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<document id="7CF3EA4A9E0240EDF758E326271BD8F5" ID-DOI="10.11646/zootaxa.5538.4.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="14239597" ID-ZooBank="A9538D6B-FB28-44B3-B29F-6C5FD79FBAF8" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732829322446" checkinUser="plazi" docAuthor="Park, Jaekook &amp; Jung, Sunghoon" docDate="2024" docId="03BEC82B6922FF83FF0DFE3F13D310F3" docLanguage="en" docName="zootaxa.5538.4.5.pdf" docOrigin="Zootaxa 5538 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5538.4.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Toyoides bilineata Park &amp; Jung 2024, comb. n." docType="treatment" docVersion="1" lastPageNumber="378" masterDocId="FF87B0536926FF84FF9AFFE513091436" masterDocTitle="The genus Toyoides Matsumura (Hemiptera: Auchenorrhyncha: Delphacidae): new combination and lectotype designations" masterLastPageNumber="380" masterPageNumber="371" pageNumber="375" updateTime="1733172059991" updateUser="felipe" zenodo-license-document="CLOSED">
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<mods:title id="E0914DA633A13B837FA063628312B2F7">The genus Toyoides Matsumura (Hemiptera: Auchenorrhyncha: Delphacidae): new combination and lectotype designations</mods:title>
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<mods:namePart id="770543C3624B5FD32339FB5B7C3F3373">Park, Jaekook</mods:namePart>
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<mods:affiliation id="CC069C04299EE8264E5ED7B0F779D6B1">Laboratory of Systematic Entomology, Department of Applied Biology, College of Agriculture &amp; Life Sciences, Chungnam National University, Daejeon, Republic of Korea. &amp; Department of Smart Agriculture Systems, College of Agriculture &amp; Life Sciences, Chungnam National University, Daejeon, Republic of Korea.</mods:affiliation>
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<mods:namePart id="0BC07B65FDB2A015FB387B586B3917ED">Jung, Sunghoon</mods:namePart>
<mods:affiliation id="1F2647FE062E02712614EC9861C53D5B">Laboratory of Systematic Entomology, Department of Applied Biology, College of Agriculture &amp; Life Sciences, Chungnam National University, Daejeon, Republic of Korea. &amp; Department of Smart Agriculture Systems, College of Agriculture &amp; Life Sciences, Chungnam National University, Daejeon, Republic of Korea.</mods:affiliation>
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<subSubSection id="C30D2AB66922FF80FF0DFE3F11D515C2" box="[151,732,474,501]" pageId="4" pageNumber="375" type="nomenclature">
<paragraph id="8BA8793D6922FF80FF0DFE3F11D515C2" blockId="4.[151,732,474,536]" box="[151,732,474,501]" pageId="4" pageNumber="375">
<emphasis id="B963A52F6922FF80FF0DFE3F11D515C2" bold="true" box="[151,732,474,501]" pageId="4" pageNumber="375">
<taxonomicName id="4C1702BE6922FF80FF0DFE3F116315C2" authority="(Matsumura, 1935)" authorityName="Park &amp; Jung" authorityYear="2024" baseAuthorityName="Matsumura" baseAuthorityYear="1935" box="[151,618,474,501]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="bilineata" status="comb. nov.">
<emphasis id="B963A52F6922FF80FF0DFE3F127815C3" bold="true" box="[151,369,474,501]" italics="true" pageId="4" pageNumber="375">Toyoides bilineata</emphasis>
(
<bibRefCitation id="EF8604CC6922FF80FE1AFE3F116815C2" author="Matsumura, S." box="[384,609,474,500]" pageId="4" pageNumber="375" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25018546922FF80FDEBFE3F11D515C2" box="[625,732,474,500]" pageId="4" pageNumber="375" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66922FF80FF0DFE1B12C2162E" box="[151,459,510,536]" pageId="4" pageNumber="375" type="description">
<paragraph id="8BA8793D6922FF80FF0DFE1B12C2162E" blockId="4.[151,732,474,536]" box="[151,459,510,536]" pageId="4" pageNumber="375">
(
<figureCitation id="132C65B86922FF80FF05FE1B1216162E" box="[159,287,510,536]" captionStart="FIGURE 2" captionStartId="4.[151,250,1653,1677]" captionTargetBox="[151,1431,678,1626]" captionTargetId="figure-252@4.[151,1436,669,1635]" captionTargetPageId="4" captionText="FIGURE 2. Lectotype and paralectotype of Toyoides bilineata Matsumura comb. n. A. habitus, dorsal view; B. lectotype, habitus, lateral view; C. ditto, frons, ventral view; D. ditto, right hind leg with spur; E. lectotype and paralectotype labels. Abbreviations: f, front; b, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239601" httpUri="https://zenodo.org/record/14239601/files/figure.png" pageId="4" pageNumber="375">Figs 2AD</figureCitation>
,
<figureCitation id="132C65B86922FF80FEB0FE1B1278162E" box="[298,369,510,536]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="4" pageNumber="375">3AG</figureCitation>
,
<figureCitation id="132C65B86922FF80FEE7FE1B12CA162E" box="[381,451,510,536]" captionStart="FIGURE 4" captionStartId="6.[151,250,1530,1554]" captionTargetBox="[158,1429,188,1499]" captionTargetId="figure-20@6.[151,1436,181,1506]" captionTargetPageId="6" captionText="FIGURE 4. Male genitalia of Toyoides bilineata Matsumura comb. n. A. genitalia, caudal view; B. ditto, lateral view; C. anal segment, caudal view; D. ditto, lateral view; E. pygofer, caudal view; F. ditto, lateral view; G. genital styles, caudoventral view; H. ditto, lateral view; I. aedeagus with suspensorium, left-lateral view; J. aedeagus without suspensorium, dorsal view; K. suspensorium. Scale bar: AB, EF: 0.5 mm; CD, GH: 0.3 mm; IK: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239605" httpUri="https://zenodo.org/record/14239605/files/figure.png" pageId="4" pageNumber="375">4AK</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66922FF80FF0DFDA6111B164F" pageId="4" pageNumber="375" type="reference_group">
<paragraph id="8BA8793D6922FF80FF0DFDA61125166C" blockId="4.[151,556,578,633]" box="[151,556,578,603]" pageId="4" pageNumber="375">
<treatmentCitationGroup id="AB075E136922FF80FF0DFDA61125166C" box="[151,556,578,603]" pageId="4" pageNumber="375">
<treatmentCitation id="0AB65F2C6922FF80FF0DFDA6112E166C" author="Matsumura, S." box="[151,551,578,603]" page="77" pageId="4" pageNumber="375" year="1935">
<taxonomicName id="4C1702BE6922FF80FF0DFDA6112E166C" ID-CoL="6FYTL" authority="Matsumura, 1935: 77" authorityName="Matsumura" authorityPageNumber="77" authorityYear="1935" box="[151,551,578,603]" class="Insecta" family="Delphacidae" genus="Epunka" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="bilineata">
<emphasis id="B963A52F6922FF80FF0DFDA6124C166D" box="[151,325,579,603]" italics="true" pageId="4" pageNumber="375">Epunka bilineata</emphasis>
