE8A8F8E4A3917708" box="[1235,1265,1807,1833]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">61</figureCitation>
<figureCitation id="130A2A04FFE48C0DE8A8F8E4A3917708" box="[1235,1265,1807,1833]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="5" pageNumber="531">61</figureCitation>
,
<figureCitation id="130A2A04FFE48C0DE886F8E4A27D7708" box="[1277,1309,1807,1833]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="5" pageNumber="531">76</figureCitation>
<figureCitation id="130A2A04FFE48C0DE886F8E4A27D7708" box="[1277,1309,1807,1833]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804671" httpUri="https://zenodo.org/record/14804671/files/figure.png" pageId="5" pageNumber="531">76</figureCitation>
).
</paragraph>
<paragraph id="8B8E3681FFE48C0DECBCF8D9A24D776C" blockId="5.[151,1437,151,1941]" box="[199,1325,1842,1869]" pageId="5" pageNumber="531">
@ -408,7 +409,7 @@ to the state of
, all in
<collectingCountry id="F3267611FFE48C0DEDD3F890A68C77B4" box="[424,492,1915,1941]" name="Brazil" pageId="5" pageNumber="531">Brazil</collectingCountry>
(
<figureCitation id="130A2A04FFE48C0DED80F890A51F77B4" box="[507,639,1915,1941]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." pageId="5" pageNumber="531">Figs 78–81</figureCitation>
<figureCitation id="130A2A04FFE48C0DED80F890A51F77B4" box="[507,639,1915,1941]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14804675" httpUri="https://zenodo.org/record/14804675/files/figure.png" pageId="5" pageNumber="531">Figs 78–81</figureCitation>
).
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99
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<figureCitation id="130A2A04FFE78C0EECECFCD5A7997379" box="[151,249,830,856]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." figureDoi="http://doi.org/10.5281/zenodo.14804631" httpUri="https://zenodo.org/record/14804631/files/figure.png" pageId="6" pageNumber="532">Figs 1, 3</figureCitation>
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<figureCitation id="130A2A04FFE78C0EED7DFCD5A6527379" box="[262,306,830,856]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-20@3.[167,1419,181,1919]" captionTargetPageId="3" captionText="FIGURES 7–12. 7, 9 P. cabruca sp. nov. (male, IBSP 47126); 8, 12 P. caete sp. nov. (male, IBSP 249602); 10–11 P. cacau sp. nov. (10 male IBSP 275288, 11 female IBSP 275471). 7–10 Tarsus claws, left leg, prolateral (7 leg I, 8 leg II, 9 leg III, 10 leg IV); 11 Opisthosoma, epiandrium, epiandrous spigots, ventral; 12 Opisthosoma colulus, ventral." pageId="6" pageNumber="532">7, 9</figureCitation>
<figureCitation id="130A2A04FFE78C0EED7DFCD5A6527379" box="[262,306,830,856]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-20@3.[167,1419,181,1919]" captionTargetPageId="3" captionText="FIGURES 7–12. 7, 9 P. cabruca sp. nov. (male, IBSP 47126); 8, 12 P. caete sp. nov. (male, IBSP 249602); 10–11 P. cacau sp. nov. (10 male IBSP 275288, 11 female IBSP 275471). 7–10 Tarsus claws, left leg, prolateral (7 leg I, 8 leg II, 9 leg III, 10 leg IV); 11 Opisthosoma, epiandrium, epiandrous spigots, ventral; 12 Opisthosoma colulus, ventral." figureDoi="http://doi.org/10.5281/zenodo.14804635" httpUri="https://zenodo.org/record/14804635/files/figure.png" pageId="6" pageNumber="532">7, 9</figureCitation>
,
<figureCitation id="130A2A04FFE78C0EED45FCD5A6E77379" box="[318,391,830,856]" captionStart-0="FIGURES 17–20" captionStart-1="FIGURES 21–25" captionStart-2="FIGURES 26–31" captionStartId-0="7.[151,264,1949,1973]" captionStartId-1="8.[151,264,1366,1390]" captionStartId-2="9.[151,264,1450,1474]" captionTargetBox-0="[200,1384,429,1911]" captionTargetBox-1="[180,1417,194,1329]" captionTargetBox-2="[180,1415,186,1411]" captionTargetId-0="figure-125@7.[189,1397,417,1925]" captionTargetId-1="figure-16@8.[166,1422,181,1342]" captionTargetId-2="figure-20@9.[162,1425,181,1425]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURES 17–20. P. cabruca sp. nov., habitus. 17–18 Male, IBSP 47105 (17 dorsal, 18 ventral); 19–20 Female, IBSP 47125 (19 dorsal, 20 ventral). Scale lines: 2 mm." captionText-1="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." captionText-2="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="6" pageNumber="532">17–31</figureCitation>
<figureCitation id="130A2A04FFE78C0EED45FCD5A6E77379" box="[318,391,830,856]" captionStart-0="FIGURES 17–20" captionStart-1="FIGURES 21–25" captionStart-2="FIGURES 26–31" captionStartId-0="7.[151,264,1949,1973]" captionStartId-1="8.[151,264,1366,1390]" captionStartId-2="9.[151,264,1450,1474]" captionTargetBox-0="[200,1384,429,1911]" captionTargetBox-1="[180,1417,194,1329]" captionTargetBox-2="[180,1415,186,1411]" captionTargetId-0="figure-125@7.[189,1397,417,1925]" captionTargetId-1="figure-16@8.[166,1422,181,1342]" captionTargetId-2="figure-20@9.[162,1425,181,1425]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURES 17–20. P. cabruca sp. nov., habitus. 17–18 Male, IBSP 47105 (17 dorsal, 18 ventral); 19–20 Female, IBSP 47125 (19 dorsal, 20 ventral). Scale lines: 2 mm." captionText-1="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." captionText-2="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi-0="http://doi.org/10.5281/zenodo.14804639" figureDoi-1="http://doi.org/10.5281/zenodo.14804643" figureDoi-2="http://doi.org/10.5281/zenodo.14804645" httpUri-0="https://zenodo.org/record/14804639/files/figure.png" httpUri-1="https://zenodo.org/record/14804643/files/figure.png" httpUri-2="https://zenodo.org/record/14804645/files/figure.png" pageId="6" pageNumber="532">17–31</figureCitation>
,
<figureCitation id="130A2A04FFE78C0EEDE9FCD5A6CE7379" box="[402,430,830,856]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." pageId="6" pageNumber="532">78</figureCitation>
<figureCitation id="130A2A04FFE78C0EEDE9FCD5A6CE7379" box="[402,430,830,856]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14804675" httpUri="https://zenodo.org/record/14804675/files/figure.png" pageId="6" pageNumber="532">78</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32B650AFFE78C0EECECFC6DA57A7475" pageId="6" pageNumber="532" type="materials_examined">
@ -204,11 +205,11 @@ resemble those of
<taxonomicNameLabel id="A27657E8FFE78C0EE86FFB21A31074C5" box="[1044,1136,1226,1252]" pageId="6" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="130A2A04FFE78C0EE8FAFB21A26374C5" box="[1153,1283,1226,1252]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="6" pageNumber="532">Figs 41–46</figureCitation>
<figureCitation id="130A2A04FFE78C0EE8FAFB21A26374C5" box="[1153,1283,1226,1252]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="6" pageNumber="532">Figs 41–46</figureCitation>
) by the male palps with single branched RTA (two or three branches in the other species) and female epigyne with MS partially covering LL and anterior groove projecting posteriorly over MS (LL partially covering MS and anterior groove not projecting over MS in the other species). They are distinguished from those of the latter species by the male palps with MA cup-shaped, with strongly indented margin (
<figureCitation id="130A2A04FFE78C0EEE94FAB1A4267555" box="[751,838,1370,1396]" captionStart="FIGURES 21–25" captionStartId="8.[151,264,1366,1390]" captionTargetBox="[180,1417,194,1329]" captionTargetId="figure-16@8.[166,1422,181,1342]" captionTargetPageId="8" captionText="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." pageId="6" pageNumber="532">Figs 22</figureCitation>
<figureCitation id="130A2A04FFE78C0EEE94FAB1A4267555" box="[751,838,1370,1396]" captionStart="FIGURES 21–25" captionStartId="8.[151,264,1366,1390]" captionTargetBox="[180,1417,194,1329]" captionTargetId="figure-16@8.[166,1422,181,1342]" captionTargetPageId="8" captionText="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." figureDoi="http://doi.org/10.5281/zenodo.14804643" httpUri="https://zenodo.org/record/14804643/files/figure.png" pageId="6" pageNumber="532">Figs 22</figureCitation>
,
<figureCitation id="130A2A04FFE78C0EEF2AFAB1A40E7555" box="[849,878,1370,1396]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="6" pageNumber="532">27</figureCitation>
<figureCitation id="130A2A04FFE78C0EEF2AFAB1A40E7555" box="[849,878,1370,1396]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="6" pageNumber="532">27</figureCitation>
) (small, knob-shaped in
<taxonomicName id="4C314D02FFE78C0EE8F8FAB0A3817555" authorityName="Rheims" authorityYear="2025" box="[1155,1249,1371,1396]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="532" phylum="Arthropoda" rank="species" species="cacau" status="sp. nov.">
<emphasis id="B945EA93FFE78C0EE8F8FAB0A3817555" box="[1155,1249,1371,1396]" italics="true" pageId="6" pageNumber="532">P. cacau</emphasis>
@ -217,9 +218,9 @@ resemble those of
<taxonomicNameLabel id="A27657E8FFE78C0EE893FAB1A2217555" box="[1256,1345,1370,1396]" pageId="6" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and by the epigyne with LL with posterior triangular projections and MS not surpassing the posterior margin of LL (
<figureCitation id="130A2A04FFE78C0EE910FA95A7D5759D" captionStart="FIGURES 21–25" captionStartId="8.[151,264,1366,1390]" captionTargetBox="[180,1417,194,1329]" captionTargetId="figure-16@8.[166,1422,181,1342]" captionTargetPageId="8" captionText="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." pageId="6" pageNumber="532">Figs 24</figureCitation>
<figureCitation id="130A2A04FFE78C0EE910FA95A7D5759D" captionStart="FIGURES 21–25" captionStartId="8.[151,264,1366,1390]" captionTargetBox="[180,1417,194,1329]" captionTargetId="figure-16@8.[166,1422,181,1342]" captionTargetPageId="8" captionText="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." figureDoi="http://doi.org/10.5281/zenodo.14804643" httpUri="https://zenodo.org/record/14804643/files/figure.png" pageId="6" pageNumber="532">Figs 24</figureCitation>
,
<figureCitation id="130A2A04FFE78C0EECBAFA49A7BE759D" box="[193,222,1442,1468]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="6" pageNumber="532">29</figureCitation>
<figureCitation id="130A2A04FFE78C0EECBAFA49A7BE759D" box="[193,222,1442,1468]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="6" pageNumber="532">29</figureCitation>
) (LL with posterior margin smooth and MS surpassing the posterior margin of LL in
<taxonomicName id="4C314D02FFE78C0EE8E1FA48A399759D" authorityName="Rheims" authorityYear="2025" box="[1178,1273,1443,1468]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="532" phylum="Arthropoda" rank="species" species="cacau" status="sp. nov.">
<emphasis id="B945EA93FFE78C0EE8E1FA48A399759D" box="[1178,1273,1443,1468]" italics="true" pageId="6" pageNumber="532">P. cacau</emphasis>
@ -238,11 +239,11 @@ Description. Male (
):
</emphasis>
Coloration as in
<figureCitation id="130A2A04FFE78C0EEE95FA2DA41075C1" box="[750,880,1478,1504]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1949,1973]" captionTargetBox="[200,1384,429,1911]" captionTargetId="figure-125@7.[189,1397,417,1925]" captionTargetPageId="7" captionText="FIGURES 17–20. P. cabruca sp. nov., habitus. 17–18 Male, IBSP 47105 (17 dorsal, 18 ventral); 19–20 Female, IBSP 47125 (19 dorsal, 20 ventral). Scale lines: 2 mm." pageId="6" pageNumber="532">Figs 17–18</figureCitation>
<figureCitation id="130A2A04FFE78C0EEE95FA2DA41075C1" box="[750,880,1478,1504]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1949,1973]" captionTargetBox="[200,1384,429,1911]" captionTargetId="figure-125@7.[189,1397,417,1925]" captionTargetPageId="7" captionText="FIGURES 17–20. P. cabruca sp. nov., habitus. 17–18 Male, IBSP 47105 (17 dorsal, 18 ventral); 19–20 Female, IBSP 47125 (19 dorsal, 20 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804639" httpUri="https://zenodo.org/record/14804639/files/figure.png" pageId="6" pageNumber="532">Figs 17–18</figureCitation>
. Total length 13.2. Dorsal shield of prosoma: 6.0 long, 5.1 wide. Opisthosoma: 6.7 long, 4.0 wide. Eyes: diameters: 0.36, 0.30, 0.20, 0.30; interdistances: 0.30, 0.16, 0.54, 0.52, 0.24, 0.14. Legs (12/43): I: 19.2 (5.7, 3.0, 4.9, 4.4, 1.2); II: 18.4 (5.5, 2.9, 4.6, 4.2, 1.2); III: 15.0 (4.7, 2.4, 3.4, 3.4, 1.1); IV: 18.4 (5.6, 2.1, 4.3, 4.9, 1.5). Spination follows the generic pattern, with tibiae I–II with three pairs of ventral spines. Palp: VTA absent; RTA single branched, shaped as an upside-down shoe in retrolateral view and with distal margin medially depressed in ventral view; ST visible between 9–12 o’clock in ventral view; spermophore with pronounced undulation between 6–7 o’clock, close to embolus base; TBE long, with rounded distal point; TBC laminar, roughly as wide as long with rounded tip and more sclerotized margins; C hood three times longer than wide, strongly concave in retrolateral view; tip truncated and curved; E filiform, arising from tegulum at 8:30 o’clock position; MA roughly as wide as long, widest distally (
<figureCitation id="130A2A04FFE78C0EE875F8E1A3EF7705" box="[1038,1167,1802,1828]" captionStart="FIGURES 21–25" captionStartId="8.[151,264,1366,1390]" captionTargetBox="[180,1417,194,1329]" captionTargetId="figure-16@8.[166,1422,181,1342]" captionTargetPageId="8" captionText="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." pageId="6" pageNumber="532">Figs 21–23</figureCitation>
<figureCitation id="130A2A04FFE78C0EE875F8E1A3EF7705" box="[1038,1167,1802,1828]" captionStart="FIGURES 21–25" captionStartId="8.[151,264,1366,1390]" captionTargetBox="[180,1417,194,1329]" captionTargetId="figure-16@8.[166,1422,181,1342]" captionTargetPageId="8" captionText="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." figureDoi="http://doi.org/10.5281/zenodo.14804643" httpUri="https://zenodo.org/record/14804643/files/figure.png" pageId="6" pageNumber="532">Figs 21–23</figureCitation>
,
<figureCitation id="130A2A04FFE78C0EE8E0F8E1A3857705" box="[1179,1253,1802,1828]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="6" pageNumber="532">26–28</figureCitation>
<figureCitation id="130A2A04FFE78C0EE8E0F8E1A3857705" box="[1179,1253,1802,1828]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="6" pageNumber="532">26–28</figureCitation>
).
</paragraph>
<paragraph id="8B8E3681FFE78C0FECBCF8C5A4C0713C" blockId="6.[151,1437,901,2008]" lastBlockId="7.[151,1436,151,393]" lastPageId="7" lastPageNumber="533" pageId="6" pageNumber="532">
@ -252,17 +253,17 @@ Female (IBSP 41116,
):
</emphasis>
Coloration as in
<figureCitation id="130A2A04FFE78C0EEE85F8C5A41E7769" box="[766,894,1838,1864]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1949,1973]" captionTargetBox="[200,1384,429,1911]" captionTargetId="figure-125@7.[189,1397,417,1925]" captionTargetPageId="7" captionText="FIGURES 17–20. P. cabruca sp. nov., habitus. 17–18 Male, IBSP 47105 (17 dorsal, 18 ventral); 19–20 Female, IBSP 47125 (19 dorsal, 20 ventral). Scale lines: 2 mm." pageId="6" pageNumber="532">Figs 19–20</figureCitation>
<figureCitation id="130A2A04FFE78C0EEE85F8C5A41E7769" box="[766,894,1838,1864]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1949,1973]" captionTargetBox="[200,1384,429,1911]" captionTargetId="figure-125@7.[189,1397,417,1925]" captionTargetPageId="7" captionText="FIGURES 17–20. P. cabruca sp. nov., habitus. 17–18 Male, IBSP 47105 (17 dorsal, 18 ventral); 19–20 Female, IBSP 47125 (19 dorsal, 20 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804639" httpUri="https://zenodo.org/record/14804639/files/figure.png" pageId="6" pageNumber="532">Figs 19–20</figureCitation>
. Total length 13.6. Dorsal shield of prosoma: 6.1 long, 5.0 wide. Opisthosoma: 7.3 long, 4.2 wide. Eyes: diameters: 0.36, 0.32, 0.22, 0.30; interdistances: 0.30, 0.20, 0.50, 0.52, 0.20, 0.08. Legs (4123): I: 16.8 (5.1, 2.9, 4.2, 3.5, 1.1); II: 16.4 (5.1, 2.8, 4.0, 3.4, 1.1); III: 13.4 (4.3, 2.2, 2.9, 2.9, 1.1); IV: 17.0 (5.1, 2.1, 3.8, 4.6, 1.4). Spination follows the generic pattern except tibiae I–II: v2-2-2- 2, metatarsus I: p1-1-0; tibia III: p0-1-0; r0-1-0; femur IV: r1-1-1. Epigyne: EF slightly longer than wide; with one long lateral MAB on each side (
<figureCitation id="130A2A04FFE68C0FED87FF7CA52F7090" box="[508,591,151,177]" captionStart="FIGURES 21–25" captionStartId="8.[151,264,1366,1390]" captionTargetBox="[180,1417,194,1329]" captionTargetId="figure-16@8.[166,1422,181,1342]" captionTargetPageId="8" captionText="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." pageId="7" pageNumber="533">Fig. 24</figureCitation>
<figureCitation id="130A2A04FFE68C0FED87FF7CA52F7090" box="[508,591,151,177]" captionStart="FIGURES 21–25" captionStartId="8.[151,264,1366,1390]" captionTargetBox="[180,1417,194,1329]" captionTargetId="figure-16@8.[166,1422,181,1342]" captionTargetPageId="8" captionText="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." figureDoi="http://doi.org/10.5281/zenodo.14804643" httpUri="https://zenodo.org/record/14804643/files/figure.png" pageId="7" pageNumber="533">Fig. 24</figureCitation>
), not always distinct; LL rounded, with lateral triangular projections almost three times longer than wide; MS oval, wider than long; ag forming a triangular elevated area over the entire length of MS (
<figureCitation id="130A2A04FFE68C0FEC91FF34A62070D8" box="[234,320,223,249]" captionStart="FIGURES 21–25" captionStartId="8.[151,264,1366,1390]" captionTargetBox="[180,1417,194,1329]" captionTargetId="figure-16@8.[166,1422,181,1342]" captionTargetPageId="8" captionText="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." pageId="7" pageNumber="533">Figs 24</figureCitation>
<figureCitation id="130A2A04FFE68C0FEC91FF34A62070D8" box="[234,320,223,249]" captionStart="FIGURES 21–25" captionStartId="8.[151,264,1366,1390]" captionTargetBox="[180,1417,194,1329]" captionTargetId="figure-16@8.[166,1422,181,1342]" captionTargetPageId="8" captionText="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." figureDoi="http://doi.org/10.5281/zenodo.14804643" httpUri="https://zenodo.org/record/14804643/files/figure.png" pageId="7" pageNumber="533">Figs 24</figureCitation>
,
<figureCitation id="130A2A04FFE68C0FED30FF34A60B70D8" box="[331,363,223,249]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="7" pageNumber="533">29</figureCitation>
<figureCitation id="130A2A04FFE68C0FED30FF34A60B70D8" box="[331,363,223,249]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="7" pageNumber="533">29</figureCitation>
). Vulva: IDS with Dh irregularly shaped connecting to Ds posteriorly; Ds irregularly shaped with many chambers visible by transparency; FD laterad (
<figureCitation id="130A2A04FFE68C0FEE9CFEE8A45E713C" box="[743,830,258,285]" captionStart="FIGURES 21–25" captionStartId="8.[151,264,1366,1390]" captionTargetBox="[180,1417,194,1329]" captionTargetId="figure-16@8.[166,1422,181,1342]" captionTargetPageId="8" captionText="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." pageId="7" pageNumber="533">Figs 25</figureCitation>
<figureCitation id="130A2A04FFE68C0FEE9CFEE8A45E713C" box="[743,830,258,285]" captionStart="FIGURES 21–25" captionStartId="8.[151,264,1366,1390]" captionTargetBox="[180,1417,194,1329]" captionTargetId="figure-16@8.[166,1422,181,1342]" captionTargetPageId="8" captionText="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." figureDoi="http://doi.org/10.5281/zenodo.14804643" httpUri="https://zenodo.org/record/14804643/files/figure.png" pageId="7" pageNumber="533">Figs 25</figureCitation>
,
<figureCitation id="130A2A04FFE68C0FEF31FEE8A4F4713C" box="[842,916,259,285]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="7" pageNumber="533">30–31</figureCitation>
<figureCitation id="130A2A04FFE68C0FEF31FEE8A4F4713C" box="[842,916,259,285]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="7" pageNumber="533">30–31</figureCitation>
).
</paragraph>
<paragraph id="8B8E3681FFE68C0FECBCFECDA4D57144" blockId="7.[151,1436,151,393]" pageId="7" pageNumber="533">
@ -278,7 +279,7 @@ Known only from the state of
, northeastern
<collectingCountry id="F3267611FFE68C0FEFE1FE84A4BE71A8" box="[922,990,367,393]" name="Brazil" pageId="7" pageNumber="533">Brazil</collectingCountry>
(
<figureCitation id="130A2A04FFE68C0FEF96FE84A32271A8" box="[1005,1090,367,393]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." pageId="7" pageNumber="533">Fig. 78</figureCitation>
<figureCitation id="130A2A04FFE68C0FEF96FE84A32271A8" box="[1005,1090,367,393]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14804675" httpUri="https://zenodo.org/record/14804675/files/figure.png" pageId="7" pageNumber="533">Fig. 78</figureCitation>
).
</paragraph>
</subSubSection>

107
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<paragraph id="8B8E3681FFE98C00ECECF9E3A6CD7603" blockId="8.[151,429,1508,1570]" box="[151,429,1544,1570]" pageId="8" pageNumber="534">
<figureCitation id="130A2A04FFE98C00ECECF9E3A7BE7603" box="[151,222,1544,1570]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." pageId="8" pageNumber="534">Figs 6</figureCitation>
<figureCitation id="130A2A04FFE98C00ECECF9E3A7BE7603" box="[151,222,1544,1570]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." figureDoi="http://doi.org/10.5281/zenodo.14804631" httpUri="https://zenodo.org/record/14804631/files/figure.png" pageId="8" pageNumber="534">Figs 6</figureCitation>
,
<figureCitation id="130A2A04FFE98C00EC91F9E3A6527603" box="[234,306,1544,1570]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-20@3.[167,1419,181,1919]" captionTargetPageId="3" captionText="FIGURES 7–12. 7, 9 P. cabruca sp. nov. (male, IBSP 47126); 8, 12 P. caete sp. nov. (male, IBSP 249602); 10–11 P. cacau sp. nov. (10 male IBSP 275288, 11 female IBSP 275471). 7–10 Tarsus claws, left leg, prolateral (7 leg I, 8 leg II, 9 leg III, 10 leg IV); 11 Opisthosoma, epiandrium, epiandrous spigots, ventral; 12 Opisthosoma colulus, ventral." pageId="8" pageNumber="534">10–11</figureCitation>
<figureCitation id="130A2A04FFE98C00EC91F9E3A6527603" box="[234,306,1544,1570]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-20@3.[167,1419,181,1919]" captionTargetPageId="3" captionText="FIGURES 7–12. 7, 9 P. cabruca sp. nov. (male, IBSP 47126); 8, 12 P. caete sp. nov. (male, IBSP 249602); 10–11 P. cacau sp. nov. (10 male IBSP 275288, 11 female IBSP 275471). 7–10 Tarsus claws, left leg, prolateral (7 leg I, 8 leg II, 9 leg III, 10 leg IV); 11 Opisthosoma, epiandrium, epiandrous spigots, ventral; 12 Opisthosoma colulus, ventral." figureDoi="http://doi.org/10.5281/zenodo.14804635" httpUri="https://zenodo.org/record/14804635/files/figure.png" pageId="8" pageNumber="534">10–11</figureCitation>
,
<figureCitation id="130A2A04FFE98C00ED46F9E3A6E67603" box="[317,390,1544,1570]" captionStart-0="FIGURES 32–35" captionStart-1="FIGURES 36–40" captionStart-2="FIGURES 41–46" captionStartId-0="10.[151,264,1928,1952]" captionStartId-1="11.[151,264,1815,1839]" captionStartId-2="12.[151,264,1746,1770]" captionTargetBox-0="[154,1424,395,1890]" captionTargetBox-1="[195,1392,576,1779]" captionTargetBox-2="[184,1396,501,1710]" captionTargetId-0="figure-95@10.[152,1434,381,1905]" captionTargetId-1="figure-193@11.[180,1408,561,1792]" captionTargetId-2="figure-167@12.[181,1406,489,1722]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURES 32–35. P. cacau sp. nov., habitus. 32–33 Male, IBSP 47114 (32 dorsal, 33 ventral); 34–35 Female, IBSP 47106 (34 dorsal, 35 ventral). Scale lines: 2 mm." captionText-1="FIGURES 36–40. P. cacau sp. nov. 36–38 Male (IBSP 47114), left palp (36 prolateral, 37 ventral, 38 retrolateral); 39–40 Female (IBSP 275186), genitalia (39 epigyne, ventral, 40 vulva, dorsal). Scale lines: 1mm." captionText-2="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="8" pageNumber="534">32–46</figureCitation>
<figureCitation id="130A2A04FFE98C00ED46F9E3A6E67603" box="[317,390,1544,1570]" captionStart-0="FIGURES 32–35" captionStart-1="FIGURES 36–40" captionStart-2="FIGURES 41–46" captionStartId-0="10.[151,264,1928,1952]" captionStartId-1="11.[151,264,1815,1839]" captionStartId-2="12.[151,264,1746,1770]" captionTargetBox-0="[154,1424,395,1890]" captionTargetBox-1="[195,1392,576,1779]" captionTargetBox-2="[184,1396,501,1710]" captionTargetId-0="figure-95@10.[152,1434,381,1905]" captionTargetId-1="figure-193@11.[180,1408,561,1792]" captionTargetId-2="figure-167@12.[181,1406,489,1722]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURES 32–35. P. cacau sp. nov., habitus. 32–33 Male, IBSP 47114 (32 dorsal, 33 ventral); 34–35 Female, IBSP 47106 (34 dorsal, 35 ventral). Scale lines: 2 mm." captionText-1="FIGURES 36–40. P. cacau sp. nov. 36–38 Male (IBSP 47114), left palp (36 prolateral, 37 ventral, 38 retrolateral); 39–40 Female (IBSP 275186), genitalia (39 epigyne, ventral, 40 vulva, dorsal). Scale lines: 1mm." captionText-2="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi-0="http://doi.org/10.5281/zenodo.14804647" figureDoi-1="http://doi.org/10.5281/zenodo.14804651" figureDoi-2="http://doi.org/10.5281/zenodo.14804653" httpUri-0="https://zenodo.org/record/14804647/files/figure.png" httpUri-1="https://zenodo.org/record/14804651/files/figure.png" httpUri-2="https://zenodo.org/record/14804653/files/figure.png" pageId="8" pageNumber="534">32–46</figureCitation>
,
<figureCitation id="130A2A04FFE98C00EDEAF9E3A6CD7603" box="[401,429,1544,1570]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." pageId="8" pageNumber="534">79</figureCitation>
<figureCitation id="130A2A04FFE98C00EDEAF9E3A6CD7603" box="[401,429,1544,1570]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14804675" httpUri="https://zenodo.org/record/14804675/files/figure.png" pageId="8" pageNumber="534">79</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32B650AFFE98C00ECECF9BBA4CC77D7" pageId="8" pageNumber="534" type="materials_examined">
@ -293,7 +294,7 @@ leg. (
.
</paragraph>
</subSubSection>
<caption id="DF4E6609FFE88C01ECECFA41A6897657" pageId="9" pageNumber="535" startId="9.[151,264,1450,1474]" targetBox="[180,1415,186,1411]" targetPageId="9" targetType="figure">
<caption id="DF4E6609FFE88C01ECECFA41A6897657" ID-DOI="http://doi.org/10.5281/zenodo.14804645" ID-Zenodo-Dep="14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="9" pageNumber="535" startId="9.[151,264,1450,1474]" targetBox="[180,1415,186,1411]" targetPageId="9" targetType="figure">
<paragraph id="8B8E3681FFE88C01ECECFA41A6897657" blockId="9.[151,1437,1450,1654]" pageId="9" pageNumber="535">
<emphasis id="B945EA93FFE88C01ECECFA41A63675E3" bold="true" box="[151,342,1450,1474]" pageId="9" pageNumber="535">FIGURES 26–31.</emphasis>
<taxonomicName id="4C314D02FFE88C01ED26FA40A6AA75E3" authorityName="Rheims" authorityYear="2025" box="[349,458,1450,1474]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="535" phylum="Arthropoda" rank="species" species="cabruca" status="sp. nov.">
@ -343,13 +344,13 @@ resemble those of
<taxonomicNameLabel id="A27657E8FFE88C01E86FF897A31077B7" box="[1044,1136,1916,1942]" pageId="9" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="130A2A04FFE88C01E8FAF897A26377B7" box="[1153,1283,1916,1942]" captionStart-0="FIGURES 21–25" captionStart-1="FIGURES 26–31" captionStartId-0="8.[151,264,1366,1390]" captionStartId-1="9.[151,264,1450,1474]" captionTargetBox-0="[180,1417,194,1329]" captionTargetBox-1="[180,1415,186,1411]" captionTargetId-0="figure-16@8.[166,1422,181,1342]" captionTargetId-1="figure-20@9.[162,1425,181,1425]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." captionText-1="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="9" pageNumber="535">Figs 21–31</figureCitation>
<figureCitation id="130A2A04FFE88C01E8FAF897A26377B7" box="[1153,1283,1916,1942]" captionStart-0="FIGURES 21–25" captionStart-1="FIGURES 26–31" captionStartId-0="8.[151,264,1366,1390]" captionStartId-1="9.[151,264,1450,1474]" captionTargetBox-0="[180,1417,194,1329]" captionTargetBox-1="[180,1415,186,1411]" captionTargetId-0="figure-16@8.[166,1422,181,1342]" captionTargetId-1="figure-20@9.[162,1425,181,1425]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 21–25. P. cabruca sp. nov. 21–23 Male (IBSP 47105), left palp (21 prolateral, 22 ventral, 23 retrolateral); 24–25 Female (IBSP 47125), genitalia (24 epigyne, ventral; 25 vulva, dorsal). Scale lines: 1 mm. MAB—muscle attachment bands." captionText-1="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi-0="http://doi.org/10.5281/zenodo.14804643" figureDoi-1="http://doi.org/10.5281/zenodo.14804645" httpUri-0="https://zenodo.org/record/14804643/files/figure.png" httpUri-1="https://zenodo.org/record/14804645/files/figure.png" pageId="9" pageNumber="535">Figs 21–31</figureCitation>
) by the male palps with single branched RTA (two or three branches in the other species) and female epigyne with MS partially covering LL and anterior groove projecting posteriorly over MS (LL partially covering MS and anterior groove not projecting over MS in the other species). They are distinguished from those of the latter species by the male palps with MA, small, knob-shaped (
<figureCitation id="130A2A04FFEB8C02ED8AFF50A51170F4" box="[497,625,186,213]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1390,1414]" captionTargetBox="[151,1435,181,1366]" captionTargetId="figure-16@4.[151,1435,181,1366]" captionTargetPageId="4" captionText="FIGURES 13–16. 13–14 P. caete sp. nov.; 15–16 P. cacau sp. nov. 13 Left male palp, ventral (IBSP 91751); 14 Right palp, embolus, detail of tip, ventral, white arrow points to embolus opening (IBSP 91752); 15 Right palp, bulb, ventral (IBSP 275228); 16 Right palp, bulb, detail of median apophysis, ventral (IBSP 275228)." pageId="10" pageNumber="536">Figs 15–16</figureCitation>
<figureCitation id="130A2A04FFEB8C02ED8AFF50A51170F4" box="[497,625,186,213]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1390,1414]" captionTargetBox="[151,1435,181,1366]" captionTargetId="figure-16@4.[151,1435,181,1366]" captionTargetPageId="4" captionText="FIGURES 13–16. 13–14 P. caete sp. nov.; 15–16 P. cacau sp. nov. 13 Left male palp, ventral (IBSP 91751); 14 Right palp, embolus, detail of tip, ventral, white arrow points to embolus opening (IBSP 91752); 15 Right palp, bulb, ventral (IBSP 275228); 16 Right palp, bulb, detail of median apophysis, ventral (IBSP 275228)." figureDoi="http://doi.org/10.5281/zenodo.14804637" httpUri="https://zenodo.org/record/14804637/files/figure.png" pageId="10" pageNumber="536">Figs 15–16</figureCitation>
,
<figureCitation id="130A2A04FFEB8C02EE07FF50A5FA70F4" box="[636,666,187,213]" captionStart="FIGURES 36–40" captionStartId="11.[151,264,1815,1839]" captionTargetBox="[195,1392,576,1779]" captionTargetId="figure-193@11.[180,1408,561,1792]" captionTargetPageId="11" captionText="FIGURES 36–40. P. cacau sp. nov. 36–38 Male (IBSP 47114), left palp (36 prolateral, 37 ventral, 38 retrolateral); 39–40 Female (IBSP 275186), genitalia (39 epigyne, ventral, 40 vulva, dorsal). Scale lines: 1mm." pageId="10" pageNumber="536">37</figureCitation>
<figureCitation id="130A2A04FFEB8C02EE07FF50A5FA70F4" box="[636,666,187,213]" captionStart="FIGURES 36–40" captionStartId="11.[151,264,1815,1839]" captionTargetBox="[195,1392,576,1779]" captionTargetId="figure-193@11.[180,1408,561,1792]" captionTargetPageId="11" captionText="FIGURES 36–40. P. cacau sp. nov. 36–38 Male (IBSP 47114), left palp (36 prolateral, 37 ventral, 38 retrolateral); 39–40 Female (IBSP 275186), genitalia (39 epigyne, ventral, 40 vulva, dorsal). Scale lines: 1mm." figureDoi="http://doi.org/10.5281/zenodo.14804651" httpUri="https://zenodo.org/record/14804651/files/figure.png" pageId="10" pageNumber="536">37</figureCitation>
,
<figureCitation id="130A2A04FFEB8C02EEDEFF50A5A270F4" box="[677,706,187,213]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="10" pageNumber="536">42</figureCitation>
<figureCitation id="130A2A04FFEB8C02EEDEFF50A5A270F4" box="[677,706,187,213]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="10" pageNumber="536">42</figureCitation>
) (cup-shaped, with strongly indented margin
<taxonomicName id="4C314D02FFEB8C02E8C6FF50A25470F4" authorityName="Rheims" authorityYear="2025" box="[1213,1332,187,213]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="536" phylum="Arthropoda" rank="species" species="cabruca" status="sp. nov.">
<emphasis id="B945EA93FFEB8C02E8C6FF50A25470F4" box="[1213,1332,187,213]" italics="true" pageId="10" pageNumber="536">P. cabruca</emphasis>
@ -358,9 +359,9 @@ resemble those of
<taxonomicNameLabel id="A27657E8FFEB8C02E941FF50A2F370F4" box="[1338,1427,187,213]" pageId="10" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and by the epigyne with LL with posterior margin smooth and MS surpassing the posterior margin of LL (
<figureCitation id="130A2A04FFEB8C02E93BFF34A2F770D8" box="[1344,1431,223,249]" captionStart="FIGURES 36–40" captionStartId="11.[151,264,1815,1839]" captionTargetBox="[195,1392,576,1779]" captionTargetId="figure-193@11.[180,1408,561,1792]" captionTargetPageId="11" captionText="FIGURES 36–40. P. cacau sp. nov. 36–38 Male (IBSP 47114), left palp (36 prolateral, 37 ventral, 38 retrolateral); 39–40 Female (IBSP 275186), genitalia (39 epigyne, ventral, 40 vulva, dorsal). Scale lines: 1mm." pageId="10" pageNumber="536">Figs 39</figureCitation>
<figureCitation id="130A2A04FFEB8C02E93BFF34A2F770D8" box="[1344,1431,223,249]" captionStart="FIGURES 36–40" captionStartId="11.[151,264,1815,1839]" captionTargetBox="[195,1392,576,1779]" captionTargetId="figure-193@11.[180,1408,561,1792]" captionTargetPageId="11" captionText="FIGURES 36–40. P. cacau sp. nov. 36–38 Male (IBSP 47114), left palp (36 prolateral, 37 ventral, 38 retrolateral); 39–40 Female (IBSP 275186), genitalia (39 epigyne, ventral, 40 vulva, dorsal). Scale lines: 1mm." figureDoi="http://doi.org/10.5281/zenodo.14804651" httpUri="https://zenodo.org/record/14804651/files/figure.png" pageId="10" pageNumber="536">Figs 39</figureCitation>
,
<figureCitation id="130A2A04FFEB8C02ECECFEE8A7D4713C" box="[151,180,259,285]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="10" pageNumber="536">44</figureCitation>
<figureCitation id="130A2A04FFEB8C02ECECFEE8A7D4713C" box="[151,180,259,285]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="10" pageNumber="536">44</figureCitation>
) (LL with posterior triangular projections and MS not surpassing the posterior margin of LL in
<taxonomicName id="4C314D02FFEB8C02E881FEE8A213713C" authorityName="Rheims" authorityYear="2025" box="[1274,1395,259,285]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="536" phylum="Arthropoda" rank="species" species="cabruca" status="sp. nov.">
<emphasis id="B945EA93FFEB8C02E881FEE8A213713C" box="[1274,1395,259,285]" italics="true" pageId="10" pageNumber="536">P. cabruca</emphasis>
@ -371,7 +372,7 @@ resemble those of
).
</paragraph>
</subSubSection>
<caption id="DF4E6609FFEB8C02ECECF863A57977E5" pageId="10" pageNumber="536" startId="10.[151,264,1928,1952]" targetBox="[154,1424,395,1890]" targetPageId="10" targetType="figure">
<caption id="DF4E6609FFEB8C02ECECF863A57977E5" ID-DOI="http://doi.org/10.5281/zenodo.14804647" ID-Zenodo-Dep="14804647" httpUri="https://zenodo.org/record/14804647/files/figure.png" pageId="10" pageNumber="536" startId="10.[151,264,1928,1952]" targetBox="[154,1424,395,1890]" targetPageId="10" targetType="figure">
<paragraph id="8B8E3681FFEB8C02ECECF863A57977E5" blockId="10.[151,1436,1928,1989]" pageId="10" pageNumber="536">
<emphasis id="B945EA93FFEB8C02ECECF863A6347781" bold="true" box="[151,340,1928,1952]" pageId="10" pageNumber="536">FIGURES 32–35.</emphasis>
<taxonomicName id="4C314D02FFEB8C02ED21F862A6CF7781" authorityName="Rheims" authorityYear="2025" box="[346,431,1929,1952]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="536" phylum="Arthropoda" rank="species" species="cacau" status="sp. nov.">
@ -391,18 +392,18 @@ Description. Male (
):
</emphasis>
Coloration as in
<figureCitation id="130A2A04FFEA8C03EE95FF7CA4107090" box="[750,880,151,177]" captionStart="FIGURES 32–35" captionStartId="10.[151,264,1928,1952]" captionTargetBox="[154,1424,395,1890]" captionTargetId="figure-95@10.[152,1434,381,1905]" captionTargetPageId="10" captionText="FIGURES 32–35. P. cacau sp. nov., habitus. 32–33 Male, IBSP 47114 (32 dorsal, 33 ventral); 34–35 Female, IBSP 47106 (34 dorsal, 35 ventral). Scale lines: 2 mm." pageId="11" pageNumber="537">Figs 32–33</figureCitation>
<figureCitation id="130A2A04FFEA8C03EE95FF7CA4107090" box="[750,880,151,177]" captionStart="FIGURES 32–35" captionStartId="10.[151,264,1928,1952]" captionTargetBox="[154,1424,395,1890]" captionTargetId="figure-95@10.[152,1434,381,1905]" captionTargetPageId="10" captionText="FIGURES 32–35. P. cacau sp. nov., habitus. 32–33 Male, IBSP 47114 (32 dorsal, 33 ventral); 34–35 Female, IBSP 47106 (34 dorsal, 35 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804647" httpUri="https://zenodo.org/record/14804647/files/figure.png" pageId="11" pageNumber="537">Figs 32–33</figureCitation>
. Total length 10.3. Dorsal shield of prosoma: 4.7 long, 3.8 wide. Opisthosoma: 5.1 long, 2.9 wide. Eyes: diameters:0.35, 0.27, 0.22, 0.26; interdistances: 0.24, 0.10, 0.38, 0.39, 0.15, 0.09. Legs (1423): I: 15.0 (4.5, 2.3, 4.0, 3.3, 0.9); II: 14.7 (4.6, 2.3, 3.8, 3.1, 0.9); III: 12.3 (3.9, 1.9, 2.9, 2.8, 0.8); IV: 14.6 (4.4, 1.6, 3.5, 4.0, 1.1). Spination follows the generic pattern, with tibia I–II bearing six pairs of ventral spines. Palp: VTA small, triangular; RTA single branched, laminar, resembling a dog’s head in retrolateral view and curled ventrally (
<figureCitation id="130A2A04FFEA8C03EE30FEA0A5C07144" box="[587,672,331,357]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." pageId="11" pageNumber="537">Fig. 27</figureCitation>
<figureCitation id="130A2A04FFEA8C03EE30FEA0A5C07144" box="[587,672,331,357]" captionStart="FIGURES 26–31" captionStartId="9.[151,264,1450,1474]" captionTargetBox="[180,1415,186,1411]" captionTargetId="figure-20@9.[162,1425,181,1425]" captionTargetPageId="9" captionText="FIGURES 26–31. P. cabruca sp. nov. 26–28 Male (IBSP 47105), left palp (26 prolateral, 27 ventral, 28 retrolateral); 29–31 Female (IBSP 41116) genitalia (29 epigyne dorsal, 30 vulva ventral, 31 schematic course of IDS). Scale lines 1 mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projections close to conductor base; TBE—tegular projection close to embolus base." figureDoi="http://doi.org/10.5281/zenodo.14804645" httpUri="https://zenodo.org/record/14804645/files/figure.png" pageId="11" pageNumber="537">Fig. 27</figureCitation>
); ST visible between 3–4 o’clock in retrolateral view; spermophore with retrolateral undulation (between 3–4 o’clock); TBE long, with an acute point; TBC laminar, three times longer than wide, slightly pointed at tip; C hood four times longer than wide, tip curled, ventrally resembling a beanie; E arising from tegulum at 7 o’clock position, narrowing subdistally to a filiform tip, with a filiform subdistal projection; MA knob-like (
<figureCitation id="130A2A04FFEA8C03EDBFFE30A52571D4" box="[452,581,474,501]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1390,1414]" captionTargetBox="[151,1435,181,1366]" captionTargetId="figure-16@4.[151,1435,181,1366]" captionTargetPageId="4" captionText="FIGURES 13–16. 13–14 P. caete sp. nov.; 15–16 P. cacau sp. nov. 13 Left male palp, ventral (IBSP 91751); 14 Right palp, embolus, detail of tip, ventral, white arrow points to embolus opening (IBSP 91752); 15 Right palp, bulb, ventral (IBSP 275228); 16 Right palp, bulb, detail of median apophysis, ventral (IBSP 275228)." pageId="11" pageNumber="537">Figs 15–16</figureCitation>
<figureCitation id="130A2A04FFEA8C03EDBFFE30A52571D4" box="[452,581,474,501]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1390,1414]" captionTargetBox="[151,1435,181,1366]" captionTargetId="figure-16@4.[151,1435,181,1366]" captionTargetPageId="4" captionText="FIGURES 13–16. 13–14 P. caete sp. nov.; 15–16 P. cacau sp. nov. 13 Left male palp, ventral (IBSP 91751); 14 Right palp, embolus, detail of tip, ventral, white arrow points to embolus opening (IBSP 91752); 15 Right palp, bulb, ventral (IBSP 275228); 16 Right palp, bulb, detail of median apophysis, ventral (IBSP 275228)." figureDoi="http://doi.org/10.5281/zenodo.14804637" httpUri="https://zenodo.org/record/14804637/files/figure.png" pageId="11" pageNumber="537">Figs 15–16</figureCitation>
,
<figureCitation id="130A2A04FFEA8C03EE2AFE30A5F971D4" box="[593,665,475,501]" captionStart="FIGURES 36–40" captionStartId="11.[151,264,1815,1839]" captionTargetBox="[195,1392,576,1779]" captionTargetId="figure-193@11.[180,1408,561,1792]" captionTargetPageId="11" captionText="FIGURES 36–40. P. cacau sp. nov. 36–38 Male (IBSP 47114), left palp (36 prolateral, 37 ventral, 38 retrolateral); 39–40 Female (IBSP 275186), genitalia (39 epigyne, ventral, 40 vulva, dorsal). Scale lines: 1mm." pageId="11" pageNumber="537">36–38</figureCitation>
<figureCitation id="130A2A04FFEA8C03EE2AFE30A5F971D4" box="[593,665,475,501]" captionStart="FIGURES 36–40" captionStartId="11.[151,264,1815,1839]" captionTargetBox="[195,1392,576,1779]" captionTargetId="figure-193@11.[180,1408,561,1792]" captionTargetPageId="11" captionText="FIGURES 36–40. P. cacau sp. nov. 36–38 Male (IBSP 47114), left palp (36 prolateral, 37 ventral, 38 retrolateral); 39–40 Female (IBSP 275186), genitalia (39 epigyne, ventral, 40 vulva, dorsal). Scale lines: 1mm." figureDoi="http://doi.org/10.5281/zenodo.14804651" httpUri="https://zenodo.org/record/14804651/files/figure.png" pageId="11" pageNumber="537">36–38</figureCitation>
,
<figureCitation id="130A2A04FFEA8C03EEDEFE30A59071D4" box="[677,752,475,501]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="11" pageNumber="537">41–43</figureCitation>
<figureCitation id="130A2A04FFEA8C03EEDEFE30A59071D4" box="[677,752,475,501]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="11" pageNumber="537">41–43</figureCitation>
).
</paragraph>
<caption id="DF4E6609FFEA8C03ECECF8FCA34F7775" pageId="11" pageNumber="537" startId="11.[151,264,1815,1839]" targetBox="[195,1392,576,1779]" targetPageId="11" targetType="figure">
<caption id="DF4E6609FFEA8C03ECECF8FCA34F7775" ID-DOI="http://doi.org/10.5281/zenodo.14804651" ID-Zenodo-Dep="14804651" httpUri="https://zenodo.org/record/14804651/files/figure.png" pageId="11" pageNumber="537" startId="11.[151,264,1815,1839]" targetBox="[195,1392,576,1779]" targetPageId="11" targetType="figure">
<paragraph id="8B8E3681FFEA8C03ECECF8FCA34F7775" blockId="11.[151,1437,1815,1876]" pageId="11" pageNumber="537">
<emphasis id="B945EA93FFEA8C03ECECF8FCA637770E" bold="true" box="[151,343,1815,1839]" pageId="11" pageNumber="537">FIGURES 36–40.</emphasis>
<taxonomicName id="4C314D02FFEA8C03ED1BF8F3A6D7770E" authorityName="Rheims" authorityYear="2025" box="[352,439,1816,1839]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="537" phylum="Arthropoda" rank="species" species="cacau" status="sp. nov.">
@ -421,18 +422,18 @@ Female (IBSP 47112,
):
</emphasis>
Coloration as in
<figureCitation id="130A2A04FFEA8C03EE84F84DA41F77E0" box="[767,895,1958,1985]" captionStart="FIGURES 32–35" captionStartId="10.[151,264,1928,1952]" captionTargetBox="[154,1424,395,1890]" captionTargetId="figure-95@10.[152,1434,381,1905]" captionTargetPageId="10" captionText="FIGURES 32–35. P. cacau sp. nov., habitus. 32–33 Male, IBSP 47114 (32 dorsal, 33 ventral); 34–35 Female, IBSP 47106 (34 dorsal, 35 ventral). Scale lines: 2 mm." pageId="11" pageNumber="537">Figs 34–35</figureCitation>
<figureCitation id="130A2A04FFEA8C03EE84F84DA41F77E0" box="[767,895,1958,1985]" captionStart="FIGURES 32–35" captionStartId="10.[151,264,1928,1952]" captionTargetBox="[154,1424,395,1890]" captionTargetId="figure-95@10.[152,1434,381,1905]" captionTargetPageId="10" captionText="FIGURES 32–35. P. cacau sp. nov., habitus. 32–33 Male, IBSP 47114 (32 dorsal, 33 ventral); 34–35 Female, IBSP 47106 (34 dorsal, 35 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804647" httpUri="https://zenodo.org/record/14804647/files/figure.png" pageId="11" pageNumber="537">Figs 34–35</figureCitation>
. Total length 13.4. Dorsal shield of prosoma: 5.1 long, 4.1 wide. Opisthosoma: 8.0 long, 4.9 wide. Eyes: diameters: 0.37, 0.30, 0.20, 0.30; interdistances: 0.25, 0.14,
</paragraph>
<paragraph id="8B8E3681FFED8C04ECECFF7CA5507144" blockId="12.[151,1437,150,465]" pageId="12" pageNumber="538">
0.46, 0.45, 0.20, 0.10. Legs (4123): I: 14.2 (4.2, 2.5, 3.8, 2.8, 0.9); II: 13.8 (4.2, 2.4, 3.5, 2.8, 0.9); III: 11.8 (3.8, 2.0, 2.6, 2.5, 0.9); IV: 14.9 (4.5, 1.8, 3.4, 4.0, 1.2). Spination follows the generic pattern with tibiae I–II as in male. Epigyne: EF roughly as long as wide, widest posteriorly, with one long lateral MAB on each side; LL posteriorly smooth, rounded; MS as long as wide; ag forming a triangular elevated area over the entire length of MS (
<figureCitation id="130A2A04FFED8C04E93BFEE8A2F7713C" box="[1344,1431,259,285]" captionStart="FIGURES 36–40" captionStartId="11.[151,264,1815,1839]" captionTargetBox="[195,1392,576,1779]" captionTargetId="figure-193@11.[180,1408,561,1792]" captionTargetPageId="11" captionText="FIGURES 36–40. P. cacau sp. nov. 36–38 Male (IBSP 47114), left palp (36 prolateral, 37 ventral, 38 retrolateral); 39–40 Female (IBSP 275186), genitalia (39 epigyne, ventral, 40 vulva, dorsal). Scale lines: 1mm." pageId="12" pageNumber="538">Figs 39</figureCitation>
<figureCitation id="130A2A04FFED8C04E93BFEE8A2F7713C" box="[1344,1431,259,285]" captionStart="FIGURES 36–40" captionStartId="11.[151,264,1815,1839]" captionTargetBox="[195,1392,576,1779]" captionTargetId="figure-193@11.[180,1408,561,1792]" captionTargetPageId="11" captionText="FIGURES 36–40. P. cacau sp. nov. 36–38 Male (IBSP 47114), left palp (36 prolateral, 37 ventral, 38 retrolateral); 39–40 Female (IBSP 275186), genitalia (39 epigyne, ventral, 40 vulva, dorsal). Scale lines: 1mm." figureDoi="http://doi.org/10.5281/zenodo.14804651" httpUri="https://zenodo.org/record/14804651/files/figure.png" pageId="12" pageNumber="538">Figs 39</figureCitation>
,
<figureCitation id="130A2A04FFED8C04ECECFECCA7D77160" box="[151,183,295,321]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="12" pageNumber="538">44</figureCitation>
<figureCitation id="130A2A04FFED8C04ECECFECCA7D77160" box="[151,183,295,321]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="12" pageNumber="538">44</figureCitation>
). Vulva: IDS with Dh kidney-shaped connecting ventrally with Ds by long sclerotized, narrow duct; Ds roughly oval; FD posteriad (
<figureCitation id="130A2A04FFED8C04ED0DFEA0A6AE7144" box="[374,462,331,357]" captionStart="FIGURES 36–40" captionStartId="11.[151,264,1815,1839]" captionTargetBox="[195,1392,576,1779]" captionTargetId="figure-193@11.[180,1408,561,1792]" captionTargetPageId="11" captionText="FIGURES 36–40. P. cacau sp. nov. 36–38 Male (IBSP 47114), left palp (36 prolateral, 37 ventral, 38 retrolateral); 39–40 Female (IBSP 275186), genitalia (39 epigyne, ventral, 40 vulva, dorsal). Scale lines: 1mm." pageId="12" pageNumber="538">Figs 40</figureCitation>
<figureCitation id="130A2A04FFED8C04ED0DFEA0A6AE7144" box="[374,462,331,357]" captionStart="FIGURES 36–40" captionStartId="11.[151,264,1815,1839]" captionTargetBox="[195,1392,576,1779]" captionTargetId="figure-193@11.[180,1408,561,1792]" captionTargetPageId="11" captionText="FIGURES 36–40. P. cacau sp. nov. 36–38 Male (IBSP 47114), left palp (36 prolateral, 37 ventral, 38 retrolateral); 39–40 Female (IBSP 275186), genitalia (39 epigyne, ventral, 40 vulva, dorsal). Scale lines: 1mm." figureDoi="http://doi.org/10.5281/zenodo.14804651" httpUri="https://zenodo.org/record/14804651/files/figure.png" pageId="12" pageNumber="538">Figs 40</figureCitation>
,
<figureCitation id="130A2A04FFED8C04EDA2FEA1A5447144" box="[473,548,330,357]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." pageId="12" pageNumber="538">45–46</figureCitation>
<figureCitation id="130A2A04FFED8C04EDA2FEA1A5447144" box="[473,548,330,357]" captionStart="FIGURES 41–46" captionStartId="12.[151,264,1746,1770]" captionTargetBox="[184,1396,501,1710]" captionTargetId="figure-167@12.[181,1406,489,1722]" captionTargetPageId="12" captionText="FIGURES 41–46. P. cacau sp. nov. 41–43 Male (IBSP 47109), left palp (41 prolateral, 42 ventral, 43 retrolateral); 44–46 Female (IBSP 47112), genitalia (44 epigyne ventral, 45 vulva dorsal, 46 schematic course of IDS). Scale lines: 1mm. ag— anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment bands; MS— median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; SeP—subapical embolic projection; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to embolus base; VTA— ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804653" httpUri="https://zenodo.org/record/14804653/files/figure.png" pageId="12" pageNumber="538">45–46</figureCitation>
).
</paragraph>
<paragraph id="8B8E3681FFED8C04ECBCFE85A37A718C" blockId="12.[151,1437,150,465]" pageId="12" pageNumber="538">
@ -450,7 +451,7 @@ and
, northeastern
<collectingCountry id="F3267611FFED8C04E855FE5CA31271F0" box="[1070,1138,439,465]" name="Brazil" pageId="12" pageNumber="538">Brazil</collectingCountry>
(
<figureCitation id="130A2A04FFED8C04E8F9FE5CA3B671F0" box="[1154,1238,439,465]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." pageId="12" pageNumber="538">Fig. 79</figureCitation>
<figureCitation id="130A2A04FFED8C04E8F9FE5CA3B671F0" box="[1154,1238,439,465]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14804675" httpUri="https://zenodo.org/record/14804675/files/figure.png" pageId="12" pageNumber="538">Fig. 79</figureCitation>
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103
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@ -1,43 +1,44 @@
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@ -54,13 +55,13 @@
</subSubSection>
<subSubSection id="C32B650AFFEC8C05ECECFF50A56270F4" box="[151,514,187,214]" pageId="13" pageNumber="539" type="description">
<paragraph id="8B8E3681FFEC8C05ECECFF50A56270F4" blockId="13.[151,514,151,214]" box="[151,514,187,214]" pageId="13" pageNumber="539">
<figureCitation id="130A2A04FFEC8C05ECECFF50A67770F7" box="[151,279,187,214]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." pageId="13" pageNumber="539">Figs 2, 4–5</figureCitation>
<figureCitation id="130A2A04FFEC8C05ECECFF50A67770F7" box="[151,279,187,214]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-16@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 3 P. cabruca sp. nov. (male, IBSP 47126); 2, 4–5 P. caete sp. nov. (male, IBSP 234826); 6 P. cacau sp. nov. (female, IBSP 275471). 1 Left chelicerae, ventral; 2 Left endite, serrula; 3 Left leg I, metatarsus, trilobate membrane, dorsal; 4 Left leg I, tarsus, tricobothrium; 5 Left leg I, tarsus, tarsal organ; 6 Left palp, tarsus, claw, prolateral." figureDoi="http://doi.org/10.5281/zenodo.14804631" httpUri="https://zenodo.org/record/14804631/files/figure.png" pageId="13" pageNumber="539">Figs 2, 4–5</figureCitation>
,
<figureCitation id="130A2A04FFEC8C05ED59FF50A6E770F4" box="[290,391,187,213]" captionStart-0="FIGURES 7–12" captionStart-1="FIGURES 13–16" captionStartId-0="3.[151,264,1943,1967]" captionStartId-1="4.[151,264,1390,1414]" captionTargetBox-0="[167,1419,181,1919]" captionTargetBox-1="[151,1435,181,1366]" captionTargetId-0="figure-20@3.[167,1419,181,1919]" captionTargetId-1="figure-16@4.[151,1435,181,1366]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 7–12. 7, 9 P. cabruca sp. nov. (male, IBSP 47126); 8, 12 P. caete sp. nov. (male, IBSP 249602); 10–11 P. cacau sp. nov. (10 male IBSP 275288, 11 female IBSP 275471). 7–10 Tarsus claws, left leg, prolateral (7 leg I, 8 leg II, 9 leg III, 10 leg IV); 11 Opisthosoma, epiandrium, epiandrous spigots, ventral; 12 Opisthosoma colulus, ventral." captionText-1="FIGURES 13–16. 13–14 P. caete sp. nov.; 15–16 P. cacau sp. nov. 13 Left male palp, ventral (IBSP 91751); 14 Right palp, embolus, detail of tip, ventral, white arrow points to embolus opening (IBSP 91752); 15 Right palp, bulb, ventral (IBSP 275228); 16 Right palp, bulb, detail of median apophysis, ventral (IBSP 275228)." pageId="13" pageNumber="539">8, 12–14</figureCitation>
<figureCitation id="130A2A04FFEC8C05ED59FF50A6E770F4" box="[290,391,187,213]" captionStart-0="FIGURES 7–12" captionStart-1="FIGURES 13–16" captionStartId-0="3.[151,264,1943,1967]" captionStartId-1="4.[151,264,1390,1414]" captionTargetBox-0="[167,1419,181,1919]" captionTargetBox-1="[151,1435,181,1366]" captionTargetId-0="figure-20@3.[167,1419,181,1919]" captionTargetId-1="figure-16@4.[151,1435,181,1366]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 7–12. 7, 9 P. cabruca sp. nov. (male, IBSP 47126); 8, 12 P. caete sp. nov. (male, IBSP 249602); 10–11 P. cacau sp. nov. (10 male IBSP 275288, 11 female IBSP 275471). 7–10 Tarsus claws, left leg, prolateral (7 leg I, 8 leg II, 9 leg III, 10 leg IV); 11 Opisthosoma, epiandrium, epiandrous spigots, ventral; 12 Opisthosoma colulus, ventral." captionText-1="FIGURES 13–16. 13–14 P. caete sp. nov.; 15–16 P. cacau sp. nov. 13 Left male palp, ventral (IBSP 91751); 14 Right palp, embolus, detail of tip, ventral, white arrow points to embolus opening (IBSP 91752); 15 Right palp, bulb, ventral (IBSP 275228); 16 Right palp, bulb, detail of median apophysis, ventral (IBSP 275228)." figureDoi-0="http://doi.org/10.5281/zenodo.14804635" figureDoi-1="http://doi.org/10.5281/zenodo.14804637" httpUri-0="https://zenodo.org/record/14804635/files/figure.png" httpUri-1="https://zenodo.org/record/14804637/files/figure.png" pageId="13" pageNumber="539">8, 12–14</figureCitation>
,
<figureCitation id="130A2A04FFEC8C05EDE9FF50A6BB70F4" box="[402,475,187,213]" captionStart-0="FIGURES 47–50" captionStart-1="FIGURES 51–55" captionStart-2="FIGURES 56–62" captionStartId-0="13.[151,264,1967,1991]" captionStartId-1="14.[151,264,1345,1369]" captionStartId-2="15.[151,260,1836,1860]" captionTargetBox-0="[218,1372,646,1931]" captionTargetBox-1="[169,1413,195,1307]" captionTargetBox-2="[165,1420,701,1803]" captionTargetId-0="figure-183@13.[203,1384,633,1944]" captionTargetId-1="figure-16@14.[158,1428,181,1320]" captionTargetId-2="figure-252@15.[158,1430,698,1811]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="FIGURES 47–50. P. caete sp. nov., habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm." captionText-1="FIGURES 51–55. P. caete sp. nov. 51–53 Male (IBSP 135153), left palp (51 prolateral, 52 ventral, 53 retrolateral); 54–55 Female (IBSP 234826), genitalia (54 epigyne ventral, 55 vulva dorsal). Scale lines: 1 mm." captionText-2="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="13" pageNumber="539">47–62</figureCitation>
<figureCitation id="130A2A04FFEC8C05EDE9FF50A6BB70F4" box="[402,475,187,213]" captionStart-0="FIGURES 47–50" captionStart-1="FIGURES 51–55" captionStart-2="FIGURES 56–62" captionStartId-0="13.[151,264,1967,1991]" captionStartId-1="14.[151,264,1345,1369]" captionStartId-2="15.[151,260,1836,1860]" captionTargetBox-0="[218,1372,646,1931]" captionTargetBox-1="[169,1413,195,1307]" captionTargetBox-2="[165,1420,701,1803]" captionTargetId-0="figure-183@13.[203,1384,633,1944]" captionTargetId-1="figure-16@14.[158,1428,181,1320]" captionTargetId-2="figure-252@15.[158,1430,698,1811]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="FIGURES 47–50. P. caete sp. nov., habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm." captionText-1="FIGURES 51–55. P. caete sp. nov. 51–53 Male (IBSP 135153), left palp (51 prolateral, 52 ventral, 53 retrolateral); 54–55 Female (IBSP 234826), genitalia (54 epigyne ventral, 55 vulva dorsal). Scale lines: 1 mm." captionText-2="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi-0="http://doi.org/10.5281/zenodo.14804657" figureDoi-1="http://doi.org/10.5281/zenodo.14804659" figureDoi-2="http://doi.org/10.5281/zenodo.14804663" httpUri-0="https://zenodo.org/record/14804657/files/figure.png" httpUri-1="https://zenodo.org/record/14804659/files/figure.png" httpUri-2="https://zenodo.org/record/14804663/files/figure.png" pageId="13" pageNumber="539">47–62</figureCitation>
,
<figureCitation id="130A2A04FFEC8C05ED9DFF50A56270F4" box="[486,514,187,213]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." pageId="13" pageNumber="539">80</figureCitation>
<figureCitation id="130A2A04FFEC8C05ED9DFF50A56270F4" box="[486,514,187,213]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14804675" httpUri="https://zenodo.org/record/14804675/files/figure.png" pageId="13" pageNumber="539">80</figureCitation>
</paragraph>
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@ -252,7 +253,7 @@ leg. (
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.
