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<mods:title id="34459F93891F9C0B18430BC8A08E140E">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<treatment ID-GBIF-Taxon="181753573" LSID="urn:lsid:plazi:treatment:02249EDF73C059D1843C3DE96E998ED5" httpUri="http://treatment.plazi.org/id/02249EDF73C059D1843C3DE96E998ED5" lastPageNumber="145" pageId="0" pageNumber="145">
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<treatment id="02249EDF73C059D1843C3DE96E998ED5" ID-GBIF-Taxon="181753573" LSID="urn:lsid:plazi:treatment:02249EDF73C059D1843C3DE96E998ED5" httpUri="http://treatment.plazi.org/id/02249EDF73C059D1843C3DE96E998ED5" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
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<paragraph id="F57323D5F4BF387FAEB4E4A1F52DCF9B" pageId="0" pageNumber="145">
Genus
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<paragraph pageId="0" pageNumber="145">Type species.</paragraph>
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<taxonomicName class="Arachnida" family="Zodariidae" genus="Enyo" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Enyo nitida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitida">
<emphasis italics="true" pageId="0" pageNumber="145">Enyo nitida</emphasis>
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<paragraph id="AA3D977C13206B9747DA3EFED6760182" pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph id="7F8D3C2D3C5A9A833515AD0FAD217570" pageId="0" pageNumber="145">
<taxonomicName id="115C472EE2D87AE4E48D6A38CFE76810" class="Arachnida" family="Zodariidae" genus="Enyo" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Enyo nitida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="117FDD52C239BE7B18D49F660B263B6A" italics="true" pageId="0" pageNumber="145">Enyo nitida</emphasis>
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Audouin, 1826 from Egypt.
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<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">
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<paragraph id="C64B8E8ED26644473A6B18383A27FA40" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph id="770B3FA50E3FE105D288BF2C86B81558" pageId="0" pageNumber="145">
With 177 named species, this is the largest genus within
<taxonomicName authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName>
<taxonomicName id="DFBEA76ADD13A99871546F92434385E4" authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName>
(
<bibRefCitation author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC 2021</bibRefCitation>
<bibRefCitation id="C05AB38B7C6D7AF1D0F69EE51548D4D8" author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC 2021</bibRefCitation>
). Based on the copulatory organs of the species currently considered in this genus,
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
<taxonomicName id="B3FA8F757BAE53D3B79F767ECACF0022" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="515E6BA3F5881C9037F4ADCF4569B76B" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
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does not appear to be monophyletic. There are only two species that are morphologically similar to the generotype:
<taxonomicName lsidName="Z. luctuosum" pageId="0" pageNumber="145" rank="species" species="luctuosum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. luctuosum</emphasis>
<taxonomicName id="DC0AE2A849AC5F3D8C255AB65F217583" lsidName="Z. luctuosum" pageId="0" pageNumber="145" rank="species" species="luctuosum">
<emphasis id="F48A6AB60F66F3BBE2A2DEB115D5537E" italics="true" pageId="0" pageNumber="145">Z. luctuosum</emphasis>
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(O. Pickard-Cambridge, 1872) and
<taxonomicName lsidName="Z. lutipes" pageId="0" pageNumber="145" rank="species" species="lutipes">
<emphasis italics="true" pageId="0" pageNumber="145">Z. lutipes</emphasis>
<taxonomicName id="0A42897BDB14512B2D3DDC403E6EEB0D" lsidName="Z. lutipes" pageId="0" pageNumber="145" rank="species" species="lutipes">
<emphasis id="49C06D8660EA049914C72E95F23C302F" italics="true" pageId="0" pageNumber="145">Z. lutipes</emphasis>
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(O. Pickard-Cambridge, 1872), from the eastern Mediterranean (east of Tunisia to Iran).
</paragraph>
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<subSubSection pageId="0" pageNumber="145" type="materials_examined">
<paragraph pageId="0" pageNumber="145">Comparative material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion expers" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="expers">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion expers</emphasis>
<subSubSection id="AFC0DECB1E432B5AC42F55A6D1EF941C" pageId="0" pageNumber="145" type="materials_examined">
<paragraph id="FDA07F83F6FE446D984509F47C3CDDC8" pageId="0" pageNumber="145">Comparative material.</paragraph>
<paragraph id="9E9F4963AA4FB08881BFC4A3E8A72F4E" pageId="0" pageNumber="145">
<taxonomicName id="3B29D01F09ED3D6BA6BBB59E99944192" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion expers" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="expers">
<emphasis id="D1292117F21BA97FB2AF0697ABDC4782" italics="true" pageId="0" pageNumber="145">Zodarion expers</emphasis>
</taxonomicName>
(O. Pickard-Cambridge, 1876),
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from
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(Figs
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<figureCitation id="C0FD670EA72F7D67F5A8E954F92A0F02" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19D-F</figureCitation>
,
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. SEM images of the male palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, B, D ventral C, E retrolateral F dorsoretrolateral. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis, Ra - retrolateral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145">25D-F</figureCitation>
<figureCitation id="C66B6858C629A09A41F52678942D56D4" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. SEM images of the male palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, B, D ventral C, E retrolateral F dorsoretrolateral. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis, Ra - retrolateral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145">25D-F</figureCitation>
) and
<taxonomicName baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion nitidum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitidum">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion nitidum</emphasis>
<taxonomicName id="7146295478D7D1F6E9244C53097055C6" baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion nitidum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitidum">
<emphasis id="0E8CC7027D40D607B0E85F92C8447757" italics="true" pageId="0" pageNumber="145">Zodarion nitidum</emphasis>
</taxonomicName>
(Audouin, 1826):
<specimenCount type="male">1♂</specimenCount>
<specimenCount id="155930A6EE02471FB027C119F53F5F27" type="male">1♂</specimenCount>
from
<collectingCountry name="Israel">Israel</collectingCountry>
<collectingCountry id="8B71DF92A1B97C49683FFE5DF75A1EF7" name="Israel">Israel</collectingCountry>
(Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. SEM images of the male palps of Zodarion nitidum (A-C) and Z. bactrianum (D-F) A, D ventral B, E retrolateral C dorsal F retrodorsal. Abbreviations: Co - conductor, Eb - embolar base, Em - embolus, Ma - median apophysis, Mc - claw of median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process, Tu - tutaculum, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure28" httpUri="https://binary.pensoft.net/fig/536905" pageId="0" pageNumber="145">28A-C</figureCitation>
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)
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</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="0FADA4FBE19B27EE4A69AFF650F35BD3" pageId="0" pageNumber="145" type="comments">
<paragraph id="164E39919DF6E328EF23608B29B61DAF" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph id="8C34E0AB325456F06198E651D00EBDFC" pageId="0" pageNumber="145">
Until recently,
<taxonomicName lsidName="Z. expers" pageId="0" pageNumber="145" rank="species" species="expers">
<emphasis italics="true" pageId="0" pageNumber="145">Z. expers</emphasis>
<taxonomicName id="BB865E18A2720E571E982147199500BE" lsidName="Z. expers" pageId="0" pageNumber="145" rank="species" species="expers">
<emphasis id="C71D61FAE2379104D1B2F2C9B85CED09" italics="true" pageId="0" pageNumber="145">Z. expers</emphasis>
</taxonomicName>
was placed in
<taxonomicName authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Ranops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ranops" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Ranops</emphasis>
<taxonomicName id="A6858C0602702035FA45B5A15E759F5D" authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Ranops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ranops" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="91EE6D153201F2F4EB13F397B9C8CB33" italics="true" pageId="0" pageNumber="145">Ranops</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jocqué">Jocque</normalizedToken>
<normalizedToken id="C848D4BC653BD01B5E7FFAE7BE307045" originalValue="Jocqué">Jocque</normalizedToken>
, 1991, but now it is placed in
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
<taxonomicName id="1C7D7790739EBA24233509A649D2BFA2" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="E9327B15B34E680F8EBDB863DC117FD1" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName>
. It differs from the generotype of
<taxonomicName authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Ranops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ranops" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Ranops</emphasis>
<taxonomicName id="C67344BB633707FF4126777440A68D12" authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Ranops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ranops" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="78F8CB800925D0FC93C169CE540C2C0A" italics="true" pageId="0" pageNumber="145">Ranops</emphasis>
</taxonomicName>
, but the copulatory organs are also very different from those of
<taxonomicName lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
<taxonomicName id="24BFF4C6E6F2596F0726A0F138E158B5" lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis id="DDB00CC34670707472BD2151C274A103" italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
</taxonomicName>
, and most likely it represents a separate genus. Although it is absent in Iran or Turkmenistan, we have provided figures of this species to illustrate the differences with the generotype of
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
<taxonomicName id="90E2A520B9CCACA16748ADC34DDD3C31" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="B4A13C221605FAB2545AE40877D8F3D9" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName>
. Furthermore, although
<taxonomicName lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
<taxonomicName id="0836D45646F457A4FA4B25B0DE5823C6" lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis id="E009DD5F9048FB07135881C4BF4DD618" italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
</taxonomicName>
has not been recorded in the current study area, because it is the type species of the genus, we have provided illustartions for this species as well, to demonstrate its differences with other
<taxonomicName authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName>
<taxonomicName id="ADED0C1D71FB722A1EB3020E9E7265B0" authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName>
genera in Iran and Turkmenistan, as well as with other species currently placed in
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
<taxonomicName id="E1B59239019B906425258E8D7E3FB10B" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="CF42DDAA6AC98587437FB890BF576FD4" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName>
.
</paragraph>

View file

@ -1,229 +1,229 @@
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<taxonomicName LSID="http://zoobank.org/9DEB48F2-08A4-482E-AEFF-405D9AAAA248" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila" status="sp. nov.">Pax leila</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">Figs 5C, D</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">, 8</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">, 9</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">, 10F-J</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
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<paragraph id="51B3461CAB9433631D2937A742852474" pageId="0" pageNumber="145">
<taxonomicName id="2AE6673BC1401BE60225688D592ECDDC" ID-CoL="98YZ7" LSID="http://zoobank.org/9DEB48F2-08A4-482E-AEFF-405D9AAAA248" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila" status="sp. nov.">Pax leila</taxonomicName>
<taxonomicNameLabel id="95FDE61E3CAE8BD7840F68654D255665" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="5A2C9A11B4F60C5FCF176709C3DC9950" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">Figs 5C, D</figureCitation>
<figureCitation id="A1D4B4C4F1ADB760BFACBDC250ABBE47" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">, 8</figureCitation>
<figureCitation id="5DFA95A2A776C7D0F6DAA863ED8DA71F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">, 9</figureCitation>
<figureCitation id="739E51E13DAD97771AC0B89180FB47B8" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">, 10F-J</figureCitation>
<figureCitation id="6E481C54DBF5EFEEFC3EFFFEB5E9FFD0" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="FA777627746326AA3AA1688C50FB90C2" pageId="0" pageNumber="145" type="type material">
<paragraph id="C070B08018403A6BCB6B0F55E791C79A" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="0D2D9333DCC2EE79B57E480508A4DE18" pageId="0" pageNumber="145">
<emphasis id="F45790FB34322E8B205E86073A8E878D" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ and
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis>
<emphasis id="6CAB6126CA078587D15355101B173BC3" bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis>
1♀ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Fars Province</emphasis>
<emphasis id="76BA2AF8D2599E86CE055840E33F8B70" italics="true" pageId="0" pageNumber="145">Fars Province</emphasis>
: road to Yasuj,
<geoCoordinate degrees="30" direction="north" minutes="28" orientation="latitude" precision="925" value="30.466667">30°28'N</geoCoordinate>
<geoCoordinate id="B54ABE06A265D8118A97710001BA370B" degrees="30" direction="north" minutes="28" orientation="latitude" precision="925" value="30.466667">30°28'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="30" orientation="longitude" precision="925" value="51.5">51°30'E</geoCoordinate>
<geoCoordinate id="011833FB1319D5E8D350FC766577CD85" degrees="51" direction="east" minutes="30" orientation="longitude" precision="925" value="51.5">51°30'E</geoCoordinate>
, 25.05.1974 (A. Senglet).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet is a feminine given name in the Persian language, meaning &quot;daughter of the night&quot;. Noun in apposition.</paragraph>
<subSubSection id="4ED5441780D1374CB0C48EF52CA86BDB" pageId="0" pageNumber="145" type="etymology">
<paragraph id="D51D30BA473B006B9373EC1F5A99CE7A" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="4E51DEEBF696DA81FE5F3D1F1F15AF38" pageId="0" pageNumber="145">The specific epithet is a feminine given name in the Persian language, meaning &quot;daughter of the night&quot;. Noun in apposition.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="4C6E036F3BB0C13E9083C2750C2F436D" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="D01350BE9F66B49B71DF8D83E107406A" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="C84CD4E99DE8B4FB544C491CA84F5848" pageId="0" pageNumber="145">
The new species differs from all congeners by having a pair of spine-like cymbial outgrowths (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">8</figureCitation>
<figureCitation id="98B655D74E64C078F1168E438F370CA2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">8</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">9C, E</figureCitation>
<figureCitation id="652433934DEE6D723C2CD5E44A2EF15D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">9C, E</figureCitation>
) (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="CF42DCA06FA17C8D0709A0AB601ADBE7" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
lacking) and a triangular epigynal median plate (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="27F6AC7D2B5202B236A7BE366003E581" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
absent or rectangular).
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 8.</emphasis>
<caption id="C448ADE7E1135D19E0E32EA8E1C922C2" doi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145" start="Figure 8" startId="F8">
<paragraph id="D37620CFE35199C30125B543D8469984" pageId="0" pageNumber="145">
<emphasis id="7EAE1F8E57B854F60C4CED2234D6CD54" bold="true" pageId="0" pageNumber="145">Figure 8.</emphasis>
Male palp of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila">
<emphasis italics="true" pageId="0" pageNumber="145">Pax leila</emphasis>
<taxonomicName id="9C4DADAE36CA6C14EEB7798770C346F6" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila">
<emphasis id="0B305F4FD2CE5A844A834C54087FBC90" italics="true" pageId="0" pageNumber="145">Pax leila</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
<emphasis id="52AF5D2BF2EF266E2235D2B4A4480CE5" bold="true" pageId="0" pageNumber="145">A</emphasis>
proventral
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
<emphasis id="165B7CC20AE8F37D1DFF9F13EC7F596D" bold="true" pageId="0" pageNumber="145">B</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
<emphasis id="F3B1907D8EC0DAC76B1DC8ED6BCD3BBF" bold="true" pageId="0" pageNumber="145">C</emphasis>
ventroretrolateral
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
<emphasis id="CDFDB57750CA1A33167FE39424DA8B99" bold="true" pageId="0" pageNumber="145">D</emphasis>
prolateral
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
<emphasis id="4DC1B840D415D6338D0ADE753EC6C5CE" bold="true" pageId="0" pageNumber="145">E</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
<emphasis id="746417865834C56F4B14565D44FF79E5" bold="true" pageId="0" pageNumber="145">F</emphasis>
retrolateral. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
<subSubSection id="331289FAAEE58BC5B91C32D445903BF1" pageId="0" pageNumber="145" type="description">
<paragraph id="C5369750918E42EC43D2ECB09A88FAEE" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="644A209423ED7521A6DAAB2F56708660" pageId="0" pageNumber="145">
<emphasis id="2806484CF0771CBE32C9FD1F9EEBFFFB" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5C</figureCitation>
<figureCitation id="8B110704F23D11AA905653FDBBFBF08B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5C</figureCitation>
. Total length 4.42. Carapace 2.07 long, 1.03 wide at pars cephalica, 1.46 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.09, ALE: 0.09, PME: 0.06, PLE: 0.09, PME-PME: 0.07. Carapace, sternum, labium, chelicerae and maxillae reddish brown, without any pattern. Chelicera with 2 promarginal teeth. Legs yellow, with few spines and without annulations. Abdomen dark brown, dorsally with large scutum covering 2/3 of the abdomen. Spinnerets pale, uniform in color. Measurements of legs: I: 4.51 (1.21, 0.50, 1.06, 0.93, 0.81), II: 3.89 (1.04, 0.52, 0.79, 0.83, 0.71), III: 3.62 (0.98, 0.50, 0.69, 0.85, 0.60), IV: 4.99 (1.33, 0.58, 1.05, 1.33, 0.70).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="19CADD542F51ABE823B88F014581B96F" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">8A-F</figureCitation>
<figureCitation id="45BF8077FE0548D1BCA6D47FA3F42655" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">8A-F</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">9A-E</figureCitation>
<figureCitation id="35F512F44E81CEA4B087B0280C75F515" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">9A-E</figureCitation>
. Tibia wider than long, with short apophysis; cymbium with 2 strong spine-like baso-posterior outgrowths (
<emphasis italics="true" pageId="0" pageNumber="145">Cp</emphasis>
<emphasis id="B11777083988408CD5AAECF53B9943C0" italics="true" pageId="0" pageNumber="145">Cp</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="145">Cr</emphasis>
<emphasis id="EC1C8CBEA463F34317A5E9DE75B3F760" italics="true" pageId="0" pageNumber="145">Cr</emphasis>
), lateral fold lacking; embolus long, with robust triangular outgrowth near base (
<emphasis italics="true" pageId="0" pageNumber="145">Se</emphasis>
<emphasis id="1D1AB9F10151F8EA863238A1BE6557F0" italics="true" pageId="0" pageNumber="145">Se</emphasis>
); median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="2F34EE9F12F3E8BFD1282E2BCCC25080" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) very large, with 2 claw-like extensions; conductor small and strongly sclerotized.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 9.</emphasis>
<caption id="AB5F3B3E382925E0FFCED6505E966E61" doi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145" start="Figure 9" startId="F9">
<paragraph id="D9F0744620903DD21D67AC88FC4BFC9C" pageId="0" pageNumber="145">
<emphasis id="49DB0536F2B6BFE2349C8BF91C60E2DC" bold="true" pageId="0" pageNumber="145">Figure 9.</emphasis>
SEM images of the male palp of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila">
<emphasis italics="true" pageId="0" pageNumber="145">Pax leila</emphasis>
<taxonomicName id="A54BA2C6573765749463CBD9A0F67663" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila">
<emphasis id="4F50C05BAE0A78165AD08B978857EC93" italics="true" pageId="0" pageNumber="145">Pax leila</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
<emphasis id="3F9916DBB667F91B49C837670A83B4D6" bold="true" pageId="0" pageNumber="145">A</emphasis>
apicoventral
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
<emphasis id="4E30F043477EF83D7E191B75F426B0FB" bold="true" pageId="0" pageNumber="145">B</emphasis>
ventroretrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
<emphasis id="E0CE82CAE995D85A0A75C36858801F9A" bold="true" pageId="0" pageNumber="145">C</emphasis>
prodorsal
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
<emphasis id="20F1FE821B4DBDEC8DF54D356D1F1EBB" bold="true" pageId="0" pageNumber="145">D</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
<emphasis id="FF5B27286CDDF0ADC258D0EEF7C58D19" bold="true" pageId="0" pageNumber="145">E</emphasis>
retrodorsal. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Cp</emphasis>
<emphasis id="A079B523888F130DC7BF015E41C13A45" italics="true" pageId="0" pageNumber="145">Cp</emphasis>
- prolateral cymbial process,
<emphasis italics="true" pageId="0" pageNumber="145">Cr</emphasis>
<emphasis id="DEE48351EB109074B5381A71F561E781" italics="true" pageId="0" pageNumber="145">Cr</emphasis>
- retrolateral cymbial process,
<emphasis italics="true" pageId="0" pageNumber="145">Da</emphasis>
<emphasis id="579CEBD106D87CCC235830E4CB563A0A" italics="true" pageId="0" pageNumber="145">Da</emphasis>
- dorsal apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="711B0A4EED352E17F7A04EB4013B71A3" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Se</emphasis>
<emphasis id="3352366A9EC96F193065E824CDA97085" italics="true" pageId="0" pageNumber="145">Se</emphasis>
- spine of embolus base. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="942FED49D47F484ED2761D73697F641B" pageId="0" pageNumber="145">
<emphasis id="114C8D63694102AF62B9ED89D14E282B" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5D</figureCitation>
<figureCitation id="0CCFBC148C0036178772F63F1509504A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5D</figureCitation>
. Total length 4.95. Carapace 1.97 long, 1.00 wide at pars cephalica, 1.31 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.07, ALE: 0.08, PME: 0.06, PLE: 0.07, PME-PME: 0.08. Coloration as in male, with lighter abdomen lacking a scutum. Measurements of legs: I: 4.25 (1.21, 0.50, 0.99, 0.84, 0.71), II: 3.58 (0.94, 0.52, 0.74, 0.74, 0.64), III: 3.47 (0.90, 0.51, 0.70, 0.81, 0.55), IV: 4.71 (1.18, 0.53, 1.05, 1.21, 0.74).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="68AA08A588CA6B32860A9EF6E7699DC2" pageId="0" pageNumber="145">
Epigyne as in Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">10F-J</figureCitation>
<figureCitation id="48C364A8470D174C535E92B421B88A45" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">10F-J</figureCitation>
. Epigyne trilobate, with triangular median plate, posteriorly as wide as lateral lobes; copulatory openings slit-like, broad; copulatory ducts wide, as wide as receptacles; receptacles globular, separated by less than their radii.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 10.</emphasis>
<caption id="37185AAEE15756DCCB2FA4E70BC21A99" doi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145" start="Figure 10" startId="F10">
<paragraph id="B9B355F792377E7AD6083EF71BD87A19" pageId="0" pageNumber="145">
<emphasis id="B1B4D3E39976DE0177B82AB01AE26C27" bold="true" pageId="0" pageNumber="145">Figure 10.</emphasis>
Epigynes of
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<emphasis italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
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<emphasis id="C2DDF008B5B86FA9F0F6431782F1F901" italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
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sp. nov. (
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<emphasis id="459C89D9239C9FF21C7A2C6A02FED11A" bold="true" pageId="0" pageNumber="145">A-E</emphasis>
) and
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<emphasis italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
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<emphasis id="2246B5D492F3C8E7D3AE0D0323B509C5" italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
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sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">F-J</emphasis>
<emphasis id="EF9BB29F2C2203253F286F8A4DFABC94" bold="true" pageId="0" pageNumber="145">F-J</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, F</emphasis>
<emphasis id="0D874B34DDA34980B6BB3BA459935F4A" bold="true" pageId="0" pageNumber="145">A, F</emphasis>
intact, ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, J</emphasis>
<emphasis id="7DDF6F8A2F76A296832DA3B079272E3B" bold="true" pageId="0" pageNumber="145">B, J</emphasis>
posterior
<emphasis bold="true" pageId="0" pageNumber="145">C, G</emphasis>
<emphasis id="9D4A03E34AC3F83D104F4EB2CC10A63C" bold="true" pageId="0" pageNumber="145">C, G</emphasis>
macerated, ventral
<emphasis bold="true" pageId="0" pageNumber="145">D, E, H, I</emphasis>
<emphasis id="8CEA8F5430C18508630D9ACD4608DC17" bold="true" pageId="0" pageNumber="145">D, E, H, I</emphasis>
macerated, dorsal. Scale bars: 0.2 mm.
</paragraph>
</caption>
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<subSubSection pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="E2BF68B596F34C6894A43572B0168470" pageId="0" pageNumber="145" type="comments">
<paragraph id="2785376854BC6704A1D632D63FF70B48" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph id="E36EC4EA81645698AD413ECE8BA6A072" pageId="0" pageNumber="145">
Although this species is rather similar in general appearance to the generotype and the other species known from Iran, the bulb and cymbium conformation is very different from those of other species considered in
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<emphasis italics="true" pageId="0" pageNumber="145">Pax</emphasis>
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<emphasis id="F084086178C6FAD3DD2BF150E9FCA8AE" italics="true" pageId="0" pageNumber="145">Pax</emphasis>
</taxonomicName>
and most likely belongs to an undescribed genus.
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<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="D5107F62298DDB995517B1CE34077CA2" pageId="0" pageNumber="145" type="distribution">
<paragraph id="81FDC14758040DF14F4BA504FE4F71D8" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="09AD0C2B35941441BC7654607FCA8E26" pageId="0" pageNumber="145">
Known only from the type locality in Fars Province, southwestern Iran (Fig.
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<figureCitation id="394150865ED8F5BDB3ED207BF2F3353E" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). This is the easternmost record of the genus
<taxonomicName authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Pax</emphasis>
<taxonomicName id="0F202F09DCB601B369C896E48AFCFD18" authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="E1A395D5243E1516E653F3F45C6B91FB" italics="true" pageId="0" pageNumber="145">Pax</emphasis>
</taxonomicName>
.
</paragraph>

View file

@ -1,79 +1,79 @@
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<taxonomicName id="6E655B50FDDC7D13C10D4CB7B6FE958B" authorityName="Marusik" authorityYear="2011" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus gromovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="gromovi">Trygetus gromovi</taxonomicName>
Marusik, 2011: 30, f. 1-7 (♀).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in turkmenistan">
<paragraph pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="3074E72F660EC25345CF1C9EEA2F87FE" pageId="0" pageNumber="145" type="records in turkmenistan">
<paragraph id="61ABDF858F43021EFEDF064FC1ABB03A" pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph>
<paragraph id="D72974DC930EDD9ED1E45BBC15FB3BB9" pageId="0" pageNumber="145">
Mary (
<bibRefCitation author="Marusik, YM" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="145" pagination="29 - 31" refId="B12" refString="Marusik, YM, 2011. A new Trygetus species from Central Asia (Araneae: Zodariidae). Turkish Journal of Zoology 35 (1): 29 - 31" title="A new Trygetus species from Central Asia (Araneae: Zodariidae)." volume="35" year="2011">Marusik 2011</bibRefCitation>
<bibRefCitation id="FB6B2745719C75A97EE377BDEE468756" author="Marusik, YM" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="145" pagination="29 - 31" refId="B12" refString="Marusik, YM, 2011. A new Trygetus species from Central Asia (Araneae: Zodariidae). Turkish Journal of Zoology 35 (1): 29 - 31" title="A new Trygetus species from Central Asia (Araneae: Zodariidae)." volume="35" year="2011">Marusik 2011</bibRefCitation>
) (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
<figureCitation id="5D032D22073501C8B091AA6F3B0397A1" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
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<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">Known only from the type locality in Turkmenistan.</paragraph>
<subSubSection id="0266F1A1CABB45A4D4109C03F1AC4661" pageId="0" pageNumber="145" type="distribution">
<paragraph id="CE753CF9EE233B9E2B1BB959B7B3B655" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="0F8416A2B23B8D32D4024BC8789C6BB9" pageId="0" pageNumber="145">Known only from the type locality in Turkmenistan.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,96 +1,96 @@
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Subfamily
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Lachesaninae
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, 1991
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<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
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<paragraph id="8FC7885249691573CC3E27FFBF1D239A" pageId="0" pageNumber="145">Comments.</paragraph>
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The following four genera are currently considered in this subfamily, with the number of species known from each in parentheses:
<taxonomicName authorityName="Jocque" authorityYear="1995" class="Arachnida" family="Zodariidae" genus="Australutica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Australutica" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Australutica</emphasis>
<taxonomicName id="9520716391FE27C95A939CDD22E9A562" authorityName="Jocque" authorityYear="1995" class="Arachnida" family="Zodariidae" genus="Australutica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Australutica" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="581DC5283C641EB91E4429A8506BA164" italics="true" pageId="0" pageNumber="145">Australutica</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jocqué">Jocque</normalizedToken>
<normalizedToken id="458A0C161B9CD9DDFE6F0B81A9835AEB" originalValue="Jocqué">Jocque</normalizedToken>
, 1995 (6),
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<emphasis italics="true" pageId="0" pageNumber="145">Antillorena</emphasis>
<taxonomicName id="D64901D39C095F053FDC32B5033D7C8C" authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Antillorena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Antillorena" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="1C29E9DD39E3B9022C122A3CF4D4BB48" italics="true" pageId="0" pageNumber="145">Antillorena</emphasis>
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<normalizedToken originalValue="Jocqué">Jocque</normalizedToken>
<normalizedToken id="2D7DB7F34F47D8689D3F751D071FCDB3" originalValue="Jocqué">Jocque</normalizedToken>
, 1991 (4),
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<emphasis italics="true" pageId="0" pageNumber="145">Lachesana</emphasis>
<taxonomicName id="617723A4125A3442503EADED15127A2A" authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="146E7BEBEC7FE8C48B6BF700204D1970" italics="true" pageId="0" pageNumber="145">Lachesana</emphasis>
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Strand, 1932 (8) and
<taxonomicName authorityName="Marx" authorityYear="1891" class="Arachnida" family="Zodariidae" genus="Lutica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lutica" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Lutica</emphasis>
<taxonomicName id="958CC45898F36197CD8B6B1DD6CA33F8" authorityName="Marx" authorityYear="1891" class="Arachnida" family="Zodariidae" genus="Lutica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lutica" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="ADDA1DE3FEF6DD35F1BF5BDC24851CD3" italics="true" pageId="0" pageNumber="145">Lutica</emphasis>
</taxonomicName>
Marx, 1891 (4).
<taxonomicName authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana</emphasis>
<taxonomicName id="9B555FF05628E7089B010B61A478057C" authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="D0C4D7AAEEE8A5384B25FB1332E05544" italics="true" pageId="0" pageNumber="145">Lachesana</emphasis>
</taxonomicName>
is the only one restricted to the western Palaearctic (
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2008
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;
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).
</paragraph>
</subSubSection>

View file

@ -1,132 +1,132 @@
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<treatment ID-GBIF-Taxon="181753565" LSID="urn:lsid:zoobank.org:act:DB25B91A-E827-4482-A2D4-1EC1D5012BF2" httpUri="http://treatment.plazi.org/id/3027AB9446A65E6DA6A64FCE757D5270" lastPageNumber="145" pageId="0" pageNumber="145">
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<taxonomicName LSID="http://zoobank.org/DB25B91A-E827-4482-A2D4-1EC1D5012BF2" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium ovtchinnikovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ovtchinnikovi" status="sp. nov.">Acanthinozodium ovtchinnikovi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">Figs 22D, E, G-I</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
<treatment id="3027AB9446A65E6DA6A64FCE757D5270" ID-GBIF-Taxon="181753565" LSID="urn:lsid:zoobank.org:act:DB25B91A-E827-4482-A2D4-1EC1D5012BF2" httpUri="http://treatment.plazi.org/id/3027AB9446A65E6DA6A64FCE757D5270" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection id="D8888DE4918D27180DDF7120B35EB92E" pageId="0" pageNumber="145" type="nomenclature">
<paragraph id="4F1CEBFC651B3D50C630ADC873666C82" pageId="0" pageNumber="145">
<taxonomicName id="A0125E350D632CAD4E4D14979A439BDE" ID-CoL="98YZ2" LSID="http://zoobank.org/DB25B91A-E827-4482-A2D4-1EC1D5012BF2" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium ovtchinnikovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ovtchinnikovi" status="sp. nov.">Acanthinozodium ovtchinnikovi</taxonomicName>
<taxonomicNameLabel id="05C956309A0F2AE1BAF5905AF798D31C" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="9D061F030F4119B5C324E76ED23EB9F7" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">Figs 22D, E, G-I</figureCitation>
<figureCitation id="600256BE01A5C1B229BA16F11EA9FF59" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="CB59F8FBEDC5E5E5B9A1CB9088EB9EB2" pageId="0" pageNumber="145" type="type material">
<paragraph id="623E39AFC6E956EB7EC88A73E0AFA7C3" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="90F8615E100DED7C7C991BB4F1254331" pageId="0" pageNumber="145">
<emphasis id="73159058B830D65A259D44BB834B3034" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ palp (ZMMU), Turkmenistan:
<emphasis italics="true" pageId="0" pageNumber="145">Mary Region</emphasis>
<emphasis id="A29F533C148205F07D57F20BDA5DD1F8" italics="true" pageId="0" pageNumber="145">Mary Region</emphasis>
: Kushka Dist., ca 18 km S of Kyzyldzhar Kordon, ca 1 km ESE of Eroilandaz,
<geoCoordinate degrees="35" direction="north" minutes="39" orientation="latitude" precision="925" value="35.65">35°39'N</geoCoordinate>
<geoCoordinate id="B9A6E69750D8777423AE246B8FE36D65" degrees="35" direction="north" minutes="39" orientation="latitude" precision="925" value="35.65">35°39'N</geoCoordinate>
,
<geoCoordinate degrees="61" direction="east" minutes="50" orientation="longitude" precision="925" value="61.833332">61°50'E</geoCoordinate>
<geoCoordinate id="2729AD70DFBEBE64EF435916B8D0D0FD" degrees="61" direction="east" minutes="50" orientation="longitude" precision="925" value="61.833332">61°50'E</geoCoordinate>
, 7.04.2002 (A.V. Gromov).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The new species is named after our late colleague Sergei V. Ovtchinnikov who made important contributions to the study of Central Asian spiders.</paragraph>
<subSubSection id="68AD5E69E842F14E6CDB114BA4D8BBE4" pageId="0" pageNumber="145" type="etymology">
<paragraph id="E133D69E2A620177BFECE4DE4C2537AC" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="A99CF2D1248B8BF692CF458C1B9DAF58" pageId="0" pageNumber="145">The new species is named after our late colleague Sergei V. Ovtchinnikov who made important contributions to the study of Central Asian spiders.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="A4C620B1A5DE69CBA620E3A802147FEE" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="CEB685AB86427B19AE6F51D217391B1F" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="0DA0B7F31A8097FCC0F33F0AAAA27AF5" pageId="0" pageNumber="145">
The new species differs from the congeners in the region (except
<taxonomicName lsidName="A. parmida" pageId="0" pageNumber="145" rank="species" species="parmida">
<emphasis italics="true" pageId="0" pageNumber="145">A. parmida</emphasis>
<taxonomicName id="CBBC3264EE5E193D05F721C49FB67D32" lsidName="A. parmida" pageId="0" pageNumber="145" rank="species" species="parmida">
<emphasis id="FF2C66D3A1D738111E31D82071A0F53A" italics="true" pageId="0" pageNumber="145">A. parmida</emphasis>
</taxonomicName>
sp. nov.) by the small size of the palp (cymbium 0.28 long,
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="B645E842BDA028DD48B30F0707B30B54" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
&gt; 0.6) and the unique embolus which has a long furrow with a serrate ventral margin (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22E</figureCitation>
<figureCitation id="B8D74EFF48D8F9C8E626104F54C779DA" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22E</figureCitation>
). It is closely related to
<taxonomicName lsidName="A. parmida" pageId="0" pageNumber="145" rank="species" species="parmida">
<emphasis italics="true" pageId="0" pageNumber="145">A. parmida</emphasis>
<taxonomicName id="64E563C84CB5AF027166162CB42C4E73" lsidName="A. parmida" pageId="0" pageNumber="145" rank="species" species="parmida">
<emphasis id="CD957D44EC34174285DE77CB592CC3CE" italics="true" pageId="0" pageNumber="145">A. parmida</emphasis>
</taxonomicName>
sp. nov. from central Iran, from which it can be differentiated by having a longer than wide bulb (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="29BCB8C483DA28958D8EEE5CABE299D6" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
almost as long as wide), relatively longer cymbium (length/width ratio 1.86 vs. 1.4), and a different shape of median apophysis (anterior portion larger than posterior one,
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="E5225822C94D1EC226666459A6686BAA" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
opposite; cf. Figs
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22G</figureCitation>
<figureCitation id="F14A9CA0A06EBAC231FBC018524D31A7" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22G</figureCitation>
and
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19B</figureCitation>
<figureCitation id="05EEB41F374DBEE2F4F03E8DE2F4CF9C" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
<subSubSection id="7C682029776913AF75F7D9C5798A15D2" pageId="0" pageNumber="145" type="description">
<paragraph id="CFEFA68E68832D40380E02A331A4D7CB" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="EBB0325CC799DD619E4C7506913D5AD1" pageId="0" pageNumber="145">
<emphasis id="E48CE970C73C7F0B789E3BC21BF90EF6" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Body missing. It is assumed to be a very small zodariid based on the size of the palp.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="F95B4F0F8DE9B6040A0F91E01CDA776B" pageId="0" pageNumber="145">
Palp as in Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22D, E, G-I</figureCitation>
<figureCitation id="5A0D28334E1AD39B9271AF1734D4214F" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22D, E, G-I</figureCitation>
. Ventral tibial apophysis lacking; RTA almost triangular, slightly longer than tibia; cymbium longer than wide; sperm duct tracking margin of tegulum, lacking any turns; median apophysis longer than wide, with anterior part larger than posterior part; embolus originates at about 8:00 o'clock position with its terminal 2/3 having a longitudinal furrow (
<emphasis italics="true" pageId="0" pageNumber="145">Er</emphasis>
<emphasis id="4305FA83C3234FC00687B573DFE48A5A" italics="true" pageId="0" pageNumber="145">Er</emphasis>
) with a finely serrated ventral margin.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="6F816B27751EAC062B03E68EED95E2FB" pageId="0" pageNumber="145">
<emphasis id="C2FB76EE65788440B7F4FEEEDD25A9E2" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="notes">
<paragraph pageId="0" pageNumber="145">Note.</paragraph>
<paragraph pageId="0" pageNumber="145">The sample collected by Gromov contained three males. The palp of one specimen was dissected and imaged with a SEM in 2005. Then, all three specimens were given to Ovtchinnikov who was planning to revise this group. After his death, his collection was transferred to Almaty, Kazakhstan by Alexander Gromov and is now inaccessible.</paragraph>
<subSubSection id="4EBC796BACE286CF8385164728A838F8" pageId="0" pageNumber="145" type="notes">
<paragraph id="86283D0F22B28CD4F41EABCFCDD87E48" pageId="0" pageNumber="145">Note.</paragraph>
<paragraph id="8969DA625D0310C5234095DACE874564" pageId="0" pageNumber="145">The sample collected by Gromov contained three males. The palp of one specimen was dissected and imaged with a SEM in 2005. Then, all three specimens were given to Ovtchinnikov who was planning to revise this group. After his death, his collection was transferred to Almaty, Kazakhstan by Alexander Gromov and is now inaccessible.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="EB8C00CB3818D0402BC36464F5C3A82E" pageId="0" pageNumber="145" type="distribution">
<paragraph id="3E305FB09ACA7BF817416655EA36782E" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="FA524AC25D13F726BFD1E98C794CB3CE" pageId="0" pageNumber="145">
Known only from the type locality in Mary Region, southeastern Turkmenistan (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
<figureCitation id="12F060B80CFBCDDDC4667E0812296523" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,99 +1,99 @@
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Zodarion buettikeri (Ono &amp;
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<normalizedToken id="BBD88E646188A112E3FC0E7EB751385D" originalValue="Jocqué">Jocque</normalizedToken>
, 1986)
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">Fig. 33</figureCitation>
<figureCitation id="DBFEF7F02E2A0C667FADA3F13AB6AA20" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">Fig. 33</figureCitation>
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<taxonomicName class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium buettikeri" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="buettikeri">Acanthinozodium buettikeri</taxonomicName>
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<taxonomicName id="07FF28A531FB12F0DB8B5FDB9EED3472" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium buettikeri" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="buettikeri">Acanthinozodium buettikeri</taxonomicName>
Ono &amp;
<normalizedToken originalValue="Jocqué">Jocque</normalizedToken>
<normalizedToken id="2FF1BF596483418B5C328A277E5F5E19" originalValue="Jocqué">Jocque</normalizedToken>
, 1986: 7, f. 1-4 (♂♀).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName baseAuthorityName="Ono &amp; Jocque" baseAuthorityYear="1986" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion buettikeri" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="buettikeri">Zodarion buettikeri</taxonomicName>
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<taxonomicName id="1E3DC52D0B336E3435B605586A1E95BE" baseAuthorityName="Ono &amp; Jocque" baseAuthorityYear="1986" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion buettikeri" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="buettikeri">Zodarion buettikeri</taxonomicName>
:
<bibRefCitation author="Levy, G" journalOrPublisher="Israel Journal of Zoology" pageId="0" pageNumber="145" pagination="67 - 110" refId="B11" refString="Levy, G, 1992. The spider genera Palaestina, Trygetus, Zodarion and Ranops (Araneae, Zodariidae) in Israel with annotations on species of the Middle East. Israel Journal of Zoology 38: 67 - 110" title="The spider genera Palaestina, Trygetus, Zodarion and Ranops (Araneae, Zodariidae) in Israel with annotations on species of the Middle East." volume="38" year="1992">Levy 1992</bibRefCitation>
<bibRefCitation id="BB60855DEFFB308C0A30CFC73CE877CD" author="Levy, G" journalOrPublisher="Israel Journal of Zoology" pageId="0" pageNumber="145" pagination="67 - 110" refId="B11" refString="Levy, G, 1992. The spider genera Palaestina, Trygetus, Zodarion and Ranops (Araneae, Zodariidae) in Israel with annotations on species of the Middle East. Israel Journal of Zoology 38: 67 - 110" title="The spider genera Palaestina, Trygetus, Zodarion and Ranops (Araneae, Zodariidae) in Israel with annotations on species of the Middle East." volume="38" year="1992">Levy 1992</bibRefCitation>
: 85;
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation>
<bibRefCitation id="E8D2C3684478B782709DE58D26127A8E" DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation>
: 69, f. 3E-F (♂).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in iran">
<paragraph pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="CF9988D900DA81B5B291DE9A98DD5E90" pageId="0" pageNumber="145" type="records in iran">
<paragraph id="135008EA9F56B237E9E441A8C990CD5C" pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph id="A7FA9406077087A40C4FACE4C30CCB37" pageId="0" pageNumber="145">
Bushehr (
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation>
<bibRefCitation id="96B8362315094D379D11DA6E70E026C0" DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation>
) (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
<figureCitation id="F36E06291829CA93177A573EF7A41B76" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">Saudi Arabia, Iran.</paragraph>
<subSubSection id="7D834D27B35E0533B61F8C3D13FEE84D" pageId="0" pageNumber="145" type="distribution">
<paragraph id="B33B1042ADB2C4CD1BE621B41BF85DA5" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="D45C34882484722D2D2EF54E499EB049" pageId="0" pageNumber="145">Saudi Arabia, Iran.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="6E2A682E08CD963B18C1D34C2902316A" pageId="0" pageNumber="145" type="comments">
<paragraph id="8FE338D1FA9D7540295CE3C05F10E599" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph id="B43C42C664D5009D79F244D181E3573E" pageId="0" pageNumber="145">
Since the Iranian material of this species has already been illustrated in
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. (2017)</bibRefCitation>
<bibRefCitation id="C199319132749F820FF69B6C66896B1E" DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. (2017)</bibRefCitation>
, we are not providing new figures for it.
</paragraph>
</subSubSection>