<bibRefCitation id="EF8604CC6922FF80FED1FDA6112E166C" author="Matsumura, S." box="[331,551,578,602]" pageId="4" pageNumber="375" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935: 77</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA8793D6922FF80FF0DFD84111B164F" blockId="4.[151,556,578,633]" box="[151,530,609,633]" pageId="4" pageNumber="375">
<treatmentCitationGroup id="AB075E136922FF80FF0DFD84111B164F" box="[151,530,609,633]" pageId="4" pageNumber="375">
<treatmentCitation id="0AB65F2C6922FF80FF0DFD841107164F" author="Ishihara, T." box="[151,526,609,633]" page="34" pageId="4" pageNumber="375" year="1949">
<taxonomicName id="4C1702BE6922FF80FF0DFD841107164F" authority="Ishihara, 1949: 34" authorityName="Ishihara" authorityPageNumber="34" authorityYear="1949" box="[151,526,609,633]" class="Insecta" family="Delphacidae" genus="Euidella" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="bilineata">
<emphasis id="B963A52F6922FF80FF0DFD841244164F" box="[151,333,609,633]" italics="true" pageId="4" pageNumber="375">Euidella bilineata</emphasis>
<bibRefCitation id="EF8604CC6922FF80FECEFD841107164F" author="Ishihara, T." box="[340,526,609,633]" pageId="4" pageNumber="375" pagination="1 - 102" refId="ref5544" refString="Ishihara, T. (1949) Revision of the Araeopidae of Japan, Ryukyu Island and Formosa (Hemiptera). Scientific Reports of the Matsuyama Agricultural College, 2, 1 - 102." type="journal article" year="1949">Ishihara, 1949: 34</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
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<caption id="DF6829B56922FF80FF0DF990107912E3" ID-DOI="http://doi.org/10.5281/zenodo.14239601" ID-Zenodo-Dep="14239601" httpUri="https://zenodo.org/record/14239601/files/figure.png" pageId="4" pageNumber="375" startId="4.[151,250,1653,1677]" targetBox="[151,1431,678,1626]" targetPageId="4" targetType="figure">
<paragraph id="8BA8793D6922FF80FF0DF990107912E3" blockId="4.[151,1437,1653,1750]" pageId="4" pageNumber="375">
<emphasis id="B963A52F6922FF80FF0DF990121F12BB" bold="true" box="[151,278,1653,1677]" pageId="4" pageNumber="375">FIGURE 2.</emphasis>
Lectotype and paralectotype of
<taxonomicName id="4C1702BE6922FF80FDF6F99010AD12BB" authority="Matsumura" authorityName="Matsumura" authorityYear="2024" baseAuthorityName="Park &amp; Jung" baseAuthorityYear="2024" box="[620,932,1653,1677]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="bilineata" status="comb. nov.">
<emphasis id="B963A52F6922FF80FDF6F990102112BB" box="[620,808,1653,1677]" italics="true" pageId="4" pageNumber="375">Toyoides bilineata</emphasis>
Matsumura
</taxonomicName>
<emphasis id="B963A52F6922FF80FC34F990170412BB" bold="true" box="[942,1037,1653,1677]" pageId="4" pageNumber="375">
<taxonomicNameLabel id="A25018546922FF80FC34F990170412BB" box="[942,1037,1653,1677]" pageId="4" pageNumber="375" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
A. habitus, dorsal view; B. lectotype, habitus, lateral view; C.
<emphasis id="B963A52F6922FF80FE00F97C12C11287" box="[410,456,1689,1713]" italics="true" pageId="4" pageNumber="375">ditto</emphasis>
, frons, ventral view; D.
<emphasis id="B963A52F6922FF80FD55F97C11F41287" box="[719,765,1689,1713]" italics="true" pageId="4" pageNumber="375">ditto</emphasis>
, right hind leg with spur; E. lectotype and paralectotype labels. Abbreviations:
<emphasis id="B963A52F6922FF80FEAEF958123212E3" box="[308,315,1725,1749]" italics="true" pageId="4" pageNumber="375">f</emphasis>
, front;
<emphasis id="B963A52F6922FF80FE1CF958129A12E3" box="[390,403,1725,1749]" italics="true" pageId="4" pageNumber="375">b</emphasis>
, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C30D2AB66922FF80FF0DF8C216951398" pageId="4" pageNumber="375" type="diagnosis">
<paragraph id="8BA8793D6922FF80FF0DF8C216951398" blockId="4.[151,1436,1831,1966]" pageId="4" pageNumber="375">
<emphasis id="B963A52F6922FF80FF0DF8C2121B1377" bold="true" box="[151,274,1831,1857]" pageId="4" pageNumber="375">Diagnosis.</emphasis>
Recognized by body light brown; tegmina with fuscous stripes visibly; vertex with blackish marking medially; pronotum, mesonotum and prosternum with a pair of parallel blackish markings (
<figureCitation id="132C65B86922FF80FB33F8AE16241350" box="[1193,1325,1867,1894]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="4" pageNumber="375">Figs 3AG</figureCitation>
); tegmina smoothly curved (
<figureCitation id="132C65B86922FF80FEFAF88A12B013BC" box="[352,441,1903,1930]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="4" pageNumber="375">Fig. 3D</figureCitation>
); frons with median carinae forked above level of lower margin of eyes (
<figureCitation id="132C65B86922FF80FB46F88A163B13BC" box="[1244,1330,1903,1930]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="4" pageNumber="375">Fig. 3E</figureCitation>
); pygofer without medioventral structures (
<figureCitation id="132C65B86922FF80FD88F87611861398" box="[530,655,1939,1966]" captionStart="FIGURE 4" captionStartId="6.[151,250,1530,1554]" captionTargetBox="[158,1429,188,1499]" captionTargetId="figure-20@6.[151,1436,181,1506]" captionTargetPageId="6" captionText="FIGURE 4. Male genitalia of Toyoides bilineata Matsumura comb. n. A. genitalia, caudal view; B. ditto, lateral view; C. anal segment, caudal view; D. ditto, lateral view; E. pygofer, caudal view; F. ditto, lateral view; G. genital styles, caudoventral view; H. ditto, lateral view; I. aedeagus with suspensorium, left-lateral view; J. aedeagus without suspensorium, dorsal view; K. suspensorium. Scale bar: AB, EF: 0.5 mm; CD, GH: 0.3 mm; IK: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239605" httpUri="https://zenodo.org/record/14239605/files/figure.png" pageId="4" pageNumber="375">Figs 4EF</figureCitation>
); aedeagus branched medially, division with small spinule (
<figureCitation id="132C65B86922FF80FAA4F87616861398" box="[1342,1423,1939,1966]" captionStart="FIGURE 4" captionStartId="6.[151,250,1530,1554]" captionTargetBox="[158,1429,188,1499]" captionTargetId="figure-20@6.[151,1436,181,1506]" captionTargetPageId="6" captionText="FIGURE 4. Male genitalia of Toyoides bilineata Matsumura comb. n. A. genitalia, caudal view; B. ditto, lateral view; C. anal segment, caudal view; D. ditto, lateral view; E. pygofer, caudal view; F. ditto, lateral view; G. genital styles, caudoventral view; H. ditto, lateral view; I. aedeagus with suspensorium, left-lateral view; J. aedeagus without suspensorium, dorsal view; K. suspensorium. Scale bar: AB, EF: 0.5 mm; CD, GH: 0.3 mm; IK: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239605" httpUri="https://zenodo.org/record/14239605/files/figure.png" pageId="4" pageNumber="375">Fig. 4I</figureCitation>
).