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<paragraph id="8B8E3681FFEC8C05ECECF844A52177CA" blockId="13.[151,1437,1967,2028]" pageId="13" pageNumber="539">
<emphasis id="B945EA93FFEC8C05ECECF844A63577E6" bold="true" box="[151,341,1967,1991]" pageId="13" pageNumber="539">FIGURES 47–50.</emphasis>
<taxonomicName id="4C314D02FFEC8C05ED27F85BA6CB77E6" authorityName="Rheims" authorityYear="2025" box="[348,427,1968,1991]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="539" phylum="Arthropoda" rank="species" species="caete" status="sp. nov.">
@ -264,7 +265,7 @@ leg. (
, habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm.
</paragraph>
</caption>
<caption id="DF4E6609FFEF8C06ECECFAAAA349755D" pageId="14" pageNumber="540" startId="14.[151,264,1345,1369]" targetBox="[169,1413,195,1307]" targetPageId="14" targetType="figure">
<caption id="DF4E6609FFEF8C06ECECFAAAA349755D" ID-DOI="http://doi.org/10.5281/zenodo.14804659" ID-Zenodo-Dep="14804659" httpUri="https://zenodo.org/record/14804659/files/figure.png" pageId="14" pageNumber="540" startId="14.[151,264,1345,1369]" targetBox="[169,1413,195,1307]" targetPageId="14" targetType="figure">
<paragraph id="8B8E3681FFEF8C06ECECFAAAA349755D" blockId="14.[151,1436,1345,1405]" pageId="14" pageNumber="540">
<emphasis id="B945EA93FFEF8C06ECECFAAAA6377578" bold="true" box="[151,343,1345,1369]" pageId="14" pageNumber="540">FIGURES 51–55.</emphasis>
<taxonomicName id="4C314D02FFEF8C06ED24FAA9A6CF7578" authorityName="Rheims" authorityYear="2025" box="[351,431,1346,1369]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="caete" status="sp. nov.">
@ -801,7 +802,7 @@ leg. (
.
</paragraph>
</subSubSection>
<caption id="DF4E6609FFEE8C07ECECF8C7A2F377D9" pageId="15" pageNumber="541" startId="15.[151,260,1836,1860]" targetBox="[165,1420,701,1803]" targetPageId="15" targetType="figure">
<caption id="DF4E6609FFEE8C07ECECF8C7A2F377D9" ID-DOI="http://doi.org/10.5281/zenodo.14804663" ID-Zenodo-Dep="14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="15" pageNumber="541" startId="15.[151,260,1836,1860]" targetBox="[165,1420,701,1803]" targetPageId="15" targetType="figure">
<paragraph id="8B8E3681FFEE8C07ECECF8C7A2F377D9" blockId="15.[151,1437,1836,2040]" pageId="15" pageNumber="541">
<emphasis id="B945EA93FFEE8C07ECECF8C7A62C7765" bold="true" box="[151,332,1836,1860]" pageId="15" pageNumber="541">FIGURES 56–62.</emphasis>
<taxonomicName id="4C314D02FFEE8C07ED2AF8C6A6FA7765" authorityName="Rheims" authorityYear="2025" box="[337,410,1837,1860]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="caete" status="sp. nov.">
@ -837,13 +838,13 @@ resemble those
<taxonomicNameLabel id="A27657E8FFF18C18EFA4FEE8A359713C" box="[991,1081,259,285]" pageId="16" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="130A2A04FFF18C18E833FEE8A3A8713C" box="[1096,1224,259,285]" captionStart-0="FIGURES 67–71" captionStart-1="FIGURES 72–77" captionStartId-0="18.[151,264,1441,1465]" captionStartId-1="19.[151,264,1677,1701]" captionTargetBox-0="[166,1421,219,1407]" captionTargetBox-1="[172,1413,494,1645]" captionTargetId-0="figure-16@18.[152,1435,201,1418]" captionTargetId-1="figure-134@19.[159,1428,489,1654]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." captionText-1="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="16" pageNumber="542">Figs 67–77</figureCitation>
<figureCitation id="130A2A04FFF18C18E833FEE8A3A8713C" box="[1096,1224,259,285]" captionStart-0="FIGURES 67–71" captionStart-1="FIGURES 72–77" captionStartId-0="18.[151,264,1441,1465]" captionStartId-1="19.[151,264,1677,1701]" captionTargetBox-0="[166,1421,219,1407]" captionTargetBox-1="[172,1413,494,1645]" captionTargetId-0="figure-16@18.[152,1435,201,1418]" captionTargetId-1="figure-134@19.[159,1428,489,1654]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." captionText-1="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi-0="http://doi.org/10.5281/zenodo.14804669" figureDoi-1="http://doi.org/10.5281/zenodo.14804671" httpUri-0="https://zenodo.org/record/14804669/files/figure.png" httpUri-1="https://zenodo.org/record/14804671/files/figure.png" pageId="16" pageNumber="542">Figs 67–77</figureCitation>
) by the male palps with RTA bearing more than one branch (single branched in the other species) and by the female epigyne with ag straight or slightly recurved, not projecting over the median septum (projecting in the other species). Males are distinguished from those of the latter species by the RTA with three branches (
<figureCitation id="130A2A04FFF18C18E86FFE84A30671A8" box="[1044,1126,366,393]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="16" pageNumber="542">Fig. 58</figureCitation>
<figureCitation id="130A2A04FFF18C18E86FFE84A30671A8" box="[1044,1126,366,393]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="16" pageNumber="542">Fig. 58</figureCitation>
) and embolus tip complex, with three projections (
<figureCitation id="130A2A04FFF18C18EDD8FE78A69C718C" box="[419,508,403,429]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1390,1414]" captionTargetBox="[151,1435,181,1366]" captionTargetId="figure-16@4.[151,1435,181,1366]" captionTargetPageId="4" captionText="FIGURES 13–16. 13–14 P. caete sp. nov.; 15–16 P. cacau sp. nov. 13 Left male palp, ventral (IBSP 91751); 14 Right palp, embolus, detail of tip, ventral, white arrow points to embolus opening (IBSP 91752); 15 Right palp, bulb, ventral (IBSP 275228); 16 Right palp, bulb, detail of median apophysis, ventral (IBSP 275228)." pageId="16" pageNumber="542">Figs 14</figureCitation>
<figureCitation id="130A2A04FFF18C18EDD8FE78A69C718C" box="[419,508,403,429]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1390,1414]" captionTargetBox="[151,1435,181,1366]" captionTargetId="figure-16@4.[151,1435,181,1366]" captionTargetPageId="4" captionText="FIGURES 13–16. 13–14 P. caete sp. nov.; 15–16 P. cacau sp. nov. 13 Left male palp, ventral (IBSP 91751); 14 Right palp, embolus, detail of tip, ventral, white arrow points to embolus opening (IBSP 91752); 15 Right palp, bulb, ventral (IBSP 275228); 16 Right palp, bulb, detail of median apophysis, ventral (IBSP 275228)." figureDoi="http://doi.org/10.5281/zenodo.14804637" httpUri="https://zenodo.org/record/14804637/files/figure.png" pageId="16" pageNumber="542">Figs 14</figureCitation>
,
<figureCitation id="130A2A04FFF18C18EE70FE79A548718C" box="[523,552,402,429]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="16" pageNumber="542">59</figureCitation>
<figureCitation id="130A2A04FFF18C18EE70FE79A548718C" box="[523,552,402,429]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="16" pageNumber="542">59</figureCitation>
) (RTA with two branches and embolus tip filiform in
<taxonomicName id="4C314D02FFF18C18E8E4FE78A246718D" authorityName="Rheims" authorityYear="2025" box="[1183,1318,403,428]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="542" phylum="Arthropoda" rank="species" species="tacomare" status="sp. nov.">
<emphasis id="B945EA93FFF18C18E8E4FE78A246718D" box="[1183,1318,403,428]" italics="true" pageId="16" pageNumber="542">P. tacomare</emphasis>
@ -852,9 +853,9 @@ resemble those
<taxonomicNameLabel id="A27657E8FFF18C18E94BFE78A2EC718C" box="[1328,1420,403,429]" pageId="16" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). Females are distinguished by the MS longer than wide with posterior margin at level with posterior margin of LL (
<figureCitation id="130A2A04FFF18C18ECE4FE30A79671D4" box="[159,246,474,501]" captionStart="FIGURES 51–55" captionStartId="14.[151,264,1345,1369]" captionTargetBox="[169,1413,195,1307]" captionTargetId="figure-16@14.[158,1428,181,1320]" captionTargetPageId="14" captionText="FIGURES 51–55. P. caete sp. nov. 51–53 Male (IBSP 135153), left palp (51 prolateral, 52 ventral, 53 retrolateral); 54–55 Female (IBSP 234826), genitalia (54 epigyne ventral, 55 vulva dorsal). Scale lines: 1 mm." pageId="16" pageNumber="542">Figs 54</figureCitation>
<figureCitation id="130A2A04FFF18C18ECE4FE30A79671D4" box="[159,246,474,501]" captionStart="FIGURES 51–55" captionStartId="14.[151,264,1345,1369]" captionTargetBox="[169,1413,195,1307]" captionTargetId="figure-16@14.[158,1428,181,1320]" captionTargetPageId="14" captionText="FIGURES 51–55. P. caete sp. nov. 51–53 Male (IBSP 135153), left palp (51 prolateral, 52 ventral, 53 retrolateral); 54–55 Female (IBSP 234826), genitalia (54 epigyne ventral, 55 vulva dorsal). Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14804659" httpUri="https://zenodo.org/record/14804659/files/figure.png" pageId="16" pageNumber="542">Figs 54</figureCitation>
,
<figureCitation id="130A2A04FFF18C18ED79FE30A67F71D4" box="[258,287,475,501]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="16" pageNumber="542">60</figureCitation>
<figureCitation id="130A2A04FFF18C18ED79FE30A67F71D4" box="[258,287,475,501]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="16" pageNumber="542">60</figureCitation>
) (MS wider than long with posterior margin reaching slightly more than half LL length in
<taxonomicName id="4C314D02FFF18C18E963FE30A2FD71D5" authorityName="Rheims" authorityYear="2025" box="[1304,1437,475,500]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="542" phylum="Arthropoda" rank="species" species="tacomare" status="sp. nov.">
<emphasis id="B945EA93FFF18C18E963FE30A2FD71D5" box="[1304,1437,475,500]" italics="true" pageId="16" pageNumber="542">P. tacomare</emphasis>
@ -873,25 +874,25 @@ Description. Male (
):
</emphasis>
Coloration as in
<figureCitation id="130A2A04FFF18C18EE8FFDC8A417721C" box="[756,887,547,573]" captionStart="FIGURES 47–50" captionStartId="13.[151,264,1967,1991]" captionTargetBox="[218,1372,646,1931]" captionTargetId="figure-183@13.[203,1384,633,1944]" captionTargetPageId="13" captionText="FIGURES 47–50. P. caete sp. nov., habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm." pageId="16" pageNumber="542">Figs 47–48</figureCitation>
<figureCitation id="130A2A04FFF18C18EE8FFDC8A417721C" box="[756,887,547,573]" captionStart="FIGURES 47–50" captionStartId="13.[151,264,1967,1991]" captionTargetBox="[218,1372,646,1931]" captionTargetId="figure-183@13.[203,1384,633,1944]" captionTargetPageId="13" captionText="FIGURES 47–50. P. caete sp. nov., habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804657" httpUri="https://zenodo.org/record/14804657/files/figure.png" pageId="16" pageNumber="542">Figs 47–48</figureCitation>
. Total length 8.9. Dorsal shield of prosoma: 4.1 long, 3.4 wide. Opisthosoma: 4.6 long, 2.4 wide. Eyes: diameters: 0.29, 0.25, 0.15, 0.24; interdistances: 0.34, 0.09, 0.48, 0.36, 0.23, 0.12. Legs (1243): I: 13.2 (4.0, 2.0, 3.3, 3.0, 0.9); II: 12.6 (3.9, 2.0, 3.0, 2.9, 0.8, 12.6); III: 10.5 (3.4, 1.7, 2.3, 2.4, 0.7); IV: 13.1 (3.9, 1.5, 3.0, 3.6, 1.1). Spination follows the generic pattern except tibiae I–II: d0-1-0; v2-2-2-2-2. Palp: VTA large triangular; RTA with three acute branches with rounded tips in retrolateral view and sub squared in ventral view; ST visible between 3–5 o’clock in retrolateral view; spermophore with undulation between 11–12 o’clock; TBE hyaline, fanning out at tip; TBC reduced to a small sclerotized elevation; C hood four times longer than wide, tongue-like in ventral view; E arising from tegulum at 7 o’clock position, tip complex bearing a leaf-like projection at the embolus opening, between a long prong-like projection and a slender, laminar projection; MA slightly wider than long, widest distally, with slightly medially depressed margin (
<figureCitation id="130A2A04FFF18C18E81AFC8CA38273A0" box="[1121,1250,870,897]" captionStart="FIGURES 51–55" captionStartId="14.[151,264,1345,1369]" captionTargetBox="[169,1413,195,1307]" captionTargetId="figure-16@14.[158,1428,181,1320]" captionTargetPageId="14" captionText="FIGURES 51–55. P. caete sp. nov. 51–53 Male (IBSP 135153), left palp (51 prolateral, 52 ventral, 53 retrolateral); 54–55 Female (IBSP 234826), genitalia (54 epigyne ventral, 55 vulva dorsal). Scale lines: 1 mm." pageId="16" pageNumber="542">Figs 51–53</figureCitation>
<figureCitation id="130A2A04FFF18C18E81AFC8CA38273A0" box="[1121,1250,870,897]" captionStart="FIGURES 51–55" captionStartId="14.[151,264,1345,1369]" captionTargetBox="[169,1413,195,1307]" captionTargetId="figure-16@14.[158,1428,181,1320]" captionTargetPageId="14" captionText="FIGURES 51–55. P. caete sp. nov. 51–53 Male (IBSP 135153), left palp (51 prolateral, 52 ventral, 53 retrolateral); 54–55 Female (IBSP 234826), genitalia (54 epigyne ventral, 55 vulva dorsal). Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14804659" httpUri="https://zenodo.org/record/14804659/files/figure.png" pageId="16" pageNumber="542">Figs 51–53</figureCitation>
,
<figureCitation id="130A2A04FFF18C18E895FC8DA25873A0" box="[1262,1336,870,897]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="16" pageNumber="542">56–59</figureCitation>
<figureCitation id="130A2A04FFF18C18E895FC8DA25873A0" box="[1262,1336,870,897]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="16" pageNumber="542">56–59</figureCitation>
).
</paragraph>
<paragraph id="8B8E3681FFF18C18ECBCFC60A5E2745C" blockId="16.[151,1437,151,1293]" pageId="16" pageNumber="542">
<emphasis id="B945EA93FFF18C18ECBCFC60A6B77385" bold="true" box="[199,471,906,933]" pageId="16" pageNumber="542">Female (IBSP 135183):</emphasis>
Coloration as in
<figureCitation id="130A2A04FFF18C18EEEDFC60A4767384" box="[662,790,906,933]" captionStart="FIGURES 47–50" captionStartId="13.[151,264,1967,1991]" captionTargetBox="[218,1372,646,1931]" captionTargetId="figure-183@13.[203,1384,633,1944]" captionTargetPageId="13" captionText="FIGURES 47–50. P. caete sp. nov., habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm." pageId="16" pageNumber="542">Figs 49–50</figureCitation>
<figureCitation id="130A2A04FFF18C18EEEDFC60A4767384" box="[662,790,906,933]" captionStart="FIGURES 47–50" captionStartId="13.[151,264,1967,1991]" captionTargetBox="[218,1372,646,1931]" captionTargetId="figure-183@13.[203,1384,633,1944]" captionTargetPageId="13" captionText="FIGURES 47–50. P. caete sp. nov., habitus. 47–48 Male, UFMG 10316 (47 dorsal, 48 ventral); 49–50 Female, UFMG 7199 (49 dorsal, 50 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804657" httpUri="https://zenodo.org/record/14804657/files/figure.png" pageId="16" pageNumber="542">Figs 49–50</figureCitation>
. Total length 10.4. Dorsal shield of prosoma: 4.7 long, 4.1 wide. Opisthosoma: 5.6 long, 3.5 wide. Eyes: diameters: 0.30, 0.30, 0.17, 0.25; interdistances: 0.40, 0.23, 0.60, 0.55, 0.27, 0.18. Legs (4213): I: 12.7 (3.8, 2.2, 3.3, 2.6, 0.8); II: 13.1 (4.1, 2.3, 3.2, 2.7, 0.8); III: 10.4 (3.4, 1.7, 2.4, 2.2, 0.7); IV: 13.4 (4.0, 1.7, 3.0, 3.6, 1.1). Epigyne: EF as wide as long, with elongate MAB on each side; LL with small lateral projections on posterior margin; MS longer than wide with posterior margin slightly curved; ag straight or slightly recurved (
<figureCitation id="130A2A04FFF18C18ED1BFBD4A6D67478" box="[352,438,1086,1113]" captionStart="FIGURES 51–55" captionStartId="14.[151,264,1345,1369]" captionTargetBox="[169,1413,195,1307]" captionTargetId="figure-16@14.[158,1428,181,1320]" captionTargetPageId="14" captionText="FIGURES 51–55. P. caete sp. nov. 51–53 Male (IBSP 135153), left palp (51 prolateral, 52 ventral, 53 retrolateral); 54–55 Female (IBSP 234826), genitalia (54 epigyne ventral, 55 vulva dorsal). Scale lines: 1 mm." pageId="16" pageNumber="542">Figs 54</figureCitation>
<figureCitation id="130A2A04FFF18C18ED1BFBD4A6D67478" box="[352,438,1086,1113]" captionStart="FIGURES 51–55" captionStartId="14.[151,264,1345,1369]" captionTargetBox="[169,1413,195,1307]" captionTargetId="figure-16@14.[158,1428,181,1320]" captionTargetPageId="14" captionText="FIGURES 51–55. P. caete sp. nov. 51–53 Male (IBSP 135153), left palp (51 prolateral, 52 ventral, 53 retrolateral); 54–55 Female (IBSP 234826), genitalia (54 epigyne ventral, 55 vulva dorsal). Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14804659" httpUri="https://zenodo.org/record/14804659/files/figure.png" pageId="16" pageNumber="542">Figs 54</figureCitation>
,
<figureCitation id="130A2A04FFF18C18EDBBFBD4A6807478" box="[448,480,1087,1113]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="16" pageNumber="542">60</figureCitation>
<figureCitation id="130A2A04FFF18C18EDBBFBD4A6807478" box="[448,480,1087,1113]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="16" pageNumber="542">60</figureCitation>
). Vulva: IDS with Dh much longer than wide connecting to Ds laterally; Ds irregularly shaped; FD antero-laterad (
<figureCitation id="130A2A04FFF18C18EDB2FB88A540745C" box="[457,544,1122,1149]" captionStart="FIGURES 51–55" captionStartId="14.[151,264,1345,1369]" captionTargetBox="[169,1413,195,1307]" captionTargetId="figure-16@14.[158,1428,181,1320]" captionTargetPageId="14" captionText="FIGURES 51–55. P. caete sp. nov. 51–53 Male (IBSP 135153), left palp (51 prolateral, 52 ventral, 53 retrolateral); 54–55 Female (IBSP 234826), genitalia (54 epigyne ventral, 55 vulva dorsal). Scale lines: 1 mm." pageId="16" pageNumber="542">Figs 55</figureCitation>
<figureCitation id="130A2A04FFF18C18EDB2FB88A540745C" box="[457,544,1122,1149]" captionStart="FIGURES 51–55" captionStartId="14.[151,264,1345,1369]" captionTargetBox="[169,1413,195,1307]" captionTargetId="figure-16@14.[158,1428,181,1320]" captionTargetPageId="14" captionText="FIGURES 51–55. P. caete sp. nov. 51–53 Male (IBSP 135153), left palp (51 prolateral, 52 ventral, 53 retrolateral); 54–55 Female (IBSP 234826), genitalia (54 epigyne ventral, 55 vulva dorsal). Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14804659" httpUri="https://zenodo.org/record/14804659/files/figure.png" pageId="16" pageNumber="542">Figs 55</figureCitation>
,
<figureCitation id="130A2A04FFF18C18EE57FB88A516745C" box="[556,630,1123,1149]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="16" pageNumber="542">61–62</figureCitation>
<figureCitation id="130A2A04FFF18C18EE57FB88A516745C" box="[556,630,1123,1149]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="16" pageNumber="542">61–62</figureCitation>
).
</paragraph>
<paragraph id="8B8E3681FFF18C18ECBCFB6DA37874E4" blockId="16.[151,1437,151,1293]" pageId="16" pageNumber="542">
@ -921,7 +922,7 @@ Known from the states of
and
<collectingRegion id="49F5F863FFF18C18EDE1FB18A55E752C" box="[410,574,1267,1293]" country="Brazil" name="Santa Catarina" pageId="16" pageNumber="542">Santa Catarina</collectingRegion>
(
<figureCitation id="130A2A04FFF18C18EE36FB18A5C2752C" box="[589,674,1267,1293]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." pageId="16" pageNumber="542">Fig. 79</figureCitation>
<figureCitation id="130A2A04FFF18C18EE36FB18A5C2752C" box="[589,674,1267,1293]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14804675" httpUri="https://zenodo.org/record/14804675/files/figure.png" pageId="16" pageNumber="542">Fig. 79</figureCitation>
).
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</subSubSection>

97
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@ -1,45 +1,46 @@
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<treatment id="03988797FFF18C1BECECFAB4A21671A8" ID-DOI="http://doi.org/10.5281/zenodo.14804796" ID-Zenodo-Dep="14804796" LSID="urn:lsid:plazi:treatment:03988797FFF18C1BECECFAB4A21671A8" httpUri="http://treatment.plazi.org/id/03988797FFF18C1BECECFAB4A21671A8" lastPageId="19" lastPageNumber="545" pageId="16" pageNumber="542">
<subSubSection id="C32B650AFFF18C18ECECFAB4A6DF7558" box="[151,447,1375,1401]" pageId="16" pageNumber="542" type="nomenclature">
<paragraph id="8B8E3681FFF18C18ECECFAB4A6DF7558" blockId="16.[151,447,1375,1437]" box="[151,447,1375,1401]" pageId="16" pageNumber="542">
<heading id="D0C681EDFFF18C18ECECFAB4A6DF7558" bold="true" box="[151,447,1375,1401]" fontSize="11" level="1" pageId="16" pageNumber="542" reason="1">
@ -54,9 +55,9 @@
</subSubSection>
<subSubSection id="C32B650AFFF18C18ECECFA68A65E75BC" box="[151,318,1411,1437]" pageId="16" pageNumber="542" type="description">
<paragraph id="8B8E3681FFF18C18ECECFA68A65E75BC" blockId="16.[151,447,1375,1437]" box="[151,318,1411,1437]" pageId="16" pageNumber="542">
<figureCitation id="130A2A04FFF18C18ECECFA68A67775BC" box="[151,279,1411,1437]" captionStart-0="FIGURES 63–66" captionStart-1="FIGURES 67–71" captionStart-2="FIGURES 72–77" captionStartId-0="17.[151,264,1654,1678]" captionStartId-1="18.[151,264,1441,1465]" captionStartId-2="19.[151,264,1677,1701]" captionTargetBox-0="[166,1415,192,1616]" captionTargetBox-1="[166,1421,219,1407]" captionTargetBox-2="[172,1413,494,1645]" captionTargetId-0="figure-20@17.[165,1422,181,1629]" captionTargetId-1="figure-16@18.[152,1435,201,1418]" captionTargetId-2="figure-134@19.[159,1428,489,1654]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionText-0="FIGURES 63–66. P. tacomare sp. nov., habitus. 63–64 Male, IBSP 97195 (63 dorsal, 64 ventral); 65–66 Female, UFMG 9488 (65 dorsal, 66 ventral). Scale lines: 2 mm." captionText-1="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." captionText-2="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="16" pageNumber="542">Figs 63–77</figureCitation>
<figureCitation id="130A2A04FFF18C18ECECFA68A67775BC" box="[151,279,1411,1437]" captionStart-0="FIGURES 63–66" captionStart-1="FIGURES 67–71" captionStart-2="FIGURES 72–77" captionStartId-0="17.[151,264,1654,1678]" captionStartId-1="18.[151,264,1441,1465]" captionStartId-2="19.[151,264,1677,1701]" captionTargetBox-0="[166,1415,192,1616]" captionTargetBox-1="[166,1421,219,1407]" captionTargetBox-2="[172,1413,494,1645]" captionTargetId-0="figure-20@17.[165,1422,181,1629]" captionTargetId-1="figure-16@18.[152,1435,201,1418]" captionTargetId-2="figure-134@19.[159,1428,489,1654]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionText-0="FIGURES 63–66. P. tacomare sp. nov., habitus. 63–64 Male, IBSP 97195 (63 dorsal, 64 ventral); 65–66 Female, UFMG 9488 (65 dorsal, 66 ventral). Scale lines: 2 mm." captionText-1="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." captionText-2="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi-0="http://doi.org/10.5281/zenodo.14804665" figureDoi-1="http://doi.org/10.5281/zenodo.14804669" figureDoi-2="http://doi.org/10.5281/zenodo.14804671" httpUri-0="https://zenodo.org/record/14804665/files/figure.png" httpUri-1="https://zenodo.org/record/14804669/files/figure.png" httpUri-2="https://zenodo.org/record/14804671/files/figure.png" pageId="16" pageNumber="542">Figs 63–77</figureCitation>
,
<figureCitation id="130A2A04FFF18C18ED59FA68A65E75BC" box="[290,318,1411,1437]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." pageId="16" pageNumber="542">81</figureCitation>
<figureCitation id="130A2A04FFF18C18ED59FA68A65E75BC" box="[290,318,1411,1437]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14804675" httpUri="https://zenodo.org/record/14804675/files/figure.png" pageId="16" pageNumber="542">81</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32B650AFFF18C18ECECFA20A79D776C" pageId="16" pageNumber="542" type="materials_examined">
@ -292,7 +293,7 @@ to southern
, composed mainly of Atlantic Forest, where this species occurs; noun in apposition.
</paragraph>
</subSubSection>
<caption id="DF4E6609FFF08C19ECECF99DA5217693" pageId="17" pageNumber="543" startId="17.[151,264,1654,1678]" targetBox="[166,1415,192,1616]" targetPageId="17" targetType="figure">
<caption id="DF4E6609FFF08C19ECECF99DA5217693" ID-DOI="http://doi.org/10.5281/zenodo.14804665" ID-Zenodo-Dep="14804665" httpUri="https://zenodo.org/record/14804665/files/figure.png" pageId="17" pageNumber="543" startId="17.[151,264,1654,1678]" targetBox="[166,1415,192,1616]" targetPageId="17" targetType="figure">
<paragraph id="8B8E3681FFF08C19ECECF99DA5217693" blockId="17.[151,1436,1654,1714]" pageId="17" pageNumber="543">
<emphasis id="B945EA93FFF08C19ECECF99DA63476AF" bold="true" box="[151,340,1654,1678]" pageId="17" pageNumber="543">FIGURES 63–66.</emphasis>
<taxonomicName id="4C314D02FFF08C19ED21F99CA6B176AF" authorityName="Rheims" authorityYear="2025" box="[346,465,1655,1678]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="543" phylum="Arthropoda" rank="species" species="tacomare" status="sp. nov.">
@ -322,11 +323,11 @@ resemble those
<taxonomicNameLabel id="A27657E8FFF08C19EFA4F8EFA359773F" box="[991,1081,1796,1822]" pageId="17" pageNumber="543" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="130A2A04FFF08C19E833F8EFA3A8773E" box="[1096,1224,1796,1823]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="17" pageNumber="543">Figs 56–62</figureCitation>
<figureCitation id="130A2A04FFF08C19E833F8EFA3A8773E" box="[1096,1224,1796,1823]" captionStart="FIGURES 56–62" captionStartId="15.[151,260,1836,1860]" captionTargetBox="[165,1420,701,1803]" captionTargetId="figure-252@15.[158,1430,698,1811]" captionTargetPageId="15" captionText="FIGURES 56–62. P.caete sp. nov. 56–59 Male (IBSP 16504), left palp (56 prolateral, 57 ventral, 58 retrolateral, 59 detail of embolus, black arrow points to embolus opening); 60–62 Female (IBSP 41116) genitalia (60 epigyne dorsal, 61 vulva ventral, 62 schematic course of IDS). Scale lines 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E— embolus; FD—fertilization duct; Hc—curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804663" httpUri="https://zenodo.org/record/14804663/files/figure.png" pageId="17" pageNumber="543">Figs 56–62</figureCitation>
) by the male palps with RTA bearing more than one branch (single branched in the other species) and by the female epigyne with ag straight or slightly recurved, not projecting over the median septum (projecting in the other species). Males are distinguished from those of the latter species by the RTA with two branches and embolus tip filiform (
<figureCitation id="130A2A04FFF08C19E96DF89BA2F877AB" box="[1302,1432,1904,1930]" captionStart="FIGURES 67–71" captionStartId="18.[151,264,1441,1465]" captionTargetBox="[166,1421,219,1407]" captionTargetId="figure-16@18.[152,1435,201,1418]" captionTargetPageId="18" captionText="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." pageId="17" pageNumber="543">Figs 68–69</figureCitation>
<figureCitation id="130A2A04FFF08C19E96DF89BA2F877AB" box="[1302,1432,1904,1930]" captionStart="FIGURES 67–71" captionStartId="18.[151,264,1441,1465]" captionTargetBox="[166,1421,219,1407]" captionTargetId="figure-16@18.[152,1435,201,1418]" captionTargetPageId="18" captionText="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14804669" httpUri="https://zenodo.org/record/14804669/files/figure.png" pageId="17" pageNumber="543">Figs 68–69</figureCitation>
,
<figureCitation id="130A2A04FFF08C19ECECF87FA7BF778F" box="[151,223,1940,1966]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="17" pageNumber="543">73–74</figureCitation>
<figureCitation id="130A2A04FFF08C19ECECF87FA7BF778F" box="[151,223,1940,1966]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804671" httpUri="https://zenodo.org/record/14804671/files/figure.png" pageId="17" pageNumber="543">73–74</figureCitation>
) (RTA with three branches and embolus tip complex, with projections in
<taxonomicName id="4C314D02FFF08C19E843F87EA3F2778F" authorityName="Rheims" authorityYear="2025" box="[1080,1170,1941,1966]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="543" phylum="Arthropoda" rank="species" species="caete" status="sp. nov.">
<emphasis id="B945EA93FFF08C19E843F87EA3F2778F" box="[1080,1170,1941,1966]" italics="true" pageId="17" pageNumber="543">P. caete</emphasis>
@ -335,9 +336,9 @@ resemble those
<taxonomicNameLabel id="A27657E8FFF08C19E8E7F87FA398778F" box="[1180,1272,1940,1966]" pageId="17" pageNumber="543" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). Females are distinguished by the MS wider than long with posterior margin reaching slightly more than half LL length (
<figureCitation id="130A2A04FFF08C19E93AF853A2F877F3" box="[1345,1432,1976,2002]" captionStart="FIGURES 67–71" captionStartId="18.[151,264,1441,1465]" captionTargetBox="[166,1421,219,1407]" captionTargetId="figure-16@18.[152,1435,201,1418]" captionTargetPageId="18" captionText="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." pageId="17" pageNumber="543">Figs 70</figureCitation>
<figureCitation id="130A2A04FFF08C19E93AF853A2F877F3" box="[1345,1432,1976,2002]" captionStart="FIGURES 67–71" captionStartId="18.[151,264,1441,1465]" captionTargetBox="[166,1421,219,1407]" captionTargetId="figure-16@18.[152,1435,201,1418]" captionTargetPageId="18" captionText="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14804669" httpUri="https://zenodo.org/record/14804669/files/figure.png" pageId="17" pageNumber="543">Figs 70</figureCitation>
,
<figureCitation id="130A2A04FFF08C19ECECF837A7D477D6" box="[151,180,2012,2039]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="17" pageNumber="543">75</figureCitation>
<figureCitation id="130A2A04FFF08C19ECECF837A7D477D6" box="[151,180,2012,2039]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804671" httpUri="https://zenodo.org/record/14804671/files/figure.png" pageId="17" pageNumber="543">75</figureCitation>
) (MS longer than wide with posterior margin at level with posterior margin of LL in
<taxonomicName id="4C314D02FFF08C19E815F836A3A477D7" authorityName="Rheims" authorityYear="2025" box="[1134,1220,2013,2038]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="543" phylum="Arthropoda" rank="species" species="caete" status="sp. nov.">
<emphasis id="B945EA93FFF08C19E815F836A3A477D7" box="[1134,1220,2013,2038]" italics="true" pageId="17" pageNumber="543">P. caete</emphasis>
@ -348,7 +349,7 @@ resemble those
).
</paragraph>
</subSubSection>
<caption id="DF4E6609FFF38C1AECECFA4AA37D75FC" pageId="18" pageNumber="544" startId="18.[151,264,1441,1465]" targetBox="[166,1421,219,1407]" targetPageId="18" targetType="figure">
<caption id="DF4E6609FFF38C1AECECFA4AA37D75FC" ID-DOI="http://doi.org/10.5281/zenodo.14804669" ID-Zenodo-Dep="14804669" httpUri="https://zenodo.org/record/14804669/files/figure.png" pageId="18" pageNumber="544" startId="18.[151,264,1441,1465]" targetBox="[166,1421,219,1407]" targetPageId="18" targetType="figure">
<paragraph id="8B8E3681FFF38C1AECECFA4AA37D75FC" blockId="18.[151,1436,1441,1501]" pageId="18" pageNumber="544">
<emphasis id="B945EA93FFF38C1AECECFA4AA6357598" bold="true" box="[151,341,1441,1465]" pageId="18" pageNumber="544">FIGURES 67–71.</emphasis>
<taxonomicName id="4C314D02FFF38C1AED27FA49A6B47598" authorityName="Rheims" authorityYear="2025" box="[348,468,1442,1465]" class="Arachnida" family="Sparassidae" genus="Papiri" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="544" phylum="Arthropoda" rank="species" species="tacomare" status="sp. nov.">
@ -368,11 +369,11 @@ Description. Male (
):
</emphasis>
Coloration as in
<figureCitation id="130A2A04FFF38C1AEE95F9C4A4117668" box="[750,881,1583,1609]" captionStart="FIGURES 63–66" captionStartId="17.[151,264,1654,1678]" captionTargetBox="[166,1415,192,1616]" captionTargetId="figure-20@17.[165,1422,181,1629]" captionTargetPageId="17" captionText="FIGURES 63–66. P. tacomare sp. nov., habitus. 63–64 Male, IBSP 97195 (63 dorsal, 64 ventral); 65–66 Female, UFMG 9488 (65 dorsal, 66 ventral). Scale lines: 2 mm." pageId="18" pageNumber="544">Figs 63–64</figureCitation>
<figureCitation id="130A2A04FFF38C1AEE95F9C4A4117668" box="[750,881,1583,1609]" captionStart="FIGURES 63–66" captionStartId="17.[151,264,1654,1678]" captionTargetBox="[166,1415,192,1616]" captionTargetId="figure-20@17.[165,1422,181,1629]" captionTargetPageId="17" captionText="FIGURES 63–66. P. tacomare sp. nov., habitus. 63–64 Male, IBSP 97195 (63 dorsal, 64 ventral); 65–66 Female, UFMG 9488 (65 dorsal, 66 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804665" httpUri="https://zenodo.org/record/14804665/files/figure.png" pageId="18" pageNumber="544">Figs 63–64</figureCitation>
. Total length 12.2. Dorsal shield of prosoma: 6.0 long, 5.0 wide. Opisthosoma: 6.0 long, 3.7 wide. Eyes: diameters: 0.42, 0.35, 0.22, 0.33; interdistances: 0.30, 0.20, 0.50, 0.55, 0.25, 0.13. Legs (4/123): I: 17.4 (5.2, 2.8, 4.5, 3.8, 1.1); II: 16.6 (5.2, 2.6, 4.1, 3.7, 1.0); III: 13.6 (4.4, 2.2, 3.0, 3.1, 0.9); IV: 17.4 (5.1, 2.0, 4.0, 4.9, 1.4). Spination follows the generic pattern except tibiae I–II: v2-2-2- 2-2; tibia III: d0; metatarsus II: p1-1-0; metatarsus III: p1-1-0. Palp: VTA small, triangular; RTA with two branches with pointed tips in retrolateral view and curled in ventral view; ST not visible in ventral view; spermophore with pronounced undulation between 6–7 o’clock, close to embolus base; TBE long, with pointed tip; TBC reduced to small sclerotized area; C hood four times longer than wide, twisted in ventral view; E arising from tegulum at 7 o’clock position, gradually narrowing towards filiform tip; MA cup-shaped, distally rounded, longer than wide (
<figureCitation id="130A2A04FFF38C1AE911F8A4A78077AC" captionStart="FIGURES 67–71" captionStartId="18.[151,264,1441,1465]" captionTargetBox="[166,1421,219,1407]" captionTargetId="figure-16@18.[152,1435,201,1418]" captionTargetPageId="18" captionText="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." pageId="18" pageNumber="544">Figs 67–69</figureCitation>
<figureCitation id="130A2A04FFF38C1AE911F8A4A78077AC" captionStart="FIGURES 67–71" captionStartId="18.[151,264,1441,1465]" captionTargetBox="[166,1421,219,1407]" captionTargetId="figure-16@18.[152,1435,201,1418]" captionTargetPageId="18" captionText="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14804669" httpUri="https://zenodo.org/record/14804669/files/figure.png" pageId="18" pageNumber="544">Figs 67–69</figureCitation>
,
<figureCitation id="130A2A04FFF38C1AEC90F898A65677AC" box="[235,310,1907,1933]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="18" pageNumber="544">72–74</figureCitation>
<figureCitation id="130A2A04FFF38C1AEC90F898A65677AC" box="[235,310,1907,1933]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804671" httpUri="https://zenodo.org/record/14804671/files/figure.png" pageId="18" pageNumber="544">72–74</figureCitation>
).
</paragraph>
<paragraph id="8B8E3681FFF38C1BECBCF87CA5F07160" blockId="18.[151,1437,1583,2042]" lastBlockId="19.[151,1436,151,393]" lastPageId="19" lastPageNumber="545" pageId="18" pageNumber="544">
@ -382,15 +383,15 @@ Female (IBSP 85206,
):
</emphasis>
Coloration as in
<figureCitation id="130A2A04FFF38C1AEF7BF87CA4E07793" box="[768,896,1943,1970]" captionStart="FIGURES 63–66" captionStartId="17.[151,264,1654,1678]" captionTargetBox="[166,1415,192,1616]" captionTargetId="figure-20@17.[165,1422,181,1629]" captionTargetPageId="17" captionText="FIGURES 63–66. P. tacomare sp. nov., habitus. 63–64 Male, IBSP 97195 (63 dorsal, 64 ventral); 65–66 Female, UFMG 9488 (65 dorsal, 66 ventral). Scale lines: 2 mm." pageId="18" pageNumber="544">Figs 65–66</figureCitation>
<figureCitation id="130A2A04FFF38C1AEF7BF87CA4E07793" box="[768,896,1943,1970]" captionStart="FIGURES 63–66" captionStartId="17.[151,264,1654,1678]" captionTargetBox="[166,1415,192,1616]" captionTargetId="figure-20@17.[165,1422,181,1629]" captionTargetPageId="17" captionText="FIGURES 63–66. P. tacomare sp. nov., habitus. 63–64 Male, IBSP 97195 (63 dorsal, 64 ventral); 65–66 Female, UFMG 9488 (65 dorsal, 66 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14804665" httpUri="https://zenodo.org/record/14804665/files/figure.png" pageId="18" pageNumber="544">Figs 65–66</figureCitation>
. Total length 12.0. Dorsal shield of prosoma: 5.3 long, 4.3 wide. Opisthosoma: 6.5 long, 3.5 wide. Eyes: diameters: 0.35, 0.30, 0.20, 0.27; interdistances: 0.34, 0.22, 0.56, 0.50, 0.26, 0.15. Legs (4123): I: 13.5 (4.0, 2.5, 3.4, 2.7, 0.9); II: 13.3 (4.1, 2.4, 3.1, 2.8, 0.9); III: 11.1 (3.5, 2.0, 2.4, 2.4, 0.8); IV: 13.8 (4.0, 1.6, 3.2, 3.9, 1.1). Spination follows the generic pattern except femur II: r0-0-1; femur III: p1-0-0; r1-0-0; tibiae I–II: v2-2-2-2-2. Epigyne: EF roughly as wide as long, widest medially with one long MAB on each side; LL with small triangular projections laterally on posterior margin; MS wider than long; ag straight or recurved (
<figureCitation id="130A2A04FFF28C1BEDFEFEE8A6BE713C" box="[389,478,259,285]" captionStart="FIGURES 67–71" captionStartId="18.[151,264,1441,1465]" captionTargetBox="[166,1421,219,1407]" captionTargetId="figure-16@18.[152,1435,201,1418]" captionTargetPageId="18" captionText="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." pageId="19" pageNumber="545">Figs 70</figureCitation>
<figureCitation id="130A2A04FFF28C1BEDFEFEE8A6BE713C" box="[389,478,259,285]" captionStart="FIGURES 67–71" captionStartId="18.[151,264,1441,1465]" captionTargetBox="[166,1421,219,1407]" captionTargetId="figure-16@18.[152,1435,201,1418]" captionTargetPageId="18" captionText="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14804669" httpUri="https://zenodo.org/record/14804669/files/figure.png" pageId="19" pageNumber="545">Figs 70</figureCitation>
,
<figureCitation id="130A2A04FFF28C1BED91FEE9A56A713C" box="[490,522,258,285]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="19" pageNumber="545">75</figureCitation>
<figureCitation id="130A2A04FFF28C1BED91FEE9A56A713C" box="[490,522,258,285]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804671" httpUri="https://zenodo.org/record/14804671/files/figure.png" pageId="19" pageNumber="545">75</figureCitation>
). Vulva: IDS with Dh with two chambers connecting to Ds by elongate duct; Ds rounded; FD antero-laterad (
<figureCitation id="130A2A04FFF28C1BEDACFECCA54E7160" box="[471,558,295,321]" captionStart="FIGURES 67–71" captionStartId="18.[151,264,1441,1465]" captionTargetBox="[166,1421,219,1407]" captionTargetId="figure-16@18.[152,1435,201,1418]" captionTargetPageId="18" captionText="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." pageId="19" pageNumber="545">Figs 71</figureCitation>
<figureCitation id="130A2A04FFF28C1BEDACFECCA54E7160" box="[471,558,295,321]" captionStart="FIGURES 67–71" captionStartId="18.[151,264,1441,1465]" captionTargetBox="[166,1421,219,1407]" captionTargetId="figure-16@18.[152,1435,201,1418]" captionTargetPageId="18" captionText="FIGURES 67–71. P. tacomare sp. nov. 67–69 Male (IBSP 97195), left palp (67 prolateral, 68 ventral, 69 retrolateral); 70–71 Female (IBSP 91735), genitalia (70 epigyne dorsal, 71 vulva ventral). Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14804669" httpUri="https://zenodo.org/record/14804669/files/figure.png" pageId="19" pageNumber="545">Figs 71</figureCitation>
,
<figureCitation id="130A2A04FFF28C1BEE41FECCA5E47160" box="[570,644,295,321]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." pageId="19" pageNumber="545">76–77</figureCitation>
<figureCitation id="130A2A04FFF28C1BEE41FECCA5E47160" box="[570,644,295,321]" captionStart="FIGURES 72–77" captionStartId="19.[151,264,1677,1701]" captionTargetBox="[172,1413,494,1645]" captionTargetId="figure-134@19.[159,1428,489,1654]" captionTargetPageId="19" captionText="FIGURES 72–77. P. tacomare sp. nov. 72–74 Male (IBSP 97195), left palp (72 prolateral, 73 ventral, 74 retrolateral); Female (IBSP 85206), genitalia (75 epigyne dorsal, 76 vulva ventral, 77 schematic course of IDS). Scale lines: 1 mm. ag—anterior groove; C—conductor; Dh—hyaline part of IDS; Ds—sclerotized part of IDS; E—embolus; FD—fertilization duct; Hc— curved end of conductor hood; LL—lateral lobe; MA—median apophysis; MAB—muscle attachment band; MS—median septum; RbP—cymbial retrobasal process; RTA—retrolateral tibial apophysis; ST—subtegulum; TBC—tegular projection close to conductor base; TBE—tegular projection close to conductor base; VTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.14804671" httpUri="https://zenodo.org/record/14804671/files/figure.png" pageId="19" pageNumber="545">76–77</figureCitation>
).
</paragraph>
<paragraph id="8B8E3681FFF28C1BECBCFEA1A2767144" blockId="19.[151,1436,151,393]" box="[199,1302,330,357]" pageId="19" pageNumber="545">
@ -412,7 +413,7 @@ and
, northeastern
<collectingCountry id="F3267611FFF28C1BE8BAFE84A26571A8" box="[1217,1285,367,393]" name="Brazil" pageId="19" pageNumber="545">Brazil</collectingCountry>
(
<figureCitation id="130A2A04FFF28C1BE96EFE84A20871A8" box="[1301,1384,367,393]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." pageId="19" pageNumber="545">Fig. 81</figureCitation>
<figureCitation id="130A2A04FFF28C1BE96EFE84A20871A8" box="[1301,1384,367,393]" captionStart="FIGURES 78–81" captionStartId="20.[151,264,1217,1241]" captionTargetBox="[189,1398,181,1193]" captionTargetId="figure-16@20.[189,1398,181,1193]" captionTargetPageId="20" captionText="FIGURES 78–81. Map of distribution records. 78 P. cabruca sp. nov.; 79 P. cacau sp. nov.; 80 P. caete sp. nov.; 81 P. tacomare sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14804675" httpUri="https://zenodo.org/record/14804675/files/figure.png" pageId="19" pageNumber="545">Fig. 81</figureCitation>
).
</paragraph>
</subSubSection>

75
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@ -1,51 +1,52 @@
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<figureCitation id="132B2A3C9509FF9DFD043A2D2D5F0021" box="[751,814,1895,1919]" captionStart="FIGURE 3" captionStartId="23.[151,250,1843,1867]" captionTargetBox="[215,1373,626,1820]" captionTargetId="figure-112@23.[215,1373,626,1820]" captionTargetPageId="23" captionText="FIGURE 3. Plesionika tenuipes (Smith, 1881). Female (ov.) 15 mm CL, 12 miles off North Guanta, Anzoátegui, 10°25’00”N 64°35’00”W (MMM-Crus-256)." pageId="14" pageNumber="415">Fig. 3</figureCitation>
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<figureCitation id="132B2A3C950FFF9BFCC63A832D4A00BF" box="[813,827,1993,2017]" captionStart="FIGURE 2" captionStartId="22.[151,250,1978,2002]" captionTargetBox="[151,1436,1289,1954]" captionTargetId="figure-381@22.[151,1436,1289,1954]" captionTargetPageId="22" captionText="FIGURE 2. Plesionika martia (A. Milne-Edwards, 1883). Female (ov.), 58 mm CL, 25 miles off N Los Testigos Archipelago, 11°45’00”N 63°05’00”W, 807–878 m depth, 31.viii.1988, (MMM-Crus-444)." pageId="8" pageNumber="409">2</figureCitation>
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<figureCitation id="132B2A3C950FFF9BFCAC3A832D2500BF" box="[839,852,1993,2017]" captionStart="FIGURE 4" captionStartId="26.[151,250,1662,1686]" captionTargetBox="[151,1436,181,1638]" captionTargetId="figure-21@26.[151,1436,181,1638]" captionTargetPageId="26" captionText="FIGURE 4. Glyphocrangon alispina Chace, 1939. Female 27.2 mm CL, off N Los Testigos Islands, 11°45’00”N 63° 5’00”W (MMM-Crus-0246)." figureDoi="http://doi.org/10.5281/zenodo.14804543" httpUri="https://zenodo.org/record/14804543/files/figure.png" pageId="8" pageNumber="409">4</figureCitation>
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(
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<figureCitation id="132B2A3C9513FF87FF7438412E910278" box="[159,224,1291,1318]" captionStart="FIGURE 1" captionStartId="21.[151,250,1667,1691]" captionTargetBox="[151,1436,181,1643]" captionTargetId="figure-17@21.[151,1436,181,1643]" captionTargetPageId="21" captionText="FIGURE 1. Plesionika longicauda: a) male (9.68 mm CL) 17 miles NE Orinoco Delta, 10°25’00”N 64°35’00”W, (MMM- Crus-261). b) specimen from Chichiriviche de la Costa, La Guaira, 60 m. Photography Humberto Ramirez." figureDoi="http://doi.org/10.5281/zenodo.14804537" httpUri="https://zenodo.org/record/14804537/files/figure.png" pageId="20" pageNumber="421">Fig. 1</figureCitation>
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<bibRefCitation id="EF814B489513FF87FE9938812CFE02BD" author="Lemaitre, R. &amp; Gore, R. H." box="[370,655,1483,1507]" pageId="20" pageNumber="421" pagination="382 - 390" refId="ref21525" refString="Lemaitre, R. &amp; Gore, R. H. (1988) Redescription, ecological observations, and distribution of the caridean shrimp Plesionika escatilis (Stimpson, 1860). Proceedings of the Biological Society of Washington, 101, 382-390." type="journal article" year="1988">Lemaitre &amp; Gore 1988: 383</bibRefCitation>
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<figureCitation id="132B2A3C9513FF87FD7138812CAE02BD" box="[666,735,1483,1507]" captionStart="FIGURE 1" captionStartId="21.[151,250,1667,1691]" captionTargetBox="[151,1436,181,1643]" captionTargetId="figure-17@21.[151,1436,181,1643]" captionTargetPageId="21" captionText="FIGURE 1. Plesionika longicauda: a) male (9.68 mm CL) 17 miles NE Orinoco Delta, 10°25’00”N 64°35’00”W, (MMM- Crus-261). b) specimen from Chichiriviche de la Costa, La Guaira, 60 m. Photography Humberto Ramirez." figureDoi="http://doi.org/10.5281/zenodo.14804537" httpUri="https://zenodo.org/record/14804537/files/figure.png" pageId="20" pageNumber="421">Figs. 1</figureCitation>
,
<figureCitation id="132B2A3C9513FF87FD0038812C8802BD" box="[747,761,1483,1507]" captionStart="FIGURE 2" captionStartId="22.[151,250,1978,2002]" captionTargetBox="[151,1436,1289,1954]" captionTargetId="figure-381@22.[151,1436,1289,1954]" captionTargetPageId="22" captionText="FIGURE 2. Plesionika martia (A. Milne-Edwards, 1883). Female (ov.), 58 mm CL, 25 miles off N Los Testigos Archipelago, 11°45’00”N 63°05’00”W, 807–878 m depth, 31.viii.1988, (MMM-Crus-444)." pageId="20" pageNumber="421">2</figureCitation>
<figureCitation id="132B2A3C9513FF87FD0038812C8802BD" box="[747,761,1483,1507]" captionStart="FIGURE 2" captionStartId="22.[151,250,1978,2002]" captionTargetBox="[151,1436,1289,1954]" captionTargetId="figure-381@22.[151,1436,1289,1954]" captionTargetPageId="22" captionText="FIGURE 2. Plesionika martia (A. Milne-Edwards, 1883). Female (ov.), 58 mm CL, 25 miles off N Los Testigos Archipelago, 11°45’00”N 63°05’00”W, 807–878 m depth, 31.viii.1988, (MMM-Crus-444)." figureDoi="http://doi.org/10.5281/zenodo.14804539" httpUri="https://zenodo.org/record/14804539/files/figure.png" pageId="20" pageNumber="421">2</figureCitation>
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<figureCitation id="132B2A3C9513FF87FCEF38812D4D02BC" box="[772,828,1483,1507]" captionStart="FIGURE 3" captionStartId="23.[151,250,1843,1867]" captionTargetBox="[215,1373,626,1820]" captionTargetId="figure-112@23.[215,1373,626,1820]" captionTargetPageId="23" captionText="FIGURE 3. Plesionika tenuipes (Smith, 1881). Female (ov.) 15 mm CL, 12 miles off North Guanta, Anzoátegui, 10°25’00”N 64°35’00”W (MMM-Crus-256)." pageId="20" pageNumber="421">
<figureCitation id="132B2A3C9513FF87FCEF38812D4D02BC" box="[772,828,1483,1507]" captionStart="FIGURE 3" captionStartId="23.[151,250,1843,1867]" captionTargetBox="[215,1373,626,1820]" captionTargetId="figure-112@23.[215,1373,626,1820]" captionTargetPageId="23" captionText="FIGURE 3. Plesionika tenuipes (Smith, 1881). Female (ov.) 15 mm CL, 12 miles off North Guanta, Anzoátegui, 10°25’00”N 64°35’00”W (MMM-Crus-256)." figureDoi="http://doi.org/10.5281/zenodo.14804541" httpUri="https://zenodo.org/record/14804541/files/figure.png" pageId="20" pageNumber="421">
3A
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J
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<figureCitation id="132B2A3C9513FF87FCAC38812D2502BD" box="[839,852,1483,1507]" captionStart="FIGURE 4" captionStartId="26.[151,250,1662,1686]" captionTargetBox="[151,1436,181,1638]" captionTargetId="figure-21@26.[151,1436,181,1638]" captionTargetPageId="26" captionText="FIGURE 4. Glyphocrangon alispina Chace, 1939. Female 27.2 mm CL, off N Los Testigos Islands, 11°45’00”N 63° 5’00”W (MMM-Crus-0246)." pageId="20" pageNumber="421">4</figureCitation>
<figureCitation id="132B2A3C9513FF87FCAC38812D2502BD" box="[839,852,1483,1507]" captionStart="FIGURE 4" captionStartId="26.[151,250,1662,1686]" captionTargetBox="[151,1436,181,1638]" captionTargetId="figure-21@26.[151,1436,181,1638]" captionTargetPageId="26" captionText="FIGURE 4. Glyphocrangon alispina Chace, 1939. Female 27.2 mm CL, off N Los Testigos Islands, 11°45’00”N 63° 5’00”W (MMM-Crus-0246)." figureDoi="http://doi.org/10.5281/zenodo.14804543" httpUri="https://zenodo.org/record/14804543/files/figure.png" pageId="20" pageNumber="421">4</figureCitation>
(non
<bibRefCitation id="EF814B489513FF87FC6538812A5A02BD" author="Stimpson, W." box="[910,1067,1483,1507]" pageId="20" pageNumber="421" pagination="22 - 47" refId="ref24101" refString="Stimpson, W. (1860) Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republic Federata missa, Cadwaladore Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars VIII, Crustacea Macrura. Proceedings of the Academy of Natural Sciences of Philadelphia, 12, 22-47 [91 - 116 in separate]." type="journal article" year="1860">Stimpson 1860</bibRefCitation>
).
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; present studio).