View file

@ -1,147 +1,147 @@
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<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">Figs 26D</figureCitation>
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
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<subSubSection id="A3BCB1B11328530D6A322CFB5ECFB5B0" pageId="0" pageNumber="145" type="nomenclature">
<paragraph id="3EF98410216CEC071107F7CA92A8CC2C" pageId="0" pageNumber="145">
<taxonomicName id="0FE2E3659FF00E71A09299F118F1CC6D" ID-CoL="98YZB" LSID="http://zoobank.org/4E56657F-4A5F-48F2-A3B4-E306128A62E6" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus susianus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="susianus" status="sp. nov.">Trygetus susianus</taxonomicName>
<taxonomicNameLabel id="A95A7F71AC860B352E6081E79781AD78" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="48D40055C363690865CB0D3C8F904705" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">Figs 26D</figureCitation>
<figureCitation id="E69100EB89559B4873175A59F211ACAF" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Epigynes of Parazodarion raddei (A-D) and Trygetus susianus sp. nov. (E-H) A, E intact, ventral B, F-H macerated, dorsal C, D macerated, ventral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145">, 27E-H</figureCitation>
<figureCitation id="0FBF980B8BA605BA2ABDCAA7B65CBCB9" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="reference_group">
<paragraph pageId="0" pageNumber="145">
<taxonomicName authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus jacksoni" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="jacksoni" status="sp. nov.">Trygetus jacksoni</taxonomicName>
<subSubSection id="E2D11B060B305AB246E5DCDFB1F748B6" pageId="0" pageNumber="145" type="reference_group">
<paragraph id="31F4F98DD3303AD0B988F484FCD97864" pageId="0" pageNumber="145">
<taxonomicName id="4420B8C7DFE811BF39B4FC6CE3799C44" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus jacksoni" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="jacksoni" status="sp. nov.">Trygetus jacksoni</taxonomicName>
:
<bibRefCitation author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. 2018</bibRefCitation>
<bibRefCitation id="3021A7ED0779DD9BEDE3FB07A36B64A2" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. 2018</bibRefCitation>
: 188 (♀, misidentified).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="84A74D2ABA0E17B3DBE60D63FFA3CB28" pageId="0" pageNumber="145" type="type material">
<paragraph id="E06F65BCA04E4A369E28D7D089D1FC18" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="802D6FD00579F0C3F09FB0CAD473190B" pageId="0" pageNumber="145">
<emphasis id="DC770590FA6CFDFA8DCD91FD074C77EC" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♀ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Khuzestan Province</emphasis>
<emphasis id="AF1DDCC1FB84103EF2952CA0F81401C6" italics="true" pageId="0" pageNumber="145">Khuzestan Province</emphasis>
: north of Andimeshk,
<geoCoordinate degrees="32" direction="north" minutes="41" orientation="latitude" precision="925" value="32.683334">32°41'N</geoCoordinate>
<geoCoordinate id="D03E165C63A4637CFFAE02E1712E2B1C" degrees="32" direction="north" minutes="41" orientation="latitude" precision="925" value="32.683334">32°41'N</geoCoordinate>
,
<geoCoordinate degrees="48" direction="east" minutes="15" orientation="longitude" precision="925" value="48.25">48°15'E</geoCoordinate>
<geoCoordinate id="11342565ACD0FB2784456B9B1A54BA20" degrees="48" direction="east" minutes="15" orientation="longitude" precision="925" value="48.25">48°15'E</geoCoordinate>
, 17.05.1974 (A. Senglet).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">This species is named after Susa, one of the most important cities of the Ancient Near East. It is located in the lower Zagros Mountains, about 250 km east of the Tigris River, between the Karkheh and Dez Rivers, in what is currently the Khuzestan Province of Iran.</paragraph>
<subSubSection id="504E233995CFBAF92EAA8B2AEA14A06A" pageId="0" pageNumber="145" type="etymology">
<paragraph id="7AC54454E20E7D0DD643EC5BAEB40C09" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="FAA8A3E7E32F7C4048CF05EF65DD3AE5" pageId="0" pageNumber="145">This species is named after Susa, one of the most important cities of the Ancient Near East. It is located in the lower Zagros Mountains, about 250 km east of the Tigris River, between the Karkheh and Dez Rivers, in what is currently the Khuzestan Province of Iran.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="2D0B20BE488F9094DEF26117A1019537" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="F62FBEC78D88CEE616E9646CA418011C" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="762936A75A2EB20F99B4C0FA769ECE1C" pageId="0" pageNumber="145">
The epigyne of the new species is very similar to that of
<taxonomicName lsidName="T. gromovi" pageId="0" pageNumber="145" rank="species" species="gromovi">
<emphasis italics="true" pageId="0" pageNumber="145">T. gromovi</emphasis>
<taxonomicName id="A37827C56581D3D0B7D086A4B6C630B6" lsidName="T. gromovi" pageId="0" pageNumber="145" rank="species" species="gromovi">
<emphasis id="C51F255739488B4057D73651B45F5AC5" italics="true" pageId="0" pageNumber="145">T. gromovi</emphasis>
</taxonomicName>
. They differ by the presence of a lateral extension of the receptacle and the lack of sclerotized lateral margins in the new species (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="6C9E3FE1821A1B8608FE11B227A60CC7" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
absent and present, respectively).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female</emphasis>
<subSubSection id="7F306A9B2920F7F391727D57F22289E1" pageId="0" pageNumber="145" type="description">
<paragraph id="A109B64C0E7520850728A7A792D15FA9" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="F7A3246B2B9AAB9F47C3D3671A612F20" pageId="0" pageNumber="145">
<emphasis id="B8C19FED2F4ADBE1A3052367184755B8" bold="true" pageId="0" pageNumber="145">Female</emphasis>
(specimen partially bleached). Habitus as in Fig.
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">26D</figureCitation>
<figureCitation id="01D4F5030418AD0C689643C115E198B8" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">26D</figureCitation>
. Total length 2.07. Carapace 0.96 long, 0.53 wide at pars cephalica, 0.69 wide at pars thoracica. Eye sizes: AME: 0.08, ALE: 0.05, PLE: 0.04. Carapace, sternum, labium, chelicerae and maxillae yellowish brown, without any pattern. Legs slightly lighter than carapace, without spines and without annulations. Abdomen grayish, darker dorsally. Spinnerets pale, uniform in color. Measurements of legs: I: 2.19 (0.61, 0.29, 0.44, 0.44, 0.41), II: 1.98 (0.58, 0.28, 0.39, 0.36, 0.37), III: 1.90 (0.52, 0.26, 0.32, 0.40, 0.40), IV: 2.50 (0.67, 0.28, 0.48, 0.63, 0.44).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="AFDE0792F6F4C7E492ACC00FA25653BB" pageId="0" pageNumber="145">
Epigyne as in Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Epigynes of Parazodarion raddei (A-D) and Trygetus susianus sp. nov. (E-H) A, E intact, ventral B, F-H macerated, dorsal C, D macerated, ventral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145">27E-H</figureCitation>
<figureCitation id="C0E736EF700D1DEB6C302A2A404F7538" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Epigynes of Parazodarion raddei (A-D) and Trygetus susianus sp. nov. (E-H) A, E intact, ventral B, F-H macerated, dorsal C, D macerated, ventral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145">27E-H</figureCitation>
. Epigyne medially with posteriorly diverging sclerotization, laterally with translucent fertilization ducts; receptacles almost round, separated by less than their diameters, each subdivided into a large, weakly sclerotized part and a smaller, more heavily sclerotized part.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145" start="Figure 27" startId="F27">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 27.</emphasis>
<caption id="F3679EEFF180C068D1CED1590333FC8C" doi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145" start="Figure 27" startId="F27">
<paragraph id="2D26797137DE03E48A325541C2024E39" pageId="0" pageNumber="145">
<emphasis id="49655540CA742A0A8CD3838D7C0D012E" bold="true" pageId="0" pageNumber="145">Figure 27.</emphasis>
Epigynes of
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion raddei" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei">
<emphasis italics="true" pageId="0" pageNumber="145">Parazodarion raddei</emphasis>
<taxonomicName id="20DE22DE7D4E552D156D079DAF75B61C" baseAuthorityName="Simon" baseAuthorityYear="1889" family="Zodariidae" genus="Parazodarion" kingdom="Animalia" lsidName="Parazodarion raddei" order="Aranei" pageId="0" pageNumber="145" rank="species" species="raddei">
<emphasis id="7E0AFE395F7B1ED9C0D5AF9EDB1366BE" italics="true" pageId="0" pageNumber="145">Parazodarion raddei</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="145">A-D</emphasis>
<emphasis id="C715F552A6BF7579867DA882EA734E5B" bold="true" pageId="0" pageNumber="145">A-D</emphasis>
) and
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus susianus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="susianus">
<emphasis italics="true" pageId="0" pageNumber="145">Trygetus susianus</emphasis>
<taxonomicName id="94BEDA40FAED9F6E8F0B3F6ADDAB2FD2" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus susianus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="susianus">
<emphasis id="1AF011167D88E5A4F8599A7B42BDE0C8" italics="true" pageId="0" pageNumber="145">Trygetus susianus</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">E-H</emphasis>
<emphasis id="F3CDB45AD1FC32BDC26FE788DB277A9E" bold="true" pageId="0" pageNumber="145">E-H</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, E</emphasis>
<emphasis id="A00A9E3B00E508831D8CCB3F84512A31" bold="true" pageId="0" pageNumber="145">A, E</emphasis>
intact, ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, F-H</emphasis>
<emphasis id="B82CB123549762110DEF91A67CCAB280" bold="true" pageId="0" pageNumber="145">B, F-H</emphasis>
macerated, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">C, D</emphasis>
<emphasis id="80207104489B8E8D8651664D3FCAD40D" bold="true" pageId="0" pageNumber="145">C, D</emphasis>
macerated, ventral. Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
<paragraph id="3BCCB640FBA7FDBDF3142D3817D4A754" pageId="0" pageNumber="145">
<emphasis id="E3C9FA4D06C9E6B7F8CBDA9C065DE7AA" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="D3B41B8C68B48840D127CE85ACF7766D" pageId="0" pageNumber="145" type="distribution">
<paragraph id="14032BB7B8B3F0A2CDEC4CE2DA2EDE63" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="66A318A4FC1970F922BAEAD3B76BF95F" pageId="0" pageNumber="145">
Known from two localities in Khuzestan Province, southwestern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
<figureCitation id="2ADC834748329E3497CD5250189B3CA4" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,278 +1,278 @@
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<mods:namePart>Zamani, Alireza</mods:namePart>
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<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
<mods:nameIdentifier type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
<mods:namePart id="55D20788F1CAD38D31E0B60E3158A2E5">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="F74B586DEDB4CA22B86B7E9D410C6C27">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
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</mods:role>
<mods:namePart>Marusik, Yuri M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4499-5148</mods:nameIdentifier>
<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
<mods:namePart id="45C95BC9740DEE534BB5C8930AFE646D">Marusik, Yuri M.</mods:namePart>
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<mods:affiliation id="667152E583F848652679AAD8BB39F815">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<mods:title id="177FA9CD5696095AF9C8B8FEFCC89DCC">ZooKeys</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>1035</mods:number>
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<treatment ID-GBIF-Taxon="181753571" LSID="urn:lsid:zoobank.org:act:D52287DC-6ED8-4E56-B435-F5CB56650469" httpUri="http://treatment.plazi.org/id/3D04C9134C3357AF827C326761DF8161" lastPageNumber="145" pageId="0" pageNumber="145">
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/D52287DC-6ED8-4E56-B435-F5CB56650469" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita" status="sp. nov.">Pax ellipita</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">Figs 5A, B</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male palp of Pax ellipita sp. nov. A ventral B, C retrolateral D, E retrodorsal and prolateral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. SEM images of the male palp of Pax ellipita sp. nov. A retrolateral B anteroretrolateral C proventral D posterodorsal E ventral. Abbreviations: E 1, E 2 - pro- and retrolateral cymbial extensions, Eb - embolar base, Em - embolus, Cf - cymbial fold, Co - conductor, Ma - median apophysis, Ra - retrolateral apophysis, Va - ventral apophysis. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145">, 7</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">, 10A-E</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
<treatment id="3D04C9134C3357AF827C326761DF8161" ID-GBIF-Taxon="181753571" LSID="urn:lsid:zoobank.org:act:D52287DC-6ED8-4E56-B435-F5CB56650469" httpUri="http://treatment.plazi.org/id/3D04C9134C3357AF827C326761DF8161" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Storeninae">
<subSubSection id="B8EF491F9B8E49D577FAE4C3EB164330" pageId="0" pageNumber="145" type="nomenclature">
<paragraph id="5DD65436067242D7DFE59FF67CF05956" pageId="0" pageNumber="145">
<taxonomicName id="4787CDF2B73D0908C79E07BD5ACD8B19" ID-CoL="98YZ5" LSID="http://zoobank.org/D52287DC-6ED8-4E56-B435-F5CB56650469" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita" status="sp. nov.">Pax ellipita</taxonomicName>
<taxonomicNameLabel id="4E1D86037F50AEE3366F396D40ED649D" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="84E29CF178B42E2C4341CFD98F1102B1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">Figs 5A, B</figureCitation>
<figureCitation id="A3ED547FA882A6F33E2077169CCDDB1C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male palp of Pax ellipita sp. nov. A ventral B, C retrolateral D, E retrodorsal and prolateral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145">, 6</figureCitation>
<figureCitation id="07EBA79E1AE5DB94B5B7E919E764F54F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. SEM images of the male palp of Pax ellipita sp. nov. A retrolateral B anteroretrolateral C proventral D posterodorsal E ventral. Abbreviations: E 1, E 2 - pro- and retrolateral cymbial extensions, Eb - embolar base, Em - embolus, Cf - cymbial fold, Co - conductor, Ma - median apophysis, Ra - retrolateral apophysis, Va - ventral apophysis. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145">, 7</figureCitation>
<figureCitation id="77A312715848D1E3C25B030D6E8C70B6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">, 10A-E</figureCitation>
<figureCitation id="1FEC49BC8BAE262A375CC0EC4A253A9B" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="EC14C324732A0F42DA3E4EB31D04D8DD" pageId="0" pageNumber="145" type="type material">
<paragraph id="900A73ED5341BDDDCEBFFE04204251B5" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="46A8702CB5303F32A7C93754C5C15ACE" pageId="0" pageNumber="145">
<emphasis id="5BF8A099EF7F1194B6988DE55BBD246C" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Kermanshah Province</emphasis>
<emphasis id="869DB96280192DA74EFDBE6A61A6976E" italics="true" pageId="0" pageNumber="145">Kermanshah Province</emphasis>
: north of Kermanshah,
<geoCoordinate degrees="34" direction="north" minutes="28" orientation="latitude" precision="925" value="34.466667">34°28'N</geoCoordinate>
<geoCoordinate id="BEFF1B1B545027CEA9F09FA9A6311117" degrees="34" direction="north" minutes="28" orientation="latitude" precision="925" value="34.466667">34°28'N</geoCoordinate>
,
<geoCoordinate degrees="47" direction="east" minutes="00" orientation="longitude" precision="925" value="47.0">47°00'E</geoCoordinate>
<geoCoordinate id="E7AD95E7ED71201D36E84507D766B717" degrees="47" direction="east" minutes="00" orientation="longitude" precision="925" value="47.0">47°00'E</geoCoordinate>
, 18.06.1975 (A. Senglet).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="materials_examined">
<paragraph pageId="0" pageNumber="145">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<materialsCitation ID-GBIF-Occurrence="3093634308" collectingDate="1977-04-11" collectorName="B. Prazan" country="Iran" latitude="32.85" location="river Saimareh" longLatPrecision="1208" longitude="47.933334" municipality="Pol-e Tang" specimenCount="1" specimenCount-female="1" stateProvince="Lorestan Province">
<collectingCountry name="Iran">Iran</collectingCountry>
<subSubSection id="05CC8E29FFE0718000E9FD0CA084BFD4" pageId="0" pageNumber="145" type="materials_examined">
<paragraph id="DF76CD51F0481726BE6DE8DC5E65931F" pageId="0" pageNumber="145">Additional material.</paragraph>
<paragraph id="58124689C0592D5E511534EAD839DF1B" pageId="0" pageNumber="145">
<materialsCitation id="85A1406004A9103FC355D151ADB55CAC" ID-GBIF-Occurrence="3093634308" collectingDate="1977-04-11" collectorName="B. Prazan" country="Iran" latitude="32.85" location="river Saimareh" longLatPrecision="1208" longitude="47.933334" municipality="Pol-e Tang" specimenCount="1" specimenCount-female="1" stateProvince="Lorestan Province">
<collectingCountry id="114FB4C83AB694210D7E96E47F46A3FB" name="Iran">Iran</collectingCountry>
:
<emphasis italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Lorestan">Lorestan Province</collectingRegion>
<emphasis id="6B8F74AA2E103A05F163B285727F2376" italics="true" pageId="0" pageNumber="145">
<collectingRegion id="0986DAEC94B11F67DD13D77455D1EC82" country="Iran" name="Lorestan">Lorestan Province</collectingRegion>
</emphasis>
:
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="AE619007CD3437A402C2CA24AAA37F63" type="female">1♀</specimenCount>
(NMP),
<collectingMunicipality>Pol-e Tang</collectingMunicipality>
<collectingMunicipality id="478FAF9D98D3524414C00C370D4D7177">Pol-e Tang</collectingMunicipality>
,
<locationDeviation location="Andimeshk">
<quantity metricMagnitude="4" metricUnit="m" metricValue="6.0" unit="km" value="60.0">60 km</quantity>
<locationDeviation id="5F71721AD0CA0389EDFB4B86EAC252E0" location="Andimeshk">
<quantity id="B359E673989041A749B5377713E2E46E" metricMagnitude="4" metricUnit="m" metricValue="6.0" unit="km" value="60.0">60 km</quantity>
NW of Andimeshk
</locationDeviation>
,
<geoCoordinate degrees="32" direction="north" minutes="51" orientation="latitude" precision="925" value="32.85">32°51'N</geoCoordinate>
<geoCoordinate id="BD04B59C4D28FF919B25D29FFC19755D" degrees="32" direction="north" minutes="51" orientation="latitude" precision="925" value="32.85">32°51'N</geoCoordinate>
,
<geoCoordinate degrees="47" direction="east" minutes="56" orientation="longitude" precision="925" value="47.933334">47°56'E</geoCoordinate>
<geoCoordinate id="432531442F29D18302C181D03A7FFED9" degrees="47" direction="east" minutes="56" orientation="longitude" precision="925" value="47.933334">47°56'E</geoCoordinate>
, near the
<location LSID="urn:lsid:plazi:treatment:3D04C9134C3357AF827C326761DF8161:71B04B1A5985BF7FCAC1BD1C1C6158A6" country="Iran" latitude="32.85" longLatPrecision="1208" longitude="47.933334" municipality="Pol-e Tang" name="river Saimareh" stateProvince="Lorestan Province">river Saimareh</location>
<location id="71B04B1A5985BF7FCAC1BD1C1C6158A6" LSID="urn:lsid:plazi:treatment:3D04C9134C3357AF827C326761DF8161:71B04B1A5985BF7FCAC1BD1C1C6158A6" country="Iran" latitude="32.85" longLatPrecision="1208" longitude="47.933334" municipality="Pol-e Tang" name="river Saimareh" stateProvince="Lorestan Province">river Saimareh</location>
,
<collectingDate value="1977-04-11">11.04.1977</collectingDate>
<collectingDate id="0CB8554777EE6D42E577014CAEF3CBB2" value="1977-04-11">11.04.1977</collectingDate>
(
<collectorName>
<collectorName id="C1948061F7D6881F5B8160D89AD5F5BE">
B.
<normalizedToken originalValue="Pražan">Prazan</normalizedToken>
<normalizedToken id="53E7B174EB3073FE68C387C839F3D8C0" originalValue="Pražan">Prazan</normalizedToken>
</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet refers to Ellipi, an ancient kingdom located on the western side of the Zagros Mountains, between Babylonia at the west, Media at the north-east, Mannae at the north and Elam at the south.</paragraph>
<subSubSection id="A103B63F8C68DF75C3FF07DE3F2AB00B" pageId="0" pageNumber="145" type="etymology">
<paragraph id="B51FC1D3977CD5F14E3E44BAB9C482A9" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="6E862DD21B7A359C739BD5146F77E816" pageId="0" pageNumber="145">The specific epithet refers to Ellipi, an ancient kingdom located on the western side of the Zagros Mountains, between Babylonia at the west, Media at the north-east, Mannae at the north and Elam at the south.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="CDFA2F69DD4B693153B39847D94170EF" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="8568E266A8898EADA433C7D798811955" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="C03EA28D5896E613695CFFB916D01EB7" pageId="0" pageNumber="145">
The new species differs from
<taxonomicName lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila">
<emphasis italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
<taxonomicName id="F3E9162C5481726F5FFCA7DF1FF81475" lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila">
<emphasis id="5197A674C74CD9B07F1C953AE1B37517" italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
</taxonomicName>
sp. nov., the only other
<taxonomicName authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Pax</emphasis>
<taxonomicName id="1BB9CDB0DCB2A3A29F2BCED1BCB659F9" authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="F6F3D674A5E18706E8EE0709A72A2E26" italics="true" pageId="0" pageNumber="145">Pax</emphasis>
</taxonomicName>
species known from Iran, by the cymbium lacking a horn-like outgrowth (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="174E826A1043CF3A2B0543810074F778" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
present), by having a deep lateral fold of the cymbium (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="A428345B1C8A606390895F0EC1600D29" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
lacking) and the epigyne with a rectangular median plate (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="0F151811F4905EBBF5B734C49D8C4DEC" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
triangular). The male of
<taxonomicName lsidName="P. ellipita" pageId="0" pageNumber="145" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">P. ellipita</emphasis>
<taxonomicName id="405092E4B45B69D231A40DFAE8F6A544" lsidName="P. ellipita" pageId="0" pageNumber="145" rank="species" species="ellipita">
<emphasis id="0EB747DFAC90C8D80BAE9F84D3AA0176" italics="true" pageId="0" pageNumber="145">P. ellipita</emphasis>
</taxonomicName>
sp. nov. differs from the rest of the species by having the RTA shorter than the tibia (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="45589CE9127F3D6E5511F48DA7FA511A" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
longer), while the female differs by the trilobate posterior margin of the epigyne (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="8F7C44620E890CA23F944A29F3241F82" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
solid) and copulatory openings located posteriorly (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="6C8F3F4F40F57860B43A1F3DE04AD463" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
anteriorly).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
<subSubSection id="8C9D177F02B41C6199E5B3C180C1B2BB" pageId="0" pageNumber="145" type="description">
<paragraph id="B4D807EE5EC17DFD087A40C2F6A0B558" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="F9467B9BCC7A13150DA0F14B355B0FF8" pageId="0" pageNumber="145">
<emphasis id="DC418155A2CF8826BA96ABC53DF7A815" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5B</figureCitation>
<figureCitation id="F88D73B81895251271656C4A65C29AFD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5B</figureCitation>
. Total length 5.07. Carapace 2.43 long, 1.27 wide at pars cephalica, 1.65 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.11, ALE: 0.10, PME: 0.08, PLE: 0.10, PME-PME: 0.09. Carapace, sternum, labium, chelicerae and maxillae reddish brown, without any pattern. Chelicera with 2 promarginal teeth. Legs yellow, with few spines and without annulations. Abdomen dark brown, dorsally with large scutum covering almost the entire abdomen. Spinnerets pale, uniform in color. Measurements of legs: I: 5.49 (1.48, 0.62, 1.22, 1.10, 1.07), II: 4.88 (1.29, 0.60, 1.04, 1.05, 0.90), III: 4.74 (1.27, 0.61, 0.90, 1.19, 0.77), IV: 6.71 (1.80, 0.68, 1.39, 1.80, 1.04).
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 5.</emphasis>
<caption id="1DC138CDBEB1D6E579DDF4B26BB63867" doi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145" start="Figure 5" startId="F5">
<paragraph id="F7F808C62584DE5F191AE85269AB3914" pageId="0" pageNumber="145">
<emphasis id="9940F099B881D44282F9AEE37B2C42F4" bold="true" pageId="0" pageNumber="145">Figure 5.</emphasis>
Dorsal habitus of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
<taxonomicName id="5FFE42A9781C9199C508FFB4D0B2A41F" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
<emphasis id="2A884F0D0F6E6E78A631BAF5F3726F3D" italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A, B</emphasis>
<emphasis id="DF876837BE669C53744D6008FC5E86BD" bold="true" pageId="0" pageNumber="145">A, B</emphasis>
) and
<taxonomicName lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila">
<emphasis italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
<taxonomicName id="99147058216493ADA9462D732B0E2B70" lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila">
<emphasis id="3CED58E8C8CC22DDE719017F373F4258" italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">C, D</emphasis>
<emphasis id="CABE0C977B91978C3DBBD5387839E3CD" bold="true" pageId="0" pageNumber="145">C, D</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis>
<emphasis id="F77B7947986D25019AF3A881FC0621E6" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
females
<emphasis bold="true" pageId="0" pageNumber="145">B, C</emphasis>
<emphasis id="5669683DE18BB9909C847773EB83F81D" bold="true" pageId="0" pageNumber="145">B, C</emphasis>
males.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<paragraph id="DA59EFB89B5F0757AEAD0D36CA3BAA78" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male palp of Pax ellipita sp. nov. A ventral B, C retrolateral D, E retrodorsal and prolateral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145">6A-E</figureCitation>
<figureCitation id="439D1B46A3E9BA3526CDA723CE4C95E3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male palp of Pax ellipita sp. nov. A ventral B, C retrolateral D, E retrodorsal and prolateral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145">6A-E</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. SEM images of the male palp of Pax ellipita sp. nov. A retrolateral B anteroretrolateral C proventral D posterodorsal E ventral. Abbreviations: E 1, E 2 - pro- and retrolateral cymbial extensions, Eb - embolar base, Em - embolus, Cf - cymbial fold, Co - conductor, Ma - median apophysis, Ra - retrolateral apophysis, Va - ventral apophysis. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145">7A-E</figureCitation>
<figureCitation id="19A7CA425DE92DF2AD059A5E9B69C5D4" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. SEM images of the male palp of Pax ellipita sp. nov. A retrolateral B anteroretrolateral C proventral D posterodorsal E ventral. Abbreviations: E 1, E 2 - pro- and retrolateral cymbial extensions, Eb - embolar base, Em - embolus, Cf - cymbial fold, Co - conductor, Ma - median apophysis, Ra - retrolateral apophysis, Va - ventral apophysis. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145">7A-E</figureCitation>
. Femur slightly shorter than cymbium, 3.3 times longer than wide; patella globular; tibia wider than long with ventral apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis>
<emphasis id="3E865AAA7C2FCD45DB60AFE1CD33DDB7" italics="true" pageId="0" pageNumber="145">Va</emphasis>
) and short RTA bifurcated at the tip (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
<emphasis id="39F9E498225F863711F1CC4F90E3C8D0" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
); cymbium with large fold (
<emphasis italics="true" pageId="0" pageNumber="145">Cf</emphasis>
<emphasis id="97205EAD6FA40E9B507E111BE158956E" italics="true" pageId="0" pageNumber="145">Cf</emphasis>
) and 2 extensions, pro- and retrolateral (
<emphasis italics="true" pageId="0" pageNumber="145">E1</emphasis>
<emphasis id="54F1E04DCF1F89AB9F8704D87410DAA2" italics="true" pageId="0" pageNumber="145">E1</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="145">E2</emphasis>
<emphasis id="76FA5A6A59257E0233CE78C7089B09C0" italics="true" pageId="0" pageNumber="145">E2</emphasis>
); bulb with large lamella, as long as bulb, tapering to the tip; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="39288C442791F17C00BAD1F097851766" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) long, about 5 times longer than wide; conductor (
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis>
<emphasis id="4684AD63BEA5EEB6DF1F145CD0D4F9DE" italics="true" pageId="0" pageNumber="145">Co</emphasis>
) heavily sclerotized; embolus (
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
<emphasis id="982D429F9366AE39C7638F3B759294FF" italics="true" pageId="0" pageNumber="145">Em</emphasis>
) long, originates at 6
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
<normalizedToken id="68FB6C65B0090DAE047EE3E933892715" originalValue="oclock">o'clock</normalizedToken>
position.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 6.</emphasis>
<caption id="EF95E638690874A842A19B3F5147AB61" doi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145" start="Figure 6" startId="F6">
<paragraph id="4EFBD814475B1B8ADEF951125AFACDEC" pageId="0" pageNumber="145">
<emphasis id="E5E3FC8C67B350B38806645DBB3F9959" bold="true" pageId="0" pageNumber="145">Figure 6.</emphasis>
Male palp of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
<taxonomicName id="AF19F40617E25C36F85435B3DB26AA1C" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
<emphasis id="1A246894F03470C4EA88996A5EEB2142" italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
<emphasis id="CD12809AEA2312D730F93FEBA62A02CD" bold="true" pageId="0" pageNumber="145">A</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, C</emphasis>
<emphasis id="C1FB719A163E1078B7FF55EA6C7B75FB" bold="true" pageId="0" pageNumber="145">B, C</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">D, E</emphasis>
<emphasis id="6C4363BDB1ABD99D0466C079F6F18928" bold="true" pageId="0" pageNumber="145">D, E</emphasis>
retrodorsal and prolateral. Scale bars: 0.2 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 7.</emphasis>
<caption id="17199827B2FC18A0AAF0D0FE7D19CC73" doi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145" start="Figure 7" startId="F7">
<paragraph id="2DBA9ABF4D31C32527132B6B8D68836D" pageId="0" pageNumber="145">
<emphasis id="82182B536DADBF5FE219589DEEC203A0" bold="true" pageId="0" pageNumber="145">Figure 7.</emphasis>
SEM images of the male palp of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
<taxonomicName id="EDCF2A1FEAB00118B48D93D845F25721" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
<emphasis id="442A1278412D2991517BA066E4D71AEB" italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
<emphasis id="918FBBB333B7CC46625038276D2CC4C9" bold="true" pageId="0" pageNumber="145">A</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
<emphasis id="528D4500E8EB46EA73AAF86614D8CF57" bold="true" pageId="0" pageNumber="145">B</emphasis>
anteroretrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
<emphasis id="8271ED8609C131C03AEBB9CF32CF71B6" bold="true" pageId="0" pageNumber="145">C</emphasis>
proventral
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
<emphasis id="9850714E0E19C46B51388E2A7DEAAF93" bold="true" pageId="0" pageNumber="145">D</emphasis>
posterodorsal
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
<emphasis id="1F4680C995C71FF1BF205303C8C581DE" bold="true" pageId="0" pageNumber="145">E</emphasis>
ventral. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">E1</emphasis>
<emphasis id="6FD455BF9FE64CF90088FCE919E3A4CE" italics="true" pageId="0" pageNumber="145">E1</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="145">E2</emphasis>
<emphasis id="F5E456EBF18C20F2D749F243DFCC1988" italics="true" pageId="0" pageNumber="145">E2</emphasis>
- pro- and retrolateral cymbial extensions,
<emphasis italics="true" pageId="0" pageNumber="145">Eb</emphasis>
<emphasis id="73F44A3591D0F848AAADC2E43DBCABE8" italics="true" pageId="0" pageNumber="145">Eb</emphasis>
- embolar base,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
<emphasis id="8914E1CB388CFA311B12599B98E5B6EE" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Cf</emphasis>
<emphasis id="FCBB04742C43E8E1425F5D5BE16154CA" italics="true" pageId="0" pageNumber="145">Cf</emphasis>
- cymbial fold,
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis>
<emphasis id="F8AB0DE353115714D315D7D58D37219E" italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="665D67F229A17A3CE9265F7FDF34BA56" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
<emphasis id="1AB049F4EA75B6D1E8D6F373DE692B3E" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis>
<emphasis id="57D21FA36F097285395F6FAA56859437" italics="true" pageId="0" pageNumber="145">Va</emphasis>
- ventral apophysis. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="72A015393BA1AE5F020C0066547762A9" pageId="0" pageNumber="145">
<emphasis id="15F98AD0B5C468FE520E43E0D5F7A39A" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5A</figureCitation>
<figureCitation id="B3B8BD19BCB6CF8B443FD98ECDEF1A80" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5A</figureCitation>
. Total length 5.33. Carapace 2.18 long, 1.08 wide at pars cephalica, 1.51 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.06, ALE: 0.09, PME: 0.07, PLE: 0.09, PME-PME: 0.08. Coloration as in male, with lighter abdomen lacking a scutum. Measurements of legs: I: 5.36 (1.52, 0.60, 1.15, 1.09, 1.00), II: 4.76 (1.32, 0.61, 0.99, 1.04, 0.80), III: 4.53 (1.23, 0.60, 0.89, 1.11, 0.70), IV: 5.78 (1.42, 0.65, 1.38, 1.53, 0.80).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="3056C5EDBA73D20EC189F68B32DDFBC7" pageId="0" pageNumber="145">
Epigyne as in Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">10A-E</figureCitation>
<figureCitation id="BB633367CAF7C63B7DBC36FC87AB6411" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">10A-E</figureCitation>
. Trilobate, wider than long; median plate rectangular (with subparallel lateral margins), ca. 3.6 times thinner than lateral lobe; receptacles oval, wider than long, touching each other.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">Because the female specimen was not collected together with the male, its assignment to this species is tentative and shall be confirmed when both sexes are collected together.</paragraph>
<subSubSection id="1463D875C6E761EAABF14C813291E7A6" pageId="0" pageNumber="145" type="comments">
<paragraph id="30AE261AD5D57CC7B0DA59FA00FFDFAE" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph id="546F50C8ABB5B6EDCA88C54C6AB26D1A" pageId="0" pageNumber="145">Because the female specimen was not collected together with the male, its assignment to this species is tentative and shall be confirmed when both sexes are collected together.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="35D7C190BC4558823E9907D38154835B" pageId="0" pageNumber="145" type="distribution">
<paragraph id="AF1CD7107E797C8720D28EB8FD63AF4C" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="D8F0C6555133DF602D8C0C63843CB8ED" pageId="0" pageNumber="145">
Known from the listed localities in Kermanshah and Lorestan provinces, western Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
<figureCitation id="24678029B1CC77DD848DB4A51676B0D8" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,162 +1,162 @@
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<treatment ID-GBIF-Taxon="181753564" LSID="urn:lsid:zoobank.org:act:92A3E44F-1EC2-4246-A21E-CAEED957E5BE" httpUri="http://treatment.plazi.org/id/440E5E09922D5E9C8DE00B36BC3DA4CB" lastPageNumber="145" pageId="0" pageNumber="145">
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/92A3E44F-1EC2-4246-A21E-CAEED957E5BE" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parysatis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parysatis" status="sp. nov.">Acanthinozodium parysatis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11F, G</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">, 13F-I</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">, 15A-C</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">, 17D-F</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
<treatment id="440E5E09922D5E9C8DE00B36BC3DA4CB" ID-GBIF-Taxon="181753564" LSID="urn:lsid:zoobank.org:act:92A3E44F-1EC2-4246-A21E-CAEED957E5BE" httpUri="http://treatment.plazi.org/id/440E5E09922D5E9C8DE00B36BC3DA4CB" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection id="710C1096B1E6359288B4D30B22A78067" pageId="0" pageNumber="145" type="nomenclature">
<paragraph id="C3D6F054C8830C5E4C079B6D81F410E9" pageId="0" pageNumber="145">
<taxonomicName id="B0CF97B07CA1BB44A90A5D8C8E9B2249" ID-CoL="98YYZ" LSID="http://zoobank.org/92A3E44F-1EC2-4246-A21E-CAEED957E5BE" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parysatis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parysatis" status="sp. nov.">Acanthinozodium parysatis</taxonomicName>
<taxonomicNameLabel id="F3D56AAD7FFA81BAF42B4ED9B8F40702" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="16C8113191A4D31363E2DA799FF022E1" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11F, G</figureCitation>
<figureCitation id="8977A59A092409F205C8E15536F19661" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">, 13F-I</figureCitation>
<figureCitation id="759AA99EBA06D17C16169D3652250537" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">, 15A-C</figureCitation>
<figureCitation id="FAFA5C11B574C9993100C82D0DC6AE95" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">, 17D-F</figureCitation>
<figureCitation id="579B00F5B1542548955E1113085100DA" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="E4E57C35F9435247E6485E166E20F4C9" pageId="0" pageNumber="145" type="type material">
<paragraph id="C80CF9CF534A30FBB38A511CD2D4E563" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="01354254F13C81F772016A0E38275BCD" pageId="0" pageNumber="145">
<emphasis id="BA0486FADF27D48CD641D3642D2E10E3" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Ardabil Province</emphasis>
<emphasis id="A95D99A2435668CDA6F66DACF7FB037C" italics="true" pageId="0" pageNumber="145">Ardabil Province</emphasis>
: Kivi Pain,
<geoCoordinate degrees="37" direction="north" minutes="41" orientation="latitude" precision="925" value="37.683334">37°41'N</geoCoordinate>
<geoCoordinate id="78AB146C1718E9F7404819B0B32A427F" degrees="37" direction="north" minutes="41" orientation="latitude" precision="925" value="37.683334">37°41'N</geoCoordinate>
,
<geoCoordinate degrees="48" direction="east" minutes="21" orientation="longitude" precision="925" value="48.35">48°21'E</geoCoordinate>
<geoCoordinate id="25685867874E2E161EB1D162C3E64A9F" degrees="48" direction="east" minutes="21" orientation="longitude" precision="925" value="48.35">48°21'E</geoCoordinate>
, 09.06.1975 (A. Senglet).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
<emphasis id="92B61AF3E8499BF0F73D19CE31654251" bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
: 1♂ (MHNG), same data as holotype; 3♀1♂ palp (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Qazvin Province</emphasis>
<emphasis id="2F87DA422F92C032966BF2F8F10E2AEC" italics="true" pageId="0" pageNumber="145">Qazvin Province</emphasis>
: Shahrak,
<geoCoordinate degrees="36" direction="north" minutes="25" orientation="latitude" precision="925" value="36.416668">36°25'N</geoCoordinate>
<geoCoordinate id="E0B47AD025A6F9DD61BFDB02D48D779A" degrees="36" direction="north" minutes="25" orientation="latitude" precision="925" value="36.416668">36°25'N</geoCoordinate>
,
<geoCoordinate degrees="50" direction="east" minutes="30" orientation="longitude" precision="925" value="50.5">50°30'E</geoCoordinate>
<geoCoordinate id="7D8E80872B9A1120AE266C8D59B6439B" degrees="50" direction="east" minutes="30" orientation="longitude" precision="925" value="50.5">50°30'E</geoCoordinate>
, 02.07.1975 (A. Senglet).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="A34E98BD1C225DB0158C24996C6B97FC" pageId="0" pageNumber="145" type="etymology">
<paragraph id="F08FF252E7D44D5D13D8AF387CFB97E5" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="06CD9B99D8C91855B6B1C42408917877" pageId="0" pageNumber="145">
The specific epithet is an ancient Persian feminine name, meaning
<normalizedToken originalValue="“fairy-like”">&quot;fairy-like&quot;</normalizedToken>
<normalizedToken id="31E46F1118ED8BE76FFF0C38179D999E" originalValue="“fairy-like”">&quot;fairy-like&quot;</normalizedToken>
. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="990F482248FA94F3BDD9B9663E06BBBE" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="7E576DCD11E03AF8048BCE89BE79BA28" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="1FDFE37AB113D307907F212B59530A27" pageId="0" pageNumber="145">
The male palp of
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
<taxonomicName id="92C774392E21B76DFFCF0DC5241C3D48" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis id="C83ABA4657307891CF5FC27956BA2897" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName>
sp. nov. is similar to that of
<taxonomicName lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
<taxonomicName id="ACB5569446C1C0741D46845A1B4650E2" lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis id="151A5E8C2372E3B253F6D97311253EE1" italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
</taxonomicName>
sp. nov. by the overall shape of the median apophysis and RTA but differs in lacking a retrolateral extension on the median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="F771AC476CBDF63411F87BD6F7778669" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
present) and by having a shorter, stouter RTA (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="A4A37C802A0F5C6DBA5D6733D4FD5A2B" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
longer and tapering). The epigyne of the new species is most similar to that of
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
<taxonomicName id="491CCEE06597B3E74934C9D35C639FA8" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis id="848C1DAD3B45628398DC41FF17D2E2E3" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName>
sp. nov. in having a small anterior hood and thin furrow between the lateral lobes but differs by the relative length of the anterior part of the fovea being shorter than the bursae in
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
<taxonomicName id="61EA29D866B339409A5D965EFC68776D" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis id="5809651E95D21BB24EE707AD11161FAB" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="1973EBC66AB4FBB5507C6F75759AEB3E" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
as long as bursae).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis>
<subSubSection id="45A77B456D0EF09C7EA08C278F42A4B4" pageId="0" pageNumber="145" type="description">
<paragraph id="1C4FBD733AD12FE285EEAED15AE1E793" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="FA14FAC4ACB233964430B5B7A96F3B05" pageId="0" pageNumber="145">
<emphasis id="8C4A7B130C158834F76370F399923DD9" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11F</figureCitation>
<figureCitation id="9599B7FD5FD6ABA6798F4BC92305C07F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11F</figureCitation>
. Total length 2.23. Carapace 1.03 long, 0.46 wide at pars cephalica, 0.80 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.10, ALE: 0.07, PME: 0.06, PLE: 0.05, PME-PME: 0.10. Carapace, sternum, labium, chelicerae and maxillae yellowish. Legs yellowish, slightly darker at femora, without annulations. Abdomen dorsally grayish, pale ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 2.98+missing tarsus (0.95, 0.37, 0.80, 0.86, missing), II: 3.13 (0.77, 0.34, 0.66, 0.79, 0.57), III: 3.03 (0.76, 0.35, 0.58, 0.86, 0.48), IV: 3.44 (1.00, 0.30, 0.73, 0.85, 0.56).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="2FE005722D4D99DF9E855D94695FFDFF" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13F-I</figureCitation>
<figureCitation id="68F2997448A20660AE9B2880D7DD653E" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13F-I</figureCitation>
,
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">15A-C</figureCitation>
<figureCitation id="8CCD482CE6AC281582C13C3DD31F6E4D" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">15A-C</figureCitation>
. RTA (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
<emphasis id="4D4BFDD561F06B5D3781AB64C0FE0989" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
) relatively short and stout, with a finger-like projection dorsally; tegulum with posterior process, sperm duct almost straight retrolaterally, and gently bent along prolateral side; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="3C8FFC1F0B895439C5D3C23F41E350C4" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) wider than long; embolus broad basally, originating at about the 7
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
<normalizedToken id="FF9DDF36E5487C3A7137544BAFEE5783" originalValue="oclock">o'clock</normalizedToken>
position; embolus proper thin, slightly twisted apically.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="BCA1250AEFFA58DE11D1B61CBA4D44C8" pageId="0" pageNumber="145">
<emphasis id="9BF32961AF48833828A91473B6738BE7" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11G</figureCitation>
<figureCitation id="13C7454F66C63617786B917045215864" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11G</figureCitation>
. Total length 3.68. Carapace 1.34 long, 0.60 wide at pars cephalica, 0.94 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.09, ALE: 0.08, PME: 0.07, PLE: 0.07, PME-PME: 0.16. Coloration generally as in male. Abdomen dorsally with a pale median band occupying half of the
<normalizedToken originalValue="abdomens">abdomen's</normalizedToken>
<normalizedToken id="ED366AC177A790178F6AC06B53C5490D" originalValue="abdomens">abdomen's</normalizedToken>
length. Measurements of legs: I: 3.79 (0.90, 0.40, 0.84, 1.01, 0.64), II: 2.90+missing tarsus (0.94, 0.36, 0.74, 0.86, missing), III: 3.48 (0.88, 0.40, 0.65, 1.03, 0.52), IV: 5.09 (1.33, 0.48, 1.13, 1.47, 0.68).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="C2BD51BA32818908C58F75BC2248D7ED" pageId="0" pageNumber="145">
Epigyne as in Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">17D-F</figureCitation>
<figureCitation id="8AEF5553FB3E4C20833A538C6CDDCF7E" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">17D-F</figureCitation>
. Fovea triangular and small anterior hood present; bursae round, separated by less than one of their radii; receptacles smaller than bursae, triangular, separated by more than 3 diameters.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="F0656C9F1D57EB41FCBCDB4FAEFA514C" pageId="0" pageNumber="145" type="distribution">
<paragraph id="B1D2D450F40816836FC1A08C326FCC9C" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="DADD1DB87BFB4BE7D0285FBDF71A11B2" pageId="0" pageNumber="145">
Known only from the listed localities in Ardabil and Qazvin provinces, northern and northwestern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
<figureCitation id="7C6319360F031C736F87DE4423CD4627" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,80 +1,80 @@
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Genus
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<paragraph pageId="0" pageNumber="145">Type species.</paragraph>
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<emphasis italics="true" pageId="0" pageNumber="145">Palaestina sexoculata</emphasis>
<subSubSection id="4225875D165B52587AB5302DCD17B8A5" pageId="0" pageNumber="145" type="type species">
<paragraph id="C5F6C0FDFA69213EFCD438D12BECCC40" pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph id="BAA33531A4A8519017F0121489AE6154" pageId="0" pageNumber="145">
<emphasis id="E118E56D3F3BB975B766C071E6161012" italics="true" pageId="0" pageNumber="145">Palaestina sexoculata</emphasis>
O. Pickard-Cambridge, 1872.
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<subSubSection pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="D342005E66B0AB1523B7BEA0B38F0933" pageId="0" pageNumber="145" type="comments">
<paragraph id="98B70148162BFF81634390B0536C41FB" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph id="95FFFA5517439B969B827BD56C2B6DD8" pageId="0" pageNumber="145">
Small genus with seven named species distributed from Morocco to Turkmenistan (
<bibRefCitation author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC 2021</bibRefCitation>
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).
<taxonomicName authorityName="Simon" authorityYear="1882" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Trygetus</emphasis>
<taxonomicName id="6D5D5A83E1D646320C719631BABDF95F" authorityName="Simon" authorityYear="1882" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="161162C68967A0FB17DEDC2AB4C04583" italics="true" pageId="0" pageNumber="145">Trygetus</emphasis>
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differs from all other
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<taxonomicName id="8FECBC5F65D88E493E228EAC478E636C" authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName>
from the region by having only six eyes (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="54D599146C4B8B9781C670BC12A39892" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
eight eyes).
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View file

@ -1,256 +1,256 @@
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18E-J</figureCitation>
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">, 23A-C</figureCitation>
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">, 26E</figureCitation>
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A) and Z. talyschicum (B, C) A, B ventral C ventroretrolateral. Abbreviations: Tm - median tegular process, Tp - prolateral tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145">, 29A</figureCitation>
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">, 30A-E</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
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<subSubSection id="1D6DA4437D3858492767754384B713FB" pageId="0" pageNumber="145" type="nomenclature">
<paragraph id="E473E90F8921936BBFE2B84C5B05CB01" pageId="0" pageNumber="145">
<taxonomicName id="7023E6A2F9451297FC86C8258A779A78" ID-CoL="98YYY" LSID="http://zoobank.org/61538C61-2E8A-4F21-8289-B9129E5E4965" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum" status="sp. nov.">Acanthinozodium elburzicum</taxonomicName>
<taxonomicNameLabel id="395B023899DE4FA4338A5CECCC86C3E7" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="4C029FE81556B1EAEA47FB549AE0A965" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18E-J</figureCitation>
<figureCitation id="584DB1B2F16FF64505E054CEB2A27027" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">, 21D-F</figureCitation>
<figureCitation id="C1546E0A71419358DF879FD971DB6709" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">, 23A-C</figureCitation>
<figureCitation id="BC31179AFAF5928B552DC95378D80C0B" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">, 26E</figureCitation>
<figureCitation id="9CA97665CFE184DD20B27D81F89ECC29" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A) and Z. talyschicum (B, C) A, B ventral C ventroretrolateral. Abbreviations: Tm - median tegular process, Tp - prolateral tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145">, 29A</figureCitation>
<figureCitation id="CF7138F7C18FDEDB7FA4997166DAF58C" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">, 30A-E</figureCitation>
<figureCitation id="E242524D9F8D2CB21563728105A81B31" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="07308D6308FDEDA492C38A2650E62B68" pageId="0" pageNumber="145" type="type material">
<paragraph id="F606FBA3E43ACC5A377F34392DF084D0" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="EBA2979CFE1E57E2B3216E6CADFD67B1" pageId="0" pageNumber="145">
<emphasis id="CC7F71C239AB57514439A7A94E773BD6" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Tehran Province</emphasis>
<emphasis id="96E5BDF818EBDEEA05798DB5D8488C82" italics="true" pageId="0" pageNumber="145">Tehran Province</emphasis>
: Jamshidieh Mts.,
<geoCoordinate degrees="35" direction="north" minutes="49" orientation="latitude" precision="925" value="35.816666">35°49'N</geoCoordinate>
<geoCoordinate id="23C501F238E454D8CAE637FF0E289560" degrees="35" direction="north" minutes="49" orientation="latitude" precision="925" value="35.816666">35°49'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="27" orientation="longitude" precision="925" value="51.45">51°27'E</geoCoordinate>
<geoCoordinate id="BCB2C90BE131202CF857756174C80D55" degrees="51" direction="east" minutes="27" orientation="longitude" precision="925" value="51.45">51°27'E</geoCoordinate>
, 05.2015 (A. Zamani).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
<emphasis id="D74E407E95E5E4477087FC2563F3982E" bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
: 1♀ (MHNG), Pardisan Park,
<geoCoordinate degrees="35" direction="north" minutes="44" orientation="latitude" precision="925" value="35.733334">35°44'N</geoCoordinate>
<geoCoordinate id="E52D035BA041019E308DB89674199BD6" degrees="35" direction="north" minutes="44" orientation="latitude" precision="925" value="35.733334">35°44'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="21" orientation="longitude" precision="925" value="51.35">51°21'E</geoCoordinate>
<geoCoordinate id="B04774AEA13526854C63F4221BA170D6" degrees="51" direction="east" minutes="21" orientation="longitude" precision="925" value="51.35">51°21'E</geoCoordinate>
, 05.2015 (A. Zamani); 5♂2♀1j. (MMUE), Latian Dam,
<geoCoordinate degrees="35" direction="north" minutes="48" orientation="latitude" precision="925" value="35.8">35°48'N</geoCoordinate>
<geoCoordinate id="40F37F5E735EF70D0BEE18C1ED071F57" degrees="35" direction="north" minutes="48" orientation="latitude" precision="925" value="35.8">35°48'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="08" orientation="longitude" precision="925" value="51.133335">51°08'E</geoCoordinate>
<geoCoordinate id="D0BA311D1F38119233FB1CDF95050A3B" degrees="51" direction="east" minutes="08" orientation="longitude" precision="925" value="51.133335">51°08'E</geoCoordinate>
, 06-19.06.2000 (Y.M. Marusik); 3♂1♀ (MMUE), Plant Protection Institute,
<geoCoordinate degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate>
<geoCoordinate id="D46550EA77C95011C92CB5AB86DBA381" degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="24" orientation="longitude" precision="925" value="51.4">51°24'E</geoCoordinate>
<geoCoordinate id="822BC5231985F4E929BC55D0B20AED46" degrees="51" direction="east" minutes="24" orientation="longitude" precision="925" value="51.4">51°24'E</geoCoordinate>
, 07-22.06.2000 (Y.M. Marusik).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet refers to the Elburz Mountain Range, in which the type locality of the new species is situated.</paragraph>
<subSubSection id="F8916519217126D033753EF36FB45AC6" pageId="0" pageNumber="145" type="etymology">
<paragraph id="CB642AE001CF090E7A17E3A035C92B64" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="25F1CBE3207C58F396E66BE2B90A9AA1" pageId="0" pageNumber="145">The specific epithet refers to the Elburz Mountain Range, in which the type locality of the new species is situated.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="9320EA437358EE5F65D8F97D435D892F" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="081A97DEF09E5A2484C2D84B9F57980C" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="749D61135B05659A5E1694F6474B1F4B" pageId="0" pageNumber="145">
The male of the new species is similar to
<taxonomicName lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
<taxonomicName id="E87FD4685DCCFE50917B0F45F75EA6C5" lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis id="7AE864ED682160F396AB8FE7E38D9154" italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
</taxonomicName>
sp. nov. by the similar shape of the RTA and they both have a small retrolateral indentation apically on the conductor, but differs by the shape of the median apophysis, which is with a distinctly larger prolateral outgrowth. The female of the new species is most similar to
<taxonomicName lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
<taxonomicName id="A723BEF55FE15DB80B0C1B606C874DB0" lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis id="BD26450319666D6673C3A4AC8AD86412" italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
</taxonomicName>
sp. nov. but differs by the epigynal hood being wider than long (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="4F11D00D21B503DF07EBA4CA64F504CA" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
longer than wide) and receptacles separated by about 4 diameters (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="1C01DD9D15C324E1D24AC553D63CEAF1" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
ca. 2.5).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis>
<subSubSection id="39DFF3643BA8C32BDD2006B4472632CB" pageId="0" pageNumber="145" type="description">
<paragraph id="9C96C772F983C9A460EFB6D522CD2D4B" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="B90ED37053D338F366C0CABE012CF7B0" pageId="0" pageNumber="145">
<emphasis id="B6A390A47BB7CB7EC04E84629D051C41" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18F-H</figureCitation>
<figureCitation id="1F2CCBE7963C8D15BCF04AE19888B61D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18F-H</figureCitation>
(holotype: 18F). Total length 4.30. Carapace 2.08 long, 0.97 wide at pars cephalica, 1.49 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.14, ALE: 0.12, PME: 0.08, PLE: 0.09, PME-PME: 0.15. Carapace dark brown, lighter in pars cephalica, with irregular dark patches and lines. Sternum, labium and maxillae light brown. Chelicera (Fig.
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">26E</figureCitation>
<figureCitation id="183FBEB2670163B7579AC3A1D5B02BCA" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">26E</figureCitation>
) dark brown, with one retromarginal tooth. Legs slightly lighter than carapace, without annulations. Abdomen dorsally black and with large scutum; ventrally grayish, without any pattern. Spinnerets pale, uniform in color. Measurements of legs: I: 7.52 (1.79, 0.71, 1.75, 2.03, 1.24), II: 6.79 (1.87, 0.69, 1.40, 1.83, 1.00), III: 6.90 (1.78, 0.74, 1.40, 2.06, 0.92), IV: 9.19 (2.48, 0.83, 2.15, 2.84, 0.89).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="0DB44EBAC7B0F53D0B92F2BB1A72A65E" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">21D-F</figureCitation>
<figureCitation id="FA9DD1CFBD204BF2D6A8A9A2D4388570" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">21D-F</figureCitation>
,
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">23A-C</figureCitation>
<figureCitation id="2B33F7E88A321A4A311DD30B1142321C" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">23A-C</figureCitation>
,
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A) and Z. talyschicum (B, C) A, B ventral C ventroretrolateral. Abbreviations: Tm - median tegular process, Tp - prolateral tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145">29A</figureCitation>
<figureCitation id="676309995796844BEC863B6331E7DDDC" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A) and Z. talyschicum (B, C) A, B ventral C ventroretrolateral. Abbreviations: Tm - median tegular process, Tp - prolateral tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145">29A</figureCitation>
. RTA (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
<emphasis id="F30A7D59E729E35CFC444D7766CFEBF7" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
) long and conical, with a small projection apically; ventral apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis>
<emphasis id="B60A025450910F8C2375DB0CCCA0FEC4" italics="true" pageId="0" pageNumber="145">Va</emphasis>
) small and conical; tegulum with posterior process; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="2BB883B7F43C577E6573718BD8BEE40A" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) wider than long, with outgrowths on all four sides, the triangular prolateral one the largest; embolus (
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
<emphasis id="6E5C53A4A8FD7692DF30EAFCD609E63A" italics="true" pageId="0" pageNumber="145">Em</emphasis>
) originating at about the 7
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
<normalizedToken id="D09D6AE6C103A9CF954A04CF3E7F7853" originalValue="oclock">o'clock</normalizedToken>
position; embolus proper thin and steadily curving.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145" start="Figure 21" startId="F21">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 21.</emphasis>
<caption id="FD0299D2E0D123CDBA7F7F8F442B9C62" doi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145" start="Figure 21" startId="F21">
<paragraph id="D0C1A14DA7DC86FA6122E40463CA5A49" pageId="0" pageNumber="145">
<emphasis id="7D2517EE328FAD4E27F6695D4AFA0FC9" bold="true" pageId="0" pageNumber="145">Figure 21.</emphasis>
Male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium dorsa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium dorsa</emphasis>
<taxonomicName id="A36D0ED871E6FCC911BD6C9F9FD3B7ED" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium dorsa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dorsa">
<emphasis id="E723928E768CE7EDC12F58AEE885208B" italics="true" pageId="0" pageNumber="145">Acanthinozodium dorsa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
<emphasis id="B446F6C1E5BDFAB43EC809A7E2C9DE42" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and
<taxonomicName lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis>
<taxonomicName id="23DC8CD2B924E7CA9C4CA1D13304DB9E" lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum">
<emphasis id="F4BFF92F33B9E38896C6F08200AD376C" italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
<emphasis id="78C0803222DEEE0620E5C2B586D7E761" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis>
<emphasis id="273ABC0AC32DC4CE12D5ACDA8ACBF13C" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, E</emphasis>
<emphasis id="5A13DC765DE318E68F54CEE14FFBE607" bold="true" pageId="0" pageNumber="145">B, E</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, F</emphasis>
<emphasis id="35A81FCB17E98629C9EFBD53615B1306" bold="true" pageId="0" pageNumber="145">C, F</emphasis>
dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145" start="Figure 22" startId="F22">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 22.</emphasis>
<caption id="57137E8BBB07B68EB05BB4B5CD0010EA" doi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145" start="Figure 22" startId="F22">
<paragraph id="D00B7F15A9C19E20F49547E10DF8DB96" pageId="0" pageNumber="145">
<emphasis id="71C18DED5B8D9050ED552DBE06B14510" bold="true" pageId="0" pageNumber="145">Figure 22.</emphasis>
SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium armita</emphasis>
<taxonomicName id="EF049C19027CA8C6CAFD0F238A458F2D" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita">
<emphasis id="AAC0ACEB3F6CB319E8E2E53CB202FBF4" italics="true" pageId="0" pageNumber="145">Acanthinozodium armita</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C, F</emphasis>
<emphasis id="3EDC37CA45B74E7B2A6FFCFCA472F4CD" bold="true" pageId="0" pageNumber="145">A-C, F</emphasis>
) and
<taxonomicName lsidName="A. ovtchinnikovi" pageId="0" pageNumber="145" rank="species" species="ovtchinnikovi">
<emphasis italics="true" pageId="0" pageNumber="145">A. ovtchinnikovi</emphasis>
<taxonomicName id="05100D9169462A4D940A986302F5083F" lsidName="A. ovtchinnikovi" pageId="0" pageNumber="145" rank="species" species="ovtchinnikovi">
<emphasis id="AA16193EFEA282FFA7CCBBDD42F1FFBE" italics="true" pageId="0" pageNumber="145">A. ovtchinnikovi</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D, E, G-I</emphasis>
<emphasis id="E6ECE5ED7C0497E5A411408789920177" bold="true" pageId="0" pageNumber="145">D, E, G-I</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, H</emphasis>
<emphasis id="17BF74C7D2E23B6C755389FDF8CA2810" bold="true" pageId="0" pageNumber="145">A, H</emphasis>
retroventral and retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">B, C, G</emphasis>
<emphasis id="63E2C82DB4EDD0D6BBADCD0BE69B5420" bold="true" pageId="0" pageNumber="145">B, C, G</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
<emphasis id="AC291749C22F70F3A3963B240140F13F" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
proventral, apicoproventral and apicoventral
<emphasis bold="true" pageId="0" pageNumber="145">I</emphasis>
<emphasis id="F2A871B60682A7AC2E14EA826E8D4C44" bold="true" pageId="0" pageNumber="145">I</emphasis>
retrolateral tibial and cymbial groove. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Cg</emphasis>
<emphasis id="01F5318B65CD310AE1099B1B6F88A808" italics="true" pageId="0" pageNumber="145">Cg</emphasis>
- cymbial groove,
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis>
<emphasis id="61633E5DA08084AA722EE0A8ED808597" italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
<emphasis id="F7FAD983499BB03AF0267B165486E679" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Er</emphasis>
<emphasis id="7D21BFF826D0CFA33E34BEECF27EC1EC" italics="true" pageId="0" pageNumber="145">Er</emphasis>
-embolar ridge,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="FB3DCA943A77C7FAF078CF760D2C6039" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis. Scale bars: 0.1 mm, unless stated otherwise.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145" start="Figure 23" startId="F23">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 23.</emphasis>
<caption id="B902C497AE657AD99572C81724BC3FA7" doi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145" start="Figure 23" startId="F23">
<paragraph id="35B1D993267AC875B7C514CE04E1D181" pageId="0" pageNumber="145">
<emphasis id="B91B3C98DE41BB3200BEA86E34A86D35" bold="true" pageId="0" pageNumber="145">Figure 23.</emphasis>
SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis>
<taxonomicName id="93AAD109AFF9C0E0FDF6446EAAC5439F" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum">
<emphasis id="A6E50972C2E924D50098EB727C744D63" italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
<emphasis id="8679548182CE2C3892293B2F5B8FEB59" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and
<taxonomicName lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
<taxonomicName id="0F49720ABABE31B140127428A8E1ED33" lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis id="DED5A56804CB8F14DC5E549E53D72244" italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
<emphasis id="181307FBA7B6183BCF071CEF02506BF8" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis>
<emphasis id="D82E4596E186A24A212E8356F78BBF34" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, F</emphasis>
<emphasis id="EE63ADFCC1909A78F7AA1FA1EF2EAD47" bold="true" pageId="0" pageNumber="145">B, F</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, E</emphasis>
<emphasis id="D0A77B3E7B181DC443170E7D06213193" bold="true" pageId="0" pageNumber="145">C, E</emphasis>
dorsoretrolateral. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Cg</emphasis>
<emphasis id="AF6142E31FC2B60229C4E0BE99DBCE19" italics="true" pageId="0" pageNumber="145">Cg</emphasis>
- cymbial groove,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
<emphasis id="168BBC39282E9B7FC6C871F3E9ED2932" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="A68391B813E10C141E4672DD77EC285B" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
<emphasis id="61D9AB1DC163A99F1DC1FE98C91C5841" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral tibial apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis>
<emphasis id="BA8DA243AE84501EC08FD9867084947A" italics="true" pageId="0" pageNumber="145">Va</emphasis>
- ventral apophysis. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="9E76A0D88821D9FE99D5094CB3650B6F" pageId="0" pageNumber="145">
<emphasis id="3FCDB9674FEDD74F16934D7AAD2AED7E" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18E, I, J</figureCitation>
<figureCitation id="3C1AEA71E60652EAC3A6A0AC93445058" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18E, I, J</figureCitation>
. Total length 4.95. Carapace 1.95 long, 0.96 wide at pars cephalica, 1.40 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.15, ALE: 0.12, PME: 0.08, PLE: 0.11, PME-PME: 0.15. Coloration as in male, with paler abdomen lacking a scutum. Measurements of legs: I: 6.54 (1.50, 0.70, 1.44, 1.76, 1.14), II: 6.27 (1.59, 0.67, 1.33, 1.67, 1.01), III: 6.23 (1.60, 0.71, 1.21, 1.74, 0.97), IV: 8.11 (2.04, 0.78, 1.81, 2.46, 1.02).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="10EF5E79F3675EC7C4DC52329CA11209" pageId="0" pageNumber="145">
Epigyne as in Fig.
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">30A-E</figureCitation>
<figureCitation id="83462842097947057366789C1BA0D156" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">30A-E</figureCitation>
. Epigynal plater over 3 times wider than long, lacking fovea; anterior hood wider than long; receptacles with rounded posterior parts, separated by about 4 diameters.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="BB59AEA44A54F966CB5675AB8CAA633A" pageId="0" pageNumber="145" type="distribution">
<paragraph id="5C5237BB14756546E8F1048FB6EC87BB" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="CC29874BFBF5564357F8D36F7FBD0272" pageId="0" pageNumber="145">
Known from the listed localities in Tehran Province, northern Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
<figureCitation id="359050A6D28A92256041F999BCBCA561" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
).
</paragraph>
</subSubSection>

