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<mods:title id="E3BE84BC5A391F14909F3307CACEA4E4">A new record of a sea star, Henricia aspera Fisher, 1906 (Asteroidea: Spinulosida: Echinasteridae) from Jeju Island, Korea</mods:title>
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<mods:title id="0A5EAA40D43E20D0411B595652C90D0B">A new record of a sea star, Henricia aspera Fisher, 1906 (Asteroidea: Spinulosida: Echinasteridae) from Jeju Island, Korea</mods:title>
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<emphasis id="99CA6B37ED09FFB0FF1AFACBFEA1FAB5" bold="true" box="[144,282,1370,1394]" pageId="2" pageNumber="156">Description.</emphasis>
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Arms five, long, semi-cylindrical, swollen arm base, gradually tapering to tips; disc small (
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<figureCitation id="3385ABA0ED09FFB0FD19FAEAFF1FFA74" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." pageId="2" pageNumber="156">Fig. 1A, B</figureCitation>
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<figureCitation id="3385ABA0ED09FFB0FD19FAEAFF1FFA74" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." figureDoi="http://doi.org/10.5281/zenodo.13139803" httpUri="https://zenodo.org/record/13139803/files/figure.png" pageId="2" pageNumber="156">Fig. 1A, B</figureCitation>
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). Abactinal paxillae scattered, containing three to seven small abactinal spinelets, covered with thin integument (
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<figureCitation id="3385ABA0ED09FFB0FF12FA4AFF54FA33" box="[152,239,1499,1524]" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." pageId="2" pageNumber="156">Fig. 1C</figureCitation>
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<figureCitation id="3385ABA0ED09FFB0FF12FA4AFF54FA33" box="[152,239,1499,1524]" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." figureDoi="http://doi.org/10.5281/zenodo.13139803" httpUri="https://zenodo.org/record/13139803/files/figure.png" pageId="2" pageNumber="156">Fig. 1C</figureCitation>
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). Denuded abactinal spines broad at base and rapidly tapering from middle part to tip (
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<figureCitation id="3385ABA0ED09FFB0FDB7FA6DFD3CF9D3" box="[573,647,1532,1556]" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." pageId="2" pageNumber="156">Fig. 1J</figureCitation>
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<figureCitation id="3385ABA0ED09FFB0FDB7FA6DFD3CF9D3" box="[573,647,1532,1556]" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." figureDoi="http://doi.org/10.5281/zenodo.13139803" httpUri="https://zenodo.org/record/13139803/files/figure.png" pageId="2" pageNumber="156">Fig. 1J</figureCitation>
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). Papular areas wide with irregular shapes, containing two to nine papulae in an area; some areas bearing small ossicles (
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<figureCitation id="3385ABA0ED09FFB0FD48F9ADFF6FF9B2" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." pageId="2" pageNumber="156">Fig. 1C, H</figureCitation>
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<figureCitation id="3385ABA0ED09FFB0FD48F9ADFF6FF9B2" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." figureDoi="http://doi.org/10.5281/zenodo.13139803" httpUri="https://zenodo.org/record/13139803/files/figure.png" pageId="2" pageNumber="156">Fig. 1C, H</figureCitation>
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). Denuded abactinal skeleton open meshed, reticulated, comprised of lobe-shaped and crescent-shaped plates; some skeleton forming semi-rounded plates (
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<figureCitation id="3385ABA0ED09FFB0FD48F90CFF08F912" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." pageId="2" pageNumber="156">Fig. 1G</figureCitation>
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<figureCitation id="3385ABA0ED09FFB0FD48F90CFF08F912" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." figureDoi="http://doi.org/10.5281/zenodo.13139803" httpUri="https://zenodo.org/record/13139803/files/figure.png" pageId="2" pageNumber="156">Fig. 1G</figureCitation>