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<emphasis id="B964EAAB9512FF86FF7C3BC92F6401C5" bold="true" box="[151,277,1667,1691]" pageId="21" pageNumber="422">FIGURE 1.</emphasis>
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was seen and photographed inside a cave at approximately
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in Chichiriviche de la Costa, La Guaira state (
<figureCitation id="132B2A3C9512FF86FD413A5B2C880075" box="[682,761,1809,1835]" captionStart="FIGURE 1" captionStartId="21.[151,250,1667,1691]" captionTargetBox="[151,1436,181,1643]" captionTargetId="figure-17@21.[151,1436,181,1643]" captionTargetPageId="21" captionText="FIGURE 1. Plesionika longicauda: a) male (9.68 mm CL) 17 miles NE Orinoco Delta, 10°25’00”N 64°35’00”W, (MMM- Crus-261). b) specimen from Chichiriviche de la Costa, La Guaira, 60 m. Photography Humberto Ramirez." pageId="21" pageNumber="422">Fig. 1b</figureCitation>
<figureCitation id="132B2A3C9512FF86FD413A5B2C880075" box="[682,761,1809,1835]" captionStart="FIGURE 1" captionStartId="21.[151,250,1667,1691]" captionTargetBox="[151,1436,181,1643]" captionTargetId="figure-17@21.[151,1436,181,1643]" captionTargetPageId="21" captionText="FIGURE 1. Plesionika longicauda: a) male (9.68 mm CL) 17 miles NE Orinoco Delta, 10°25’00”N 64°35’00”W, (MMM- Crus-261). b) specimen from Chichiriviche de la Costa, La Guaira, 60 m. Photography Humberto Ramirez." figureDoi="http://doi.org/10.5281/zenodo.14804537" httpUri="https://zenodo.org/record/14804537/files/figure.png" pageId="21" pageNumber="422">Fig. 1b</figureCitation>
) and represents the first reports of this species for
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(
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<figureCitation id="13462A38FFA0B342FF4CFB77FEBFFB43" box="[159,289,1195,1221]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804809" httpUri="https://zenodo.org/record/14804809/files/figure.png" pageId="8" pageNumber="470">Figs 21–22</figureCitation>
,
<figureCitation id="13462A38FFA0B342FEFFFB77FEEBFB43" box="[300,373,1195,1221]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1191,1215]" captionTargetBox="[156,1430,186,1162]" captionTargetId="figure-15@10.[151,1435,181,1167]" captionTargetPageId="10" captionText="FIGURES 30–34. Stenoscelis longisetosus. 30, aedeagus, dorsal view; 31, ditto, lateral view; 32, tegmen; 33, sternite VIII &amp; IX; 34, spermatheca. Scale bar: 0.20 mm." pageId="8" pageNumber="470">30–34</figureCitation>
<figureCitation id="13462A38FFA0B342FEFFFB77FEEBFB43" box="[300,373,1195,1221]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1191,1215]" captionTargetBox="[156,1430,186,1162]" captionTargetId="figure-15@10.[151,1435,181,1167]" captionTargetPageId="10" captionText="FIGURES 30–34. Stenoscelis longisetosus. 30, aedeagus, dorsal view; 31, ditto, lateral view; 32, tegmen; 33, sternite VIII &amp; IX; 34, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804813" httpUri="https://zenodo.org/record/14804813/files/figure.png" pageId="8" pageNumber="470">30–34</figureCitation>
,
<figureCitation id="13462A38FFA0B342FE53FB77FE02FB43" box="[384,412,1195,1221]" captionStart="FIGURE 68–73" captionStartId="21.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-92@21.[151,1435,309,1165]" captionTargetPageId="21" captionText="FIGURE 68–73. Heads of Stenoscelis spp. 68, Stenoscelis acutipennis; 69, Stenoscelis brevithoracicus sp. nov.; 70, Stenoscelis cavirostris sp. nov., male; 71, ditto, female; 72, Stenoscelis longisetosus; 73, Stenoscelis aceri. Scale bar: 0.50 mm." pageId="8" pageNumber="470">72</figureCitation>
<figureCitation id="13462A38FFA0B342FE53FB77FE02FB43" box="[384,412,1195,1221]" captionStart="FIGURE 68–73" captionStartId="21.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-92@21.[151,1435,309,1165]" captionTargetPageId="21" captionText="FIGURE 68–73. Heads of Stenoscelis spp. 68, Stenoscelis acutipennis; 69, Stenoscelis brevithoracicus sp. nov.; 70, Stenoscelis cavirostris sp. nov., male; 71, ditto, female; 72, Stenoscelis longisetosus; 73, Stenoscelis aceri. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804827" httpUri="https://zenodo.org/record/14804827/files/figure.png" pageId="8" pageNumber="470">72</figureCitation>
&amp;
<figureCitation id="13462A38FFA0B342FE13FB77FE43FB43" box="[448,477,1195,1221]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." pageId="8" pageNumber="470">81</figureCitation>
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)
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@ -157,13 +158,13 @@ in the upper margin of scrobe being directed to middle of eye, the elytral setae
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in the following character states: elytra shorter (1.6–1.7 times as long as wide); apex of elytra conjointly rounded (
<figureCitation id="13462A38FFA0B342FC51F985FC4AF9F5" box="[898,980,1625,1651]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." pageId="8" pageNumber="470">Fig. 81</figureCitation>
<figureCitation id="13462A38FFA0B342FC51F985FC4AF9F5" box="[898,980,1625,1651]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804831" httpUri="https://zenodo.org/record/14804831/files/figure.png" pageId="8" pageNumber="470">Fig. 81</figureCitation>
), obtusely angulated in lateral view (
<figureCitation id="13462A38FFA0B342FABCF985FF29F911" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." pageId="8" pageNumber="470">Fig. 22</figureCitation>
<figureCitation id="13462A38FFA0B342FABCF985FF29F911" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804809" httpUri="https://zenodo.org/record/14804809/files/figure.png" pageId="8" pageNumber="470">Fig. 22</figureCitation>
); aedeagal body 1.4 times as long as wide, 1.0 times as long as apodeme (
<figureCitation id="13462A38FFA0B342FC3EF9A1FBEEF911" box="[1005,1136,1661,1687]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1191,1215]" captionTargetBox="[156,1430,186,1162]" captionTargetId="figure-15@10.[151,1435,181,1167]" captionTargetPageId="10" captionText="FIGURES 30–34. Stenoscelis longisetosus. 30, aedeagus, dorsal view; 31, ditto, lateral view; 32, tegmen; 33, sternite VIII &amp; IX; 34, spermatheca. Scale bar: 0.20 mm." pageId="8" pageNumber="470">Figs 30–31</figureCitation>
<figureCitation id="13462A38FFA0B342FC3EF9A1FBEEF911" box="[1005,1136,1661,1687]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1191,1215]" captionTargetBox="[156,1430,186,1162]" captionTargetId="figure-15@10.[151,1435,181,1167]" captionTargetPageId="10" captionText="FIGURES 30–34. Stenoscelis longisetosus. 30, aedeagus, dorsal view; 31, ditto, lateral view; 32, tegmen; 33, sternite VIII &amp; IX; 34, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804813" httpUri="https://zenodo.org/record/14804813/files/figure.png" pageId="8" pageNumber="470">Figs 30–31</figureCitation>
); spermatheca with collum developed and ramus not developed (
<figureCitation id="13462A38FFA0B342FDEAF97DFD10F93D" box="[569,654,1697,1723]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1191,1215]" captionTargetBox="[156,1430,186,1162]" captionTargetId="figure-15@10.[151,1435,181,1167]" captionTargetPageId="10" captionText="FIGURES 30–34. Stenoscelis longisetosus. 30, aedeagus, dorsal view; 31, ditto, lateral view; 32, tegmen; 33, sternite VIII &amp; IX; 34, spermatheca. Scale bar: 0.20 mm." pageId="8" pageNumber="470">Fig. 34</figureCitation>
<figureCitation id="13462A38FFA0B342FDEAF97DFD10F93D" box="[569,654,1697,1723]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1191,1215]" captionTargetBox="[156,1430,186,1162]" captionTargetId="figure-15@10.[151,1435,181,1167]" captionTargetPageId="10" captionText="FIGURES 30–34. Stenoscelis longisetosus. 30, aedeagus, dorsal view; 31, ditto, lateral view; 32, tegmen; 33, sternite VIII &amp; IX; 34, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804813" httpUri="https://zenodo.org/record/14804813/files/figure.png" pageId="8" pageNumber="470">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
@ -176,7 +177,7 @@ Body length
</paragraph>
<paragraph id="8BC236BDFFA0B342FF14F8D2FBAEF83E" blockId="8.[151,1437,1517,2012]" pageId="8" pageNumber="470">Rostrum 0.6–0.7 times as long as wide; dorsum convex; dorsal surface reticulate, rugosely punctate; punctures small, separated by interspace approximately 1–2 times as long as diameter; upper margin of scrobe directed to middle of eye; epistome with six long setae basally, with anterior margin obtusely projected at right side. Head with dorsal surface reticulate, smoothly punctate; punctures small, separated by interspace approximately 2–4 times as long as diameter; interocular area with a shallow fovea; eye round, hardly convex.</paragraph>
<paragraph id="8BC236BDFFA0B342FF14F81EFAC9F85D" blockId="8.[151,1437,1517,2012]" box="[199,1367,1985,2012]" pageId="8" pageNumber="470">Antennae with club subcircular, slightly wider than long; funicle 7-segmented, slightly longer than club.</paragraph>
<caption id="DF026635FFA1B343FF44FBF5FDC9FBE3" pageId="9" pageNumber="471" startId="9.[151,264,1065,1089]" targetBox="[157,1430,187,1036]" targetPageId="9" targetType="figure">
<caption id="DF026635FFA1B343FF44FBF5FDC9FBE3" ID-DOI="http://doi.org/10.5281/zenodo.14804811" ID-Zenodo-Dep="14804811" httpUri="https://zenodo.org/record/14804811/files/figure.png" pageId="9" pageNumber="471" startId="9.[151,264,1065,1089]" targetBox="[157,1430,187,1036]" targetPageId="9" targetType="figure">
<paragraph id="8BC236BDFFA1B343FF44FBF5FDC9FBE3" blockId="9.[151,1437,1065,1126]" pageId="9" pageNumber="471">
<emphasis id="B909EAAFFFA1B343FF44FBF5FECDFBC7" bold="true" box="[151,339,1065,1089]" pageId="9" pageNumber="471">FIGURES 25–29.</emphasis>
<taxonomicName id="4C7D4D3EFFA1B343FE8BFBF5FDB1FBC7" box="[344,559,1065,1089]" class="Insecta" family="Curculionidae" genus="Stenoscelis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="471" phylum="Arthropoda" rank="species" species="cavirostri">
@ -191,11 +192,11 @@ s
<paragraph id="8BC236BDFFA1B343FF14FAE4FE82F9CC" blockId="9.[151,1437,1194,2035]" pageId="9" pageNumber="471">Elytra 1.6–1.7 times as long as wide, 1.1 times as wide as and 2.1–2.3 times as long as prothorax, subparallel, conjointly rounded at apex; apex obtusely angulated in lateral view; striae rather deep, with medium punctures separated by interspace approximately 1–2 times as long as diameter; apical part of striae moderated, not dilated or reticulate; second stria distinctly separated from base; intervals somewhat convex, with an irregular row of small puncture; base of second to third or fourth interval rugosely punctate-granulate; apical half of ninth interval and tenth interval carinate; granules of intervals arranged in a single row, indistinct basally, distinct and conical on declivity; setae on declivity arranged in a single row, each behind a granule, distinct and more than twice as long as granules.</paragraph>
<paragraph id="8BC236BDFFA1B343FF14F98FFC33F97A" blockId="9.[151,1437,1194,2035]" pageId="9" pageNumber="471">
Aedeagal body (
<figureCitation id="13462A38FFA1B343FE56F98FFD92F9E8" box="[389,524,1619,1646]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1191,1215]" captionTargetBox="[156,1430,186,1162]" captionTargetId="figure-15@10.[151,1435,181,1167]" captionTargetPageId="10" captionText="FIGURES 30–34. Stenoscelis longisetosus. 30, aedeagus, dorsal view; 31, ditto, lateral view; 32, tegmen; 33, sternite VIII &amp; IX; 34, spermatheca. Scale bar: 0.20 mm." pageId="9" pageNumber="471">Figs 30, 31</figureCitation>
<figureCitation id="13462A38FFA1B343FE56F98FFD92F9E8" box="[389,524,1619,1646]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1191,1215]" captionTargetBox="[156,1430,186,1162]" captionTargetId="figure-15@10.[151,1435,181,1167]" captionTargetPageId="10" captionText="FIGURES 30–34. Stenoscelis longisetosus. 30, aedeagus, dorsal view; 31, ditto, lateral view; 32, tegmen; 33, sternite VIII &amp; IX; 34, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804813" httpUri="https://zenodo.org/record/14804813/files/figure.png" pageId="9" pageNumber="471">Figs 30, 31</figureCitation>
) 1.4 times as long as wide, 1.0 times as long as apodeme, moderately arcuate laterally, gradually narrowed and weakly prolonged apically; apodeme attached sublaterally; endophallus with a pair of sclerite on subapical part; tegmen (
<figureCitation id="13462A38FFA1B343FD56F946FD47F932" box="[645,729,1690,1716]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1191,1215]" captionTargetBox="[156,1430,186,1162]" captionTargetId="figure-15@10.[151,1435,181,1167]" captionTargetPageId="10" captionText="FIGURES 30–34. Stenoscelis longisetosus. 30, aedeagus, dorsal view; 31, ditto, lateral view; 32, tegmen; 33, sternite VIII &amp; IX; 34, spermatheca. Scale bar: 0.20 mm." pageId="9" pageNumber="471">Fig. 32</figureCitation>
<figureCitation id="13462A38FFA1B343FD56F946FD47F932" box="[645,729,1690,1716]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1191,1215]" captionTargetBox="[156,1430,186,1162]" captionTargetId="figure-15@10.[151,1435,181,1167]" captionTargetPageId="10" captionText="FIGURES 30–34. Stenoscelis longisetosus. 30, aedeagus, dorsal view; 31, ditto, lateral view; 32, tegmen; 33, sternite VIII &amp; IX; 34, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804813" httpUri="https://zenodo.org/record/14804813/files/figure.png" pageId="9" pageNumber="471">Fig. 32</figureCitation>
) without dorsal part and paramere, with distinct manubrium; spiculum gastrale (
<figureCitation id="13462A38FFA1B343FEB9F962FE24F95E" box="[362,442,1726,1752]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1191,1215]" captionTargetBox="[156,1430,186,1162]" captionTargetId="figure-15@10.[151,1435,181,1167]" captionTargetPageId="10" captionText="FIGURES 30–34. Stenoscelis longisetosus. 30, aedeagus, dorsal view; 31, ditto, lateral view; 32, tegmen; 33, sternite VIII &amp; IX; 34, spermatheca. Scale bar: 0.20 mm." pageId="9" pageNumber="471">Fig. 33</figureCitation>
<figureCitation id="13462A38FFA1B343FEB9F962FE24F95E" box="[362,442,1726,1752]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1191,1215]" captionTargetBox="[156,1430,186,1162]" captionTargetId="figure-15@10.[151,1435,181,1167]" captionTargetPageId="10" captionText="FIGURES 30–34. Stenoscelis longisetosus. 30, aedeagus, dorsal view; 31, ditto, lateral view; 32, tegmen; 33, sternite VIII &amp; IX; 34, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804813" httpUri="https://zenodo.org/record/14804813/files/figure.png" pageId="9" pageNumber="471">Fig. 33</figureCitation>
) gradually curved to right side apically, not expanded apically; sternite VIII divided, with several long setae apically; without spinulate apical membranous area.
</paragraph>
<paragraph id="8BC236BDFFA1B343FF14F8D9FE8CF8C4" blockId="9.[151,1437,1194,2035]" pageId="9" pageNumber="471">Spiculum ventral not bifurcate at base; spermatheca with cornu falciform; collum well developed; ramus not developed.</paragraph>
@ -233,7 +234,7 @@ of
<collectingRegion id="49B9F85FFFA1B343FA96F86BFF50F875" country="Japan" name="Nagano" pageId="9" pageNumber="471">Nagano Pref.</collectingRegion>
</paragraph>
</subSubSection>
<caption id="DF026635FFA2B340FF44FB7BFDA3FB64" pageId="10" pageNumber="472" startId="10.[151,264,1191,1215]" targetBox="[156,1430,186,1162]" targetPageId="10" targetType="figure">
<caption id="DF026635FFA2B340FF44FB7BFDA3FB64" ID-DOI="http://doi.org/10.5281/zenodo.14804813" ID-Zenodo-Dep="14804813" httpUri="https://zenodo.org/record/14804813/files/figure.png" pageId="10" pageNumber="472" startId="10.[151,264,1191,1215]" targetBox="[156,1430,186,1162]" targetPageId="10" targetType="figure">
<paragraph id="8BC236BDFFA2B340FF44FB7BFDA3FB64" blockId="10.[151,1437,1191,1251]" pageId="10" pageNumber="472">
<emphasis id="B909EAAFFFA2B340FF44FB7BFECBFB39" bold="true" box="[151,341,1191,1215]" pageId="10" pageNumber="472">FIGURES 30–34.</emphasis>
<taxonomicName id="4C7D4D3EFFA2B340FE8FFB7BFDCFFB39" authorityName="Konishi" authorityYear="1956" box="[348,593,1191,1215]" class="Insecta" family="Curculionidae" genus="Stenoscelis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="472" phylum="Arthropoda" rank="species" species="longisetosus">
@ -254,7 +255,7 @@ of
</taxonomicName>
</emphasis>
(
<figureCitation id="13462A38FFA2B340FCFEFAE9FC5AFAC9" box="[813,964,1333,1359]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." pageId="10" pageNumber="472">Figs 21 &amp; 22</figureCitation>
<figureCitation id="13462A38FFA2B340FCFEFAE9FC5AFAC9" box="[813,964,1333,1359]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804809" httpUri="https://zenodo.org/record/14804809/files/figure.png" pageId="10" pageNumber="472">Figs 21 &amp; 22</figureCitation>
): “Pref
<collectingRegion id="49B9F85FFFA2B340FBCBFAE9FBD5FAC9" box="[1048,1099,1333,1359]" country="Japan" name="Gifu" pageId="10" pageNumber="472">Gifu</collectingRegion>
/

83
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@ -57,13 +58,13 @@
<subSubSection id="C3676536FFA3B341FF44F97FFE7BF93B" box="[151,485,1698,1725]" pageId="11" pageNumber="473" type="description">
<paragraph id="8BC236BDFFA3B341FF44F97FFE7BF93B" blockId="11.[151,805,1626,1725]" box="[151,485,1698,1725]" pageId="11" pageNumber="473">
(
<figureCitation id="13462A38FFA3B341FF4CF97FFEBEF93B" box="[159,288,1699,1725]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." pageId="11" pageNumber="473">Figs 23–24</figureCitation>
<figureCitation id="13462A38FFA3B341FF4CF97FFEBEF93B" box="[159,288,1699,1725]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804809" httpUri="https://zenodo.org/record/14804809/files/figure.png" pageId="11" pageNumber="473">Figs 23–24</figureCitation>
,
<figureCitation id="13462A38FFA3B341FEFFF97EFEEBF93B" box="[300,373,1698,1725]" captionStart="FIGURES 35–39" captionStartId="12.[151,264,1129,1153]" captionTargetBox="[157,1430,187,1099]" captionTargetId="figure-15@12.[151,1435,181,1105]" captionTargetPageId="12" captionText="FIGURES 35–39. Stenoscelis aceri. 35, aedeagus, dorsal view; 36, ditto, lateral view; 37, tegmen; 38, sternite VIII &amp; IX; 39, spermatheca. Scale bar: 0.20 mm." pageId="11" pageNumber="473">35–39</figureCitation>
<figureCitation id="13462A38FFA3B341FEFFF97EFEEBF93B" box="[300,373,1698,1725]" captionStart="FIGURES 35–39" captionStartId="12.[151,264,1129,1153]" captionTargetBox="[157,1430,187,1099]" captionTargetId="figure-15@12.[151,1435,181,1105]" captionTargetPageId="12" captionText="FIGURES 35–39. Stenoscelis aceri. 35, aedeagus, dorsal view; 36, ditto, lateral view; 37, tegmen; 38, sternite VIII &amp; IX; 39, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804815" httpUri="https://zenodo.org/record/14804815/files/figure.png" pageId="11" pageNumber="473">35–39</figureCitation>
,
<figureCitation id="13462A38FFA3B341FE53F97FFE02F93B" box="[384,412,1699,1725]" captionStart="FIGURE 68–73" captionStartId="21.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-92@21.[151,1435,309,1165]" captionTargetPageId="21" captionText="FIGURE 68–73. Heads of Stenoscelis spp. 68, Stenoscelis acutipennis; 69, Stenoscelis brevithoracicus sp. nov.; 70, Stenoscelis cavirostris sp. nov., male; 71, ditto, female; 72, Stenoscelis longisetosus; 73, Stenoscelis aceri. Scale bar: 0.50 mm." pageId="11" pageNumber="473">73</figureCitation>
<figureCitation id="13462A38FFA3B341FE53F97FFE02F93B" box="[384,412,1699,1725]" captionStart="FIGURE 68–73" captionStartId="21.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-92@21.[151,1435,309,1165]" captionTargetPageId="21" captionText="FIGURE 68–73. Heads of Stenoscelis spp. 68, Stenoscelis acutipennis; 69, Stenoscelis brevithoracicus sp. nov.; 70, Stenoscelis cavirostris sp. nov., male; 71, ditto, female; 72, Stenoscelis longisetosus; 73, Stenoscelis aceri. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804827" httpUri="https://zenodo.org/record/14804827/files/figure.png" pageId="11" pageNumber="473">73</figureCitation>
&amp;
<figureCitation id="13462A38FFA3B341FE13F97FFE43F93B" box="[448,477,1699,1725]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." pageId="11" pageNumber="473">82</figureCitation>
<figureCitation id="13462A38FFA3B341FE13F97FFE43F93B" box="[448,477,1699,1725]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804831" httpUri="https://zenodo.org/record/14804831/files/figure.png" pageId="11" pageNumber="473">82</figureCitation>
)
</paragraph>
</subSubSection>
@ -108,17 +109,17 @@ This species is very similar to
<emphasis id="B909EAAFFFA3B341FD5BF890FC8CF8E0" box="[648,786,1868,1894]" italics="true" pageId="11" pageNumber="473">longisetosus</emphasis>
</taxonomicName>
, but differs in the following character states: elytra longer (1.8–1.9 times as long as wide); apex of elytra separately produced in a small process at apex (
<figureCitation id="13462A38FFA3B341FB0CF8ACFAABF80C" box="[1247,1333,1904,1930]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." pageId="11" pageNumber="473">Fig. 82</figureCitation>
<figureCitation id="13462A38FFA3B341FB0CF8ACFAABF80C" box="[1247,1333,1904,1930]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804831" httpUri="https://zenodo.org/record/14804831/files/figure.png" pageId="11" pageNumber="473">Fig. 82</figureCitation>
), acutely angulated in lateral view (
<figureCitation id="13462A38FFA3B341FE12F848FD89F828" box="[449,535,1940,1966]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." pageId="11" pageNumber="473">Fig. 24</figureCitation>
<figureCitation id="13462A38FFA3B341FE12F848FD89F828" box="[449,535,1940,1966]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804809" httpUri="https://zenodo.org/record/14804809/files/figure.png" pageId="11" pageNumber="473">Fig. 24</figureCitation>
); aedeagal body 1.7 times as long as wide, 1.4 times as long as apodeme (
<figureCitation id="13462A38FFA3B341FAB9F848FF7CF855" captionStart="FIGURES 35–39" captionStartId="12.[151,264,1129,1153]" captionTargetBox="[157,1430,187,1099]" captionTargetId="figure-15@12.[151,1435,181,1105]" captionTargetPageId="12" captionText="FIGURES 35–39. Stenoscelis aceri. 35, aedeagus, dorsal view; 36, ditto, lateral view; 37, tegmen; 38, sternite VIII &amp; IX; 39, spermatheca. Scale bar: 0.20 mm." pageId="11" pageNumber="473">Figs 35–36</figureCitation>
<figureCitation id="13462A38FFA3B341FAB9F848FF7CF855" captionStart="FIGURES 35–39" captionStartId="12.[151,264,1129,1153]" captionTargetBox="[157,1430,187,1099]" captionTargetId="figure-15@12.[151,1435,181,1105]" captionTargetPageId="12" captionText="FIGURES 35–39. Stenoscelis aceri. 35, aedeagus, dorsal view; 36, ditto, lateral view; 37, tegmen; 38, sternite VIII &amp; IX; 39, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804815" httpUri="https://zenodo.org/record/14804815/files/figure.png" pageId="11" pageNumber="473">Figs 35–36</figureCitation>
); spermatheca with collum and ramus developed (
<figureCitation id="13462A38FFA3B341FCC0F864FCF9F854" box="[787,871,1976,2002]" captionStart="FIGURES 35–39" captionStartId="12.[151,264,1129,1153]" captionTargetBox="[157,1430,187,1099]" captionTargetId="figure-15@12.[151,1435,181,1105]" captionTargetPageId="12" captionText="FIGURES 35–39. Stenoscelis aceri. 35, aedeagus, dorsal view; 36, ditto, lateral view; 37, tegmen; 38, sternite VIII &amp; IX; 39, spermatheca. Scale bar: 0.20 mm." pageId="11" pageNumber="473">Fig. 39</figureCitation>
<figureCitation id="13462A38FFA3B341FCC0F864FCF9F854" box="[787,871,1976,2002]" captionStart="FIGURES 35–39" captionStartId="12.[151,264,1129,1153]" captionTargetBox="[157,1430,187,1099]" captionTargetId="figure-15@12.[151,1435,181,1105]" captionTargetPageId="12" captionText="FIGURES 35–39. Stenoscelis aceri. 35, aedeagus, dorsal view; 36, ditto, lateral view; 37, tegmen; 38, sternite VIII &amp; IX; 39, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804815" httpUri="https://zenodo.org/record/14804815/files/figure.png" pageId="11" pageNumber="473">Fig. 39</figureCitation>
).
</paragraph>
</subSubSection>
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<caption id="DF026635FFA4B346FF44FBB5FE71FB23" ID-DOI="http://doi.org/10.5281/zenodo.14804815" ID-Zenodo-Dep="14804815" httpUri="https://zenodo.org/record/14804815/files/figure.png" pageId="12" pageNumber="474" startId="12.[151,264,1129,1153]" targetBox="[157,1430,187,1099]" targetPageId="12" targetType="figure">
<paragraph id="8BC236BDFFA4B346FF44FBB5FE71FB23" blockId="12.[151,1436,1129,1189]" pageId="12" pageNumber="474">
<emphasis id="B909EAAFFFA4B346FF44FBB5FECBFB07" bold="true" box="[151,341,1129,1153]" pageId="12" pageNumber="474">FIGURES 35–39.</emphasis>
<emphasis id="B909EAAFFFA4B346FE88FBB5FD90FB07" box="[347,526,1129,1153]" italics="true" pageId="12" pageNumber="474">
@ -166,7 +167,7 @@ in
<typeStatus id="54C6881FFFA5B347FEAEFE6BFE74FE57" box="[381,490,439,465]" pageId="13" pageNumber="475" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="13462A38FFA5B347FE2AFE6BFD0DFE57" box="[505,659,439,465]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." pageId="13" pageNumber="475">Figs 23 &amp; 24</figureCitation>
<figureCitation id="13462A38FFA5B347FE2AFE6BFD0DFE57" box="[505,659,439,465]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804809" httpUri="https://zenodo.org/record/14804809/files/figure.png" pageId="13" pageNumber="475">Figs 23 &amp; 24</figureCitation>
): male (
<collectionCode id="ED6CAE78FFA5B347FD3EFE6BFCA2FE57" box="[749,828,439,465]" pageId="13" pageNumber="475">SEHU</collectionCode>
), “Maruyama /

71
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<subSubSection id="C3676536FFA5B347FF44FCC2FC57FC07" pageId="13" pageNumber="475" type="nomenclature">
@ -64,11 +65,11 @@ urn:lsid:zoobank.org:act:
<subSubSection id="C3676536FFA5B347FF44FC57FE25FC23" box="[151,443,906,933]" pageId="13" pageNumber="475" type="description">
<paragraph id="8BC236BDFFA5B347FF44FC57FE25FC23" blockId="13.[151,969,798,933]" box="[151,443,906,933]" pageId="13" pageNumber="475">
(
<figureCitation id="13462A38FFA5B347FF4CFC57FEBEFC23" box="[159,288,907,933]" captionStart="FIGURES 40–47" captionStartId="14.[151,264,1484,1508]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-15@14.[151,1435,181,1461]" captionTargetPageId="14" captionText="FIGURES 40–47. Habitus of Stenoscelis spp. 40 &amp; 41, Stenoscelis matobai sp. nov., holotype, male; 42 &amp; 43, Stenoscelis reniformis sp. nov., holotype, male; 44 &amp; 45, Stenoscelis gracilitarsis; 46 &amp; 47, Stenoscelis ogasawarensis sp. nov., holotype, male. 40, 42, 44 &amp; 46, dorsal view; 41, 43, 45 &amp; 47. lateral view. Scale bar: 10.0 mm." pageId="13" pageNumber="475">Figs 40–41</figureCitation>
<figureCitation id="13462A38FFA5B347FF4CFC57FEBEFC23" box="[159,288,907,933]" captionStart="FIGURES 40–47" captionStartId="14.[151,264,1484,1508]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-15@14.[151,1435,181,1461]" captionTargetPageId="14" captionText="FIGURES 40–47. Habitus of Stenoscelis spp. 40 &amp; 41, Stenoscelis matobai sp. nov., holotype, male; 42 &amp; 43, Stenoscelis reniformis sp. nov., holotype, male; 44 &amp; 45, Stenoscelis gracilitarsis; 46 &amp; 47, Stenoscelis ogasawarensis sp. nov., holotype, male. 40, 42, 44 &amp; 46, dorsal view; 41, 43, 45 &amp; 47. lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804817" httpUri="https://zenodo.org/record/14804817/files/figure.png" pageId="13" pageNumber="475">Figs 40–41</figureCitation>
,
<figureCitation id="13462A38FFA5B347FEFFFC56FEECFC23" box="[300,370,906,933]" captionStart="FIGURES 48 –52" captionStartId="16.[151,264,1278,1302]" captionTargetBox="[156,1430,186,1249]" captionTargetId="figure-15@16.[151,1435,181,1254]" captionTargetPageId="16" captionText="FIGURES 48 –52. Stenoscelis matobai sp. nov. 48, aedeagus, dorsal view; 49, ditto, lateral view; 50, tegmen; 51, sternite VIII &amp; IX; 52, spermatheca. Arrows indicate the spatulate projection of sclerotized structure. Scale bar: 0.20 mm." pageId="13" pageNumber="475">48–52</figureCitation>
<figureCitation id="13462A38FFA5B347FEFFFC56FEECFC23" box="[300,370,906,933]" captionStart="FIGURES 48 –52" captionStartId="16.[151,264,1278,1302]" captionTargetBox="[156,1430,186,1249]" captionTargetId="figure-15@16.[151,1435,181,1254]" captionTargetPageId="16" captionText="FIGURES 48 –52. Stenoscelis matobai sp. nov. 48, aedeagus, dorsal view; 49, ditto, lateral view; 50, tegmen; 51, sternite VIII &amp; IX; 52, spermatheca. Arrows indicate the spatulate projection of sclerotized structure. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804819" httpUri="https://zenodo.org/record/14804819/files/figure.png" pageId="13" pageNumber="475">48–52</figureCitation>
&amp;
<figureCitation id="13462A38FFA5B347FE45FC57FE2DFC23" box="[406,435,907,933]" captionStart="FIGURE 74–77" captionStartId="22.[151,250,1490,1514]" captionTargetBox="[151,1436,181,1466]" captionTargetId="figure-15@22.[151,1436,181,1466]" captionTargetPageId="22" captionText="FIGURE 74–77. Heads of Stenoscelis spp. 74, Stenoscelis matobai sp. nov.; 75, Stenoscelis reniformis sp. nov.; 76, Stenoscelis gracilitarsis; 77, Stenoscelis ogasawarensis sp. nov.. Scale bar: 0.50 mm." pageId="13" pageNumber="475">74</figureCitation>
<figureCitation id="13462A38FFA5B347FE45FC57FE2DFC23" box="[406,435,907,933]" captionStart="FIGURE 74–77" captionStartId="22.[151,250,1490,1514]" captionTargetBox="[151,1436,181,1466]" captionTargetId="figure-15@22.[151,1436,181,1466]" captionTargetPageId="22" captionText="FIGURE 74–77. Heads of Stenoscelis spp. 74, Stenoscelis matobai sp. nov.; 75, Stenoscelis reniformis sp. nov.; 76, Stenoscelis gracilitarsis; 77, Stenoscelis ogasawarensis sp. nov.. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804829" httpUri="https://zenodo.org/record/14804829/files/figure.png" pageId="13" pageNumber="475">74</figureCitation>
)
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@ -111,7 +112,7 @@ Body length
<paragraph id="8BC236BDFFA5B347FF14FA83FEA9FA63" blockId="13.[151,1437,978,1977]" pageId="13" pageNumber="475">Prothorax 0.8 times as long as wide, widest before base, gradually narrowed anteriorly, weakly sinuate laterally; anterior margin weakly sinuate in middle or not; subapical constriction distinct on lateral side, obsolete on dorsal side; disc reticulate, rugosely punctate; punctures medium, separated by interspace almost as long as or narrower than diameter.</paragraph>
<paragraph id="8BC236BDFFA5B347FF14FA33FA81F967" blockId="13.[151,1437,978,1977]" pageId="13" pageNumber="475">Elytra 1.6–1.7 times as long as wide, 1.0 times as wide as and 1.9–2.0 times as long as prothorax, subparallel, conjointly rounded at apex; apex obtusely angulated in lateral view; striae rather deep, with medium punctures separated by interspace approximately 1–2 times as long as diameter; apical part of striae moderated, not dilated or reticulate; second stria distinctly separated from base; intervals somewhat convex, with an irregular row of small puncture; base of second to third or fourth interval rugosely punctate-granulate; apical half of ninth interval and tenth interval carinate; granules of intervals arranged in a single row, indistinct basally, distinct and conical on declivity; setae on declivity arranged irregularly in one or two rows, as long as or shorter than granules.</paragraph>
<paragraph id="8BC236BDFFA5B347FF14F937FDE6F83F" blockId="13.[151,1437,978,1977]" pageId="13" pageNumber="475">Aedeagal body 1.4 times as long as wide, 0.8 times as long as apodeme, subparallel, rapidly narrowed and weakly prolonged apically; apodeme attached laterally; endophallus with a large sclerotized structure on dorsal side of subapical part; the structure suborbicular, flat, with a large spatulate projection on dorsal side, with long hairs, with a separated part basally; tegmen without dorsal part and paramere, with distinct manubrium; spiculum gastrale gradually curved to right side apically, not expanded apically; sternite VIII divided, with several long setae apically; without spinulate apical membranous area.</paragraph>
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<paragraph id="8BC236BDFFA6B344FF44FA10FC63F9AA" blockId="14.[151,1437,1484,1581]" pageId="14" pageNumber="476">
<emphasis id="B909EAAFFFA6B344FF44FA10FEC9FA63" bold="true" box="[151,343,1484,1509]" pageId="14" pageNumber="476">FIGURES 40–47.</emphasis>
Habitus of
@ -177,7 +178,7 @@ specimens emerged from a dead broad-leaved tree (Isao Matoba pers. comm.).
<typeStatus id="54C6881FFFA6B344FE5BF88CFE6BF8EC" box="[392,501,1872,1898]" pageId="14" pageNumber="476" type="holotype">Holotype</typeStatus>
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(
<figureCitation id="13462A38FFA6B344FDD9F88CFD2BF8EC" box="[522,693,1872,1898]" captionStart="FIGURES 40–47" captionStartId="14.[151,264,1484,1508]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-15@14.[151,1435,181,1461]" captionTargetPageId="14" captionText="FIGURES 40–47. Habitus of Stenoscelis spp. 40 &amp; 41, Stenoscelis matobai sp. nov., holotype, male; 42 &amp; 43, Stenoscelis reniformis sp. nov., holotype, male; 44 &amp; 45, Stenoscelis gracilitarsis; 46 &amp; 47, Stenoscelis ogasawarensis sp. nov., holotype, male. 40, 42, 44 &amp; 46, dorsal view; 41, 43, 45 &amp; 47. lateral view. Scale bar: 10.0 mm." pageId="14" pageNumber="476">Figs 40 &amp; 41</figureCitation>
<figureCitation id="13462A38FFA6B344FDD9F88CFD2BF8EC" box="[522,693,1872,1898]" captionStart="FIGURES 40–47" captionStartId="14.[151,264,1484,1508]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-15@14.[151,1435,181,1461]" captionTargetPageId="14" captionText="FIGURES 40–47. Habitus of Stenoscelis spp. 40 &amp; 41, Stenoscelis matobai sp. nov., holotype, male; 42 &amp; 43, Stenoscelis reniformis sp. nov., holotype, male; 44 &amp; 45, Stenoscelis gracilitarsis; 46 &amp; 47, Stenoscelis ogasawarensis sp. nov., holotype, male. 40, 42, 44 &amp; 46, dorsal view; 41, 43, 45 &amp; 47. lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804817" httpUri="https://zenodo.org/record/14804817/files/figure.png" pageId="14" pageNumber="476">Figs 40 &amp; 41</figureCitation>
): male (
<collectionCode id="ED6CAE78FFA6B344FCC9F88CFCFFF8EC" box="[794,865,1872,1898]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="14" pageNumber="476" type="Herbarium">KUM</collectionCode>
), “Yarabu-dake, Ishigaki Is., Japan,

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<mods:namePart id="2633C5A2AB9DEF0465878D94B42E03D0">Inoue, Shûgo</mods:namePart>
<mods:affiliation id="3CB9786CC7DB377BDB703F97D50744AA">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka 819 - 0395, Japan</mods:affiliation>
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<mods:date id="A64DA32455B059D11C7F6AB28E6825C0">2025</mods:date>
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<treatment id="03D487ABFFA7B35BFF44FD0AFB59FD77" LSID="urn:lsid:plazi:treatment:03D487ABFFA7B35BFF44FD0AFB59FD77" httpUri="http://treatment.plazi.org/id/03D487ABFFA7B35BFF44FD0AFB59FD77" lastPageId="17" lastPageNumber="479" pageId="15" pageNumber="477">
<subSubSection id="C3676536FFA7B345FF44FD0AFC23FCBF" pageId="15" pageNumber="477" type="nomenclature">
@ -64,13 +65,13 @@ urn:lsid:zoobank.org:act:
<subSubSection id="C3676536FFA7B345FF44FC9FFE66FC19" pageId="15" pageNumber="477" type="description">
<paragraph id="8BC236BDFFA7B345FF44FC9FFE7AFCDB" blockId="15.[151,957,726,861]" box="[151,484,834,861]" pageId="15" pageNumber="477">
(
<figureCitation id="13462A38FFA7B345FF4CFC9FFEBEFCDB" box="[159,288,835,861]" captionStart="FIGURES 40–47" captionStartId="14.[151,264,1484,1508]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-15@14.[151,1435,181,1461]" captionTargetPageId="14" captionText="FIGURES 40–47. Habitus of Stenoscelis spp. 40 &amp; 41, Stenoscelis matobai sp. nov., holotype, male; 42 &amp; 43, Stenoscelis reniformis sp. nov., holotype, male; 44 &amp; 45, Stenoscelis gracilitarsis; 46 &amp; 47, Stenoscelis ogasawarensis sp. nov., holotype, male. 40, 42, 44 &amp; 46, dorsal view; 41, 43, 45 &amp; 47. lateral view. Scale bar: 10.0 mm." pageId="15" pageNumber="477">Figs 42–43</figureCitation>
<figureCitation id="13462A38FFA7B345FF4CFC9FFEBEFCDB" box="[159,288,835,861]" captionStart="FIGURES 40–47" captionStartId="14.[151,264,1484,1508]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-15@14.[151,1435,181,1461]" captionTargetPageId="14" captionText="FIGURES 40–47. Habitus of Stenoscelis spp. 40 &amp; 41, Stenoscelis matobai sp. nov., holotype, male; 42 &amp; 43, Stenoscelis reniformis sp. nov., holotype, male; 44 &amp; 45, Stenoscelis gracilitarsis; 46 &amp; 47, Stenoscelis ogasawarensis sp. nov., holotype, male. 40, 42, 44 &amp; 46, dorsal view; 41, 43, 45 &amp; 47. lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804817" httpUri="https://zenodo.org/record/14804817/files/figure.png" pageId="15" pageNumber="477">Figs 42–43</figureCitation>
,
<figureCitation id="13462A38FFA7B345FEFFFC9EFEEBFCDB" box="[300,373,834,861]" captionStart="FIGURES 53–57" captionStartId="17.[151,264,1708,1732]" captionTargetBox="[156,1430,782,1680]" captionTargetId="figure-281@17.[151,1435,777,1685]" captionTargetPageId="17" captionText="FIGURES 53–57. Stenoscelis reniformis sp. nov. 53, aedeagus, dorsal view; 54, ditto, lateral view; 55, tegmen; 56, sternite VIII &amp; IX; 57, spermatheca. Scale bar: 0.20 mm." pageId="15" pageNumber="477">53–57</figureCitation>
<figureCitation id="13462A38FFA7B345FEFFFC9EFEEBFCDB" box="[300,373,834,861]" captionStart="FIGURES 53–57" captionStartId="17.[151,264,1708,1732]" captionTargetBox="[156,1430,782,1680]" captionTargetId="figure-281@17.[151,1435,777,1685]" captionTargetPageId="17" captionText="FIGURES 53–57. Stenoscelis reniformis sp. nov. 53, aedeagus, dorsal view; 54, ditto, lateral view; 55, tegmen; 56, sternite VIII &amp; IX; 57, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804821" httpUri="https://zenodo.org/record/14804821/files/figure.png" pageId="15" pageNumber="477">53–57</figureCitation>
,
<figureCitation id="13462A38FFA7B345FE53FC9EFE02FCDB" box="[384,412,834,861]" captionStart="FIGURE 74–77" captionStartId="22.[151,250,1490,1514]" captionTargetBox="[151,1436,181,1466]" captionTargetId="figure-15@22.[151,1436,181,1466]" captionTargetPageId="22" captionText="FIGURE 74–77. Heads of Stenoscelis spp. 74, Stenoscelis matobai sp. nov.; 75, Stenoscelis reniformis sp. nov.; 76, Stenoscelis gracilitarsis; 77, Stenoscelis ogasawarensis sp. nov.. Scale bar: 0.50 mm." pageId="15" pageNumber="477">75</figureCitation>
<figureCitation id="13462A38FFA7B345FE53FC9EFE02FCDB" box="[384,412,834,861]" captionStart="FIGURE 74–77" captionStartId="22.[151,250,1490,1514]" captionTargetBox="[151,1436,181,1466]" captionTargetId="figure-15@22.[151,1436,181,1466]" captionTargetPageId="22" captionText="FIGURE 74–77. Heads of Stenoscelis spp. 74, Stenoscelis matobai sp. nov.; 75, Stenoscelis reniformis sp. nov.; 76, Stenoscelis gracilitarsis; 77, Stenoscelis ogasawarensis sp. nov.. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804829" httpUri="https://zenodo.org/record/14804829/files/figure.png" pageId="15" pageNumber="477">75</figureCitation>
&amp;
<figureCitation id="13462A38FFA7B345FE13FC9FFE43FCDB" box="[448,477,835,861]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." pageId="15" pageNumber="477">83</figureCitation>
<figureCitation id="13462A38FFA7B345FE13FC9FFE43FCDB" box="[448,477,835,861]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804831" httpUri="https://zenodo.org/record/14804831/files/figure.png" pageId="15" pageNumber="477">83</figureCitation>
)
</paragraph>
<paragraph id="8BC236BDFFA7B345FF44FC5BFE66FC19" blockId="15.[151,504,903,927]" box="[151,504,903,927]" pageId="15" pageNumber="477">
@ -108,7 +109,7 @@ is very similar to
<emphasis id="B909EAAFFFA7B345FBA2FC1BFA88FC66" box="[1137,1302,967,992]" italics="true" pageId="15" pageNumber="477">ogasawarensis</emphasis>
</taxonomicName>
, but can be distinguished from them by the sclerotized structure on the subapical part of endophallus being reniform and convex ventrally (
<figureCitation id="13462A38FFA7B345FED9FBD3FE18FBAF" box="[266,390,1038,1065]" captionStart="FIGURES 53–57" captionStartId="17.[151,264,1708,1732]" captionTargetBox="[156,1430,782,1680]" captionTargetId="figure-281@17.[151,1435,777,1685]" captionTargetPageId="17" captionText="FIGURES 53–57. Stenoscelis reniformis sp. nov. 53, aedeagus, dorsal view; 54, ditto, lateral view; 55, tegmen; 56, sternite VIII &amp; IX; 57, spermatheca. Scale bar: 0.20 mm." pageId="15" pageNumber="477">Fig. 53–54</figureCitation>
<figureCitation id="13462A38FFA7B345FED9FBD3FE18FBAF" box="[266,390,1038,1065]" captionStart="FIGURES 53–57" captionStartId="17.[151,264,1708,1732]" captionTargetBox="[156,1430,782,1680]" captionTargetId="figure-281@17.[151,1435,777,1685]" captionTargetPageId="17" captionText="FIGURES 53–57. Stenoscelis reniformis sp. nov. 53, aedeagus, dorsal view; 54, ditto, lateral view; 55, tegmen; 56, sternite VIII &amp; IX; 57, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804821" httpUri="https://zenodo.org/record/14804821/files/figure.png" pageId="15" pageNumber="477">Fig. 53–54</figureCitation>
) (flat and ovate in the other species).
</paragraph>
</subSubSection>
@ -126,7 +127,7 @@ Body length
<paragraph id="8BC236BDFFA7B345FF14F903FD41F82B" blockId="15.[151,1437,966,2037]" pageId="15" pageNumber="477">Aedeagal body 1.4 times as long as wide, 0.8 times as long as apodeme, subparallel, rapidly narrowed and weakly prolonged apically; apodeme attached laterally; endophallus with a large sclerotized structure on dorsal side of subapical part; the structure reniform, convex ventrally, with a pair of short projection on dorsal side, with short hairs, without separated part basally; tegmen without dorsal part and paramere, with distinct manubrium; spiculum gastrale gradually curved to right side apically, not expanded apically; sternite VIII divided, with several long setae apically; without spinulate apical membranous area.</paragraph>
<paragraph id="8BC236BDFFA7B345FF14F86BFE8CF873" blockId="15.[151,1437,966,2037]" pageId="15" pageNumber="477">Spiculum ventral not bifurcate at base; spermatheca with cornu falciform; collum not developed; ramus not developed.</paragraph>
</subSubSection>
<caption id="DF026635FFB8B35AFF44FB22FB78FABC" pageId="16" pageNumber="478" startId="16.[151,264,1278,1302]" targetBox="[156,1430,186,1249]" targetPageId="16" targetType="figure">
<caption id="DF026635FFB8B35AFF44FB22FB78FABC" ID-DOI="http://doi.org/10.5281/zenodo.14804819" ID-Zenodo-Dep="14804819" httpUri="https://zenodo.org/record/14804819/files/figure.png" pageId="16" pageNumber="478" startId="16.[151,264,1278,1302]" targetBox="[156,1430,186,1249]" targetPageId="16" targetType="figure">
<paragraph id="8BC236BDFFB8B35AFF44FB22FB78FABC" blockId="16.[151,1437,1278,1339]" pageId="16" pageNumber="478">
<emphasis id="B909EAAFFFB8B35AFF44FB22FEC4FA90" bold="true" box="[151,346,1278,1302]" pageId="16" pageNumber="478">FIGURES 48 –52.</emphasis>
<taxonomicName id="4C7D4D3EFFB8B35AFEB3FB23FDB4FA91" authorityName="Inoue" authorityYear="2025" box="[352,554,1279,1303]" class="Insecta" family="Curculionidae" genus="Stenoscelis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="478" phylum="Arthropoda" rank="species" species="matobai" status="sp. nov.">
@ -149,13 +150,13 @@ Is., and Iriomote Is.
<paragraph id="8BC236BDFFB8B35AFF14FA09FCE7F9B1" blockId="16.[151,1437,1421,2023]" pageId="16" pageNumber="478">
<emphasis id="B909EAAFFFB8B35AFF14FA09FE1CFA69" bold="true" box="[199,386,1493,1519]" pageId="16" pageNumber="478">Biological note.</emphasis>
The adults were collected from the tunnels bored in dead trees (
<figureCitation id="13462A38FFB8B35AFBABFA09FA89FA69" box="[1144,1303,1493,1519]" captionStart="FIGURE 84–89" captionStartId="24.[151,250,1649,1673]" captionTargetBox="[151,1435,181,1625]" captionTargetId="figure-15@24.[151,1435,181,1625]" captionTargetPageId="24" captionText="FIGURE 84–89. Habitats of Stenoscelis spp. 84, dead broad-leaved tree in Daisenzan-rindô, Ôita Pref., on which the adults of Stenoscelis acutipennis sp. nov. were found; 85, fallen branch of broad-leaved tree with tunnels in Mt. Hikosan, Fukuoka Pref., in which the adults of Stenoscelis brevithoracicus sp. nov. were found; 86, standing dead broad-leaved tree in Jôyama, Fukuoka Pref., in which the adults of Stenoscelis reniformis sp. nov. were found; 87, adult inside the same tree of Fig. 86; 88, fallen branch of broad-leaved tree with tunnels in the same locality of Fig. 86, in which the adults of S. reniformis were found; 89, larva inside the same branch of Fig. 88." pageId="16" pageNumber="478">Figs 86 &amp; 87</figureCitation>
<figureCitation id="13462A38FFB8B35AFBABFA09FA89FA69" box="[1144,1303,1493,1519]" captionStart="FIGURE 84–89" captionStartId="24.[151,250,1649,1673]" captionTargetBox="[151,1435,181,1625]" captionTargetId="figure-15@24.[151,1435,181,1625]" captionTargetPageId="24" captionText="FIGURE 84–89. Habitats of Stenoscelis spp. 84, dead broad-leaved tree in Daisenzan-rindô, Ôita Pref., on which the adults of Stenoscelis acutipennis sp. nov. were found; 85, fallen branch of broad-leaved tree with tunnels in Mt. Hikosan, Fukuoka Pref., in which the adults of Stenoscelis brevithoracicus sp. nov. were found; 86, standing dead broad-leaved tree in Jôyama, Fukuoka Pref., in which the adults of Stenoscelis reniformis sp. nov. were found; 87, adult inside the same tree of Fig. 86; 88, fallen branch of broad-leaved tree with tunnels in the same locality of Fig. 86, in which the adults of S. reniformis were found; 89, larva inside the same branch of Fig. 88." figureDoi="http://doi.org/10.5281/zenodo.14804833" httpUri="https://zenodo.org/record/14804833/files/figure.png" pageId="16" pageNumber="478">Figs 86 &amp; 87</figureCitation>
) and fallen branches of broad-leaved trees (
<figureCitation id="13462A38FFB8B35AFDDBFA25FDC2F995" box="[520,604,1529,1555]" captionStart="FIGURE 84–89" captionStartId="24.[151,250,1649,1673]" captionTargetBox="[151,1435,181,1625]" captionTargetId="figure-15@24.[151,1435,181,1625]" captionTargetPageId="24" captionText="FIGURE 84–89. Habitats of Stenoscelis spp. 84, dead broad-leaved tree in Daisenzan-rindô, Ôita Pref., on which the adults of Stenoscelis acutipennis sp. nov. were found; 85, fallen branch of broad-leaved tree with tunnels in Mt. Hikosan, Fukuoka Pref., in which the adults of Stenoscelis brevithoracicus sp. nov. were found; 86, standing dead broad-leaved tree in Jôyama, Fukuoka Pref., in which the adults of Stenoscelis reniformis sp. nov. were found; 87, adult inside the same tree of Fig. 86; 88, fallen branch of broad-leaved tree with tunnels in the same locality of Fig. 86, in which the adults of S. reniformis were found; 89, larva inside the same branch of Fig. 88." pageId="16" pageNumber="478">Fig. 88</figureCitation>
<figureCitation id="13462A38FFB8B35AFDDBFA25FDC2F995" box="[520,604,1529,1555]" captionStart="FIGURE 84–89" captionStartId="24.[151,250,1649,1673]" captionTargetBox="[151,1435,181,1625]" captionTargetId="figure-15@24.[151,1435,181,1625]" captionTargetPageId="24" captionText="FIGURE 84–89. Habitats of Stenoscelis spp. 84, dead broad-leaved tree in Daisenzan-rindô, Ôita Pref., on which the adults of Stenoscelis acutipennis sp. nov. were found; 85, fallen branch of broad-leaved tree with tunnels in Mt. Hikosan, Fukuoka Pref., in which the adults of Stenoscelis brevithoracicus sp. nov. were found; 86, standing dead broad-leaved tree in Jôyama, Fukuoka Pref., in which the adults of Stenoscelis reniformis sp. nov. were found; 87, adult inside the same tree of Fig. 86; 88, fallen branch of broad-leaved tree with tunnels in the same locality of Fig. 86, in which the adults of S. reniformis were found; 89, larva inside the same branch of Fig. 88." figureDoi="http://doi.org/10.5281/zenodo.14804833" httpUri="https://zenodo.org/record/14804833/files/figure.png" pageId="16" pageNumber="478">Fig. 88</figureCitation>
) in Jôyama,
<collectingRegion id="49B9F85FFFB8B35AFD27FA25FC08F995" box="[756,918,1529,1555]" country="Japan" name="Fukuoka" pageId="16" pageNumber="478">Fukuoka Pref.</collectingRegion>
The larvae were also found inside the fallen branch in Jôyama, from which the adults were collected (
<figureCitation id="13462A38FFB8B35AFCCBF9C1FCF3F9B1" box="[792,877,1565,1591]" captionStart="FIGURE 84–89" captionStartId="24.[151,250,1649,1673]" captionTargetBox="[151,1435,181,1625]" captionTargetId="figure-15@24.[151,1435,181,1625]" captionTargetPageId="24" captionText="FIGURE 84–89. Habitats of Stenoscelis spp. 84, dead broad-leaved tree in Daisenzan-rindô, Ôita Pref., on which the adults of Stenoscelis acutipennis sp. nov. were found; 85, fallen branch of broad-leaved tree with tunnels in Mt. Hikosan, Fukuoka Pref., in which the adults of Stenoscelis brevithoracicus sp. nov. were found; 86, standing dead broad-leaved tree in Jôyama, Fukuoka Pref., in which the adults of Stenoscelis reniformis sp. nov. were found; 87, adult inside the same tree of Fig. 86; 88, fallen branch of broad-leaved tree with tunnels in the same locality of Fig. 86, in which the adults of S. reniformis were found; 89, larva inside the same branch of Fig. 88." pageId="16" pageNumber="478">Fig. 89</figureCitation>
<figureCitation id="13462A38FFB8B35AFCCBF9C1FCF3F9B1" box="[792,877,1565,1591]" captionStart="FIGURE 84–89" captionStartId="24.[151,250,1649,1673]" captionTargetBox="[151,1435,181,1625]" captionTargetId="figure-15@24.[151,1435,181,1625]" captionTargetPageId="24" captionText="FIGURE 84–89. Habitats of Stenoscelis spp. 84, dead broad-leaved tree in Daisenzan-rindô, Ôita Pref., on which the adults of Stenoscelis acutipennis sp. nov. were found; 85, fallen branch of broad-leaved tree with tunnels in Mt. Hikosan, Fukuoka Pref., in which the adults of Stenoscelis brevithoracicus sp. nov. were found; 86, standing dead broad-leaved tree in Jôyama, Fukuoka Pref., in which the adults of Stenoscelis reniformis sp. nov. were found; 87, adult inside the same tree of Fig. 86; 88, fallen branch of broad-leaved tree with tunnels in the same locality of Fig. 86, in which the adults of S. reniformis were found; 89, larva inside the same branch of Fig. 88." figureDoi="http://doi.org/10.5281/zenodo.14804833" httpUri="https://zenodo.org/record/14804833/files/figure.png" pageId="16" pageNumber="478">Fig. 89</figureCitation>
).
</paragraph>
</subSubSection>
@ -173,7 +174,7 @@ The new species is named after its reniform sclerotized structure of endophallus
<typeStatus id="54C6881FFFB8B35AFE51F9B9FE71F9F9" box="[386,495,1637,1663]" pageId="16" pageNumber="478" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="13462A38FFB8B35AFDD2F9B9FD3CF9F9" box="[513,674,1637,1663]" captionStart="FIGURES 40–47" captionStartId="14.[151,264,1484,1508]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-15@14.[151,1435,181,1461]" captionTargetPageId="14" captionText="FIGURES 40–47. Habitus of Stenoscelis spp. 40 &amp; 41, Stenoscelis matobai sp. nov., holotype, male; 42 &amp; 43, Stenoscelis reniformis sp. nov., holotype, male; 44 &amp; 45, Stenoscelis gracilitarsis; 46 &amp; 47, Stenoscelis ogasawarensis sp. nov., holotype, male. 40, 42, 44 &amp; 46, dorsal view; 41, 43, 45 &amp; 47. lateral view. Scale bar: 10.0 mm." pageId="16" pageNumber="478">Figs 42 &amp; 43</figureCitation>
<figureCitation id="13462A38FFB8B35AFDD2F9B9FD3CF9F9" box="[513,674,1637,1663]" captionStart="FIGURES 40–47" captionStartId="14.[151,264,1484,1508]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-15@14.[151,1435,181,1461]" captionTargetPageId="14" captionText="FIGURES 40–47. Habitus of Stenoscelis spp. 40 &amp; 41, Stenoscelis matobai sp. nov., holotype, male; 42 &amp; 43, Stenoscelis reniformis sp. nov., holotype, male; 44 &amp; 45, Stenoscelis gracilitarsis; 46 &amp; 47, Stenoscelis ogasawarensis sp. nov., holotype, male. 40, 42, 44 &amp; 46, dorsal view; 41, 43, 45 &amp; 47. lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804817" httpUri="https://zenodo.org/record/14804817/files/figure.png" pageId="16" pageNumber="478">Figs 42 &amp; 43</figureCitation>
): male (
<collectionCode id="ED6CAE78FFB8B35AFCD1F9B9FCD7F9F9" box="[770,841,1637,1663]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="16" pageNumber="478" type="Herbarium">KUM</collectionCode>
), “

85
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@ -64,13 +65,13 @@ urn:lsid:zoobank.org:act:
<subSubSection id="C3676536FFADB34FFF44FEDFFE57FE9B" box="[151,457,259,286]" pageId="5" pageNumber="467" type="description">
<paragraph id="8BC236BDFFADB34FFF44FEDFFE57FE9B" blockId="5.[151,988,151,286]" box="[151,457,259,286]" pageId="5" pageNumber="467">
(
<figureCitation id="13462A38FFADB34FFF4CFEDFFE9AFE98" box="[159,260,259,286]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-28@3.[151,1435,181,1465]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Stenoscelis spp. 1 &amp; 2, Stenoscelis acutipennis, male; 3 &amp; 4, ditto, female; 5 &amp; 6, Stenoscelis brevithoracicus sp. nov., holotype, male. 1, 3, &amp; 5, dorsal view; 2, 4 &amp; 6, lateral view. Scale bar: 10.0 mm." pageId="5" pageNumber="467">Figs 5–6</figureCitation>
<figureCitation id="13462A38FFADB34FFF4CFEDFFE9AFE98" box="[159,260,259,286]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-28@3.[151,1435,181,1465]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Stenoscelis spp. 1 &amp; 2, Stenoscelis acutipennis, male; 3 &amp; 4, ditto, female; 5 &amp; 6, Stenoscelis brevithoracicus sp. nov., holotype, male. 1, 3, &amp; 5, dorsal view; 2, 4 &amp; 6, lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804801" httpUri="https://zenodo.org/record/14804801/files/figure.png" pageId="5" pageNumber="467">Figs 5–6</figureCitation>
,
<figureCitation id="13462A38FFADB34FFEC3FEDFFEC6FE9B" box="[272,344,259,285]" captionStart="FIGURES 12–16" captionStartId="5.[151,264,1346,1370]" captionTargetBox="[156,1430,459,1317]" captionTargetId="figure-93@5.[151,1435,453,1323]" captionTargetPageId="5" captionText="FIGURES 12–16. Stenoscelis brevithoracicus sp. nov. 12, aedeagus, dorsal view; 13, ditto, lateral view; 14, tegmen; 15, sternite VIII &amp; IX; 16, spermatheca. Scale bar: 0.20 mm." pageId="5" pageNumber="467">12–16</figureCitation>
<figureCitation id="13462A38FFADB34FFEC3FEDFFEC6FE9B" box="[272,344,259,285]" captionStart="FIGURES 12–16" captionStartId="5.[151,264,1346,1370]" captionTargetBox="[156,1430,459,1317]" captionTargetId="figure-93@5.[151,1435,453,1323]" captionTargetPageId="5" captionText="FIGURES 12–16. Stenoscelis brevithoracicus sp. nov. 12, aedeagus, dorsal view; 13, ditto, lateral view; 14, tegmen; 15, sternite VIII &amp; IX; 16, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804805" httpUri="https://zenodo.org/record/14804805/files/figure.png" pageId="5" pageNumber="467">12–16</figureCitation>
,
<figureCitation id="13462A38FFADB34FFEB7FEDFFE1EFE9B" box="[356,384,259,285]" captionStart="FIGURE 68–73" captionStartId="21.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-92@21.[151,1435,309,1165]" captionTargetPageId="21" captionText="FIGURE 68–73. Heads of Stenoscelis spp. 68, Stenoscelis acutipennis; 69, Stenoscelis brevithoracicus sp. nov.; 70, Stenoscelis cavirostris sp. nov., male; 71, ditto, female; 72, Stenoscelis longisetosus; 73, Stenoscelis aceri. Scale bar: 0.50 mm." pageId="5" pageNumber="467">69</figureCitation>
<figureCitation id="13462A38FFADB34FFEB7FEDFFE1EFE9B" box="[356,384,259,285]" captionStart="FIGURE 68–73" captionStartId="21.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-92@21.[151,1435,309,1165]" captionTargetPageId="21" captionText="FIGURE 68–73. Heads of Stenoscelis spp. 68, Stenoscelis acutipennis; 69, Stenoscelis brevithoracicus sp. nov.; 70, Stenoscelis cavirostris sp. nov., male; 71, ditto, female; 72, Stenoscelis longisetosus; 73, Stenoscelis aceri. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804827" httpUri="https://zenodo.org/record/14804827/files/figure.png" pageId="5" pageNumber="467">69</figureCitation>
&amp;
<figureCitation id="13462A38FFADB34FFE77FEDFFE5FFE9B" box="[420,449,259,285]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." pageId="5" pageNumber="467">79</figureCitation>
<figureCitation id="13462A38FFADB34FFE77FEDFFE5FFE9B" box="[420,449,259,285]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804831" httpUri="https://zenodo.org/record/14804831/files/figure.png" pageId="5" pageNumber="467">79</figureCitation>
)
</paragraph>
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@ -81,13 +82,13 @@ urn:lsid:zoobank.org:act:
<emphasis id="B909EAAFFFADB34FFECBFE97FDD4FEE3" box="[280,586,331,357]" italics="true" pageId="5" pageNumber="467">Stenoscelis brevithoracicus</emphasis>
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can be distinguished from the other Japanese congeners by the combination of the following character states: upper margin of scrobe directed to middle of eye (
<figureCitation id="13462A38FFADB34FFB8BFEB3FB00FE0F" box="[1112,1182,367,393]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-28@3.[151,1435,181,1465]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Stenoscelis spp. 1 &amp; 2, Stenoscelis acutipennis, male; 3 &amp; 4, ditto, female; 5 &amp; 6, Stenoscelis brevithoracicus sp. nov., holotype, male. 1, 3, &amp; 5, dorsal view; 2, 4 &amp; 6, lateral view. Scale bar: 10.0 mm." pageId="5" pageNumber="467">Fig. 6</figureCitation>
<figureCitation id="13462A38FFADB34FFB8BFEB3FB00FE0F" box="[1112,1182,367,393]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-28@3.[151,1435,181,1465]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Stenoscelis spp. 1 &amp; 2, Stenoscelis acutipennis, male; 3 &amp; 4, ditto, female; 5 &amp; 6, Stenoscelis brevithoracicus sp. nov., holotype, male. 1, 3, &amp; 5, dorsal view; 2, 4 &amp; 6, lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804801" httpUri="https://zenodo.org/record/14804801/files/figure.png" pageId="5" pageNumber="467">Fig. 6</figureCitation>
); setae on intervals of elytra arranged irregularly, as long as or shorter than granules on declivity (
<figureCitation id="13462A38FFADB34FFC30FE4FFBA9FE2B" box="[995,1079,403,429]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." pageId="5" pageNumber="467">Fig. 79</figureCitation>
<figureCitation id="13462A38FFADB34FFC30FE4FFBA9FE2B" box="[995,1079,403,429]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804831" httpUri="https://zenodo.org/record/14804831/files/figure.png" pageId="5" pageNumber="467">Fig. 79</figureCitation>
).