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@ -1,72 +1,72 @@
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<paragraph id="F85315FAEC9896D9AD2E0A7107581AEE" pageId="0" pageNumber="145">
Genus
<taxonomicName LSID="5E0B26BF-C08A-592C-A10A-3AA52539B4A6" authority="Levy, 1990" authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Pax Levy, 1990</taxonomicName>
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<paragraph pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName class="Arachnida" family="Zodariidae" genus="Habronestes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Habronestes libani" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="libani">
<emphasis italics="true" pageId="0" pageNumber="145">Habronestes libani</emphasis>
<subSubSection id="14CCFAA886F98A222C26D9F216BF1F03" pageId="0" pageNumber="145" type="type species">
<paragraph id="8C8FEE0B79EB33851CCCCE67A758EC72" pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph id="4BD7A0187B2F8F3712CA0A8702E54E8B" pageId="0" pageNumber="145">
<taxonomicName id="335B70F12A999C702EC374F86A79D1F5" class="Arachnida" family="Zodariidae" genus="Habronestes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Habronestes libani" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="libani">
<emphasis id="C6EBF2AFF57E5A78A94A509EC1488888" italics="true" pageId="0" pageNumber="145">Habronestes libani</emphasis>
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Simon, 1873 from Lebanon.
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<subSubSection pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">This is a small genus with seven species distributed exclusively in the Middle East from Turkey and Israel to Iran (including the two new species described below, representing the first record of this genus in this country). Members of this genus can be easily distinguished from all other zodariids found in the study area by the ovoid carapace lacking a distinct separation between the cephalic and thoracic parts and also by males having a modified cymbium.</paragraph>
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<paragraph id="FC9228D6F4183657C9F1A9B58FB3AD1D" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph id="BD02FA6309CA2DE8187EDD77717D9D78" pageId="0" pageNumber="145">This is a small genus with seven species distributed exclusively in the Middle East from Turkey and Israel to Iran (including the two new species described below, representing the first record of this genus in this country). Members of this genus can be easily distinguished from all other zodariids found in the study area by the ovoid carapace lacking a distinct separation between the cephalic and thoracic parts and also by males having a modified cymbium.</paragraph>
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@ -1,88 +1,88 @@
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Lachesana insensibilis
<normalizedToken originalValue="Jocqué">Jocque</normalizedToken>
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, 1991
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">Fig. 32</figureCitation>
<figureCitation id="62957599DA8D28E63724AF60F663B3C8" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">Fig. 32</figureCitation>
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<taxonomicName authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana insensibilis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="insensibilis">Lachesana insensibilis</taxonomicName>
<normalizedToken originalValue="Jocqué">Jocque</normalizedToken>
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<taxonomicName id="148BB59CA1563475AEC6EFE2E83F2EA1" authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana insensibilis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="insensibilis">Lachesana insensibilis</taxonomicName>
<normalizedToken id="83FC5F6560FCB0E9B285E4135D9786C1" originalValue="Jocqué">Jocque</normalizedToken>
, 1991: 37, f. 59 (♂).
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<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">This species is known from a single taxonomic entry, and unfortunately, the original description does not provide illustrations of the lateral view of the palp, making the identification of specimens from outside of the type locality (Saudi Arabia) highly questionable.</paragraph>
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<paragraph id="2418C39222C050063B3A290DBDBD5DE7" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph id="7A5CAE97F41D27BAEBF3C0C77685582A" pageId="0" pageNumber="145">This species is known from a single taxonomic entry, and unfortunately, the original description does not provide illustrations of the lateral view of the palp, making the identification of specimens from outside of the type locality (Saudi Arabia) highly questionable.</paragraph>
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<paragraph pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="C7409797704D008E971A3E26A4D86E6E" pageId="0" pageNumber="145" type="records in iran">
<paragraph id="9B3AC66511BF0E97E52BE5E29CFF5EE2" pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph id="88BE480B27D5DD48C43639A372498205" pageId="0" pageNumber="145">
Khuzestan (
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<bibRefCitation id="8B50ECA77788999DBB99F3E76BB3DCA6" DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation>
) (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
<figureCitation id="1CE0FE83D792ECEF6FC2953BB9410B9B" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">Saudi Arabia, Israel, Iran, United Arab Emirates.</paragraph>
<subSubSection id="431A32A8495DF11341309420760A57C9" pageId="0" pageNumber="145" type="distribution">
<paragraph id="9BAD67502F4B9467436B591DD60DB362" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="93BD20F3081D1AE601B610E25A4ADB7E" pageId="0" pageNumber="145">Saudi Arabia, Israel, Iran, United Arab Emirates.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,127 +1,127 @@
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18L</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20I-K</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">, 24A, B</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
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<taxonomicNameLabel id="040313C731AE3DF581AAFD2CA0EDE3C5" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="DAD062AD45364CCD940C701302BE218A" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18L</figureCitation>
<figureCitation id="0D0F1645F4EC62BCF555A1D04C0BACFB" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20I-K</figureCitation>
<figureCitation id="E661810D463FB1DDAC14E77DEFF1B5AC" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">, 24A, B</figureCitation>
<figureCitation id="EEDD38CD2522B27A94ADA1DB50BF5EF2" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="126DE4794FF8DFA770B5041B115D1E55" pageId="0" pageNumber="145" type="type material">
<paragraph id="299E6266591B4A488BA7A2C510ADA9F9" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="93FCD052AF78D6326A8A392D4EFD84DF" pageId="0" pageNumber="145">
<emphasis id="D915FAEB8715EB80555B94611900512C" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ and
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis>
<emphasis id="F82470FE6DAC851514DF769849ECB73A" bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis>
♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Kermanshah Province</emphasis>
<emphasis id="EC75688354FF031223784BEE25A56F93" italics="true" pageId="0" pageNumber="145">Kermanshah Province</emphasis>
: Mahi Dasht,
<geoCoordinate degrees="34" direction="north" minutes="14" orientation="latitude" precision="925" value="34.233334">34°14'N</geoCoordinate>
<geoCoordinate id="8F29C6074C569DB08DE2194CD1AEC2E3" degrees="34" direction="north" minutes="14" orientation="latitude" precision="925" value="34.233334">34°14'N</geoCoordinate>
,
<geoCoordinate degrees="46" direction="east" minutes="42" orientation="longitude" precision="925" value="46.7">46°42'E</geoCoordinate>
<geoCoordinate id="2B6926513B59C25680848C5816A5EFF6" degrees="46" direction="east" minutes="42" orientation="longitude" precision="925" value="46.7">46°42'E</geoCoordinate>
, 29.06.1974 (A. Senglet).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet is a Kurdish feminine name, meaning &quot;bright like the moon&quot;. Noun in apposition.</paragraph>
<subSubSection id="8733BA9D9D073538D126F1C70BCEB4F7" pageId="0" pageNumber="145" type="etymology">
<paragraph id="3B42208E74F1FF50076EF9DEA47986D7" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="E7D8B02ECE1EDDBF947CD7736654E66F" pageId="0" pageNumber="145">The specific epithet is a Kurdish feminine name, meaning &quot;bright like the moon&quot;. Noun in apposition.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="6706EDBA731ED3ED4403069EA2170B89" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="EE0E1FDF20C1394526CFCCFD0CC36BA3" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="36BC012433206032A9A8A21F52198277" pageId="0" pageNumber="145">
The new species is very similar to
<taxonomicName lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
<taxonomicName id="6CD6673F081AFAB9B582D8C675E21C80" lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis id="27A558AAB17F90D549B662BC630EA197" italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
</taxonomicName>
sp. nov. by the shape of the RTA, the ventral tibial apophysis and the curvature of the embolus but differs by the shape of the median apophysis (cf. Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20D</figureCitation>
<figureCitation id="734359D0490911A8DB61AC7CADD86DCF" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20D</figureCitation>
and
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20J</figureCitation>
<figureCitation id="1F6BCC64ED66CCB27AEEA26B19FF3CEE" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20J</figureCitation>
) and the conductor lacking a small retrolateral indentation apically (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="72667BB81FB3CF8DAEE0BD89A196A50C" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
present).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="6AE023028C9724141F429456C59A8AC6" pageId="0" pageNumber="145" type="description">
<paragraph id="0F999683C187ADD458BEED6371687330" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="C677F2EC2940470E183406C629F2F37B" pageId="0" pageNumber="145">
Male (holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18L</figureCitation>
<figureCitation id="758F3CD253C54B4FAB1388428BC98AD0" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18L</figureCitation>
. Total length 3.05. Carapace 1.47 long, 0.66 wide at pars cephalica, 1.10 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.12, ALE: 0.10, PME: 0.07, PLE: 0.08, PME-PME: 0.12. Carapace, sternum, labium, chelicerae and maxillae brown; carapace with irregular patterns. Chelicera with one tooth. Legs yellowish brown. Abdomen dorsally black, grayish ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 5.52 (1.50, 0.47, 1.23, 1.42, 0.90), II: 4.91 (1.15, 0.50, 1.11, 1.30, 0.85), III: 4.62 (1.10, 0.45, 0.94, 1.43, 0.70), Fe IV: 1.80, rest of the segments missing.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="BC183F9C496A195F581334CF06370A2E" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20I-K</figureCitation>
<figureCitation id="968637A44FB3CC7DDD30B02F9CCC8881" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20I-K</figureCitation>
,
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">24A, B</figureCitation>
<figureCitation id="2E46D77AD61BAB4110F3B5C67C08C342" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">24A, B</figureCitation>
. RTA long and conical, with a small projection apically; tegulum with posterior process (
<emphasis italics="true" pageId="0" pageNumber="145">Tp</emphasis>
<emphasis id="6ACF847A37FC908E9E6FEDF0590DB651" italics="true" pageId="0" pageNumber="145">Tp</emphasis>
); ventral apophysis relatively large and conical; median apophysis wider than long, with a large bifurcated outgrowth prolaterally; embolus broad basally, originating at about the 7
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
<normalizedToken id="1610F06C25E3C13B1A2C3A05F485F8CA" originalValue="oclock">o'clock</normalizedToken>
position; embolus proper thin and steadily curving.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="170D3B28A032E164A487276DE78227E3" pageId="0" pageNumber="145">
<emphasis id="135B6803A23F7892BA551ABC2C17DFA3" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="197F25C70BEF4F887BD1056B15434275" pageId="0" pageNumber="145" type="distribution">
<paragraph id="0D6DBE4409D891C74B0A5BB48F43166F" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="B2C8FF4BF83436C1639B87023B5F44EC" pageId="0" pageNumber="145">
Known only from the type locality in Kermanshah Province, western Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
<figureCitation id="E453F15120DC340FB72128190E058AEA" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,83 +1,83 @@
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<taxonomicName LSID="6A815D86-982B-505E-8552-F43F3F2F2A9E" authority="(Nenilin &amp; Fet, 1985)" baseAuthorityName="Nenilin &amp; Fet" baseAuthorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum sytchevskajae" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="sytchevskajae">Zodariellum sytchevskajae (Nenilin &amp; Fet, 1985)</taxonomicName>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">Fig. 33</figureCitation>
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<taxonomicName id="E77E621A2A474BCE967A434AF255B924" ID-CoL="93S8P" LSID="6A815D86-982B-505E-8552-F43F3F2F2A9E" authority="(Nenilin &amp; Fet, 1985)" baseAuthorityName="Nenilin &amp; Fet" baseAuthorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum sytchevskajae" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="sytchevskajae">Zodariellum sytchevskajae (Nenilin &amp; Fet, 1985)</taxonomicName>
<figureCitation id="77A731E733858A272188027F1FD5FA42" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">Fig. 33</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Nenilin &amp; Fet" authorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion sytchevskajae" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="sytchevskajae">Zodarion sytchevskajae</taxonomicName>
<subSubSection id="0A77439F56FBDE111946958E01EDB43B" pageId="0" pageNumber="145" type="reference_group">
<paragraph id="CBEDDDCCCA043A00D77CFAC80F86C568" pageId="0" pageNumber="145">
<taxonomicName id="5DA1603558B6CB82FD331B8E69EBCB8A" authorityName="Nenilin &amp; Fet" authorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion sytchevskajae" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="sytchevskajae">Zodarion sytchevskajae</taxonomicName>
Nenilin &amp; Fet, 1985: 619, f. 5-8, 10 (♂♀).
</paragraph>
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<subSubSection pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">This species is known from the original description only.</paragraph>
<subSubSection id="35A2C2D1008AFA2729611681CE0C71AB" pageId="0" pageNumber="145" type="comments">
<paragraph id="05E7278A417F74EB41FDCB6CC09040EF" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph id="83BDCF3D6B4E8545DD87F559D1063953" pageId="0" pageNumber="145">This species is known from the original description only.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in turkmenistan">
<paragraph pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="784D29C55893E13E425708751D5FA926" pageId="0" pageNumber="145" type="records in turkmenistan">
<paragraph id="4BC601174795DA58A698DD50DA3165FE" pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph>
<paragraph id="7C5C3D084BA319304133505CE8D13241" pageId="0" pageNumber="145">
Ahal, Lebap (
<bibRefCitation author="Nenilin, AB" journalOrPublisher="Zoologicheskiĭ Zhurnal" pageId="0" pageNumber="145" pagination="618 - 620" refId="B17" refString="Nenilin, AB, Fet, VY, 1985. New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia. Zoologicheskiĭ Zhurnal 64: 618 - 620" title="New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia." volume="64" year="1985">Nenilin and Fet 1985</bibRefCitation>
<bibRefCitation id="77A504DD5E298C04EC035B8FE1A91CC3" author="Nenilin, AB" journalOrPublisher="Zoologicheskiĭ Zhurnal" pageId="0" pageNumber="145" pagination="618 - 620" refId="B17" refString="Nenilin, AB, Fet, VY, 1985. New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia. Zoologicheskiĭ Zhurnal 64: 618 - 620" title="New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia." volume="64" year="1985">Nenilin and Fet 1985</bibRefCitation>
) (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
<figureCitation id="0E7E4B4B633AB112992D8FE8192B519A" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
).
</paragraph>
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<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">Turkmenistan.</paragraph>
<subSubSection id="C8FEF5E864B2C4177EB8FCF7C901D58C" pageId="0" pageNumber="145" type="distribution">
<paragraph id="D5253D37DE193C7A64B984A35276AFBA" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="284C96BC32F277CCF3AF51E082371766" pageId="0" pageNumber="145">Turkmenistan.</paragraph>
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<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">Figs 26A-C</figureCitation>
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Epigynes of Parazodarion raddei (A-D) and Trygetus susianus sp. nov. (E-H) A, E intact, ventral B, F-H macerated, dorsal C, D macerated, ventral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145">, 27A-D</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
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<figureCitation id="DDB3A7F0597EFE566FB143B7A85CD5E3" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">Figs 26A-C</figureCitation>
<figureCitation id="6433C66F2F39906454E3BF7FC9AEC1D8" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Epigynes of Parazodarion raddei (A-D) and Trygetus susianus sp. nov. (E-H) A, E intact, ventral B, F-H macerated, dorsal C, D macerated, ventral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145">, 27A-D</figureCitation>
<figureCitation id="BB936EA2C50217DDA4A160C67250DFC1" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="reference_group">
<paragraph pageId="0" pageNumber="145">
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion raddei" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei">Parazodarion raddei</taxonomicName>
<subSubSection id="A54CCD8A75F94B6502AA855958115F19" pageId="0" pageNumber="145" type="reference_group">
<paragraph id="302FDF512D4B230592281B3C8BBB62A7" pageId="0" pageNumber="145">
<taxonomicName id="6731F357A928CAED9BFB039DF5A58D59" baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion raddei" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei">Parazodarion raddei</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.2478/v10058-009-0020-z" author="Ovtchinnikov, SV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="145" pagination="471 - 474" refId="B19" refString="Ovtchinnikov, SV, Ahmad, B, Gurko, VO, 2009. Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia. Vestnik Zoologii 43 (5): 471 - 474, DOI: https://doi.org/10.2478/v10058-009-0020-z" title="Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia." url="https://doi.org/10.2478/v10058-009-0020-z" volume="43" year="2009">Ovtchinnikov et al. 2009</bibRefCitation>
<bibRefCitation id="A3B19CD2BE64EB2F1827073A49A38D1D" DOI="https://doi.org/10.2478/v10058-009-0020-z" author="Ovtchinnikov, SV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="145" pagination="471 - 474" refId="B19" refString="Ovtchinnikov, SV, Ahmad, B, Gurko, VO, 2009. Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia. Vestnik Zoologii 43 (5): 471 - 474, DOI: https://doi.org/10.2478/v10058-009-0020-z" title="Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia." url="https://doi.org/10.2478/v10058-009-0020-z" volume="43" year="2009">Ovtchinnikov et al. 2009</bibRefCitation>
: 471, f. 1.1-6 (♂♀).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion raddei" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei">Parazodarion raddei</taxonomicName>
<paragraph id="8A001BE69001B2BAEAC52E397595DE2D" pageId="0" pageNumber="145">
<taxonomicName id="BD515B266358C3B8866838D118A8873C" baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion raddei" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei">Parazodarion raddei</taxonomicName>
For the complete list of references see
<bibRefCitation author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC (2021)</bibRefCitation>
<bibRefCitation id="35DE8EE2716FBD97D292DB2E5B853BEF" author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC (2021)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="materials_examined">
<paragraph pageId="0" pageNumber="145">Material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<materialsCitation ID-GBIF-Occurrence="3093634306" collectingDate="1975-06-16" collectorName="A. Senglet" country="Iran" elevation="2600" latitude="34.733334" location="Hamedan Province" longLatPrecision="1197" longitude="48.783333" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Hamedan Province">
<collectingCountry name="Iran">Iran</collectingCountry>
<subSubSection id="E244968BCEE0CF4D65EA207DDA71814A" pageId="0" pageNumber="145" type="materials_examined">
<paragraph id="410E113E69E1A6E4627F948CE8C9A3F8" pageId="0" pageNumber="145">Material.</paragraph>
<paragraph id="6EE5587EC0D625303C320F20DE5A8160" pageId="0" pageNumber="145">
<materialsCitation id="D389C4F3571AB72460E18D5E46854904" ID-GBIF-Occurrence="3093634306" collectingDate="1975-06-16" collectorName="A. Senglet" country="Iran" elevation="2600" latitude="34.733334" location="Hamedan Province" longLatPrecision="1197" longitude="48.783333" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Hamedan Province">
<collectingCountry id="C0D734EA38415EB57AEB8041F0D44C5A" name="Iran">Iran</collectingCountry>
:
<emphasis italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Hamadan">Hamedan Province</collectingRegion>
<emphasis id="3E503A6D9738B9958EDF0B81C8C2CB6A" italics="true" pageId="0" pageNumber="145">
<collectingRegion id="685C403C6B295859C957CDE6C073F57B" country="Iran" name="Hamadan">Hamedan Province</collectingRegion>
</emphasis>
:
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">3♀</specimenCount>
<specimenCount id="647FC74E0BA2CE6FA2AEEF62C6F3C520" type="male">1♂</specimenCount>
<specimenCount id="A167E860FCF6ACE8C25DF5C87E92A73E" type="female">3♀</specimenCount>
(MHNG), around
<collectingRegion country="Iran" name="Hamadan">Hamedan</collectingRegion>
<collectingRegion id="098538C1A695C3FEF052150A2961CC74" country="Iran" name="Hamadan">Hamedan</collectingRegion>
,
<geoCoordinate degrees="34" direction="north" minutes="44" orientation="latitude" precision="925" value="34.733334">34°44'N</geoCoordinate>
<geoCoordinate id="7C75A0F829DC9350A5ABD9958BE61C9A" degrees="34" direction="north" minutes="44" orientation="latitude" precision="925" value="34.733334">34°44'N</geoCoordinate>
,
<geoCoordinate degrees="48" direction="east" minutes="47" orientation="longitude" precision="925" value="48.783333">48°47'E</geoCoordinate>
<geoCoordinate id="66D89399D60D3C419D85096DE21D7A6D" degrees="48" direction="east" minutes="47" orientation="longitude" precision="925" value="48.783333">48°47'E</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2600 m</elevation>
<quantity id="5017699E4F3723889E6C4FF0C0D6DC0E" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">
<elevation id="66443E2833EEED4B41F4AC7F15FD7659" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2600 m</elevation>
</quantity>
,
<collectingDate value="1975-06-16">16.06.1975</collectingDate>
<collectingDate id="9161C150BEF5B26CFEA65F2A47C5F34E" value="1975-06-16">16.06.1975</collectingDate>
(
<collectorName>A. Senglet</collectorName>
<collectorName id="8FCA931AFC8C9E49B765ECA5E8FD1683">A. Senglet</collectorName>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3093634304" collectingDate="1974-07-02" collectorName="A. Senglet" country="Iran" latitude="34.85" location="Aliabad" longLatPrecision="1197" longitude="48.2" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Hamedan">
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">2♀</specimenCount>
<materialsCitation id="405D1F290C37F40F717C49E8166C72F8" ID-GBIF-Occurrence="3093634304" collectingDate="1974-07-02" collectorName="A. Senglet" country="Iran" latitude="34.85" location="Aliabad" longLatPrecision="1197" longitude="48.2" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Hamedan">
<specimenCount id="3E3BFB512B41AFEDB696B449480BA2C2" type="male">1♂</specimenCount>
<specimenCount id="8B798197F6BC0BA2B31FBA2421AC58F6" type="female">2♀</specimenCount>
(MHNG),
<location LSID="urn:lsid:plazi:treatment:6BA12FA2395B570193927134C049CA76:51B9CCABD6DF0BB85FE5DB52E7473A31" country="Iran" latitude="34.85" longLatPrecision="1197" longitude="48.2" name="Aliabad" stateProvince="Hamedan">Aliabad</location>
<location id="51B9CCABD6DF0BB85FE5DB52E7473A31" LSID="urn:lsid:plazi:treatment:6BA12FA2395B570193927134C049CA76:51B9CCABD6DF0BB85FE5DB52E7473A31" country="Iran" latitude="34.85" longLatPrecision="1197" longitude="48.2" name="Aliabad" stateProvince="Hamedan">Aliabad</location>
,
<geoCoordinate degrees="34" direction="north" minutes="51" orientation="latitude" precision="925" value="34.85">34°51'N</geoCoordinate>
<geoCoordinate id="F1E007AB69025CE1E944A1BF0C47FFE4" degrees="34" direction="north" minutes="51" orientation="latitude" precision="925" value="34.85">34°51'N</geoCoordinate>
,
<geoCoordinate degrees="48" direction="east" minutes="12" orientation="longitude" precision="925" value="48.2">48°12'E</geoCoordinate>
<geoCoordinate id="53FC3B7AF407E15AEF8310400DD42893" degrees="48" direction="east" minutes="12" orientation="longitude" precision="925" value="48.2">48°12'E</geoCoordinate>
,
<collectingDate value="1974-07-02">02.07.1974</collectingDate>
<collectingDate id="59EEC32FE6649381975F739B9B375D91" value="1974-07-02">02.07.1974</collectingDate>
(
<collectorName>A. Senglet</collectorName>
<collectorName id="B90FAE11AEFB7D8DC1A6934AAAF21AC9">A. Senglet</collectorName>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3093634301" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Iran" location="Tehran Province" specimenCount="12" specimenCount-female="5" specimenCount-male="7" stateProvince="Tehran Province">
<emphasis italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Tehran">Tehran Province</collectingRegion>
<materialsCitation id="2FB29C7D04BAD0F07F4DEE7F87F9C779" ID-GBIF-Occurrence="3093634301" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Iran" location="Tehran Province" specimenCount="12" specimenCount-female="5" specimenCount-male="7" stateProvince="Tehran Province">
<emphasis id="07F1480B0AFAE52AE9EA143D4411D22B" italics="true" pageId="0" pageNumber="145">
<collectingRegion id="B8D89F72E621D958D4E537BB2D77869C" country="Iran" name="Tehran">Tehran Province</collectingRegion>
</emphasis>
:
<specimenCount type="male">7♂</specimenCount>
<specimenCount type="female">5♀</specimenCount>
<specimenCount id="01D9292E8F3700C7725AE706F5A37449" type="male">7♂</specimenCount>
<specimenCount id="285286A30BB8617FC269BE4B1F0BE9E5" type="female">5♀</specimenCount>
(ZMMU),
<quantity metricMagnitude="4" metricUnit="m" metricValue="8.0" unit="km" value="80.0">80 km</quantity>
<quantity id="E9B29F6B8375D7F06561AD4B377EA742" metricMagnitude="4" metricUnit="m" metricValue="8.0" unit="km" value="80.0">80 km</quantity>
E of
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3093634313" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Iran" isTailEnumeration="true" location="Tehran" specimenCount="1" stateProvince="Tehran">
<collectingRegion country="Iran" name="Tehran">Tehran</collectingRegion>
<materialsCitation id="847A7EE2DFBA8E7ED0149655283FDA56" ID-GBIF-Occurrence="3093634313" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Iran" isTailEnumeration="true" location="Tehran" specimenCount="1" stateProvince="Tehran">
<collectingRegion id="9BF0D811207BF446C53229D3B591EDFC" country="Iran" name="Tehran">Tehran</collectingRegion>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3093634305" collectingDate="2000-06-15" collectorName="Y. M. Marusik, F. Mozaffarian" country="Iran" latitude="35.666668" location="Aroo" longLatPrecision="1191" longitude="52.45" municipality="Damavand area" specimenCount="1" stateProvince="Tehran">
<collectingMunicipality>Damavand area</collectingMunicipality>
<materialsCitation id="498252D9D0F2A543CF51EE32A354EA60" ID-GBIF-Occurrence="3093634305" collectingDate="2000-06-15" collectorName="Y. M. Marusik, F. Mozaffarian" country="Iran" latitude="35.666668" location="Aroo" longLatPrecision="1191" longitude="52.45" municipality="Damavand area" specimenCount="1" stateProvince="Tehran">
<collectingMunicipality id="90D53C2EC39C3AD600C1DA74C980FC88">Damavand area</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:6BA12FA2395B570193927134C049CA76:493F0732B732A456CB82CA23484035E1" country="Iran" latitude="35.666668" longLatPrecision="1191" longitude="52.45" municipality="Damavand area" name="Aroo" stateProvince="Tehran">Aroo</location>
<location id="493F0732B732A456CB82CA23484035E1" LSID="urn:lsid:plazi:treatment:6BA12FA2395B570193927134C049CA76:493F0732B732A456CB82CA23484035E1" country="Iran" latitude="35.666668" longLatPrecision="1191" longitude="52.45" municipality="Damavand area" name="Aroo" stateProvince="Tehran">Aroo</location>
,
<geoCoordinate degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate>
<geoCoordinate id="5344DEB640DF7D256AEA0EA77418550F" degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate>
,
<geoCoordinate degrees="52" direction="east" minutes="27" orientation="longitude" precision="925" value="52.45">52°27'E</geoCoordinate>
<geoCoordinate id="4B708A745A5BC4B618FDA5EDA34D2C3A" degrees="52" direction="east" minutes="27" orientation="longitude" precision="925" value="52.45">52°27'E</geoCoordinate>
,
<collectingDate value="2000-06-15">15.06.2000</collectingDate>
<collectingDate id="E0838B13B4549F8BAFB96F0992CC964D" value="2000-06-15">15.06.2000</collectingDate>
(
<collectorName>Y.M. Marusik</collectorName>
<collectorName id="A2B77B8CB0D3ABB20B67B6A3974E9A16">Y.M. Marusik</collectorName>
&amp;
<collectorName>F. Mozaffarian</collectorName>
<collectorName id="002AF8DD71BD1B6E535301B45B808F9E">F. Mozaffarian</collectorName>
)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145" start="Figure 26" startId="F26">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 26.</emphasis>
<caption id="79078603D0BB70011D02C36E8C276DF7" doi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145" start="Figure 26" startId="F26">
<paragraph id="6397AE631761A593F5DE53E29335823C" pageId="0" pageNumber="145">
<emphasis id="74290CE0C29141EF4D334EDE75F377EA" bold="true" pageId="0" pageNumber="145">Figure 26.</emphasis>
Males of
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion raddei" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei">
<emphasis italics="true" pageId="0" pageNumber="145">Parazodarion raddei</emphasis>
<taxonomicName id="53888DAD3B26F74971BE194646EC67CE" baseAuthorityName="Simon" baseAuthorityYear="1889" family="Zodariidae" genus="Parazodarion" kingdom="Animalia" lsidName="Parazodarion raddei" order="Aranei" pageId="0" pageNumber="145" rank="species" species="raddei">
<emphasis id="BA2B8D1A20B1511C9B6B0F8FBFAB9BE9" italics="true" pageId="0" pageNumber="145">Parazodarion raddei</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
<emphasis id="87377BBFFDC88971BF7053C50D4462DE" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis>
<taxonomicName id="011C6F53169E88E8CC49E484D6CF6EFE" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum">
<emphasis id="358C7F2FCD8BCA15F8179C831EAD725C" italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
<emphasis id="FB46C6C08C818740010538D535AF4FE4" bold="true" pageId="0" pageNumber="145">E</emphasis>
) and female of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus susianus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="susianus">
<emphasis italics="true" pageId="0" pageNumber="145">Trygetus susianus</emphasis>
<taxonomicName id="FCA9836621A214A2BA802BC116AD3AC1" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus susianus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="susianus">
<emphasis id="C78F4B0A00CDC821421CDF759FCC58BE" italics="true" pageId="0" pageNumber="145">Trygetus susianus</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
<emphasis id="9F6FC6D31C359C233E4DE3FDC0EA9508" bold="true" pageId="0" pageNumber="145">D</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis>
<emphasis id="45F0C679842D9474C9B0CD48FDE899DD" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
habitus, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">B, C</emphasis>
<emphasis id="E54984AB214FB1CE00579F53F7A4A043" bold="true" pageId="0" pageNumber="145">B, C</emphasis>
palp, ventral and retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
<emphasis id="495E8E2EE9E0A691F161B3AD9F3A6B78" bold="true" pageId="0" pageNumber="145">E</emphasis>
chelicerae, ventral. Scale bars:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
<quantity id="83CA1B1261E4053A156C8EE21DD7993E" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
, unless stated otherwise.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in iran">
<paragraph pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="97C576AF1F265137BE68ADB647195D9E" pageId="0" pageNumber="145" type="records in iran">
<paragraph id="83EED1989E8ED1C75855F70FB9B5DC56" pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph id="CA77D57D33F2884AE5F4BE1D928365BA" pageId="0" pageNumber="145">
Alborz, Isfahan, Qom, Razavi Khorasan, Sistan and Baluchistan, West Azerbaijan, Yazd, Zanjan (
<bibRefCitation DOI="https://doi.org/10.2478/v10058-009-0020-z" author="Ovtchinnikov, SV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="145" pagination="471 - 474" refId="B19" refString="Ovtchinnikov, SV, Ahmad, B, Gurko, VO, 2009. Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia. Vestnik Zoologii 43 (5): 471 - 474, DOI: https://doi.org/10.2478/v10058-009-0020-z" title="Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia." url="https://doi.org/10.2478/v10058-009-0020-z" volume="43" year="2009">Ovtchinnikov et al. 2009</bibRefCitation>
<bibRefCitation id="2D27BF154405077BEB65A8A256CE9308" DOI="https://doi.org/10.2478/v10058-009-0020-z" author="Ovtchinnikov, SV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="145" pagination="471 - 474" refId="B19" refString="Ovtchinnikov, SV, Ahmad, B, Gurko, VO, 2009. Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia. Vestnik Zoologii 43 (5): 471 - 474, DOI: https://doi.org/10.2478/v10058-009-0020-z" title="Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia." url="https://doi.org/10.2478/v10058-009-0020-z" volume="43" year="2009">Ovtchinnikov et al. 2009</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3906/zoo-1307-16" author="Hosseini, M" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="145" pagination="437 - 443" refId="B6" refString="Hosseini, M, Mirshamsi, O, Kashefi, R, Fekrat, L, 2014. A contribution to the knowledge of spiders in wheat fields of Khorasan-e-Razavi Province, Iran. Turkish Journal of Zoology 38 (4): 437 - 443, DOI: https://doi.org/10.3906/zoo-1307-16" title="A contribution to the knowledge of spiders in wheat fields of Khorasan-e-Razavi Province, Iran." url="https://doi.org/10.3906/zoo-1307-16" volume="38" year="2014">Hosseini et al. 2014</bibRefCitation>
<bibRefCitation id="1E53F4ED3D3A6D3A52CADCCB1765D0AF" DOI="https://doi.org/10.3906/zoo-1307-16" author="Hosseini, M" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="145" pagination="437 - 443" refId="B6" refString="Hosseini, M, Mirshamsi, O, Kashefi, R, Fekrat, L, 2014. A contribution to the knowledge of spiders in wheat fields of Khorasan-e-Razavi Province, Iran. Turkish Journal of Zoology 38 (4): 437 - 443, DOI: https://doi.org/10.3906/zoo-1307-16" title="A contribution to the knowledge of spiders in wheat fields of Khorasan-e-Razavi Province, Iran." url="https://doi.org/10.3906/zoo-1307-16" volume="38" year="2014">Hosseini et al. 2014</bibRefCitation>
;
<bibRefCitation author="Sadeghi, H" journalOrPublisher="Biharean Biologist" pageId="0" pageNumber="145" pagination="4 - 7" refId="B23" refString="Sadeghi, H, Ahmadi, M, Zamani, A, Jabaleh, I, 2016. A study on the spider fauna of Dargaz and Kalat Counties in Razavi Khorasan Province, Iran (Arachnida: Araneae). Biharean Biologist 10 (1): 4 - 7" title="A study on the spider fauna of Dargaz and Kalat Counties in Razavi Khorasan Province, Iran (Arachnida: Araneae)." volume="10" year="2016">Sadeghi et al. 2016</bibRefCitation>
<bibRefCitation id="A78F2147FC9B808A67BE3B30D73E2597" author="Sadeghi, H" journalOrPublisher="Biharean Biologist" pageId="0" pageNumber="145" pagination="4 - 7" refId="B23" refString="Sadeghi, H, Ahmadi, M, Zamani, A, Jabaleh, I, 2016. A study on the spider fauna of Dargaz and Kalat Counties in Razavi Khorasan Province, Iran (Arachnida: Araneae). Biharean Biologist 10 (1): 4 - 7" title="A study on the spider fauna of Dargaz and Kalat Counties in Razavi Khorasan Province, Iran (Arachnida: Araneae)." volume="10" year="2016">Sadeghi et al. 2016</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation>
<bibRefCitation id="DD0CF3E1BB756B0FB081A4B18E7EFDCA" DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation>
;
<bibRefCitation author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. 2018</bibRefCitation>
<bibRefCitation id="5BF1018429C6E25350DDEF31648D8EDC" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. 2018</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.67.67" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="67 - 75" refId="B28" refString="Zamani, A, Marusik, YM, 2018. The first report on the spider fauna (Arachnida: Araneae) of the Lut Desert, Iran. Acta Arachnologica 67 (2): 67 - 75, DOI: https://doi.org/10.2476/asjaa.67.67" title="The first report on the spider fauna (Arachnida: Araneae) of the Lut Desert, Iran." url="https://doi.org/10.2476/asjaa.67.67" volume="67" year="2018">Zamani and Marusik 2018</bibRefCitation>
<bibRefCitation id="E789439F10D51B604C37BC87A72816BB" DOI="https://doi.org/10.2476/asjaa.67.67" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="67 - 75" refId="B28" refString="Zamani, A, Marusik, YM, 2018. The first report on the spider fauna (Arachnida: Araneae) of the Lut Desert, Iran. Acta Arachnologica 67 (2): 67 - 75, DOI: https://doi.org/10.2476/asjaa.67.67" title="The first report on the spider fauna (Arachnida: Araneae) of the Lut Desert, Iran." url="https://doi.org/10.2476/asjaa.67.67" volume="67" year="2018">Zamani and Marusik 2018</bibRefCitation>
). New records for Hamedan and Tehran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
<figureCitation id="36370E523861F15B8D52BF34178CE7BD" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in turkmenistan">
<paragraph pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="6462EFA650493E3213A318B36233EF1F" pageId="0" pageNumber="145" type="records in turkmenistan">
<paragraph id="50DECEDC4FA9829F075E657AF8C72546" pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph>
<paragraph id="5B84CF0C11B7A0F4BE7CBAEC112F2172" pageId="0" pageNumber="145">
Ahal, Archman (=
<normalizedToken originalValue="Arçman">Arcman</normalizedToken>
<normalizedToken id="31C02D60722A9267F6E78CB2F7D6A39E" originalValue="Arçman">Arcman</normalizedToken>
), Ashgabat, Balkan, Lebap, Mary (
<bibRefCitation author="Simon, E" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="145" pagination="373 - 386" refId="B24" refString="Simon, E, 1889. Arachnidae transcaspicae ab ill. Dr. G. Radde, Dr. A. Walter et A. Conchin inventae (annis 1886-1887). Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 39: 373 - 386" title="Arachnidae transcaspicae ab ill. Dr. G. Radde, Dr. A. Walter et A. Conchin inventae (annis 1886 - 1887)." volume="39" year="1889">Simon 1889</bibRefCitation>
<bibRefCitation id="9BF8EC7BF7F09C86EE6953E63FECDCB6" author="Simon, E" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="145" pagination="373 - 386" refId="B24" refString="Simon, E, 1889. Arachnidae transcaspicae ab ill. Dr. G. Radde, Dr. A. Walter et A. Conchin inventae (annis 1886-1887). Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 39: 373 - 386" title="Arachnidae transcaspicae ab ill. Dr. G. Radde, Dr. A. Walter et A. Conchin inventae (annis 1886 - 1887)." volume="39" year="1889">Simon 1889</bibRefCitation>
;
<bibRefCitation author="Vlassov, JP" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" pagination="247 - 258" refId="B25" refString="Vlassov, JP, Sytshevskaja, VI, 1937. The spiders of the neighbourhood of Ashkhabad. Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja 9: 247 - 258" title="The spiders of the neighbourhood of Ashkhabad." volume="9" year="1937">Vlassov and Sytshevskaja 1937</bibRefCitation>
<bibRefCitation id="ABD70865005F05A55F31E8A5FF298A67" author="Vlassov, JP" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" pagination="247 - 258" refId="B25" refString="Vlassov, JP, Sytshevskaja, VI, 1937. The spiders of the neighbourhood of Ashkhabad. Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja 9: 247 - 258" title="The spiders of the neighbourhood of Ashkhabad." volume="9" year="1937">Vlassov and Sytshevskaja 1937</bibRefCitation>
;
<bibRefCitation author="Ovtsharenko, VI" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="0" pageNumber="145" pagination="442 - 447" refId="B20" refString="Ovtsharenko, VI, Fet, VY, 1980. Fauna and ecology of spiders (Aranei) of Badhyz (Turkmenian SSR). Entomologicheskoe Obozrenie 59: 442 - 447" title="Fauna and ecology of spiders (Aranei) of Badhyz (Turkmenian SSR)." volume="59" year="1980">Ovtsharenko and Fet 1980</bibRefCitation>
<bibRefCitation id="8CC21B300DFDF875763DB72677234E5C" author="Ovtsharenko, VI" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="0" pageNumber="145" pagination="442 - 447" refId="B20" refString="Ovtsharenko, VI, Fet, VY, 1980. Fauna and ecology of spiders (Aranei) of Badhyz (Turkmenian SSR). Entomologicheskoe Obozrenie 59: 442 - 447" title="Fauna and ecology of spiders (Aranei) of Badhyz (Turkmenian SSR)." volume="59" year="1980">Ovtsharenko and Fet 1980</bibRefCitation>
;
<bibRefCitation author="Fet, VY" journalOrPublisher="Senckenbergiana Biologica" pageId="0" pageNumber="145" refId="B4" refString="Fet, VY, 1985. [The ecological distribution of spiders in Syunt Khassardagh Reserve]. In: [The Vegetation and Animal Life of Western Kopetdagh]. Ashkhabad, 271-277. [in Russian]" title="[The ecological distribution of spiders in Syunt Khassardagh Reserve]. In: [The Vegetation and Animal Life of Western Kopetdagh]. Ashkhabad, 271 - 277. [in Russian]" year="1985">Fet 1985</bibRefCitation>
<bibRefCitation id="0806912218825B1BD7FF114007E3839C" author="Fet, VY" journalOrPublisher="Senckenbergiana Biologica" pageId="0" pageNumber="145" refId="B4" refString="Fet, VY, 1985. [The ecological distribution of spiders in Syunt Khassardagh Reserve]. In: [The Vegetation and Animal Life of Western Kopetdagh]. Ashkhabad, 271-277. [in Russian]" title="[The ecological distribution of spiders in Syunt Khassardagh Reserve]. In: [The Vegetation and Animal Life of Western Kopetdagh]. Ashkhabad, 271 - 277. [in Russian]" year="1985">Fet 1985</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1007/978-94-011-1116-4_30" author="Mikhailov, KG" editor="Fet, V" journalOrPublisher="Kluwer Academic Publisher Netherlands" pageId="0" pageNumber="145" pagination="499 - 524" refId="B16" refString="Mikhailov, KG, Fet, V, 1994. Fauna and zoogeography of spiders (Aranei) of Turkmenistan. In: Fet, V, Atamuradov, KI, Eds., Biogeography and Ecology of Turkmenistan. Kluwer Academic Publisher Netherlands: 499 - 524, DOI: https://doi.org/10.1007/978-94-011-1116-4_30" title="Fauna and zoogeography of spiders (Aranei) of Turkmenistan." url="https://doi.org/10.1007/978-94-011-1116-4_30" volumeTitle="Biogeography and Ecology of Turkmenistan." year="1994">Mikhailov and Fet 1994</bibRefCitation>
<bibRefCitation id="22D36A319827560B3129DE24675D3868" DOI="https://doi.org/10.1007/978-94-011-1116-4_30" author="Mikhailov, KG" editor="Fet, V" journalOrPublisher="Kluwer Academic Publisher Netherlands" pageId="0" pageNumber="145" pagination="499 - 524" refId="B16" refString="Mikhailov, KG, Fet, V, 1994. Fauna and zoogeography of spiders (Aranei) of Turkmenistan. In: Fet, V, Atamuradov, KI, Eds., Biogeography and Ecology of Turkmenistan. Kluwer Academic Publisher Netherlands: 499 - 524, DOI: https://doi.org/10.1007/978-94-011-1116-4_30" title="Fauna and zoogeography of spiders (Aranei) of Turkmenistan." url="https://doi.org/10.1007/978-94-011-1116-4_30" volumeTitle="Biogeography and Ecology of Turkmenistan." year="1994">Mikhailov and Fet 1994</bibRefCitation>
;
<bibRefCitation author="Mikhailov, KG" journalOrPublisher="Zoological Museum, Moscow State University Moscow" pageId="0" pageNumber="145" refId="B14" refString="Mikhailov, KG, 1997. Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei). Zoological Museum, Moscow State University Moscow" title="Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei)." year="1997">Mikhailov 1997</bibRefCitation>
<bibRefCitation id="E74E917CC92DD771F14BDCF9A2787EB9" author="Mikhailov, KG" journalOrPublisher="Zoological Museum, Moscow State University Moscow" pageId="0" pageNumber="145" refId="B14" refString="Mikhailov, KG, 1997. Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei). Zoological Museum, Moscow State University Moscow" title="Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei)." year="1997">Mikhailov 1997</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.2478/v10058-009-0020-z" author="Ovtchinnikov, SV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="145" pagination="471 - 474" refId="B19" refString="Ovtchinnikov, SV, Ahmad, B, Gurko, VO, 2009. Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia. Vestnik Zoologii 43 (5): 471 - 474, DOI: https://doi.org/10.2478/v10058-009-0020-z" title="Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia." url="https://doi.org/10.2478/v10058-009-0020-z" volume="43" year="2009">Ovtchinnikov et al. 2009</bibRefCitation>
<bibRefCitation id="CFD90574522250FE61E79EBBF2923298" DOI="https://doi.org/10.2478/v10058-009-0020-z" author="Ovtchinnikov, SV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="145" pagination="471 - 474" refId="B19" refString="Ovtchinnikov, SV, Ahmad, B, Gurko, VO, 2009. Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia. Vestnik Zoologii 43 (5): 471 - 474, DOI: https://doi.org/10.2478/v10058-009-0020-z" title="Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia." url="https://doi.org/10.2478/v10058-009-0020-z" volume="43" year="2009">Ovtchinnikov et al. 2009</bibRefCitation>
) (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
<figureCitation id="976ED5E46F45C5950E611DA32479E1F3" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">From United Arab Emirates to Kazakhstan and Afghanistan in the east.</paragraph>
<subSubSection id="0EEEB13E047CAA9EE25F90B140FA93CC" pageId="0" pageNumber="145" type="distribution">
<paragraph id="DCA3B6F4DEBC16FEBD789A21427946EF" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="B87F88B6FCD637FA7C3D37EFE632227D" pageId="0" pageNumber="145">From United Arab Emirates to Kazakhstan and Afghanistan in the east.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,158 +1,158 @@
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:title id="ED62FFFDD789E12CDFCE54EE14C8CDEB">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:namePart>Zamani, Alireza</mods:namePart>
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<mods:namePart>Marusik, Yuri M.</mods:namePart>
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<mods:affiliation id="D9C7C12EDC219E3DC0A33444BBE7E658">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<treatment ID-GBIF-Taxon="181753555" LSID="urn:lsid:zoobank.org:act:3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" httpUri="http://treatment.plazi.org/id/75080B2F5CE955999C9274B037E542BE" lastPageNumber="145" pageId="0" pageNumber="145">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">Figs 1A-C</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">, 3A-E</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">, 4I</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
<treatment id="75080B2F5CE955999C9274B037E542BE" ID-GBIF-Taxon="181753555" LSID="urn:lsid:zoobank.org:act:3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" httpUri="http://treatment.plazi.org/id/75080B2F5CE955999C9274B037E542BE" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Lachesaninae">
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<paragraph id="0652F93F8648253B38C99090BAA5A5F4" pageId="0" pageNumber="145">
<taxonomicName id="FD2D5B0ACF69683FA3B9F0CD925AA485" ID-CoL="98YYV" LSID="http://zoobank.org/3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis" status="sp. nov.">Lachesana kavirensis</taxonomicName>
<taxonomicNameLabel id="6F9266FD992B194DB159817F468C6961" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="1EBCAC53707311EA5F299A866C33C333" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">Figs 1A-C</figureCitation>
<figureCitation id="7D8D8367E734BC9A24BF4370318190D5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">, 3A-E</figureCitation>
<figureCitation id="3620ED69DCD6F826F7E6B13D8DD38725" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">, 4I</figureCitation>
<figureCitation id="4E0449A87159DED2C316FDAF4582E224" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="5C8A2B03B4B9250026BF42C87C0DBA01" pageId="0" pageNumber="145" type="type material">
<paragraph id="40EEE7B48C74BE9FB6263AD1CBAC29F8" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="362367AE0476F7FE6F33BA14E2207BA5" pageId="0" pageNumber="145">
<emphasis id="5F294A06E0007CB5FDC8EFF7F5F77BAA" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Qom Province</emphasis>
<emphasis id="063C512755E185B0C5D495855D7160F1" italics="true" pageId="0" pageNumber="145">Qom Province</emphasis>
: Dasht-e Masileh,
<geoCoordinate degrees="34" direction="north" minutes="47" orientation="latitude" precision="925" value="34.783333">34°47'N</geoCoordinate>
<geoCoordinate id="729F8F592B18821BCE57EF5B16A2E7A7" degrees="34" direction="north" minutes="47" orientation="latitude" precision="925" value="34.783333">34°47'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="11" orientation="longitude" precision="925" value="51.183334">51°11'E</geoCoordinate>
<geoCoordinate id="3284D87A06E53983F4B8EA878A52C191" degrees="51" direction="east" minutes="11" orientation="longitude" precision="925" value="51.183334">51°11'E</geoCoordinate>
, 11.2017 (M. Mirghazanfari).
</paragraph>
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<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="43A9C13ED5E96F9550DE1616D3789377" pageId="0" pageNumber="145" type="etymology">
<paragraph id="1E6E5A1FB6B6229D70CCDE963B31603C" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="ECD674A6220A7BF754CEACBB2AE0814D" pageId="0" pageNumber="145">
The specific epithet is derived from
<normalizedToken originalValue="Kavir">'Kavir'</normalizedToken>
<normalizedToken id="3320C6189828C831E44BB2136D4B772C" originalValue="Kavir">'Kavir'</normalizedToken>
, a Persian word for desert, referring to the occurrence of the species in the central deserts of Iran.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="1F81D003D25F702331063A2445B67BBA" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="C0899A83683D2FC5E6470B7DDE141D49" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="8D39B779133E88E64E924F91A5735C1E" pageId="0" pageNumber="145">
The male of
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<emphasis italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
<taxonomicName id="AFE5CE602157642213A101E334CCD8C6" lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis id="1EF70A5F5F24B1B44FE46320B724BC43" italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
</taxonomicName>
sp. nov. can be distinguished from the similar
<taxonomicName lsidName="L. dyachkovi" pageId="0" pageNumber="145" rank="species" species="dyachkovi">
<emphasis italics="true" pageId="0" pageNumber="145">L. dyachkovi</emphasis>
<taxonomicName id="C1B7658179D4C32CA663F6574DF32EE3" lsidName="L. dyachkovi" pageId="0" pageNumber="145" rank="species" species="dyachkovi">
<emphasis id="3A7180F9435650C08A0F9936F76B1254" italics="true" pageId="0" pageNumber="145">L. dyachkovi</emphasis>
</taxonomicName>
Fomichev &amp; Marusik, 2019 by having a gently bent cheliceral fang (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="9E6831C9F200F912EBC9329F42D2B5F0" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
bent over 90°, cf. Fig.
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and fig. 15 in
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.28.4.08" author="Fomichev, AA" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="556 - 561" refId="B5" refString="Fomichev, AA, Marusik, YM, 2019. A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan. Arthropoda Selecta 28 (4): 556 - 561, DOI: https://doi.org/10.15298/arthsel.28.4.08" title="A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan." url="https://doi.org/10.15298/arthsel.28.4.08" volume="28" year="2019">Fomichev and Marusik 2019</bibRefCitation>
<bibRefCitation id="0301D11C0FA22D61052F96E758AE6C5E" DOI="https://doi.org/10.15298/arthsel.28.4.08" author="Fomichev, AA" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="556 - 561" refId="B5" refString="Fomichev, AA, Marusik, YM, 2019. A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan. Arthropoda Selecta 28 (4): 556 - 561, DOI: https://doi.org/10.15298/arthsel.28.4.08" title="A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan." url="https://doi.org/10.15298/arthsel.28.4.08" volume="28" year="2019">Fomichev and Marusik 2019</bibRefCitation>
). The shape of the RTA is also different from other species of the region (stalk with a gentle curve ventrally and tip straightly pointed; cf. Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4I</figureCitation>
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and Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4F-H, J</figureCitation>
<figureCitation id="1E0695FF0E08863C57682C767F4B6717" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4F-H, J</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 1.</emphasis>
<caption id="6E02DF173E7CCAA75D1160FD8E3AE8BC" doi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145" start="Figure 1" startId="F1">
<paragraph id="70A7462D885D40811A3429CF85B87B7E" pageId="0" pageNumber="145">
<emphasis id="89C53FD05330D9D2077A9D7083CAF455" bold="true" pageId="0" pageNumber="145">Figure 1.</emphasis>
Habitus and chelicerae of male
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana kavirensis</emphasis>
<taxonomicName id="6F067D0A69C2959CB658B05CE5A33D62" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis">
<emphasis id="983FFCDC3921BDEFC278449D0B8D47F9" italics="true" pageId="0" pageNumber="145">Lachesana kavirensis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
<emphasis id="F48042E2001767800412B211E5F7AA9A" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and
<taxonomicName lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
<taxonomicName id="29138CFD20DAA6C163A6C315F04A635F" lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis id="E0445D307571C5E8230322D4C98CB5EB" italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
<emphasis id="4524C988A95A19F56DA00C5EF7023E96" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, B, D, E</emphasis>
<emphasis id="94212D215E4FB0E23E79327BDEF48081" bold="true" pageId="0" pageNumber="145">A, B, D, E</emphasis>
habitus, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">C, F</emphasis>
<emphasis id="F32153E33FD758F95B2139BEEA32BC42" bold="true" pageId="0" pageNumber="145">C, F</emphasis>
chelicerae, ventral.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
<subSubSection id="CAADF2ECB101A1BC983F8F3B31A3CD1C" pageId="0" pageNumber="145" type="description">
<paragraph id="13CDC11543B4E7C5463C451ABCD70CCC" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="2259A8733FDE5B904D5BF6F639983779" pageId="0" pageNumber="145">
<emphasis id="C221BC61D6EDF4B20A3FF93023EFCA67" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1A, B</figureCitation>
<figureCitation id="A679570090A25AC627FE334E723AF8DB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1A, B</figureCitation>
. Total length 11.55. Carapace 5.80 long, 2.60 wide at pars cephalica, 3.75 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.23, ALE: 0.20, PME: 0.17, PLE: 0.23, PME-PME: 0.14. Carapace, sternum, labium, chelicerae and maxillae light brown. Chelicera (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1C</figureCitation>
<figureCitation id="E0CE6D685589F3867E1F35B4E4479CBE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1C</figureCitation>
) with dense, black setae and one promarginal tooth; fangs gently curved, almost straight. Legs the same color as carapace, without annulations and with numerous spines. Abdomen pale, dorsally with a longitudinal dark stripe and several small spots. Spinnerets slightly lighter than abdomen, uniform in color. Measurements of legs: I: 15.49 (4.47, 1.88, 3.33, 3.29, 2.52), II: 15.03 (4.20, 1.79, 3.02, 3.50, 2.52), III: 15.78 (4.08, 1.96, 2.48, 4.71, 2.55), IV: 18.30 (4.58, 2.07, 3.20, 5.24, 3.21).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="304E1DF7FAA0F3E0AA1213524791DAFC" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3A-E</figureCitation>
<figureCitation id="81D0B86A58E887F9B50E48105E1A8DA9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3A-E</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4I</figureCitation>
<figureCitation id="E48303C1CB0CE47198597D0A987CE857" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4I</figureCitation>
. RTA long, over 1.2 times longer than tibia, its widest part less than 2 times wider than stalk; cymbium with 2 spines in mesal proximal part; bulb elongate, 1.3 times longer than wide; base of embolus large, 1/3 of the tegulum's length; embolus filamentous, lying in chute formed by conductor.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="8D0F520E623C595BDB6A4BF97F263548" pageId="0" pageNumber="145">
<emphasis id="3E3506B042F14DB6D86A3B4879E5FC4D" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="93138FA7613F60626D020CBFA0315E7F" pageId="0" pageNumber="145" type="distribution">
<paragraph id="349676EA43DCC9D6FE7D20BC5FDAF437" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="F8D01D058C16801AC1AB2064130704D7" pageId="0" pageNumber="145">
Known only from the type locality in Qom Province, northern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
<figureCitation id="4821E25AC0408CC2280B594D11AC825A" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,92 +1,92 @@
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<figureCitation id="5242B22207A8B094F2ABC477E98A30AD" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">Fig. 33</figureCitation>
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:
<bibRefCitation author="Bosmans, R" journalOrPublisher="Contributions to Natural History" pageId="0" pageNumber="145" pagination="211 - 295" refId="B1" refString="Bosmans, R, 2009. Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). Contributions to Natural History 12: 211 - 295" title="Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae)." volume="12" year="2009">Bosmans 2009</bibRefCitation>
<bibRefCitation id="B4D612A916BCB3894A8E0B473FEECA0C" author="Bosmans, R" journalOrPublisher="Contributions to Natural History" pageId="0" pageNumber="145" pagination="211 - 295" refId="B1" refString="Bosmans, R, 2009. Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). Contributions to Natural History 12: 211 - 295" title="Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae)." volume="12" year="2009">Bosmans 2009</bibRefCitation>
: 281, f. 184-185, 194-195 (♂♀).
</paragraph>
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<taxonomicName baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion lutipes" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="lutipes">Zodarion lutipes</taxonomicName>
<paragraph id="82A33F71C7681F7F281EABF729421A6D" pageId="0" pageNumber="145">
<taxonomicName id="0A2595A380CA604883195AE127F61944" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion lutipes" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="lutipes">Zodarion lutipes</taxonomicName>
For the complete list of references see
<bibRefCitation author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC (2021)</bibRefCitation>
<bibRefCitation id="B9F140603BD741E771DE97C925926507" author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC (2021)</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in iran">
<paragraph pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="C367FCCD3AEA043D79EC0F81A6EDFD11" pageId="0" pageNumber="145" type="records in iran">
<paragraph id="526EE248414511F3FBF1CD00411D67CD" pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph id="D8B0BDCCB18956560BBC702D343B97D5" pageId="0" pageNumber="145">
Kermanshah (
<bibRefCitation author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. 2018</bibRefCitation>
<bibRefCitation id="C3FE159D89E8DE58294D4A8605106C46" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. 2018</bibRefCitation>
) (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
<figureCitation id="A7FDFB48732D4B197C09EF6154C606FA" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">Cyprus, Israel, Lebanon, Jordan, Iran.</paragraph>
<subSubSection id="8BECA0E90DC1BF62AE0A6CCD601AD5DE" pageId="0" pageNumber="145" type="distribution">
<paragraph id="FED0E0824A846302B36454FBC67ED893" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="E17A484BCC7B356DAA73281F1CC5EDBB" pageId="0" pageNumber="145">Cyprus, Israel, Lebanon, Jordan, Iran.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="8B14EB2B27A30EFECB92DF3FDA613D8F" pageId="0" pageNumber="145" type="comments">
<paragraph id="0A4B28D550755EB70FD75067633729A1" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph id="CA2DDD2CB6D56872CFA4E5214B9983F2" pageId="0" pageNumber="145">
Unfortunately, we were not able to re-examine the material studied by
<bibRefCitation author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. (2018)</bibRefCitation>
<bibRefCitation id="CF57EAF2568E85D11B31A44276F031B5" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. (2018)</bibRefCitation>
, therefore, no figures are provided for this species.
</paragraph>
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<mods:identifier id="4C72F111915E644E4439DDBDFCAAA768" type="Pensoft-UUID">55CAE096B4A451D88EA433175454DA3F</mods:identifier>
</mods:mods>
<subSection pageId="0" pageNumber="145" type="taxonomy">
<treatment ID-GBIF-Taxon="181753574" LSID="urn:lsid:plazi:treatment:8837EB822336513D8735EAA812AFC392" httpUri="http://treatment.plazi.org/id/8837EB822336513D8735EAA812AFC392" lastPageNumber="145" pageId="0" pageNumber="145">
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145">
<taxonomicName LSID="8837EB82-2336-513D-8735-EAA812AFC392" authority="Dunin &amp; Nenilin, 1987" authorityName="Dunin &amp; Nenilin" authorityYear="1987" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion talyschicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyschicum">Zodarion talyschicum Dunin &amp; Nenilin, 1987</taxonomicName>
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A) and Z. talyschicum (B, C) A, B ventral C ventroretrolateral. Abbreviations: Tm - median tegular process, Tp - prolateral tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145">Figs 29B, C</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
<subSection id="48D53AB326946B3B236BB260F90E85A5" pageId="0" pageNumber="145" type="taxonomy">
<treatment id="8837EB822336513D8735EAA812AFC392" ID-GBIF-Taxon="181753574" LSID="urn:lsid:plazi:treatment:8837EB822336513D8735EAA812AFC392" httpUri="http://treatment.plazi.org/id/8837EB822336513D8735EAA812AFC392" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection id="5F7894CD2F9A79BF0299F44D9573262E" pageId="0" pageNumber="145" type="nomenclature">
<paragraph id="C49AD006B9C5B4BA221D5823C27E8057" pageId="0" pageNumber="145">
<taxonomicName id="B55996BC3F78FF7E622466B8052040D4" ID-CoL="5D6DJ" LSID="8837EB82-2336-513D-8735-EAA812AFC392" authority="Dunin &amp; Nenilin, 1987" authorityName="Dunin &amp; Nenilin" authorityYear="1987" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion talyschicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyschicum">Zodarion talyschicum Dunin &amp; Nenilin, 1987</taxonomicName>
<figureCitation id="44C503980295675AB54B16A69219F0E5" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A) and Z. talyschicum (B, C) A, B ventral C ventroretrolateral. Abbreviations: Tm - median tegular process, Tp - prolateral tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145">Figs 29B, C</figureCitation>
<figureCitation id="DE85063C2E72A4959679DC0BDD5A04F5" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="reference_group">
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Dunin &amp; Nenilin" authorityYear="1987" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion talyschicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyschicum">Zodarion talyschicum</taxonomicName>
<subSubSection id="00E29607B5A138FFFA84A86496690E76" pageId="0" pageNumber="145" type="reference_group">
<paragraph id="A45B0D3D0049819ACCC297F5687614F1" pageId="0" pageNumber="145">
<taxonomicName id="C028A07D6BC4F19026CABFF529CFBF12" authorityName="Dunin &amp; Nenilin" authorityYear="1987" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion talyschicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyschicum">Zodarion talyschicum</taxonomicName>
Dunin &amp; Nenilin, 1987: 196, f. 14-18 (♂♀).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Dunin &amp; Nenilin" authorityYear="1987" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion talyschicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyschicum">Zodarion talyschicum</taxonomicName>
<paragraph id="68249AE9672CEF8DB20885309D7BC936" pageId="0" pageNumber="145">
<taxonomicName id="B805D890C67F1D392FD83D8F723F87F8" authorityName="Dunin &amp; Nenilin" authorityYear="1987" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion talyschicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyschicum">Zodarion talyschicum</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.13156/arac.2020.18.6.569" author="Zamani, A" journalOrPublisher="Arachnology" pageId="0" pageNumber="145" pagination="569 - 591" refId="B27" refString="Zamani, A, Dimitrov, D, Weiss, I, Alimohammadi, S, Rafiei-Jahed, R, Esyunin, SL, Moradmand, M, Chatzaki, M, Marusik, YM, 2020. New data on the spider fauna of Iran (Arachnida: Araneae), Part VII. Arachnology 18 (6): 569 - 591, DOI: https://doi.org/10.13156/arac.2020.18.6.569" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part VII." url="https://doi.org/10.13156/arac.2020.18.6.569" volume="18" year="2020">Zamani et al. 2020</bibRefCitation>
<bibRefCitation id="1BDD8C49D2601A97B6EB8743D4805669" DOI="https://doi.org/10.13156/arac.2020.18.6.569" author="Zamani, A" journalOrPublisher="Arachnology" pageId="0" pageNumber="145" pagination="569 - 591" refId="B27" refString="Zamani, A, Dimitrov, D, Weiss, I, Alimohammadi, S, Rafiei-Jahed, R, Esyunin, SL, Moradmand, M, Chatzaki, M, Marusik, YM, 2020. New data on the spider fauna of Iran (Arachnida: Araneae), Part VII. Arachnology 18 (6): 569 - 591, DOI: https://doi.org/10.13156/arac.2020.18.6.569" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part VII." url="https://doi.org/10.13156/arac.2020.18.6.569" volume="18" year="2020">Zamani et al. 2020</bibRefCitation>
: 589, f. 12D-F (♂♀).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="materials_examined">
<paragraph pageId="0" pageNumber="145">Material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<materialsCitation ID-GBIF-Occurrence="3093634311" collectingDate="1973-09-06" collectorName="A. Senglet" country="Iran" latitude="37.0" location="Bidjar" longLatPrecision="1184" longitude="49.566666" specimenCount="1" specimenCount-male="1" stateProvince="Gilan Province">
<collectingCountry name="Iran">Iran</collectingCountry>
<subSubSection id="B098585A59BEC6C939D33DBE7A5C0D0C" pageId="0" pageNumber="145" type="materials_examined">
<paragraph id="BAD750EA429AC774BA460FE9C484EDD3" pageId="0" pageNumber="145">Material.</paragraph>
<paragraph id="E360AE9081D538116A018FBEDBD59C97" pageId="0" pageNumber="145">
<materialsCitation id="7E3EA6BDA19801311AE12FB54AD29617" ID-GBIF-Occurrence="3093634311" collectingDate="1973-09-06" collectorName="A. Senglet" country="Iran" latitude="37.0" location="Bidjar" longLatPrecision="1184" longitude="49.566666" specimenCount="1" specimenCount-male="1" stateProvince="Gilan Province">
<collectingCountry id="1E5453AD2CBC9B0E2325933E384D279D" name="Iran">Iran</collectingCountry>
:
<emphasis italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Gilan">Gilan Province</collectingRegion>
<emphasis id="2003BD1A1725748A14825A832185E181" italics="true" pageId="0" pageNumber="145">
<collectingRegion id="5B32214C778D4A1172055BCF7001E577" country="Iran" name="Gilan">Gilan Province</collectingRegion>
</emphasis>
:
<specimenCount type="male">1♂</specimenCount>
<specimenCount id="70200D3D491253AC663D58820BD9327E" type="male">1♂</specimenCount>
(MHNG),
<location LSID="urn:lsid:plazi:treatment:8837EB822336513D8735EAA812AFC392:58CC290B857581F03D504C1A89C2FF28" country="Iran" latitude="37.0" longLatPrecision="1184" longitude="49.566666" name="Bidjar" stateProvince="Gilan Province">Bidjar</location>
<location id="58CC290B857581F03D504C1A89C2FF28" LSID="urn:lsid:plazi:treatment:8837EB822336513D8735EAA812AFC392:58CC290B857581F03D504C1A89C2FF28" country="Iran" latitude="37.0" longLatPrecision="1184" longitude="49.566666" name="Bidjar" stateProvince="Gilan Province">Bidjar</location>
,
<geoCoordinate degrees="37" direction="north" minutes="00" orientation="latitude" precision="925" value="37.0">37°00'N</geoCoordinate>
<geoCoordinate id="1E95CBE10F802AAF35340CD8003FC0D0" degrees="37" direction="north" minutes="00" orientation="latitude" precision="925" value="37.0">37°00'N</geoCoordinate>
,
<geoCoordinate degrees="49" direction="east" minutes="34" orientation="longitude" precision="925" value="49.566666">49°34'E</geoCoordinate>
<geoCoordinate id="2910219365D53F585F057F91038B559E" degrees="49" direction="east" minutes="34" orientation="longitude" precision="925" value="49.566666">49°34'E</geoCoordinate>
,
<collectingDate value="1973-09-06">06.09.1973</collectingDate>
<collectingDate id="C94C009DADAFB836AD2AC475E85065BC" value="1973-09-06">06.09.1973</collectingDate>
(
<collectorName>A. Senglet</collectorName>
<collectorName id="41F1F5B6D52210318BDC8F16C49127A2">A. Senglet</collectorName>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3093634314" collectingDate="1973-07-20" collectorName="A. Senglet" country="Iran" latitude="36.733334" location="Naharkhoran" longLatPrecision="1185" longitude="54.483334" municipality="Gorgan" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Golestan Province">
<emphasis italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Golestan">Golestan Province</collectingRegion>
<materialsCitation id="B0B137871BAFC5C96B2A4A8D809314C4" ID-GBIF-Occurrence="3093634314" collectingDate="1973-07-20" collectorName="A. Senglet" country="Iran" latitude="36.733334" location="Naharkhoran" longLatPrecision="1185" longitude="54.483334" municipality="Gorgan" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Golestan Province">
<emphasis id="F94BA5E5CB4516BAD56209D9A4411210" italics="true" pageId="0" pageNumber="145">
<collectingRegion id="55996B9A1F0899BA51B1AAE67BF4085E" country="Iran" name="Golestan">Golestan Province</collectingRegion>
</emphasis>
:
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">3♀</specimenCount>
<specimenCount id="7FBEE7FD0B816FA323C5BE322ADFF60C" type="male">1♂</specimenCount>
<specimenCount id="1B3ABE6E631FAD5C5CD439C23C518A25" type="female">3♀</specimenCount>
(MHNG),
<collectingMunicipality>Gorgan</collectingMunicipality>
<collectingMunicipality id="9FD6C8B771D0EB4502EEF8B5EF16EAAA">Gorgan</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:8837EB822336513D8735EAA812AFC392:6E2DA9D776891C7A93329FD8CBDC4692" country="Iran" latitude="36.733334" longLatPrecision="1185" longitude="54.483334" municipality="Gorgan" name="Naharkhoran" stateProvince="Golestan Province">Naharkhoran</location>
<location id="6E2DA9D776891C7A93329FD8CBDC4692" LSID="urn:lsid:plazi:treatment:8837EB822336513D8735EAA812AFC392:6E2DA9D776891C7A93329FD8CBDC4692" country="Iran" latitude="36.733334" longLatPrecision="1185" longitude="54.483334" municipality="Gorgan" name="Naharkhoran" stateProvince="Golestan Province">Naharkhoran</location>
,
<geoCoordinate degrees="36" direction="north" minutes="44" orientation="latitude" precision="925" value="36.733334">36°44'N</geoCoordinate>
<geoCoordinate id="8513F498D98AF19A25E5C51049DFFC00" degrees="36" direction="north" minutes="44" orientation="latitude" precision="925" value="36.733334">36°44'N</geoCoordinate>
,
<geoCoordinate degrees="54" direction="east" minutes="29" orientation="longitude" precision="925" value="54.483334">54°29'E</geoCoordinate>
<geoCoordinate id="B40CC35B3A34B85F5AA4098611AA3701" degrees="54" direction="east" minutes="29" orientation="longitude" precision="925" value="54.483334">54°29'E</geoCoordinate>
,
<collectingDate value="1973-07-20">20.07.1973</collectingDate>
<collectingDate id="474C02B19517031C28284873AF720634" value="1973-07-20">20.07.1973</collectingDate>
(
<collectorName>A. Senglet</collectorName>
<collectorName id="8367590D5B8CDDC255D85157CE55A653">A. Senglet</collectorName>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3093634315" collectingDate="1972-04-26" collectorName="G. Pretzmann" country="Iran" location="10 km SW of Shahpasand" specimenCount="1" specimenCount-male="1" stateProvince="Hamedan Province">
<emphasis italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Hamadan">Hamedan Province</collectingRegion>
<materialsCitation id="A0694ABAAB8E4C8CC3B3B8F2931B198A" ID-GBIF-Occurrence="3093634315" collectingDate="1972-04-26" collectorName="G. Pretzmann" country="Iran" location="10 km SW of Shahpasand" specimenCount="1" specimenCount-male="1" stateProvince="Hamedan Province">
<emphasis id="1107DD41BCD75258A8E197DBBBED4B1D" italics="true" pageId="0" pageNumber="145">
<collectingRegion id="50EC0810C97FF242D66F7EE452858AD0" country="Iran" name="Hamadan">Hamedan Province</collectingRegion>
</emphasis>
:
<specimenCount type="male">1♂</specimenCount>
<specimenCount id="BD510DBC998BE5D99D5C3F0AC9BD9AEB" type="male">1♂</specimenCount>
(NHMW),
<location LSID="urn:lsid:plazi:treatment:8837EB822336513D8735EAA812AFC392:73633F66EE6431725B7961B53D455C81" country="Iran" name="10 km SW of Shahpasand" stateProvince="Hamedan Province">
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
<location id="73633F66EE6431725B7961B53D455C81" LSID="urn:lsid:plazi:treatment:8837EB822336513D8735EAA812AFC392:73633F66EE6431725B7961B53D455C81" country="Iran" name="10 km SW of Shahpasand" stateProvince="Hamedan Province">
<quantity id="DB231006CF5CFD55A070FD3BB7670C9F" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
SW of Shahpasand
</location>
,
<collectingDate value="1972-04-26">26.04.1972</collectingDate>
<collectingDate id="F9833F874D3EE6C1C8CC412293CBA9E6" value="1972-04-26">26.04.1972</collectingDate>
(
<collectorName>G. Pretzmann</collectorName>
<collectorName id="0DB7590360BE868CE99DD11CE3C69533">G. Pretzmann</collectorName>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3093634309" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Iran" location="Mazandaran Province" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mazandaran Province">
<emphasis italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Mazandaran">Mazandaran Province</collectingRegion>
<materialsCitation id="F44C02A6526C00283D966CBF5D61ABE4" ID-GBIF-Occurrence="3093634309" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Iran" location="Mazandaran Province" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mazandaran Province">
<emphasis id="E8A97B59073FF05FD0209BB611D5C9E0" italics="true" pageId="0" pageNumber="145">
<collectingRegion id="4B97FA36557F469DA4AE8436295B0F40" country="Iran" name="Mazandaran">Mazandaran Province</collectingRegion>
</emphasis>
:
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="0E86C68E17E233B459A950D027E5AFFF" type="male">1♂</specimenCount>
<specimenCount id="D06DCB4C28CD8B9E7273A05BDB93FACA" type="female">1♀</specimenCount>
1sub
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3093634302" collectingDate="2000-06-10" collectorName="Y. M. Marusik" country="Iran" elevation="2000" latitude="36.616665" location="Barseh Vil." longLatPrecision="1186" longitude="50.683334" specimenCount="♂" stateProvince="Mazandaran Province">
<specimenCount></specimenCount>
<materialsCitation id="44443BDD62B1A01E4476AEB5D899EAA9" ID-GBIF-Occurrence="3093634302" collectingDate="2000-06-10" collectorName="Y. M. Marusik" country="Iran" elevation="2000" latitude="36.616665" location="Barseh Vil." longLatPrecision="1186" longitude="50.683334" specimenCount="♂" stateProvince="Mazandaran Province">
<specimenCount id="D8707C2DF1EF9CCB439AF75A0B1DE699"></specimenCount>
(ZMMU),
<location LSID="urn:lsid:plazi:treatment:8837EB822336513D8735EAA812AFC392:72B1CC6B2B79F194485F7485FEC6F85B" country="Iran" latitude="36.616665" longLatPrecision="1186" longitude="50.683334" name="Barseh Vil." stateProvince="Mazandaran Province">Barseh Vil.</location>
<location id="72B1CC6B2B79F194485F7485FEC6F85B" LSID="urn:lsid:plazi:treatment:8837EB822336513D8735EAA812AFC392:72B1CC6B2B79F194485F7485FEC6F85B" country="Iran" latitude="36.616665" longLatPrecision="1186" longitude="50.683334" name="Barseh Vil." stateProvince="Mazandaran Province">Barseh Vil.</location>
,
<geoCoordinate degrees="36" direction="north" minutes="37" orientation="latitude" precision="925" value="36.616665">36°37'N</geoCoordinate>
<geoCoordinate id="451A7F205FA6EC64C4995268EF3ECB74" degrees="36" direction="north" minutes="37" orientation="latitude" precision="925" value="36.616665">36°37'N</geoCoordinate>
,
<geoCoordinate degrees="50" direction="east" minutes="41" orientation="longitude" precision="925" value="50.683334">50°41'E</geoCoordinate>
<geoCoordinate id="3D1FDB7DEA0DBA1826FCD9AE081C1626" degrees="50" direction="east" minutes="41" orientation="longitude" precision="925" value="50.683334">50°41'E</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">2000 m</elevation>
<quantity id="4A0CE853E4E48F7A75D09049D2B8C755" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">
<elevation id="AB276021A9F47BADA547264E761C1BDF" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<collectingDate value="2000-06-10">10.06.2000</collectingDate>
<collectingDate id="36E2C5FB5D8078462EC7B8AD6434FB94" value="2000-06-10">10.06.2000</collectingDate>
(
<collectorName>Y.M. Marusik</collectorName>
<collectorName id="F642838F5F703025DCA7D684106B5464">Y.M. Marusik</collectorName>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3093634312" collectingDate="1974-07-26" collectorName="A. Senglet" country="Iran" latitude="37.483334" location="Bojnurd" longLatPrecision="1181" longitude="57.433334" specimenCount="1" specimenCount-male="1" stateProvince="North Khorasan Province">
<emphasis italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Khorasan-e Shomali">North Khorasan Province</collectingRegion>
<materialsCitation id="6ADD158CEA9431BCC8CFA275646C842F" ID-GBIF-Occurrence="3093634312" collectingDate="1974-07-26" collectorName="A. Senglet" country="Iran" latitude="37.483334" location="Bojnurd" longLatPrecision="1181" longitude="57.433334" specimenCount="1" specimenCount-male="1" stateProvince="North Khorasan Province">
<emphasis id="1298E7F34EEEAE565DFDC6B6739387B1" italics="true" pageId="0" pageNumber="145">
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:
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(MHNG),
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(
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.
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<paragraph pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="94B0E954FFE85CC24E556DC0AA3387BB" pageId="0" pageNumber="145" type="records in iran">
<paragraph id="CF12A06414CA9F7000F53C4DB00C3B67" pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph id="71EE795F53F87E4AA95000F81D5E38C3" pageId="0" pageNumber="145">
Golestan (
<bibRefCitation DOI="https://doi.org/10.13156/arac.2020.18.6.569" author="Zamani, A" journalOrPublisher="Arachnology" pageId="0" pageNumber="145" pagination="569 - 591" refId="B27" refString="Zamani, A, Dimitrov, D, Weiss, I, Alimohammadi, S, Rafiei-Jahed, R, Esyunin, SL, Moradmand, M, Chatzaki, M, Marusik, YM, 2020. New data on the spider fauna of Iran (Arachnida: Araneae), Part VII. Arachnology 18 (6): 569 - 591, DOI: https://doi.org/10.13156/arac.2020.18.6.569" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part VII." url="https://doi.org/10.13156/arac.2020.18.6.569" volume="18" year="2020">Zamani et al. 2020</bibRefCitation>
<bibRefCitation id="511EDA6FD63E757062DD8CC2F27E7BFA" DOI="https://doi.org/10.13156/arac.2020.18.6.569" author="Zamani, A" journalOrPublisher="Arachnology" pageId="0" pageNumber="145" pagination="569 - 591" refId="B27" refString="Zamani, A, Dimitrov, D, Weiss, I, Alimohammadi, S, Rafiei-Jahed, R, Esyunin, SL, Moradmand, M, Chatzaki, M, Marusik, YM, 2020. New data on the spider fauna of Iran (Arachnida: Araneae), Part VII. Arachnology 18 (6): 569 - 591, DOI: https://doi.org/10.13156/arac.2020.18.6.569" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part VII." url="https://doi.org/10.13156/arac.2020.18.6.569" volume="18" year="2020">Zamani et al. 2020</bibRefCitation>
). New records for Gilan, Hamedan, Mazandaran and North Khorasan, with the latter representing the easternmost record in the whole range (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
<figureCitation id="50EF9BFACB746FC85AF2A483B719DA3C" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">Azerbaijan, Iran.</paragraph>
<subSubSection id="935C5223B1CBD69A5EF27AE39EE78109" pageId="0" pageNumber="145" type="distribution">
<paragraph id="81E1B4C0245468E6D27BC7970A9A6AFE" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="ED2B9F6EFD90B88960889EA8E46F463B" pageId="0" pageNumber="145">Azerbaijan, Iran.</paragraph>
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</treatment>
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View file

