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<mods:title id="BB7648F9AFF85B0F570755C243021875">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:namePart>Zamani, Alireza</mods:namePart>
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<treatment ID-GBIF-Taxon="181753554" LSID="urn:lsid:zoobank.org:act:F3F99EF7-B5F4-42CB-AD3B-2404007AFBA2" httpUri="http://treatment.plazi.org/id/C18D05F563465EA48B4B75AB8EAF40A2" lastPageNumber="145" pageId="0" pageNumber="145">
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<taxonomicName LSID="http://zoobank.org/F3F99EF7-B5F4-42CB-AD3B-2404007AFBA2" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana perseus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="perseus" status="sp. nov.">Lachesana perseus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">Figs 1D-F</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live habitus of male Lachesana perseus sp. nov. Photo: A. Zamani." figureDoi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">, 3F-J</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">, 4J</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
<treatment id="C18D05F563465EA48B4B75AB8EAF40A2" ID-GBIF-Taxon="181753554" LSID="urn:lsid:zoobank.org:act:F3F99EF7-B5F4-42CB-AD3B-2404007AFBA2" httpUri="http://treatment.plazi.org/id/C18D05F563465EA48B4B75AB8EAF40A2" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Lachesaninae">
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<paragraph id="BBA2982FCDCD950D2F7CCCB5A1685745" pageId="0" pageNumber="145">
<taxonomicName id="C33C25152DD2967F0B586564437BF9E6" ID-CoL="98YYX" LSID="http://zoobank.org/F3F99EF7-B5F4-42CB-AD3B-2404007AFBA2" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana perseus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="perseus" status="sp. nov.">Lachesana perseus</taxonomicName>
<taxonomicNameLabel id="61613BBFC11310B8911AF37D0CDC64C1" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation id="C001E61BA894AD3CE3460CF357CD9EA3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">Figs 1D-F</figureCitation>
<figureCitation id="D5AC0B550EC5C290E8143EA1B5065D0E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live habitus of male Lachesana perseus sp. nov. Photo: A. Zamani." figureDoi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145">, 2</figureCitation>
<figureCitation id="F238BFACD8FDCE897EA02FD10FDC58BE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">, 3F-J</figureCitation>
<figureCitation id="AC2DE1B1721AC498E801F8B75B3239AC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">, 4J</figureCitation>
<figureCitation id="B178CF70A9DBB419C034AB91486F9D5A" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
<subSubSection id="87671BD4CA74B642AAF06F69A7D87615" pageId="0" pageNumber="145" type="type material">
<paragraph id="576B6D34BF7062F3EF6CB299B3EA7490" pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph id="5B661A38C023F1E6C784EB5654A2377F" pageId="0" pageNumber="145">
<emphasis id="596DCE962F379C0A97CC14DC5AE80F0A" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ and
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis>
<emphasis id="C2FC2A19C29202E1F0029695B3E7B6FF" bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis>
1♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Alborz Province</emphasis>
<emphasis id="59AB063C8F3979DA6FD4AF4837E32D0A" italics="true" pageId="0" pageNumber="145">Alborz Province</emphasis>
: Eshtehard, Jaru,
<geoCoordinate degrees="35" direction="north" minutes="44" orientation="latitude" precision="925" value="35.733334">35°44'N</geoCoordinate>
<geoCoordinate id="5F6C02A3209AD75805951C9A43E85196" degrees="35" direction="north" minutes="44" orientation="latitude" precision="925" value="35.733334">35°44'N</geoCoordinate>
,
<geoCoordinate degrees="50" direction="east" minutes="35" orientation="longitude" precision="925" value="50.583332">50°35'E</geoCoordinate>
<geoCoordinate id="D9AC6272898100BCD28564661B0972E8" degrees="50" direction="east" minutes="35" orientation="longitude" precision="925" value="50.583332">50°35'E</geoCoordinate>
, 10.2018 (A. Zamani).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet refers to the legendary founder of Mycenae and of the Perseid dynasty in Greek mythology. Noun in apposition.</paragraph>
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<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 2.</emphasis>
<subSubSection id="F7BBBD96BA50651BF1DB4A0498E931EC" pageId="0" pageNumber="145" type="etymology">
<paragraph id="23B37499EDB9B3EB05827BCC7BB81175" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="EA3780F47B3CC52CFE115FF1FF9B3E2D" pageId="0" pageNumber="145">The specific epithet refers to the legendary founder of Mycenae and of the Perseid dynasty in Greek mythology. Noun in apposition.</paragraph>
<caption id="5FB1607EB3265A211A3A8617F4AAA8A4" doi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145" start="Figure 2" startId="F2">
