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<mods:titleInfo> <mods:titleInfo id="2395D094FD08DB16E675C667B8CFE280">
<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title> <mods:title id="34459F93891F9C0B18430BC8A08E140E">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:namePart>Zamani, Alireza</mods:namePart> <mods:namePart id="5DCAEDDD51F55C0230567504371D58B6">Zamani, Alireza</mods:namePart>
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<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation> <mods:affiliation id="BD17B663E76AA73150AA05A44B39CB55">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
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<mods:namePart>Marusik, Yuri M.</mods:namePart> <mods:namePart id="29C03CB1D9B6A22C936F7E24453615C9">Marusik, Yuri M.</mods:namePart>
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation> <mods:affiliation id="B5650E5E3C067A3AF4CA6EA206615630">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<treatment ID-GBIF-Taxon="181753573" LSID="urn:lsid:plazi:treatment:02249EDF73C059D1843C3DE96E998ED5" httpUri="http://treatment.plazi.org/id/02249EDF73C059D1843C3DE96E998ED5" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="02249EDF73C059D1843C3DE96E998ED5" ID-GBIF-Taxon="181753573" LSID="urn:lsid:plazi:treatment:02249EDF73C059D1843C3DE96E998ED5" httpUri="http://treatment.plazi.org/id/02249EDF73C059D1843C3DE96E998ED5" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="20827C5608BC70AFE03991121EA42FBB" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="F57323D5F4BF387FAEB4E4A1F52DCF9B" pageId="0" pageNumber="145">
Genus Genus
<taxonomicName LSID="02249EDF-73C0-59D1-843C-3DE96E998ED5" authority="Walckenaer, 1826" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Zodarion Walckenaer, 1826</taxonomicName> <taxonomicName id="47A03107E1710609DCE974C709322D57" ID-CoL="8B2B" LSID="02249EDF-73C0-59D1-843C-3DE96E998ED5" authority="Walckenaer, 1826" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Zodarion Walckenaer, 1826</taxonomicName>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type species"> <subSubSection id="0C756A4F82E62D458828E26411AEA910" pageId="0" pageNumber="145" type="type species">
<paragraph pageId="0" pageNumber="145">Type species.</paragraph> <paragraph id="AA3D977C13206B9747DA3EFED6760182" pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="7F8D3C2D3C5A9A833515AD0FAD217570" pageId="0" pageNumber="145">
<taxonomicName class="Arachnida" family="Zodariidae" genus="Enyo" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Enyo nitida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitida"> <taxonomicName id="115C472EE2D87AE4E48D6A38CFE76810" class="Arachnida" family="Zodariidae" genus="Enyo" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Enyo nitida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitida">
<emphasis italics="true" pageId="0" pageNumber="145">Enyo nitida</emphasis> <emphasis id="117FDD52C239BE7B18D49F660B263B6A" italics="true" pageId="0" pageNumber="145">Enyo nitida</emphasis>
</taxonomicName> </taxonomicName>
Audouin, 1826 from Egypt. Audouin, 1826 from Egypt.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comments"> <subSubSection id="FECE839F907964B1787D21132432E37C" pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="C64B8E8ED26644473A6B18383A27FA40" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="770B3FA50E3FE105D288BF2C86B81558" pageId="0" pageNumber="145">
With 177 named species, this is the largest genus within With 177 named species, this is the largest genus within
<taxonomicName authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName> <taxonomicName id="DFBEA76ADD13A99871546F92434385E4" authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName>
( (
<bibRefCitation author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC 2021</bibRefCitation> <bibRefCitation id="C05AB38B7C6D7AF1D0F69EE51548D4D8" author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC 2021</bibRefCitation>
). Based on the copulatory organs of the species currently considered in this genus, ). Based on the copulatory organs of the species currently considered in this genus,
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="B3FA8F757BAE53D3B79F767ECACF0022" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis> <emphasis id="515E6BA3F5881C9037F4ADCF4569B76B" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName> </taxonomicName>
does not appear to be monophyletic. There are only two species that are morphologically similar to the generotype: does not appear to be monophyletic. There are only two species that are morphologically similar to the generotype:
<taxonomicName lsidName="Z. luctuosum" pageId="0" pageNumber="145" rank="species" species="luctuosum"> <taxonomicName id="DC0AE2A849AC5F3D8C255AB65F217583" lsidName="Z. luctuosum" pageId="0" pageNumber="145" rank="species" species="luctuosum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. luctuosum</emphasis> <emphasis id="F48A6AB60F66F3BBE2A2DEB115D5537E" italics="true" pageId="0" pageNumber="145">Z. luctuosum</emphasis>
</taxonomicName> </taxonomicName>
(O. Pickard-Cambridge, 1872) and (O. Pickard-Cambridge, 1872) and
<taxonomicName lsidName="Z. lutipes" pageId="0" pageNumber="145" rank="species" species="lutipes"> <taxonomicName id="0A42897BDB14512B2D3DDC403E6EEB0D" lsidName="Z. lutipes" pageId="0" pageNumber="145" rank="species" species="lutipes">
<emphasis italics="true" pageId="0" pageNumber="145">Z. lutipes</emphasis> <emphasis id="49C06D8660EA049914C72E95F23C302F" italics="true" pageId="0" pageNumber="145">Z. lutipes</emphasis>
</taxonomicName> </taxonomicName>
(O. Pickard-Cambridge, 1872), from the eastern Mediterranean (east of Tunisia to Iran). (O. Pickard-Cambridge, 1872), from the eastern Mediterranean (east of Tunisia to Iran).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="materials_examined"> <subSubSection id="AFC0DECB1E432B5AC42F55A6D1EF941C" pageId="0" pageNumber="145" type="materials_examined">
<paragraph pageId="0" pageNumber="145">Comparative material.</paragraph> <paragraph id="FDA07F83F6FE446D984509F47C3CDDC8" pageId="0" pageNumber="145">Comparative material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="9E9F4963AA4FB08881BFC4A3E8A72F4E" pageId="0" pageNumber="145">
<taxonomicName class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion expers" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="expers"> <taxonomicName id="3B29D01F09ED3D6BA6BBB59E99944192" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion expers" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="expers">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion expers</emphasis> <emphasis id="D1292117F21BA97FB2AF0697ABDC4782" italics="true" pageId="0" pageNumber="145">Zodarion expers</emphasis>
</taxonomicName> </taxonomicName>
(O. Pickard-Cambridge, 1876), (O. Pickard-Cambridge, 1876),
<materialsCitation ID-GBIF-Occurrence="3093634307" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Israel" location="Israel" specimenCount="2" specimenCount-male="2"> <materialsCitation id="4D435058BD88808C86C253D33C016488" ID-GBIF-Occurrence="3093634307" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Israel" location="Israel" specimenCount="2" specimenCount-male="2">
<specimenCount type="male">1♂</specimenCount> <specimenCount id="5071CE3BD50261CB86F0EF22CA7BDAD9" type="male">1♂</specimenCount>
from from
<collectingCountry name="Israel">Israel</collectingCountry> <collectingCountry id="0DD4215C26A66F76E65A44991DB195A6" name="Israel">Israel</collectingCountry>
(Figs (Figs
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19D-F</figureCitation> <figureCitation id="C0FD670EA72F7D67F5A8E954F92A0F02" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19D-F</figureCitation>
, ,
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. SEM images of the male palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, B, D ventral C, E retrolateral F dorsoretrolateral. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis, Ra - retrolateral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145">25D-F</figureCitation> <figureCitation id="C66B6858C629A09A41F52678942D56D4" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. SEM images of the male palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, B, D ventral C, E retrolateral F dorsoretrolateral. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis, Ra - retrolateral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145">25D-F</figureCitation>
) and ) and
<taxonomicName baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion nitidum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitidum"> <taxonomicName id="7146295478D7D1F6E9244C53097055C6" baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion nitidum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitidum">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion nitidum</emphasis> <emphasis id="0E8CC7027D40D607B0E85F92C8447757" italics="true" pageId="0" pageNumber="145">Zodarion nitidum</emphasis>
</taxonomicName> </taxonomicName>
(Audouin, 1826): (Audouin, 1826):
<specimenCount type="male">1♂</specimenCount> <specimenCount id="155930A6EE02471FB027C119F53F5F27" type="male">1♂</specimenCount>
from from
<collectingCountry name="Israel">Israel</collectingCountry> <collectingCountry id="8B71DF92A1B97C49683FFE5DF75A1EF7" name="Israel">Israel</collectingCountry>
(Fig. (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. SEM images of the male palps of Zodarion nitidum (A-C) and Z. bactrianum (D-F) A, D ventral B, E retrolateral C dorsal F retrodorsal. Abbreviations: Co - conductor, Eb - embolar base, Em - embolus, Ma - median apophysis, Mc - claw of median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process, Tu - tutaculum, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure28" httpUri="https://binary.pensoft.net/fig/536905" pageId="0" pageNumber="145">28A-C</figureCitation> <figureCitation id="7B50D0951E658D5C0DBE12A183D27F8F" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. SEM images of the male palps of Zodarion nitidum (A-C) and Z. bactrianum (D-F) A, D ventral B, E retrolateral C dorsal F retrodorsal. Abbreviations: Co - conductor, Eb - embolar base, Em - embolus, Ma - median apophysis, Mc - claw of median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process, Tu - tutaculum, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure28" httpUri="https://binary.pensoft.net/fig/536905" pageId="0" pageNumber="145">28A-C</figureCitation>
) )
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</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="comments"> <subSubSection id="0FADA4FBE19B27EE4A69AFF650F35BD3" pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="164E39919DF6E328EF23608B29B61DAF" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="8C34E0AB325456F06198E651D00EBDFC" pageId="0" pageNumber="145">
Until recently, Until recently,
<taxonomicName lsidName="Z. expers" pageId="0" pageNumber="145" rank="species" species="expers"> <taxonomicName id="BB865E18A2720E571E982147199500BE" lsidName="Z. expers" pageId="0" pageNumber="145" rank="species" species="expers">
<emphasis italics="true" pageId="0" pageNumber="145">Z. expers</emphasis> <emphasis id="C71D61FAE2379104D1B2F2C9B85CED09" italics="true" pageId="0" pageNumber="145">Z. expers</emphasis>
</taxonomicName> </taxonomicName>
was placed in was placed in
<taxonomicName authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Ranops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ranops" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="A6858C0602702035FA45B5A15E759F5D" authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Ranops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ranops" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Ranops</emphasis> <emphasis id="91EE6D153201F2F4EB13F397B9C8CB33" italics="true" pageId="0" pageNumber="145">Ranops</emphasis>
</taxonomicName> </taxonomicName>
<normalizedToken originalValue="Jocqué">Jocque</normalizedToken> <normalizedToken id="C848D4BC653BD01B5E7FFAE7BE307045" originalValue="Jocqué">Jocque</normalizedToken>
, 1991, but now it is placed in , 1991, but now it is placed in
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="1C7D7790739EBA24233509A649D2BFA2" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis> <emphasis id="E9327B15B34E680F8EBDB863DC117FD1" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName> </taxonomicName>
. It differs from the generotype of . It differs from the generotype of
<taxonomicName authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Ranops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ranops" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="C67344BB633707FF4126777440A68D12" authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Ranops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ranops" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Ranops</emphasis> <emphasis id="78F8CB800925D0FC93C169CE540C2C0A" italics="true" pageId="0" pageNumber="145">Ranops</emphasis>
</taxonomicName> </taxonomicName>
, but the copulatory organs are also very different from those of , but the copulatory organs are also very different from those of
<taxonomicName lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum"> <taxonomicName id="24BFF4C6E6F2596F0726A0F138E158B5" lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis> <emphasis id="DDB00CC34670707472BD2151C274A103" italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
</taxonomicName> </taxonomicName>
, and most likely it represents a separate genus. Although it is absent in Iran or Turkmenistan, we have provided figures of this species to illustrate the differences with the generotype of , and most likely it represents a separate genus. Although it is absent in Iran or Turkmenistan, we have provided figures of this species to illustrate the differences with the generotype of
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="90E2A520B9CCACA16748ADC34DDD3C31" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis> <emphasis id="B4A13C221605FAB2545AE40877D8F3D9" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName> </taxonomicName>
. Furthermore, although . Furthermore, although
<taxonomicName lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum"> <taxonomicName id="0836D45646F457A4FA4B25B0DE5823C6" lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis> <emphasis id="E009DD5F9048FB07135881C4BF4DD618" italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
</taxonomicName> </taxonomicName>
has not been recorded in the current study area, because it is the type species of the genus, we have provided illustartions for this species as well, to demonstrate its differences with other has not been recorded in the current study area, because it is the type species of the genus, we have provided illustartions for this species as well, to demonstrate its differences with other
<taxonomicName authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName> <taxonomicName id="ADED0C1D71FB722A1EB3020E9E7265B0" authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName>
genera in Iran and Turkmenistan, as well as with other species currently placed in genera in Iran and Turkmenistan, as well as with other species currently placed in
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="E1B59239019B906425258E8D7E3FB10B" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis> <emphasis id="CF42DDAA6AC98587437FB890BF576FD4" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName> </taxonomicName>
. .
</paragraph> </paragraph>

264
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@ -1,229 +1,229 @@
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<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="95FDE61E3CAE8BD7840F68654D255665" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">Figs 5C, D</figureCitation> <figureCitation id="5A2C9A11B4F60C5FCF176709C3DC9950" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">Figs 5C, D</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">, 8</figureCitation> <figureCitation id="A1D4B4C4F1ADB760BFACBDC250ABBE47" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">, 8</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">, 9</figureCitation> <figureCitation id="5DFA95A2A776C7D0F6DAA863ED8DA71F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">, 9</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">, 10F-J</figureCitation> <figureCitation id="739E51E13DAD97771AC0B89180FB47B8" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">, 10F-J</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation> <figureCitation id="6E481C54DBF5EFEEFC3EFFFEB5E9FFD0" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="FA777627746326AA3AA1688C50FB90C2" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="C070B08018403A6BCB6B0F55E791C79A" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="0D2D9333DCC2EE79B57E480508A4DE18" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="F45790FB34322E8B205E86073A8E878D" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ and ♂ and
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis> <emphasis id="6CAB6126CA078587D15355101B173BC3" bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis>
1♀ (MHNG), Iran: 1♀ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Fars Province</emphasis> <emphasis id="76BA2AF8D2599E86CE055840E33F8B70" italics="true" pageId="0" pageNumber="145">Fars Province</emphasis>
: road to Yasuj, : road to Yasuj,
<geoCoordinate degrees="30" direction="north" minutes="28" orientation="latitude" precision="925" value="30.466667">30°28'N</geoCoordinate> <geoCoordinate id="B54ABE06A265D8118A97710001BA370B" degrees="30" direction="north" minutes="28" orientation="latitude" precision="925" value="30.466667">30°28'N</geoCoordinate>
, ,
<geoCoordinate degrees="51" direction="east" minutes="30" orientation="longitude" precision="925" value="51.5">51°30'E</geoCoordinate> <geoCoordinate id="011833FB1319D5E8D350FC766577CD85" degrees="51" direction="east" minutes="30" orientation="longitude" precision="925" value="51.5">51°30'E</geoCoordinate>
, 25.05.1974 (A. Senglet). , 25.05.1974 (A. Senglet).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="4ED5441780D1374CB0C48EF52CA86BDB" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="D51D30BA473B006B9373EC1F5A99CE7A" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet is a feminine given name in the Persian language, meaning &quot;daughter of the night&quot;. Noun in apposition.</paragraph> <paragraph id="4E51DEEBF696DA81FE5F3D1F1F15AF38" pageId="0" pageNumber="145">The specific epithet is a feminine given name in the Persian language, meaning &quot;daughter of the night&quot;. Noun in apposition.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="4C6E036F3BB0C13E9083C2750C2F436D" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="D01350BE9F66B49B71DF8D83E107406A" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="C84CD4E99DE8B4FB544C491CA84F5848" pageId="0" pageNumber="145">
The new species differs from all congeners by having a pair of spine-like cymbial outgrowths (Figs The new species differs from all congeners by having a pair of spine-like cymbial outgrowths (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">8</figureCitation> <figureCitation id="98B655D74E64C078F1168E438F370CA2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">8</figureCitation>
, ,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">9C, E</figureCitation> <figureCitation id="652433934DEE6D723C2CD5E44A2EF15D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">9C, E</figureCitation>
) ( ) (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="CF42DCA06FA17C8D0709A0AB601ADBE7" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
lacking) and a triangular epigynal median plate ( lacking) and a triangular epigynal median plate (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="27F6AC7D2B5202B236A7BE366003E581" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
absent or rectangular). absent or rectangular).
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145" start="Figure 8" startId="F8"> <caption id="C448ADE7E1135D19E0E32EA8E1C922C2" doi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="145"> <paragraph id="D37620CFE35199C30125B543D8469984" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 8.</emphasis> <emphasis id="7EAE1F8E57B854F60C4CED2234D6CD54" bold="true" pageId="0" pageNumber="145">Figure 8.</emphasis>
Male palp of Male palp of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila"> <taxonomicName id="9C4DADAE36CA6C14EEB7798770C346F6" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila">
<emphasis italics="true" pageId="0" pageNumber="145">Pax leila</emphasis> <emphasis id="0B305F4FD2CE5A844A834C54087FBC90" italics="true" pageId="0" pageNumber="145">Pax leila</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis> <emphasis id="52AF5D2BF2EF266E2235D2B4A4480CE5" bold="true" pageId="0" pageNumber="145">A</emphasis>
proventral proventral
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis> <emphasis id="165B7CC20AE8F37D1DFF9F13EC7F596D" bold="true" pageId="0" pageNumber="145">B</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis> <emphasis id="F3B1907D8EC0DAC76B1DC8ED6BCD3BBF" bold="true" pageId="0" pageNumber="145">C</emphasis>
ventroretrolateral ventroretrolateral
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis> <emphasis id="CDFDB57750CA1A33167FE39424DA8B99" bold="true" pageId="0" pageNumber="145">D</emphasis>
prolateral prolateral
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis> <emphasis id="4DC1B840D415D6338D0ADE753EC6C5CE" bold="true" pageId="0" pageNumber="145">E</emphasis>
dorsal dorsal
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis> <emphasis id="746417865834C56F4B14565D44FF79E5" bold="true" pageId="0" pageNumber="145">F</emphasis>
retrolateral. Scale bars: 0.2 mm, unless stated otherwise. retrolateral. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph> </paragraph>
</caption> </caption>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="331289FAAEE58BC5B91C32D445903BF1" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="C5369750918E42EC43D2ECB09A88FAEE" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="644A209423ED7521A6DAAB2F56708660" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis> <emphasis id="2806484CF0771CBE32C9FD1F9EEBFFFB" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Habitus as in Fig. Habitus as in Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5C</figureCitation> <figureCitation id="8B110704F23D11AA905653FDBBFBF08B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5C</figureCitation>
. Total length 4.42. Carapace 2.07 long, 1.03 wide at pars cephalica, 1.46 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.09, ALE: 0.09, PME: 0.06, PLE: 0.09, PME-PME: 0.07. Carapace, sternum, labium, chelicerae and maxillae reddish brown, without any pattern. Chelicera with 2 promarginal teeth. Legs yellow, with few spines and without annulations. Abdomen dark brown, dorsally with large scutum covering 2/3 of the abdomen. Spinnerets pale, uniform in color. Measurements of legs: I: 4.51 (1.21, 0.50, 1.06, 0.93, 0.81), II: 3.89 (1.04, 0.52, 0.79, 0.83, 0.71), III: 3.62 (0.98, 0.50, 0.69, 0.85, 0.60), IV: 4.99 (1.33, 0.58, 1.05, 1.33, 0.70). . Total length 4.42. Carapace 2.07 long, 1.03 wide at pars cephalica, 1.46 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.09, ALE: 0.09, PME: 0.06, PLE: 0.09, PME-PME: 0.07. Carapace, sternum, labium, chelicerae and maxillae reddish brown, without any pattern. Chelicera with 2 promarginal teeth. Legs yellow, with few spines and without annulations. Abdomen dark brown, dorsally with large scutum covering 2/3 of the abdomen. Spinnerets pale, uniform in color. Measurements of legs: I: 4.51 (1.21, 0.50, 1.06, 0.93, 0.81), II: 3.89 (1.04, 0.52, 0.79, 0.83, 0.71), III: 3.62 (0.98, 0.50, 0.69, 0.85, 0.60), IV: 4.99 (1.33, 0.58, 1.05, 1.33, 0.70).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="19CADD542F51ABE823B88F014581B96F" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">8A-F</figureCitation> <figureCitation id="45BF8077FE0548D1BCA6D47FA3F42655" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">8A-F</figureCitation>
, ,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">9A-E</figureCitation> <figureCitation id="35F512F44E81CEA4B087B0280C75F515" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">9A-E</figureCitation>
. Tibia wider than long, with short apophysis; cymbium with 2 strong spine-like baso-posterior outgrowths ( . Tibia wider than long, with short apophysis; cymbium with 2 strong spine-like baso-posterior outgrowths (
<emphasis italics="true" pageId="0" pageNumber="145">Cp</emphasis> <emphasis id="B11777083988408CD5AAECF53B9943C0" italics="true" pageId="0" pageNumber="145">Cp</emphasis>
, ,
<emphasis italics="true" pageId="0" pageNumber="145">Cr</emphasis> <emphasis id="EC1C8CBEA463F34317A5E9DE75B3F760" italics="true" pageId="0" pageNumber="145">Cr</emphasis>
), lateral fold lacking; embolus long, with robust triangular outgrowth near base ( ), lateral fold lacking; embolus long, with robust triangular outgrowth near base (
<emphasis italics="true" pageId="0" pageNumber="145">Se</emphasis> <emphasis id="1D1AB9F10151F8EA863238A1BE6557F0" italics="true" pageId="0" pageNumber="145">Se</emphasis>
); median apophysis ( ); median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="2F34EE9F12F3E8BFD1282E2BCCC25080" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) very large, with 2 claw-like extensions; conductor small and strongly sclerotized. ) very large, with 2 claw-like extensions; conductor small and strongly sclerotized.
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145" start="Figure 9" startId="F9"> <caption id="AB5F3B3E382925E0FFCED6505E966E61" doi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="145"> <paragraph id="D9F0744620903DD21D67AC88FC4BFC9C" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 9.</emphasis> <emphasis id="49DB0536F2B6BFE2349C8BF91C60E2DC" bold="true" pageId="0" pageNumber="145">Figure 9.</emphasis>
SEM images of the male palp of SEM images of the male palp of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila"> <taxonomicName id="A54BA2C6573765749463CBD9A0F67663" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila">
<emphasis italics="true" pageId="0" pageNumber="145">Pax leila</emphasis> <emphasis id="4F50C05BAE0A78165AD08B978857EC93" italics="true" pageId="0" pageNumber="145">Pax leila</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis> <emphasis id="3F9916DBB667F91B49C837670A83B4D6" bold="true" pageId="0" pageNumber="145">A</emphasis>
apicoventral apicoventral
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis> <emphasis id="4E30F043477EF83D7E191B75F426B0FB" bold="true" pageId="0" pageNumber="145">B</emphasis>
ventroretrolateral ventroretrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis> <emphasis id="E0CE82CAE995D85A0A75C36858801F9A" bold="true" pageId="0" pageNumber="145">C</emphasis>
prodorsal prodorsal
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis> <emphasis id="20F1FE821B4DBDEC8DF54D356D1F1EBB" bold="true" pageId="0" pageNumber="145">D</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis> <emphasis id="FF5B27286CDDF0ADC258D0EEF7C58D19" bold="true" pageId="0" pageNumber="145">E</emphasis>
retrodorsal. Abbreviations: retrodorsal. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Cp</emphasis> <emphasis id="A079B523888F130DC7BF015E41C13A45" italics="true" pageId="0" pageNumber="145">Cp</emphasis>
- prolateral cymbial process, - prolateral cymbial process,
<emphasis italics="true" pageId="0" pageNumber="145">Cr</emphasis> <emphasis id="DEE48351EB109074B5381A71F561E781" italics="true" pageId="0" pageNumber="145">Cr</emphasis>
- retrolateral cymbial process, - retrolateral cymbial process,
<emphasis italics="true" pageId="0" pageNumber="145">Da</emphasis> <emphasis id="579CEBD106D87CCC235830E4CB563A0A" italics="true" pageId="0" pageNumber="145">Da</emphasis>
- dorsal apophysis, - dorsal apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="711B0A4EED352E17F7A04EB4013B71A3" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis, - median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Se</emphasis> <emphasis id="3352366A9EC96F193065E824CDA97085" italics="true" pageId="0" pageNumber="145">Se</emphasis>
- spine of embolus base. Scale bars: 0.1 mm. - spine of embolus base. Scale bars: 0.1 mm.
</paragraph> </paragraph>
</caption> </caption>
<paragraph pageId="0" pageNumber="145"> <paragraph id="942FED49D47F484ED2761D73697F641B" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="114C8D63694102AF62B9ED89D14E282B" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig. Habitus as in Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5D</figureCitation> <figureCitation id="0CCFBC148C0036178772F63F1509504A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5D</figureCitation>
. Total length 4.95. Carapace 1.97 long, 1.00 wide at pars cephalica, 1.31 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.07, ALE: 0.08, PME: 0.06, PLE: 0.07, PME-PME: 0.08. Coloration as in male, with lighter abdomen lacking a scutum. Measurements of legs: I: 4.25 (1.21, 0.50, 0.99, 0.84, 0.71), II: 3.58 (0.94, 0.52, 0.74, 0.74, 0.64), III: 3.47 (0.90, 0.51, 0.70, 0.81, 0.55), IV: 4.71 (1.18, 0.53, 1.05, 1.21, 0.74). . Total length 4.95. Carapace 1.97 long, 1.00 wide at pars cephalica, 1.31 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.07, ALE: 0.08, PME: 0.06, PLE: 0.07, PME-PME: 0.08. Coloration as in male, with lighter abdomen lacking a scutum. Measurements of legs: I: 4.25 (1.21, 0.50, 0.99, 0.84, 0.71), II: 3.58 (0.94, 0.52, 0.74, 0.74, 0.64), III: 3.47 (0.90, 0.51, 0.70, 0.81, 0.55), IV: 4.71 (1.18, 0.53, 1.05, 1.21, 0.74).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="68AA08A588CA6B32860A9EF6E7699DC2" pageId="0" pageNumber="145">
Epigyne as in Fig. Epigyne as in Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">10F-J</figureCitation> <figureCitation id="48C364A8470D174C535E92B421B88A45" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">10F-J</figureCitation>
. Epigyne trilobate, with triangular median plate, posteriorly as wide as lateral lobes; copulatory openings slit-like, broad; copulatory ducts wide, as wide as receptacles; receptacles globular, separated by less than their radii. . Epigyne trilobate, with triangular median plate, posteriorly as wide as lateral lobes; copulatory openings slit-like, broad; copulatory ducts wide, as wide as receptacles; receptacles globular, separated by less than their radii.
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145" start="Figure 10" startId="F10"> <caption id="37185AAEE15756DCCB2FA4E70BC21A99" doi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="145"> <paragraph id="B9B355F792377E7AD6083EF71BD87A19" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 10.</emphasis> <emphasis id="B1B4D3E39976DE0177B82AB01AE26C27" bold="true" pageId="0" pageNumber="145">Figure 10.</emphasis>
Epigynes of Epigynes of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita"> <taxonomicName id="2FE37A265797936D5130B198CF4CC489" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis> <emphasis id="C2DDF008B5B86FA9F0F6431782F1F901" italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-E</emphasis> <emphasis id="459C89D9239C9FF21C7A2C6A02FED11A" bold="true" pageId="0" pageNumber="145">A-E</emphasis>
) and ) and
<taxonomicName lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila"> <taxonomicName id="F7A1C8605D9F21BA15C06FFF13FC46B1" lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila">
<emphasis italics="true" pageId="0" pageNumber="145">P. leila</emphasis> <emphasis id="2246B5D492F3C8E7D3AE0D0323B509C5" italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">F-J</emphasis> <emphasis id="EF9BB29F2C2203253F286F8A4DFABC94" bold="true" pageId="0" pageNumber="145">F-J</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, F</emphasis> <emphasis id="0D874B34DDA34980B6BB3BA459935F4A" bold="true" pageId="0" pageNumber="145">A, F</emphasis>
intact, ventral intact, ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, J</emphasis> <emphasis id="7DDF6F8A2F76A296832DA3B079272E3B" bold="true" pageId="0" pageNumber="145">B, J</emphasis>
posterior posterior
<emphasis bold="true" pageId="0" pageNumber="145">C, G</emphasis> <emphasis id="9D4A03E34AC3F83D104F4EB2CC10A63C" bold="true" pageId="0" pageNumber="145">C, G</emphasis>
macerated, ventral macerated, ventral
<emphasis bold="true" pageId="0" pageNumber="145">D, E, H, I</emphasis> <emphasis id="8CEA8F5430C18508630D9ACD4608DC17" bold="true" pageId="0" pageNumber="145">D, E, H, I</emphasis>
macerated, dorsal. Scale bars: 0.2 mm. macerated, dorsal. Scale bars: 0.2 mm.
</paragraph> </paragraph>
</caption> </caption>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comments"> <subSubSection id="E2BF68B596F34C6894A43572B0168470" pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="2785376854BC6704A1D632D63FF70B48" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="E36EC4EA81645698AD413ECE8BA6A072" pageId="0" pageNumber="145">
Although this species is rather similar in general appearance to the generotype and the other species known from Iran, the bulb and cymbium conformation is very different from those of other species considered in Although this species is rather similar in general appearance to the generotype and the other species known from Iran, the bulb and cymbium conformation is very different from those of other species considered in
<taxonomicName authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="26A98999BC44AEFBEE5E892EC4859534" authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Pax</emphasis> <emphasis id="F084086178C6FAD3DD2BF150E9FCA8AE" italics="true" pageId="0" pageNumber="145">Pax</emphasis>
</taxonomicName> </taxonomicName>
and most likely belongs to an undescribed genus. and most likely belongs to an undescribed genus.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="D5107F62298DDB995517B1CE34077CA2" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="81FDC14758040DF14F4BA504FE4F71D8" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="09AD0C2B35941441BC7654607FCA8E26" pageId="0" pageNumber="145">
Known only from the type locality in Fars Province, southwestern Iran (Fig. Known only from the type locality in Fars Province, southwestern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation> <figureCitation id="394150865ED8F5BDB3ED207BF2F3353E" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). This is the easternmost record of the genus ). This is the easternmost record of the genus
<taxonomicName authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="0F202F09DCB601B369C896E48AFCFD18" authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Pax</emphasis> <emphasis id="E1A395D5243E1516E653F3F45C6B91FB" italics="true" pageId="0" pageNumber="145">Pax</emphasis>
</taxonomicName> </taxonomicName>
. .
</paragraph> </paragraph>

106
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<taxonomicName LSID="0C488F61-89B4-58B2-8A39-7260E487A64B" authority="Marusik, 2011" authorityName="Marusik" authorityYear="2011" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus gromovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="gromovi">Trygetus gromovi Marusik, 2011</taxonomicName> <taxonomicName id="1E010666500E115E68179A7E8EBBE535" ID-CoL="7CXYT" LSID="0C488F61-89B4-58B2-8A39-7260E487A64B" authority="Marusik, 2011" authorityName="Marusik" authorityYear="2011" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus gromovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="gromovi">Trygetus gromovi Marusik, 2011</taxonomicName>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">Fig. 32</figureCitation> <figureCitation id="5F2D2950D4DE034B44B490667480DBC5" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">Fig. 32</figureCitation>
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="reference_group"> <subSubSection id="5668C69EA0AE98BE82B448A9F8D0798F" pageId="0" pageNumber="145" type="reference_group">
<paragraph pageId="0" pageNumber="145"> <paragraph id="6971B16786C1BC6FA5A2DA64665CEDED" pageId="0" pageNumber="145">
<taxonomicName authorityName="Marusik" authorityYear="2011" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus gromovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="gromovi">Trygetus gromovi</taxonomicName> <taxonomicName id="6E655B50FDDC7D13C10D4CB7B6FE958B" authorityName="Marusik" authorityYear="2011" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus gromovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="gromovi">Trygetus gromovi</taxonomicName>
Marusik, 2011: 30, f. 1-7 (♀). Marusik, 2011: 30, f. 1-7 (♀).
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="records in turkmenistan"> <subSubSection id="3074E72F660EC25345CF1C9EEA2F87FE" pageId="0" pageNumber="145" type="records in turkmenistan">
<paragraph pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph> <paragraph id="61ABDF858F43021EFEDF064FC1ABB03A" pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="D72974DC930EDD9ED1E45BBC15FB3BB9" pageId="0" pageNumber="145">
Mary ( Mary (
<bibRefCitation author="Marusik, YM" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="145" pagination="29 - 31" refId="B12" refString="Marusik, YM, 2011. A new Trygetus species from Central Asia (Araneae: Zodariidae). Turkish Journal of Zoology 35 (1): 29 - 31" title="A new Trygetus species from Central Asia (Araneae: Zodariidae)." volume="35" year="2011">Marusik 2011</bibRefCitation> <bibRefCitation id="FB6B2745719C75A97EE377BDEE468756" author="Marusik, YM" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="145" pagination="29 - 31" refId="B12" refString="Marusik, YM, 2011. A new Trygetus species from Central Asia (Araneae: Zodariidae). Turkish Journal of Zoology 35 (1): 29 - 31" title="A new Trygetus species from Central Asia (Araneae: Zodariidae)." volume="35" year="2011">Marusik 2011</bibRefCitation>
) (Fig. ) (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation> <figureCitation id="5D032D22073501C8B091AA6F3B0397A1" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="0266F1A1CABB45A4D4109C03F1AC4661" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="CE753CF9EE233B9E2B1BB959B7B3B655" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">Known only from the type locality in Turkmenistan.</paragraph> <paragraph id="0F8416A2B23B8D32D4024BC8789C6BB9" pageId="0" pageNumber="145">Known only from the type locality in Turkmenistan.</paragraph>
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</treatment> </treatment>
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Subfamily Subfamily
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Lachesaninae Lachesaninae
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, 1991 , 1991
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<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="8FC7885249691573CC3E27FFBF1D239A" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="7B8BAC6945875998522D359F00CDEE21" pageId="0" pageNumber="145">
The following four genera are currently considered in this subfamily, with the number of species known from each in parentheses: The following four genera are currently considered in this subfamily, with the number of species known from each in parentheses:
<taxonomicName authorityName="Jocque" authorityYear="1995" class="Arachnida" family="Zodariidae" genus="Australutica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Australutica" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="9520716391FE27C95A939CDD22E9A562" authorityName="Jocque" authorityYear="1995" class="Arachnida" family="Zodariidae" genus="Australutica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Australutica" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Australutica</emphasis> <emphasis id="581DC5283C641EB91E4429A8506BA164" italics="true" pageId="0" pageNumber="145">Australutica</emphasis>
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, 1995 (6), , 1995 (6),
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<emphasis italics="true" pageId="0" pageNumber="145">Antillorena</emphasis> <emphasis id="1C29E9DD39E3B9022C122A3CF4D4BB48" italics="true" pageId="0" pageNumber="145">Antillorena</emphasis>
</taxonomicName> </taxonomicName>
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, 1991 (4), , 1991 (4),
<taxonomicName authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="617723A4125A3442503EADED15127A2A" authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana</emphasis> <emphasis id="146E7BEBEC7FE8C48B6BF700204D1970" italics="true" pageId="0" pageNumber="145">Lachesana</emphasis>
</taxonomicName> </taxonomicName>
Strand, 1932 (8) and Strand, 1932 (8) and
<taxonomicName authorityName="Marx" authorityYear="1891" class="Arachnida" family="Zodariidae" genus="Lutica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lutica" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="958CC45898F36197CD8B6B1DD6CA33F8" authorityName="Marx" authorityYear="1891" class="Arachnida" family="Zodariidae" genus="Lutica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lutica" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Lutica</emphasis> <emphasis id="ADDA1DE3FEF6DD35F1BF5BDC24851CD3" italics="true" pageId="0" pageNumber="145">Lutica</emphasis>
</taxonomicName> </taxonomicName>
Marx, 1891 (4). Marx, 1891 (4).
<taxonomicName authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="9B555FF05628E7089B010B61A478057C" authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana</emphasis> <emphasis id="D0C4D7AAEEE8A5384B25FB1332E05544" italics="true" pageId="0" pageNumber="145">Lachesana</emphasis>
</taxonomicName> </taxonomicName>
is the only one restricted to the western Palaearctic ( is the only one restricted to the western Palaearctic (
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; ;
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). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

166
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<taxonomicName LSID="http://zoobank.org/DB25B91A-E827-4482-A2D4-1EC1D5012BF2" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium ovtchinnikovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ovtchinnikovi" status="sp. nov.">Acanthinozodium ovtchinnikovi</taxonomicName> <taxonomicName id="A0125E350D632CAD4E4D14979A439BDE" ID-CoL="98YZ2" LSID="http://zoobank.org/DB25B91A-E827-4482-A2D4-1EC1D5012BF2" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium ovtchinnikovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ovtchinnikovi" status="sp. nov.">Acanthinozodium ovtchinnikovi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="05C956309A0F2AE1BAF5905AF798D31C" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">Figs 22D, E, G-I</figureCitation> <figureCitation id="9D061F030F4119B5C324E76ED23EB9F7" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">Figs 22D, E, G-I</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation> <figureCitation id="600256BE01A5C1B229BA16F11EA9FF59" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="CB59F8FBEDC5E5E5B9A1CB9088EB9EB2" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="623E39AFC6E956EB7EC88A73E0AFA7C3" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="90F8615E100DED7C7C991BB4F1254331" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="73159058B830D65A259D44BB834B3034" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ palp (ZMMU), Turkmenistan: ♂ palp (ZMMU), Turkmenistan:
<emphasis italics="true" pageId="0" pageNumber="145">Mary Region</emphasis> <emphasis id="A29F533C148205F07D57F20BDA5DD1F8" italics="true" pageId="0" pageNumber="145">Mary Region</emphasis>
: Kushka Dist., ca 18 km S of Kyzyldzhar Kordon, ca 1 km ESE of Eroilandaz, : Kushka Dist., ca 18 km S of Kyzyldzhar Kordon, ca 1 km ESE of Eroilandaz,
<geoCoordinate degrees="35" direction="north" minutes="39" orientation="latitude" precision="925" value="35.65">35°39'N</geoCoordinate> <geoCoordinate id="B9A6E69750D8777423AE246B8FE36D65" degrees="35" direction="north" minutes="39" orientation="latitude" precision="925" value="35.65">35°39'N</geoCoordinate>
, ,
<geoCoordinate degrees="61" direction="east" minutes="50" orientation="longitude" precision="925" value="61.833332">61°50'E</geoCoordinate> <geoCoordinate id="2729AD70DFBEBE64EF435916B8D0D0FD" degrees="61" direction="east" minutes="50" orientation="longitude" precision="925" value="61.833332">61°50'E</geoCoordinate>
, 7.04.2002 (A.V. Gromov). , 7.04.2002 (A.V. Gromov).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="68AD5E69E842F14E6CDB114BA4D8BBE4" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="E133D69E2A620177BFECE4DE4C2537AC" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The new species is named after our late colleague Sergei V. Ovtchinnikov who made important contributions to the study of Central Asian spiders.</paragraph> <paragraph id="A99CF2D1248B8BF692CF458C1B9DAF58" pageId="0" pageNumber="145">The new species is named after our late colleague Sergei V. Ovtchinnikov who made important contributions to the study of Central Asian spiders.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="A4C620B1A5DE69CBA620E3A802147FEE" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="CEB685AB86427B19AE6F51D217391B1F" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="0DA0B7F31A8097FCC0F33F0AAAA27AF5" pageId="0" pageNumber="145">
The new species differs from the congeners in the region (except The new species differs from the congeners in the region (except
<taxonomicName lsidName="A. parmida" pageId="0" pageNumber="145" rank="species" species="parmida"> <taxonomicName id="CBBC3264EE5E193D05F721C49FB67D32" lsidName="A. parmida" pageId="0" pageNumber="145" rank="species" species="parmida">
<emphasis italics="true" pageId="0" pageNumber="145">A. parmida</emphasis> <emphasis id="FF2C66D3A1D738111E31D82071A0F53A" italics="true" pageId="0" pageNumber="145">A. parmida</emphasis>
</taxonomicName> </taxonomicName>
sp. nov.) by the small size of the palp (cymbium 0.28 long, sp. nov.) by the small size of the palp (cymbium 0.28 long,
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="B645E842BDA028DD48B30F0707B30B54" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
&gt; 0.6) and the unique embolus which has a long furrow with a serrate ventral margin (Fig. &gt; 0.6) and the unique embolus which has a long furrow with a serrate ventral margin (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22E</figureCitation> <figureCitation id="B8D74EFF48D8F9C8E626104F54C779DA" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22E</figureCitation>
). It is closely related to ). It is closely related to
<taxonomicName lsidName="A. parmida" pageId="0" pageNumber="145" rank="species" species="parmida"> <taxonomicName id="64E563C84CB5AF027166162CB42C4E73" lsidName="A. parmida" pageId="0" pageNumber="145" rank="species" species="parmida">
<emphasis italics="true" pageId="0" pageNumber="145">A. parmida</emphasis> <emphasis id="CD957D44EC34174285DE77CB592CC3CE" italics="true" pageId="0" pageNumber="145">A. parmida</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. from central Iran, from which it can be differentiated by having a longer than wide bulb ( sp. nov. from central Iran, from which it can be differentiated by having a longer than wide bulb (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="29BCB8C483DA28958D8EEE5CABE299D6" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
almost as long as wide), relatively longer cymbium (length/width ratio 1.86 vs. 1.4), and a different shape of median apophysis (anterior portion larger than posterior one, almost as long as wide), relatively longer cymbium (length/width ratio 1.86 vs. 1.4), and a different shape of median apophysis (anterior portion larger than posterior one,
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="E5225822C94D1EC226666459A6686BAA" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
opposite; cf. Figs opposite; cf. Figs
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22G</figureCitation> <figureCitation id="F14A9CA0A06EBAC231FBC018524D31A7" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22G</figureCitation>
and and
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19B</figureCitation> <figureCitation id="05EEB41F374DBEE2F4F03E8DE2F4CF9C" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19B</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="7C682029776913AF75F7D9C5798A15D2" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="CFEFA68E68832D40380E02A331A4D7CB" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="EBB0325CC799DD619E4C7506913D5AD1" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis> <emphasis id="E48CE970C73C7F0B789E3BC21BF90EF6" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Body missing. It is assumed to be a very small zodariid based on the size of the palp. Body missing. It is assumed to be a very small zodariid based on the size of the palp.
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="F95B4F0F8DE9B6040A0F91E01CDA776B" pageId="0" pageNumber="145">
Palp as in Fig. Palp as in Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22D, E, G-I</figureCitation> <figureCitation id="5A0D28334E1AD39B9271AF1734D4214F" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22D, E, G-I</figureCitation>
. Ventral tibial apophysis lacking; RTA almost triangular, slightly longer than tibia; cymbium longer than wide; sperm duct tracking margin of tegulum, lacking any turns; median apophysis longer than wide, with anterior part larger than posterior part; embolus originates at about 8:00 o'clock position with its terminal 2/3 having a longitudinal furrow ( . Ventral tibial apophysis lacking; RTA almost triangular, slightly longer than tibia; cymbium longer than wide; sperm duct tracking margin of tegulum, lacking any turns; median apophysis longer than wide, with anterior part larger than posterior part; embolus originates at about 8:00 o'clock position with its terminal 2/3 having a longitudinal furrow (
<emphasis italics="true" pageId="0" pageNumber="145">Er</emphasis> <emphasis id="4305FA83C3234FC00687B573DFE48A5A" italics="true" pageId="0" pageNumber="145">Er</emphasis>
) with a finely serrated ventral margin. ) with a finely serrated ventral margin.
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="6F816B27751EAC062B03E68EED95E2FB" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="C2FB76EE65788440B7F4FEEEDD25A9E2" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown. Unknown.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="notes"> <subSubSection id="4EBC796BACE286CF8385164728A838F8" pageId="0" pageNumber="145" type="notes">
<paragraph pageId="0" pageNumber="145">Note.</paragraph> <paragraph id="86283D0F22B28CD4F41EABCFCDD87E48" pageId="0" pageNumber="145">Note.</paragraph>
<paragraph pageId="0" pageNumber="145">The sample collected by Gromov contained three males. The palp of one specimen was dissected and imaged with a SEM in 2005. Then, all three specimens were given to Ovtchinnikov who was planning to revise this group. After his death, his collection was transferred to Almaty, Kazakhstan by Alexander Gromov and is now inaccessible.</paragraph> <paragraph id="8969DA625D0310C5234095DACE874564" pageId="0" pageNumber="145">The sample collected by Gromov contained three males. The palp of one specimen was dissected and imaged with a SEM in 2005. Then, all three specimens were given to Ovtchinnikov who was planning to revise this group. After his death, his collection was transferred to Almaty, Kazakhstan by Alexander Gromov and is now inaccessible.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="EB8C00CB3818D0402BC36464F5C3A82E" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="3E305FB09ACA7BF817416655EA36782E" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="FA524AC25D13F726BFD1E98C794CB3CE" pageId="0" pageNumber="145">
Known only from the type locality in Mary Region, southeastern Turkmenistan (Fig. Known only from the type locality in Mary Region, southeastern Turkmenistan (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation> <figureCitation id="12F060B80CFBCDDDC4667E0812296523" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

126
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<taxonomicName LSID="349A2817-76CA-5C7C-B644-621BA3CB3027" authority="(Ono &amp; Jocque, 1986)" baseAuthorityName="Ono &amp; Jocque" baseAuthorityYear="1986" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion buettikeri" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="buettikeri"> <taxonomicName id="64F7B8A0FB427044BCD250D1E675F08C" ID-CoL="5D69F" LSID="349A2817-76CA-5C7C-B644-621BA3CB3027" authority="(Ono &amp; Jocque, 1986)" baseAuthorityName="Ono &amp; Jocque" baseAuthorityYear="1986" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion buettikeri" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="buettikeri">
Zodarion buettikeri (Ono &amp; Zodarion buettikeri (Ono &amp;
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, 1986) , 1986)
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">Fig. 33</figureCitation> <figureCitation id="DBFEF7F02E2A0C667FADA3F13AB6AA20" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">Fig. 33</figureCitation>
</paragraph> </paragraph>
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<paragraph pageId="0" pageNumber="145"> <paragraph id="154D2604E5F1449A4F348BEB4EA1F417" pageId="0" pageNumber="145">
<taxonomicName class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium buettikeri" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="buettikeri">Acanthinozodium buettikeri</taxonomicName> <taxonomicName id="07FF28A531FB12F0DB8B5FDB9EED3472" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium buettikeri" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="buettikeri">Acanthinozodium buettikeri</taxonomicName>
Ono &amp; Ono &amp;
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, 1986: 7, f. 1-4 (♂♀). , 1986: 7, f. 1-4 (♂♀).
</paragraph> </paragraph>
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<taxonomicName baseAuthorityName="Ono &amp; Jocque" baseAuthorityYear="1986" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion buettikeri" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="buettikeri">Zodarion buettikeri</taxonomicName> <taxonomicName id="1E3DC52D0B336E3435B605586A1E95BE" baseAuthorityName="Ono &amp; Jocque" baseAuthorityYear="1986" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion buettikeri" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="buettikeri">Zodarion buettikeri</taxonomicName>
: :
<bibRefCitation author="Levy, G" journalOrPublisher="Israel Journal of Zoology" pageId="0" pageNumber="145" pagination="67 - 110" refId="B11" refString="Levy, G, 1992. The spider genera Palaestina, Trygetus, Zodarion and Ranops (Araneae, Zodariidae) in Israel with annotations on species of the Middle East. Israel Journal of Zoology 38: 67 - 110" title="The spider genera Palaestina, Trygetus, Zodarion and Ranops (Araneae, Zodariidae) in Israel with annotations on species of the Middle East." volume="38" year="1992">Levy 1992</bibRefCitation> <bibRefCitation id="BB60855DEFFB308C0A30CFC73CE877CD" author="Levy, G" journalOrPublisher="Israel Journal of Zoology" pageId="0" pageNumber="145" pagination="67 - 110" refId="B11" refString="Levy, G, 1992. The spider genera Palaestina, Trygetus, Zodarion and Ranops (Araneae, Zodariidae) in Israel with annotations on species of the Middle East. Israel Journal of Zoology 38: 67 - 110" title="The spider genera Palaestina, Trygetus, Zodarion and Ranops (Araneae, Zodariidae) in Israel with annotations on species of the Middle East." volume="38" year="1992">Levy 1992</bibRefCitation>
: 85; : 85;
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation> <bibRefCitation id="E8D2C3684478B782709DE58D26127A8E" DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation>
: 69, f. 3E-F (♂). : 69, f. 3E-F (♂).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in iran"> <subSubSection id="CF9988D900DA81B5B291DE9A98DD5E90" pageId="0" pageNumber="145" type="records in iran">
<paragraph pageId="0" pageNumber="145">Records in Iran.</paragraph> <paragraph id="135008EA9F56B237E9E441A8C990CD5C" pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="A7FA9406077087A40C4FACE4C30CCB37" pageId="0" pageNumber="145">
Bushehr ( Bushehr (
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation> <bibRefCitation id="96B8362315094D379D11DA6E70E026C0" DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation>
) (Fig. ) (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation> <figureCitation id="F36E06291829CA93177A573EF7A41B76" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="7D834D27B35E0533B61F8C3D13FEE84D" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="B33B1042ADB2C4CD1BE621B41BF85DA5" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">Saudi Arabia, Iran.</paragraph> <paragraph id="D45C34882484722D2D2EF54E499EB049" pageId="0" pageNumber="145">Saudi Arabia, Iran.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comments"> <subSubSection id="6E2A682E08CD963B18C1D34C2902316A" pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="8FE338D1FA9D7540295CE3C05F10E599" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="B43C42C664D5009D79F244D181E3573E" pageId="0" pageNumber="145">
Since the Iranian material of this species has already been illustrated in Since the Iranian material of this species has already been illustrated in
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. (2017)</bibRefCitation> <bibRefCitation id="C199319132749F820FF69B6C66896B1E" DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. (2017)</bibRefCitation>
, we are not providing new figures for it. , we are not providing new figures for it.
</paragraph> </paragraph>
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180
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<treatment ID-GBIF-Taxon="181753568" LSID="urn:lsid:zoobank.org:act:4E56657F-4A5F-48F2-A3B4-E306128A62E6" httpUri="http://treatment.plazi.org/id/363BA2902296556F82E4535F556248F1" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="363BA2902296556F82E4535F556248F1" ID-GBIF-Taxon="181753568" LSID="urn:lsid:zoobank.org:act:4E56657F-4A5F-48F2-A3B4-E306128A62E6" httpUri="http://treatment.plazi.org/id/363BA2902296556F82E4535F556248F1" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="A3BCB1B11328530D6A322CFB5ECFB5B0" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="3EF98410216CEC071107F7CA92A8CC2C" pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/4E56657F-4A5F-48F2-A3B4-E306128A62E6" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus susianus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="susianus" status="sp. nov.">Trygetus susianus</taxonomicName> <taxonomicName id="0FE2E3659FF00E71A09299F118F1CC6D" ID-CoL="98YZB" LSID="http://zoobank.org/4E56657F-4A5F-48F2-A3B4-E306128A62E6" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus susianus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="susianus" status="sp. nov.">Trygetus susianus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="A95A7F71AC860B352E6081E79781AD78" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">Figs 26D</figureCitation> <figureCitation id="48D40055C363690865CB0D3C8F904705" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">Figs 26D</figureCitation>
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Epigynes of Parazodarion raddei (A-D) and Trygetus susianus sp. nov. (E-H) A, E intact, ventral B, F-H macerated, dorsal C, D macerated, ventral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145">, 27E-H</figureCitation> <figureCitation id="E69100EB89559B4873175A59F211ACAF" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Epigynes of Parazodarion raddei (A-D) and Trygetus susianus sp. nov. (E-H) A, E intact, ventral B, F-H macerated, dorsal C, D macerated, ventral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145">, 27E-H</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation> <figureCitation id="0FBF980B8BA605BA2ABDCAA7B65CBCB9" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="reference_group"> <subSubSection id="E2D11B060B305AB246E5DCDFB1F748B6" pageId="0" pageNumber="145" type="reference_group">
<paragraph pageId="0" pageNumber="145"> <paragraph id="31F4F98DD3303AD0B988F484FCD97864" pageId="0" pageNumber="145">
<taxonomicName authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus jacksoni" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="jacksoni" status="sp. nov.">Trygetus jacksoni</taxonomicName> <taxonomicName id="4420B8C7DFE811BF39B4FC6CE3799C44" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus jacksoni" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="jacksoni" status="sp. nov.">Trygetus jacksoni</taxonomicName>
: :
<bibRefCitation author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. 2018</bibRefCitation> <bibRefCitation id="3021A7ED0779DD9BEDE3FB07A36B64A2" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. 2018</bibRefCitation>
: 188 (♀, misidentified). : 188 (♀, misidentified).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="84A74D2ABA0E17B3DBE60D63FFA3CB28" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="E06F65BCA04E4A369E28D7D089D1FC18" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="802D6FD00579F0C3F09FB0CAD473190B" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="DC770590FA6CFDFA8DCD91FD074C77EC" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♀ (MHNG), Iran: ♀ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Khuzestan Province</emphasis> <emphasis id="AF1DDCC1FB84103EF2952CA0F81401C6" italics="true" pageId="0" pageNumber="145">Khuzestan Province</emphasis>
: north of Andimeshk, : north of Andimeshk,
<geoCoordinate degrees="32" direction="north" minutes="41" orientation="latitude" precision="925" value="32.683334">32°41'N</geoCoordinate> <geoCoordinate id="D03E165C63A4637CFFAE02E1712E2B1C" degrees="32" direction="north" minutes="41" orientation="latitude" precision="925" value="32.683334">32°41'N</geoCoordinate>
, ,
<geoCoordinate degrees="48" direction="east" minutes="15" orientation="longitude" precision="925" value="48.25">48°15'E</geoCoordinate> <geoCoordinate id="11342565ACD0FB2784456B9B1A54BA20" degrees="48" direction="east" minutes="15" orientation="longitude" precision="925" value="48.25">48°15'E</geoCoordinate>
, 17.05.1974 (A. Senglet). , 17.05.1974 (A. Senglet).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="504E233995CFBAF92EAA8B2AEA14A06A" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="7AC54454E20E7D0DD643EC5BAEB40C09" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">This species is named after Susa, one of the most important cities of the Ancient Near East. It is located in the lower Zagros Mountains, about 250 km east of the Tigris River, between the Karkheh and Dez Rivers, in what is currently the Khuzestan Province of Iran.</paragraph> <paragraph id="FAA8A3E7E32F7C4048CF05EF65DD3AE5" pageId="0" pageNumber="145">This species is named after Susa, one of the most important cities of the Ancient Near East. It is located in the lower Zagros Mountains, about 250 km east of the Tigris River, between the Karkheh and Dez Rivers, in what is currently the Khuzestan Province of Iran.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="2D0B20BE488F9094DEF26117A1019537" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="F62FBEC78D88CEE616E9646CA418011C" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="762936A75A2EB20F99B4C0FA769ECE1C" pageId="0" pageNumber="145">
The epigyne of the new species is very similar to that of The epigyne of the new species is very similar to that of
<taxonomicName lsidName="T. gromovi" pageId="0" pageNumber="145" rank="species" species="gromovi"> <taxonomicName id="A37827C56581D3D0B7D086A4B6C630B6" lsidName="T. gromovi" pageId="0" pageNumber="145" rank="species" species="gromovi">
<emphasis italics="true" pageId="0" pageNumber="145">T. gromovi</emphasis> <emphasis id="C51F255739488B4057D73651B45F5AC5" italics="true" pageId="0" pageNumber="145">T. gromovi</emphasis>
</taxonomicName> </taxonomicName>
. They differ by the presence of a lateral extension of the receptacle and the lack of sclerotized lateral margins in the new species ( . They differ by the presence of a lateral extension of the receptacle and the lack of sclerotized lateral margins in the new species (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="6C9E3FE1821A1B8608FE11B227A60CC7" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
absent and present, respectively). absent and present, respectively).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="7F306A9B2920F7F391727D57F22289E1" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="A109B64C0E7520850728A7A792D15FA9" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="F7A3246B2B9AAB9F47C3D3671A612F20" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female</emphasis> <emphasis id="B8C19FED2F4ADBE1A3052367184755B8" bold="true" pageId="0" pageNumber="145">Female</emphasis>
(specimen partially bleached). Habitus as in Fig. (specimen partially bleached). Habitus as in Fig.
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">26D</figureCitation> <figureCitation id="01D4F5030418AD0C689643C115E198B8" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">26D</figureCitation>
. Total length 2.07. Carapace 0.96 long, 0.53 wide at pars cephalica, 0.69 wide at pars thoracica. Eye sizes: AME: 0.08, ALE: 0.05, PLE: 0.04. Carapace, sternum, labium, chelicerae and maxillae yellowish brown, without any pattern. Legs slightly lighter than carapace, without spines and without annulations. Abdomen grayish, darker dorsally. Spinnerets pale, uniform in color. Measurements of legs: I: 2.19 (0.61, 0.29, 0.44, 0.44, 0.41), II: 1.98 (0.58, 0.28, 0.39, 0.36, 0.37), III: 1.90 (0.52, 0.26, 0.32, 0.40, 0.40), IV: 2.50 (0.67, 0.28, 0.48, 0.63, 0.44). . Total length 2.07. Carapace 0.96 long, 0.53 wide at pars cephalica, 0.69 wide at pars thoracica. Eye sizes: AME: 0.08, ALE: 0.05, PLE: 0.04. Carapace, sternum, labium, chelicerae and maxillae yellowish brown, without any pattern. Legs slightly lighter than carapace, without spines and without annulations. Abdomen grayish, darker dorsally. Spinnerets pale, uniform in color. Measurements of legs: I: 2.19 (0.61, 0.29, 0.44, 0.44, 0.41), II: 1.98 (0.58, 0.28, 0.39, 0.36, 0.37), III: 1.90 (0.52, 0.26, 0.32, 0.40, 0.40), IV: 2.50 (0.67, 0.28, 0.48, 0.63, 0.44).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="AFDE0792F6F4C7E492ACC00FA25653BB" pageId="0" pageNumber="145">
Epigyne as in Fig. Epigyne as in Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Epigynes of Parazodarion raddei (A-D) and Trygetus susianus sp. nov. (E-H) A, E intact, ventral B, F-H macerated, dorsal C, D macerated, ventral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145">27E-H</figureCitation> <figureCitation id="C0E736EF700D1DEB6C302A2A404F7538" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Epigynes of Parazodarion raddei (A-D) and Trygetus susianus sp. nov. (E-H) A, E intact, ventral B, F-H macerated, dorsal C, D macerated, ventral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145">27E-H</figureCitation>
. Epigyne medially with posteriorly diverging sclerotization, laterally with translucent fertilization ducts; receptacles almost round, separated by less than their diameters, each subdivided into a large, weakly sclerotized part and a smaller, more heavily sclerotized part. . Epigyne medially with posteriorly diverging sclerotization, laterally with translucent fertilization ducts; receptacles almost round, separated by less than their diameters, each subdivided into a large, weakly sclerotized part and a smaller, more heavily sclerotized part.
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145" start="Figure 27" startId="F27"> <caption id="F3679EEFF180C068D1CED1590333FC8C" doi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145" start="Figure 27" startId="F27">
<paragraph pageId="0" pageNumber="145"> <paragraph id="2D26797137DE03E48A325541C2024E39" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 27.</emphasis> <emphasis id="49655540CA742A0A8CD3838D7C0D012E" bold="true" pageId="0" pageNumber="145">Figure 27.</emphasis>
Epigynes of Epigynes of
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion raddei" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei"> <taxonomicName id="20DE22DE7D4E552D156D079DAF75B61C" baseAuthorityName="Simon" baseAuthorityYear="1889" family="Zodariidae" genus="Parazodarion" kingdom="Animalia" lsidName="Parazodarion raddei" order="Aranei" pageId="0" pageNumber="145" rank="species" species="raddei">
<emphasis italics="true" pageId="0" pageNumber="145">Parazodarion raddei</emphasis> <emphasis id="7E0AFE395F7B1ED9C0D5AF9EDB1366BE" italics="true" pageId="0" pageNumber="145">Parazodarion raddei</emphasis>
</taxonomicName> </taxonomicName>
( (
<emphasis bold="true" pageId="0" pageNumber="145">A-D</emphasis> <emphasis id="C715F552A6BF7579867DA882EA734E5B" bold="true" pageId="0" pageNumber="145">A-D</emphasis>
) and ) and
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus susianus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="susianus"> <taxonomicName id="94BEDA40FAED9F6E8F0B3F6ADDAB2FD2" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus susianus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="susianus">
<emphasis italics="true" pageId="0" pageNumber="145">Trygetus susianus</emphasis> <emphasis id="1AF011167D88E5A4F8599A7B42BDE0C8" italics="true" pageId="0" pageNumber="145">Trygetus susianus</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">E-H</emphasis> <emphasis id="F3CDB45AD1FC32BDC26FE788DB277A9E" bold="true" pageId="0" pageNumber="145">E-H</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, E</emphasis> <emphasis id="A00A9E3B00E508831D8CCB3F84512A31" bold="true" pageId="0" pageNumber="145">A, E</emphasis>
intact, ventral intact, ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, F-H</emphasis> <emphasis id="B82CB123549762110DEF91A67CCAB280" bold="true" pageId="0" pageNumber="145">B, F-H</emphasis>
macerated, dorsal macerated, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">C, D</emphasis> <emphasis id="80207104489B8E8D8651664D3FCAD40D" bold="true" pageId="0" pageNumber="145">C, D</emphasis>
macerated, ventral. Scale bars: 0.2 mm. macerated, ventral. Scale bars: 0.2 mm.
</paragraph> </paragraph>
</caption> </caption>
<paragraph pageId="0" pageNumber="145"> <paragraph id="3BCCB640FBA7FDBDF3142D3817D4A754" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis> <emphasis id="E3C9FA4D06C9E6B7F8CBDA9C065DE7AA" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Unknown. Unknown.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="D3B41B8C68B48840D127CE85ACF7766D" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="14032BB7B8B3F0A2CDEC4CE2DA2EDE63" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="66A318A4FC1970F922BAEAD3B76BF95F" pageId="0" pageNumber="145">
Known from two localities in Khuzestan Province, southwestern Iran (Fig. Known from two localities in Khuzestan Province, southwestern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation> <figureCitation id="2ADC834748329E3497CD5250189B3CA4" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