</paragraph>
</subSubSection>
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<paragraph id="8BA8793D6923FF81FF0DFAA91122119D" blockId="5.[151,1437,1356,1452]" pageId="5" pageNumber="376">
<emphasis id="B963A52F6923FF81FF0DFAA9121D1152" bold="true" box="[151,276,1356,1380]" pageId="5" pageNumber="376">FIGURE 3.</emphasis>
Habitus of
<taxonomicName id="4C1702BE6923FF81FE17FAA911C81155" authority="Matsumura" authorityName="Matsumura" authorityYear="2024" baseAuthorityName="Park &amp; Jung" baseAuthorityYear="2024" box="[397,705,1356,1380]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="bilineata" status="comb. nov.">
<emphasis id="B963A52F6923FF81FE17FAA9114E1152" box="[397,583,1356,1380]" italics="true" pageId="5" pageNumber="376">Toyoides bilineata</emphasis>
Matsumura
</taxonomicName>
<emphasis id="B963A52F6923FF81FD53FAA9102F1152" bold="true" box="[713,806,1356,1380]" pageId="5" pageNumber="376">
<taxonomicNameLabel id="A25018546923FF81FD53FAA9102F1152" box="[713,806,1356,1380]" pageId="5" pageNumber="376" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
A. female habitus, dorsal view; B. male habitus, dorsal view; C.
<emphasis id="B963A52F6923FF81FF2FFA9513EA11BE" box="[181,227,1392,1416]" italics="true" pageId="5" pageNumber="376">ditto</emphasis>
, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G.
<emphasis id="B963A52F6923FF81FB7FFA95161A11BE" box="[1253,1299,1392,1416]" italics="true" pageId="5" pageNumber="376">ditto</emphasis>
, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C30D2AB66923FF83FF5DFA1B12C516B3" lastPageId="7" lastPageNumber="378" pageId="5" pageNumber="376" type="description">
<paragraph id="8BA8793D6923FF83FF5DFA1B119E15BF" blockId="5.[151,1437,1534,2029]" lastBlockId="7.[151,1437,151,1221]" lastPageId="7" lastPageNumber="378" pageId="5" pageNumber="376">
<emphasis id="B963A52F6923FF81FF5DFA1B12B5122E" bold="true" box="[199,444,1534,1560]" pageId="5" pageNumber="376">Redescription. Male.</emphasis>
COLORATION:General coloration of body light brown(
<figureCitation id="132C65B86923FF81FBA9FA1B17B9122E" box="[1075,1200,1534,1560]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="5" pageNumber="376">Figs 3AC</figureCitation>
).Vertex concolorous, with background, lateral carinae slightly darker with fuscous marking medially (
<figureCitation id="132C65B86923FF81FB8AF9C7176D120A" box="[1040,1124,1570,1596]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="5" pageNumber="376">Fig. 3F</figureCitation>
). Pronotum and mesonotum concolorous, pronotum, mesonotum and prosternum with one pair of parallel blackish markings, mesonotum with longitudinal fuscous stripes developed inside lateral carinae (
<figureCitation id="132C65B86923FF81FCADF98F10BA12B2" box="[823,947,1642,1668]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="5" pageNumber="376">Figs 3FG</figureCitation>
). Frons concolorous, lateral carinae fuscous. Gena darker than background; compound eyes dark brown (
<figureCitation id="132C65B86923FF81FCB5F96B1083129E" box="[815,906,1678,1704]" captionStart="FIGURE 2" captionStartId="4.[151,250,1653,1677]" captionTargetBox="[151,1431,678,1626]" captionTargetId="figure-252@4.[151,1436,669,1635]" captionTargetPageId="4" captionText="FIGURE 2. Lectotype and paralectotype of Toyoides bilineata Matsumura comb. n. A. habitus, dorsal view; B. lectotype, habitus, lateral view; C. ditto, frons, ventral view; D. ditto, right hind leg with spur; E. lectotype and paralectotype labels. Abbreviations: f, front; b, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239601" httpUri="https://zenodo.org/record/14239601/files/figure.png" pageId="5" pageNumber="376">Figs 2C</figureCitation>
,
<figureCitation id="132C65B86923FF81FC0FF96B10B1129E" box="[917,952,1678,1704]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="5" pageNumber="376">3E</figureCitation>
). Tegmina light brown, with fuscous stripes on Pcu+A1, CuA and Sc+R reaching anterior and posterior margins; veins slightly darker than background (
<figureCitation id="132C65B86923FF81FAF5F95713B412C6" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="5" pageNumber="376">Fig. 3D</figureCitation>
). Legs slightly darker than background, spines and spur teeth black (
<figureCitation id="132C65B86923FF81FC29F933170412C6" box="[947,1037,1750,1776]" captionStart="FIGURE 2" captionStartId="4.[151,250,1653,1677]" captionTargetBox="[151,1431,678,1626]" captionTargetId="figure-252@4.[151,1436,669,1635]" captionTargetPageId="4" captionText="FIGURE 2. Lectotype and paralectotype of Toyoides bilineata Matsumura comb. n. A. habitus, dorsal view; B. lectotype, habitus, lateral view; C. ditto, frons, ventral view; D. ditto, right hind leg with spur; E. lectotype and paralectotype labels. Abbreviations: f, front; b, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239601" httpUri="https://zenodo.org/record/14239601/files/figure.png" pageId="5" pageNumber="376">Fig. 2D</figureCitation>
). Abdomen light brown. SURFACE AND VESTITURE: Tegmina and body partly covered with thin wax layer, irregular and rough. STRUCTURE: Macropterous species, slender.
<emphasis id="B963A52F6923FF81FD9BF8FB1141130E" bold="true" box="[513,584,1822,1848]" pageId="5" pageNumber="376">Head.</emphasis>
Head with compound eyes semicircular, narrower than pronotum; vertex at base flat, longer than width at midline, lateral and median carina well-developed (
<figureCitation id="132C65B86923FF81FBE8F8A717FA136A" box="[1138,1267,1858,1884]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="5" pageNumber="376">Figs 3FG</figureCitation>
). Frons widest at basal level of compound eyes, converging narrowly to basal, median carinae visible; postclypeus with median carina. Antenna clavated, first segment distinctly shorter than second segment almost two times longer than first segment; flagellum fine, bristle-like, as long as pedicel (
<figureCitation id="132C65B86923FF81FC81F84B107D13FE" box="[795,884,1966,1992]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="5" pageNumber="376">Fig. 3E</figureCitation>
).
