Add updates up until 2024-11-29 17:06:45
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:title id="FF255353734DED2D98DC3B95A1995253">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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<mods:namePart>Zamani, Alireza</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8084-9666</mods:nameIdentifier>
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<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
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<mods:nameIdentifier type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A1CFA4FCAA6A3F295A2710B4782C7B13">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="82592BE4002A51F2687556C6DC5E1153">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
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<mods:namePart>Marusik, Yuri M.</mods:namePart>
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia & Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<mods:namePart id="40F30FCADF0160790787DD721DB1793C">Marusik, Yuri M.</mods:namePart>
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<treatment ID-GBIF-Taxon="181753572" LSID="urn:lsid:zoobank.org:act:9DEB48F2-08A4-482E-AEFF-405D9AAAA248" httpUri="http://treatment.plazi.org/id/09834123329357E49008CCADEDB4806A" lastPageNumber="145" pageId="0" pageNumber="145">
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<taxonomicName LSID="http://zoobank.org/9DEB48F2-08A4-482E-AEFF-405D9AAAA248" authority="Zamani & Marusik, 2021" authorityName="Zamani & Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila" status="sp. nov.">Pax leila</taxonomicName>
|
||||
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">Figs 5C, D</figureCitation>
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">, 8</figureCitation>
|
||||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">, 9</figureCitation>
|
||||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">, 10F-J</figureCitation>
|
||||
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
|
||||
<treatment id="09834123329357E49008CCADEDB4806A" ID-GBIF-Taxon="181753572" LSID="urn:lsid:zoobank.org:act:9DEB48F2-08A4-482E-AEFF-405D9AAAA248" httpUri="http://treatment.plazi.org/id/09834123329357E49008CCADEDB4806A" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Storeninae">
|
||||
<subSubSection id="A548E98B5BF32F05558326154E8C36D9" pageId="0" pageNumber="145" type="nomenclature">
|
||||
<paragraph id="51B3461CAB9433631D2937A742852474" pageId="0" pageNumber="145">
|
||||
<taxonomicName id="2AE6673BC1401BE60225688D592ECDDC" ID-CoL="98YZ7" LSID="http://zoobank.org/9DEB48F2-08A4-482E-AEFF-405D9AAAA248" authority="Zamani & Marusik, 2021" authorityName="Zamani & Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila" status="sp. nov.">Pax leila</taxonomicName>
|
||||
<taxonomicNameLabel id="95FDE61E3CAE8BD7840F68654D255665" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
|
||||
<figureCitation id="5A2C9A11B4F60C5FCF176709C3DC9950" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">Figs 5C, D</figureCitation>
|
||||
<figureCitation id="A1D4B4C4F1ADB760BFACBDC250ABBE47" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">, 8</figureCitation>
|
||||
<figureCitation id="5DFA95A2A776C7D0F6DAA863ED8DA71F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">, 9</figureCitation>
|
||||
<figureCitation id="739E51E13DAD97771AC0B89180FB47B8" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">, 10F-J</figureCitation>
|
||||
<figureCitation id="6E481C54DBF5EFEEFC3EFFFEB5E9FFD0" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="145" type="type material">
|
||||
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="145">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
|
||||
<subSubSection id="FA777627746326AA3AA1688C50FB90C2" pageId="0" pageNumber="145" type="type material">
|
||||
<paragraph id="C070B08018403A6BCB6B0F55E791C79A" pageId="0" pageNumber="145">Type material.</paragraph>
|
||||
<paragraph id="0D2D9333DCC2EE79B57E480508A4DE18" pageId="0" pageNumber="145">
|
||||
<emphasis id="F45790FB34322E8B205E86073A8E878D" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
|
||||
♂ and
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||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis>
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<emphasis id="6CAB6126CA078587D15355101B173BC3" bold="true" italics="true" pageId="0" pageNumber="145">paratype</emphasis>
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1♀ (MHNG), Iran:
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<emphasis italics="true" pageId="0" pageNumber="145">Fars Province</emphasis>
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<emphasis id="76BA2AF8D2599E86CE055840E33F8B70" italics="true" pageId="0" pageNumber="145">Fars Province</emphasis>
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||||
: road to Yasuj,
|
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<geoCoordinate degrees="30" direction="north" minutes="28" orientation="latitude" precision="925" value="30.466667">30°28'N</geoCoordinate>
|
||||
<geoCoordinate id="B54ABE06A265D8118A97710001BA370B" degrees="30" direction="north" minutes="28" orientation="latitude" precision="925" value="30.466667">30°28'N</geoCoordinate>