312
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@ -1,278 +1,278 @@
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title> <mods:title id="03885FA4023CF9ABB64939CAC395BBC3">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<treatment ID-GBIF-Taxon="181753571" LSID="urn:lsid:zoobank.org:act:D52287DC-6ED8-4E56-B435-F5CB56650469" httpUri="http://treatment.plazi.org/id/3D04C9134C3357AF827C326761DF8161" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="3D04C9134C3357AF827C326761DF8161" ID-GBIF-Taxon="181753571" LSID="urn:lsid:zoobank.org:act:D52287DC-6ED8-4E56-B435-F5CB56650469" httpUri="http://treatment.plazi.org/id/3D04C9134C3357AF827C326761DF8161" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Storeninae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="B8EF491F9B8E49D577FAE4C3EB164330" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="5DD65436067242D7DFE59FF67CF05956" pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/D52287DC-6ED8-4E56-B435-F5CB56650469" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita" status="sp. nov.">Pax ellipita</taxonomicName> <taxonomicName id="4787CDF2B73D0908C79E07BD5ACD8B19" ID-CoL="98YZ5" LSID="http://zoobank.org/D52287DC-6ED8-4E56-B435-F5CB56650469" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita" status="sp. nov.">Pax ellipita</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="4E1D86037F50AEE3366F396D40ED649D" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">Figs 5A, B</figureCitation> <figureCitation id="84E29CF178B42E2C4341CFD98F1102B1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">Figs 5A, B</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male palp of Pax ellipita sp. nov. A ventral B, C retrolateral D, E retrodorsal and prolateral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145">, 6</figureCitation> <figureCitation id="A3ED547FA882A6F33E2077169CCDDB1C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male palp of Pax ellipita sp. nov. A ventral B, C retrolateral D, E retrodorsal and prolateral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. SEM images of the male palp of Pax ellipita sp. nov. A retrolateral B anteroretrolateral C proventral D posterodorsal E ventral. Abbreviations: E 1, E 2 - pro- and retrolateral cymbial extensions, Eb - embolar base, Em - embolus, Cf - cymbial fold, Co - conductor, Ma - median apophysis, Ra - retrolateral apophysis, Va - ventral apophysis. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145">, 7</figureCitation> <figureCitation id="07EBA79E1AE5DB94B5B7E919E764F54F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. SEM images of the male palp of Pax ellipita sp. nov. A retrolateral B anteroretrolateral C proventral D posterodorsal E ventral. Abbreviations: E 1, E 2 - pro- and retrolateral cymbial extensions, Eb - embolar base, Em - embolus, Cf - cymbial fold, Co - conductor, Ma - median apophysis, Ra - retrolateral apophysis, Va - ventral apophysis. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145">, 7</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">, 10A-E</figureCitation> <figureCitation id="77A312715848D1E3C25B030D6E8C70B6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">, 10A-E</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation> <figureCitation id="1FEC49BC8BAE262A375CC0EC4A253A9B" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="EC14C324732A0F42DA3E4EB31D04D8DD" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="900A73ED5341BDDDCEBFFE04204251B5" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="46A8702CB5303F32A7C93754C5C15ACE" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="5BF8A099EF7F1194B6988DE55BBD246C" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran: ♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Kermanshah Province</emphasis> <emphasis id="869DB96280192DA74EFDBE6A61A6976E" italics="true" pageId="0" pageNumber="145">Kermanshah Province</emphasis>
: north of Kermanshah, : north of Kermanshah,
<geoCoordinate degrees="34" direction="north" minutes="28" orientation="latitude" precision="925" value="34.466667">34°28'N</geoCoordinate> <geoCoordinate id="BEFF1B1B545027CEA9F09FA9A6311117" degrees="34" direction="north" minutes="28" orientation="latitude" precision="925" value="34.466667">34°28'N</geoCoordinate>
, ,
<geoCoordinate degrees="47" direction="east" minutes="00" orientation="longitude" precision="925" value="47.0">47°00'E</geoCoordinate> <geoCoordinate id="E7AD95E7ED71201D36E84507D766B717" degrees="47" direction="east" minutes="00" orientation="longitude" precision="925" value="47.0">47°00'E</geoCoordinate>
, 18.06.1975 (A. Senglet). , 18.06.1975 (A. Senglet).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="materials_examined"> <subSubSection id="05CC8E29FFE0718000E9FD0CA084BFD4" pageId="0" pageNumber="145" type="materials_examined">
<paragraph pageId="0" pageNumber="145">Additional material.</paragraph> <paragraph id="DF76CD51F0481726BE6DE8DC5E65931F" pageId="0" pageNumber="145">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="58124689C0592D5E511534EAD839DF1B" pageId="0" pageNumber="145">
<materialsCitation ID-GBIF-Occurrence="3093634308" collectingDate="1977-04-11" collectorName="B. Prazan" country="Iran" latitude="32.85" location="river Saimareh" longLatPrecision="1208" longitude="47.933334" municipality="Pol-e Tang" specimenCount="1" specimenCount-female="1" stateProvince="Lorestan Province"> <materialsCitation id="85A1406004A9103FC355D151ADB55CAC" ID-GBIF-Occurrence="3093634308" collectingDate="1977-04-11" collectorName="B. Prazan" country="Iran" latitude="32.85" location="river Saimareh" longLatPrecision="1208" longitude="47.933334" municipality="Pol-e Tang" specimenCount="1" specimenCount-female="1" stateProvince="Lorestan Province">
<collectingCountry name="Iran">Iran</collectingCountry> <collectingCountry id="114FB4C83AB694210D7E96E47F46A3FB" name="Iran">Iran</collectingCountry>
: :
<emphasis italics="true" pageId="0" pageNumber="145"> <emphasis id="6B8F74AA2E103A05F163B285727F2376" italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Lorestan">Lorestan Province</collectingRegion> <collectingRegion id="0986DAEC94B11F67DD13D77455D1EC82" country="Iran" name="Lorestan">Lorestan Province</collectingRegion>
</emphasis> </emphasis>
: :
<specimenCount type="female">1♀</specimenCount> <specimenCount id="AE619007CD3437A402C2CA24AAA37F63" type="female">1♀</specimenCount>
(NMP), (NMP),
<collectingMunicipality>Pol-e Tang</collectingMunicipality> <collectingMunicipality id="478FAF9D98D3524414C00C370D4D7177">Pol-e Tang</collectingMunicipality>
, ,
<locationDeviation location="Andimeshk"> <locationDeviation id="5F71721AD0CA0389EDFB4B86EAC252E0" location="Andimeshk">
<quantity metricMagnitude="4" metricUnit="m" metricValue="6.0" unit="km" value="60.0">60 km</quantity> <quantity id="B359E673989041A749B5377713E2E46E" metricMagnitude="4" metricUnit="m" metricValue="6.0" unit="km" value="60.0">60 km</quantity>
NW of Andimeshk NW of Andimeshk
</locationDeviation> </locationDeviation>
, ,
<geoCoordinate degrees="32" direction="north" minutes="51" orientation="latitude" precision="925" value="32.85">32°51'N</geoCoordinate> <geoCoordinate id="BD04B59C4D28FF919B25D29FFC19755D" degrees="32" direction="north" minutes="51" orientation="latitude" precision="925" value="32.85">32°51'N</geoCoordinate>
, ,
<geoCoordinate degrees="47" direction="east" minutes="56" orientation="longitude" precision="925" value="47.933334">47°56'E</geoCoordinate> <geoCoordinate id="432531442F29D18302C181D03A7FFED9" degrees="47" direction="east" minutes="56" orientation="longitude" precision="925" value="47.933334">47°56'E</geoCoordinate>
, near the , near the
<location LSID="urn:lsid:plazi:treatment:3D04C9134C3357AF827C326761DF8161:71B04B1A5985BF7FCAC1BD1C1C6158A6" country="Iran" latitude="32.85" longLatPrecision="1208" longitude="47.933334" municipality="Pol-e Tang" name="river Saimareh" stateProvince="Lorestan Province">river Saimareh</location> <location id="71B04B1A5985BF7FCAC1BD1C1C6158A6" LSID="urn:lsid:plazi:treatment:3D04C9134C3357AF827C326761DF8161:71B04B1A5985BF7FCAC1BD1C1C6158A6" country="Iran" latitude="32.85" longLatPrecision="1208" longitude="47.933334" municipality="Pol-e Tang" name="river Saimareh" stateProvince="Lorestan Province">river Saimareh</location>
, ,
<collectingDate value="1977-04-11">11.04.1977</collectingDate> <collectingDate id="0CB8554777EE6D42E577014CAEF3CBB2" value="1977-04-11">11.04.1977</collectingDate>
( (
<collectorName> <collectorName id="C1948061F7D6881F5B8160D89AD5F5BE">
B. B.
<normalizedToken originalValue="Pražan">Prazan</normalizedToken> <normalizedToken id="53E7B174EB3073FE68C387C839F3D8C0" originalValue="Pražan">Prazan</normalizedToken>
</collectorName> </collectorName>
) )
</materialsCitation> </materialsCitation>
. .
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="A103B63F8C68DF75C3FF07DE3F2AB00B" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="B51FC1D3977CD5F14E3E44BAB9C482A9" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet refers to Ellipi, an ancient kingdom located on the western side of the Zagros Mountains, between Babylonia at the west, Media at the north-east, Mannae at the north and Elam at the south.</paragraph> <paragraph id="6E862DD21B7A359C739BD5146F77E816" pageId="0" pageNumber="145">The specific epithet refers to Ellipi, an ancient kingdom located on the western side of the Zagros Mountains, between Babylonia at the west, Media at the north-east, Mannae at the north and Elam at the south.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="CDFA2F69DD4B693153B39847D94170EF" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="8568E266A8898EADA433C7D798811955" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="C03EA28D5896E613695CFFB916D01EB7" pageId="0" pageNumber="145">
The new species differs from The new species differs from
<taxonomicName lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila"> <taxonomicName id="F3E9162C5481726F5FFCA7DF1FF81475" lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila">
<emphasis italics="true" pageId="0" pageNumber="145">P. leila</emphasis> <emphasis id="5197A674C74CD9B07F1C953AE1B37517" italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
</taxonomicName> </taxonomicName>
sp. nov., the only other sp. nov., the only other
<taxonomicName authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="1BB9CDB0DCB2A3A29F2BCED1BCB659F9" authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Pax</emphasis> <emphasis id="F6F3D674A5E18706E8EE0709A72A2E26" italics="true" pageId="0" pageNumber="145">Pax</emphasis>
</taxonomicName> </taxonomicName>
species known from Iran, by the cymbium lacking a horn-like outgrowth ( species known from Iran, by the cymbium lacking a horn-like outgrowth (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="174E826A1043CF3A2B0543810074F778" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
present), by having a deep lateral fold of the cymbium ( present), by having a deep lateral fold of the cymbium (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="A428345B1C8A606390895F0EC1600D29" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
lacking) and the epigyne with a rectangular median plate ( lacking) and the epigyne with a rectangular median plate (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="0F151811F4905EBBF5B734C49D8C4DEC" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
triangular). The male of triangular). The male of
<taxonomicName lsidName="P. ellipita" pageId="0" pageNumber="145" rank="species" species="ellipita"> <taxonomicName id="405092E4B45B69D231A40DFAE8F6A544" lsidName="P. ellipita" pageId="0" pageNumber="145" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">P. ellipita</emphasis> <emphasis id="0EB747DFAC90C8D80BAE9F84D3AA0176" italics="true" pageId="0" pageNumber="145">P. ellipita</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. differs from the rest of the species by having the RTA shorter than the tibia ( sp. nov. differs from the rest of the species by having the RTA shorter than the tibia (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="45589CE9127F3D6E5511F48DA7FA511A" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
longer), while the female differs by the trilobate posterior margin of the epigyne ( longer), while the female differs by the trilobate posterior margin of the epigyne (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="8F7C44620E890CA23F944A29F3241F82" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
solid) and copulatory openings located posteriorly ( solid) and copulatory openings located posteriorly (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="6C8F3F4F40F57860B43A1F3DE04AD463" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
anteriorly). anteriorly).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="8C9D177F02B41C6199E5B3C180C1B2BB" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="B4D807EE5EC17DFD087A40C2F6A0B558" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="F9467B9BCC7A13150DA0F14B355B0FF8" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis> <emphasis id="DC418155A2CF8826BA96ABC53DF7A815" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Habitus as in Fig. Habitus as in Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5B</figureCitation> <figureCitation id="F88D73B81895251271656C4A65C29AFD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5B</figureCitation>
. Total length 5.07. Carapace 2.43 long, 1.27 wide at pars cephalica, 1.65 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.11, ALE: 0.10, PME: 0.08, PLE: 0.10, PME-PME: 0.09. Carapace, sternum, labium, chelicerae and maxillae reddish brown, without any pattern. Chelicera with 2 promarginal teeth. Legs yellow, with few spines and without annulations. Abdomen dark brown, dorsally with large scutum covering almost the entire abdomen. Spinnerets pale, uniform in color. Measurements of legs: I: 5.49 (1.48, 0.62, 1.22, 1.10, 1.07), II: 4.88 (1.29, 0.60, 1.04, 1.05, 0.90), III: 4.74 (1.27, 0.61, 0.90, 1.19, 0.77), IV: 6.71 (1.80, 0.68, 1.39, 1.80, 1.04). . Total length 5.07. Carapace 2.43 long, 1.27 wide at pars cephalica, 1.65 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.11, ALE: 0.10, PME: 0.08, PLE: 0.10, PME-PME: 0.09. Carapace, sternum, labium, chelicerae and maxillae reddish brown, without any pattern. Chelicera with 2 promarginal teeth. Legs yellow, with few spines and without annulations. Abdomen dark brown, dorsally with large scutum covering almost the entire abdomen. Spinnerets pale, uniform in color. Measurements of legs: I: 5.49 (1.48, 0.62, 1.22, 1.10, 1.07), II: 4.88 (1.29, 0.60, 1.04, 1.05, 0.90), III: 4.74 (1.27, 0.61, 0.90, 1.19, 0.77), IV: 6.71 (1.80, 0.68, 1.39, 1.80, 1.04).
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145" start="Figure 5" startId="F5"> <caption id="1DC138CDBEB1D6E579DDF4B26BB63867" doi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="145"> <paragraph id="F7F808C62584DE5F191AE85269AB3914" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 5.</emphasis> <emphasis id="9940F099B881D44282F9AEE37B2C42F4" bold="true" pageId="0" pageNumber="145">Figure 5.</emphasis>
Dorsal habitus of Dorsal habitus of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita"> <taxonomicName id="5FFE42A9781C9199C508FFB4D0B2A41F" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis> <emphasis id="2A884F0D0F6E6E78A631BAF5F3726F3D" italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A, B</emphasis> <emphasis id="DF876837BE669C53744D6008FC5E86BD" bold="true" pageId="0" pageNumber="145">A, B</emphasis>
) and ) and
<taxonomicName lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila"> <taxonomicName id="99147058216493ADA9462D732B0E2B70" lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila">
<emphasis italics="true" pageId="0" pageNumber="145">P. leila</emphasis> <emphasis id="3CED58E8C8CC22DDE719017F373F4258" italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">C, D</emphasis> <emphasis id="CABE0C977B91978C3DBBD5387839E3CD" bold="true" pageId="0" pageNumber="145">C, D</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis> <emphasis id="F77B7947986D25019AF3A881FC0621E6" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
females females
<emphasis bold="true" pageId="0" pageNumber="145">B, C</emphasis> <emphasis id="5669683DE18BB9909C847773EB83F81D" bold="true" pageId="0" pageNumber="145">B, C</emphasis>
males. males.
</paragraph> </paragraph>
</caption> </caption>
<paragraph pageId="0" pageNumber="145"> <paragraph id="DA59EFB89B5F0757AEAD0D36CA3BAA78" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male palp of Pax ellipita sp. nov. A ventral B, C retrolateral D, E retrodorsal and prolateral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145">6A-E</figureCitation> <figureCitation id="439D1B46A3E9BA3526CDA723CE4C95E3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male palp of Pax ellipita sp. nov. A ventral B, C retrolateral D, E retrodorsal and prolateral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145">6A-E</figureCitation>
, ,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. SEM images of the male palp of Pax ellipita sp. nov. A retrolateral B anteroretrolateral C proventral D posterodorsal E ventral. Abbreviations: E 1, E 2 - pro- and retrolateral cymbial extensions, Eb - embolar base, Em - embolus, Cf - cymbial fold, Co - conductor, Ma - median apophysis, Ra - retrolateral apophysis, Va - ventral apophysis. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145">7A-E</figureCitation> <figureCitation id="19A7CA425DE92DF2AD059A5E9B69C5D4" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. SEM images of the male palp of Pax ellipita sp. nov. A retrolateral B anteroretrolateral C proventral D posterodorsal E ventral. Abbreviations: E 1, E 2 - pro- and retrolateral cymbial extensions, Eb - embolar base, Em - embolus, Cf - cymbial fold, Co - conductor, Ma - median apophysis, Ra - retrolateral apophysis, Va - ventral apophysis. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145">7A-E</figureCitation>
. Femur slightly shorter than cymbium, 3.3 times longer than wide; patella globular; tibia wider than long with ventral apophysis ( . Femur slightly shorter than cymbium, 3.3 times longer than wide; patella globular; tibia wider than long with ventral apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis> <emphasis id="3E865AAA7C2FCD45DB60AFE1CD33DDB7" italics="true" pageId="0" pageNumber="145">Va</emphasis>
) and short RTA bifurcated at the tip ( ) and short RTA bifurcated at the tip (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis> <emphasis id="39F9E498225F863711F1CC4F90E3C8D0" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
); cymbium with large fold ( ); cymbium with large fold (
<emphasis italics="true" pageId="0" pageNumber="145">Cf</emphasis> <emphasis id="97205EAD6FA40E9B507E111BE158956E" italics="true" pageId="0" pageNumber="145">Cf</emphasis>
) and 2 extensions, pro- and retrolateral ( ) and 2 extensions, pro- and retrolateral (
<emphasis italics="true" pageId="0" pageNumber="145">E1</emphasis> <emphasis id="54F1E04DCF1F89AB9F8704D87410DAA2" italics="true" pageId="0" pageNumber="145">E1</emphasis>
, ,
<emphasis italics="true" pageId="0" pageNumber="145">E2</emphasis> <emphasis id="76FA5A6A59257E0233CE78C7089B09C0" italics="true" pageId="0" pageNumber="145">E2</emphasis>
); bulb with large lamella, as long as bulb, tapering to the tip; median apophysis ( ); bulb with large lamella, as long as bulb, tapering to the tip; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="39288C442791F17C00BAD1F097851766" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) long, about 5 times longer than wide; conductor ( ) long, about 5 times longer than wide; conductor (
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis> <emphasis id="4684AD63BEA5EEB6DF1F145CD0D4F9DE" italics="true" pageId="0" pageNumber="145">Co</emphasis>
) heavily sclerotized; embolus ( ) heavily sclerotized; embolus (
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis> <emphasis id="982D429F9366AE39C7638F3B759294FF" italics="true" pageId="0" pageNumber="145">Em</emphasis>
) long, originates at 6 ) long, originates at 6
<normalizedToken originalValue="o’clock">o'clock</normalizedToken> <normalizedToken id="68FB6C65B0090DAE047EE3E933892715" originalValue="o’clock">o'clock</normalizedToken>
position. position.
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145" start="Figure 6" startId="F6"> <caption id="EF95E638690874A842A19B3F5147AB61" doi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="145"> <paragraph id="4EFBD814475B1B8ADEF951125AFACDEC" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 6.</emphasis> <emphasis id="E5E3FC8C67B350B38806645DBB3F9959" bold="true" pageId="0" pageNumber="145">Figure 6.</emphasis>
Male palp of Male palp of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita"> <taxonomicName id="AF19F40617E25C36F85435B3DB26AA1C" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis> <emphasis id="1A246894F03470C4EA88996A5EEB2142" italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis> <emphasis id="CD12809AEA2312D730F93FEBA62A02CD" bold="true" pageId="0" pageNumber="145">A</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, C</emphasis> <emphasis id="C1FB719A163E1078B7FF55EA6C7B75FB" bold="true" pageId="0" pageNumber="145">B, C</emphasis>
retrolateral retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">D, E</emphasis> <emphasis id="6C4363BDB1ABD99D0466C079F6F18928" bold="true" pageId="0" pageNumber="145">D, E</emphasis>
retrodorsal and prolateral. Scale bars: 0.2 mm. retrodorsal and prolateral. Scale bars: 0.2 mm.
</paragraph> </paragraph>
</caption> </caption>
<caption doi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145" start="Figure 7" startId="F7"> <caption id="17199827B2FC18A0AAF0D0FE7D19CC73" doi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="145"> <paragraph id="2DBA9ABF4D31C32527132B6B8D68836D" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 7.</emphasis> <emphasis id="82182B536DADBF5FE219589DEEC203A0" bold="true" pageId="0" pageNumber="145">Figure 7.</emphasis>
SEM images of the male palp of SEM images of the male palp of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita"> <taxonomicName id="EDCF2A1FEAB00118B48D93D845F25721" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis> <emphasis id="442A1278412D2991517BA066E4D71AEB" italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis> <emphasis id="918FBBB333B7CC46625038276D2CC4C9" bold="true" pageId="0" pageNumber="145">A</emphasis>
retrolateral retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis> <emphasis id="528D4500E8EB46EA73AAF86614D8CF57" bold="true" pageId="0" pageNumber="145">B</emphasis>
anteroretrolateral anteroretrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis> <emphasis id="8271ED8609C131C03AEBB9CF32CF71B6" bold="true" pageId="0" pageNumber="145">C</emphasis>
proventral proventral
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis> <emphasis id="9850714E0E19C46B51388E2A7DEAAF93" bold="true" pageId="0" pageNumber="145">D</emphasis>
posterodorsal posterodorsal
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis> <emphasis id="1F4680C995C71FF1BF205303C8C581DE" bold="true" pageId="0" pageNumber="145">E</emphasis>
ventral. Abbreviations: ventral. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">E1</emphasis> <emphasis id="6FD455BF9FE64CF90088FCE919E3A4CE" italics="true" pageId="0" pageNumber="145">E1</emphasis>
, ,
<emphasis italics="true" pageId="0" pageNumber="145">E2</emphasis> <emphasis id="F5E456EBF18C20F2D749F243DFCC1988" italics="true" pageId="0" pageNumber="145">E2</emphasis>
- pro- and retrolateral cymbial extensions, - pro- and retrolateral cymbial extensions,
<emphasis italics="true" pageId="0" pageNumber="145">Eb</emphasis> <emphasis id="73F44A3591D0F848AAADC2E43DBCABE8" italics="true" pageId="0" pageNumber="145">Eb</emphasis>
- embolar base, - embolar base,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis> <emphasis id="8914E1CB388CFA311B12599B98E5B6EE" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus, - embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Cf</emphasis> <emphasis id="FCBB04742C43E8E1425F5D5BE16154CA" italics="true" pageId="0" pageNumber="145">Cf</emphasis>
- cymbial fold, - cymbial fold,
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis> <emphasis id="F8AB0DE353115714D315D7D58D37219E" italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor, - conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="665D67F229A17A3CE9265F7FDF34BA56" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis, - median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis> <emphasis id="1AB049F4EA75B6D1E8D6F373DE692B3E" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral apophysis, - retrolateral apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis> <emphasis id="57D21FA36F097285395F6FAA56859437" italics="true" pageId="0" pageNumber="145">Va</emphasis>
- ventral apophysis. Scale bar: 0.1 mm. - ventral apophysis. Scale bar: 0.1 mm.
</paragraph> </paragraph>
</caption> </caption>
<paragraph pageId="0" pageNumber="145"> <paragraph id="72A015393BA1AE5F020C0066547762A9" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="15F98AD0B5C468FE520E43E0D5F7A39A" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig. Habitus as in Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5A</figureCitation> <figureCitation id="B3B8BD19BCB6CF8B443FD98ECDEF1A80" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5A</figureCitation>
. Total length 5.33. Carapace 2.18 long, 1.08 wide at pars cephalica, 1.51 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.06, ALE: 0.09, PME: 0.07, PLE: 0.09, PME-PME: 0.08. Coloration as in male, with lighter abdomen lacking a scutum. Measurements of legs: I: 5.36 (1.52, 0.60, 1.15, 1.09, 1.00), II: 4.76 (1.32, 0.61, 0.99, 1.04, 0.80), III: 4.53 (1.23, 0.60, 0.89, 1.11, 0.70), IV: 5.78 (1.42, 0.65, 1.38, 1.53, 0.80). . Total length 5.33. Carapace 2.18 long, 1.08 wide at pars cephalica, 1.51 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.06, ALE: 0.09, PME: 0.07, PLE: 0.09, PME-PME: 0.08. Coloration as in male, with lighter abdomen lacking a scutum. Measurements of legs: I: 5.36 (1.52, 0.60, 1.15, 1.09, 1.00), II: 4.76 (1.32, 0.61, 0.99, 1.04, 0.80), III: 4.53 (1.23, 0.60, 0.89, 1.11, 0.70), IV: 5.78 (1.42, 0.65, 1.38, 1.53, 0.80).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="3056C5EDBA73D20EC189F68B32DDFBC7" pageId="0" pageNumber="145">
Epigyne as in Fig. Epigyne as in Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">10A-E</figureCitation> <figureCitation id="BB633367CAF7C63B7DBC36FC87AB6411" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">10A-E</figureCitation>
. Trilobate, wider than long; median plate rectangular (with subparallel lateral margins), ca. 3.6 times thinner than lateral lobe; receptacles oval, wider than long, touching each other. . Trilobate, wider than long; median plate rectangular (with subparallel lateral margins), ca. 3.6 times thinner than lateral lobe; receptacles oval, wider than long, touching each other.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comments"> <subSubSection id="1463D875C6E761EAABF14C813291E7A6" pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="30AE261AD5D57CC7B0DA59FA00FFDFAE" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">Because the female specimen was not collected together with the male, its assignment to this species is tentative and shall be confirmed when both sexes are collected together.</paragraph> <paragraph id="546F50C8ABB5B6EDCA88C54C6AB26D1A" pageId="0" pageNumber="145">Because the female specimen was not collected together with the male, its assignment to this species is tentative and shall be confirmed when both sexes are collected together.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="35D7C190BC4558823E9907D38154835B" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="AF1CD7107E797C8720D28EB8FD63AF4C" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="D8F0C6555133DF602D8C0C63843CB8ED" pageId="0" pageNumber="145">
Known from the listed localities in Kermanshah and Lorestan provinces, western Iran (Fig. Known from the listed localities in Kermanshah and Lorestan provinces, western Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation> <figureCitation id="24678029B1CC77DD848DB4A51676B0D8" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

194
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title> <mods:title id="45A174980AA2739C8605A55C306B17F7">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation> <mods:affiliation id="B4383A2B208D41CD3C98685665622F9D">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="710C1096B1E6359288B4D30B22A78067" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="C3D6F054C8830C5E4C079B6D81F410E9" pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/92A3E44F-1EC2-4246-A21E-CAEED957E5BE" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parysatis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parysatis" status="sp. nov.">Acanthinozodium parysatis</taxonomicName> <taxonomicName id="B0CF97B07CA1BB44A90A5D8C8E9B2249" ID-CoL="98YYZ" LSID="http://zoobank.org/92A3E44F-1EC2-4246-A21E-CAEED957E5BE" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parysatis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parysatis" status="sp. nov.">Acanthinozodium parysatis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="F3D56AAD7FFA81BAF42B4ED9B8F40702" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11F, G</figureCitation> <figureCitation id="16C8113191A4D31363E2DA799FF022E1" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11F, G</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">, 13F-I</figureCitation> <figureCitation id="8977A59A092409F205C8E15536F19661" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">, 13F-I</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">, 15A-C</figureCitation> <figureCitation id="759AA99EBA06D17C16169D3652250537" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">, 15A-C</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">, 17D-F</figureCitation> <figureCitation id="FAFA5C11B574C9993100C82D0DC6AE95" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">, 17D-F</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation> <figureCitation id="579B00F5B1542548955E1113085100DA" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="E4E57C35F9435247E6485E166E20F4C9" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="C80CF9CF534A30FBB38A511CD2D4E563" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="01354254F13C81F772016A0E38275BCD" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="BA0486FADF27D48CD641D3642D2E10E3" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran: ♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Ardabil Province</emphasis> <emphasis id="A95D99A2435668CDA6F66DACF7FB037C" italics="true" pageId="0" pageNumber="145">Ardabil Province</emphasis>
: Kivi Pain, : Kivi Pain,
<geoCoordinate degrees="37" direction="north" minutes="41" orientation="latitude" precision="925" value="37.683334">37°41'N</geoCoordinate> <geoCoordinate id="78AB146C1718E9F7404819B0B32A427F" degrees="37" direction="north" minutes="41" orientation="latitude" precision="925" value="37.683334">37°41'N</geoCoordinate>
, ,
<geoCoordinate degrees="48" direction="east" minutes="21" orientation="longitude" precision="925" value="48.35">48°21'E</geoCoordinate> <geoCoordinate id="25685867874E2E161EB1D162C3E64A9F" degrees="48" direction="east" minutes="21" orientation="longitude" precision="925" value="48.35">48°21'E</geoCoordinate>
, 09.06.1975 (A. Senglet). , 09.06.1975 (A. Senglet).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis> <emphasis id="92B61AF3E8499BF0F73D19CE31654251" bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
: 1♂ (MHNG), same data as holotype; 3♀1♂ palp (MHNG), Iran: : 1♂ (MHNG), same data as holotype; 3♀1♂ palp (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Qazvin Province</emphasis> <emphasis id="2F87DA422F92C032966BF2F8F10E2AEC" italics="true" pageId="0" pageNumber="145">Qazvin Province</emphasis>
: Shahrak, : Shahrak,
<geoCoordinate degrees="36" direction="north" minutes="25" orientation="latitude" precision="925" value="36.416668">36°25'N</geoCoordinate> <geoCoordinate id="E0B47AD025A6F9DD61BFDB02D48D779A" degrees="36" direction="north" minutes="25" orientation="latitude" precision="925" value="36.416668">36°25'N</geoCoordinate>
, ,
<geoCoordinate degrees="50" direction="east" minutes="30" orientation="longitude" precision="925" value="50.5">50°30'E</geoCoordinate> <geoCoordinate id="7D8E80872B9A1120AE266C8D59B6439B" degrees="50" direction="east" minutes="30" orientation="longitude" precision="925" value="50.5">50°30'E</geoCoordinate>
, 02.07.1975 (A. Senglet). , 02.07.1975 (A. Senglet).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="A34E98BD1C225DB0158C24996C6B97FC" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="F08FF252E7D44D5D13D8AF387CFB97E5" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="06CD9B99D8C91855B6B1C42408917877" pageId="0" pageNumber="145">
The specific epithet is an ancient Persian feminine name, meaning The specific epithet is an ancient Persian feminine name, meaning
<normalizedToken originalValue="“fairy-like”">&quot;fairy-like&quot;</normalizedToken> <normalizedToken id="31E46F1118ED8BE76FFF0C38179D999E" originalValue="“fairy-like”">&quot;fairy-like&quot;</normalizedToken>
. Noun in apposition. . Noun in apposition.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="990F482248FA94F3BDD9B9663E06BBBE" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="7E576DCD11E03AF8048BCE89BE79BA28" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="1FDFE37AB113D307907F212B59530A27" pageId="0" pageNumber="145">
The male palp of The male palp of
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis"> <taxonomicName id="92C774392E21B76DFFCF0DC5241C3D48" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis> <emphasis id="C83ABA4657307891CF5FC27956BA2897" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. is similar to that of sp. nov. is similar to that of
<taxonomicName lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha"> <taxonomicName id="ACB5569446C1C0741D46845A1B4650E2" lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">A. niusha</emphasis> <emphasis id="151A5E8C2372E3B253F6D97311253EE1" italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. by the overall shape of the median apophysis and RTA but differs in lacking a retrolateral extension on the median apophysis ( sp. nov. by the overall shape of the median apophysis and RTA but differs in lacking a retrolateral extension on the median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="F771AC476CBDF63411F87BD6F7778669" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
present) and by having a shorter, stouter RTA ( present) and by having a shorter, stouter RTA (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="A4A37C802A0F5C6DBA5D6733D4FD5A2B" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
longer and tapering). The epigyne of the new species is most similar to that of longer and tapering). The epigyne of the new species is most similar to that of
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa"> <taxonomicName id="491CCEE06597B3E74934C9D35C639FA8" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis> <emphasis id="848C1DAD3B45628398DC41FF17D2E2E3" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. in having a small anterior hood and thin furrow between the lateral lobes but differs by the relative length of the anterior part of the fovea being shorter than the bursae in sp. nov. in having a small anterior hood and thin furrow between the lateral lobes but differs by the relative length of the anterior part of the fovea being shorter than the bursae in
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis"> <taxonomicName id="61EA29D866B339409A5D965EFC68776D" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis> <emphasis id="5809651E95D21BB24EE707AD11161FAB" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="1973EBC66AB4FBB5507C6F75759AEB3E" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
as long as bursae). as long as bursae).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="45A77B456D0EF09C7EA08C278F42A4B4" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="1C4FBD733AD12FE285EEAED15AE1E793" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="FA14FAC4ACB233964430B5B7A96F3B05" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis> <emphasis id="8C4A7B130C158834F76370F399923DD9" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig. (holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11F</figureCitation> <figureCitation id="9599B7FD5FD6ABA6798F4BC92305C07F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11F</figureCitation>
. Total length 2.23. Carapace 1.03 long, 0.46 wide at pars cephalica, 0.80 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.10, ALE: 0.07, PME: 0.06, PLE: 0.05, PME-PME: 0.10. Carapace, sternum, labium, chelicerae and maxillae yellowish. Legs yellowish, slightly darker at femora, without annulations. Abdomen dorsally grayish, pale ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 2.98+missing tarsus (0.95, 0.37, 0.80, 0.86, missing), II: 3.13 (0.77, 0.34, 0.66, 0.79, 0.57), III: 3.03 (0.76, 0.35, 0.58, 0.86, 0.48), IV: 3.44 (1.00, 0.30, 0.73, 0.85, 0.56). . Total length 2.23. Carapace 1.03 long, 0.46 wide at pars cephalica, 0.80 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.10, ALE: 0.07, PME: 0.06, PLE: 0.05, PME-PME: 0.10. Carapace, sternum, labium, chelicerae and maxillae yellowish. Legs yellowish, slightly darker at femora, without annulations. Abdomen dorsally grayish, pale ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 2.98+missing tarsus (0.95, 0.37, 0.80, 0.86, missing), II: 3.13 (0.77, 0.34, 0.66, 0.79, 0.57), III: 3.03 (0.76, 0.35, 0.58, 0.86, 0.48), IV: 3.44 (1.00, 0.30, 0.73, 0.85, 0.56).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="2FE005722D4D99DF9E855D94695FFDFF" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13F-I</figureCitation> <figureCitation id="68F2997448A20660AE9B2880D7DD653E" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13F-I</figureCitation>
, ,
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">15A-C</figureCitation> <figureCitation id="8CCD482CE6AC281582C13C3DD31F6E4D" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">15A-C</figureCitation>
. RTA ( . RTA (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis> <emphasis id="4D4BFDD561F06B5D3781AB64C0FE0989" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
) relatively short and stout, with a finger-like projection dorsally; tegulum with posterior process, sperm duct almost straight retrolaterally, and gently bent along prolateral side; median apophysis ( ) relatively short and stout, with a finger-like projection dorsally; tegulum with posterior process, sperm duct almost straight retrolaterally, and gently bent along prolateral side; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="3C8FFC1F0B895439C5D3C23F41E350C4" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) wider than long; embolus broad basally, originating at about the 7 ) wider than long; embolus broad basally, originating at about the 7
<normalizedToken originalValue="o’clock">o'clock</normalizedToken> <normalizedToken id="FF9DDF36E5487C3A7137544BAFEE5783" originalValue="o’clock">o'clock</normalizedToken>
position; embolus proper thin, slightly twisted apically. position; embolus proper thin, slightly twisted apically.
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="BCA1250AEFFA58DE11D1B61CBA4D44C8" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="9BF32961AF48833828A91473B6738BE7" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig. Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11G</figureCitation> <figureCitation id="13C7454F66C63617786B917045215864" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11G</figureCitation>
. Total length 3.68. Carapace 1.34 long, 0.60 wide at pars cephalica, 0.94 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.09, ALE: 0.08, PME: 0.07, PLE: 0.07, PME-PME: 0.16. Coloration generally as in male. Abdomen dorsally with a pale median band occupying half of the . Total length 3.68. Carapace 1.34 long, 0.60 wide at pars cephalica, 0.94 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.09, ALE: 0.08, PME: 0.07, PLE: 0.07, PME-PME: 0.16. Coloration generally as in male. Abdomen dorsally with a pale median band occupying half of the
<normalizedToken originalValue="abdomen’s">abdomen's</normalizedToken> <normalizedToken id="ED366AC177A790178F6AC06B53C5490D" originalValue="abdomen’s">abdomen's</normalizedToken>
length. Measurements of legs: I: 3.79 (0.90, 0.40, 0.84, 1.01, 0.64), II: 2.90+missing tarsus (0.94, 0.36, 0.74, 0.86, missing), III: 3.48 (0.88, 0.40, 0.65, 1.03, 0.52), IV: 5.09 (1.33, 0.48, 1.13, 1.47, 0.68). length. Measurements of legs: I: 3.79 (0.90, 0.40, 0.84, 1.01, 0.64), II: 2.90+missing tarsus (0.94, 0.36, 0.74, 0.86, missing), III: 3.48 (0.88, 0.40, 0.65, 1.03, 0.52), IV: 5.09 (1.33, 0.48, 1.13, 1.47, 0.68).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="C2BD51BA32818908C58F75BC2248D7ED" pageId="0" pageNumber="145">
Epigyne as in Fig. Epigyne as in Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">17D-F</figureCitation> <figureCitation id="8AEF5553FB3E4C20833A538C6CDDCF7E" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">17D-F</figureCitation>
. Fovea triangular and small anterior hood present; bursae round, separated by less than one of their radii; receptacles smaller than bursae, triangular, separated by more than 3 diameters. . Fovea triangular and small anterior hood present; bursae round, separated by less than one of their radii; receptacles smaller than bursae, triangular, separated by more than 3 diameters.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="F0656C9F1D57EB41FCBCDB4FAEFA514C" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="B1D2D450F40816836FC1A08C326FCC9C" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="DADD1DB87BFB4BE7D0285FBDF71A11B2" pageId="0" pageNumber="145">
Known only from the listed localities in Ardabil and Qazvin provinces, northern and northwestern Iran (Fig. Known only from the listed localities in Ardabil and Qazvin provinces, northern and northwestern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation> <figureCitation id="7C6319360F031C736F87DE4423CD4627" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

106
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<treatment ID-GBIF-Taxon="181753557" LSID="urn:lsid:plazi:treatment:44378935A45655B4B53C83F1B267BCA3" httpUri="http://treatment.plazi.org/id/44378935A45655B4B53C83F1B267BCA3" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="44378935A45655B4B53C83F1B267BCA3" ID-GBIF-Taxon="181753557" LSID="urn:lsid:plazi:treatment:44378935A45655B4B53C83F1B267BCA3" httpUri="http://treatment.plazi.org/id/44378935A45655B4B53C83F1B267BCA3" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
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<paragraph pageId="0" pageNumber="145"> <paragraph id="D22AA37E3FB2795368BCF3A41969F00E" pageId="0" pageNumber="145">
Genus Genus
<taxonomicName LSID="44378935-A456-55B4-B53C-83F1B267BCA3" authority="Simon, 1882" authorityName="Simon" authorityYear="1882" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Trygetus Simon, 1882</taxonomicName> <taxonomicName id="080C7D5194E576B13BF80EA8C522CB67" ID-CoL="8398" LSID="44378935-A456-55B4-B53C-83F1B267BCA3" authority="Simon, 1882" authorityName="Simon" authorityYear="1882" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Trygetus Simon, 1882</taxonomicName>
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<subSubSection pageId="0" pageNumber="145" type="type species"> <subSubSection id="4225875D165B52587AB5302DCD17B8A5" pageId="0" pageNumber="145" type="type species">
<paragraph pageId="0" pageNumber="145">Type species.</paragraph> <paragraph id="C5F6C0FDFA69213EFCD438D12BECCC40" pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="BAA33531A4A8519017F0121489AE6154" pageId="0" pageNumber="145">
<emphasis italics="true" pageId="0" pageNumber="145">Palaestina sexoculata</emphasis> <emphasis id="E118E56D3F3BB975B766C071E6161012" italics="true" pageId="0" pageNumber="145">Palaestina sexoculata</emphasis>
O. Pickard-Cambridge, 1872. O. Pickard-Cambridge, 1872.
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="comments"> <subSubSection id="D342005E66B0AB1523B7BEA0B38F0933" pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="98B70148162BFF81634390B0536C41FB" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="95FFFA5517439B969B827BD56C2B6DD8" pageId="0" pageNumber="145">
Small genus with seven named species distributed from Morocco to Turkmenistan ( Small genus with seven named species distributed from Morocco to Turkmenistan (
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). ).
<taxonomicName authorityName="Simon" authorityYear="1882" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="6D5D5A83E1D646320C719631BABDF95F" authorityName="Simon" authorityYear="1882" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Trygetus</emphasis> <emphasis id="161162C68967A0FB17DEDC2AB4C04583" italics="true" pageId="0" pageNumber="145">Trygetus</emphasis>
</taxonomicName> </taxonomicName>
differs from all other differs from all other
<taxonomicName authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName> <taxonomicName id="8FECBC5F65D88E493E228EAC478E636C" authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName>
from the region by having only six eyes ( from the region by having only six eyes (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="54D599146C4B8B9781C670BC12A39892" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
eight eyes). eight eyes).
</paragraph> </paragraph>
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292
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2021" docId="46CFFF9DFBBB518D9F9802AC3F372F3E" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Acanthinozodium elburzicum Zamani &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="61538C61-2E8A-4F21-8289-B9129E5E4965" docUuidSource="ZooBank" docVersion="4" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1668150222111" updateUser="ExternalLinkService"> <document ID-CLB-Dataset="7157" ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2021" docId="46CFFF9DFBBB518D9F9802AC3F372F3E" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Acanthinozodium elburzicum Zamani &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="61538C61-2E8A-4F21-8289-B9129E5E4965" docUuidSource="ZooBank" docVersion="6" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1732670661048" updateUser="ExternalLinkService">
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title> <mods:title id="A38019F11B117BA127B88E498DF540AE">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:namePart>Zamani, Alireza</mods:namePart> <mods:namePart id="E7856979368A2687B2EBE3BED43E78FC">Zamani, Alireza</mods:namePart>
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<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation> <mods:affiliation id="A251374753B04F3F052A6809169541FA">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation> <mods:affiliation id="E3715C06174D4A1179E56FC4C7172D13">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<treatment ID-GBIF-Taxon="181753563" LSID="urn:lsid:zoobank.org:act:61538C61-2E8A-4F21-8289-B9129E5E4965" httpUri="http://treatment.plazi.org/id/46CFFF9DFBBB518D9F9802AC3F372F3E" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="46CFFF9DFBBB518D9F9802AC3F372F3E" ID-GBIF-Taxon="181753563" LSID="urn:lsid:zoobank.org:act:61538C61-2E8A-4F21-8289-B9129E5E4965" httpUri="http://treatment.plazi.org/id/46CFFF9DFBBB518D9F9802AC3F372F3E" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="1D6DA4437D3858492767754384B713FB" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="E473E90F8921936BBFE2B84C5B05CB01" pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/61538C61-2E8A-4F21-8289-B9129E5E4965" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum" status="sp. nov.">Acanthinozodium elburzicum</taxonomicName> <taxonomicName id="7023E6A2F9451297FC86C8258A779A78" ID-CoL="98YYY" LSID="http://zoobank.org/61538C61-2E8A-4F21-8289-B9129E5E4965" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum" status="sp. nov.">Acanthinozodium elburzicum</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="395B023899DE4FA4338A5CECCC86C3E7" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18E-J</figureCitation> <figureCitation id="4C029FE81556B1EAEA47FB549AE0A965" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18E-J</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">, 21D-F</figureCitation> <figureCitation id="584DB1B2F16FF64505E054CEB2A27027" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">, 21D-F</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">, 23A-C</figureCitation> <figureCitation id="C1546E0A71419358DF879FD971DB6709" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">, 23A-C</figureCitation>
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">, 26E</figureCitation> <figureCitation id="BC31179AFAF5928B552DC95378D80C0B" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">, 26E</figureCitation>
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A) and Z. talyschicum (B, C) A, B ventral C ventroretrolateral. Abbreviations: Tm - median tegular process, Tp - prolateral tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145">, 29A</figureCitation> <figureCitation id="9CA97665CFE184DD20B27D81F89ECC29" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A) and Z. talyschicum (B, C) A, B ventral C ventroretrolateral. Abbreviations: Tm - median tegular process, Tp - prolateral tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145">, 29A</figureCitation>
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">, 30A-E</figureCitation> <figureCitation id="CF7138F7C18FDEDB7FA4997166DAF58C" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">, 30A-E</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation> <figureCitation id="E242524D9F8D2CB21563728105A81B31" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="07308D6308FDEDA492C38A2650E62B68" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="F606FBA3E43ACC5A377F34392DF084D0" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="EBA2979CFE1E57E2B3216E6CADFD67B1" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="CC7F71C239AB57514439A7A94E773BD6" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran: ♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Tehran Province</emphasis> <emphasis id="96E5BDF818EBDEEA05798DB5D8488C82" italics="true" pageId="0" pageNumber="145">Tehran Province</emphasis>
: Jamshidieh Mts., : Jamshidieh Mts.,
<geoCoordinate degrees="35" direction="north" minutes="49" orientation="latitude" precision="925" value="35.816666">35°49'N</geoCoordinate> <geoCoordinate id="23C501F238E454D8CAE637FF0E289560" degrees="35" direction="north" minutes="49" orientation="latitude" precision="925" value="35.816666">35°49'N</geoCoordinate>
, ,
<geoCoordinate degrees="51" direction="east" minutes="27" orientation="longitude" precision="925" value="51.45">51°27'E</geoCoordinate> <geoCoordinate id="BCB2C90BE131202CF857756174C80D55" degrees="51" direction="east" minutes="27" orientation="longitude" precision="925" value="51.45">51°27'E</geoCoordinate>
, 05.2015 (A. Zamani). , 05.2015 (A. Zamani).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis> <emphasis id="D74E407E95E5E4477087FC2563F3982E" bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
: 1♀ (MHNG), Pardisan Park, : 1♀ (MHNG), Pardisan Park,
<geoCoordinate degrees="35" direction="north" minutes="44" orientation="latitude" precision="925" value="35.733334">35°44'N</geoCoordinate> <geoCoordinate id="E52D035BA041019E308DB89674199BD6" degrees="35" direction="north" minutes="44" orientation="latitude" precision="925" value="35.733334">35°44'N</geoCoordinate>
, ,
<geoCoordinate degrees="51" direction="east" minutes="21" orientation="longitude" precision="925" value="51.35">51°21'E</geoCoordinate> <geoCoordinate id="B04774AEA13526854C63F4221BA170D6" degrees="51" direction="east" minutes="21" orientation="longitude" precision="925" value="51.35">51°21'E</geoCoordinate>
, 05.2015 (A. Zamani); 5♂2♀1j. (MMUE), Latian Dam, , 05.2015 (A. Zamani); 5♂2♀1j. (MMUE), Latian Dam,
<geoCoordinate degrees="35" direction="north" minutes="48" orientation="latitude" precision="925" value="35.8">35°48'N</geoCoordinate> <geoCoordinate id="40F37F5E735EF70D0BEE18C1ED071F57" degrees="35" direction="north" minutes="48" orientation="latitude" precision="925" value="35.8">35°48'N</geoCoordinate>
, ,
<geoCoordinate degrees="51" direction="east" minutes="08" orientation="longitude" precision="925" value="51.133335">51°08'E</geoCoordinate> <geoCoordinate id="D0BA311D1F38119233FB1CDF95050A3B" degrees="51" direction="east" minutes="08" orientation="longitude" precision="925" value="51.133335">51°08'E</geoCoordinate>
, 06-19.06.2000 (Y.M. Marusik); 3♂1♀ (MMUE), Plant Protection Institute, , 06-19.06.2000 (Y.M. Marusik); 3♂1♀ (MMUE), Plant Protection Institute,
<geoCoordinate degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate> <geoCoordinate id="D46550EA77C95011C92CB5AB86DBA381" degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate>
, ,
<geoCoordinate degrees="51" direction="east" minutes="24" orientation="longitude" precision="925" value="51.4">51°24'E</geoCoordinate> <geoCoordinate id="822BC5231985F4E929BC55D0B20AED46" degrees="51" direction="east" minutes="24" orientation="longitude" precision="925" value="51.4">51°24'E</geoCoordinate>
, 07-22.06.2000 (Y.M. Marusik). , 07-22.06.2000 (Y.M. Marusik).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="F8916519217126D033753EF36FB45AC6" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="CB642AE001CF090E7A17E3A035C92B64" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet refers to the Elburz Mountain Range, in which the type locality of the new species is situated.</paragraph> <paragraph id="25F1CBE3207C58F396E66BE2B90A9AA1" pageId="0" pageNumber="145">The specific epithet refers to the Elburz Mountain Range, in which the type locality of the new species is situated.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="9320EA437358EE5F65D8F97D435D892F" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="081A97DEF09E5A2484C2D84B9F57980C" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="749D61135B05659A5E1694F6474B1F4B" pageId="0" pageNumber="145">
The male of the new species is similar to The male of the new species is similar to
<taxonomicName lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara"> <taxonomicName id="E87FD4685DCCFE50917B0F45F75EA6C5" lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">A. diara</emphasis> <emphasis id="7AE864ED682160F396AB8FE7E38D9154" italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. by the similar shape of the RTA and they both have a small retrolateral indentation apically on the conductor, but differs by the shape of the median apophysis, which is with a distinctly larger prolateral outgrowth. The female of the new species is most similar to sp. nov. by the similar shape of the RTA and they both have a small retrolateral indentation apically on the conductor, but differs by the shape of the median apophysis, which is with a distinctly larger prolateral outgrowth. The female of the new species is most similar to
<taxonomicName lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa"> <taxonomicName id="A723BEF55FE15DB80B0C1B606C874DB0" lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis> <emphasis id="BD26450319666D6673C3A4AC8AD86412" italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. but differs by the epigynal hood being wider than long ( sp. nov. but differs by the epigynal hood being wider than long (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="4F11D00D21B503DF07EBA4CA64F504CA" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
longer than wide) and receptacles separated by about 4 diameters ( longer than wide) and receptacles separated by about 4 diameters (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="1C01DD9D15C324E1D24AC553D63CEAF1" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
ca. 2.5). ca. 2.5).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="39DFF3643BA8C32BDD2006B4472632CB" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="9C96C772F983C9A460EFB6D522CD2D4B" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="B90ED37053D338F366C0CABE012CF7B0" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis> <emphasis id="B6A390A47BB7CB7EC04E84629D051C41" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig. (holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18F-H</figureCitation> <figureCitation id="1F2CCBE7963C8D15BCF04AE19888B61D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18F-H</figureCitation>
(holotype: 18F). Total length 4.30. Carapace 2.08 long, 0.97 wide at pars cephalica, 1.49 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.14, ALE: 0.12, PME: 0.08, PLE: 0.09, PME-PME: 0.15. Carapace dark brown, lighter in pars cephalica, with irregular dark patches and lines. Sternum, labium and maxillae light brown. Chelicera (Fig. (holotype: 18F). Total length 4.30. Carapace 2.08 long, 0.97 wide at pars cephalica, 1.49 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.14, ALE: 0.12, PME: 0.08, PLE: 0.09, PME-PME: 0.15. Carapace dark brown, lighter in pars cephalica, with irregular dark patches and lines. Sternum, labium and maxillae light brown. Chelicera (Fig.
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">26E</figureCitation> <figureCitation id="183FBEB2670163B7579AC3A1D5B02BCA" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">26E</figureCitation>
) dark brown, with one retromarginal tooth. Legs slightly lighter than carapace, without annulations. Abdomen dorsally black and with large scutum; ventrally grayish, without any pattern. Spinnerets pale, uniform in color. Measurements of legs: I: 7.52 (1.79, 0.71, 1.75, 2.03, 1.24), II: 6.79 (1.87, 0.69, 1.40, 1.83, 1.00), III: 6.90 (1.78, 0.74, 1.40, 2.06, 0.92), IV: 9.19 (2.48, 0.83, 2.15, 2.84, 0.89). ) dark brown, with one retromarginal tooth. Legs slightly lighter than carapace, without annulations. Abdomen dorsally black and with large scutum; ventrally grayish, without any pattern. Spinnerets pale, uniform in color. Measurements of legs: I: 7.52 (1.79, 0.71, 1.75, 2.03, 1.24), II: 6.79 (1.87, 0.69, 1.40, 1.83, 1.00), III: 6.90 (1.78, 0.74, 1.40, 2.06, 0.92), IV: 9.19 (2.48, 0.83, 2.15, 2.84, 0.89).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="0DB44EBAC7B0F53D0B92F2BB1A72A65E" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">21D-F</figureCitation> <figureCitation id="FA9DD1CFBD204BF2D6A8A9A2D4388570" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">21D-F</figureCitation>
, ,
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">23A-C</figureCitation> <figureCitation id="2B33F7E88A321A4A311DD30B1142321C" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">23A-C</figureCitation>
, ,
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A) and Z. talyschicum (B, C) A, B ventral C ventroretrolateral. Abbreviations: Tm - median tegular process, Tp - prolateral tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145">29A</figureCitation> <figureCitation id="676309995796844BEC863B6331E7DDDC" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A) and Z. talyschicum (B, C) A, B ventral C ventroretrolateral. Abbreviations: Tm - median tegular process, Tp - prolateral tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145">29A</figureCitation>
. RTA ( . RTA (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis> <emphasis id="F30A7D59E729E35CFC444D7766CFEBF7" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
) long and conical, with a small projection apically; ventral apophysis ( ) long and conical, with a small projection apically; ventral apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis> <emphasis id="B60A025450910F8C2375DB0CCCA0FEC4" italics="true" pageId="0" pageNumber="145">Va</emphasis>
) small and conical; tegulum with posterior process; median apophysis ( ) small and conical; tegulum with posterior process; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="2BB883B7F43C577E6573718BD8BEE40A" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) wider than long, with outgrowths on all four sides, the triangular prolateral one the largest; embolus ( ) wider than long, with outgrowths on all four sides, the triangular prolateral one the largest; embolus (
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis> <emphasis id="6E5C53A4A8FD7692DF30EAFCD609E63A" italics="true" pageId="0" pageNumber="145">Em</emphasis>
) originating at about the 7 ) originating at about the 7
<normalizedToken originalValue="o’clock">o'clock</normalizedToken> <normalizedToken id="D09D6AE6C103A9CF954A04CF3E7F7853" originalValue="o’clock">o'clock</normalizedToken>
position; embolus proper thin and steadily curving. position; embolus proper thin and steadily curving.
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145" start="Figure 21" startId="F21"> <caption id="FD0299D2E0D123CDBA7F7F8F442B9C62" doi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145" start="Figure 21" startId="F21">
<paragraph pageId="0" pageNumber="145"> <paragraph id="D0C1A14DA7DC86FA6122E40463CA5A49" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 21.</emphasis> <emphasis id="7D2517EE328FAD4E27F6695D4AFA0FC9" bold="true" pageId="0" pageNumber="145">Figure 21.</emphasis>
Male palps of Male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium dorsa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dorsa"> <taxonomicName id="A36D0ED871E6FCC911BD6C9F9FD3B7ED" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium dorsa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium dorsa</emphasis> <emphasis id="E723928E768CE7EDC12F58AEE885208B" italics="true" pageId="0" pageNumber="145">Acanthinozodium dorsa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis> <emphasis id="B446F6C1E5BDFAB43EC809A7E2C9DE42" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and ) and
<taxonomicName lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum"> <taxonomicName id="23DC8CD2B924E7CA9C4CA1D13304DB9E" lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis> <emphasis id="F4BFF92F33B9E38896C6F08200AD376C" italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis> <emphasis id="78C0803222DEEE0620E5C2B586D7E761" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis> <emphasis id="273ABC0AC32DC4CE12D5ACDA8ACBF13C" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, E</emphasis> <emphasis id="5A13DC765DE318E68F54CEE14FFBE607" bold="true" pageId="0" pageNumber="145">B, E</emphasis>
retrolateral retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, F</emphasis> <emphasis id="35A81FCB17E98629C9EFBD53615B1306" bold="true" pageId="0" pageNumber="145">C, F</emphasis>
dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm. dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm.
</paragraph> </paragraph>
</caption> </caption>
<caption doi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145" start="Figure 22" startId="F22"> <caption id="57137E8BBB07B68EB05BB4B5CD0010EA" doi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145" start="Figure 22" startId="F22">
<paragraph pageId="0" pageNumber="145"> <paragraph id="D00B7F15A9C19E20F49547E10DF8DB96" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 22.</emphasis> <emphasis id="71C18DED5B8D9050ED552DBE06B14510" bold="true" pageId="0" pageNumber="145">Figure 22.</emphasis>
SEM images of the male palps of SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita"> <taxonomicName id="EF049C19027CA8C6CAFD0F238A458F2D" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium armita</emphasis> <emphasis id="AAC0ACEB3F6CB319E8E2E53CB202FBF4" italics="true" pageId="0" pageNumber="145">Acanthinozodium armita</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C, F</emphasis> <emphasis id="3EDC37CA45B74E7B2A6FFCFCA472F4CD" bold="true" pageId="0" pageNumber="145">A-C, F</emphasis>
) and ) and
<taxonomicName lsidName="A. ovtchinnikovi" pageId="0" pageNumber="145" rank="species" species="ovtchinnikovi"> <taxonomicName id="05100D9169462A4D940A986302F5083F" lsidName="A. ovtchinnikovi" pageId="0" pageNumber="145" rank="species" species="ovtchinnikovi">
<emphasis italics="true" pageId="0" pageNumber="145">A. ovtchinnikovi</emphasis> <emphasis id="AA16193EFEA282FFA7CCBBDD42F1FFBE" italics="true" pageId="0" pageNumber="145">A. ovtchinnikovi</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D, E, G-I</emphasis> <emphasis id="E6ECE5ED7C0497E5A411408789920177" bold="true" pageId="0" pageNumber="145">D, E, G-I</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, H</emphasis> <emphasis id="17BF74C7D2E23B6C755389FDF8CA2810" bold="true" pageId="0" pageNumber="145">A, H</emphasis>
retroventral and retrolateral retroventral and retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">B, C, G</emphasis> <emphasis id="63E2C82DB4EDD0D6BBADCD0BE69B5420" bold="true" pageId="0" pageNumber="145">B, C, G</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis> <emphasis id="AC291749C22F70F3A3963B240140F13F" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
proventral, apicoproventral and apicoventral proventral, apicoproventral and apicoventral
<emphasis bold="true" pageId="0" pageNumber="145">I</emphasis> <emphasis id="F2A871B60682A7AC2E14EA826E8D4C44" bold="true" pageId="0" pageNumber="145">I</emphasis>
retrolateral tibial and cymbial groove. Abbreviations: retrolateral tibial and cymbial groove. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Cg</emphasis> <emphasis id="01F5318B65CD310AE1099B1B6F88A808" italics="true" pageId="0" pageNumber="145">Cg</emphasis>
- cymbial groove, - cymbial groove,
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis> <emphasis id="61633E5DA08084AA722EE0A8ED808597" italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor, - conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis> <emphasis id="F7FAD983499BB03AF0267B165486E679" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus, - embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Er</emphasis> <emphasis id="7D21BFF826D0CFA33E34BEECF27EC1EC" italics="true" pageId="0" pageNumber="145">Er</emphasis>
-embolar ridge, -embolar ridge,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="FB3DCA943A77C7FAF078CF760D2C6039" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis. Scale bars: 0.1 mm, unless stated otherwise. - median apophysis. Scale bars: 0.1 mm, unless stated otherwise.
</paragraph> </paragraph>
</caption> </caption>
<caption doi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145" start="Figure 23" startId="F23"> <caption id="B902C497AE657AD99572C81724BC3FA7" doi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145" start="Figure 23" startId="F23">
<paragraph pageId="0" pageNumber="145"> <paragraph id="35B1D993267AC875B7C514CE04E1D181" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 23.</emphasis> <emphasis id="B91B3C98DE41BB3200BEA86E34A86D35" bold="true" pageId="0" pageNumber="145">Figure 23.</emphasis>
SEM images of the male palps of SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum"> <taxonomicName id="93AAD109AFF9C0E0FDF6446EAAC5439F" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis> <emphasis id="A6E50972C2E924D50098EB727C744D63" italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis> <emphasis id="8679548182CE2C3892293B2F5B8FEB59" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and ) and
<taxonomicName lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara"> <taxonomicName id="0F49720ABABE31B140127428A8E1ED33" lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">A. diara</emphasis> <emphasis id="DED5A56804CB8F14DC5E549E53D72244" italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis> <emphasis id="181307FBA7B6183BCF071CEF02506BF8" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis> <emphasis id="D82E4596E186A24A212E8356F78BBF34" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, F</emphasis> <emphasis id="EE63ADFCC1909A78F7AA1FA1EF2EAD47" bold="true" pageId="0" pageNumber="145">B, F</emphasis>
retrolateral retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, E</emphasis> <emphasis id="D0A77B3E7B181DC443170E7D06213193" bold="true" pageId="0" pageNumber="145">C, E</emphasis>
dorsoretrolateral. Abbreviations: dorsoretrolateral. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Cg</emphasis> <emphasis id="AF6142E31FC2B60229C4E0BE99DBCE19" italics="true" pageId="0" pageNumber="145">Cg</emphasis>
- cymbial groove, - cymbial groove,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis> <emphasis id="168BBC39282E9B7FC6C871F3E9ED2932" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus, - embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="A68391B813E10C141E4672DD77EC285B" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis, - median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis> <emphasis id="61D9AB1DC163A99F1DC1FE98C91C5841" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral tibial apophysis, - retrolateral tibial apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis> <emphasis id="BA8DA243AE84501EC08FD9867084947A" italics="true" pageId="0" pageNumber="145">Va</emphasis>
- ventral apophysis. Scale bars: 0.1 mm. - ventral apophysis. Scale bars: 0.1 mm.
</paragraph> </paragraph>
</caption> </caption>
<paragraph pageId="0" pageNumber="145"> <paragraph id="9E76A0D88821D9FE99D5094CB3650B6F" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="3FCDB9674FEDD74F16934D7AAD2AED7E" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig. Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18E, I, J</figureCitation> <figureCitation id="3C1AEA71E60652EAC3A6A0AC93445058" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18E, I, J</figureCitation>
. Total length 4.95. Carapace 1.95 long, 0.96 wide at pars cephalica, 1.40 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.15, ALE: 0.12, PME: 0.08, PLE: 0.11, PME-PME: 0.15. Coloration as in male, with paler abdomen lacking a scutum. Measurements of legs: I: 6.54 (1.50, 0.70, 1.44, 1.76, 1.14), II: 6.27 (1.59, 0.67, 1.33, 1.67, 1.01), III: 6.23 (1.60, 0.71, 1.21, 1.74, 0.97), IV: 8.11 (2.04, 0.78, 1.81, 2.46, 1.02). . Total length 4.95. Carapace 1.95 long, 0.96 wide at pars cephalica, 1.40 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.15, ALE: 0.12, PME: 0.08, PLE: 0.11, PME-PME: 0.15. Coloration as in male, with paler abdomen lacking a scutum. Measurements of legs: I: 6.54 (1.50, 0.70, 1.44, 1.76, 1.14), II: 6.27 (1.59, 0.67, 1.33, 1.67, 1.01), III: 6.23 (1.60, 0.71, 1.21, 1.74, 0.97), IV: 8.11 (2.04, 0.78, 1.81, 2.46, 1.02).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="10EF5E79F3675EC7C4DC52329CA11209" pageId="0" pageNumber="145">
Epigyne as in Fig. Epigyne as in Fig.
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">30A-E</figureCitation> <figureCitation id="83462842097947057366789C1BA0D156" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">30A-E</figureCitation>
. Epigynal plater over 3 times wider than long, lacking fovea; anterior hood wider than long; receptacles with rounded posterior parts, separated by about 4 diameters. . Epigynal plater over 3 times wider than long, lacking fovea; anterior hood wider than long; receptacles with rounded posterior parts, separated by about 4 diameters.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="BB59AEA44A54F966CB5675AB8CAA633A" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="5C5237BB14756546E8F1048FB6EC87BB" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="CC29874BFBF5564357F8D36F7FBD0272" pageId="0" pageNumber="145">
Known from the listed localities in Tehran Province, northern Iran (Fig. Known from the listed localities in Tehran Province, northern Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation> <figureCitation id="359050A6D28A92256041F999BCBCA561" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