<emphasis id="B963A52F6923FF81FC13F84B10E013FE" bold="true" box="[905,1001,1966,1992]" pageId="5" pageNumber="376">Thorax.</emphasis>
Pronotum narrow, length shorter than width; anterior margins convex, posterior margins slightly concave at midline; lateral carinae not reaching posterior margins. Mesonotum as long as broad, posterior margins triangular, longer than vertex; lateral carinae reaching posterior margins (
<figureCitation id="132C65B86920FF82FEF5F91112E61339" box="[367,495,1780,1807]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="6" pageNumber="377">Figs 3FG</figureCitation>
). Tegmina elongated, posterior margin of abdomen almost reaching half-length of tegmina, curved to dorsally; with tubercles along veins regularly; Sc+R branched at almost division of CuP (
<figureCitation id="132C65B86920FF82FAF5F8FD13B41361" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="6" pageNumber="377">Fig. 3D</figureCitation>
). Metatibial spur shorter than basitarsus, bearing 2832 distinct black-tipped teeth on trailing edge (
<figureCitation id="132C65B86920FF82FAAEF8D916991361" box="[1332,1424,1852,1879]" captionStart="FIGURE 2" captionStartId="4.[151,250,1653,1677]" captionTargetBox="[151,1431,678,1626]" captionTargetId="figure-252@4.[151,1436,669,1635]" captionTargetPageId="4" captionText="FIGURE 2. Lectotype and paralectotype of Toyoides bilineata Matsumura comb. n. A. habitus, dorsal view; B. lectotype, habitus, lateral view; C. ditto, frons, ventral view; D. ditto, right hind leg with spur; E. lectotype and paralectotype labels. Abbreviations: f, front; b, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239601" httpUri="https://zenodo.org/record/14239601/files/figure.png" pageId="6" pageNumber="377">Fig. 2D</figureCitation>
).
<emphasis id="B963A52F6920FF82FF0DF88413FC134D" bold="true" box="[151,245,1889,1915]" pageId="6" pageNumber="377">Female.</emphasis>
STRUCTURE: Same as male in general features, length distinctly longer than male (
<figureCitation id="132C65B86920FF82FB29F8851604134D" box="[1203,1293,1888,1915]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="6" pageNumber="377">Fig. 3A</figureCitation>
).
<emphasis id="B963A52F6920FF82FABBF8851695134C" bold="true" box="[1313,1436,1888,1914]" pageId="6" pageNumber="377">Abdomen.</emphasis>
GENITALIA: Pygofer in lateral view roughly quadrate, narrowed dorsally, angulated; anterior margin flat, caudal margin without teeth or processes (
<figureCitation id="132C65B86920FF82FD81F84D117813F5" box="[539,625,1960,1987]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="6" pageNumber="377">Fig. 3E</figureCitation>
). In caudal view, pygofer opening simple, margins carinated; opening for genital styles compressed-oval (
<figureCitation id="132C65B86920FF82FE61F829115913D1" box="[507,592,1996,2023]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="6" pageNumber="377">Fig. 3F</figureCitation>
). genital styles forceps-shaped; converging apically (
<figureCitation id="132C65B86920FF82FB07F829161413D1" box="[1181,1309,1996,2023]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="6" pageNumber="377">Figs 3GH</figureCitation>
). Aedeagus hook-shaped, robust and blunt; widest before midlength (in lateral view); dividing medially with small spinule; rounded at each apex, almost sclerotized but inner margins of upper appendage membranous; upper appendage curved to ventrally, narrower to median portion but apex expanded (
<figureCitation id="132C65B86921FF83FC3DFF3A10FE14CF" box="[935,1015,223,249]" captionStart="FIGURE 4" captionStartId="6.[151,250,1530,1554]" captionTargetBox="[158,1429,188,1499]" captionTargetId="figure-20@6.[151,1436,181,1506]" captionTargetPageId="6" captionText="FIGURE 4. Male genitalia of Toyoides bilineata Matsumura comb. n. A. genitalia, caudal view; B. ditto, lateral view; C. anal segment, caudal view; D. ditto, lateral view; E. pygofer, caudal view; F. ditto, lateral view; G. genital styles, caudoventral view; H. ditto, lateral view; I. aedeagus with suspensorium, left-lateral view; J. aedeagus without suspensorium, dorsal view; K. suspensorium. Scale bar: AB, EF: 0.5 mm; CD, GH: 0.3 mm; IK: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239605" httpUri="https://zenodo.org/record/14239605/files/figure.png" pageId="7" pageNumber="378">Fig. 4I</figureCitation>
); lower appendage simple, short and blunt; in dorsal view, slightly twisted and bilobed with margins strongly serrated (
<figureCitation id="132C65B86921FF83FBB4FEE61776152B" box="[1070,1151,259,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1530,1554]" captionTargetBox="[158,1429,188,1499]" captionTargetId="figure-20@6.[151,1436,181,1506]" captionTargetPageId="6" captionText="FIGURE 4. Male genitalia of Toyoides bilineata Matsumura comb. n. A. genitalia, caudal view; B. ditto, lateral view; C. anal segment, caudal view; D. ditto, lateral view; E. pygofer, caudal view; F. ditto, lateral view; G. genital styles, caudoventral view; H. ditto, lateral view; I. aedeagus with suspensorium, left-lateral view; J. aedeagus without suspensorium, dorsal view; K. suspensorium. Scale bar: AB, EF: 0.5 mm; CD, GH: 0.3 mm; IK: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239605" httpUri="https://zenodo.org/record/14239605/files/figure.png" pageId="7" pageNumber="378">Fig. 4J</figureCitation>
). Aedeagus tapering with suspensorium at basal (
<figureCitation id="132C65B86921FF83FE06FEC212E41577" box="[412,493,295,321]" captionStart="FIGURE 4" captionStartId="6.[151,250,1530,1554]" captionTargetBox="[158,1429,188,1499]" captionTargetId="figure-20@6.[151,1436,181,1506]" captionTargetPageId="6" captionText="FIGURE 4. Male genitalia of Toyoides bilineata Matsumura comb. n. A. genitalia, caudal view; B. ditto, lateral view; C. anal segment, caudal view; D. ditto, lateral view; E. pygofer, caudal view; F. ditto, lateral view; G. genital styles, caudoventral view; H. ditto, lateral view; I. aedeagus with suspensorium, left-lateral view; J. aedeagus without suspensorium, dorsal view; K. suspensorium. Scale bar: AB, EF: 0.5 mm; CD, GH: 0.3 mm; IK: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239605" httpUri="https://zenodo.org/record/14239605/files/figure.png" pageId="7" pageNumber="378">Fig. 4J</figureCitation>
). Suspensorium ovate, ring-like; longer than wide (
<figureCitation id="132C65B86921FF83FBB6FEC2178E1577" box="[1068,1159,295,321]" captionStart="FIGURE 4" captionStartId="6.[151,250,1530,1554]" captionTargetBox="[158,1429,188,1499]" captionTargetId="figure-20@6.[151,1436,181,1506]" captionTargetPageId="6" captionText="FIGURE 4. Male genitalia of Toyoides bilineata Matsumura comb. n. A. genitalia, caudal view; B. ditto, lateral view; C. anal segment, caudal view; D. ditto, lateral view; E. pygofer, caudal view; F. ditto, lateral view; G. genital styles, caudoventral view; H. ditto, lateral view; I. aedeagus with suspensorium, left-lateral view; J. aedeagus without suspensorium, dorsal view; K. suspensorium. Scale bar: AB, EF: 0.5 mm; CD, GH: 0.3 mm; IK: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239605" httpUri="https://zenodo.org/record/14239605/files/figure.png" pageId="7" pageNumber="378">Fig. 4K</figureCitation>
). Anal segment with one pair of spines pointed curved ventrally (
<figureCitation id="132C65B86921FF83FDCBFEAE11D91553" box="[593,720,331,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1530,1554]" captionTargetBox="[158,1429,188,1499]" captionTargetId="figure-20@6.[151,1436,181,1506]" captionTargetPageId="6" captionText="FIGURE 4. Male genitalia of Toyoides bilineata Matsumura comb. n. A. genitalia, caudal view; B. ditto, lateral view; C. anal segment, caudal view; D. ditto, lateral view; E. pygofer, caudal view; F. ditto, lateral view; G. genital styles, caudoventral view; H. ditto, lateral view; I. aedeagus with suspensorium, left-lateral view; J. aedeagus without suspensorium, dorsal view; K. suspensorium. Scale bar: AB, EF: 0.5 mm; CD, GH: 0.3 mm; IK: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239605" httpUri="https://zenodo.org/record/14239605/files/figure.png" pageId="7" pageNumber="378">Figs 4CD</figureCitation>
). See
<bibRefCitation id="EF8604CC6921FF83FC8BFEAE10E81553" author="Matsumura, S." box="[785,993,330,357]" pageId="7" pageNumber="378" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura (1935)</bibRefCitation>
for original description and
<bibRefCitation id="EF8604CC6921FF83FA87FEAE126915BF" author="Matsumura, S. &amp; Ishihara, T." pageId="7" pageNumber="378" pagination="59 - 76" refId="ref5610" refString="Matsumura, S. &amp; Ishihara, T. (1945) Species novae vel cognitae Araeopidarum Imperii Japonici (Hemiptera). Mushi, 16, 59 - 76." type="journal article" year="1945">Matsumura &amp; Ishihara (1945)</bibRefCitation>
for genital illustration also.