|
||||
,
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||||
<geoCoordinate degrees="51" direction="east" minutes="30" orientation="longitude" precision="925" value="51.5">51°30'E</geoCoordinate>
|
||||
<geoCoordinate id="011833FB1319D5E8D350FC766577CD85" degrees="51" direction="east" minutes="30" orientation="longitude" precision="925" value="51.5">51°30'E</geoCoordinate>
|
||||
, 25.05.1974 (A. Senglet).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="145" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="145">The specific epithet is a feminine given name in the Persian language, meaning "daughter of the night". Noun in apposition.</paragraph>
|
||||
<subSubSection id="4ED5441780D1374CB0C48EF52CA86BDB" pageId="0" pageNumber="145" type="etymology">
|
||||
<paragraph id="D51D30BA473B006B9373EC1F5A99CE7A" pageId="0" pageNumber="145">Etymology.</paragraph>
|
||||
<paragraph id="4E51DEEBF696DA81FE5F3D1F1F15AF38" pageId="0" pageNumber="145">The specific epithet is a feminine given name in the Persian language, meaning "daughter of the night". Noun in apposition.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="145">
|
||||
<subSubSection id="4C6E036F3BB0C13E9083C2750C2F436D" pageId="0" pageNumber="145" type="diagnosis">
|
||||
<paragraph id="D01350BE9F66B49B71DF8D83E107406A" pageId="0" pageNumber="145">Diagnosis.</paragraph>
|
||||
<paragraph id="C84CD4E99DE8B4FB544C491CA84F5848" pageId="0" pageNumber="145">
|
||||
The new species differs from all congeners by having a pair of spine-like cymbial outgrowths (Figs
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">8</figureCitation>
|
||||
<figureCitation id="98B655D74E64C078F1168E438F370CA2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">8</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">9C, E</figureCitation>
|
||||
<figureCitation id="652433934DEE6D723C2CD5E44A2EF15D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">9C, E</figureCitation>
|
||||
) (
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
|
||||
<emphasis id="CF42DCA06FA17C8D0709A0AB601ADBE7" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
|
||||
lacking) and a triangular epigynal median plate (
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
|
||||
<emphasis id="27F6AC7D2B5202B236A7BE366003E581" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
|
||||
absent or rectangular).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145" start="Figure 8" startId="F8">
|
||||
<paragraph pageId="0" pageNumber="145">
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 8.</emphasis>
|
||||
<caption id="C448ADE7E1135D19E0E32EA8E1C922C2" doi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145" start="Figure 8" startId="F8">
|
||||
<paragraph id="D37620CFE35199C30125B543D8469984" pageId="0" pageNumber="145">
|
||||
<emphasis id="7EAE1F8E57B854F60C4CED2234D6CD54" bold="true" pageId="0" pageNumber="145">Figure 8.</emphasis>
|
||||
Male palp of
|
||||
<taxonomicName authorityName="Zamani & Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila">
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">Pax leila</emphasis>
|
||||
<taxonomicName id="9C4DADAE36CA6C14EEB7798770C346F6" authorityName="Zamani & Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila">
|
||||
<emphasis id="0B305F4FD2CE5A844A834C54087FBC90" italics="true" pageId="0" pageNumber="145">Pax leila</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||||
<emphasis id="52AF5D2BF2EF266E2235D2B4A4480CE5" bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||||
proventral
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
|
||||
<emphasis id="165B7CC20AE8F37D1DFF9F13EC7F596D" bold="true" pageId="0" pageNumber="145">B</emphasis>
|
||||
ventral
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
|
||||
<emphasis id="F3B1907D8EC0DAC76B1DC8ED6BCD3BBF" bold="true" pageId="0" pageNumber="145">C</emphasis>
|
||||
ventroretrolateral
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
|
||||
<emphasis id="CDFDB57750CA1A33167FE39424DA8B99" bold="true" pageId="0" pageNumber="145">D</emphasis>
|
||||
prolateral
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
|
||||
<emphasis id="4DC1B840D415D6338D0ADE753EC6C5CE" bold="true" pageId="0" pageNumber="145">E</emphasis>
|
||||
dorsal
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
|
||||
<emphasis id="746417865834C56F4B14565D44FF79E5" bold="true" pageId="0" pageNumber="145">F</emphasis>
|
||||
retrolateral. Scale bars: 0.2 mm, unless stated otherwise.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="145" type="description">
|
||||
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="145">
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
|
||||
<subSubSection id="331289FAAEE58BC5B91C32D445903BF1" pageId="0" pageNumber="145" type="description">
|
||||
<paragraph id="C5369750918E42EC43D2ECB09A88FAEE" pageId="0" pageNumber="145">Description.</paragraph>
|
||||
<paragraph id="644A209423ED7521A6DAAB2F56708660" pageId="0" pageNumber="145">
|
||||
<emphasis id="2806484CF0771CBE32C9FD1F9EEBFFFB" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
|
||||
Habitus as in Fig.