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<treatment ID-GBIF-Taxon="181753552" LSID="urn:lsid:plazi:treatment:5E0B26BFC08A592CA10A3AA52539B4A6" httpUri="http://treatment.plazi.org/id/5E0B26BFC08A592CA10A3AA52539B4A6" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="5E0B26BFC08A592CA10A3AA52539B4A6" ID-GBIF-Taxon="181753552" LSID="urn:lsid:plazi:treatment:5E0B26BFC08A592CA10A3AA52539B4A6" httpUri="http://treatment.plazi.org/id/5E0B26BFC08A592CA10A3AA52539B4A6" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Storeninae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="8D40E30F2AA16F3747D7F8D6EDB7F11E" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="F85315FAEC9896D9AD2E0A7107581AEE" pageId="0" pageNumber="145">
Genus Genus
<taxonomicName LSID="5E0B26BF-C08A-592C-A10A-3AA52539B4A6" authority="Levy, 1990" authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Pax Levy, 1990</taxonomicName> <taxonomicName id="70AE80238B2F210F857777938765E159" ID-CoL="6JSM" LSID="5E0B26BF-C08A-592C-A10A-3AA52539B4A6" authority="Levy, 1990" authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Pax Levy, 1990</taxonomicName>
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<subSubSection pageId="0" pageNumber="145" type="type species"> <subSubSection id="14CCFAA886F98A222C26D9F216BF1F03" pageId="0" pageNumber="145" type="type species">
<paragraph pageId="0" pageNumber="145">Type species.</paragraph> <paragraph id="8C8FEE0B79EB33851CCCCE67A758EC72" pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="4BD7A0187B2F8F3712CA0A8702E54E8B" pageId="0" pageNumber="145">
<taxonomicName class="Arachnida" family="Zodariidae" genus="Habronestes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Habronestes libani" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="libani"> <taxonomicName id="335B70F12A999C702EC374F86A79D1F5" class="Arachnida" family="Zodariidae" genus="Habronestes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Habronestes libani" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="libani">
<emphasis italics="true" pageId="0" pageNumber="145">Habronestes libani</emphasis> <emphasis id="C6EBF2AFF57E5A78A94A509EC1488888" italics="true" pageId="0" pageNumber="145">Habronestes libani</emphasis>
</taxonomicName> </taxonomicName>
Simon, 1873 from Lebanon. Simon, 1873 from Lebanon.
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="comments"> <subSubSection id="9B5FDEDA8032FAC478F4426F32DFD446" pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="FC9228D6F4183657C9F1A9B58FB3AD1D" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">This is a small genus with seven species distributed exclusively in the Middle East from Turkey and Israel to Iran (including the two new species described below, representing the first record of this genus in this country). Members of this genus can be easily distinguished from all other zodariids found in the study area by the ovoid carapace lacking a distinct separation between the cephalic and thoracic parts and also by males having a modified cymbium.</paragraph> <paragraph id="BD02FA6309CA2DE8187EDD77717D9D78" pageId="0" pageNumber="145">This is a small genus with seven species distributed exclusively in the Middle East from Turkey and Israel to Iran (including the two new species described below, representing the first record of this genus in this country). Members of this genus can be easily distinguished from all other zodariids found in the study area by the ovoid carapace lacking a distinct separation between the cephalic and thoracic parts and also by males having a modified cymbium.</paragraph>
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<treatment ID-GBIF-Taxon="181753556" LSID="urn:lsid:plazi:treatment:667578A987BD58158C0574BA9775A7AD" httpUri="http://treatment.plazi.org/id/667578A987BD58158C0574BA9775A7AD" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="667578A987BD58158C0574BA9775A7AD" ID-GBIF-Taxon="181753556" LSID="urn:lsid:plazi:treatment:667578A987BD58158C0574BA9775A7AD" httpUri="http://treatment.plazi.org/id/667578A987BD58158C0574BA9775A7AD" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Lachesaninae">
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<taxonomicName LSID="667578A9-87BD-5815-8C05-74BA9775A7AD" authority="Jocque, 1991" authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana insensibilis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="insensibilis"> <taxonomicName id="9E9CC44F1E634D980DE47D61369B7D10" ID-CoL="3RNBY" LSID="667578A9-87BD-5815-8C05-74BA9775A7AD" authority="Jocque, 1991" authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana insensibilis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="insensibilis">
Lachesana insensibilis Lachesana insensibilis
<normalizedToken originalValue="Jocqué">Jocque</normalizedToken> <normalizedToken id="AEF9DFAF82A2B32ED70895F5C3224E70" originalValue="Jocqué">Jocque</normalizedToken>
, 1991 , 1991
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">Fig. 32</figureCitation> <figureCitation id="62957599DA8D28E63724AF60F663B3C8" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">Fig. 32</figureCitation>
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="reference_group"> <subSubSection id="BC27098151102B0C2412F53906B9D1DB" pageId="0" pageNumber="145" type="reference_group">
<paragraph pageId="0" pageNumber="145"> <paragraph id="531C008E5457289C4CFC1CB1D50D3A48" pageId="0" pageNumber="145">
<taxonomicName authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana insensibilis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="insensibilis">Lachesana insensibilis</taxonomicName> <taxonomicName id="148BB59CA1563475AEC6EFE2E83F2EA1" authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana insensibilis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="insensibilis">Lachesana insensibilis</taxonomicName>
<normalizedToken originalValue="Jocqué">Jocque</normalizedToken> <normalizedToken id="83FC5F6560FCB0E9B285E4135D9786C1" originalValue="Jocqué">Jocque</normalizedToken>
, 1991: 37, f. 59 (♂). , 1991: 37, f. 59 (♂).
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="comments"> <subSubSection id="88AA5B64A789A1157C14438350B53F10" pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="2418C39222C050063B3A290DBDBD5DE7" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">This species is known from a single taxonomic entry, and unfortunately, the original description does not provide illustrations of the lateral view of the palp, making the identification of specimens from outside of the type locality (Saudi Arabia) highly questionable.</paragraph> <paragraph id="7A5CAE97F41D27BAEBF3C0C77685582A" pageId="0" pageNumber="145">This species is known from a single taxonomic entry, and unfortunately, the original description does not provide illustrations of the lateral view of the palp, making the identification of specimens from outside of the type locality (Saudi Arabia) highly questionable.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in iran"> <subSubSection id="C7409797704D008E971A3E26A4D86E6E" pageId="0" pageNumber="145" type="records in iran">
<paragraph pageId="0" pageNumber="145">Records in Iran.</paragraph> <paragraph id="9B3AC66511BF0E97E52BE5E29CFF5EE2" pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="88BE480B27D5DD48C43639A372498205" pageId="0" pageNumber="145">
Khuzestan ( Khuzestan (
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation> <bibRefCitation id="8B50ECA77788999DBB99F3E76BB3DCA6" DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation>
) (Fig. ) (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation> <figureCitation id="1CE0FE83D792ECEF6FC2953BB9410B9B" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). ).
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="431A32A8495DF11341309420760A57C9" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="9BAD67502F4B9467436B591DD60DB362" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">Saudi Arabia, Israel, Iran, United Arab Emirates.</paragraph> <paragraph id="93BD20F3081D1AE601B610E25A4ADB7E" pageId="0" pageNumber="145">Saudi Arabia, Israel, Iran, United Arab Emirates.</paragraph>
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</treatment> </treatment>
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2021" docId="675497C0692754C18A481685DC73BB2B" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Acanthinozodium masa Zamani &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="C50A0614-8086-4BCE-B6D8-8B9E40318781" docUuidSource="ZooBank" docVersion="4" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1668150222111" updateUser="ExternalLinkService"> <document ID-CLB-Dataset="7157" ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2021" docId="675497C0692754C18A481685DC73BB2B" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Acanthinozodium masa Zamani &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="C50A0614-8086-4BCE-B6D8-8B9E40318781" docUuidSource="ZooBank" docVersion="6" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1732670661048" updateUser="ExternalLinkService">
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title> <mods:title id="192D25F99EB4E6482B3D14ED0063C86B">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:namePart>Zamani, Alireza</mods:namePart> <mods:namePart id="630B1C225A22ED7B91689C6CEA7D174F">Zamani, Alireza</mods:namePart>
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation> <mods:affiliation id="288AB4A7107B7AAFE83387EAE769C098">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<treatment ID-GBIF-Taxon="181753562" LSID="urn:lsid:zoobank.org:act:C50A0614-8086-4BCE-B6D8-8B9E40318781" httpUri="http://treatment.plazi.org/id/675497C0692754C18A481685DC73BB2B" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="675497C0692754C18A481685DC73BB2B" ID-GBIF-Taxon="181753562" LSID="urn:lsid:zoobank.org:act:C50A0614-8086-4BCE-B6D8-8B9E40318781" httpUri="http://treatment.plazi.org/id/675497C0692754C18A481685DC73BB2B" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="96FA338A63E2107A4D81954188D68AC1" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="C04FC6BBFFFFA125B132B2DBE99EB252" pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/C50A0614-8086-4BCE-B6D8-8B9E40318781" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium masa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="masa" status="sp. nov.">Acanthinozodium masa</taxonomicName> <taxonomicName id="C0678E1C83CAA724CBA8122381DC5B8C" ID-CoL="98YZ3" LSID="http://zoobank.org/C50A0614-8086-4BCE-B6D8-8B9E40318781" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium masa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="masa" status="sp. nov.">Acanthinozodium masa</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="040313C731AE3DF581AAFD2CA0EDE3C5" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18L</figureCitation> <figureCitation id="DAD062AD45364CCD940C701302BE218A" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18L</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20I-K</figureCitation> <figureCitation id="0D0F1645F4EC62BCF555A1D04C0BACFB" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20I-K</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">, 24A, B</figureCitation> <figureCitation id="E661810D463FB1DDAC14E77DEFF1B5AC" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">, 24A, B</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation> <figureCitation id="EEDD38CD2522B27A94ADA1DB50BF5EF2" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="126DE4794FF8DFA770B5041B115D1E55" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="299E6266591B4A488BA7A2C510ADA9F9" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="93FCD052AF78D6326A8A392D4EFD84DF" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="D915FAEB8715EB80555B94611900512C" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ and ♂ and
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis> <emphasis id="F82470FE6DAC851514DF769849ECB73A" bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis>
♂ (MHNG), Iran: ♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Kermanshah Province</emphasis> <emphasis id="EC75688354FF031223784BEE25A56F93" italics="true" pageId="0" pageNumber="145">Kermanshah Province</emphasis>
: Mahi Dasht, : Mahi Dasht,
<geoCoordinate degrees="34" direction="north" minutes="14" orientation="latitude" precision="925" value="34.233334">34°14'N</geoCoordinate> <geoCoordinate id="8F29C6074C569DB08DE2194CD1AEC2E3" degrees="34" direction="north" minutes="14" orientation="latitude" precision="925" value="34.233334">34°14'N</geoCoordinate>
, ,
<geoCoordinate degrees="46" direction="east" minutes="42" orientation="longitude" precision="925" value="46.7">46°42'E</geoCoordinate> <geoCoordinate id="2B6926513B59C25680848C5816A5EFF6" degrees="46" direction="east" minutes="42" orientation="longitude" precision="925" value="46.7">46°42'E</geoCoordinate>
, 29.06.1974 (A. Senglet). , 29.06.1974 (A. Senglet).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="8733BA9D9D073538D126F1C70BCEB4F7" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="3B42208E74F1FF50076EF9DEA47986D7" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet is a Kurdish feminine name, meaning &quot;bright like the moon&quot;. Noun in apposition.</paragraph> <paragraph id="E7D8B02ECE1EDDBF947CD7736654E66F" pageId="0" pageNumber="145">The specific epithet is a Kurdish feminine name, meaning &quot;bright like the moon&quot;. Noun in apposition.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="6706EDBA731ED3ED4403069EA2170B89" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="EE0E1FDF20C1394526CFCCFD0CC36BA3" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="36BC012433206032A9A8A21F52198277" pageId="0" pageNumber="145">
The new species is very similar to The new species is very similar to
<taxonomicName lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara"> <taxonomicName id="6CD6673F081AFAB9B582D8C675E21C80" lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">A. diara</emphasis> <emphasis id="27A558AAB17F90D549B662BC630EA197" italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. by the shape of the RTA, the ventral tibial apophysis and the curvature of the embolus but differs by the shape of the median apophysis (cf. Fig. sp. nov. by the shape of the RTA, the ventral tibial apophysis and the curvature of the embolus but differs by the shape of the median apophysis (cf. Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20D</figureCitation> <figureCitation id="734359D0490911A8DB61AC7CADD86DCF" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20D</figureCitation>
and and
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20J</figureCitation> <figureCitation id="1F6BCC64ED66CCB27AEEA26B19FF3CEE" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20J</figureCitation>
) and the conductor lacking a small retrolateral indentation apically ( ) and the conductor lacking a small retrolateral indentation apically (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="72667BB81FB3CF8DAEE0BD89A196A50C" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
present). present).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="6AE023028C9724141F429456C59A8AC6" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="0F999683C187ADD458BEED6371687330" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="C677F2EC2940470E183406C629F2F37B" pageId="0" pageNumber="145">
Male (holotype). Habitus as in Fig. Male (holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18L</figureCitation> <figureCitation id="758F3CD253C54B4FAB1388428BC98AD0" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18L</figureCitation>
. Total length 3.05. Carapace 1.47 long, 0.66 wide at pars cephalica, 1.10 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.12, ALE: 0.10, PME: 0.07, PLE: 0.08, PME-PME: 0.12. Carapace, sternum, labium, chelicerae and maxillae brown; carapace with irregular patterns. Chelicera with one tooth. Legs yellowish brown. Abdomen dorsally black, grayish ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 5.52 (1.50, 0.47, 1.23, 1.42, 0.90), II: 4.91 (1.15, 0.50, 1.11, 1.30, 0.85), III: 4.62 (1.10, 0.45, 0.94, 1.43, 0.70), Fe IV: 1.80, rest of the segments missing. . Total length 3.05. Carapace 1.47 long, 0.66 wide at pars cephalica, 1.10 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.12, ALE: 0.10, PME: 0.07, PLE: 0.08, PME-PME: 0.12. Carapace, sternum, labium, chelicerae and maxillae brown; carapace with irregular patterns. Chelicera with one tooth. Legs yellowish brown. Abdomen dorsally black, grayish ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 5.52 (1.50, 0.47, 1.23, 1.42, 0.90), II: 4.91 (1.15, 0.50, 1.11, 1.30, 0.85), III: 4.62 (1.10, 0.45, 0.94, 1.43, 0.70), Fe IV: 1.80, rest of the segments missing.
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="BC183F9C496A195F581334CF06370A2E" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20I-K</figureCitation> <figureCitation id="968637A44FB3CC7DDD30B02F9CCC8881" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20I-K</figureCitation>
, ,
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">24A, B</figureCitation> <figureCitation id="2E46D77AD61BAB4110F3B5C67C08C342" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">24A, B</figureCitation>
. RTA long and conical, with a small projection apically; tegulum with posterior process ( . RTA long and conical, with a small projection apically; tegulum with posterior process (
<emphasis italics="true" pageId="0" pageNumber="145">Tp</emphasis> <emphasis id="6ACF847A37FC908E9E6FEDF0590DB651" italics="true" pageId="0" pageNumber="145">Tp</emphasis>
); ventral apophysis relatively large and conical; median apophysis wider than long, with a large bifurcated outgrowth prolaterally; embolus broad basally, originating at about the 7 ); ventral apophysis relatively large and conical; median apophysis wider than long, with a large bifurcated outgrowth prolaterally; embolus broad basally, originating at about the 7
<normalizedToken originalValue="o’clock">o'clock</normalizedToken> <normalizedToken id="1610F06C25E3C13B1A2C3A05F485F8CA" originalValue="o’clock">o'clock</normalizedToken>
position; embolus proper thin and steadily curving. position; embolus proper thin and steadily curving.
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="170D3B28A032E164A487276DE78227E3" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="135B6803A23F7892BA551ABC2C17DFA3" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown. Unknown.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="197F25C70BEF4F887BD1056B15434275" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="0D6DBE4409D891C74B0A5BB48F43166F" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="B2C8FF4BF83436C1639B87023B5F44EC" pageId="0" pageNumber="145">
Known only from the type locality in Kermanshah Province, western Iran (Fig. Known only from the type locality in Kermanshah Province, western Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation> <figureCitation id="E453F15120DC340FB72128190E058AEA" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

112
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<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="8ABD690DF811128674AAC6026A55C0AA" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="8959566D43979EDF6FFF5E019DB9FF82" pageId="0" pageNumber="145">
<taxonomicName LSID="6A815D86-982B-505E-8552-F43F3F2F2A9E" authority="(Nenilin &amp; Fet, 1985)" baseAuthorityName="Nenilin &amp; Fet" baseAuthorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum sytchevskajae" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="sytchevskajae">Zodariellum sytchevskajae (Nenilin &amp; Fet, 1985)</taxonomicName> <taxonomicName id="E77E621A2A474BCE967A434AF255B924" ID-CoL="93S8P" LSID="6A815D86-982B-505E-8552-F43F3F2F2A9E" authority="(Nenilin &amp; Fet, 1985)" baseAuthorityName="Nenilin &amp; Fet" baseAuthorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum sytchevskajae" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="sytchevskajae">Zodariellum sytchevskajae (Nenilin &amp; Fet, 1985)</taxonomicName>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">Fig. 33</figureCitation> <figureCitation id="77A731E733858A272188027F1FD5FA42" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">Fig. 33</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="reference_group"> <subSubSection id="0A77439F56FBDE111946958E01EDB43B" pageId="0" pageNumber="145" type="reference_group">
<paragraph pageId="0" pageNumber="145"> <paragraph id="CBEDDDCCCA043A00D77CFAC80F86C568" pageId="0" pageNumber="145">
<taxonomicName authorityName="Nenilin &amp; Fet" authorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion sytchevskajae" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="sytchevskajae">Zodarion sytchevskajae</taxonomicName> <taxonomicName id="5DA1603558B6CB82FD331B8E69EBCB8A" authorityName="Nenilin &amp; Fet" authorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion sytchevskajae" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="sytchevskajae">Zodarion sytchevskajae</taxonomicName>
Nenilin &amp; Fet, 1985: 619, f. 5-8, 10 (♂♀). Nenilin &amp; Fet, 1985: 619, f. 5-8, 10 (♂♀).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comments"> <subSubSection id="35A2C2D1008AFA2729611681CE0C71AB" pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="05E7278A417F74EB41FDCB6CC09040EF" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">This species is known from the original description only.</paragraph> <paragraph id="83BDCF3D6B4E8545DD87F559D1063953" pageId="0" pageNumber="145">This species is known from the original description only.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in turkmenistan"> <subSubSection id="784D29C55893E13E425708751D5FA926" pageId="0" pageNumber="145" type="records in turkmenistan">
<paragraph pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph> <paragraph id="4BC601174795DA58A698DD50DA3165FE" pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="7C5C3D084BA319304133505CE8D13241" pageId="0" pageNumber="145">
Ahal, Lebap ( Ahal, Lebap (
<bibRefCitation author="Nenilin, AB" journalOrPublisher="Zoologicheskiĭ Zhurnal" pageId="0" pageNumber="145" pagination="618 - 620" refId="B17" refString="Nenilin, AB, Fet, VY, 1985. New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia. Zoologicheskiĭ Zhurnal 64: 618 - 620" title="New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia." volume="64" year="1985">Nenilin and Fet 1985</bibRefCitation> <bibRefCitation id="77A504DD5E298C04EC035B8FE1A91CC3" author="Nenilin, AB" journalOrPublisher="Zoologicheskiĭ Zhurnal" pageId="0" pageNumber="145" pagination="618 - 620" refId="B17" refString="Nenilin, AB, Fet, VY, 1985. New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia. Zoologicheskiĭ Zhurnal 64: 618 - 620" title="New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia." volume="64" year="1985">Nenilin and Fet 1985</bibRefCitation>
) (Fig. ) (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation> <figureCitation id="0E7E4B4B633AB112992D8FE8192B519A" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="C8FEF5E864B2C4177EB8FCF7C901D58C" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="D5253D37DE193C7A64B984A35276AFBA" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">Turkmenistan.</paragraph> <paragraph id="284C96BC32F277CCF3AF51E082371766" pageId="0" pageNumber="145">Turkmenistan.</paragraph>
</subSubSection> </subSubSection>
</treatment> </treatment>
</document> </document>

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<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">Figs 26A-C</figureCitation> <figureCitation id="DDB3A7F0597EFE566FB143B7A85CD5E3" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">Figs 26A-C</figureCitation>
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Epigynes of Parazodarion raddei (A-D) and Trygetus susianus sp. nov. (E-H) A, E intact, ventral B, F-H macerated, dorsal C, D macerated, ventral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145">, 27A-D</figureCitation> <figureCitation id="6433C66F2F39906454E3BF7FC9AEC1D8" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Epigynes of Parazodarion raddei (A-D) and Trygetus susianus sp. nov. (E-H) A, E intact, ventral B, F-H macerated, dorsal C, D macerated, ventral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure27" httpUri="https://binary.pensoft.net/fig/536904" pageId="0" pageNumber="145">, 27A-D</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation> <figureCitation id="BB936EA2C50217DDA4A160C67250DFC1" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="reference_group"> <subSubSection id="A54CCD8A75F94B6502AA855958115F19" pageId="0" pageNumber="145" type="reference_group">
<paragraph pageId="0" pageNumber="145"> <paragraph id="302FDF512D4B230592281B3C8BBB62A7" pageId="0" pageNumber="145">
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion raddei" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei">Parazodarion raddei</taxonomicName> <taxonomicName id="6731F357A928CAED9BFB039DF5A58D59" baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion raddei" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei">Parazodarion raddei</taxonomicName>
: :
<bibRefCitation DOI="https://doi.org/10.2478/v10058-009-0020-z" author="Ovtchinnikov, SV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="145" pagination="471 - 474" refId="B19" refString="Ovtchinnikov, SV, Ahmad, B, Gurko, VO, 2009. Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia. Vestnik Zoologii 43 (5): 471 - 474, DOI: https://doi.org/10.2478/v10058-009-0020-z" title="Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia." url="https://doi.org/10.2478/v10058-009-0020-z" volume="43" year="2009">Ovtchinnikov et al. 2009</bibRefCitation> <bibRefCitation id="A3B19CD2BE64EB2F1827073A49A38D1D" DOI="https://doi.org/10.2478/v10058-009-0020-z" author="Ovtchinnikov, SV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="145" pagination="471 - 474" refId="B19" refString="Ovtchinnikov, SV, Ahmad, B, Gurko, VO, 2009. Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia. Vestnik Zoologii 43 (5): 471 - 474, DOI: https://doi.org/10.2478/v10058-009-0020-z" title="Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia." url="https://doi.org/10.2478/v10058-009-0020-z" volume="43" year="2009">Ovtchinnikov et al. 2009</bibRefCitation>
: 471, f. 1.1-6 (♂♀). : 471, f. 1.1-6 (♂♀).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="8A001BE69001B2BAEAC52E397595DE2D" pageId="0" pageNumber="145">
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion raddei" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei">Parazodarion raddei</taxonomicName> <taxonomicName id="BD515B266358C3B8866838D118A8873C" baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion raddei" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei">Parazodarion raddei</taxonomicName>
For the complete list of references see For the complete list of references see
<bibRefCitation author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC (2021)</bibRefCitation> <bibRefCitation id="35DE8EE2716FBD97D292DB2E5B853BEF" author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC (2021)</bibRefCitation>
. .
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="materials_examined"> <subSubSection id="E244968BCEE0CF4D65EA207DDA71814A" pageId="0" pageNumber="145" type="materials_examined">
<paragraph pageId="0" pageNumber="145">Material.</paragraph> <paragraph id="410E113E69E1A6E4627F948CE8C9A3F8" pageId="0" pageNumber="145">Material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="6EE5587EC0D625303C320F20DE5A8160" pageId="0" pageNumber="145">
<materialsCitation ID-GBIF-Occurrence="3093634306" collectingDate="1975-06-16" collectorName="A. Senglet" country="Iran" elevation="2600" latitude="34.733334" location="Hamedan Province" longLatPrecision="1197" longitude="48.783333" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Hamedan Province"> <materialsCitation id="D389C4F3571AB72460E18D5E46854904" ID-GBIF-Occurrence="3093634306" collectingDate="1975-06-16" collectorName="A. Senglet" country="Iran" elevation="2600" latitude="34.733334" location="Hamedan Province" longLatPrecision="1197" longitude="48.783333" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Hamedan Province">
<collectingCountry name="Iran">Iran</collectingCountry> <collectingCountry id="C0D734EA38415EB57AEB8041F0D44C5A" name="Iran">Iran</collectingCountry>
: :
<emphasis italics="true" pageId="0" pageNumber="145"> <emphasis id="3E503A6D9738B9958EDF0B81C8C2CB6A" italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Hamadan">Hamedan Province</collectingRegion> <collectingRegion id="685C403C6B295859C957CDE6C073F57B" country="Iran" name="Hamadan">Hamedan Province</collectingRegion>
</emphasis> </emphasis>
: :
<specimenCount type="male">1♂</specimenCount> <specimenCount id="647FC74E0BA2CE6FA2AEEF62C6F3C520" type="male">1♂</specimenCount>
<specimenCount type="female">3♀</specimenCount> <specimenCount id="A167E860FCF6ACE8C25DF5C87E92A73E" type="female">3♀</specimenCount>
(MHNG), around (MHNG), around
<collectingRegion country="Iran" name="Hamadan">Hamedan</collectingRegion> <collectingRegion id="098538C1A695C3FEF052150A2961CC74" country="Iran" name="Hamadan">Hamedan</collectingRegion>
, ,
<geoCoordinate degrees="34" direction="north" minutes="44" orientation="latitude" precision="925" value="34.733334">34°44'N</geoCoordinate> <geoCoordinate id="7C75A0F829DC9350A5ABD9958BE61C9A" degrees="34" direction="north" minutes="44" orientation="latitude" precision="925" value="34.733334">34°44'N</geoCoordinate>
, ,
<geoCoordinate degrees="48" direction="east" minutes="47" orientation="longitude" precision="925" value="48.783333">48°47'E</geoCoordinate> <geoCoordinate id="66D89399D60D3C419D85096DE21D7A6D" degrees="48" direction="east" minutes="47" orientation="longitude" precision="925" value="48.783333">48°47'E</geoCoordinate>
, ,
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0"> <quantity id="5017699E4F3723889E6C4FF0C0D6DC0E" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2600 m</elevation> <elevation id="66443E2833EEED4B41F4AC7F15FD7659" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2600 m</elevation>
</quantity> </quantity>
, ,
<collectingDate value="1975-06-16">16.06.1975</collectingDate> <collectingDate id="9161C150BEF5B26CFEA65F2A47C5F34E" value="1975-06-16">16.06.1975</collectingDate>
( (
<collectorName>A. Senglet</collectorName> <collectorName id="8FCA931AFC8C9E49B765ECA5E8FD1683">A. Senglet</collectorName>
) )
</materialsCitation> </materialsCitation>
; ;
<materialsCitation ID-GBIF-Occurrence="3093634304" collectingDate="1974-07-02" collectorName="A. Senglet" country="Iran" latitude="34.85" location="Aliabad" longLatPrecision="1197" longitude="48.2" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Hamedan"> <materialsCitation id="405D1F290C37F40F717C49E8166C72F8" ID-GBIF-Occurrence="3093634304" collectingDate="1974-07-02" collectorName="A. Senglet" country="Iran" latitude="34.85" location="Aliabad" longLatPrecision="1197" longitude="48.2" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Hamedan">
<specimenCount type="male">1♂</specimenCount> <specimenCount id="3E3BFB512B41AFEDB696B449480BA2C2" type="male">1♂</specimenCount>
<specimenCount type="female">2♀</specimenCount> <specimenCount id="8B798197F6BC0BA2B31FBA2421AC58F6" type="female">2♀</specimenCount>
(MHNG), (MHNG),
<location LSID="urn:lsid:plazi:treatment:6BA12FA2395B570193927134C049CA76:51B9CCABD6DF0BB85FE5DB52E7473A31" country="Iran" latitude="34.85" longLatPrecision="1197" longitude="48.2" name="Aliabad" stateProvince="Hamedan">Aliabad</location> <location id="51B9CCABD6DF0BB85FE5DB52E7473A31" LSID="urn:lsid:plazi:treatment:6BA12FA2395B570193927134C049CA76:51B9CCABD6DF0BB85FE5DB52E7473A31" country="Iran" latitude="34.85" longLatPrecision="1197" longitude="48.2" name="Aliabad" stateProvince="Hamedan">Aliabad</location>
, ,
<geoCoordinate degrees="34" direction="north" minutes="51" orientation="latitude" precision="925" value="34.85">34°51'N</geoCoordinate> <geoCoordinate id="F1E007AB69025CE1E944A1BF0C47FFE4" degrees="34" direction="north" minutes="51" orientation="latitude" precision="925" value="34.85">34°51'N</geoCoordinate>
, ,
<geoCoordinate degrees="48" direction="east" minutes="12" orientation="longitude" precision="925" value="48.2">48°12'E</geoCoordinate> <geoCoordinate id="53FC3B7AF407E15AEF8310400DD42893" degrees="48" direction="east" minutes="12" orientation="longitude" precision="925" value="48.2">48°12'E</geoCoordinate>
, ,
<collectingDate value="1974-07-02">02.07.1974</collectingDate> <collectingDate id="59EEC32FE6649381975F739B9B375D91" value="1974-07-02">02.07.1974</collectingDate>
( (
<collectorName>A. Senglet</collectorName> <collectorName id="B90FAE11AEFB7D8DC1A6934AAAF21AC9">A. Senglet</collectorName>
) )
</materialsCitation> </materialsCitation>
; ;
<materialsCitation ID-GBIF-Occurrence="3093634301" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Iran" location="Tehran Province" specimenCount="12" specimenCount-female="5" specimenCount-male="7" stateProvince="Tehran Province"> <materialsCitation id="2FB29C7D04BAD0F07F4DEE7F87F9C779" ID-GBIF-Occurrence="3093634301" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Iran" location="Tehran Province" specimenCount="12" specimenCount-female="5" specimenCount-male="7" stateProvince="Tehran Province">
<emphasis italics="true" pageId="0" pageNumber="145"> <emphasis id="07F1480B0AFAE52AE9EA143D4411D22B" italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Tehran">Tehran Province</collectingRegion> <collectingRegion id="B8D89F72E621D958D4E537BB2D77869C" country="Iran" name="Tehran">Tehran Province</collectingRegion>
</emphasis> </emphasis>
: :
<specimenCount type="male">7♂</specimenCount> <specimenCount id="01D9292E8F3700C7725AE706F5A37449" type="male">7♂</specimenCount>
<specimenCount type="female">5♀</specimenCount> <specimenCount id="285286A30BB8617FC269BE4B1F0BE9E5" type="female">5♀</specimenCount>
(ZMMU), (ZMMU),
<quantity metricMagnitude="4" metricUnit="m" metricValue="8.0" unit="km" value="80.0">80 km</quantity> <quantity id="E9B29F6B8375D7F06561AD4B377EA742" metricMagnitude="4" metricUnit="m" metricValue="8.0" unit="km" value="80.0">80 km</quantity>
E of E of
</materialsCitation> </materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3093634313" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Iran" isTailEnumeration="true" location="Tehran" specimenCount="1" stateProvince="Tehran"> <materialsCitation id="847A7EE2DFBA8E7ED0149655283FDA56" ID-GBIF-Occurrence="3093634313" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Iran" isTailEnumeration="true" location="Tehran" specimenCount="1" stateProvince="Tehran">
<collectingRegion country="Iran" name="Tehran">Tehran</collectingRegion> <collectingRegion id="9BF0D811207BF446C53229D3B591EDFC" country="Iran" name="Tehran">Tehran</collectingRegion>
</materialsCitation> </materialsCitation>
, ,
<materialsCitation ID-GBIF-Occurrence="3093634305" collectingDate="2000-06-15" collectorName="Y. M. Marusik, F. Mozaffarian" country="Iran" latitude="35.666668" location="Aroo" longLatPrecision="1191" longitude="52.45" municipality="Damavand area" specimenCount="1" stateProvince="Tehran"> <materialsCitation id="498252D9D0F2A543CF51EE32A354EA60" ID-GBIF-Occurrence="3093634305" collectingDate="2000-06-15" collectorName="Y. M. Marusik, F. Mozaffarian" country="Iran" latitude="35.666668" location="Aroo" longLatPrecision="1191" longitude="52.45" municipality="Damavand area" specimenCount="1" stateProvince="Tehran">
<collectingMunicipality>Damavand area</collectingMunicipality> <collectingMunicipality id="90D53C2EC39C3AD600C1DA74C980FC88">Damavand area</collectingMunicipality>
, ,
<location LSID="urn:lsid:plazi:treatment:6BA12FA2395B570193927134C049CA76:493F0732B732A456CB82CA23484035E1" country="Iran" latitude="35.666668" longLatPrecision="1191" longitude="52.45" municipality="Damavand area" name="Aroo" stateProvince="Tehran">Aroo</location> <location id="493F0732B732A456CB82CA23484035E1" LSID="urn:lsid:plazi:treatment:6BA12FA2395B570193927134C049CA76:493F0732B732A456CB82CA23484035E1" country="Iran" latitude="35.666668" longLatPrecision="1191" longitude="52.45" municipality="Damavand area" name="Aroo" stateProvince="Tehran">Aroo</location>
, ,
<geoCoordinate degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate> <geoCoordinate id="5344DEB640DF7D256AEA0EA77418550F" degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate>
, ,
<geoCoordinate degrees="52" direction="east" minutes="27" orientation="longitude" precision="925" value="52.45">52°27'E</geoCoordinate> <geoCoordinate id="4B708A745A5BC4B618FDA5EDA34D2C3A" degrees="52" direction="east" minutes="27" orientation="longitude" precision="925" value="52.45">52°27'E</geoCoordinate>
, ,
<collectingDate value="2000-06-15">15.06.2000</collectingDate> <collectingDate id="E0838B13B4549F8BAFB96F0992CC964D" value="2000-06-15">15.06.2000</collectingDate>
( (
<collectorName>Y.M. Marusik</collectorName> <collectorName id="A2B77B8CB0D3ABB20B67B6A3974E9A16">Y.M. Marusik</collectorName>
&amp; &amp;
<collectorName>F. Mozaffarian</collectorName> <collectorName id="002AF8DD71BD1B6E535301B45B808F9E">F. Mozaffarian</collectorName>
) )
</materialsCitation> </materialsCitation>
. .
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145" start="Figure 26" startId="F26"> <caption id="79078603D0BB70011D02C36E8C276DF7" doi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145" start="Figure 26" startId="F26">
<paragraph pageId="0" pageNumber="145"> <paragraph id="6397AE631761A593F5DE53E29335823C" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 26.</emphasis> <emphasis id="74290CE0C29141EF4D334EDE75F377EA" bold="true" pageId="0" pageNumber="145">Figure 26.</emphasis>
Males of Males of
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion raddei" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei"> <taxonomicName id="53888DAD3B26F74971BE194646EC67CE" baseAuthorityName="Simon" baseAuthorityYear="1889" family="Zodariidae" genus="Parazodarion" kingdom="Animalia" lsidName="Parazodarion raddei" order="Aranei" pageId="0" pageNumber="145" rank="species" species="raddei">
<emphasis italics="true" pageId="0" pageNumber="145">Parazodarion raddei</emphasis> <emphasis id="BA2B8D1A20B1511C9B6B0F8FBFAB9BE9" italics="true" pageId="0" pageNumber="145">Parazodarion raddei</emphasis>
</taxonomicName> </taxonomicName>
( (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis> <emphasis id="87377BBFFDC88971BF7053C50D4462DE" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and ) and
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum"> <taxonomicName id="011C6F53169E88E8CC49E484D6CF6EFE" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis> <emphasis id="358C7F2FCD8BCA15F8179C831EAD725C" italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis> <emphasis id="FB46C6C08C818740010538D535AF4FE4" bold="true" pageId="0" pageNumber="145">E</emphasis>
) and female of ) and female of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus susianus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="susianus"> <taxonomicName id="FCA9836621A214A2BA802BC116AD3AC1" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus susianus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="susianus">
<emphasis italics="true" pageId="0" pageNumber="145">Trygetus susianus</emphasis> <emphasis id="C78F4B0A00CDC821421CDF759FCC58BE" italics="true" pageId="0" pageNumber="145">Trygetus susianus</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis> <emphasis id="9F6FC6D31C359C233E4DE3FDC0EA9508" bold="true" pageId="0" pageNumber="145">D</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis> <emphasis id="45F0C679842D9474C9B0CD48FDE899DD" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
habitus, dorsal habitus, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">B, C</emphasis> <emphasis id="E54984AB214FB1CE00579F53F7A4A043" bold="true" pageId="0" pageNumber="145">B, C</emphasis>
palp, ventral and retrolateral palp, ventral and retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis> <emphasis id="495E8E2EE9E0A691F161B3AD9F3A6B78" bold="true" pageId="0" pageNumber="145">E</emphasis>
chelicerae, ventral. Scale bars: chelicerae, ventral. Scale bars:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity> <quantity id="83CA1B1261E4053A156C8EE21DD7993E" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
, unless stated otherwise. , unless stated otherwise.
</paragraph> </paragraph>
</caption> </caption>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in iran"> <subSubSection id="97C576AF1F265137BE68ADB647195D9E" pageId="0" pageNumber="145" type="records in iran">
<paragraph pageId="0" pageNumber="145">Records in Iran.</paragraph> <paragraph id="83EED1989E8ED1C75855F70FB9B5DC56" pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="CA77D57D33F2884AE5F4BE1D928365BA" pageId="0" pageNumber="145">
Alborz, Isfahan, Qom, Razavi Khorasan, Sistan and Baluchistan, West Azerbaijan, Yazd, Zanjan ( Alborz, Isfahan, Qom, Razavi Khorasan, Sistan and Baluchistan, West Azerbaijan, Yazd, Zanjan (
<bibRefCitation DOI="https://doi.org/10.2478/v10058-009-0020-z" author="Ovtchinnikov, SV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="145" pagination="471 - 474" refId="B19" refString="Ovtchinnikov, SV, Ahmad, B, Gurko, VO, 2009. Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia. Vestnik Zoologii 43 (5): 471 - 474, DOI: https://doi.org/10.2478/v10058-009-0020-z" title="Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia." url="https://doi.org/10.2478/v10058-009-0020-z" volume="43" year="2009">Ovtchinnikov et al. 2009</bibRefCitation> <bibRefCitation id="2D27BF154405077BEB65A8A256CE9308" DOI="https://doi.org/10.2478/v10058-009-0020-z" author="Ovtchinnikov, SV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="145" pagination="471 - 474" refId="B19" refString="Ovtchinnikov, SV, Ahmad, B, Gurko, VO, 2009. Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia. Vestnik Zoologii 43 (5): 471 - 474, DOI: https://doi.org/10.2478/v10058-009-0020-z" title="Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia." url="https://doi.org/10.2478/v10058-009-0020-z" volume="43" year="2009">Ovtchinnikov et al. 2009</bibRefCitation>
; ;
<bibRefCitation DOI="https://doi.org/10.3906/zoo-1307-16" author="Hosseini, M" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="145" pagination="437 - 443" refId="B6" refString="Hosseini, M, Mirshamsi, O, Kashefi, R, Fekrat, L, 2014. A contribution to the knowledge of spiders in wheat fields of Khorasan-e-Razavi Province, Iran. Turkish Journal of Zoology 38 (4): 437 - 443, DOI: https://doi.org/10.3906/zoo-1307-16" title="A contribution to the knowledge of spiders in wheat fields of Khorasan-e-Razavi Province, Iran." url="https://doi.org/10.3906/zoo-1307-16" volume="38" year="2014">Hosseini et al. 2014</bibRefCitation> <bibRefCitation id="1E53F4ED3D3A6D3A52CADCCB1765D0AF" DOI="https://doi.org/10.3906/zoo-1307-16" author="Hosseini, M" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="145" pagination="437 - 443" refId="B6" refString="Hosseini, M, Mirshamsi, O, Kashefi, R, Fekrat, L, 2014. A contribution to the knowledge of spiders in wheat fields of Khorasan-e-Razavi Province, Iran. Turkish Journal of Zoology 38 (4): 437 - 443, DOI: https://doi.org/10.3906/zoo-1307-16" title="A contribution to the knowledge of spiders in wheat fields of Khorasan-e-Razavi Province, Iran." url="https://doi.org/10.3906/zoo-1307-16" volume="38" year="2014">Hosseini et al. 2014</bibRefCitation>
; ;
<bibRefCitation author="Sadeghi, H" journalOrPublisher="Biharean Biologist" pageId="0" pageNumber="145" pagination="4 - 7" refId="B23" refString="Sadeghi, H, Ahmadi, M, Zamani, A, Jabaleh, I, 2016. A study on the spider fauna of Dargaz and Kalat Counties in Razavi Khorasan Province, Iran (Arachnida: Araneae). Biharean Biologist 10 (1): 4 - 7" title="A study on the spider fauna of Dargaz and Kalat Counties in Razavi Khorasan Province, Iran (Arachnida: Araneae)." volume="10" year="2016">Sadeghi et al. 2016</bibRefCitation> <bibRefCitation id="A78F2147FC9B808A67BE3B30D73E2597" author="Sadeghi, H" journalOrPublisher="Biharean Biologist" pageId="0" pageNumber="145" pagination="4 - 7" refId="B23" refString="Sadeghi, H, Ahmadi, M, Zamani, A, Jabaleh, I, 2016. A study on the spider fauna of Dargaz and Kalat Counties in Razavi Khorasan Province, Iran (Arachnida: Araneae). Biharean Biologist 10 (1): 4 - 7" title="A study on the spider fauna of Dargaz and Kalat Counties in Razavi Khorasan Province, Iran (Arachnida: Araneae)." volume="10" year="2016">Sadeghi et al. 2016</bibRefCitation>
; ;
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation> <bibRefCitation id="DD0CF3E1BB756B0FB081A4B18E7EFDCA" DOI="https://doi.org/10.2476/asjaa.66.55" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="55 - 71" refId="B29" refString="Zamani, A, Mirshamsi, O, Dolejs, P, Marusik, YM, Esyunin, SL, Hula, V, Ponel, P, 2017. New data on the spider fauna of Iran (Arachnida: Araneae), Part IV. Acta Arachnologica 66 (2): 55 - 71, DOI: https://doi.org/10.2476/asjaa.66.55" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part IV." url="https://doi.org/10.2476/asjaa.66.55" volume="66" year="2017">Zamani et al. 2017</bibRefCitation>
; ;
<bibRefCitation author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. 2018</bibRefCitation> <bibRefCitation id="5BF1018429C6E25350DDEF31648D8EDC" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. 2018</bibRefCitation>
; ;
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.67.67" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="67 - 75" refId="B28" refString="Zamani, A, Marusik, YM, 2018. The first report on the spider fauna (Arachnida: Araneae) of the Lut Desert, Iran. Acta Arachnologica 67 (2): 67 - 75, DOI: https://doi.org/10.2476/asjaa.67.67" title="The first report on the spider fauna (Arachnida: Araneae) of the Lut Desert, Iran." url="https://doi.org/10.2476/asjaa.67.67" volume="67" year="2018">Zamani and Marusik 2018</bibRefCitation> <bibRefCitation id="E789439F10D51B604C37BC87A72816BB" DOI="https://doi.org/10.2476/asjaa.67.67" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" pagination="67 - 75" refId="B28" refString="Zamani, A, Marusik, YM, 2018. The first report on the spider fauna (Arachnida: Araneae) of the Lut Desert, Iran. Acta Arachnologica 67 (2): 67 - 75, DOI: https://doi.org/10.2476/asjaa.67.67" title="The first report on the spider fauna (Arachnida: Araneae) of the Lut Desert, Iran." url="https://doi.org/10.2476/asjaa.67.67" volume="67" year="2018">Zamani and Marusik 2018</bibRefCitation>
). New records for Hamedan and Tehran (Fig. ). New records for Hamedan and Tehran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation> <figureCitation id="36370E523861F15B8D52BF34178CE7BD" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in turkmenistan"> <subSubSection id="6462EFA650493E3213A318B36233EF1F" pageId="0" pageNumber="145" type="records in turkmenistan">
<paragraph pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph> <paragraph id="50DECEDC4FA9829F075E657AF8C72546" pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="5B84CF0C11B7A0F4BE7CBAEC112F2172" pageId="0" pageNumber="145">
Ahal, Archman (= Ahal, Archman (=
<normalizedToken originalValue="Arçman">Arcman</normalizedToken> <normalizedToken id="31C02D60722A9267F6E78CB2F7D6A39E" originalValue="Arçman">Arcman</normalizedToken>
), Ashgabat, Balkan, Lebap, Mary ( ), Ashgabat, Balkan, Lebap, Mary (
<bibRefCitation author="Simon, E" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="145" pagination="373 - 386" refId="B24" refString="Simon, E, 1889. Arachnidae transcaspicae ab ill. Dr. G. Radde, Dr. A. Walter et A. Conchin inventae (annis 1886-1887). Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 39: 373 - 386" title="Arachnidae transcaspicae ab ill. Dr. G. Radde, Dr. A. Walter et A. Conchin inventae (annis 1886 - 1887)." volume="39" year="1889">Simon 1889</bibRefCitation> <bibRefCitation id="9BF8EC7BF7F09C86EE6953E63FECDCB6" author="Simon, E" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="145" pagination="373 - 386" refId="B24" refString="Simon, E, 1889. Arachnidae transcaspicae ab ill. Dr. G. Radde, Dr. A. Walter et A. Conchin inventae (annis 1886-1887). Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 39: 373 - 386" title="Arachnidae transcaspicae ab ill. Dr. G. Radde, Dr. A. Walter et A. Conchin inventae (annis 1886 - 1887)." volume="39" year="1889">Simon 1889</bibRefCitation>
; ;
<bibRefCitation author="Vlassov, JP" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" pagination="247 - 258" refId="B25" refString="Vlassov, JP, Sytshevskaja, VI, 1937. The spiders of the neighbourhood of Ashkhabad. Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja 9: 247 - 258" title="The spiders of the neighbourhood of Ashkhabad." volume="9" year="1937">Vlassov and Sytshevskaja 1937</bibRefCitation> <bibRefCitation id="ABD70865005F05A55F31E8A5FF298A67" author="Vlassov, JP" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" pagination="247 - 258" refId="B25" refString="Vlassov, JP, Sytshevskaja, VI, 1937. The spiders of the neighbourhood of Ashkhabad. Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja 9: 247 - 258" title="The spiders of the neighbourhood of Ashkhabad." volume="9" year="1937">Vlassov and Sytshevskaja 1937</bibRefCitation>
; ;
<bibRefCitation author="Ovtsharenko, VI" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="0" pageNumber="145" pagination="442 - 447" refId="B20" refString="Ovtsharenko, VI, Fet, VY, 1980. Fauna and ecology of spiders (Aranei) of Badhyz (Turkmenian SSR). Entomologicheskoe Obozrenie 59: 442 - 447" title="Fauna and ecology of spiders (Aranei) of Badhyz (Turkmenian SSR)." volume="59" year="1980">Ovtsharenko and Fet 1980</bibRefCitation> <bibRefCitation id="8CC21B300DFDF875763DB72677234E5C" author="Ovtsharenko, VI" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="0" pageNumber="145" pagination="442 - 447" refId="B20" refString="Ovtsharenko, VI, Fet, VY, 1980. Fauna and ecology of spiders (Aranei) of Badhyz (Turkmenian SSR). Entomologicheskoe Obozrenie 59: 442 - 447" title="Fauna and ecology of spiders (Aranei) of Badhyz (Turkmenian SSR)." volume="59" year="1980">Ovtsharenko and Fet 1980</bibRefCitation>
; ;
<bibRefCitation author="Fet, VY" journalOrPublisher="Senckenbergiana Biologica" pageId="0" pageNumber="145" refId="B4" refString="Fet, VY, 1985. [The ecological distribution of spiders in Syunt Khassardagh Reserve]. In: [The Vegetation and Animal Life of Western Kopetdagh]. Ashkhabad, 271-277. [in Russian]" title="[The ecological distribution of spiders in Syunt Khassardagh Reserve]. In: [The Vegetation and Animal Life of Western Kopetdagh]. Ashkhabad, 271 - 277. [in Russian]" year="1985">Fet 1985</bibRefCitation> <bibRefCitation id="0806912218825B1BD7FF114007E3839C" author="Fet, VY" journalOrPublisher="Senckenbergiana Biologica" pageId="0" pageNumber="145" refId="B4" refString="Fet, VY, 1985. [The ecological distribution of spiders in Syunt Khassardagh Reserve]. In: [The Vegetation and Animal Life of Western Kopetdagh]. Ashkhabad, 271-277. [in Russian]" title="[The ecological distribution of spiders in Syunt Khassardagh Reserve]. In: [The Vegetation and Animal Life of Western Kopetdagh]. Ashkhabad, 271 - 277. [in Russian]" year="1985">Fet 1985</bibRefCitation>
; ;
<bibRefCitation DOI="https://doi.org/10.1007/978-94-011-1116-4_30" author="Mikhailov, KG" editor="Fet, V" journalOrPublisher="Kluwer Academic Publisher Netherlands" pageId="0" pageNumber="145" pagination="499 - 524" refId="B16" refString="Mikhailov, KG, Fet, V, 1994. Fauna and zoogeography of spiders (Aranei) of Turkmenistan. In: Fet, V, Atamuradov, KI, Eds., Biogeography and Ecology of Turkmenistan. Kluwer Academic Publisher Netherlands: 499 - 524, DOI: https://doi.org/10.1007/978-94-011-1116-4_30" title="Fauna and zoogeography of spiders (Aranei) of Turkmenistan." url="https://doi.org/10.1007/978-94-011-1116-4_30" volumeTitle="Biogeography and Ecology of Turkmenistan." year="1994">Mikhailov and Fet 1994</bibRefCitation> <bibRefCitation id="22D36A319827560B3129DE24675D3868" DOI="https://doi.org/10.1007/978-94-011-1116-4_30" author="Mikhailov, KG" editor="Fet, V" journalOrPublisher="Kluwer Academic Publisher Netherlands" pageId="0" pageNumber="145" pagination="499 - 524" refId="B16" refString="Mikhailov, KG, Fet, V, 1994. Fauna and zoogeography of spiders (Aranei) of Turkmenistan. In: Fet, V, Atamuradov, KI, Eds., Biogeography and Ecology of Turkmenistan. Kluwer Academic Publisher Netherlands: 499 - 524, DOI: https://doi.org/10.1007/978-94-011-1116-4_30" title="Fauna and zoogeography of spiders (Aranei) of Turkmenistan." url="https://doi.org/10.1007/978-94-011-1116-4_30" volumeTitle="Biogeography and Ecology of Turkmenistan." year="1994">Mikhailov and Fet 1994</bibRefCitation>
; ;
<bibRefCitation author="Mikhailov, KG" journalOrPublisher="Zoological Museum, Moscow State University Moscow" pageId="0" pageNumber="145" refId="B14" refString="Mikhailov, KG, 1997. Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei). Zoological Museum, Moscow State University Moscow" title="Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei)." year="1997">Mikhailov 1997</bibRefCitation> <bibRefCitation id="E74E917CC92DD771F14BDCF9A2787EB9" author="Mikhailov, KG" journalOrPublisher="Zoological Museum, Moscow State University Moscow" pageId="0" pageNumber="145" refId="B14" refString="Mikhailov, KG, 1997. Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei). Zoological Museum, Moscow State University Moscow" title="Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei)." year="1997">Mikhailov 1997</bibRefCitation>
; ;
<bibRefCitation DOI="https://doi.org/10.2478/v10058-009-0020-z" author="Ovtchinnikov, SV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="145" pagination="471 - 474" refId="B19" refString="Ovtchinnikov, SV, Ahmad, B, Gurko, VO, 2009. Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia. Vestnik Zoologii 43 (5): 471 - 474, DOI: https://doi.org/10.2478/v10058-009-0020-z" title="Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia." url="https://doi.org/10.2478/v10058-009-0020-z" volume="43" year="2009">Ovtchinnikov et al. 2009</bibRefCitation> <bibRefCitation id="CFD90574522250FE61E79EBBF2923298" DOI="https://doi.org/10.2478/v10058-009-0020-z" author="Ovtchinnikov, SV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="145" pagination="471 - 474" refId="B19" refString="Ovtchinnikov, SV, Ahmad, B, Gurko, VO, 2009. Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia. Vestnik Zoologii 43 (5): 471 - 474, DOI: https://doi.org/10.2478/v10058-009-0020-z" title="Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia." url="https://doi.org/10.2478/v10058-009-0020-z" volume="43" year="2009">Ovtchinnikov et al. 2009</bibRefCitation>
) (Fig. ) (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation> <figureCitation id="976ED5E46F45C5950E611DA32479E1F3" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="0EEEB13E047CAA9EE25F90B140FA93CC" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="DCA3B6F4DEBC16FEBD789A21427946EF" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">From United Arab Emirates to Kazakhstan and Afghanistan in the east.</paragraph> <paragraph id="B87F88B6FCD637FA7C3D37EFE632227D" pageId="0" pageNumber="145">From United Arab Emirates to Kazakhstan and Afghanistan in the east.</paragraph>
</subSubSection> </subSubSection>
</treatment> </treatment>
</document> </document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2021" docId="75080B2F5CE955999C9274B037E542BE" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Lachesana kavirensis Zamani &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" docUuidSource="ZooBank" docVersion="4" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1668150222111" updateUser="ExternalLinkService"> <document ID-CLB-Dataset="7157" ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2021" docId="75080B2F5CE955999C9274B037E542BE" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Lachesana kavirensis Zamani &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" docUuidSource="ZooBank" docVersion="6" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1732670661048" updateUser="ExternalLinkService">
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title> <mods:title id="ED62FFFDD789E12CDFCE54EE14C8CDEB">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:namePart>Zamani, Alireza</mods:namePart> <mods:namePart id="F6CD8B773B0BB31C55D91A0AA31C437A">Zamani, Alireza</mods:namePart>
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation> <mods:affiliation id="D9C7C12EDC219E3DC0A33444BBE7E658">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<treatment ID-GBIF-Taxon="181753555" LSID="urn:lsid:zoobank.org:act:3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" httpUri="http://treatment.plazi.org/id/75080B2F5CE955999C9274B037E542BE" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="75080B2F5CE955999C9274B037E542BE" ID-GBIF-Taxon="181753555" LSID="urn:lsid:zoobank.org:act:3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" httpUri="http://treatment.plazi.org/id/75080B2F5CE955999C9274B037E542BE" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Lachesaninae">
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<paragraph pageId="0" pageNumber="145"> <paragraph id="0652F93F8648253B38C99090BAA5A5F4" pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis" status="sp. nov.">Lachesana kavirensis</taxonomicName> <taxonomicName id="FD2D5B0ACF69683FA3B9F0CD925AA485" ID-CoL="98YYV" LSID="http://zoobank.org/3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis" status="sp. nov.">Lachesana kavirensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="6F9266FD992B194DB159817F468C6961" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">Figs 1A-C</figureCitation> <figureCitation id="1EBCAC53707311EA5F299A866C33C333" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">Figs 1A-C</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">, 3A-E</figureCitation> <figureCitation id="7D8D8367E734BC9A24BF4370318190D5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">, 3A-E</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">, 4I</figureCitation> <figureCitation id="3620ED69DCD6F826F7E6B13D8DD38725" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">, 4I</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation> <figureCitation id="4E0449A87159DED2C316FDAF4582E224" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="5C8A2B03B4B9250026BF42C87C0DBA01" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="40EEE7B48C74BE9FB6263AD1CBAC29F8" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="362367AE0476F7FE6F33BA14E2207BA5" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="5F294A06E0007CB5FDC8EFF7F5F77BAA" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran: ♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Qom Province</emphasis> <emphasis id="063C512755E185B0C5D495855D7160F1" italics="true" pageId="0" pageNumber="145">Qom Province</emphasis>
: Dasht-e Masileh, : Dasht-e Masileh,
<geoCoordinate degrees="34" direction="north" minutes="47" orientation="latitude" precision="925" value="34.783333">34°47'N</geoCoordinate> <geoCoordinate id="729F8F592B18821BCE57EF5B16A2E7A7" degrees="34" direction="north" minutes="47" orientation="latitude" precision="925" value="34.783333">34°47'N</geoCoordinate>
, ,
<geoCoordinate degrees="51" direction="east" minutes="11" orientation="longitude" precision="925" value="51.183334">51°11'E</geoCoordinate> <geoCoordinate id="3284D87A06E53983F4B8EA878A52C191" degrees="51" direction="east" minutes="11" orientation="longitude" precision="925" value="51.183334">51°11'E</geoCoordinate>
, 11.2017 (M. Mirghazanfari). , 11.2017 (M. Mirghazanfari).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="43A9C13ED5E96F9550DE1616D3789377" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="1E6E5A1FB6B6229D70CCDE963B31603C" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="ECD674A6220A7BF754CEACBB2AE0814D" pageId="0" pageNumber="145">
The specific epithet is derived from The specific epithet is derived from
<normalizedToken originalValue="‘Kavir’">'Kavir'</normalizedToken> <normalizedToken id="3320C6189828C831E44BB2136D4B772C" originalValue="‘Kavir’">'Kavir'</normalizedToken>
, a Persian word for desert, referring to the occurrence of the species in the central deserts of Iran. , a Persian word for desert, referring to the occurrence of the species in the central deserts of Iran.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="1F81D003D25F702331063A2445B67BBA" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="C0899A83683D2FC5E6470B7DDE141D49" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="8D39B779133E88E64E924F91A5735C1E" pageId="0" pageNumber="145">
The male of The male of
<taxonomicName lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis"> <taxonomicName id="AFE5CE602157642213A101E334CCD8C6" lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis> <emphasis id="1EF70A5F5F24B1B44FE46320B724BC43" italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. can be distinguished from the similar sp. nov. can be distinguished from the similar
<taxonomicName lsidName="L. dyachkovi" pageId="0" pageNumber="145" rank="species" species="dyachkovi"> <taxonomicName id="C1B7658179D4C32CA663F6574DF32EE3" lsidName="L. dyachkovi" pageId="0" pageNumber="145" rank="species" species="dyachkovi">
<emphasis italics="true" pageId="0" pageNumber="145">L. dyachkovi</emphasis> <emphasis id="3A7180F9435650C08A0F9936F76B1254" italics="true" pageId="0" pageNumber="145">L. dyachkovi</emphasis>
</taxonomicName> </taxonomicName>
Fomichev &amp; Marusik, 2019 by having a gently bent cheliceral fang ( Fomichev &amp; Marusik, 2019 by having a gently bent cheliceral fang (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="9E6831C9F200F912EBC9329F42D2B5F0" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
bent over 90°, cf. Fig. bent over 90°, cf. Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1C</figureCitation> <figureCitation id="B5D684590E0E33B03A34BE6FD95620C5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1C</figureCitation>
and fig. 15 in and fig. 15 in
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.28.4.08" author="Fomichev, AA" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="556 - 561" refId="B5" refString="Fomichev, AA, Marusik, YM, 2019. A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan. Arthropoda Selecta 28 (4): 556 - 561, DOI: https://doi.org/10.15298/arthsel.28.4.08" title="A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan." url="https://doi.org/10.15298/arthsel.28.4.08" volume="28" year="2019">Fomichev and Marusik 2019</bibRefCitation> <bibRefCitation id="0301D11C0FA22D61052F96E758AE6C5E" DOI="https://doi.org/10.15298/arthsel.28.4.08" author="Fomichev, AA" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="556 - 561" refId="B5" refString="Fomichev, AA, Marusik, YM, 2019. A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan. Arthropoda Selecta 28 (4): 556 - 561, DOI: https://doi.org/10.15298/arthsel.28.4.08" title="A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan." url="https://doi.org/10.15298/arthsel.28.4.08" volume="28" year="2019">Fomichev and Marusik 2019</bibRefCitation>
). The shape of the RTA is also different from other species of the region (stalk with a gentle curve ventrally and tip straightly pointed; cf. Fig. ). The shape of the RTA is also different from other species of the region (stalk with a gentle curve ventrally and tip straightly pointed; cf. Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4I</figureCitation> <figureCitation id="F3E50994048DE8ACA551316E63BF6DF9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4I</figureCitation>
and Fig. and Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4F-H, J</figureCitation> <figureCitation id="1E0695FF0E08863C57682C767F4B6717" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4F-H, J</figureCitation>
). ).
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145" start="Figure 1" startId="F1"> <caption id="6E02DF173E7CCAA75D1160FD8E3AE8BC" doi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="145"> <paragraph id="70A7462D885D40811A3429CF85B87B7E" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 1.</emphasis> <emphasis id="89C53FD05330D9D2077A9D7083CAF455" bold="true" pageId="0" pageNumber="145">Figure 1.</emphasis>
Habitus and chelicerae of male Habitus and chelicerae of male
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis"> <taxonomicName id="6F067D0A69C2959CB658B05CE5A33D62" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana kavirensis</emphasis> <emphasis id="983FFCDC3921BDEFC278449D0B8D47F9" italics="true" pageId="0" pageNumber="145">Lachesana kavirensis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis> <emphasis id="F48042E2001767800412B211E5F7AA9A" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and ) and
<taxonomicName lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus"> <taxonomicName id="29138CFD20DAA6C163A6C315F04A635F" lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis italics="true" pageId="0" pageNumber="145">L. perseus</emphasis> <emphasis id="E0445D307571C5E8230322D4C98CB5EB" italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis> <emphasis id="4524C988A95A19F56DA00C5EF7023E96" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, B, D, E</emphasis> <emphasis id="94212D215E4FB0E23E79327BDEF48081" bold="true" pageId="0" pageNumber="145">A, B, D, E</emphasis>
habitus, dorsal habitus, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">C, F</emphasis> <emphasis id="F32153E33FD758F95B2139BEEA32BC42" bold="true" pageId="0" pageNumber="145">C, F</emphasis>
chelicerae, ventral. chelicerae, ventral.
</paragraph> </paragraph>
</caption> </caption>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="CAADF2ECB101A1BC983F8F3B31A3CD1C" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="13CDC11543B4E7C5463C451ABCD70CCC" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="2259A8733FDE5B904D5BF6F639983779" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis> <emphasis id="C221BC61D6EDF4B20A3FF93023EFCA67" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Habitus as in Fig. Habitus as in Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1A, B</figureCitation> <figureCitation id="A679570090A25AC627FE334E723AF8DB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1A, B</figureCitation>
. Total length 11.55. Carapace 5.80 long, 2.60 wide at pars cephalica, 3.75 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.23, ALE: 0.20, PME: 0.17, PLE: 0.23, PME-PME: 0.14. Carapace, sternum, labium, chelicerae and maxillae light brown. Chelicera (Fig. . Total length 11.55. Carapace 5.80 long, 2.60 wide at pars cephalica, 3.75 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.23, ALE: 0.20, PME: 0.17, PLE: 0.23, PME-PME: 0.14. Carapace, sternum, labium, chelicerae and maxillae light brown. Chelicera (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1C</figureCitation> <figureCitation id="E0CE6D685589F3867E1F35B4E4479CBE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1C</figureCitation>
) with dense, black setae and one promarginal tooth; fangs gently curved, almost straight. Legs the same color as carapace, without annulations and with numerous spines. Abdomen pale, dorsally with a longitudinal dark stripe and several small spots. Spinnerets slightly lighter than abdomen, uniform in color. Measurements of legs: I: 15.49 (4.47, 1.88, 3.33, 3.29, 2.52), II: 15.03 (4.20, 1.79, 3.02, 3.50, 2.52), III: 15.78 (4.08, 1.96, 2.48, 4.71, 2.55), IV: 18.30 (4.58, 2.07, 3.20, 5.24, 3.21). ) with dense, black setae and one promarginal tooth; fangs gently curved, almost straight. Legs the same color as carapace, without annulations and with numerous spines. Abdomen pale, dorsally with a longitudinal dark stripe and several small spots. Spinnerets slightly lighter than abdomen, uniform in color. Measurements of legs: I: 15.49 (4.47, 1.88, 3.33, 3.29, 2.52), II: 15.03 (4.20, 1.79, 3.02, 3.50, 2.52), III: 15.78 (4.08, 1.96, 2.48, 4.71, 2.55), IV: 18.30 (4.58, 2.07, 3.20, 5.24, 3.21).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="304E1DF7FAA0F3E0AA1213524791DAFC" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3A-E</figureCitation> <figureCitation id="81D0B86A58E887F9B50E48105E1A8DA9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3A-E</figureCitation>
, ,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4I</figureCitation> <figureCitation id="E48303C1CB0CE47198597D0A987CE857" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4I</figureCitation>
. RTA long, over 1.2 times longer than tibia, its widest part less than 2 times wider than stalk; cymbium with 2 spines in mesal proximal part; bulb elongate, 1.3 times longer than wide; base of embolus large, 1/3 of the tegulum's length; embolus filamentous, lying in chute formed by conductor. . RTA long, over 1.2 times longer than tibia, its widest part less than 2 times wider than stalk; cymbium with 2 spines in mesal proximal part; bulb elongate, 1.3 times longer than wide; base of embolus large, 1/3 of the tegulum's length; embolus filamentous, lying in chute formed by conductor.
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="8D0F520E623C595BDB6A4BF97F263548" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="3E3506B042F14DB6D86A3B4879E5FC4D" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown. Unknown.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="93138FA7613F60626D020CBFA0315E7F" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="349676EA43DCC9D6FE7D20BC5FDAF437" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="F8D01D058C16801AC1AB2064130704D7" pageId="0" pageNumber="145">
Known only from the type locality in Qom Province, northern Iran (Fig. Known only from the type locality in Qom Province, northern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation> <figureCitation id="4821E25AC0408CC2280B594D11AC825A" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