</paragraph>
<caption id="DF6829B56920FF82FF0DFA1F10CE1248" ID-DOI="http://doi.org/10.5281/zenodo.14239605" ID-Zenodo-Dep="14239605" httpUri="https://zenodo.org/record/14239605/files/figure.png" pageId="6" pageNumber="377" startId="6.[151,250,1530,1554]" targetBox="[158,1429,188,1499]" targetPageId="6" targetType="figure">
<paragraph id="8BA8793D6920FF82FF0DFA1F10CE1248" blockId="6.[151,1437,1530,1663]" pageId="6" pageNumber="377">
<emphasis id="B963A52F6920FF82FF0DFA1F121A1224" bold="true" box="[151,275,1530,1554]" pageId="6" pageNumber="377">FIGURE 4.</emphasis>
Male genitalia of
<taxonomicName id="4C1702BE6920FF82FE54FA1F100B1224" authority="Matsumura" authorityName="Matsumura" authorityYear="2024" baseAuthorityName="Park &amp; Jung" baseAuthorityYear="2024" box="[462,770,1530,1554]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="bilineata" status="comb. nov.">
<emphasis id="B963A52F6920FF82FE54FA1F11811224" box="[462,648,1530,1554]" italics="true" pageId="6" pageNumber="377">Toyoides bilineata</emphasis>
Matsumura
</taxonomicName>
<emphasis id="B963A52F6920FF82FC93FA1F106C1224" bold="true" box="[777,869,1530,1554]" pageId="6" pageNumber="377">
<taxonomicNameLabel id="A25018546920FF82FC93FA1F106C1224" box="[777,869,1530,1554]" pageId="6" pageNumber="377" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
A. genitalia, caudal view; B.
<emphasis id="B963A52F6920FF82FB0EFA1F17CB1224" box="[1172,1218,1530,1554]" italics="true" pageId="6" pageNumber="377">ditto</emphasis>
, lateral view; C. anal segment, caudal view; D.
<emphasis id="B963A52F6920FF82FE06F9FB12C31200" box="[412,458,1566,1590]" italics="true" pageId="6" pageNumber="377">ditto</emphasis>
, lateral view; E. pygofer, caudal view; F.
<emphasis id="B963A52F6920FF82FCF4F9FB10951200" box="[878,924,1566,1590]" italics="true" pageId="6" pageNumber="377">ditto</emphasis>
, lateral view; G. genital styles, caudoventral view; H.
<emphasis id="B963A52F6920FF82FF23F9A713EE126C" box="[185,231,1602,1626]" italics="true" pageId="6" pageNumber="377">ditto</emphasis>
, lateral view; I. aedeagus with suspensorium, left-lateral view; J.
<emphasis id="B963A52F6920FF82FC3AF9A710D5126C" box="[928,988,1602,1626]" italics="true" pageId="6" pageNumber="377">aedea</emphasis>
gus without suspensorium, dorsal view; K. suspensorium. Scale bar: AB, EF: 0.5 mm; CD, GH: 0.3 mm; IK: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BA8793D6921FF83FF5DFE7612C516B3" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="378">
<emphasis id="B963A52F6921FF83FF5DFE7612E7159A" bold="true" box="[199,494,402,429]" pageId="7" pageNumber="378">Measurements (in mm).</emphasis>
Male (n=4) / Female (n=11). Body length (including tegmina): 6.20-6.22/6.20- 6.53; body length (excluding tegmina): 3.37-3.40/3.50-3.60; body width: 1.37-1.40/1.52-1.55; head length: 0.51- 0.54/0.55-0.59; head width (including eyes): 0.93-0.95/0.94-0.97; first antennal segment length: 0.22-0.23/0.28- 0.33; second antennal segment length: 0.40-0.41/0.43-0.46; vertex length: 0.36-0.38/0.38-0.41; vertex width (basal width): 0.33-0.35/0.35-0.38; frons length: 0.78-0.79/0.79-0.82; frons width: 0.31-0.32/0.32-0.35; pronotum length: 0.39-0.40/0.40-0.43; pronotum width: 1.03-1.04/1.06-1.09; mesonotum length: 0.57-0.59/0.59-0.61; mesonotum width: 0.90-0.91/0.92-0.94.