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5C</figureCitation>
|
||||
<figureCitation id="8B110704F23D11AA905653FDBBFBF08B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5C</figureCitation>
|
||||
. Total length 4.42. Carapace 2.07 long, 1.03 wide at pars cephalica, 1.46 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.09, ALE: 0.09, PME: 0.06, PLE: 0.09, PME-PME: 0.07. Carapace, sternum, labium, chelicerae and maxillae reddish brown, without any pattern. Chelicera with 2 promarginal teeth. Legs yellow, with few spines and without annulations. Abdomen dark brown, dorsally with large scutum covering 2/3 of the abdomen. Spinnerets pale, uniform in color. Measurements of legs: I: 4.51 (1.21, 0.50, 1.06, 0.93, 0.81), II: 3.89 (1.04, 0.52, 0.79, 0.83, 0.71), III: 3.62 (0.98, 0.50, 0.69, 0.85, 0.60), IV: 4.99 (1.33, 0.58, 1.05, 1.33, 0.70).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="145">
|
||||
<paragraph id="19CADD542F51ABE823B88F014581B96F" pageId="0" pageNumber="145">
|
||||
Palp as in Figs
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">8A-F</figureCitation>
|
||||
<figureCitation id="45BF8077FE0548D1BCA6D47FA3F42655" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male palp of Pax leila sp. nov. A proventral B ventral C ventroretrolateral D prolateral E dorsal F retrolateral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure8" httpUri="https://binary.pensoft.net/fig/536885" pageId="0" pageNumber="145">8A-F</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">9A-E</figureCitation>
|
||||
<figureCitation id="35F512F44E81CEA4B087B0280C75F515" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of the male palp of Pax leila sp. nov. A apicoventral B ventroretrolateral C prodorsal D ventral E retrodorsal. Abbreviations: Cp - prolateral cymbial process, Cr - retrolateral cymbial process, Da - dorsal apophysis, Ma - median apophysis, Se - spine of embolus base. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145">9A-E</figureCitation>
|
||||
. Tibia wider than long, with short apophysis; cymbium with 2 strong spine-like baso-posterior outgrowths (
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">Cp</emphasis>
|
||||
<emphasis id="B11777083988408CD5AAECF53B9943C0" italics="true" pageId="0" pageNumber="145">Cp</emphasis>
|
||||
,
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">Cr</emphasis>
|
||||
<emphasis id="EC1C8CBEA463F34317A5E9DE75B3F760" italics="true" pageId="0" pageNumber="145">Cr</emphasis>
|
||||
), lateral fold lacking; embolus long, with robust triangular outgrowth near base (
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">Se</emphasis>
|
||||
<emphasis id="1D1AB9F10151F8EA863238A1BE6557F0" italics="true" pageId="0" pageNumber="145">Se</emphasis>
|
||||
); median apophysis (
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
|
||||
<emphasis id="2F34EE9F12F3E8BFD1282E2BCCC25080" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
|
||||
) very large, with 2 claw-like extensions; conductor small and strongly sclerotized.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145" start="Figure 9" startId="F9">
|
||||
<paragraph pageId="0" pageNumber="145">
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 9.</emphasis>
|
||||
<caption id="AB5F3B3E382925E0FFCED6505E966E61" doi="10.3897/zookeys.1035.65767.figure9" httpUri="https://binary.pensoft.net/fig/536886" pageId="0" pageNumber="145" start="Figure 9" startId="F9">
|
||||
<paragraph id="D9F0744620903DD21D67AC88FC4BFC9C" pageId="0" pageNumber="145">
|
||||
<emphasis id="49DB0536F2B6BFE2349C8BF91C60E2DC" bold="true" pageId="0" pageNumber="145">Figure 9.</emphasis>
|
||||