122
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">Fig. 33</figureCitation> <figureCitation id="5242B22207A8B094F2ABC477E98A30AD" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">Fig. 33</figureCitation>
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: :
<bibRefCitation author="Bosmans, R" journalOrPublisher="Contributions to Natural History" pageId="0" pageNumber="145" pagination="211 - 295" refId="B1" refString="Bosmans, R, 2009. Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). Contributions to Natural History 12: 211 - 295" title="Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae)." volume="12" year="2009">Bosmans 2009</bibRefCitation> <bibRefCitation id="B4D612A916BCB3894A8E0B473FEECA0C" author="Bosmans, R" journalOrPublisher="Contributions to Natural History" pageId="0" pageNumber="145" pagination="211 - 295" refId="B1" refString="Bosmans, R, 2009. Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). Contributions to Natural History 12: 211 - 295" title="Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae)." volume="12" year="2009">Bosmans 2009</bibRefCitation>
: 281, f. 184-185, 194-195 (♂♀). : 281, f. 184-185, 194-195 (♂♀).
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<taxonomicName baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion lutipes" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="lutipes">Zodarion lutipes</taxonomicName> <taxonomicName id="0A2595A380CA604883195AE127F61944" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion lutipes" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="lutipes">Zodarion lutipes</taxonomicName>
For the complete list of references see For the complete list of references see
<bibRefCitation author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC (2021)</bibRefCitation> <bibRefCitation id="B9F140603BD741E771DE97C925926507" author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC (2021)</bibRefCitation>
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="records in iran"> <subSubSection id="C367FCCD3AEA043D79EC0F81A6EDFD11" pageId="0" pageNumber="145" type="records in iran">
<paragraph pageId="0" pageNumber="145">Records in Iran.</paragraph> <paragraph id="526EE248414511F3FBF1CD00411D67CD" pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="D8B0BDCCB18956560BBC702D343B97D5" pageId="0" pageNumber="145">
Kermanshah ( Kermanshah (
<bibRefCitation author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. 2018</bibRefCitation> <bibRefCitation id="C3FE159D89E8DE58294D4A8605106C46" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. 2018</bibRefCitation>
) (Fig. ) (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation> <figureCitation id="A7FDFB48732D4B197C09EF6154C606FA" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="8BECA0E90DC1BF62AE0A6CCD601AD5DE" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="FED0E0824A846302B36454FBC67ED893" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">Cyprus, Israel, Lebanon, Jordan, Iran.</paragraph> <paragraph id="E17A484BCC7B356DAA73281F1CC5EDBB" pageId="0" pageNumber="145">Cyprus, Israel, Lebanon, Jordan, Iran.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comments"> <subSubSection id="8B14EB2B27A30EFECB92DF3FDA613D8F" pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="0A4B28D550755EB70FD75067633729A1" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="CA2DDD2CB6D56872CFA4E5214B9983F2" pageId="0" pageNumber="145">
Unfortunately, we were not able to re-examine the material studied by Unfortunately, we were not able to re-examine the material studied by
<bibRefCitation author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. (2018)</bibRefCitation> <bibRefCitation id="CF57EAF2568E85D11B31A44276F031B5" author="Zamani, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="145" refId="B30" refString="Zamani, A, Mirshamsi, O, Kashani, GM, Karami, L, 2018. New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13(2, for 2017): 183-197." title="New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics 13 (2, for 2017): 183 - 197." year="2018">Zamani et al. (2018)</bibRefCitation>
, therefore, no figures are provided for this species. , therefore, no figures are provided for this species.
</paragraph> </paragraph>
</subSubSection> </subSubSection>

226
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation> <mods:affiliation id="BFCE586D6E467F10BF117692D268CC58">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<subSection pageId="0" pageNumber="145" type="taxonomy"> <subSection id="48D53AB326946B3B236BB260F90E85A5" pageId="0" pageNumber="145" type="taxonomy">
<treatment ID-GBIF-Taxon="181753574" LSID="urn:lsid:plazi:treatment:8837EB822336513D8735EAA812AFC392" httpUri="http://treatment.plazi.org/id/8837EB822336513D8735EAA812AFC392" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="8837EB822336513D8735EAA812AFC392" ID-GBIF-Taxon="181753574" LSID="urn:lsid:plazi:treatment:8837EB822336513D8735EAA812AFC392" httpUri="http://treatment.plazi.org/id/8837EB822336513D8735EAA812AFC392" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="5F7894CD2F9A79BF0299F44D9573262E" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="C49AD006B9C5B4BA221D5823C27E8057" pageId="0" pageNumber="145">
<taxonomicName LSID="8837EB82-2336-513D-8735-EAA812AFC392" authority="Dunin &amp; Nenilin, 1987" authorityName="Dunin &amp; Nenilin" authorityYear="1987" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion talyschicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyschicum">Zodarion talyschicum Dunin &amp; Nenilin, 1987</taxonomicName> <taxonomicName id="B55996BC3F78FF7E622466B8052040D4" ID-CoL="5D6DJ" LSID="8837EB82-2336-513D-8735-EAA812AFC392" authority="Dunin &amp; Nenilin, 1987" authorityName="Dunin &amp; Nenilin" authorityYear="1987" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion talyschicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyschicum">Zodarion talyschicum Dunin &amp; Nenilin, 1987</taxonomicName>
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A) and Z. talyschicum (B, C) A, B ventral C ventroretrolateral. Abbreviations: Tm - median tegular process, Tp - prolateral tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145">Figs 29B, C</figureCitation> <figureCitation id="44C503980295675AB54B16A69219F0E5" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A) and Z. talyschicum (B, C) A, B ventral C ventroretrolateral. Abbreviations: Tm - median tegular process, Tp - prolateral tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145">Figs 29B, C</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation> <figureCitation id="DE85063C2E72A4959679DC0BDD5A04F5" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="reference_group"> <subSubSection id="00E29607B5A138FFFA84A86496690E76" pageId="0" pageNumber="145" type="reference_group">
<paragraph pageId="0" pageNumber="145"> <paragraph id="A45B0D3D0049819ACCC297F5687614F1" pageId="0" pageNumber="145">
<taxonomicName authorityName="Dunin &amp; Nenilin" authorityYear="1987" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion talyschicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyschicum">Zodarion talyschicum</taxonomicName> <taxonomicName id="C028A07D6BC4F19026CABFF529CFBF12" authorityName="Dunin &amp; Nenilin" authorityYear="1987" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion talyschicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyschicum">Zodarion talyschicum</taxonomicName>
Dunin &amp; Nenilin, 1987: 196, f. 14-18 (♂♀). Dunin &amp; Nenilin, 1987: 196, f. 14-18 (♂♀).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="68249AE9672CEF8DB20885309D7BC936" pageId="0" pageNumber="145">
<taxonomicName authorityName="Dunin &amp; Nenilin" authorityYear="1987" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion talyschicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyschicum">Zodarion talyschicum</taxonomicName> <taxonomicName id="B805D890C67F1D392FD83D8F723F87F8" authorityName="Dunin &amp; Nenilin" authorityYear="1987" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion talyschicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyschicum">Zodarion talyschicum</taxonomicName>
: :
<bibRefCitation DOI="https://doi.org/10.13156/arac.2020.18.6.569" author="Zamani, A" journalOrPublisher="Arachnology" pageId="0" pageNumber="145" pagination="569 - 591" refId="B27" refString="Zamani, A, Dimitrov, D, Weiss, I, Alimohammadi, S, Rafiei-Jahed, R, Esyunin, SL, Moradmand, M, Chatzaki, M, Marusik, YM, 2020. New data on the spider fauna of Iran (Arachnida: Araneae), Part VII. Arachnology 18 (6): 569 - 591, DOI: https://doi.org/10.13156/arac.2020.18.6.569" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part VII." url="https://doi.org/10.13156/arac.2020.18.6.569" volume="18" year="2020">Zamani et al. 2020</bibRefCitation> <bibRefCitation id="1BDD8C49D2601A97B6EB8743D4805669" DOI="https://doi.org/10.13156/arac.2020.18.6.569" author="Zamani, A" journalOrPublisher="Arachnology" pageId="0" pageNumber="145" pagination="569 - 591" refId="B27" refString="Zamani, A, Dimitrov, D, Weiss, I, Alimohammadi, S, Rafiei-Jahed, R, Esyunin, SL, Moradmand, M, Chatzaki, M, Marusik, YM, 2020. New data on the spider fauna of Iran (Arachnida: Araneae), Part VII. Arachnology 18 (6): 569 - 591, DOI: https://doi.org/10.13156/arac.2020.18.6.569" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part VII." url="https://doi.org/10.13156/arac.2020.18.6.569" volume="18" year="2020">Zamani et al. 2020</bibRefCitation>
: 589, f. 12D-F (♂♀). : 589, f. 12D-F (♂♀).
</paragraph> </paragraph>
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<paragraph pageId="0" pageNumber="145">Material.</paragraph> <paragraph id="BAD750EA429AC774BA460FE9C484EDD3" pageId="0" pageNumber="145">Material.</paragraph>
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<collectingCountry name="Iran">Iran</collectingCountry> <collectingCountry id="1E5453AD2CBC9B0E2325933E384D279D" name="Iran">Iran</collectingCountry>
: :
<emphasis italics="true" pageId="0" pageNumber="145"> <emphasis id="2003BD1A1725748A14825A832185E181" italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Gilan">Gilan Province</collectingRegion> <collectingRegion id="5B32214C778D4A1172055BCF7001E577" country="Iran" name="Gilan">Gilan Province</collectingRegion>
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(MHNG), (MHNG),
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, ,
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; ;
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<emphasis italics="true" pageId="0" pageNumber="145"> <emphasis id="F94BA5E5CB4516BAD56209D9A4411210" italics="true" pageId="0" pageNumber="145">
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</emphasis> </emphasis>
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<specimenCount type="female">3♀</specimenCount> <specimenCount id="1B3ABE6E631FAD5C5CD439C23C518A25" type="female">3♀</specimenCount>
(MHNG), (MHNG),
<collectingMunicipality>Gorgan</collectingMunicipality> <collectingMunicipality id="9FD6C8B771D0EB4502EEF8B5EF16EAAA">Gorgan</collectingMunicipality>
, ,
<location LSID="urn:lsid:plazi:treatment:8837EB822336513D8735EAA812AFC392:6E2DA9D776891C7A93329FD8CBDC4692" country="Iran" latitude="36.733334" longLatPrecision="1185" longitude="54.483334" municipality="Gorgan" name="Naharkhoran" stateProvince="Golestan Province">Naharkhoran</location> <location id="6E2DA9D776891C7A93329FD8CBDC4692" LSID="urn:lsid:plazi:treatment:8837EB822336513D8735EAA812AFC392:6E2DA9D776891C7A93329FD8CBDC4692" country="Iran" latitude="36.733334" longLatPrecision="1185" longitude="54.483334" municipality="Gorgan" name="Naharkhoran" stateProvince="Golestan Province">Naharkhoran</location>
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<emphasis italics="true" pageId="0" pageNumber="145"> <emphasis id="1107DD41BCD75258A8E197DBBBED4B1D" italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Hamadan">Hamedan Province</collectingRegion> <collectingRegion id="50EC0810C97FF242D66F7EE452858AD0" country="Iran" name="Hamadan">Hamedan Province</collectingRegion>
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<specimenCount type="male">1♂</specimenCount> <specimenCount id="BD510DBC998BE5D99D5C3F0AC9BD9AEB" type="male">1♂</specimenCount>
(NHMW), (NHMW),
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<quantity metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity> <quantity id="DB231006CF5CFD55A070FD3BB7670C9F" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
SW of Shahpasand SW of Shahpasand
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, ,
<collectingDate value="1972-04-26">26.04.1972</collectingDate> <collectingDate id="F9833F874D3EE6C1C8CC412293CBA9E6" value="1972-04-26">26.04.1972</collectingDate>
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; ;
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<collectingRegion country="Iran" name="Mazandaran">Mazandaran Province</collectingRegion> <collectingRegion id="4B97FA36557F469DA4AE8436295B0F40" country="Iran" name="Mazandaran">Mazandaran Province</collectingRegion>
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. .
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="records in iran"> <subSubSection id="94B0E954FFE85CC24E556DC0AA3387BB" pageId="0" pageNumber="145" type="records in iran">
<paragraph pageId="0" pageNumber="145">Records in Iran.</paragraph> <paragraph id="CF12A06414CA9F7000F53C4DB00C3B67" pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="71EE795F53F87E4AA95000F81D5E38C3" pageId="0" pageNumber="145">
Golestan ( Golestan (
<bibRefCitation DOI="https://doi.org/10.13156/arac.2020.18.6.569" author="Zamani, A" journalOrPublisher="Arachnology" pageId="0" pageNumber="145" pagination="569 - 591" refId="B27" refString="Zamani, A, Dimitrov, D, Weiss, I, Alimohammadi, S, Rafiei-Jahed, R, Esyunin, SL, Moradmand, M, Chatzaki, M, Marusik, YM, 2020. New data on the spider fauna of Iran (Arachnida: Araneae), Part VII. Arachnology 18 (6): 569 - 591, DOI: https://doi.org/10.13156/arac.2020.18.6.569" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part VII." url="https://doi.org/10.13156/arac.2020.18.6.569" volume="18" year="2020">Zamani et al. 2020</bibRefCitation> <bibRefCitation id="511EDA6FD63E757062DD8CC2F27E7BFA" DOI="https://doi.org/10.13156/arac.2020.18.6.569" author="Zamani, A" journalOrPublisher="Arachnology" pageId="0" pageNumber="145" pagination="569 - 591" refId="B27" refString="Zamani, A, Dimitrov, D, Weiss, I, Alimohammadi, S, Rafiei-Jahed, R, Esyunin, SL, Moradmand, M, Chatzaki, M, Marusik, YM, 2020. New data on the spider fauna of Iran (Arachnida: Araneae), Part VII. Arachnology 18 (6): 569 - 591, DOI: https://doi.org/10.13156/arac.2020.18.6.569" title="New data on the spider fauna of Iran (Arachnida: Araneae), Part VII." url="https://doi.org/10.13156/arac.2020.18.6.569" volume="18" year="2020">Zamani et al. 2020</bibRefCitation>
). New records for Gilan, Hamedan, Mazandaran and North Khorasan, with the latter representing the easternmost record in the whole range (Fig. ). New records for Gilan, Hamedan, Mazandaran and North Khorasan, with the latter representing the easternmost record in the whole range (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation> <figureCitation id="50EF9BFACB746FC85AF2A483B719DA3C" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
). ).
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<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="81E1B4C0245468E6D27BC7970A9A6AFE" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">Azerbaijan, Iran.</paragraph> <paragraph id="ED2B9F6EFD90B88960889EA8E46F463B" pageId="0" pageNumber="145">Azerbaijan, Iran.</paragraph>
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<paragraph pageId="0" pageNumber="145"> <paragraph id="83A16D3B98F2C81C3F2C7805E36DDCBC" pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/DA859B09-12D2-4370-A6B6-93C1BF274678" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium niusha" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="niusha" status="sp. nov.">Acanthinozodium niusha</taxonomicName> <taxonomicName id="E594F0B8027AF80396A85C58077F827C" ID-CoL="98YZC" LSID="http://zoobank.org/DA859B09-12D2-4370-A6B6-93C1BF274678" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium niusha" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="niusha" status="sp. nov.">Acanthinozodium niusha</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="772616B96D34DE75FC7B7C1845427471" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11C-E</figureCitation> <figureCitation id="5C6F4D845EAD9FC1F2AC67839E5A2DD7" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11C-E</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">, 13A-E</figureCitation> <figureCitation id="EA4A8DAD9F3E09AE017C195A9C3099CB" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">, 13A-E</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">, 14G, H</figureCitation> <figureCitation id="85789A7CC8DC17BBFA09663C0AB9B58F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">, 14G, H</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">, 15D-F</figureCitation> <figureCitation id="852FD67D9347720F26BB7D97CBE292FC" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">, 15D-F</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">, 17A-C</figureCitation> <figureCitation id="9357D749088B644DE2BF9B0E7D3ABE1C" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">, 17A-C</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation> <figureCitation id="1FD718CA57FBFF05BF9C4FB9BC8B508F" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="773554C7B364C4C15299410429D1E1F3" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="19F82786918DA1F6652D6353E832869D" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="679516DC484ED555CCC0A6AD2AF01855" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="8CCEC5022FB6DC429F61C40D4E6716E5" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran: ♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Markazi Province</emphasis> <emphasis id="D9814A07E46909EBA029D06DFFEA65BE" italics="true" pageId="0" pageNumber="145">Markazi Province</emphasis>
: Shazand, : Shazand,
<geoCoordinate degrees="33" direction="north" minutes="55" orientation="latitude" precision="925" value="33.916668">33°55'N</geoCoordinate> <geoCoordinate id="98AB602EF3431F313B6D4227E8DF0226" degrees="33" direction="north" minutes="55" orientation="latitude" precision="925" value="33.916668">33°55'N</geoCoordinate>
, ,
<geoCoordinate degrees="49" direction="east" minutes="24" orientation="longitude" precision="925" value="49.4">49°24'E</geoCoordinate> <geoCoordinate id="84075F0A5F7D51B2CA91F2F048F67A2D" degrees="49" direction="east" minutes="24" orientation="longitude" precision="925" value="49.4">49°24'E</geoCoordinate>
, 11.04.2015 (A. Zamani). , 11.04.2015 (A. Zamani).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis> <emphasis id="E12578E7F03BE1E02D5DBCCD274A1AF6" bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
: 5♂3♀ (MHNG), same data as holotype; : 5♂3♀ (MHNG), same data as holotype;
<emphasis italics="true" pageId="0" pageNumber="145">Isfahan Province</emphasis> <emphasis id="7344859D1980DCE8167417A6B3780F90" italics="true" pageId="0" pageNumber="145">Isfahan Province</emphasis>
: 1♂1♀ (MHNG), Riz-e Landjan, : 1♂1♀ (MHNG), Riz-e Landjan,
<geoCoordinate degrees="32" direction="north" minutes="24" orientation="latitude" precision="925" value="32.4">32°24'N</geoCoordinate> <geoCoordinate id="1749420FF805491633D66D7A14286B06" degrees="32" direction="north" minutes="24" orientation="latitude" precision="925" value="32.4">32°24'N</geoCoordinate>
, ,
<geoCoordinate degrees="51" direction="east" minutes="19" orientation="longitude" precision="925" value="51.316666">51°19'E</geoCoordinate> <geoCoordinate id="1B6CA51454FEA96BF6739EF29BD9F680" degrees="51" direction="east" minutes="19" orientation="longitude" precision="925" value="51.316666">51°19'E</geoCoordinate>
, 11.08.1973 (A. Senglet); , 11.08.1973 (A. Senglet);
<emphasis italics="true" pageId="0" pageNumber="145">Fars Province</emphasis> <emphasis id="C0099BEBE31E7435D31B281609496967" italics="true" pageId="0" pageNumber="145">Fars Province</emphasis>
: 1♂1♀ (MHNG), Izad Khast, : 1♂1♀ (MHNG), Izad Khast,
<geoCoordinate degrees="31" direction="north" minutes="31" orientation="latitude" precision="925" value="31.516666">31°31'N</geoCoordinate> <geoCoordinate id="2A62568DE8A0CC6CE8F93EE65393F766" degrees="31" direction="north" minutes="31" orientation="latitude" precision="925" value="31.516666">31°31'N</geoCoordinate>
, ,
<geoCoordinate degrees="52" direction="east" minutes="08" orientation="longitude" precision="925" value="52.133335">52°08'E</geoCoordinate> <geoCoordinate id="DF802E175788AFD890E23F67B2FAB343" degrees="52" direction="east" minutes="08" orientation="longitude" precision="925" value="52.133335">52°08'E</geoCoordinate>
, 12.06.1974 (A. Senglet). , 12.06.1974 (A. Senglet).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="DAF26C7E580B689E27983FAF60F025B3" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="81663E6C5D4D24E7B4454793EAE0FEF1" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet is a Persian feminine name meaning &quot;good listener&quot;. Noun in apposition.</paragraph> <paragraph id="9B1E8A14008B8371EBFF5884B4709378" pageId="0" pageNumber="145">The specific epithet is a Persian feminine name meaning &quot;good listener&quot;. Noun in apposition.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="2DE591BB9AEE26064419358FB0CA8707" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="662D9FABD61B81FB5344988063F47ED2" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="C8F4A18C32CBC7BD8A93C6EA870B83FF" pageId="0" pageNumber="145">
The male of the new species is most similar to The male of the new species is most similar to
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis"> <taxonomicName id="ECB9F8E89D27D4F556E73ADC70A001F8" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis> <emphasis id="B2B4C8AC4B0E1CA04F1892FDF52F0E29" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. from which it differs by a sharply tapering RTA, shorter tip of the cymbium (cf. Fig. sp. nov. from which it differs by a sharply tapering RTA, shorter tip of the cymbium (cf. Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13A</figureCitation> <figureCitation id="BAFF7BF4D85BA3ABCDFEED882FC3CA59" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13A</figureCitation>
and and
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13F</figureCitation> <figureCitation id="652DDB644C820FCF988BCDD85BBB805A" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13F</figureCitation>
), the angle of embolus and the claw of the median apophysis. Female of ), the angle of embolus and the claw of the median apophysis. Female of
<taxonomicName lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha"> <taxonomicName id="403958DEB399F0754DA70374BB78DC3C" lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">A. niusha</emphasis> <emphasis id="1CDED917EE133F9408AC019077645CC3" italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. is similar to sp. nov. is similar to
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa"> <taxonomicName id="F18FDA697E71365AEC9C44A0E4B59FE2" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis> <emphasis id="6ED35F1653FDC06712944DF2C8103DE4" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. by having a similar anterior hood but differs by the longer furrow leading to the hood (as long as receptacle sp. nov. by having a similar anterior hood but differs by the longer furrow leading to the hood (as long as receptacle
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="AFC64046F704C011A9126505999E4EE0" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
1.5 times shorter) and less separated receptacles. 1.5 times shorter) and less separated receptacles.
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145" start="Figure 13" startId="F13"> <caption id="E69FD088AE0608560359FDDAA58AE99E" doi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="145"> <paragraph id="128C1BC78583456F04FED0E2DBB2D4D8" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 13.</emphasis> <emphasis id="ADF2639395D6E3CB5B85599A45F0E23F" bold="true" pageId="0" pageNumber="145">Figure 13.</emphasis>
Male palps of Male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium niusha" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="niusha"> <taxonomicName id="BFA2EBD102BE1115C255CF57654FDBAB" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium niusha" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium niusha</emphasis> <emphasis id="2BF089909DD6EE4721C097E1D128B226" italics="true" pageId="0" pageNumber="145">Acanthinozodium niusha</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-E</emphasis> <emphasis id="0CD884D84EB32A044777E5CF51D55BFF" bold="true" pageId="0" pageNumber="145">A-E</emphasis>
) and ) and
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis"> <taxonomicName id="007B7EA9F715CFAF7C06E1F3568ABE95" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis> <emphasis id="28442C0E98866FFB100641C5107502D1" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">F-I</emphasis> <emphasis id="8CE29022413715E23CEB093254451A89" bold="true" pageId="0" pageNumber="145">F-I</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, D, G, F</emphasis> <emphasis id="8EB596FFAD6405C53C6DF18E8405DC06" bold="true" pageId="0" pageNumber="145">A, D, G, F</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, H</emphasis> <emphasis id="8EAA69130BE44E85412710637CBD4548" bold="true" pageId="0" pageNumber="145">B, H</emphasis>
retrolateral retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, I</emphasis> <emphasis id="1ED02DEBB44BE075F2B411DEEFD8708F" bold="true" pageId="0" pageNumber="145">C, I</emphasis>
dorsal dorsal
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis> <emphasis id="C1E5FE42CBB2966273EE9B740B40A1C5" bold="true" pageId="0" pageNumber="145">E</emphasis>
dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise. dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph> </paragraph>
</caption> </caption>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="397B7531233C101BD44629B14C270AD1" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="E5AE8A372C63317333DBAAD64B3C2C43" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="54F24A1347143A17E64B9962BCE388C9" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis> <emphasis id="B7A410A364E9F0A689A4E9C5CB710C05" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig. (holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11C, D</figureCitation> <figureCitation id="E8663A405C0669D7981576E3047C37A4" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11C, D</figureCitation>
. Total length 2.09. Carapace 0.94 long, 0.46 wide at pars cephalica, 0.69 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.10, ALE: 0.07, PME: 0.06, PLE: 0.06, PME-PME: 0.12. Carapace, sternum, labium, chelicerae and maxillae yellowish; carapace with irregular dark patterns. Chelicera with one retromarginal tooth. Legs yellowish, without annulations. Abdomen dorsally black, grayish ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 1.67+missing tarsus (0.74, 0.31, 0.62, missing), II: 2.56 (0.66, 0.30, 0.52, 0.59, 0.49), III: 2.43 (0.60, 0.33, 0.46, 0.59, 0.45), IV: 3.43 (0.90, 0.31, 0.81, 0.89, 0.52). . Total length 2.09. Carapace 0.94 long, 0.46 wide at pars cephalica, 0.69 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.10, ALE: 0.07, PME: 0.06, PLE: 0.06, PME-PME: 0.12. Carapace, sternum, labium, chelicerae and maxillae yellowish; carapace with irregular dark patterns. Chelicera with one retromarginal tooth. Legs yellowish, without annulations. Abdomen dorsally black, grayish ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 1.67+missing tarsus (0.74, 0.31, 0.62, missing), II: 2.56 (0.66, 0.30, 0.52, 0.59, 0.49), III: 2.43 (0.60, 0.33, 0.46, 0.59, 0.45), IV: 3.43 (0.90, 0.31, 0.81, 0.89, 0.52).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="1F71DF334B658AB6742BB92C4DC6E3E3" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13A-E</figureCitation> <figureCitation id="346B1056219D832350BE7C61B95022DC" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male palps of Acanthinozodium niusha sp. nov. (A-E) and A. parysatis sp. nov. (F-I) A, D, G, F ventral B, H retrolateral C, I dorsal E dorsoretrolateral. Arrow on C pointing to cymbial groove. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure13" httpUri="https://binary.pensoft.net/fig/536890" pageId="0" pageNumber="145">13A-E</figureCitation>
, ,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14G, H</figureCitation> <figureCitation id="876337F2AE6424309EE6F8509B97A039" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14G, H</figureCitation>
, ,
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">15D-F</figureCitation> <figureCitation id="4700B662A66B2410DD93292CFA13A93E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of the male palps of Acanthinozodium parysatis sp. nov. (A-C) and A. niusha sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsoretrolateral and dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145">15D-F</figureCitation>
. RTA ( . RTA (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis> <emphasis id="CF198E27A8431795BF9B84344373E339" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
) as long as wide and as long as tibia, basal part very wide, sharply tapering, tip bent ventrally; cymbium 1.5 times longer than wide; posterior part of tegulum with broad and indistinct projection postero-prolaterally; median apophysis ( ) as long as wide and as long as tibia, basal part very wide, sharply tapering, tip bent ventrally; cymbium 1.5 times longer than wide; posterior part of tegulum with broad and indistinct projection postero-prolaterally; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="4824F75DA06E27969E602DE2462998CE" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) with massive base, much larger than the claw of median apophysis; sperm duct gradually tracking along margin of tegulum; tip of embolus ( ) with massive base, much larger than the claw of median apophysis; sperm duct gradually tracking along margin of tegulum; tip of embolus (
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis> <emphasis id="8548FEBFE1D14EA0186BA6CC854F70A9" italics="true" pageId="0" pageNumber="145">Em</emphasis>
) straight and lacking any processes or tubercles. ) straight and lacking any processes or tubercles.
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145" start="Figure 14" startId="F14"> <caption id="558EED32F920BDB5DB3AEEECF23039FA" doi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="145"> <paragraph id="5C01E8BAE72639CD8C847CE55EEF5293" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 14.</emphasis> <emphasis id="A7AE04F1E4632EC479DE98095701B74A" bold="true" pageId="0" pageNumber="145">Figure 14.</emphasis>
SEM images of the male palps of SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa"> <taxonomicName id="4583400910C60D0A2492F48F346CDDF0" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis> <emphasis id="D75CEDF4818AD26E6B612A038440EBD0" italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis> <emphasis id="117C9245161D06B13F09B66C52418643" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
), ),
<taxonomicName lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani"> <taxonomicName id="6D24C2B2587DA2923C97AFBA23A6D114" lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis italics="true" pageId="0" pageNumber="145">A. sorani</emphasis> <emphasis id="9A421C5C487E8CBCB82A039FB2AAF118" italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis> <emphasis id="EFD011C237EA83CE5C528A55ACFA2A68" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
), ),
<taxonomicName lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha"> <taxonomicName id="9F04E21CEF91B66B9BDF220FB1397BAF" lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">A. niusha</emphasis> <emphasis id="332846E4458418AFFB16965BE0516184" italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">G, H</emphasis> <emphasis id="25B4458B32A8BA4EB27A76E77646C14D" bold="true" pageId="0" pageNumber="145">G, H</emphasis>
) and ) and
<taxonomicName lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa"> <taxonomicName id="BE1F841A3C2DE950436BF475E975A624" lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis> <emphasis id="F318A1749C37C601ADD6E888F12BD81F" italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">I</emphasis> <emphasis id="A920C43DDE3B4695A595E92D8BC8DA4E" bold="true" pageId="0" pageNumber="145">I</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, C, D, F, I</emphasis> <emphasis id="5F4FBE4437FE622EBBDDB59653CE775C" bold="true" pageId="0" pageNumber="145">A, C, D, F, I</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, E</emphasis> <emphasis id="CAA5F1A1586CFD2B9E5112D4C5DB7BDB" bold="true" pageId="0" pageNumber="145">B, E</emphasis>
retrolateral retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">G, H</emphasis> <emphasis id="815B0007C7C984161E49CF3D1DE22D90" bold="true" pageId="0" pageNumber="145">G, H</emphasis>
cymbial groove, dorsal. Abbreviations: cymbial groove, dorsal. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Cg</emphasis> <emphasis id="18E1F734337B1E185A438C8285F5FA14" italics="true" pageId="0" pageNumber="145">Cg</emphasis>
- cymbial groove, - cymbial groove,
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis> <emphasis id="ECCB6A7DA89211BDBE643930AB1D1A17" italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor, - conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis> <emphasis id="56267A4F25D2B39B6DD5600523B0FD3B" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus, - embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Ep</emphasis> <emphasis id="3678A563FC1AEEC6B1DEF1DF80A54C99" italics="true" pageId="0" pageNumber="145">Ep</emphasis>
- embolic process, - embolic process,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="26E04396C32BB55B0CD83766E684345A" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis, - median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis> <emphasis id="710BD691A0176DB1784E67B235D98F02" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral tibial apophysis, - retrolateral tibial apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Tp</emphasis> <emphasis id="3506F0BC39E9BED926E2F9951E5E7F51" italics="true" pageId="0" pageNumber="145">Tp</emphasis>
- tegular process. Scale bars: 0.1 mm, unless stated otherwise. - tegular process. Scale bars: 0.1 mm, unless stated otherwise.
</paragraph> </paragraph>
</caption> </caption>
<caption doi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145" start="Figure 15" startId="F15"> <caption id="4618E753572298948B753463BA1CF1C3" doi="10.3897/zookeys.1035.65767.figure15" httpUri="https://binary.pensoft.net/fig/536892" pageId="0" pageNumber="145" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="145"> <paragraph id="C4144F250047F587DBDD253EF33EFFDD" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 15.</emphasis> <emphasis id="34341633D1B6AAAA80342F36B9D23E15" bold="true" pageId="0" pageNumber="145">Figure 15.</emphasis>
SEM images of the male palps of SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parysatis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parysatis"> <taxonomicName id="0959B9F06C3A2BC85CCC0D2D77E6471A" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parysatis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium parysatis</emphasis> <emphasis id="10B2D196E66484CBB3022CB5C101AFE7" italics="true" pageId="0" pageNumber="145">Acanthinozodium parysatis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis> <emphasis id="4909BC6A29EAEC5FD94CF21A1CC65D4D" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and ) and
<taxonomicName lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha"> <taxonomicName id="99DACC53332A46BB2E363A1E4CAC69D4" lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">A. niusha</emphasis> <emphasis id="B2F8D0DADE0B32E9B4BFD963B952E585" italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis> <emphasis id="B6A1DC5D63066F789C74318DBA53E931" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis> <emphasis id="124038BCF5AF3D97D497762FBB59C9FD" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, E</emphasis> <emphasis id="9C592968A0EDF7B8558B3A971FDDCB71" bold="true" pageId="0" pageNumber="145">B, E</emphasis>
retrolateral retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, F</emphasis> <emphasis id="99B3DDB5D91CF456E8AED81EAA55E243" bold="true" pageId="0" pageNumber="145">C, F</emphasis>
dorsoretrolateral and dorsal. Abbreviations: dorsoretrolateral and dorsal. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Cg</emphasis> <emphasis id="137D124C2DBA8776136B925FA2898B6C" italics="true" pageId="0" pageNumber="145">Cg</emphasis>
- cymbial groove, - cymbial groove,
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis> <emphasis id="6A0B8A321EF87847052A34A42F4D148B" italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor, - conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis> <emphasis id="32717E028130CE0039F61335F8822316" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus, - embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="DAF766E00BE403C6D3737B2FBD9E8452" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis, - median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis> <emphasis id="57AA06C318FD299C83029BA72808EA24" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral tibial apophysis, - retrolateral tibial apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis> <emphasis id="45785417A85D7692A129BB33E411DB29" italics="true" pageId="0" pageNumber="145">Va</emphasis>
- ventral apophysis. Scale bars: 0.1 mm. - ventral apophysis. Scale bars: 0.1 mm.
</paragraph> </paragraph>
</caption> </caption>
<caption doi="10.3897/zookeys.1035.65767.figure16" httpUri="https://binary.pensoft.net/fig/536893" pageId="0" pageNumber="145" start="Figure 16" startId="F16"> <caption id="603D79D238830DFA5470520D56DADF73" doi="10.3897/zookeys.1035.65767.figure16" httpUri="https://binary.pensoft.net/fig/536893" pageId="0" pageNumber="145" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="145"> <paragraph id="8799FAE0730B56C8B7991D24F659C6AF" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 16.</emphasis> <emphasis id="6AB47E1977FB9D95768DCF5D16031A47" bold="true" pageId="0" pageNumber="145">Figure 16.</emphasis>
Epigyne of Epigyne of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa"> <taxonomicName id="1891F47FCAE64E23055D416AD9B44C60" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis> <emphasis id="AA9504AC7697BBAA440E9227B3827003" italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis> <emphasis id="46024445CE06CDB4D9F764B944B71B6A" bold="true" pageId="0" pageNumber="145">A</emphasis>
posteroventral posteroventral
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis> <emphasis id="649D986595EE325D3FABD5B3F064DBC1" bold="true" pageId="0" pageNumber="145">B</emphasis>
posterior posterior
<emphasis bold="true" pageId="0" pageNumber="145">C, D</emphasis> <emphasis id="D356F865C2BCE96F73859B43F5EE9F27" bold="true" pageId="0" pageNumber="145">C, D</emphasis>
ventral. Scale bar: 0.2 mm. ventral. Scale bar: 0.2 mm.
</paragraph> </paragraph>
</caption> </caption>
<paragraph pageId="0" pageNumber="145"> <paragraph id="FC8F8A8F32B4F71A2442BA3EB4D0F2DE" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="94A6D579B3FC1E4A6AF8725AF7ECCA73" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig. Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11E</figureCitation> <figureCitation id="D483E1911A9CFABB850225E4D4E14EC2" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11E</figureCitation>
. Total length 2.10. Carapace 0.86 long, 0.44 wide at pars cephalica, 0.69 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.07, ALE: 0.06, PME: 0.05, PLE: 0.05, PME-PME: 0.10. Coloration as in male. Measurements of legs: I: 2.68 (0.73, 0.30, 0.59, 0.61, 0.45), II: 2.58 (0.74, 0.24, 0.53, 0.58, 0.49), III: 2.32 (0.59, 0.29, 0.50, 0.55, 0.39), IV: 3.45 (0.89, 0.33, 0.80, 0.92, 0.51). . Total length 2.10. Carapace 0.86 long, 0.44 wide at pars cephalica, 0.69 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.07, ALE: 0.06, PME: 0.05, PLE: 0.05, PME-PME: 0.10. Coloration as in male. Measurements of legs: I: 2.68 (0.73, 0.30, 0.59, 0.61, 0.45), II: 2.58 (0.74, 0.24, 0.53, 0.58, 0.49), III: 2.32 (0.59, 0.29, 0.50, 0.55, 0.39), IV: 3.45 (0.89, 0.33, 0.80, 0.92, 0.51).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="44A54F81748E904207DEBB16FC7BC5CF" pageId="0" pageNumber="145">
Epigyne as in Fig. Epigyne as in Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">17A-C</figureCitation> <figureCitation id="EDE5490914C279263DD3C5D7B79F1BA7" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">17A-C</figureCitation>
. Fovea about 3 times wider than long; hood as wide as fovea, bursae ovoid, weakly sclerotized, with fine pores (Fig. . Fovea about 3 times wider than long; hood as wide as fovea, bursae ovoid, weakly sclerotized, with fine pores (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">17C</figureCitation> <figureCitation id="72697654BD085BF89B7B110B0F6DB69C" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ventral view of epigynes of Acanthinozodium niusha sp. nov. (A-C) and A. parysatis sp. nov. (D-F). Scale bar: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145">17C</figureCitation>
); receptacles round, spaced by over 5 diameters. ); receptacles round, spaced by over 5 diameters.
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145" start="Figure 17" startId="F17"> <caption id="BE62E9BC05399FFFBFA88E4D7C3798F5" doi="10.3897/zookeys.1035.65767.figure17" httpUri="https://binary.pensoft.net/fig/536894" pageId="0" pageNumber="145" start="Figure 17" startId="F17">
<paragraph pageId="0" pageNumber="145"> <paragraph id="A67FADEA663E574667952EAC0A776B88" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 17.</emphasis> <emphasis id="0B1A27F53247A5891D5939DE9F36AA4B" bold="true" pageId="0" pageNumber="145">Figure 17.</emphasis>
Ventral view of epigynes of Ventral view of epigynes of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium niusha" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="niusha"> <taxonomicName id="5F3F74B56787F98133E20916F66DB755" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium niusha" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium niusha</emphasis> <emphasis id="8DF0428BC841ACC56197C0AE9582866C" italics="true" pageId="0" pageNumber="145">Acanthinozodium niusha</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis> <emphasis id="415916AD278B51AE6F6A4F557F546716" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and ) and
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis"> <taxonomicName id="58BB2D116985E45AE09CEF281CA565FA" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis> <emphasis id="3FE21E04161166348C4D871DB6316266" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis> <emphasis id="40E97B2A26EDD02AD9BB813B5280CCD5" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
). Scale bar: 0.2 mm, unless stated otherwise. ). Scale bar: 0.2 mm, unless stated otherwise.
</paragraph> </paragraph>
</caption> </caption>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="2E9E80ABE7B9854C3FDE9F8C7A810DC3" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="5A360DA5166A8F402D74B955B8913936" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="08E6675554BCD6BD47F019E165C519E8" pageId="0" pageNumber="145">
Known only from the listed localities in Fars, Isfahan and Markazi provinces, central Iran (Fig. Known only from the listed localities in Fars, Isfahan and Markazi provinces, central Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation> <figureCitation id="123AE02DCE2B7BF5112A54FECF92247E" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