</paragraph>
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<emphasis id="B909EAAFFFADB34FFF44FA9EFEC6FADC" bold="true" box="[151,344,1346,1370]" pageId="5" pageNumber="467">FIGURES 12–16.</emphasis>
<taxonomicName id="4C7D4D3EFFADB34FFEB2FA9FFDE4FADD" authorityName="Inoue" authorityYear="2025" box="[353,634,1347,1371]" class="Insecta" family="Curculionidae" genus="Stenoscelis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="467" phylum="Arthropoda" rank="species" species="brevithoracicus" status="sp. nov.">
@ -112,16 +113,16 @@ Body length
<paragraph id="8BC236BDFFADB34CFF14F85DFAB1FE9B" blockId="5.[151,1436,1489,2019]" lastBlockId="6.[151,1437,151,1509]" lastPageId="6" lastPageNumber="468" pageId="5" pageNumber="467">Elytra 1.8–1.9 times as long as wide, 1.1 times as wide as and 2.4–2.6 times as long as prothorax, subparallel, conjointly rounded at apex; apex obtusely angulated in lateral view; striae rather deep, with medium punctures separated by interspace approximately 1–2 times as long as diameter; apical part of striae moderated, not dilated or reticulate; second stria distinctly separated from base; intervals somewhat convex, with an irregular row of small puncture; base of second to third or fourth interval rugosely punctate-granulate; apical half of ninth interval and tenth interval carinate; granules of intervals arranged in a single row, indistinct basally, distinct and conical on declivity; setae on declivity arranged irregularly in two or three rows, as long as or shorter than granules.</paragraph>
<paragraph id="8BC236BDFFAEB34CFF14FEFBFB46FE57" blockId="6.[151,1437,151,1509]" pageId="6" pageNumber="468">
Aedeagal body (
<figureCitation id="13462A38FFAEB34CFE56FEFBFD94FEC7" box="[389,522,295,321]" captionStart="FIGURES 12–16" captionStartId="5.[151,264,1346,1370]" captionTargetBox="[156,1430,459,1317]" captionTargetId="figure-93@5.[151,1435,453,1323]" captionTargetPageId="5" captionText="FIGURES 12–16. Stenoscelis brevithoracicus sp. nov. 12, aedeagus, dorsal view; 13, ditto, lateral view; 14, tegmen; 15, sternite VIII &amp; IX; 16, spermatheca. Scale bar: 0.20 mm." pageId="6" pageNumber="468">Figs 12, 13</figureCitation>
<figureCitation id="13462A38FFAEB34CFE56FEFBFD94FEC7" box="[389,522,295,321]" captionStart="FIGURES 12–16" captionStartId="5.[151,264,1346,1370]" captionTargetBox="[156,1430,459,1317]" captionTargetId="figure-93@5.[151,1435,453,1323]" captionTargetPageId="5" captionText="FIGURES 12–16. Stenoscelis brevithoracicus sp. nov. 12, aedeagus, dorsal view; 13, ditto, lateral view; 14, tegmen; 15, sternite VIII &amp; IX; 16, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804805" httpUri="https://zenodo.org/record/14804805/files/figure.png" pageId="6" pageNumber="468">Figs 12, 13</figureCitation>
) 2.0 times as long as wide, 1.5 times as long as apodeme, subparallel, rapidly narrowed and weakly prolonged apically; apodeme attached laterally; endophallus with a pair of spinulate sclerite on subapical part and a U-shaped sclerite on basal part; tegmen (
<figureCitation id="13462A38FFAEB34CFCAEFEB3FC4EFE0F" box="[893,976,367,393]" captionStart="FIGURES 12–16" captionStartId="5.[151,264,1346,1370]" captionTargetBox="[156,1430,459,1317]" captionTargetId="figure-93@5.[151,1435,453,1323]" captionTargetPageId="5" captionText="FIGURES 12–16. Stenoscelis brevithoracicus sp. nov. 12, aedeagus, dorsal view; 13, ditto, lateral view; 14, tegmen; 15, sternite VIII &amp; IX; 16, spermatheca. Scale bar: 0.20 mm." pageId="6" pageNumber="468">Fig. 14</figureCitation>
<figureCitation id="13462A38FFAEB34CFCAEFEB3FC4EFE0F" box="[893,976,367,393]" captionStart="FIGURES 12–16" captionStartId="5.[151,264,1346,1370]" captionTargetBox="[156,1430,459,1317]" captionTargetId="figure-93@5.[151,1435,453,1323]" captionTargetPageId="5" captionText="FIGURES 12–16. Stenoscelis brevithoracicus sp. nov. 12, aedeagus, dorsal view; 13, ditto, lateral view; 14, tegmen; 15, sternite VIII &amp; IX; 16, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804805" httpUri="https://zenodo.org/record/14804805/files/figure.png" pageId="6" pageNumber="468">Fig. 14</figureCitation>
) without dorsal part and paramere, with distinct manubrium; spiculum gastrale (
<figureCitation id="13462A38FFAEB34CFDB0FE4FFD26FE2B" box="[611,696,402,429]" captionStart="FIGURES 12–16" captionStartId="5.[151,264,1346,1370]" captionTargetBox="[156,1430,459,1317]" captionTargetId="figure-93@5.[151,1435,453,1323]" captionTargetPageId="5" captionText="FIGURES 12–16. Stenoscelis brevithoracicus sp. nov. 12, aedeagus, dorsal view; 13, ditto, lateral view; 14, tegmen; 15, sternite VIII &amp; IX; 16, spermatheca. Scale bar: 0.20 mm." pageId="6" pageNumber="468">Fig. 15</figureCitation>
<figureCitation id="13462A38FFAEB34CFDB0FE4FFD26FE2B" box="[611,696,402,429]" captionStart="FIGURES 12–16" captionStartId="5.[151,264,1346,1370]" captionTargetBox="[156,1430,459,1317]" captionTargetId="figure-93@5.[151,1435,453,1323]" captionTargetPageId="5" captionText="FIGURES 12–16. Stenoscelis brevithoracicus sp. nov. 12, aedeagus, dorsal view; 13, ditto, lateral view; 14, tegmen; 15, sternite VIII &amp; IX; 16, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804805" httpUri="https://zenodo.org/record/14804805/files/figure.png" pageId="6" pageNumber="468">Fig. 15</figureCitation>
) gradually curved to right side apically, not expanded apically; sternite VIII divided, with several long setae apically; without spinulate apical membranous area.
</paragraph>
<paragraph id="8BC236BDFFAEB34CFF14FE07FE16FD9F" blockId="6.[151,1437,151,1509]" pageId="6" pageNumber="468">
Spiculum ventral not bifurcate at base; spermatheca (
<figureCitation id="13462A38FFAEB34CFCF7FE07FCE9FE73" box="[804,887,475,501]" captionStart="FIGURES 12–16" captionStartId="5.[151,264,1346,1370]" captionTargetBox="[156,1430,459,1317]" captionTargetId="figure-93@5.[151,1435,453,1323]" captionTargetPageId="5" captionText="FIGURES 12–16. Stenoscelis brevithoracicus sp. nov. 12, aedeagus, dorsal view; 13, ditto, lateral view; 14, tegmen; 15, sternite VIII &amp; IX; 16, spermatheca. Scale bar: 0.20 mm." pageId="6" pageNumber="468">Fig. 16</figureCitation>
<figureCitation id="13462A38FFAEB34CFCF7FE07FCE9FE73" box="[804,887,475,501]" captionStart="FIGURES 12–16" captionStartId="5.[151,264,1346,1370]" captionTargetBox="[156,1430,459,1317]" captionTargetId="figure-93@5.[151,1435,453,1323]" captionTargetPageId="5" captionText="FIGURES 12–16. Stenoscelis brevithoracicus sp. nov. 12, aedeagus, dorsal view; 13, ditto, lateral view; 14, tegmen; 15, sternite VIII &amp; IX; 16, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804805" httpUri="https://zenodo.org/record/14804805/files/figure.png" pageId="6" pageNumber="468">Fig. 16</figureCitation>
) with cornu falciform; collum rather developed; ramus not developed.
</paragraph>
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@ -134,7 +135,7 @@ Spiculum ventral not bifurcate at base; spermatheca (
<paragraph id="8BC236BDFFAEB34CFF14FD9AFC2DFD03" blockId="6.[151,1437,151,1509]" pageId="6" pageNumber="468">
<emphasis id="B909EAAFFFAEB34CFF14FD9AFEE1FDE7" bold="true" box="[199,383,582,609]" pageId="6" pageNumber="468">Biological note.</emphasis>
The adults were collected from tunnels bored in the fallen branch of a broad-leaved tree (
<figureCitation id="13462A38FFAEB34CFABCFD9BFF2AFD03" captionStart="FIGURE 84–89" captionStartId="24.[151,250,1649,1673]" captionTargetBox="[151,1435,181,1625]" captionTargetId="figure-15@24.[151,1435,181,1625]" captionTargetPageId="24" captionText="FIGURE 84–89. Habitats of Stenoscelis spp. 84, dead broad-leaved tree in Daisenzan-rindô, Ôita Pref., on which the adults of Stenoscelis acutipennis sp. nov. were found; 85, fallen branch of broad-leaved tree with tunnels in Mt. Hikosan, Fukuoka Pref., in which the adults of Stenoscelis brevithoracicus sp. nov. were found; 86, standing dead broad-leaved tree in Jôyama, Fukuoka Pref., in which the adults of Stenoscelis reniformis sp. nov. were found; 87, adult inside the same tree of Fig. 86; 88, fallen branch of broad-leaved tree with tunnels in the same locality of Fig. 86, in which the adults of S. reniformis were found; 89, larva inside the same branch of Fig. 88." pageId="6" pageNumber="468">Fig. 85</figureCitation>
<figureCitation id="13462A38FFAEB34CFABCFD9BFF2AFD03" captionStart="FIGURE 84–89" captionStartId="24.[151,250,1649,1673]" captionTargetBox="[151,1435,181,1625]" captionTargetId="figure-15@24.[151,1435,181,1625]" captionTargetPageId="24" captionText="FIGURE 84–89. Habitats of Stenoscelis spp. 84, dead broad-leaved tree in Daisenzan-rindô, Ôita Pref., on which the adults of Stenoscelis acutipennis sp. nov. were found; 85, fallen branch of broad-leaved tree with tunnels in Mt. Hikosan, Fukuoka Pref., in which the adults of Stenoscelis brevithoracicus sp. nov. were found; 86, standing dead broad-leaved tree in Jôyama, Fukuoka Pref., in which the adults of Stenoscelis reniformis sp. nov. were found; 87, adult inside the same tree of Fig. 86; 88, fallen branch of broad-leaved tree with tunnels in the same locality of Fig. 86, in which the adults of S. reniformis were found; 89, larva inside the same branch of Fig. 88." figureDoi="http://doi.org/10.5281/zenodo.14804833" httpUri="https://zenodo.org/record/14804833/files/figure.png" pageId="6" pageNumber="468">Fig. 85</figureCitation>
) at Mt. Hikosan,
<collectingRegion id="49B9F85FFFAEB34CFEAAFDB7FDCBFD03" box="[377,597,619,645]" country="Japan" name="Fukuoka" pageId="6" pageNumber="468">Fukuoka Prefecture</collectingRegion>
(Naomichi Tsuji pers. comm.).

87
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@ -1,44 +1,45 @@
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(
<figureCitation id="13462A38FFAEB34CFF4CF97FFEBEF93B" box="[159,288,1699,1725]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." pageId="6" pageNumber="468">Figs 17–20</figureCitation>
<figureCitation id="13462A38FFAEB34CFF4CF97FFEBEF93B" box="[159,288,1699,1725]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804809" httpUri="https://zenodo.org/record/14804809/files/figure.png" pageId="6" pageNumber="468">Figs 17–20</figureCitation>
,
<figureCitation id="13462A38FFAEB34CFEFFF97EFEEBF93B" box="[300,373,1698,1725]" captionStart="FIGURES 25–29" captionStartId="9.[151,264,1065,1089]" captionTargetBox="[157,1430,187,1036]" captionTargetId="figure-28@9.[151,1435,181,1041]" captionTargetPageId="9" captionText="FIGURES 25–29. Stenoscelis cavirostris sp. nov. 25, aedeagus, dorsal view; 26, ditto, lateral view; 27, tegmen; 28, sternite VIII &amp; IX; 29, spermatheca. Scale bar: 0.20 mm." pageId="6" pageNumber="468">25–29</figureCitation>
<figureCitation id="13462A38FFAEB34CFEFFF97EFEEBF93B" box="[300,373,1698,1725]" captionStart="FIGURES 25–29" captionStartId="9.[151,264,1065,1089]" captionTargetBox="[157,1430,187,1036]" captionTargetId="figure-28@9.[151,1435,181,1041]" captionTargetPageId="9" captionText="FIGURES 25–29. Stenoscelis cavirostris sp. nov. 25, aedeagus, dorsal view; 26, ditto, lateral view; 27, tegmen; 28, sternite VIII &amp; IX; 29, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804811" httpUri="https://zenodo.org/record/14804811/files/figure.png" pageId="6" pageNumber="468">25–29</figureCitation>
,
<figureCitation id="13462A38FFAEB34CFE53F97FFE5BF93B" box="[384,453,1699,1725]" captionStart="FIGURE 68–73" captionStartId="21.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-92@21.[151,1435,309,1165]" captionTargetPageId="21" captionText="FIGURE 68–73. Heads of Stenoscelis spp. 68, Stenoscelis acutipennis; 69, Stenoscelis brevithoracicus sp. nov.; 70, Stenoscelis cavirostris sp. nov., male; 71, ditto, female; 72, Stenoscelis longisetosus; 73, Stenoscelis aceri. Scale bar: 0.50 mm." pageId="6" pageNumber="468">70–71</figureCitation>
<figureCitation id="13462A38FFAEB34CFE53F97FFE5BF93B" box="[384,453,1699,1725]" captionStart="FIGURE 68–73" captionStartId="21.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-92@21.[151,1435,309,1165]" captionTargetPageId="21" captionText="FIGURE 68–73. Heads of Stenoscelis spp. 68, Stenoscelis acutipennis; 69, Stenoscelis brevithoracicus sp. nov.; 70, Stenoscelis cavirostris sp. nov., male; 71, ditto, female; 72, Stenoscelis longisetosus; 73, Stenoscelis aceri. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804827" httpUri="https://zenodo.org/record/14804827/files/figure.png" pageId="6" pageNumber="468">70–71</figureCitation>
&amp;
<figureCitation id="13462A38FFAEB34CFE39F97FFD99F93B" box="[490,519,1699,1725]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." pageId="6" pageNumber="468">80</figureCitation>
<figureCitation id="13462A38FFAEB34CFE39F97FFD99F93B" box="[490,519,1699,1725]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804831" httpUri="https://zenodo.org/record/14804831/files/figure.png" pageId="6" pageNumber="468">80</figureCitation>
)
</paragraph>
</subSubSection>
@ -81,11 +82,11 @@ urn:lsid:zoobank.org:act:
<emphasis id="B909EAAFFFAEB34CFECAF937FD8BF882" box="[281,533,1771,1797]" italics="true" pageId="6" pageNumber="468">Stenoscelis cavirostris</emphasis>
</taxonomicName>
can be distinguished from the other congeners by the sexual dimorphism of the rostrum (dorsally concave in male and convex in female) (
<figureCitation id="13462A38FFAEB34CFCF0F8D3FC3FF8AF" box="[803,929,1807,1833]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." pageId="6" pageNumber="468">Figs 18, 20</figureCitation>
<figureCitation id="13462A38FFAEB34CFCF0F8D3FC3FF8AF" box="[803,929,1807,1833]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804809" httpUri="https://zenodo.org/record/14804809/files/figure.png" pageId="6" pageNumber="468">Figs 18, 20</figureCitation>
&amp;
<figureCitation id="13462A38FFAEB34CFC17F8D3FB92F8AF" box="[964,1036,1807,1833]" captionStart="FIGURE 68–73" captionStartId="21.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-92@21.[151,1435,309,1165]" captionTargetPageId="21" captionText="FIGURE 68–73. Heads of Stenoscelis spp. 68, Stenoscelis acutipennis; 69, Stenoscelis brevithoracicus sp. nov.; 70, Stenoscelis cavirostris sp. nov., male; 71, ditto, female; 72, Stenoscelis longisetosus; 73, Stenoscelis aceri. Scale bar: 0.50 mm." pageId="6" pageNumber="468">70–71</figureCitation>
<figureCitation id="13462A38FFAEB34CFC17F8D3FB92F8AF" box="[964,1036,1807,1833]" captionStart="FIGURE 68–73" captionStartId="21.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-92@21.[151,1435,309,1165]" captionTargetPageId="21" captionText="FIGURE 68–73. Heads of Stenoscelis spp. 68, Stenoscelis acutipennis; 69, Stenoscelis brevithoracicus sp. nov.; 70, Stenoscelis cavirostris sp. nov., male; 71, ditto, female; 72, Stenoscelis longisetosus; 73, Stenoscelis aceri. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804827" httpUri="https://zenodo.org/record/14804827/files/figure.png" pageId="6" pageNumber="468">70–71</figureCitation>
) and the apical part of striae widely dilated and reticulate, without distinct punctures (
<figureCitation id="13462A38FFAEB34CFD11F8EFFC89F8CB" box="[706,791,1843,1869]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." pageId="6" pageNumber="468">Fig. 80</figureCitation>
<figureCitation id="13462A38FFAEB34CFD11F8EFFC89F8CB" box="[706,791,1843,1869]" captionStart="FIGURE 78–83" captionStartId="23.[151,250,1621,1645]" captionTargetBox="[151,1435,741,1597]" captionTargetId="figure-247@23.[151,1435,741,1597]" captionTargetPageId="23" captionText="FIGURE 78–83. Apical part of elytra of Stenoscelis spp. 78, Stenoscelis acutipennis; 79, Stenoscelis brevithoracicus sp. nov.; 80, Stenoscelis cavirostris sp. nov., male; 81, Stenoscelis longisetosus; 82, Stenoscelis aceri; 83. Stenoscelis reniformis sp. nov. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804831" httpUri="https://zenodo.org/record/14804831/files/figure.png" pageId="6" pageNumber="468">Fig. 80</figureCitation>
).
</paragraph>
</subSubSection>
@ -95,7 +96,7 @@ can be distinguished from the other congeners by the sexual dimorphism of the ro
Body length 2.4–3.0 mm; black, rather shiny; mouth parts, antennae, tarsi, and apical part of tibiae reddish brown; setae yellowish brown.
</paragraph>
<paragraph id="8BC236BDFFAEB34DFF14F843FC47FF7F" blockId="6.[151,1436,1770,2013]" lastBlockId="7.[151,1437,151,249]" lastPageId="7" lastPageNumber="469" pageId="6" pageNumber="468">Rostrum 0.6 times as long as wide; dorsum concave in male, convex in female; dorsal surface reticulate, smoothly punctate; punctures minute, separated by interspace much longer than diameters; upper margin of scrobe directed to middle of eye; epistome with six long setae basally, with anterior margin obtusely projected at right side. Head with dorsal surface reticulate, smoothly punctate; punctures minute, separated by interspace much longer than diameter; interocular area with a shallow fovea; eye round, hardly convex.</paragraph>
<caption id="DF026635FFAFB34DFF44F9E5FDF5F91F" pageId="7" pageNumber="469" startId="7.[151,264,1593,1617]" targetBox="[151,1435,273,1570]" targetPageId="7" targetType="figure">
<caption id="DF026635FFAFB34DFF44F9E5FDF5F91F" ID-DOI="http://doi.org/10.5281/zenodo.14804809" ID-Zenodo-Dep="14804809" httpUri="https://zenodo.org/record/14804809/files/figure.png" pageId="7" pageNumber="469" startId="7.[151,264,1593,1617]" targetBox="[151,1435,273,1570]" targetPageId="7" targetType="figure">
<paragraph id="8BC236BDFFAFB34DFF44F9E5FDF5F91F" blockId="7.[151,1437,1593,1690]" pageId="7" pageNumber="469">
<emphasis id="B909EAAFFFAFB34DFF44F9E5FECAF9D7" bold="true" box="[151,340,1593,1617]" pageId="7" pageNumber="469">FIGURES 17–24.</emphasis>
Habitus of
@ -126,16 +127,16 @@ spp.
<paragraph id="8BC236BDFFAFB342FF14F87CFBC8FEE3" blockId="7.[151,1437,1772,2014]" lastBlockId="8.[151,1437,151,1041]" lastPageId="8" lastPageNumber="470" pageId="7" pageNumber="469">Elytra 1.8 times as long as wide, 1.0–1.1 times as wide as and 2.1–2.4 times as long as prothorax, subparallel, conjointly rounded at apex; apex obtusely angulated in lateral view; striae rather shallow, with medium punctures separated by interspace almost as long as diameter; apical part of striae widely dilated and reticulate, without distinct punctures; apical part of intervals becoming indistinct as striae becoming dilated; second stria distinctly separated from base; intervals somewhat convex, with an irregular row of small puncture; base of second to third or fourth interval rugosely punctate-granulate; apical half of ninth interval and tenth interval carinate; granules of intervals arranged in a single row, indistinct basally, distinct and conical on declivity; setae on declivity arranged in a single row, each behind a granule, distinct and more than twice as long as granules.</paragraph>
<paragraph id="8BC236BDFFA0B342FF14FEB3FC33FD9F" blockId="8.[151,1437,151,1041]" pageId="8" pageNumber="470">
Aedeagal body (
<figureCitation id="13462A38FFA0B342FE56FEB3FD92FE0F" box="[389,524,366,393]" captionStart="FIGURES 25–29" captionStartId="9.[151,264,1065,1089]" captionTargetBox="[157,1430,187,1036]" captionTargetId="figure-28@9.[151,1435,181,1041]" captionTargetPageId="9" captionText="FIGURES 25–29. Stenoscelis cavirostris sp. nov. 25, aedeagus, dorsal view; 26, ditto, lateral view; 27, tegmen; 28, sternite VIII &amp; IX; 29, spermatheca. Scale bar: 0.20 mm." pageId="8" pageNumber="470">Figs 25, 26</figureCitation>
<figureCitation id="13462A38FFA0B342FE56FEB3FD92FE0F" box="[389,524,366,393]" captionStart="FIGURES 25–29" captionStartId="9.[151,264,1065,1089]" captionTargetBox="[157,1430,187,1036]" captionTargetId="figure-28@9.[151,1435,181,1041]" captionTargetPageId="9" captionText="FIGURES 25–29. Stenoscelis cavirostris sp. nov. 25, aedeagus, dorsal view; 26, ditto, lateral view; 27, tegmen; 28, sternite VIII &amp; IX; 29, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804811" httpUri="https://zenodo.org/record/14804811/files/figure.png" pageId="8" pageNumber="470">Figs 25, 26</figureCitation>
) 2.0 times as long as wide, 2.0 times as long as apodeme, moderately arcuate laterally, gradually narrowed and moderately prolonged apically; apodeme attached sublaterally; endophallus with a pair of sclerite on subapical part; tegmen (
<figureCitation id="13462A38FFA0B342FD42FE6BFD7AFE57" box="[657,740,439,465]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." pageId="8" pageNumber="470">Fig. 22</figureCitation>
<figureCitation id="13462A38FFA0B342FD42FE6BFD7AFE57" box="[657,740,439,465]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804809" httpUri="https://zenodo.org/record/14804809/files/figure.png" pageId="8" pageNumber="470">Fig. 22</figureCitation>
) without dorsal part and paramere, with distinct manubrium; spiculum gastrale (
<figureCitation id="13462A38FFA0B342FEB9FE07FE24FE73" box="[362,442,475,501]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." pageId="8" pageNumber="470">Fig. 23</figureCitation>
<figureCitation id="13462A38FFA0B342FEB9FE07FE24FE73" box="[362,442,475,501]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804809" httpUri="https://zenodo.org/record/14804809/files/figure.png" pageId="8" pageNumber="470">Fig. 23</figureCitation>
) gradually curved to right side apically, not expanded apically; sternite VIII divided, with several long setae apically; without spinulate apical membranous area.
</paragraph>
<paragraph id="8BC236BDFFA0B342FF14FDFFFE26FDE7" blockId="8.[151,1437,151,1041]" pageId="8" pageNumber="470">
Spiculum ventral not bifurcate at base; spermatheca (
<figureCitation id="13462A38FFA0B342FCC9FDFFFCF2FDBB" box="[794,876,547,573]" captionStart="FIGURES 25–29" captionStartId="9.[151,264,1065,1089]" captionTargetBox="[157,1430,187,1036]" captionTargetId="figure-28@9.[151,1435,181,1041]" captionTargetPageId="9" captionText="FIGURES 25–29. Stenoscelis cavirostris sp. nov. 25, aedeagus, dorsal view; 26, ditto, lateral view; 27, tegmen; 28, sternite VIII &amp; IX; 29, spermatheca. Scale bar: 0.20 mm." pageId="8" pageNumber="470">Fig. 29</figureCitation>
<figureCitation id="13462A38FFA0B342FCC9FDFFFCF2FDBB" box="[794,876,547,573]" captionStart="FIGURES 25–29" captionStartId="9.[151,264,1065,1089]" captionTargetBox="[157,1430,187,1036]" captionTargetId="figure-28@9.[151,1435,181,1041]" captionTargetPageId="9" captionText="FIGURES 25–29. Stenoscelis cavirostris sp. nov. 25, aedeagus, dorsal view; 26, ditto, lateral view; 27, tegmen; 28, sternite VIII &amp; IX; 29, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804811" httpUri="https://zenodo.org/record/14804811/files/figure.png" pageId="8" pageNumber="470">Fig. 29</figureCitation>
) with cornu falciform; collum slightly developed; ramus slightly developed.
</paragraph>
</subSubSection>
@ -166,7 +167,7 @@ The new species is named after its dorsally concave rostrum in males.
<typeStatus id="54C6881FFFA0B342FE42FD27FE60FC93" box="[401,510,763,789]" pageId="8" pageNumber="470" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="13462A38FFA0B342FDCBFD27FD51FC93" box="[536,719,763,789]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." pageId="8" pageNumber="470">Figs 17 &amp; 18</figureCitation>
<figureCitation id="13462A38FFA0B342FDCBFD27FD51FC93" box="[536,719,763,789]" captionStart="FIGURES 17–24" captionStartId="7.[151,264,1593,1617]" captionTargetBox="[151,1435,273,1570]" captionTargetId="figure-73@7.[151,1435,273,1570]" captionTargetPageId="7" captionText="FIGURES 17–24. Habitus of Stenoscelis spp. 17 &amp; 18, Stenoscelis cavirostris sp. nov., holotype, male; 19 &amp; 20, ditto, female; 21 &amp; 22, Stenoscelis longisetosus, holotype, male; 23 &amp; 24, Stenoscelis aceri, holotype, male. 17, 19, 21 &amp; 23, dorsal view; 18, 20, 22 &amp; 24, lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804809" httpUri="https://zenodo.org/record/14804809/files/figure.png" pageId="8" pageNumber="470">Figs 17 &amp; 18</figureCitation>
): male (
<collectionCode id="ED6CAE78FFA0B342FCECFD27FC18FC93" box="[831,902,763,789]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="8" pageNumber="470" type="Herbarium">KUM</collectionCode>
), “

77
data/03/D4/87/03D487ABFFBCB35DFF44FE06FF78FD2F.xml
View file

@ -1,44 +1,45 @@
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<mods:title id="A95879141F40F29974AD2FE7842F8FEB">Revision of the genus Stenoscelis Wollaston (Coleoptera: Curculionidae: Cossoninae) of Japan</mods:title>
<document id="162B999A83F1EBD4C521CC768C94BA34" ID-DOI="10.11646/zootaxa.5583.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14804799" ID-ZooBank="E33B14CF-09AC-4EE4-9D5A-3C87056B6C38" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738699118123" checkinUser="plazi" docAuthor="Inoue, Shûgo" docDate="2025" docId="03D487ABFFBCB35DFF44FE06FF78FD2F" docLanguage="en" docName="zootaxa.5583.3.3.pdf" docOrigin="Zootaxa 5583 (3)" docSource="https://doi.org/10.11646/zootaxa.5583.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stenoscelis ogasawarensis Inoue 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="485" masterDocId="FFEDFFD3FFA8B34AFFD3FFDCFF9EFF86" masterDocTitle="Revision of the genus Stenoscelis Wollaston (Coleoptera: Curculionidae: Cossoninae) of Japan" masterLastPageNumber="490" masterPageNumber="462" pageNumber="482" updateTime="1738699772878" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CEB1FDF262097E2CCC68942155676987">Revision of the genus Stenoscelis Wollaston (Coleoptera: Curculionidae: Cossoninae) of Japan</mods:title>
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<mods:namePart id="0B0FBABE6F94C4F117B4EF8938280714">Inoue, Shûgo</mods:namePart>
<mods:affiliation id="38FEDF565D4554CC24ED60569383160D">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka 819 - 0395, Japan</mods:affiliation>
<mods:namePart id="E7CFD17493937449A1F306ACDE59DACF">Inoue, Shûgo</mods:namePart>
<mods:affiliation id="839F31E99C5A43094A65487A181D2FAF">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka 819 - 0395, Japan</mods:affiliation>
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<subSubSection id="C3676536FFBCB35EFF44FE06FC21FDBB" pageId="20" pageNumber="482" type="nomenclature">
@ -64,11 +65,11 @@ urn:lsid:zoobank.org:act:
<subSubSection id="C3676536FFBCB35EFF44FD9BFE25FDE7" box="[151,443,583,609]" pageId="20" pageNumber="482" type="description">
<paragraph id="8BC236BDFFBCB35EFF44FD9BFE25FDE7" blockId="20.[151,959,474,609]" box="[151,443,583,609]" pageId="20" pageNumber="482">
(
<figureCitation id="13462A38FFBCB35EFF4CFD9BFEBEFDE7" box="[159,288,583,609]" captionStart="FIGURES 40–47" captionStartId="14.[151,264,1484,1508]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-15@14.[151,1435,181,1461]" captionTargetPageId="14" captionText="FIGURES 40–47. Habitus of Stenoscelis spp. 40 &amp; 41, Stenoscelis matobai sp. nov., holotype, male; 42 &amp; 43, Stenoscelis reniformis sp. nov., holotype, male; 44 &amp; 45, Stenoscelis gracilitarsis; 46 &amp; 47, Stenoscelis ogasawarensis sp. nov., holotype, male. 40, 42, 44 &amp; 46, dorsal view; 41, 43, 45 &amp; 47. lateral view. Scale bar: 10.0 mm." pageId="20" pageNumber="482">Figs 46, 47</figureCitation>
<figureCitation id="13462A38FFBCB35EFF4CFD9BFEBEFDE7" box="[159,288,583,609]" captionStart="FIGURES 40–47" captionStartId="14.[151,264,1484,1508]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-15@14.[151,1435,181,1461]" captionTargetPageId="14" captionText="FIGURES 40–47. Habitus of Stenoscelis spp. 40 &amp; 41, Stenoscelis matobai sp. nov., holotype, male; 42 &amp; 43, Stenoscelis reniformis sp. nov., holotype, male; 44 &amp; 45, Stenoscelis gracilitarsis; 46 &amp; 47, Stenoscelis ogasawarensis sp. nov., holotype, male. 40, 42, 44 &amp; 46, dorsal view; 41, 43, 45 &amp; 47. lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804817" httpUri="https://zenodo.org/record/14804817/files/figure.png" pageId="20" pageNumber="482">Figs 46, 47</figureCitation>
,
<figureCitation id="13462A38FFBCB35EFEFFFD9BFEECFDE7" box="[300,370,583,609]" captionStart-0="FIGURES 58–62" captionStart-1="FIGURES 63–67" captionStartId-0="19.[151,264,1751,1775]" captionStartId-1="20.[151,264,1783,1807]" captionTargetBox-0="[156,1430,746,1723]" captionTargetBox-1="[156,1430,783,1755]" captionTargetId-0="figure-286@19.[151,1435,741,1728]" captionTargetId-1="figure-199@20.[151,1435,777,1760]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 58–62. Stenoscelis gracilitarsis. 58, aedeagus, dorsal view; 59, ditto, lateral view; 60, tegmen; 61, sternite VIII &amp; IX; 62, spermatheca. Scale bar: 0.20 mm." captionText-1="FIGURES 63–67. Stenoscelis ogasawarensis sp. nov. 63, aedeagus, dorsal view; 64, ditto, lateral view; 65, tegmen; 66, sternite VIII &amp; IX; 67, spermatheca. Scale bar: 0.20 mm." pageId="20" pageNumber="482">62–67</figureCitation>
<figureCitation id="13462A38FFBCB35EFEFFFD9BFEECFDE7" box="[300,370,583,609]" captionStart-0="FIGURES 58–62" captionStart-1="FIGURES 63–67" captionStartId-0="19.[151,264,1751,1775]" captionStartId-1="20.[151,264,1783,1807]" captionTargetBox-0="[156,1430,746,1723]" captionTargetBox-1="[156,1430,783,1755]" captionTargetId-0="figure-286@19.[151,1435,741,1728]" captionTargetId-1="figure-199@20.[151,1435,777,1760]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 58–62. Stenoscelis gracilitarsis. 58, aedeagus, dorsal view; 59, ditto, lateral view; 60, tegmen; 61, sternite VIII &amp; IX; 62, spermatheca. Scale bar: 0.20 mm." captionText-1="FIGURES 63–67. Stenoscelis ogasawarensis sp. nov. 63, aedeagus, dorsal view; 64, ditto, lateral view; 65, tegmen; 66, sternite VIII &amp; IX; 67, spermatheca. Scale bar: 0.20 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14804823" figureDoi-1="http://doi.org/10.5281/zenodo.14804825" httpUri-0="https://zenodo.org/record/14804823/files/figure.png" httpUri-1="https://zenodo.org/record/14804825/files/figure.png" pageId="20" pageNumber="482">62–67</figureCitation>
&amp;
<figureCitation id="13462A38FFBCB35EFE45FD9BFE2DFDE7" box="[406,435,583,609]" captionStart="FIGURE 74–77" captionStartId="22.[151,250,1490,1514]" captionTargetBox="[151,1436,181,1466]" captionTargetId="figure-15@22.[151,1436,181,1466]" captionTargetPageId="22" captionText="FIGURE 74–77. Heads of Stenoscelis spp. 74, Stenoscelis matobai sp. nov.; 75, Stenoscelis reniformis sp. nov.; 76, Stenoscelis gracilitarsis; 77, Stenoscelis ogasawarensis sp. nov.. Scale bar: 0.50 mm." pageId="20" pageNumber="482">77</figureCitation>
<figureCitation id="13462A38FFBCB35EFE45FD9BFE2DFDE7" box="[406,435,583,609]" captionStart="FIGURE 74–77" captionStartId="22.[151,250,1490,1514]" captionTargetBox="[151,1436,181,1466]" captionTargetId="figure-15@22.[151,1436,181,1466]" captionTargetPageId="22" captionText="FIGURE 74–77. Heads of Stenoscelis spp. 74, Stenoscelis matobai sp. nov.; 75, Stenoscelis reniformis sp. nov.; 76, Stenoscelis gracilitarsis; 77, Stenoscelis ogasawarensis sp. nov.. Scale bar: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.14804829" httpUri="https://zenodo.org/record/14804829/files/figure.png" pageId="20" pageNumber="482">77</figureCitation>
)
</paragraph>
</subSubSection>
@ -97,11 +98,11 @@ is very similar to
<emphasis id="B909EAAFFFBCB35EFB5DFD53FA89FD2F" box="[1166,1303,655,681]" italics="true" pageId="20" pageNumber="482">gracilitarsis</emphasis>
</taxonomicName>
, but can be distinguished from them by the sclerotized structure on the subapical part of endophallus being ovate, flat, with a pair of long projections dorsally, and separated basally (
<figureCitation id="13462A38FFBCB35EFCDAFD0BFC16FD77" box="[777,904,727,753]" captionStart="FIGURES 63–67" captionStartId="20.[151,264,1783,1807]" captionTargetBox="[156,1430,783,1755]" captionTargetId="figure-199@20.[151,1435,777,1760]" captionTargetPageId="20" captionText="FIGURES 63–67. Stenoscelis ogasawarensis sp. nov. 63, aedeagus, dorsal view; 64, ditto, lateral view; 65, tegmen; 66, sternite VIII &amp; IX; 67, spermatheca. Scale bar: 0.20 mm." pageId="20" pageNumber="482">Fig. 63–64</figureCitation>
<figureCitation id="13462A38FFBCB35EFCDAFD0BFC16FD77" box="[777,904,727,753]" captionStart="FIGURES 63–67" captionStartId="20.[151,264,1783,1807]" captionTargetBox="[156,1430,783,1755]" captionTargetId="figure-199@20.[151,1435,777,1760]" captionTargetPageId="20" captionText="FIGURES 63–67. Stenoscelis ogasawarensis sp. nov. 63, aedeagus, dorsal view; 64, ditto, lateral view; 65, tegmen; 66, sternite VIII &amp; IX; 67, spermatheca. Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14804825" httpUri="https://zenodo.org/record/14804825/files/figure.png" pageId="20" pageNumber="482">Fig. 63–64</figureCitation>
).
</paragraph>
</subSubSection>
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<caption id="DF026635FFBCB35EFF44F92BFD17F8B5" ID-DOI="http://doi.org/10.5281/zenodo.14804825" ID-Zenodo-Dep="14804825" httpUri="https://zenodo.org/record/14804825/files/figure.png" pageId="20" pageNumber="482" startId="20.[151,264,1783,1807]" targetBox="[156,1430,783,1755]" targetPageId="20" targetType="figure">
<paragraph id="8BC236BDFFBCB35EFF44F92BFD17F8B5" blockId="20.[151,1437,1783,1844]" pageId="20" pageNumber="482">
<emphasis id="B909EAAFFFBCB35EFF44F92BFECDF896" bold="true" box="[151,339,1783,1808]" pageId="20" pageNumber="482">FIGURES 63–67.</emphasis>
<taxonomicName id="4C7D4D3EFFBCB35EFE8BF924FDFAF889" authorityName="Inoue" authorityYear="2025" box="[344,612,1784,1808]" class="Insecta" family="Curculionidae" genus="Stenoscelis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="482" phylum="Arthropoda" rank="species" species="ogasawarensis" status="sp. nov.">
@ -121,7 +122,7 @@ Body length
; black, rather shiny; mouth parts, antennae, tarsi, and apical part of tibiae reddish brown; setae yellowish brown.
</paragraph>
<paragraph id="8BC236BDFFBCB35FFF14F813FB0CFE9B" blockId="20.[151,1436,1926,2025]" lastBlockId="21.[151,1437,151,285]" lastPageId="21" lastPageNumber="483" pageId="20" pageNumber="482">Rostrum 0.6 times as long as wide; dorsum convex; dorsal surface reticulate, rugosely punctate; punctures small, separated by interspace approximately 1–2 times as long as diameter; upper margin of scrobe directed to lower margin of eye; epistome with six long setae basally, with anterior margin obtusely projected at right side. Head with dorsal surface reticulate, smoothly punctate; punctures small, separated by interspace approximately 2–4 times as long as diameter; interocular area with a shallow fovea; eye round, hardly convex.</paragraph>
<caption id="DF026635FFBDB35FFF44FB78FAB4FB66" pageId="21" pageNumber="483" startId="21.[151,250,1188,1212]" targetBox="[151,1435,309,1165]" targetPageId="21" targetType="figure">
<caption id="DF026635FFBDB35FFF44FB78FAB4FB66" ID-DOI="http://doi.org/10.5281/zenodo.14804827" ID-Zenodo-Dep="14804827" httpUri="https://zenodo.org/record/14804827/files/figure.png" pageId="21" pageNumber="483" startId="21.[151,250,1188,1212]" targetBox="[151,1435,309,1165]" targetPageId="21" targetType="figure">
<paragraph id="8BC236BDFFBDB35FFF44FB78FAB4FB66" blockId="21.[151,1437,1188,1249]" pageId="21" pageNumber="483">
<emphasis id="B909EAAFFFBDB35FFF44FB78FEDBFB3B" bold="true" box="[151,325,1188,1213]" pageId="21" pageNumber="483">FIGURE 68–73.</emphasis>
Heads of
@ -164,7 +165,7 @@ spp.
<paragraph id="8BC236BDFFBDB35FFF14F93FFDE6F837" blockId="21.[151,1437,1331,2041]" pageId="21" pageNumber="483">Aedeagal body 1.7 times as long as wide, 0.8 times as long as apodeme, subparallel, gradually narrowed and weakly prolonged apically; apodeme attached laterally; endophallus with a large sclerotized structure on dorsal side of subapical part; the structure ovate, flat, with a pair of long projection on dorsal side, with long hairs, with a separated part basally; tegmen without dorsal part and paramere, with distinct manubrium; spiculum gastrale abruptly curved to right side apically, not expanded apically; sternite VIII divided, with several long setae apically; without spinulate apical membranous area.</paragraph>
<paragraph id="8BC236BDFFBDB35FFF14F867FE8CF87F" blockId="21.[151,1437,1331,2041]" pageId="21" pageNumber="483">Spiculum ventral not bifurcate at base; spermatheca with cornu falciform; collum not developed; ramus not developed.</paragraph>
</subSubSection>
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<caption id="DF026635FFBEB35CFF44FA0EFC1DF988" ID-DOI="http://doi.org/10.5281/zenodo.14804829" ID-Zenodo-Dep="14804829" httpUri="https://zenodo.org/record/14804829/files/figure.png" pageId="22" pageNumber="484" startId="22.[151,250,1490,1514]" targetBox="[151,1436,181,1466]" targetPageId="22" targetType="figure">
<paragraph id="8BC236BDFFBEB35CFF44FA0EFC1DF988" blockId="22.[151,1437,1490,1550]" pageId="22" pageNumber="484">
<emphasis id="B909EAAFFFBEB35CFF44FA0EFEDBFA6C" bold="true" box="[151,325,1490,1514]" pageId="22" pageNumber="484">FIGURE 74–77.</emphasis>
Heads of
@ -241,7 +242,7 @@ locality, the Ogasawara Islands.
<typeStatus id="54C6881FFFBEB35CFE50F92CFE6EF88C" box="[387,496,1776,1802]" pageId="22" pageNumber="484" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="13462A38FFBEB35CFDD0F92CFD39F88C" box="[515,679,1776,1802]" captionStart="FIGURES 40–47" captionStartId="14.[151,264,1484,1508]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-15@14.[151,1435,181,1461]" captionTargetPageId="14" captionText="FIGURES 40–47. Habitus of Stenoscelis spp. 40 &amp; 41, Stenoscelis matobai sp. nov., holotype, male; 42 &amp; 43, Stenoscelis reniformis sp. nov., holotype, male; 44 &amp; 45, Stenoscelis gracilitarsis; 46 &amp; 47, Stenoscelis ogasawarensis sp. nov., holotype, male. 40, 42, 44 &amp; 46, dorsal view; 41, 43, 45 &amp; 47. lateral view. Scale bar: 10.0 mm." pageId="22" pageNumber="484">Figs 46 &amp; 47</figureCitation>
<figureCitation id="13462A38FFBEB35CFDD0F92CFD39F88C" box="[515,679,1776,1802]" captionStart="FIGURES 40–47" captionStartId="14.[151,264,1484,1508]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-15@14.[151,1435,181,1461]" captionTargetPageId="14" captionText="FIGURES 40–47. Habitus of Stenoscelis spp. 40 &amp; 41, Stenoscelis matobai sp. nov., holotype, male; 42 &amp; 43, Stenoscelis reniformis sp. nov., holotype, male; 44 &amp; 45, Stenoscelis gracilitarsis; 46 &amp; 47, Stenoscelis ogasawarensis sp. nov., holotype, male. 40, 42, 44 &amp; 46, dorsal view; 41, 43, 45 &amp; 47. lateral view. Scale bar: 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14804817" httpUri="https://zenodo.org/record/14804817/files/figure.png" pageId="22" pageNumber="484">Figs 46 &amp; 47</figureCitation>
): male (
<collectionCode id="ED6CAE78FFBEB35CFCDBF92CFCD1F88C" box="[776,847,1776,1802]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="22" pageNumber="484" type="Herbarium">KUM</collectionCode>
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75
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<mods:title id="C0179931A2175A4D66A191A0F69AB2EB">Discovery of an endemism hotspot of freshwater mussels (Bivalvia: Unionidae) in Assam, with a description of two new genera</mods:title>
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<mods:namePart id="22A1CDC680D9EC5F224F13ADAD54EB1C">Bolotov, Ivan N.</mods:namePart>
<mods:affiliation id="56AB9BCD704C2370F193F04DE66E28BD">N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Av. 20, 163020 Arkhangelsk, Russia &amp; SSC / IUCN-Mollusc Specialist Group, Species Survival Commission, International Union for Conservation of Nature, CB 2 3 QZ Cambridge, United Kingdom</mods:affiliation>
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<mods:namePart id="5FD0CFD7F09D3D2DBF42A904D6484726">Sonowal, Jyotish</mods:namePart>
<mods:affiliation id="1DCDF876E88C42908DF9D0CCCE38D49C">Centre for Biotechnology and Bioinformatics, Dibrugarh University, 786004 Dibrugarh, Assam, India</mods:affiliation>
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<mods:namePart id="ECBDD1CA548E41C9BAF994575D4D312E">Kardong, Devid</mods:namePart>
<mods:affiliation id="D6AA4642E775C26D705BE79CAC445924">Department of Life Sciences, Dibrugarh University, 786004 Dibrugarh, Assam, India</mods:affiliation>
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<mods:namePart id="BC67102AE1B3CD61292B71476EAF27EE">Pasupuleti, Rajeev</mods:namePart>
<mods:affiliation id="F86598FB43D15EBE381C50C50A9D5938">Institute of Molecular Biotechnology (IMBT), Technical University of Graz, Petersgasse 14, 8010 Graz, Austria</mods:affiliation>
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<mods:namePart id="B6A6EF25F53BF4A0C857E6EDD7A0CDC2">Subba Rao, Nalluri V.</mods:namePart>
<mods:affiliation id="247CD97B03C83FABE059E591E0CFB27D">Hyderabad, India (retired)</mods:affiliation>
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<mods:namePart id="AE0EC4F6D49F6B8B3E1934D145DA7C24">Unnikrishnan, Suresh Kumar</mods:namePart>
<mods:affiliation id="7A596FCEB6E30FE17E6DBF281EF7EAE5">Regional Facility for DNA Fingerprinting (RFDF), Rajiv Gandhi Centre for Biotechnology (RGCB), 695014 Thiruvananthapuram, Kerala, India</mods:affiliation>
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<mods:namePart id="29A830DB23F1FB8D3A6082386CF8E08E">Gofarov, Mikhail Y.</mods:namePart>
<mods:affiliation id="956DD34C974407B21E4A5617377FC961">N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Av. 20, 163020 Arkhangelsk, Russia</mods:affiliation>
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<mods:namePart id="B37A5244E60A1906AE871375AEAB4C6B">Kondakov, Alexander V.</mods:namePart>
<mods:affiliation id="2F56E3848D05E68146AD258836C25A7D">N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Av. 20, 163020 Arkhangelsk, Russia</mods:affiliation>
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<mods:namePart id="92D1654126B37F74816AC18F4BA10E51">Konopleva, Ekaterina S.</mods:namePart>
<mods:affiliation id="B26CF14C149434E9EE2C3B498A3F3FA3">N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Av. 20, 163020 Arkhangelsk, Russia</mods:affiliation>
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<mods:namePart id="9771F7ABA0B6CDF7C354D6523AED611D">Lyubas, Artem A.</mods:namePart>
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<mods:namePart id="AC0997507EDB5A71C3A7A90DEB3438C0">Vikhrev, Ilya V.</mods:namePart>
<mods:affiliation id="95C8524A03D1BB0895A8D21E28D523D9">N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Av. 20, 163020 Arkhangelsk, Russia &amp; SSC / IUCN-Mollusc Specialist Group, Species Survival Commission, International Union for Conservation of Nature, CB 2 3 QZ Cambridge, United Kingdom</mods:affiliation>
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Genus
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<emphasis id="23F4EAF9FF84FFE0FF35F9D104256F76" bold="true" box="[200,292,1574,1601]" italics="true" pageId="5" pageNumber="6">Indonaia</emphasis>
<bibRefCitation id="75114B1AFF84FFE0FED4F9D004C16F77" author="Prashad B" box="[297,448,1574,1601]" pageId="5" pageNumber="6" pagination="143 - 8" refId="ref15383" refString="Prashad B. The marsupium and glochidium of some Unionidae and on the Indian species hitherto assigned to the genus Nodularia. Records of the Indian Museum 1918; 15: 143 - 8." type="journal article" year="1918">Prashad, 1918</bibRefCitation>
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(
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species:
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<emphasis id="23F4EAF9FF84FFE0FDA6F9D107EC6F76" bold="true" box="[603,749,1574,1601]" italics="true" pageId="5" pageNumber="6">Unio caeruleus</emphasis>
Lea, 1831
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; by original designation) (
<bibRefCitation id="75114B1AFF84FFE0FDD9F9B107B46F57" author="Prashad B" box="[548,693,1606,1632]" pageId="5" pageNumber="6" pagination="143 - 8" refId="ref15383" refString="Prashad B. The marsupium and glochidium of some Unionidae and on the Indian species hitherto assigned to the genus Nodularia. Records of the Indian Museum 1918; 15: 143 - 8." type="journal article" year="1918">Prashad 1918</bibRefCitation>
).
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<paragraph id="113F36EBFF84FFE0FE88F99804F66FBE" blockId="5.[373,503,1647,1674]" box="[373,503,1647,1674]" pageId="5" pageNumber="6">
<emphasis id="23F4EAF9FF84FFE0FE88F99804F66FBE" bold="true" box="[373,503,1647,1674]" pageId="5" pageNumber="6">
(
<figureCitation id="89BB2A6EFF84FFE0FE82F98704ED6FBD" box="[383,492,1648,1674]" captionStart="Figure 3" captionStartId="7.[113,178,1711,1735]" captionTargetBox="[114,1458,144,1683]" captionTargetId="figure-69@7.[114,1458,144,1683]" captionTargetPageId="7" captionText="Figure 3. Shells of Indochinellini species from Assam (outer view of the left valve and inner view of the right valve).(A–H) Indonaia corbis (Hanley, 1856) stat. rev.: topotypes IX-BDH-1 (A), IS-TLM-3 (B), IA-GEL-5 (C), X2-SES (D), X1-SES (E), I2-SES-2 (F), TZ-BDH-2 (G); holotype of Trapezoideus dhanushori Annandale and Prashad, 1921 with its labels (ZSI M11962/2), Dhanushori Stream at a distance of about a mile from Dimapur, Assam (H). (I–K) Pseudoscabies scobina (Hanley, 1856) gen.et comb. nov.: topotypes SC-SES-1 (I), IO-BDH-5 (J), and IO-BDH-4 (K). Scale bars = 10 mm. The shell vouchers of the new samples are deposited in FBRC ZSI (Supporting Information, Dataset S1). Photos: J. Sonowal and D. Kardong (A–G, I–K); A.Mukhopadhyaya and S. Sajan (H)." pageId="5" pageNumber="6">Fig. 3A–H</figureCitation>
)
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<paragraph id="113F36EBFF84FFE0FF8CF95006836F55" blockId="5.[111,764,1702,1978]" lastBlockId="5.[809,1460,1359,1634]" pageId="5" pageNumber="6">
<emphasis id="23F4EAF9FF84FFE0FF8CF95005E36F89" box="[113,226,1703,1726]" italics="true" pageId="5" pageNumber="6">Comments:</emphasis>
This genus contains 12 accepted species.Based on our new phylogenetic and morphological data, we consider
<taxonomicName id="D6804D68FF84FFE0FD5FF931044A6FCA" authority="(Hanley, 1856)" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Hanley" baseAuthorityYear="1856" class="Bivalvia" family="Unionidae" genus="Indonaia" kingdom="Animalia" order="Unionida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="corbis" status="stat. nov.">
<emphasis id="23F4EAF9FF84FFE0FD5FF93105A86FCA" italics="true" pageId="5" pageNumber="6">Indonaia corbis</emphasis>
(
<bibRefCitation id="75114B1AFF84FFE0FF48F91204426FCA" author="Hanley S" box="[181,323,1765,1789]" pageId="5" pageNumber="6" refId="ref14239" refString="Hanley S. An Illustrated and Descriptive Catalogue of Recent Bivalve Shells. London: Williams and Norgate, 1856." type="book" year="1856">Hanley, 1856</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="38C75782FF84FFE0FEB2F91104A36FCA" box="[335,418,1766,1789]" pageId="5" pageNumber="6">stat. rev.</taxonomicNameLabel>
as an accepted species endemic to the Brahmaputra River basin (
<tableCitation id="5C020350FF84FFE0FE5FF8F304ED6E2B" box="[418,492,1796,1820]" captionStart="Table 1" captionStartId="4.[129,184,143,167]" captionTargetBox="[128,1473,189,1998]" captionText="Table 1. Checklist of freshwater mussel species (Unionidae:Parreysiinae) recorded from the Brahmaputra River basin in Assam, India." pageId="5" pageNumber="6">Table 1</tableCitation>
;
<figureCitation id="89BB2A6EFF84FFE0FE05F8F307606E2B" box="[504,609,1796,1820]" captionStart="Figure 3" captionStartId="7.[113,178,1711,1735]" captionTargetBox="[114,1458,144,1683]" captionTargetId="figure-69@7.[114,1458,144,1683]" captionTargetPageId="7" captionText="Figure 3. Shells of Indochinellini species from Assam (outer view of the left valve and inner view of the right valve).(A–H) Indonaia corbis (Hanley, 1856) stat. rev.: topotypes IX-BDH-1 (A), IS-TLM-3 (B), IA-GEL-5 (C), X2-SES (D), X1-SES (E), I2-SES-2 (F), TZ-BDH-2 (G); holotype of Trapezoideus dhanushori Annandale and Prashad, 1921 with its labels (ZSI M11962/2), Dhanushori Stream at a distance of about a mile from Dimapur, Assam (H). (I–K) Pseudoscabies scobina (Hanley, 1856) gen.et comb. nov.: topotypes SC-SES-1 (I), IO-BDH-5 (J), and IO-BDH-4 (K). Scale bars = 10 mm. The shell vouchers of the new samples are deposited in FBRC ZSI (Supporting Information, Dataset S1). Photos: J. Sonowal and D. Kardong (A–G, I–K); A.Mukhopadhyaya and S. Sajan (H)." pageId="5" pageNumber="6">Fig. 3A–H</figureCitation>
). Additionally, two nominal species are transferred here from
<taxonomicName id="D6804D68FF84FFE0FDABF8D307AF6E0B" authorityName="Prashad" authorityYear="1918" box="[598,686,1828,1852]" class="Bivalvia" family="Unionidae" genus="Indonaia" kingdom="Animalia" order="Unionida" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF84FFE0FDABF8D307AF6E0B" box="[598,686,1828,1852]" italics="true" pageId="5" pageNumber="6">Indonaia</emphasis>
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to new genera (see below). Earlier, we separated three species groups in this genus: (i) the
<taxonomicName id="D6804D68FF84FFE0FECCF89504816E4D" authorityName="Imsong and Murali" authorityYear="2023" box="[305,384,1890,1914]" class="Bivalvia" family="Unionidae" genus="Indonaia" kingdom="Animalia" order="Unionida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="caerulea">
<emphasis id="23F4EAF9FF84FFE0FECCF89504816E4D" box="[305,384,1890,1914]" italics="true" pageId="5" pageNumber="6">caerulea</emphasis>
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-group; (ii) the
<emphasis id="23F4EAF9FF84FFE0FDD9F89507806E4D" box="[548,641,1890,1914]" italics="true" pageId="5" pageNumber="6">cylindrica</emphasis>
-group; and (iii) the
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Bolotov
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2022a
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).