<paragraph id="84F050CFCFE1A02C353FA05AEF059E8B" pageId="0" pageNumber="145">
<emphasis id="F28A0BF5C5E14491EB477286B4533FEA" bold="true" pageId="0" pageNumber="145">Figure 2.</emphasis>
Live habitus of male
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana perseus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="perseus">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana perseus</emphasis>
<taxonomicName id="4B73561D9058BB4DBDA2DFF5BA9FB1E8" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana perseus" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="perseus">
<emphasis id="348A24E0AE32407E3D19C771DB43BCE1" italics="true" pageId="0" pageNumber="145">Lachesana perseus</emphasis>
</taxonomicName>
sp. nov. Photo: A. Zamani.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="D3C8686AC86D0A2A76334FAB8B713654" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="1867A39548B62E39870A5278289F4ABC" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="CCC88A0E0DA2B33BA791848E3EFDB688" pageId="0" pageNumber="145">
The new species differs from the similar
<taxonomicName lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
<taxonomicName id="F9FF47EFD8655668402C8AA573876293" lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis id="1C2BF81EE2AEFF95398C842F63411ADF" italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
</taxonomicName>
sp. nov. by the lack of spines in the proximal mesal part of the cymbium (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="19905CC5A9CADFC049A5D93A3568B50B" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
present; cf. Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3C</figureCitation>
<figureCitation id="BA5CFC78108A5251F3376253A21EF5B0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3C</figureCitation>
and
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3H</figureCitation>
<figureCitation id="4F90C4BFA2F9717A812FF7F4D17B36A3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3H</figureCitation>
) as well as by the RTA with the terminal half almost as wide as the tibia (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
<emphasis id="B15485D65F37F0F8620A09913F302C83" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
1.5 times thinner). The shape of the RTA readily distinguishes this species from congeners in the region (stalk abruptly bent basally and tip gently curved; cf. Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4J</figureCitation>
<figureCitation id="CDE4FAF8955D141F79CE0D5DF3613CFB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4J</figureCitation>
and
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4F-I</figureCitation>
<figureCitation id="F86AF3682A06AFE011C0996E5671A9E0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4F-I</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 3.</emphasis>
<caption id="650E2FF432E1235F78C6962C76E711A8" doi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145" start="Figure 3" startId="F3">
<paragraph id="026255F77C6678A7CB3A4CBCAF741D85" pageId="0" pageNumber="145">
<emphasis id="35A66CB4C877C71249C357D8729094A0" bold="true" pageId="0" pageNumber="145">Figure 3.</emphasis>
Male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana kavirensis</emphasis>
<taxonomicName id="CAD3834CEDDF22D8721D1B322464633F" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis">
<emphasis id="7C46872DF48D5AF5264D1D4282C4029F" italics="true" pageId="0" pageNumber="145">Lachesana kavirensis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-E</emphasis>
<emphasis id="ABE0F8F69321D7569651A884CBD048D1" bold="true" pageId="0" pageNumber="145">A-E</emphasis>
) and
<taxonomicName lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
<taxonomicName id="B1F33A7F38F7F04C992B82D8C93F4F70" lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis id="44FB43EDDEF00979BB20EF01CD1F1C53" italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">F-J</emphasis>
<emphasis id="F954BC4F68BE8C099D96B9AD26FBF8FF" bold="true" pageId="0" pageNumber="145">F-J</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, B, G, F</emphasis>
<emphasis id="95DA4D1687553A2F488B59F9DF6B687A" bold="true" pageId="0" pageNumber="145">A, B, G, F</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, H</emphasis>
<emphasis id="4CCECB1D574E73FF823E6FBC3B6A25FB" bold="true" pageId="0" pageNumber="145">C, H</emphasis>
prolateral
<emphasis bold="true" pageId="0" pageNumber="145">D, I</emphasis>
<emphasis id="956B9E31A0DA1D9E0965CB122ABC13FA" bold="true" pageId="0" pageNumber="145">D, I</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">E, J</emphasis>
<emphasis id="4D303E1E3C0C3E5B00F39F9F0C7A159D" bold="true" pageId="0" pageNumber="145">E, J</emphasis>
dorsal.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 4.</emphasis>
<caption id="588C3E20AD42F9763CECA1A3226281BF" doi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145" start="Figure 4" startId="F4">