180
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title> <mods:title id="F013EB37938C37D32EEF6D743D216267">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation> <mods:affiliation id="23AA9F61202447333060F358D9BE36D8">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<taxonomicName LSID="http://zoobank.org/CDA6AFFA-3C76-4F74-8BC4-03B5DF32B011" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium dorsa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dorsa" status="sp. nov.">Acanthinozodium dorsa</taxonomicName> <taxonomicName id="45A84BB2FE00BC4F7B0B48D43E4DC16E" ID-CoL="98YZ8" LSID="http://zoobank.org/CDA6AFFA-3C76-4F74-8BC4-03B5DF32B011" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium dorsa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dorsa" status="sp. nov.">Acanthinozodium dorsa</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="6339900F21E600E6B13EEFB8A6B5E7C6" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">Figs 14I</figureCitation> <figureCitation id="EE4517A49F6653B17A83B5E5E4D3A2CF" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">Figs 14I</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">, 18C, D</figureCitation> <figureCitation id="69CF368F83A661F8C20461645FA6BB7C" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">, 18C, D</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">, 21A-C</figureCitation> <figureCitation id="1F70D1BF7BF69588F0FBF8A1E47A7F8B" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">, 21A-C</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">, 24C, D</figureCitation> <figureCitation id="51D4BE3EA6250244595B9B22E871A91A" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">, 24C, D</figureCitation>
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">, 30F-I</figureCitation> <figureCitation id="60906CC1085FF33520132EBBB8EB31C4" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">, 30F-I</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation> <figureCitation id="E7D8B54444D77D475B2C1E286D3B4FA0" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="B65CC97E4EFBCAB56797A8E7B1FE3D4B" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="C0B4B900174070655BD1AC5769D5B054" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="EF259E45F9D744B645A1FE722A4F2BBC" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="BFAF80273BC4DD8192A76AC4CF49AE61" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ and ♂ and
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">paratypes</emphasis> <emphasis id="AD35EBEB3729962DD275F789D54FFD64" bold="true" italics="true" pageId="0" pageNumber="145">paratypes</emphasis>
26♂16♀ (MMUE), Iran: 26♂16♀ (MMUE), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Fars Province</emphasis> <emphasis id="2B21E91E1DD32089A4A1CBD25CB9FFD1" italics="true" pageId="0" pageNumber="145">Fars Province</emphasis>
: 50 km NE of Shiraz, Bamoo reserve area, : 50 km NE of Shiraz, Bamoo reserve area,
<geoCoordinate degrees="29" direction="north" minutes="45" orientation="latitude" precision="925" value="29.75">29°45'N</geoCoordinate> <geoCoordinate id="010498D29753719949516BDF7BA49DA4" degrees="29" direction="north" minutes="45" orientation="latitude" precision="925" value="29.75">29°45'N</geoCoordinate>
, ,
<geoCoordinate degrees="52" direction="east" minutes="45" orientation="longitude" precision="925" value="52.75">52°45'E</geoCoordinate> <geoCoordinate id="B84D0A73461FBCC756CA0A79F8041F83" degrees="52" direction="east" minutes="45" orientation="longitude" precision="925" value="52.75">52°45'E</geoCoordinate>
, 18-28.05.2000 (Y.M. Marusik). , 18-28.05.2000 (Y.M. Marusik).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="93A12A63BBACC6BCE43F833207DA5648" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="198AD5B68DD5BBE6E7269AF40E8F49FE" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="8FA70457CFC8C21C75F3E13C6DD420C2" pageId="0" pageNumber="145">
The specific epithet is a Persian feminine name meaning The specific epithet is a Persian feminine name meaning
<normalizedToken originalValue="“precious”">&quot;precious&quot;</normalizedToken> <normalizedToken id="B113536B9ECA4B9803F70DEF51EA808C" originalValue="“precious”">&quot;precious&quot;</normalizedToken>
. Noun in apposition. . Noun in apposition.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="688294B0C30373D8B690ED812982BAA2" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="71F0F4CAB0B9B2EFF9A2AE5F0DE932AD" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="5E5F1EAE90B3855DCC8E03AAE208A459" pageId="0" pageNumber="145">
The male of the new species differs from all congeners in the region by the prolateral outgrowth of the median apophysis projecting ventrally ( The male of the new species differs from all congeners in the region by the prolateral outgrowth of the median apophysis projecting ventrally (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="A3A29DF4EE93D5D792FFC509E7B0BAE6" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
not projecting) and the long and broad RTA, apically twisted and lacking outgrowths ( not projecting) and the long and broad RTA, apically twisted and lacking outgrowths (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="1D81A94787466F357FBCF8F3B5E98FF2" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
with 1-2 outgrowths). The female is most similar to with 1-2 outgrowths). The female is most similar to
<taxonomicName lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum"> <taxonomicName id="56805E07ACD1D8E3250C693B515A6A9D" lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis> <emphasis id="AF7B3EB1C7AEA7E225E6E808DD502094" italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. but differs by the epigynal hood being longer than wide ( sp. nov. but differs by the epigynal hood being longer than wide (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="69BB360C13A436724585775D4C3021EE" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
wider than long) and less separated receptacles (ca. 2.5 times of their diameter wider than long) and less separated receptacles (ca. 2.5 times of their diameter
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="27313F3CF1D961F0E0C766E00563D3BA" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
4). 4).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="AA5B9DE56EEB949AD39B3279198221BD" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="10679ACBD87F23C3F3C08E244250732A" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="A6008B5FB2B3C3C1517ED51AC4DF8343" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis> <emphasis id="58C35D8E24CC77CB7490855D20B958C5" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig. (holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18C</figureCitation> <figureCitation id="F3696593E83F2644B1431627985F396A" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18C</figureCitation>
. Total length 3.85. Carapace 1.87 long, 0.89 wide at pars cephalica, 1.39 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.15, ALE: 0.12, PME: 0.09, PLE: 0.09, PME-PME: 0.20. Carapace yellowish brown, darker at pars cephalica. Sternum, labium, maxillae and chelicerae light brown. Legs yellowish, without annulations. Abdomen dark brown dorsally, pale ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 8.20 (2.02, 0.68, 1.90, 2.19, 1.41), II: 6.89 (1.68, 0.68, 1.49, 1.99, 1.05), III: 7.23 (1.88, 0.64, 1.45, 2.13, 1.13), IV: 8.71 (2.18, 0.62, 2.11, 2.70, 1.10). . Total length 3.85. Carapace 1.87 long, 0.89 wide at pars cephalica, 1.39 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.15, ALE: 0.12, PME: 0.09, PLE: 0.09, PME-PME: 0.20. Carapace yellowish brown, darker at pars cephalica. Sternum, labium, maxillae and chelicerae light brown. Legs yellowish, without annulations. Abdomen dark brown dorsally, pale ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 8.20 (2.02, 0.68, 1.90, 2.19, 1.41), II: 6.89 (1.68, 0.68, 1.49, 1.99, 1.05), III: 7.23 (1.88, 0.64, 1.45, 2.13, 1.13), IV: 8.71 (2.18, 0.62, 2.11, 2.70, 1.10).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="01F0BB98BAE0A36DCCA0600C968749B4" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14I</figureCitation> <figureCitation id="E82EBAF9E44382B567B915D936D2AB55" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14I</figureCitation>
, ,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">21A-C</figureCitation> <figureCitation id="CA91C17795970543D37ADD8DA3D516DB" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">21A-C</figureCitation>
, ,
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">24C, D</figureCitation> <figureCitation id="660120BDCA880A60FF8F70F98F9D1F80" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">24C, D</figureCitation>
. RTA long and broad, almost as long as the bulb, twisting ventrally toward the apex; ventral apophysis small and finger shaped; tegulum with posterior process; median apophysis ( . RTA long and broad, almost as long as the bulb, twisting ventrally toward the apex; ventral apophysis small and finger shaped; tegulum with posterior process; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="D92350EBD8EDFCD63040CDD5C72F8A48" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) with a distinct prolateral projection, winding ventrally to a blunt tip; embolus ( ) with a distinct prolateral projection, winding ventrally to a blunt tip; embolus (
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis> <emphasis id="AE75D71D0C4F567D0857E566C48A63BD" italics="true" pageId="0" pageNumber="145">Em</emphasis>
) originating at about the 8:30 ) originating at about the 8:30
<normalizedToken originalValue="o’clock">o'clock</normalizedToken> <normalizedToken id="3C5841B01DABF95D9EF09DE4C53B39CE" originalValue="o’clock">o'clock</normalizedToken>
position; embolus proper thin and slightly curving near the base and apex. position; embolus proper thin and slightly curving near the base and apex.
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="2BF9FF218194797C3D716BFB2E39CA03" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="0A6D7BC6F65A4CFFFE5160DE668AB314" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig. Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18D</figureCitation> <figureCitation id="3B4B3D7C66BE4161AC06A783A7D25146" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18D</figureCitation>
. Total length 4.46. Carapace 1.93 long, 0.96 wide at pars cephalica, 1.46 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.15, ALE: 0.12, PME: 0.09, PLE: 0.10, PME-PME: 0.15. Coloration as in male. Measurements of legs: I: 6.58 (1.39, 0.56, 1.52, 1.88, 1.23), II: 6.49 (1.62, 0.69, 1.34, 1.77, 1.07), III: 6.24 (1.71, 0.63, 1.25, 1.85, 0.80), IV: 8.45 (2.24, 0.72, 1.90, 2.60, 0.99). . Total length 4.46. Carapace 1.93 long, 0.96 wide at pars cephalica, 1.46 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.15, ALE: 0.12, PME: 0.09, PLE: 0.10, PME-PME: 0.15. Coloration as in male. Measurements of legs: I: 6.58 (1.39, 0.56, 1.52, 1.88, 1.23), II: 6.49 (1.62, 0.69, 1.34, 1.77, 1.07), III: 6.24 (1.71, 0.63, 1.25, 1.85, 0.80), IV: 8.45 (2.24, 0.72, 1.90, 2.60, 0.99).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="520A533923B51E0F0813930CAE781A8F" pageId="0" pageNumber="145">
Epigyne as in Fig. Epigyne as in Fig.
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">30F-I</figureCitation> <figureCitation id="7A5675D2831C41AA62DC956D9CA3E112" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">30F-I</figureCitation>
. Epigynal plate over 3 times wider than long; anterior hood ca. 2 times longer than wide; receptacles subrectangular, separated by about 2.5 times of their widths. . Epigynal plate over 3 times wider than long; anterior hood ca. 2 times longer than wide; receptacles subrectangular, separated by about 2.5 times of their widths.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="A23DF2336A2DCB1FA7015BD496BBDDE4" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="A491033BBA0C78C4577C82069327092D" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="CCE45017157CE8AEB79AAEECFAEE9D00" pageId="0" pageNumber="145">
Known from the type locality in Fars Province, southern Iran (Fig. Known from the type locality in Fars Province, southern Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation> <figureCitation id="03F8305F8E084BD35D2E5C7D042B2A88" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

324
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<paragraph pageId="0" pageNumber="145"> <paragraph id="3BE4B5963B860E17C0F32D8894B9C24E" pageId="0" pageNumber="145">
Genus Genus
<taxonomicName LSID="A4551295-2A34-50C5-8AD2-BFB995F9BB5C" authority="Andreeva &amp; Tyshchenko, 1968" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Zodariellum Andreeva &amp; Tyshchenko, 1968</taxonomicName> <taxonomicName id="29789D14DF663A196519B6669CDE60F8" ID-CoL="63WNV" LSID="A4551295-2A34-50C5-8AD2-BFB995F9BB5C" authority="Andreeva &amp; Tyshchenko, 1968" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Zodariellum Andreeva &amp; Tyshchenko, 1968</taxonomicName>
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<paragraph pageId="0" pageNumber="145">Type species.</paragraph> <paragraph id="4DF85C2DEA0BBDA5663C0D6C0C23E2EF" pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="4EB5F9F06F71E5B0AAA25C304C14D269" pageId="0" pageNumber="145">
<taxonomicName authorityName="Andreeva &amp; Tyschchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum surprisum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="surprisum"> <taxonomicName id="2D01E732987195C6C2B821F5B7900BF9" authorityName="Andreeva &amp; Tyschchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum surprisum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="surprisum">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum surprisum</emphasis> <emphasis id="5AEF65A2BE75420532AA365266C3B8FD" italics="true" pageId="0" pageNumber="145">Zodariellum surprisum</emphasis>
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Andreeva &amp; Tyshchenko, 1968 from Tajikistan. Andreeva &amp; Tyshchenko, 1968 from Tajikistan.
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<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="358A3696D3CD96960E049733B7899CDA" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="F777C31DC6AE542BE531205C6217F301" pageId="0" pageNumber="145">
The genus was described as monotypic and synonymized with The genus was described as monotypic and synonymized with
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<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis> <emphasis id="B9D022FC689466850B2657C47097ABAB" italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis>
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by by
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(1991) (1991)
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. .
<bibRefCitation author="Marusik, YM" journalOrPublisher="Reichenbachia" pageId="0" pageNumber="145" pagination="39 - 48" refId="B13" refString="Marusik, YM, Koponen, S, 2001. Spiders of the family Zodariidae from Mongolia (Arachnida: Araneae). Reichenbachia 34: 39 - 48" title="Spiders of the family Zodariidae from Mongolia (Arachnida: Araneae)." volume="34" year="2001">Marusik and Koponen (2001)</bibRefCitation> <bibRefCitation id="99D35395B1776453C0C41A6F70D5DC94" author="Marusik, YM" journalOrPublisher="Reichenbachia" pageId="0" pageNumber="145" pagination="39 - 48" refId="B13" refString="Marusik, YM, Koponen, S, 2001. Spiders of the family Zodariidae from Mongolia (Arachnida: Araneae). Reichenbachia 34: 39 - 48" title="Spiders of the family Zodariidae from Mongolia (Arachnida: Araneae)." volume="34" year="2001">Marusik and Koponen (2001)</bibRefCitation>
resurrected resurrected
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="8EB8AF6CDACC550D323A835948E2A150" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis> <emphasis id="FA01546FAE862F59655C428D0EED274E" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName> </taxonomicName>
, described two new species from Mongolia ( , described two new species from Mongolia (
<taxonomicName lsidName="Z. schmidti" pageId="0" pageNumber="145" rank="species" species="schmidti"> <taxonomicName id="B7252A9989CB15C6A3D7BAC7378BE11E" lsidName="Z. schmidti" pageId="0" pageNumber="145" rank="species" species="schmidti">
<emphasis italics="true" pageId="0" pageNumber="145">Z. schmidti</emphasis> <emphasis id="D5C95A41833040D3690A3A71990F1547" italics="true" pageId="0" pageNumber="145">Z. schmidti</emphasis>
</taxonomicName> </taxonomicName>
Marusik &amp; Koponen, 2001 and Marusik &amp; Koponen, 2001 and
<taxonomicName lsidName="Z. mongolicum" pageId="0" pageNumber="145" rank="species" species="mongolicum"> <taxonomicName id="1DD9B133D7BF233C6E3F567F2B38B4D2" lsidName="Z. mongolicum" pageId="0" pageNumber="145" rank="species" species="mongolicum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. mongolicum</emphasis> <emphasis id="E03BE334BD04C160CCF94D007BAD7E75" italics="true" pageId="0" pageNumber="145">Z. mongolicum</emphasis>
</taxonomicName> </taxonomicName>
Marusik &amp; Koponen, 2001), and transferred eight species to it from Marusik &amp; Koponen, 2001), and transferred eight species to it from
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="9E19BFC15FAAF20622A43394878F3D27" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis> <emphasis id="5A7BFE50F082DC7743D8E123429D47AE" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName> </taxonomicName>
: :
<taxonomicName lsidName="Z. asiaticum" pageId="0" pageNumber="145" rank="species" species="asiaticum"> <taxonomicName id="BFA487994611047AD3CFBD40127E9C3E" lsidName="Z. asiaticum" pageId="0" pageNumber="145" rank="species" species="asiaticum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. asiaticum</emphasis> <emphasis id="A4D6DCE7BF88E8DB3B87E2A11086843C" italics="true" pageId="0" pageNumber="145">Z. asiaticum</emphasis>
</taxonomicName> </taxonomicName>
(Tyshchenko, 1970), (Tyshchenko, 1970),
<taxonomicName lsidName="Z. bekuzini" pageId="0" pageNumber="145" rank="species" species="bekuzini"> <taxonomicName id="148FB0A36E75FB6FA77F4DB04AC9B111" lsidName="Z. bekuzini" pageId="0" pageNumber="145" rank="species" species="bekuzini">
<emphasis italics="true" pageId="0" pageNumber="145">Z. bekuzini</emphasis> <emphasis id="4A347225E723E531CBBBB0D70BCCEE2C" italics="true" pageId="0" pageNumber="145">Z. bekuzini</emphasis>
</taxonomicName> </taxonomicName>
(Nenilin, 1985), (Nenilin, 1985),
<taxonomicName lsidName="Z. chaoyangense" pageId="0" pageNumber="145" rank="species" species="chaoyangense"> <taxonomicName id="1CD22B07215BAFC5589DC45022FA6B98" lsidName="Z. chaoyangense" pageId="0" pageNumber="145" rank="species" species="chaoyangense">
<emphasis italics="true" pageId="0" pageNumber="145">Z. chaoyangense</emphasis> <emphasis id="18F202D96D4AA0BD2384B26535564DEA" italics="true" pageId="0" pageNumber="145">Z. chaoyangense</emphasis>
</taxonomicName> </taxonomicName>
(Zhu &amp; Zhu, 1983), (Zhu &amp; Zhu, 1983),
<taxonomicName lsidName="Z. continentalis" pageId="0" pageNumber="145" rank="species" species="continentalis"> <taxonomicName id="94B973D5A8E3A2148F8C691524BC21A6" lsidName="Z. continentalis" pageId="0" pageNumber="145" rank="species" species="continentalis">
<emphasis italics="true" pageId="0" pageNumber="145">Z. continentalis</emphasis> <emphasis id="10E7B307F6A626B6806DCF2CF4A98506" italics="true" pageId="0" pageNumber="145">Z. continentalis</emphasis>
</taxonomicName> </taxonomicName>
(Andreeva &amp; Tyshchenko, 1968), (Andreeva &amp; Tyshchenko, 1968),
<taxonomicName lsidName="Z. furcum" pageId="0" pageNumber="145" rank="species" species="furcum"> <taxonomicName id="F03CC005FA2B53E177F89B9C0313F253" lsidName="Z. furcum" pageId="0" pageNumber="145" rank="species" species="furcum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. furcum</emphasis> <emphasis id="FA648D639482A7B4872BA6BB211F02C4" italics="true" pageId="0" pageNumber="145">Z. furcum</emphasis>
</taxonomicName> </taxonomicName>
(Zhu, 1988), (Zhu, 1988),
<taxonomicName lsidName="Z. proszynskii" pageId="0" pageNumber="145" rank="species" species="proszynskii"> <taxonomicName id="1EFD09E8DAD63F5F8A6F05BA4E8437E2" lsidName="Z. proszynskii" pageId="0" pageNumber="145" rank="species" species="proszynskii">
<emphasis italics="true" pageId="0" pageNumber="145">Z. proszynskii</emphasis> <emphasis id="CCEC5C872B1483361BF58AA042D5A3C0" italics="true" pageId="0" pageNumber="145">Z. proszynskii</emphasis>
</taxonomicName> </taxonomicName>
(Nenilin &amp; Fet, 1985), (Nenilin &amp; Fet, 1985),
<taxonomicName lsidName="Z. nenilini" pageId="0" pageNumber="145" rank="species" species="nenilini"> <taxonomicName id="D542BF9C9399D82F2AF97F138640929D" lsidName="Z. nenilini" pageId="0" pageNumber="145" rank="species" species="nenilini">
<emphasis italics="true" pageId="0" pageNumber="145">Z. nenilini</emphasis> <emphasis id="FAEC0527C4CC3A236FE409B7908F0A90" italics="true" pageId="0" pageNumber="145">Z. nenilini</emphasis>
</taxonomicName> </taxonomicName>
(Eskov, 1996) and (Eskov, 1996) and
<taxonomicName lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae"> <taxonomicName id="D9EC927FAFC615279CB821AA06500CBE" lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae">
<emphasis italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis> <emphasis id="227BE13D5297CA49C3438C7E2AA4F166" italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis>
</taxonomicName> </taxonomicName>
(Nenilin &amp; Fet, 1985). (Nenilin &amp; Fet, 1985).
<bibRefCitation DOI="https://doi.org/10.23885/1814-3326-2007-3-2-87-95" author="Ponomarev, AV" journalOrPublisher="Caucasian Entomological Bulletin" pageId="0" pageNumber="145" pagination="87 - 95" refId="B22" refString="Ponomarev, AV, 2007. New taxa of spiders (Aranei) from the south of Russia and western Kazakhstan. Caucasian Entomological Bulletin 3: 87 - 95, DOI: https://doi.org/10.23885/1814-3326-2007-3-2-87-95" title="New taxa of spiders (Aranei) from the south of Russia and western Kazakhstan." url="https://doi.org/10.23885/1814-3326-2007-3-2-87-95" volume="3" year="2007">Ponomarev (2007)</bibRefCitation> <bibRefCitation id="A30D9C43592FF2E55CF3826812ECF4B1" DOI="https://doi.org/10.23885/1814-3326-2007-3-2-87-95" author="Ponomarev, AV" journalOrPublisher="Caucasian Entomological Bulletin" pageId="0" pageNumber="145" pagination="87 - 95" refId="B22" refString="Ponomarev, AV, 2007. New taxa of spiders (Aranei) from the south of Russia and western Kazakhstan. Caucasian Entomological Bulletin 3: 87 - 95, DOI: https://doi.org/10.23885/1814-3326-2007-3-2-87-95" title="New taxa of spiders (Aranei) from the south of Russia and western Kazakhstan." url="https://doi.org/10.23885/1814-3326-2007-3-2-87-95" volume="3" year="2007">Ponomarev (2007)</bibRefCitation>
described two more species in the genus ( described two more species in the genus (
<taxonomicName lsidName="Z. volgouralensis" pageId="0" pageNumber="145" rank="species" species="volgouralensis"> <taxonomicName id="23FF01975A96A880C67F06722E936A7C" lsidName="Z. volgouralensis" pageId="0" pageNumber="145" rank="species" species="volgouralensis">
<emphasis italics="true" pageId="0" pageNumber="145">Z. volgouralensis</emphasis> <emphasis id="D973E79C49D0030DCA790F7252BCD74A" italics="true" pageId="0" pageNumber="145">Z. volgouralensis</emphasis>
</taxonomicName> </taxonomicName>
Ponomarev, 2007 and Z. Ponomarev, 2007 and Z.
<taxonomicName baseAuthorityName="Ponomarev" baseAuthorityYear="2007" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion inderensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="inderensis"> <taxonomicName id="4856273C586BDD293AF9D5AC46761545" baseAuthorityName="Ponomarev" baseAuthorityYear="2007" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion inderensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="inderensis">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion inderensis</emphasis> <emphasis id="3DE2692364DBE061645968DE3FB61FF6" italics="true" pageId="0" pageNumber="145">Zodarion inderensis</emphasis>
</taxonomicName> </taxonomicName>
Ponomarev, 2007). Ponomarev, 2007).
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2015.114" author="Jocque, R" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="145" pagination="1 - 23" refId="B10" refString="Jocque, R, Henrard, A, 2015. Revalidation of Acanthinozodium Denis, 1966 with description of three new species and discovery of a remarkable male palpal character (Araneae, Zodariidae). European Journal of Taxonomy 114: 1 - 23, DOI: https://doi.org/10.5852/ejt.2015.114" title="Revalidation of Acanthinozodium Denis, 1966 with description of three new species and discovery of a remarkable male palpal character (Araneae, Zodariidae)." url="https://doi.org/10.5852/ejt.2015.114" volume="114" year="2015"> <bibRefCitation id="8BCC38D8D3454217D253F47976C4800C" DOI="https://doi.org/10.5852/ejt.2015.114" author="Jocque, R" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="145" pagination="1 - 23" refId="B10" refString="Jocque, R, Henrard, A, 2015. Revalidation of Acanthinozodium Denis, 1966 with description of three new species and discovery of a remarkable male palpal character (Araneae, Zodariidae). European Journal of Taxonomy 114: 1 - 23, DOI: https://doi.org/10.5852/ejt.2015.114" title="Revalidation of Acanthinozodium Denis, 1966 with description of three new species and discovery of a remarkable male palpal character (Araneae, Zodariidae)." url="https://doi.org/10.5852/ejt.2015.114" volume="114" year="2015">
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and Henrard (2015) and Henrard (2015)
</bibRefCitation> </bibRefCitation>
transferred all transferred all
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="38B28B0E0AFF716DE673FA8739715CCD" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis> <emphasis id="1ACF2187A2B3021CACAC38631985C685" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName> </taxonomicName>
species, except for the generotype into species, except for the generotype into
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="03F9C89CBCC315FA2189EBB6FED1EC2E" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis> <emphasis id="C44B43832A2DC1607BE5F773D8EA8448" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName> </taxonomicName>
. .
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="E98445EF6AD65C9E71E2FFB7FA1A4CFE" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="4BF635674201B268E4A3BF353DD7CDBB" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="A967295BE666987B1708EA09156747A8" pageId="0" pageNumber="145">
All All
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="D6123F41D1288F3130BE72D32913EDD1" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis> <emphasis id="0289EA87C5CCA253CE652EA9C746F87B" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName> </taxonomicName>
spp. are clearly different from the generotype of spp. are clearly different from the generotype of
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="598D48D856E9404FD3EAE7C8B36271BB" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis> <emphasis id="9EF0171210A9406CCD0225D77BB6C1F7" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName> </taxonomicName>
, ,
<taxonomicName lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum"> <taxonomicName id="3CDF7E30FFA6DAB3058A158383FEF8D7" lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis> <emphasis id="D9875EE444D968D70D050C3745FB0E90" italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
</taxonomicName> </taxonomicName>
(Audouin, 1826), and the species considered in the genus by the following: 1) the cymbium has a tutaculum ( (Audouin, 1826), and the species considered in the genus by the following: 1) the cymbium has a tutaculum (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="FEE1125D6C12685507B4D986384E33B7" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
lacking in lacking in
<taxonomicName lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum"> <taxonomicName id="52977E492B56B7E6AAF01F2B388431A1" lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis> <emphasis id="1B35F7F71949D24FB9C8902C65B504E0" italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
</taxonomicName> </taxonomicName>
and other species groups); 2) filamentous embolus starting at the 5 and other species groups); 2) filamentous embolus starting at the 5
<normalizedToken originalValue="o’clock">o'clock</normalizedToken> <normalizedToken id="7B33C5079961EA1269553798B464DFC3" originalValue="o’clock">o'clock</normalizedToken>
position ( position (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="B1E11F2E6121E15E43D37B7DEBF6F8AC" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
6 - in 6 - in
<taxonomicName lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum"> <taxonomicName id="BEE32C8DEE03891A3E2FEDE115B0F33B" lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis> <emphasis id="D34ABEF5D31277D0DFC84F1127793C3D" italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
</taxonomicName> </taxonomicName>
, and many species groups of , and many species groups of
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="F31CCBB719F3ED504878548C2D80D06E" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis> <emphasis id="36F987A5A7237AE258F904B206ECE474" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName> </taxonomicName>
sensu lato lack the filamentous embolus, or they are not at 5 sensu lato lack the filamentous embolus, or they are not at 5
<normalizedToken originalValue="o’clock">o'clock</normalizedToken> <normalizedToken id="7658EB7BEC4EECDB879CBCC45930C7C5" originalValue="o’clock">o'clock</normalizedToken>
position); 3) the absence of a ventral tibial apophysis ( position); 3) the absence of a ventral tibial apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="0BEF2BCBBF7CFEDF546E4D73F62ACDAE" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
present in the generotype) and 4) the shape of the RTA: one arm longer than wide, with claw or wart-like outgrowth posteriorly from the tip ( present in the generotype) and 4) the shape of the RTA: one arm longer than wide, with claw or wart-like outgrowth posteriorly from the tip (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="8F5471931B60BBF6272C773E2B3F2A9E" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
wider than long, with 3 branches in the generotype and various shapes in other species groups). Females of the two generotypes have long and twisted copulatory ducts, but they are converging in wider than long, with 3 branches in the generotype and various shapes in other species groups). Females of the two generotypes have long and twisted copulatory ducts, but they are converging in
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="2D8EF4CC205E097B4CD5017C12737EA3" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis> <emphasis id="A80792BC5A7F196F69BFDDCE4689D8BE" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName> </taxonomicName>
and diverging in and diverging in
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="0848021D6A6789567121C786B9D18CDA" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis> <emphasis id="FD39C2AB515362704B070FC2A597BEE6" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName> </taxonomicName>
. .
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="composition"> <subSubSection id="6CA0AA08FA938F9D9C0D17F4AADDFA8E" pageId="0" pageNumber="145" type="composition">
<paragraph pageId="0" pageNumber="145">Composition.</paragraph> <paragraph id="72356A76374DF583D445033E0986E6B1" pageId="0" pageNumber="145">Composition.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="8157756323FB47EB1AE1AEB007C01CE9" pageId="0" pageNumber="145">
We consider the following species in We consider the following species in
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="4738A1E7EAC26100E2A30E50B4968B27" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis> <emphasis id="0DC49F6D5673D39C30268DC5B3BE445A" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName> </taxonomicName>
because they have similar male palps and epigynes: because they have similar male palps and epigynes:
<taxonomicName lsidName="Z. asiaticum" pageId="0" pageNumber="145" rank="species" species="asiaticum"> <taxonomicName id="810A736A78654DC19637A42337643037" lsidName="Z. asiaticum" pageId="0" pageNumber="145" rank="species" species="asiaticum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. asiaticum</emphasis> <emphasis id="5F398D4CA461719A6DBD2AB9A604512B" italics="true" pageId="0" pageNumber="145">Z. asiaticum</emphasis>
</taxonomicName> </taxonomicName>
(Tyshchenko, 1970) comb. res., (Tyshchenko, 1970) comb. res.,
<taxonomicName lsidName="Z. bactrianum" pageId="0" pageNumber="145" rank="species" species="bactrianum"> <taxonomicName id="D50312D270F2EA31F2103B754F8608C9" lsidName="Z. bactrianum" pageId="0" pageNumber="145" rank="species" species="bactrianum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. bactrianum</emphasis> <emphasis id="CD389B00DBDD1FD8CD4D79D00A826B9B" italics="true" pageId="0" pageNumber="145">Z. bactrianum</emphasis>
</taxonomicName> </taxonomicName>
(Kroneberg, 1875) comb. nov. (ex. (Kroneberg, 1875) comb. nov. (ex.
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="14630A44662DBC032B551A465E807409" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis> <emphasis id="BB9E8C430B203DA7415ABE38733B8293" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName> </taxonomicName>
), ),
<taxonomicName lsidName="Z. bekuzini" pageId="0" pageNumber="145" rank="species" species="bekuzini"> <taxonomicName id="2A71D8A8AD8FC51672374C9BEB38D46C" lsidName="Z. bekuzini" pageId="0" pageNumber="145" rank="species" species="bekuzini">
<emphasis italics="true" pageId="0" pageNumber="145">Z. bekuzini</emphasis> <emphasis id="EA1D7D775A47530EEB382CD0D6BD7CB0" italics="true" pageId="0" pageNumber="145">Z. bekuzini</emphasis>
</taxonomicName> </taxonomicName>
(Nenilin, 1985) comb. res., (Nenilin, 1985) comb. res.,
<taxonomicName lsidName="Z. chaoyangense" pageId="0" pageNumber="145" rank="species" species="chaoyangense"> <taxonomicName id="5929EFC0508BE5C2DA1E8B590C739422" lsidName="Z. chaoyangense" pageId="0" pageNumber="145" rank="species" species="chaoyangense">
<emphasis italics="true" pageId="0" pageNumber="145">Z. chaoyangense</emphasis> <emphasis id="0E92F91D083B7C0F424EA31397A0842F" italics="true" pageId="0" pageNumber="145">Z. chaoyangense</emphasis>
</taxonomicName> </taxonomicName>
(Zhu &amp; Zhu, 1983) comb. res., (Zhu &amp; Zhu, 1983) comb. res.,
<taxonomicName lsidName="Z. continentalis" pageId="0" pageNumber="145" rank="species" species="continentalis"> <taxonomicName id="5B70A84ED7C1945BAB89E2DDFEC3AEF2" lsidName="Z. continentalis" pageId="0" pageNumber="145" rank="species" species="continentalis">
<emphasis italics="true" pageId="0" pageNumber="145">Z. continentalis</emphasis> <emphasis id="4A2D65E27825BD0EBD8FBF0CEDADF87C" italics="true" pageId="0" pageNumber="145">Z. continentalis</emphasis>
</taxonomicName> </taxonomicName>
(Andreeva &amp; Tyshchenko, 1968) comb. res., (Andreeva &amp; Tyshchenko, 1968) comb. res.,
<taxonomicName lsidName="Z. furcum" pageId="0" pageNumber="145" rank="species" species="furcum"> <taxonomicName id="92D4307BAE6F0324EC8A3EEF33C40B00" lsidName="Z. furcum" pageId="0" pageNumber="145" rank="species" species="furcum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. furcum</emphasis> <emphasis id="1083D531DABACB112123374D200C7E33" italics="true" pageId="0" pageNumber="145">Z. furcum</emphasis>
</taxonomicName> </taxonomicName>
(Zhu, 1988) comb. res., (Zhu, 1988) comb. res.,
<taxonomicName lsidName="Z. mongolicum" pageId="0" pageNumber="145" rank="species" species="mongolicum"> <taxonomicName id="8D88FF0365AF83744728A1025FC3BEF8" lsidName="Z. mongolicum" pageId="0" pageNumber="145" rank="species" species="mongolicum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. mongolicum</emphasis> <emphasis id="65A78DA8F9C198F82D1542EB39A1FE39" italics="true" pageId="0" pageNumber="145">Z. mongolicum</emphasis>
</taxonomicName> </taxonomicName>
Marusik &amp; Koponen, 2001 comb. res., Marusik &amp; Koponen, 2001 comb. res.,
<taxonomicName lsidName="Z. proszynskii" pageId="0" pageNumber="145" rank="species" species="proszynskii"> <taxonomicName id="CA1B456B3FC1503B0EB7438E2FA4C2B0" lsidName="Z. proszynskii" pageId="0" pageNumber="145" rank="species" species="proszynskii">
<emphasis italics="true" pageId="0" pageNumber="145">Z. proszynskii</emphasis> <emphasis id="0CA171070C1C9BC2F96E2C8ED3B7AC1C" italics="true" pageId="0" pageNumber="145">Z. proszynskii</emphasis>
</taxonomicName> </taxonomicName>
(Nenilin &amp; Fet, 1985) comb. res., (Nenilin &amp; Fet, 1985) comb. res.,
<taxonomicName lsidName="Z. nenilini" pageId="0" pageNumber="145" rank="species" species="nenilini"> <taxonomicName id="E03163312858C183FA32CE583BD098C9" lsidName="Z. nenilini" pageId="0" pageNumber="145" rank="species" species="nenilini">
<emphasis italics="true" pageId="0" pageNumber="145">Z. nenilini</emphasis> <emphasis id="7AEA432356D257D06752D6817DA9E574" italics="true" pageId="0" pageNumber="145">Z. nenilini</emphasis>
</taxonomicName> </taxonomicName>
(Eskov, 1996) comb. res., (Eskov, 1996) comb. res.,
<taxonomicName lsidName="Z. surprisum" pageId="0" pageNumber="145" rank="species" species="surprisum"> <taxonomicName id="8E40DA0199DD68EE13981BCCB81F7150" lsidName="Z. surprisum" pageId="0" pageNumber="145" rank="species" species="surprisum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. surprisum</emphasis> <emphasis id="10E251D91B1CB7252717AFCA41916A48" italics="true" pageId="0" pageNumber="145">Z. surprisum</emphasis>
</taxonomicName> </taxonomicName>
Andreeva &amp; Tyshchenko, 1968 comb. res., Andreeva &amp; Tyshchenko, 1968 comb. res.,
<taxonomicName lsidName="Z. schmidti" pageId="0" pageNumber="145" rank="species" species="schmidti"> <taxonomicName id="2C4842E1EFC397405C3F0F4A60C04565" lsidName="Z. schmidti" pageId="0" pageNumber="145" rank="species" species="schmidti">
<emphasis italics="true" pageId="0" pageNumber="145">Z. schmidti</emphasis> <emphasis id="174FFDFE516DA8A9D3619A36C5EA6E6D" italics="true" pageId="0" pageNumber="145">Z. schmidti</emphasis>
</taxonomicName> </taxonomicName>
Marusik &amp; Koponen, 2001 comb. res., Marusik &amp; Koponen, 2001 comb. res.,
<taxonomicName lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae"> <taxonomicName id="038DC46BA5C7CAD9A638FC7D24226BD7" lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae">
<emphasis italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis> <emphasis id="01D5A555B179886BA13A416F8C4003E9" italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis>
</taxonomicName> </taxonomicName>
(Nenilin &amp; Fet, 1985) comb. res. and (Nenilin &amp; Fet, 1985) comb. res. and
<taxonomicName lsidName="Z. volgouralensis" pageId="0" pageNumber="145" rank="species" species="volgouralensis"> <taxonomicName id="0B5189F80440873C22A01FA609E226CA" lsidName="Z. volgouralensis" pageId="0" pageNumber="145" rank="species" species="volgouralensis">
<emphasis italics="true" pageId="0" pageNumber="145">Z. volgouralensis</emphasis> <emphasis id="76706D7C8746D0E4252E630CCCD95E56" italics="true" pageId="0" pageNumber="145">Z. volgouralensis</emphasis>
</taxonomicName> </taxonomicName>
Ponomarev, 2007 comb. res. Taking into account the shape of the epigyne of Ponomarev, 2007 comb. res. Taking into account the shape of the epigyne of
<taxonomicName baseAuthorityName="Ponomarev" baseAuthorityYear="2007" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion inderensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="inderensis"> <taxonomicName id="8A8EBBD4B968158E24FDA6F6D6B2AA57" baseAuthorityName="Ponomarev" baseAuthorityYear="2007" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion inderensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="inderensis">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion inderensis</emphasis> <emphasis id="99EC6FE0D52F9535AD6387A36F5E6F41" italics="true" pageId="0" pageNumber="145">Zodarion inderensis</emphasis>
</taxonomicName> </taxonomicName>
(Ponomarev, 2007), originally placed in (Ponomarev, 2007), originally placed in
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="1EB27874DFA4AE1B9B9DCCDAED421C2F" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis> <emphasis id="2162C6B142F1446FB0B6ACDDA0835B92" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName> </taxonomicName>
, we do not restore the original combination for this species. , we do not restore the original combination for this species.
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="7AB0087D4DB1506A4E51FBBE376BA172" pageId="0" pageNumber="145">
To illustrate the conformation of the male palp in To illustrate the conformation of the male palp in
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="A7554EA2F0D28AD5BCE6834EEC2F0C3B" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis> <emphasis id="A8B2FAD159B835D0D2D1608484B549E6" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName> </taxonomicName>
(male specimens are lacking among the material studied in Iran and Turkmenistan), we have provided figures of (male specimens are lacking among the material studied in Iran and Turkmenistan), we have provided figures of
<taxonomicName lsidName="Z. bactrianum" pageId="0" pageNumber="145" rank="species" species="bactrianum"> <taxonomicName id="CDF952C88D227E09FD47CC9A87C309F8" lsidName="Z. bactrianum" pageId="0" pageNumber="145" rank="species" species="bactrianum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. bactrianum</emphasis> <emphasis id="914B272215198AC030B1F9FE7EC9DF66" italics="true" pageId="0" pageNumber="145">Z. bactrianum</emphasis>
</taxonomicName> </taxonomicName>
, a species previously known from the original description only and previously considered in , a species previously known from the original description only and previously considered in
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="921082B4914F8D7A0DB13467C98514FF" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis> <emphasis id="D993536F652A83AD22B3AC11A3D122C5" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName> </taxonomicName>
. .
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="2C0B11E5D6F75E74389E6D78ED321014" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="55AFB533211D364DE843AEF53515D1DB" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="7E043D3E7DEA177C98C2628E278B6712" pageId="0" pageNumber="145">
Western Russia, Iran, Central Asia to northern China ( Western Russia, Iran, Central Asia to northern China (
<bibRefCitation author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC 2021</bibRefCitation> <bibRefCitation id="FB17B5AFE1A217077EF3291DB010CB75" author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC 2021</bibRefCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

178
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@ -1,144 +1,144 @@
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title> <mods:title id="1404225D79280986BA49840E2CFB995A">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:namePart>Zamani, Alireza</mods:namePart> <mods:namePart id="C8C2155A382D4D9250C012F9B26F870A">Zamani, Alireza</mods:namePart>
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<mods:title>ZooKeys</mods:title> <mods:title id="BB3D7DAA5F093E3BA665A632DD50354D">ZooKeys</mods:title>
</mods:titleInfo> </mods:titleInfo>
<mods:part> <mods:part id="9CABFFEFCBD6535C62A64412A3854762">
<mods:date>2021</mods:date> <mods:date id="1D8602529529D00B38376036CFC8694E">2021</mods:date>
<mods:detail type="pubDate"> <mods:detail id="02F6087001770DCD58D0CA8B2E56873F" type="pubDate">
<mods:number>2021-04-27</mods:number> <mods:number id="D51AC3701325FACD72231025C33A3E3E">2021-04-27</mods:number>
</mods:detail> </mods:detail>
<mods:detail type="volume"> <mods:detail id="366209A25B234F33E052E3567D98E073" type="volume">
<mods:number>1035</mods:number> <mods:number id="1FD689B4D6BC2162C79F9B8A3D049271">1035</mods:number>
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<mods:start>145</mods:start> <mods:start id="596DF0DE78C27E15B660955AFF11A447">145</mods:start>
<mods:end>193</mods:end> <mods:end id="C01D5E045EA6D68A08A12D4FAECC788E">193</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1035.65767</mods:url> <mods:url id="39F603CF4E456487156C2DA773B6D437">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:url>
</mods:location> </mods:location>
<mods:classification>journal article</mods:classification> <mods:classification id="DBA1E2B3B2FFBFDB54476B8C807C6089">journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:identifier> <mods:identifier id="40DE8EC713D1E359928082FC3600BCE5" type="DOI">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1035-145</mods:identifier> <mods:identifier id="92C745B6A94757D3E482F23CE994281C" type="Pensoft-Pub">1313-2970-1035-145</mods:identifier>
<mods:identifier type="ZooBank">ADD73A162DB34873890B1E490C2C8841</mods:identifier> <mods:identifier id="86529C7E756AD15D4612E7D0DC16B6D2" type="ZooBank">ADD73A162DB34873890B1E490C2C8841</mods:identifier>
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</mods:mods> </mods:mods>
<treatment ID-GBIF-Taxon="181753559" LSID="urn:lsid:zoobank.org:act:CD502D82-8DBC-4E66-9626-D9CD571D1DAC" httpUri="http://treatment.plazi.org/id/A5D289752A415787BF60F44464C2C3A0" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="A5D289752A415787BF60F44464C2C3A0" ID-GBIF-Taxon="181753559" LSID="urn:lsid:zoobank.org:act:CD502D82-8DBC-4E66-9626-D9CD571D1DAC" httpUri="http://treatment.plazi.org/id/A5D289752A415787BF60F44464C2C3A0" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="2D22F1CE9081C3A9194934530461B2DC" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="BDE896B7FC898034D6E4BBCAB040CAD7" pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/CD502D82-8DBC-4E66-9626-D9CD571D1DAC" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium sorani" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="sorani" status="sp. nov.">Acanthinozodium sorani</taxonomicName> <taxonomicName id="A9F49AE7D137CDDDDA0AF17AAB6BE718" ID-CoL="98YZ9" LSID="http://zoobank.org/CD502D82-8DBC-4E66-9626-D9CD571D1DAC" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium sorani" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="sorani" status="sp. nov.">Acanthinozodium sorani</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="12DFAB6D8E980852F4F20F5632A5C00F" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11H</figureCitation> <figureCitation id="5FC245150B6E41A15113CC7D24FD99D6" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11H</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">, 12D-F</figureCitation> <figureCitation id="4629C352535227328416B351A30C8D81" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">, 12D-F</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">, 14D-F</figureCitation> <figureCitation id="298DF6A56B5F4D140BC9192571F4DD77" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">, 14D-F</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation> <figureCitation id="813DFC9CAA79BDE96F8B90CF66E49860" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="C4D34920BA2A34C41743CA0180AB13B6" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="6E997BC10EF880A4431D37563C1F4C77" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="E1E0508A17D91B41979F0FAE919ED1E6" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="0877B8020B6DB32CCB067FAFE4F3B880" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran: ♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Kurdistan Province</emphasis> <emphasis id="435175963B8A39EA52B6D8D0FED16DA0" italics="true" pageId="0" pageNumber="145">Kurdistan Province</emphasis>
: Santeh, : Santeh,
<geoCoordinate degrees="36" direction="north" minutes="11" orientation="latitude" precision="925" value="36.183334">36°11'N</geoCoordinate> <geoCoordinate id="829F44C5FC03ADE7EA1A15141B7CC970" degrees="36" direction="north" minutes="11" orientation="latitude" precision="925" value="36.183334">36°11'N</geoCoordinate>
, ,
<geoCoordinate degrees="46" direction="east" minutes="32" orientation="longitude" precision="925" value="46.533333">46°32'E</geoCoordinate> <geoCoordinate id="9E0DB2F54B14A30B94769D39A37B52E4" degrees="46" direction="east" minutes="32" orientation="longitude" precision="925" value="46.533333">46°32'E</geoCoordinate>
, 23.06.1975 (A. Senglet). , 23.06.1975 (A. Senglet).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis> <emphasis id="9C74D44EA1AA3D1874F9ED91BCC96F3D" bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
: 6♂ (MHNG), Marivan, 5.2017 (A. Zamani); : 6♂ (MHNG), Marivan, 5.2017 (A. Zamani);
<emphasis italics="true" pageId="0" pageNumber="145">East Azerbaijan Province</emphasis> <emphasis id="B9AD9BC4C5902F0D7A6C5C224A691448" italics="true" pageId="0" pageNumber="145">East Azerbaijan Province</emphasis>
: 2♂ (MHNG), north of Bonati, : 2♂ (MHNG), north of Bonati,
<geoCoordinate degrees="37" direction="north" minutes="26" orientation="latitude" precision="925" value="37.433334">37°26'N</geoCoordinate> <geoCoordinate id="DDDBE6C89D829F94353C4A871F5B1665" degrees="37" direction="north" minutes="26" orientation="latitude" precision="925" value="37.433334">37°26'N</geoCoordinate>
, ,
<geoCoordinate degrees="45" direction="east" minutes="57" orientation="longitude" precision="925" value="45.95">45°57'E</geoCoordinate> <geoCoordinate id="AEDFA5D4B24F6085CF237E36B9F6CBD5" degrees="45" direction="east" minutes="57" orientation="longitude" precision="925" value="45.95">45°57'E</geoCoordinate>
, 04.06.1975 (A. Senglet). , 04.06.1975 (A. Senglet).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="87052E0E516946B6834C7842DC0AFADB" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="376C20998795A962A0915D800D538E18" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet refers to a dialect or a language of the Kurdish languages that is spoken in Iraq, mainly in Iraqi Kurdistan, as well as the Kurdistan Province, Kermanshah Province, and West Azerbaijan Province of western Iran. Noun in apposition.</paragraph> <paragraph id="7F94C9F421F6BF28F98D5146B1946F44" pageId="0" pageNumber="145">The specific epithet refers to a dialect or a language of the Kurdish languages that is spoken in Iraq, mainly in Iraqi Kurdistan, as well as the Kurdistan Province, Kermanshah Province, and West Azerbaijan Province of western Iran. Noun in apposition.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="0AA59162C407DC240A27AF45BA08D3B7" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="0E6405A5FEEF6D60085774D290B3E776" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="0000906DCF35EAD893C49C777F9BC868" pageId="0" pageNumber="145">
The new species is most similar to The new species is most similar to
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa"> <taxonomicName id="4E50676DFBA5849D99512D563466AE0A" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis> <emphasis id="346EAAFD709B7D2315C227657B8873D3" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. but differs by embolic anterior process which is spine-like and thinner than the embolus proper in sp. nov. but differs by embolic anterior process which is spine-like and thinner than the embolus proper in
<taxonomicName lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani"> <taxonomicName id="A4622FBC907189BB7CF6FCA7ABE211AA" lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis italics="true" pageId="0" pageNumber="145">A. sorani</emphasis> <emphasis id="A720B855ABDA697AFEAB71670233B17E" italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="9908DA81651034FA8676DB826D64BEFF" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
broad) and also by the relative length of the RTA, longer than bulb in broad) and also by the relative length of the RTA, longer than bulb in
<taxonomicName lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani"> <taxonomicName id="0D75CD8D4E67752713DE09C2C403761F" lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis italics="true" pageId="0" pageNumber="145">A. sorani</emphasis> <emphasis id="D95136398D76A70F0F251F92B8D7923D" italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="79CDC3D47DBB239665CB989DF53D2CD9" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
shorter). shorter).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="58D95CF25C0AF8A9FB760D259632D431" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="F2EF2EF187DCCB8A4FE483297B729D6B" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="F1155469D0366DE40EE0D7DF83E78497" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis> <emphasis id="04F59D364A06BCBF2E82E251AD11A127" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig. (holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11H</figureCitation> <figureCitation id="0AC8439490E0F197BD8FB220F40E1CF1" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11H</figureCitation>
. Total length 2.11. Carapace 1.00 long, 0.46 wide at pars cephalica, 0.72 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.09, ALE: 0.07, PME: 0.06, PLE: 0.06, PME-PME: 0.12. Carapace, sternum, labium, chelicerae and maxillae yellowish brown; carapace with irregular dark patterns. Chelicera with one retromarginal tooth. Legs yellowish, dark brown at femora I and II, without annulations. Abdomen dorsally black, grayish ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 3.87 (0.97, 0.37, 0.89, 0.91, 0.64), II: 3.47 (0.92, 0.32, 0.72, 0.88, 0.63), Fe III: 0.88, other segments missing, IV: 4.51 (1.06, 0.36, 1.12, 1.33, 0.64). . Total length 2.11. Carapace 1.00 long, 0.46 wide at pars cephalica, 0.72 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.09, ALE: 0.07, PME: 0.06, PLE: 0.06, PME-PME: 0.12. Carapace, sternum, labium, chelicerae and maxillae yellowish brown; carapace with irregular dark patterns. Chelicera with one retromarginal tooth. Legs yellowish, dark brown at femora I and II, without annulations. Abdomen dorsally black, grayish ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 3.87 (0.97, 0.37, 0.89, 0.91, 0.64), II: 3.47 (0.92, 0.32, 0.72, 0.88, 0.63), Fe III: 0.88, other segments missing, IV: 4.51 (1.06, 0.36, 1.12, 1.33, 0.64).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="86149AA310DB114BB87081B7BEFE4405" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">12D-F</figureCitation> <figureCitation id="B7101A84EED58561041991442EB1BFF8" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">12D-F</figureCitation>
, ,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14D-F</figureCitation> <figureCitation id="C394C273C3AE3A6C7F75B8090BC20384" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14D-F</figureCitation>
. RTA ( . RTA (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis> <emphasis id="13DDA5A0A6477490D53DD9D27D92E59D" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
) long (longer than bulb) and thin (about 6 times longer than wide), slightly bent; tegulum with posterior process ( ) long (longer than bulb) and thin (about 6 times longer than wide), slightly bent; tegulum with posterior process (
<emphasis italics="true" pageId="0" pageNumber="145">Tp</emphasis> <emphasis id="10959BAFEF44E962594396AE5CC86D62" italics="true" pageId="0" pageNumber="145">Tp</emphasis>
), sperm duct almost straight retrolaterally, and gently bent along prolateral side; median apophysis ( ), sperm duct almost straight retrolaterally, and gently bent along prolateral side; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="E4DD7A13B8E094B2C59B7B2EBE42DA87" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) wider than long; embolus ( ) wider than long; embolus (
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis> <emphasis id="C45BEFBA5C602BF31D34AFC546980825" italics="true" pageId="0" pageNumber="145">Em</emphasis>
) broad basally, bifurcated terminally, with spine-like anterior process; embolus proper thin and straight, with small tubercle. ) broad basally, bifurcated terminally, with spine-like anterior process; embolus proper thin and straight, with small tubercle.
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="7A7318BA15F00A4B52BAA61AF1B361FB" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="B53C23E786ECED8F93E28B5E3CFEC69D" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown. Unknown.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="93D1F09C05E5E2A8426D56F30E3DDB22" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="7D195F0C8AEB4E1FB58F24FD44238736" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="291829E58DE713E0D77F2CAE99108AB8" pageId="0" pageNumber="145">
Known only from the listed localities in Kurdistan and East Azerbaijan provinces, western and northwestern Iran (Fig. Known only from the listed localities in Kurdistan and East Azerbaijan provinces, western and northwestern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation> <figureCitation id="146E84245C0E7F6B94600A5A63A7603D" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

104
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<treatment ID-GBIF-Taxon="181753540" LSID="urn:lsid:plazi:treatment:AA4D61BB05A75F3299A814AB2E95E88E" httpUri="http://treatment.plazi.org/id/AA4D61BB05A75F3299A814AB2E95E88E" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="AA4D61BB05A75F3299A814AB2E95E88E" ID-GBIF-Taxon="181753540" LSID="urn:lsid:plazi:treatment:AA4D61BB05A75F3299A814AB2E95E88E" httpUri="http://treatment.plazi.org/id/AA4D61BB05A75F3299A814AB2E95E88E" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="D75A839AC1DA163BE02638909A66E0BA" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="824A291020B6C94E22F0A75693255A62" pageId="0" pageNumber="145">
Genus Genus
<taxonomicName LSID="AA4D61BB-05A7-5F32-99A8-14AB2E95E88E" authority="Denis, 1966" authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Acanthinozodium Denis, 1966</taxonomicName> <taxonomicName id="DDF89E628F972CE37D34237CF2661BF1" ID-CoL="M2C" ID-ENA="2033106" LSID="AA4D61BB-05A7-5F32-99A8-14AB2E95E88E" authority="Denis, 1966" authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Acanthinozodium Denis, 1966</taxonomicName>
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<subSubSection pageId="0" pageNumber="145" type="type species"> <subSubSection id="1860FCBD08DDD6A8CE4D4EA461B03E37" pageId="0" pageNumber="145" type="type species">
<paragraph pageId="0" pageNumber="145">Type species.</paragraph> <paragraph id="487965BA535E9E84945B9F751828F91C" pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="01786BC77BEE50A7F145592E581FC321" pageId="0" pageNumber="145">
<taxonomicName authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium spinulosum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="spinulosum"> <taxonomicName id="FFDB8D9549E139BEC8BC36E0EF7A7507" authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium spinulosum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="spinulosum">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium spinulosum</emphasis> <emphasis id="2F41EC0EBBC2130285E1B889DEDE0CC7" italics="true" pageId="0" pageNumber="145">Acanthinozodium spinulosum</emphasis>
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Denis, 1966 from Libya. Denis, 1966 from Libya.
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="comments"> <subSubSection id="B3B38E83CD43D52415BAAFA2197E26F2" pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="62A8EA1E95DF6B6464D05AC2A1FB0136" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="53E4AC9082E9C12ED69FCDD209E67F3E" pageId="0" pageNumber="145">
Twelve species are currently considered in this genus which are known from the Maghreb and adjacent countries in the south and in Socotra; previously, it was not recorded from Iran ( Twelve species are currently considered in this genus which are known from the Maghreb and adjacent countries in the south and in Socotra; previously, it was not recorded from Iran (
<bibRefCitation author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC 2021</bibRefCitation> <bibRefCitation id="AC198906881A306CA1279A05972B9438" author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC 2021</bibRefCitation>
). Although the male of the type species is unknown, ). Although the male of the type species is unknown,
<taxonomicName authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="E39293DC763E6357AB317579BBA8DCE6" authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis> <emphasis id="86C2CC832D4E55813D72DC0F30602FFF" italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis>
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comprises species having a unique gland located in the conical pit of the cymbium. The 11 species occurring in Iran and the one from Turkmenistan have the same conical pit and are thus placed in this genus; however, the male palpal tibia and particularly the bulbs are different from those of the species occurring in the Maghreb and surrounding countries. Further revisions are needed to clarify the taxonomy and composition of this group, which are beyond the scope of this paper. comprises species having a unique gland located in the conical pit of the cymbium. The 11 species occurring in Iran and the one from Turkmenistan have the same conical pit and are thus placed in this genus; however, the male palpal tibia and particularly the bulbs are different from those of the species occurring in the Maghreb and surrounding countries. Further revisions are needed to clarify the taxonomy and composition of this group, which are beyond the scope of this paper.
</paragraph> </paragraph>