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belongs to the first group that is well resolved in both
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-based and combined phylogenies (
<figureCitation id="89BB2A6EFF84FFE0FBB7FAB801836C50" box="[1098,1154,1359,1383]" captionStart="Figure 2" captionStartId="6.[129,194,1839,1863]" captionTargetBox="[254,1349,144,1811]" captionTargetId="figure-8@6.[254,1349,144,1811]" captionTargetPageId="6" captionText="Figure 2. Maximum likelihood (IQ-TREE v.1.6.12) phylogeny of the Parreysiinae based on the combined alignment of the two mitochondrial and one nuclear gene fragments (three codons of COI + 16S rRNA + 28S rRNA). The labels of species recorded from Assam (Brahmaputra River basin) are red.The scale bar indicates the branch lengths.The information on DNA sequences is presented in the Supporting Information, Dataset S2. Outgroup taxa are not shown." pageId="5" pageNumber="6">Fig. 2</figureCitation>
; Supporting Information,
<figureCitation id="89BB2A6EFF84FFE0FA70FAB806456CB0" captionStart="Figure 1" captionStartId="2.[129,194,1112,1136]" captionTargetBox="[128,1475,144,1084]" captionTargetId="figure-317@2.[128,1475,144,1084]" captionTargetPageId="2" captionText="Figure 1. Map of the Indian subcontinent with location of the study region (the frame) (A) and enlarged map of the study region in northeastern India (B). Assam is indicated by green filling (A) and cross-hatching (B). Green circles with numbers indicate the sampling localities in Assam:Burhidihing River, Dibrugarh (1); Gelua River, Dhemaji (2); an oxbow lake of Dhansiri River, Golaghat (3); old channel/pond, Telam River basin, Telam, Dhemaji (4); an oxbow lake of Tongani River (Tongani beel), Dhemaji (5); Dhansiri River, Golaghat (6); Sesa River, Dibrugarh (7); and a pond, Sutar Gaon, Kamrup (Rural) District (8). The maps were created using ESRI ArcGIS 10 software (www.esri. com/arcgis); the topographic base of the maps was created with Natural Earth Free Vector and Raster Map Data (www.naturalearthdata.com), Global Self-consistent Hierarchical High-resolution Geography (https://www.soest.hawaii.edu/wessel/gshhg), GEBCO_2022 (https://www. gebco.net/data_and_products/historical_data_sets/#gebco_2022), and Global Edge-matched Subnational Boundaries (https://fieldmaps. io). Maps: Mikhail Yu. Gofarov." pageId="5" pageNumber="6">Fig. S1</figureCitation>
). The
<emphasis id="23F4EAF9FF84FFE0FC79FA9806E36CB0" box="[900,994,1391,1415]" italics="true" pageId="5" pageNumber="6">cylindrica</emphasis>
-group is supported by the
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phylogeny but is unresolved (non-monophyletic) in the three-locus combined tree. However, this group contains two
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<emphasis id="23F4EAF9FF84FFE0FB4FFA59000D6CF2" box="[1202,1292,1454,1477]" italics="true" pageId="5" pageNumber="6">Parreysia</emphasis>
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-like species that could be distinguished from other
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<emphasis id="23F4EAF9FF84FFE0FB6FFA3A01EB6CD2" box="[1170,1258,1485,1509]" italics="true" pageId="5" pageNumber="6">Indonaia</emphasis>
species
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by having a thick, massive, ovate shell (
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Bolotov
<emphasis id="23F4EAF9FF84FFE0FB6FFA1A01C16F33" box="[1170,1216,1516,1540]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
2022a
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) and seems to be a pair of sister-species rather than two unrelated taxa. This issue needs to be checked in the future using an expanded set of DNA markers.
</paragraph>
</subSubSection>
</treatment>
</document>

434
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Genus
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</emphasis>
<subSubSection id="599A6560FF84FFE0FBCEF90B01DF6E6C" pageId="5" pageNumber="6" type="description">
<paragraph id="113F36EBFF84FFE0FBCEF90B01AA6E21" blockId="5.[810,1459,1788,1883]" box="[1075,1195,1788,1815]" pageId="5" pageNumber="6">
<emphasis id="23F4EAF9FF84FFE0FBCEF90B01AA6E21" bold="true" box="[1075,1195,1788,1815]" pageId="5" pageNumber="6">
(
<figureCitation id="89BB2A6EFF84FFE0FBC0F90A01A16E20" box="[1085,1184,1789,1815]" captionStart="Figure 3" captionStartId="7.[113,178,1711,1735]" captionTargetBox="[114,1458,144,1683]" captionTargetId="figure-69@7.[114,1458,144,1683]" captionTargetPageId="7" captionText="Figure 3. Shells of Indochinellini species from Assam (outer view of the left valve and inner view of the right valve).(A–H) Indonaia corbis (Hanley, 1856) stat. rev.: topotypes IX-BDH-1 (A), IS-TLM-3 (B), IA-GEL-5 (C), X2-SES (D), X1-SES (E), I2-SES-2 (F), TZ-BDH-2 (G); holotype of Trapezoideus dhanushori Annandale and Prashad, 1921 with its labels (ZSI M11962/2), Dhanushori Stream at a distance of about a mile from Dimapur, Assam (H). (I–K) Pseudoscabies scobina (Hanley, 1856) gen.et comb. nov.: topotypes SC-SES-1 (I), IO-BDH-5 (J), and IO-BDH-4 (K). Scale bars = 10 mm. The shell vouchers of the new samples are deposited in FBRC ZSI (Supporting Information, Dataset S1). Photos: J. Sonowal and D. Kardong (A–G, I–K); A.Mukhopadhyaya and S. Sajan (H)." pageId="5" pageNumber="6">Fig. 3I–K</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="113F36EBFF84FFE0FCD7F8D401DF6E6C" blockId="5.[810,1459,1788,1883]" pageId="5" pageNumber="6">
LSID: https://zoobank.org/
<uri id="65113AE9FF84FFE0FB8AF8D401DF6E6C" pageId="5" pageNumber="6">
urn:lsid:zoobank.org:act:
<uuid id="65260C3EFF84FFE0FA86F8D401DF6E6C" pageId="5" pageNumber="6">6366 F102-F465-4FBD-BFF2-D8BF8FEE87C7</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="599A6560FF84FFEFFCD7F87506E26EE1" lastPageId="10" lastPageNumber="11" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="113F36EBFF84FFEFFCD7F87506E26EE1" blockId="5.[810,1458,1922,1977]" lastBlockId="10.[825,1475,1888,2006]" lastPageId="10" lastPageNumber="11" pageId="5" pageNumber="6">
<emphasis id="23F4EAF9FF84FFE0FCD7F87501086EAD" box="[810,1033,1922,1946]" italics="true" pageId="5" pageNumber="6">Differential diagnosis:</emphasis>
Phylogenetically, the new genus is the sister-lineage of
<taxonomicName id="D6804D68FF84FFE0FC2BF85601316E8E" authorityName="Prashad" authorityYear="1918" box="[982,1072,1953,1977]" class="Bivalvia" family="Unionidae" genus="Indonaia" kingdom="Animalia" order="Unionida" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF84FFE0FC2BF85601316E8E" box="[982,1072,1953,1977]" italics="true" pageId="5" pageNumber="6">Indonaia</emphasis>
</taxonomicName>
. However, it does not correspond to the latter genus by having a specific, strongly-developed sculpture over the shell disc, i.e. V- or W-shaped ridges at the middle of the shell, and waved and arcuate ridges at the anterior and posterior parts of the shell, respectively, while corresponding depressions display the same pattern in the inner side of the shell. Conchologically, the new genus is similar to the genera
<taxonomicName id="D6804D68FF8BFFEFFED7F897077D6E4F" authority="Rochebrune, 1882" authorityName="Rochebrune" authorityYear="1882" box="[298,636,1888,1912]" class="Bivalvia" family="Unionidae" genus="Harmandia" kingdom="Animalia" order="Unionida" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8BFFEFFED7F89704A16E4F" box="[298,416,1888,1912]" italics="true" pageId="10" pageNumber="11">Harmandia</emphasis>
Rochebrune, 1882
</taxonomicName>
and
<taxonomicName id="D6804D68FF8BFFEFFD3DF89705F66EAF" authority="Haas, 1911" authorityName="Haas" authorityYear="1911" class="Bivalvia" family="Unionidae" genus="Scabies" kingdom="Animalia" order="Unionida" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8BFFEFFD3DF89706086E4F" box="[704,777,1888,1912]" italics="true" pageId="10" pageNumber="11">Scabies</emphasis>
Haas, 1911
</taxonomicName>
(
<taxonomicName id="D6804D68FF8BFFEFFEFBF88804A26EA0" box="[262,419,1919,1943]" class="Bivalvia" family="Unionidae" kingdom="Animalia" order="Unionida" pageId="10" pageNumber="11" phylum="Mollusca" rank="tribe" tribe="Indochinellini">Indochinellini</taxonomicName>
) but could be distinguished from these taxa by having a rectangular V- or W-shaped sculpture at the middle part of the shell. Furthermore,
<taxonomicName id="D6804D68FF8BFFEFFD9AF84906E06E4F" authority="(Gmelin, 1791)" baseAuthorityName="Gmelin" baseAuthorityYear="1791" box="[615,993,1888,2006]" class="Bivalvia" family="Unionidae" genus="Indonaia" kingdom="Animalia" order="Unionida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="rugosa">
<emphasis id="23F4EAF9FF8BFFEFFD9AF849060A6EE1" box="[615,779,1982,2006]" italics="true" pageId="10" pageNumber="11">Indonaia rugosa</emphasis>
(Gmelin, 1791)
</taxonomicName>
remotely resembles the new genus by having a few W-shaped ridges near the umbo but these ridges are less developed, less prominent, and do not spread over the dorsal half of the shell.
</paragraph>
</subSubSection>
<caption id="45FF6663FF87FFE3FF7CF8D807896EAC" pageId="6" pageNumber="7" startId="6.[129,194,1839,1863]" targetBox="[254,1349,144,1811]" targetPageId="6" targetType="figure">
<paragraph id="113F36EBFF87FFE3FF7CF8D807896EAC" blockId="6.[129,1466,1839,1947]" pageId="6" pageNumber="7">
<emphasis id="23F4EAF9FF87FFE3FF7CF8D805D96E70" bold="true" box="[129,216,1839,1863]" pageId="6" pageNumber="7">Figure 2.</emphasis>
Maximum likelihood (IQ-TREE v.1.6.12) phylogeny of the
<taxonomicName id="D6804D68FF87FFE3FCEDF8D806826E70" authorityName="Henderson" authorityYear="1935" box="[784,899,1839,1863]" class="Bivalvia" family="Unionidae" kingdom="Animalia" order="Unionida" pageId="6" pageNumber="7" phylum="Mollusca" rank="subFamily" subFamily="Parreysiinae">Parreysiinae</taxonomicName>
based on the combined alignment of the two mitochondrial and one nuclear gene fragments (three codons of
<emphasis id="23F4EAF9FF87FFE3FDACF8BB077B6E54" box="[593,634,1868,1891]" italics="true" pageId="6" pageNumber="7">COI</emphasis>
+ 16S rRNA + 28S rRNA). The labels of species recorded from Assam (Brahmaputra River basin) are red. The scale bar indicates the branch lengths. The information on DNA sequences is presented in the Supporting Information, Dataset S2. Outgroup taxa are not shown.
</paragraph>
</caption>
<caption id="45FF6663FF86FFE2FF8CF958069E6E64" pageId="7" pageNumber="8" startId="7.[113,178,1711,1735]" targetBox="[114,1458,144,1683]" targetPageId="7" targetType="figure">
<paragraph id="113F36EBFF86FFE2FF8CF958069E6E64" blockId="7.[113,1455,1711,1875]" pageId="7" pageNumber="8">
<emphasis id="23F4EAF9FF86FFE2FF8CF95805C86FF0" bold="true" box="[113,201,1711,1735]" pageId="7" pageNumber="8">Figure 3.</emphasis>
Shells of
<taxonomicName id="D6804D68FF86FFE2FEDFF95804A96FF0" box="[290,424,1711,1735]" class="Bivalvia" family="Unionidae" kingdom="Animalia" order="Unionida" pageId="7" pageNumber="8" phylum="Mollusca" rank="tribe" tribe="Indochinellini">Indochinellini</taxonomicName>
species from Assam (outer view of the left valve and inner view of the right valve). (A–H)
<taxonomicName id="D6804D68FF86FFE2FB04F95804026FD4" authority="(Hanley, 1856)" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Hanley" baseAuthorityYear="1856" class="Bivalvia" family="Unionidae" genus="Indonaia" kingdom="Animalia" order="Unionida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="corbis" status="stat. nov.">
<emphasis id="23F4EAF9FF86FFE2FB04F95800826FF0" box="[1273,1411,1711,1735]" italics="true" pageId="7" pageNumber="8">Indonaia corbis</emphasis>
(
<bibRefCitation id="75114B1AFF86FFE2FF85F93C05FA6FD4" author="Hanley S" box="[120,251,1739,1763]" pageId="7" pageNumber="8" refId="ref14239" refString="Hanley S. An Illustrated and Descriptive Catalogue of Recent Bivalve Shells. London: Williams and Norgate, 1856." type="book" year="1856">Hanley, 1856</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="38C75782FF86FFE2FEF5F93B04596FD4" box="[264,344,1740,1763]" pageId="7" pageNumber="8">stat. rev.</taxonomicNameLabel>
: topotypes IX-BDH-1 (A), IS-TLM-3 (B), IA-GEL-5 (C), X2-SES (D), X1-SES (E), I2-SES-2 (F), TZ-BDH-2 (G); holotype of
<taxonomicName id="D6804D68FF86FFE2FF19F91007E46FC8" authority="Annandale and Prashad, 1921" authorityName="Annandale and Prashad" authorityYear="1921" box="[228,741,1767,1791]" class="Bivalvia" family="Unionidae" genus="Trapezoideus" kingdom="Animalia" order="Unionida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="dhanushori">
<emphasis id="23F4EAF9FF86FFE2FF19F91004C66FC8" box="[228,455,1767,1791]" italics="true" pageId="7" pageNumber="8">Trapezoideus dhanushori</emphasis>
<bibRefCitation id="75114B1AFF86FFE2FE31F91007E46FC8" author="Annandale N &amp; Prashad B" box="[460,741,1767,1791]" pageId="7" pageNumber="8" pagination="529 - 631" refId="ref12662" refString="Annandale N, Prashad B. XXVIII. The aquatic and amphibious Mollusca of Manipur. Records of the Indian Museum 1921; 22: 529 - 631." type="journal article" year="1921">Annandale and Prashad, 1921</bibRefCitation>
</taxonomicName>
with its labels (ZSI M11962/2),
<taxonomicName id="D6804D68FF86FFE2FBE2F91001D86FC8" authority="Stream" authorityName="Stream" box="[1055,1241,1767,1791]" class="Bivalvia" family="Unionidae" genus="Trapezoideus" kingdom="Animalia" order="Unionida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="dhanushori">Dhanushori Stream</taxonomicName>
at a distance of about a mile from Dimapur, Assam (H). (I–K)
<taxonomicName id="D6804D68FF86FFE2FE0EF8F4064C6E2C" authority="(Hanley, 1856)" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Hanley" baseAuthorityYear="1856" box="[499,845,1795,1819]" class="Bivalvia" family="Unionidae" genus="Pseudoscabies" higherTaxonomySource="CoL" kingdom="Animalia" order="Unionida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="scobina" status="gen. et comb. nov.">
<emphasis id="23F4EAF9FF86FFE2FE0EF8F407B66E2C" box="[499,695,1795,1819]" italics="true" pageId="7" pageNumber="8">Pseudoscabies scobina</emphasis>
(
<bibRefCitation id="75114B1AFF86FFE2FD3EF8F406476E2C" author="Hanley S" box="[707,838,1795,1819]" pageId="7" pageNumber="8" refId="ref14239" refString="Hanley S. An Illustrated and Descriptive Catalogue of Recent Bivalve Shells. London: Williams and Norgate, 1856." type="book" year="1856">Hanley, 1856</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="38C75782FF86FFE2FCAFF8F301016E2C" box="[850,1024,1795,1820]" pageId="7" pageNumber="8" rank="species">gen. et comb. nov.</taxonomicNameLabel>
: topotypes SC-SES-1 (I), IO-BDH-5 (J), and IO-BDH-4 (K). Scale bars = 10 mm. The shell vouchers of the new samples are deposited in FBRC ZSI (Supporting Information, Dataset S1). Photos: J. Sonowal and D. Kardong (A–G, I–K); A. Mukhopadhyaya and S. Sajan (H).
</paragraph>
</caption>
<caption id="45FF6663FF89FFEDFF7CF8DB001C6EAF" pageId="8" pageNumber="9" startId="8.[129,194,1836,1860]" targetBox="[174,1429,144,1808]" targetPageId="8" targetType="figure">
<paragraph id="113F36EBFF89FFEDFF7CF8DB001C6EAF" blockId="8.[129,1468,1836,1944]" pageId="8" pageNumber="9">
<emphasis id="23F4EAF9FF89FFEDFF7CF8DB05D96E73" bold="true" box="[129,216,1836,1860]" pageId="8" pageNumber="9">Figure 4.</emphasis>
Shells of
<taxonomicName id="D6804D68FF89FFEDFECCF8DB04BC6E73" authorityName="Modell" authorityYear="1942" box="[305,445,1836,1860]" class="Bivalvia" family="Unionidae" kingdom="Animalia" order="Unionida" pageId="8" pageNumber="9" phylum="Mollusca" rank="tribe" tribe="Lamellidentini">Lamellidentini</taxonomicName>
species from Assam (sequenced specimens; outer view of the left valve and inner view of the right valve). A,
<taxonomicName id="D6804D68FF89FFEDFF7CF8BF04E96E57" authority="(Benson, 1836)" baseAuthorityName="Benson" baseAuthorityYear="1836" box="[129,488,1864,1888]" class="Bivalvia" family="Unionidae" genus="Balwantia" kingdom="Animalia" order="Unionida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="soleniformis">
<emphasis id="23F4EAF9FF89FFEDFF7CF8BF044D6E57" box="[129,332,1864,1888]" italics="true" pageId="8" pageNumber="9">Balwantia soleniformis</emphasis>
(
<bibRefCitation id="75114B1AFF89FFEDFEA5F8BF04E16E57" author="Benson WH" box="[344,480,1864,1888]" pageId="8" pageNumber="9" pagination="741 - 50" refId="ref12750" refString="Benson WH. Descriptive catalogue of a collection of land and fresh-water shells, chiefly contained in the museum of the Asiatic Society. Journal of the Asiatic Society of Bengal 1836; 5: 741 - 50." type="journal article" year="1836">Benson, 1836</bibRefCitation>
)
</taxonomicName>
, sample DLS-DU BS-BDH-1. B–G,
<taxonomicName id="D6804D68FF89FFEDFCC3F8BF01BC6E57" authority="(Benson, 1862)" baseAuthorityName="Benson" baseAuthorityYear="1862" box="[830,1213,1864,1888]" class="Bivalvia" family="Unionidae" genus="Lamellidens" kingdom="Animalia" order="Unionida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="jenkinsianus">
<emphasis id="23F4EAF9FF89FFEDFCC3F8BF011F6E57" box="[830,1054,1864,1888]" italics="true" pageId="8" pageNumber="9">Lamellidens jenkinsianus</emphasis>
(
<bibRefCitation id="75114B1AFF89FFEDFBD7F8BF01B46E57" author="Benson WH" box="[1066,1205,1864,1888]" pageId="8" pageNumber="9" pagination="184 - 95" refId="ref12789" refString="Benson WH. Descriptions of Indian and Burmese species of the genus Unio, Retz. Annals and Magazine of Natural History 1862; 10: 184 - 95. https: // doi. org / 10.1080 / 00222936208681307" type="journal article" year="1862">Benson, 1862</bibRefCitation>
)
</taxonomicName>
: topotypes LJ-NKM-2 (B), LJ-NKM-3 (C), L5-NKM-1 (D), L4-NKM-1 (E), L5-NKM-4 (F), and L4-NKM-2 (
<emphasis id="23F4EAF9FF89FFEDFC64F89306AA6E4B" bold="true" box="[921,939,1892,1916]" pageId="8" pageNumber="9">G</emphasis>
). Scale bars = 10 mm. The shell vouchers of the new samples are deposited in FBRC ZSI and DLS-DU (Supporting Information, Dataset S1). Photos: J. Sonowal and D. Kardong.
</paragraph>
</caption>
<caption id="45FF6663FF88FFECFF8CF8DC062C6E83" pageId="9" pageNumber="10" startId="9.[113,178,1835,1859]" targetBox="[159,1413,144,1808]" targetPageId="9" targetType="figure">
<paragraph id="113F36EBFF88FFECFF8CF8DC00876E74" blockId="9.[113,1450,1835,1972]" box="[113,1414,1835,1860]" pageId="9" pageNumber="10">
<emphasis id="23F4EAF9FF88FFECFF8CF8DC05C86E73" bold="true" box="[113,201,1835,1860]" pageId="9" pageNumber="10">Figure 5.</emphasis>
Shells of
<taxonomicName id="D6804D68FF88FFECFEDFF8DC079D6E73" authority="Preston, 1912" authorityName="Preston" authorityYear="1912" box="[290,668,1835,1860]" class="Bivalvia" family="Unionidae" genus="Lamellidens" kingdom="Animalia" order="Unionida" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="nongyangensis">
<emphasis id="23F4EAF9FF88FFECFEDFF8DC07146E74" box="[290,533,1835,1859]" italics="true" pageId="9" pageNumber="10">Lamellidens nongyangensis</emphasis>
<bibRefCitation id="75114B1AFF88FFECFDE4F8DB079D6E73" author="Preston HB" box="[537,668,1836,1860]" pageId="9" pageNumber="10" pagination="279 - 308" refId="ref15608" refString="Preston HB. A catalogue of the Asiatic naiades in the collection of the Indian Museum, Calcutta, with descriptions of new species. Records of the Indian Museum 1912; 7: 279 - 308." type="journal article" year="1912">Preston, 1912</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="D6804D68FF88FFECFD5AF8DC063B6E74" authorityName="Modell" authorityYear="1942" box="[679,826,1835,1859]" class="Bivalvia" family="Unionidae" kingdom="Animalia" order="Unionida" pageId="9" pageNumber="10" phylum="Mollusca" rank="tribe" tribe="Lamellidentini">Lamellidentini</taxonomicName>
) (outer view of the left valve and inner view of the right valve).
</paragraph>
<paragraph id="113F36EBFF88FFECFF8CF8BF062C6E83" blockId="9.[113,1450,1835,1972]" pageId="9" pageNumber="10">A, holotype ZSI 5068/1 with its label, Lake of No Return (=Nongyang Lake), 27.2192°N, 96.1439°E, Irrawaddy Basin, Myanmar. B–G, sequenced specimens from Assam: samples L1-TLM-3 (B), L2-TLM-2 (C), LL-TNG-4 (D), LN-TNG-3 (E), LU-DHN-2 (F), and LJ-TLM-1 (G). Scale bars = 10 mm. The shell vouchers of the new samples are deposited in FBRC ZSI (Supporting Information, Dataset S1). Photos: N.V. Subba Rao and R. Pasupuleti (A); J. Sonowal and D. Kardong (B–G).</paragraph>
</caption>
<caption id="45FF6663FF8BFFEFFF7CF94A074C6E71" pageId="10" pageNumber="11" startId="10.[129,194,1725,1749]" targetBox="[129,1473,144,1698]" targetPageId="10" targetType="figure">
<paragraph id="113F36EBFF8BFFEFFF7CF94A074C6E71" blockId="10.[129,1472,1725,1862]" pageId="10" pageNumber="11">
<emphasis id="23F4EAF9FF8BFFEFFF7CF94A05D96FE2" bold="true" box="[129,216,1725,1749]" pageId="10" pageNumber="11">Figure 6.</emphasis>
Shells of
<taxonomicName id="D6804D68FF8BFFEFFECCF94904946FE1" authorityName="Henderson" authorityYear="1935" box="[305,405,1726,1750]" class="Bivalvia" family="Unionidae" kingdom="Animalia" order="Unionida" pageId="10" pageNumber="11" phylum="Mollusca" rank="tribe" tribe="Parreysiini">Parreysiini</taxonomicName>
species from Assam (outer view of the left valve and inner view of the right valve). A–C,
<taxonomicName id="D6804D68FF8BFFEFFB2BF94904136FC6" authority="(Hanley, 1856)" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Hanley" baseAuthorityYear="1856" class="Bivalvia" family="Unionidae" genus="Assamnaia" higherTaxonomySource="CoL" kingdom="Animalia" order="Unionida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="involuta" status="gen. et comb. nov.">
<emphasis id="23F4EAF9FF8BFFEFFB2BF94900896FE1" box="[1238,1416,1726,1750]" italics="true" pageId="10" pageNumber="11">Assamnaia involuta</emphasis>
(
<bibRefCitation id="75114B1AFF8BFFEFFF7AF92D040B6FC6" author="Hanley S" box="[135,266,1753,1778]" pageId="10" pageNumber="11" refId="ref14239" refString="Hanley S. An Illustrated and Descriptive Catalogue of Recent Bivalve Shells. London: Williams and Norgate, 1856." type="book" year="1856">Hanley, 1856</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="38C75782FF8BFFEFFEEAF92D04C56FC6" box="[279,452,1754,1778]" pageId="10" pageNumber="11" rank="species">gen. et comb. nov.</taxonomicNameLabel>
: topotypes II-DHN-1 (A), II-DHN-2 (B), and DLS-DU II-D (dead shell; non-sequenced) (C). D–I,
<taxonomicName id="D6804D68FF8BFFEFFF7CF90104BC6E39" authority="(Benson, 1862)" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Benson" baseAuthorityYear="1862" box="[129,445,1782,1806]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="marcens" status="stat. nov.">
<emphasis id="23F4EAF9FF8BFFEFFF7CF90104236E3A" box="[129,290,1782,1805]" italics="true" pageId="10" pageNumber="11">Parreysia marcens</emphasis>
(
<bibRefCitation id="75114B1AFF8BFFEFFED3F90104B76E39" author="Benson WH" box="[302,438,1782,1806]" pageId="10" pageNumber="11" pagination="184 - 95" refId="ref12789" refString="Benson WH. Descriptions of Indian and Burmese species of the genus Unio, Retz. Annals and Magazine of Natural History 1862; 10: 184 - 95. https: // doi. org / 10.1080 / 00222936208681307" type="journal article" year="1862">Benson, 1862</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="38C75782FF8BFFEFFE3FF90107126E3A" box="[450,531,1782,1805]" pageId="10" pageNumber="11">stat. rev.</taxonomicNameLabel>
: topotypes PS-SES-5 (D), PS-SES-1 (E), P1-SES-4 (F), PC-SES-1 (G), PSm-SES-2 (H), and PF-BDH-4 (I). Scale bars = 10 mm. The shell vouchers of the new samples are deposited in FBRC ZSI and DLS-DU (Supporting Information, Dataset S1). Photos: J. Sonowal and D. Kardong.
</paragraph>
</caption>
<caption id="45FF6663FF8AFFEEFF8DFAE107BC6C8D" pageId="11" pageNumber="12" startId="11.[112,177,1302,1326]" targetBox="[114,1458,161,1274]" targetPageId="11" targetType="figure">
<paragraph id="113F36EBFF8AFFEEFF8DFAE107BC6C8D" blockId="11.[112,1448,1302,1466]" pageId="11" pageNumber="12">
<emphasis id="23F4EAF9FF8AFFEEFF8DFAE105C96C18" bold="true" box="[112,200,1302,1327]" pageId="11" pageNumber="12">Figure 7.</emphasis>
Habitats of little-known freshwater mussel species in Assam. A, collecting locality of
<taxonomicName id="D6804D68FF8AFFEEFC1BFAE001996C19" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Hanley" baseAuthorityYear="1856" box="[998,1176,1302,1326]" class="Bivalvia" family="Unionidae" genus="Assamnaia" higherTaxonomySource="CoL" kingdom="Animalia" order="Unionida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="involuta" status="gen. et comb. nov.">
<emphasis id="23F4EAF9FF8AFFEEFC1BFAE001996C19" box="[998,1176,1302,1326]" italics="true" pageId="11" pageNumber="12">Assamnaia involuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38C75782FF8AFFEEFB60FAE0004B6C19" box="[1181,1354,1302,1327]" pageId="11" pageNumber="12" rank="species">gen. et comb. nov.</taxonomicNameLabel>
: a shallow oxbow lake with silt and clay bottom substrate, Dhansiri River valley, Golaghat, 26 March 2022. B, collecting locality of
<taxonomicName id="D6804D68FF8AFFEEFB2CFAC5009C6C7D" baseAuthorityName="Benson" baseAuthorityYear="1836" box="[1233,1437,1330,1354]" class="Bivalvia" family="Unionidae" genus="Balwantia" kingdom="Animalia" order="Unionida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="soleniformis">
<emphasis id="23F4EAF9FF8AFFEEFB2CFAC5009C6C7D" box="[1233,1437,1330,1354]" italics="true" pageId="11" pageNumber="12">Balwantia soleniformis</emphasis>
</taxonomicName>
,
<taxonomicName id="D6804D68FF8AFFEEFF8CFAB905FD6C51" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Hanley" baseAuthorityYear="1856" box="[113,252,1358,1382]" class="Bivalvia" family="Unionidae" genus="Indonaia" kingdom="Animalia" order="Unionida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="corbis">
<emphasis id="23F4EAF9FF8AFFEEFF8CFAB905FD6C51" box="[113,252,1358,1382]" italics="true" pageId="11" pageNumber="12">Indonaia corbis</emphasis>
</taxonomicName>
,
<taxonomicName id="D6804D68FF8AFFEEFEFBFAB904CB6C51" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Hanley" baseAuthorityYear="1856" box="[262,458,1358,1382]" class="Bivalvia" family="Unionidae" genus="Pseudoscabies" higherTaxonomySource="CoL" kingdom="Animalia" order="Unionida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="scobina" status="gen. et comb. nov.">
<emphasis id="23F4EAF9FF8AFFEEFEFBFAB904CB6C51" box="[262,458,1358,1382]" italics="true" pageId="11" pageNumber="12">Pseudoscabies scobina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38C75782FF8AFFEEFE32FAB8077B6C51" box="[463,634,1358,1383]" pageId="11" pageNumber="12" rank="species">gen. et comb. nov.</taxonomicNameLabel>
, and
<taxonomicName id="D6804D68FF8AFFEEFD57FAB8064A6C51" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Benson" baseAuthorityYear="1862" box="[682,843,1359,1382]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="marcens">
<emphasis id="23F4EAF9FF8AFFEEFD57FAB8064A6C51" box="[682,843,1359,1382]" italics="true" pageId="11" pageNumber="12">Parreysia marcens</emphasis>
</taxonomicName>
: Burhidihing River, Dibrugarh, 22 February 2022. The white arrow shows the river cliff, whose bottom portion houses numerous
<taxonomicName id="D6804D68FF8AFFEEFD0EFA9C06756CB5" baseAuthorityName="Benson" baseAuthorityYear="1836" box="[755,884,1386,1410]" class="Bivalvia" family="Unionidae" genus="Balwantia" kingdom="Animalia" order="Unionida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="soleniformis">
<emphasis id="23F4EAF9FF8AFFEEFD0EFA9C06756CB5" box="[755,884,1386,1410]" italics="true" pageId="11" pageNumber="12">B. soleniformis</emphasis>
</taxonomicName>
burrows; the river cliff is mostly composed by firm ground (hard clay). C, close-up view of an aggregation of
<taxonomicName id="D6804D68FF8AFFEEFDBEFA7007C56CA9" baseAuthorityName="Benson" baseAuthorityYear="1836" box="[579,708,1414,1438]" class="Bivalvia" family="Unionidae" genus="Balwantia" kingdom="Animalia" order="Unionida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="soleniformis">
<emphasis id="23F4EAF9FF8AFFEEFDBEFA7007C56CA9" box="[579,708,1414,1438]" italics="true" pageId="11" pageNumber="12">B. soleniformis</emphasis>
</taxonomicName>
burrows in the river cliff, 22 February 2022. Living mussels were extracted from submerged burrows. Photos: J. Sonowal and D. Kardong.
</paragraph>
</caption>
<subSubSection id="599A6560FF8AFFEEFF8CFA1C06BF6F57" pageId="11" pageNumber="12" type="description">
<paragraph id="113F36EBFF8AFFEEFF8CFA1C06BF6F57" blockId="11.[112,764,1514,1977]" lastBlockId="11.[810,1459,1514,1632]" pageId="11" pageNumber="12">
<emphasis id="23F4EAF9FF8AFFEEFF8CFA1C05E96F35" box="[113,232,1515,1538]" italics="true" pageId="11" pageNumber="12">Description:</emphasis>
Shell thin, rather compressed; anterior margin broadly rounded; posterior margin slightly narrower, with small wing; dorsal margin slightly convex, its posterior part angular, umbo not prominent; ventral margin slightly convex. Umbo slightly shifted to the anterior part of the shell. Periostracum brown, sometimes with green markings on ridges at the posterior half of the shell. Nacre bluish white, sometimes with goldish spots. The shell is covered by peculiar sculpture: repeating V- or W-shaped ridges at the middle of the shell, surrounded by waved ridges at the anterior part of the shell and arcuate ridges at the posterior part of the shell. This sculpture is clearly visible in the inner side of the shell by depressions, the pattern of which corresponds to ridges on the outer side. Pseudocardinal teeth rather short, representing shallow, smooth ridges, two teeth in right valve and one tooth in left valve. Lateral teeth elongated, slightly curved, lamella-like, one tooth in right valve and two teeth in left valve. Anterior adductor muscle scar ovate, shallow; posterior adductor muscle scar ovate or round, very shallow, sometimes weakly visible.
</paragraph>
</subSubSection>
<subSubSection id="599A6560FF8AFFEEFCD7F98001086E3B" pageId="11" pageNumber="12" type="etymology">
<paragraph id="113F36EBFF8AFFEEFCD7F98001086E3B" blockId="11.[810,1460,1655,1804]" pageId="11" pageNumber="12">
<emphasis id="23F4EAF9FF8AFFEEFCD7F98006986FB8" box="[810,921,1655,1679]" italics="true" pageId="11" pageNumber="12">Etymology:</emphasis>
The name of this genus comprises of two words: ‘pseudo’ (translingual prefix, meaning ‘appears like a true but is false’) and ‘scabies’ [reference to the genus
<taxonomicName id="D6804D68FF8AFFEEFAF8F941004A6FF9" authorityName="Haas" authorityYear="1911" box="[1285,1355,1718,1742]" class="Bivalvia" family="Unionidae" genus="Scabies" kingdom="Animalia" order="Unionida" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8AFFEEFAF8F941004A6FF9" box="[1285,1355,1718,1742]" italics="true" pageId="11" pageNumber="12">Scabies</emphasis>
</taxonomicName>
, in which it was placed due to a similar shell sculpture (
<bibRefCitation id="75114B1AFF8AFFEEFAFFF92200AC6FDA" author="Subba Rao NV" box="[1282,1453,1749,1773]" pageId="11" pageNumber="12" refId="ref15978" refString="Subba Rao NV. Handbook. Freshwater Molluscs of India. Calcutta: Zoological Survey of India, 1989." type="book" year="1989">Subba Rao 1989</bibRefCitation>
,
<bibRefCitation id="75114B1AFF8AFFEEFCD7F90306EF6E3B" author="Sonowal J &amp; Puzari M &amp; Kardong D" box="[810,1006,1780,1804]" pageId="11" pageNumber="12" pagination="18237 - 46" refId="ref15885" refString="Sonowal J, Puzari M, Kardong D. Diversity of freshwater molluscs from the upper Brahmaputra Basin, Assam, India. Journal of Threatened Taxa 2021; 13: 18237 - 46. https: // doi. org / 10.11609 / jott. 7144.13.5.18237 - 18246" type="journal article" year="2021">
Sonowal
<emphasis id="23F4EAF9FF8AFFEEFC7AF90206B76E3B" box="[903,950,1780,1804]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2021
</bibRefCitation>
)].
</paragraph>
</subSubSection>
<subSubSection id="599A6560FF8AFFEEFCD7F8DB06F76E54" pageId="11" pageNumber="12" type="distribution">
<paragraph id="113F36EBFF8AFFEEFCD7F8DB06F76E54" blockId="11.[810,1459,1836,1891]" pageId="11" pageNumber="12">
<emphasis id="23F4EAF9FF8AFFEEFCD7F8DB06A96E73" box="[810,936,1836,1860]" italics="true" pageId="11" pageNumber="12">Distribution:</emphasis>
Endemic to the Brahmaputra River basin, north-eastern
<collectingCountry id="6997767BFF8AFFEEFC46F8BC06F26E54" box="[955,1011,1867,1891]" name="India" pageId="11" pageNumber="12">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="599A6560FF8AFFE9FCD7F87707E969F0" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="biology_ecology">
<paragraph id="113F36EBFF8AFFE9FCD7F87707E969F0" blockId="11.[810,1460,1920,1975]" lastBlockId="12.[129,777,144,199]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="23F4EAF9FF8AFFEEFCD7F87706866EAF" box="[810,903,1920,1944]" italics="true" pageId="11" pageNumber="12">Habitats:</emphasis>
<typeStatus id="CE3B8849FF8AFFEEFC70F87706FA6EAF" box="[909,1019,1920,1944]" pageId="11" pageNumber="12" type="topotype">Topotypes</typeStatus>
of
<taxonomicName id="D6804D68FF8AFFEEFBE3F87701F26EAF" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Hanley" baseAuthorityYear="1856" box="[1054,1267,1920,1944]" class="Bivalvia" family="Unionidae" genus="Pseudoscabies" higherTaxonomySource="CoL" kingdom="Animalia" order="Unionida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="scobina" status="gen. et comb. nov.">
<emphasis id="23F4EAF9FF8AFFEEFBE3F87701F26EAF" box="[1054,1267,1920,1944]" italics="true" pageId="11" pageNumber="12">Pseudoscabies scobina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38C75782FF8AFFEEFB04F87700B56EA0" box="[1273,1460,1920,1944]" pageId="11" pageNumber="12" rank="species">gen. et comb. nov.</taxonomicNameLabel>
(
<emphasis id="23F4EAF9FF8AFFEEFCC8F85706496E80" box="[821,840,1952,1975]" italics="true" pageId="11" pageNumber="12">N</emphasis>
=
<specimenCount id="0786FD62FF8AFFEEFC98F86806E56E80" box="[869,996,1951,1975]" count="8" pageId="11" pageNumber="12" type="generic">8 specimens</specimenCount>
) were collected from the Sesa and Burhidihing rivers in
<collectingRegion id="D344F809FF8DFFE9FF26FF67041E699F" box="[219,287,144,168]" country="India" name="Assam" pageId="12" pageNumber="13">Assam</collectingRegion>
(
<figureCitation id="89BB2A6EFF8DFFE9FECDFF6704786990" box="[304,377,144,168]" captionStart="Figure 7" captionStartId="11.[112,177,1302,1326]" captionTargetBox="[114,1458,161,1274]" captionTargetId="figure-269@11.[114,1458,144,1274]" captionTargetPageId="11" captionText="Figure 7. Habitats of little-known freshwater mussel species in Assam.A, collecting locality of Assamnaia involuta gen. et comb.nov.: a shallow oxbow lake with silt and clay bottom substrate, Dhansiri River valley, Golaghat, 26 March 2022. B, collecting locality of Balwantia soleniformis, Indonaia corbis, Pseudoscabies scobina gen. et comb.nov., and Parreysia marcens: Burhidihing River, Dibrugarh, 22 February 2022. The white arrow shows the river cliff, whose bottom portion houses numerous B. soleniformis burrows; the river cliff is mostly composed by firm ground (hard clay). C, close-up view of an aggregation of B. soleniformis burrows in the river cliff,22 February 2022. Living mussels were extracted from submerged burrows.Photos: J. Sonowal and D. Kardong." pageId="12" pageNumber="13">Fig. 7B</figureCitation>
). The species was found together with
<taxonomicName id="D6804D68FF8DFFE9FF7FFF47042E69F0" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Benson" baseAuthorityYear="1862" box="[130,303,176,199]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="marcens">
<emphasis id="23F4EAF9FF8DFFE9FF7FFF47042E69F0" box="[130,303,176,199]" italics="true" pageId="12" pageNumber="13">Parreysia marcens</emphasis>
</taxonomicName>
,
<taxonomicName id="D6804D68FF8DFFE9FEC7FF5804CE69F0" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Hanley" baseAuthorityYear="1856" box="[314,463,175,199]" class="Bivalvia" family="Unionidae" genus="Indonaia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="corbis">
<emphasis id="23F4EAF9FF8DFFE9FEC7FF5804CE69F0" box="[314,463,175,199]" italics="true" pageId="12" pageNumber="13">Indonaia corbis</emphasis>
</taxonomicName>
, and
<taxonomicName id="D6804D68FF8DFFE9FDFBFF5807E369F0" baseAuthorityName="Benson" baseAuthorityYear="1836" box="[518,738,175,199]" class="Bivalvia" family="Unionidae" genus="Balwantia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="soleniformis">
<emphasis id="23F4EAF9FF8DFFE9FDFBFF5807E369F0" box="[518,738,175,199]" italics="true" pageId="12" pageNumber="13">Balwantia soleniformis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="599A6560FF8DFFE9FF7CFF1E042B68AA" pageId="12" pageNumber="13" type="discussion">
<paragraph id="113F36EBFF8DFFE9FF7CFF1E042B68AA" blockId="12.[129,783,233,413]" pageId="12" pageNumber="13">
<emphasis id="23F4EAF9FF8DFFE9FF7CFF1E05F06837" box="[129,241,233,256]" italics="true" pageId="12" pageNumber="13">Comments:</emphasis>
Here, we transfer the nominal species
<taxonomicName id="D6804D68FF8DFFE9FD74FF1E05CA6817" authority="Hanley" authorityName="Hanley" authorityYear="1856" class="Bivalvia" family="Unionidae" genus="Unio" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="scobina">
<emphasis id="23F4EAF9FF8DFFE9FD74FF1E06086836" box="[649,777,233,257]" italics="true" pageId="12" pageNumber="13">Unio scobina</emphasis>
Hanley
</taxonomicName>
from
<taxonomicName id="D6804D68FF8DFFE9FEF6FEFF04626817" authorityName="Prashad" authorityYear="1918" box="[267,355,264,288]" class="Bivalvia" family="Unionidae" genus="Indonaia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FEF6FEFF04626817" box="[267,355,264,288]" italics="true" pageId="12" pageNumber="13">Indonaia</emphasis>
</taxonomicName>
to
<taxonomicName id="D6804D68FF8DFFE9FE78FEFF070A6817" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" box="[389,523,264,288]" class="Bivalvia" family="Unionidae" genus="Pseudoscabies" higherTaxonomySource="CoL" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="23F4EAF9FF8DFFE9FE78FEFF070A6817" box="[389,523,264,288]" italics="true" pageId="12" pageNumber="13">Pseudoscabies</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38C75782FF8DFFE9FDECFEFF076C6828" box="[529,621,264,288]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
and propose
<taxonomicName id="D6804D68FF8DFFE9FD07FEFE05C86808" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Unio" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="scobina" status="gen. et comb. nov.">
<emphasis id="23F4EAF9FF8DFFE9FD07FEFE05C86808" italics="true" pageId="12" pageNumber="13">P. scobina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38C75782FF8DFFE9FF32FEDF048E6808" box="[207,399,295,320]" pageId="12" pageNumber="13" rank="species">gen. et comb. nov.</taxonomicNameLabel>
; its conchological form with broadly rounded V-shaped sculpture (e.g. see
<figureCitation id="89BB2A6EFF8DFFE9FDEEFEB007636868" box="[531,610,327,351]" captionStart="Figure 3" captionStartId="7.[113,178,1711,1735]" captionTargetBox="[114,1458,144,1683]" captionTargetId="figure-69@7.[114,1458,144,1683]" captionTargetPageId="7" captionText="Figure 3. Shells of Indochinellini species from Assam (outer view of the left valve and inner view of the right valve).(A–H) Indonaia corbis (Hanley, 1856) stat. rev.: topotypes IX-BDH-1 (A), IS-TLM-3 (B), IA-GEL-5 (C), X2-SES (D), X1-SES (E), I2-SES-2 (F), TZ-BDH-2 (G); holotype of Trapezoideus dhanushori Annandale and Prashad, 1921 with its labels (ZSI M11962/2), Dhanushori Stream at a distance of about a mile from Dimapur, Assam (H). (I–K) Pseudoscabies scobina (Hanley, 1856) gen.et comb. nov.: topotypes SC-SES-1 (I), IO-BDH-5 (J), and IO-BDH-4 (K). Scale bars = 10 mm. The shell vouchers of the new samples are deposited in FBRC ZSI (Supporting Information, Dataset S1). Photos: J. Sonowal and D. Kardong (A–G, I–K); A.Mukhopadhyaya and S. Sajan (H)." pageId="12" pageNumber="13">Fig. 3K</figureCitation>
) was described as
<taxonomicName id="D6804D68FF8DFFE9FF61FE9004A46849" authority="Hanley, 1856" authorityName="Hanley" authorityYear="1856" box="[156,421,358,382]" class="Bivalvia" family="Unionidae" genus="Unio" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="radula">
<emphasis id="23F4EAF9FF8DFFE9FF61FE9004156849" box="[156,276,358,382]" italics="true" pageId="12" pageNumber="13">Unio radula</emphasis>
<bibRefCitation id="75114B1AFF8DFFE9FEE1FE9104A46849" author="Hanley S" box="[284,421,358,382]" pageId="12" pageNumber="13" refId="ref14239" refString="Hanley S. An Illustrated and Descriptive Catalogue of Recent Bivalve Shells. London: Williams and Norgate, 1856." type="book" year="1856">Hanley, 1856</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="D6804D68FF8DFFE9FE3BFE91072A6849" authorityName="Conrad" authorityYear="1853" box="[454,555,358,382]" class="Bivalvia" family="Unionidae" genus="Nodularia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FE3BFE91072A6849" box="[454,555,358,382]" italics="true" pageId="12" pageNumber="13">Nodularia</emphasis>
</taxonomicName>
(sec.
<emphasis id="23F4EAF9FF8DFFE9FD96FE9107CA6849" box="[619,715,358,382]" italics="true" pageId="12" pageNumber="13">Radiatula</emphasis>
)
<taxonomicName id="D6804D68FF8DFFE9FD23FE91041E68AA" authority="Simpson, 1900" authorityName="Simpson" authorityYear="1900" class="Bivalvia" family="Unionidae" genus="Nodularia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="lima">
<emphasis id="23F4EAF9FF8DFFE9FD23FE91060B6849" box="[734,778,358,382]" italics="true" pageId="12" pageNumber="13">lima</emphasis>
<bibRefCitation id="75114B1AFF8DFFE9FF7CFE72041E68AA" author="Simpson CT" box="[129,287,389,413]" pageId="12" pageNumber="13" pagination="501 - 1044" refId="ref15743" refString="Simpson CT. Synopsis of the naiades, or pearly fresh-water mussels. Proceedings of the United States National Museum 1900; 22: 501 - 1044. https: // doi. org / 10.5479 / si. 00963801.22 - 1205.501" type="journal article" year="1900">Simpson, 1900</bibRefCitation>
</taxonomicName>
].
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="73183FB01C86DC3BD54EB990EF770F06">Discovery of an endemism hotspot of freshwater mussels (Bivalvia: Unionidae) in Assam, with a description of two new genera</mods:title>
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<mods:namePart id="117E08448326D42BC89FEDF76E926E44">Bolotov, Ivan N.</mods:namePart>
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Genus
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<emphasis id="23F4EAF9FF8DFFE9FC20F97A013A6F90" bold="true" box="[989,1083,1677,1703]" italics="true" pageId="12" pageNumber="13">Parreysia</emphasis>
<bibRefCitation id="75114B1AFF8DFFE9FBC2F97A01D36F90" author="Conrad TA" box="[1087,1234,1677,1703]" pageId="12" pageNumber="13" pagination="243 - 69" refId="ref13680" refString="Conrad TA. A synopsis of the family of Naiades of North America, with notes, and a table of some of the genera and sub-genera of the family, according to their geographical distribution, and descriptions of genera and sub-genera. Proceedings of the Academy of Natural Sciences of Philadelphia 1853; 6: 243 - 69." type="journal article" year="1853">Conrad, 1853</bibRefCitation>
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(
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species:
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<emphasis id="23F4EAF9FF8DFFE9FCA2F95B01216FF0" bold="true" box="[863,1056,1708,1735]" italics="true" pageId="12" pageNumber="13">Unio multidentatus</emphasis>
Philippi, 1847
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[=
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<emphasis id="23F4EAF9FF8DFFE9FB25F95B009D6FF0" bold="true" box="[1240,1436,1708,1735]" italics="true" pageId="12" pageNumber="13">Parreysia corrugata</emphasis>
(O.F.
<bibRefCitation id="75114B1AFF8DFFE9FC91F93C06FE6FD2" author="Muller OF" box="[876,1023,1739,1765]" pageId="12" pageNumber="13" pagination="1 - 214" refId="ref14939" refString="Muller OF. Testacea. Vermivm Terrestrium et Fluviatilium, Seu Animalium Infusoriorum, Helminthicorum et Testaceorum, non Marinorum, Succincta Historia 1774; 2: 1 - 214." type="journal article" year="1774">Müller, 1774</bibRefCitation>
)
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]; by original designation) (
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).
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<emphasis id="23F4EAF9FF8DFFE9FBBDF90B01BA6E21" bold="true" box="[1088,1211,1788,1815]" pageId="12" pageNumber="13">
(
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)
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<emphasis id="23F4EAF9FF8DFFE9FCC4F8C906A66E62" box="[825,935,1854,1877]" italics="true" pageId="12" pageNumber="13">Comments:</emphasis>
This genus contains eight recent species. Four nominal species belonging to the genus
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were described from
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:
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<emphasis id="23F4EAF9FF8DFFE9FC7AF88A01096EA3" box="[903,1032,1916,1940]" italics="true" pageId="12" pageNumber="13">Unio favidens</emphasis>
var.
<emphasis id="23F4EAF9FF8DFFE9FBD2F88A017C6EA3" box="[1071,1149,1917,1940]" italics="true" pageId="12" pageNumber="13">marcens</emphasis>
<bibRefCitation id="75114B1AFF8DFFE9FB7EF88B000E6EA3" author="Benson WH" box="[1155,1295,1916,1940]" pageId="12" pageNumber="13" pagination="184 - 95" refId="ref12789" refString="Benson WH. Descriptions of Indian and Burmese species of the genus Unio, Retz. Annals and Magazine of Natural History 1862; 10: 184 - 95. https: // doi. org / 10.1080 / 00222936208681307" type="journal article" year="1862">Benson, 1862</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="D6804D68FF8DFFE9FAE9F88A06DD6E84" authority="Theobald, 1873" authorityName="Theobald" authorityYear="1873" class="Bivalvia" family="Unionidae" genus="Unio" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="gowhattensis">
<emphasis id="23F4EAF9FF8DFFE9FAE9F88A00C06EA3" box="[1300,1473,1916,1940]" italics="true" pageId="12" pageNumber="13">Unio gowhattensis</emphasis>
<bibRefCitation id="75114B1AFF8DFFE9FCC4F86C06DD6E84" author="Theobald W" box="[825,988,1947,1971]" pageId="12" pageNumber="13" pagination="207 - 9" refId="ref16043" refString="Theobald W. Descriptions of new species of Unionidae. Journal of the Asiatic Society of Bengal 1873; 42: 207 - 9." type="journal article" year="1873">Theobald, 1873</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="D6804D68FF8DFFE9FC15F86B00BF6E84" authority="Preston, 1912" authorityName="Preston" authorityYear="1912" box="[1000,1470,1947,1971]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="annandalei" subGenus="Parreysia">
<emphasis id="23F4EAF9FF8DFFE9FC15F86B01406E84" box="[1000,1089,1948,1971]" italics="true" pageId="12" pageNumber="13">Parreysia</emphasis>
(
<emphasis id="23F4EAF9FF8DFFE9FBACF86B01AB6E84" box="[1105,1194,1948,1971]" italics="true" pageId="12" pageNumber="13">Parreysia</emphasis>
)
<emphasis id="23F4EAF9FF8DFFE9FB41F86C00266E84" box="[1212,1319,1947,1971]" italics="true" pageId="12" pageNumber="13">annandalei</emphasis>
<bibRefCitation id="75114B1AFF8DFFE9FAD3F86B00BF6E84" author="Preston HB" box="[1326,1470,1947,1971]" pageId="12" pageNumber="13" pagination="279 - 308" refId="ref15608" refString="Preston HB. A catalogue of the Asiatic naiades in the collection of the Indian Museum, Calcutta, with descriptions of new species. Records of the Indian Museum 1912; 7: 279 - 308." type="journal article" year="1912">Preston, 1912</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="D6804D68FF8DFFE9FC9BF84C00406EE4" authority="Preston, 1912" authorityName="Preston" authorityYear="1912" box="[870,1345,1979,2003]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="variety" species="favidens" variety="assamensis">
<emphasis id="23F4EAF9FF8DFFE9FC9BF84C01156EE4" box="[870,1044,1979,2003]" italics="true" pageId="12" pageNumber="13">Parreysia favidens</emphasis>
var.
<emphasis id="23F4EAF9FF8DFFE9FBBAF84C01AE6EE5" box="[1095,1199,1979,2002]" italics="true" pageId="12" pageNumber="13">assamensis</emphasis>
<bibRefCitation id="75114B1AFF8DFFE9FB49F84C00406EE4" author="Preston HB" box="[1204,1345,1979,2003]" pageId="12" pageNumber="13" pagination="279 - 308" refId="ref15608" refString="Preston HB. A catalogue of the Asiatic naiades in the collection of the Indian Museum, Calcutta, with descriptions of new species. Records of the Indian Museum 1912; 7: 279 - 308." type="journal article" year="1912">Preston, 1912</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="75114B1AFF8DFFE8FAAFF84C05A5699F" author="Subba Rao NV" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" refId="ref15978" refString="Subba Rao NV. Handbook. Freshwater Molluscs of India. Calcutta: Zoological Survey of India, 1989." type="book" year="1989">Subba Rao 1989</bibRefCitation>
,
<bibRefCitation id="75114B1AFF8CFFE8FF4DFF670479699F" author="Bolotov IN &amp; Pasupuleti R &amp; Subba Rao NV" box="[176,376,144,168]" pageId="13" pageNumber="14" refId="ref13300" refString="Bolotov IN, Pasupuleti R, Subba Rao NV et al. Oriental freshwater mussels arose in East Gondwana and arrived to Asia on the Indian Plate and Burma Terrane. Scientific Reports 2022 a; 12: 1518. https: // doi. org / 10.1038 / s 41598 - 022 - 05257 - 0" type="journal volume" year="2022">
Bolotov
<emphasis id="23F4EAF9FF8CFFE8FEFBFF660434699F" box="[262,309,144,168]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2022a
</bibRefCitation>
). These species were separated on the basis of minor morphological differences in shell shape (
<bibRefCitation id="75114B1AFF8CFFE8FD53FF4705A569D0" author="Benson WH" pageId="13" pageNumber="14" pagination="184 - 95" refId="ref12789" refString="Benson WH. Descriptions of Indian and Burmese species of the genus Unio, Retz. Annals and Magazine of Natural History 1862; 10: 184 - 95. https: // doi. org / 10.1080 / 00222936208681307" type="journal article" year="1862">Benson 1862</bibRefCitation>
,
<bibRefCitation id="75114B1AFF8CFFE8FF4CFF38044A69D0" author="Theobald W" box="[177,331,207,231]" pageId="13" pageNumber="14" pagination="207 - 9" refId="ref16043" refString="Theobald W. Descriptions of new species of Unionidae. Journal of the Asiatic Society of Bengal 1873; 42: 207 - 9." type="journal article" year="1873">Theobald 1873</bibRefCitation>
,
<bibRefCitation id="75114B1AFF8CFFE8FEABFF3804DF69D0" author="Preston HB" box="[342,478,207,231]" pageId="13" pageNumber="14" pagination="279 - 308" refId="ref15608" refString="Preston HB. A catalogue of the Asiatic naiades in the collection of the Indian Museum, Calcutta, with descriptions of new species. Records of the Indian Museum 1912; 7: 279 - 308." type="journal article" year="1912">Preston 1912</bibRefCitation>
). Our phylogenetic data reveal that there is a single
<taxonomicName id="D6804D68FF8CFFE8FE93FF1807146831" box="[366,533,238,262]" pageId="13" pageNumber="14">
<emphasis id="23F4EAF9FF8CFFE8FE93FF1804C66831" box="[366,455,239,262]" italics="true" pageId="13" pageNumber="14">Parreysia</emphasis>
species
</taxonomicName>
in
<collectingRegion id="D344F809FF8CFFE8FDC9FF1907766831" box="[564,631,238,262]" country="India" name="Assam" pageId="13" pageNumber="14">Assam</collectingRegion>
(Supporting Information,
<figureCitation id="89BB2A6EFF8CFFE8FF04FEFA04416812" box="[249,320,269,293]" captionStart="Figure 1" captionStartId="2.[129,194,1112,1136]" captionTargetBox="[128,1475,144,1084]" captionTargetId="figure-317@2.[128,1475,144,1084]" captionTargetPageId="2" captionText="Figure 1. Map of the Indian subcontinent with location of the study region (the frame) (A) and enlarged map of the study region in northeastern India (B). Assam is indicated by green filling (A) and cross-hatching (B). Green circles with numbers indicate the sampling localities in Assam:Burhidihing River, Dibrugarh (1); Gelua River, Dhemaji (2); an oxbow lake of Dhansiri River, Golaghat (3); old channel/pond, Telam River basin, Telam, Dhemaji (4); an oxbow lake of Tongani River (Tongani beel), Dhemaji (5); Dhansiri River, Golaghat (6); Sesa River, Dibrugarh (7); and a pond, Sutar Gaon, Kamrup (Rural) District (8). The maps were created using ESRI ArcGIS 10 software (www.esri. com/arcgis); the topographic base of the maps was created with Natural Earth Free Vector and Raster Map Data (www.naturalearthdata.com), Global Self-consistent Hierarchical High-resolution Geography (https://www.soest.hawaii.edu/wessel/gshhg), GEBCO_2022 (https://www. gebco.net/data_and_products/historical_data_sets/#gebco_2022), and Global Edge-matched Subnational Boundaries (https://fieldmaps. io). Maps: Mikhail Yu. Gofarov." pageId="13" pageNumber="14">Fig. S1</figureCitation>
), the conchological varieties of which correspond to the four nominal species, listed above. Here, we consider
<taxonomicName id="D6804D68FF8CFFE8FF57FEBA07056853" authority="(Benson, 1862)" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Benson" baseAuthorityYear="1862" box="[170,516,332,356]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="marcens" status="stat. nov.">
<emphasis id="23F4EAF9FF8CFFE8FF57FEBA04586853" box="[170,345,333,356]" italics="true" pageId="13" pageNumber="14">Parreysia marcens</emphasis>
(
<bibRefCitation id="75114B1AFF8CFFE8FE95FEBB04FD6853" author="Benson WH" box="[360,508,332,356]" pageId="13" pageNumber="14" pagination="184 - 95" refId="ref12789" refString="Benson WH. Descriptions of Indian and Burmese species of the genus Unio, Retz. Annals and Magazine of Natural History 1862; 10: 184 - 95. https: // doi. org / 10.1080 / 00222936208681307" type="journal article" year="1862">Benson, 1862</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="38C75782FF8CFFE8FDF6FEBB075E6854" box="[523,607,332,355]" pageId="13" pageNumber="14">stat. rev.</taxonomicNameLabel>
as an accepted species based on new DNA sequences and morphological data from
<collectingRegion id="D344F809FF8CFFE8FF5AFE7C05ED6894" box="[167,236,395,419]" country="India" name="Assam" pageId="13" pageNumber="14">Assam</collectingRegion>
, while three other taxa described from this region are considered its synonyms (
<tableCitation id="5C020350FF8CFFE8FE84FE5D04C268F5" box="[377,451,426,450]" captionStart="Table 1" captionStartId="4.[129,184,143,167]" captionTargetBox="[128,1473,189,1998]" captionText="Table 1. Checklist of freshwater mussel species (Unionidae:Parreysiinae) recorded from the Brahmaputra River basin in Assam, India." pageId="13" pageNumber="14">Table 1</tableCitation>
). This species has large genetic distance from its congeners (mean uncorrected
<emphasis id="23F4EAF9FF8CFFE8FDB5FE3D077568D6" box="[584,628,458,481]" italics="true" pageId="13" pageNumber="14">COI</emphasis>
p-distance of 7.3–8.3%) (Supporting Information,
<tableCitation id="5C020350FF8CFFE8FE19FE1E073B6B36" box="[484,570,489,513]" captionStart="Table 1" captionStartId="4.[129,184,143,167]" captionTargetBox="[128,1473,189,1998]" captionText="Table 1. Checklist of freshwater mussel species (Unionidae:Parreysiinae) recorded from the Brahmaputra River basin in Assam, India." pageId="13" pageNumber="14">Table S1</tableCitation>
).