<paragraph id="A6A7A4CD1F4CBAC06ED810D46474E752" pageId="0" pageNumber="145">
<emphasis id="B9502AFC8FB772A4EDBB022CD71000F7" bold="true" pageId="0" pageNumber="145">Figure 4.</emphasis>
Male palps of
<taxonomicName class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana dyachkovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dyachkovi">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana dyachkovi</emphasis>
<taxonomicName id="D27736380EE4C0AB3C76D7ADCCBE1B46" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana dyachkovi" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dyachkovi">
<emphasis id="38362931D4499FD40FEBB63ACEF3A407" italics="true" pageId="0" pageNumber="145">Lachesana dyachkovi</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="145">A-C, F</emphasis>
<emphasis id="3EDA2D76F7969BB3208532ED6D3BEDF6" bold="true" pageId="0" pageNumber="145">A-C, F</emphasis>
),
<taxonomicName lsidName="L. tarabaevi" pageId="0" pageNumber="145" rank="species" species="tarabaevi">
<emphasis italics="true" pageId="0" pageNumber="145">L. tarabaevi</emphasis>
<taxonomicName id="9131979BBA2C374022B84869146E484C" lsidName="L. tarabaevi" pageId="0" pageNumber="145" rank="species" species="tarabaevi">
<emphasis id="A89A0CA90DE3ECEB716E3B3E6F607108" italics="true" pageId="0" pageNumber="145">L. tarabaevi</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="145">D, E, G</emphasis>
<emphasis id="CE501965A218224D23C9750C650B984A" bold="true" pageId="0" pageNumber="145">D, E, G</emphasis>
),
<taxonomicName lsidName="L. blackwalli" pageId="0" pageNumber="145" rank="species" species="blackwalli">
<emphasis italics="true" pageId="0" pageNumber="145">L. blackwalli</emphasis>
<taxonomicName id="F338C72C67A30B7A5FD7ECECBD9B9375" lsidName="L. blackwalli" pageId="0" pageNumber="145" rank="species" species="blackwalli">
<emphasis id="06B0530BE966B2CFB550DAA511AC782A" italics="true" pageId="0" pageNumber="145">L. blackwalli</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis>
<emphasis id="50D3418A5467E5909F6A82CDEEC57093" bold="true" pageId="0" pageNumber="145">H</emphasis>
),
<taxonomicName lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
<taxonomicName id="7D186755AE93311598C33D1435AE6671" lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis id="7AA444B6117E9DD15CB13BF77D3BC90C" italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">I</emphasis>
<emphasis id="9397F1F9DFAE66F852AF06FD73789FE2" bold="true" pageId="0" pageNumber="145">I</emphasis>
) and
<taxonomicName lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
<taxonomicName id="7994865710A2D22E22E1F9A07C8A139E" lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis id="75822E06EAE3B4BEC2A510D996BC28C2" italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">J</emphasis>
<emphasis id="7C8EB94C80D63B9DA0018C035F7826D5" bold="true" pageId="0" pageNumber="145">J</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis>
<emphasis id="16195BAD1A8BE33A1566FE8DA7753759" bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, C, E</emphasis>
<emphasis id="03C3EBADB84D3515EDD4719D059BC0ED" bold="true" pageId="0" pageNumber="145">B, C, E</emphasis>
dorsal, prolateral and retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">F-J</emphasis>
<emphasis id="30CA170CCCCC699B297BBE8F1139313C" bold="true" pageId="0" pageNumber="145">F-J</emphasis>
retrolateral tibial apophysis
<emphasis bold="true" pageId="0" pageNumber="145">A-G</emphasis>
<emphasis id="07853A964F99B5622D0E38C073A6E3B6" bold="true" pageId="0" pageNumber="145">A-G</emphasis>
reproduced after
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.28.4.08" author="Fomichev, AA" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="556 - 561" refId="B5" refString="Fomichev, AA, Marusik, YM, 2019. A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan. Arthropoda Selecta 28 (4): 556 - 561, DOI: https://doi.org/10.15298/arthsel.28.4.08" title="A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan." url="https://doi.org/10.15298/arthsel.28.4.08" volume="28" year="2019">Fomichev and Marusik (2019)</bibRefCitation>
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis>
<bibRefCitation id="18D230A92D52BB7499C2BA592102C376" DOI="https://doi.org/10.15298/arthsel.28.4.08" author="Fomichev, AA" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="556 - 561" refId="B5" refString="Fomichev, AA, Marusik, YM, 2019. A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan. Arthropoda Selecta 28 (4): 556 - 561, DOI: https://doi.org/10.15298/arthsel.28.4.08" title="A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan." url="https://doi.org/10.15298/arthsel.28.4.08" volume="28" year="2019">Fomichev and Marusik (2019)</bibRefCitation>
<emphasis id="AF2C07ACB48CFE9CC6CF62BC73FB8FCA" bold="true" pageId="0" pageNumber="145">H</emphasis>
illustrated after