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). Madreporite sunken, situated near margin of disk, circular in form, and bearing spines similar as adjacent spines (
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<figureCitation id="3385ABA0ED09FFB0FF68F96FFE8DF8D1" box="[226,310,1790,1814]" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." pageId="2" pageNumber="156">Fig. 1F</figureCitation>
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<figureCitation id="3385ABA0ED09FFB0FF68F96FFE8DF8D1" box="[226,310,1790,1814]" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." figureDoi="http://doi.org/10.5281/zenodo.13139803" httpUri="https://zenodo.org/record/13139803/files/figure.png" pageId="2" pageNumber="156">Fig. 1F</figureCitation>
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). Marginal plates well discernible. Denuded actinal skeleton showing three series of plates (Inferomarginal, intermarginal, and ventrolateral) which diverging near arm tip, leaving intermarginal plates wide spaces. Inferomarginal plates imbricated, elongated cross-shaped, reaching three-fourth length of arm. Intermarginal plate consisting of lobe-shaped plates, with small roundshape ossicles, reaching near half of arm. Ventrolateral plates rounded cross-shaped, showing larger size compared to adjacent plates; narrow papular areas with one or two papulae (
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<figureCitation id="3385ABA0ED09FFB0FC32FCE7FBB9FC48" box="[952,1026,886,911]" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." pageId="2" pageNumber="156">Fig. 1I</figureCitation>
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<figureCitation id="3385ABA0ED09FFB0FC32FCE7FBB9FC48" box="[952,1026,886,911]" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." figureDoi="http://doi.org/10.5281/zenodo.13139803" httpUri="https://zenodo.org/record/13139803/files/figure.png" pageId="2" pageNumber="156">Fig. 1I</figureCitation>
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). Adambulacral armature composed of four to eight short, robust spinelets, inner two or three spatulate-shape, larger spines, arranged in two transverse rows or in a zigzag row (
|
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<figureCitation id="3385ABA0ED09FFB0FBBAFC46FB3FFC28" box="[1072,1156,983,1007]" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." pageId="2" pageNumber="156">Fig. 1D</figureCitation>
|
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<figureCitation id="3385ABA0ED09FFB0FBBAFC46FB3FFC28" box="[1072,1156,983,1007]" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." figureDoi="http://doi.org/10.5281/zenodo.13139803" httpUri="https://zenodo.org/record/13139803/files/figure.png" pageId="2" pageNumber="156">Fig. 1D</figureCitation>
|
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). Oral plate bearing two spatulate spines (
|
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<figureCitation id="3385ABA0ED09FFB0FC53FC66FB93FBD7" box="[985,1064,1015,1040]" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." pageId="2" pageNumber="156">Fig. 1E</figureCitation>
|
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<figureCitation id="3385ABA0ED09FFB0FC53FC66FB93FBD7" box="[985,1064,1015,1040]" captionStart="Fig" captionStartId="1.[176,209,1849,1871]" captionTargetBox="[176,1446,257,1812]" captionTargetId="figure-12@1.[176,1446,255,1827]" captionTargetPageId="1" captionText="Fig. 1. Henricia aspera. A. abactinal side; B. actinal side; C. abactinal paxillae; D, K. adambulacral spines; E. oral part; F. madreporite; G. abactinal skeleton; H. papulae (arrows); I. actinal skeleton: inferomarginal plates (i); intermarginal plates(in), ventrolateral plates (v), adambulacral plates (a); and J. abactinal spines. Scale bars:A, B=2 cm, C-I=1 mm, J=100 μm, K =500 μm (J, K, SEM images)." figureDoi="http://doi.org/10.5281/zenodo.13139803" httpUri="https://zenodo.org/record/13139803/files/figure.png" pageId="2" pageNumber="156">Fig. 1E</figureCitation>
|
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). Furrow spine single.
|
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</paragraph>
|
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<paragraph id="AB01B725ED09FFB0FCA3FB89FB7EFBF7" blockId="2.[809,1414,757,1975]" box="[809,1221,1048,1072]" pageId="2" pageNumber="156">
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