@ -1,309 +1,309 @@
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11C-E</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">, 13A-E</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">, 14G, H</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">, 15D-F</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">, 17A-C</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
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<taxonomicNameLabel id="772616B96D34DE75FC7B7C1845427471" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="5C6F4D845EAD9FC1F2AC67839E5A2DD7" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11C-E</figureCitation>
<figureCitation id="EA4A8DAD9F3E09AE017C195A9C3099CB" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">, 13A-E</figureCitation>
<figureCitation id="85789A7CC8DC17BBFA09663C0AB9B58F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">, 14G, H</figureCitation>
<figureCitation id="852FD67D9347720F26BB7D97CBE292FC" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">, 15D-F</figureCitation>
<figureCitation id="9357D749088B644DE2BF9B0E7D3ABE1C" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">, 17A-C</figureCitation>
<figureCitation id="1FD718CA57FBFF05BF9C4FB9BC8B508F" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="773554C7B364C4C15299410429D1E1F3" pageId="0" pageNumber="145" type="type material">
<paragraph id="19F82786918DA1F6652D6353E832869D" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="679516DC484ED555CCC0A6AD2AF01855" pageId="0" pageNumber="145">
<emphasis id="8CCEC5022FB6DC429F61C40D4E6716E5" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Markazi Province</emphasis>
<emphasis id="D9814A07E46909EBA029D06DFFEA65BE" italics="true" pageId="0" pageNumber="145">Markazi Province</emphasis>
: Shazand,
<geoCoordinate degrees="33" direction="north" minutes="55" orientation="latitude" precision="925" value="33.916668">33°55'N</geoCoordinate>
<geoCoordinate id="98AB602EF3431F313B6D4227E8DF0226" degrees="33" direction="north" minutes="55" orientation="latitude" precision="925" value="33.916668">33°55'N</geoCoordinate>
,
<geoCoordinate degrees="49" direction="east" minutes="24" orientation="longitude" precision="925" value="49.4">49°24'E</geoCoordinate>
<geoCoordinate id="84075F0A5F7D51B2CA91F2F048F67A2D" degrees="49" direction="east" minutes="24" orientation="longitude" precision="925" value="49.4">49°24'E</geoCoordinate>
, 11.04.2015 (A. Zamani).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
<emphasis id="E12578E7F03BE1E02D5DBCCD274A1AF6" bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
: 5♂3♀ (MHNG), same data as holotype;
<emphasis italics="true" pageId="0" pageNumber="145">Isfahan Province</emphasis>
<emphasis id="7344859D1980DCE8167417A6B3780F90" italics="true" pageId="0" pageNumber="145">Isfahan Province</emphasis>
: 1♂1♀ (MHNG), Riz-e Landjan,
<geoCoordinate degrees="32" direction="north" minutes="24" orientation="latitude" precision="925" value="32.4">32°24'N</geoCoordinate>
<geoCoordinate id="1749420FF805491633D66D7A14286B06" degrees="32" direction="north" minutes="24" orientation="latitude" precision="925" value="32.4">32°24'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="19" orientation="longitude" precision="925" value="51.316666">51°19'E</geoCoordinate>
<geoCoordinate id="1B6CA51454FEA96BF6739EF29BD9F680" degrees="51" direction="east" minutes="19" orientation="longitude" precision="925" value="51.316666">51°19'E</geoCoordinate>
, 11.08.1973 (A. Senglet);
<emphasis italics="true" pageId="0" pageNumber="145">Fars Province</emphasis>
<emphasis id="C0099BEBE31E7435D31B281609496967" italics="true" pageId="0" pageNumber="145">Fars Province</emphasis>
: 1♂1♀ (MHNG), Izad Khast,
<geoCoordinate degrees="31" direction="north" minutes="31" orientation="latitude" precision="925" value="31.516666">31°31'N</geoCoordinate>
<geoCoordinate id="2A62568DE8A0CC6CE8F93EE65393F766" degrees="31" direction="north" minutes="31" orientation="latitude" precision="925" value="31.516666">31°31'N</geoCoordinate>
,
<geoCoordinate degrees="52" direction="east" minutes="08" orientation="longitude" precision="925" value="52.133335">52°08'E</geoCoordinate>
<geoCoordinate id="DF802E175788AFD890E23F67B2FAB343" degrees="52" direction="east" minutes="08" orientation="longitude" precision="925" value="52.133335">52°08'E</geoCoordinate>
, 12.06.1974 (A. Senglet).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet is a Persian feminine name meaning &quot;good listener&quot;. Noun in apposition.</paragraph>
<subSubSection id="DAF26C7E580B689E27983FAF60F025B3" pageId="0" pageNumber="145" type="etymology">
<paragraph id="81663E6C5D4D24E7B4454793EAE0FEF1" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="9B1E8A14008B8371EBFF5884B4709378" pageId="0" pageNumber="145">The specific epithet is a Persian feminine name meaning &quot;good listener&quot;. Noun in apposition.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="2DE591BB9AEE26064419358FB0CA8707" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="662D9FABD61B81FB5344988063F47ED2" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="C8F4A18C32CBC7BD8A93C6EA870B83FF" pageId="0" pageNumber="145">
The male of the new species is most similar to
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
<taxonomicName id="ECB9F8E89D27D4F556E73ADC70A001F8" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis id="B2B4C8AC4B0E1CA04F1892FDF52F0E29" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName>
sp. nov. from which it differs by a sharply tapering RTA, shorter tip of the cymbium (cf. Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13A</figureCitation>
<figureCitation id="BAFF7BF4D85BA3ABCDFEED882FC3CA59" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13A</figureCitation>
and
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13F</figureCitation>
<figureCitation id="652DDB644C820FCF988BCDD85BBB805A" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13F</figureCitation>
), the angle of embolus and the claw of the median apophysis. Female of
<taxonomicName lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
<taxonomicName id="403958DEB399F0754DA70374BB78DC3C" lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis id="1CDED917EE133F9408AC019077645CC3" italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
</taxonomicName>
sp. nov. is similar to
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
<taxonomicName id="F18FDA697E71365AEC9C44A0E4B59FE2" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis id="6ED35F1653FDC06712944DF2C8103DE4" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName>
sp. nov. by having a similar anterior hood but differs by the longer furrow leading to the hood (as long as receptacle
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="AFC64046F704C011A9126505999E4EE0" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
1.5 times shorter) and less separated receptacles.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 13.</emphasis>
<caption id="E69FD088AE0608560359FDDAA58AE99E" doi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145" start="Figure 13" startId="F13">
<paragraph id="128C1BC78583456F04FED0E2DBB2D4D8" pageId="0" pageNumber="145">
<emphasis id="ADF2639395D6E3CB5B85599A45F0E23F" bold="true" pageId="0" pageNumber="145">Figure 13.</emphasis>
Male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium niusha" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium niusha</emphasis>
<taxonomicName id="BFA2EBD102BE1115C255CF57654FDBAB" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium niusha" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="niusha">
<emphasis id="2BF089909DD6EE4721C097E1D128B226" italics="true" pageId="0" pageNumber="145">Acanthinozodium niusha</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-E</emphasis>
<emphasis id="0CD884D84EB32A044777E5CF51D55BFF" bold="true" pageId="0" pageNumber="145">A-E</emphasis>
) and
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
<taxonomicName id="007B7EA9F715CFAF7C06E1F3568ABE95" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis id="28442C0E98866FFB100641C5107502D1" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">F-I</emphasis>
<emphasis id="8CE29022413715E23CEB093254451A89" bold="true" pageId="0" pageNumber="145">F-I</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D, G, F</emphasis>
<emphasis id="8EB596FFAD6405C53C6DF18E8405DC06" bold="true" pageId="0" pageNumber="145">A, D, G, F</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, H</emphasis>
<emphasis id="8EAA69130BE44E85412710637CBD4548" bold="true" pageId="0" pageNumber="145">B, H</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, I</emphasis>
<emphasis id="1ED02DEBB44BE075F2B411DEEFD8708F" bold="true" pageId="0" pageNumber="145">C, I</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
<emphasis id="C1E5FE42CBB2966273EE9B740B40A1C5" bold="true" pageId="0" pageNumber="145">E</emphasis>
dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis>
<subSubSection id="397B7531233C101BD44629B14C270AD1" pageId="0" pageNumber="145" type="description">
<paragraph id="E5AE8A372C63317333DBAAD64B3C2C43" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="54F24A1347143A17E64B9962BCE388C9" pageId="0" pageNumber="145">
<emphasis id="B7A410A364E9F0A689A4E9C5CB710C05" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11C, D</figureCitation>
<figureCitation id="E8663A405C0669D7981576E3047C37A4" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11C, D</figureCitation>
. Total length 2.09. Carapace 0.94 long, 0.46 wide at pars cephalica, 0.69 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.10, ALE: 0.07, PME: 0.06, PLE: 0.06, PME-PME: 0.12. Carapace, sternum, labium, chelicerae and maxillae yellowish; carapace with irregular dark patterns. Chelicera with one retromarginal tooth. Legs yellowish, without annulations. Abdomen dorsally black, grayish ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 1.67+missing tarsus (0.74, 0.31, 0.62, missing), II: 2.56 (0.66, 0.30, 0.52, 0.59, 0.49), III: 2.43 (0.60, 0.33, 0.46, 0.59, 0.45), IV: 3.43 (0.90, 0.31, 0.81, 0.89, 0.52).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="1F71DF334B658AB6742BB92C4DC6E3E3" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13A-E</figureCitation>
<figureCitation id="346B1056219D832350BE7C61B95022DC" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13A-E</figureCitation>
,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14G, H</figureCitation>
<figureCitation id="876337F2AE6424309EE6F8509B97A039" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14G, H</figureCitation>
,
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">15D-F</figureCitation>
<figureCitation id="4700B662A66B2410DD93292CFA13A93E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">15D-F</figureCitation>
. RTA (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
<emphasis id="CF198E27A8431795BF9B84344373E339" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
) as long as wide and as long as tibia, basal part very wide, sharply tapering, tip bent ventrally; cymbium 1.5 times longer than wide; posterior part of tegulum with broad and indistinct projection postero-prolaterally; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="4824F75DA06E27969E602DE2462998CE" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) with massive base, much larger than the claw of median apophysis; sperm duct gradually tracking along margin of tegulum; tip of embolus (
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
<emphasis id="8548FEBFE1D14EA0186BA6CC854F70A9" italics="true" pageId="0" pageNumber="145">Em</emphasis>
) straight and lacking any processes or tubercles.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 14.</emphasis>
<caption id="558EED32F920BDB5DB3AEEECF23039FA" doi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145" start="Figure 14" startId="F14">
<paragraph id="5C01E8BAE72639CD8C847CE55EEF5293" pageId="0" pageNumber="145">
<emphasis id="A7AE04F1E4632EC479DE98095701B74A" bold="true" pageId="0" pageNumber="145">Figure 14.</emphasis>
SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis>
<taxonomicName id="4583400910C60D0A2492F48F346CDDF0" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa">
<emphasis id="D75CEDF4818AD26E6B612A038440EBD0" italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
<emphasis id="117C9245161D06B13F09B66C52418643" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
),
<taxonomicName lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
<taxonomicName id="6D24C2B2587DA2923C97AFBA23A6D114" lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis id="9A421C5C487E8CBCB82A039FB2AAF118" italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
<emphasis id="EFD011C237EA83CE5C528A55ACFA2A68" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
),
<taxonomicName lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
<taxonomicName id="9F04E21CEF91B66B9BDF220FB1397BAF" lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis id="332846E4458418AFFB16965BE0516184" italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">G, H</emphasis>
<emphasis id="25B4458B32A8BA4EB27A76E77646C14D" bold="true" pageId="0" pageNumber="145">G, H</emphasis>
) and
<taxonomicName lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
<taxonomicName id="BE1F841A3C2DE950436BF475E975A624" lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis id="F318A1749C37C601ADD6E888F12BD81F" italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">I</emphasis>
<emphasis id="A920C43DDE3B4695A595E92D8BC8DA4E" bold="true" pageId="0" pageNumber="145">I</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, C, D, F, I</emphasis>
<emphasis id="5F4FBE4437FE622EBBDDB59653CE775C" bold="true" pageId="0" pageNumber="145">A, C, D, F, I</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, E</emphasis>
<emphasis id="CAA5F1A1586CFD2B9E5112D4C5DB7BDB" bold="true" pageId="0" pageNumber="145">B, E</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">G, H</emphasis>
<emphasis id="815B0007C7C984161E49CF3D1DE22D90" bold="true" pageId="0" pageNumber="145">G, H</emphasis>
cymbial groove, dorsal. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Cg</emphasis>
<emphasis id="18E1F734337B1E185A438C8285F5FA14" italics="true" pageId="0" pageNumber="145">Cg</emphasis>
- cymbial groove,
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis>
<emphasis id="ECCB6A7DA89211BDBE643930AB1D1A17" italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
<emphasis id="56267A4F25D2B39B6DD5600523B0FD3B" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Ep</emphasis>
<emphasis id="3678A563FC1AEEC6B1DEF1DF80A54C99" italics="true" pageId="0" pageNumber="145">Ep</emphasis>
- embolic process,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="26E04396C32BB55B0CD83766E684345A" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
<emphasis id="710BD691A0176DB1784E67B235D98F02" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral tibial apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Tp</emphasis>
<emphasis id="3506F0BC39E9BED926E2F9951E5E7F51" italics="true" pageId="0" pageNumber="145">Tp</emphasis>
- tegular process. Scale bars: 0.1 mm, unless stated otherwise.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 15.</emphasis>
<caption id="4618E753572298948B753463BA1CF1C3" doi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145" start="Figure 15" startId="F15">
<paragraph id="C4144F250047F587DBDD253EF33EFFDD" pageId="0" pageNumber="145">
<emphasis id="34341633D1B6AAAA80342F36B9D23E15" bold="true" pageId="0" pageNumber="145">Figure 15.</emphasis>
SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parysatis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium parysatis</emphasis>
<taxonomicName id="0959B9F06C3A2BC85CCC0D2D77E6471A" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parysatis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parysatis">
<emphasis id="10B2D196E66484CBB3022CB5C101AFE7" italics="true" pageId="0" pageNumber="145">Acanthinozodium parysatis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
<emphasis id="4909BC6A29EAEC5FD94CF21A1CC65D4D" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and
<taxonomicName lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
<taxonomicName id="99DACC53332A46BB2E363A1E4CAC69D4" lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis id="B2F8D0DADE0B32E9B4BFD963B952E585" italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
<emphasis id="B6A1DC5D63066F789C74318DBA53E931" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis>
<emphasis id="124038BCF5AF3D97D497762FBB59C9FD" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, E</emphasis>
<emphasis id="9C592968A0EDF7B8558B3A971FDDCB71" bold="true" pageId="0" pageNumber="145">B, E</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, F</emphasis>
<emphasis id="99B3DDB5D91CF456E8AED81EAA55E243" bold="true" pageId="0" pageNumber="145">C, F</emphasis>
dorsoretrolateral and dorsal. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Cg</emphasis>
<emphasis id="137D124C2DBA8776136B925FA2898B6C" italics="true" pageId="0" pageNumber="145">Cg</emphasis>
- cymbial groove,
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis>
<emphasis id="6A0B8A321EF87847052A34A42F4D148B" italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
<emphasis id="32717E028130CE0039F61335F8822316" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="DAF766E00BE403C6D3737B2FBD9E8452" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
<emphasis id="57AA06C318FD299C83029BA72808EA24" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral tibial apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis>
<emphasis id="45785417A85D7692A129BB33E411DB29" italics="true" pageId="0" pageNumber="145">Va</emphasis>
- ventral apophysis. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure16" httpUri="https://binary.pensoft.net/fig/536893" pageId="0" pageNumber="145" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 16.</emphasis>
<caption id="603D79D238830DFA5470520D56DADF73" doi="10.3897/zookeys.1035.65767.figure16" httpUri="https://binary.pensoft.net/fig/536893" pageId="0" pageNumber="145" start="Figure 16" startId="F16">
<paragraph id="8799FAE0730B56C8B7991D24F659C6AF" pageId="0" pageNumber="145">
<emphasis id="6AB47E1977FB9D95768DCF5D16031A47" bold="true" pageId="0" pageNumber="145">Figure 16.</emphasis>
Epigyne of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis>
<taxonomicName id="1891F47FCAE64E23055D416AD9B44C60" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa">
<emphasis id="AA9504AC7697BBAA440E9227B3827003" italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
<emphasis id="46024445CE06CDB4D9F764B944B71B6A" bold="true" pageId="0" pageNumber="145">A</emphasis>
posteroventral
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
<emphasis id="649D986595EE325D3FABD5B3F064DBC1" bold="true" pageId="0" pageNumber="145">B</emphasis>
posterior
<emphasis bold="true" pageId="0" pageNumber="145">C, D</emphasis>
<emphasis id="D356F865C2BCE96F73859B43F5EE9F27" bold="true" pageId="0" pageNumber="145">C, D</emphasis>
ventral. Scale bar: 0.2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="FC8F8A8F32B4F71A2442BA3EB4D0F2DE" pageId="0" pageNumber="145">
<emphasis id="94A6D579B3FC1E4A6AF8725AF7ECCA73" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11E</figureCitation>
<figureCitation id="D483E1911A9CFABB850225E4D4E14EC2" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11E</figureCitation>
. Total length 2.10. Carapace 0.86 long, 0.44 wide at pars cephalica, 0.69 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.07, ALE: 0.06, PME: 0.05, PLE: 0.05, PME-PME: 0.10. Coloration as in male. Measurements of legs: I: 2.68 (0.73, 0.30, 0.59, 0.61, 0.45), II: 2.58 (0.74, 0.24, 0.53, 0.58, 0.49), III: 2.32 (0.59, 0.29, 0.50, 0.55, 0.39), IV: 3.45 (0.89, 0.33, 0.80, 0.92, 0.51).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="44A54F81748E904207DEBB16FC7BC5CF" pageId="0" pageNumber="145">
Epigyne as in Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">17A-C</figureCitation>
<figureCitation id="EDE5490914C279263DD3C5D7B79F1BA7" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">17A-C</figureCitation>
. Fovea about 3 times wider than long; hood as wide as fovea, bursae ovoid, weakly sclerotized, with fine pores (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">17C</figureCitation>
<figureCitation id="72697654BD085BF89B7B110B0F6DB69C" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">17C</figureCitation>
); receptacles round, spaced by over 5 diameters.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145" start="Figure 17" startId="F17">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 17.</emphasis>
<caption id="BE62E9BC05399FFFBFA88E4D7C3798F5" doi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145" start="Figure 17" startId="F17">
<paragraph id="A67FADEA663E574667952EAC0A776B88" pageId="0" pageNumber="145">
<emphasis id="0B1A27F53247A5891D5939DE9F36AA4B" bold="true" pageId="0" pageNumber="145">Figure 17.</emphasis>
Ventral view of epigynes of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium niusha" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium niusha</emphasis>
<taxonomicName id="5F3F74B56787F98133E20916F66DB755" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium niusha" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="niusha">
<emphasis id="8DF0428BC841ACC56197C0AE9582866C" italics="true" pageId="0" pageNumber="145">Acanthinozodium niusha</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
<emphasis id="415916AD278B51AE6F6A4F557F546716" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
<taxonomicName id="58BB2D116985E45AE09CEF281CA565FA" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis id="3FE21E04161166348C4D871DB6316266" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
<emphasis id="40E97B2A26EDD02AD9BB813B5280CCD5" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
). Scale bar: 0.2 mm, unless stated otherwise.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="2E9E80ABE7B9854C3FDE9F8C7A810DC3" pageId="0" pageNumber="145" type="distribution">
<paragraph id="5A360DA5166A8F402D74B955B8913936" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="08E6675554BCD6BD47F019E165C519E8" pageId="0" pageNumber="145">
Known only from the listed localities in Fars, Isfahan and Markazi provinces, central Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
<figureCitation id="123AE02DCE2B7BF5112A54FECF92247E" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,147 +1,147 @@
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<treatment ID-GBIF-Taxon="181753560" LSID="urn:lsid:zoobank.org:act:CDA6AFFA-3C76-4F74-8BC4-03B5DF32B011" httpUri="http://treatment.plazi.org/id/A34DB4A249F35C48A7A670818BA31422" lastPageNumber="145" pageId="0" pageNumber="145">
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<taxonomicName LSID="http://zoobank.org/CDA6AFFA-3C76-4F74-8BC4-03B5DF32B011" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium dorsa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dorsa" status="sp. nov.">Acanthinozodium dorsa</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">Figs 14I</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">, 18C, D</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">, 21A-C</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">, 24C, D</figureCitation>
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">, 30F-I</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
<treatment id="A34DB4A249F35C48A7A670818BA31422" ID-GBIF-Taxon="181753560" LSID="urn:lsid:zoobank.org:act:CDA6AFFA-3C76-4F74-8BC4-03B5DF32B011" httpUri="http://treatment.plazi.org/id/A34DB4A249F35C48A7A670818BA31422" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection id="649E242CF8E209403DEEB96CBA1242F0" pageId="0" pageNumber="145" type="nomenclature">
<paragraph id="D8BFE674C7223D20D76BF2D56CD83FD4" pageId="0" pageNumber="145">
<taxonomicName id="45A84BB2FE00BC4F7B0B48D43E4DC16E" ID-CoL="98YZ8" LSID="http://zoobank.org/CDA6AFFA-3C76-4F74-8BC4-03B5DF32B011" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium dorsa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dorsa" status="sp. nov.">Acanthinozodium dorsa</taxonomicName>
<taxonomicNameLabel id="6339900F21E600E6B13EEFB8A6B5E7C6" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="EE4517A49F6653B17A83B5E5E4D3A2CF" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">Figs 14I</figureCitation>
<figureCitation id="69CF368F83A661F8C20461645FA6BB7C" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">, 18C, D</figureCitation>
<figureCitation id="1F70D1BF7BF69588F0FBF8A1E47A7F8B" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">, 21A-C</figureCitation>
<figureCitation id="51D4BE3EA6250244595B9B22E871A91A" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">, 24C, D</figureCitation>
<figureCitation id="60906CC1085FF33520132EBBB8EB31C4" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">, 30F-I</figureCitation>
<figureCitation id="E7D8B54444D77D475B2C1E286D3B4FA0" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="B65CC97E4EFBCAB56797A8E7B1FE3D4B" pageId="0" pageNumber="145" type="type material">
<paragraph id="C0B4B900174070655BD1AC5769D5B054" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="EF259E45F9D744B645A1FE722A4F2BBC" pageId="0" pageNumber="145">
<emphasis id="BFAF80273BC4DD8192A76AC4CF49AE61" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ and
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">paratypes</emphasis>
<emphasis id="AD35EBEB3729962DD275F789D54FFD64" bold="true" italics="true" pageId="0" pageNumber="145">paratypes</emphasis>
26♂16♀ (MMUE), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Fars Province</emphasis>
<emphasis id="2B21E91E1DD32089A4A1CBD25CB9FFD1" italics="true" pageId="0" pageNumber="145">Fars Province</emphasis>
: 50 km NE of Shiraz, Bamoo reserve area,
<geoCoordinate degrees="29" direction="north" minutes="45" orientation="latitude" precision="925" value="29.75">29°45'N</geoCoordinate>
<geoCoordinate id="010498D29753719949516BDF7BA49DA4" degrees="29" direction="north" minutes="45" orientation="latitude" precision="925" value="29.75">29°45'N</geoCoordinate>
,
<geoCoordinate degrees="52" direction="east" minutes="45" orientation="longitude" precision="925" value="52.75">52°45'E</geoCoordinate>
<geoCoordinate id="B84D0A73461FBCC756CA0A79F8041F83" degrees="52" direction="east" minutes="45" orientation="longitude" precision="925" value="52.75">52°45'E</geoCoordinate>
, 18-28.05.2000 (Y.M. Marusik).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="93A12A63BBACC6BCE43F833207DA5648" pageId="0" pageNumber="145" type="etymology">
<paragraph id="198AD5B68DD5BBE6E7269AF40E8F49FE" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="8FA70457CFC8C21C75F3E13C6DD420C2" pageId="0" pageNumber="145">
The specific epithet is a Persian feminine name meaning
<normalizedToken originalValue="“precious”">&quot;precious&quot;</normalizedToken>
<normalizedToken id="B113536B9ECA4B9803F70DEF51EA808C" originalValue="“precious”">&quot;precious&quot;</normalizedToken>
. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="688294B0C30373D8B690ED812982BAA2" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="71F0F4CAB0B9B2EFF9A2AE5F0DE932AD" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="5E5F1EAE90B3855DCC8E03AAE208A459" pageId="0" pageNumber="145">
The male of the new species differs from all congeners in the region by the prolateral outgrowth of the median apophysis projecting ventrally (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="A3A29DF4EE93D5D792FFC509E7B0BAE6" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
not projecting) and the long and broad RTA, apically twisted and lacking outgrowths (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="1D81A94787466F357FBCF8F3B5E98FF2" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
with 1-2 outgrowths). The female is most similar to
<taxonomicName lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis>
<taxonomicName id="56805E07ACD1D8E3250C693B515A6A9D" lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum">
<emphasis id="AF7B3EB1C7AEA7E225E6E808DD502094" italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis>
</taxonomicName>
sp. nov. but differs by the epigynal hood being longer than wide (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="69BB360C13A436724585775D4C3021EE" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
wider than long) and less separated receptacles (ca. 2.5 times of their diameter
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="27313F3CF1D961F0E0C766E00563D3BA" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
4).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis>
<subSubSection id="AA5B9DE56EEB949AD39B3279198221BD" pageId="0" pageNumber="145" type="description">
<paragraph id="10679ACBD87F23C3F3C08E244250732A" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="A6008B5FB2B3C3C1517ED51AC4DF8343" pageId="0" pageNumber="145">
<emphasis id="58C35D8E24CC77CB7490855D20B958C5" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18C</figureCitation>
<figureCitation id="F3696593E83F2644B1431627985F396A" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18C</figureCitation>
. Total length 3.85. Carapace 1.87 long, 0.89 wide at pars cephalica, 1.39 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.15, ALE: 0.12, PME: 0.09, PLE: 0.09, PME-PME: 0.20. Carapace yellowish brown, darker at pars cephalica. Sternum, labium, maxillae and chelicerae light brown. Legs yellowish, without annulations. Abdomen dark brown dorsally, pale ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 8.20 (2.02, 0.68, 1.90, 2.19, 1.41), II: 6.89 (1.68, 0.68, 1.49, 1.99, 1.05), III: 7.23 (1.88, 0.64, 1.45, 2.13, 1.13), IV: 8.71 (2.18, 0.62, 2.11, 2.70, 1.10).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="01F0BB98BAE0A36DCCA0600C968749B4" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14I</figureCitation>
<figureCitation id="E82EBAF9E44382B567B915D936D2AB55" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14I</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">21A-C</figureCitation>
<figureCitation id="CA91C17795970543D37ADD8DA3D516DB" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">21A-C</figureCitation>
,
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">24C, D</figureCitation>
<figureCitation id="660120BDCA880A60FF8F70F98F9D1F80" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">24C, D</figureCitation>
. RTA long and broad, almost as long as the bulb, twisting ventrally toward the apex; ventral apophysis small and finger shaped; tegulum with posterior process; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="D92350EBD8EDFCD63040CDD5C72F8A48" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) with a distinct prolateral projection, winding ventrally to a blunt tip; embolus (
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
<emphasis id="AE75D71D0C4F567D0857E566C48A63BD" italics="true" pageId="0" pageNumber="145">Em</emphasis>
) originating at about the 8:30
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
<normalizedToken id="3C5841B01DABF95D9EF09DE4C53B39CE" originalValue="oclock">o'clock</normalizedToken>
position; embolus proper thin and slightly curving near the base and apex.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="2BF9FF218194797C3D716BFB2E39CA03" pageId="0" pageNumber="145">
<emphasis id="0A6D7BC6F65A4CFFFE5160DE668AB314" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18D</figureCitation>
<figureCitation id="3B4B3D7C66BE4161AC06A783A7D25146" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18D</figureCitation>
. Total length 4.46. Carapace 1.93 long, 0.96 wide at pars cephalica, 1.46 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.15, ALE: 0.12, PME: 0.09, PLE: 0.10, PME-PME: 0.15. Coloration as in male. Measurements of legs: I: 6.58 (1.39, 0.56, 1.52, 1.88, 1.23), II: 6.49 (1.62, 0.69, 1.34, 1.77, 1.07), III: 6.24 (1.71, 0.63, 1.25, 1.85, 0.80), IV: 8.45 (2.24, 0.72, 1.90, 2.60, 0.99).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="520A533923B51E0F0813930CAE781A8F" pageId="0" pageNumber="145">
Epigyne as in Fig.
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">30F-I</figureCitation>
<figureCitation id="7A5675D2831C41AA62DC956D9CA3E112" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">30F-I</figureCitation>
. Epigynal plate over 3 times wider than long; anterior hood ca. 2 times longer than wide; receptacles subrectangular, separated by about 2.5 times of their widths.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="A23DF2336A2DCB1FA7015BD496BBDDE4" pageId="0" pageNumber="145" type="distribution">
<paragraph id="A491033BBA0C78C4577C82069327092D" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="CCE45017157CE8AEB79AAEECFAEE9D00" pageId="0" pageNumber="145">
Known from the type locality in Fars Province, southern Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
<figureCitation id="03F8305F8E084BD35D2E5C7D042B2A88" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,301 +1,301 @@
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<mods:identifier id="F417DE23613138D674FEB12E7BBE5124" type="DOI">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:identifier>
<mods:identifier id="7A004C6141933504B03202BD5A7D8CBF" type="Pensoft-Pub">1313-2970-1035-145</mods:identifier>
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</mods:mods>
<treatment ID-GBIF-Taxon="181753546" LSID="urn:lsid:plazi:treatment:A45512952A3450C58AD2BFB995F9BB5C" httpUri="http://treatment.plazi.org/id/A45512952A3450C58AD2BFB995F9BB5C" lastPageNumber="145" pageId="0" pageNumber="145">
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145">
<treatment id="A45512952A3450C58AD2BFB995F9BB5C" ID-GBIF-Taxon="181753546" LSID="urn:lsid:plazi:treatment:A45512952A3450C58AD2BFB995F9BB5C" httpUri="http://treatment.plazi.org/id/A45512952A3450C58AD2BFB995F9BB5C" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection id="DD0438CE2C349EAE262022E407DE5A94" pageId="0" pageNumber="145" type="nomenclature">
<paragraph id="3BE4B5963B860E17C0F32D8894B9C24E" pageId="0" pageNumber="145">
Genus
<taxonomicName LSID="A4551295-2A34-50C5-8AD2-BFB995F9BB5C" authority="Andreeva &amp; Tyshchenko, 1968" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Zodariellum Andreeva &amp; Tyshchenko, 1968</taxonomicName>
<taxonomicName id="29789D14DF663A196519B6669CDE60F8" ID-CoL="63WNV" LSID="A4551295-2A34-50C5-8AD2-BFB995F9BB5C" authority="Andreeva &amp; Tyshchenko, 1968" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Zodariellum Andreeva &amp; Tyshchenko, 1968</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type species">
<paragraph pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Andreeva &amp; Tyschchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum surprisum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="surprisum">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum surprisum</emphasis>
<subSubSection id="0C72A0543BC8E555E790CE8240643C31" pageId="0" pageNumber="145" type="type species">
<paragraph id="4DF85C2DEA0BBDA5663C0D6C0C23E2EF" pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph id="4EB5F9F06F71E5B0AAA25C304C14D269" pageId="0" pageNumber="145">
<taxonomicName id="2D01E732987195C6C2B821F5B7900BF9" authorityName="Andreeva &amp; Tyschchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum surprisum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="surprisum">
<emphasis id="5AEF65A2BE75420532AA365266C3B8FD" italics="true" pageId="0" pageNumber="145">Zodariellum surprisum</emphasis>
</taxonomicName>
Andreeva &amp; Tyshchenko, 1968 from Tajikistan.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="05A5681F3630245476663252C7FB73E9" pageId="0" pageNumber="145" type="comments">
<paragraph id="358A3696D3CD96960E049733B7899CDA" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph id="F777C31DC6AE542BE531205C6217F301" pageId="0" pageNumber="145">
The genus was described as monotypic and synonymized with
<taxonomicName authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis>
<taxonomicName id="5A8640E24BDF2BA516375B8F7A59BD5F" authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="B9D022FC689466850B2657C47097ABAB" italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis>
</taxonomicName>
by
<bibRefCitation author="Jocque, R" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="145" pagination="1 - 160" refId="B7" refString="Jocque, R, 1991. A generic revision of the spider family Zodariidae (Araneae). Bulletin of the American Museum of Natural History 201: 1 - 160" title="A generic revision of the spider family Zodariidae (Araneae)." volume="201" year="1991">
<normalizedToken originalValue="Jocqué">Jocque</normalizedToken>
<bibRefCitation id="C404E62D33720B9122997C0FF6A5FFC2" author="Jocque, R" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="145" pagination="1 - 160" refId="B7" refString="Jocque, R, 1991. A generic revision of the spider family Zodariidae (Araneae). Bulletin of the American Museum of Natural History 201: 1 - 160" title="A generic revision of the spider family Zodariidae (Araneae)." volume="201" year="1991">
<normalizedToken id="1833466D146B1C2DD2BFB06EDF39D7B3" originalValue="Jocqué">Jocque</normalizedToken>
(1991)
</bibRefCitation>
.
<bibRefCitation author="Marusik, YM" journalOrPublisher="Reichenbachia" pageId="0" pageNumber="145" pagination="39 - 48" refId="B13" refString="Marusik, YM, Koponen, S, 2001. Spiders of the family Zodariidae from Mongolia (Arachnida: Araneae). Reichenbachia 34: 39 - 48" title="Spiders of the family Zodariidae from Mongolia (Arachnida: Araneae)." volume="34" year="2001">Marusik and Koponen (2001)</bibRefCitation>
<bibRefCitation id="99D35395B1776453C0C41A6F70D5DC94" author="Marusik, YM" journalOrPublisher="Reichenbachia" pageId="0" pageNumber="145" pagination="39 - 48" refId="B13" refString="Marusik, YM, Koponen, S, 2001. Spiders of the family Zodariidae from Mongolia (Arachnida: Araneae). Reichenbachia 34: 39 - 48" title="Spiders of the family Zodariidae from Mongolia (Arachnida: Araneae)." volume="34" year="2001">Marusik and Koponen (2001)</bibRefCitation>
resurrected
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
<taxonomicName id="8EB8AF6CDACC550D323A835948E2A150" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FA01546FAE862F59655C428D0EED274E" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName>
, described two new species from Mongolia (
<taxonomicName lsidName="Z. schmidti" pageId="0" pageNumber="145" rank="species" species="schmidti">
<emphasis italics="true" pageId="0" pageNumber="145">Z. schmidti</emphasis>
<taxonomicName id="B7252A9989CB15C6A3D7BAC7378BE11E" lsidName="Z. schmidti" pageId="0" pageNumber="145" rank="species" species="schmidti">
<emphasis id="D5C95A41833040D3690A3A71990F1547" italics="true" pageId="0" pageNumber="145">Z. schmidti</emphasis>
</taxonomicName>
Marusik &amp; Koponen, 2001 and
<taxonomicName lsidName="Z. mongolicum" pageId="0" pageNumber="145" rank="species" species="mongolicum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. mongolicum</emphasis>
<taxonomicName id="1DD9B133D7BF233C6E3F567F2B38B4D2" lsidName="Z. mongolicum" pageId="0" pageNumber="145" rank="species" species="mongolicum">
<emphasis id="E03BE334BD04C160CCF94D007BAD7E75" italics="true" pageId="0" pageNumber="145">Z. mongolicum</emphasis>
</taxonomicName>
Marusik &amp; Koponen, 2001), and transferred eight species to it from
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
<taxonomicName id="9E19BFC15FAAF20622A43394878F3D27" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="5A7BFE50F082DC7743D8E123429D47AE" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName>
:
<taxonomicName lsidName="Z. asiaticum" pageId="0" pageNumber="145" rank="species" species="asiaticum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. asiaticum</emphasis>
<taxonomicName id="BFA487994611047AD3CFBD40127E9C3E" lsidName="Z. asiaticum" pageId="0" pageNumber="145" rank="species" species="asiaticum">
<emphasis id="A4D6DCE7BF88E8DB3B87E2A11086843C" italics="true" pageId="0" pageNumber="145">Z. asiaticum</emphasis>
</taxonomicName>
(Tyshchenko, 1970),
<taxonomicName lsidName="Z. bekuzini" pageId="0" pageNumber="145" rank="species" species="bekuzini">
<emphasis italics="true" pageId="0" pageNumber="145">Z. bekuzini</emphasis>
<taxonomicName id="148FB0A36E75FB6FA77F4DB04AC9B111" lsidName="Z. bekuzini" pageId="0" pageNumber="145" rank="species" species="bekuzini">
<emphasis id="4A347225E723E531CBBBB0D70BCCEE2C" italics="true" pageId="0" pageNumber="145">Z. bekuzini</emphasis>
</taxonomicName>
(Nenilin, 1985),
<taxonomicName lsidName="Z. chaoyangense" pageId="0" pageNumber="145" rank="species" species="chaoyangense">
<emphasis italics="true" pageId="0" pageNumber="145">Z. chaoyangense</emphasis>
<taxonomicName id="1CD22B07215BAFC5589DC45022FA6B98" lsidName="Z. chaoyangense" pageId="0" pageNumber="145" rank="species" species="chaoyangense">
<emphasis id="18F202D96D4AA0BD2384B26535564DEA" italics="true" pageId="0" pageNumber="145">Z. chaoyangense</emphasis>
</taxonomicName>
(Zhu &amp; Zhu, 1983),
<taxonomicName lsidName="Z. continentalis" pageId="0" pageNumber="145" rank="species" species="continentalis">
<emphasis italics="true" pageId="0" pageNumber="145">Z. continentalis</emphasis>
<taxonomicName id="94B973D5A8E3A2148F8C691524BC21A6" lsidName="Z. continentalis" pageId="0" pageNumber="145" rank="species" species="continentalis">
<emphasis id="10E7B307F6A626B6806DCF2CF4A98506" italics="true" pageId="0" pageNumber="145">Z. continentalis</emphasis>
</taxonomicName>
(Andreeva &amp; Tyshchenko, 1968),
<taxonomicName lsidName="Z. furcum" pageId="0" pageNumber="145" rank="species" species="furcum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. furcum</emphasis>
<taxonomicName id="F03CC005FA2B53E177F89B9C0313F253" lsidName="Z. furcum" pageId="0" pageNumber="145" rank="species" species="furcum">
<emphasis id="FA648D639482A7B4872BA6BB211F02C4" italics="true" pageId="0" pageNumber="145">Z. furcum</emphasis>
</taxonomicName>
(Zhu, 1988),
<taxonomicName lsidName="Z. proszynskii" pageId="0" pageNumber="145" rank="species" species="proszynskii">
<emphasis italics="true" pageId="0" pageNumber="145">Z. proszynskii</emphasis>
<taxonomicName id="1EFD09E8DAD63F5F8A6F05BA4E8437E2" lsidName="Z. proszynskii" pageId="0" pageNumber="145" rank="species" species="proszynskii">
<emphasis id="CCEC5C872B1483361BF58AA042D5A3C0" italics="true" pageId="0" pageNumber="145">Z. proszynskii</emphasis>
</taxonomicName>
(Nenilin &amp; Fet, 1985),
<taxonomicName lsidName="Z. nenilini" pageId="0" pageNumber="145" rank="species" species="nenilini">
<emphasis italics="true" pageId="0" pageNumber="145">Z. nenilini</emphasis>
<taxonomicName id="D542BF9C9399D82F2AF97F138640929D" lsidName="Z. nenilini" pageId="0" pageNumber="145" rank="species" species="nenilini">
<emphasis id="FAEC0527C4CC3A236FE409B7908F0A90" italics="true" pageId="0" pageNumber="145">Z. nenilini</emphasis>
</taxonomicName>
(Eskov, 1996) and
<taxonomicName lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae">
<emphasis italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis>
<taxonomicName id="D9EC927FAFC615279CB821AA06500CBE" lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae">
<emphasis id="227BE13D5297CA49C3438C7E2AA4F166" italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis>
</taxonomicName>
(Nenilin &amp; Fet, 1985).
<bibRefCitation DOI="https://doi.org/10.23885/1814-3326-2007-3-2-87-95" author="Ponomarev, AV" journalOrPublisher="Caucasian Entomological Bulletin" pageId="0" pageNumber="145" pagination="87 - 95" refId="B22" refString="Ponomarev, AV, 2007. New taxa of spiders (Aranei) from the south of Russia and western Kazakhstan. Caucasian Entomological Bulletin 3: 87 - 95, DOI: https://doi.org/10.23885/1814-3326-2007-3-2-87-95" title="New taxa of spiders (Aranei) from the south of Russia and western Kazakhstan." url="https://doi.org/10.23885/1814-3326-2007-3-2-87-95" volume="3" year="2007">Ponomarev (2007)</bibRefCitation>
<bibRefCitation id="A30D9C43592FF2E55CF3826812ECF4B1" DOI="https://doi.org/10.23885/1814-3326-2007-3-2-87-95" author="Ponomarev, AV" journalOrPublisher="Caucasian Entomological Bulletin" pageId="0" pageNumber="145" pagination="87 - 95" refId="B22" refString="Ponomarev, AV, 2007. New taxa of spiders (Aranei) from the south of Russia and western Kazakhstan. Caucasian Entomological Bulletin 3: 87 - 95, DOI: https://doi.org/10.23885/1814-3326-2007-3-2-87-95" title="New taxa of spiders (Aranei) from the south of Russia and western Kazakhstan." url="https://doi.org/10.23885/1814-3326-2007-3-2-87-95" volume="3" year="2007">Ponomarev (2007)</bibRefCitation>
described two more species in the genus (
<taxonomicName lsidName="Z. volgouralensis" pageId="0" pageNumber="145" rank="species" species="volgouralensis">
<emphasis italics="true" pageId="0" pageNumber="145">Z. volgouralensis</emphasis>
<taxonomicName id="23FF01975A96A880C67F06722E936A7C" lsidName="Z. volgouralensis" pageId="0" pageNumber="145" rank="species" species="volgouralensis">
<emphasis id="D973E79C49D0030DCA790F7252BCD74A" italics="true" pageId="0" pageNumber="145">Z. volgouralensis</emphasis>
</taxonomicName>
Ponomarev, 2007 and Z.
<taxonomicName baseAuthorityName="Ponomarev" baseAuthorityYear="2007" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion inderensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="inderensis">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion inderensis</emphasis>
<taxonomicName id="4856273C586BDD293AF9D5AC46761545" baseAuthorityName="Ponomarev" baseAuthorityYear="2007" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion inderensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="inderensis">
<emphasis id="3DE2692364DBE061645968DE3FB61FF6" italics="true" pageId="0" pageNumber="145">Zodarion inderensis</emphasis>
</taxonomicName>
Ponomarev, 2007).
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2015.114" author="Jocque, R" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="145" pagination="1 - 23" refId="B10" refString="Jocque, R, Henrard, A, 2015. Revalidation of Acanthinozodium Denis, 1966 with description of three new species and discovery of a remarkable male palpal character (Araneae, Zodariidae). European Journal of Taxonomy 114: 1 - 23, DOI: https://doi.org/10.5852/ejt.2015.114" title="Revalidation of Acanthinozodium Denis, 1966 with description of three new species and discovery of a remarkable male palpal character (Araneae, Zodariidae)." url="https://doi.org/10.5852/ejt.2015.114" volume="114" year="2015">
<normalizedToken originalValue="Jocqué">Jocque</normalizedToken>
<bibRefCitation id="8BCC38D8D3454217D253F47976C4800C" DOI="https://doi.org/10.5852/ejt.2015.114" author="Jocque, R" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="145" pagination="1 - 23" refId="B10" refString="Jocque, R, Henrard, A, 2015. Revalidation of Acanthinozodium Denis, 1966 with description of three new species and discovery of a remarkable male palpal character (Araneae, Zodariidae). European Journal of Taxonomy 114: 1 - 23, DOI: https://doi.org/10.5852/ejt.2015.114" title="Revalidation of Acanthinozodium Denis, 1966 with description of three new species and discovery of a remarkable male palpal character (Araneae, Zodariidae)." url="https://doi.org/10.5852/ejt.2015.114" volume="114" year="2015">
<normalizedToken id="9C414B0D3F332C7580362F270E681B49" originalValue="Jocqué">Jocque</normalizedToken>
and Henrard (2015)
</bibRefCitation>
transferred all
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
<taxonomicName id="38B28B0E0AFF716DE673FA8739715CCD" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="1ACF2187A2B3021CACAC38631985C685" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName>
species, except for the generotype into
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
<taxonomicName id="03F9C89CBCC315FA2189EBB6FED1EC2E" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="C44B43832A2DC1607BE5F773D8EA8448" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="E98445EF6AD65C9E71E2FFB7FA1A4CFE" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="4BF635674201B268E4A3BF353DD7CDBB" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="A967295BE666987B1708EA09156747A8" pageId="0" pageNumber="145">
All
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
<taxonomicName id="D6123F41D1288F3130BE72D32913EDD1" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="0289EA87C5CCA253CE652EA9C746F87B" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName>
spp. are clearly different from the generotype of
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
<taxonomicName id="598D48D856E9404FD3EAE7C8B36271BB" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="9EF0171210A9406CCD0225D77BB6C1F7" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
<taxonomicName id="3CDF7E30FFA6DAB3058A158383FEF8D7" lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis id="D9875EE444D968D70D050C3745FB0E90" italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
</taxonomicName>
(Audouin, 1826), and the species considered in the genus by the following: 1) the cymbium has a tutaculum (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="FEE1125D6C12685507B4D986384E33B7" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
lacking in
<taxonomicName lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
<taxonomicName id="52977E492B56B7E6AAF01F2B388431A1" lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis id="1B35F7F71949D24FB9C8902C65B504E0" italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
</taxonomicName>
and other species groups); 2) filamentous embolus starting at the 5
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
<normalizedToken id="7B33C5079961EA1269553798B464DFC3" originalValue="oclock">o'clock</normalizedToken>
position (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="B1E11F2E6121E15E43D37B7DEBF6F8AC" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
6 - in
<taxonomicName lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
<taxonomicName id="BEE32C8DEE03891A3E2FEDE115B0F33B" lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis id="D34ABEF5D31277D0DFC84F1127793C3D" italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
</taxonomicName>
, and many species groups of
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
<taxonomicName id="F31CCBB719F3ED504878548C2D80D06E" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="36F987A5A7237AE258F904B206ECE474" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName>
sensu lato lack the filamentous embolus, or they are not at 5
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
<normalizedToken id="7658EB7BEC4EECDB879CBCC45930C7C5" originalValue="oclock">o'clock</normalizedToken>
position); 3) the absence of a ventral tibial apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="0BEF2BCBBF7CFEDF546E4D73F62ACDAE" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
present in the generotype) and 4) the shape of the RTA: one arm longer than wide, with claw or wart-like outgrowth posteriorly from the tip (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="8F5471931B60BBF6272C773E2B3F2A9E" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
wider than long, with 3 branches in the generotype and various shapes in other species groups). Females of the two generotypes have long and twisted copulatory ducts, but they are converging in
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
<taxonomicName id="2D8EF4CC205E097B4CD5017C12737EA3" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="A80792BC5A7F196F69BFDDCE4689D8BE" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName>
and diverging in
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
<taxonomicName id="0848021D6A6789567121C786B9D18CDA" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FD39C2AB515362704B070FC2A597BEE6" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="composition">
<paragraph pageId="0" pageNumber="145">Composition.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="6CA0AA08FA938F9D9C0D17F4AADDFA8E" pageId="0" pageNumber="145" type="composition">
<paragraph id="72356A76374DF583D445033E0986E6B1" pageId="0" pageNumber="145">Composition.</paragraph>
<paragraph id="8157756323FB47EB1AE1AEB007C01CE9" pageId="0" pageNumber="145">
We consider the following species in
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
<taxonomicName id="4738A1E7EAC26100E2A30E50B4968B27" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="0DC49F6D5673D39C30268DC5B3BE445A" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName>
because they have similar male palps and epigynes:
<taxonomicName lsidName="Z. asiaticum" pageId="0" pageNumber="145" rank="species" species="asiaticum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. asiaticum</emphasis>
<taxonomicName id="810A736A78654DC19637A42337643037" lsidName="Z. asiaticum" pageId="0" pageNumber="145" rank="species" species="asiaticum">
<emphasis id="5F398D4CA461719A6DBD2AB9A604512B" italics="true" pageId="0" pageNumber="145">Z. asiaticum</emphasis>
</taxonomicName>
(Tyshchenko, 1970) comb. res.,
<taxonomicName lsidName="Z. bactrianum" pageId="0" pageNumber="145" rank="species" species="bactrianum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. bactrianum</emphasis>
<taxonomicName id="D50312D270F2EA31F2103B754F8608C9" lsidName="Z. bactrianum" pageId="0" pageNumber="145" rank="species" species="bactrianum">
<emphasis id="CD389B00DBDD1FD8CD4D79D00A826B9B" italics="true" pageId="0" pageNumber="145">Z. bactrianum</emphasis>
</taxonomicName>
(Kroneberg, 1875) comb. nov. (ex.
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
<taxonomicName id="14630A44662DBC032B551A465E807409" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="BB9E8C430B203DA7415ABE38733B8293" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName>
),
<taxonomicName lsidName="Z. bekuzini" pageId="0" pageNumber="145" rank="species" species="bekuzini">
<emphasis italics="true" pageId="0" pageNumber="145">Z. bekuzini</emphasis>
<taxonomicName id="2A71D8A8AD8FC51672374C9BEB38D46C" lsidName="Z. bekuzini" pageId="0" pageNumber="145" rank="species" species="bekuzini">
<emphasis id="EA1D7D775A47530EEB382CD0D6BD7CB0" italics="true" pageId="0" pageNumber="145">Z. bekuzini</emphasis>
</taxonomicName>
(Nenilin, 1985) comb. res.,
<taxonomicName lsidName="Z. chaoyangense" pageId="0" pageNumber="145" rank="species" species="chaoyangense">
<emphasis italics="true" pageId="0" pageNumber="145">Z. chaoyangense</emphasis>
<taxonomicName id="5929EFC0508BE5C2DA1E8B590C739422" lsidName="Z. chaoyangense" pageId="0" pageNumber="145" rank="species" species="chaoyangense">
<emphasis id="0E92F91D083B7C0F424EA31397A0842F" italics="true" pageId="0" pageNumber="145">Z. chaoyangense</emphasis>
</taxonomicName>
(Zhu &amp; Zhu, 1983) comb. res.,
<taxonomicName lsidName="Z. continentalis" pageId="0" pageNumber="145" rank="species" species="continentalis">
<emphasis italics="true" pageId="0" pageNumber="145">Z. continentalis</emphasis>
<taxonomicName id="5B70A84ED7C1945BAB89E2DDFEC3AEF2" lsidName="Z. continentalis" pageId="0" pageNumber="145" rank="species" species="continentalis">
<emphasis id="4A2D65E27825BD0EBD8FBF0CEDADF87C" italics="true" pageId="0" pageNumber="145">Z. continentalis</emphasis>
</taxonomicName>
(Andreeva &amp; Tyshchenko, 1968) comb. res.,
<taxonomicName lsidName="Z. furcum" pageId="0" pageNumber="145" rank="species" species="furcum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. furcum</emphasis>
<taxonomicName id="92D4307BAE6F0324EC8A3EEF33C40B00" lsidName="Z. furcum" pageId="0" pageNumber="145" rank="species" species="furcum">
<emphasis id="1083D531DABACB112123374D200C7E33" italics="true" pageId="0" pageNumber="145">Z. furcum</emphasis>
</taxonomicName>
(Zhu, 1988) comb. res.,
<taxonomicName lsidName="Z. mongolicum" pageId="0" pageNumber="145" rank="species" species="mongolicum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. mongolicum</emphasis>
<taxonomicName id="8D88FF0365AF83744728A1025FC3BEF8" lsidName="Z. mongolicum" pageId="0" pageNumber="145" rank="species" species="mongolicum">
<emphasis id="65A78DA8F9C198F82D1542EB39A1FE39" italics="true" pageId="0" pageNumber="145">Z. mongolicum</emphasis>
</taxonomicName>
Marusik &amp; Koponen, 2001 comb. res.,
<taxonomicName lsidName="Z. proszynskii" pageId="0" pageNumber="145" rank="species" species="proszynskii">
<emphasis italics="true" pageId="0" pageNumber="145">Z. proszynskii</emphasis>
<taxonomicName id="CA1B456B3FC1503B0EB7438E2FA4C2B0" lsidName="Z. proszynskii" pageId="0" pageNumber="145" rank="species" species="proszynskii">
<emphasis id="0CA171070C1C9BC2F96E2C8ED3B7AC1C" italics="true" pageId="0" pageNumber="145">Z. proszynskii</emphasis>
</taxonomicName>
(Nenilin &amp; Fet, 1985) comb. res.,
<taxonomicName lsidName="Z. nenilini" pageId="0" pageNumber="145" rank="species" species="nenilini">
<emphasis italics="true" pageId="0" pageNumber="145">Z. nenilini</emphasis>
<taxonomicName id="E03163312858C183FA32CE583BD098C9" lsidName="Z. nenilini" pageId="0" pageNumber="145" rank="species" species="nenilini">
<emphasis id="7AEA432356D257D06752D6817DA9E574" italics="true" pageId="0" pageNumber="145">Z. nenilini</emphasis>
</taxonomicName>
(Eskov, 1996) comb. res.,
<taxonomicName lsidName="Z. surprisum" pageId="0" pageNumber="145" rank="species" species="surprisum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. surprisum</emphasis>
<taxonomicName id="8E40DA0199DD68EE13981BCCB81F7150" lsidName="Z. surprisum" pageId="0" pageNumber="145" rank="species" species="surprisum">
<emphasis id="10E251D91B1CB7252717AFCA41916A48" italics="true" pageId="0" pageNumber="145">Z. surprisum</emphasis>
</taxonomicName>
Andreeva &amp; Tyshchenko, 1968 comb. res.,
<taxonomicName lsidName="Z. schmidti" pageId="0" pageNumber="145" rank="species" species="schmidti">
<emphasis italics="true" pageId="0" pageNumber="145">Z. schmidti</emphasis>
<taxonomicName id="2C4842E1EFC397405C3F0F4A60C04565" lsidName="Z. schmidti" pageId="0" pageNumber="145" rank="species" species="schmidti">
<emphasis id="174FFDFE516DA8A9D3619A36C5EA6E6D" italics="true" pageId="0" pageNumber="145">Z. schmidti</emphasis>
</taxonomicName>
Marusik &amp; Koponen, 2001 comb. res.,
<taxonomicName lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae">
<emphasis italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis>
<taxonomicName id="038DC46BA5C7CAD9A638FC7D24226BD7" lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae">
<emphasis id="01D5A555B179886BA13A416F8C4003E9" italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis>
</taxonomicName>
(Nenilin &amp; Fet, 1985) comb. res. and
<taxonomicName lsidName="Z. volgouralensis" pageId="0" pageNumber="145" rank="species" species="volgouralensis">
<emphasis italics="true" pageId="0" pageNumber="145">Z. volgouralensis</emphasis>
<taxonomicName id="0B5189F80440873C22A01FA609E226CA" lsidName="Z. volgouralensis" pageId="0" pageNumber="145" rank="species" species="volgouralensis">
<emphasis id="76706D7C8746D0E4252E630CCCD95E56" italics="true" pageId="0" pageNumber="145">Z. volgouralensis</emphasis>
</taxonomicName>
Ponomarev, 2007 comb. res. Taking into account the shape of the epigyne of
<taxonomicName baseAuthorityName="Ponomarev" baseAuthorityYear="2007" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion inderensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="inderensis">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion inderensis</emphasis>
<taxonomicName id="8A8EBBD4B968158E24FDA6F6D6B2AA57" baseAuthorityName="Ponomarev" baseAuthorityYear="2007" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion inderensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="inderensis">
<emphasis id="99EC6FE0D52F9535AD6387A36F5E6F41" italics="true" pageId="0" pageNumber="145">Zodarion inderensis</emphasis>
</taxonomicName>
(Ponomarev, 2007), originally placed in
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
<taxonomicName id="1EB27874DFA4AE1B9B9DCCDAED421C2F" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="2162C6B142F1446FB0B6ACDDA0835B92" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName>
, we do not restore the original combination for this species.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="7AB0087D4DB1506A4E51FBBE376BA172" pageId="0" pageNumber="145">
To illustrate the conformation of the male palp in
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
<taxonomicName id="A7554EA2F0D28AD5BCE6834EEC2F0C3B" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="A8B2FAD159B835D0D2D1608484B549E6" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName>
(male specimens are lacking among the material studied in Iran and Turkmenistan), we have provided figures of
<taxonomicName lsidName="Z. bactrianum" pageId="0" pageNumber="145" rank="species" species="bactrianum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. bactrianum</emphasis>
<taxonomicName id="CDF952C88D227E09FD47CC9A87C309F8" lsidName="Z. bactrianum" pageId="0" pageNumber="145" rank="species" species="bactrianum">
<emphasis id="914B272215198AC030B1F9FE7EC9DF66" italics="true" pageId="0" pageNumber="145">Z. bactrianum</emphasis>
</taxonomicName>
, a species previously known from the original description only and previously considered in
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
<taxonomicName id="921082B4914F8D7A0DB13467C98514FF" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="D993536F652A83AD22B3AC11A3D122C5" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="2C0B11E5D6F75E74389E6D78ED321014" pageId="0" pageNumber="145" type="distribution">
<paragraph id="55AFB533211D364DE843AEF53515D1DB" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="7E043D3E7DEA177C98C2628E278B6712" pageId="0" pageNumber="145">
Western Russia, Iran, Central Asia to northern China (
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<bibRefCitation id="FB17B5AFE1A217077EF3291DB010CB75" author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC 2021</bibRefCitation>
).
</paragraph>
</subSubSection>