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.29.3.06" author="Oezkuetuek, RS" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="334 - 338" refId="B21" refString="Oezkuetuek, RS, Yagmur, EA, Guecel, S, Shafaie, S, Oezden, Oe, Kunt, KB, 2020. New records of the East Mediterranean Lachesana (Aranei: Zodariidae), with description of a new species. Arthropoda Selecta 29 (3): 334 - 338, DOI: https://doi.org/10.15298/arthsel.29.3.06" title="New records of the East Mediterranean Lachesana (Aranei: Zodariidae), with description of a new species." url="https://doi.org/10.15298/arthsel.29.3.06" volume="29" year="2020">
<normalizedToken originalValue="Özkütük">Oezkuetuek</normalizedToken>
<bibRefCitation id="21537E01B4EB997BB5FA093770DC8530" DOI="https://doi.org/10.15298/arthsel.29.3.06" author="Oezkuetuek, RS" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="334 - 338" refId="B21" refString="Oezkuetuek, RS, Yagmur, EA, Guecel, S, Shafaie, S, Oezden, Oe, Kunt, KB, 2020. New records of the East Mediterranean Lachesana (Aranei: Zodariidae), with description of a new species. Arthropoda Selecta 29 (3): 334 - 338, DOI: https://doi.org/10.15298/arthsel.29.3.06" title="New records of the East Mediterranean Lachesana (Aranei: Zodariidae), with description of a new species." url="https://doi.org/10.15298/arthsel.29.3.06" volume="29" year="2020">
<normalizedToken id="47091E4EE797D80FD5B55CC4BDD1E17C" originalValue="Özkütük">Oezkuetuek</normalizedToken>
et al. (2020)
</bibRefCitation>
. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis>
<subSubSection id="7DD39D0C6D50FB64360E66C80F2FC700" pageId="0" pageNumber="145" type="description">
<paragraph id="55D69154750B663AA75DC2002CB98808" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="0304E9E183716FE27F9A7861B7AB9AD7" pageId="0" pageNumber="145">
<emphasis id="C166BD5B5854D7F83BAF18BBB12FC48A" bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1D, E</figureCitation>
<figureCitation id="5B83CF800F2598DDE76BE26DE4C8E73F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1D, E</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live habitus of male Lachesana perseus sp. nov. Photo: A. Zamani." figureDoi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145">2</figureCitation>
<figureCitation id="F440B607D5D6394BC41565849427DB94" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live habitus of male Lachesana perseus sp. nov. Photo: A. Zamani." figureDoi="10.3897/zookeys.1035.65767.figure2" httpUri="https://binary.pensoft.net/fig/536879" pageId="0" pageNumber="145">2</figureCitation>
. Total length 10.50. Carapace 5.60 long, 2.60 wide at pars cephalica, 3.88 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.19, ALE: 0.19, PME: 0.16, PLE: 0.19, PME-PME: 0.18. Carapace, sternum, labium, chelicerae and maxillae light brown. Chelicera (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1F</figureCitation>
<figureCitation id="AF24280057BC0A00E04CB022E2589ACE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1F</figureCitation>
) with dense, black setae and one promarginal tooth; fangs curved. Legs the same color as carapace, without annulations and with numerous spines. Abdomen pale, dorsally with a longitudinal dark stripe and several large spots. Spinnerets grayish, uniform in color. Measurements of legs: I: 16.42 (4.69, 2.02, 3.61, 3.56, 2.54), II: 15.48 (4.22, 2.01, 3.19, 3.53, 2.53), III: 16.25 (4.46, 1.96, 2.51, 4.66, 2.66), IV: 19.39 (4.87, 2.32, 3.29, 5.66, 3.25).
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="9516173D2BFC2942A4C7EDB07C862D94" pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3F-J</figureCitation>
<figureCitation id="18B9EF623E50DC3572D19CB0D9A4AD52" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3F-J</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4J</figureCitation>
<figureCitation id="0AEA5A9D78300C46729788416DC388B3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4J</figureCitation>
. Tibial apophysis more than 1.2 times longer than tibia, widest part over 2 times wider than stalk; cymbium lacking spines in proximal mesal part; bulb elongate, 1.3 times longer than wide; base of embolus large, 1/3 of the tegulum's length; embolus filamentous, lying in chute formed by conductor.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
<paragraph id="A5183832E79F7D3898AB82AE58E125CB" pageId="0" pageNumber="145">
<emphasis id="8C73054740359E8ABB48768F6C76EEDD" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="5629D4B1DA20B13B4946C451DE16D0CC" pageId="0" pageNumber="145" type="distribution">
<paragraph id="222616A642345930D5BB0A41A0DA34B3" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="571ADD87054897BA34C3655C2A7A8834" pageId="0" pageNumber="145">
Known only from the type locality in Alborz Province, northern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
<figureCitation id="67C10143A0799B0656F0F1845C3F7A5C" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>