SEM images of the male palp of
|
||||
<taxonomicName authorityName="Zamani & Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila">
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">Pax leila</emphasis>
|
||||
<taxonomicName id="A54BA2C6573765749463CBD9A0F67663" authorityName="Zamani & Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax leila" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leila">
|
||||
<emphasis id="4F50C05BAE0A78165AD08B978857EC93" italics="true" pageId="0" pageNumber="145">Pax leila</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||||
<emphasis id="3F9916DBB667F91B49C837670A83B4D6" bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||||
apicoventral
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
|
||||
<emphasis id="4E30F043477EF83D7E191B75F426B0FB" bold="true" pageId="0" pageNumber="145">B</emphasis>
|
||||
ventroretrolateral
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
|
||||
<emphasis id="E0CE82CAE995D85A0A75C36858801F9A" bold="true" pageId="0" pageNumber="145">C</emphasis>
|
||||
prodorsal
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
|
||||
<emphasis id="20F1FE821B4DBDEC8DF54D356D1F1EBB" bold="true" pageId="0" pageNumber="145">D</emphasis>
|
||||
ventral
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
|
||||
<emphasis id="FF5B27286CDDF0ADC258D0EEF7C58D19" bold="true" pageId="0" pageNumber="145">E</emphasis>
|
||||
retrodorsal. Abbreviations:
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">Cp</emphasis>
|
||||
<emphasis id="A079B523888F130DC7BF015E41C13A45" italics="true" pageId="0" pageNumber="145">Cp</emphasis>
|
||||
- prolateral cymbial process,
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">Cr</emphasis>
|
||||
<emphasis id="DEE48351EB109074B5381A71F561E781" italics="true" pageId="0" pageNumber="145">Cr</emphasis>
|
||||
- retrolateral cymbial process,
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">Da</emphasis>
|
||||
<emphasis id="579CEBD106D87CCC235830E4CB563A0A" italics="true" pageId="0" pageNumber="145">Da</emphasis>
|
||||
- dorsal apophysis,
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
|
||||
<emphasis id="711B0A4EED352E17F7A04EB4013B71A3" italics="true" pageId="0" pageNumber="145">Ma</emphasis>
|
||||
- median apophysis,
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">Se</emphasis>
|
||||
<emphasis id="3352366A9EC96F193065E824CDA97085" italics="true" pageId="0" pageNumber="145">Se</emphasis>
|
||||
- spine of embolus base. Scale bars: 0.1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph pageId="0" pageNumber="145">
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
|
||||
<paragraph id="942FED49D47F484ED2761D73697F641B" pageId="0" pageNumber="145">
|
||||
<emphasis id="114C8D63694102AF62B9ED89D14E282B" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
|
||||
Habitus as in Fig.
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5D</figureCitation>
|
||||
<figureCitation id="0CCFBC148C0036178772F63F1509504A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5D</figureCitation>
|
||||
. Total length 4.95. Carapace 1.97 long, 1.00 wide at pars cephalica, 1.31 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.07, ALE: 0.08, PME: 0.06, PLE: 0.07, PME-PME: 0.08. Coloration as in male, with lighter abdomen lacking a scutum. Measurements of legs: I: 4.25 (1.21, 0.50, 0.99, 0.84, 0.71), II: 3.58 (0.94, 0.52, 0.74, 0.74, 0.64), III: 3.47 (0.90, 0.51, 0.70, 0.81, 0.55), IV: 4.71 (1.18, 0.53, 1.05, 1.21, 0.74).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="145">
|
||||
<paragraph id="68AA08A588CA6B32860A9EF6E7699DC2" pageId="0" pageNumber="145">
|
||||
Epigyne as in Fig.