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66921FF83FF5DFD6B1035170F" pageId="7" pageNumber="378" type="materials_examined">
<paragraph id="8BA8793D6921FF83FF5DFD6B1035170F" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="378">
<emphasis id="B963A52F6921FF83FF5DFD6B12B4169E" bold="true" box="[199,445,654,680]" pageId="7" pageNumber="378">Material examined. [</emphasis>
<materialsCitation id="3B7F73606921FF83FE27FD6B17E1169F" box="[445,1256,654,681]" collectingDate="1924-08-03" collectionCode="SEHU" collectorName="S. Matsumura" country="Japan" location="Sapporo" pageId="7" pageNumber="378" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="B963A52F6921FF83FE27FD6B1192169F" bold="true" box="[445,667,654,681]" pageId="7" pageNumber="378">
<collectionCode id="ED06E1F86921FF83FE27FD6B1107169E" box="[445,526,654,680]" collectionName="SEHU" pageId="7" pageNumber="378">SEHU</collectionCode>
]
<typeStatus id="54ACC79F6921FF83FD81FD6A119F169F" box="[539,662,655,681]" pageId="7" pageNumber="378" type="lectotype">Lectotype</typeStatus>
:
</emphasis>
<specimenCount id="9D11B2B46921FF83FD3BFD6A11CF169F" box="[673,710,655,681]" count="1" pageId="7" pageNumber="378" type="male">1♂</specimenCount>
,
<location id="8EC82FE66921FF83FD48FD6A103B169F" LSID="urn:lsid:plazi:treatment:03BEC82B6922FF83FF0DFE3F13D310F3:8EC82FE66921FF83FD48FD6A103B169F" box="[722,818,655,681]" country="Japan" name="Sapporo" pageId="7" pageNumber="378">Sapporo</location>
,
<collectingCountry id="F30039AD6921FF83FCA6FD6A1076169F" box="[828,895,655,681]" name="Japan" pageId="7" pageNumber="378">Japan</collectingCountry>
;
<date id="FFA95FFD6921FF83FC10FD6A1718169F" box="[906,1041,655,681]" pageId="7" pageNumber="378" value="1924-08-03">
<collectingDate id="EFEDA6156921FF83FC10FD6A1718169F" box="[906,1041,655,681]" pageId="7" pageNumber="378" value="1924-08-03">3.VIII.1924</collectingDate>
</date>
,
<collectorName id="26E21CEB6921FF83FB86FD6A17B0169F" box="[1052,1209,655,681]" pageId="7" pageNumber="378">S. Matsumura</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B7F73606921FF83FB74FD6A110816FB" collectingDate="1924-08-03" collectionCode="SEHU" collectorName="S. Matsumura" country="Japan" location="Sapporo" pageId="7" pageNumber="378" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<emphasis id="B963A52F6921FF83FB74FD6A1695169F" bold="true" box="[1262,1436,655,681]" pageId="7" pageNumber="378">
<typeStatus id="54ACC79F6921FF83FB74FD6A169E169F" box="[1262,1431,655,681]" pageId="7" pageNumber="378" type="paralectotype">Paralectotype</typeStatus>
:
</emphasis>
<specimenCount id="9D11B2B46921FF83FF0DFD5613B316FB" box="[151,186,691,717]" count="1" pageId="7" pageNumber="378" type="female">1♀</specimenCount>
, same data as lectotype male
</materialsCitation>
;
<materialsCitation id="3B7F73606921FF83FD91FD56176016FB" box="[523,1129,691,717]" collectingDate="1924-08-09" collectorName="S. Matsumura" country="Japan" location="Sapporo" pageId="7" pageNumber="378" specimenCount="2" specimenCount-female="2" typeStatus="paralectotype">
<specimenCount id="9D11B2B46921FF83FD91FD56114C16FB" box="[523,581,691,717]" count="2" pageId="7" pageNumber="378" type="female">2♀♀</specimenCount>
,
<location id="8EC82FE66921FF83FDCBFD5611B816FB" LSID="urn:lsid:plazi:treatment:03BEC82B6922FF83FF0DFE3F13D310F3:8EC82FE66921FF83FDCBFD5611B816FB" box="[593,689,691,717]" country="Japan" name="Sapporo" pageId="7" pageNumber="378">Sapporo</location>
,
<collectingCountry id="F30039AD6921FF83FD21FD5611F716FB" box="[699,766,691,717]" name="Japan" pageId="7" pageNumber="378">Japan</collectingCountry>
;
<date id="FFA95FFD6921FF83FC93FD56109916FB" box="[777,912,691,717]" pageId="7" pageNumber="378" value="1924-08-09">
<collectingDate id="EFEDA6156921FF83FC93FD56109916FB" box="[777,912,691,717]" pageId="7" pageNumber="378" value="1924-08-09">9.VIII.1924</collectingDate>
</date>
,
<collectorName id="26E21CEB6921FF83FC00FD56173E16FB" box="[922,1079,691,717]" pageId="7" pageNumber="378">S. Matsumura</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7F73606921FF83FBEEFD56110C16C7" collectingDate="1918-08-04" collectorName="S. Matsumura" country="Japan" location="Sapporo" pageId="7" pageNumber="378" specimenCount="8" specimenCount-female="7" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="9D11B2B46921FF83FBEEFD56179016FB" box="[1140,1177,691,717]" count="1" pageId="7" pageNumber="378" type="male">1♂</specimenCount>
,
<specimenCount id="9D11B2B46921FF83FB3FFD5617D716FB" box="[1189,1246,691,717]" count="7" pageId="7" pageNumber="378" type="female">7♀♀</specimenCount>
,
<location id="8EC82FE66921FF83FB70FD56164316FB" LSID="urn:lsid:plazi:treatment:03BEC82B6922FF83FF0DFE3F13D310F3:8EC82FE66921FF83FB70FD56164316FB" box="[1258,1354,691,717]" country="Japan" name="Sapporo" pageId="7" pageNumber="378">Sapporo</location>
,
<collectingCountry id="F30039AD6921FF83FACFFD56169116FB" box="[1365,1432,691,717]" name="Japan" pageId="7" pageNumber="378">Japan</collectingCountry>
;
<date id="FFA95FFD6921FF83FF0DFD32121716C7" box="[151,286,727,753]" pageId="7" pageNumber="378" value="1918-08-04">
<collectingDate id="EFEDA6156921FF83FF0DFD32121716C7" box="[151,286,727,753]" pageId="7" pageNumber="378" value="1918-08-04">4.VIII.1918</collectingDate>
</date>
,
<collectorName id="26E21CEB6921FF83FEB7FD3212C616C7" box="[301,463,727,753]" pageId="7" pageNumber="378">S. Matsumura</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7F73606921FF83FD8FFD33179516C7" box="[533,1180,726,753]" collectingDate="1918-08-12" collectorName="S. Matsumura" country="Japan" location="Sapporo" pageId="7" pageNumber="378" specimenCount="5" specimenCount-female="5" typeStatus="paralectotype">
<specimenCount id="9D11B2B46921FF83FD8FFD33114716C7" box="[533,590,726,753]" count="5" pageId="7" pageNumber="378" type="female">5♀♀</specimenCount>
,
<location id="8EC82FE66921FF83FDC4FD3211B716C7" LSID="urn:lsid:plazi:treatment:03BEC82B6922FF83FF0DFE3F13D310F3:8EC82FE66921FF83FDC4FD3211B716C7" box="[606,702,727,753]" country="Japan" name="Sapporo" pageId="7" pageNumber="378">Sapporo</location>
,
<collectingCountry id="F30039AD6921FF83FD54FD32101816C7" box="[718,785,727,753]" name="Japan" pageId="7" pageNumber="378">Japan</collectingCountry>
;
<date id="FFA95FFD6921FF83FCBAFD3210BC16C7" box="[800,949,727,753]" pageId="7" pageNumber="378" value="1918-08-12">
<collectingDate id="EFEDA6156921FF83FCBAFD3210BC16C7" box="[800,949,727,753]" pageId="7" pageNumber="378" value="1918-08-12">12.VIII.1918</collectingDate>
</date>
,
<collectorName id="26E21CEB6921FF83FC5EFD32176C16C7" box="[964,1125,727,753]" pageId="7" pageNumber="378">S. Matsumura</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7F73606921FF83FB31FD331035170F" collectingDate="2023-07-28" collectingMethod="sweeping on Phacelurus latifolius" collectionCode="CNU" collectorName="J. K. Park &amp; G. H. Jang &amp; E. J. Kim &amp; J. H. Jeon &amp; Hamzah M." country="South Korea" location="Janggok-dong" municipality="Siheung-si" pageId="7" pageNumber="378" specimenCount="13" specimenCount-female="10" specimenCount-male="3" stateProvince="Gyeonggi-do" typeStatus="paralectotype">
<emphasis id="B963A52F6921FF83FB31FD3317F316C6" bold="true" box="[1195,1274,726,752]" pageId="7" pageNumber="378">
[
<collectionCode id="ED06E1F86921FF83FB29FD3317FD16C6" box="[1203,1268,726,752]" collectionName="CNU" country="China" httpUri="http://grbio.org/cool/kyjm-bn0n" name="Capital Normal University, College of Life Sciences" pageId="7" pageNumber="378">CNU</collectionCode>
]
</emphasis>
<specimenCount id="9D11B2B46921FF83FA9FFD32164916C7" box="[1285,1344,727,753]" count="3" pageId="7" pageNumber="378" type="male">3♂♂</specimenCount>
,
<specimenCount id="9D11B2B46921FF83FAD5FD32169E16C7" box="[1359,1431,727,753]" count="10" pageId="7" pageNumber="378" type="female">10♀♀</specimenCount>
,
<location id="8EC82FE66921FF83FF0DFD1E12301723" LSID="urn:lsid:plazi:treatment:03BEC82B6922FF83FF0DFE3F13D310F3:8EC82FE66921FF83FF0DFD1E12301723" box="[151,313,763,789]" country="South Korea" municipality="Siheung-si" name="Janggok-dong" pageId="7" pageNumber="378" stateProvince="Gyeonggi-do">Janggok-dong</location>
,
<collectingMunicipality id="6BCCE3476921FF83FEDCFD1E12C81723" box="[326,449,763,789]" pageId="7" pageNumber="378">Siheung-si</collectingMunicipality>
,
<collectingRegion id="49D3B7DF6921FF83FE57FD1E116D1723" box="[461,612,763,789]" country="South Korea" name="Gyeonggi" pageId="7" pageNumber="378">Gyeonggi-do</collectingRegion>
.