512
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title> <mods:title id="AEEBA60F6CB7F7FBA49B92F69BA92787">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:namePart>Zamani, Alireza</mods:namePart> <mods:namePart id="053EFADAEF22FE73AD3F6B706B45B4DE">Zamani, Alireza</mods:namePart>
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation> <mods:affiliation id="1AA2C79309D8E25645BA0E63A1DB77CB">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<treatment ID-GBIF-Taxon="181753547" LSID="urn:lsid:plazi:treatment:B038AB8CF4435894B2AD946067C7B0E2" httpUri="http://treatment.plazi.org/id/B038AB8CF4435894B2AD946067C7B0E2" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="B038AB8CF4435894B2AD946067C7B0E2" ID-GBIF-Taxon="181753547" LSID="urn:lsid:plazi:treatment:B038AB8CF4435894B2AD946067C7B0E2" httpUri="http://treatment.plazi.org/id/B038AB8CF4435894B2AD946067C7B0E2" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="065C40270184C30CC5857CB40E25144B" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="0D8E25883E2FC29C4265881C63C1B186" pageId="0" pageNumber="145">
<taxonomicName LSID="B038AB8C-F443-5894-B2AD-946067C7B0E2" authority="(Nenilin &amp; Fet, 1985)" baseAuthorityName="Nenilin &amp; Fet" baseAuthorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum proszynskii" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="proszynskii">Zodariellum proszynskii (Nenilin &amp; Fet, 1985)</taxonomicName> <taxonomicName id="5DEC716A3D5E853E5D22C9B3545200F3" ID-CoL="93S8H" LSID="B038AB8C-F443-5894-B2AD-946067C7B0E2" authority="(Nenilin &amp; Fet, 1985)" baseAuthorityName="Nenilin &amp; Fet" baseAuthorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum proszynskii" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="proszynskii">Zodariellum proszynskii (Nenilin &amp; Fet, 1985)</taxonomicName>
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Epigyne of Zodariellum proszynskii A intact, ventral B, C macerated, ventral and dorsal. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure31" httpUri="https://binary.pensoft.net/fig/536908" pageId="0" pageNumber="145">Figs 31A-C</figureCitation> <figureCitation id="2FA6313CF6CB770F57FE1768587D7C28" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Epigyne of Zodariellum proszynskii A intact, ventral B, C macerated, ventral and dorsal. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure31" httpUri="https://binary.pensoft.net/fig/536908" pageId="0" pageNumber="145">Figs 31A-C</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation> <figureCitation id="FC7CCDCA4C468F0EEC54D39A250E4C80" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="reference_group"> <subSubSection id="F3AF752B3C6F723094ED72254653A813" pageId="0" pageNumber="145" type="reference_group">
<paragraph pageId="0" pageNumber="145"> <paragraph id="258F1103C3D07711F0BECF28A74F1C87" pageId="0" pageNumber="145">
<taxonomicName authorityName="Nenilin &amp; Fet" authorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion proszynskii" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="proszynskii">Zodarion proszynskii</taxonomicName> <taxonomicName id="B61A0B357509825B349E40E3AC98E807" authorityName="Nenilin &amp; Fet" authorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion proszynskii" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="proszynskii">Zodarion proszynskii</taxonomicName>
Nenilin &amp; Fet, 1985: 618, f. 1-4, 9 (♂♀). Nenilin &amp; Fet, 1985: 618, f. 1-4, 9 (♂♀).
</paragraph> </paragraph>
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<paragraph pageId="0" pageNumber="145">Material.</paragraph> <paragraph id="BE211A4BE27DCA6E7789D583A7A536FC" pageId="0" pageNumber="145">Material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="56B070913768170295150B901A047174" pageId="0" pageNumber="145">
<materialsCitation ID-GBIF-Occurrence="3093634303" collectingDate="1974-07-15" collectorName="A. Senglet" country="Iran" latitude="37.2" location="Qouchan" longLatPrecision="1183" longitude="58.483334" specimenCount="1" specimenCount-female="1" stateProvince="Razavi Khorasan Province"> <materialsCitation id="2974A522B42F5A68D0F378CB72FADE45" ID-GBIF-Occurrence="3093634303" collectingDate="1974-07-15" collectorName="A. Senglet" country="Iran" latitude="37.2" location="Qouchan" longLatPrecision="1183" longitude="58.483334" specimenCount="1" specimenCount-female="1" stateProvince="Razavi Khorasan Province">
<collectingCountry name="Iran">Iran</collectingCountry> <collectingCountry id="A5ECDE5C129F411DAFBE1342EDEB43F4" name="Iran">Iran</collectingCountry>
: :
<emphasis italics="true" pageId="0" pageNumber="145"> <emphasis id="3CE245AF767381E565D9F415B155A175" italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Khorasan-e Razavi">Razavi Khorasan Province</collectingRegion> <collectingRegion id="B86B700E3EC868FEC4E215B916F63F09" country="Iran" name="Khorasan-e Razavi">Razavi Khorasan Province</collectingRegion>
</emphasis> </emphasis>
: :
<specimenCount type="female">1♀</specimenCount> <specimenCount id="3A2DA40BCE0BA4A30797B348768CBCEC" type="female">1♀</specimenCount>
(MHNG), (MHNG),
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, ,
<geoCoordinate degrees="37" direction="north" minutes="12" orientation="latitude" precision="925" value="37.2">37°12'N</geoCoordinate> <geoCoordinate id="090360DB7A814F02877F45B35ABC58D2" degrees="37" direction="north" minutes="12" orientation="latitude" precision="925" value="37.2">37°12'N</geoCoordinate>
, ,
<geoCoordinate degrees="58" direction="east" minutes="29" orientation="longitude" precision="925" value="58.483334">58°29'E</geoCoordinate> <geoCoordinate id="D0083CA125A3FCD79BF6853A5529B394" degrees="58" direction="east" minutes="29" orientation="longitude" precision="925" value="58.483334">58°29'E</geoCoordinate>
, ,
<collectingDate value="1974-07-15">15.07.1974</collectingDate> <collectingDate id="E797D3B6DE9B9A4BF095816E05E8C9D1" value="1974-07-15">15.07.1974</collectingDate>
( (
<collectorName>A. Senglet</collectorName> <collectorName id="8A1F9ADA68A6C2FAF8B325729887584F">A. Senglet</collectorName>
) )
</materialsCitation> </materialsCitation>
. .
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="materials_examined"> <subSubSection id="B6DD170F031D68BED0BD934E9474F984" pageId="0" pageNumber="145" type="materials_examined">
<paragraph pageId="0" pageNumber="145">Comparative material.</paragraph> <paragraph id="79B7A7E351F923E08E8B058243CC244B" pageId="0" pageNumber="145">Comparative material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="C2064391A61D2BDF151B33390A57FC51" pageId="0" pageNumber="145">
<taxonomicName class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum bactrianum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bactrianum"> <taxonomicName id="5C2AA32BB0F293F9217DF2BB3FFB2D0C" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum bactrianum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bactrianum">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum bactrianum</emphasis> <emphasis id="B91713F5B958F3F98628F6075CDC5095" italics="true" pageId="0" pageNumber="145">Zodariellum bactrianum</emphasis>
</taxonomicName> </taxonomicName>
(Kroneberg, 1875): (Kroneberg, 1875):
<materialsCitation ID-GBIF-Occurrence="3093634310" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Tajikistan" location="Tajikistan" specimenCount="1" specimenCount-male="1"> <materialsCitation id="FFE28AF4412845EEB3A1A4955A6D706D" ID-GBIF-Occurrence="3093634310" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Tajikistan" location="Tajikistan" specimenCount="1" specimenCount-male="1">
<specimenCount type="male">1♂</specimenCount> <specimenCount id="EEBF734050F03C082A45002FE78ACB4A" type="male">1♂</specimenCount>
from from
<collectingCountry name="Tajikistan">Tajikistan</collectingCountry> <collectingCountry id="42A23B2E20A6E17E428B12F0C1A7D2EB" name="Tajikistan">Tajikistan</collectingCountry>
(Fig. (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. SEM images of the male palps of Zodarion nitidum (A-C) and Z. bactrianum (D-F) A, D ventral B, E retrolateral C dorsal F retrodorsal. Abbreviations: Co - conductor, Eb - embolar base, Em - embolus, Ma - median apophysis, Mc - claw of median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process, Tu - tutaculum, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure28" httpUri="https://binary.pensoft.net/fig/536905" pageId="0" pageNumber="145">28D-F</figureCitation> <figureCitation id="AE7E9CC0F4F6DFF9B65B5C7C3F340801" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. SEM images of the male palps of Zodarion nitidum (A-C) and Z. bactrianum (D-F) A, D ventral B, E retrolateral C dorsal F retrodorsal. Abbreviations: Co - conductor, Eb - embolar base, Em - embolus, Ma - median apophysis, Mc - claw of median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process, Tu - tutaculum, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure28" httpUri="https://binary.pensoft.net/fig/536905" pageId="0" pageNumber="145">28D-F</figureCitation>
) )
</materialsCitation> </materialsCitation>
. .
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure28" httpUri="https://binary.pensoft.net/fig/536905" pageId="0" pageNumber="145" start="Figure 28" startId="F28"> <caption id="A37110105D6D5A517B82A90FCD498AAF" doi="10.3897/zookeys.1035.65767.figure28" httpUri="https://binary.pensoft.net/fig/536905" pageId="0" pageNumber="145" start="Figure 28" startId="F28">
<paragraph pageId="0" pageNumber="145"> <paragraph id="14524D96915F4C8BBBB1FC06793BEE8B" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 28.</emphasis> <emphasis id="BBD4DCECCCBDF1327DD739C59FAE7108" bold="true" pageId="0" pageNumber="145">Figure 28.</emphasis>
SEM images of the male palps of SEM images of the male palps of
<taxonomicName baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion nitidum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitidum"> <taxonomicName id="B2CA97C5376C721B563B2D80CE2E262C" baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion nitidum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitidum">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion nitidum</emphasis> <emphasis id="BE9E0C278465CCA44D1C2963C16EEE7C" italics="true" pageId="0" pageNumber="145">Zodarion nitidum</emphasis>
</taxonomicName> </taxonomicName>
( (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis> <emphasis id="4BC60731C8EAFB87A3B817ABD0582D2E" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and ) and
<taxonomicName lsidName="Z. bactrianum" pageId="0" pageNumber="145" rank="species" species="bactrianum"> <taxonomicName id="4FA2E473D77CC73381EC4A3BB2C521D8" lsidName="Z. bactrianum" pageId="0" pageNumber="145" rank="species" species="bactrianum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. bactrianum</emphasis> <emphasis id="1FEB7BDC330EC2103652F2E205B0ABE7" italics="true" pageId="0" pageNumber="145">Z. bactrianum</emphasis>
</taxonomicName> </taxonomicName>
( (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis> <emphasis id="65ACF3E08D743F140DDD2DA0D669652A" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis> <emphasis id="47C4B640CE418A0EC705E38E6A238174" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, E</emphasis> <emphasis id="9C129D844D3156F9D51C78D8F6F6DD46" bold="true" pageId="0" pageNumber="145">B, E</emphasis>
retrolateral retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis> <emphasis id="9CC76B23E8FC1519DD6C147F4EE4A2BB" bold="true" pageId="0" pageNumber="145">C</emphasis>
dorsal dorsal
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis> <emphasis id="A15E01EB324E81930ECD4E395AB4DB7A" bold="true" pageId="0" pageNumber="145">F</emphasis>
retrodorsal. Abbreviations: retrodorsal. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis> <emphasis id="2DFED0F9FFDE01FFEB7E4C37F781CDC9" italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor, - conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Eb</emphasis> <emphasis id="9B9936BF02ED4965D93151BE0BAB5BCA" italics="true" pageId="0" pageNumber="145">Eb</emphasis>
- embolar base, - embolar base,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis> <emphasis id="F5FFE9D11CE2D9976E2AB579968AD4EF" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus, - embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="5A62B95EC61A713DB3777E8BC190CC10" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis, - median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Mc</emphasis> <emphasis id="360A838B0D367D04BDB7F7B764BEAECF" italics="true" pageId="0" pageNumber="145">Mc</emphasis>
- claw of median apophysis, - claw of median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis> <emphasis id="42241431956C30BE0EEA602E03A20D11" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral tibial apophysis, - retrolateral tibial apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Tp</emphasis> <emphasis id="FEF5BA9081A2036F2D3F81D7033FB9CF" italics="true" pageId="0" pageNumber="145">Tp</emphasis>
- tegular process, - tegular process,
<emphasis italics="true" pageId="0" pageNumber="145">Tu</emphasis> <emphasis id="BDE8C90E696AE78DBBED5D23BC823082" italics="true" pageId="0" pageNumber="145">Tu</emphasis>
- tutaculum, - tutaculum,
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis> <emphasis id="FF24DE124C1BBEEA2C6F4ABA1A44F4E8" italics="true" pageId="0" pageNumber="145">Va</emphasis>
- ventral apophysis. Scale bars: - ventral apophysis. Scale bars:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity> <quantity id="C010CCF98F74A4EDC7C127D3AB33F8C3" metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity>
. .
</paragraph> </paragraph>
</caption> </caption>
<caption doi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145" start="Figure 29" startId="F29"> <caption id="02961ECC6F8D7970A8A2008339324ED8" doi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145" start="Figure 29" startId="F29">
<paragraph pageId="0" pageNumber="145"> <paragraph id="04BBC98FF5973822CFC9EB9DA4EEF4E9" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 29.</emphasis> <emphasis id="3FA4067DE935BB43E057C9912A844061" bold="true" pageId="0" pageNumber="145">Figure 29.</emphasis>
SEM images of the male palps of SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum"> <taxonomicName id="5D2EDF3229BFBD18B2749109B7691B4B" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis> <emphasis id="D206B147F6DB3B102A57DFAB3D356CC2" italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis> <emphasis id="63E12EF4E8C8572866B47C446A934C3C" bold="true" pageId="0" pageNumber="145">A</emphasis>
) and ) and
<taxonomicName lsidName="Z. talyschicum" pageId="0" pageNumber="145" rank="species" species="talyschicum"> <taxonomicName id="F6BF0FEEE8C6837F136E6A80EC9DBECB" lsidName="Z. talyschicum" pageId="0" pageNumber="145" rank="species" species="talyschicum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. talyschicum</emphasis> <emphasis id="32474EC651636895FC9198CF3B23F382" italics="true" pageId="0" pageNumber="145">Z. talyschicum</emphasis>
</taxonomicName> </taxonomicName>
( (
<emphasis bold="true" pageId="0" pageNumber="145">B, C</emphasis> <emphasis id="E508410255FCF0FB099F9D7B1BC6FC49" bold="true" pageId="0" pageNumber="145">B, C</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, B</emphasis> <emphasis id="F3C9CC09EB7BB251F45988F1B7786946" bold="true" pageId="0" pageNumber="145">A, B</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis> <emphasis id="5C4EF35A90AD0F5F1DB58B4E37ABA031" bold="true" pageId="0" pageNumber="145">C</emphasis>
ventroretrolateral. Abbreviations: ventroretrolateral. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Tm</emphasis> <emphasis id="EC6FFFD67ACC912B2D49545F9DFCE3B0" italics="true" pageId="0" pageNumber="145">Tm</emphasis>
- median tegular process, - median tegular process,
<emphasis italics="true" pageId="0" pageNumber="145">Tp</emphasis> <emphasis id="82E68C50C8C7051D26CE0398E10518EB" italics="true" pageId="0" pageNumber="145">Tp</emphasis>
- prolateral tegular process. Scale bars: - prolateral tegular process. Scale bars:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity> <quantity id="FA39E8A330657C075E1B2CF5727277F0" metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity>
. .
</paragraph> </paragraph>
</caption> </caption>
<caption doi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145" start="Figure 30" startId="F30"> <caption id="6FE083E86D8C10CB3FB7A60B6ABC1232" doi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145" start="Figure 30" startId="F30">
<paragraph pageId="0" pageNumber="145"> <paragraph id="3E735EECB3392C52D9C88F1470AAC0D2" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 30.</emphasis> <emphasis id="8C9F0DF34D7B0CF2E841440BDFE4473C" bold="true" pageId="0" pageNumber="145">Figure 30.</emphasis>
Epigynes of Epigynes of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum"> <taxonomicName id="462E650C4654020456E4968B6062B0D2" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis> <emphasis id="9D0074896D8DE04163DBC285DC01236B" italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-E</emphasis> <emphasis id="44592BA1F270071FD8EBFE4851B937A0" bold="true" pageId="0" pageNumber="145">A-E</emphasis>
) and ) and
<taxonomicName lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa"> <taxonomicName id="4743C6BFE99CD197A131ABC0A8EA525C" lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis> <emphasis id="8A265E4EF5717A032DFC8BBB17CAA52E" italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">F-I</emphasis> <emphasis id="67BE33FACF8F754B4C9BE0DCDBB3B0B9" bold="true" pageId="0" pageNumber="145">F-I</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, F</emphasis> <emphasis id="86EA4B3BBDC4BB1909680AA4D3A6316E" bold="true" pageId="0" pageNumber="145">A, F</emphasis>
intact, ventral intact, ventral
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis> <emphasis id="72884519B37A980333C3850FC1D4E28B" bold="true" pageId="0" pageNumber="145">B</emphasis>
macerated, ventral macerated, ventral
<emphasis bold="true" pageId="0" pageNumber="145">C-E, G-I</emphasis> <emphasis id="EAB4BFFE72834745CBBFAD836E0F0788" bold="true" pageId="0" pageNumber="145">C-E, G-I</emphasis>
macerated, dorsal. Scale bars: macerated, dorsal. Scale bars:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity> <quantity id="1B330561076FA86BCA49479ADFEBC0F2" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
. .
</paragraph> </paragraph>
</caption> </caption>
<caption doi="10.3897/zookeys.1035.65767.figure31" httpUri="https://binary.pensoft.net/fig/536908" pageId="0" pageNumber="145" start="Figure 31" startId="F31"> <caption id="F682A2CD384715561E60C8918D38EDC3" doi="10.3897/zookeys.1035.65767.figure31" httpUri="https://binary.pensoft.net/fig/536908" pageId="0" pageNumber="145" start="Figure 31" startId="F31">
<paragraph pageId="0" pageNumber="145"> <paragraph id="AE099FB0E3171C78DCA7320BAC418C90" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 31.</emphasis> <emphasis id="9C89E2EE8988B7F804510DF418666E07" bold="true" pageId="0" pageNumber="145">Figure 31.</emphasis>
Epigyne of Epigyne of
<taxonomicName baseAuthorityName="Nenilin &amp; Fet" baseAuthorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum proszynskii" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="proszynskii"> <taxonomicName id="7643F354CBDE560CEA7A5A5E6991F07E" baseAuthorityName="Nenilin &amp; Fet" baseAuthorityYear="1985" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum proszynskii" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="proszynskii">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum proszynskii</emphasis> <emphasis id="6F43BBFC49C6BC587E761E9FB5A9064F" italics="true" pageId="0" pageNumber="145">Zodariellum proszynskii</emphasis>
</taxonomicName> </taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis> <emphasis id="87A10CB463DDCDC191C8F27CF9C53A36" bold="true" pageId="0" pageNumber="145">A</emphasis>
intact, ventral intact, ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, C</emphasis> <emphasis id="C14CC2040E3585AA1FF879F29567C45B" bold="true" pageId="0" pageNumber="145">B, C</emphasis>
macerated, ventral and dorsal. Scale bar: macerated, ventral and dorsal. Scale bar:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity> <quantity id="89A50D1F649D28E7F8A875FF0F1CAD25" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
. .
</paragraph> </paragraph>
</caption> </caption>
<caption doi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145" start="Figure 32" startId="F32"> <caption id="75FD0E76FAD73E7236DDE4CB33E81D07" doi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145" start="Figure 32" startId="F32">
<paragraph pageId="0" pageNumber="145"> <paragraph id="C6CED1DD8409E023F07064EEF32D7B0F" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 32.</emphasis> <emphasis id="9AB37816EE751D9D5EA1C2A455DDBA45" bold="true" pageId="0" pageNumber="145">Figure 32.</emphasis>
Distribution records of Distribution records of
<taxonomicName authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="71DB44ED5814BFAE86548468729CC05D" authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis> <emphasis id="5A22EA0BBEF0145F14B06D0131D5AA74" italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis>
</taxonomicName> </taxonomicName>
spp. (partim spp. (partim
<emphasis bold="true" pageId="0" pageNumber="145">1-4, 11</emphasis> <emphasis id="81A08B5F014F628B826F153598C1CF25" bold="true" pageId="0" pageNumber="145">1-4, 11</emphasis>
), ),
<taxonomicName authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="35FFEF9903A573F12246A4A7F7CE2DDF" authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana</emphasis> <emphasis id="7D5B7FCFF87341DDB2F033F43CE762E6" italics="true" pageId="0" pageNumber="145">Lachesana</emphasis>
</taxonomicName> </taxonomicName>
spp. ( spp. (
<emphasis bold="true" pageId="0" pageNumber="145">5-7</emphasis> <emphasis id="870862C3910843AA57FAADAD5A004D46" bold="true" pageId="0" pageNumber="145">5-7</emphasis>
), ),
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Zodariidae" genus="Parazodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazodarion raddei" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei"> <taxonomicName id="C0DDDFC8F9E4669E2EA0CA7BDCCB4483" baseAuthorityName="Simon" baseAuthorityYear="1889" family="Zodariidae" genus="Parazodarion" kingdom="Animalia" lsidName="Parazodarion raddei" order="Aranei" pageId="0" pageNumber="145" rank="species" species="raddei">
<emphasis italics="true" pageId="0" pageNumber="145">Parazodarion raddei</emphasis> <emphasis id="C59F19C79CB7BEC631E0CC70494D8803" italics="true" pageId="0" pageNumber="145">Parazodarion raddei</emphasis>
</taxonomicName> </taxonomicName>
( (
<emphasis bold="true" pageId="0" pageNumber="145">8</emphasis> <emphasis id="DD21FD6AE4598FE4249EFB6FB40123B7" bold="true" pageId="0" pageNumber="145">8</emphasis>
), ),
<taxonomicName authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="5AF7FD8D1A107A41BACEB296C888C4ED" authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Pax</emphasis> <emphasis id="3DCF4FC4B366B0CB0BBF613006DBFB0E" italics="true" pageId="0" pageNumber="145">Pax</emphasis>
</taxonomicName> </taxonomicName>
spp. ( spp. (
<emphasis bold="true" pageId="0" pageNumber="145">9, 10</emphasis> <emphasis id="741AAD243AA1BA741E59884DF6F79DE5" bold="true" pageId="0" pageNumber="145">9, 10</emphasis>
) and ) and
<taxonomicName authorityName="Simon" authorityYear="1882" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="F763A8D50B3AFF13CF2695BACA8578A0" authorityName="Simon" authorityYear="1882" class="Arachnida" family="Zodariidae" genus="Trygetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygetus" order="Aranei" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Trygetus</emphasis> <emphasis id="E197407341744D795D6005E5246F56C0" italics="true" pageId="0" pageNumber="145">Trygetus</emphasis>
</taxonomicName> </taxonomicName>
spp. ( spp. (
<emphasis bold="true" pageId="0" pageNumber="145">11, 12</emphasis> <emphasis id="FC435744E68E39616B6668DA98C483CA" bold="true" pageId="0" pageNumber="145">11, 12</emphasis>
) in ) in
<collectingCountry name="Iran">Iran</collectingCountry> <collectingCountry id="C6C101F2511106F71F9E67E424B8748D" name="Iran">Iran</collectingCountry>
and and
<collectingCountry name="Turkmenistan">Turkmenistan</collectingCountry> <collectingCountry id="BEC7080193562EC3B2B47AFBF6D15E4B" name="Turkmenistan">Turkmenistan</collectingCountry>
: :
<emphasis bold="true" pageId="0" pageNumber="145">1</emphasis> <emphasis id="DB2114804E5BB5100CF363D9280DFCF3" bold="true" pageId="0" pageNumber="145">1</emphasis>
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa"> <taxonomicName id="1EB42CD8DF6766E36EB5B70A4B598243" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis> <emphasis id="74600BD3CA4D97F52782640F06B315D2" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">2</emphasis> <emphasis id="D35CD5E4DC3BF6E83DF53DE87667ED50" bold="true" pageId="0" pageNumber="145">2</emphasis>
<taxonomicName lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha"> <taxonomicName id="3666701D3718AB02B83B84033B561DCC" lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">A. niusha</emphasis> <emphasis id="9FBBA4E4F038584094117D753928504F" italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">3</emphasis> <emphasis id="58D8F1D099F4D9B41A2FB3D4DFB1BEC4" bold="true" pageId="0" pageNumber="145">3</emphasis>
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis"> <taxonomicName id="12E75191FE2FF83681014866CEB1B402" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis> <emphasis id="F09BCB72DB99FD5596674D2F8B65879E" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">4</emphasis> <emphasis id="866CFF4BC29C4CC689E176510330465A" bold="true" pageId="0" pageNumber="145">4</emphasis>
<taxonomicName lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani"> <taxonomicName id="6267A066F09872E930245F60E3EBE9CF" lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis italics="true" pageId="0" pageNumber="145">A. sorani</emphasis> <emphasis id="C505A1778391468723842DB35B34DD11" italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<specimenCount type="generic"> <specimenCount id="4ED81429E213297064ACD374043BE90D" type="generic">
<emphasis bold="true" pageId="0" pageNumber="145">5</emphasis> <emphasis id="E723FAC7AA690BBF9947BDB10382EC2A" bold="true" pageId="0" pageNumber="145">5</emphasis>
<taxonomicName lsidName="L. insensibilis" pageId="0" pageNumber="145" rank="species" species="insensibilis"> <taxonomicName id="BED4FE9EBFFA6791C97301F5AEB2AD2C" lsidName="L. insensibilis" pageId="0" pageNumber="145" rank="species" species="insensibilis">
<emphasis italics="true" pageId="0" pageNumber="145">L. insensibilis</emphasis> <emphasis id="599CFBCBEF27A7BF64971E4DAFCACF98" italics="true" pageId="0" pageNumber="145">L. insensibilis</emphasis>
</taxonomicName> </taxonomicName>
</specimenCount> </specimenCount>
<specimenCount type="generic"> <specimenCount id="B0F2BDD72ED130195004664F2C655E04" type="generic">
<emphasis bold="true" pageId="0" pageNumber="145">6</emphasis> <emphasis id="2D78638266D3E3FD9969EDED98186234" bold="true" pageId="0" pageNumber="145">6</emphasis>
<taxonomicName lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis"> <taxonomicName id="8473EA687AC6E5B16CC1DD9D1A9FA157" lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis> <emphasis id="34D0FE4F10BE963CD03DC3784E240787" italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
</taxonomicName> </taxonomicName>
</specimenCount> </specimenCount>
sp. nov. sp. nov.
<specimenCount type="generic"> <specimenCount id="77EE2F57283B6CED66F1D88FA4EE4676" type="generic">
<emphasis bold="true" pageId="0" pageNumber="145">7</emphasis> <emphasis id="0FD296EE84E9AC57C1DD887FE23E0790" bold="true" pageId="0" pageNumber="145">7</emphasis>
<taxonomicName lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus"> <taxonomicName id="ED1DD43B22E2839196AA6FC636392FF0" lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis italics="true" pageId="0" pageNumber="145">L. perseus</emphasis> <emphasis id="6B579A69EBC87AB56F4E1B6C34B9A0D5" italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
</taxonomicName> </taxonomicName>
</specimenCount> </specimenCount>
sp. nov. sp. nov.
<specimenCount type="generic"> <specimenCount id="5C50BB8E0AAC1CB72C4768885B996BA2" type="generic">
<emphasis bold="true" pageId="0" pageNumber="145">8</emphasis> <emphasis id="87072DE9A6CE21056933628B2CC92D6D" bold="true" pageId="0" pageNumber="145">8</emphasis>
<taxonomicName lsidName="P. raddei" pageId="0" pageNumber="145" rank="species" species="raddei"> <taxonomicName id="3D448C0E9179A704511C8B25D9E0C70C" lsidName="P. raddei" pageId="0" pageNumber="145" rank="species" species="raddei">
<emphasis italics="true" pageId="0" pageNumber="145">P. raddei</emphasis> <emphasis id="1EF0A2CA3C061C038B5080A2A7CA9EA6" italics="true" pageId="0" pageNumber="145">P. raddei</emphasis>
</taxonomicName> </taxonomicName>
</specimenCount> </specimenCount>
<specimenCount type="generic"> <specimenCount id="6F0C84994DF62CC670307B4B3373B937" type="generic">
<emphasis bold="true" pageId="0" pageNumber="145">9</emphasis> <emphasis id="30BC5BCE08A50BD0832ED60118EFCD8F" bold="true" pageId="0" pageNumber="145">9</emphasis>
<taxonomicName lsidName="P. ellipita" pageId="0" pageNumber="145" rank="species" species="ellipita"> <taxonomicName id="25371C293CAEF5D4EB46BBFC697D367B" lsidName="P. ellipita" pageId="0" pageNumber="145" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">P. ellipita</emphasis> <emphasis id="8594ABD5A4CB139385D21C257B1194FE" italics="true" pageId="0" pageNumber="145">P. ellipita</emphasis>
</taxonomicName> </taxonomicName>
</specimenCount> </specimenCount>
sp. nov. sp. nov.
<specimenCount type="generic"> <specimenCount id="AC2EA74861E0F1783752F3A5A74A4C2F" type="generic">
<emphasis bold="true" pageId="0" pageNumber="145">10</emphasis> <emphasis id="24EAF3B05C74FEDB351C4C92DDBEC454" bold="true" pageId="0" pageNumber="145">10</emphasis>
<taxonomicName lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila"> <taxonomicName id="D6585C63332E7EBAC1AD78C60320B09F" lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila">
<emphasis italics="true" pageId="0" pageNumber="145">P. leila</emphasis> <emphasis id="BC95DC2986114351A15359ED908F1FF5" italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
</taxonomicName> </taxonomicName>
</specimenCount> </specimenCount>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">11</emphasis> <emphasis id="22EFC43923B26EABF9CC13B52B3F4A70" bold="true" pageId="0" pageNumber="145">11</emphasis>
<taxonomicName lsidName="A. ovtchinnikovi" pageId="0" pageNumber="145" rank="species" species="ovtchinnikovi"> <taxonomicName id="B13424CEA8F6C1E718842CE8A7E8ADB3" lsidName="A. ovtchinnikovi" pageId="0" pageNumber="145" rank="species" species="ovtchinnikovi">
<emphasis italics="true" pageId="0" pageNumber="145">A. ovtchinnikovi</emphasis> <emphasis id="A84C2522F45852E3157D499597B971CB" italics="true" pageId="0" pageNumber="145">A. ovtchinnikovi</emphasis>
</taxonomicName> </taxonomicName>
sp. nov., sp. nov.,
<taxonomicName lsidName="T. gromovi" pageId="0" pageNumber="145" rank="species" species="gromovi"> <taxonomicName id="265F174DF793F03DE1C41635F3551BAA" lsidName="T. gromovi" pageId="0" pageNumber="145" rank="species" species="gromovi">
<emphasis italics="true" pageId="0" pageNumber="145">T. gromovi</emphasis> <emphasis id="97DF6F11AC90263F0012E11CF5473CDC" italics="true" pageId="0" pageNumber="145">T. gromovi</emphasis>
</taxonomicName> </taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="145">12</emphasis> <emphasis id="96752460472E3806335BBE81BF80AD58" bold="true" pageId="0" pageNumber="145">12</emphasis>
<taxonomicName lsidName="T. susianus" pageId="0" pageNumber="145" rank="species" species="susianus"> <taxonomicName id="E5B7AB2824AAEDFF72C3E82E4FAB3704" lsidName="T. susianus" pageId="0" pageNumber="145" rank="species" species="susianus">
<emphasis italics="true" pageId="0" pageNumber="145">T. susianus</emphasis> <emphasis id="A50ABCB6203AE65327CDCF40E569BBB5" italics="true" pageId="0" pageNumber="145">T. susianus</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
</paragraph> </paragraph>
</caption> </caption>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comments"> <subSubSection id="D15536E1F48D40385F5D20F76E3442FA" pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="FAE8B22F9E8C7F3442DAE1F576A4CB03" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="4CABE2A7DC4C602557E59420DDF33FCC" pageId="0" pageNumber="145">
This species was previously known from the original description only. It differs from the similar This species was previously known from the original description only. It differs from the similar
<taxonomicName lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae"> <taxonomicName id="22DF70C58918AC0517E12CDFE064037B" lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae">
<emphasis italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis> <emphasis id="7C6308E63CAFBB1517CCCA6F31BB0E6F" italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis>
</taxonomicName> </taxonomicName>
by the shape of the male palpal tibia and the epigyne (see by the shape of the male palpal tibia and the epigyne (see
<bibRefCitation author="Nenilin, AB" journalOrPublisher="Zoologicheskiĭ Zhurnal" pageId="0" pageNumber="145" pagination="618 - 620" refId="B17" refString="Nenilin, AB, Fet, VY, 1985. New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia. Zoologicheskiĭ Zhurnal 64: 618 - 620" title="New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia." volume="64" year="1985">Nenilin and Fet 1985</bibRefCitation> <bibRefCitation id="F30F1C5EF91985ECF82672745D194205" author="Nenilin, AB" journalOrPublisher="Zoologicheskiĭ Zhurnal" pageId="0" pageNumber="145" pagination="618 - 620" refId="B17" refString="Nenilin, AB, Fet, VY, 1985. New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia. Zoologicheskiĭ Zhurnal 64: 618 - 620" title="New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia." volume="64" year="1985">Nenilin and Fet 1985</bibRefCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in iran"> <subSubSection id="6F824BA35F9E676337D1FB6236317FBA" pageId="0" pageNumber="145" type="records in iran">
<paragraph pageId="0" pageNumber="145">Records in Iran.</paragraph> <paragraph id="F231D1CF1F254E7F66C50C3AFDFFF372" pageId="0" pageNumber="145">Records in Iran.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="BAD21158921768D04A23564DAEBA985D" pageId="0" pageNumber="145">
Razavi Khorasan (current data) (Fig. Razavi Khorasan (current data) (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation> <figureCitation id="075B469ACE0539D17ACD00730C2B0CC0" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="records in turkmenistan"> <subSubSection id="5B0D4D407E5FA6086E0C9B221340775B" pageId="0" pageNumber="145" type="records in turkmenistan">
<paragraph pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph> <paragraph id="DBABCD03D0F554F7633EEA89A2B91437" pageId="0" pageNumber="145">Records in Turkmenistan.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="5AE24525BBC6DBA1F222C33A5326F4C2" pageId="0" pageNumber="145">
Balkan ( Balkan (
<bibRefCitation author="Nenilin, AB" journalOrPublisher="Zoologicheskiĭ Zhurnal" pageId="0" pageNumber="145" pagination="618 - 620" refId="B17" refString="Nenilin, AB, Fet, VY, 1985. New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia. Zoologicheskiĭ Zhurnal 64: 618 - 620" title="New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia." volume="64" year="1985">Nenilin and Fet 1985</bibRefCitation> <bibRefCitation id="A7112A588ADED7A60EA2762F4F935961" author="Nenilin, AB" journalOrPublisher="Zoologicheskiĭ Zhurnal" pageId="0" pageNumber="145" pagination="618 - 620" refId="B17" refString="Nenilin, AB, Fet, VY, 1985. New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia. Zoologicheskiĭ Zhurnal 64: 618 - 620" title="New species of spiders of the genus Zodarion (Aranei, Zodariidae) from Turkmenia." volume="64" year="1985">Nenilin and Fet 1985</bibRefCitation>
) (Fig. ) (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation> <figureCitation id="2BFB4F0EF462613D5832514B0397ECC6" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
). ).
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145" start="Figure 33" startId="F33"> <caption id="C02EA5C7EE2BA0CA035B7CD8BFD57687" doi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145" start="Figure 33" startId="F33">
<paragraph pageId="0" pageNumber="145"> <paragraph id="C10B1A57E7F7007C741C89969EA513C7" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 33.</emphasis> <emphasis id="257DB7163396D3697DCE985B1884E20E" bold="true" pageId="0" pageNumber="145">Figure 33.</emphasis>
Distribution records of Distribution records of
<taxonomicName authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="1A9CEE7147358B41918BD842798D8A72" authorityName="Denis" authorityYear="1966" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis> <emphasis id="8534AE684AC3105DDDF95AE5777DA3C7" italics="true" pageId="0" pageNumber="145">Acanthinozodium</emphasis>
</taxonomicName> </taxonomicName>
spp. (partim spp. (partim
<emphasis bold="true" pageId="0" pageNumber="145">1-3, 5, 6, 8, 9</emphasis> <emphasis id="64E76E00C7AFCF71F4E350AC26701DB4" bold="true" pageId="0" pageNumber="145">1-3, 5, 6, 8, 9</emphasis>
), ),
<taxonomicName authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="B640AE8465ADFED68986FDDA2333DD5D" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodariellum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis> <emphasis id="84BC7DBA62FACA5A418DBF26FC6C9933" italics="true" pageId="0" pageNumber="145">Zodariellum</emphasis>
</taxonomicName> </taxonomicName>
spp. ( spp. (
<emphasis bold="true" pageId="0" pageNumber="145">10, 11</emphasis> <emphasis id="450B07E8564820B99EF080EB11AD866B" bold="true" pageId="0" pageNumber="145">10, 11</emphasis>
) and ) and
<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="B86DF6BB78E04CEED67515A7FC70E167" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis> <emphasis id="C77936BA4222503B64E7B2C432F7F388" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
</taxonomicName> </taxonomicName>
spp. ( spp. (
<emphasis bold="true" pageId="0" pageNumber="145">4, 7, 12</emphasis> <emphasis id="D696FC34DF279C19569CC705F327B8D5" bold="true" pageId="0" pageNumber="145">4, 7, 12</emphasis>
) in Iran and Turkmenistan: ) in Iran and Turkmenistan:
<emphasis bold="true" pageId="0" pageNumber="145">1</emphasis> <emphasis id="D1E46D99494C80A4C4280C4801B2EC92" bold="true" pageId="0" pageNumber="145">1</emphasis>
<taxonomicName lsidName="A. armita" pageId="0" pageNumber="145" rank="species" species="armita"> <taxonomicName id="EBD3C22BF61BBFA2F5F1774FA771A46A" lsidName="A. armita" pageId="0" pageNumber="145" rank="species" species="armita">
<emphasis italics="true" pageId="0" pageNumber="145">A. armita</emphasis> <emphasis id="A226C1B5AC5CF119CAC91B7FBD68970E" italics="true" pageId="0" pageNumber="145">A. armita</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">2</emphasis> <emphasis id="DBF530EBE6409B8E918E895CB8AB7F4B" bold="true" pageId="0" pageNumber="145">2</emphasis>
<taxonomicName lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara"> <taxonomicName id="DD22DB2C7345B278810FE8ABB844454A" lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">A. diara</emphasis> <emphasis id="9DC450ADE5D660E3F5F25C8CEC465872" italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">3</emphasis> <emphasis id="89E897D72CE64FBA78462E3A7622A59F" bold="true" pageId="0" pageNumber="145">3</emphasis>
<taxonomicName lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum"> <taxonomicName id="01102B8306887501804EBF8BF457597E" lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis> <emphasis id="FD767A0F50A6ABE1770DBF70CB8B7DC4" italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">4</emphasis> <emphasis id="EFCB10AE29A525B87650C1A8CEA76A8F" bold="true" pageId="0" pageNumber="145">4</emphasis>
<taxonomicName lsidName="Z. buettikeri" pageId="0" pageNumber="145" rank="species" species="buettikeri"> <taxonomicName id="02A8FB5731B5B6538D22B39CFCF6AB3F" lsidName="Z. buettikeri" pageId="0" pageNumber="145" rank="species" species="buettikeri">
<emphasis italics="true" pageId="0" pageNumber="145">Z. buettikeri</emphasis> <emphasis id="D825ED10312B149B85D55593EC00FA48" italics="true" pageId="0" pageNumber="145">Z. buettikeri</emphasis>
</taxonomicName> </taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="145">5</emphasis> <emphasis id="8E8764624C4CF4A8F4878AADBAFF834C" bold="true" pageId="0" pageNumber="145">5</emphasis>
<taxonomicName lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa"> <taxonomicName id="B4D7CAA0987DA27A5FD573864D08366A" lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis> <emphasis id="C0E2F378CD05C967E5D2A2B8B29E845F" italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">6</emphasis> <emphasis id="C8F5EBFE48FA37F6ECAEB08EE8BD0D42" bold="true" pageId="0" pageNumber="145">6</emphasis>
<taxonomicName lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana"> <taxonomicName id="C85FB6195DC43D4E0D86F310F07CC358" lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana">
<emphasis italics="true" pageId="0" pageNumber="145">A. kiana</emphasis> <emphasis id="94A6CBB39208BD3435909B0F2AD4E031" italics="true" pageId="0" pageNumber="145">A. kiana</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">7</emphasis> <emphasis id="25F3F0EF54FAB225F41EE16A38307F38" bold="true" pageId="0" pageNumber="145">7</emphasis>
<taxonomicName lsidName="Z. lutipes" pageId="0" pageNumber="145" rank="species" species="lutipes"> <taxonomicName id="DE2885C6EE53B0B4DB6326D79DACDD77" lsidName="Z. lutipes" pageId="0" pageNumber="145" rank="species" species="lutipes">
<emphasis italics="true" pageId="0" pageNumber="145">Z. lutipes</emphasis> <emphasis id="0128A9DFCD3A58828521A64421D947A6" italics="true" pageId="0" pageNumber="145">Z. lutipes</emphasis>
</taxonomicName> </taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="145">8</emphasis> <emphasis id="B43833DB70828684D8F1EDE3E6BE7AF6" bold="true" pageId="0" pageNumber="145">8</emphasis>
<taxonomicName lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa"> <taxonomicName id="DB1BE73D2E6FE2AF6D724AA5BC6E2582" lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis italics="true" pageId="0" pageNumber="145">A. masa</emphasis> <emphasis id="3DB04AFC0117CD8C0A06F9410C67124F" italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">9</emphasis> <emphasis id="69CED20E95AC29EC18AEC87840168737" bold="true" pageId="0" pageNumber="145">9</emphasis>
<taxonomicName lsidName="A. parmida" pageId="0" pageNumber="145" rank="species" species="parmida"> <taxonomicName id="B7FC225D770FEDE6806B1F7291B169B4" lsidName="A. parmida" pageId="0" pageNumber="145" rank="species" species="parmida">
<emphasis italics="true" pageId="0" pageNumber="145">A. parmida</emphasis> <emphasis id="5C218968B99E991647C2207EBB8B2256" italics="true" pageId="0" pageNumber="145">A. parmida</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">10</emphasis> <emphasis id="E66B65D1141768D698D3FF7FD3DA3BDA" bold="true" pageId="0" pageNumber="145">10</emphasis>
<taxonomicName lsidName="Z. proszynskii" pageId="0" pageNumber="145" rank="species" species="proszynskii"> <taxonomicName id="64F9CB278EBA055B4F87C68B39B78056" lsidName="Z. proszynskii" pageId="0" pageNumber="145" rank="species" species="proszynskii">
<emphasis italics="true" pageId="0" pageNumber="145">Z. proszynskii</emphasis> <emphasis id="8DAD4E4ED3E71F600DB60516305F2595" italics="true" pageId="0" pageNumber="145">Z. proszynskii</emphasis>
</taxonomicName> </taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="145">11</emphasis> <emphasis id="95184A3018014774B2C858E318A0148C" bold="true" pageId="0" pageNumber="145">11</emphasis>
<taxonomicName lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae"> <taxonomicName id="BDD48FC9D148E28E4051FF87566B9B6B" lsidName="Z. sytchevskajae" pageId="0" pageNumber="145" rank="species" species="sytchevskajae">
<emphasis italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis> <emphasis id="B3DC8FC08E48632B3C0E0F01F1D96C5F" italics="true" pageId="0" pageNumber="145">Z. sytchevskajae</emphasis>
</taxonomicName> </taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="145">12</emphasis> <emphasis id="C6D95EF15AD5AE93D613AF191868F066" bold="true" pageId="0" pageNumber="145">12</emphasis>
<taxonomicName lsidName="Z. talyschicum" pageId="0" pageNumber="145" rank="species" species="talyschicum"> <taxonomicName id="F8E9462709171B4E6FC196EB5A5BAE77" lsidName="Z. talyschicum" pageId="0" pageNumber="145" rank="species" species="talyschicum">
<emphasis italics="true" pageId="0" pageNumber="145">Z. talyschicum</emphasis> <emphasis id="9BE1C78A5D4D8AE45ADE4727230DF1AB" italics="true" pageId="0" pageNumber="145">Z. talyschicum</emphasis>
</taxonomicName> </taxonomicName>
. .
</paragraph> </paragraph>
</caption> </caption>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="054985D7426EAE74BCD88EE83DFB3F39" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="4F6E731070FB4647AFAC842A3B8A32A3" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">Previously known only from the type locality in Turkmenistan. This is a new record for Iran, representing the southernmost record in the known genus range.</paragraph> <paragraph id="7077C86C135DA2DF4543C3B924458997" pageId="0" pageNumber="145">Previously known only from the type locality in Turkmenistan. This is a new record for Iran, representing the southernmost record in the known genus range.</paragraph>
</subSubSection> </subSubSection>
</treatment> </treatment>
</document> </document>

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@ -1,167 +1,167 @@
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title> <mods:title id="F0E0EBF3225110B8F17CAB53FEF4E557">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<taxonomicName LSID="http://zoobank.org/57AB4566-6EB3-4ACF-A4CD-E69F22ECD172" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parmida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parmida" status="sp. nov.">Acanthinozodium parmida</taxonomicName> <taxonomicName id="E1103A13784A9440E3F4539726AF5B87" ID-CoL="98YYS" LSID="http://zoobank.org/57AB4566-6EB3-4ACF-A4CD-E69F22ECD172" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parmida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parmida" status="sp. nov.">Acanthinozodium parmida</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="0673BCD421D4DEF6F6A46FAD3198C5D1" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">Figs 19A-C</figureCitation> <figureCitation id="46152494F35F26FF83BB34DDEF2B98E3" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">Figs 19A-C</figureCitation>
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. SEM images of the male palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, B, D ventral C, E retrolateral F dorsoretrolateral. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis, Ra - retrolateral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145">, 25A-C</figureCitation> <figureCitation id="5989CA90BC92E92DCB963E8428CCF013" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. SEM images of the male palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, B, D ventral C, E retrolateral F dorsoretrolateral. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis, Ra - retrolateral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145">, 25A-C</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation> <figureCitation id="142455B99D9CC093B11EB466C433CD63" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="A9E9EF8445391010562F1233592FC8D2" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="E53D4ED3210F20FB8A038CFB323B1106" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="D036F9649138050D1B7F13C467EC18BB" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="60589CFCED3A8D0E8FB66D874035987F" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran: ♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Isfahan Province</emphasis> <emphasis id="1B37337B477954F82E18447965F701F7" italics="true" pageId="0" pageNumber="145">Isfahan Province</emphasis>
: Qamsar and Barzok Protected Area, 55 km SW of Qamsar, 14 km NE Kamoo, near the road of Gargash observatory, : Qamsar and Barzok Protected Area, 55 km SW of Qamsar, 14 km NE Kamoo, near the road of Gargash observatory,
<geoCoordinate degrees="33" direction="north" minutes="37" orientation="latitude" precision="925" value="33.616665">33°37'N</geoCoordinate> <geoCoordinate id="78D414568B991BD2A03D7441EC0B654C" degrees="33" direction="north" minutes="37" orientation="latitude" precision="925" value="33.616665">33°37'N</geoCoordinate>
, ,
<geoCoordinate degrees="51" direction="east" minutes="19" orientation="longitude" precision="925" value="51.316666">51°19'E</geoCoordinate> <geoCoordinate id="7AB141CCC27D4182ADBC475316C2F85F" degrees="51" direction="east" minutes="19" orientation="longitude" precision="925" value="51.316666">51°19'E</geoCoordinate>
, 2710 m, 19.05.2016 (P. Ponel). , 2710 m, 19.05.2016 (P. Ponel).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="23D9F20F1CD144EFD85DABE5CAA2DF30" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="F172A8668452A98C7775553471C14566" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet refers to a Persian princess, the only daughter of Bardiya (Smerdis), son of Cyrus the Great. Noun in apposition.</paragraph> <paragraph id="0A01ABD223F0F410131CDA6922775C1C" pageId="0" pageNumber="145">The specific epithet refers to a Persian princess, the only daughter of Bardiya (Smerdis), son of Cyrus the Great. Noun in apposition.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="714F434CEA413D905F4E34C8C92B8672" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="95CBD790C464C26368BBD1D1F121E2C3" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="52A834F51748C6C1EF4BB3C48F654E5D" pageId="0" pageNumber="145">
The new species differs from the congeners in the region by its smaller size, by having a dorsal scutum, and by the very short tip (free part) of embolus having a long furrow with a serrate ventral margin. It is closely related to The new species differs from the congeners in the region by its smaller size, by having a dorsal scutum, and by the very short tip (free part) of embolus having a long furrow with a serrate ventral margin. It is closely related to
<taxonomicName lsidName="A. ovtchinnikovi" pageId="0" pageNumber="145" rank="species" species="ovtchinnikovi"> <taxonomicName id="A41D3763C15B697528383FD4605570FF" lsidName="A. ovtchinnikovi" pageId="0" pageNumber="145" rank="species" species="ovtchinnikovi">
<emphasis italics="true" pageId="0" pageNumber="145">A. ovtchinnikovi</emphasis> <emphasis id="986052AB780596F88524C47F3579DDA1" italics="true" pageId="0" pageNumber="145">A. ovtchinnikovi</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. from southeastern Turkmenistan, from which it can be differentiated by having an almost as long as wide bulb ( sp. nov. from southeastern Turkmenistan, from which it can be differentiated by having an almost as long as wide bulb (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="B65B4858E391103C4B4C401BD31286D6" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
longer than wide), shorter cymbium (length/width ratio 1.4 vs. 1.86), a different shape of median apophysis (posterior portion larger than anterior one, vs. opposite) and the different position of embolic base (6:30 o'clock, longer than wide), shorter cymbium (length/width ratio 1.4 vs. 1.86), a different shape of median apophysis (posterior portion larger than anterior one, vs. opposite) and the different position of embolic base (6:30 o'clock,
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="951FB32D642C89A82A00BD91EFFA017E" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
8:00 o'clock; cf. Figs 8:00 o'clock; cf. Figs
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19B</figureCitation> <figureCitation id="A8784FC3EA376B8598ACE44483ECDECA" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19B</figureCitation>
and and
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22G</figureCitation> <figureCitation id="6CFA98CE10A0A6FA11F983A48F3954B4" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22G</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="66B5D79D8BBF87878ED9145E6351E792" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="C6C91FDE1742031C3674C2D052A2F15E" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="0858E77A05CF7D99AFCDF9D531041E84" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis> <emphasis id="09EA750BC0F26C83AFC59D7111F31719" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Habitus as in Fig. Habitus as in Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19A</figureCitation> <figureCitation id="0D677BA00F155DE5574EC53F6CEFC595" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19A</figureCitation>
. Total length 1.75. Carapace 0.83 long, 0.42 wide at pars cephalica, 0.60 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.10, ALE: 0.06, PME: 0.06, PLE: 0.05, PME-PME: 0.10. Carapace, sternum, labium, chelicerae and maxillae yellowish; carapace with irregular dark patterns. Chelicera with one tooth. Legs yellowish, without annulations. Abdomen dorsally black, covered with scutum, ventrally grayish. Spinnerets pale, uniform in color. Measurements of legs: I: 1.89 (0.42, 0.23, 0.41, 0.41, 0.42), II: 1.64 (0.40, 0.23, 0.31, 0.33, 0.37), III: 1.54 (0.41, 0.21, 0.32, 0.29, 0.31), IV: 2.35 (0.60, 0.26, 0.52, 0.59, 0.38). . Total length 1.75. Carapace 0.83 long, 0.42 wide at pars cephalica, 0.60 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.10, ALE: 0.06, PME: 0.06, PLE: 0.05, PME-PME: 0.10. Carapace, sternum, labium, chelicerae and maxillae yellowish; carapace with irregular dark patterns. Chelicera with one tooth. Legs yellowish, without annulations. Abdomen dorsally black, covered with scutum, ventrally grayish. Spinnerets pale, uniform in color. Measurements of legs: I: 1.89 (0.42, 0.23, 0.41, 0.41, 0.42), II: 1.64 (0.40, 0.23, 0.31, 0.33, 0.37), III: 1.54 (0.41, 0.21, 0.32, 0.29, 0.31), IV: 2.35 (0.60, 0.26, 0.52, 0.59, 0.38).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="56289EB124928399D6F12EA7B3AFEE76" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19B, C</figureCitation> <figureCitation id="D1E445885C260A9295F5CF2971181671" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19B, C</figureCitation>
, ,
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. SEM images of the male palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, B, D ventral C, E retrolateral F dorsoretrolateral. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis, Ra - retrolateral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145">25A-C</figureCitation> <figureCitation id="565B1981F25433CCCCC7E7653B1D0713" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. SEM images of the male palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, B, D ventral C, E retrolateral F dorsoretrolateral. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis, Ra - retrolateral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145">25A-C</figureCitation>
. Ventral tibial apophysis lacking, RTA ( . Ventral tibial apophysis lacking, RTA (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis> <emphasis id="8318C6DD46805B001B87B79A1CF59BEB" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
) almost triangular, slightly longer than tibia; cymbium very broad; tegulum slightly wider than long; sperm duct tracking margin of tegulum, lacking any turns; median apophysis ( ) almost triangular, slightly longer than tibia; cymbium very broad; tegulum slightly wider than long; sperm duct tracking margin of tegulum, lacking any turns; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="43A41C1CF3C2D54526BC7EBAA247E4AA" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) relatively small, less than half of tegulum's height, posterior part larger than anterior; embolus very short, shorter than median apophysis and conductor ( ) relatively small, less than half of tegulum's height, posterior part larger than anterior; embolus very short, shorter than median apophysis and conductor (
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis> <emphasis id="43FA70018ACFD834ADA5BD66574912D8" italics="true" pageId="0" pageNumber="145">Co</emphasis>
), embolus originating at about 6:30 o'clock position with its terminal 2/3 having a longitudinal furrow ( ), embolus originating at about 6:30 o'clock position with its terminal 2/3 having a longitudinal furrow (
<emphasis italics="true" pageId="0" pageNumber="145">Er</emphasis> <emphasis id="66334AC3D5716844DEAACE6064E51BF8" italics="true" pageId="0" pageNumber="145">Er</emphasis>
) with a finely serrated ventral margin. ) with a finely serrated ventral margin.
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145" start="Figure 25" startId="F25"> <caption id="A0B0E3FF0F739B1DE2593493E13EC08B" doi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145" start="Figure 25" startId="F25">
<paragraph pageId="0" pageNumber="145"> <paragraph id="6D9CAB18535DA365E7D1C641EB66734B" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 25.</emphasis> <emphasis id="18BFFC9503B457A841C1D09286E9033B" bold="true" pageId="0" pageNumber="145">Figure 25.</emphasis>
SEM images of the male palps of SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parmida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parmida"> <taxonomicName id="5D68A24C7A81F7C7990AC811DCEB937F" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parmida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parmida">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium parmida</emphasis> <emphasis id="D326AF4D01728BD9DFE5542BF3FE247B" italics="true" pageId="0" pageNumber="145">Acanthinozodium parmida</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis> <emphasis id="71EF152D0CC8C0F6DE99BE4F7306AED4" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and ) and
<taxonomicName class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion expers" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="expers"> <taxonomicName id="9A045FF7A07615A38487421F6875BDCA" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion expers" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="expers">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion expers</emphasis> <emphasis id="301138A24C49BD40ACB48701E17E8DBE" italics="true" pageId="0" pageNumber="145">Zodarion expers</emphasis>
</taxonomicName> </taxonomicName>
( (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis> <emphasis id="37C27760999B4E922614A838D0102014" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, B, D</emphasis> <emphasis id="D012CCABFF87AB297A962E3BDDFB3160" bold="true" pageId="0" pageNumber="145">A, B, D</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">C, E</emphasis> <emphasis id="B8451FA4FB25553D2BAE0401B0D50AA6" bold="true" pageId="0" pageNumber="145">C, E</emphasis>
retrolateral retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis> <emphasis id="816F4C51676DE6FD7B58F87038793F28" bold="true" pageId="0" pageNumber="145">F</emphasis>
dorsoretrolateral. Abbreviations: dorsoretrolateral. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Cg</emphasis> <emphasis id="A7B9B0D8B5FA166B084805197F3571C3" italics="true" pageId="0" pageNumber="145">Cg</emphasis>
- cymbial groove, - cymbial groove,
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis> <emphasis id="47B4A478042A4E918D256A93C93F0357" italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor, - conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis> <emphasis id="B6EF05B1FD8ADBD812DA7F9CD1E119E4" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus, - embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Er</emphasis> <emphasis id="27CB80EAB52957480399399D1E7C6643" italics="true" pageId="0" pageNumber="145">Er</emphasis>
- embolar ridge, - embolar ridge,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="C5BF2D618686F2A5E565F55C7AD028A2" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis, - median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis> <emphasis id="649862489E38C5C0C44A4EF4F52D1624" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral tibial apophysis. Scale bars: 0.1 mm. - retrolateral tibial apophysis. Scale bars: 0.1 mm.
</paragraph> </paragraph>
</caption> </caption>
<paragraph pageId="0" pageNumber="145"> <paragraph id="F257324C018D436D7D8CE85E1C83EC3C" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="58BACFE6CBAD8AE12E74D627D571B0B6" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown. Unknown.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="60A33943DE0F6088028E0A1BD6BBD784" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="76D7EE9D714437847E757BF9C0DFC9B3" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="E6CA2241CA65DB642D870089CFA29331" pageId="0" pageNumber="145">
Known from the type locality in Isfahan Province, central Iran (Fig. Known from the type locality in Isfahan Province, central Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation> <figureCitation id="3D82C5B5D8B0F7CB4A1BB7A157CBCF38" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