<taxonomicName id="D6804D68FF8CFFE8FDB2FE1E07FB6B37" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Benson" baseAuthorityYear="1862" box="[591,762,489,512]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="marcens">
<emphasis id="23F4EAF9FF8CFFE8FDB2FE1E07FB6B37" box="[591,762,489,512]" italics="true" pageId="13" pageNumber="14">Parreysia marcens</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="D6804D68FF8CFFE8FF03FDFF04676B17" box="[254,358,520,544]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="keralaensis">
<emphasis id="23F4EAF9FF8CFFE8FF03FDFF04676B17" box="[254,358,520,544]" italics="true" pageId="13" pageNumber="14">keralaensis</emphasis>
</taxonomicName>
-group of the genus, being a sister-lineage of
<taxonomicName id="D6804D68FF8CFFE8FF70FDDF072D6B08" authority="Bolotov et al., 2022 a" authorityName="Bolotov" authorityYear="2022" box="[141,556,551,575]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="keralaensis">
<emphasis id="23F4EAF9FF8CFFE8FF70FDDF04546B08" box="[141,341,551,575]" italics="true" pageId="13" pageNumber="14">Parreysia keralaensis</emphasis>
<bibRefCitation id="75114B1AFF8CFFE8FEA6FDD0072D6B08" author="Bolotov IN &amp; Pasupuleti R &amp; Subba Rao NV" box="[347,556,551,575]" pageId="13" pageNumber="14" refId="ref13300" refString="Bolotov IN, Pasupuleti R, Subba Rao NV et al. Oriental freshwater mussels arose in East Gondwana and arrived to Asia on the Indian Plate and Burma Terrane. Scientific Reports 2022 a; 12: 1518. https: // doi. org / 10.1038 / s 41598 - 022 - 05257 - 0" type="journal volume" year="2022">
Bolotov
<emphasis id="23F4EAF9FF8CFFE8FE49FDDF04E36B08" box="[436,482,551,575]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
, 2022a
</bibRefCitation>
</taxonomicName>
from south-western
<collectingCountry id="6997767BFF8CFFE8FF8EFDB005AB6B68" box="[115,170,583,607]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
. Our occurrence and sequence datasets indicate that
<taxonomicName id="D6804D68FF8CFFE8FF8CFD90041F6B49" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Benson" baseAuthorityYear="1862" box="[113,286,615,638]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="marcens">
<emphasis id="23F4EAF9FF8CFFE8FF8CFD90041F6B49" box="[113,286,615,638]" italics="true" pageId="13" pageNumber="14">Parreysia marcens</emphasis>
</taxonomicName>
is endemic to the Brahmaputra River basin and that this species inhabits a broad range of habitats (Supporting Information, Dataset S1). This species was not recorded beyond the Brahmaputra Basin. Published records of other
<taxonomicName id="D6804D68FF8CFFE8FD8FFD3205966BCC" pageId="13" pageNumber="14">
<emphasis id="23F4EAF9FF8CFFE8FD8FFD3207CA6BEB" box="[626,715,709,732]" italics="true" pageId="13" pageNumber="14">Parreysia</emphasis>
species
</taxonomicName>
from
<collectingRegion id="D344F809FF8CFFE8FF29FD1304166BCB" box="[212,279,740,764]" country="India" name="Assam" pageId="13" pageNumber="14">Assam</collectingRegion>
(
<bibRefCitation id="75114B1AFF8CFFE8FED5FD1404D16BCC" author="Subba Rao NV" box="[296,464,739,763]" pageId="13" pageNumber="14" refId="ref15978" refString="Subba Rao NV. Handbook. Freshwater Molluscs of India. Calcutta: Zoological Survey of India, 1989." type="book" year="1989">Subba Rao 1989</bibRefCitation>
,
<bibRefCitation id="75114B1AFF8CFFE8FE26FD14079E6BCC" author="Sonowal J &amp; Puzari M &amp; Kardong D" box="[475,671,739,763]" pageId="13" pageNumber="14" pagination="18237 - 46" refId="ref15885" refString="Sonowal J, Puzari M, Kardong D. Diversity of freshwater molluscs from the upper Brahmaputra Basin, Assam, India. Journal of Threatened Taxa 2021; 13: 18237 - 46. https: // doi. org / 10.11609 / jott. 7144.13.5.18237 - 18246" type="journal article" year="2021">
Sonowal
<emphasis id="23F4EAF9FF8CFFE8FDC5FD1307676BCC" box="[568,614,739,763]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2021
</bibRefCitation>
,
<bibRefCitation id="75114B1AFF8CFFE8FD57FD1405E46A2C" author="Bolotov IN &amp; Pasupuleti R &amp; Subba Rao NV" pageId="13" pageNumber="14" refId="ref13300" refString="Bolotov IN, Pasupuleti R, Subba Rao NV et al. Oriental freshwater mussels arose in East Gondwana and arrived to Asia on the Indian Plate and Burma Terrane. Scientific Reports 2022 a; 12: 1518. https: // doi. org / 10.1038 / s 41598 - 022 - 05257 - 0" type="journal volume" year="2022">
Bolotov
<emphasis id="23F4EAF9FF8CFFE8FF8CFCF405A06A2C" box="[113,161,771,795]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2022a
</bibRefCitation>
) should be considered as belonging to
<taxonomicName id="D6804D68FF8CFFE8FD84FCF407DF6A2D" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Benson" baseAuthorityYear="1862" box="[633,734,771,794]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="marcens">
<emphasis id="23F4EAF9FF8CFFE8FD84FCF407DF6A2D" box="[633,734,771,794]" italics="true" pageId="13" pageNumber="14">P. marcens</emphasis>
</taxonomicName>
. A published
<emphasis id="23F4EAF9FF8CFFE8FF21FCD504096A0E" box="[220,264,802,825]" italics="true" pageId="13" pageNumber="14">COI</emphasis>
sequence of
<taxonomicName id="D6804D68FF8CFFE8FE6FFCD404EA6A0D" authorityName="Conrad" authorityYear="1853" box="[402,491,803,826]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8CFFE8FE6FFCD404EA6A0D" box="[402,491,803,826]" italics="true" pageId="13" pageNumber="14">Parreysia</emphasis>
</taxonomicName>
from
<collectingRegion id="D344F809FF8CFFE8FDD4FCD507886A0D" box="[553,649,802,826]" country="India" name="Nagaland" pageId="13" pageNumber="14">Nagaland</collectingRegion>
(GenBank acc. No. OM074301;
<bibRefCitation id="75114B1AFF8CFFE8FEB0FCB507466A6E" author="Imsong L &amp; Murali M" box="[333,583,833,858]" pageId="13" pageNumber="14" pagination="29 - 38" refId="ref14427" refString="Imsong L, Murali M. Efficacy of molecular identification using DNA barcoding technique of freshwater edible molluscs from Nagaland, India. Uttar Pradesh Journal of Zoology 2023; 44: 29 - 38. https: // doi. org / 10.56557 / upjoz / 2023 / v 44 i 203643" type="journal article" year="2023">Imsong and Murali 2023</bibRefCitation>
) that was erroneously identified as
<taxonomicName id="D6804D68FF8CFFE8FEC9FC9604F16A4F" baseAuthorityName="O. F. Muller" baseAuthorityYear="1774" box="[308,496,865,888]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="corrugata">
<emphasis id="23F4EAF9FF8CFFE8FEC9FC9604F16A4F" box="[308,496,865,888]" italics="true" pageId="13" pageNumber="14">Parreysia corrugata</emphasis>
</taxonomicName>
(see:
<bibRefCitation id="75114B1AFF8CFFE8FDCDFC9607F16A4E" author="Bolotov IN &amp; Konopleva ES &amp; Vikhrev IV" box="[560,752,865,889]" pageId="13" pageNumber="14" refId="ref13586" refString="Bolotov IN, Konopleva ES, Vikhrev IV et al. An alarming tendency towards freshwater mussel misidentification in scientific works may bias endangered and invasive species management. Aquatic Conservation: Marine and Freshwater Ecosystems 2024; 34: e 4061. https: // doi. org / 10.1002 / aqc. 4061" type="journal volume" year="2024">
Bolotov
<emphasis id="23F4EAF9FF8CFFE8FD78FC9607B76A4E" box="[645,694,865,889]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2024
</bibRefCitation>
) also belongs to
<taxonomicName id="D6804D68FF8CFFE8FEEBFC7604C76AAF" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Benson" baseAuthorityYear="1862" box="[278,454,897,920]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="marcens">
<emphasis id="23F4EAF9FF8CFFE8FEEBFC7604C76AAF" box="[278,454,897,920]" italics="true" pageId="13" pageNumber="14">Parreysia marcens</emphasis>
</taxonomicName>
(
<tableCitation id="5C020350FF8CFFE8FE24FC7707266AAF" box="[473,551,896,920]" captionStart="Table 1" captionStartId="4.[129,184,143,167]" captionTargetBox="[128,1473,189,1998]" captionText="Table 1. Checklist of freshwater mussel species (Unionidae:Parreysiinae) recorded from the Brahmaputra River basin in Assam, India." pageId="13" pageNumber="14">Table 1</tableCitation>
). This conclusion is based on the molecular genetic data (Supporting Information,
<figureCitation id="89BB2A6EFF8CFFE8FF8CFC4805B86AE0" box="[113,185,959,983]" captionStart="Figure 1" captionStartId="2.[129,194,1112,1136]" captionTargetBox="[128,1475,144,1084]" captionTargetId="figure-317@2.[128,1475,144,1084]" captionTargetPageId="2" captionText="Figure 1. Map of the Indian subcontinent with location of the study region (the frame) (A) and enlarged map of the study region in northeastern India (B). Assam is indicated by green filling (A) and cross-hatching (B). Green circles with numbers indicate the sampling localities in Assam:Burhidihing River, Dibrugarh (1); Gelua River, Dhemaji (2); an oxbow lake of Dhansiri River, Golaghat (3); old channel/pond, Telam River basin, Telam, Dhemaji (4); an oxbow lake of Tongani River (Tongani beel), Dhemaji (5); Dhansiri River, Golaghat (6); Sesa River, Dibrugarh (7); and a pond, Sutar Gaon, Kamrup (Rural) District (8). The maps were created using ESRI ArcGIS 10 software (www.esri. com/arcgis); the topographic base of the maps was created with Natural Earth Free Vector and Raster Map Data (www.naturalearthdata.com), Global Self-consistent Hierarchical High-resolution Geography (https://www.soest.hawaii.edu/wessel/gshhg), GEBCO_2022 (https://www. gebco.net/data_and_products/historical_data_sets/#gebco_2022), and Global Edge-matched Subnational Boundaries (https://fieldmaps. io). Maps: Mikhail Yu. Gofarov." pageId="13" pageNumber="14">Fig. S1</figureCitation>
) and published image of the shell (
<bibRefCitation id="75114B1AFF8CFFE8FDCEFC4805A56AC1" author="Imsong L &amp; Murali M" pageId="13" pageNumber="14" pagination="29 - 38" refId="ref14427" refString="Imsong L, Murali M. Efficacy of molecular identification using DNA barcoding technique of freshwater edible molluscs from Nagaland, India. Uttar Pradesh Journal of Zoology 2023; 44: 29 - 38. https: // doi. org / 10.56557 / upjoz / 2023 / v 44 i 203643" type="journal article" year="2023">Imsong and Murali 2023</bibRefCitation>
: fig. 1G).
</paragraph>
<paragraph id="113F36EBFF8CFFE8FF70FC0A04D26CBA" blockId="13.[112,764,144,1421]" pageId="13" pageNumber="14">
Additionally,
<taxonomicName id="D6804D68FF8CFFE8FEE1FC0907A66D22" authority="(Benson, 1862)" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Benson" baseAuthorityYear="1862" box="[284,679,1021,1045]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="smaragdites">
<emphasis id="23F4EAF9FF8CFFE8FEE1FC0904F56D22" box="[284,500,1021,1045]" italics="true" pageId="13" pageNumber="14">Parreysia smaragdites</emphasis>
(
<bibRefCitation id="75114B1AFF8CFFE8FDFBFC0A079E6D22" author="Benson WH" box="[518,671,1021,1045]" pageId="13" pageNumber="14" pagination="184 - 95" refId="ref12789" refString="Benson WH. Descriptions of Indian and Burmese species of the genus Unio, Retz. Annals and Magazine of Natural History 1862; 10: 184 - 95. https: // doi. org / 10.1080 / 00222936208681307" type="journal article" year="1862">Benson, 1862</bibRefCitation>
)
</taxonomicName>
is here considered a putative accepted species based on morphological features such as a rather thin shell and prominent, conical umbo (see:
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Preece
<emphasis id="23F4EAF9FF8CFFE8FF01FBAB04316D44" box="[252,304,1115,1139]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2022
</bibRefCitation>
: fig. 111A). Comparative analysis of a
<typeStatus id="CE3B8849FF8CFFE8FF79FB8C05D46DA4" box="[132,213,1147,1171]" pageId="13" pageNumber="14" type="syntype">syntype</typeStatus>
of
<taxonomicName id="D6804D68FF8CFFE8FF07FB8C04856DA4" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Benson" baseAuthorityYear="1862" box="[250,388,1147,1171]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="smaragdites">
<emphasis id="23F4EAF9FF8CFFE8FF07FB8C04856DA4" box="[250,388,1147,1171]" italics="true" pageId="13" pageNumber="14">P. smaragdites</emphasis>
</taxonomicName>
(UMZC I.103360.A) (
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Preece
<emphasis id="23F4EAF9FF8CFFE8FD37FB8C07FA6DA4" box="[714,763,1147,1171]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2022
</bibRefCitation>
) and the
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of
<taxonomicName id="D6804D68FF8CFFE8FE6EFB6C04466DE6" authority="Hanley and Theobald, 1872" authorityName="Hanley and Theobald" authorityYear="1872" class="Bivalvia" family="Unionidae" genus="Unio" kingdom="Animalia" order="Unionida" pageId="13" pageNumber="14" phylum="Mollusca" rank="variety" species="corrugatus" variety="fragilis">
<emphasis id="23F4EAF9FF8CFFE8FE6EFB6C07336D85" box="[403,562,1179,1202]" italics="true" pageId="13" pageNumber="14">Unio corrugatus</emphasis>
var.
<emphasis id="23F4EAF9FF8CFFE8FD9AFB6D07A96D85" box="[615,680,1178,1202]" italics="true" pageId="13" pageNumber="14">fragilis</emphasis>
<bibRefCitation id="75114B1AFF8CFFE8FD4DFB6D04466DE6" author="Hanley S &amp; Theobald W" pageId="13" pageNumber="14" refId="ref14260" refString="Hanley S, Theobald W. Var. fragilis. Conchologia Indica: Illustrations of the Land and Freshwater Shells of British India 1872; 3: 21, pl. 45, fig. 4." type="journal volume" year="1872">Hanley and Theobald, 1872</bibRefCitation>
</taxonomicName>
(NHMUK 1968651; unavailable name) from an unknown type locality indicates that these taxa are conspecific (
<tableCitation id="5C020350FF8CFFE8FF08FB0F04406C27" box="[245,321,1272,1296]" captionStart="Table 1" captionStartId="4.[129,184,143,167]" captionTargetBox="[128,1473,189,1998]" captionText="Table 1. Checklist of freshwater mussel species (Unionidae:Parreysiinae) recorded from the Brahmaputra River basin in Assam, India." pageId="13" pageNumber="14">Table 1</tableCitation>
). It is likely that the latter shell also originated from the Brahmaputra River basin. We were unable to find living specimens of
<taxonomicName id="D6804D68FF8CFFE8FEBDFAC004C86C78" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Benson" baseAuthorityYear="1862" box="[320,457,1335,1359]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="smaragdites">
<emphasis id="23F4EAF9FF8CFFE8FEBDFAC004C86C78" box="[320,457,1335,1359]" italics="true" pageId="13" pageNumber="14">P. smaragdites</emphasis>
</taxonomicName>
in
<collectingRegion id="D344F809FF8CFFE8FE17FAC007316C78" box="[490,560,1335,1359]" country="India" name="Assam" pageId="13" pageNumber="14">Assam</collectingRegion>
, and the identity of this interesting species needs to be checked by means of a DNA sequencing approach in the future.
</paragraph>
</subSubSection>
</treatment>
</document>

263
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Genus
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(
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<emphasis id="23F4EAF9FF8DFFE9FE8CFD9C07186BB2" bold="true" box="[369,537,619,646]" pageId="12" pageNumber="13">
(
<figureCitation id="89BB2A6EFF8DFFE9FE86FD9C04C86BB2" box="[379,457,619,645]" captionStart="Figure 4" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[174,1429,144,1808]" captionTargetId="figure-8@8.[174,1429,144,1808]" captionTargetPageId="8" captionText="Figure 4. Shells of Lamellidentini species from Assam (sequenced specimens; outer view of the left valve and inner view of the right valve).A, Balwantia soleniformis (Benson, 1836), sample DLS-DU BS-BDH-1. B–G, Lamellidens jenkinsianus (Benson, 1862): topotypes LJ-NKM-2 (B), LJ-NKM-3 (C), L5-NKM-1 (D), L4-NKM-1 (E), L5-NKM-4 (F), and L4-NKM-2 (G). Scale bars = 10 mm. The shell vouchers of the new samples are deposited in FBRC ZSI and DLS-DU (Supporting Information, Dataset S1). Photos:J.Sonowal and D. Kardong." pageId="12" pageNumber="13">Figs 4A</figureCitation>
,
<figureCitation id="89BB2A6EFF8DFFE9FE29FD9B070F6BB2" box="[468,526,619,646]" captionStart="Figure 7" captionStartId="11.[112,177,1302,1326]" captionTargetBox="[114,1458,161,1274]" captionTargetId="figure-269@11.[114,1458,144,1274]" captionTargetPageId="11" captionText="Figure 7. Habitats of little-known freshwater mussel species in Assam.A, collecting locality of Assamnaia involuta gen. et comb.nov.: a shallow oxbow lake with silt and clay bottom substrate, Dhansiri River valley, Golaghat, 26 March 2022. B, collecting locality of Balwantia soleniformis, Indonaia corbis, Pseudoscabies scobina gen. et comb.nov., and Parreysia marcens: Burhidihing River, Dibrugarh, 22 February 2022. The white arrow shows the river cliff, whose bottom portion houses numerous B. soleniformis burrows; the river cliff is mostly composed by firm ground (hard clay). C, close-up view of an aggregation of B. soleniformis burrows in the river cliff,22 February 2022. Living mussels were extracted from submerged burrows.Photos: J. Sonowal and D. Kardong." pageId="12" pageNumber="13">7B, C</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="599A6560FF8DFFE9FF7CFD5B06086D6C" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="113F36EBFF8DFFE9FF7CFD5B06086D6C" blockId="12.[129,778,684,1116]" pageId="12" pageNumber="13">
<emphasis id="23F4EAF9FF8DFFE9FF7CFD5B04BE6BF3" box="[129,447,684,708]" italics="true" pageId="12" pageNumber="13">
Differential diagnosis:
<taxonomicName id="D6804D68FF8DFFE9FEA1FD5B04BE6BF3" authorityName="Prashad" authorityYear="1919" box="[348,447,684,708]" class="Bivalvia" family="Unionidae" genus="Balwantia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">Balwantia</taxonomicName>
</emphasis>
is the sister-lineage of the genus
<taxonomicName id="D6804D68FF8DFFE9FF7FFD3C05F66BD4" box="[130,247,715,739]" class="Bivalvia" family="Unionidae" genus="Trapezidens" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FF7FFD3C05F66BD4" box="[130,247,715,739]" italics="true" pageId="12" pageNumber="13">Trapezidens</emphasis>
</taxonomicName>
but it cannot be placed in the latter genus due to a combination of specific morphological characters: ultra-elongated, narrow shell (vs. ovate or rhomboid, broad) and reduced pseudocardinal teeth (vs. well-developed, massive). Conchologically,
<taxonomicName id="D6804D68FF8DFFE9FEC0FCBE04A16A56" authorityName="Prashad" authorityYear="1919" box="[317,416,841,865]" class="Bivalvia" family="Unionidae" genus="Balwantia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FEC0FCBE04A16A56" box="[317,416,841,865]" italics="true" pageId="12" pageNumber="13">Balwantia</emphasis>
</taxonomicName>
resembles other ultra-elongated genera of freshwater mussels such as
<taxonomicName id="D6804D68FF8DFFE9FDE8FC9F06046AB7" authority="Conrad, 1869" authorityName="Conrad" authorityYear="1869" box="[533,773,872,896]" class="Bivalvia" family="Unionidae" genus="Solenaia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FDE8FC9F07666AB7" box="[533,615,872,896]" italics="true" pageId="12" pageNumber="13">Solenaia</emphasis>
Conrad, 1869
</taxonomicName>
,
<taxonomicName id="D6804D68FF8DFFE9FF7CFC7004E16AA8" authority="Bolotov et al., 2021" authorityName="Bolotov" authorityYear="2021" box="[129,480,903,927]" class="Bivalvia" family="Unionidae" genus="Sinosolenaia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FF7CFC7005F86AA8" box="[129,249,903,927]" italics="true" pageId="12" pageNumber="13">Sinosolenaia</emphasis>
<bibRefCitation id="75114B1AFF8DFFE9FEF8FC7004E16AA8" author="Bolotov IN &amp; Kondakov AV &amp; Konopleva ES" box="[261,480,903,927]" pageId="12" pageNumber="13" pagination="1 - 6" refId="ref13249" refString="Bolotov IN, Kondakov AV, Konopleva ES et al. A new genus of ultra-elongate freshwater mussels from Vietnam and eastern China (Bivalvia: Unionidae). Ecologica Montenegrina 2021; 39: 1 - 6. https: // doi. org / 10.37828 / em. 2021.39.1" type="journal article" year="2021">
Bolotov
<emphasis id="23F4EAF9FF8DFFE9FE9EFC7F04926AA8" box="[355,403,903,927]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
., 2021
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="D6804D68FF8DFFE9FE0CFC7006046AA8" authority="Conrad, 1853" authorityName="Conrad" authorityYear="1853" box="[497,773,903,927]" class="Bivalvia" family="Unionidae" genus="Lanceolaria" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FE0CFC7007626AA8" box="[497,611,903,927]" italics="true" pageId="12" pageNumber="13">Lanceolaria</emphasis>
<bibRefCitation id="75114B1AFF8DFFE9FD8DFC7006046AA8" author="Conrad TA" box="[624,773,903,927]" pageId="12" pageNumber="13" pagination="243 - 69" refId="ref13680" refString="Conrad TA. A synopsis of the family of Naiades of North America, with notes, and a table of some of the genera and sub-genera of the family, according to their geographical distribution, and descriptions of genera and sub-genera. Proceedings of the Academy of Natural Sciences of Philadelphia 1853; 6: 243 - 69." type="journal article" year="1853">Conrad, 1853</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="D6804D68FF8DFFE9FF7CFC5004E96A88" authority="Huang and Wu, 2019" authorityName="Huang and Wu" authorityYear="2019" box="[129,488,935,959]" class="Bivalvia" family="Unionidae" genus="Parvasolenaia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FF7CFC5004066A88" box="[129,263,935,959]" italics="true" pageId="12" pageNumber="13">Parvasolenaia</emphasis>
Huang and Wu, 2019
</taxonomicName>
, and
<taxonomicName id="D6804D68FF8DFFE9FDE3FC5005B66AE9" authority="Lee et al., 2020" authorityName="Lee" authorityYear="2020" class="Bivalvia" family="Unionidae" genus="Koreosolenaia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FDE3FC5007A56A88" box="[542,676,935,959]" italics="true" pageId="12" pageNumber="13">Koreosolenaia</emphasis>
Lee
<emphasis id="23F4EAF9FF8DFFE9FD2BFC5007FF6A88" box="[726,766,935,959]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
., 2020
</taxonomicName>
. External similarity between these genera reflects convergent evolution and most of them cannot be distinguished based on conchological features alone (
<bibRefCitation id="75114B1AFF8DFFE9FE2FFBF2078E6D2A" author="Bolotov IN &amp; Kondakov AV &amp; Konopleva ES" box="[466,655,1029,1053]" pageId="12" pageNumber="13" pagination="1 - 6" refId="ref13249" refString="Bolotov IN, Kondakov AV, Konopleva ES et al. A new genus of ultra-elongate freshwater mussels from Vietnam and eastern China (Bivalvia: Unionidae). Ecologica Montenegrina 2021; 39: 1 - 6. https: // doi. org / 10.37828 / em. 2021.39.1" type="journal article" year="2021">
Bolotov
<emphasis id="23F4EAF9FF8DFFE9FDD5FBF207566D2A" box="[552,599,1029,1053]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2021
</bibRefCitation>
).
<taxonomicName id="D6804D68FF8DFFE9FD5BFBF206086D2A" authorityName="Prashad" authorityYear="1919" box="[678,777,1029,1053]" class="Bivalvia" family="Unionidae" genus="Balwantia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FD5BFBF206086D2A" box="[678,777,1029,1053]" italics="true" pageId="12" pageNumber="13">Balwantia</emphasis>
</taxonomicName>
differs from these genera by having unhooked glochidia and tetragenous brooding mode (
<bibRefCitation id="75114B1AFF8DFFE9FE58FBB4072C6D6C" author="Prashad B" box="[421,557,1091,1115]" pageId="12" pageNumber="13" pagination="289 - 96" refId="ref15418" refString="Prashad B. Studies on the anatomy of Indian Mollusca. The soft parts of some Indian Unionidae. Records of the Indian Museum 1919; 16: 289 - 96." type="journal article" year="1919">Prashad 1919</bibRefCitation>
,
<bibRefCitation id="75114B1AFF8DFFE9FDCAFBB407F86D6C" author="Bolotov IN &amp; Pasupuleti R &amp; Subba Rao NV" box="[567,761,1091,1115]" pageId="12" pageNumber="13" refId="ref13300" refString="Bolotov IN, Pasupuleti R, Subba Rao NV et al. Oriental freshwater mussels arose in East Gondwana and arrived to Asia on the Indian Plate and Burma Terrane. Scientific Reports 2022 a; 12: 1518. https: // doi. org / 10.1038 / s 41598 - 022 - 05257 - 0" type="journal volume" year="2022">
Bolotov
<emphasis id="23F4EAF9FF8DFFE9FD76FBB307B96D6C" box="[651,696,1091,1115]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2022a
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="599A6560FF8DFFE9FF7CFB8A04BB6CB8" pageId="12" pageNumber="13" type="discussion">
<paragraph id="113F36EBFF8DFFE9FF7CFB8A04BB6CB8" blockId="12.[128,779,1149,1423]" pageId="12" pageNumber="13">
<emphasis id="23F4EAF9FF8DFFE9FF7CFB8A05F06DA3" box="[129,241,1149,1172]" italics="true" pageId="12" pageNumber="13">Comments:</emphasis>
The genus is monotypic and contains a single species having an ultra-elongated shell,
<taxonomicName id="D6804D68FF8DFFE9FE42FB6A074B6D83" authorityName="Benson" authorityYear="1836" box="[447,586,1180,1204]" class="Bivalvia" family="Unionidae" genus="Anodonta" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="soleniformis">
<emphasis id="23F4EAF9FF8DFFE9FE42FB6A074B6D83" box="[447,586,1180,1204]" italics="true" pageId="12" pageNumber="13">B. soleniformis</emphasis>
</taxonomicName>
(
<tableCitation id="5C020350FF8DFFE9FDA7FB6B07A56D83" box="[602,676,1180,1204]" captionStart="Table 1" captionStartId="4.[129,184,143,167]" captionTargetBox="[128,1473,189,1998]" captionText="Table 1. Checklist of freshwater mussel species (Unionidae:Parreysiinae) recorded from the Brahmaputra River basin in Assam, India." pageId="12" pageNumber="13">Table 1</tableCitation>
;
<figureCitation id="89BB2A6EFF8DFFE9FD4DFB6B07F86D83" box="[688,761,1180,1204]" captionStart="Figure 4" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[174,1429,144,1808]" captionTargetId="figure-8@8.[174,1429,144,1808]" captionTargetPageId="8" captionText="Figure 4. Shells of Lamellidentini species from Assam (sequenced specimens; outer view of the left valve and inner view of the right valve).A, Balwantia soleniformis (Benson, 1836), sample DLS-DU BS-BDH-1. B–G, Lamellidens jenkinsianus (Benson, 1862): topotypes LJ-NKM-2 (B), LJ-NKM-3 (C), L5-NKM-1 (D), L4-NKM-1 (E), L5-NKM-4 (F), and L4-NKM-2 (G). Scale bars = 10 mm. The shell vouchers of the new samples are deposited in FBRC ZSI and DLS-DU (Supporting Information, Dataset S1). Photos:J.Sonowal and D. Kardong." pageId="12" pageNumber="13">Fig. 4A</figureCitation>
). Our new phylogenetic results support the morphology-based assumption that this genus belongs to the
<taxonomicName id="D6804D68FF8DFFE9FDBCFB2C07BC6DC4" authorityName="Henderson" authorityYear="1935" box="[577,701,1243,1267]" class="Bivalvia" family="Unionidae" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="subFamily" subFamily="Parreysiinae">Parreysiinae</taxonomicName>
due to the presence of unhooked glochidia and tetragenous brooding mode (
<bibRefCitation id="75114B1AFF8DFFE9FF30FAEE048B6C06" author="Pfeiffer JM &amp; Graf DL &amp; Cummings KS" box="[205,394,1305,1329]" pageId="12" pageNumber="13" pagination="394 - 470" refId="ref15331" refString="Pfeiffer JM, Graf DL, Cummings KS et al. Taxonomic revision of a radiation of South-East Asian freshwater mussels (Unionidae: Gonideinae: Contradentini + Rectidentini). Invertebrate Systematics 2021; 35: 394 - 470. https: // doi. org / 10.1071 / is 20044" type="journal article" year="2021">
Pfeiffer
<emphasis id="23F4EAF9FF8DFFE9FEE2FAED044E6C06" box="[287,335,1305,1329]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2021
</bibRefCitation>
,
<bibRefCitation id="75114B1AFF8DFFE9FE6BFAEE07656C06" author="Bolotov IN &amp; Pasupuleti R &amp; Subba Rao NV" box="[406,612,1305,1329]" pageId="12" pageNumber="13" refId="ref13300" refString="Bolotov IN, Pasupuleti R, Subba Rao NV et al. Oriental freshwater mussels arose in East Gondwana and arrived to Asia on the Indian Plate and Burma Terrane. Scientific Reports 2022 a; 12: 1518. https: // doi. org / 10.1038 / s 41598 - 022 - 05257 - 0" type="journal volume" year="2022">
Bolotov
<emphasis id="23F4EAF9FF8DFFE9FE12FAED071E6C06" box="[495,543,1305,1329]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2022a
</bibRefCitation>
). We also show that it is a representative of the tribe
<taxonomicName id="D6804D68FF8DFFE9FDEFFACE07AA6C66" authorityName="Modell" authorityYear="1942" box="[530,683,1337,1361]" class="Bivalvia" family="Unionidae" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="tribe" tribe="Lamellidentini">Lamellidentini</taxonomicName>
, whereas
<taxonomicName id="D6804D68FF8DFFE9FF7CFAAF05E46C47" authorityName="Prashad" authorityYear="1919" box="[129,229,1368,1392]" class="Bivalvia" family="Unionidae" genus="Balwantia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FF7CFAAF05E46C47" box="[129,229,1368,1392]" italics="true" pageId="12" pageNumber="13">Balwantia</emphasis>
</taxonomicName>
was placed as incertae sedis
<taxonomicName id="D6804D68FF8DFFE9FDF8FAAF07806C47" authorityName="Henderson" authorityYear="1935" box="[517,641,1368,1392]" class="Bivalvia" family="Unionidae" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="subFamily" subFamily="Parreysiinae">Parreysiinae</taxonomicName>
in our earlier revision (
<bibRefCitation id="75114B1AFF8DFFE9FF1FFA8004AB6CB8" author="Bolotov IN &amp; Pasupuleti R &amp; Subba Rao NV" box="[226,426,1399,1423]" pageId="12" pageNumber="13" refId="ref13300" refString="Bolotov IN, Pasupuleti R, Subba Rao NV et al. Oriental freshwater mussels arose in East Gondwana and arrived to Asia on the Indian Plate and Burma Terrane. Scientific Reports 2022 a; 12: 1518. https: // doi. org / 10.1038 / s 41598 - 022 - 05257 - 0" type="journal volume" year="2022">
Bolotov
<emphasis id="23F4EAF9FF8DFFE9FEC5FA8F04666CB8" box="[312,359,1399,1423]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2022a
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:mods id="58A93F7DAF531E446524A0BB9F8728C4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="0DEFCEAF0AF6FFE4C06F799712768B53">Discovery of an endemism hotspot of freshwater mussels (Bivalvia: Unionidae) in Assam, with a description of two new genera</mods:title>
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<mods:roleTerm id="A1DC12C5D64F572EF7EC0681E4937DE1">Author</mods:roleTerm>
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<mods:namePart id="F9F7BC22D3E2D9E1083EA875B6996E4E">Bolotov, Ivan N.</mods:namePart>
<mods:affiliation id="461BFD80DB778066FC218F159B7A3127">N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Av. 20, 163020 Arkhangelsk, Russia &amp; SSC / IUCN-Mollusc Specialist Group, Species Survival Commission, International Union for Conservation of Nature, CB 2 3 QZ Cambridge, United Kingdom</mods:affiliation>
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<mods:namePart id="F713362E98800644977112F3AD020A7E">Sonowal, Jyotish</mods:namePart>
<mods:affiliation id="A67BE220237B5F1E72270CA955A02877">Centre for Biotechnology and Bioinformatics, Dibrugarh University, 786004 Dibrugarh, Assam, India</mods:affiliation>
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<mods:namePart id="31495FE2384A7783F2B8CB54B8E9D841">Kardong, Devid</mods:namePart>
<mods:affiliation id="2D07F92AC8895733CF363C2A17C09D56">Department of Life Sciences, Dibrugarh University, 786004 Dibrugarh, Assam, India</mods:affiliation>
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<mods:namePart id="DACBF23879212549A0E413403855F4F2">Pasupuleti, Rajeev</mods:namePart>
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<mods:namePart id="67AD1A075BACB86B5BF81FD566419D82">Subba Rao, Nalluri V.</mods:namePart>
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<mods:affiliation id="54DB3EFA21060B62C70F9BEFD4C85038">N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Av. 20, 163020 Arkhangelsk, Russia</mods:affiliation>
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<mods:affiliation id="8DABC4491D860B4C989756627145936A">N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Av. 20, 163020 Arkhangelsk, Russia</mods:affiliation>
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<mods:affiliation id="EFE5DC02ADFBD82F5970A6AEBD0DF6F0">N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Av. 20, 163020 Arkhangelsk, Russia</mods:affiliation>
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<mods:affiliation id="AA0D5B4704CAA231CB94442E81F01500">N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Av. 20, 163020 Arkhangelsk, Russia &amp; SSC / IUCN-Mollusc Specialist Group, Species Survival Commission, International Union for Conservation of Nature, CB 2 3 QZ Cambridge, United Kingdom</mods:affiliation>
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<mods:title id="76892BE4BCD06B41AE909EAAF293B095">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="BD050BCDE747F5F461795760622F66C3">
<mods:date id="10FBFCF0F7708DD3CDE2D69FB3DEDF7A">2024</mods:date>
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<mods:title id="65FD40ACB81F69371C4A5DDF3398544B">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="8C19D486C99B22871B7B1BDACC2D6E7D">2024-05-17</mods:number>
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<mods:number id="81D6088C28E2D722827DF4A1A34BECB5">202</mods:number>
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<subSubSection id="599A6560FF8DFFE9FF36F8FE07386E74" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="113F36EBFF8DFFE9FF36F8FE07386E74" blockId="12.[186,720,1801,1891]" pageId="12" pageNumber="13">
Genus
<taxonomicName id="D6804D68FF8DFFE9FEE8F8FE04CC6E74" ID-CoL="CCS3Y" authority="Bolotov, Sonowal, Kardong, Pasupuleti and Subba Rao" authorityName="Bolotov, Sonowal, Kardong, Pasupuleti and Subba Rao" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Assamnaia" higherTaxonomySource="CoL" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="23F4EAF9FF8DFFE9FEE8F8FE04936E14" bold="true" box="[277,402,1801,1827]" italics="true" pageId="12" pageNumber="13">Assamnaia</emphasis>
Bolotov, Sonowal, Kardong, Pasupuleti and Subba Rao
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<taxonomicNameLabel id="38C75782FF8DFFE9FE2FF8DE07386E74" box="[466,569,1833,1859]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="599A6560FF8DFFE9FDC3F8DF075F6E55" pageId="12" pageNumber="13" type="type_taxon">
<paragraph id="113F36EBFF8DFFE9FDC3F8DF075F6E55" blockId="12.[186,720,1801,1891]" pageId="12" pageNumber="13">
(
<typeStatus id="CE3B8849FF8DFFE9FDB8F8DF07766E75" box="[581,631,1832,1858]" pageId="12" pageNumber="13">type</typeStatus>
species:
<taxonomicName id="D6804D68FF8DFFE9FED1F8BF07506E55" ID-CoL="7TJQ9" authority="Hanley, 1856" authorityName="Hanley" authorityYear="1856" box="[300,593,1864,1891]" class="Bivalvia" family="Unionidae" genus="Unio" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="involutus">
<emphasis id="23F4EAF9FF8DFFE9FED1F8BF04BC6E54" bold="true" box="[300,445,1864,1891]" italics="true" pageId="12" pageNumber="13">Unio involutus</emphasis>
<bibRefCitation id="75114B1AFF8DFFE9FE3FF8BF07506E55" author="Hanley S" box="[450,593,1864,1890]" pageId="12" pageNumber="13" refId="ref14239" refString="Hanley S. An Illustrated and Descriptive Catalogue of Recent Bivalve Shells. London: Williams and Norgate, 1856." type="book" year="1856">Hanley, 1856</bibRefCitation>
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).
</paragraph>
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<subSubSection id="599A6560FF8DFFE9FE92F88E06966A85" pageId="12" pageNumber="13" type="description">
<paragraph id="113F36EBFF8DFFE9FE92F88E071D6EA4" blockId="12.[129,778,1913,2007]" box="[367,540,1913,1940]" pageId="12" pageNumber="13">
<emphasis id="23F4EAF9FF8DFFE9FE92F88E071D6EA4" bold="true" box="[367,540,1913,1940]" pageId="12" pageNumber="13">
(
<figureCitation id="89BB2A6EFF8DFFE9FE84F88E04E96EA4" box="[377,488,1913,1939]" captionStart="Figure 6" captionStartId="10.[129,194,1725,1749]" captionTargetBox="[129,1473,144,1698]" captionTargetId="figure-79@10.[129,1473,144,1698]" captionTargetPageId="10" captionText="Figure 6. Shells of Parreysiini species from Assam (outer view of the left valve and inner view of the right valve). A–C, Assamnaia involuta (Hanley, 1856) gen. et comb.nov.: topotypes II-DHN-1 (A), II-DHN-2 (B), and DLS-DU II-D (dead shell; non-sequenced) (C). D–I, Parreysia marcens (Benson, 1862) stat.rev.: topotypes PS-SES-5 (D), PS-SES-1 (E), P1-SES-4 (F), PC-SES-1 (G), PSm-SES-2 (H), and PF-BDH-4 (I). Scale bars = 10 mm.The shell vouchers of the new samples are deposited in FBRC ZSI and DLS-DU (Supporting Information, Dataset S1). Photos:J.Sonowal and D. Kardong." pageId="12" pageNumber="13">Figs 6A–C</figureCitation>
,
<figureCitation id="89BB2A6EFF8DFFE9FE0FF88E07116EA4" box="[498,528,1913,1939]" captionStart="Figure 7" captionStartId="11.[112,177,1302,1326]" captionTargetBox="[114,1458,161,1274]" captionTargetId="figure-269@11.[114,1458,144,1274]" captionTargetPageId="11" captionText="Figure 7. Habitats of little-known freshwater mussel species in Assam.A, collecting locality of Assamnaia involuta gen. et comb.nov.: a shallow oxbow lake with silt and clay bottom substrate, Dhansiri River valley, Golaghat, 26 March 2022. B, collecting locality of Balwantia soleniformis, Indonaia corbis, Pseudoscabies scobina gen. et comb.nov., and Parreysia marcens: Burhidihing River, Dibrugarh, 22 February 2022. The white arrow shows the river cliff, whose bottom portion houses numerous B. soleniformis burrows; the river cliff is mostly composed by firm ground (hard clay). C, close-up view of an aggregation of B. soleniformis burrows in the river cliff,22 February 2022. Living mussels were extracted from submerged burrows.Photos: J. Sonowal and D. Kardong." pageId="12" pageNumber="13">7A</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="113F36EBFF8DFFE9FF7CF85707316EE0" blockId="12.[129,778,1913,2007]" pageId="12" pageNumber="13">
LSID: https://zoobank.org/
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urn:lsid:zoobank.org:act:
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<paragraph id="113F36EBFF8DFFE9FCC4FF67000E6B08" blockId="12.[825,1477,144,576]" pageId="12" pageNumber="13">
<emphasis id="23F4EAF9FF8DFFE9FCC4FF670117699F" box="[825,1046,144,168]" italics="true" pageId="12" pageNumber="13">Differential diagnosis:</emphasis>
From a phylogenetic perspective, the new genus is the sister-lineage of
<taxonomicName id="D6804D68FF8DFFE9FB65FF4701F369F0" authorityName="Conrad" authorityYear="1853" box="[1176,1266,176,199]" class="Bivalvia" family="Unionidae" genus="Parreysia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FB65FF4701F369F0" box="[1176,1266,176,199]" italics="true" pageId="12" pageNumber="13">Parreysia</emphasis>
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but it could be distinguished from the latter genus by having a combination of the following characters: thin, fragile shell (vs. thick, massive), thin, lamella-like pseudocardinal teeth (vs. thick, massive, strongly indented), almost invisible adductor muscle scars (vs. deep, well visible). Among the Asian representatives of the subfamily, the new genus is more similar to
<taxonomicName id="D6804D68FF8DFFE9FB23FE9C005268B4" authorityName="Simpson" authorityYear="1900" box="[1246,1363,363,387]" class="Bivalvia" family="Unionidae" genus="Lamellidens" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FB23FE9C005268B4" box="[1246,1363,363,387]" italics="true" pageId="12" pageNumber="13">Lamellidens</emphasis>
</taxonomicName>
but differs from it by having a much broader and larger umbo, longer and thinner pseudocardinal teeth, and the central position of umbo. Conchologically, the East African species
<taxonomicName id="D6804D68FF8DFFE9FB02FE3E011A6B37" authority="Mandahl-Barth, 1954" authorityName="Mandahl-Barth" authorityYear="1954" class="Bivalvia" family="Unionidae" genus="Coelatura" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="cridlandi">
<emphasis id="23F4EAF9FF8DFFE9FB02FE3E00C368D6" box="[1279,1474,457,481]" italics="true" pageId="12" pageNumber="13">Coelatura cridlandi</emphasis>
Mandahl-Barth, 1954
</taxonomicName>
(
<taxonomicName id="D6804D68FF8DFFE9FBD3FE1E01A86B36" authorityName="Modell" authorityYear="1942" box="[1070,1193,489,513]" class="Bivalvia" family="Unionidae" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="tribe" tribe="Coelaturini">Coelaturini</taxonomicName>
) that is endemic to Lake Victoria looks similar to the new genus but this external appearance most likely reflects convergent evolution.
</paragraph>
<paragraph id="113F36EBFF8DFFE9FCC4FD9606966A85" blockId="12.[825,1476,609,946]" pageId="12" pageNumber="13">
<emphasis id="23F4EAF9FF8DFFE9FCC4FD9606B16B4F" box="[825,944,609,632]" italics="true" pageId="12" pageNumber="13">Description:</emphasis>
Shell thin, very inflated; anterior margin broadly rounded; posterior margin triangular, with narrowly rounded apex; dorsal margin convex, dominated by massive, broad, very prominent umbo; ventral margin broadly convex. Umbo situated in the middle of the shell. Periostracum glossy, light green, usually with dirty yellow patches and bands of long growth lines. Pseudocardinal teeth elongated, narrow, thin, lamella-like, two teeth in right valve and one tooth in left valve. Lateral teeth elongated, lamella-like, one tooth in right valve and two teeth in left valve. Anterior and posterior adductor muscle scars almost invisible.
</paragraph>
</subSubSection>
<subSubSection id="599A6560FF8DFFE9FCC4FC2406C96D7E" pageId="12" pageNumber="13" type="etymology">
<paragraph id="113F36EBFF8DFFE9FCC4FC2406C96D7E" blockId="12.[825,1475,979,1097]" pageId="12" pageNumber="13">
<emphasis id="23F4EAF9FF8DFFE9FCC4FC2406A96ADC" box="[825,936,979,1003]" italics="true" pageId="12" pageNumber="13">Etymology:</emphasis>
The name of this genus is derived from two words: ‘Assam’ (Indian state, in which it ranges) and ‘
<emphasis id="23F4EAF9FF8DFFE9FADFFC04004C6D3D" box="[1314,1357,1011,1034]" italics="true" pageId="12" pageNumber="13">naia</emphasis>
’ (a nymph in Latin, reference to the genus
<taxonomicName id="D6804D68FF8DFFE9FB83FBE501D46D1D" authorityName="Prashad" authorityYear="1918" box="[1150,1237,1042,1066]" class="Bivalvia" family="Unionidae" genus="Indonaia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FB83FBE501D46D1D" box="[1150,1237,1042,1066]" italics="true" pageId="12" pageNumber="13">Indonaia</emphasis>
</taxonomicName>
, in which it was placed traditionally).
</paragraph>
</subSubSection>
<subSubSection id="599A6560FF8DFFE9FCC4FB9C01C16D95" pageId="12" pageNumber="13" type="distribution">
<paragraph id="113F36EBFF8DFFE9FCC4FB9C01C16D95" blockId="12.[825,1474,1131,1186]" pageId="12" pageNumber="13">
<emphasis id="23F4EAF9FF8DFFE9FCC4FB9C06B66DB4" box="[825,951,1131,1155]" italics="true" pageId="12" pageNumber="13">Distribution:</emphasis>
Endemic to the Brahmaputra River basin in
<collectingCountry id="6997767BFF8DFFE9FA70FB9C00C36DB4" box="[1421,1474,1131,1155]" name="India" pageId="12" pageNumber="13">India</collectingCountry>
and to the Surma River in
<collectingCountry id="6997767BFF8DFFE9FBB8FB7D01BD6D95" box="[1093,1212,1162,1186]" name="Bangladesh" pageId="12" pageNumber="13">Bangladesh</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="599A6560FF8DFFE9FCC4FB3400B16C4F" pageId="12" pageNumber="13" type="biology_ecology">
<paragraph id="113F36EBFF8DFFE9FCC4FB3400B16C4F" blockId="12.[824,1475,1219,1400]" pageId="12" pageNumber="13">
<emphasis id="23F4EAF9FF8DFFE9FCC4FB3406976DEC" box="[825,918,1219,1243]" italics="true" pageId="12" pageNumber="13">Habitats:</emphasis>
<materialsCitation id="A1E83CB6FF8DFFE9FC61FB3300AD6C4F" county="Dhansiri River" location="This" municipality="Kaziranga National Park" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Assam" typeStatus="Topotypes">
<typeStatus id="CE3B8849FF8DFFE9FC61FB33010B6DEB" box="[924,1034,1220,1244]" pageId="12" pageNumber="13" type="topotype">Topotypes</typeStatus>
of
<taxonomicName id="D6804D68FF8DFFE9FBD0FB3300936DEC" authority="(Hanley, 1856)" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Hanley" baseAuthorityYear="1856" box="[1069,1426,1219,1243]" class="Bivalvia" family="Unionidae" genus="Assamnaia" higherTaxonomySource="CoL" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="involuta" status="gen. et comb. nov.">
<emphasis id="23F4EAF9FF8DFFE9FBD0FB3301EF6DEC" box="[1069,1262,1219,1243]" italics="true" pageId="12" pageNumber="13">Assamnaia involuta</emphasis>
(
<bibRefCitation id="75114B1AFF8DFFE9FB07FB3400886DEC" author="Hanley S" box="[1274,1417,1219,1243]" pageId="12" pageNumber="13" refId="ref14239" refString="Hanley S. An Illustrated and Descriptive Catalogue of Recent Bivalve Shells. London: Williams and Norgate, 1856." type="book" year="1856">Hanley, 1856</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="38C75782FF8DFFE9FA65FB3306C56DCD" pageId="12" pageNumber="13" rank="species">gen. et comb. nov.</taxonomicNameLabel>
(
<emphasis id="23F4EAF9FF8DFFE9FC2EFB1406E76DCD" box="[979,998,1251,1274]" italics="true" pageId="12" pageNumber="13">N</emphasis>
= 3 sequenced specimens and one dead shell) were collected from an oxbow lake of the
<collectingCounty id="F85E4E67FF8DFFE9FB25FAF5006C6C2D" box="[1240,1389,1282,1306]" pageId="12" pageNumber="13">Dhansiri River</collectingCounty>
, close to the
<collectingMunicipality id="F15BAC91FF8DFFE9FCA3FAD601556C0E" box="[862,1108,1313,1337]" pageId="12" pageNumber="13">Kaziranga National Park</collectingMunicipality>
in
<collectingRegion id="D344F809FF8DFFE9FB8CFAD501B46C0D" box="[1137,1205,1314,1338]" country="India" name="Assam" pageId="12" pageNumber="13">Assam</collectingRegion>
(
<figureCitation id="89BB2A6EFF8DFFE9FB39FAD6000D6C0D" box="[1220,1292,1313,1338]" captionStart="Figure 7" captionStartId="11.[112,177,1302,1326]" captionTargetBox="[114,1458,161,1274]" captionTargetId="figure-269@11.[114,1458,144,1274]" captionTargetPageId="11" captionText="Figure 7. Habitats of little-known freshwater mussel species in Assam.A, collecting locality of Assamnaia involuta gen. et comb.nov.: a shallow oxbow lake with silt and clay bottom substrate, Dhansiri River valley, Golaghat, 26 March 2022. B, collecting locality of Balwantia soleniformis, Indonaia corbis, Pseudoscabies scobina gen. et comb.nov., and Parreysia marcens: Burhidihing River, Dibrugarh, 22 February 2022. The white arrow shows the river cliff, whose bottom portion houses numerous B. soleniformis burrows; the river cliff is mostly composed by firm ground (hard clay). C, close-up view of an aggregation of B. soleniformis burrows in the river cliff,22 February 2022. Living mussels were extracted from submerged burrows.Photos: J. Sonowal and D. Kardong." pageId="12" pageNumber="13">Fig. 7A</figureCitation>
).
<location id="145F6030FF8DFFE9FADCFAD6004C6C0E" LSID="urn:lsid:plazi:treatment:992987FDFF8DFFE9FF36F8FE00936F5A:145F6030FF8DFFE9FADCFAD6004C6C0E" box="[1313,1357,1313,1337]" county="Dhansiri River" municipality="Kaziranga National Park" name="This" pageId="12" pageNumber="13" stateProvince="Assam">This</location>
water body is connected with the main river channel during the rainy season.
<location id="145F6030FF8DFFE9FCC5FA97065E6C4F" LSID="urn:lsid:plazi:treatment:992987FDFF8DFFE9FF36F8FE00936F5A:145F6030FF8DFFE9FCC5FA97065E6C4F" box="[824,863,1376,1400]" county="Dhansiri River" municipality="Kaziranga National Park" name="The" pageId="12" pageNumber="13" stateProvince="Assam">The</location>
mussels were sampled from silt and clay bottom substrate
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="599A6560FF8DFFE9FCC4FA6D00936F5A" pageId="12" pageNumber="13" type="discussion">
<paragraph id="113F36EBFF8DFFE9FCC4FA6D00936F5A" blockId="12.[824,1475,1433,1646]" pageId="12" pageNumber="13">
<emphasis id="23F4EAF9FF8DFFE9FCC4FA6D06A86C86" box="[825,937,1434,1457]" italics="true" pageId="12" pageNumber="13">Comments:</emphasis>
Here, we transfer the nominal species
<taxonomicName id="D6804D68FF8DFFE9FAC9FA6D06826CE6" authority="Hanley" authorityName="Hanley" authorityYear="1856" class="Bivalvia" family="Unionidae" genus="Unio" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="involutus">
<emphasis id="23F4EAF9FF8DFFE9FAC9FA6D00C26C86" box="[1332,1475,1433,1457]" italics="true" pageId="12" pageNumber="13">Unio involutus</emphasis>
Hanley
</taxonomicName>
from
<taxonomicName id="D6804D68FF8DFFE9FC35FA4E01216CE6" authorityName="Prashad" authorityYear="1918" box="[968,1056,1465,1489]" class="Bivalvia" family="Unionidae" genus="Indonaia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FC35FA4E01216CE6" box="[968,1056,1465,1489]" italics="true" pageId="12" pageNumber="13">Indonaia</emphasis>
</taxonomicName>
(the
<taxonomicName id="D6804D68FF8DFFE9FBA3FA4E01AC6CE6" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Hanley" baseAuthorityYear="1856" box="[1118,1197,1465,1489]" class="Bivalvia" family="Unionidae" genus="Assamnaia" higherTaxonomySource="CoL" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="involuta">
<emphasis id="23F4EAF9FF8DFFE9FBA3FA4E01AC6CE6" box="[1118,1197,1465,1489]" italics="true" pageId="12" pageNumber="13">involuta</emphasis>
</taxonomicName>
-group) to
<taxonomicName id="D6804D68FF8DFFE9FADEFA4E008E6CE7" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" box="[1315,1423,1465,1488]" class="Bivalvia" family="Unionidae" genus="Assamnaia" higherTaxonomySource="CoL" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="23F4EAF9FF8DFFE9FADEFA4E008E6CE7" box="[1315,1423,1465,1488]" italics="true" pageId="12" pageNumber="13">Assamnaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38C75782FF8DFFE9FA65FA4E06646CD8" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
and propose
<taxonomicName id="D6804D68FF8DFFE9FBFFFA2F01756CC7" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Hanley" baseAuthorityYear="1856" box="[1026,1140,1496,1520]" class="Bivalvia" family="Unionidae" genus="Assamnaia" higherTaxonomySource="CoL" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="involuta" status="gen. et comb. nov.">
<emphasis id="23F4EAF9FF8DFFE9FBFFFA2F01756CC7" box="[1026,1140,1496,1520]" italics="true" pageId="12" pageNumber="13">A. involuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38C75782FF8DFFE9FB7DFA2F004D6CD8" box="[1152,1356,1496,1520]" pageId="12" pageNumber="13" rank="species">gen. et comb. nov.</taxonomicNameLabel>
The other member of the
<taxonomicName id="D6804D68FF8DFFE9FC33FA00011C6F38" authorityName="Bolotov &amp; Sonowal &amp; Kardong &amp; Pasupuleti &amp; Subba Rao &amp; Unnikrishnan &amp; Gofarov &amp; Kondakov &amp; Konopleva &amp; Lyubas &amp; Vikhrev" authorityYear="2024" baseAuthorityName="Hanley" baseAuthorityYear="1856" box="[974,1053,1527,1551]" class="Bivalvia" family="Unionidae" genus="Assamnaia" higherTaxonomySource="CoL" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="involuta">
<emphasis id="23F4EAF9FF8DFFE9FC33FA00011C6F38" box="[974,1053,1527,1551]" italics="true" pageId="12" pageNumber="13">involuta</emphasis>
</taxonomicName>
-group,
<taxonomicName id="D6804D68FF8DFFE9FB96FA00008C6F38" authority="(Lea, 1831)" baseAuthorityName="Lea" baseAuthorityYear="1831" box="[1131,1421,1527,1551]" class="Bivalvia" family="Unionidae" genus="Indonaia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="olivaria">
<emphasis id="23F4EAF9FF8DFFE9FB96FA0000106F38" box="[1131,1297,1527,1551]" italics="true" pageId="12" pageNumber="13">Indonaia olivaria</emphasis>
(Lea, 1831)
</taxonomicName>
from the Ganges River, needs future study. In the absence of available DNA sequences, we prefer to consider it as an unassigned-togroup (incertae sedis) representative of the genus
<taxonomicName id="D6804D68FF8DFFE9FAC9F9A2008D6F5A" authorityName="Prashad" authorityYear="1918" box="[1332,1420,1621,1645]" class="Bivalvia" family="Unionidae" genus="Indonaia" kingdom="Animalia" order="Unionida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="23F4EAF9FF8DFFE9FAC9F9A2008D6F5A" box="[1332,1420,1621,1645]" italics="true" pageId="12" pageNumber="13">Indonaia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

171
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<mods:title id="7875FBFECF8A1A41C55C1022AED8DB67">Discovery of an endemism hotspot of freshwater mussels (Bivalvia: Unionidae) in Assam, with a description of two new genera</mods:title>
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Genus
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<emphasis id="23F4EAF9FF8DFFE9FEEDFA59048B6CFE" bold="true" box="[272,394,1454,1481]" italics="true" pageId="12" pageNumber="13">Lamellidens</emphasis>
<bibRefCitation id="75114B1AFF8DFFE9FE73FA58072C6CFE" author="Simpson CT" box="[398,557,1455,1481]" pageId="12" pageNumber="13" pagination="501 - 1044" refId="ref15743" refString="Simpson CT. Synopsis of the naiades, or pearly fresh-water mussels. Proceedings of the United States National Museum 1900; 22: 501 - 1044. https: // doi. org / 10.5479 / si. 00963801.22 - 1205.501" type="journal article" year="1900">Simpson, 1900</bibRefCitation>
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(
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Lamarck, 1819
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).
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(
<figureCitation id="89BB2A6EFF8DFFE9FE9AF9E804D86F0E" box="[359,473,1567,1593]" captionStart="Figure 4" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[174,1429,144,1808]" captionTargetId="figure-8@8.[174,1429,144,1808]" captionTargetPageId="8" captionText="Figure 4. Shells of Lamellidentini species from Assam (sequenced specimens; outer view of the left valve and inner view of the right valve).A, Balwantia soleniformis (Benson, 1836), sample DLS-DU BS-BDH-1. B–G, Lamellidens jenkinsianus (Benson, 1862): topotypes LJ-NKM-2 (B), LJ-NKM-3 (C), L5-NKM-1 (D), L4-NKM-1 (E), L5-NKM-4 (F), and L4-NKM-2 (G). Scale bars = 10 mm. The shell vouchers of the new samples are deposited in FBRC ZSI and DLS-DU (Supporting Information, Dataset S1). Photos:J.Sonowal and D. Kardong." pageId="12" pageNumber="13">Figs 4B–G</figureCitation>
,
<figureCitation id="89BB2A6EFF8DFFE9FE1FF9E807226F0E" box="[482,547,1567,1593]" captionStart="Figure 5" captionStartId="9.[113,178,1835,1859]" captionTargetBox="[159,1413,144,1808]" captionTargetId="figure-7@9.[159,1413,144,1808]" captionTargetPageId="9" captionText="Figure 5. Shells of Lamellidens nongyangensis Preston, 1912 (Lamellidentini) (outer view of the left valve and inner view of the right valve). A, holotype ZSI 5068/1 with its label, Lake of No Return (=Nongyang Lake), 27.2192°N, 96.1439°E, Irrawaddy Basin, Myanmar. B–G, sequenced specimens from Assam: samples L1-TLM-3 (B), L2-TLM-2 (C), LL-TNG-4 (D), LN-TNG-3 (E), LU-DHN-2 (F), and LJ-TLM-1 (G). Scale bars = 10 mm. The shell vouchers of the new samples are deposited in FBRC ZSI (Supporting Information, Dataset S1). Photos: N.V.Subba Rao and R. Pasupuleti (A); J. Sonowal and D. Kardong (B–G)." pageId="12" pageNumber="13">5A–G</figureCitation>
)
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<emphasis id="23F4EAF9FF8DFFE9FF7CF99705F06F40" box="[129,241,1632,1655]" italics="true" pageId="12" pageNumber="13">Comments:</emphasis>
This genus contains 13 recent species (
<bibRefCitation id="75114B1AFF8DFFE9FD79F9A805C16FA0" author="Bolotov IN &amp; Pasupuleti R &amp; Subba Rao NV" pageId="12" pageNumber="13" refId="ref13300" refString="Bolotov IN, Pasupuleti R, Subba Rao NV et al. Oriental freshwater mussels arose in East Gondwana and arrived to Asia on the Indian Plate and Burma Terrane. Scientific Reports 2022 a; 12: 1518. https: // doi. org / 10.1038 / s 41598 - 022 - 05257 - 0" type="journal volume" year="2022">
Bolotov
<emphasis id="23F4EAF9FF8DFFE9FD26F997060B6F40" box="[731,778,1631,1655]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2022a
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, b). Two widespread and one putative local species were recorded from
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(
<tableCitation id="5C020350FF8DFFE9FE91F96904B76F81" box="[364,438,1694,1718]" captionStart="Table 1" captionStartId="4.[129,184,143,167]" captionTargetBox="[128,1473,189,1998]" captionText="Table 1. Checklist of freshwater mussel species (Unionidae:Parreysiinae) recorded from the Brahmaputra River basin in Assam, India." pageId="12" pageNumber="13">Table 1</tableCitation>
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(2).