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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:title id="1404225D79280986BA49840E2CFB995A">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:namePart>Zamani, Alireza</mods:namePart>
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<mods:namePart>Marusik, Yuri M.</mods:namePart>
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<treatment ID-GBIF-Taxon="181753559" LSID="urn:lsid:zoobank.org:act:CD502D82-8DBC-4E66-9626-D9CD571D1DAC" httpUri="http://treatment.plazi.org/id/A5D289752A415787BF60F44464C2C3A0" lastPageNumber="145" pageId="0" pageNumber="145">
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<paragraph pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/CD502D82-8DBC-4E66-9626-D9CD571D1DAC" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium sorani" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="sorani" status="sp. nov.">Acanthinozodium sorani</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11H</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">, 12D-F</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">, 14D-F</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
<treatment id="A5D289752A415787BF60F44464C2C3A0" ID-GBIF-Taxon="181753559" LSID="urn:lsid:zoobank.org:act:CD502D82-8DBC-4E66-9626-D9CD571D1DAC" httpUri="http://treatment.plazi.org/id/A5D289752A415787BF60F44464C2C3A0" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection id="2D22F1CE9081C3A9194934530461B2DC" pageId="0" pageNumber="145" type="nomenclature">
<paragraph id="BDE896B7FC898034D6E4BBCAB040CAD7" pageId="0" pageNumber="145">
<taxonomicName id="A9F49AE7D137CDDDDA0AF17AAB6BE718" ID-CoL="98YZ9" LSID="http://zoobank.org/CD502D82-8DBC-4E66-9626-D9CD571D1DAC" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium sorani" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="sorani" status="sp. nov.">Acanthinozodium sorani</taxonomicName>
<taxonomicNameLabel id="12DFAB6D8E980852F4F20F5632A5C00F" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="5FC245150B6E41A15113CC7D24FD99D6" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11H</figureCitation>
<figureCitation id="4629C352535227328416B351A30C8D81" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">, 12D-F</figureCitation>
<figureCitation id="298DF6A56B5F4D140BC9192571F4DD77" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">, 14D-F</figureCitation>
<figureCitation id="813DFC9CAA79BDE96F8B90CF66E49860" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="C4D34920BA2A34C41743CA0180AB13B6" pageId="0" pageNumber="145" type="type material">
<paragraph id="6E997BC10EF880A4431D37563C1F4C77" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="E1E0508A17D91B41979F0FAE919ED1E6" pageId="0" pageNumber="145">
<emphasis id="0877B8020B6DB32CCB067FAFE4F3B880" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Kurdistan Province</emphasis>
<emphasis id="435175963B8A39EA52B6D8D0FED16DA0" italics="true" pageId="0" pageNumber="145">Kurdistan Province</emphasis>
: Santeh,
<geoCoordinate degrees="36" direction="north" minutes="11" orientation="latitude" precision="925" value="36.183334">36°11'N</geoCoordinate>
<geoCoordinate id="829F44C5FC03ADE7EA1A15141B7CC970" degrees="36" direction="north" minutes="11" orientation="latitude" precision="925" value="36.183334">36°11'N</geoCoordinate>
,
<geoCoordinate degrees="46" direction="east" minutes="32" orientation="longitude" precision="925" value="46.533333">46°32'E</geoCoordinate>
<geoCoordinate id="9E0DB2F54B14A30B94769D39A37B52E4" degrees="46" direction="east" minutes="32" orientation="longitude" precision="925" value="46.533333">46°32'E</geoCoordinate>
, 23.06.1975 (A. Senglet).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
<emphasis id="9C74D44EA1AA3D1874F9ED91BCC96F3D" bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
: 6♂ (MHNG), Marivan, 5.2017 (A. Zamani);
<emphasis italics="true" pageId="0" pageNumber="145">East Azerbaijan Province</emphasis>
<emphasis id="B9AD9BC4C5902F0D7A6C5C224A691448" italics="true" pageId="0" pageNumber="145">East Azerbaijan Province</emphasis>
: 2♂ (MHNG), north of Bonati,
<geoCoordinate degrees="37" direction="north" minutes="26" orientation="latitude" precision="925" value="37.433334">37°26'N</geoCoordinate>
<geoCoordinate id="DDDBE6C89D829F94353C4A871F5B1665" degrees="37" direction="north" minutes="26" orientation="latitude" precision="925" value="37.433334">37°26'N</geoCoordinate>
,
<geoCoordinate degrees="45" direction="east" minutes="57" orientation="longitude" precision="925" value="45.95">45°57'E</geoCoordinate>
<geoCoordinate id="AEDFA5D4B24F6085CF237E36B9F6CBD5" degrees="45" direction="east" minutes="57" orientation="longitude" precision="925" value="45.95">45°57'E</geoCoordinate>
, 04.06.1975 (A. Senglet).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet refers to a dialect or a language of the Kurdish languages that is spoken in Iraq, mainly in Iraqi Kurdistan, as well as the Kurdistan Province, Kermanshah Province, and West Azerbaijan Province of western Iran. Noun in apposition.</paragraph>
<subSubSection id="87052E0E516946B6834C7842DC0AFADB" pageId="0" pageNumber="145" type="etymology">
<paragraph id="376C20998795A962A0915D800D538E18" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="7F94C9F421F6BF28F98D5146B1946F44" pageId="0" pageNumber="145">The specific epithet refers to a dialect or a language of the Kurdish languages that is spoken in Iraq, mainly in Iraqi Kurdistan, as well as the Kurdistan Province, Kermanshah Province, and West Azerbaijan Province of western Iran. Noun in apposition.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="0AA59162C407DC240A27AF45BA08D3B7" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="0E6405A5FEEF6D60085774D290B3E776" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="0000906DCF35EAD893C49C777F9BC868" pageId="0" pageNumber="145">
The new species is most similar to
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
<taxonomicName id="4E50676DFBA5849D99512D563466AE0A" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis id="346EAAFD709B7D2315C227657B8873D3" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName>
sp. nov. but differs by embolic anterior process which is spine-like and thinner than the embolus proper in
<taxonomicName lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
<taxonomicName id="A4622FBC907189BB7CF6FCA7ABE211AA" lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis id="A720B855ABDA697AFEAB71670233B17E" italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
</taxonomicName>
sp. nov. (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="9908DA81651034FA8676DB826D64BEFF" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
broad) and also by the relative length of the RTA, longer than bulb in
<taxonomicName lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
<taxonomicName id="0D75CD8D4E67752713DE09C2C403761F" lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis id="D95136398D76A70F0F251F92B8D7923D" italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
</taxonomicName>
sp. nov. (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="79CDC3D47DBB239665CB989DF53D2CD9" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
shorter).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis>
<subSubSection id="58D95CF25C0AF8A9FB760D259632D431" pageId="0" pageNumber="145" type="description">
<paragraph id="F2EF2EF187DCCB8A4FE483297B729D6B" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="F1155469D0366DE40EE0D7DF83E78497" pageId="0" pageNumber="145">
<emphasis id="04F59D364A06BCBF2E82E251AD11A127" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11H</figureCitation>
<figureCitation id="0AC8439490E0F197BD8FB220F40E1CF1" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11H</figureCitation>
. Total length 2.11. Carapace 1.00 long, 0.46 wide at pars cephalica, 0.72 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.09, ALE: 0.07, PME: 0.06, PLE: 0.06, PME-PME: 0.12. Carapace, sternum, labium, chelicerae and maxillae yellowish brown; carapace with irregular dark patterns. Chelicera with one retromarginal tooth. Legs yellowish, dark brown at femora I and II, without annulations. Abdomen dorsally black, grayish ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 3.87 (0.97, 0.37, 0.89, 0.91, 0.64), II: 3.47 (0.92, 0.32, 0.72, 0.88, 0.63), Fe III: 0.88, other segments missing, IV: 4.51 (1.06, 0.36, 1.12, 1.33, 0.64).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="86149AA310DB114BB87081B7BEFE4405" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">12D-F</figureCitation>
<figureCitation id="B7101A84EED58561041991442EB1BFF8" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">12D-F</figureCitation>
,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14D-F</figureCitation>
<figureCitation id="C394C273C3AE3A6C7F75B8090BC20384" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14D-F</figureCitation>
. RTA (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
<emphasis id="13DDA5A0A6477490D53DD9D27D92E59D" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
) long (longer than bulb) and thin (about 6 times longer than wide), slightly bent; tegulum with posterior process (
<emphasis italics="true" pageId="0" pageNumber="145">Tp</emphasis>
<emphasis id="10959BAFEF44E962594396AE5CC86D62" italics="true" pageId="0" pageNumber="145">Tp</emphasis>
), sperm duct almost straight retrolaterally, and gently bent along prolateral side; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="E4DD7A13B8E094B2C59B7B2EBE42DA87" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) wider than long; embolus (
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
<emphasis id="C45BEFBA5C602BF31D34AFC546980825" italics="true" pageId="0" pageNumber="145">Em</emphasis>
) broad basally, bifurcated terminally, with spine-like anterior process; embolus proper thin and straight, with small tubercle.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="7A7318BA15F00A4B52BAA61AF1B361FB" pageId="0" pageNumber="145">
<emphasis id="B53C23E786ECED8F93E28B5E3CFEC69D" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="93D1F09C05E5E2A8426D56F30E3DDB22" pageId="0" pageNumber="145" type="distribution">
<paragraph id="7D195F0C8AEB4E1FB58F24FD44238736" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="291829E58DE713E0D77F2CAE99108AB8" pageId="0" pageNumber="145">
Known only from the listed localities in Kurdistan and East Azerbaijan provinces, western and northwestern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
<figureCitation id="146E84245C0E7F6B94600A5A63A7603D" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,77 +1,77 @@
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<subSubSection pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145">
<treatment id="AA4D61BB05A75F3299A814AB2E95E88E" ID-GBIF-Taxon="181753540" LSID="urn:lsid:plazi:treatment:AA4D61BB05A75F3299A814AB2E95E88E" httpUri="http://treatment.plazi.org/id/AA4D61BB05A75F3299A814AB2E95E88E" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
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<paragraph id="824A291020B6C94E22F0A75693255A62" pageId="0" pageNumber="145">
Genus
<taxonomicName LSID="AA4D61BB-05A7-5F32-99A8-14AB2E95E88E" authority="Denis, 1966" authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Acanthinozodium Denis, 1966</taxonomicName>
<taxonomicName id="DDF89E628F972CE37D34237CF2661BF1" ID-CoL="M2C" ID-ENA="2033106" LSID="AA4D61BB-05A7-5F32-99A8-14AB2E95E88E" authority="Denis, 1966" authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Acanthinozodium Denis, 1966</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="145" type="type species">
<paragraph pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium spinulosum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="spinulosum">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium spinulosum</emphasis>
<subSubSection id="1860FCBD08DDD6A8CE4D4EA461B03E37" pageId="0" pageNumber="145" type="type species">
<paragraph id="487965BA535E9E84945B9F751828F91C" pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph id="01786BC77BEE50A7F145592E581FC321" pageId="0" pageNumber="145">
<taxonomicName id="FFDB8D9549E139BEC8BC36E0EF7A7507" authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium spinulosum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="spinulosum">
<emphasis id="2F41EC0EBBC2130285E1B889DEDE0CC7" italics="true" pageId="0" pageNumber="145">Acanthinozodium spinulosum</emphasis>
</taxonomicName>
Denis, 1966 from Libya.
</paragraph>
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<subSubSection pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="B3B38E83CD43D52415BAAFA2197E26F2" pageId="0" pageNumber="145" type="comments">
<paragraph id="62A8EA1E95DF6B6464D05AC2A1FB0136" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph id="53E4AC9082E9C12ED69FCDD209E67F3E" pageId="0" pageNumber="145">
Twelve species are currently considered in this genus which are known from the Maghreb and adjacent countries in the south and in Socotra; previously, it was not recorded from Iran (
<bibRefCitation author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC 2021</bibRefCitation>
<bibRefCitation id="AC198906881A306CA1279A05972B9438" author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC 2021</bibRefCitation>
). Although the male of the type species is unknown,
<taxonomicName authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis>
<taxonomicName id="E39293DC763E6357AB317579BBA8DCE6" authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="86C2CC832D4E55813D72DC0F30602FFF" italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis>
</taxonomicName>
comprises species having a unique gland located in the conical pit of the cymbium. The 11 species occurring in Iran and the one from Turkmenistan have the same conical pit and are thus placed in this genus; however, the male palpal tibia and particularly the bulbs are different from those of the species occurring in the Maghreb and surrounding countries. Further revisions are needed to clarify the taxonomy and composition of this group, which are beyond the scope of this paper.
</paragraph>