258
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2021" docId="C18D05F563465EA48B4B75AB8EAF40A2" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Lachesana perseus Zamani &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="F3F99EF7-B5F4-42CB-AD3B-2404007AFBA2" docUuidSource="ZooBank" docVersion="4" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1668150222111" updateUser="ExternalLinkService"> <document ID-CLB-Dataset="7157" ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2021" docId="C18D05F563465EA48B4B75AB8EAF40A2" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Lachesana perseus Zamani &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="F3F99EF7-B5F4-42CB-AD3B-2404007AFBA2" docUuidSource="ZooBank" docVersion="6" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1732670661048" updateUser="ExternalLinkService">
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title> <mods:title id="BB7648F9AFF85B0F570755C243021875">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:namePart>Zamani, Alireza</mods:namePart> <mods:namePart id="161DB7506B52B02D5EFB32AF9379B538">Zamani, Alireza</mods:namePart>
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<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation> <mods:affiliation id="01E4EA145F8F5FF05BE102E80F01C444">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation> <mods:affiliation id="39F791D7D2FCE6F0C6CC1D0DADF8472D">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<treatment ID-GBIF-Taxon="181753554" LSID="urn:lsid:zoobank.org:act:F3F99EF7-B5F4-42CB-AD3B-2404007AFBA2" httpUri="http://treatment.plazi.org/id/C18D05F563465EA48B4B75AB8EAF40A2" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="C18D05F563465EA48B4B75AB8EAF40A2" ID-GBIF-Taxon="181753554" LSID="urn:lsid:zoobank.org:act:F3F99EF7-B5F4-42CB-AD3B-2404007AFBA2" httpUri="http://treatment.plazi.org/id/C18D05F563465EA48B4B75AB8EAF40A2" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Lachesaninae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="845C1BA24484C1BB004502B50DEEE3BE" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="BBA2982FCDCD950D2F7CCCB5A1685745" pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/F3F99EF7-B5F4-42CB-AD3B-2404007AFBA2" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana perseus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="perseus" status="sp. nov.">Lachesana perseus</taxonomicName> <taxonomicName id="C33C25152DD2967F0B586564437BF9E6" ID-CoL="98YYX" LSID="http://zoobank.org/F3F99EF7-B5F4-42CB-AD3B-2404007AFBA2" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana perseus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="perseus" status="sp. nov.">Lachesana perseus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="61613BBFC11310B8911AF37D0CDC64C1" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">Figs 1D-F</figureCitation> <figureCitation id="C001E61BA894AD3CE3460CF357CD9EA3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">Figs 1D-F</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live habitus of male Lachesana perseus sp. nov. Photo: A. Zamani." figureDoi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145">, 2</figureCitation> <figureCitation id="D5AC0B550EC5C290E8143EA1B5065D0E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live habitus of male Lachesana perseus sp. nov. Photo: A. Zamani." figureDoi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">, 3F-J</figureCitation> <figureCitation id="F238BFACD8FDCE897EA02FD10FDC58BE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">, 3F-J</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">, 4J</figureCitation> <figureCitation id="AC2DE1B1721AC498E801F8B75B3239AC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">, 4J</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation> <figureCitation id="B178CF70A9DBB419C034AB91486F9D5A" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="87671BD4CA74B642AAF06F69A7D87615" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="576B6D34BF7062F3EF6CB299B3EA7490" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="5B661A38C023F1E6C784EB5654A2377F" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="596DCE962F379C0A97CC14DC5AE80F0A" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ and ♂ and
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis> <emphasis id="C2FC2A19C29202E1F0029695B3E7B6FF" bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis>
1♂ (MHNG), Iran: 1♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Alborz Province</emphasis> <emphasis id="59AB063C8F3979DA6FD4AF4837E32D0A" italics="true" pageId="0" pageNumber="145">Alborz Province</emphasis>
: Eshtehard, Jaru, : Eshtehard, Jaru,
<geoCoordinate degrees="35" direction="north" minutes="44" orientation="latitude" precision="925" value="35.733334">35°44'N</geoCoordinate> <geoCoordinate id="5F6C02A3209AD75805951C9A43E85196" degrees="35" direction="north" minutes="44" orientation="latitude" precision="925" value="35.733334">35°44'N</geoCoordinate>
, ,
<geoCoordinate degrees="50" direction="east" minutes="35" orientation="longitude" precision="925" value="50.583332">50°35'E</geoCoordinate> <geoCoordinate id="D9AC6272898100BCD28564661B0972E8" degrees="50" direction="east" minutes="35" orientation="longitude" precision="925" value="50.583332">50°35'E</geoCoordinate>
, 10.2018 (A. Zamani). , 10.2018 (A. Zamani).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="F7BBBD96BA50651BF1DB4A0498E931EC" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="23B37499EDB9B3EB05827BCC7BB81175" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet refers to the legendary founder of Mycenae and of the Perseid dynasty in Greek mythology. Noun in apposition.</paragraph> <paragraph id="EA3780F47B3CC52CFE115FF1FF9B3E2D" pageId="0" pageNumber="145">The specific epithet refers to the legendary founder of Mycenae and of the Perseid dynasty in Greek mythology. Noun in apposition.</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145" start="Figure 2" startId="F2"> <caption id="5FB1607EB3265A211A3A8617F4AAA8A4" doi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="145"> <paragraph id="84F050CFCFE1A02C353FA05AEF059E8B" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 2.</emphasis> <emphasis id="F28A0BF5C5E14491EB477286B4533FEA" bold="true" pageId="0" pageNumber="145">Figure 2.</emphasis>
Live habitus of male Live habitus of male
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana perseus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="perseus"> <taxonomicName id="4B73561D9058BB4DBDA2DFF5BA9FB1E8" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana perseus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="perseus">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana perseus</emphasis> <emphasis id="348A24E0AE32407E3D19C771DB43BCE1" italics="true" pageId="0" pageNumber="145">Lachesana perseus</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. Photo: A. Zamani. sp. nov. Photo: A. Zamani.
</paragraph> </paragraph>
</caption> </caption>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="D3C8686AC86D0A2A76334FAB8B713654" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="1867A39548B62E39870A5278289F4ABC" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="CCC88A0E0DA2B33BA791848E3EFDB688" pageId="0" pageNumber="145">
The new species differs from the similar The new species differs from the similar
<taxonomicName lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis"> <taxonomicName id="F9FF47EFD8655668402C8AA573876293" lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis> <emphasis id="1C2BF81EE2AEFF95398C842F63411ADF" italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. by the lack of spines in the proximal mesal part of the cymbium ( sp. nov. by the lack of spines in the proximal mesal part of the cymbium (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="19905CC5A9CADFC049A5D93A3568B50B" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
present; cf. Fig. present; cf. Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3C</figureCitation> <figureCitation id="BA5CFC78108A5251F3376253A21EF5B0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3C</figureCitation>
and and
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3H</figureCitation> <figureCitation id="4F90C4BFA2F9717A812FF7F4D17B36A3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3H</figureCitation>
) as well as by the RTA with the terminal half almost as wide as the tibia ( ) as well as by the RTA with the terminal half almost as wide as the tibia (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="B15485D65F37F0F8620A09913F302C83" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
1.5 times thinner). The shape of the RTA readily distinguishes this species from congeners in the region (stalk abruptly bent basally and tip gently curved; cf. Fig. 1.5 times thinner). The shape of the RTA readily distinguishes this species from congeners in the region (stalk abruptly bent basally and tip gently curved; cf. Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4J</figureCitation> <figureCitation id="CDE4FAF8955D141F79CE0D5DF3613CFB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4J</figureCitation>
and and
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4F-I</figureCitation> <figureCitation id="F86AF3682A06AFE011C0996E5671A9E0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4F-I</figureCitation>
). ).
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145" start="Figure 3" startId="F3"> <caption id="650E2FF432E1235F78C6962C76E711A8" doi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="145"> <paragraph id="026255F77C6678A7CB3A4CBCAF741D85" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 3.</emphasis> <emphasis id="35A66CB4C877C71249C357D8729094A0" bold="true" pageId="0" pageNumber="145">Figure 3.</emphasis>
Male palps of Male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis"> <taxonomicName id="CAD3834CEDDF22D8721D1B322464633F" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana kavirensis</emphasis> <emphasis id="7C46872DF48D5AF5264D1D4282C4029F" italics="true" pageId="0" pageNumber="145">Lachesana kavirensis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-E</emphasis> <emphasis id="ABE0F8F69321D7569651A884CBD048D1" bold="true" pageId="0" pageNumber="145">A-E</emphasis>
) and ) and
<taxonomicName lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus"> <taxonomicName id="B1F33A7F38F7F04C992B82D8C93F4F70" lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis italics="true" pageId="0" pageNumber="145">L. perseus</emphasis> <emphasis id="44FB43EDDEF00979BB20EF01CD1F1C53" italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">F-J</emphasis> <emphasis id="F954BC4F68BE8C099D96B9AD26FBF8FF" bold="true" pageId="0" pageNumber="145">F-J</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, B, G, F</emphasis> <emphasis id="95DA4D1687553A2F488B59F9DF6B687A" bold="true" pageId="0" pageNumber="145">A, B, G, F</emphasis>
retrolateral retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, H</emphasis> <emphasis id="4CCECB1D574E73FF823E6FBC3B6A25FB" bold="true" pageId="0" pageNumber="145">C, H</emphasis>
prolateral prolateral
<emphasis bold="true" pageId="0" pageNumber="145">D, I</emphasis> <emphasis id="956B9E31A0DA1D9E0965CB122ABC13FA" bold="true" pageId="0" pageNumber="145">D, I</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">E, J</emphasis> <emphasis id="4D303E1E3C0C3E5B00F39F9F0C7A159D" bold="true" pageId="0" pageNumber="145">E, J</emphasis>
dorsal. dorsal.
</paragraph> </paragraph>
</caption> </caption>
<caption doi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145" start="Figure 4" startId="F4"> <caption id="588C3E20AD42F9763CECA1A3226281BF" doi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="145"> <paragraph id="A6A7A4CD1F4CBAC06ED810D46474E752" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 4.</emphasis> <emphasis id="B9502AFC8FB772A4EDBB022CD71000F7" bold="true" pageId="0" pageNumber="145">Figure 4.</emphasis>
Male palps of Male palps of
<taxonomicName class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana dyachkovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dyachkovi"> <taxonomicName id="D27736380EE4C0AB3C76D7ADCCBE1B46" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana dyachkovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dyachkovi">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana dyachkovi</emphasis> <emphasis id="38362931D4499FD40FEBB63ACEF3A407" italics="true" pageId="0" pageNumber="145">Lachesana dyachkovi</emphasis>
</taxonomicName> </taxonomicName>
( (
<emphasis bold="true" pageId="0" pageNumber="145">A-C, F</emphasis> <emphasis id="3EDA2D76F7969BB3208532ED6D3BEDF6" bold="true" pageId="0" pageNumber="145">A-C, F</emphasis>
), ),
<taxonomicName lsidName="L. tarabaevi" pageId="0" pageNumber="145" rank="species" species="tarabaevi"> <taxonomicName id="9131979BBA2C374022B84869146E484C" lsidName="L. tarabaevi" pageId="0" pageNumber="145" rank="species" species="tarabaevi">
<emphasis italics="true" pageId="0" pageNumber="145">L. tarabaevi</emphasis> <emphasis id="A89A0CA90DE3ECEB716E3B3E6F607108" italics="true" pageId="0" pageNumber="145">L. tarabaevi</emphasis>
</taxonomicName> </taxonomicName>
( (
<emphasis bold="true" pageId="0" pageNumber="145">D, E, G</emphasis> <emphasis id="CE501965A218224D23C9750C650B984A" bold="true" pageId="0" pageNumber="145">D, E, G</emphasis>
), ),
<taxonomicName lsidName="L. blackwalli" pageId="0" pageNumber="145" rank="species" species="blackwalli"> <taxonomicName id="F338C72C67A30B7A5FD7ECECBD9B9375" lsidName="L. blackwalli" pageId="0" pageNumber="145" rank="species" species="blackwalli">
<emphasis italics="true" pageId="0" pageNumber="145">L. blackwalli</emphasis> <emphasis id="06B0530BE966B2CFB550DAA511AC782A" italics="true" pageId="0" pageNumber="145">L. blackwalli</emphasis>
</taxonomicName> </taxonomicName>
( (
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis> <emphasis id="50D3418A5467E5909F6A82CDEEC57093" bold="true" pageId="0" pageNumber="145">H</emphasis>
), ),
<taxonomicName lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis"> <taxonomicName id="7D186755AE93311598C33D1435AE6671" lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis> <emphasis id="7AA444B6117E9DD15CB13BF77D3BC90C" italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">I</emphasis> <emphasis id="9397F1F9DFAE66F852AF06FD73789FE2" bold="true" pageId="0" pageNumber="145">I</emphasis>
) and ) and
<taxonomicName lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus"> <taxonomicName id="7994865710A2D22E22E1F9A07C8A139E" lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis italics="true" pageId="0" pageNumber="145">L. perseus</emphasis> <emphasis id="75822E06EAE3B4BEC2A510D996BC28C2" italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">J</emphasis> <emphasis id="7C8EB94C80D63B9DA0018C035F7826D5" bold="true" pageId="0" pageNumber="145">J</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis> <emphasis id="16195BAD1A8BE33A1566FE8DA7753759" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, C, E</emphasis> <emphasis id="03C3EBADB84D3515EDD4719D059BC0ED" bold="true" pageId="0" pageNumber="145">B, C, E</emphasis>
dorsal, prolateral and retrolateral dorsal, prolateral and retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">F-J</emphasis> <emphasis id="30CA170CCCCC699B297BBE8F1139313C" bold="true" pageId="0" pageNumber="145">F-J</emphasis>
retrolateral tibial apophysis retrolateral tibial apophysis
<emphasis bold="true" pageId="0" pageNumber="145">A-G</emphasis> <emphasis id="07853A964F99B5622D0E38C073A6E3B6" bold="true" pageId="0" pageNumber="145">A-G</emphasis>
reproduced after reproduced after
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.28.4.08" author="Fomichev, AA" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="556 - 561" refId="B5" refString="Fomichev, AA, Marusik, YM, 2019. A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan. Arthropoda Selecta 28 (4): 556 - 561, DOI: https://doi.org/10.15298/arthsel.28.4.08" title="A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan." url="https://doi.org/10.15298/arthsel.28.4.08" volume="28" year="2019">Fomichev and Marusik (2019)</bibRefCitation> <bibRefCitation id="18D230A92D52BB7499C2BA592102C376" DOI="https://doi.org/10.15298/arthsel.28.4.08" author="Fomichev, AA" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="556 - 561" refId="B5" refString="Fomichev, AA, Marusik, YM, 2019. A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan. Arthropoda Selecta 28 (4): 556 - 561, DOI: https://doi.org/10.15298/arthsel.28.4.08" title="A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan." url="https://doi.org/10.15298/arthsel.28.4.08" volume="28" year="2019">Fomichev and Marusik (2019)</bibRefCitation>
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis> <emphasis id="AF2C07ACB48CFE9CC6CF62BC73FB8FCA" bold="true" pageId="0" pageNumber="145">H</emphasis>
illustrated after illustrated after
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.29.3.06" author="Oezkuetuek, RS" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="334 - 338" refId="B21" refString="Oezkuetuek, RS, Yagmur, EA, Guecel, S, Shafaie, S, Oezden, Oe, Kunt, KB, 2020. New records of the East Mediterranean Lachesana (Aranei: Zodariidae), with description of a new species. Arthropoda Selecta 29 (3): 334 - 338, DOI: https://doi.org/10.15298/arthsel.29.3.06" title="New records of the East Mediterranean Lachesana (Aranei: Zodariidae), with description of a new species." url="https://doi.org/10.15298/arthsel.29.3.06" volume="29" year="2020"> <bibRefCitation id="21537E01B4EB997BB5FA093770DC8530" DOI="https://doi.org/10.15298/arthsel.29.3.06" author="Oezkuetuek, RS" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="334 - 338" refId="B21" refString="Oezkuetuek, RS, Yagmur, EA, Guecel, S, Shafaie, S, Oezden, Oe, Kunt, KB, 2020. New records of the East Mediterranean Lachesana (Aranei: Zodariidae), with description of a new species. Arthropoda Selecta 29 (3): 334 - 338, DOI: https://doi.org/10.15298/arthsel.29.3.06" title="New records of the East Mediterranean Lachesana (Aranei: Zodariidae), with description of a new species." url="https://doi.org/10.15298/arthsel.29.3.06" volume="29" year="2020">
<normalizedToken originalValue="Özkütük">Oezkuetuek</normalizedToken> <normalizedToken id="47091E4EE797D80FD5B55CC4BDD1E17C" originalValue="Özkütük">Oezkuetuek</normalizedToken>
et al. (2020) et al. (2020)
</bibRefCitation> </bibRefCitation>
. Scale bars: 0.2 mm, unless stated otherwise. . Scale bars: 0.2 mm, unless stated otherwise.
</paragraph> </paragraph>
</caption> </caption>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="7DD39D0C6D50FB64360E66C80F2FC700" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="55D69154750B663AA75DC2002CB98808" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="0304E9E183716FE27F9A7861B7AB9AD7" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis> <emphasis id="C166BD5B5854D7F83BAF18BBB12FC48A" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Figs (holotype). Habitus as in Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1D, E</figureCitation> <figureCitation id="5B83CF800F2598DDE76BE26DE4C8E73F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1D, E</figureCitation>
, ,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live habitus of male Lachesana perseus sp. nov. Photo: A. Zamani." figureDoi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145">2</figureCitation> <figureCitation id="F440B607D5D6394BC41565849427DB94" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live habitus of male Lachesana perseus sp. nov. Photo: A. Zamani." figureDoi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145">2</figureCitation>
. Total length 10.50. Carapace 5.60 long, 2.60 wide at pars cephalica, 3.88 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.19, ALE: 0.19, PME: 0.16, PLE: 0.19, PME-PME: 0.18. Carapace, sternum, labium, chelicerae and maxillae light brown. Chelicera (Fig. . Total length 10.50. Carapace 5.60 long, 2.60 wide at pars cephalica, 3.88 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.19, ALE: 0.19, PME: 0.16, PLE: 0.19, PME-PME: 0.18. Carapace, sternum, labium, chelicerae and maxillae light brown. Chelicera (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1F</figureCitation> <figureCitation id="AF24280057BC0A00E04CB022E2589ACE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1F</figureCitation>
) with dense, black setae and one promarginal tooth; fangs curved. Legs the same color as carapace, without annulations and with numerous spines. Abdomen pale, dorsally with a longitudinal dark stripe and several large spots. Spinnerets grayish, uniform in color. Measurements of legs: I: 16.42 (4.69, 2.02, 3.61, 3.56, 2.54), II: 15.48 (4.22, 2.01, 3.19, 3.53, 2.53), III: 16.25 (4.46, 1.96, 2.51, 4.66, 2.66), IV: 19.39 (4.87, 2.32, 3.29, 5.66, 3.25). ) with dense, black setae and one promarginal tooth; fangs curved. Legs the same color as carapace, without annulations and with numerous spines. Abdomen pale, dorsally with a longitudinal dark stripe and several large spots. Spinnerets grayish, uniform in color. Measurements of legs: I: 16.42 (4.69, 2.02, 3.61, 3.56, 2.54), II: 15.48 (4.22, 2.01, 3.19, 3.53, 2.53), III: 16.25 (4.46, 1.96, 2.51, 4.66, 2.66), IV: 19.39 (4.87, 2.32, 3.29, 5.66, 3.25).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="9516173D2BFC2942A4C7EDB07C862D94" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3F-J</figureCitation> <figureCitation id="18B9EF623E50DC3572D19CB0D9A4AD52" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3F-J</figureCitation>
, ,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4J</figureCitation> <figureCitation id="0AEA5A9D78300C46729788416DC388B3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4J</figureCitation>
. Tibial apophysis more than 1.2 times longer than tibia, widest part over 2 times wider than stalk; cymbium lacking spines in proximal mesal part; bulb elongate, 1.3 times longer than wide; base of embolus large, 1/3 of the tegulum's length; embolus filamentous, lying in chute formed by conductor. . Tibial apophysis more than 1.2 times longer than tibia, widest part over 2 times wider than stalk; cymbium lacking spines in proximal mesal part; bulb elongate, 1.3 times longer than wide; base of embolus large, 1/3 of the tegulum's length; embolus filamentous, lying in chute formed by conductor.
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="A5183832E79F7D3898AB82AE58E125CB" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="8C73054740359E8ABB48768F6C76EEDD" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown. Unknown.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="5629D4B1DA20B13B4946C451DE16D0CC" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="222616A642345930D5BB0A41A0DA34B3" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="571ADD87054897BA34C3655C2A7A8834" pageId="0" pageNumber="145">
Known only from the type locality in Alborz Province, northern Iran (Fig. Known only from the type locality in Alborz Province, northern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation> <figureCitation id="67C10143A0799B0656F0F1845C3F7A5C" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

206
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<treatment ID-GBIF-Taxon="181753543" LSID="urn:lsid:zoobank.org:act:AFBD4904-D586-4FD5-A261-EC34D20EB469" httpUri="http://treatment.plazi.org/id/C61523CF4B07574DA528FA84A2573392" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="C61523CF4B07574DA528FA84A2573392" ID-GBIF-Taxon="181753543" LSID="urn:lsid:zoobank.org:act:AFBD4904-D586-4FD5-A261-EC34D20EB469" httpUri="http://treatment.plazi.org/id/C61523CF4B07574DA528FA84A2573392" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
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<paragraph pageId="0" pageNumber="145"> <paragraph id="A0C21398B47D8ADD8FFBDFFE0FD1BBF2" pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/AFBD4904-D586-4FD5-A261-EC34D20EB469" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium diara" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="diara" status="sp. nov.">Acanthinozodium diara</taxonomicName> <taxonomicName id="AC3012AFA5CFF65E1311C9896FA52472" ID-CoL="98YZ4" LSID="http://zoobank.org/AFBD4904-D586-4FD5-A261-EC34D20EB469" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium diara" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="diara" status="sp. nov.">Acanthinozodium diara</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="CE2C346A552F3E3D47D2A6F0832907B8" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18B</figureCitation> <figureCitation id="0AE384378257BA454D92A368C7BFD922" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18B</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20D-F</figureCitation> <figureCitation id="A0118C2A434C9328B338707E55D6E2CB" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20D-F</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">, 23D-F</figureCitation> <figureCitation id="BC248247A28BE2EAAC6CBBF671A7B474" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">, 23D-F</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation> <figureCitation id="14B3017DAC152629DAA25D3FD6A97C41" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="B0CE2196FEFD079EC83FFA08172025B2" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="A54AD4488F7219AC9550E07612376EFC" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="868E5E6C8A3B1B3F16ED2AF93854E429" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="888AF076A1F20B7BF3ACF22BE08579AE" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran: ♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Ilam Province</emphasis> <emphasis id="14586DDC272E57DEFC11019199C4162F" italics="true" pageId="0" pageNumber="145">Ilam Province</emphasis>
: Dizgaran, : Dizgaran,
<geoCoordinate degrees="33" direction="north" minutes="44" orientation="latitude" precision="925" value="33.733334">33°44'N</geoCoordinate> <geoCoordinate id="BC423E6AEE3B64AFBD8B413FF876E413" degrees="33" direction="north" minutes="44" orientation="latitude" precision="925" value="33.733334">33°44'N</geoCoordinate>
, ,
<geoCoordinate degrees="46" direction="east" minutes="59" orientation="longitude" precision="925" value="46.983334">46°59'E</geoCoordinate> <geoCoordinate id="28692CB0353B832047A50FBA2D05E496" degrees="46" direction="east" minutes="59" orientation="longitude" precision="925" value="46.983334">46°59'E</geoCoordinate>
, 16.5.1974 (A. Senglet). , 16.5.1974 (A. Senglet).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratype</emphasis> <emphasis id="26880B9F7B6FADE54BCE9738B98CD032" bold="true" italics="true" pageId="0" pageNumber="145">Paratype</emphasis>
♂ (MMUE), IRAN: ♂ (MMUE), IRAN:
<emphasis italics="true" pageId="0" pageNumber="145">Lorestan Province</emphasis> <emphasis id="81B3901D817D85D659921FFEF57CFC67" italics="true" pageId="0" pageNumber="145">Lorestan Province</emphasis>
: Dorood, 31.7.2011 (S. Zaruni). : Dorood, 31.7.2011 (S. Zaruni).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="B19243988F1CC56C7286A51D85428265" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="4B23365B69A24AB7561E99C58C59B0F8" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="9FD6169358D0C6E859C79178355AA3BE" pageId="0" pageNumber="145">
The specific epithet is a Persian feminine name meaning The specific epithet is a Persian feminine name meaning
<normalizedToken originalValue="“motherland”">&quot;motherland&quot;</normalizedToken> <normalizedToken id="C1F4439045A81D57E5652D898D30EC29" originalValue="“motherland”">&quot;motherland&quot;</normalizedToken>
. Noun in apposition. . Noun in apposition.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="16CE91F586ED480AE8501AEFD29C206E" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="0E6F10DDDDF71ACA2659187529DD2245" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="2E6992972796AC9E41846D9A4177B800" pageId="0" pageNumber="145">
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium diara" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="diara"> <taxonomicName id="00918FF2A5AA338E474537A78D48A237" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium diara" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium diara</emphasis> <emphasis id="CF1B8542DE497B1E2500D1CC3C5D0B34" italics="true" pageId="0" pageNumber="145">Acanthinozodium diara</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. is very similar to sp. nov. is very similar to
<taxonomicName lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa"> <taxonomicName id="A2D7F337B2859B592B07E591D836E23F" lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis italics="true" pageId="0" pageNumber="145">A. masa</emphasis> <emphasis id="7A9F52CFBDFD50DB85B5FFC017F9D901" italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. by the shape of the RTA, the ventral tibial apophysis and the curvature of the embolus but differs by the shape of the median apophysis (cf. Fig. sp. nov. by the shape of the RTA, the ventral tibial apophysis and the curvature of the embolus but differs by the shape of the median apophysis (cf. Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20D</figureCitation> <figureCitation id="F82B0365F2A9294BA0F71AAE29390B5D" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20D</figureCitation>
and and
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20J</figureCitation> <figureCitation id="10E1596C16FF2681A3DCF34F8156F9F1" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20J</figureCitation>
) and the conductor having a small retrolateral indentation apically ( ) and the conductor having a small retrolateral indentation apically (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="1522E2BEFBDA9A07972D3556D0765CF4" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
without indentation). without indentation).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="1652BC0AFC3CEAF48E44D0D530C798F0" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="1D930A5301CCEB567EE6582003EB1B1F" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="4AD75DB6CC54AF8578D8B4CAD37E0D08" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis> <emphasis id="EE56903B074FFD7E0A718DC2DDE00E17" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig. (holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18B</figureCitation> <figureCitation id="57FE2AF270D676C74F9099E63D427BE0" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18B</figureCitation>
. Total length 4.70. Carapace 2.18 long, 0.92 wide at pars cephalica, 1.52 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.19, ALE: 0.12, PME: 0.10, PLE: 0.12, PME-PME: 0.25. Carapace dark brown, with irregular dark patches and lines. Sternum, labium and maxillae light brown. Chelicera dark brown, with one retromarginal tooth. Legs light brown, without annulations. Abdomen black, without any pattern; slightly lighter ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 9.49 (2.38, 0.71, 2.16, 2.76, 1.48), II: 8.76 (2.10, 0.78, 1.98, 2.50, 1.40), III: 8.36 (2.11, 0.70, 1.69, 2.73, 1.13), IV: 11.42 (3.00, 0.81, 2.60, 3.59, 1.42). . Total length 4.70. Carapace 2.18 long, 0.92 wide at pars cephalica, 1.52 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.19, ALE: 0.12, PME: 0.10, PLE: 0.12, PME-PME: 0.25. Carapace dark brown, with irregular dark patches and lines. Sternum, labium and maxillae light brown. Chelicera dark brown, with one retromarginal tooth. Legs light brown, without annulations. Abdomen black, without any pattern; slightly lighter ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 9.49 (2.38, 0.71, 2.16, 2.76, 1.48), II: 8.76 (2.10, 0.78, 1.98, 2.50, 1.40), III: 8.36 (2.11, 0.70, 1.69, 2.73, 1.13), IV: 11.42 (3.00, 0.81, 2.60, 3.59, 1.42).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="15F1AFC2B46CADA0B98D729EAA0B83E1" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20D-F</figureCitation> <figureCitation id="C03CC2A2A382EC8817F89A4A1512F3EF" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20D-F</figureCitation>
, ,
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">23D-F</figureCitation> <figureCitation id="3D49112824A69546CB4A52A5B94580AD" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">23D-F</figureCitation>
. RTA long and conical, with a small projection apically (Fig. . RTA long and conical, with a small projection apically (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20E</figureCitation> <figureCitation id="9FC303AD89C3A31453DA8F5DCFB503AC" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20E</figureCitation>
); tegulum with posterior process; ventral apophysis small and conical; median apophysis almost as long as wide, with outgrowths on all four sides, posterior one largest; embolus broad basally, originating at about the 7 ); tegulum with posterior process; ventral apophysis small and conical; median apophysis almost as long as wide, with outgrowths on all four sides, posterior one largest; embolus broad basally, originating at about the 7
<normalizedToken originalValue="o’clock">o'clock</normalizedToken> <normalizedToken id="FC1A120D7414BEC54918C90BA3378603" originalValue="o’clock">o'clock</normalizedToken>
position; embolus proper thin and steadily curving. position; embolus proper thin and steadily curving.
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145" start="Figure 20" startId="F20"> <caption id="976E7D1B5A5660979B38B882A3B2BF24" doi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145" start="Figure 20" startId="F20">
<paragraph pageId="0" pageNumber="145"> <paragraph id="5BE4B760FCE50992A0B7C980C42B051B" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 20.</emphasis> <emphasis id="C4C01CCE1E1F8B176D67FDFE81BBAB5E" bold="true" pageId="0" pageNumber="145">Figure 20.</emphasis>
Male palps of Male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita"> <taxonomicName id="58C3481789AD175352CCD1C356DD1C4D" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium armita</emphasis> <emphasis id="8EC769C0022BCDA6EA99A17BBBCB94F1" italics="true" pageId="0" pageNumber="145">Acanthinozodium armita</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis> <emphasis id="91A02158D9F44BF3764C8CEB74460053" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
), ),
<taxonomicName lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara"> <taxonomicName id="A6BB2AE41768A126C3ABEF7B83A6601E" lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">A. diara</emphasis> <emphasis id="28DC94BF852692D24AAC57D7751F381C" italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis> <emphasis id="04ACC5CE025FA9C2E28712756FC32478" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
), ),
<taxonomicName lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana"> <taxonomicName id="F68280C70455E20A16F98C170E92045F" lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana">
<emphasis italics="true" pageId="0" pageNumber="145">A. kiana</emphasis> <emphasis id="070FB45F2C9FED9F4CC75E69934EEBB2" italics="true" pageId="0" pageNumber="145">A. kiana</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">G, H</emphasis> <emphasis id="0B97FCA3048452981F08598FEEFEBCF6" bold="true" pageId="0" pageNumber="145">G, H</emphasis>
) and ) and
<taxonomicName lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa"> <taxonomicName id="D0F2263C72DF88D20A2A1CDC6892C064" lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis italics="true" pageId="0" pageNumber="145">A. masa</emphasis> <emphasis id="D2F31508A0DD688C78825E8D2D83ABC4" italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">I-K</emphasis> <emphasis id="C68DCF14C55440E74F71B93765FCDB1C" bold="true" pageId="0" pageNumber="145">I-K</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, D, G, J</emphasis> <emphasis id="19C20669F8B74299E851D3549EE1E9F4" bold="true" pageId="0" pageNumber="145">A, D, G, J</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, E, H, I</emphasis> <emphasis id="67CCF3E3BE2D69048D6A4219E1575631" bold="true" pageId="0" pageNumber="145">B, E, H, I</emphasis>
retrolateral retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, F, K</emphasis> <emphasis id="ECDC123DBA045D60A8B669F2C773645B" bold="true" pageId="0" pageNumber="145">C, F, K</emphasis>
dorsal. Scale bars: 0.2 mm, unless stated otherwise. dorsal. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph> </paragraph>
</caption> </caption>
<paragraph pageId="0" pageNumber="145"> <paragraph id="F64B20D5B49E90451A35AF6B9A49CA4F" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="9AFAF06F6D563902946DC985AA8C7212" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown. Unknown.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="55F8A6E4E68DBC8219230A4E1DD30660" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="C9A07C783C998ABEC624D99754E70634" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="BEBCD9D6C8BE9FFF834285D872CADE88" pageId="0" pageNumber="145">
Known from the listed localities in Ilam and Lorestan provinces, western Iran (Fig. Known from the listed localities in Ilam and Lorestan provinces, western Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation> <figureCitation id="A84BE9C8A8627D736BD13F861759540D" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

286
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2021" docId="D5FCAC79232E53A39DAF2BF008656B1E" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Acanthinozodium atrisa Zamani &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="7BBBC164-8239-4F57-9F44-4F5DE93F8C24" docUuidSource="ZooBank" docVersion="4" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1668150222111" updateUser="ExternalLinkService"> <document ID-CLB-Dataset="7157" ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2021" docId="D5FCAC79232E53A39DAF2BF008656B1E" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Acanthinozodium atrisa Zamani &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="7BBBC164-8239-4F57-9F44-4F5DE93F8C24" docUuidSource="ZooBank" docVersion="6" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1732670661048" updateUser="ExternalLinkService">
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title> <mods:title id="29DF60A3DEF71544F2449213C4EF96EF">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:namePart>Zamani, Alireza</mods:namePart> <mods:namePart id="A4720F5F02CACD0048BFC2EFB76A81A0">Zamani, Alireza</mods:namePart>
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<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation> <mods:affiliation id="93429F24E2E04B0BC23CC2D4129FA2A3">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation> <mods:affiliation id="523D491B9C78AC73E30458045E093A7F">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<treatment ID-GBIF-Taxon="181753544" LSID="urn:lsid:zoobank.org:act:7BBBC164-8239-4F57-9F44-4F5DE93F8C24" httpUri="http://treatment.plazi.org/id/D5FCAC79232E53A39DAF2BF008656B1E" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="D5FCAC79232E53A39DAF2BF008656B1E" ID-GBIF-Taxon="181753544" LSID="urn:lsid:zoobank.org:act:7BBBC164-8239-4F57-9F44-4F5DE93F8C24" httpUri="http://treatment.plazi.org/id/D5FCAC79232E53A39DAF2BF008656B1E" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="DA58599AD2FC7D559BE324FD970941D1" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="59001C7E25FB05C3C6F100F950DF7D98" pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/7BBBC164-8239-4F57-9F44-4F5DE93F8C24" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa" status="sp. nov.">Acanthinozodium atrisa</taxonomicName> <taxonomicName id="FC477103F54CF71F4F21B9645C748A90" ID-CoL="98YYW" LSID="http://zoobank.org/7BBBC164-8239-4F57-9F44-4F5DE93F8C24" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa" status="sp. nov.">Acanthinozodium atrisa</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="6CAE441758F079C3A6E6C54DF9CCD0D1" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11A, B</figureCitation> <figureCitation id="855A7F30DD367648F17313DC41783521" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">Figs 11A, B</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">, 12A-C</figureCitation> <figureCitation id="C978D18B4BF3EA8A409A7CB007BD1A32" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">, 12A-C</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">, 14A-C</figureCitation> <figureCitation id="A0D684CF55DA0BF26EC00D79FA97CB43" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">, 14A-C</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Epigyne of Acanthinozodium atrisa sp. nov. A posteroventral B posterior C, D ventral. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure16" httpUri="https://binary.pensoft.net/fig/536893" pageId="0" pageNumber="145">, 16A-D</figureCitation> <figureCitation id="A06518477021E8E2540878ECC15C31B8" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Epigyne of Acanthinozodium atrisa sp. nov. A posteroventral B posterior C, D ventral. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure16" httpUri="https://binary.pensoft.net/fig/536893" pageId="0" pageNumber="145">, 16A-D</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation> <figureCitation id="CEC31E19211931C32FAC31202401F865" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
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<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="811E99740F897D905397A1037F4CB5BA" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="37B694769B0327A17CC7F206D11DD7B9" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="124F2C6A32EF599D1F066CB41F394C82" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran: ♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Tehran Province</emphasis> <emphasis id="AA9E731DE8F1B9D6A934EC2601C17069" italics="true" pageId="0" pageNumber="145">Tehran Province</emphasis>
: Jamshidieh Mts., : Jamshidieh Mts.,
<geoCoordinate degrees="35" direction="north" minutes="49" orientation="latitude" precision="925" value="35.816666">35°49'N</geoCoordinate> <geoCoordinate id="C2A4A6416863210704824F920903ED83" degrees="35" direction="north" minutes="49" orientation="latitude" precision="925" value="35.816666">35°49'N</geoCoordinate>
, ,
<geoCoordinate degrees="51" direction="east" minutes="27" orientation="longitude" precision="925" value="51.45">51°27'E</geoCoordinate> <geoCoordinate id="20327AABEDED0D415476500C10E67A77" degrees="51" direction="east" minutes="27" orientation="longitude" precision="925" value="51.45">51°27'E</geoCoordinate>
, 05.2015 (A. Zamani). , 05.2015 (A. Zamani).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis> <emphasis id="C0C8EF3B490CC62341EF0E668C9EFD55" bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
: 1♂2♀ (MHNG), same data as holotype; 5♂3♀ (MMUE), Latian Dam, : 1♂2♀ (MHNG), same data as holotype; 5♂3♀ (MMUE), Latian Dam,
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, ,
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, 6-19.06.2000 (Y.M. Marusik); 1♂30♀1j. (MMUE), Plant Protection Institute, , 6-19.06.2000 (Y.M. Marusik); 1♂30♀1j. (MMUE), Plant Protection Institute,
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, ,
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, 7-22.06.2000 (Y.M. Marusik); 2♀ (MMUE), 5 km north of Tehran, Tochal Mts., , 7-22.06.2000 (Y.M. Marusik); 2♀ (MMUE), 5 km north of Tehran, Tochal Mts.,
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, ,
<geoCoordinate degrees="51" direction="east" minutes="20" orientation="longitude" precision="925" value="51.333332">51°20'E</geoCoordinate> <geoCoordinate id="15BE8E3739F35BD16F2E398D48B40BA3" degrees="51" direction="east" minutes="20" orientation="longitude" precision="925" value="51.333332">51°20'E</geoCoordinate>
, 2000-2900 m, 16.06.2000 (Y.M. Marusik); 9♀1j. (ZMMU), 80 km east of Tehran, Damavand area, Aroo, , 2000-2900 m, 16.06.2000 (Y.M. Marusik); 9♀1j. (ZMMU), 80 km east of Tehran, Damavand area, Aroo,
<geoCoordinate degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate> <geoCoordinate id="8CD06789976DF9FA9A7C5800E185F034" degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate>
, ,
<geoCoordinate degrees="52" direction="east" minutes="27" orientation="longitude" precision="925" value="52.45">52°27'E</geoCoordinate> <geoCoordinate id="457E98F3E667FC7FEDCB6733DCB76B46" degrees="52" direction="east" minutes="27" orientation="longitude" precision="925" value="52.45">52°27'E</geoCoordinate>
, 15.06.2000 (Y.M. Marusik &amp; F. Mozaffarian); , 15.06.2000 (Y.M. Marusik &amp; F. Mozaffarian);
<emphasis italics="true" pageId="0" pageNumber="145">Qazvin Province</emphasis> <emphasis id="48DA8893C675FBAC2A79DA2B1BF860D7" italics="true" pageId="0" pageNumber="145">Qazvin Province</emphasis>
: 1♂2♀ (MHNG), Agha Baba, : 1♂2♀ (MHNG), Agha Baba,
<geoCoordinate degrees="36" direction="north" minutes="19" orientation="latitude" precision="925" value="36.316666">36°19'N</geoCoordinate> <geoCoordinate id="1B15D7693DF32DC44992196947CD4DD8" degrees="36" direction="north" minutes="19" orientation="latitude" precision="925" value="36.316666">36°19'N</geoCoordinate>
, ,
<geoCoordinate degrees="49" direction="east" minutes="49" orientation="longitude" precision="925" value="49.816666">49°49'E</geoCoordinate> <geoCoordinate id="98C054FEC141A40B5D470395C6E65253" degrees="49" direction="east" minutes="49" orientation="longitude" precision="925" value="49.816666">49°49'E</geoCoordinate>
, 06.07.1974 (A. Senglet); 1♂1♀ (MHNG), Tarazan, Lowshan, , 06.07.1974 (A. Senglet); 1♂1♀ (MHNG), Tarazan, Lowshan,
<geoCoordinate degrees="36" direction="north" minutes="28" orientation="latitude" precision="925" value="36.466667">36°28'N</geoCoordinate> <geoCoordinate id="03F73198B34AF28F1FA12D57F0F57D65" degrees="36" direction="north" minutes="28" orientation="latitude" precision="925" value="36.466667">36°28'N</geoCoordinate>
, ,
<geoCoordinate degrees="49" direction="east" minutes="31" orientation="longitude" precision="925" value="49.516666">49°31'E</geoCoordinate> <geoCoordinate id="43D80C2E627B9AFB6439513B8603E5F1" degrees="49" direction="east" minutes="31" orientation="longitude" precision="925" value="49.516666">49°31'E</geoCoordinate>
, 08.08.1974 (A. Senglet). , 08.08.1974 (A. Senglet).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="001956B52441282BB7FE4004592C7A52" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="7FA4ACFE34F2DE09B38636A323C52FB5" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet is a Persian feminine name meaning &quot;queen of fire&quot;. Noun in apposition.</paragraph> <paragraph id="837F99CB067C887F1408FDCFF1247C3C" pageId="0" pageNumber="145">The specific epithet is a Persian feminine name meaning &quot;queen of fire&quot;. Noun in apposition.</paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="959D44240DB1ED47BBD1689463115614" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="76AA9E641E860D24E332D71CA0CC923B" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="99B4F89220FB9A2D6957F9078FBCAAD1" pageId="0" pageNumber="145">
The new species has an abdominal pattern similar to that of The new species has an abdominal pattern similar to that of
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis"> <taxonomicName id="3E31F2BD1CCEF3D7AD2E6093E8449CB3" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis> <emphasis id="1FA0E65A559A69A4AFB0106111EEA6F3" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. (dark abdomen with posterior median white stripe; less distinct in males) but differs by having a dark pars cephalica ( sp. nov. (dark abdomen with posterior median white stripe; less distinct in males) but differs by having a dark pars cephalica (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="70E5C94CB2BFA5CC0242AFF599113677" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
pale). The male palps of the two species differ by the RTA being thin and more than twice as long as the tibia in pale). The male palps of the two species differ by the RTA being thin and more than twice as long as the tibia in
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa"> <taxonomicName id="AD4729341B155ECA006FE0F41E6D9B6D" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis> <emphasis id="9F5AE418E8C6C1B8CA38260A4DAB230E" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="A7377E2676461395CE2A50814A22A2DB" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
broad and as long as the tibia). Judging by the shape of the male palp, broad and as long as the tibia). Judging by the shape of the male palp,
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa"> <taxonomicName id="915CCCFAE5E3C5724D43BFAA8199E4DD" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis> <emphasis id="208A6ED24F102C5772872AA3D2C9EE76" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. is probably closely related to sp. nov. is probably closely related to
<taxonomicName lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani"> <taxonomicName id="485757E298B005D8A454E67970D6C77E" lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis italics="true" pageId="0" pageNumber="145">A. sorani</emphasis> <emphasis id="545145733BE8415AEE23CA75A4C90995" italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
</taxonomicName> </taxonomicName>
sp. nov., as both have a long RTA, a similarly shaped median apophysis, a posterior tegular process and an embolus with an anterior process. The two species differ by the shape of the embolic process ( sp. nov., as both have a long RTA, a similarly shaped median apophysis, a posterior tegular process and an embolus with an anterior process. The two species differ by the shape of the embolic process (
<emphasis italics="true" pageId="0" pageNumber="145">Ep</emphasis> <emphasis id="4B29EB2B7D0CFC5F6FCFED221CC9074C" italics="true" pageId="0" pageNumber="145">Ep</emphasis>
), wider than the embolus and gently bent at the tip in ), wider than the embolus and gently bent at the tip in
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa"> <taxonomicName id="C480775EC359673EAD485FDDC6AFD858" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis> <emphasis id="69AD8C937F06C2F46469459DAF06EBC3" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="7CA67ADC502462FA711BA6086DD0CE77" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
spine-like). The epigyne of spine-like). The epigyne of
<taxonomicName lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa"> <taxonomicName id="A8DE1FEBA98F1B267A937FCCCBCCC500" lsidName="A. atrisa" pageId="0" pageNumber="145" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis> <emphasis id="BC243AFFF0636DED9693C6CF29C1D0C9" italics="true" pageId="0" pageNumber="145">A. atrisa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. is most similar to that of sp. nov. is most similar to that of
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis"> <taxonomicName id="887F94A4F8CA54857CD9130DA2B9E03C" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis> <emphasis id="AABA7D2457B8463550C6EE754FF83FFD" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. in having a small anterior hood and a thin furrow between the lateral lobes but differs by the furrow being shorter than the bursa copulatrix ( sp. nov. in having a small anterior hood and a thin furrow between the lateral lobes but differs by the furrow being shorter than the bursa copulatrix (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="C31D84EBB4ED7829AD7D88B1F87CEF81" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
the same length as bursa). the same length as bursa).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="0CB2A26A3BE45DD9D506F56187810305" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="1D97C22490EB7A991DECEDCC0875113F" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="1D124B8296807B3F82D11EBCD7372B58" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis> <emphasis id="AF1E32F09914FEE87FB892A1ECE73304" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig. (holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11A</figureCitation> <figureCitation id="75BE483F4CA7038B45D017468BBC7BEA" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11A</figureCitation>
. Total length 1.78. Carapace 0.81 long, 0.39 wide at pars cephalica, 0.62 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.07, ALE: 0.06, PME: 0.05, PLE: 0.05, PME-PME: 0.10. Carapace, sternum, labium, chelicerae and maxillae yellowish; carapace with irregular dark brown patterns. Chelicera with retromarginal tooth. Legs yellowish, without annulations. Abdomen dorsally black with longitudinal pale median band and grayish ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 2.39 (0.61, 0.25, 0.52, 0.53, 0.48), II: 2.07 (0.54, 0.26, 0.45, 0.46, 0.36), III: 2.06 (0.57, 0.25, 0.37, 0.48, 0.39), IV: 2.86 (0.69, 0.29, 0.64, 0.77, 0.47). . Total length 1.78. Carapace 0.81 long, 0.39 wide at pars cephalica, 0.62 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.07, ALE: 0.06, PME: 0.05, PLE: 0.05, PME-PME: 0.10. Carapace, sternum, labium, chelicerae and maxillae yellowish; carapace with irregular dark brown patterns. Chelicera with retromarginal tooth. Legs yellowish, without annulations. Abdomen dorsally black with longitudinal pale median band and grayish ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 2.39 (0.61, 0.25, 0.52, 0.53, 0.48), II: 2.07 (0.54, 0.26, 0.45, 0.46, 0.36), III: 2.06 (0.57, 0.25, 0.37, 0.48, 0.39), IV: 2.86 (0.69, 0.29, 0.64, 0.77, 0.47).
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145" start="Figure 11" startId="F11"> <caption id="B39431BB8A4CFD52E84007F25A661BB7" doi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="145"> <paragraph id="A289F7BC27B9CF1F33E38A205E1A9467" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 11.</emphasis> <emphasis id="E43C30826F069826423704E8C02E6DF5" bold="true" pageId="0" pageNumber="145">Figure 11.</emphasis>
Habitus of Habitus of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa"> <taxonomicName id="FF514550E611F5000B6030B3276EFBF9" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis> <emphasis id="AF9220CD5BC1E7DC928A227FAEA0D88B" italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A, B</emphasis> <emphasis id="48B826D5014E89A7ABC1D3A72C2E5680" bold="true" pageId="0" pageNumber="145">A, B</emphasis>
) )
<taxonomicName lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha"> <taxonomicName id="E667D6034B409194786C1A1611D9C933" lsidName="A. niusha" pageId="0" pageNumber="145" rank="species" species="niusha">
<emphasis italics="true" pageId="0" pageNumber="145">A. niusha</emphasis> <emphasis id="35C01EE83AF5438F9D9C4644C7D9394C" italics="true" pageId="0" pageNumber="145">A. niusha</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">C-E</emphasis> <emphasis id="4905A3995EC64ED1CB50B64A1C4424F0" bold="true" pageId="0" pageNumber="145">C-E</emphasis>
), ),
<taxonomicName lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis"> <taxonomicName id="BFAD611CD0FDA3FC3C6E94225253EC95" lsidName="A. parysatis" pageId="0" pageNumber="145" rank="species" species="parysatis">
<emphasis italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis> <emphasis id="23412446932EFE6ABD8D13F6284C3BE7" italics="true" pageId="0" pageNumber="145">A. parysatis</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">F, G</emphasis> <emphasis id="CBAB16C23EA6A61D6A902E334F8354A1" bold="true" pageId="0" pageNumber="145">F, G</emphasis>
) and ) and
<taxonomicName lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani"> <taxonomicName id="376B701740644B9AC2BDF88056F44E3F" lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis italics="true" pageId="0" pageNumber="145">A. sorani</emphasis> <emphasis id="473C93AD0C6EFA4CFCC8D8488BC9E022" italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis> <emphasis id="8607E09476CB12E223AFDE51174303F1" bold="true" pageId="0" pageNumber="145">H</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, C, F, H</emphasis> <emphasis id="6FF08D0B602AF8A7ACD50F5E81805F95" bold="true" pageId="0" pageNumber="145">A, C, F, H</emphasis>
males, dorsal males, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis> <emphasis id="CA16909374FB2DE890E07B9D1929326D" bold="true" pageId="0" pageNumber="145">D</emphasis>
male, ventral male, ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, G, E</emphasis> <emphasis id="4E73A2FBC70B0D15A6FEB98B4F7DBADA" bold="true" pageId="0" pageNumber="145">B, G, E</emphasis>
females, dorsal. Scale bars: 0.2 mm, unless stated otherwise. females, dorsal. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph> </paragraph>
</caption> </caption>
<paragraph pageId="0" pageNumber="145"> <paragraph id="C2E3BC64C7B5B88157C3B2D9410F8854" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">12A-C</figureCitation> <figureCitation id="7A014041EF2A2D28137E3ADD03295A88" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male palps of Acanthinozodium atrisa sp. nov. (A-C) and A. sorani sp. nov. (D-F) A, D ventral B, C, F retrolateral E dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145">12A-C</figureCitation>
, ,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14A-C</figureCitation> <figureCitation id="0EF9E8FFC3A37CC00D87C10EF89AC5FB" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM images of the male palps of Acanthinozodium atrisa sp. nov. (A-C), A. sorani sp. nov. (D-F), A. niusha sp. nov. (G, H) and A. dorsa sp. nov. (I) A, C, D, F, I ventral B, E retrolateral G, H cymbial groove, dorsal. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Ep - embolic process, Ma - median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure14" httpUri="https://binary.pensoft.net/fig/536891" pageId="0" pageNumber="145">14A-C</figureCitation>
. RTA ( . RTA (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis> <emphasis id="FAA8EF256A3F761844665353EA48135A" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
) long (almost as long as bulb) and thin (7 times longer than wide), slightly bent; tegulum with posterior process, sperm duct almost straight retrolaterally, and gently bent along prolateral side; median apophysis wider than long; embolus ( ) long (almost as long as bulb) and thin (7 times longer than wide), slightly bent; tegulum with posterior process, sperm duct almost straight retrolaterally, and gently bent along prolateral side; median apophysis wider than long; embolus (
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis> <emphasis id="6D6CCAC09A98210E3FDB0C45033E90CF" italics="true" pageId="0" pageNumber="145">Em</emphasis>
) broad basally, originating at about the 7 ) broad basally, originating at about the 7
<normalizedToken originalValue="o’clock">o'clock</normalizedToken> <normalizedToken id="22EF041EB99C0BF09088D6891C139927" originalValue="o’clock">o'clock</normalizedToken>
position, bifurcated terminally, with broad anterior process; embolus proper thin and straight, with small tubercle. position, bifurcated terminally, with broad anterior process; embolus proper thin and straight, with small tubercle.
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145" start="Figure 12" startId="F12"> <caption id="DE27AF243D7D0361839AFFAF6CE5559D" doi="10.3897/zookeys.1035.65767.figure12" httpUri="https://binary.pensoft.net/fig/536889" pageId="0" pageNumber="145" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="145"> <paragraph id="0E651A63255E198B3738B9A65B1356FF" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 12.</emphasis> <emphasis id="800F79475A9D06F75ADC7DF7B43955A8" bold="true" pageId="0" pageNumber="145">Figure 12.</emphasis>
Male palps of Male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa"> <taxonomicName id="D55A075781F4F62909F0AC399B4EEF05" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium atrisa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="atrisa">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis> <emphasis id="9ABB158DDAFA850ED15A8313AC8981B7" italics="true" pageId="0" pageNumber="145">Acanthinozodium atrisa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis> <emphasis id="6722C5241DC5A02AD38F5B2C4441C6DA" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and ) and
<taxonomicName lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani"> <taxonomicName id="66166BA8BEAA74A3F1F452DF545253F8" lsidName="A. sorani" pageId="0" pageNumber="145" rank="species" species="sorani">
<emphasis italics="true" pageId="0" pageNumber="145">A. sorani</emphasis> <emphasis id="85891AA643D85205039AF67CA00277AB" italics="true" pageId="0" pageNumber="145">A. sorani</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis> <emphasis id="08067CF6EF3AEA924BCF5253239A52E9" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis> <emphasis id="044DDE9011F942812EB56392169174A1" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, C, F</emphasis> <emphasis id="0B8C8BA940D7AED07A3CC25F206414C1" bold="true" pageId="0" pageNumber="145">B, C, F</emphasis>
retrolateral retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis> <emphasis id="FFF2647DDFDCDC18E80146F379A71915" bold="true" pageId="0" pageNumber="145">E</emphasis>
dorsal. Scale bars: 0.2 mm. dorsal. Scale bars: 0.2 mm.
</paragraph> </paragraph>
</caption> </caption>
<paragraph pageId="0" pageNumber="145"> <paragraph id="2148FA7A601E8C70B860132D2FEEDA84" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="EF314CA62F08D3A6688B5C6367B48683" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig. Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11B</figureCitation> <figureCitation id="EFBAA1E60593AE5F690725E954CDCA63" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Acanthinozodium atrisa sp. nov. (A, B) A. niusha sp. nov. (C-E), A. parysatis sp. nov. (F, G) and A. sorani sp. nov. (H) A, C, F, H males, dorsal D male, ventral B, G, E females, dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure11" httpUri="https://binary.pensoft.net/fig/536888" pageId="0" pageNumber="145">11B</figureCitation>
. Total length 2.38. Carapace 0.87 long, 0.39 wide at pars cephalica, 0.59 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.07, ALE: 0.06, PME: 0.05, PLE: 0.05, PME-PME: 0.10. Coloration as in male, with paler carapace and less prominent abdominal median stripe. Measurements of legs: I: 2.36 (0.71, 0.24, 0.45, 0.51, 0.45), II: 2.18 (0.56, 0.28, 0.42, 0.48, 0.44), III: 2.16 (0.54, 0.27, 0.42, 0.51, 0.42), Fe IV: 0.76, rest of the segments missing. . Total length 2.38. Carapace 0.87 long, 0.39 wide at pars cephalica, 0.59 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.07, ALE: 0.06, PME: 0.05, PLE: 0.05, PME-PME: 0.10. Coloration as in male, with paler carapace and less prominent abdominal median stripe. Measurements of legs: I: 2.36 (0.71, 0.24, 0.45, 0.51, 0.45), II: 2.18 (0.56, 0.28, 0.42, 0.48, 0.44), III: 2.16 (0.54, 0.27, 0.42, 0.51, 0.42), Fe IV: 0.76, rest of the segments missing.
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="1EA2C4F6F180467D26963D739A1CBCC5" pageId="0" pageNumber="145">
Epigyne as in Fig. Epigyne as in Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Epigyne of Acanthinozodium atrisa sp. nov. A posteroventral B posterior C, D ventral. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure16" httpUri="https://binary.pensoft.net/fig/536893" pageId="0" pageNumber="145">16A-D</figureCitation> <figureCitation id="1F21B260E9E07D3520F81A986C6A63E9" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Epigyne of Acanthinozodium atrisa sp. nov. A posteroventral B posterior C, D ventral. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure16" httpUri="https://binary.pensoft.net/fig/536893" pageId="0" pageNumber="145">16A-D</figureCitation>
. Fovea triangular, and small anterior hood present; bursae subtriangular, wider anteriorly, separated by less than one of their radii; receptacles smaller than bursae, ovoid, separated by more than 3 diameters. . Fovea triangular, and small anterior hood present; bursae subtriangular, wider anteriorly, separated by less than one of their radii; receptacles smaller than bursae, ovoid, separated by more than 3 diameters.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="0721DCD3DF6E21A81C0B5B0CA79AA9B0" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="C0298897008790FCB13B756900880874" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="876C630757CE5B2CA843E488A162BE20" pageId="0" pageNumber="145">
Known only from the listed localities in Tehran and Qazvin provinces, northern Iran (Fig. Known only from the listed localities in Tehran and Qazvin provinces, northern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation> <figureCitation id="0BBC674903C4EE2787AE7DD0C93DDD18" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