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species:
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; M.
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<emphasis id="1BE5EAAEDA53C230FCAFFA51BDC3AD46" box="[809,925,1529,1550]" italics="true" pageId="40" pageNumber="533">Remarks:</emphasis>
The only Recent species,
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was examined anatomically by
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and proved to be similar to
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H.
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&amp; A.
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<smallCapsWord id="2FC8A060DA53C230FB37FABFBB5CAE67" baselines="1322,1322" box="[1201,1282,1303,1328]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Adams" pageId="40" pageNumber="533">ADAMS</smallCapsWord>
, 1853
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(2).
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<subSubSection id="618B6537DA53C230FCAFFAE9BBCCAE1D" box="[809,1426,1344,1366]" pageId="40" pageNumber="533" type="materials_examined">
<paragraph id="292E36BCDA53C230FCAFFAE9BBCCAE1D" blockId="40.[809,1426,1344,1366]" box="[809,1426,1344,1366]" pageId="40" pageNumber="533">
<emphasis id="1BE5EAAEDA53C230FCAFFAE9BDE9AE1D" box="[809,951,1344,1366]" italics="true" pageId="40" pageNumber="533">
<typeStatus id="F62A881EDA53C230FCAFFAE9BD00AE1E" box="[809,862,1345,1366]" pageId="40" pageNumber="533">Type</typeStatus>
species:
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<taxonomicName id="EE914D3FDA53C230FC3AFAE8BB35AE1D" authority="G. B. Sowerby I, 1833" authorityName="G. B. Sowerby I" authorityYear="1833" box="[956,1387,1344,1366]" class="Gastropoda" family="Ancillariidae" genus="Eburna" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="533" phylum="Mollusca" rank="species" species="australis">
<emphasis id="1BE5EAAEDA53C230FC3AFAE8BA22AE1D" box="[956,1148,1344,1365]" italics="true" pageId="40" pageNumber="533">Eburna australis</emphasis>
G.B. Sowerby I, 1833
</taxonomicName>
; M.
</paragraph>
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</treatment>
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176
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<heading id="726681D0DA53C230FCF3FE9EBB1BAA07" box="[885,1349,310,335]" centered="true" fontSize="9" level="2" pageId="40" pageNumber="533" reason="2">
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<smallCapsWord id="2FC8A060DA53C230FC4AFE9FBA05AA06" baselines="328,329" box="[972,1115,311,334]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Pseudoliva" pageId="40" pageNumber="533">
<emphasis id="1BE5EAAEDA53C230FC4AFE9FBA05AA06" box="[972,1115,311,334]" italics="true" pageId="40" pageNumber="533">PSEUDOLIVA</emphasis>
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<smallCapsWord id="2FC8A060DA53C230FBE4FE9EBA87AA06" baselines="329,329" box="[1122,1241,310,335]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Swainson" pageId="40" pageNumber="533">SWAINSON</smallCapsWord>
, 1840
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(1)
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<subSubSection id="618B6537DA53C230FCAFFEF7BABDA90D" pageId="40" pageNumber="533" type="materials_examined">
<paragraph id="292E36BCDA53C230FCAFFEF7BB5EAADA" blockId="40.[809,1424,350,403]" pageId="40" pageNumber="533">
<emphasis id="1BE5EAAEDA53C230FCAFFEF7BD96AA3C" box="[809,968,351,372]" italics="true" pageId="40" pageNumber="533">
<typeStatus id="F62A881EDA53C230FCAFFEF7BD3FAA3C" box="[809,865,351,372]" pageId="40" pageNumber="533">Type</typeStatus>
species:
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<taxonomicName id="EE914D3FDA53C230FC55FEF7BBCEAA3C" authority="Dillwyn, 1817" authorityName="Dillwyn" authorityYear="1817" box="[979,1424,350,372]" class="Gastropoda" family="Buccinidae" genus="Buccinum" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="533" phylum="Mollusca" rank="species" species="plumbeum">
<emphasis id="1BE5EAAEDA53C230FC55FEF7BA87AA3B" box="[979,1241,350,372]" italics="true" pageId="40" pageNumber="533">Buccinum plumbeum</emphasis>
Dillwyn, 1817
</taxonomicName>
(=
<taxonomicName id="EE914D3FDA53C230FCC1FED6BA9EAADA" authority="Gmelin, 1791" authorityName="Gmelin" authorityYear="1791" box="[839,1216,381,403]" class="Gastropoda" family="Buccinidae" genus="Buccinum" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="533" phylum="Mollusca" rank="species" species="crassum">
<emphasis id="1BE5EAAEDA53C230FCC1FED6BA7EAADB" box="[839,1056,382,403]" italics="true" pageId="40" pageNumber="533">Buccinum crassum</emphasis>
Gmelin, 1791
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); OD.
</paragraph>
<paragraph id="292E36BCDA53C230FCAFFE71BDF9AAB8" blockId="40.[809,935,473,496]" box="[809,935,473,496]" pageId="40" pageNumber="533">
<emphasis id="1BE5EAAEDA53C230FCAFFE71BDF9AAB8" box="[809,935,473,496]" italics="true" pageId="40" pageNumber="533">Synonyms</emphasis>
</paragraph>
<paragraph id="292E36BCDA53C230FCAFFDBABABDA90D" blockId="40.[809,1425,529,735]" pageId="40" pageNumber="533">
<taxonomicName id="EE914D3FDA53C230FCAFFDBABA8AA96E" authority="G. B. Sowerby III, 1890" authorityName="G. B. Sowerby III" authorityYear="1890" box="[809,1236,529,551]" class="Microsporea" genus="Mariona" kingdom="Protozoa" pageId="40" pageNumber="533" phylum="Microsporidia" rank="genus">
<emphasis id="1BE5EAAEDA53C230FCAFFDBABDC8A96F" box="[809,918,530,551]" italics="true" pageId="40" pageNumber="533">Mariona</emphasis>
G.B. Sowerby III, 1890
</taxonomicName>
.
<typeStatus id="F62A881EDA53C230FB62FDBABB7FA96F" box="[1252,1313,530,551]" pageId="40" pageNumber="533">Type</typeStatus>
species:
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<emphasis id="1BE5EAAEDA53C230FCCDFD98BA43A90D" box="[843,1053,560,581]" italics="true" pageId="40" pageNumber="533">Pseudoliva ancilla</emphasis>
Hanley, 1859
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; M.
</paragraph>
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<subSubSection id="618B6537DA53C230FCAFFDE6BAA6AF28" pageId="40" pageNumber="533" type="description">
<paragraph id="292E36BCDA53C230FCAFFDE6BA63A9E9" blockId="40.[809,1425,529,735]" pageId="40" pageNumber="533">
<taxonomicName id="EE914D3FDA53C230FCAFFDE6BA30A92C" authority="Melvill, 1903" authorityName="Melvill" authorityYear="1903" box="[809,1134,590,612]" class="Gastropoda" family="Pseudolividae" genus="Sylvanocochlis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="533" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA53C230FCAFFDE6BD8FA92B" box="[809,977,590,611]" italics="true" pageId="40" pageNumber="533">Sylvanocochlis</emphasis>
Melvill, 1903
</taxonomicName>
.
<typeStatus id="F62A881EDA53C230FBFEFDE7BAEEA92C" box="[1144,1200,591,612]" pageId="40" pageNumber="533">Type</typeStatus>
species:
<taxonomicName id="EE914D3FDA53C230FA93FDE6BA65A9CB" authority="Hanley, 1859" authorityName="Hanley" authorityYear="1859" class="Gastropoda" family="Pseudolividae" genus="Pseudoliva" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="533" phylum="Mollusca" rank="species" species="ancilla">
<emphasis id="1BE5EAAEDA53C230FA93FDE6BDC5A9CA" italics="true" pageId="40" pageNumber="533">Pseudoliva ancilla</emphasis>
Hanley, 1859
</taxonomicName>
; OD; Invalid: junior objective synonym of
<taxonomicName id="EE914D3FDA53C230FC52FD24BA68A9E9" authorityName="G. B. Sowerby III" authorityYear="1890" box="[980,1078,652,673]" class="Microsporea" genus="Mariona" kingdom="Protozoa" pageId="40" pageNumber="533" phylum="Microsporidia" rank="genus">
<emphasis id="1BE5EAAEDA53C230FC52FD24BA68A9E9" box="[980,1078,652,673]" italics="true" pageId="40" pageNumber="533">Mariona</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="292E36BCDA53C230FCAFFD02BAD9A997" blockId="40.[809,1425,529,735]" pageId="40" pageNumber="533">
<taxonomicName id="EE914D3FDA53C230FCAFFD02BA2EA988" authority="P. Fischer, 1884" authorityName="P. Fischer" authorityYear="1884" box="[809,1136,682,704]" class="Gastropoda" family="Pseudolividae" genus="Fulmentum" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="533" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA53C230FCAFFD02BDEFA9F7" box="[809,945,682,703]" italics="true" pageId="40" pageNumber="533">Fulmentum</emphasis>
P. Fischer, 1884
</taxonomicName>
.
<typeStatus id="F62A881EDA53C230FBFDFD03BAEAA988" box="[1147,1204,683,704]" pageId="40" pageNumber="533">Type</typeStatus>
species:
<taxonomicName id="EE914D3FDA53C230FA9AFD03BA03A996" authority="Rang, 1832" authorityName="Rang" authorityYear="1832" class="Gastropoda" family="Buccinidae" genus="Buccinum" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="533" phylum="Mollusca" rank="species" species="sepimentum">
<emphasis id="1BE5EAAEDA53C230FA9AFD03BD8BA997" italics="true" pageId="40" pageNumber="533">Buccinum sepimentum</emphasis>
Rang, 1832
</taxonomicName>
; M.
</paragraph>
<paragraph id="292E36BCDA53C230FCAFFCB0BAA6AF28" blockId="40.[809,1425,792,1120]" pageId="40" pageNumber="533">
<emphasis id="1BE5EAAEDA53C230FCAFFCB0BDFEA865" box="[809,928,792,813]" italics="true" pageId="40" pageNumber="533">Remarks:</emphasis>
Besides size, the distinction between
<taxonomicName id="EE914D3FDA53C230FCAFFC9FBDE2A804" authorityName=", Swainson" authorityYear="1840" box="[809,956,822,844]" class="Gastropoda" family="Pseudolividae" genus="Pseudoliva" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="533" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA53C230FCAFFC9FBDE2A804" box="[809,956,822,844]" italics="true" pageId="40" pageNumber="533">Pseudoliva</emphasis>
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and
<taxonomicName id="EE914D3FDA53C230FB8AFC9FBAF8A804" authorityName="P. Fischer" authorityYear="1884" box="[1036,1190,822,844]" class="Gastropoda" family="Pseudolividae" genus="Fulmentum" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="533" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA53C230FB8AFC9FBAF8A804" box="[1036,1190,822,844]" italics="true" pageId="40" pageNumber="533">Fulmentum</emphasis>
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is in the better development of the plate-like parietal ridge (
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), which divides the aperture of
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have rather similar and characteristic bicuspid lateral teeth, differing in this from all other pseudolivids that have unicuspid lateral teeth. We consider the conchological differences insufficient for separation of two genera.
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.
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species:
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<taxonomicName id="EE914D3FDA53C230FC46F971BB7BADA6" authority="Verco, 1909" authorityName="Verco" authorityYear="1909" box="[960,1317,1753,1775]" class="Gastropoda" family="Olividae" genus="Olivella" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="533" phylum="Mollusca" rank="species" species="adiorygma">
<emphasis id="1BE5EAAEDA53C230FC46F971BAC4ADA6" box="[960,1178,1753,1774]" italics="true" pageId="40" pageNumber="533">Olivella adiorygma</emphasis>
<bibRefCitation id="4D004B4DDA53C230FB27F972BB7BADA6" author="Verco JC" box="[1185,1317,1753,1775]" pageId="40" pageNumber="533" pagination="293 - 342" refId="ref34966" refString="Verco JC. 1909. Notes on south Australian marine mollusca, with descriptions of new species. Part XII. Transactions of the Royal Society of South Australia 33: 293-342." type="journal article" year="1909">Verco, 1909</bibRefCitation>
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; OD.
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The position of this monotypical genus is unclear. Overall, it has an olivoid shell with nonchannelled sutures, a very simplified pication plate and, according to the original illustration of the
<typeStatus id="F62A881EDA52C231FD5FFF6EBD55AB93" box="[729,779,198,219]" pageId="41" pageNumber="534">type</typeStatus>
species (
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: pl. 25, figs 3, 4), it lacks an anterior band. The last whorl is lifting towards the aperture, a character not known in any other
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. The species is known only from empty shells.
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has suggested that this may be a columbellid, a hypothesis that we find convincing.
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[=
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Mawe, 1828
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]; SD,
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.
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Remarks:
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is conchologically rather variable, and a number of (sub)genera have been proposed. Only three species were included in our molecular analysis and, based on this limited dataset, we cannot make any conclusion on the possible subgeneric division of the genus. In the following, we list the classification proposed by
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with some additions and corrections.
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genus:
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<emphasis id="1BE5EAAEDA58C23BFC40F9D4BA17ADD9" box="[966,1097,1660,1681]" italics="true" pageId="35" pageNumber="528">Calyptoliva</emphasis>
<bibRefCitation id="4D004B4DDA58C23BFBC9F9D4BB3DADD9" author="Kantor YI &amp; Bouchet P" box="[1103,1379,1660,1682]" pageId="35" pageNumber="528" pagination="22 - 73" refId="ref32753" refString="Kantor YI, Bouchet P. 2007. Out of Australia: Belloliva (Neogastropoda: Olividae) in the Coral Sea and New Caledonia. American Malacological Bulletin 22: 22-73." type="journal article" year="2007">Kantor &amp; Bouchet, 2007</bibRefCitation>
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<emphasis id="1BE5EAAEDA58C23BFCBDF913BDECAD98" box="[827,946,1723,1744]" italics="true" pageId="35" pageNumber="528">Diagnosis:</emphasis>
Plication plate either smooth or with weak spiral plicae that extend to columella. Primary callus very thin, overlaying suture. Anterior band very thin, hardly discernible, very slightly raised above the shell cloak, uniform. Filament channel absent. Anterior mantle tentacle absent. Posterior mantle lobe moderately or well developed and without anterior mantle tentacle. Head with small, broadly spaced, dorso-ventrally compressed flaps; eyes present or absent. Operculum present. Radula with three teeth per transverse row; central tooth medium broad, tricuspid. Lateral teeth hook-shaped. Stomach with narrow and long posterior mixing area.
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, 1789
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(1)
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<typeStatus id="F62A881EDA59C23AFF17FE13BE98AA98" box="[145,198,443,464]" pageId="34" pageNumber="527">Type</typeStatus>
species:
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<emphasis id="1BE5EAAEDA59C23AFEAFFE12BFEDAA87" box="[297,435,442,463]" italics="true" pageId="34" pageNumber="527">Voluta oliva</emphasis>
Linnaeus, 1758
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; subsequent monotypy by
<bibRefCitation id="4D004B4DDA59C23AFEAFFE71BF8DAAA6" author="Lamarck JB" box="[297,467,473,495]" pageId="34" pageNumber="527" pagination="63 - 91" refId="ref33317" refString="Lamarck JB. 1799. Prodrome d'une nouvelle classification des coquilles; Comprenant une redaction appropriee des caracteres generiques, et l'etablissement d'un grand nombre de genres nouveaux. Memoires de la Societe d'Histoire naturelle de Paris An 7: 63-91." type="journal article" year="1799">Lamarck, 1799</bibRefCitation>
.
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<paragraph id="292E36BCDA59C23AFF17FDB0BF45AFFA" blockId="34.[145,762,536,1539]" pageId="34" pageNumber="527">
<emphasis id="1BE5EAAEDA59C23AFF17FDB0BF5FA965" box="[145,257,536,557]" italics="true" pageId="34" pageNumber="527">Remarks:</emphasis>
The genus
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<emphasis id="1BE5EAAEDA59C23AFE09FDB0BF8EA965" box="[399,464,536,557]" italics="true" pageId="34" pageNumber="527">Oliva</emphasis>
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is rather heterogeneous and a number of subgenera have been proposed, some of them even raised to full genus (
<bibRefCitation id="4D004B4DDA59C23AFD01FDFDBF40A9C1" author="Petuch EJ &amp; Myers RF" pageId="34" pageNumber="527" refId="ref34139" refString="Petuch EJ, Myers RF. 2014. Molluscan communities of the Florida Keys and adjacent areas: their ecology and biodiversity. Boca Raton / London / New York: CERF." type="book" year="2014">Petuch &amp; Myers, 2014</bibRefCitation>
).
<bibRefCitation id="4D004B4DDA59C23AFEB2FDDCBC0DA9C2" author="Petuch EJ &amp; Sargent D" box="[308,595,628,650]" pageId="34" pageNumber="527" refId="ref34172" refString="Petuch EJ, Sargent D. 1986. Atlas of the living olive shells of the world. Charlottesville: SERF." type="book" year="1986">Petuch &amp; Sargent (1986)</bibRefCitation>
recognized 19 subgenera, of which five were monotypic.
<bibRefCitation id="4D004B4DDA59C23AFD0FFD3ABF33A98E" author="Tursch B &amp; Greifeneder D" pageId="34" pageNumber="527" refId="ref34932" refString="Tursch B, Greifeneder D. 2001. Oliva shells. The genus Oliva and the species problem. Italy: L'Informatore Piceno; Costa Rica: Bosque BMT, S. A." type="book" year="2001">Tursch &amp; Greifeneder (2001)</bibRefCitation>
discussed briefly their status but abstained from the use of subgenera, since in many cases they did not find clear conchological distinctions between them. On the contrary,
<bibRefCitation id="4D004B4DDA59C23AFD94FCA6BCA7A86A" author="Hunon C &amp; Horau A &amp; Robain A" box="[530,761,781,803]" pageId="34" pageNumber="527" refId="ref32532" refString="Hunon C, Horau A, Robain A. 2009. Olividae (Mollusca, Gastropoda). A complete survey of Recent species of the genus Oliva. Xenophora: Conchbooks." type="book" year="2009">
Hunon
<emphasis id="1BE5EAAEDA59C23AFDEDFCA6BCF8A86A" box="[619,678,781,803]" italics="true" pageId="34" pageNumber="527">et al.</emphasis>
(2009)
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used as valid most of the subgenera that had been proposed earlier, while still leaving some species in
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<emphasis id="1BE5EAAEDA59C23AFF37FCC1BEACA836" box="[177,242,873,894]" italics="true" pageId="34" pageNumber="527">Oliva</emphasis>
</taxonomicName>
(e.g.
<taxonomicName id="EE914D3FDA59C23AFEB2FCC1BE92A8D6" authority="Greifeneder &amp; Blocher, 1985" authorityName="Greifeneder &amp; Blocher" authorityYear="1985" class="Gastropoda" family="Olividae" genus="Oliva" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="species" species="lacanientai">
<emphasis id="1BE5EAAEDA59C23AFEB2FCC1BF83A836" box="[308,477,873,894]" italics="true" pageId="34" pageNumber="527">O. lacanientai</emphasis>
Greifeneder &amp; Blöcher, 1985
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). Our COI molecular analysis demonstrates that at least some of the subgenera in their current usage are paraphyletic (
<figureCitation id="B1AA2A39DA59C23AFEF5FC6DBFE4A892" box="[371,442,965,987]" captionStart="Figure 2" captionStartId="17.[163,241,1605,1627]" captionTargetBox="[227,1374,196,1564]" captionTargetId="figure-94@17.[226,1378,195,1566]" captionTargetPageId="17" captionText="Figure 2. Phylogenetic tree of Oliva and outgroup olivoidean taxa obtained with Bayesian analysis of COI gene sequences. Only supported nodes with posterior probability values between 0.98 and 1.00 are marked on the tree with circles. The names of the corresponding subgenera (after Petuch &amp; Sargent, 1986; Hunon et al., 2009) are given after the species names. Ac, Acutoliva; An, Annulatoilva; C, Carmione; G, Galeola; Mi, Miniaceoliva; Mu, Musteoliva; O, Omogymna; Ol, Oliva; P, Parvoliva; R, Rufoliva." pageId="34" pageNumber="527">Fig. 2</figureCitation>
– abbreviated subgenera indicated in bold after species names). Nevertheless, we foresee that, with increased taxon sampling in molecular phylogenies, a number of (sub)genera will be found to be justified. However, with the molecular data currently available, we simply list in the following the accepted subgenera, but abstain from using them.
</paragraph>
<paragraph id="292E36BCDA59C23AFF2FFB12BF24AD4A" blockId="34.[145,762,536,1539]" pageId="34" pageNumber="527">
The species delimitation approach would suggest that
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<emphasis id="1BE5EAAEDA59C23AFF4EFB71BF6AAFA7" box="[200,308,1241,1263]" italics="true" pageId="34" pageNumber="527">O. sericea</emphasis>
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and
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<emphasis id="1BE5EAAEDA59C23AFEE8FB71BFA8AFA7" box="[366,502,1241,1263]" italics="true" pageId="34" pageNumber="527">O. miniacea</emphasis>
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would be a single species. However, because they are represented by a single specimen each in our analysis and because the two species are distinct conchologically, we remained conservative and did not change the status of the species. Furthermore,
<taxonomicName id="EE914D3FDA59C23AFEB6FADABFA7AECF" authorityName="Greifeneder &amp; Blocher" authorityYear="1985" box="[304,505,1394,1415]" class="Gastropoda" family="Olividae" genus="Oliva" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="species" species="lacanientai">
<emphasis id="1BE5EAAEDA59C23AFEB6FADABF31AECF" box="[304,367,1394,1415]" italics="true" pageId="34" pageNumber="527">Oliva</emphasis>
‘
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’ shows great intraspecific variability and probably represents the species complex. We abstained from description of new species, since the species level taxonomy was beyond the scope of the current paper.
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<emphasis id="1BE5EAAEDA5BC238FF17F92CBF5BADD1" box="[145,261,1668,1689]" italics="true" pageId="32" pageNumber="525">Diagnosis:</emphasis>
Shell variable, from broadly oval to narrowly fusiform, with high last whorl and diameter of the spire whorls expanding slowly or rapidly. Siphonal canal either absent (in which case the shell is strongly notched) or very short, unnotched. Shell sometimes covered with thick periostracum, but in most cases not and then shell surface glossy. Protoconch multispiral or paucispiral. In all families except
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, anterior end of the shell strongly callused and bearing an anterior band. Callus differently pronounced, sometimes limited to inner apertural lip, but at other times extending on significant part of the last whorl and spire, or even completely covering the shell (some Ancillidae). Radula with five to three teeth per transverse row.
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; OD.
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<emphasis id="1BE5EAAEDA5CC23FFCBDF947BDF7AC4C" box="[827,937,1775,1796]" italics="true" pageId="39" pageNumber="532">Remarks:</emphasis>
The anatomy of
<taxonomicName id="EE914D3FDA5CC23FFBF5F947BB3CAC4D" authorityName="Bouchet &amp; Vermeij" authorityYear="1998" box="[1139,1378,1775,1797]" class="Gastropoda" family="Benthobiidae" genus="Fusulculus" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="species" species="crenatus">
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(own observations) is similar to that of
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<emphasis id="1BE5EAAEDA5CC23FFB63F8A6BB3EAC6B" box="[1253,1376,1806,1827]" italics="true" pageId="39" pageNumber="532">Benthobia</emphasis>
spp.
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, especially the anterior foregut. The mid-oesophagus is glandular, as in
<taxonomicName id="EE914D3FDA53C230FEEEFF6DBF80AB92" baseAuthorityName="Barnard" baseAuthorityYear="1960" box="[360,478,197,218]" class="Gastropoda" family="Benthobiidae" genus="Benthobia" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="533" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA53C230FEEEFF6DBF80AB92" box="[360,478,197,218]" italics="true" pageId="40" pageNumber="533">Benthobia</emphasis>
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, although the glandular epithelium is more pronounced in the former. The synapomorphy is the presence of a single accessory salivary gland, terminating in a very large muscular bulb. Similarly, in both genera, the anteriormost part of the osphradium lacks lamellae on the left side.
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133
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genus:
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<taxonomicName id="EE914D3FDA5CC23FFC40FF44BAEBAA49" authority="Dall, 1889" authorityName="Dall" authorityYear="1889" box="[966,1205,236,258]" class="Gastropoda" family="Benthobiidae" genus="Benthobia" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA5CC23FFC40FF44BA64AA49" box="[966,1082,236,257]" italics="true" pageId="39" pageNumber="532">Benthobia</emphasis>
<bibRefCitation id="4D004B4DDA5CC23FFBC7FF44BAEBAA49" author="Dall WH" box="[1089,1205,236,258]" pageId="39" pageNumber="532" pagination="1 - 492" refId="ref31751" refString="Dall WH. 1889. Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80) .... Report on the Mollusca. Part II. Gastropoda and Scaphopoda. Bulletin of the Museum of Comparative Zoology, Harvard 18: 1-492." type="journal article" year="1889">Dall, 1889</bibRefCitation>
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Shell covered by periostracum, ovoid to biconical, with high or very high last whorl and medium-broad to broadly ovate aperture. Siphonal canal very short and broad, not notched. Anterior band not pronounced. Olivoid groove present, terminating by a short denticle on the lip. Plication plate either weakly pronounced, limited to columella, smooth (
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<emphasis id="1BE5EAAEDA5CC23FFCC2FDAABDE1A95F" box="[836,959,514,535]" italics="true" pageId="39" pageNumber="532">Benthobia</emphasis>
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) or absent (
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). Primary spire callus limited to inner lip, extending up to parietal wall. Spiral sculpture either absent (
<taxonomicName id="EE914D3FDA5CC23FFB78FD97BB26A91C" authorityName="Dall" authorityYear="1889" box="[1278,1400,575,596]" class="Gastropoda" family="Benthobiidae" genus="Benthobia" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA5CC23FFB78FD97BB26A91C" box="[1278,1400,575,596]" italics="true" pageId="39" pageNumber="532">Benthobia</emphasis>
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) or consisting of rather distinct spiral cords (
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). Suture simple, adpressed.
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Foot with narrow propodium, lacking parapodia. Operculum large, with terminal nucleus. Radula with three teeth per transverse row. Central tooth either tricuspid (
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;
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) or multicuspid (
<taxonomicName id="EE914D3FDA5CC23FFBE6FCBEBABFA863" authorityName="Dall" authorityYear="1889" box="[1120,1249,790,811]" class="Gastropoda" family="Benthobiidae" genus="Benthobia" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA5CC23FFBE6FCBEBABFA863" box="[1120,1249,790,811]" italics="true" pageId="39" pageNumber="532">Benthobia</emphasis>
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;
<bibRefCitation id="4D004B4DDA5CC23FFB73FCBEBBC5A863" author="Kantor YI" box="[1269,1435,790,811]" pageId="39" pageNumber="532" pagination="17 - 52" refId="ref32665" refString="Kantor YI. 1991. On the morphology and relationships of some oliviform gastropods. Ruthenica 1: 17-52." type="journal article" year="1991">Kantor, 1991</bibRefCitation>
;
<bibRefCitation id="4D004B4DDA5CC23FFCBDFC9CBDB8A801" author="Simone LRL" box="[827,998,820,842]" pageId="39" pageNumber="532" pagination="245 - 262" refId="ref34450" refString="Simone LRL. 2003. Revision of the genus Benthobia (Caenogastropoda, Pseudolividae). Journal of Molluscan Studies 69: 245-262." type="journal article" year="2003">Simone, 2003</bibRefCitation>
). Lateral teeth unicuspid. Mantle without mantle filament, with weak to moderately pronounced posterior mantle lobe. Head with long cylindrical tentacles, with (
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<emphasis id="1BE5EAAEDA5CC23FFB1BFC38BB76A8ED" box="[1181,1320,912,933]" italics="true" pageId="39" pageNumber="532">Fusulculus</emphasis>
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) or without eyes (
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<emphasis id="1BE5EAAEDA5CC23FFC37FC07BA72A88C" box="[945,1068,943,964]" italics="true" pageId="39" pageNumber="532">Benthobia</emphasis>
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). Rhynchostome shifted to the base of right tentacle. Proboscis very short when retracted, with basal buccal mass and radular odontophoral retractor passing through the nerve ring. Salivary glands acinous. Accessory salivary gland terminating with very large muscular bulb.
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132
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(1)
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; OD.
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; OD.
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In the absence of anatomical data,
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considered
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a separate genus, pointing out its close affinity to
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. The radula of
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is nearly identical to that of
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(
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)
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and there are no good reasons to consider it a separate genus. The mantle lobe (called
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by
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) is moderately pronounced and simple.
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<taxonomicName id="EE914D3FDA5CC23FFED2FCEBBCB8A810" authority="Lamarck, 1801" authorityName="Lamarck" authorityYear="1801" box="[340,742,835,857]" class="Gastropoda" family="Ancillariidae" genus="Eburna" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="species" species="flavida">
<emphasis id="1BE5EAAEDA5CC23FFED2FCEBBC44A810" box="[340,538,835,856]" italics="true" pageId="39" pageNumber="532">Eburna flavida</emphasis>
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Linnaeus, 1758
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We examined the anatomy of
<taxonomicName id="EE914D3FDA5CC23FFD2AFC08BF56A89C" baseAuthorityName="Bernardi" baseAuthorityYear="1859" class="Gastropoda" family="Ancillariidae" genus="Eburna" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="species" species="lienardi">
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(unpublished data). It is rather typical ancillariid and the genus is attributed to the family on the basis of morphology and radula (
<figureCitation id="B1AA2A39DA5CC23FFDE9FC54BC96AF59" box="[623,712,1020,1042]" captionStart="Figure 12" captionStartId="32.[144,224,1259,1281]" captionTargetBox="[352,1216,194,1220]" captionTargetId="figure-202@32.[352,1216,194,1220]" captionTargetPageId="32" captionText="Figure 12. Radulae of Pseudolividae (A–B) and Ancillariidae (C–I). (A) Pseudoliva sepimenta, MNHN IM-2013-52005. (B) Pseudoliva crassa (Gmelin, 1791), Baia do Baba, Moçâmedes, Angola, 6–20 m, coll. S. Gofas, SL 39.5 mm, AL 31.5 mm. (C) Entomoliva mirabilis New Caledonia: Canal de la Havannah, EXBODI, st. DW3787, 22°13′S, 167°06′E, 223–249 m, SL 11.2 mm. (D) Amalda contusa, MNHN IM-2009-22264. (E, F) Amalda cf. hilgendorfi richeri, MNHN IM-2007-43661. (F) enlarged lateral tooth to show bifurcation on the tip. (G) Ancillina cf. sumatrana, Madagascar, between Majunga and Cap Saint-André, MIRIKY, st. CP3278, 15°24′S, 45°56′E, 750–780 m. (H) Turrancilla sp. 2, MNHN IM-2007-31947. (I) Eburna lienardi (Bernardi, 1859), MZSP 40140, Proc. Areia Branca, RN, Brazil, 04°45′S, 36°58′W, 9 m." pageId="39" pageNumber="532">Fig. 12I</figureCitation>
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(1)
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; OD.
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species:
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†
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,
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); OD
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In addition to several fossil species from
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and recently described ones from the Paris and Aquitanian basins, the Recent species
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Strebel, 1908
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. Neither the anatomy nor the radula of
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Synonym:
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Anatomy and radular characters remain unknown; therefore, the validity of the genus is uncertain.
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<taxonomicName id="EE914D3FDA5DC23EFC1BFB68BB10AF90" ID-CoL="7NNZB" authority="H. ADAMS &amp; A. ADAMS, 1853" authorityName="H. ADAMS &amp; A. ADAMS" authorityYear="1853" box="[925,1358,1215,1240]" class="Gastropoda" family="Ancillariidae" genus="Amalda" kingdom="Animalia" order="Neogastropoda" pageId="38" pageNumber="531" phylum="Mollusca" rank="genus">
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<emphasis id="1BE5EAAEDA5DC23EFC1BFB68BDA2AF9F" box="[925,1020,1216,1239]" italics="true" pageId="38" pageNumber="531">AMALDA</emphasis>
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H.
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&amp; A.
<bibRefCitation id="4D004B4DDA5DC23EFB3EFB17BB10AF90" author="Adams H &amp; Adams A" box="[1208,1358,1215,1240]" pageId="38" pageNumber="531" refId="ref31088" refString="Adams H, Adams A. 1853. The genera of Recent Mollusca arranged according to their organisation, Vol. 1. London: Van Voorst." type="book" year="1853">
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, 1853
</bibRefCitation>
</taxonomicName>
(1)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="618B6537DA5DC23EFCAFFB41BB73AC2C" pageId="38" pageNumber="531" type="materials_examined">
<paragraph id="292E36BCDA5DC23EFCAFFB41BA7DAE54" blockId="38.[809,1424,1256,1308]" pageId="38" pageNumber="531">
<emphasis id="1BE5EAAEDA5DC23EFCAFFB41BDE0AFB5" box="[809,958,1256,1278]" italics="true" pageId="38" pageNumber="531">
<typeStatus id="F62A881EDA5DC23EFCAFFB41BD01AFB6" box="[809,863,1257,1278]" pageId="38" pageNumber="531">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="EE914D3FDA5DC23EFC41FB40BBD2AFB6" authority="Swainson, 1825" authorityName="Swainson" authorityYear="1825" box="[967,1420,1256,1278]" class="Gastropoda" family="Ancillariidae" genus="Ancillaria" kingdom="Animalia" order="Neogastropoda" pageId="38" pageNumber="531" phylum="Mollusca" rank="species" species="tankervillii">
<emphasis id="1BE5EAAEDA5DC23EFC41FB40BA94AFB5" box="[967,1226,1256,1277]" italics="true" pageId="38" pageNumber="531">Ancillaria tankervillii</emphasis>
Swainson, 1825
</taxonomicName>
; SD,
<bibRefCitation id="4D004B4DDA5DC23EFCD1FAAFBA43AE54" author="Cossmann M" box="[855,1053,1287,1308]" pageId="38" pageNumber="531" refId="ref31588" refString="Cossmann M. 1899. Essais de paleoconchologie comparee, Vol. 3. Paris: The author and Comptoir Geologique." type="book" year="1899">Cossmann (1899)</bibRefCitation>
.
</paragraph>
<paragraph id="292E36BCDA5DC23EFCAFFAEDBB73AC2C" blockId="38.[809,1425,1349,1892]" pageId="38" pageNumber="531">
<emphasis id="1BE5EAAEDA5DC23EFCAFFAEDBDC9AE12" box="[809,919,1349,1370]" italics="true" pageId="38" pageNumber="531">Remarks:</emphasis>
The species of
<taxonomicName id="EE914D3FDA5DC23EFBC9FAEDBAF2AE12" authorityName="H. ADAMS &amp; A. ADAMS" authorityYear="1853" box="[1103,1196,1349,1370]" class="Gastropoda" family="Ancillariidae" genus="Amalda" kingdom="Animalia" order="Neogastropoda" pageId="38" pageNumber="531" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA5DC23EFBC9FAEDBAF2AE12" box="[1103,1196,1349,1370]" italics="true" pageId="38" pageNumber="531">Amalda</emphasis>
</taxonomicName>
together present a wide range of shell forms and it is not surprising that several genus group names have been proposed. Our analysis covers very little of the general diversity of the group, but at least three subgenera are present in our dataset:
<taxonomicName id="EE914D3FDA5DC23EFC67FA76BA61AEBB" authorityName="H. ADAMS &amp; A. ADAMS" authorityYear="1853" box="[993,1087,1502,1523]" class="Gastropoda" family="Ancillariidae" genus="Amalda" kingdom="Animalia" order="Neogastropoda" pageId="38" pageNumber="531" phylum="Mollusca" rank="genus" sensu="stricto">
<emphasis id="1BE5EAAEDA5DC23EFC67FA76BA61AEBB" box="[993,1087,1502,1523]" italics="true" pageId="38" pageNumber="531">Amalda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00D657D5DA5DC23EFBC1FA77BA33AEBC" box="[1095,1133,1503,1524]" pageId="38" pageNumber="531" sensu="stricto">s.s.</taxonomicNameLabel>
,
<taxonomicName id="EE914D3FDA5DC23EFBFAFA77BAACAEBC" authorityName="P. Fischer" authorityYear="1883" box="[1148,1266,1503,1524]" class="Gastropoda" family="Ancillariidae" genus="Baryspira" kingdom="Animalia" order="Neogastropoda" pageId="38" pageNumber="531" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA5DC23EFBFAFA77BAACAEBC" box="[1148,1266,1503,1524]" italics="true" pageId="38" pageNumber="531">Baryspira</emphasis>
</taxonomicName>
[represented in our analysis by
<taxonomicName id="EE914D3FDA5DC23EFB94FA55BBD7AD5B" authority="(Reeve, 1864)" baseAuthorityName="Reeve" baseAuthorityYear="1864" box="[1042,1417,1533,1555]" class="Gastropoda" family="Ancillariidae" genus="Amalda" kingdom="Animalia" order="Neogastropoda" pageId="38" pageNumber="531" phylum="Mollusca" rank="species" species="contusa">
<emphasis id="1BE5EAAEDA5DC23EFB94FA55BA87AD5B" box="[1042,1241,1533,1555]" italics="true" pageId="38" pageNumber="531">Amalda contusa</emphasis>
(Reeve, 1864)
</taxonomicName>
] and
<taxonomicName id="EE914D3FDA5DC23EFCDBF9B4BD87AD79" authorityName="Cossmann" authorityYear="1899" box="[861,985,1564,1585]" class="Gastropoda" family="Ancillariidae" genus="Alocospira" kingdom="Animalia" order="Neogastropoda" pageId="38" pageNumber="531" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA5DC23EFCDBF9B4BD87AD79" box="[861,985,1564,1585]" italics="true" pageId="38" pageNumber="531">Alocospira</emphasis>
</taxonomicName>
(represented by
<taxonomicName id="EE914D3FDA5DC23EFB22F9B4BA18AD18" authority="Kilburn &amp; Bouchet, 1988" authorityName="Kilburn &amp; Bouchet" authorityYear="1988" class="Gastropoda" family="Ancillariidae" genus="Amalda" kingdom="Animalia" order="Neogastropoda" pageId="38" pageNumber="531" phylum="Mollusca" rank="species" species="bellonarum">
<emphasis id="1BE5EAAEDA5DC23EFB22F9B4BBCFAD79" box="[1188,1425,1564,1585]" italics="true" pageId="38" pageNumber="531">Amalda bellonarum</emphasis>
Kilburn &amp; Bouchet, 1988
</taxonomicName>
). Although the support of the genus itself is significant (
<quantity id="EE699B59DA5DC23EFBCCF9F1BA2EAD27" box="[1098,1136,1625,1647]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="38" pageNumber="531" unit="in" value="1.0">1 in</quantity>
BA), the genetic distances within its constituents are rather small, compared to, for example, the genus
<taxonomicName id="EE914D3FDA5DC23EFBB0F93EBAD7ADE3" authorityName="Lamarck" authorityYear="1799" box="[1078,1161,1686,1707]" class="Gastropoda" family="Ancillariidae" genus="Ancilla" kingdom="Animalia" order="Neogastropoda" pageId="38" pageNumber="531" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA5DC23EFBB0F93EBAD7ADE3" box="[1078,1161,1686,1707]" italics="true" pageId="38" pageNumber="531">Ancilla</emphasis>
</taxonomicName>
. Nevertheless, at least
<taxonomicName id="EE914D3FDA5DC23EFCAFF91DBDFDAD82" authorityName="Cossmann" authorityYear="1899" box="[809,931,1717,1738]" class="Gastropoda" family="Ancillariidae" genus="Alocospira" kingdom="Animalia" order="Neogastropoda" pageId="38" pageNumber="531" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA5DC23EFCAFF91DBDFDAD82" box="[809,931,1717,1738]" italics="true" pageId="38" pageNumber="531">Alocospira</emphasis>
</taxonomicName>
constitutes a rather long branch, and it is possible that future molecular studies will reveal a more complex and robust structuring necessitating the recognition of (sub)genera. At present we abstain from using any subgenera and provide a list of names with Recent
<typeStatus id="F62A881EDA5DC23EFC3EF8E7BDB7AC2C" box="[952,1001,1871,1892]" pageId="38" pageNumber="531">type</typeStatus>
species for future reference.
</paragraph>
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<heading id="726681D0DA5EC23DFE86FB34BCF3AFFD" box="[256,685,1180,1205]" centered="true" fontSize="9" level="2" pageId="37" pageNumber="530" reason="2">
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<taxonomicName id="EE914D3FDA5EC23DFED1FB35BCD9AFFD" ID-CoL="7P64D" authority="OLSSON, 1956" authorityName="OLSSON" authorityYear="1956" box="[343,647,1180,1205]" class="Gastropoda" family="Bellolividae" genus="Jaspidella" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="genus">
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<emphasis id="1BE5EAAEDA5EC23DFED1FB35BFBEAFFB" box="[343,480,1181,1204]" italics="true" pageId="37" pageNumber="530">JASPIDELLA</emphasis>
</smallCapsWord>
<bibRefCitation id="4D004B4DDA5EC23DFE61FB34BCD9AFFD" author="Olsson AA" box="[487,647,1180,1205]" pageId="37" pageNumber="530" pagination="155 - 225" refId="ref33854" refString="Olsson AA. 1956. Studies of the genus Olivella. Proceedings of the Academy of Natural Sciences of Philadelphia 108: 155-225." type="journal article" year="1956">
<smallCapsWord id="2FC8A060DA5EC23DFE61FB34BC1CAFFB" baselines="1199,1198" box="[487,578,1180,1205]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Olsson" pageId="37" pageNumber="530">OLSSON</smallCapsWord>
, 1956
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(2)
</heading>
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<subSubSection id="618B6537DA5EC23DFF25FB6DBF20AEB8" pageId="37" pageNumber="530" type="materials_examined">
<paragraph id="292E36BCDA5EC23DFF25FB6DBC9CAF91" blockId="37.[163,706,1220,1242]" box="[163,706,1220,1242]" pageId="37" pageNumber="530">
<emphasis id="1BE5EAAEDA5EC23DFF25FB6DBF6DAF92" box="[163,307,1221,1242]" italics="true" pageId="37" pageNumber="530">
<typeStatus id="F62A881EDA5EC23DFF25FB6DBE86AF92" box="[163,216,1221,1242]" pageId="37" pageNumber="530">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="EE914D3FDA5EC23DFEBFFB6CBCD7AF91" authority="Gmelin, 1791" authorityName="Gmelin" authorityYear="1791" box="[313,649,1220,1242]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="species" species="jaspidea">
<emphasis id="1BE5EAAEDA5EC23DFEBFFB6CBFB7AF91" box="[313,489,1220,1241]" italics="true" pageId="37" pageNumber="530">Voluta jaspidea</emphasis>
Gmelin, 1791
</taxonomicName>
; OD.
</paragraph>
<paragraph id="292E36BCDA5EC23DFF25FAABBF20AEB8" blockId="37.[163,779,1283,1520]" pageId="37" pageNumber="530">
<emphasis id="1BE5EAAEDA5EC23DFF25FAABBF50AE50" box="[163,270,1283,1304]" italics="true" pageId="37" pageNumber="530">Remarks:</emphasis>
The genus was established in the subfamily
<taxonomicName id="EE914D3FDA5EC23DFF25FA8ABF57AE70" box="[163,265,1314,1336]" class="Gastropoda" family="Olividae" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="subFamily" subFamily="Olivinae">Olivinae</taxonomicName>
on the basis of the tricuspidate rachidian tooth. The
<typeStatus id="F62A881EDA5EC23DFE9AFAE9BF13AE1E" box="[284,333,1345,1366]" pageId="37" pageNumber="530">type</typeStatus>
species from the Caribbean has a shell similar to that of
<taxonomicName id="EE914D3FDA5EC23DFEE9FAF7BF88AE3C" authorityName="Peile" authorityYear="1922" box="[367,470,1375,1396]" class="Gastropoda" family="Bellolividae" genus="Belloliva" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA5EC23DFEE9FAF7BF88AE3C" box="[367,470,1375,1396]" italics="true" pageId="37" pageNumber="530">Belloliva</emphasis>
</taxonomicName>
, as well as the operculum, also characteristic for this family. No material was available to us and we attribute
<taxonomicName id="EE914D3FDA5EC23DFD0EFA34BD54AEF9" authorityName="OLSSON" authorityYear="1956" box="[648,778,1436,1457]" class="Gastropoda" family="Bellolividae" genus="Jaspidella" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA5EC23DFD0EFA34BD54AEF9" box="[648,778,1436,1457]" italics="true" pageId="37" pageNumber="530">Jaspidella</emphasis>
</taxonomicName>
conditionally to
<taxonomicName id="EE914D3FDA5EC23DFED9FA13BFB6AE99" authorityName="Kantor &amp; Fedosov &amp; Puillandre &amp; Bonillo &amp; Bouchet" authorityYear="2017" box="[351,488,1467,1489]" class="Gastropoda" family="Bellolividae" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="family">Bellolividae</taxonomicName>
on the basis of shell and radula similarities.
</paragraph>
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<paragraph id="292E36BCDA5EC23DFF70FF6CBCE9AB95" blockId="37.[246,695,196,221]" box="[246,695,196,221]" pageId="37" pageNumber="530">
<heading id="726681D0DA5EC23DFF70FF6CBCE9AB95" box="[246,695,196,221]" centered="true" fontSize="9" level="2" pageId="37" pageNumber="530" reason="2">
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<taxonomicName id="EE914D3FDA5EC23DFEC8FF6CBCCFAB95" ID-CoL="7PB78" authority="THIELE, 1929" authorityName="Thiele" authorityYear="1929" box="[334,657,196,221]" class="Gastropoda" family="Bellolividae" genus="Olivellopsis" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="genus">
<smallCapsWord id="2FC8A060DA5EC23DFEC8FF6CBFB1AB93" baselines="213,214" box="[334,495,196,219]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Olivellopsis" pageId="37" pageNumber="530">
<emphasis id="1BE5EAAEDA5EC23DFEC8FF6CBFB1AB93" box="[334,495,196,219]" italics="true" pageId="37" pageNumber="530">OLIVELLOPSIS</emphasis>
</smallCapsWord>
<bibRefCitation id="4D004B4DDA5EC23DFE71FF6CBCCFAB95" author="Thiele J" box="[503,657,196,221]" pageId="37" pageNumber="530" refId="ref34844" refString="Thiele J. 1929. Handbuch der systematischen Weichtierkunde. Jena: Gustave Fischer." type="book" year="1929">
<smallCapsWord id="2FC8A060DA5EC23DFE71FF6CBC12AB93" baselines="215,214" box="[503,588,196,221]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Thiele" pageId="37" pageNumber="530">THIELE</smallCapsWord>
, 1929
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</taxonomicName>
(1)
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<subSubSection id="618B6537DA5EC23DFF25FF45BC0FAF17" pageId="37" pageNumber="530" type="materials_examined">
<paragraph id="292E36BCDA5EC23DFF25FF45BF5BAA68" blockId="37.[163,777,236,289]" pageId="37" pageNumber="530">
<emphasis id="1BE5EAAEDA5EC23DFF25FF45BF1BAA4A" box="[163,325,237,258]" italics="true" pageId="37" pageNumber="530">
<typeStatus id="F62A881EDA5EC23DFF25FF45BE82AA4A" box="[163,220,237,258]" pageId="37" pageNumber="530">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="EE914D3FDA5EC23DFED7FF44BE82AA69" authority="Pease 1868" authorityName="Pease" authorityYear="1868" class="Gastropoda" family="Olividae" genus="Olivella" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="species" species="simplex" subGenus="Callianax">
<emphasis id="1BE5EAAEDA5EC23DFED7FF44BCE6AA49" box="[337,696,236,257]" italics="true" pageId="37" pageNumber="530">Olivella (Callianax) simplex</emphasis>
Pease 1868
</taxonomicName>
; M.
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Synonym:
<taxonomicName id="EE914D3FDA5EC23DFE9CFE92BD59AA18" authority="Sterba &amp; Lorenz, 2005" authorityName="Sterba &amp; Lorenz" authorityYear="2005" box="[282,775,314,336]" class="Gastropoda" family="Olividae" genus="Olivella" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="subGenus" subGenus="Janaoliva">
<emphasis id="1BE5EAAEDA5EC23DFE9CFE92BF2AAA07" box="[282,372,314,335]" italics="true" pageId="37" pageNumber="530">Olivella</emphasis>
(
<emphasis id="1BE5EAAEDA5EC23DFE05FE92BFAAAA07" box="[387,500,314,335]" italics="true" pageId="37" pageNumber="530">Janaoliva</emphasis>
)
<bibRefCitation id="4D004B4DDA5EC23DFD82FE92BD59AA18" author="Sterba GHW &amp; Lorenz F" box="[516,775,314,336]" pageId="37" pageNumber="530" pagination="43 - 46" refId="ref34622" refString="Sterba GHW, Lorenz F. 2005. Olivella (Janaoliva) amoni, a new subgenus and species from the Bismark Archipelago and Indonesia (Mollusca: Gastropoda: Olivellidae). Visaya 1: 43-46." type="journal article" year="2005">Sterba &amp; Lorenz, 2005</bibRefCitation>
</taxonomicName>
,
<emphasis id="1BE5EAAEDA5EC23DFF43FEF0BF66AA25" bold="true" box="[197,312,344,365]" pageId="37" pageNumber="530">syn. nov.</emphasis>
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species:
<taxonomicName id="EE914D3FDA5EC23DFE60FEF0BF04AAC5" authority="Sterba &amp; Lorenz, 2005" authorityName="Sterba &amp; Lorenz" authorityYear="2005" class="Gastropoda" family="Olividae" genus="Olivella" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="species" species="amoni">
<emphasis id="1BE5EAAEDA5EC23DFE60FEF0BCC8AA26" box="[486,662,344,366]" italics="true" pageId="37" pageNumber="530">Olivella amoni</emphasis>
<bibRefCitation id="4D004B4DDA5EC23DFD19FEF0BF04AAC5" author="Sterba GHW &amp; Lorenz F" pageId="37" pageNumber="530" pagination="43 - 46" refId="ref34622" refString="Sterba GHW, Lorenz F. 2005. Olivella (Janaoliva) amoni, a new subgenus and species from the Bismark Archipelago and Indonesia (Mollusca: Gastropoda: Olivellidae). Visaya 1: 43-46." type="journal article" year="2005">Sterba &amp; Lorenz, 2005</bibRefCitation>
</taxonomicName>
; OD.
</paragraph>
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<emphasis id="1BE5EAAEDA5EC23DFF25FE6EBF52AA93" box="[163,268,454,475]" italics="true" pageId="37" pageNumber="530">Remarks:</emphasis>
When describing
<emphasis id="1BE5EAAEDA5EC23DFE53FE6EBC1AAA93" box="[469,580,454,475]" italics="true" pageId="37" pageNumber="530">Janaoliva</emphasis>
,
<bibRefCitation id="4D004B4DDA5EC23DFDD7FE6EBEB3AAB2" author="Sterba GHW &amp; Lorenz F" pageId="37" pageNumber="530" pagination="43 - 46" refId="ref34622" refString="Sterba GHW, Lorenz F. 2005. Olivella (Janaoliva) amoni, a new subgenus and species from the Bismark Archipelago and Indonesia (Mollusca: Gastropoda: Olivellidae). Visaya 1: 43-46." type="journal article" year="2005">Sterba &amp; Lorenz (2005)</bibRefCitation>
compared it with
<taxonomicName id="EE914D3FDA5EC23DFE4CFE4CBC76AAB1" authorityName="Swainson" authorityYear="1831" box="[458,552,484,505]" class="Gastropoda" family="Olividae" genus="Olivella" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA5EC23DFE4CFE4CBC76AAB1" box="[458,552,484,505]" italics="true" pageId="37" pageNumber="530">Olivella</emphasis>
</taxonomicName>
, correctly pointing out the very simplified structure of the plication plate, which differs from other subgenera of
<taxonomicName id="EE914D3FDA5EC23DFDC6FD8ABCC6A97F" authorityName="Swainson" authorityYear="1831" box="[576,664,546,567]" class="Gastropoda" family="Olividae" genus="Olivella" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA5EC23DFDC6FD8ABCC6A97F" box="[576,664,546,567]" italics="true" pageId="37" pageNumber="530">Olivella</emphasis>
</taxonomicName>
.
<bibRefCitation id="4D004B4DDA5EC23DFD24FD8ABF15A91D" author="Kantor YI &amp; Bouchet P" pageId="37" pageNumber="530" pagination="22 - 73" refId="ref32753" refString="Kantor YI, Bouchet P. 2007. Out of Australia: Belloliva (Neogastropoda: Olividae) in the Coral Sea and New Caledonia. American Malacological Bulletin 22: 22-73." type="journal article" year="2007">Kantor &amp; Bouchet (2007)</bibRefCitation>
overlooked the description of
<emphasis id="1BE5EAAEDA5EC23DFD1DFDE8BD55A91D" box="[667,779,576,597]" italics="true" pageId="37" pageNumber="530">Janaoliva</emphasis>
and thus did not compare
<taxonomicName id="EE914D3FDA5EC23DFE4EFDF7BC30A93D" authorityName="Sterba &amp; Lorenz" authorityYear="2005" box="[456,622,607,629]" class="Gastropoda" family="Olividae" genus="Olivella" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="species" species="amoni">
<emphasis id="1BE5EAAEDA5EC23DFE4EFDF7BC30A93D" box="[456,622,607,629]" italics="true" pageId="37" pageNumber="530">Olivella amoni</emphasis>
</taxonomicName>
with the very similar species
<taxonomicName id="EE914D3FDA5EC23DFECCFDD6BC7BA9DB" baseAuthorityName="Pease" baseAuthorityYear="1868" box="[330,549,638,659]" class="Gastropoda" family="Bellolividae" genus="Olivellopsis" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="1BE5EAAEDA5EC23DFECCFDD6BC7BA9DB" box="[330,549,638,659]" italics="true" pageId="37" pageNumber="530">Olivellopsis simplex</emphasis>
</taxonomicName>
, which they included in
<emphasis id="1BE5EAAEDA5EC23DFF44FD34BF71A9F9" box="[194,303,668,689]" italics="true" pageId="37" pageNumber="530">
<taxonomicName id="EE914D3FDA5EC23DFF44FD34BF72A9F9" authorityName="Peile" authorityYear="1922" box="[194,300,668,689]" class="Gastropoda" family="Bellolividae" genus="Belloliva" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="genus">Belloliva</taxonomicName>
.
</emphasis>
The very long branches of both
<taxonomicName id="EE914D3FDA5EC23DFD23FD34BD55A9F9" authorityName="Peile" authorityYear="1922" box="[677,779,668,689]" class="Gastropoda" family="Bellolividae" genus="Belloliva" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA5EC23DFD23FD34BD55A9F9" box="[677,779,668,689]" italics="true" pageId="37" pageNumber="530">Belloliva</emphasis>
</taxonomicName>
and
<taxonomicName id="EE914D3FDA5EC23DFF5EFD13BF3CA998" authorityName="Thiele" authorityYear="1929" box="[216,354,699,720]" class="Gastropoda" family="Bellolividae" genus="Olivellopsis" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="genus">
<emphasis id="1BE5EAAEDA5EC23DFF5EFD13BF3CA998" box="[216,354,699,720]" italics="true" pageId="37" pageNumber="530">Olivellopsis</emphasis>
</taxonomicName>
suggest that they are two separate genera. In the vicinity of Kavieng (New Ireland), the
<typeStatus id="F62A881EDA5EC23DFF25FD51BE88A846" box="[163,214,761,782]" pageId="37" pageNumber="530">type</typeStatus>
locality of
<taxonomicName id="EE914D3FDA5EC23DFED8FD51BF94A846" authorityName="Sterba &amp; Lorenz" authorityYear="2005" box="[350,458,761,782]" class="Gastropoda" family="Olividae" genus="Olivella" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="species" species="amoni">
<emphasis id="1BE5EAAEDA5EC23DFED8FD51BF94A846" box="[350,458,761,782]" italics="true" pageId="37" pageNumber="530">O. amoni</emphasis>
</taxonomicName>
, another similar, although molecularly different species was recently found. Their radulae are similar (
<figureCitation id="B1AA2A39DA5EC23DFE27FC9EBC58A803" box="[417,518,822,844]" captionStart="Figure11" captionStartId="31.[162,239,1260,1282]" captionTargetBox="[370,1234,195,1221]" captionTargetId="figure-258@31.[370,1234,195,1221]" captionTargetPageId="31" captionText="Figure11. RadulaeofBellolividae(A–C)andOlividae(E–H).(A,B) Bellolivaexquisita, NorthernNewCaledonia,MUSORSTOM 4, st. DW 184, 19°04′S, 163°27′E, 260 m (after Kantor &amp; Bouchet, 2007). (C) Olivellopsis amoni, Papua-New Guinea, Kavieng Lagoon NE corner of Nusa I., KAVIENG 2014, st. KS19, 02°34′S, 150°47,1′E, 10 m. (D) Calyptoliva bbugeae, holotype, MNHN IM-2007–38919. (E) Agaronia acuminata, MNHN IM-2013–52002, central tooth. (F) Oliva rufofulgurata Schepman, 1904, MNHN IM-2007–36132. (G) Olivella minuta, MNHN IM-2009–24350. (H) Olivella exilis, MNHN IM-2009–24367." pageId="37" pageNumber="530">Fig. 11C</figureCitation>
) and are close to the radulae of
<taxonomicName id="EE914D3FDA5EC23DFEA6FCFDBFC6A821" baseAuthorityName="Pease" baseAuthorityYear="1868" box="[288,408,852,874]" class="Gastropoda" family="Bellolividae" genus="Olivellopsis" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="1BE5EAAEDA5EC23DFEA6FCFDBFC6A821" box="[288,408,852,874]" italics="true" pageId="37" pageNumber="530">O. simplex</emphasis>
</taxonomicName>
from
<collectingCountry id="5186762CDA5EC23DFE5BFCFDBCD3A822" box="[477,653,852,874]" name="New Caledonia" pageId="37" pageNumber="530">New Caledonia</collectingCountry>
(
<bibRefCitation id="4D004B4DDA5EC23DFD1BFCFDBF1EA8C0" author="Kantor YI &amp; Bouchet P" pageId="37" pageNumber="530" pagination="22 - 73" refId="ref32753" refString="Kantor YI, Bouchet P. 2007. Out of Australia: Belloliva (Neogastropoda: Olividae) in the Coral Sea and New Caledonia. American Malacological Bulletin 22: 22-73." type="journal article" year="2007">Kantor &amp; Bouchet, 2007</bibRefCitation>
:
<figureCitation id="B1AA2A39DA5EC23DFECAFCDBBF96A8C0" box="[332,456,883,905]" captionStart="Figure 12" captionStartId="32.[144,224,1259,1281]" captionTargetBox="[352,1216,194,1220]" captionTargetId="figure-202@32.[352,1216,194,1220]" captionTargetPageId="32" captionText="Figure 12. Radulae of Pseudolividae (A–B) and Ancillariidae (C–I). (A) Pseudoliva sepimenta, MNHN IM-2013-52005. (B) Pseudoliva crassa (Gmelin, 1791), Baia do Baba, Moçâmedes, Angola, 6–20 m, coll. S. Gofas, SL 39.5 mm, AL 31.5 mm. (C) Entomoliva mirabilis New Caledonia: Canal de la Havannah, EXBODI, st. DW3787, 22°13′S, 167°06′E, 223–249 m, SL 11.2 mm. (D) Amalda contusa, MNHN IM-2009-22264. (E, F) Amalda cf. hilgendorfi richeri, MNHN IM-2007-43661. (F) enlarged lateral tooth to show bifurcation on the tip. (G) Ancillina cf. sumatrana, Madagascar, between Majunga and Cap Saint-André, MIRIKY, st. CP3278, 15°24′S, 45°56′E, 750–780 m. (H) Turrancilla sp. 2, MNHN IM-2007-31947. (I) Eburna lienardi (Bernardi, 1859), MZSP 40140, Proc. Areia Branca, RN, Brazil, 04°45′S, 36°58′W, 9 m." pageId="37" pageNumber="530">Fig. 12A–D</figureCitation>
). In the light of the new data indicating the presence of a species complex,
<taxonomicName id="EE914D3FDA5EC23DFD11FC3ABD54A8EF" baseAuthorityName="Pease" baseAuthorityYear="1868" box="[663,778,914,935]" class="Gastropoda" family="Bellolividae" genus="Olivellopsis" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="1BE5EAAEDA5EC23DFD11FC3ABD54A8EF" box="[663,778,914,935]" italics="true" pageId="37" pageNumber="530">O. simplex</emphasis>
</taxonomicName>
from
<collectingCountry id="5186762CDA5EC23DFF59FC19BFD5A88E" box="[223,395,944,966]" name="New Caledonia" pageId="37" pageNumber="530">New Caledonia</collectingCountry>
may be specifically different from
<taxonomicName id="EE914D3FDA5EC23DFF25FC67BF47A8AC" baseAuthorityName="Pease" baseAuthorityYear="1868" box="[163,281,975,996]" class="Gastropoda" family="Bellolividae" genus="Olivellopsis" kingdom="Animalia" order="Neogastropoda" pageId="37" pageNumber="530" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="1BE5EAAEDA5EC23DFF25FC67BF47A8AC" box="[163,281,975,996]" italics="true" pageId="37" pageNumber="530">O. simplex</emphasis>
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, the
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locality of which is the ‘Paumotus Islands’ (Tuamotu Archipelago),
<collectingCountry id="5186762CDA5EC23DFD91FC45BC80AF4B" box="[535,734,1005,1027]" name="French Polynesia" pageId="37" pageNumber="530">French Polynesia</collectingCountry>
. We also found very similar shells off
<collectingCountry id="5186762CDA5EC23DFDBDFBA4BC92AF6A" box="[571,716,1036,1058]" name="Madagascar" pageId="37" pageNumber="530">Madagascar</collectingCountry>
. It is possible that a whole radiation of
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will be uncovered by molecular phylogenetics.