<collectingCountry id="F30039AD6921FF83FDEBFD1E104E1723" box="[625,839,763,789]" name="South Korea" pageId="7" pageNumber="378">Republic of Korea</collectingCountry>
;
<date id="FFA95FFD6921FF83FCCEFD1E10E91723" box="[852,992,763,789]" pageId="7" pageNumber="378" value="2023-07-28">
<collectingDate id="EFEDA6156921FF83FCCEFD1E10E91723" box="[852,992,763,789]" pageId="7" pageNumber="378" value="2023-07-28">28.VII.2023</collectingDate>
</date>
, by
<collectingMethod id="5256012A6921FF83FB88FD1E169F1723" box="[1042,1430,763,789]" pageId="7" pageNumber="378">
sweeping on
<taxonomicName id="4C1702BE6921FF83FB33FD1E1005170F" authority=", J. K. Park, G. H. Jang, E. J. Kim, J. H. Jeon &amp; Hamzah M." authorityName="J. K. Park, G. H. Jang, E. J. Kim, J. H. Jeon &amp; Hamzah M." class="Liliopsida" family="Poaceae" genus="Phacelurus" kingdom="Plantae" order="Poales" pageId="7" pageNumber="378" phylum="Tracheophyta" rank="species" species="latifolius">
<emphasis id="B963A52F6921FF83FB33FD1E169F1723" box="[1193,1430,763,789]" italics="true" pageId="7" pageNumber="378">Phacelurus latifolius</emphasis>
,
<collectorName id="26E21CEB6921FF83FF0DFCFA1208170F" box="[151,257,799,825]" pageId="7" pageNumber="378">J.K. Park</collectorName>
,
<collectorName id="26E21CEB6921FF83FE97FCFA1289170F" box="[269,384,799,825]" pageId="7" pageNumber="378">G.H. Jang</collectorName>
,
<collectorName id="26E21CEB6921FF83FE11FCFA12F8170F" box="[395,497,799,825]" pageId="7" pageNumber="378">E.J. Kim</collectorName>
,
<collectorName id="26E21CEB6921FF83FE66FCFA116A170F" box="[508,611,799,825]" pageId="7" pageNumber="378">J.H. Jeon</collectorName>
&amp;
<collectorName id="26E21CEB6921FF83FD1DFCFA1005170F" box="[647,780,799,825]" pageId="7" pageNumber="378">Hamzah M.</collectorName>
</taxonomicName>
</collectingMethod>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66921FF83FF5DFCA71687176B" box="[199,1422,834,861]" pageId="7" pageNumber="378" type="distribution">
<paragraph id="8BA8793D6921FF83FF5DFCA71687176B" blockId="7.[151,1437,151,1221]" box="[199,1422,834,861]" pageId="7" pageNumber="378">
<emphasis id="B963A52F6921FF83FF5DFCA71265176A" bold="true" box="[199,364,834,860]" pageId="7" pageNumber="378">Distributions.</emphasis>
Korean Peninsula (Gyeonggi)
<superScript id="7C62D4756921FF83FD5BFCA711C01766" attach="none" box="[705,713,834,848]" fontSize="6" pageId="7" pageNumber="378">*</superScript>
,
<collectingCountry id="F30039AD6921FF83FD4DFCA6101E176B" box="[727,791,835,861]" name="Japan" pageId="7" pageNumber="378">Japan</collectingCountry>
(
<collectingRegion id="49D3B7DF6921FF83FCBFFCA61091176B" box="[805,920,835,861]" country="Japan" name="Hokkaido" pageId="7" pageNumber="378">Hokkaido</collectingRegion>
, Honshu, Kyushu, Tsushima Island) (
<figureCitation id="132C65B86921FF83FAA6FCA61689176B" box="[1340,1408,834,861]" captionStart="FIGURE 5" captionStartId="7.[151,250,1946,1970]" captionTargetBox="[151,1435,1281,1923]" captionTargetId="figure-410@7.[151,1435,1281,1923]" captionTargetPageId="7" captionText="FIGURE 5. Distribution map of the genus Toyoides species from the Korean Peninsula and neighboring countries." figureDoi="http://doi.org/10.5281/zenodo.14239607" httpUri="https://zenodo.org/record/14239607/files/figure.png" pageId="7" pageNumber="378">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66921FF83FF5DFC8212D917DB" pageId="7" pageNumber="378" type="biology_ecology">
<paragraph id="8BA8793D6921FF83FF5DFC8211F817B7" blockId="7.[151,1437,151,1221]" box="[199,753,870,897]" pageId="7" pageNumber="378">
<emphasis id="B963A52F6921FF83FF5DFC82124217B7" bold="true" box="[199,331,871,897]" pageId="7" pageNumber="378">Host plant.</emphasis>
<taxonomicName id="4C1702BE6921FF83FEC8FC8211EB17B7" authority="(Steud.) Ohwi" authorityName="Ohwi" baseAuthorityName="Steud." box="[338,738,871,897]" class="Liliopsida" family="Poaceae" genus="Phacelurus" kingdom="Plantae" order="Poales" pageId="7" pageNumber="378" phylum="Tracheophyta" rank="species" species="latifolius">
<emphasis id="B963A52F6921FF83FEC8FC82113417B7" box="[338,573,871,897]" italics="true" pageId="7" pageNumber="378">Phacelurus latifolius</emphasis>
(Steud.) Ohwi
</taxonomicName>
<superScript id="7C62D4756921FF83FD78FC8311E31742" attach="left" box="[738,746,870,884]" fontSize="6" pageId="7" pageNumber="378">*</superScript>
.