View file

@ -1,452 +1,452 @@
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<treatment ID-GBIF-Taxon="181753547" LSID="urn:lsid:plazi:treatment:B038AB8CF4435894B2AD946067C7B0E2" httpUri="http://treatment.plazi.org/id/B038AB8CF4435894B2AD946067C7B0E2" lastPageNumber="145" pageId="0" pageNumber="145">
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<taxonomicName LSID="B038AB8C-F443-5894-B2AD-946067C7B0E2" authority="(Nenilin &amp; Fet, 1985)" baseAuthorityName="Nenilin &amp; Fet" baseAuthorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum proszynskii" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="proszynskii">Zodariellum proszynskii (Nenilin &amp; Fet, 1985)</taxonomicName>
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Epigyne of Zodariellum proszynskii A intact, ventral B, C macerated, ventral and dorsal. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure31" httpUri="https://binary.pensoft.net/fig/536908" pageId="0" pageNumber="145">Figs 31A-C</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
<treatment id="B038AB8CF4435894B2AD946067C7B0E2" ID-GBIF-Taxon="181753547" LSID="urn:lsid:plazi:treatment:B038AB8CF4435894B2AD946067C7B0E2" httpUri="http://treatment.plazi.org/id/B038AB8CF4435894B2AD946067C7B0E2" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
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<taxonomicName id="5DEC716A3D5E853E5D22C9B3545200F3" ID-CoL="93S8H" LSID="B038AB8C-F443-5894-B2AD-946067C7B0E2" authority="(Nenilin &amp; Fet, 1985)" baseAuthorityName="Nenilin &amp; Fet" baseAuthorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum proszynskii" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="proszynskii">Zodariellum proszynskii (Nenilin &amp; Fet, 1985)</taxonomicName>
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<figureCitation id="FC7CCDCA4C468F0EEC54D39A250E4C80" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
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<taxonomicName authorityName="Nenilin &amp; Fet" authorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion proszynskii" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="proszynskii">Zodarion proszynskii</taxonomicName>
<subSubSection id="F3AF752B3C6F723094ED72254653A813" pageId="0" pageNumber="145" type="reference_group">
<paragraph id="258F1103C3D07711F0BECF28A74F1C87" pageId="0" pageNumber="145">
<taxonomicName id="B61A0B357509825B349E40E3AC98E807" authorityName="Nenilin &amp; Fet" authorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion proszynskii" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="proszynskii">Zodarion proszynskii</taxonomicName>
Nenilin &amp; Fet, 1985: 618, f. 1-4, 9 (♂♀).
</paragraph>
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<paragraph pageId="0" pageNumber="145">Material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<materialsCitation ID-GBIF-Occurrence="3093634303" collectingDate="1974-07-15" collectorName="A. Senglet" country="Iran" latitude="37.2" location="Qouchan" longLatPrecision="1183" longitude="58.483334" specimenCount="1" specimenCount-female="1" stateProvince="Razavi Khorasan Province">
<collectingCountry name="Iran">Iran</collectingCountry>
<subSubSection id="E729DC64A7E4BDE5C848587D9A57CD7A" pageId="0" pageNumber="145" type="materials_examined">
<paragraph id="BE211A4BE27DCA6E7789D583A7A536FC" pageId="0" pageNumber="145">Material.</paragraph>
<paragraph id="56B070913768170295150B901A047174" pageId="0" pageNumber="145">
<materialsCitation id="2974A522B42F5A68D0F378CB72FADE45" ID-GBIF-Occurrence="3093634303" collectingDate="1974-07-15" collectorName="A. Senglet" country="Iran" latitude="37.2" location="Qouchan" longLatPrecision="1183" longitude="58.483334" specimenCount="1" specimenCount-female="1" stateProvince="Razavi Khorasan Province">
<collectingCountry id="A5ECDE5C129F411DAFBE1342EDEB43F4" name="Iran">Iran</collectingCountry>
:
<emphasis italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Khorasan-e Razavi">Razavi Khorasan Province</collectingRegion>
<emphasis id="3CE245AF767381E565D9F415B155A175" italics="true" pageId="0" pageNumber="145">
<collectingRegion id="B86B700E3EC868FEC4E215B916F63F09" country="Iran" name="Khorasan-e Razavi">Razavi Khorasan Province</collectingRegion>
</emphasis>
:
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="3A2DA40BCE0BA4A30797B348768CBCEC" type="female">1♀</specimenCount>
(MHNG),
<location LSID="urn:lsid:plazi:treatment:B038AB8CF4435894B2AD946067C7B0E2:DD10D950B5255ED4C9546E1D257B3CE7" country="Iran" latitude="37.2" longLatPrecision="1183" longitude="58.483334" name="Qouchan" stateProvince="Razavi Khorasan Province">Qouchan</location>
<location id="DD10D950B5255ED4C9546E1D257B3CE7" LSID="urn:lsid:plazi:treatment:B038AB8CF4435894B2AD946067C7B0E2:DD10D950B5255ED4C9546E1D257B3CE7" country="Iran" latitude="37.2" longLatPrecision="1183" longitude="58.483334" name="Qouchan" stateProvince="Razavi Khorasan Province">Qouchan</location>
,
<geoCoordinate degrees="37" direction="north" minutes="12" orientation="latitude" precision="925" value="37.2">37°12'N</geoCoordinate>
<geoCoordinate id="090360DB7A814F02877F45B35ABC58D2" degrees="37" direction="north" minutes="12" orientation="latitude" precision="925" value="37.2">37°12'N</geoCoordinate>
,
<geoCoordinate degrees="58" direction="east" minutes="29" orientation="longitude" precision="925" value="58.483334">58°29'E</geoCoordinate>
<geoCoordinate id="D0083CA125A3FCD79BF6853A5529B394" degrees="58" direction="east" minutes="29" orientation="longitude" precision="925" value="58.483334">58°29'E</geoCoordinate>
,
<collectingDate value="1974-07-15">15.07.1974</collectingDate>
<collectingDate id="E797D3B6DE9B9A4BF095816E05E8C9D1" value="1974-07-15">15.07.1974</collectingDate>
(
<collectorName>A. Senglet</collectorName>
<collectorName id="8A1F9ADA68A6C2FAF8B325729887584F">A. Senglet</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="materials_examined">
<paragraph pageId="0" pageNumber="145">Comparative material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum bactrianum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bactrianum">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum bactrianum</emphasis>
<subSubSection id="B6DD170F031D68BED0BD934E9474F984" pageId="0" pageNumber="145" type="materials_examined">
<paragraph id="79B7A7E351F923E08E8B058243CC244B" pageId="0" pageNumber="145">Comparative material.</paragraph>
<paragraph id="C2064391A61D2BDF151B33390A57FC51" pageId="0" pageNumber="145">
<taxonomicName id="5C2AA32BB0F293F9217DF2BB3FFB2D0C" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum bactrianum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bactrianum">
<emphasis id="B91713F5B958F3F98628F6075CDC5095" italics="true" pageId="0" pageNumber="145">Zodariellum bactrianum</emphasis>
</taxonomicName>
(Kroneberg, 1875):
<materialsCitation ID-GBIF-Occurrence="3093634310" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Tajikistan" location="Tajikistan" specimenCount="1" specimenCount-male="1">
<specimenCount type="male">1♂</specimenCount>
<materialsCitation id="FFE28AF4412845EEB3A1A4955A6D706D" ID-GBIF-Occurrence="3093634310" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Tajikistan" location="Tajikistan" specimenCount="1" specimenCount-male="1">
<specimenCount id="EEBF734050F03C082A45002FE78ACB4A" type="male">1♂</specimenCount>
from
<collectingCountry name="Tajikistan">Tajikistan</collectingCountry>
<collectingCountry id="42A23B2E20A6E17E428B12F0C1A7D2EB" name="Tajikistan">Tajikistan</collectingCountry>
(Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. SEM images of the male palps of Zodarion nitidum (A-C) and Z. bactrianum (D-F) A, D ventral B, E retrolateral C dorsal F retrodorsal. Abbreviations: Co - conductor, Eb - embolar base, Em - embolus, Ma - median apophysis, Mc - claw of median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process, Tu - tutaculum, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure28" httpUri="https://binary.pensoft.net/fig/536905" pageId="0" pageNumber="145">28D-F</figureCitation>
<figureCitation id="AE7E9CC0F4F6DFF9B65B5C7C3F340801" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. SEM images of the male palps of Zodarion nitidum (A-C) and Z. bactrianum (D-F) A, D ventral B, E retrolateral C dorsal F retrodorsal. Abbreviations: Co - conductor, Eb - embolar base, Em - embolus, Ma - median apophysis, Mc - claw of median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process, Tu - tutaculum, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure28" httpUri="https://binary.pensoft.net/fig/536905" pageId="0" pageNumber="145">28D-F</figureCitation>
)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure28" httpUri="https://binary.pensoft.net/fig/536905" pageId="0" pageNumber="145" start="Figure 28" startId="F28">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 28.</emphasis>
<caption id="A37110105D6D5A517B82A90FCD498AAF" doi="10.3897/zookeys.1035.65767.figure28" httpUri="https://binary.pensoft.net/fig/536905" pageId="0" pageNumber="145" start="Figure 28" startId="F28">
<paragraph id="14524D96915F4C8BBBB1FC06793BEE8B" pageId="0" pageNumber="145">
<emphasis id="BBD4DCECCCBDF1327DD739C59FAE7108" bold="true" pageId="0" pageNumber="145">Figure 28.</emphasis>
SEM images of the male palps of
<taxonomicName baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion nitidum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitidum">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion nitidum</emphasis>
<taxonomicName id="B2CA97C5376C721B563B2D80CE2E262C" baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion nitidum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitidum">
<emphasis id="BE9E0C278465CCA44D1C2963C16EEE7C" italics="true" pageId="0" pageNumber="145">Zodarion nitidum</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
<emphasis id="4BC60731C8EAFB87A3B817ABD0582D2E" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and
<taxonomicName lsidName="Z. bactrianum" pageId="0" pageNumber="145" rank="species" species="bactrianum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. bactrianum</emphasis>
<taxonomicName id="4FA2E473D77CC73381EC4A3BB2C521D8" lsidName="Z. bactrianum" pageId="0" pageNumber="145" rank="species" species="bactrianum">
<emphasis id="1FEB7BDC330EC2103652F2E205B0ABE7" italics="true" pageId="0" pageNumber="145">Z. bactrianum</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
<emphasis id="65ACF3E08D743F140DDD2DA0D669652A" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis>
<emphasis id="47C4B640CE418A0EC705E38E6A238174" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, E</emphasis>
<emphasis id="9C129D844D3156F9D51C78D8F6F6DD46" bold="true" pageId="0" pageNumber="145">B, E</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
<emphasis id="9CC76B23E8FC1519DD6C147F4EE4A2BB" bold="true" pageId="0" pageNumber="145">C</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
<emphasis id="A15E01EB324E81930ECD4E395AB4DB7A" bold="true" pageId="0" pageNumber="145">F</emphasis>
retrodorsal. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis>
<emphasis id="2DFED0F9FFDE01FFEB7E4C37F781CDC9" italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Eb</emphasis>
<emphasis id="9B9936BF02ED4965D93151BE0BAB5BCA" italics="true" pageId="0" pageNumber="145">Eb</emphasis>
- embolar base,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
<emphasis id="F5FFE9D11CE2D9976E2AB579968AD4EF" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="5A62B95EC61A713DB3777E8BC190CC10" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Mc</emphasis>
<emphasis id="360A838B0D367D04BDB7F7B764BEAECF" italics="true" pageId="0" pageNumber="145">Mc</emphasis>
- claw of median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
<emphasis id="42241431956C30BE0EEA602E03A20D11" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral tibial apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Tp</emphasis>
<emphasis id="FEF5BA9081A2036F2D3F81D7033FB9CF" italics="true" pageId="0" pageNumber="145">Tp</emphasis>
- tegular process,
<emphasis italics="true" pageId="0" pageNumber="145">Tu</emphasis>
<emphasis id="BDE8C90E696AE78DBBED5D23BC823082" italics="true" pageId="0" pageNumber="145">Tu</emphasis>
- tutaculum,
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis>
<emphasis id="FF24DE124C1BBEEA2C6F4ABA1A44F4E8" italics="true" pageId="0" pageNumber="145">Va</emphasis>
- ventral apophysis. Scale bars:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity>
<quantity id="C010CCF98F74A4EDC7C127D3AB33F8C3" metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity>
.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145" start="Figure 29" startId="F29">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 29.</emphasis>
<caption id="02961ECC6F8D7970A8A2008339324ED8" doi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145" start="Figure 29" startId="F29">
<paragraph id="04BBC98FF5973822CFC9EB9DA4EEF4E9" pageId="0" pageNumber="145">
<emphasis id="3FA4067DE935BB43E057C9912A844061" bold="true" pageId="0" pageNumber="145">Figure 29.</emphasis>
SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis>
<taxonomicName id="5D2EDF3229BFBD18B2749109B7691B4B" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum">
<emphasis id="D206B147F6DB3B102A57DFAB3D356CC2" italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
<emphasis id="63E12EF4E8C8572866B47C446A934C3C" bold="true" pageId="0" pageNumber="145">A</emphasis>
) and
<taxonomicName lsidName="Z. talyschicum" pageId="0" pageNumber="145" rank="species" species="talyschicum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. talyschicum</emphasis>
<taxonomicName id="F6BF0FEEE8C6837F136E6A80EC9DBECB" lsidName="Z. talyschicum" pageId="0" pageNumber="145" rank="species" species="talyschicum">
<emphasis id="32474EC651636895FC9198CF3B23F382" italics="true" pageId="0" pageNumber="145">Z. talyschicum</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="145">B, C</emphasis>
<emphasis id="E508410255FCF0FB099F9D7B1BC6FC49" bold="true" pageId="0" pageNumber="145">B, C</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, B</emphasis>
<emphasis id="F3C9CC09EB7BB251F45988F1B7786946" bold="true" pageId="0" pageNumber="145">A, B</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
<emphasis id="5C4EF35A90AD0F5F1DB58B4E37ABA031" bold="true" pageId="0" pageNumber="145">C</emphasis>
ventroretrolateral. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Tm</emphasis>
<emphasis id="EC6FFFD67ACC912B2D49545F9DFCE3B0" italics="true" pageId="0" pageNumber="145">Tm</emphasis>
- median tegular process,
<emphasis italics="true" pageId="0" pageNumber="145">Tp</emphasis>
<emphasis id="82E68C50C8C7051D26CE0398E10518EB" italics="true" pageId="0" pageNumber="145">Tp</emphasis>
- prolateral tegular process. Scale bars:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity>
<quantity id="FA39E8A330657C075E1B2CF5727277F0" metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity>
.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145" start="Figure 30" startId="F30">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 30.</emphasis>
<caption id="6FE083E86D8C10CB3FB7A60B6ABC1232" doi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145" start="Figure 30" startId="F30">
<paragraph id="3E735EECB3392C52D9C88F1470AAC0D2" pageId="0" pageNumber="145">
<emphasis id="8C9F0DF34D7B0CF2E841440BDFE4473C" bold="true" pageId="0" pageNumber="145">Figure 30.</emphasis>
Epigynes of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis>
<taxonomicName id="462E650C4654020456E4968B6062B0D2" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum">
<emphasis id="9D0074896D8DE04163DBC285DC01236B" italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-E</emphasis>
<emphasis id="44592BA1F270071FD8EBFE4851B937A0" bold="true" pageId="0" pageNumber="145">A-E</emphasis>
) and
<taxonomicName lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
<taxonomicName id="4743C6BFE99CD197A131ABC0A8EA525C" lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis id="8A265E4EF5717A032DFC8BBB17CAA52E" italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">F-I</emphasis>
<emphasis id="67BE33FACF8F754B4C9BE0DCDBB3B0B9" bold="true" pageId="0" pageNumber="145">F-I</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, F</emphasis>
<emphasis id="86EA4B3BBDC4BB1909680AA4D3A6316E" bold="true" pageId="0" pageNumber="145">A, F</emphasis>
intact, ventral
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
<emphasis id="72884519B37A980333C3850FC1D4E28B" bold="true" pageId="0" pageNumber="145">B</emphasis>
macerated, ventral
<emphasis bold="true" pageId="0" pageNumber="145">C-E, G-I</emphasis>
<emphasis id="EAB4BFFE72834745CBBFAD836E0F0788" bold="true" pageId="0" pageNumber="145">C-E, G-I</emphasis>
macerated, dorsal. Scale bars:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
<quantity id="1B330561076FA86BCA49479ADFEBC0F2" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure31" httpUri="https://binary.pensoft.net/fig/536908" pageId="0" pageNumber="145" start="Figure 31" startId="F31">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 31.</emphasis>
<caption id="F682A2CD384715561E60C8918D38EDC3" doi="10.3897/zookeys.1035.65767.figure31" httpUri="https://binary.pensoft.net/fig/536908" pageId="0" pageNumber="145" start="Figure 31" startId="F31">
<paragraph id="AE099FB0E3171C78DCA7320BAC418C90" pageId="0" pageNumber="145">
<emphasis id="9C89E2EE8988B7F804510DF418666E07" bold="true" pageId="0" pageNumber="145">Figure 31.</emphasis>
Epigyne of
<taxonomicName baseAuthorityName="Nenilin &amp; Fet" baseAuthorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum proszynskii" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="proszynskii">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum proszynskii</emphasis>
<taxonomicName id="7643F354CBDE560CEA7A5A5E6991F07E" baseAuthorityName="Nenilin &amp; Fet" baseAuthorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum proszynskii" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="proszynskii">
<emphasis id="6F43BBFC49C6BC587E761E9FB5A9064F" italics="true" pageId="0" pageNumber="145">Zodariellum proszynskii</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
<emphasis id="87A10CB463DDCDC191C8F27CF9C53A36" bold="true" pageId="0" pageNumber="145">A</emphasis>
intact, ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, C</emphasis>
<emphasis id="C14CC2040E3585AA1FF879F29567C45B" bold="true" pageId="0" pageNumber="145">B, C</emphasis>
macerated, ventral and dorsal. Scale bar:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
<quantity id="89A50D1F649D28E7F8A875FF0F1CAD25" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145" start="Figure 32" startId="F32">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 32.</emphasis>
<caption id="75FD0E76FAD73E7236DDE4CB33E81D07" doi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145" start="Figure 32" startId="F32">
<paragraph id="C6CED1DD8409E023F07064EEF32D7B0F" pageId="0" pageNumber="145">
<emphasis id="9AB37816EE751D9D5EA1C2A455DDBA45" bold="true" pageId="0" pageNumber="145">Figure 32.</emphasis>
Distribution records of
<taxonomicName authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis>
<taxonomicName id="71DB44ED5814BFAE86548468729CC05D" authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="5A22EA0BBEF0145F14B06D0131D5AA74" italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis>
</taxonomicName>
spp. (partim
<emphasis bold="true" pageId="0" pageNumber="145">1-4, 11</emphasis>
<emphasis id="81A08B5F014F628B826F153598C1CF25" bold="true" pageId="0" pageNumber="145">1-4, 11</emphasis>
),
<taxonomicName authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana</emphasis>
<taxonomicName id="35FFEF9903A573F12246A4A7F7CE2DDF" authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="7D5B7FCFF87341DDB2F033F43CE762E6" italics="true" pageId="0" pageNumber="145">Lachesana</emphasis>
</taxonomicName>
spp. (
<emphasis bold="true" pageId="0" pageNumber="145">5-7</emphasis>
<emphasis id="870862C3910843AA57FAADAD5A004D46" bold="true" pageId="0" pageNumber="145">5-7</emphasis>
),
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion raddei" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei">
<emphasis italics="true" pageId="0" pageNumber="145">Parazodarion raddei</emphasis>
<taxonomicName id="C0DDDFC8F9E4669E2EA0CA7BDCCB4483" baseAuthorityName="Simon" baseAuthorityYear="1889" family="Zodariidae" genus="Parazodarion" kingdom="Animalia" lsidName="Parazodarion raddei" order="Aranei" pageId="0" pageNumber="145" rank="species" species="raddei">
<emphasis id="C59F19C79CB7BEC631E0CC70494D8803" italics="true" pageId="0" pageNumber="145">Parazodarion raddei</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="145">8</emphasis>
<emphasis id="DD21FD6AE4598FE4249EFB6FB40123B7" bold="true" pageId="0" pageNumber="145">8</emphasis>
),
<taxonomicName authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Pax</emphasis>
<taxonomicName id="5AF7FD8D1A107A41BACEB296C888C4ED" authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="3DCF4FC4B366B0CB0BBF613006DBFB0E" italics="true" pageId="0" pageNumber="145">Pax</emphasis>
</taxonomicName>
spp. (
<emphasis bold="true" pageId="0" pageNumber="145">9, 10</emphasis>
<emphasis id="741AAD243AA1BA741E59884DF6F79DE5" bold="true" pageId="0" pageNumber="145">9, 10</emphasis>
) and
<taxonomicName authorityName="Simon" authorityYear="1882" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Trygetus</emphasis>
<taxonomicName id="F763A8D50B3AFF13CF2695BACA8578A0" authorityName="Simon" authorityYear="1882" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="E197407341744D795D6005E5246F56C0" italics="true" pageId="0" pageNumber="145">Trygetus</emphasis>
</taxonomicName>
spp. (
<emphasis bold="true" pageId="0" pageNumber="145">11, 12</emphasis>
<emphasis id="FC435744E68E39616B6668DA98C483CA" bold="true" pageId="0" pageNumber="145">11, 12</emphasis>
) in
<collectingCountry name="Iran">Iran</collectingCountry>
<collectingCountry id="C6C101F2511106F71F9E67E424B8748D" name="Iran">Iran</collectingCountry>
and
<collectingCountry name="Turkmenistan">Turkmenistan</collectingCountry>
<collectingCountry id="BEC7080193562EC3B2B47AFBF6D15E4B" name="Turkmenistan">Turkmenistan</collectingCountry>
:
<emphasis bold="true" pageId="0" pageNumber="145">1</emphasis>
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
<emphasis id="DB2114804E5BB5100CF363D9280DFCF3" bold="true" pageId="0" pageNumber="145">1</emphasis>
<taxonomicName id="1EB42CD8DF6766E36EB5B70A4B598243" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis id="74600BD3CA4D97F52782640F06B315D2" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">2</emphasis>
<taxonomicName lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
<emphasis id="D35CD5E4DC3BF6E83DF53DE87667ED50" bold="true" pageId="0" pageNumber="145">2</emphasis>
<taxonomicName id="3666701D3718AB02B83B84033B561DCC" lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis id="9FBBA4E4F038584094117D753928504F" italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">3</emphasis>
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
<emphasis id="58D8F1D099F4D9B41A2FB3D4DFB1BEC4" bold="true" pageId="0" pageNumber="145">3</emphasis>
<taxonomicName id="12E75191FE2FF83681014866CEB1B402" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis id="F09BCB72DB99FD5596674D2F8B65879E" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">4</emphasis>
<taxonomicName lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
<emphasis id="866CFF4BC29C4CC689E176510330465A" bold="true" pageId="0" pageNumber="145">4</emphasis>
<taxonomicName id="6267A066F09872E930245F60E3EBE9CF" lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis id="C505A1778391468723842DB35B34DD11" italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
</taxonomicName>
sp. nov.
<specimenCount type="generic">
<emphasis bold="true" pageId="0" pageNumber="145">5</emphasis>
<taxonomicName lsidName="L. insensibilis" pageId="0" pageNumber="145" rank="species" species="insensibilis">
<emphasis italics="true" pageId="0" pageNumber="145">L. insensibilis</emphasis>
<specimenCount id="4ED81429E213297064ACD374043BE90D" type="generic">
<emphasis id="E723FAC7AA690BBF9947BDB10382EC2A" bold="true" pageId="0" pageNumber="145">5</emphasis>
<taxonomicName id="BED4FE9EBFFA6791C97301F5AEB2AD2C" lsidName="L. insensibilis" pageId="0" pageNumber="145" rank="species" species="insensibilis">
<emphasis id="599CFBCBEF27A7BF64971E4DAFCACF98" italics="true" pageId="0" pageNumber="145">L. insensibilis</emphasis>
</taxonomicName>
</specimenCount>
<specimenCount type="generic">
<emphasis bold="true" pageId="0" pageNumber="145">6</emphasis>
<taxonomicName lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
<specimenCount id="B0F2BDD72ED130195004664F2C655E04" type="generic">
<emphasis id="2D78638266D3E3FD9969EDED98186234" bold="true" pageId="0" pageNumber="145">6</emphasis>
<taxonomicName id="8473EA687AC6E5B16CC1DD9D1A9FA157" lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis id="34D0FE4F10BE963CD03DC3784E240787" italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
</taxonomicName>
</specimenCount>
sp. nov.
<specimenCount type="generic">
<emphasis bold="true" pageId="0" pageNumber="145">7</emphasis>
<taxonomicName lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
<specimenCount id="77EE2F57283B6CED66F1D88FA4EE4676" type="generic">
<emphasis id="0FD296EE84E9AC57C1DD887FE23E0790" bold="true" pageId="0" pageNumber="145">7</emphasis>
<taxonomicName id="ED1DD43B22E2839196AA6FC636392FF0" lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis id="6B579A69EBC87AB56F4E1B6C34B9A0D5" italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
</taxonomicName>
</specimenCount>
sp. nov.
<specimenCount type="generic">
<emphasis bold="true" pageId="0" pageNumber="145">8</emphasis>
<taxonomicName lsidName="P. raddei" pageId="0" pageNumber="145" rank="species" species="raddei">
<emphasis italics="true" pageId="0" pageNumber="145">P. raddei</emphasis>
<specimenCount id="5C50BB8E0AAC1CB72C4768885B996BA2" type="generic">
<emphasis id="87072DE9A6CE21056933628B2CC92D6D" bold="true" pageId="0" pageNumber="145">8</emphasis>
<taxonomicName id="3D448C0E9179A704511C8B25D9E0C70C" lsidName="P. raddei" pageId="0" pageNumber="145" rank="species" species="raddei">
<emphasis id="1EF0A2CA3C061C038B5080A2A7CA9EA6" italics="true" pageId="0" pageNumber="145">P. raddei</emphasis>
</taxonomicName>
</specimenCount>
<specimenCount type="generic">
<emphasis bold="true" pageId="0" pageNumber="145">9</emphasis>
<taxonomicName lsidName="P. ellipita" pageId="0" pageNumber="145" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">P. ellipita</emphasis>
<specimenCount id="6F0C84994DF62CC670307B4B3373B937" type="generic">
<emphasis id="30BC5BCE08A50BD0832ED60118EFCD8F" bold="true" pageId="0" pageNumber="145">9</emphasis>
<taxonomicName id="25371C293CAEF5D4EB46BBFC697D367B" lsidName="P. ellipita" pageId="0" pageNumber="145" rank="species" species="ellipita">
<emphasis id="8594ABD5A4CB139385D21C257B1194FE" italics="true" pageId="0" pageNumber="145">P. ellipita</emphasis>
</taxonomicName>
</specimenCount>
sp. nov.
<specimenCount type="generic">
<emphasis bold="true" pageId="0" pageNumber="145">10</emphasis>
<taxonomicName lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila">
<emphasis italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
<specimenCount id="AC2EA74861E0F1783752F3A5A74A4C2F" type="generic">
<emphasis id="24EAF3B05C74FEDB351C4C92DDBEC454" bold="true" pageId="0" pageNumber="145">10</emphasis>
<taxonomicName id="D6585C63332E7EBAC1AD78C60320B09F" lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila">
<emphasis id="BC95DC2986114351A15359ED908F1FF5" italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
</taxonomicName>
</specimenCount>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">11</emphasis>
<taxonomicName lsidName="A. ovtchinnikovi" pageId="0" pageNumber="145" rank="species" species="ovtchinnikovi">
<emphasis italics="true" pageId="0" pageNumber="145">A. ovtchinnikovi</emphasis>
<emphasis id="22EFC43923B26EABF9CC13B52B3F4A70" bold="true" pageId="0" pageNumber="145">11</emphasis>
<taxonomicName id="B13424CEA8F6C1E718842CE8A7E8ADB3" lsidName="A. ovtchinnikovi" pageId="0" pageNumber="145" rank="species" species="ovtchinnikovi">
<emphasis id="A84C2522F45852E3157D499597B971CB" italics="true" pageId="0" pageNumber="145">A. ovtchinnikovi</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="T. gromovi" pageId="0" pageNumber="145" rank="species" species="gromovi">
<emphasis italics="true" pageId="0" pageNumber="145">T. gromovi</emphasis>
<taxonomicName id="265F174DF793F03DE1C41635F3551BAA" lsidName="T. gromovi" pageId="0" pageNumber="145" rank="species" species="gromovi">
<emphasis id="97DF6F11AC90263F0012E11CF5473CDC" italics="true" pageId="0" pageNumber="145">T. gromovi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="145">12</emphasis>
<taxonomicName lsidName="T. susianus" pageId="0" pageNumber="145" rank="species" species="susianus">
<emphasis italics="true" pageId="0" pageNumber="145">T. susianus</emphasis>
<emphasis id="96752460472E3806335BBE81BF80AD58" bold="true" pageId="0" pageNumber="145">12</emphasis>
<taxonomicName id="E5B7AB2824AAEDFF72C3E82E4FAB3704" lsidName="T. susianus" pageId="0" pageNumber="145" rank="species" species="susianus">
<emphasis id="A50ABCB6203AE65327CDCF40E569BBB5" italics="true" pageId="0" pageNumber="145">T. susianus</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="D15536E1F48D40385F5D20F76E3442FA" pageId="0" pageNumber="145" type="comments">
<paragraph id="FAE8B22F9E8C7F3442DAE1F576A4CB03" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph id="4CABE2A7DC4C602557E59420DDF33FCC" pageId="0" pageNumber="145">
This species was previously known from the original description only. It differs from the similar
<taxonomicName lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae">
<emphasis italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis>
<taxonomicName id="22DF70C58918AC0517E12CDFE064037B" lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae">
<emphasis id="7C6308E63CAFBB1517CCCA6F31BB0E6F" italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis>
</taxonomicName>
by the shape of the male palpal tibia and the epigyne (see
<bibRefCitation author="Nenilin, AB" journalOrPublisher="Zoologicheskiĭ Zhurnal" pageId="0" pageNumber="145" pagination="618 - 620" refId="B17" refString="Nenilin, AB, Fet, VY, 1985. New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia. Zoologicheskiĭ Zhurnal 64: 618 - 620" title="New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia." volume="64" year="1985">Nenilin and Fet 1985</bibRefCitation>
<bibRefCitation id="F30F1C5EF91985ECF82672745D194205" author="Nenilin, AB" journalOrPublisher="Zoologicheskiĭ Zhurnal" pageId="0" pageNumber="145" pagination="618 - 620" refId="B17" refString="Nenilin, AB, Fet, VY, 1985. New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia. Zoologicheskiĭ Zhurnal 64: 618 - 620" title="New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia." volume="64" year="1985">Nenilin and Fet 1985</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in iran">
<paragraph pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="6F824BA35F9E676337D1FB6236317FBA" pageId="0" pageNumber="145" type="records in iran">
<paragraph id="F231D1CF1F254E7F66C50C3AFDFFF372" pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph id="BAD21158921768D04A23564DAEBA985D" pageId="0" pageNumber="145">
Razavi Khorasan (current data) (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
<figureCitation id="075B469ACE0539D17ACD00730C2B0CC0" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in turkmenistan">
<paragraph pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="5B0D4D407E5FA6086E0C9B221340775B" pageId="0" pageNumber="145" type="records in turkmenistan">
<paragraph id="DBABCD03D0F554F7633EEA89A2B91437" pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph>
<paragraph id="5AE24525BBC6DBA1F222C33A5326F4C2" pageId="0" pageNumber="145">
Balkan (
<bibRefCitation author="Nenilin, AB" journalOrPublisher="Zoologicheskiĭ Zhurnal" pageId="0" pageNumber="145" pagination="618 - 620" refId="B17" refString="Nenilin, AB, Fet, VY, 1985. New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia. Zoologicheskiĭ Zhurnal 64: 618 - 620" title="New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia." volume="64" year="1985">Nenilin and Fet 1985</bibRefCitation>
<bibRefCitation id="A7112A588ADED7A60EA2762F4F935961" author="Nenilin, AB" journalOrPublisher="Zoologicheskiĭ Zhurnal" pageId="0" pageNumber="145" pagination="618 - 620" refId="B17" refString="Nenilin, AB, Fet, VY, 1985. New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia. Zoologicheskiĭ Zhurnal 64: 618 - 620" title="New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia." volume="64" year="1985">Nenilin and Fet 1985</bibRefCitation>
) (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
<figureCitation id="2BFB4F0EF462613D5832514B0397ECC6" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145" start="Figure 33" startId="F33">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 33.</emphasis>
<caption id="C02EA5C7EE2BA0CA035B7CD8BFD57687" doi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145" start="Figure 33" startId="F33">
<paragraph id="C10B1A57E7F7007C741C89969EA513C7" pageId="0" pageNumber="145">
<emphasis id="257DB7163396D3697DCE985B1884E20E" bold="true" pageId="0" pageNumber="145">Figure 33.</emphasis>
Distribution records of
<taxonomicName authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis>
<taxonomicName id="1A9CEE7147358B41918BD842798D8A72" authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="8534AE684AC3105DDDF95AE5777DA3C7" italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis>
</taxonomicName>
spp. (partim
<emphasis bold="true" pageId="0" pageNumber="145">1-3, 5, 6, 8, 9</emphasis>
<emphasis id="64E76E00C7AFCF71F4E350AC26701DB4" bold="true" pageId="0" pageNumber="145">1-3, 5, 6, 8, 9</emphasis>
),
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
<taxonomicName id="B640AE8465ADFED68986FDDA2333DD5D" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="84BC7DBA62FACA5A418DBF26FC6C9933" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName>
spp. (
<emphasis bold="true" pageId="0" pageNumber="145">10, 11</emphasis>
<emphasis id="450B07E8564820B99EF080EB11AD866B" bold="true" pageId="0" pageNumber="145">10, 11</emphasis>
) and
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
<taxonomicName id="B86DF6BB78E04CEED67515A7FC70E167" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="C77936BA4222503B64E7B2C432F7F388" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName>
spp. (
<emphasis bold="true" pageId="0" pageNumber="145">4, 7, 12</emphasis>
<emphasis id="D696FC34DF279C19569CC705F327B8D5" bold="true" pageId="0" pageNumber="145">4, 7, 12</emphasis>
) in Iran and Turkmenistan:
<emphasis bold="true" pageId="0" pageNumber="145">1</emphasis>
<taxonomicName lsidName="A. armita" pageId="0" pageNumber="145" rank="species" species="armita">
<emphasis italics="true" pageId="0" pageNumber="145">A. armita</emphasis>
<emphasis id="D1E46D99494C80A4C4280C4801B2EC92" bold="true" pageId="0" pageNumber="145">1</emphasis>
<taxonomicName id="EBD3C22BF61BBFA2F5F1774FA771A46A" lsidName="A. armita" pageId="0" pageNumber="145" rank="species" species="armita">
<emphasis id="A226C1B5AC5CF119CAC91B7FBD68970E" italics="true" pageId="0" pageNumber="145">A. armita</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">2</emphasis>
<taxonomicName lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
<emphasis id="DBF530EBE6409B8E918E895CB8AB7F4B" bold="true" pageId="0" pageNumber="145">2</emphasis>
<taxonomicName id="DD22DB2C7345B278810FE8ABB844454A" lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis id="9DC450ADE5D660E3F5F25C8CEC465872" italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">3</emphasis>
<taxonomicName lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis>
<emphasis id="89E897D72CE64FBA78462E3A7622A59F" bold="true" pageId="0" pageNumber="145">3</emphasis>
<taxonomicName id="01102B8306887501804EBF8BF457597E" lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum">
<emphasis id="FD767A0F50A6ABE1770DBF70CB8B7DC4" italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">4</emphasis>
<taxonomicName lsidName="Z. buettikeri" pageId="0" pageNumber="145" rank="species" species="buettikeri">
<emphasis italics="true" pageId="0" pageNumber="145">Z. buettikeri</emphasis>
<emphasis id="EFCB10AE29A525B87650C1A8CEA76A8F" bold="true" pageId="0" pageNumber="145">4</emphasis>
<taxonomicName id="02A8FB5731B5B6538D22B39CFCF6AB3F" lsidName="Z. buettikeri" pageId="0" pageNumber="145" rank="species" species="buettikeri">
<emphasis id="D825ED10312B149B85D55593EC00FA48" italics="true" pageId="0" pageNumber="145">Z. buettikeri</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="145">5</emphasis>
<taxonomicName lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
<emphasis id="8E8764624C4CF4A8F4878AADBAFF834C" bold="true" pageId="0" pageNumber="145">5</emphasis>
<taxonomicName id="B4D7CAA0987DA27A5FD573864D08366A" lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis id="C0E2F378CD05C967E5D2A2B8B29E845F" italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">6</emphasis>
<taxonomicName lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana">
<emphasis italics="true" pageId="0" pageNumber="145">A. kiana</emphasis>
<emphasis id="C8F5EBFE48FA37F6ECAEB08EE8BD0D42" bold="true" pageId="0" pageNumber="145">6</emphasis>
<taxonomicName id="C85FB6195DC43D4E0D86F310F07CC358" lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana">
<emphasis id="94A6CBB39208BD3435909B0F2AD4E031" italics="true" pageId="0" pageNumber="145">A. kiana</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">7</emphasis>
<taxonomicName lsidName="Z. lutipes" pageId="0" pageNumber="145" rank="species" species="lutipes">
<emphasis italics="true" pageId="0" pageNumber="145">Z. lutipes</emphasis>
<emphasis id="25F3F0EF54FAB225F41EE16A38307F38" bold="true" pageId="0" pageNumber="145">7</emphasis>
<taxonomicName id="DE2885C6EE53B0B4DB6326D79DACDD77" lsidName="Z. lutipes" pageId="0" pageNumber="145" rank="species" species="lutipes">
<emphasis id="0128A9DFCD3A58828521A64421D947A6" italics="true" pageId="0" pageNumber="145">Z. lutipes</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="145">8</emphasis>
<taxonomicName lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
<emphasis id="B43833DB70828684D8F1EDE3E6BE7AF6" bold="true" pageId="0" pageNumber="145">8</emphasis>
<taxonomicName id="DB1BE73D2E6FE2AF6D724AA5BC6E2582" lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis id="3DB04AFC0117CD8C0A06F9410C67124F" italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">9</emphasis>
<taxonomicName lsidName="A. parmida" pageId="0" pageNumber="145" rank="species" species="parmida">
<emphasis italics="true" pageId="0" pageNumber="145">A. parmida</emphasis>
<emphasis id="69CED20E95AC29EC18AEC87840168737" bold="true" pageId="0" pageNumber="145">9</emphasis>
<taxonomicName id="B7FC225D770FEDE6806B1F7291B169B4" lsidName="A. parmida" pageId="0" pageNumber="145" rank="species" species="parmida">
<emphasis id="5C218968B99E991647C2207EBB8B2256" italics="true" pageId="0" pageNumber="145">A. parmida</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">10</emphasis>
<taxonomicName lsidName="Z. proszynskii" pageId="0" pageNumber="145" rank="species" species="proszynskii">
<emphasis italics="true" pageId="0" pageNumber="145">Z. proszynskii</emphasis>
<emphasis id="E66B65D1141768D698D3FF7FD3DA3BDA" bold="true" pageId="0" pageNumber="145">10</emphasis>
<taxonomicName id="64F9CB278EBA055B4F87C68B39B78056" lsidName="Z. proszynskii" pageId="0" pageNumber="145" rank="species" species="proszynskii">
<emphasis id="8DAD4E4ED3E71F600DB60516305F2595" italics="true" pageId="0" pageNumber="145">Z. proszynskii</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="145">11</emphasis>
<taxonomicName lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae">
<emphasis italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis>
<emphasis id="95184A3018014774B2C858E318A0148C" bold="true" pageId="0" pageNumber="145">11</emphasis>
<taxonomicName id="BDD48FC9D148E28E4051FF87566B9B6B" lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae">
<emphasis id="B3DC8FC08E48632B3C0E0F01F1D96C5F" italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="145">12</emphasis>
<taxonomicName lsidName="Z. talyschicum" pageId="0" pageNumber="145" rank="species" species="talyschicum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. talyschicum</emphasis>
<emphasis id="C6D95EF15AD5AE93D613AF191868F066" bold="true" pageId="0" pageNumber="145">12</emphasis>
<taxonomicName id="F8E9462709171B4E6FC196EB5A5BAE77" lsidName="Z. talyschicum" pageId="0" pageNumber="145" rank="species" species="talyschicum">
<emphasis id="9BE1C78A5D4D8AE45ADE4727230DF1AB" italics="true" pageId="0" pageNumber="145">Z. talyschicum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">Previously known only from the type locality in Turkmenistan. This is a new record for Iran, representing the southernmost record in the known genus range.</paragraph>
<subSubSection id="054985D7426EAE74BCD88EE83DFB3F39" pageId="0" pageNumber="145" type="distribution">
<paragraph id="4F6E731070FB4647AFAC842A3B8A32A3" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="7077C86C135DA2DF4543C3B924458997" pageId="0" pageNumber="145">Previously known only from the type locality in Turkmenistan. This is a new record for Iran, representing the southernmost record in the known genus range.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,167 +1,167 @@
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<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. SEM images of the male palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, B, D ventral C, E retrolateral F dorsoretrolateral. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis, Ra - retrolateral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145">, 25A-C</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
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<figureCitation id="46152494F35F26FF83BB34DDEF2B98E3" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">Figs 19A-C</figureCitation>
<figureCitation id="5989CA90BC92E92DCB963E8428CCF013" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. SEM images of the male palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, B, D ventral C, E retrolateral F dorsoretrolateral. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis, Ra - retrolateral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145">, 25A-C</figureCitation>
<figureCitation id="142455B99D9CC093B11EB466C433CD63" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="A9E9EF8445391010562F1233592FC8D2" pageId="0" pageNumber="145" type="type material">
<paragraph id="E53D4ED3210F20FB8A038CFB323B1106" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="D036F9649138050D1B7F13C467EC18BB" pageId="0" pageNumber="145">
<emphasis id="60589CFCED3A8D0E8FB66D874035987F" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Isfahan Province</emphasis>
<emphasis id="1B37337B477954F82E18447965F701F7" italics="true" pageId="0" pageNumber="145">Isfahan Province</emphasis>
: Qamsar and Barzok Protected Area, 55 km SW of Qamsar, 14 km NE Kamoo, near the road of Gargash observatory,
<geoCoordinate degrees="33" direction="north" minutes="37" orientation="latitude" precision="925" value="33.616665">33°37'N</geoCoordinate>
<geoCoordinate id="78D414568B991BD2A03D7441EC0B654C" degrees="33" direction="north" minutes="37" orientation="latitude" precision="925" value="33.616665">33°37'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="19" orientation="longitude" precision="925" value="51.316666">51°19'E</geoCoordinate>
<geoCoordinate id="7AB141CCC27D4182ADBC475316C2F85F" degrees="51" direction="east" minutes="19" orientation="longitude" precision="925" value="51.316666">51°19'E</geoCoordinate>
, 2710 m, 19.05.2016 (P. Ponel).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet refers to a Persian princess, the only daughter of Bardiya (Smerdis), son of Cyrus the Great. Noun in apposition.</paragraph>
<subSubSection id="23D9F20F1CD144EFD85DABE5CAA2DF30" pageId="0" pageNumber="145" type="etymology">
<paragraph id="F172A8668452A98C7775553471C14566" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="0A01ABD223F0F410131CDA6922775C1C" pageId="0" pageNumber="145">The specific epithet refers to a Persian princess, the only daughter of Bardiya (Smerdis), son of Cyrus the Great. Noun in apposition.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="714F434CEA413D905F4E34C8C92B8672" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="95CBD790C464C26368BBD1D1F121E2C3" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="52A834F51748C6C1EF4BB3C48F654E5D" pageId="0" pageNumber="145">
The new species differs from the congeners in the region by its smaller size, by having a dorsal scutum, and by the very short tip (free part) of embolus having a long furrow with a serrate ventral margin. It is closely related to
<taxonomicName lsidName="A. ovtchinnikovi" pageId="0" pageNumber="145" rank="species" species="ovtchinnikovi">
<emphasis italics="true" pageId="0" pageNumber="145">A. ovtchinnikovi</emphasis>
<taxonomicName id="A41D3763C15B697528383FD4605570FF" lsidName="A. ovtchinnikovi" pageId="0" pageNumber="145" rank="species" species="ovtchinnikovi">
<emphasis id="986052AB780596F88524C47F3579DDA1" italics="true" pageId="0" pageNumber="145">A. ovtchinnikovi</emphasis>
</taxonomicName>
sp. nov. from southeastern Turkmenistan, from which it can be differentiated by having an almost as long as wide bulb (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="B65B4858E391103C4B4C401BD31286D6" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
longer than wide), shorter cymbium (length/width ratio 1.4 vs. 1.86), a different shape of median apophysis (posterior portion larger than anterior one, vs. opposite) and the different position of embolic base (6:30 o'clock,
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="951FB32D642C89A82A00BD91EFFA017E" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
8:00 o'clock; cf. Figs
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19B</figureCitation>
<figureCitation id="A8784FC3EA376B8598ACE44483ECDECA" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19B</figureCitation>
and
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22G</figureCitation>
<figureCitation id="6CFA98CE10A0A6FA11F983A48F3954B4" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
<subSubSection id="66B5D79D8BBF87878ED9145E6351E792" pageId="0" pageNumber="145" type="description">
<paragraph id="C6C91FDE1742031C3674C2D052A2F15E" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="0858E77A05CF7D99AFCDF9D531041E84" pageId="0" pageNumber="145">
<emphasis id="09EA750BC0F26C83AFC59D7111F31719" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19A</figureCitation>
<figureCitation id="0D677BA00F155DE5574EC53F6CEFC595" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19A</figureCitation>
. Total length 1.75. Carapace 0.83 long, 0.42 wide at pars cephalica, 0.60 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.10, ALE: 0.06, PME: 0.06, PLE: 0.05, PME-PME: 0.10. Carapace, sternum, labium, chelicerae and maxillae yellowish; carapace with irregular dark patterns. Chelicera with one tooth. Legs yellowish, without annulations. Abdomen dorsally black, covered with scutum, ventrally grayish. Spinnerets pale, uniform in color. Measurements of legs: I: 1.89 (0.42, 0.23, 0.41, 0.41, 0.42), II: 1.64 (0.40, 0.23, 0.31, 0.33, 0.37), III: 1.54 (0.41, 0.21, 0.32, 0.29, 0.31), IV: 2.35 (0.60, 0.26, 0.52, 0.59, 0.38).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="56289EB124928399D6F12EA7B3AFEE76" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19B, C</figureCitation>
<figureCitation id="D1E445885C260A9295F5CF2971181671" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19B, C</figureCitation>
,
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. SEM images of the male palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, B, D ventral C, E retrolateral F dorsoretrolateral. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis, Ra - retrolateral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145">25A-C</figureCitation>
<figureCitation id="565B1981F25433CCCCC7E7653B1D0713" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. SEM images of the male palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, B, D ventral C, E retrolateral F dorsoretrolateral. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis, Ra - retrolateral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145">25A-C</figureCitation>
. Ventral tibial apophysis lacking, RTA (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
<emphasis id="8318C6DD46805B001B87B79A1CF59BEB" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
) almost triangular, slightly longer than tibia; cymbium very broad; tegulum slightly wider than long; sperm duct tracking margin of tegulum, lacking any turns; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="43A41C1CF3C2D54526BC7EBAA247E4AA" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) relatively small, less than half of tegulum's height, posterior part larger than anterior; embolus very short, shorter than median apophysis and conductor (
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis>
<emphasis id="43FA70018ACFD834ADA5BD66574912D8" italics="true" pageId="0" pageNumber="145">Co</emphasis>
), embolus originating at about 6:30 o'clock position with its terminal 2/3 having a longitudinal furrow (
<emphasis italics="true" pageId="0" pageNumber="145">Er</emphasis>
<emphasis id="66334AC3D5716844DEAACE6064E51BF8" italics="true" pageId="0" pageNumber="145">Er</emphasis>
) with a finely serrated ventral margin.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145" start="Figure 25" startId="F25">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 25.</emphasis>
<caption id="A0B0E3FF0F739B1DE2593493E13EC08B" doi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145" start="Figure 25" startId="F25">
<paragraph id="6D9CAB18535DA365E7D1C641EB66734B" pageId="0" pageNumber="145">
<emphasis id="18BFFC9503B457A841C1D09286E9033B" bold="true" pageId="0" pageNumber="145">Figure 25.</emphasis>
SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parmida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parmida">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium parmida</emphasis>
<taxonomicName id="5D68A24C7A81F7C7990AC811DCEB937F" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parmida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parmida">
<emphasis id="D326AF4D01728BD9DFE5542BF3FE247B" italics="true" pageId="0" pageNumber="145">Acanthinozodium parmida</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
<emphasis id="71EF152D0CC8C0F6DE99BE4F7306AED4" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and
<taxonomicName class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion expers" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="expers">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion expers</emphasis>
<taxonomicName id="9A045FF7A07615A38487421F6875BDCA" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion expers" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="expers">
<emphasis id="301138A24C49BD40ACB48701E17E8DBE" italics="true" pageId="0" pageNumber="145">Zodarion expers</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
<emphasis id="37C27760999B4E922614A838D0102014" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, B, D</emphasis>
<emphasis id="D012CCABFF87AB297A962E3BDDFB3160" bold="true" pageId="0" pageNumber="145">A, B, D</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">C, E</emphasis>
<emphasis id="B8451FA4FB25553D2BAE0401B0D50AA6" bold="true" pageId="0" pageNumber="145">C, E</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
<emphasis id="816F4C51676DE6FD7B58F87038793F28" bold="true" pageId="0" pageNumber="145">F</emphasis>
dorsoretrolateral. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Cg</emphasis>
<emphasis id="A7B9B0D8B5FA166B084805197F3571C3" italics="true" pageId="0" pageNumber="145">Cg</emphasis>
- cymbial groove,
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis>
<emphasis id="47B4A478042A4E918D256A93C93F0357" italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
<emphasis id="B6EF05B1FD8ADBD812DA7F9CD1E119E4" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Er</emphasis>
<emphasis id="27CB80EAB52957480399399D1E7C6643" italics="true" pageId="0" pageNumber="145">Er</emphasis>
- embolar ridge,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="C5BF2D618686F2A5E565F55C7AD028A2" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
<emphasis id="649862489E38C5C0C44A4EF4F52D1624" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral tibial apophysis. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="F257324C018D436D7D8CE85E1C83EC3C" pageId="0" pageNumber="145">
<emphasis id="58BACFE6CBAD8AE12E74D627D571B0B6" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="60A33943DE0F6088028E0A1BD6BBD784" pageId="0" pageNumber="145" type="distribution">
<paragraph id="76D7EE9D714437847E757BF9C0DFC9B3" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="E6CA2241CA65DB642D870089CFA29331" pageId="0" pageNumber="145">
Known from the type locality in Isfahan Province, central Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