106
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title> <mods:title id="20B4A1E9604BD239393401CBA45AF229">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<treatment ID-GBIF-Taxon="181753539" LSID="urn:lsid:plazi:treatment:E10481F9F28959A69088F27E63C3A2FC" httpUri="http://treatment.plazi.org/id/E10481F9F28959A69088F27E63C3A2FC" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="E10481F9F28959A69088F27E63C3A2FC" ID-GBIF-Taxon="181753539" LSID="urn:lsid:plazi:treatment:E10481F9F28959A69088F27E63C3A2FC" httpUri="http://treatment.plazi.org/id/E10481F9F28959A69088F27E63C3A2FC" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Lachesaninae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="589C14BBA8A2E6951EFC2154C5856C6A" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="8A6FD4E4A2F9B77A9DF20E08F7335CAC" pageId="0" pageNumber="145">
Genus Genus
<taxonomicName LSID="E10481F9-F289-59A6-9088-F27E63C3A2FC" authority="Strand, 1932" authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Lachesana Strand, 1932</taxonomicName> <taxonomicName id="FD25D44EA4B250C1A1AC8A2A8DF18F2D" ID-CoL="59T5" LSID="E10481F9-F289-59A6-9088-F27E63C3A2FC" authority="Strand, 1932" authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Lachesana Strand, 1932</taxonomicName>
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<subSubSection pageId="0" pageNumber="145" type="type species"> <subSubSection id="1A4E8896369E39FA1323EF794FD7AAAA" pageId="0" pageNumber="145" type="type species">
<paragraph pageId="0" pageNumber="145">Type species.</paragraph> <paragraph id="EEC6A09BF7EC03C6F68DC51FDA471EC7" pageId="0" pageNumber="145">Type species.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="8F0B21C36EB26A99F4B70150F77EE429" pageId="0" pageNumber="145">
<taxonomicName class="Arachnida" family="Zodariidae" genus="Lachesis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lachesis perversa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="perversa"> <taxonomicName id="D113CA1DFCCBB05F420B7015EFF9DD47" class="Arachnida" family="Zodariidae" genus="Lachesis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lachesis perversa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="perversa">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesis perversa</emphasis> <emphasis id="EC737EFA8A90DD1F83188BFD0107B36B" italics="true" pageId="0" pageNumber="145">Lachesis perversa</emphasis>
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Audouin, 1826 from Egypt. Audouin, 1826 from Egypt.
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="comments"> <subSubSection id="C765174A813FAC67AD8B89460B32928A" pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="0C631FD96F1A11C889C14134C471E944" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="6938AB932B22B3C71741DE9EFD000AEF" pageId="0" pageNumber="145">
<taxonomicName authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus"> <taxonomicName id="052B7E82673108C322950D4C0872A434" authorityName="Strand" authorityYear="1932" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana</emphasis> <emphasis id="48376675AE9C78D2EDE4F64CF52C18C8" italics="true" pageId="0" pageNumber="145">Lachesana</emphasis>
</taxonomicName> </taxonomicName>
spp. can easily be distinguished from other zodariids occurring in Iran and Turkmenistan by their large size (&gt;10 mm spp. can easily be distinguished from other zodariids occurring in Iran and Turkmenistan by their large size (&gt;10 mm
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="F26A5A0FA3ECCF1D4D71FB376A760063" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
smaller) and numerous spines (&gt;20 on each leg smaller) and numerous spines (&gt;20 on each leg
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="DDFF8DC62F83181772BA745076182EBC" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
lacking, or &lt;15). So far, out of the eight species known from this genus, four are known solely on the basis of male specimens. Male palps are very similar to one another and differ mostly in the proportions of the tibial apophysis and the bulb. lacking, or &lt;15). So far, out of the eight species known from this genus, four are known solely on the basis of male specimens. Male palps are very similar to one another and differ mostly in the proportions of the tibial apophysis and the bulb.
</paragraph> </paragraph>
</subSubSection> </subSubSection>

238
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title> <mods:title id="7F8DC2A0D3C577186880409FA44AEBE3">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<treatment ID-GBIF-Taxon="181753541" LSID="urn:lsid:zoobank.org:act:BCEEC5DD-E5BA-483A-A43F-DF518429CC54" httpUri="http://treatment.plazi.org/id/E852B3FAE0A657ECB5448E7C4B62F79F" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="E852B3FAE0A657ECB5448E7C4B62F79F" ID-GBIF-Taxon="181753541" LSID="urn:lsid:zoobank.org:act:BCEEC5DD-E5BA-483A-A43F-DF518429CC54" httpUri="http://treatment.plazi.org/id/E852B3FAE0A657ECB5448E7C4B62F79F" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="90F74AE85D0A1A358F70F470409EB9FF" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="820AC47B31A07809A75E1B264D971CCE" pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/BCEEC5DD-E5BA-483A-A43F-DF518429CC54" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita" status="sp. nov.">Acanthinozodium armita</taxonomicName> <taxonomicName id="14E6FF79AB961BE7ABB6F3C5838F77D2" ID-CoL="98YZ6" LSID="http://zoobank.org/BCEEC5DD-E5BA-483A-A43F-DF518429CC54" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita" status="sp. nov.">Acanthinozodium armita</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="37B2FA1E27FC19624E9BCC149F32F1E4" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18A</figureCitation> <figureCitation id="9172DF65F06A97D0DF2142A74E31BA9D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18A</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20A-C</figureCitation> <figureCitation id="F59D97F52562FC33112E3B4D6304BC4E" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20A-C</figureCitation>
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">, 22A-C, F</figureCitation> <figureCitation id="E1D4DBEFBB3EBC977317B966993E8325" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">, 22A-C, F</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation> <figureCitation id="EA1CABA9F65995AA5D084D11957131AF" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="6BA58904D5B4C54F2C78033D009D812C" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="C8990DF91A002EBA2CF3893778C66F84" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="45ED1EDFEB7D78D79CE0F738B7FFB5CA" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="097193212896CE0AB61ED5D48E33DC56" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MMUE), Iran: ♂ (MMUE), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Tehran Province</emphasis> <emphasis id="C5C6177E81F18C3D95C18627D35A45E9" italics="true" pageId="0" pageNumber="145">Tehran Province</emphasis>
: northwest of Tehran, Sardor area, : northwest of Tehran, Sardor area,
<geoCoordinate degrees="35" direction="north" minutes="50" orientation="latitude" precision="925" value="35.833332">35°50'N</geoCoordinate> <geoCoordinate id="2977E2959212D520D2F96835C8A31AB5" degrees="35" direction="north" minutes="50" orientation="latitude" precision="925" value="35.833332">35°50'N</geoCoordinate>
, ,
<geoCoordinate degrees="51" direction="east" minutes="05" orientation="longitude" precision="925" value="51.083332">51°05'E</geoCoordinate> <geoCoordinate id="207E7CA4CDC0F7DF96FE0F87AD2A45F2" degrees="51" direction="east" minutes="05" orientation="longitude" precision="925" value="51.083332">51°05'E</geoCoordinate>
, 13.06.2000 (Y.M. Marusik). , 13.06.2000 (Y.M. Marusik).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratype</emphasis> <emphasis id="7D732EA9C623B852F6183041B94E2DD3" bold="true" italics="true" pageId="0" pageNumber="145">Paratype</emphasis>
♂ (NHMW), Iran: Haji Abad, 06.1972 (G. Pretzman &amp; A. Konetschnig). ♂ (NHMW), Iran: Haji Abad, 06.1972 (G. Pretzman &amp; A. Konetschnig).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="740E1E210D5F1D2B27AF82701A04A574" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="EACBFA9EDD872D9CBE79B6D63F597A74" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="91F70C731406F93D6C0A9179B13D5A55" pageId="0" pageNumber="145">
The specific epithet is a Persian feminine name meaning The specific epithet is a Persian feminine name meaning
<normalizedToken originalValue="“righteous”">&quot;righteous&quot;</normalizedToken> <normalizedToken id="B59029E1FA0260DCC38D048624E30111" originalValue="“righteous”">&quot;righteous&quot;</normalizedToken>
, ,
<normalizedToken originalValue="“virtuous”">&quot;virtuous&quot;</normalizedToken> <normalizedToken id="7AE32738A000379F665A6D28F69694C4" originalValue="“virtuous”">&quot;virtuous&quot;</normalizedToken>
, and , and
<normalizedToken originalValue="“good”">&quot;good&quot;</normalizedToken> <normalizedToken id="7C0FB8619AE353688084E707A5C52DFD" originalValue="“good”">&quot;good&quot;</normalizedToken>
. Noun in apposition. . Noun in apposition.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="F72EAA90981A952192EA7C87525D8AF7" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="F68B8DE8CFF0795CEE985889DDE1B3E5" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="6BE109FEB64EE11E29EA34CDDCD406F5" pageId="0" pageNumber="145">
The new species differs from all congeners in the region by the short RTA having 2 claw-like outgrowths on the tip ( The new species differs from all congeners in the region by the short RTA having 2 claw-like outgrowths on the tip (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="A85CF7B423E7C7B328FF63F2B0CA23A2" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
tip tapering and lacking 2 claws) and also by the modified embolus, widening near the tip ( tip tapering and lacking 2 claws) and also by the modified embolus, widening near the tip (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="471EB4466C6022B48ECE3BC58A74D787" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
unmodified). unmodified).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="108222B0A652FFD92FBBDFA88E0E08A1" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="190471E5D4C8BB3C33C14D27FACB4834" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="2E068691B01A990DA4118284E5B2C77A" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis> <emphasis id="EFEE4B53FB13ABF1013EFF4CFB8DA975" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig. (holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18A</figureCitation> <figureCitation id="AE57D6A628C2179848530F4DC8D7F799" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18A</figureCitation>
. Total length 4.70. Carapace 2.18 long, 0.92 wide at pars cephalica, 1.52 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.19, ALE: 0.12, PME: 0.10, PLE: 0.12, PME-PME: 0.25. Carapace dark brown, with irregular dark patches and lines. Sternum, labium and maxillae light brown. Chelicera dark brown, with one tooth. Legs light brown, without annulations. Abdomen black, without any pattern; slightly lighter ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 9.49 (2.38, 0.71, 2.16, 2.76, 1.48), II: 8.76 (2.10, 0.78, 1.98, 2.50, 1.40), III: 8.36 (2.11, 0.70, 1.69, 2.73, 1.13), IV: 11.42 (3.00, 0.81, 2.60, 3.59, 1.42). . Total length 4.70. Carapace 2.18 long, 0.92 wide at pars cephalica, 1.52 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.19, ALE: 0.12, PME: 0.10, PLE: 0.12, PME-PME: 0.25. Carapace dark brown, with irregular dark patches and lines. Sternum, labium and maxillae light brown. Chelicera dark brown, with one tooth. Legs light brown, without annulations. Abdomen black, without any pattern; slightly lighter ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 9.49 (2.38, 0.71, 2.16, 2.76, 1.48), II: 8.76 (2.10, 0.78, 1.98, 2.50, 1.40), III: 8.36 (2.11, 0.70, 1.69, 2.73, 1.13), IV: 11.42 (3.00, 0.81, 2.60, 3.59, 1.42).
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145" start="Figure 18" startId="F18"> <caption id="57DB8917C4AB16507381647D6058C6F7" doi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145" start="Figure 18" startId="F18">
<paragraph pageId="0" pageNumber="145"> <paragraph id="385049E296746D457E64487539DAD115" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 18.</emphasis> <emphasis id="9D346FA1C90A7EDAEF1827580F78B979" bold="true" pageId="0" pageNumber="145">Figure 18.</emphasis>
Habitus of Habitus of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita"> <taxonomicName id="3644AF4578D521158500C7F15102C0C9" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium armita</emphasis> <emphasis id="738EF1B2AA99F942E13053D689DDF52B" italics="true" pageId="0" pageNumber="145">Acanthinozodium armita</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis> <emphasis id="CDC0B9EEC6D681F5F7539413B5409301" bold="true" pageId="0" pageNumber="145">A</emphasis>
), ),
<taxonomicName lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara"> <taxonomicName id="18F60B76E0D579B8EDEFFD8EBE7CC95D" lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">A. diara</emphasis> <emphasis id="5DFC51D5ED3E22105D602B55141AB9F1" italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis> <emphasis id="366C417DD99A2F2D6EC747A7F73412BB" bold="true" pageId="0" pageNumber="145">B</emphasis>
), ),
<taxonomicName lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa"> <taxonomicName id="812C4F17F20836AB1AFD31A3C6B2270A" lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis> <emphasis id="5CCF8162C06C51B2092A6C2F32DCFFB6" italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">C, D</emphasis> <emphasis id="77EE5C05C56CA2C85E70FC6C390E7655" bold="true" pageId="0" pageNumber="145">C, D</emphasis>
), ),
<taxonomicName lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum"> <taxonomicName id="9412D8F6EDD49219FADD3CB2D4D910E6" lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis> <emphasis id="F896D31DFB9F1B797BDB37F4B79EA1D2" italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">E-J</emphasis> <emphasis id="9812AE9AD5D403FE908CCF299F995904" bold="true" pageId="0" pageNumber="145">E-J</emphasis>
), ),
<taxonomicName lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana"> <taxonomicName id="5CF21C23BB32105993BCFF5DC0866D17" lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana">
<emphasis italics="true" pageId="0" pageNumber="145">A. kiana</emphasis> <emphasis id="042746512954FBA540F07B9C1BBF357B" italics="true" pageId="0" pageNumber="145">A. kiana</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">K</emphasis> <emphasis id="FABCC9C7FDB7471C4F2C8DD655E8F324" bold="true" pageId="0" pageNumber="145">K</emphasis>
) and ) and
<taxonomicName lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa"> <taxonomicName id="D5975806357747AA1052D3C8D42F75C1" lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis italics="true" pageId="0" pageNumber="145">A. masa</emphasis> <emphasis id="C188B9C61233F8EE871B3297809CB646" italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">L</emphasis> <emphasis id="D8247BBB1D62DA0A7BB24C3D2E1BBE2B" bold="true" pageId="0" pageNumber="145">L</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A-C, F, G, K, L</emphasis> <emphasis id="382B78F4AA902362D2D35624AEFF692D" bold="true" pageId="0" pageNumber="145">A-C, F, G, K, L</emphasis>
males, dorsal males, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis> <emphasis id="8B46D53FC2C608B26664BBDE73EFBAEB" bold="true" pageId="0" pageNumber="145">H</emphasis>
male, ventral male, ventral
<emphasis bold="true" pageId="0" pageNumber="145">D, E, I</emphasis> <emphasis id="98E432A89D49CE85AB93F35BECFDDD5A" bold="true" pageId="0" pageNumber="145">D, E, I</emphasis>
females, dorsal females, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">J</emphasis> <emphasis id="4C88CA72E6D5DA225979986B1E367B23" bold="true" pageId="0" pageNumber="145">J</emphasis>
female, ventral. Scale bars: 0.2 mm, unless stated otherwise. female, ventral. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph> </paragraph>
</caption> </caption>
<caption doi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145" start="Figure 19" startId="F19"> <caption id="1D74D68B995CC3CBF27D128C3F39A0D1" doi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145" start="Figure 19" startId="F19">
<paragraph pageId="0" pageNumber="145"> <paragraph id="6D139C3F47D06CEE150334E1C4D9B25D" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 19.</emphasis> <emphasis id="AD80F19B4F9E43FBC511B15C982E0950" bold="true" pageId="0" pageNumber="145">Figure 19.</emphasis>
Male habitus and palps of Male habitus and palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parmida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parmida"> <taxonomicName id="876786E54784BF6EF55517C3E9EE4607" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium parmida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="parmida">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium parmida</emphasis> <emphasis id="9EE42A5D5EC24D4725D7B4488A3F240B" italics="true" pageId="0" pageNumber="145">Acanthinozodium parmida</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis> <emphasis id="A90C11A744494F29627DCCDCFB162862" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and ) and
<taxonomicName class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion expers" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="expers"> <taxonomicName id="7B3677A5CF94FBB3185AABA0D2C84066" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion expers" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="expers">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion expers</emphasis> <emphasis id="39FEFE31531B45870F8349C99B61DF76" italics="true" pageId="0" pageNumber="145">Zodarion expers</emphasis>
</taxonomicName> </taxonomicName>
( (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis> <emphasis id="CEE9C2AAC8331AA831A9313D4AECF3E4" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis> <emphasis id="BC09B887BD8F3E234DF92DAB0C14E51D" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
habitus, dorsal habitus, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">B, E</emphasis> <emphasis id="BE2347C85621533CDC46F8AA9681775F" bold="true" pageId="0" pageNumber="145">B, E</emphasis>
palp, ventral palp, ventral
<emphasis bold="true" pageId="0" pageNumber="145">C, F</emphasis> <emphasis id="AEC0D0C18EF08F078CE627918BEB7C3F" bold="true" pageId="0" pageNumber="145">C, F</emphasis>
palp, retrolateral. Scale bar: 0.2 mm ( palp, retrolateral. Scale bar: 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis> <emphasis id="63E7EC074AA1754BA65D38CCC78EFEE4" bold="true" pageId="0" pageNumber="145">A</emphasis>
). ).
</paragraph> </paragraph>
</caption> </caption>
<paragraph pageId="0" pageNumber="145"> <paragraph id="EC27F418AE6C7EFF828AC919EEBB0FF0" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20A-C</figureCitation> <figureCitation id="505FD69DF2DE3866C96E340EAD9496A3" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20A-C</figureCitation>
, ,
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22A-C, F</figureCitation> <figureCitation id="D3465EBA21F9928DA71E367DE8229D16" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. SEM images of the male palps of Acanthinozodium armita sp. nov. (A-C, F) and A. ovtchinnikovi sp. nov. (D, E, G-I) A, H retroventral and retrolateral B, C, G ventral D-F proventral, apicoproventral and apicoventral I retrolateral tibial and cymbial groove. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145">22A-C, F</figureCitation>
. RTA short and stout, with 2 claw-like outgrowths on the tip; ventral apophysis absent; tegulum longer than wide; median apophysis ( . RTA short and stout, with 2 claw-like outgrowths on the tip; ventral apophysis absent; tegulum longer than wide; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="EBE6261F237E1A3693CAA9914332A7BE" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) longer than wide, with outgrowths on all four sides; embolus with a distinct ridge ( ) longer than wide, with outgrowths on all four sides; embolus with a distinct ridge (
<emphasis italics="true" pageId="0" pageNumber="145">Er</emphasis> <emphasis id="E58A9ECC0764446385E0B5A3AB9ECA8B" italics="true" pageId="0" pageNumber="145">Er</emphasis>
); embolus proper twisted basally and widened near the tip. ); embolus proper twisted basally and widened near the tip.
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="24D733B7B5CF07138FB3EC2CEDEB584E" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="8F0452C80B0282E8FB3C905E036E3B3F" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown. Unknown.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="75774EDEED2C98BFEFCCF62D755A3FD3" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="B5376CD01905E46E8ECB47F31964DD33" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="6EED1A963991BC483DE132E076E35103" pageId="0" pageNumber="145">
Known from the type locality in Tehran Province, northern Iran, and another locality (Haji Abad) which cannot be georeferenced because there are many places with this name in Iran (Fig. Known from the type locality in Tehran Province, northern Iran, and another locality (Haji Abad) which cannot be georeferenced because there are many places with this name in Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation> <figureCitation id="D97B44155E9182FE8621F2E356A5365A" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
). ).
</paragraph> </paragraph>
</subSubSection> </subSubSection>

98
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Genus Genus
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<paragraph pageId="0" pageNumber="145">Type species.</paragraph> <paragraph id="10E07E10B9F71E4D6B46DDD767F514E5" pageId="0" pageNumber="145">Type species.</paragraph>
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<taxonomicName class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion raddei" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei"> <taxonomicName id="BA1DCF19FBED91B028472E9463DAFE79" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion raddei" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="raddei">
<emphasis italics="true" pageId="0" pageNumber="145">Zodarion raddei</emphasis> <emphasis id="6B562DC4ACC0BE8823644AE08815039A" italics="true" pageId="0" pageNumber="145">Zodarion raddei</emphasis>
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Simon, 1889 from Turkmenistan. Simon, 1889 from Turkmenistan.
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<paragraph pageId="0" pageNumber="145">Comments.</paragraph> <paragraph id="BAC8500A047C8A93E9D7E9EF72076AAD" pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">Monotypic genus, differing from all other genera in the region by having an elongate cymbium, an S-shaped sperm duct and embolus, a long spine-like outgrowth of the embolic base and an epigyne with large transverse oval fovea.</paragraph> <paragraph id="26E7A0364A5EFACE2055F9F26541765F" pageId="0" pageNumber="145">Monotypic genus, differing from all other genera in the region by having an elongate cymbium, an S-shaped sperm duct and embolus, a long spine-like outgrowth of the embolic base and an epigyne with large transverse oval fovea.</paragraph>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:identifier> <mods:identifier id="77E4D81C4D9ACA232DE863F958890331" type="DOI">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1035-145</mods:identifier> <mods:identifier id="380FE025EEC012CCC79AD6944A7FB368" type="Pensoft-Pub">1313-2970-1035-145</mods:identifier>
<mods:identifier type="ZooBank">ADD73A162DB34873890B1E490C2C8841</mods:identifier> <mods:identifier id="23F50348E932A75B643C003466371D2A" type="ZooBank">ADD73A162DB34873890B1E490C2C8841</mods:identifier>
<mods:identifier type="Pensoft-UUID">55CAE096B4A451D88EA433175454DA3F</mods:identifier> <mods:identifier id="229689C7225BB27676C91CA17BDEE802" type="Pensoft-UUID">55CAE096B4A451D88EA433175454DA3F</mods:identifier>
</mods:mods> </mods:mods>
<treatment ID-GBIF-Taxon="181753542" LSID="urn:lsid:zoobank.org:act:820D70DA-633E-4CF3-B11E-3B16C6974CBC" httpUri="http://treatment.plazi.org/id/F7A70D2BB0355179BD17C0C957A48B7D" lastPageNumber="145" pageId="0" pageNumber="145"> <treatment id="F7A70D2BB0355179BD17C0C957A48B7D" ID-GBIF-Taxon="181753542" LSID="urn:lsid:zoobank.org:act:820D70DA-633E-4CF3-B11E-3B16C6974CBC" httpUri="http://treatment.plazi.org/id/F7A70D2BB0355179BD17C0C957A48B7D" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
<subSubSection pageId="0" pageNumber="145" type="nomenclature"> <subSubSection id="F5813EF32B097AE5DF2CB13F72CE56AE" pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145"> <paragraph id="ADDC701B4127FB5F5262E0FE0E7E64BA" pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/820D70DA-633E-4CF3-B11E-3B16C6974CBC" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium kiana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kiana" status="sp. nov.">Acanthinozodium kiana</taxonomicName> <taxonomicName id="2620E204C470D403397E260E9E7D3DF4" ID-CoL="98YYT" LSID="http://zoobank.org/820D70DA-633E-4CF3-B11E-3B16C6974CBC" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium kiana" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kiana" status="sp. nov.">Acanthinozodium kiana</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="EA621D1911A520FAC024C80555934EE9" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18K</figureCitation> <figureCitation id="6DF48F9C2493BA18921E7E66AE6EC346" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18K</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20G, H</figureCitation> <figureCitation id="746C2DBAACF57DD509A133F0FECE5CF5" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">, 20G, H</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">, 24E, F</figureCitation> <figureCitation id="B32AF5144791761CDF2FFA34C346C16D" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">, 24E, F</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation> <figureCitation id="D88A3176E9194E7E922EFE64BB9D3AA9" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material"> <subSubSection id="FB31BA65D335BB7679A6A140950D4E40" pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph> <paragraph id="E780596864E40442C79A8B8865D381D9" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="2670CCC3F4ED174A93E77BDC0B66F303" pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis> <emphasis id="0653DE923475CF6904AF930580184AE5" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran: ♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Kurdistan Province</emphasis> <emphasis id="297D105F9C53E06A415425796831CD82" italics="true" pageId="0" pageNumber="145">Kurdistan Province</emphasis>
: south of Divandareh, : south of Divandareh,
<geoCoordinate degrees="35" direction="north" minutes="45" orientation="latitude" precision="925" value="35.75">35°45'N</geoCoordinate> <geoCoordinate id="3203D4E2DEE6C265104AB631151B7635" degrees="35" direction="north" minutes="45" orientation="latitude" precision="925" value="35.75">35°45'N</geoCoordinate>
, ,
<geoCoordinate degrees="47" direction="east" minutes="05" orientation="longitude" precision="925" value="47.083332">47°05'E</geoCoordinate> <geoCoordinate id="AE303C15925EEEEABB47A52DA3C4DD44" degrees="47" direction="east" minutes="05" orientation="longitude" precision="925" value="47.083332">47°05'E</geoCoordinate>
, 23.06.1975 (A. Senglet). , 23.06.1975 (A. Senglet).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology"> <subSubSection id="32A35433033BABCC3A702F2BF7470A6F" pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph> <paragraph id="793C658FC57A57DD04B27C6CB91AB495" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="7DF25EF92F9C21A7AEB55104F5167F02" pageId="0" pageNumber="145">
The specific epithet is a Kurdish feminine name meaning The specific epithet is a Kurdish feminine name meaning
<normalizedToken originalValue="“nature”">&quot;nature&quot;</normalizedToken> <normalizedToken id="DACE80CAE16265ECF19CEB4DF7B7AA7B" originalValue="“nature”">&quot;nature&quot;</normalizedToken>
. Noun in apposition. . Noun in apposition.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis"> <subSubSection id="843A84C3A8008F64A66BBF2D2857BBB6" pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph> <paragraph id="AC0131B425CE09CC8D2E28CA031B23A2" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="4AF9B06270B729D2B340AB2782C6492D" pageId="0" pageNumber="145">
The new species is very similar to The new species is very similar to
<taxonomicName lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa"> <taxonomicName id="00042A2C526C6F4D9D392F0CFA248621" lsidName="A. masa" pageId="0" pageNumber="145" rank="species" species="masa">
<emphasis italics="true" pageId="0" pageNumber="145">A. masa</emphasis> <emphasis id="F54FA21D3BF9F58C32A761A2E4B13622" italics="true" pageId="0" pageNumber="145">A. masa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. by the shape of the RTA and ventral tibial apophysis but differs by the shape of the prolateral outgrowth of the median apophysis which is finger-like and triangular, terminally with a small hook ( sp. nov. by the shape of the RTA and ventral tibial apophysis but differs by the shape of the prolateral outgrowth of the median apophysis which is finger-like and triangular, terminally with a small hook (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis> <emphasis id="131D69F7914E96E36AECCAD67CA49087" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
broad and bifurcated). broad and bifurcated).
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description"> <subSubSection id="46AA43C4AF21B190F1271CE728D1D649" pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph> <paragraph id="89937DDC1CE1A1B7D86ED092A566C6D8" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="E4CB6D1D58231C1F0C04AB36075987B3" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis> <emphasis id="19389DA2D0F8E5AFF4FB8CFC7168FFD3" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Habitus as in Fig. Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18K</figureCitation> <figureCitation id="E522DF4524DB9E3FF2831781F65CDD21" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18K</figureCitation>
. Total length 3.80. Carapace 2.08 long, 1.03 wide at pars cephalica, 1.85 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.17, ALE: 0.12, PME: 0.09, PLE: 0.10, PME-PME: 0.23. Carapace light brown, darker at pars cephalica. Sternum, labium, maxillae and chelicerae yellowish. Legs yellowish, with numerous macrosetae and without annulations. Abdomen grayish dorsally, lighter ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 7.57 (2.02, 0.75, 1.67, 1.97, 1.16), II: 7.06 (1.84, 0.74, 1.50, 1.95, 1.03), III: 7.24 (1.98, 0.71, 1.47, 2.08, 1.00), IV: 9.46 (2.63, 0.78, 2.26, 2.77, 1.02). . Total length 3.80. Carapace 2.08 long, 1.03 wide at pars cephalica, 1.85 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.17, ALE: 0.12, PME: 0.09, PLE: 0.10, PME-PME: 0.23. Carapace light brown, darker at pars cephalica. Sternum, labium, maxillae and chelicerae yellowish. Legs yellowish, with numerous macrosetae and without annulations. Abdomen grayish dorsally, lighter ventrally. Spinnerets pale, uniform in color. Measurements of legs: I: 7.57 (2.02, 0.75, 1.67, 1.97, 1.16), II: 7.06 (1.84, 0.74, 1.50, 1.95, 1.03), III: 7.24 (1.98, 0.71, 1.47, 2.08, 1.00), IV: 9.46 (2.63, 0.78, 2.26, 2.77, 1.02).
</paragraph> </paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="338D61E75D0AE9EA50551CE85A6955E7" pageId="0" pageNumber="145">
Palp as in Figs Palp as in Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20G, H</figureCitation> <figureCitation id="4C30AAB6C691726A4399C19424A5F5E3" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male palps of Acanthinozodium armita sp. nov. (A-C), A. diara sp. nov. (D-F), A. kiana sp. nov. (G, H) and A. masa sp. nov. (I-K) A, D, G, J ventral B, E, H, I retrolateral C, F, K dorsal. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure20" httpUri="https://binary.pensoft.net/fig/536897" pageId="0" pageNumber="145">20G, H</figureCitation>
, ,
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">24E, F</figureCitation> <figureCitation id="78085FDEB991283367D9731EE4B758C6" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. SEM images of the male palps of Acanthinozodium masa sp. nov. (A, B), A. dorsa sp. nov. (C, D) and A. kiana sp. nov. (E, F) A, C, E ventral B, F retroventral D retrolateral. Abbreviations: Co - conductor, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis, Tp - tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145">24E, F</figureCitation>
. RTA long, with a small projection apically; ventral apophysis relatively small and conical; tegulum with posterior process; median apophysis ( . RTA long, with a small projection apically; ventral apophysis relatively small and conical; tegulum with posterior process; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="53C964630C94A14F0EE5D7662C274E6C" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) wider than long, with a distinct prolateral outgrowth which is triangular terminally and with a small hook; embolus broad basally, originating at about the 7 ) wider than long, with a distinct prolateral outgrowth which is triangular terminally and with a small hook; embolus broad basally, originating at about the 7
<normalizedToken originalValue="o’clock">o'clock</normalizedToken> <normalizedToken id="AF3CB37A6D1044BE8D3600C8442D8306" originalValue="o’clock">o'clock</normalizedToken>
position; embolus proper relatively broad and distinctly curved over the median apophysis. position; embolus proper relatively broad and distinctly curved over the median apophysis.
</paragraph> </paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145" start="Figure 24" startId="F24"> <caption id="A8CAEB55244FE3C3A9B4B479629C8FB9" doi="10.3897/zookeys.1035.65767.figure24" httpUri="https://binary.pensoft.net/fig/536901" pageId="0" pageNumber="145" start="Figure 24" startId="F24">
<paragraph pageId="0" pageNumber="145"> <paragraph id="36978201FFE5725177B69EB97DFD86FA" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 24.</emphasis> <emphasis id="E3BE34C51FBE74EC42F5A1C5E0FA9B62" bold="true" pageId="0" pageNumber="145">Figure 24.</emphasis>
SEM images of the male palps of SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium masa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="masa"> <taxonomicName id="0B49705E4776091B2F56712923396F8E" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium masa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="masa">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium masa</emphasis> <emphasis id="B8D9FEBE398732DF014F15FCD9A44B1E" italics="true" pageId="0" pageNumber="145">Acanthinozodium masa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A, B</emphasis> <emphasis id="6FE1E8D34E3E087317425599AFFF24C0" bold="true" pageId="0" pageNumber="145">A, B</emphasis>
), ),
<taxonomicName lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa"> <taxonomicName id="972F40DD663A6D6BEBA442609B1FCB06" lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis> <emphasis id="3B3EEAF7DD59809A4A082EFCC160FF75" italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">C, D</emphasis> <emphasis id="82C553B6EC10EFC0D00A7BDD57C88776" bold="true" pageId="0" pageNumber="145">C, D</emphasis>
) and ) and
<taxonomicName lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana"> <taxonomicName id="409D3411313FDB23420F1FDBE1600CBF" lsidName="A. kiana" pageId="0" pageNumber="145" rank="species" species="kiana">
<emphasis italics="true" pageId="0" pageNumber="145">A. kiana</emphasis> <emphasis id="6E1E1199E3ED8EED7D793E90E83224CE" italics="true" pageId="0" pageNumber="145">A. kiana</emphasis>
</taxonomicName> </taxonomicName>
sp. nov. ( sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">E, F</emphasis> <emphasis id="878096E70211846428074DC6C65325A5" bold="true" pageId="0" pageNumber="145">E, F</emphasis>
) )
<emphasis bold="true" pageId="0" pageNumber="145">A, C, E</emphasis> <emphasis id="345276AA707DECD0206C1A91CE822BB5" bold="true" pageId="0" pageNumber="145">A, C, E</emphasis>
ventral ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, F</emphasis> <emphasis id="DDAA6F14DFC5D819C211BD513EBEAA33" bold="true" pageId="0" pageNumber="145">B, F</emphasis>
retroventral retroventral
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis> <emphasis id="54C30348B51A4A105120AD0A15A61E7A" bold="true" pageId="0" pageNumber="145">D</emphasis>
retrolateral. Abbreviations: retrolateral. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis> <emphasis id="1FAC00ED7D21971E77F29395503FE3A9" italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor, - conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis> <emphasis id="B77D01D0F0AE611D281DFF9F283EAFC7" italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus, - embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis> <emphasis id="38D0814A97675534E9DBDD6026F520C0" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis, - median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis> <emphasis id="0AFE89752273BCE5ABE9C72D7D720D8A" italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral tibial apophysis, - retrolateral tibial apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis> <emphasis id="57D3C84D10723AD98F318F2DBDCBBA23" italics="true" pageId="0" pageNumber="145">Va</emphasis>
- ventral apophysis, - ventral apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Tp</emphasis> <emphasis id="EABCF13B4B42DA297AE67BC2E4BA7F43" italics="true" pageId="0" pageNumber="145">Tp</emphasis>
- tegular process. Scale bars: 0.1 mm. - tegular process. Scale bars: 0.1 mm.
</paragraph> </paragraph>
</caption> </caption>
<paragraph pageId="0" pageNumber="145"> <paragraph id="339AABB4376EED1D7ED4907F9DFAF0E7" pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis> <emphasis id="B6C9F3E41C3D16BA2B1FE5B4D1001112" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown. Unknown.
</paragraph> </paragraph>
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<subSubSection pageId="0" pageNumber="145" type="distribution"> <subSubSection id="28FFBE9BE2BB40890DAE80EFCF0BA66D" pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph> <paragraph id="D30AE5DDE9362C5F76E958F6EEB060A7" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145"> <paragraph id="9EAADFD9100AC83980A6300FFC410EAB" pageId="0" pageNumber="145">
Known from the type locality in Kurdistan Province, western Iran (Fig. Known from the type locality in Kurdistan Province, western Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation> <figureCitation id="5AF6863333BA68541356B7B6A9CACD21" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
). ).
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231
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<emphasis bold="true" box="[151,310,1075,1102]" italics="true" pageId="17" pageNumber="108">Duroca lenta</emphasis> <emphasis id="4297EAF09B27FFD2FF3CA17FBB36FB7C" bold="true" box="[151,310,1075,1102]" italics="true" pageId="17" pageNumber="108">Duroca lenta</emphasis>
( (
<bibRefCitation author="Evans, W. H." box="[326,479,1076,1102]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans, 1955</bibRefCitation> <bibRefCitation id="14724B139B27FFD2FEEDA17FBBDFFB7C" author="Evans, W. H." box="[326,479,1076,1102]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans, 1955</bibRefCitation>
) )
</taxonomicName> </taxonomicName>
, reinstated status , reinstated status
@ -110,84 +111,84 @@
</heading> </heading>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection pageId="17" pageNumber="108" type="description"> <subSubSection id="38F965699B27FFD2FF3CA137BEB3FACC" pageId="17" pageNumber="108" type="description">
<paragraph blockId="17.[151,1437,1148,1535]" pageId="17" pageNumber="108"> <paragraph id="705C36E29B27FFD2FF3CA137BEB3FACC" blockId="17.[151,1437,1148,1535]" pageId="17" pageNumber="108">
Genomic sequencing of primary type specimen of Genomic sequencing of primary type specimen of
<taxonomicName authority="Pl" authorityName="Pl" box="[756,988,1148,1176]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="duroca"> <taxonomicName id="B7E34D619B27FFD2FD5FA136B9DCFBAA" authority="Pl" authorityName="Pl" box="[756,988,1148,1176]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="duroca">
<emphasis box="[756,949,1148,1174]" italics="true" pageId="17" pageNumber="108">Hesperia duroca</emphasis> <emphasis id="4297EAF09B27FFD2FD5FA136B9B5FBA4" box="[756,949,1148,1174]" italics="true" pageId="17" pageNumber="108">Hesperia duroca</emphasis>
Pl Pl
</taxonomicName> </taxonomicName>
̂tz, 1882 (type locality in ̂tz, 1882 (type locality in
<collectingCountry box="[1307,1380,1148,1174]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry> <collectingCountry id="08F476729B27FFD2FAB0A137BF64FBA4" box="[1307,1380,1148,1174]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry>
: :
<collectingRegion country="Brazil" name="Rio de Janeiro" pageId="17" pageNumber="108">Rio de Janeiro</collectingRegion> <collectingRegion id="B227F8009B27FFD2FADFA137BB11FB88" country="Brazil" name="Rio de Janeiro" pageId="17" pageNumber="108">Rio de Janeiro</collectingRegion>
, NVG-15034C , NVG-15034C
<quantity box="[448,507,1184,1211]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="17" pageNumber="108" unit="in" value="12.0">12 in</quantity> <quantity id="B71B9B079B27FFD2FE6BA1EBBBFBFB88" box="[448,507,1184,1211]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="17" pageNumber="108" unit="in" value="12.0">12 in</quantity>
MFNB), the type species of MFNB), the type species of
<taxonomicName authority="Grishin, 2019" authorityName="Grishin" authorityYear="2019" box="[855,1109,1184,1210]" genus="Duroca" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="108" rank="genus"> <taxonomicName id="B7E34D619B27FFD2FCFCA1EABE55FB88" authority="Grishin, 2019" authorityName="Grishin" authorityYear="2019" box="[855,1109,1184,1210]" class="Insecta" family="Hesperiidae" genus="Duroca" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis box="[855,940,1185,1210]" italics="true" pageId="17" pageNumber="108">Duroca</emphasis> <emphasis id="4297EAF09B27FFD2FCFCA1EAB9ACFB88" box="[855,940,1185,1210]" italics="true" pageId="17" pageNumber="108">Duroca</emphasis>
Grishin, 2019 Grishin, 2019
</taxonomicName> </taxonomicName>
( (
<figureCitation box="[1128,1198,1184,1210]" captionStart="FIGURE 8" captionStartId="19.[151,250,759,783]" captionTargetBox="[153,1433,185,731]" captionTargetId="figure-21@19.[151,1436,181,735]" captionTargetPageId="19" captionText="FIGURE 8. Nuclear genome tree of selected Moncina: Phlebodes relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Cantha (purple, with species transferred from Phlebodes labeled in dark green), Duroca (violet, with Duroca caraca, comb. nov. and Duroca lenta, stat. nov. labeled in aquamarine and brown, respectively). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864290" httpUri="https://zenodo.org/record/7864290/files/figure.png" pageId="17" pageNumber="108">Fig. 8</figureCitation> <figureCitation id="E8D82A679B27FFD2FBC3A1EBBEAEFB88" box="[1128,1198,1184,1210]" captionStart="FIGURE 8" captionStartId="19.[151,250,759,783]" captionTargetBox="[153,1433,185,731]" captionTargetId="figure-21@19.[151,1436,181,735]" captionTargetPageId="19" captionText="FIGURE 8. Nuclear genome tree of selected Moncina: Phlebodes relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Cantha (purple, with species transferred from Phlebodes labeled in dark green), Duroca (violet, with Duroca caraca, comb. nov. and Duroca lenta, stat. nov. labeled in aquamarine and brown, respectively). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864290" httpUri="https://zenodo.org/record/7864290/files/figure.png" pageId="17" pageNumber="108">Fig. 8</figureCitation>
violet), and violet), and
<taxonomicName authority="Evans, 1955" authorityName="Evans" authorityYear="1955" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="lenta"> <taxonomicName id="B7E34D619B27FFD2FAE3A1EABB67FBED" authority="Evans, 1955" authorityName="Evans" authorityYear="1955" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="lenta">
<emphasis italics="true" pageId="17" pageNumber="108">Lerema lenta</emphasis> <emphasis id="4297EAF09B27FFD2FAE3A1EABACFFBEC" italics="true" pageId="17" pageNumber="108">Lerema lenta</emphasis>
<bibRefCitation author="Evans, W. H." box="[217,359,1220,1247]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans, 1955</bibRefCitation> <bibRefCitation id="14724B139B27FFD2FF72A18EBB67FBED" author="Evans, W. H." box="[217,359,1220,1247]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName> </taxonomicName>
(type locality in (type locality in
<collectingCountry box="[563,636,1220,1246]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry> <collectingCountry id="08F476729B27FFD2FD98A18FB87CFBEC" box="[563,636,1220,1246]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry>
: S„o Paulo, molecular code NHMUK0247278362 in BMNH) ( : S„o Paulo, molecular code NHMUK0247278362 in BMNH) (
<figureCitation box="[1367,1436,1220,1246]" captionStart="FIGURE 8" captionStartId="19.[151,250,759,783]" captionTargetBox="[153,1433,185,731]" captionTargetId="figure-21@19.[151,1436,181,735]" captionTargetPageId="19" captionText="FIGURE 8. Nuclear genome tree of selected Moncina: Phlebodes relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Cantha (purple, with species transferred from Phlebodes labeled in dark green), Duroca (violet, with Duroca caraca, comb. nov. and Duroca lenta, stat. nov. labeled in aquamarine and brown, respectively). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864290" httpUri="https://zenodo.org/record/7864290/files/figure.png" pageId="17" pageNumber="108">Fig. 8</figureCitation> <figureCitation id="E8D82A679B27FFD2FAFCA18FBF9CFBEC" box="[1367,1436,1220,1246]" captionStart="FIGURE 8" captionStartId="19.[151,250,759,783]" captionTargetBox="[153,1433,185,731]" captionTargetId="figure-21@19.[151,1436,181,735]" captionTargetPageId="19" captionText="FIGURE 8. Nuclear genome tree of selected Moncina: Phlebodes relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Cantha (purple, with species transferred from Phlebodes labeled in dark green), Duroca (violet, with Duroca caraca, comb. nov. and Duroca lenta, stat. nov. labeled in aquamarine and brown, respectively). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864290" httpUri="https://zenodo.org/record/7864290/files/figure.png" pageId="17" pageNumber="108">Fig. 8</figureCitation>
brown), currently a subspecies of brown), currently a subspecies of
<taxonomicName box="[534,647,1256,1282]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="duroca"> <taxonomicName id="B7E34D619B27FFD2FDBDA1A2B887FA30" box="[534,647,1256,1282]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="duroca">
<emphasis box="[534,647,1256,1282]" italics="true" pageId="17" pageNumber="108">D. duroca</emphasis> <emphasis id="4297EAF09B27FFD2FDBDA1A2B887FA30" box="[534,647,1256,1282]" italics="true" pageId="17" pageNumber="108">D. duroca</emphasis>
</taxonomicName> </taxonomicName>
, reveals 2.6% (17 bp) difference in their COI barcodes. Coupled with phenotypic differences described by , reveals 2.6% (17 bp) difference in their COI barcodes. Coupled with phenotypic differences described by
<bibRefCitation author="Evans, W. H." box="[565,718,1292,1319]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans (1955)</bibRefCitation> <bibRefCitation id="14724B139B27FFD2FD9EA046B8CEFA15" author="Evans, W. H." box="[565,718,1292,1319]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans (1955)</bibRefCitation>
, such as unmarked wings, smaller size, and more produced and pointed harpe in , such as unmarked wings, smaller size, and more produced and pointed harpe in
<taxonomicName authorityName="Evans" authorityYear="1955" box="[344,430,1328,1354]" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="lenta"> <taxonomicName id="B7E34D619B27FFD2FEF3A07ABBAEFA78" authorityName="Evans" authorityYear="1955" box="[344,430,1328,1354]" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="lenta">
<emphasis box="[344,430,1328,1354]" italics="true" pageId="17" pageNumber="108">L. lenta</emphasis> <emphasis id="4297EAF09B27FFD2FEF3A07ABBAEFA78" box="[344,430,1328,1354]" italics="true" pageId="17" pageNumber="108">L. lenta</emphasis>
</taxonomicName> </taxonomicName>
, this genetic differentiation suggests species status for , this genetic differentiation suggests species status for
<taxonomicName authority="(Evans, 1955)" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[1056,1373,1328,1355]" genus="Duroca" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="108" rank="species" species="lenta"> <taxonomicName id="B7E34D619B27FFD2FB8BA07ABF5DFA79" authority="(Evans, 1955)" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[1056,1373,1328,1355]" class="Insecta" family="Hesperiidae" genus="Duroca" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="lenta">
<emphasis box="[1056,1204,1328,1354]" italics="true" pageId="17" pageNumber="108">Duroca lenta</emphasis> <emphasis id="4297EAF09B27FFD2FB8BA07ABEB4FA78" box="[1056,1204,1328,1354]" italics="true" pageId="17" pageNumber="108">Duroca lenta</emphasis>
( (
<bibRefCitation author="Evans, W. H." box="[1220,1364,1328,1355]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans, 1955</bibRefCitation> <bibRefCitation id="14724B139B27FFD2FB6FA07BBF54FA79" author="Evans, W. H." box="[1220,1364,1328,1355]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans, 1955</bibRefCitation>
) )
</taxonomicName> </taxonomicName>
, ,
<emphasis bold="true" pageId="17" pageNumber="108">stat. rest</emphasis> <emphasis id="4297EAF09B27FFD2FAC1A07BBAC4FA5C" bold="true" pageId="17" pageNumber="108">stat. rest</emphasis>
. Furthermore, sequence comparison confirms that . Furthermore, sequence comparison confirms that
<taxonomicName authority="Schaus, 1902" authorityName="Schaus" authorityYear="1902" box="[768,1086,1364,1390]" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="elgina"> <taxonomicName id="B7E34D619B27FFD2FCABA01EBE3EFA5C" authority="Schaus, 1902" authorityName="Schaus" authorityYear="1902" box="[768,1086,1364,1390]" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="elgina">
<emphasis box="[768,930,1364,1390]" italics="true" pageId="17" pageNumber="108">Lerema elgina</emphasis> <emphasis id="4297EAF09B27FFD2FCABA01EB9A2FA5C" box="[768,930,1364,1390]" italics="true" pageId="17" pageNumber="108">Lerema elgina</emphasis>
Schaus, 1902 Schaus, 1902
</taxonomicName> </taxonomicName>
(type locality in (type locality in
<collectingCountry box="[1279,1352,1364,1390]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry> <collectingCountry id="08F476729B27FFD2FB54A01FBF48FA5C" box="[1279,1352,1364,1390]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry>
: :
<collectingRegion country="Brazil" name="Rio de Janeiro" pageId="17" pageNumber="108">Rio de Janeiro</collectingRegion> <collectingRegion id="B227F8009B27FFD2FAF9A01FBAEAFAA0" country="Brazil" name="Rio de Janeiro" pageId="17" pageNumber="108">Rio de Janeiro</collectingRegion>
) is a junior subjective synonym of ) is a junior subjective synonym of
<taxonomicName box="[627,796,1400,1426]" genus="Duroca" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="108" rank="species" species="duroca"> <taxonomicName id="B7E34D619B27FFD2FDD8A032B91CFAA0" box="[627,796,1400,1426]" class="Insecta" family="Hesperiidae" genus="Duroca" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="duroca">
<emphasis box="[627,796,1400,1426]" italics="true" pageId="17" pageNumber="108">Duroca duroca</emphasis> <emphasis id="4297EAF09B27FFD2FDD8A032B91CFAA0" box="[627,796,1400,1426]" italics="true" pageId="17" pageNumber="108">Duroca duroca</emphasis>
</taxonomicName> </taxonomicName>
. Finally, to ensure correct identification of this species, a . Finally, to ensure correct identification of this species, a
<typeStatus box="[151,238,1437,1463]" pageId="17" pageNumber="108">syntype</typeStatus> <typeStatus id="AF5888409B27FFD2FF3CA0D6BAEEFA85" box="[151,238,1437,1463]" pageId="17" pageNumber="108" type="syntype">syntype</typeStatus>
in the USNM collection bearing the following five rectangular labels, 4 in the USNM collection bearing the following five rectangular labels, 4
<superScript attach="left" box="[1033,1046,1436,1450]" fontSize="6" pageId="17" pageNumber="108">th</superScript> <superScript id="87969BAA9B27FFD2FBA2A0D7BE16FA98" attach="left" box="[1033,1046,1436,1450]" fontSize="6" pageId="17" pageNumber="108">th</superScript>
red, others white: [Nova Friburgo, | red, others white: [Nova Friburgo, |
<collectingCountry box="[164,232,1472,1498]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry> <collectingCountry id="08F476729B27FFD2FF0FA08BBAE8FAE8" box="[164,232,1472,1498]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry>
], [Collection | W. Schaus], [ ], [Collection | W. Schaus], [
<taxonomicName authorityName="Scudder" authorityYear="1872" box="[583,669,1473,1499]" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="genus">Lerema</taxonomicName> <taxonomicName id="B7E34D619B27FFD2FDECA08AB89DFAE9" authorityName="Scudder" authorityYear="1872" box="[583,669,1473,1499]" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="genus">Lerema</taxonomicName>
| eglina | type Sch.], [Type | No. 6054 | | eglina | type Sch.], [Type | No. 6054 |
<collectingCountry box="[1131,1176,1472,1499]" name="United States of America" pageId="17" pageNumber="108">U.S.</collectingCountry> <collectingCountry id="08F476729B27FFD2FBC0A08ABE98FAE8" box="[1131,1176,1472,1499]" name="United States of America" pageId="17" pageNumber="108">U.S.</collectingCountry>
N.M.], [USNMENT | {QR code} | 00913064] is herein designated as the N.M.], [USNMENT | {QR code} | 00913064] is herein designated as the
<emphasis bold="true" box="[733,840,1508,1534]" pageId="17" pageNumber="108"> <emphasis id="4297EAF09B27FFD2FD76A0AFB948FACC" bold="true" box="[733,840,1508,1534]" pageId="17" pageNumber="108">
<typeStatus box="[733,840,1508,1534]" pageId="17" pageNumber="108">lectotype</typeStatus> <typeStatus id="AF5888409B27FFD2FD76A0AFB948FACC" box="[733,840,1508,1534]" pageId="17" pageNumber="108" type="lectotype">lectotype</typeStatus>
</emphasis> </emphasis>
of of
<taxonomicName authority="Schaus, 1902" authorityName="Schaus" authorityYear="1902" box="[877,1198,1508,1534]" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="elgina"> <taxonomicName id="B7E34D619B27FFD2FCC6A0AEBEAEFACC" authority="Schaus, 1902" authorityName="Schaus" authorityYear="1902" box="[877,1198,1508,1534]" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="elgina">
<emphasis box="[877,1039,1508,1534]" italics="true" pageId="17" pageNumber="108">Lerema elgina</emphasis> <emphasis id="4297EAF09B27FFD2FCC6A0AEBE0FFACC" box="[877,1039,1508,1534]" italics="true" pageId="17" pageNumber="108">Lerema elgina</emphasis>
Schaus, 1902 Schaus, 1902
</taxonomicName> </taxonomicName>
. .

193
data/F8/4A/87/F84A87F49B27FFD2FF3CA65FB901FCD1.xml
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@ -1,155 +1,156 @@
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Sequencing of the Sequencing of the
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and and
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O. Mielke, 1992 O. Mielke, 1992
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(type locality in (type locality in
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) ( ) (
<figureCitation box="[159,226,860,886]" captionStart="FIGURE 8" captionStartId="19.[151,250,759,783]" captionTargetBox="[153,1433,185,731]" captionTargetId="figure-21@19.[151,1436,181,735]" captionTargetPageId="19" captionText="FIGURE 8. Nuclear genome tree of selected Moncina: Phlebodes relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Cantha (purple, with species transferred from Phlebodes labeled in dark green), Duroca (violet, with Duroca caraca, comb. nov. and Duroca lenta, stat. nov. labeled in aquamarine and brown, respectively). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864290" httpUri="https://zenodo.org/record/7864290/files/figure.png" pageId="17" pageNumber="108">Fig. 8</figureCitation> <figureCitation id="E8D82A679B27FFD2FF34A617BAE2FC44" box="[159,226,860,886]" captionStart="FIGURE 8" captionStartId="19.[151,250,759,783]" captionTargetBox="[153,1433,185,731]" captionTargetId="figure-21@19.[151,1436,181,735]" captionTargetPageId="19" captionText="FIGURE 8. Nuclear genome tree of selected Moncina: Phlebodes relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Cantha (purple, with species transferred from Phlebodes labeled in dark green), Duroca (violet, with Duroca caraca, comb. nov. and Duroca lenta, stat. nov. labeled in aquamarine and brown, respectively). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864290" httpUri="https://zenodo.org/record/7864290/files/figure.png" pageId="17" pageNumber="108">Fig. 8</figureCitation>
aquamarine) reveals its close relationship with aquamarine) reveals its close relationship with
<taxonomicName authority="Pl" authorityName="Pl" box="[763,985,860,888]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="duroca"> <taxonomicName id="B7E34D619B27FFD2FD50A616B9D9FC4A" authority="Pl" authorityName="Pl" box="[763,985,860,888]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="duroca">
<emphasis box="[763,950,860,886]" italics="true" pageId="17" pageNumber="108">Hesperia duroca</emphasis> <emphasis id="4297EAF09B27FFD2FD50A616B9B6FC44" box="[763,950,860,886]" italics="true" pageId="17" pageNumber="108">Hesperia duroca</emphasis>
Pl Pl
</taxonomicName> </taxonomicName>
̂tz, 1882 (type locality in ̂tz, 1882 (type locality in
<collectingCountry box="[1278,1351,860,886]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry> <collectingCountry id="08F476729B27FFD2FB55A617BF47FC44" box="[1278,1351,860,886]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry>
: :
<collectingRegion country="Brazil" name="Rio de Janeiro" pageId="17" pageNumber="108">Rio de Janeiro</collectingRegion> <collectingRegion id="B227F8009B27FFD2FAF9A617BAECFCA8" country="Brazil" name="Rio de Janeiro" pageId="17" pageNumber="108">Rio de Janeiro</collectingRegion>
), the type species of ), the type species of
<taxonomicName authority="Grishin, 2019" authorityName="Grishin" authorityYear="2019" box="[469,714,896,922]" genus="Duroca" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="108" rank="genus"> <taxonomicName id="B7E34D619B27FFD2FE7EA6CAB8CAFCA8" authority="Grishin, 2019" authorityName="Grishin" authorityYear="2019" box="[469,714,896,922]" class="Insecta" family="Hesperiidae" genus="Duroca" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis box="[469,554,897,922]" italics="true" pageId="17" pageNumber="108">Duroca</emphasis> <emphasis id="4297EAF09B27FFD2FE7EA6CAB82AFCA8" box="[469,554,897,922]" italics="true" pageId="17" pageNumber="108">Duroca</emphasis>
Grishin, 2019 Grishin, 2019
</taxonomicName> </taxonomicName>
( (
<figureCitation box="[729,795,896,922]" captionStart="FIGURE 8" captionStartId="19.[151,250,759,783]" captionTargetBox="[153,1433,185,731]" captionTargetId="figure-21@19.[151,1436,181,735]" captionTargetPageId="19" captionText="FIGURE 8. Nuclear genome tree of selected Moncina: Phlebodes relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Cantha (purple, with species transferred from Phlebodes labeled in dark green), Duroca (violet, with Duroca caraca, comb. nov. and Duroca lenta, stat. nov. labeled in aquamarine and brown, respectively). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864290" httpUri="https://zenodo.org/record/7864290/files/figure.png" pageId="17" pageNumber="108">Fig. 8</figureCitation> <figureCitation id="E8D82A679B27FFD2FD72A6CBB91BFCA8" box="[729,795,896,922]" captionStart="FIGURE 8" captionStartId="19.[151,250,759,783]" captionTargetBox="[153,1433,185,731]" captionTargetId="figure-21@19.[151,1436,181,735]" captionTargetPageId="19" captionText="FIGURE 8. Nuclear genome tree of selected Moncina: Phlebodes relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Cantha (purple, with species transferred from Phlebodes labeled in dark green), Duroca (violet, with Duroca caraca, comb. nov. and Duroca lenta, stat. nov. labeled in aquamarine and brown, respectively). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864290" httpUri="https://zenodo.org/record/7864290/files/figure.png" pageId="17" pageNumber="108">Fig. 8</figureCitation>
violet). The COI barcodes of the type specimens of these species reveal 5.4% (36 bp) difference, which is small enough even for closely related congeners, and therefore we propose violet). The COI barcodes of the type specimens of these species reveal 5.4% (36 bp) difference, which is small enough even for closely related congeners, and therefore we propose
<taxonomicName authority="(O. Mielke, 1992)" authorityName="Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin" authorityYear="2023" baseAuthorityName="O. Mielke" baseAuthorityYear="1992" box="[247,628,968,995]" genus="Duroca" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="108" rank="species" species="caraca" status="comb. nov."> <taxonomicName id="B7E34D619B27FFD2FF5CA682B874FCD0" authority="(O. Mielke, 1992)" authorityName="Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin" authorityYear="2023" baseAuthorityName="O. Mielke" baseAuthorityYear="1992" box="[247,628,968,995]" class="Insecta" family="Hesperiidae" genus="Duroca" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="caraca" status="comb. nov.">
<emphasis box="[247,416,969,994]" italics="true" pageId="17" pageNumber="108">Duroca caraca</emphasis> <emphasis id="4297EAF09B27FFD2FF5CA682BBA0FCD0" box="[247,416,969,994]" italics="true" pageId="17" pageNumber="108">Duroca caraca</emphasis>
(O. Mielke, 1992) (O. Mielke, 1992)
</taxonomicName> </taxonomicName>
, ,
<taxonomicNameLabel box="[639,769,968,995]" pageId="17" pageNumber="108" rank="species"> <taxonomicNameLabel id="59A4578B9B27FFD2FDD4A683B901FCD1" box="[639,769,968,995]" pageId="17" pageNumber="108" rank="species">
<emphasis bold="true" box="[639,763,968,994]" pageId="17" pageNumber="108">comb. nov</emphasis> <emphasis id="4297EAF09B27FFD2FDD4A683B8FBFCD0" bold="true" box="[639,763,968,994]" pageId="17" pageNumber="108">comb. nov</emphasis>
. .
</taxonomicNameLabel> </taxonomicNameLabel>
</paragraph> </paragraph>