|
||||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">10F-J</figureCitation>
|
||||
<figureCitation id="48C364A8470D174C535E92B421B88A45" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">10F-J</figureCitation>
|
||||
. Epigyne trilobate, with triangular median plate, posteriorly as wide as lateral lobes; copulatory openings slit-like, broad; copulatory ducts wide, as wide as receptacles; receptacles globular, separated by less than their radii.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145" start="Figure 10" startId="F10">
|
||||
<paragraph pageId="0" pageNumber="145">
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 10.</emphasis>
|
||||
<caption id="37185AAEE15756DCCB2FA4E70BC21A99" doi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145" start="Figure 10" startId="F10">
|
||||
<paragraph id="B9B355F792377E7AD6083EF71BD87A19" pageId="0" pageNumber="145">
|
||||
<emphasis id="B1B4D3E39976DE0177B82AB01AE26C27" bold="true" pageId="0" pageNumber="145">Figure 10.</emphasis>
|
||||
Epigynes of
|
||||
<taxonomicName authorityName="Zamani & Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
|
||||
<taxonomicName id="2FE37A265797936D5130B198CF4CC489" authorityName="Zamani & Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
|
||||
<emphasis id="C2DDF008B5B86FA9F0F6431782F1F901" italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. (
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">A-E</emphasis>
|
||||
<emphasis id="459C89D9239C9FF21C7A2C6A02FED11A" bold="true" pageId="0" pageNumber="145">A-E</emphasis>
|
||||
) and
|
||||
<taxonomicName lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila">
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
|
||||
<taxonomicName id="F7A1C8605D9F21BA15C06FFF13FC46B1" lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila">
|
||||
<emphasis id="2246B5D492F3C8E7D3AE0D0323B509C5" italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. (
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">F-J</emphasis>
|
||||
<emphasis id="EF9BB29F2C2203253F286F8A4DFABC94" bold="true" pageId="0" pageNumber="145">F-J</emphasis>
|
||||
)
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">A, F</emphasis>
|
||||
<emphasis id="0D874B34DDA34980B6BB3BA459935F4A" bold="true" pageId="0" pageNumber="145">A, F</emphasis>
|
||||
intact, ventral
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">B, J</emphasis>
|
||||
<emphasis id="7DDF6F8A2F76A296832DA3B079272E3B" bold="true" pageId="0" pageNumber="145">B, J</emphasis>
|
||||
posterior
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">C, G</emphasis>
|
||||
<emphasis id="9D4A03E34AC3F83D104F4EB2CC10A63C" bold="true" pageId="0" pageNumber="145">C, G</emphasis>
|
||||
macerated, ventral
|
||||
<emphasis bold="true" pageId="0" pageNumber="145">D, E, H, I</emphasis>
|
||||
<emphasis id="8CEA8F5430C18508630D9ACD4608DC17" bold="true" pageId="0" pageNumber="145">D, E, H, I</emphasis>
|
||||
macerated, dorsal. Scale bars: 0.2 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="145" type="comments">
|
||||
<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="145">
|
||||
<subSubSection id="E2BF68B596F34C6894A43572B0168470" pageId="0" pageNumber="145" type="comments">
|
||||
<paragraph id="2785376854BC6704A1D632D63FF70B48" pageId="0" pageNumber="145">Comments.</paragraph>
|
||||
<paragraph id="E36EC4EA81645698AD413ECE8BA6A072" pageId="0" pageNumber="145">
|
||||
Although this species is rather similar in general appearance to the generotype and the other species known from Iran, the bulb and cymbium conformation is very different from those of other species considered in
|
||||
<taxonomicName authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">Pax</emphasis>
|
||||
<taxonomicName id="26A98999BC44AEFBEE5E892EC4859534" authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="F084086178C6FAD3DD2BF150E9FCA8AE" italics="true" pageId="0" pageNumber="145">Pax</emphasis>
|
||||
</taxonomicName>
|
||||
and most likely belongs to an undescribed genus.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="145" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="145">
|
||||
<subSubSection id="D5107F62298DDB995517B1CE34077CA2" pageId="0" pageNumber="145" type="distribution">
|
||||
<paragraph id="81FDC14758040DF14F4BA504FE4F71D8" pageId="0" pageNumber="145">Distribution.</paragraph>
|
||||
<paragraph id="09AD0C2B35941441BC7654607FCA8E26" pageId="0" pageNumber="145">
|
||||
Known only from the type locality in Fars Province, southwestern Iran (Fig.
|
||||
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
|
||||
<figureCitation id="394150865ED8F5BDB3ED207BF2F3353E" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
|
||||
). This is the easternmost record of the genus
|
||||
<taxonomicName authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="145">Pax</emphasis>
|
||||
<taxonomicName id="0F202F09DCB601B369C896E48AFCFD18" authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="E1A395D5243E1516E653F3F45C6B91FB" italics="true" pageId="0" pageNumber="145">Pax</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
|
|
|
|||
Loading…
Add table
Add a link
Reference in a new issue