<figureCitation id="3D82C5B5D8B0F7CB4A1BB7A157CBCF38" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,223 +1,223 @@
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:title id="BB7648F9AFF85B0F570755C243021875">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:namePart>Zamani, Alireza</mods:namePart>
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<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
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<mods:namePart id="161DB7506B52B02D5EFB32AF9379B538">Zamani, Alireza</mods:namePart>
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<mods:namePart>Marusik, Yuri M.</mods:namePart>
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<mods:affiliation id="39F791D7D2FCE6F0C6CC1D0DADF8472D">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<treatment ID-GBIF-Taxon="181753554" LSID="urn:lsid:zoobank.org:act:F3F99EF7-B5F4-42CB-AD3B-2404007AFBA2" httpUri="http://treatment.plazi.org/id/C18D05F563465EA48B4B75AB8EAF40A2" lastPageNumber="145" pageId="0" pageNumber="145">
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<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">Figs 1D-F</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live habitus of male Lachesana perseus sp. nov. Photo: A. Zamani." figureDoi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">, 3F-J</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">, 4J</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
<treatment id="C18D05F563465EA48B4B75AB8EAF40A2" ID-GBIF-Taxon="181753554" LSID="urn:lsid:zoobank.org:act:F3F99EF7-B5F4-42CB-AD3B-2404007AFBA2" httpUri="http://treatment.plazi.org/id/C18D05F563465EA48B4B75AB8EAF40A2" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Lachesaninae">
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<paragraph id="BBA2982FCDCD950D2F7CCCB5A1685745" pageId="0" pageNumber="145">
<taxonomicName id="C33C25152DD2967F0B586564437BF9E6" ID-CoL="98YYX" LSID="http://zoobank.org/F3F99EF7-B5F4-42CB-AD3B-2404007AFBA2" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana perseus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="perseus" status="sp. nov.">Lachesana perseus</taxonomicName>
<taxonomicNameLabel id="61613BBFC11310B8911AF37D0CDC64C1" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="C001E61BA894AD3CE3460CF357CD9EA3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">Figs 1D-F</figureCitation>
<figureCitation id="D5AC0B550EC5C290E8143EA1B5065D0E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live habitus of male Lachesana perseus sp. nov. Photo: A. Zamani." figureDoi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145">, 2</figureCitation>
<figureCitation id="F238BFACD8FDCE897EA02FD10FDC58BE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">, 3F-J</figureCitation>
<figureCitation id="AC2DE1B1721AC498E801F8B75B3239AC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">, 4J</figureCitation>
<figureCitation id="B178CF70A9DBB419C034AB91486F9D5A" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="87671BD4CA74B642AAF06F69A7D87615" pageId="0" pageNumber="145" type="type material">
<paragraph id="576B6D34BF7062F3EF6CB299B3EA7490" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="5B661A38C023F1E6C784EB5654A2377F" pageId="0" pageNumber="145">
<emphasis id="596DCE962F379C0A97CC14DC5AE80F0A" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ and
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis>
<emphasis id="C2FC2A19C29202E1F0029695B3E7B6FF" bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis>
1♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Alborz Province</emphasis>
<emphasis id="59AB063C8F3979DA6FD4AF4837E32D0A" italics="true" pageId="0" pageNumber="145">Alborz Province</emphasis>
: Eshtehard, Jaru,
<geoCoordinate degrees="35" direction="north" minutes="44" orientation="latitude" precision="925" value="35.733334">35°44'N</geoCoordinate>
<geoCoordinate id="5F6C02A3209AD75805951C9A43E85196" degrees="35" direction="north" minutes="44" orientation="latitude" precision="925" value="35.733334">35°44'N</geoCoordinate>
,
<geoCoordinate degrees="50" direction="east" minutes="35" orientation="longitude" precision="925" value="50.583332">50°35'E</geoCoordinate>
<geoCoordinate id="D9AC6272898100BCD28564661B0972E8" degrees="50" direction="east" minutes="35" orientation="longitude" precision="925" value="50.583332">50°35'E</geoCoordinate>
, 10.2018 (A. Zamani).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet refers to the legendary founder of Mycenae and of the Perseid dynasty in Greek mythology. Noun in apposition.</paragraph>
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<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 2.</emphasis>
<subSubSection id="F7BBBD96BA50651BF1DB4A0498E931EC" pageId="0" pageNumber="145" type="etymology">
<paragraph id="23B37499EDB9B3EB05827BCC7BB81175" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="EA3780F47B3CC52CFE115FF1FF9B3E2D" pageId="0" pageNumber="145">The specific epithet refers to the legendary founder of Mycenae and of the Perseid dynasty in Greek mythology. Noun in apposition.</paragraph>
<caption id="5FB1607EB3265A211A3A8617F4AAA8A4" doi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145" start="Figure 2" startId="F2">
<paragraph id="84F050CFCFE1A02C353FA05AEF059E8B" pageId="0" pageNumber="145">
<emphasis id="F28A0BF5C5E14491EB477286B4533FEA" bold="true" pageId="0" pageNumber="145">Figure 2.</emphasis>
Live habitus of male
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana perseus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="perseus">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana perseus</emphasis>
<taxonomicName id="4B73561D9058BB4DBDA2DFF5BA9FB1E8" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana perseus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="perseus">
<emphasis id="348A24E0AE32407E3D19C771DB43BCE1" italics="true" pageId="0" pageNumber="145">Lachesana perseus</emphasis>
</taxonomicName>
sp. nov. Photo: A. Zamani.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="D3C8686AC86D0A2A76334FAB8B713654" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="1867A39548B62E39870A5278289F4ABC" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="CCC88A0E0DA2B33BA791848E3EFDB688" pageId="0" pageNumber="145">
The new species differs from the similar
<taxonomicName lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
<taxonomicName id="F9FF47EFD8655668402C8AA573876293" lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis id="1C2BF81EE2AEFF95398C842F63411ADF" italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
</taxonomicName>
sp. nov. by the lack of spines in the proximal mesal part of the cymbium (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="19905CC5A9CADFC049A5D93A3568B50B" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
present; cf. Fig.
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<figureCitation id="BA5CFC78108A5251F3376253A21EF5B0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3C</figureCitation>
and
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3H</figureCitation>
<figureCitation id="4F90C4BFA2F9717A812FF7F4D17B36A3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3H</figureCitation>
) as well as by the RTA with the terminal half almost as wide as the tibia (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="B15485D65F37F0F8620A09913F302C83" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
1.5 times thinner). The shape of the RTA readily distinguishes this species from congeners in the region (stalk abruptly bent basally and tip gently curved; cf. Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4J</figureCitation>
<figureCitation id="CDE4FAF8955D141F79CE0D5DF3613CFB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4J</figureCitation>
and
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4F-I</figureCitation>
<figureCitation id="F86AF3682A06AFE011C0996E5671A9E0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4F-I</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 3.</emphasis>
<caption id="650E2FF432E1235F78C6962C76E711A8" doi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145" start="Figure 3" startId="F3">
<paragraph id="026255F77C6678A7CB3A4CBCAF741D85" pageId="0" pageNumber="145">
<emphasis id="35A66CB4C877C71249C357D8729094A0" bold="true" pageId="0" pageNumber="145">Figure 3.</emphasis>
Male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana kavirensis</emphasis>
<taxonomicName id="CAD3834CEDDF22D8721D1B322464633F" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis">
<emphasis id="7C46872DF48D5AF5264D1D4282C4029F" italics="true" pageId="0" pageNumber="145">Lachesana kavirensis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-E</emphasis>
<emphasis id="ABE0F8F69321D7569651A884CBD048D1" bold="true" pageId="0" pageNumber="145">A-E</emphasis>
) and
<taxonomicName lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
<taxonomicName id="B1F33A7F38F7F04C992B82D8C93F4F70" lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis id="44FB43EDDEF00979BB20EF01CD1F1C53" italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">F-J</emphasis>
<emphasis id="F954BC4F68BE8C099D96B9AD26FBF8FF" bold="true" pageId="0" pageNumber="145">F-J</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, B, G, F</emphasis>
<emphasis id="95DA4D1687553A2F488B59F9DF6B687A" bold="true" pageId="0" pageNumber="145">A, B, G, F</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, H</emphasis>
<emphasis id="4CCECB1D574E73FF823E6FBC3B6A25FB" bold="true" pageId="0" pageNumber="145">C, H</emphasis>
prolateral
<emphasis bold="true" pageId="0" pageNumber="145">D, I</emphasis>
<emphasis id="956B9E31A0DA1D9E0965CB122ABC13FA" bold="true" pageId="0" pageNumber="145">D, I</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">E, J</emphasis>
<emphasis id="4D303E1E3C0C3E5B00F39F9F0C7A159D" bold="true" pageId="0" pageNumber="145">E, J</emphasis>
dorsal.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 4.</emphasis>
<caption id="588C3E20AD42F9763CECA1A3226281BF" doi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145" start="Figure 4" startId="F4">
<paragraph id="A6A7A4CD1F4CBAC06ED810D46474E752" pageId="0" pageNumber="145">
<emphasis id="B9502AFC8FB772A4EDBB022CD71000F7" bold="true" pageId="0" pageNumber="145">Figure 4.</emphasis>
Male palps of
<taxonomicName class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana dyachkovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dyachkovi">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana dyachkovi</emphasis>
<taxonomicName id="D27736380EE4C0AB3C76D7ADCCBE1B46" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana dyachkovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dyachkovi">
<emphasis id="38362931D4499FD40FEBB63ACEF3A407" italics="true" pageId="0" pageNumber="145">Lachesana dyachkovi</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="145">A-C, F</emphasis>
<emphasis id="3EDA2D76F7969BB3208532ED6D3BEDF6" bold="true" pageId="0" pageNumber="145">A-C, F</emphasis>
),
<taxonomicName lsidName="L. tarabaevi" pageId="0" pageNumber="145" rank="species" species="tarabaevi">
<emphasis italics="true" pageId="0" pageNumber="145">L. tarabaevi</emphasis>
<taxonomicName id="9131979BBA2C374022B84869146E484C" lsidName="L. tarabaevi" pageId="0" pageNumber="145" rank="species" species="tarabaevi">
<emphasis id="A89A0CA90DE3ECEB716E3B3E6F607108" italics="true" pageId="0" pageNumber="145">L. tarabaevi</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="145">D, E, G</emphasis>
<emphasis id="CE501965A218224D23C9750C650B984A" bold="true" pageId="0" pageNumber="145">D, E, G</emphasis>
),
<taxonomicName lsidName="L. blackwalli" pageId="0" pageNumber="145" rank="species" species="blackwalli">
<emphasis italics="true" pageId="0" pageNumber="145">L. blackwalli</emphasis>
<taxonomicName id="F338C72C67A30B7A5FD7ECECBD9B9375" lsidName="L. blackwalli" pageId="0" pageNumber="145" rank="species" species="blackwalli">
<emphasis id="06B0530BE966B2CFB550DAA511AC782A" italics="true" pageId="0" pageNumber="145">L. blackwalli</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis>
<emphasis id="50D3418A5467E5909F6A82CDEEC57093" bold="true" pageId="0" pageNumber="145">H</emphasis>
),
<taxonomicName lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
<taxonomicName id="7D186755AE93311598C33D1435AE6671" lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis id="7AA444B6117E9DD15CB13BF77D3BC90C" italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">I</emphasis>
<emphasis id="9397F1F9DFAE66F852AF06FD73789FE2" bold="true" pageId="0" pageNumber="145">I</emphasis>
) and
<taxonomicName lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
<taxonomicName id="7994865710A2D22E22E1F9A07C8A139E" lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis id="75822E06EAE3B4BEC2A510D996BC28C2" italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">J</emphasis>
<emphasis id="7C8EB94C80D63B9DA0018C035F7826D5" bold="true" pageId="0" pageNumber="145">J</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis>
<emphasis id="16195BAD1A8BE33A1566FE8DA7753759" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, C, E</emphasis>
<emphasis id="03C3EBADB84D3515EDD4719D059BC0ED" bold="true" pageId="0" pageNumber="145">B, C, E</emphasis>
dorsal, prolateral and retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">F-J</emphasis>
<emphasis id="30CA170CCCCC699B297BBE8F1139313C" bold="true" pageId="0" pageNumber="145">F-J</emphasis>
retrolateral tibial apophysis
<emphasis bold="true" pageId="0" pageNumber="145">A-G</emphasis>
<emphasis id="07853A964F99B5622D0E38C073A6E3B6" bold="true" pageId="0" pageNumber="145">A-G</emphasis>
reproduced after
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.28.4.08" author="Fomichev, AA" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="556 - 561" refId="B5" refString="Fomichev, AA, Marusik, YM, 2019. A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan. Arthropoda Selecta 28 (4): 556 - 561, DOI: https://doi.org/10.15298/arthsel.28.4.08" title="A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan." url="https://doi.org/10.15298/arthsel.28.4.08" volume="28" year="2019">Fomichev and Marusik (2019)</bibRefCitation>
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis>
<bibRefCitation id="18D230A92D52BB7499C2BA592102C376" DOI="https://doi.org/10.15298/arthsel.28.4.08" author="Fomichev, AA" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="556 - 561" refId="B5" refString="Fomichev, AA, Marusik, YM, 2019. A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan. Arthropoda Selecta 28 (4): 556 - 561, DOI: https://doi.org/10.15298/arthsel.28.4.08" title="A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan." url="https://doi.org/10.15298/arthsel.28.4.08" volume="28" year="2019">Fomichev and Marusik (2019)</bibRefCitation>
<emphasis id="AF2C07ACB48CFE9CC6CF62BC73FB8FCA" bold="true" pageId="0" pageNumber="145">H</emphasis>
illustrated after
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.29.3.06" author="Oezkuetuek, RS" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="334 - 338" refId="B21" refString="Oezkuetuek, RS, Yagmur, EA, Guecel, S, Shafaie, S, Oezden, Oe, Kunt, KB, 2020. New records of the East Mediterranean Lachesana (Aranei: Zodariidae), with description of a new species. Arthropoda Selecta 29 (3): 334 - 338, DOI: https://doi.org/10.15298/arthsel.29.3.06" title="New records of the East Mediterranean Lachesana (Aranei: Zodariidae), with description of a new species." url="https://doi.org/10.15298/arthsel.29.3.06" volume="29" year="2020">
<normalizedToken originalValue="Özkütük">Oezkuetuek</normalizedToken>
<bibRefCitation id="21537E01B4EB997BB5FA093770DC8530" DOI="https://doi.org/10.15298/arthsel.29.3.06" author="Oezkuetuek, RS" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="334 - 338" refId="B21" refString="Oezkuetuek, RS, Yagmur, EA, Guecel, S, Shafaie, S, Oezden, Oe, Kunt, KB, 2020. New records of the East Mediterranean Lachesana (Aranei: Zodariidae), with description of a new species. Arthropoda Selecta 29 (3): 334 - 338, DOI: https://doi.org/10.15298/arthsel.29.3.06" title="New records of the East Mediterranean Lachesana (Aranei: Zodariidae), with description of a new species." url="https://doi.org/10.15298/arthsel.29.3.06" volume="29" year="2020">
<normalizedToken id="47091E4EE797D80FD5B55CC4BDD1E17C" originalValue="Özkütük">Oezkuetuek</normalizedToken>
et al. (2020)
</bibRefCitation>
. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis>
<subSubSection id="7DD39D0C6D50FB64360E66C80F2FC700" pageId="0" pageNumber="145" type="description">
<paragraph id="55D69154750B663AA75DC2002CB98808" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="0304E9E183716FE27F9A7861B7AB9AD7" pageId="0" pageNumber="145">
<emphasis id="C166BD5B5854D7F83BAF18BBB12FC48A" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1D, E</figureCitation>
<figureCitation id="5B83CF800F2598DDE76BE26DE4C8E73F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1D, E</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live habitus of male Lachesana perseus sp. nov. Photo: A. Zamani." figureDoi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145">2</figureCitation>
<figureCitation id="F440B607D5D6394BC41565849427DB94" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live habitus of male Lachesana perseus sp. nov. Photo: A. Zamani." figureDoi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145">2</figureCitation>
. Total length 10.50. Carapace 5.60 long, 2.60 wide at pars cephalica, 3.88 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.19, ALE: 0.19, PME: 0.16, PLE: 0.19, PME-PME: 0.18. Carapace, sternum, labium, chelicerae and maxillae light brown. Chelicera (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1F</figureCitation>
<figureCitation id="AF24280057BC0A00E04CB022E2589ACE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1F</figureCitation>
) with dense, black setae and one promarginal tooth; fangs curved. Legs the same color as carapace, without annulations and with numerous spines. Abdomen pale, dorsally with a longitudinal dark stripe and several large spots. Spinnerets grayish, uniform in color. Measurements of legs: I: 16.42 (4.69, 2.02, 3.61, 3.56, 2.54), II: 15.48 (4.22, 2.01, 3.19, 3.53, 2.53), III: 16.25 (4.46, 1.96, 2.51, 4.66, 2.66), IV: 19.39 (4.87, 2.32, 3.29, 5.66, 3.25).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="9516173D2BFC2942A4C7EDB07C862D94" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3F-J</figureCitation>
<figureCitation id="18B9EF623E50DC3572D19CB0D9A4AD52" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3F-J</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4J</figureCitation>
<figureCitation id="0AEA5A9D78300C46729788416DC388B3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4J</figureCitation>
. Tibial apophysis more than 1.2 times longer than tibia, widest part over 2 times wider than stalk; cymbium lacking spines in proximal mesal part; bulb elongate, 1.3 times longer than wide; base of embolus large, 1/3 of the tegulum's length; embolus filamentous, lying in chute formed by conductor.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="A5183832E79F7D3898AB82AE58E125CB" pageId="0" pageNumber="145">
<emphasis id="8C73054740359E8ABB48768F6C76EEDD" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="5629D4B1DA20B13B4946C451DE16D0CC" pageId="0" pageNumber="145" type="distribution">
<paragraph id="222616A642345930D5BB0A41A0DA34B3" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="571ADD87054897BA34C3655C2A7A8834" pageId="0" pageNumber="145">
Known only from the type locality in Alborz Province, northern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
<figureCitation id="67C10143A0799B0656F0F1845C3F7A5C" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,173 +1,173 @@
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<treatment ID-GBIF-Taxon="181753543" LSID="urn:lsid:zoobank.org:act:AFBD4904-D586-4FD5-A261-EC34D20EB469" httpUri="http://treatment.plazi.org/id/C61523CF4B07574DA528FA84A2573392" lastPageNumber="145" pageId="0" pageNumber="145">
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<paragraph pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/AFBD4904-D586-4FD5-A261-EC34D20EB469" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium diara" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="diara" status="sp. nov.">Acanthinozodium diara</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18B</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20D-F</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">, 23D-F</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
<treatment id="C61523CF4B07574DA528FA84A2573392" ID-GBIF-Taxon="181753543" LSID="urn:lsid:zoobank.org:act:AFBD4904-D586-4FD5-A261-EC34D20EB469" httpUri="http://treatment.plazi.org/id/C61523CF4B07574DA528FA84A2573392" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection id="DBF9FD37EF7DB0CD8D73DDF5FF69F068" pageId="0" pageNumber="145" type="nomenclature">
<paragraph id="A0C21398B47D8ADD8FFBDFFE0FD1BBF2" pageId="0" pageNumber="145">
<taxonomicName id="AC3012AFA5CFF65E1311C9896FA52472" ID-CoL="98YZ4" LSID="http://zoobank.org/AFBD4904-D586-4FD5-A261-EC34D20EB469" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium diara" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="diara" status="sp. nov.">Acanthinozodium diara</taxonomicName>
<taxonomicNameLabel id="CE2C346A552F3E3D47D2A6F0832907B8" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="0AE384378257BA454D92A368C7BFD922" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18B</figureCitation>
<figureCitation id="A0118C2A434C9328B338707E55D6E2CB" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20D-F</figureCitation>
<figureCitation id="BC248247A28BE2EAAC6CBBF671A7B474" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">, 23D-F</figureCitation>
<figureCitation id="14B3017DAC152629DAA25D3FD6A97C41" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="B0CE2196FEFD079EC83FFA08172025B2" pageId="0" pageNumber="145" type="type material">
<paragraph id="A54AD4488F7219AC9550E07612376EFC" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="868E5E6C8A3B1B3F16ED2AF93854E429" pageId="0" pageNumber="145">
<emphasis id="888AF076A1F20B7BF3ACF22BE08579AE" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Ilam Province</emphasis>
<emphasis id="14586DDC272E57DEFC11019199C4162F" italics="true" pageId="0" pageNumber="145">Ilam Province</emphasis>
: Dizgaran,
<geoCoordinate degrees="33" direction="north" minutes="44" orientation="latitude" precision="925" value="33.733334">33°44'N</geoCoordinate>
<geoCoordinate id="BC423E6AEE3B64AFBD8B413FF876E413" degrees="33" direction="north" minutes="44" orientation="latitude" precision="925" value="33.733334">33°44'N</geoCoordinate>
,
<geoCoordinate degrees="46" direction="east" minutes="59" orientation="longitude" precision="925" value="46.983334">46°59'E</geoCoordinate>
<geoCoordinate id="28692CB0353B832047A50FBA2D05E496" degrees="46" direction="east" minutes="59" orientation="longitude" precision="925" value="46.983334">46°59'E</geoCoordinate>
, 16.5.1974 (A. Senglet).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratype</emphasis>
<emphasis id="26880B9F7B6FADE54BCE9738B98CD032" bold="true" italics="true" pageId="0" pageNumber="145">Paratype</emphasis>
♂ (MMUE), IRAN:
<emphasis italics="true" pageId="0" pageNumber="145">Lorestan Province</emphasis>
<emphasis id="81B3901D817D85D659921FFEF57CFC67" italics="true" pageId="0" pageNumber="145">Lorestan Province</emphasis>
: Dorood, 31.7.2011 (S. Zaruni).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="B19243988F1CC56C7286A51D85428265" pageId="0" pageNumber="145" type="etymology">
<paragraph id="4B23365B69A24AB7561E99C58C59B0F8" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="9FD6169358D0C6E859C79178355AA3BE" pageId="0" pageNumber="145">
The specific epithet is a Persian feminine name meaning
<normalizedToken originalValue="“motherland”">&quot;motherland&quot;</normalizedToken>
<normalizedToken id="C1F4439045A81D57E5652D898D30EC29" originalValue="“motherland”">&quot;motherland&quot;</normalizedToken>
. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium diara" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium diara</emphasis>
<subSubSection id="16CE91F586ED480AE8501AEFD29C206E" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="0E6F10DDDDF71ACA2659187529DD2245" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="2E6992972796AC9E41846D9A4177B800" pageId="0" pageNumber="145">
<taxonomicName id="00918FF2A5AA338E474537A78D48A237" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium diara" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="diara">
<emphasis id="CF1B8542DE497B1E2500D1CC3C5D0B34" italics="true" pageId="0" pageNumber="145">Acanthinozodium diara</emphasis>
</taxonomicName>
sp. nov. is very similar to
<taxonomicName lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
<taxonomicName id="A2D7F337B2859B592B07E591D836E23F" lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis id="7A9F52CFBDFD50DB85B5FFC017F9D901" italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
</taxonomicName>
sp. nov. by the shape of the RTA, the ventral tibial apophysis and the curvature of the embolus but differs by the shape of the median apophysis (cf. Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20D</figureCitation>
<figureCitation id="F82B0365F2A9294BA0F71AAE29390B5D" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20D</figureCitation>
and
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20J</figureCitation>
<figureCitation id="10E1596C16FF2681A3DCF34F8156F9F1" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20J</figureCitation>
) and the conductor having a small retrolateral indentation apically (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="1522E2BEFBDA9A07972D3556D0765CF4" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
without indentation).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis>
<subSubSection id="1652BC0AFC3CEAF48E44D0D530C798F0" pageId="0" pageNumber="145" type="description">
<paragraph id="1D930A5301CCEB567EE6582003EB1B1F" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="4AD75DB6CC54AF8578D8B4CAD37E0D08" pageId="0" pageNumber="145">
<emphasis id="EE56903B074FFD7E0A718DC2DDE00E17" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18B</figureCitation>
<figureCitation id="57FE2AF270D676C74F9099E63D427BE0" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18B</figureCitation>
. Total length 4.70. Carapace 2.18 long, 0.92 wide at pars cephalica, 1.52 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.19, ALE: 0.12, PME: 0.10, PLE: 0.12, PME-PME: 0.25. Carapace dark brown, with irregular dark patches and lines. Sternum, labium and maxillae light brown. Chelicera dark brown, with one retromarginal tooth. Legs light brown, without annulations. Abdomen black, without any pattern; slightly lighter ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 9.49 (2.38, 0.71, 2.16, 2.76, 1.48), II: 8.76 (2.10, 0.78, 1.98, 2.50, 1.40), III: 8.36 (2.11, 0.70, 1.69, 2.73, 1.13), IV: 11.42 (3.00, 0.81, 2.60, 3.59, 1.42).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="15F1AFC2B46CADA0B98D729EAA0B83E1" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20D-F</figureCitation>
<figureCitation id="C03CC2A2A382EC8817F89A4A1512F3EF" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20D-F</figureCitation>
,
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">23D-F</figureCitation>
<figureCitation id="3D49112824A69546CB4A52A5B94580AD" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">23D-F</figureCitation>
. RTA long and conical, with a small projection apically (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20E</figureCitation>
<figureCitation id="9FC303AD89C3A31453DA8F5DCFB503AC" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20E</figureCitation>
); tegulum with posterior process; ventral apophysis small and conical; median apophysis almost as long as wide, with outgrowths on all four sides, posterior one largest; embolus broad basally, originating at about the 7
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
<normalizedToken id="FC1A120D7414BEC54918C90BA3378603" originalValue="oclock">o'clock</normalizedToken>
position; embolus proper thin and steadily curving.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145" start="Figure 20" startId="F20">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 20.</emphasis>
<caption id="976E7D1B5A5660979B38B882A3B2BF24" doi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145" start="Figure 20" startId="F20">
<paragraph id="5BE4B760FCE50992A0B7C980C42B051B" pageId="0" pageNumber="145">
<emphasis id="C4C01CCE1E1F8B176D67FDFE81BBAB5E" bold="true" pageId="0" pageNumber="145">Figure 20.</emphasis>
Male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium armita</emphasis>
<taxonomicName id="58C3481789AD175352CCD1C356DD1C4D" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita">
<emphasis id="8EC769C0022BCDA6EA99A17BBBCB94F1" italics="true" pageId="0" pageNumber="145">Acanthinozodium armita</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
<emphasis id="91A02158D9F44BF3764C8CEB74460053" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
),
<taxonomicName lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
<taxonomicName id="A6BB2AE41768A126C3ABEF7B83A6601E" lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis id="28DC94BF852692D24AAC57D7751F381C" italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
<emphasis id="04ACC5CE025FA9C2E28712756FC32478" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
),
<taxonomicName lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana">
<emphasis italics="true" pageId="0" pageNumber="145">A. kiana</emphasis>
<taxonomicName id="F68280C70455E20A16F98C170E92045F" lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana">
<emphasis id="070FB45F2C9FED9F4CC75E69934EEBB2" italics="true" pageId="0" pageNumber="145">A. kiana</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">G, H</emphasis>
<emphasis id="0B97FCA3048452981F08598FEEFEBCF6" bold="true" pageId="0" pageNumber="145">G, H</emphasis>
) and
<taxonomicName lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
<taxonomicName id="D0F2263C72DF88D20A2A1CDC6892C064" lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis id="D2F31508A0DD688C78825E8D2D83ABC4" italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">I-K</emphasis>
<emphasis id="C68DCF14C55440E74F71B93765FCDB1C" bold="true" pageId="0" pageNumber="145">I-K</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D, G, J</emphasis>
<emphasis id="19C20669F8B74299E851D3549EE1E9F4" bold="true" pageId="0" pageNumber="145">A, D, G, J</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, E, H, I</emphasis>
<emphasis id="67CCF3E3BE2D69048D6A4219E1575631" bold="true" pageId="0" pageNumber="145">B, E, H, I</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, F, K</emphasis>
<emphasis id="ECDC123DBA045D60A8B669F2C773645B" bold="true" pageId="0" pageNumber="145">C, F, K</emphasis>
dorsal. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="F64B20D5B49E90451A35AF6B9A49CA4F" pageId="0" pageNumber="145">
<emphasis id="9AFAF06F6D563902946DC985AA8C7212" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="55F8A6E4E68DBC8219230A4E1DD30660" pageId="0" pageNumber="145" type="distribution">
<paragraph id="C9A07C783C998ABEC624D99754E70634" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="BEBCD9D6C8BE9FFF834285D872CADE88" pageId="0" pageNumber="145">
Known from the listed localities in Ilam and Lorestan provinces, western Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
<figureCitation id="A84BE9C8A8627D736BD13F861759540D" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,252 +1,252 @@
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<mods:namePart id="A4720F5F02CACD0048BFC2EFB76A81A0">Zamani, Alireza</mods:namePart>
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<mods:namePart>Marusik, Yuri M.</mods:namePart>
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
<mods:namePart id="2EA9547663A741A285F09394C11A829D">Marusik, Yuri M.</mods:namePart>
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<mods:affiliation id="523D491B9C78AC73E30458045E093A7F">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<mods:title id="E436BD91CD7BF89F81E4E7BED90756C3">ZooKeys</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-04-27</mods:number>
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<mods:number id="8BBB32C102139714920412196770C54A">2021-04-27</mods:number>
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<mods:number>1035</mods:number>
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<treatment ID-GBIF-Taxon="181753544" LSID="urn:lsid:zoobank.org:act:7BBBC164-8239-4F57-9F44-4F5DE93F8C24" httpUri="http://treatment.plazi.org/id/D5FCAC79232E53A39DAF2BF008656B1E" lastPageNumber="145" pageId="0" pageNumber="145">
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/7BBBC164-8239-4F57-9F44-4F5DE93F8C24" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa" status="sp. nov.">Acanthinozodium atrisa</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11A, B</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">, 12A-C</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">, 14A-C</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Epigyne of Acanthinozodium atrisa sp. nov. A posteroventral B posterior C, D ventral. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure16" httpUri="https://binary.pensoft.net/fig/536893" pageId="0" pageNumber="145">, 16A-D</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
<treatment id="D5FCAC79232E53A39DAF2BF008656B1E" ID-GBIF-Taxon="181753544" LSID="urn:lsid:zoobank.org:act:7BBBC164-8239-4F57-9F44-4F5DE93F8C24" httpUri="http://treatment.plazi.org/id/D5FCAC79232E53A39DAF2BF008656B1E" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection id="DA58599AD2FC7D559BE324FD970941D1" pageId="0" pageNumber="145" type="nomenclature">
<paragraph id="59001C7E25FB05C3C6F100F950DF7D98" pageId="0" pageNumber="145">
<taxonomicName id="FC477103F54CF71F4F21B9645C748A90" ID-CoL="98YYW" LSID="http://zoobank.org/7BBBC164-8239-4F57-9F44-4F5DE93F8C24" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa" status="sp. nov.">Acanthinozodium atrisa</taxonomicName>
<taxonomicNameLabel id="6CAE441758F079C3A6E6C54DF9CCD0D1" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="855A7F30DD367648F17313DC41783521" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11A, B</figureCitation>
<figureCitation id="C978D18B4BF3EA8A409A7CB007BD1A32" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">, 12A-C</figureCitation>
<figureCitation id="A0D684CF55DA0BF26EC00D79FA97CB43" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">, 14A-C</figureCitation>
<figureCitation id="A06518477021E8E2540878ECC15C31B8" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Epigyne of Acanthinozodium atrisa sp. nov. A posteroventral B posterior C, D ventral. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure16" httpUri="https://binary.pensoft.net/fig/536893" pageId="0" pageNumber="145">, 16A-D</figureCitation>
<figureCitation id="CEC31E19211931C32FAC31202401F865" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="2EE9EF08D63E3BA5026040C76C475BB2" pageId="0" pageNumber="145" type="type material">
<paragraph id="811E99740F897D905397A1037F4CB5BA" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="37B694769B0327A17CC7F206D11DD7B9" pageId="0" pageNumber="145">
<emphasis id="124F2C6A32EF599D1F066CB41F394C82" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Tehran Province</emphasis>
<emphasis id="AA9E731DE8F1B9D6A934EC2601C17069" italics="true" pageId="0" pageNumber="145">Tehran Province</emphasis>
: Jamshidieh Mts.,
<geoCoordinate degrees="35" direction="north" minutes="49" orientation="latitude" precision="925" value="35.816666">35°49'N</geoCoordinate>
<geoCoordinate id="C2A4A6416863210704824F920903ED83" degrees="35" direction="north" minutes="49" orientation="latitude" precision="925" value="35.816666">35°49'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="27" orientation="longitude" precision="925" value="51.45">51°27'E</geoCoordinate>
<geoCoordinate id="20327AABEDED0D415476500C10E67A77" degrees="51" direction="east" minutes="27" orientation="longitude" precision="925" value="51.45">51°27'E</geoCoordinate>
, 05.2015 (A. Zamani).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
<emphasis id="C0C8EF3B490CC62341EF0E668C9EFD55" bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
: 1♂2♀ (MHNG), same data as holotype; 5♂3♀ (MMUE), Latian Dam,
<geoCoordinate degrees="35" direction="north" minutes="48" orientation="latitude" precision="925" value="35.8">35°48'N</geoCoordinate>
<geoCoordinate id="09DDE9CC8980937C6278E4E88D7ECCAB" degrees="35" direction="north" minutes="48" orientation="latitude" precision="925" value="35.8">35°48'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="08" orientation="longitude" precision="925" value="51.133335">51°08'E</geoCoordinate>
<geoCoordinate id="EEADF44E2B72F8F5D2BB4359F4406361" degrees="51" direction="east" minutes="08" orientation="longitude" precision="925" value="51.133335">51°08'E</geoCoordinate>
, 6-19.06.2000 (Y.M. Marusik); 1♂30♀1j. (MMUE), Plant Protection Institute,
<geoCoordinate degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate>
<geoCoordinate id="EDC8C6C4804E9FFDA43C5BBF88898C0A" degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="24" orientation="longitude" precision="925" value="51.4">51°24'E</geoCoordinate>
<geoCoordinate id="E7C6B968A93B1A06AA32B9700827907D" degrees="51" direction="east" minutes="24" orientation="longitude" precision="925" value="51.4">51°24'E</geoCoordinate>
, 7-22.06.2000 (Y.M. Marusik); 2♀ (MMUE), 5 km north of Tehran, Tochal Mts.,
<geoCoordinate degrees="35" direction="north" minutes="53" orientation="latitude" precision="925" value="35.883335">35°53'N</geoCoordinate>
<geoCoordinate id="9957957A27E308EC6BEB28128B07984D" degrees="35" direction="north" minutes="53" orientation="latitude" precision="925" value="35.883335">35°53'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="20" orientation="longitude" precision="925" value="51.333332">51°20'E</geoCoordinate>
<geoCoordinate id="15BE8E3739F35BD16F2E398D48B40BA3" degrees="51" direction="east" minutes="20" orientation="longitude" precision="925" value="51.333332">51°20'E</geoCoordinate>
, 2000-2900 m, 16.06.2000 (Y.M. Marusik); 9♀1j. (ZMMU), 80 km east of Tehran, Damavand area, Aroo,
<geoCoordinate degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate>
<geoCoordinate id="8CD06789976DF9FA9A7C5800E185F034" degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate>
,
<geoCoordinate degrees="52" direction="east" minutes="27" orientation="longitude" precision="925" value="52.45">52°27'E</geoCoordinate>
<geoCoordinate id="457E98F3E667FC7FEDCB6733DCB76B46" degrees="52" direction="east" minutes="27" orientation="longitude" precision="925" value="52.45">52°27'E</geoCoordinate>
, 15.06.2000 (Y.M. Marusik &amp; F. Mozaffarian);
<emphasis italics="true" pageId="0" pageNumber="145">Qazvin Province</emphasis>
<emphasis id="48DA8893C675FBAC2A79DA2B1BF860D7" italics="true" pageId="0" pageNumber="145">Qazvin Province</emphasis>
: 1♂2♀ (MHNG), Agha Baba,
<geoCoordinate degrees="36" direction="north" minutes="19" orientation="latitude" precision="925" value="36.316666">36°19'N</geoCoordinate>
<geoCoordinate id="1B15D7693DF32DC44992196947CD4DD8" degrees="36" direction="north" minutes="19" orientation="latitude" precision="925" value="36.316666">36°19'N</geoCoordinate>
,
<geoCoordinate degrees="49" direction="east" minutes="49" orientation="longitude" precision="925" value="49.816666">49°49'E</geoCoordinate>
<geoCoordinate id="98C054FEC141A40B5D470395C6E65253" degrees="49" direction="east" minutes="49" orientation="longitude" precision="925" value="49.816666">49°49'E</geoCoordinate>
, 06.07.1974 (A. Senglet); 1♂1♀ (MHNG), Tarazan, Lowshan,
<geoCoordinate degrees="36" direction="north" minutes="28" orientation="latitude" precision="925" value="36.466667">36°28'N</geoCoordinate>
<geoCoordinate id="03F73198B34AF28F1FA12D57F0F57D65" degrees="36" direction="north" minutes="28" orientation="latitude" precision="925" value="36.466667">36°28'N</geoCoordinate>
,
<geoCoordinate degrees="49" direction="east" minutes="31" orientation="longitude" precision="925" value="49.516666">49°31'E</geoCoordinate>
<geoCoordinate id="43D80C2E627B9AFB6439513B8603E5F1" degrees="49" direction="east" minutes="31" orientation="longitude" precision="925" value="49.516666">49°31'E</geoCoordinate>
, 08.08.1974 (A. Senglet).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet is a Persian feminine name meaning &quot;queen of fire&quot;. Noun in apposition.</paragraph>
<subSubSection id="001956B52441282BB7FE4004592C7A52" pageId="0" pageNumber="145" type="etymology">
<paragraph id="7FA4ACFE34F2DE09B38636A323C52FB5" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="837F99CB067C887F1408FDCFF1247C3C" pageId="0" pageNumber="145">The specific epithet is a Persian feminine name meaning &quot;queen of fire&quot;. Noun in apposition.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="959D44240DB1ED47BBD1689463115614" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="76AA9E641E860D24E332D71CA0CC923B" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="99B4F89220FB9A2D6957F9078FBCAAD1" pageId="0" pageNumber="145">
The new species has an abdominal pattern similar to that of
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
<taxonomicName id="3E31F2BD1CCEF3D7AD2E6093E8449CB3" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis id="1FA0E65A559A69A4AFB0106111EEA6F3" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName>
sp. nov. (dark abdomen with posterior median white stripe; less distinct in males) but differs by having a dark pars cephalica (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="70E5C94CB2BFA5CC0242AFF599113677" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
pale). The male palps of the two species differ by the RTA being thin and more than twice as long as the tibia in
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
<taxonomicName id="AD4729341B155ECA006FE0F41E6D9B6D" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis id="9F5AE418E8C6C1B8CA38260A4DAB230E" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="A7377E2676461395CE2A50814A22A2DB" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
broad and as long as the tibia). Judging by the shape of the male palp,
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
<taxonomicName id="915CCCFAE5E3C5724D43BFAA8199E4DD" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis id="208A6ED24F102C5772872AA3D2C9EE76" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName>
sp. nov. is probably closely related to
<taxonomicName lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
<taxonomicName id="485757E298B005D8A454E67970D6C77E" lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis id="545145733BE8415AEE23CA75A4C90995" italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
</taxonomicName>
sp. nov., as both have a long RTA, a similarly shaped median apophysis, a posterior tegular process and an embolus with an anterior process. The two species differ by the shape of the embolic process (
<emphasis italics="true" pageId="0" pageNumber="145">Ep</emphasis>
<emphasis id="4B29EB2B7D0CFC5F6FCFED221CC9074C" italics="true" pageId="0" pageNumber="145">Ep</emphasis>
), wider than the embolus and gently bent at the tip in
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
<taxonomicName id="C480775EC359673EAD485FDDC6AFD858" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis id="69AD8C937F06C2F46469459DAF06EBC3" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="7CA67ADC502462FA711BA6086DD0CE77" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
spine-like). The epigyne of
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
<taxonomicName id="A8DE1FEBA98F1B267A937FCCCBCCC500" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis id="BC243AFFF0636DED9693C6CF29C1D0C9" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName>
sp. nov. is most similar to that of
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
<taxonomicName id="887F94A4F8CA54857CD9130DA2B9E03C" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis id="AABA7D2457B8463550C6EE754FF83FFD" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName>
sp. nov. in having a small anterior hood and a thin furrow between the lateral lobes but differs by the furrow being shorter than the bursa copulatrix (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="C31D84EBB4ED7829AD7D88B1F87CEF81" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
the same length as bursa).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis>
<subSubSection id="0CB2A26A3BE45DD9D506F56187810305" pageId="0" pageNumber="145" type="description">
<paragraph id="1D97C22490EB7A991DECEDCC0875113F" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="1D124B8296807B3F82D11EBCD7372B58" pageId="0" pageNumber="145">
<emphasis id="AF1E32F09914FEE87FB892A1ECE73304" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11A</figureCitation>
<figureCitation id="75BE483F4CA7038B45D017468BBC7BEA" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11A</figureCitation>
. Total length 1.78. Carapace 0.81 long, 0.39 wide at pars cephalica, 0.62 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.07, ALE: 0.06, PME: 0.05, PLE: 0.05, PME-PME: 0.10. Carapace, sternum, labium, chelicerae and maxillae yellowish; carapace with irregular dark brown patterns. Chelicera with retromarginal tooth. Legs yellowish, without annulations. Abdomen dorsally black with longitudinal pale median band and grayish ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 2.39 (0.61, 0.25, 0.52, 0.53, 0.48), II: 2.07 (0.54, 0.26, 0.45, 0.46, 0.36), III: 2.06 (0.57, 0.25, 0.37, 0.48, 0.39), IV: 2.86 (0.69, 0.29, 0.64, 0.77, 0.47).
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 11.</emphasis>
<caption id="B39431BB8A4CFD52E84007F25A661BB7" doi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145" start="Figure 11" startId="F11">
<paragraph id="A289F7BC27B9CF1F33E38A205E1A9467" pageId="0" pageNumber="145">
<emphasis id="E43C30826F069826423704E8C02E6DF5" bold="true" pageId="0" pageNumber="145">Figure 11.</emphasis>
Habitus of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis>
<taxonomicName id="FF514550E611F5000B6030B3276EFBF9" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa">
<emphasis id="AF9220CD5BC1E7DC928A227FAEA0D88B" italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A, B</emphasis>
<emphasis id="48B826D5014E89A7ABC1D3A72C2E5680" bold="true" pageId="0" pageNumber="145">A, B</emphasis>
)
<taxonomicName lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
<taxonomicName id="E667D6034B409194786C1A1611D9C933" lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis id="35C01EE83AF5438F9D9C4644C7D9394C" italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">C-E</emphasis>
<emphasis id="4905A3995EC64ED1CB50B64A1C4424F0" bold="true" pageId="0" pageNumber="145">C-E</emphasis>
),
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
<taxonomicName id="BFAD611CD0FDA3FC3C6E94225253EC95" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis id="23412446932EFE6ABD8D13F6284C3BE7" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">F, G</emphasis>
<emphasis id="CBAB16C23EA6A61D6A902E334F8354A1" bold="true" pageId="0" pageNumber="145">F, G</emphasis>
) and
<taxonomicName lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
<taxonomicName id="376B701740644B9AC2BDF88056F44E3F" lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis id="473C93AD0C6EFA4CFCC8D8488BC9E022" italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis>
<emphasis id="8607E09476CB12E223AFDE51174303F1" bold="true" pageId="0" pageNumber="145">H</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, C, F, H</emphasis>
<emphasis id="6FF08D0B602AF8A7ACD50F5E81805F95" bold="true" pageId="0" pageNumber="145">A, C, F, H</emphasis>
males, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
<emphasis id="CA16909374FB2DE890E07B9D1929326D" bold="true" pageId="0" pageNumber="145">D</emphasis>
male, ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, G, E</emphasis>
<emphasis id="4E73A2FBC70B0D15A6FEB98B4F7DBADA" bold="true" pageId="0" pageNumber="145">B, G, E</emphasis>
females, dorsal. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<paragraph id="C2E3BC64C7B5B88157C3B2D9410F8854" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">12A-C</figureCitation>
<figureCitation id="7A014041EF2A2D28137E3ADD03295A88" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">12A-C</figureCitation>
,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14A-C</figureCitation>
<figureCitation id="0EF9E8FFC3A37CC00D87C10EF89AC5FB" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14A-C</figureCitation>
. RTA (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
<emphasis id="FAA8EF256A3F761844665353EA48135A" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
) long (almost as long as bulb) and thin (7 times longer than wide), slightly bent; tegulum with posterior process, sperm duct almost straight retrolaterally, and gently bent along prolateral side; median apophysis wider than long; embolus (
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
<emphasis id="6D6CCAC09A98210E3FDB0C45033E90CF" italics="true" pageId="0" pageNumber="145">Em</emphasis>
) broad basally, originating at about the 7
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
<normalizedToken id="22EF041EB99C0BF09088D6891C139927" originalValue="oclock">o'clock</normalizedToken>
position, bifurcated terminally, with broad anterior process; embolus proper thin and straight, with small tubercle.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 12.</emphasis>
<caption id="DE27AF243D7D0361839AFFAF6CE5559D" doi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145" start="Figure 12" startId="F12">
<paragraph id="0E651A63255E198B3738B9A65B1356FF" pageId="0" pageNumber="145">
<emphasis id="800F79475A9D06F75ADC7DF7B43955A8" bold="true" pageId="0" pageNumber="145">Figure 12.</emphasis>
Male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis>
<taxonomicName id="D55A075781F4F62909F0AC399B4EEF05" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa">
<emphasis id="9ABB158DDAFA850ED15A8313AC8981B7" italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
<emphasis id="6722C5241DC5A02AD38F5B2C4441C6DA" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and
<taxonomicName lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
<taxonomicName id="66166BA8BEAA74A3F1F452DF545253F8" lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis id="85891AA643D85205039AF67CA00277AB" italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
<emphasis id="08067CF6EF3AEA924BCF5253239A52E9" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis>
<emphasis id="044DDE9011F942812EB56392169174A1" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, C, F</emphasis>
<emphasis id="0B8C8BA940D7AED07A3CC25F206414C1" bold="true" pageId="0" pageNumber="145">B, C, F</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
<emphasis id="FFF2647DDFDCDC18E80146F379A71915" bold="true" pageId="0" pageNumber="145">E</emphasis>
dorsal. Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="2148FA7A601E8C70B860132D2FEEDA84" pageId="0" pageNumber="145">
<emphasis id="EF314CA62F08D3A6688B5C6367B48683" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11B</figureCitation>
<figureCitation id="EFBAA1E60593AE5F690725E954CDCA63" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11B</figureCitation>
. Total length 2.38. Carapace 0.87 long, 0.39 wide at pars cephalica, 0.59 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.07, ALE: 0.06, PME: 0.05, PLE: 0.05, PME-PME: 0.10. Coloration as in male, with paler carapace and less prominent abdominal median stripe. Measurements of legs: I: 2.36 (0.71, 0.24, 0.45, 0.51, 0.45), II: 2.18 (0.56, 0.28, 0.42, 0.48, 0.44), III: 2.16 (0.54, 0.27, 0.42, 0.51, 0.42), Fe IV: 0.76, rest of the segments missing.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="1EA2C4F6F180467D26963D739A1CBCC5" pageId="0" pageNumber="145">
Epigyne as in Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Epigyne of Acanthinozodium atrisa sp. nov. A posteroventral B posterior C, D ventral. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure16" httpUri="https://binary.pensoft.net/fig/536893" pageId="0" pageNumber="145">16A-D</figureCitation>
<figureCitation id="1F21B260E9E07D3520F81A986C6A63E9" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Epigyne of Acanthinozodium atrisa sp. nov. A posteroventral B posterior C, D ventral. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure16" httpUri="https://binary.pensoft.net/fig/536893" pageId="0" pageNumber="145">16A-D</figureCitation>
. Fovea triangular, and small anterior hood present; bursae subtriangular, wider anteriorly, separated by less than one of their radii; receptacles smaller than bursae, ovoid, separated by more than 3 diameters.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="0721DCD3DF6E21A81C0B5B0CA79AA9B0" pageId="0" pageNumber="145" type="distribution">
<paragraph id="C0298897008790FCB13B756900880874" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="876C630757CE5B2CA843E488A162BE20" pageId="0" pageNumber="145">
Known only from the listed localities in Tehran and Qazvin provinces, northern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
<figureCitation id="0BBC674903C4EE2787AE7DD0C93DDD18" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>