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106
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species:
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Angas, 1877
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119
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Shell glossy, lacking periostracum, narrowly fusiform, with high to very high last whorl and narrow aperture tapering adapically. Siphonal canal absent, anterior end of shell distinctly notched. Anterior shell end with poorly defined anterior band, very slightly raised above the shell cloak. Plication plate limited to columella, with spiral plicae (
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) or not (
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). Primary spire callus absent. Narrow filament channel present, opened on all spire whorls.
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Foot with well-developed crescent-shaped propodium, subdivided by a longitudinal cleft on the dorsal side, and parapodia partially embracing the shell. Operculum present. Radula with three teeth per transverse row. Central tooth with three large, closely spaced cusps and, in
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<emphasis id="1BE5EAAEDA5FC23CFC04FA1DBDB7AE82" box="[898,1001,1461,1482]" italics="true" pageId="36" pageNumber="529">Belloliva</emphasis>
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, with distinct small cusps adjoining the major lateral cusps (
<figureCitation id="B1AA2A39DA5FC23CFBC0FA7CBAF8AEA1" box="[1094,1190,1492,1514]" captionStart="Figure11" captionStartId="31.[162,239,1260,1282]" captionTargetBox="[370,1234,195,1221]" captionTargetId="figure-258@31.[370,1234,195,1221]" captionTargetPageId="31" captionText="Figure11. RadulaeofBellolividae(A–C)andOlividae(E–H).(A,B) Bellolivaexquisita, NorthernNewCaledonia,MUSORSTOM 4, st. DW 184, 19°04′S, 163°27′E, 260 m (after Kantor &amp; Bouchet, 2007). (C) Olivellopsis amoni, Papua-New Guinea, Kavieng Lagoon NE corner of Nusa I., KAVIENG 2014, st. KS19, 02°34′S, 150°47,1′E, 10 m. (D) Calyptoliva bbugeae, holotype, MNHN IM-2007–38919. (E) Agaronia acuminata, MNHN IM-2013–52002, central tooth. (F) Oliva rufofulgurata Schepman, 1904, MNHN IM-2007–36132. (G) Olivella minuta, MNHN IM-2009–24350. (H) Olivella exilis, MNHN IM-2009–24367." pageId="36" pageNumber="529">Fig. 11B</figureCitation>
), that are indistinct or absent in
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<emphasis id="1BE5EAAEDA5FC23CFC3FFA5ABA60AD4F" box="[953,1086,1522,1543]" italics="true" pageId="36" pageNumber="529">Olivellopsis</emphasis>
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(
<figureCitation id="B1AA2A39DA5FC23CFBCBFA5ABAF2AD40" box="[1101,1196,1522,1544]" captionStart="Figure11" captionStartId="31.[162,239,1260,1282]" captionTargetBox="[370,1234,195,1221]" captionTargetId="figure-258@31.[370,1234,195,1221]" captionTargetPageId="31" captionText="Figure11. RadulaeofBellolividae(A–C)andOlividae(E–H).(A,B) Bellolivaexquisita, NorthernNewCaledonia,MUSORSTOM 4, st. DW 184, 19°04′S, 163°27′E, 260 m (after Kantor &amp; Bouchet, 2007). (C) Olivellopsis amoni, Papua-New Guinea, Kavieng Lagoon NE corner of Nusa I., KAVIENG 2014, st. KS19, 02°34′S, 150°47,1′E, 10 m. (D) Calyptoliva bbugeae, holotype, MNHN IM-2007–38919. (E) Agaronia acuminata, MNHN IM-2013–52002, central tooth. (F) Oliva rufofulgurata Schepman, 1904, MNHN IM-2007–36132. (G) Olivella minuta, MNHN IM-2009–24350. (H) Olivella exilis, MNHN IM-2009–24367." pageId="36" pageNumber="529">Fig. 11C</figureCitation>
). Mantle with mantle filament, a small anterior mantle lobe, and without anterior mantle tentacle. Head formed by dorso-ventrally compressed flaps with or without eyes, at least in
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on very short and narrow tentacle.
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104
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(1)
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; OD.
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<emphasis id="1BE5EAAEDA5FC23CFF17FDFDBEA1A922" box="[145,255,597,618]" italics="true" pageId="36" pageNumber="529">Remarks:</emphasis>
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Kantor
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is the only sequenced species and forms a long branch and the sister group cannot be recognized in our analysis. However, the genus is confidently placed in
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and differs from all other representatives of the family by the absence of a channelled suture and, correspondingly, by the absence of the mantle filament. The suture is overlaid by a very narrow primary spire callus.
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386
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<mods:role id="8DF32BE7F60FF2198672B2F0BDE31D48">
<mods:roleTerm id="43926F51A4C23608BDD5E76A25405E21">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1A2050065E2EFD765C541BABC1505DD0">Roisin, Yves</mods:namePart>
<mods:nameIdentifier id="CED43A4076187A67D2C410AB0A565F1B" type="ORCID">0000-0001-6635-3552</mods:nameIdentifier>
<mods:nameIdentifier id="D0F0BEB6D29BB0752F02971B2B5967BA" type="email">oisin@ulb.be</mods:nameIdentifier>
</mods:name>
<mods:name id="64EC94FDD932FE300046A31B91B05161" type="personal">
<mods:role id="5D7737319F7B5365F20E15FDCD047ADE">
<mods:roleTerm id="32F00E16898B31E59FE35E5FB517E7D6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BAE17385C40DED8CC73F2108A4116FCF">Wang, Menglin</mods:namePart>
<mods:affiliation id="3B701FD743A8B691FAE65232AA9F0E7E">Essig Museum of Entomology, 1101 VLSB # 4780, Berkeley, California, 94720, USA</mods:affiliation>
</mods:name>
<mods:name id="A9E394F9DD303CFD359FFE2AFE9B9CA0" type="personal">
<mods:role id="4AB1D8CEAF0E35D9A99D66A265A28D93">
<mods:roleTerm id="896701614E91F548E8891729D9A0E6D4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="69B74908D19E258A322EA6F04D6EC66E">Bourguignon, Thomas</mods:namePart>
<mods:affiliation id="F66742D2A27BEF12AF04A46E60EA9DB2">Okinawa Institute of Science &amp; Technology Graduate University, 1919 - 1 Tancha, Onna-son, 904 - 0495 Okinawa, Japan</mods:affiliation>
</mods:name>
<mods:name id="3E0DB1EE48B07A1CFB4275A2A50FDDBF" type="personal">
<mods:role id="C0B578E8F7338E85AE1ACDF0C9F333BB">
<mods:roleTerm id="EDBCF2FAAD5DD73C0C591F42ED1879DD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D6C8851E283F6F77419C1A749638BCAF">Hellemans, Simon</mods:namePart>
<mods:affiliation id="707DC765F1F7F2126CEF8C71BF9EDF89">Okinawa Institute of Science &amp; Technology Graduate University, 1919 - 1 Tancha, Onna-son, 904 - 0495 Okinawa, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6BE9E25692934E42C5D0201CE53D29F4">text</mods:typeOfResource>
<mods:relatedItem id="BEB7F07D3D0AEB60BBF883800197490D" type="host">
<mods:titleInfo id="54136107C3A1C58B025E8D6AE8F86F93">
<mods:title id="B8F35387E686053B757297CABDEC36FE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="042F1917CE896E83D83C5C32BAFE2F68">
<mods:date id="18EC37840A199AAFFDEC7D8AB52487AE">2025</mods:date>
<mods:detail id="3D612E664BC803F1929BDAFA0FD95E89" type="pubDate">
<mods:number id="5B79984C16D26F18855286495CE3D5C7">2025-02-04</mods:number>
</mods:detail>
<mods:detail id="C3F2A458DB5B0B708C9B7E3B4B2550D7" type="volume">
<mods:number id="F7DF4EE8C166B54E5E241BAED9286060">5583</mods:number>
</mods:detail>
<mods:detail id="CFC2324386B523F9F943766DF913BC8F" type="issue">
<mods:number id="529B424D61166FCFD363E508F557584F">3</mods:number>
</mods:detail>
<mods:extent id="3099817E841B72EE922D8865BA1C6BC0" unit="page">
<mods:start id="19B748DBBE2C79CD7B2966B84ADBD7CB">567</mods:start>
<mods:end id="AFE86811C8B69206FC1A38607D1CA917">578</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="0ADB2E1947C7C0B3F73C7F8687821209">
<mods:url id="1E0F2CCBEE6DD5A1FD00841BA76B195E">https://doi.org/10.11646/zootaxa.5583.3.9</mods:url>
</mods:location>
<mods:classification id="AAA084701DFD310F10DFFA68B0597809">journal article</mods:classification>
<mods:identifier id="4E4C1E6EE09BB7ED07FAF14987BF7AE8" type="DOI">10.11646/zootaxa.5583.3.9</mods:identifier>
<mods:identifier id="9C4D08321DB05EAAA86DB95AC2225D78" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4B5B49F70AD0002114500279EFE6C2C8" type="Zenodo-Dep">14804861</mods:identifier>
<mods:identifier id="B3CB201EE2A6245EB984BF72A7863C16" type="ZooBank">5CB47A59-4C83-4515-B6CF-2CE580884161</mods:identifier>
</mods:mods>
<treatment id="EF27E65EFFF4FFC949E7FB78FBCA0D1C" LSID="urn:lsid:plazi:treatment:EF27E65EFFF4FFC949E7FB78FBCA0D1C" httpUri="http://treatment.plazi.org/id/EF27E65EFFF4FFC949E7FB78FBCA0D1C" lastPageId="7" lastPageNumber="574" pageId="5" pageNumber="572">
<subSubSection id="2F9404C3FFF4FFCB49E7FB78FD7F0E96" box="[151,742,1195,1222]" pageId="5" pageNumber="572" type="nomenclature">
<paragraph id="67315748FFF4FFCB49E7FB78FD7F0E96" blockId="5.[151,742,1195,1222]" box="[151,742,1195,1222]" pageId="5" pageNumber="572">
<heading id="3C79E024FFF4FFCB49E7FB78FD7F0E96" bold="true" box="[151,742,1195,1222]" fontSize="11" level="1" pageId="5" pageNumber="572" reason="1">
<emphasis id="55FA8B5AFFF4FFCB49E7FB78FD7F0E96" bold="true" box="[151,742,1195,1222]" pageId="5" pageNumber="572">
<taxonomicName id="A08E2CCBFFF4FFCB49E7FB78FE770E95" authorityName="Scheffrahn &amp; Roisin &amp; Wang &amp; Bourguignon &amp; Hellemans" authorityYear="2025" box="[151,494,1195,1222]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="monomorphus" status="sp. nov.">
<emphasis id="55FA8B5AFFF4FFCB49E7FB78FE770E95" bold="true" box="[151,494,1195,1222]" italics="true" pageId="5" pageNumber="572">Obtusitermes monomorphus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EC93621FFF4FFCB4886FB7FFDCA0E95" box="[502,595,1196,1222]" pageId="5" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
Scheffrahn
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2F9404C3FFF4FFCB49E7FB27FAD80FCD" pageId="5" pageNumber="572" type="materials_examined">
<paragraph id="67315748FFF4FFCB49E7FB27FBCD0F5D" blockId="5.[151,1437,1267,2014]" box="[151,1108,1267,1294]" pageId="5" pageNumber="572">
<materialsCitation id="D7E65D15FFF4FFCB49E7FB27FBCD0F5D" box="[151,1108,1267,1294]" collectionCode="UFTC" county="Soldier" location="Termite Collection" municipality="University" pageId="5" pageNumber="572" specimenCode="PN153" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
<emphasis id="55FA8B5AFFF4FFCB49E7FB27FE920F5D" bold="true" box="[151,267,1268,1294]" pageId="5" pageNumber="572">
<typeStatus id="B835E9EAFFF4FFCB49E7FB27FE9F0F5D" box="[151,262,1268,1294]" pageId="5" pageNumber="572" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="8E502FC4FFF4FFCB4862FB27FEFA0F5D" box="[274,355,1268,1294]" pageId="5" pageNumber="572">Soldier</collectingCounty>
from
<collectingMunicipality id="8755CD32FFF4FFCB48D7FB27FD840F5D" box="[423,541,1268,1294]" pageId="5" pageNumber="572">University</collectingMunicipality>
of
<collectingRegion id="A54A99AAFFF4FFCB4B33FB27FD0D0F5D" box="[579,660,1268,1294]" country="United States of America" name="Florida" pageId="5" pageNumber="572">Florida</collectingRegion>
<location id="62510193FFF4FFCB4BEAFB27FCF40F5D" LSID="urn:lsid:plazi:treatment:EF27E65EFFF4FFC949E7FB78FBCA0D1C:62510193FFF4FFCB4BEAFB27FCF40F5D" box="[666,877,1268,1294]" county="Soldier" municipality="University" name="Termite Collection" pageId="5" pageNumber="572" stateProvince="Florida">Termite Collection</location>
(
<collectionCode id="019FCF8DFFF4FFCB4A0BFB27FC5F0F5D" box="[891,966,1268,1294]" pageId="5" pageNumber="572">UFTC</collectionCode>
) no.
<specimenCode id="3728FF33FFF4FFCB4A8FFB20FBC90F5D" box="[1023,1104,1267,1294]" collectionCode="PN" pageId="5" pageNumber="572">PN153</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="67315748FFF4FFCB49B7FACBFBFE0F61" blockId="5.[151,1437,1267,2014]" box="[199,1127,1303,1330]" pageId="5" pageNumber="572">
<emphasis id="55FA8B5AFFF4FFCB49B7FACBFEFF0F62" bold="true" box="[199,358,1303,1330]" pageId="5" pageNumber="572">Type locality.</emphasis>
<materialsCitation id="D7E65D15FFF4FFCB481CFACBFBF80F61" box="[364,1121,1303,1330]" collectorName="El Cope Natl. Park" country="Panama" elevation="841" latitude="8.679" location="Cocle" longLatPrecision="7" longitude="-80.5972" pageId="5" pageNumber="572" specimenCount="1" stateProvince="Cocle">
<collectingCountry id="1F9917D8FFF4FFCB481CFACBFE7E0F61" box="[364,487,1304,1330]" name="Panama" pageId="5" pageNumber="572">PANAMA</collectingCountry>
:
<collectingRegion id="A54A99AAFFF4FFCB4882FACBFDAF0F61" box="[498,566,1304,1330]" country="Panama" name="Cocle" pageId="5" pageNumber="572">Coclé</collectingRegion>
,
<collectorName id="CA7B329EFFF4FFCB4B31FACBFC810F61" box="[577,792,1304,1330]" pageId="5" pageNumber="572">El Cope Natl. Park</collectorName>
;
<geoCoordinate id="02BA318FFFF4FFCB4A54FACBFCED0F61" box="[804,884,1304,1330]" degrees="8.6790" direction="north" orientation="latitude" pageId="5" pageNumber="572" precision="5" value="8.679">8.6790</geoCoordinate>
,
<geoCoordinate id="02BA318FFFF4FFCB4A0FFAC4FC7F0F61" box="[895,998,1303,1330]" degrees="80.5972" direction="west" orientation="longitude" pageId="5" pageNumber="572" precision="5" value="-80.5972">-80.5972</geoCoordinate>
;
<elevation id="ECA3B07BFFF4FFCB4A82FACBFBF80F61" box="[1010,1121,1304,1330]" metricMagnitude="2" metricUnit="m" metricValue="8.41" pageId="5" pageNumber="572" unit="m" value="841.0">
<quantity id="A076FAADFFF4FFCB4A82FACBFBA00F61" box="[1010,1081,1304,1330]" metricMagnitude="2" metricUnit="m" metricValue="8.41" pageId="5" pageNumber="572" unit="m" value="841.0">841 m</quantity>
asl
</elevation>
</materialsCitation>
;
</paragraph>
<paragraph id="67315748FFF4FFCB49B7FAEFFAAE0F05" blockId="5.[151,1437,1267,2014]" box="[199,1335,1340,1367]" pageId="5" pageNumber="572">
<materialsCitation id="D7E65D15FFF4FFCB49B7FAEFFAAD0F05" box="[199,1332,1340,1367]" collectingDate="2005-05-30" collectionCode="UFTC" collectorName="J Chase" pageId="5" pageNumber="572" specimenCode="PN153" specimenCount="1">
<date id="13307188FFF4FFCB49B7FAEFFEFD0F04" box="[199,356,1340,1367]" pageId="5" pageNumber="572" value="2005-05-30">
<collectingDate id="03748860FFF4FFCB49B7FAEFFEFD0F04" box="[199,356,1340,1367]" pageId="5" pageNumber="572" value="2005-05-30">30-May-2005</collectingDate>
</date>
;
<collectorName id="CA7B329EFFF4FFCB4800FAEFFE5F0F05" box="[368,454,1340,1366]" pageId="5" pageNumber="572">J Chase</collectorName>
coll.;
<collectionCode id="019FCF8DFFF4FFCB4B7DFAEFFDCC0F05" box="[525,597,1340,1366]" pageId="5" pageNumber="572">UFTC</collectionCode>
<specimenCode id="3728FF33FFF4FFCB4B2CFAEFFD350F04" box="[604,684,1340,1367]" collectionCode="PN" pageId="5" pageNumber="572">PN153</specimenCode>
; many soldiers and workers taken from under horse dung
</materialsCitation>
.
</paragraph>
<paragraph id="67315748FFF4FFCB49B7FAB3FAD80FCD" blockId="5.[151,1437,1267,2014]" pageId="5" pageNumber="572">
<materialsCitation id="D7E65D15FFF4FFCB49B7FAB3FDFF0FCD" collectingDate="2010-06-05" collectionCode="R, UFTC" collectorName="Kuna Rd. Stop &amp; Scheffrahn" country="Panama" elevation="360" latitude="9.3261" location="Rd. Stop" longLatPrecision="7" longitude="78.9992" municipality="Kuna Rd." pageId="5" pageNumber="572" specimenCode="PN1415" specimenCount="1" stateProvince="Kuna Yala" typeStatus="paratype">
<emphasis id="55FA8B5AFFF4FFCB49B7FAB3FEDF0F29" bold="true" box="[199,326,1376,1402]" pageId="5" pageNumber="572">
<typeStatus id="B835E9EAFFF4FFCB49B7FAB3FED80F29" box="[199,321,1376,1402]" pageId="5" pageNumber="572" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="1F9917D8FFF4FFCB483EFAB3FE500F29" box="[334,457,1376,1402]" name="Panama" pageId="5" pageNumber="572">PANAMA</collectingCountry>
:
<collectingRegion id="A54A99AAFFF4FFCB48A5FAB3FDD30F29" box="[469,586,1376,1402]" country="Panama" name="Kuna Yala" pageId="5" pageNumber="572">Guna Yala</collectingRegion>
(formerly San Blas),
<collectorName id="CA7B329EFFF4FFCB4A32FAB3FC700F29" box="[834,1001,1376,1402]" pageId="5" pageNumber="572">
<collectingMunicipality id="8755CD32FFF4FFCB4A32FAB3FC370F29" box="[834,942,1376,1402]" pageId="5" pageNumber="572">
Kuna
<location id="62510193FFF4FFCB4AF6FAB3FC370F29" LSID="urn:lsid:plazi:treatment:EF27E65EFFF4FFC949E7FB78FBCA0D1C:62510193FFF4FFCB4AF6FAB3FC370F29" box="[902,942,1376,1402]" country="Panama" latitude="9.3261" longLatPrecision="7" longitude="78.9992" municipality="Kuna Rd." name="Rd. Stop" pageId="5" pageNumber="572" stateProvince="Kuna Yala">Rd.</location>
</collectingMunicipality>
Stop
</collectorName>
4;
<geoCoordinate id="02BA318FFFF4FFCB4D7FFAB3FBC60F29" box="[1039,1119,1376,1402]" degrees="9.3261" direction="north" orientation="latitude" pageId="5" pageNumber="572" precision="5" value="9.3261">9.3261</geoCoordinate>
,
<geoCoordinate id="02BA318FFFF4FFCB4D1BFAB3FB530F29" box="[1131,1226,1376,1402]" degrees="78.9992" direction="east" orientation="longitude" pageId="5" pageNumber="572" precision="5" value="78.9992">78.9992</geoCoordinate>
;
<elevation id="ECA3B07BFFF4FFCB4DA6FAB3FAD10F29" box="[1238,1352,1376,1402]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="572" unit="m" value="360.0">
<quantity id="A076FAADFFF4FFCB4DA6FAB3FA870F29" box="[1238,1310,1376,1402]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="572" unit="m" value="360.0">360 m</quantity>
asl
</elevation>
;
<date id="13307188FFF4FFCB4C25FA8CFF4A0FCD" pageId="5" pageNumber="572" value="2010-06-05">
<collectingDate id="03748860FFF4FFCB4C25FA8CFF4A0FCD" pageId="5" pageNumber="572" value="2010-06-05">5-Jun- 2010</collectingDate>
</date>
;
<collectionCode id="019FCF8DFFF4FFCB49AEFA57FF680FCD" box="[222,241,1412,1438]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="572">R</collectionCode>
<collectorName id="CA7B329EFFF4FFCB4988FA57FEEB0FCD" box="[248,370,1412,1438]" pageId="5" pageNumber="572">Scheffrahn</collectorName>
coll.;
<collectionCode id="019FCF8DFFF4FFCB48C9FA57FD980FCD" box="[441,513,1412,1438]" pageId="5" pageNumber="572">UFTC</collectionCode>
<specimenCode id="3728FF33FFF4FFCB4B77FA50FDFF0FCD" box="[519,614,1411,1438]" collectionCode="PN" pageId="5" pageNumber="572">PN1415</specimenCode>
</materialsCitation>
; many soldiers and workers taken from under a decaying branch.
</paragraph>
</subSubSection>
<subSubSection id="2F9404C3FFF4FFC949B7FA7BFBCA0D1C" lastPageId="7" lastPageNumber="574" pageId="5" pageNumber="572" type="etymology">
<paragraph id="67315748FFF4FFCB49B7FA7BFB5A0F91" blockId="5.[151,1437,1267,2014]" box="[199,1219,1448,1474]" pageId="5" pageNumber="572">
<emphasis id="55FA8B5AFFF4FFCB49B7FA7BFED70F91" bold="true" box="[199,334,1448,1474]" pageId="5" pageNumber="572">Etymology.</emphasis>
Named for the monomorphic soldier caste which is exclusive to
<taxonomicName id="A08E2CCBFFF4FFCB4D56FA7BFB220F91" authorityName="Snyder" authorityYear="1924" box="[1062,1211,1448,1474]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="572" phylum="Arthropoda" rank="genus">
<emphasis id="55FA8B5AFFF4FFCB4D56FA7BFB220F91" box="[1062,1211,1448,1474]" italics="true" pageId="5" pageNumber="572">Obtusitermes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="67315748FFF4FFCB49B7FA1FFE010FB5" blockId="5.[151,1437,1267,2014]" box="[199,408,1484,1510]" pageId="5" pageNumber="572">
<emphasis id="55FA8B5AFFF4FFCB49B7FA1FFE8A0FB5" bold="true" box="[199,275,1484,1510]" pageId="5" pageNumber="572">Imago</emphasis>
. Unknown.
</paragraph>
<paragraph id="67315748FFF4FFCB49B7FA3CFD660CC9" blockId="5.[151,1437,1267,2014]" pageId="5" pageNumber="572">
<emphasis id="55FA8B5AFFF4FFCB49B7FA3CFE840C5A" bold="true" box="[199,285,1519,1545]" pageId="5" pageNumber="572">Soldier</emphasis>
(
<figureCitation id="FFB54BCDFFF4FFCB485EFA23FEEC0C59" box="[302,373,1520,1546]" captionStart="FIGURE 2" captionStartId="2.[151,250,681,705]" captionTargetBox="[151,1435,179,655]" captionTargetId="figure-20@2.[151,1435,179,655]" captionTargetPageId="2" captionText="FIGURE 2. Holotype soldier of Obtusitermes monomorphus sp. nov. A. dorsal and B. lateral view of head capsule." figureDoi="http://doi.org/10.5281/zenodo.14804867" httpUri="https://zenodo.org/record/14804867/files/figure.png" pageId="5" pageNumber="572">Fig. 2</figureCitation>
,
<figureCitation id="FFB54BCDFFF4FFCB48F4FA23FE0D0C59" box="[388,404,1520,1546]" captionStart="FIGURE 6" captionStartId="5.[151,250,657,681]" captionText="FIGURE 6. Live habitus of Obtusitermes monomorphus sp. nov. (PN1415)." pageId="5" pageNumber="572">6</figureCitation>
;
<tableCitation id="2A0C62F3FFF4FFCB48D2FA23FE6F0C59" box="[418,502,1520,1546]" captionStart="TABLE 1" captionStartId="1.[151,239,1529,1553]" captionText="TABLE 1. Measurements (mm) of Obtusitermes monomorphus sp. nov. soldiers (n=5) from two colonies." pageId="5" pageNumber="572">Table 1</tableCitation>
). Monomorphic, small size. Head capsule reddish brown, nasus darker. Antennal articles and palps reddish brown. Head capsule pyriform from above, constricted at anterior one-third. Lateral view of head capsule convex behind antennal sockets. Vertex with two long setae at summit of convexity and four long setae at posterior half of vertex. Nasus broadly conical, upturned about 10°. Antennae with 11 articles; antennal article formula 2&lt;3&gt;4≤5. Mandibles with small points.
</paragraph>
<paragraph id="67315748FFF4FFCB49B7F970FF690D8D" blockId="5.[151,1437,1267,2014]" pageId="5" pageNumber="572">
<emphasis id="55FA8B5AFFF4FFCB49B7F970FEFF0CEE" bold="true" box="[199,358,1699,1725]" pageId="5" pageNumber="572">Comparisons</emphasis>
. The soldier of
<taxonomicName id="A08E2CCBFFF4FFCB4B6EF977FD7B0CED" authorityName="Scheffrahn &amp; Roisin &amp; Wang &amp; Bourguignon &amp; Hellemans" authorityYear="2025" box="[542,738,1700,1726]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="monomorphus" status="sp. nov.">
<emphasis id="55FA8B5AFFF4FFCB4B6EF977FD7B0CED" box="[542,738,1700,1726]" italics="true" pageId="5" pageNumber="572">O. monomorphus</emphasis>
</taxonomicName>
<emphasis id="55FA8B5AFFF4FFCB4B9BF977FCDF0CED" bold="true" box="[747,838,1700,1726]" pageId="5" pageNumber="572">
<taxonomicNameLabel id="4EC93621FFF4FFCB4B9BF977FCDF0CED" box="[747,838,1700,1726]" pageId="5" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the major soldiers of
<taxonomicName id="A08E2CCBFFF4FFCB4DA3F977FAC70CEE" baseAuthorityName="Snyder" baseAuthorityYear="1924" box="[1235,1374,1700,1726]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="panamae">
<emphasis id="55FA8B5AFFF4FFCB4DA3F977FAC70CEE" box="[1235,1374,1700,1726]" italics="true" pageId="5" pageNumber="572">O. panamae</emphasis>
</taxonomicName>
(
<figureCitation id="FFB54BCDFFF4FFCB4C1FF977FF450CB1" captionStart="FIGURE 8" captionStartId="7.[151,250,1162,1186]" captionTargetBox="[151,1436,179,1136]" captionTargetId="figure-18@7.[151,1436,179,1136]" captionTargetPageId="7" captionText="FIGURE 8. Soldiers of Obtusitermes panamae (Snyder, 1925) (UFTC PN1076). A. dorsal and B. lateral view of major soldier head capsule. C. dorsal and D. lateral view of minor soldier head capsule." figureDoi="http://doi.org/10.5281/zenodo.14804877" httpUri="https://zenodo.org/record/14804877/files/figure.png" pageId="5" pageNumber="572">Fig. 8A, B</figureCitation>
) and
<taxonomicName id="A08E2CCBFFF4FFCB486FF91BFE260CB1" authorityName="Cuezzo and Cancello" authorityYear="2009" box="[287,447,1736,1762]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="formosulus">
<emphasis id="55FA8B5AFFF4FFCB486FF91BFE260CB1" box="[287,447,1736,1762]" italics="true" pageId="5" pageNumber="572">O. formosulus</emphasis>
</taxonomicName>
(
<figureCitation id="FFB54BCDFFF4FFCB48A0F91BFDD20CB1" box="[464,587,1736,1762]" captionStart="FIGURE 9" captionStartId="8.[151,250,1159,1183]" captionTargetBox="[151,1436,179,1134]" captionTargetId="figure-20@8.[151,1436,179,1134]" captionTargetPageId="8" captionText="FIGURE 9. Soldiers of Obtusitermes formosulus Cuezzo and Cancello, 2009 (UFTC VZ1171). A. dorsal and B. lateral view of major soldier head capsule. C. dorsal and D. lateral view of minor soldier head capsule." figureDoi="http://doi.org/10.5281/zenodo.14804879" httpUri="https://zenodo.org/record/14804879/files/figure.png" pageId="5" pageNumber="572">Fig. 9A, B</figureCitation>
) in that
<taxonomicName id="A08E2CCBFFF4FFCB4BDEF91BFCEB0CB1" authorityName="Scheffrahn &amp; Roisin &amp; Wang &amp; Bourguignon &amp; Hellemans" authorityYear="2025" box="[686,882,1736,1762]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="monomorphus" status="sp. nov.">
<emphasis id="55FA8B5AFFF4FFCB4BDEF91BFCEB0CB1" box="[686,882,1736,1762]" italics="true" pageId="5" pageNumber="572">O. monomorphus</emphasis>
</taxonomicName>
<emphasis id="55FA8B5AFFF4FFCB4A0BF91BFC4F0CB1" bold="true" box="[891,982,1736,1762]" pageId="5" pageNumber="572">
<taxonomicNameLabel id="4EC93621FFF4FFCB4A0BF91BFC4F0CB1" box="[891,982,1736,1762]" pageId="5" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
soldier has six long setae on the vertex while the other two species have numerous scattered long and short setae. The head capsule of
<taxonomicName id="A08E2CCBFFF4FFCB4DA8F93FFA050D55" authorityName="Scheffrahn &amp; Roisin &amp; Wang &amp; Bourguignon &amp; Hellemans" authorityYear="2025" box="[1240,1436,1772,1798]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="monomorphus" status="sp. nov.">
<emphasis id="55FA8B5AFFF4FFCB4DA8F93FFA050D55" box="[1240,1436,1772,1798]" italics="true" pageId="5" pageNumber="572">O. monomorphus</emphasis>
</taxonomicName>
<emphasis id="55FA8B5AFFF4FFCB49E7F8C3FF6B0D79" bold="true" box="[151,242,1808,1834]" pageId="5" pageNumber="572">
<taxonomicNameLabel id="4EC93621FFF4FFCB49E7F8C3FF6B0D79" box="[151,242,1808,1834]" pageId="5" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is more pyriform in dorsal view than the other two species of the genus. The soldier of
<taxonomicName id="A08E2CCBFFF4FFCB4DA9F8C3FA050D79" authorityName="Scheffrahn &amp; Roisin &amp; Wang &amp; Bourguignon &amp; Hellemans" authorityYear="2025" box="[1241,1436,1808,1834]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="monomorphus" status="sp. nov.">
<emphasis id="55FA8B5AFFF4FFCB4DA9F8C3FA050D79" box="[1241,1436,1808,1834]" italics="true" pageId="5" pageNumber="572">O. monomorphus</emphasis>
</taxonomicName>
<emphasis id="55FA8B5AFFF4FFCB49E7F8E7FF690D1D" bold="true" box="[151,240,1844,1870]" pageId="5" pageNumber="572">
<taxonomicNameLabel id="4EC93621FFF4FFCB49E7F8E7FF690D1D" box="[151,240,1844,1870]" pageId="5" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the other monomorphic small nasutes of
<collectingCountry id="1F9917D8FFF4FFCB4A3AF8E7FC3A0D1D" box="[842,931,1844,1870]" name="Panama" pageId="5" pageNumber="572">Panama</collectingCountry>
in having the head capsule, antennal articles, and palps reddish brown instead of yellowish (
<figureCitation id="FFB54BCDFFF4FFCB4BD8F88BFD740D21" box="[680,749,1880,1906]" captionStart="FIGURE 7" captionStartId="6.[151,250,1814,1838]" captionTargetBox="[151,1436,179,1788]" captionTargetId="figure-20@6.[151,1436,179,1788]" captionTargetPageId="6" captionText="FIGURE 7. Other small monomorphic nasute species from Panama: dorsal and lateral views of soldier head capsule and worker enteric valve armature. A–C. Araujotermes zeteki (Snyder, 1924). D–F. Atlantitermes kirbyi (Snyder, 1926). G–I. Coatitermes clevelandi (Snyder, 1926). J–L. Ereymatermes panamensis Roisin, 1995. M–O. Subulitermes denisae Roisin, 1995." figureDoi="http://doi.org/10.5281/zenodo.14804875" httpUri="https://zenodo.org/record/14804875/files/figure.png" pageId="5" pageNumber="572">Fig. 7</figureCitation>
). The nasus of the
<taxonomicName id="A08E2CCBFFF4FFCB4AB6F88BFB100D21" authorityName="Scheffrahn &amp; Roisin &amp; Wang &amp; Bourguignon &amp; Hellemans" authorityYear="2025" box="[966,1161,1880,1906]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="monomorphus" status="sp. nov.">
<emphasis id="55FA8B5AFFF4FFCB4AB6F88BFB100D21" box="[966,1161,1880,1906]" italics="true" pageId="5" pageNumber="572">O. monomorphus</emphasis>
</taxonomicName>
<emphasis id="55FA8B5AFFF4FFCB4DE2F88BFB750D21" bold="true" box="[1170,1260,1880,1906]" pageId="5" pageNumber="572">
<taxonomicNameLabel id="4EC93621FFF4FFCB4DE2F88BFB750D21" box="[1170,1260,1880,1906]" pageId="5" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
soldier is more broadly conical with the exception of
<taxonomicName id="A08E2CCBFFF4FFCB4B33F8AFFCDE0DC5" baseAuthorityName="Snyder" baseAuthorityYear="1926" box="[579,839,1916,1942]" class="Insecta" family="Termitidae" genus="Coatitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="clevelandi">
<emphasis id="55FA8B5AFFF4FFCB4B33F8AFFCDE0DC5" box="[579,839,1916,1942]" italics="true" pageId="5" pageNumber="572">Coatitermes clevelandi</emphasis>
</taxonomicName>
(
<figureCitation id="FFB54BCDFFF4FFCB4A27F8AFFC460DC5" box="[855,991,1916,1942]" captionStart="FIGURE 7" captionStartId="6.[151,250,1814,1838]" captionTargetBox="[151,1436,179,1788]" captionTargetId="figure-20@6.[151,1436,179,1788]" captionTargetPageId="6" captionText="FIGURE 7. Other small monomorphic nasute species from Panama: dorsal and lateral views of soldier head capsule and worker enteric valve armature. A–C. Araujotermes zeteki (Snyder, 1924). D–F. Atlantitermes kirbyi (Snyder, 1926). G–I. Coatitermes clevelandi (Snyder, 1926). J–L. Ereymatermes panamensis Roisin, 1995. M–O. Subulitermes denisae Roisin, 1995." figureDoi="http://doi.org/10.5281/zenodo.14804875" httpUri="https://zenodo.org/record/14804875/files/figure.png" pageId="5" pageNumber="572">Figs. 7G, H</figureCitation>
). Among this group, only
<taxonomicName id="A08E2CCBFFF4FFCB4C77F8AFFA050DC5" authorityName="Snyder" authorityYear="1924" box="[1287,1436,1916,1942]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="572" phylum="Arthropoda" rank="genus">
<emphasis id="55FA8B5AFFF4FFCB4C77F8AFFA050DC5" box="[1287,1436,1916,1942]" italics="true" pageId="5" pageNumber="572">Obtusitermes</emphasis>
</taxonomicName>
and
<taxonomicName id="A08E2CCBFFF4FFCB49B7F873FE3E0DE9" baseAuthorityName="Snyder" baseAuthorityYear="1926" box="[199,423,1952,1978]" class="Insecta" family="Termitidae" genus="Atlantitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="kirbyi">
<emphasis id="55FA8B5AFFF4FFCB49B7F873FE3E0DE9" box="[199,423,1952,1978]" italics="true" pageId="5" pageNumber="572">Atlantitermes kirbyi</emphasis>
</taxonomicName>
(
<figureCitation id="FFB54BCDFFF4FFCB48C7F873FDA20DE9" box="[439,571,1952,1978]" captionStart="FIGURE 7" captionStartId="6.[151,250,1814,1838]" captionTargetBox="[151,1436,179,1788]" captionTargetId="figure-20@6.[151,1436,179,1788]" captionTargetPageId="6" captionText="FIGURE 7. Other small monomorphic nasute species from Panama: dorsal and lateral views of soldier head capsule and worker enteric valve armature. A–C. Araujotermes zeteki (Snyder, 1924). D–F. Atlantitermes kirbyi (Snyder, 1926). G–I. Coatitermes clevelandi (Snyder, 1926). J–L. Ereymatermes panamensis Roisin, 1995. M–O. Subulitermes denisae Roisin, 1995." figureDoi="http://doi.org/10.5281/zenodo.14804875" httpUri="https://zenodo.org/record/14804875/files/figure.png" pageId="5" pageNumber="572">Figs. 7D, E</figureCitation>
) have six long setae on the head capsule versus dozens or more on the other species.
</paragraph>
<caption id="33F107C0FFF7FFC849E7F8C5FAA90D25" ID-DOI="http://doi.org/10.5281/zenodo.14804875" ID-Zenodo-Dep="14804875" httpUri="https://zenodo.org/record/14804875/files/figure.png" pageId="6" pageNumber="573" startId="6.[151,250,1814,1838]" targetBox="[151,1436,179,1788]" targetPageId="6" targetType="figure">
<paragraph id="67315748FFF7FFC849E7F8C5FAA90D25" blockId="6.[151,1436,1814,1911]" pageId="6" pageNumber="573">
<emphasis id="55FA8B5AFFF7FFC849E7F8C5FE8B0D7C" bold="true" box="[151,274,1814,1839]" pageId="6" pageNumber="573">FIGURE 7.</emphasis>
Other small monomorphic nasute species from Panama: dorsal and lateral views of soldier head capsule and worker enteric valve armature.
<emphasis id="55FA8B5AFFF7FFC848F7F8E9FE260D01" bold="true" box="[391,447,1850,1874]" pageId="6" pageNumber="573">A–C.</emphasis>
<taxonomicName id="A08E2CCBFFF7FFC848B6F8E8FCA80D00" authority="(Snyder, 1924)" baseAuthorityName="Snyder" baseAuthorityYear="1924" box="[454,817,1850,1875]" class="Insecta" family="Termitidae" genus="Araujotermes" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="zeteki">
<emphasis id="55FA8B5AFFF7FFC848B6F8E8FD090D01" box="[454,656,1850,1874]" italics="true" pageId="6" pageNumber="573">Araujotermes zeteki</emphasis>
(Snyder, 1924)
</taxonomicName>
.
<emphasis id="55FA8B5AFFF7FFC84A4BF8E8FCF40D00" bold="true" box="[827,877,1851,1875]" pageId="6" pageNumber="573">D–F.</emphasis>
<taxonomicName id="A08E2CCBFFF7FFC84A04F8E9FB790D01" authority="(Snyder, 1926)" baseAuthorityName="Snyder" baseAuthorityYear="1926" box="[884,1248,1850,1874]" class="Insecta" family="Termitidae" genus="Atlantitermes" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="kirbyi">
<emphasis id="55FA8B5AFFF7FFC84A04F8E9FBA70D01" box="[884,1086,1850,1874]" italics="true" pageId="6" pageNumber="573">Atlantitermes kirbyi</emphasis>
(
<bibRefCitation id="031F2AB9FFF7FFC84D3CF8E9FB410D01" author="Snyder, T. E." box="[1100,1240,1850,1874]" pageId="6" pageNumber="573" pagination="7 - 16" refId="ref5028" refString="Snyder, T. E. (1926) Five new termites from Panama and Costa Rica. Proceedings of the Entomological Society of Washington, 28, 7-16." type="journal article" year="1926">Snyder, 1926</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="55FA8B5AFFF7FFC84D99F8E9FA830D01" bold="true" box="[1257,1306,1850,1874]" pageId="6" pageNumber="573">G–I.</emphasis>
<taxonomicName id="A08E2CCBFFF7FFC84C50F8E8FE390D25" authority="(Snyder, 1926)" baseAuthorityName="Snyder" baseAuthorityYear="1926" class="Insecta" family="Termitidae" genus="Coatitermes" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="clevelandi">
<emphasis id="55FA8B5AFFF7FFC84C50F8E8FF660D25" italics="true" pageId="6" pageNumber="573">Coatitermes clevelandi</emphasis>
(
<bibRefCitation id="031F2AB9FFF7FFC8487DF88DFE010D25" author="Snyder, T. E." box="[269,408,1886,1910]" pageId="6" pageNumber="573" pagination="7 - 16" refId="ref5028" refString="Snyder, T. E. (1926) Five new termites from Panama and Costa Rica. Proceedings of the Entomological Society of Washington, 28, 7-16." type="journal article" year="1926">Snyder, 1926</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="55FA8B5AFFF7FFC848D9F88CFE430D24" bold="true" box="[425,474,1887,1911]" pageId="6" pageNumber="573">J–L.</emphasis>
<taxonomicName id="A08E2CCBFFF7FFC84890F88CFCE20D25" authority="Roisin, 1995" authorityName="Roisin" authorityYear="1995" box="[480,891,1886,1910]" class="Insecta" family="Termitidae" genus="Ereymatermes" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="55FA8B5AFFF7FFC84890F88CFD690D25" box="[480,752,1887,1910]" italics="true" pageId="6" pageNumber="573">Ereymatermes panamensis</emphasis>
<bibRefCitation id="031F2AB9FFF7FFC84B86F88DFCE20D25" author="Roisin, Y." box="[758,891,1886,1910]" pageId="6" pageNumber="573" pagination="283 - 300" refId="ref4780" refString="Roisin, Y. (1995) Humivorous nasute termites (Isoptera: Nasutitermitinae) from the Panama Canal area. Belgian Journal of Zoology, 125, 283-300." type="journal article" year="1995">Roisin, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="55FA8B5AFFF7FFC84AF4F88DFC5A0D25" bold="true" box="[900,963,1886,1910]" pageId="6" pageNumber="573">M–O.</emphasis>
<taxonomicName id="A08E2CCBFFF7FFC84AB9F88DFAB50D25" authority="Roisin, 1995" authorityName="Roisin" authorityYear="1995" box="[969,1324,1886,1910]" class="Insecta" family="Termitidae" genus="Subulitermes" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="denisae">
<emphasis id="55FA8B5AFFF7FFC84AB9F88DFB380D25" box="[969,1185,1886,1910]" italics="true" pageId="6" pageNumber="573">Subulitermes denisae</emphasis>
<bibRefCitation id="031F2AB9FFF7FFC84DD7F88DFAB50D25" author="Roisin, Y." box="[1191,1324,1886,1910]" pageId="6" pageNumber="573" pagination="283 - 300" refId="ref4780" refString="Roisin, Y. (1995) Humivorous nasute termites (Isoptera: Nasutitermitinae) from the Panama Canal area. Belgian Journal of Zoology, 125, 283-300." type="journal article" year="1995">Roisin, 1995</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="33F107C0FFF6FFC949E7FB59FC1A0E95" ID-DOI="http://doi.org/10.5281/zenodo.14804877" ID-Zenodo-Dep="14804877" httpUri="https://zenodo.org/record/14804877/files/figure.png" pageId="7" pageNumber="574" startId="7.[151,250,1162,1186]" targetBox="[151,1436,179,1136]" targetPageId="7" targetType="figure">
<paragraph id="67315748FFF6FFC949E7FB59FC1A0E95" blockId="7.[151,1436,1162,1223]" pageId="7" pageNumber="574">
<emphasis id="55FA8B5AFFF6FFC949E7FB59FE8A0EF1" bold="true" box="[151,275,1162,1186]" pageId="7" pageNumber="574">FIGURE 8.</emphasis>
Soldiers of
<taxonomicName id="A08E2CCBFFF6FFC948FEFB59FC500EF1" authority="(Snyder, 1925) (UFTC PN 1076)" authorityName="UFTC PN" authorityYear="1076" baseAuthorityName="Snyder" baseAuthorityYear="1925" box="[398,969,1162,1187]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="panamae">
<emphasis id="55FA8B5AFFF6FFC948FEFB59FDE10EF1" box="[398,632,1162,1186]" italics="true" pageId="7" pageNumber="574">Obtusitermes panamae</emphasis>
(
<bibRefCitation id="031F2AB9FFF6FFC94BF5FB59FC970EF1" author="Snyder, T. E." box="[645,782,1162,1186]" pageId="7" pageNumber="574" pagination="179 - 193" refId="ref4995" refString="Snyder, T. E. (1925) New termites and hitherto unknown castes from the Canal Zone, Panama. Journal of Agricultural Research, 29, 179-193." type="journal article" year="1925">Snyder, 1925</bibRefCitation>
) (UFTC PN1076)
</taxonomicName>
.
<emphasis id="55FA8B5AFFF6FFC94AA3FB59FC720EF1" bold="true" box="[979,1003,1162,1186]" pageId="7" pageNumber="574">A.</emphasis>
dorsal and
<emphasis id="55FA8B5AFFF6FFC94D10FB58FBEE0EF0" bold="true" box="[1120,1143,1163,1187]" pageId="7" pageNumber="574">B.</emphasis>
lateral view of major soldier head capsule.
<emphasis id="55FA8B5AFFF6FFC94856FB7DFEA70E95" bold="true" box="[294,318,1198,1222]" pageId="7" pageNumber="574">C.</emphasis>
dorsal and
<emphasis id="55FA8B5AFFF6FFC948C4FB7CFE540E94" bold="true" box="[436,461,1199,1223]" pageId="7" pageNumber="574">D.</emphasis>
lateral view of minor soldier head capsule.
</paragraph>
</caption>
<paragraph id="67315748FFF6FFC949B7FACAFDB80CC8" blockId="7.[151,1437,1304,1871]" pageId="7" pageNumber="574">
<emphasis id="55FA8B5AFFF6FFC949B7FACAFEBD0F60" bold="true" box="[199,292,1305,1331]" pageId="7" pageNumber="574">Worker</emphasis>
(
<figureCitation id="FFB54BCDFFF6FFC94843FACAFE060F61" box="[307,415,1304,1331]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="2.[151,250,1633,1657]" captionStartId-1="3.[151,250,1671,1695]" captionStartId-2="4.[151,250,1945,1969]" captionStartId-3="5.[151,250,657,681]" captionTargetBox-0="[151,1435,763,1596]" captionTargetBox-1="[536,1052,739,1645]" captionTargetBox-2="[151,1436,379,1917]" captionTargetId-0="figure-41@2.[151,1435,763,1607]" captionTargetId-1="figure-224@3.[536,1052,739,1645]" captionTargetId-2="figure-107@4.[151,1436,378,1918]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 3. Worker of Obtusitermes monomorphus sp. nov. A. lateral and B. dorsal view of head capsule. C. habitus. D. dorsal view of mandibles (M1+2 = first and second marginal teeth, M3 = third marginal tooth. Black arrow = cutting edge, gray arrow = molar prominence)." captionText-1="FIGURE 4. Worker gut of Obtusitermes monomorphus sp. nov. A–D: dorsal, right, ventral, and left views, respectively. Abbreviations:C=crop, M=mesenteron, P1a=proximal 1st proctodeal segment, P1b=distal 1st proctodeal segment, P2a= proximal enteric valve, P2b distal enteric valve, P3=3rd proctodeal segment, P4=4th proctodeal segment, and P5=rectum." captionText-2="FIGURE 5. Enteric valve of Obtusitermes monomorphus sp. nov. A. Proximal enteric valve tube (P2a) and distal enteric valve tube (P2b) with respective spiny cushions superimposed. B. Distal spiny cushions and C. proximal spiny cushions. D. Phase contrast of a single proximal cushion. E. Entire P2 with food particles removed." captionText-3="FIGURE 6. Live habitus of Obtusitermes monomorphus sp. nov. (PN1415)." figureDoi-0="http://doi.org/10.5281/zenodo.14804869" figureDoi-1="http://doi.org/10.5281/zenodo.14804871" figureDoi-2="http://doi.org/10.5281/zenodo.14804873" httpUri-0="https://zenodo.org/record/14804869/files/figure.png" httpUri-1="https://zenodo.org/record/14804871/files/figure.png" httpUri-2="https://zenodo.org/record/14804873/files/figure.png" pageId="7" pageNumber="574">Figs. 3–6</figureCitation>
,
<figureCitation id="FFB54BCDFFF6FFC948DBFACAFE530F60" box="[427,458,1305,1331]" captionStart="FIGURE 10" captionStartId="8.[151,250,1979,2003]" captionTargetBox="[164,1423,1241,1954]" captionTargetId="figure-57@8.[164,1423,1241,1954]" captionTargetPageId="8" captionText="FIGURE 10. Worker enteric valve (P2) of Obtusitermes panamae (Snyder, 1925) (UFTC PN1076, A.) and O. formosulus Cuezzo and Cancello, 2009 (UFTC VZ1171, B.). Cellulose particles removed." figureDoi="http://doi.org/10.5281/zenodo.14804881" httpUri="https://zenodo.org/record/14804881/files/figure.png" pageId="7" pageNumber="574">10</figureCitation>
;
<tableCitation id="2A0C62F3FFF6FFC948A5FACAFDB20F60" box="[469,555,1305,1331]" captionStart="TABLE 2" captionStartId="2.[151,239,1777,1801]" captionText="TABLE 2. Measurements (mm) of Obtusitermes monomorphus sp. nov. workers (n=5) from two colonies." pageId="7" pageNumber="574">Table 2</tableCitation>
). Monomorphic. Head capsule yellowish brown; vertex with concavity behind postclypeus; postclypeus slightly inflated.Antennae with 12 articles; antennal article formula 2&gt;3≥4=5. Mesenteron and P1 tubular, mixed segment lacking; P1 divided near middle by slight darkened constriction. Enteric valve armature very long, tubular, curved 180°; armature at both ends; EVA seating simple. Enteric valve armature composed of two rings separated by nearly
<quantity id="A076FAADFFF6FFC94BE6FA7AFD430F90" box="[662,730,1449,1475]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="574" unit="mm" value="1.0">1 mm</quantity>
of smooth cuticle. Proximal EVA ring at the terminus of P1 consists of six triangular cushions with their apex directed distally with food stream. Each cushion contains about 20 to 30 scales terminating in short spines. Distal ring at insertion of P3 consists of six cushions alternating in pairs of triangular and elongate scaled cushions; scale texture as in the proximal ring. Left mandible with apical and first marginal teeth equal; molar prominence small; distal margin of the M1+2 forms a long, slightly concave cutting edge longer than apical tooth distance to molar prominence. Right mandible with apical tooth narrower than first marginal. Molar plates with ridges.
</paragraph>
<paragraph id="67315748FFF6FFC949B7F977FBCA0D1C" blockId="7.[151,1437,1304,1871]" pageId="7" pageNumber="574">
<emphasis id="55FA8B5AFFF6FFC949B7F977FEF40CED" bold="true" box="[199,365,1700,1726]" pageId="7" pageNumber="574">Comparisons.</emphasis>
The worker head capsule width of
<taxonomicName id="A08E2CCBFFF6FFC94A7CF976FC480CEC" authorityName="Scheffrahn &amp; Roisin &amp; Wang &amp; Bourguignon &amp; Hellemans" authorityYear="2025" box="[780,977,1701,1727]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="monomorphus" status="sp. nov.">
<emphasis id="55FA8B5AFFF6FFC94A7CF976FC480CEC" box="[780,977,1701,1727]" italics="true" pageId="7" pageNumber="574">O. monomorphus</emphasis>
</taxonomicName>
<emphasis id="55FA8B5AFFF6FFC94AABF976FBAE0CEC" bold="true" box="[987,1079,1701,1727]" pageId="7" pageNumber="574">
<taxonomicNameLabel id="4EC93621FFF6FFC94AABF976FBAE0CEC" box="[987,1079,1701,1727]" pageId="7" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is somewhat larger (0.95–
<quantity id="A076FAADFFF6FFC94C1BF977FF5D0CB0" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="574" unit="mm" value="0.85">0.85 mm</quantity>
) than the major worker of
<taxonomicName id="A08E2CCBFFF6FFC94882F91AFDE20CB1" authorityName="UFTC PN" authorityYear="1076" baseAuthorityName="Snyder" baseAuthorityYear="1925" box="[498,635,1737,1763]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="panamae">
<emphasis id="55FA8B5AFFF6FFC94882F91AFDE20CB1" box="[498,635,1737,1763]" italics="true" pageId="7" pageNumber="574">O. panamae</emphasis>
</taxonomicName>
(
<quantity id="A076FAADFFF6FFC94BF9F91BFCA90CB0" box="[649,816,1736,1763]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" metricValueMax="7.8" metricValueMin="6.4" pageId="7" pageNumber="574" unit="mm" value="0.71" valueMax="0.78" valueMin="0.64">0.64–0.78 mm</quantity>
, UFTC PN1076; 0.60–0.62, Snyder 1924) or the major worker of
<taxonomicName id="A08E2CCBFFF6FFC9487BF93EFE300D54" authorityName="Cuezzo and Cancello" authorityYear="2009" box="[267,425,1773,1799]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="formosulus">
<emphasis id="55FA8B5AFFF6FFC9487BF93EFE300D54" box="[267,425,1773,1799]" italics="true" pageId="7" pageNumber="574">O. formosulus</emphasis>
</taxonomicName>
(
<quantity id="A076FAADFFF6FFC948C8F93FFDC70D54" box="[440,606,1772,1799]" metricMagnitude="-4" metricUnit="m" metricValue="7.749999999999999" metricValueMax="8.5" metricValueMin="7.0" pageId="7" pageNumber="574" unit="mm" value="0.7749999999999999" valueMax="0.85" valueMin="0.7">0.70–0.85 mm</quantity>
,
<bibRefCitation id="031F2AB9FFF6FFC94B19F93EFC1E0D54" author="Cuezzo, A. C. &amp; Cancello, E. M." box="[617,903,1773,1799]" pageId="7" pageNumber="574" pagination="61 - 68" refId="ref4044" refString="Cuezzo, A. C. &amp; Cancello, E. M. (2009) A new species of Obtusitermes (Isoptera, Termitidae, Nasutitermitinae) from South America. Zootaxa, 1993, 61-68. https://doi.org/10.11646/zootaxa.1993.1.6" type="journal article" year="2009">Cuezzo &amp; Cancello 2009</bibRefCitation>
). Mandibular dentition of all three are identical. The worker hindgut of
<taxonomicName id="A08E2CCBFFF6FFC948E9F8C2FDFD0D78" box="[409,612,1809,1835]" class="Insecta" family="Termitidae" genus="Obtusitermes" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="55FA8B5AFFF6FFC948E9F8C2FDB70D78" box="[409,558,1809,1835]" italics="true" pageId="7" pageNumber="574">Obtusitermes</emphasis>
spp.
</taxonomicName>
is unique among termites in having the P1 joined to the P3 by a very long and tubular P2 with a curvature of 180° and weak armature at both ends (
<figureCitation id="FFB54BCDFFF6FFC94ABCF8E6FB820D1C" box="[972,1051,1844,1871]" captionStart="FIGURE 5" captionStartId="4.[151,250,1945,1969]" captionTargetBox="[151,1436,379,1917]" captionTargetId="figure-107@4.[151,1436,378,1918]" captionTargetPageId="4" captionText="FIGURE 5. Enteric valve of Obtusitermes monomorphus sp. nov. A. Proximal enteric valve tube (P2a) and distal enteric valve tube (P2b) with respective spiny cushions superimposed. B. Distal spiny cushions and C. proximal spiny cushions. D. Phase contrast of a single proximal cushion. E. Entire P2 with food particles removed." figureDoi="http://doi.org/10.5281/zenodo.14804873" httpUri="https://zenodo.org/record/14804873/files/figure.png" pageId="7" pageNumber="574">Figs. 5</figureCitation>
,
<figureCitation id="FFB54BCDFFF6FFC94D57F8E6FBDF0D1C" box="[1063,1094,1845,1871]" captionStart="FIGURE 10" captionStartId="8.[151,250,1979,2003]" captionTargetBox="[164,1423,1241,1954]" captionTargetId="figure-57@8.[164,1423,1241,1954]" captionTargetPageId="8" captionText="FIGURE 10. Worker enteric valve (P2) of Obtusitermes panamae (Snyder, 1925) (UFTC PN1076, A.) and O. formosulus Cuezzo and Cancello, 2009 (UFTC VZ1171, B.). Cellulose particles removed." figureDoi="http://doi.org/10.5281/zenodo.14804881" httpUri="https://zenodo.org/record/14804881/files/figure.png" pageId="7" pageNumber="574">10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>