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@ -1,79 +1,79 @@
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Genus
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<paragraph pageId="0" pageNumber="145">Type species.</paragraph>
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Audouin, 1826 from Egypt.
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<emphasis italics="true" pageId="0" pageNumber="145">Lachesana</emphasis>
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spp. can easily be distinguished from other zodariids occurring in Iran and Turkmenistan by their large size (&gt;10 mm
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smaller) and numerous spines (&gt;20 on each leg
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<emphasis id="DDFF8DC62F83181772BA745076182EBC" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
lacking, or &lt;15). So far, out of the eight species known from this genus, four are known solely on the basis of male specimens. Male palps are very similar to one another and differ mostly in the proportions of the tibial apophysis and the bulb.
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<mods:identifier id="F2683822E803CAA8EC9A58C5D4D03A48" type="DOI">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:identifier>
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<treatment ID-GBIF-Taxon="181753541" LSID="urn:lsid:zoobank.org:act:BCEEC5DD-E5BA-483A-A43F-DF518429CC54" httpUri="http://treatment.plazi.org/id/E852B3FAE0A657ECB5448E7C4B62F79F" lastPageNumber="145" pageId="0" pageNumber="145">
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/BCEEC5DD-E5BA-483A-A43F-DF518429CC54" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita" status="sp. nov.">Acanthinozodium armita</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18A</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20A-C</figureCitation>
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">, 22A-C, F</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
<treatment id="E852B3FAE0A657ECB5448E7C4B62F79F" ID-GBIF-Taxon="181753541" LSID="urn:lsid:zoobank.org:act:BCEEC5DD-E5BA-483A-A43F-DF518429CC54" httpUri="http://treatment.plazi.org/id/E852B3FAE0A657ECB5448E7C4B62F79F" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection id="90F74AE85D0A1A358F70F470409EB9FF" pageId="0" pageNumber="145" type="nomenclature">
<paragraph id="820AC47B31A07809A75E1B264D971CCE" pageId="0" pageNumber="145">
<taxonomicName id="14E6FF79AB961BE7ABB6F3C5838F77D2" ID-CoL="98YZ6" LSID="http://zoobank.org/BCEEC5DD-E5BA-483A-A43F-DF518429CC54" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita" status="sp. nov.">Acanthinozodium armita</taxonomicName>
<taxonomicNameLabel id="37B2FA1E27FC19624E9BCC149F32F1E4" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="9172DF65F06A97D0DF2142A74E31BA9D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18A</figureCitation>
<figureCitation id="F59D97F52562FC33112E3B4D6304BC4E" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20A-C</figureCitation>
<figureCitation id="E1D4DBEFBB3EBC977317B966993E8325" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">, 22A-C, F</figureCitation>
<figureCitation id="EA1CABA9F65995AA5D084D11957131AF" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="6BA58904D5B4C54F2C78033D009D812C" pageId="0" pageNumber="145" type="type material">
<paragraph id="C8990DF91A002EBA2CF3893778C66F84" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="45ED1EDFEB7D78D79CE0F738B7FFB5CA" pageId="0" pageNumber="145">
<emphasis id="097193212896CE0AB61ED5D48E33DC56" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MMUE), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Tehran Province</emphasis>
<emphasis id="C5C6177E81F18C3D95C18627D35A45E9" italics="true" pageId="0" pageNumber="145">Tehran Province</emphasis>
: northwest of Tehran, Sardor area,
<geoCoordinate degrees="35" direction="north" minutes="50" orientation="latitude" precision="925" value="35.833332">35°50'N</geoCoordinate>
<geoCoordinate id="2977E2959212D520D2F96835C8A31AB5" degrees="35" direction="north" minutes="50" orientation="latitude" precision="925" value="35.833332">35°50'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="05" orientation="longitude" precision="925" value="51.083332">51°05'E</geoCoordinate>
<geoCoordinate id="207E7CA4CDC0F7DF96FE0F87AD2A45F2" degrees="51" direction="east" minutes="05" orientation="longitude" precision="925" value="51.083332">51°05'E</geoCoordinate>
, 13.06.2000 (Y.M. Marusik).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratype</emphasis>
<emphasis id="7D732EA9C623B852F6183041B94E2DD3" bold="true" italics="true" pageId="0" pageNumber="145">Paratype</emphasis>
♂ (NHMW), Iran: Haji Abad, 06.1972 (G. Pretzman &amp; A. Konetschnig).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="740E1E210D5F1D2B27AF82701A04A574" pageId="0" pageNumber="145" type="etymology">
<paragraph id="EACBFA9EDD872D9CBE79B6D63F597A74" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="91F70C731406F93D6C0A9179B13D5A55" pageId="0" pageNumber="145">
The specific epithet is a Persian feminine name meaning
<normalizedToken originalValue="“righteous”">&quot;righteous&quot;</normalizedToken>
<normalizedToken id="B59029E1FA0260DCC38D048624E30111" originalValue="“righteous”">&quot;righteous&quot;</normalizedToken>
,
<normalizedToken originalValue="“virtuous”">&quot;virtuous&quot;</normalizedToken>
<normalizedToken id="7AE32738A000379F665A6D28F69694C4" originalValue="“virtuous”">&quot;virtuous&quot;</normalizedToken>
, and
<normalizedToken originalValue="“good”">&quot;good&quot;</normalizedToken>
<normalizedToken id="7C0FB8619AE353688084E707A5C52DFD" originalValue="“good”">&quot;good&quot;</normalizedToken>
. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="F72EAA90981A952192EA7C87525D8AF7" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="F68B8DE8CFF0795CEE985889DDE1B3E5" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="6BE109FEB64EE11E29EA34CDDCD406F5" pageId="0" pageNumber="145">
The new species differs from all congeners in the region by the short RTA having 2 claw-like outgrowths on the tip (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="A85CF7B423E7C7B328FF63F2B0CA23A2" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
tip tapering and lacking 2 claws) and also by the modified embolus, widening near the tip (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="471EB4466C6022B48ECE3BC58A74D787" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
unmodified).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis>
<subSubSection id="108222B0A652FFD92FBBDFA88E0E08A1" pageId="0" pageNumber="145" type="description">
<paragraph id="190471E5D4C8BB3C33C14D27FACB4834" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="2E068691B01A990DA4118284E5B2C77A" pageId="0" pageNumber="145">
<emphasis id="EFEE4B53FB13ABF1013EFF4CFB8DA975" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18A</figureCitation>
<figureCitation id="AE57D6A628C2179848530F4DC8D7F799" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18A</figureCitation>
. Total length 4.70. Carapace 2.18 long, 0.92 wide at pars cephalica, 1.52 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.19, ALE: 0.12, PME: 0.10, PLE: 0.12, PME-PME: 0.25. Carapace dark brown, with irregular dark patches and lines. Sternum, labium and maxillae light brown. Chelicera dark brown, with one tooth. Legs light brown, without annulations. Abdomen black, without any pattern; slightly lighter ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 9.49 (2.38, 0.71, 2.16, 2.76, 1.48), II: 8.76 (2.10, 0.78, 1.98, 2.50, 1.40), III: 8.36 (2.11, 0.70, 1.69, 2.73, 1.13), IV: 11.42 (3.00, 0.81, 2.60, 3.59, 1.42).
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145" start="Figure 18" startId="F18">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 18.</emphasis>
<caption id="57DB8917C4AB16507381647D6058C6F7" doi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145" start="Figure 18" startId="F18">
<paragraph id="385049E296746D457E64487539DAD115" pageId="0" pageNumber="145">
<emphasis id="9D346FA1C90A7EDAEF1827580F78B979" bold="true" pageId="0" pageNumber="145">Figure 18.</emphasis>
Habitus of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium armita</emphasis>
<taxonomicName id="3644AF4578D521158500C7F15102C0C9" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita">
<emphasis id="738EF1B2AA99F942E13053D689DDF52B" italics="true" pageId="0" pageNumber="145">Acanthinozodium armita</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
<emphasis id="CDC0B9EEC6D681F5F7539413B5409301" bold="true" pageId="0" pageNumber="145">A</emphasis>
),
<taxonomicName lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
<taxonomicName id="18F60B76E0D579B8EDEFFD8EBE7CC95D" lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis id="5DFC51D5ED3E22105D602B55141AB9F1" italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
<emphasis id="366C417DD99A2F2D6EC747A7F73412BB" bold="true" pageId="0" pageNumber="145">B</emphasis>
),
<taxonomicName lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
<taxonomicName id="812C4F17F20836AB1AFD31A3C6B2270A" lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis id="5CCF8162C06C51B2092A6C2F32DCFFB6" italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">C, D</emphasis>
<emphasis id="77EE5C05C56CA2C85E70FC6C390E7655" bold="true" pageId="0" pageNumber="145">C, D</emphasis>
),
<taxonomicName lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis>
<taxonomicName id="9412D8F6EDD49219FADD3CB2D4D910E6" lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum">
<emphasis id="F896D31DFB9F1B797BDB37F4B79EA1D2" italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">E-J</emphasis>
<emphasis id="9812AE9AD5D403FE908CCF299F995904" bold="true" pageId="0" pageNumber="145">E-J</emphasis>
),
<taxonomicName lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana">
<emphasis italics="true" pageId="0" pageNumber="145">A. kiana</emphasis>
<taxonomicName id="5CF21C23BB32105993BCFF5DC0866D17" lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana">
<emphasis id="042746512954FBA540F07B9C1BBF357B" italics="true" pageId="0" pageNumber="145">A. kiana</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">K</emphasis>
<emphasis id="FABCC9C7FDB7471C4F2C8DD655E8F324" bold="true" pageId="0" pageNumber="145">K</emphasis>
) and
<taxonomicName lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
<taxonomicName id="D5975806357747AA1052D3C8D42F75C1" lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis id="C188B9C61233F8EE871B3297809CB646" italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">L</emphasis>
<emphasis id="D8247BBB1D62DA0A7BB24C3D2E1BBE2B" bold="true" pageId="0" pageNumber="145">L</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A-C, F, G, K, L</emphasis>
<emphasis id="382B78F4AA902362D2D35624AEFF692D" bold="true" pageId="0" pageNumber="145">A-C, F, G, K, L</emphasis>
males, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis>
<emphasis id="8B46D53FC2C608B26664BBDE73EFBAEB" bold="true" pageId="0" pageNumber="145">H</emphasis>
male, ventral
<emphasis bold="true" pageId="0" pageNumber="145">D, E, I</emphasis>
<emphasis id="98E432A89D49CE85AB93F35BECFDDD5A" bold="true" pageId="0" pageNumber="145">D, E, I</emphasis>
females, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">J</emphasis>
<emphasis id="4C88CA72E6D5DA225979986B1E367B23" bold="true" pageId="0" pageNumber="145">J</emphasis>
female, ventral. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145" start="Figure 19" startId="F19">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 19.</emphasis>
<caption id="1D74D68B995CC3CBF27D128C3F39A0D1" doi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145" start="Figure 19" startId="F19">
<paragraph id="6D139C3F47D06CEE150334E1C4D9B25D" pageId="0" pageNumber="145">
<emphasis id="AD80F19B4F9E43FBC511B15C982E0950" bold="true" pageId="0" pageNumber="145">Figure 19.</emphasis>
Male habitus and palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parmida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parmida">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium parmida</emphasis>
<taxonomicName id="876786E54784BF6EF55517C3E9EE4607" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parmida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parmida">
<emphasis id="9EE42A5D5EC24D4725D7B4488A3F240B" italics="true" pageId="0" pageNumber="145">Acanthinozodium parmida</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
<emphasis id="A90C11A744494F29627DCCDCFB162862" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and
<taxonomicName class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion expers" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="expers">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion expers</emphasis>
<taxonomicName id="7B3677A5CF94FBB3185AABA0D2C84066" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion expers" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="expers">
<emphasis id="39FEFE31531B45870F8349C99B61DF76" italics="true" pageId="0" pageNumber="145">Zodarion expers</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
<emphasis id="CEE9C2AAC8331AA831A9313D4AECF3E4" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis>
<emphasis id="BC09B887BD8F3E234DF92DAB0C14E51D" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
habitus, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">B, E</emphasis>
<emphasis id="BE2347C85621533CDC46F8AA9681775F" bold="true" pageId="0" pageNumber="145">B, E</emphasis>
palp, ventral
<emphasis bold="true" pageId="0" pageNumber="145">C, F</emphasis>
<emphasis id="AEC0D0C18EF08F078CE627918BEB7C3F" bold="true" pageId="0" pageNumber="145">C, F</emphasis>
palp, retrolateral. Scale bar: 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
<emphasis id="63E7EC074AA1754BA65D38CCC78EFEE4" bold="true" pageId="0" pageNumber="145">A</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<paragraph id="EC27F418AE6C7EFF828AC919EEBB0FF0" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20A-C</figureCitation>
<figureCitation id="505FD69DF2DE3866C96E340EAD9496A3" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20A-C</figureCitation>
,
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22A-C, F</figureCitation>
<figureCitation id="D3465EBA21F9928DA71E367DE8229D16" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22A-C, F</figureCitation>
. RTA short and stout, with 2 claw-like outgrowths on the tip; ventral apophysis absent; tegulum longer than wide; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="EBE6261F237E1A3693CAA9914332A7BE" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) longer than wide, with outgrowths on all four sides; embolus with a distinct ridge (
<emphasis italics="true" pageId="0" pageNumber="145">Er</emphasis>
<emphasis id="E58A9ECC0764446385E0B5A3AB9ECA8B" italics="true" pageId="0" pageNumber="145">Er</emphasis>
); embolus proper twisted basally and widened near the tip.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="24D733B7B5CF07138FB3EC2CEDEB584E" pageId="0" pageNumber="145">
<emphasis id="8F0452C80B0282E8FB3C905E036E3B3F" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="75774EDEED2C98BFEFCCF62D755A3FD3" pageId="0" pageNumber="145" type="distribution">
<paragraph id="B5376CD01905E46E8ECB47F31964DD33" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="6EED1A963991BC483DE132E076E35103" pageId="0" pageNumber="145">
Known from the type locality in Tehran Province, northern Iran, and another locality (Haji Abad) which cannot be georeferenced because there are many places with this name in Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
<figureCitation id="D97B44155E9182FE8621F2E356A5365A" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,72 +1,72 @@
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<treatment id="F64B37BC569252D1AA17139AE84C73BD" ID-GBIF-Taxon="181753538" LSID="urn:lsid:plazi:treatment:F64B37BC569252D1AA17139AE84C73BD" httpUri="http://treatment.plazi.org/id/F64B37BC569252D1AA17139AE84C73BD" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
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<paragraph id="04B7991DDB65E5DA3D27883FDC7671B1" pageId="0" pageNumber="145">
Genus
<taxonomicName LSID="F64B37BC-5692-52D1-AA17-139AE84C73BD" authority="Ovtchinnikov, Ahmad &amp; Gurko, 2009" authorityName="Ovtchinnikov, Ahmad &amp; Gurko" authorityYear="2009" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Parazodarion Ovtchinnikov, Ahmad &amp; Gurko, 2009</taxonomicName>
<taxonomicName id="266B95BCD45A2692E41EB30135C49811" ID-CoL="6HHK" LSID="F64B37BC-5692-52D1-AA17-139AE84C73BD" authority="Ovtchinnikov, Ahmad &amp; Gurko, 2009" authorityName="Ovtchinnikov, Ahmad &amp; Gurko" authorityYear="2009" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Parazodarion Ovtchinnikov, Ahmad &amp; Gurko, 2009</taxonomicName>
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<subSubSection pageId="0" pageNumber="145" type="type species">
<paragraph pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion raddei" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion raddei</emphasis>
<subSubSection id="43A1D19B8486EA5CA7F5C311F1BDB8FA" pageId="0" pageNumber="145" type="type species">
<paragraph id="10E07E10B9F71E4D6B46DDD767F514E5" pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph id="7DFBAFF73089EDF24DB7DF905E401D06" pageId="0" pageNumber="145">
<taxonomicName id="BA1DCF19FBED91B028472E9463DAFE79" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion raddei" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei">
<emphasis id="6B562DC4ACC0BE8823644AE08815039A" italics="true" pageId="0" pageNumber="145">Zodarion raddei</emphasis>
</taxonomicName>
Simon, 1889 from Turkmenistan.
</paragraph>
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<subSubSection pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">Monotypic genus, differing from all other genera in the region by having an elongate cymbium, an S-shaped sperm duct and embolus, a long spine-like outgrowth of the embolic base and an epigyne with large transverse oval fovea.</paragraph>
<subSubSection id="9EEA65EA4CC8907C04B195016BFE9D1B" pageId="0" pageNumber="145" type="comments">
<paragraph id="BAC8500A047C8A93E9D7E9EF72076AAD" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph id="26E7A0364A5EFACE2055F9F26541765F" pageId="0" pageNumber="145">Monotypic genus, differing from all other genera in the region by having an elongate cymbium, an S-shaped sperm duct and embolus, a long spine-like outgrowth of the embolic base and an epigyne with large transverse oval fovea.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,168 +1,168 @@
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<mods:title id="3E51B8D6E1766A0838AB38336CB6C469">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<treatment ID-GBIF-Taxon="181753542" LSID="urn:lsid:zoobank.org:act:820D70DA-633E-4CF3-B11E-3B16C6974CBC" httpUri="http://treatment.plazi.org/id/F7A70D2BB0355179BD17C0C957A48B7D" lastPageNumber="145" pageId="0" pageNumber="145">
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<taxonomicName LSID="http://zoobank.org/820D70DA-633E-4CF3-B11E-3B16C6974CBC" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium kiana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kiana" status="sp. nov.">Acanthinozodium kiana</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18K</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20G, H</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">, 24E, F</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
<treatment id="F7A70D2BB0355179BD17C0C957A48B7D" ID-GBIF-Taxon="181753542" LSID="urn:lsid:zoobank.org:act:820D70DA-633E-4CF3-B11E-3B16C6974CBC" httpUri="http://treatment.plazi.org/id/F7A70D2BB0355179BD17C0C957A48B7D" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection id="F5813EF32B097AE5DF2CB13F72CE56AE" pageId="0" pageNumber="145" type="nomenclature">
<paragraph id="ADDC701B4127FB5F5262E0FE0E7E64BA" pageId="0" pageNumber="145">
<taxonomicName id="2620E204C470D403397E260E9E7D3DF4" ID-CoL="98YYT" LSID="http://zoobank.org/820D70DA-633E-4CF3-B11E-3B16C6974CBC" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium kiana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kiana" status="sp. nov.">Acanthinozodium kiana</taxonomicName>
<taxonomicNameLabel id="EA621D1911A520FAC024C80555934EE9" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="6DF48F9C2493BA18921E7E66AE6EC346" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18K</figureCitation>
<figureCitation id="746C2DBAACF57DD509A133F0FECE5CF5" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20G, H</figureCitation>
<figureCitation id="B32AF5144791761CDF2FFA34C346C16D" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">, 24E, F</figureCitation>
<figureCitation id="D88A3176E9194E7E922EFE64BB9D3AA9" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="FB31BA65D335BB7679A6A140950D4E40" pageId="0" pageNumber="145" type="type material">
<paragraph id="E780596864E40442C79A8B8865D381D9" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="2670CCC3F4ED174A93E77BDC0B66F303" pageId="0" pageNumber="145">
<emphasis id="0653DE923475CF6904AF930580184AE5" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Kurdistan Province</emphasis>
<emphasis id="297D105F9C53E06A415425796831CD82" italics="true" pageId="0" pageNumber="145">Kurdistan Province</emphasis>
: south of Divandareh,
<geoCoordinate degrees="35" direction="north" minutes="45" orientation="latitude" precision="925" value="35.75">35°45'N</geoCoordinate>
<geoCoordinate id="3203D4E2DEE6C265104AB631151B7635" degrees="35" direction="north" minutes="45" orientation="latitude" precision="925" value="35.75">35°45'N</geoCoordinate>
,
<geoCoordinate degrees="47" direction="east" minutes="05" orientation="longitude" precision="925" value="47.083332">47°05'E</geoCoordinate>
<geoCoordinate id="AE303C15925EEEEABB47A52DA3C4DD44" degrees="47" direction="east" minutes="05" orientation="longitude" precision="925" value="47.083332">47°05'E</geoCoordinate>
, 23.06.1975 (A. Senglet).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="32A35433033BABCC3A702F2BF7470A6F" pageId="0" pageNumber="145" type="etymology">
<paragraph id="793C658FC57A57DD04B27C6CB91AB495" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="7DF25EF92F9C21A7AEB55104F5167F02" pageId="0" pageNumber="145">
The specific epithet is a Kurdish feminine name meaning
<normalizedToken originalValue="“nature”">&quot;nature&quot;</normalizedToken>
<normalizedToken id="DACE80CAE16265ECF19CEB4DF7B7AA7B" originalValue="“nature”">&quot;nature&quot;</normalizedToken>
. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="843A84C3A8008F64A66BBF2D2857BBB6" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="AC0131B425CE09CC8D2E28CA031B23A2" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="4AF9B06270B729D2B340AB2782C6492D" pageId="0" pageNumber="145">
The new species is very similar to
<taxonomicName lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
<taxonomicName id="00042A2C526C6F4D9D392F0CFA248621" lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis id="F54FA21D3BF9F58C32A761A2E4B13622" italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
</taxonomicName>
sp. nov. by the shape of the RTA and ventral tibial apophysis but differs by the shape of the prolateral outgrowth of the median apophysis which is finger-like and triangular, terminally with a small hook (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="131D69F7914E96E36AECCAD67CA49087" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
broad and bifurcated).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
<subSubSection id="46AA43C4AF21B190F1271CE728D1D649" pageId="0" pageNumber="145" type="description">
<paragraph id="89937DDC1CE1A1B7D86ED092A566C6D8" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="E4CB6D1D58231C1F0C04AB36075987B3" pageId="0" pageNumber="145">
<emphasis id="19389DA2D0F8E5AFF4FB8CFC7168FFD3" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Habitus as in Fig.
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<figureCitation id="E522DF4524DB9E3FF2831781F65CDD21" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18K</figureCitation>
. Total length 3.80. Carapace 2.08 long, 1.03 wide at pars cephalica, 1.85 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.17, ALE: 0.12, PME: 0.09, PLE: 0.10, PME-PME: 0.23. Carapace light brown, darker at pars cephalica. Sternum, labium, maxillae and chelicerae yellowish. Legs yellowish, with numerous macrosetae and without annulations. Abdomen grayish dorsally, lighter ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 7.57 (2.02, 0.75, 1.67, 1.97, 1.16), II: 7.06 (1.84, 0.74, 1.50, 1.95, 1.03), III: 7.24 (1.98, 0.71, 1.47, 2.08, 1.00), IV: 9.46 (2.63, 0.78, 2.26, 2.77, 1.02).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="338D61E75D0AE9EA50551CE85A6955E7" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20G, H</figureCitation>
<figureCitation id="4C30AAB6C691726A4399C19424A5F5E3" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20G, H</figureCitation>
,
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">24E, F</figureCitation>
<figureCitation id="78085FDEB991283367D9731EE4B758C6" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">24E, F</figureCitation>
. RTA long, with a small projection apically; ventral apophysis relatively small and conical; tegulum with posterior process; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="53C964630C94A14F0EE5D7662C274E6C" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) wider than long, with a distinct prolateral outgrowth which is triangular terminally and with a small hook; embolus broad basally, originating at about the 7
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
<normalizedToken id="AF3CB37A6D1044BE8D3600C8442D8306" originalValue="oclock">o'clock</normalizedToken>
position; embolus proper relatively broad and distinctly curved over the median apophysis.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145" start="Figure 24" startId="F24">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 24.</emphasis>
<caption id="A8CAEB55244FE3C3A9B4B479629C8FB9" doi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145" start="Figure 24" startId="F24">
<paragraph id="36978201FFE5725177B69EB97DFD86FA" pageId="0" pageNumber="145">
<emphasis id="E3BE34C51FBE74EC42F5A1C5E0FA9B62" bold="true" pageId="0" pageNumber="145">Figure 24.</emphasis>
SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium masa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="masa">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium masa</emphasis>
<taxonomicName id="0B49705E4776091B2F56712923396F8E" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium masa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="masa">
<emphasis id="B8D9FEBE398732DF014F15FCD9A44B1E" italics="true" pageId="0" pageNumber="145">Acanthinozodium masa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A, B</emphasis>
<emphasis id="6FE1E8D34E3E087317425599AFFF24C0" bold="true" pageId="0" pageNumber="145">A, B</emphasis>
),
<taxonomicName lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
<taxonomicName id="972F40DD663A6D6BEBA442609B1FCB06" lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis id="3B3EEAF7DD59809A4A082EFCC160FF75" italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">C, D</emphasis>
<emphasis id="82C553B6EC10EFC0D00A7BDD57C88776" bold="true" pageId="0" pageNumber="145">C, D</emphasis>
) and
<taxonomicName lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana">
<emphasis italics="true" pageId="0" pageNumber="145">A. kiana</emphasis>
<taxonomicName id="409D3411313FDB23420F1FDBE1600CBF" lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana">
<emphasis id="6E1E1199E3ED8EED7D793E90E83224CE" italics="true" pageId="0" pageNumber="145">A. kiana</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">E, F</emphasis>
<emphasis id="878096E70211846428074DC6C65325A5" bold="true" pageId="0" pageNumber="145">E, F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, C, E</emphasis>
<emphasis id="345276AA707DECD0206C1A91CE822BB5" bold="true" pageId="0" pageNumber="145">A, C, E</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, F</emphasis>
<emphasis id="DDAA6F14DFC5D819C211BD513EBEAA33" bold="true" pageId="0" pageNumber="145">B, F</emphasis>
retroventral
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
<emphasis id="54C30348B51A4A105120AD0A15A61E7A" bold="true" pageId="0" pageNumber="145">D</emphasis>
retrolateral. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis>
<emphasis id="1FAC00ED7D21971E77F29395503FE3A9" italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
<emphasis id="B77D01D0F0AE611D281DFF9F283EAFC7" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
<emphasis id="38D0814A97675534E9DBDD6026F520C0" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
<emphasis id="0AFE89752273BCE5ABE9C72D7D720D8A" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral tibial apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis>
<emphasis id="57D3C84D10723AD98F318F2DBDCBBA23" italics="true" pageId="0" pageNumber="145">Va</emphasis>
- ventral apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Tp</emphasis>
<emphasis id="EABCF13B4B42DA297AE67BC2E4BA7F43" italics="true" pageId="0" pageNumber="145">Tp</emphasis>
- tegular process. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="339AABB4376EED1D7ED4907F9DFAF0E7" pageId="0" pageNumber="145">
<emphasis id="B6C9F3E41C3D16BA2B1FE5B4D1001112" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="28FFBE9BE2BB40890DAE80EFCF0BA66D" pageId="0" pageNumber="145" type="distribution">
<paragraph id="D30AE5DDE9362C5F76E958F6EEB060A7" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="9EAADFD9100AC83980A6300FFC410EAB" pageId="0" pageNumber="145">
Known from the type locality in Kurdistan Province, western Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
<figureCitation id="5AF6863333BA68541356B7B6A9CACD21" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,108 +1,109 @@
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<emphasis bold="true" box="[151,310,1075,1102]" italics="true" pageId="17" pageNumber="108">Duroca lenta</emphasis>
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<emphasis id="4297EAF09B27FFD2FF3CA17FBB36FB7C" bold="true" box="[151,310,1075,1102]" italics="true" pageId="17" pageNumber="108">Duroca lenta</emphasis>
(
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, reinstated status
@ -110,84 +111,84 @@
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<paragraph id="705C36E29B27FFD2FF3CA137BEB3FACC" blockId="17.[151,1437,1148,1535]" pageId="17" pageNumber="108">
Genomic sequencing of primary type specimen of
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<emphasis box="[756,949,1148,1174]" italics="true" pageId="17" pageNumber="108">Hesperia duroca</emphasis>
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<emphasis id="4297EAF09B27FFD2FD5FA136B9B5FBA4" box="[756,949,1148,1174]" italics="true" pageId="17" pageNumber="108">Hesperia duroca</emphasis>
Pl
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̂tz, 1882 (type locality in
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:
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, NVG-15034C
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MFNB), the type species of
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<emphasis box="[855,940,1185,1210]" italics="true" pageId="17" pageNumber="108">Duroca</emphasis>
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<emphasis id="4297EAF09B27FFD2FCFCA1EAB9ACFB88" box="[855,940,1185,1210]" italics="true" pageId="17" pageNumber="108">Duroca</emphasis>
Grishin, 2019
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(
<figureCitation box="[1128,1198,1184,1210]" captionStart="FIGURE 8" captionStartId="19.[151,250,759,783]" captionTargetBox="[153,1433,185,731]" captionTargetId="figure-21@19.[151,1436,181,735]" captionTargetPageId="19" captionText="FIGURE 8. Nuclear genome tree of selected Moncina: Phlebodes relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Cantha (purple, with species transferred from Phlebodes labeled in dark green), Duroca (violet, with Duroca caraca, comb. nov. and Duroca lenta, stat. nov. labeled in aquamarine and brown, respectively). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864290" httpUri="https://zenodo.org/record/7864290/files/figure.png" pageId="17" pageNumber="108">Fig. 8</figureCitation>
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violet), and
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<emphasis italics="true" pageId="17" pageNumber="108">Lerema lenta</emphasis>
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<emphasis id="4297EAF09B27FFD2FAE3A1EABACFFBEC" italics="true" pageId="17" pageNumber="108">Lerema lenta</emphasis>
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(type locality in
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: S„o Paulo, molecular code NHMUK0247278362 in BMNH) (
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brown), currently a subspecies of
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<emphasis box="[534,647,1256,1282]" italics="true" pageId="17" pageNumber="108">D. duroca</emphasis>
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<emphasis id="4297EAF09B27FFD2FDBDA1A2B887FA30" box="[534,647,1256,1282]" italics="true" pageId="17" pageNumber="108">D. duroca</emphasis>
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, reveals 2.6% (17 bp) difference in their COI barcodes. Coupled with phenotypic differences described by
<bibRefCitation author="Evans, W. H." box="[565,718,1292,1319]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans (1955)</bibRefCitation>
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, such as unmarked wings, smaller size, and more produced and pointed harpe in
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<emphasis box="[344,430,1328,1354]" italics="true" pageId="17" pageNumber="108">L. lenta</emphasis>
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, this genetic differentiation suggests species status for
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<emphasis box="[1056,1204,1328,1354]" italics="true" pageId="17" pageNumber="108">Duroca lenta</emphasis>
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<emphasis id="4297EAF09B27FFD2FB8BA07ABEB4FA78" box="[1056,1204,1328,1354]" italics="true" pageId="17" pageNumber="108">Duroca lenta</emphasis>
(
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)
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,
<emphasis bold="true" pageId="17" pageNumber="108">stat. rest</emphasis>
<emphasis id="4297EAF09B27FFD2FAC1A07BBAC4FA5C" bold="true" pageId="17" pageNumber="108">stat. rest</emphasis>
. Furthermore, sequence comparison confirms that
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<emphasis box="[768,930,1364,1390]" italics="true" pageId="17" pageNumber="108">Lerema elgina</emphasis>
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(type locality in
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<emphasis id="4297EAF09B27FFD2FDD8A032B91CFAA0" box="[627,796,1400,1426]" italics="true" pageId="17" pageNumber="108">Duroca duroca</emphasis>
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. Finally, to ensure correct identification of this species, a
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<typeStatus id="AF5888409B27FFD2FF3CA0D6BAEEFA85" box="[151,238,1437,1463]" pageId="17" pageNumber="108" type="syntype">syntype</typeStatus>
in the USNM collection bearing the following five rectangular labels, 4
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red, others white: [Nova Friburgo, |
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], [Collection | W. Schaus], [
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<taxonomicName id="B7E34D619B27FFD2FDECA08AB89DFAE9" authorityName="Scudder" authorityYear="1872" box="[583,669,1473,1499]" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="genus">Lerema</taxonomicName>
| eglina | type Sch.], [Type | No. 6054 |
<collectingCountry box="[1131,1176,1472,1499]" name="United States of America" pageId="17" pageNumber="108">U.S.</collectingCountry>
<collectingCountry id="08F476729B27FFD2FBC0A08ABE98FAE8" box="[1131,1176,1472,1499]" name="United States of America" pageId="17" pageNumber="108">U.S.</collectingCountry>
N.M.], [USNMENT | {QR code} | 00913064] is herein designated as the
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</emphasis>
of
<taxonomicName authority="Schaus, 1902" authorityName="Schaus" authorityYear="1902" box="[877,1198,1508,1534]" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="elgina">
<emphasis box="[877,1039,1508,1534]" italics="true" pageId="17" pageNumber="108">Lerema elgina</emphasis>
<taxonomicName id="B7E34D619B27FFD2FCC6A0AEBEAEFACC" authority="Schaus, 1902" authorityName="Schaus" authorityYear="1902" box="[877,1198,1508,1534]" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="elgina">
<emphasis id="4297EAF09B27FFD2FCC6A0AEBE0FFACC" box="[877,1039,1508,1534]" italics="true" pageId="17" pageNumber="108">Lerema elgina</emphasis>
Schaus, 1902
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.

View file

@ -1,155 +1,156 @@
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<taxonomicName authority="(O. Mielke, 1992)" authorityName="Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin" authorityYear="2023" baseAuthorityName="O. Mielke" baseAuthorityYear="1992" box="[151,538,788,814]" family="Hesperiidae" genus="Duroca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" rank="species" species="caraca" status="new combination">
<emphasis bold="true" box="[151,323,788,814]" italics="true" pageId="17" pageNumber="108">Duroca caraca</emphasis>
<treatment id="F84A87F49B27FFD2FF3CA65FB901FCD1" ID-DOI="http://doi.org/10.5281/zenodo.7864342" ID-GBIF-Taxon="209569563" ID-Zenodo-Dep="7864342" LSID="urn:lsid:plazi:treatment:F84A87F49B27FFD2FF3CA65FB901FCD1" httpUri="http://treatment.plazi.org/id/F84A87F49B27FFD2FF3CA65FB901FCD1" lastPageNumber="108" pageId="17" pageNumber="108" scope_family="Hesperiidae" scope_order="Lepidoptera" scope_subTribe="Moncina">
<subSubSection id="38F965699B27FFD2FF3CA65FB8F3FC1C" box="[151,755,788,814]" pageId="17" pageNumber="108" type="nomenclature">
<paragraph id="705C36E29B27FFD2FF3CA65FB8F3FC1C" blockId="17.[151,1437,788,995]" box="[151,755,788,814]" pageId="17" pageNumber="108">
<heading id="2B14818E9B27FFD2FF3CA65FB8F3FC1C" bold="true" box="[151,755,788,814]" fontSize="11" level="1" pageId="17" pageNumber="108" reason="1">
<emphasis id="4297EAF09B27FFD2FF3CA65FB8F3FC1C" bold="true" box="[151,755,788,814]" pageId="17" pageNumber="108">
<taxonomicName id="B7E34D619B27FFD2FF3CA65FB81AFC1C" authority="(O. Mielke, 1992)" authorityName="Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin" authorityYear="2023" baseAuthorityName="O. Mielke" baseAuthorityYear="1992" box="[151,538,788,814]" class="Insecta" family="Hesperiidae" genus="Duroca" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="caraca" status="comb. nov.">
<emphasis id="4297EAF09B27FFD2FF3CA65FBB43FC1C" bold="true" box="[151,323,788,814]" italics="true" pageId="17" pageNumber="108">Duroca caraca</emphasis>
(O. Mielke, 1992)
</taxonomicName>
,
<taxonomicNameLabel box="[551,755,788,814]" pageId="17" pageNumber="108" rank="species">new combination</taxonomicNameLabel>
<taxonomicNameLabel id="59A4578B9B27FFD2FD8CA65FB8F3FC1C" box="[551,755,788,814]" pageId="17" pageNumber="108" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="108" type="description">
<paragraph blockId="17.[151,1437,788,995]" pageId="17" pageNumber="108">
<subSubSection id="38F965699B27FFD2FF3CA673B901FCD1" pageId="17" pageNumber="108" type="description">
<paragraph id="705C36E29B27FFD2FF3CA673B901FCD1" blockId="17.[151,1437,788,995]" pageId="17" pageNumber="108">
Sequencing of the
<typeStatus box="[361,459,824,850]" pageId="17" pageNumber="108">holotype</typeStatus>
<typeStatus id="AF5888409B27FFD2FEC2A673BBCBFC60" box="[361,459,824,850]" pageId="17" pageNumber="108" type="holotype">holotype</typeStatus>
and
<typeStatus box="[513,603,824,850]" pageId="17" pageNumber="108">allotype</typeStatus>
<typeStatus id="AF5888409B27FFD2FDAAA673B85BFC60" box="[513,603,824,850]" pageId="17" pageNumber="108" type="allotype">allotype</typeStatus>
of
<taxonomicName authority="O. Mielke, 1992" authorityName="O. Mielke" authorityYear="1992" box="[641,1002,824,851]" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="caraca">
<emphasis box="[641,811,825,850]" italics="true" pageId="17" pageNumber="108">Lerema caraca</emphasis>
<taxonomicName id="B7E34D619B27FFD2FD2AA672B9EAFC60" authority="O. Mielke, 1992" authorityName="O. Mielke" authorityYear="1992" box="[641,1002,824,851]" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="caraca">
<emphasis id="4297EAF09B27FFD2FD2AA672B92BFC60" box="[641,811,825,850]" italics="true" pageId="17" pageNumber="108">Lerema caraca</emphasis>
O. Mielke, 1992
</taxonomicName>
(type locality in
<collectingCountry box="[1193,1266,824,850]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry>
<collectingCountry id="08F476729B27FFD2FB02A673BEF2FC60" box="[1193,1266,824,850]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry>
:
<collectingRegion box="[1277,1429,824,850]" country="Brazil" name="Minas Gerais" pageId="17" pageNumber="108">Minas Gerais</collectingRegion>
<collectingRegion id="B227F8009B27FFD2FB56A673BF95FC60" box="[1277,1429,824,850]" country="Brazil" name="Minas Gerais" pageId="17" pageNumber="108">Minas Gerais</collectingRegion>
) (
<figureCitation box="[159,226,860,886]" captionStart="FIGURE 8" captionStartId="19.[151,250,759,783]" captionTargetBox="[153,1433,185,731]" captionTargetId="figure-21@19.[151,1436,181,735]" captionTargetPageId="19" captionText="FIGURE 8. Nuclear genome tree of selected Moncina: Phlebodes relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Cantha (purple, with species transferred from Phlebodes labeled in dark green), Duroca (violet, with Duroca caraca, comb. nov. and Duroca lenta, stat. nov. labeled in aquamarine and brown, respectively). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864290" httpUri="https://zenodo.org/record/7864290/files/figure.png" pageId="17" pageNumber="108">Fig. 8</figureCitation>
<figureCitation id="E8D82A679B27FFD2FF34A617BAE2FC44" box="[159,226,860,886]" captionStart="FIGURE 8" captionStartId="19.[151,250,759,783]" captionTargetBox="[153,1433,185,731]" captionTargetId="figure-21@19.[151,1436,181,735]" captionTargetPageId="19" captionText="FIGURE 8. Nuclear genome tree of selected Moncina: Phlebodes relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Cantha (purple, with species transferred from Phlebodes labeled in dark green), Duroca (violet, with Duroca caraca, comb. nov. and Duroca lenta, stat. nov. labeled in aquamarine and brown, respectively). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864290" httpUri="https://zenodo.org/record/7864290/files/figure.png" pageId="17" pageNumber="108">Fig. 8</figureCitation>
aquamarine) reveals its close relationship with
<taxonomicName authority="Pl" authorityName="Pl" box="[763,985,860,888]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="duroca">
<emphasis box="[763,950,860,886]" italics="true" pageId="17" pageNumber="108">Hesperia duroca</emphasis>
<taxonomicName id="B7E34D619B27FFD2FD50A616B9D9FC4A" authority="Pl" authorityName="Pl" box="[763,985,860,888]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="duroca">
<emphasis id="4297EAF09B27FFD2FD50A616B9B6FC44" box="[763,950,860,886]" italics="true" pageId="17" pageNumber="108">Hesperia duroca</emphasis>
Pl
</taxonomicName>
̂tz, 1882 (type locality in
<collectingCountry box="[1278,1351,860,886]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry>
<collectingCountry id="08F476729B27FFD2FB55A617BF47FC44" box="[1278,1351,860,886]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry>
:
<collectingRegion country="Brazil" name="Rio de Janeiro" pageId="17" pageNumber="108">Rio de Janeiro</collectingRegion>
<collectingRegion id="B227F8009B27FFD2FAF9A617BAECFCA8" country="Brazil" name="Rio de Janeiro" pageId="17" pageNumber="108">Rio de Janeiro</collectingRegion>
), the type species of
<taxonomicName authority="Grishin, 2019" authorityName="Grishin" authorityYear="2019" box="[469,714,896,922]" genus="Duroca" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="108" rank="genus">
<emphasis box="[469,554,897,922]" italics="true" pageId="17" pageNumber="108">Duroca</emphasis>
<taxonomicName id="B7E34D619B27FFD2FE7EA6CAB8CAFCA8" authority="Grishin, 2019" authorityName="Grishin" authorityYear="2019" box="[469,714,896,922]" class="Insecta" family="Hesperiidae" genus="Duroca" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="4297EAF09B27FFD2FE7EA6CAB82AFCA8" box="[469,554,897,922]" italics="true" pageId="17" pageNumber="108">Duroca</emphasis>
Grishin, 2019
</taxonomicName>
(
<figureCitation box="[729,795,896,922]" captionStart="FIGURE 8" captionStartId="19.[151,250,759,783]" captionTargetBox="[153,1433,185,731]" captionTargetId="figure-21@19.[151,1436,181,735]" captionTargetPageId="19" captionText="FIGURE 8. Nuclear genome tree of selected Moncina: Phlebodes relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Cantha (purple, with species transferred from Phlebodes labeled in dark green), Duroca (violet, with Duroca caraca, comb. nov. and Duroca lenta, stat. nov. labeled in aquamarine and brown, respectively). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864290" httpUri="https://zenodo.org/record/7864290/files/figure.png" pageId="17" pageNumber="108">Fig. 8</figureCitation>
<figureCitation id="E8D82A679B27FFD2FD72A6CBB91BFCA8" box="[729,795,896,922]" captionStart="FIGURE 8" captionStartId="19.[151,250,759,783]" captionTargetBox="[153,1433,185,731]" captionTargetId="figure-21@19.[151,1436,181,735]" captionTargetPageId="19" captionText="FIGURE 8. Nuclear genome tree of selected Moncina: Phlebodes relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Cantha (purple, with species transferred from Phlebodes labeled in dark green), Duroca (violet, with Duroca caraca, comb. nov. and Duroca lenta, stat. nov. labeled in aquamarine and brown, respectively). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864290" httpUri="https://zenodo.org/record/7864290/files/figure.png" pageId="17" pageNumber="108">Fig. 8</figureCitation>
violet). The COI barcodes of the type specimens of these species reveal 5.4% (36 bp) difference, which is small enough even for closely related congeners, and therefore we propose
<taxonomicName authority="(O. Mielke, 1992)" authorityName="Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin" authorityYear="2023" baseAuthorityName="O. Mielke" baseAuthorityYear="1992" box="[247,628,968,995]" genus="Duroca" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="108" rank="species" species="caraca" status="comb. nov.">
<emphasis box="[247,416,969,994]" italics="true" pageId="17" pageNumber="108">Duroca caraca</emphasis>
<taxonomicName id="B7E34D619B27FFD2FF5CA682B874FCD0" authority="(O. Mielke, 1992)" authorityName="Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin" authorityYear="2023" baseAuthorityName="O. Mielke" baseAuthorityYear="1992" box="[247,628,968,995]" class="Insecta" family="Hesperiidae" genus="Duroca" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="caraca" status="comb. nov.">
<emphasis id="4297EAF09B27FFD2FF5CA682BBA0FCD0" box="[247,416,969,994]" italics="true" pageId="17" pageNumber="108">Duroca caraca</emphasis>
(O. Mielke, 1992)
</taxonomicName>
,
<taxonomicNameLabel box="[639,769,968,995]" pageId="17" pageNumber="108" rank="species">
<emphasis bold="true" box="[639,763,968,994]" pageId="17" pageNumber="108">comb. nov</emphasis>
<taxonomicNameLabel id="59A4578B9B27FFD2FDD4A683B901FCD1" box="[639,769,968,995]" pageId="17" pageNumber="108" rank="species">
<emphasis id="4297EAF09B27FFD2FDD4A683B8FBFCD0" bold="true" box="[639,763,968,994]" pageId="17" pageNumber="108">comb. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>