</paragraph>
<paragraph id="8BA8793D6921FF83FF5DFC6F12D917DB" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="378">
<emphasis id="B963A52F6921FF83FF5DFC6F122E1792" bold="true" box="[199,295,906,932]" pageId="7" pageNumber="378">Biology.</emphasis>
The
<typeStatus id="54ACC79F6921FF83FEFAFC6E12991793" box="[352,400,907,933]" pageId="7" pageNumber="378">type</typeStatus>
locality is
<collectingRegion id="49D3B7DF6921FF83FD90FC6E11F91793" box="[522,752,907,933]" country="Japan" name="Hokkaido" pageId="7" pageNumber="378">Hokkaido prefecture</collectingRegion>
in
<collectingCountry id="F30039AD6921FF83FC8EFC6E105E1793" box="[788,855,907,933]" name="Japan" pageId="7" pageNumber="378">Japan</collectingCountry>
, shortly mentioned biology as follows; “collected in August by the author at Sapporo, but the male is rare”.
<location id="8EC82FE66921FF83FC9BFC4A101117FF" LSID="urn:lsid:plazi:treatment:03BEC82B6922FF83FF0DFE3F13D310F3:8EC82FE66921FF83FC9BFC4A101117FF" box="[769,792,943,969]" country="Japan" name="In" pageId="7" pageNumber="378" stateProvince="Hokkaido">In</location>
this study, it was observed this species feed on halophyte grass,
<taxonomicName id="4C1702BE6921FF83FF45FC3612C017DB" box="[223,457,979,1005]" class="Liliopsida" family="Poaceae" genus="Phacelurus" kingdom="Plantae" order="Poales" pageId="7" pageNumber="378" phylum="Tracheophyta" rank="species" species="latifolius">
<emphasis id="B963A52F6921FF83FF45FC3612C017DB" box="[223,457,979,1005]" italics="true" pageId="7" pageNumber="378">Phacelurus latifolius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66921FF83FF5DFC1213D310F3" pageId="7" pageNumber="378" type="discussion">
<paragraph id="8BA8793D6921FF83FF5DFC1213D310F3" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="378">
<emphasis id="B963A52F6921FF83FF5DFC1212321027" bold="true" box="[199,315,1015,1041]" pageId="7" pageNumber="378">Remarks.</emphasis>
<bibRefCitation id="EF8604CC6921FF83FEDCFC1212E21027" author="Ishihara, T." box="[326,491,1015,1041]" pageId="7" pageNumber="378" pagination="1 - 102" refId="ref5544" refString="Ishihara, T. (1949) Revision of the Araeopidae of Japan, Ryukyu Island and Formosa (Hemiptera). Scientific Reports of the Matsuyama Agricultural College, 2, 1 - 102." type="journal article" year="1949">Ishihara(1949)</bibRefCitation>
considered
<emphasis id="B963A52F6921FF83FDE1FC1211DA1027" box="[635,723,1015,1041]" italics="true" pageId="7" pageNumber="378">Toyoide</emphasis>
s as a synonym of the genus
<taxonomicName id="4C1702BE6921FF83FBABFC12178B1027" box="[1073,1154,1015,1041]" genus="Euidell" pageId="7" pageNumber="378" rank="genus">
<emphasis id="B963A52F6921FF83FBABFC12178B1027" box="[1073,1154,1015,1041]" italics="true" pageId="7" pageNumber="378">Euidell</emphasis>
</taxonomicName>
a; however, more recent publications have not accepted this classification (
<bibRefCitation id="EF8604CC6921FF83FD5EFBFE10481003" author="Nast, J." box="[708,833,1051,1077]" pageId="7" pageNumber="378" refId="ref5641" refString="Nast, J. (1972) Palaearctic Auchenorrhyncha (Homoptera). Polish Scientific publishers, Warszawa, 550 pp." type="book" year="1972">Nast, 1972</bibRefCitation>
;
<bibRefCitation id="EF8604CC6921FF83FCD6FBFE17891003" author="Hayashi, M. &amp; Fujinuma, S." box="[844,1152,1051,1077]" pageId="7" pageNumber="378" pagination="323 - 355" refId="ref5480" refString="Hayashi, M. &amp; Fujinuma, S. (2016) Part Fulgoromorpha. In: Entomological Society of Japan (2016) Catalogue of the insects of Japan. Vol. 4. Paraneoptera (Psocodea, Thysanoptera, Hemiptera). 4. Editorial Committee of Catalogue of the Insects of Japan, 4, pp. 323 - 355." type="journal article" year="2016">Hayashi &amp; Fujinuma, 2016</bibRefCitation>
). The type materials were not designated in the protologue (
<bibRefCitation id="EF8604CC6921FF83FD82FBDA11EF106F" author="Matsumura, S." box="[536,742,1086,1113]" pageId="7" pageNumber="378" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935</bibRefCitation>
). Here, we designated the
<typeStatus id="54ACC79F6921FF83FB8DFBDA1777106F" box="[1047,1150,1087,1113]" pageId="7" pageNumber="378" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="54ACC79F6921FF83FB22FBDA1646106F" box="[1208,1359,1087,1113]" pageId="7" pageNumber="378" type="paralectotype">paralectotype</typeStatus>
which are deposited in SEHU. This species has not been redescribed since the original description. It is treated as a junior synonym of the genu
<emphasis id="B963A52F6921FF83FE18FB6212D61097" box="[386,479,1159,1185]" italics="true" pageId="7" pageNumber="378">s Toyoid</emphasis>
es due to the similarity in habitus and genital characters, which are redescribed in this study.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,228 @@
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<subSubSection id="C30D2AB66925FF87FF0DFE66116315AB" box="[151,618,387,414]" pageId="3" pageNumber="374" type="nomenclature">
<paragraph id="8BA8793D6925FF87FF0DFE66116315AB" blockId="3.[151,618,387,449]" box="[151,618,387,414]" pageId="3" pageNumber="374">
<heading id="D0E0CE516925FF87FF0DFE66116315AB" bold="true" box="[151,618,387,414]" fontSize="11" level="1" pageId="3" pageNumber="374" reason="1">
<taxonomicName id="4C1702BE6925FF87FF0DFE66116315AB" ID-CoL="57N2L" authority="Matsumura, 1935" authorityName="Matsumura" authorityYear="1935" box="[151,618,387,414]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="albipennis">
<emphasis id="B963A52F6925FF87FF0DFE66116315AB" bold="true" box="[151,618,387,414]" pageId="3" pageNumber="374">
<emphasis id="B963A52F6925FF87FF0DFE66128C15A8" bold="true" box="[151,389,387,414]" italics="true" pageId="3" pageNumber="374">Toyoides albipennis</emphasis>
<bibRefCitation id="EF8604CC6925FF87FE16FE66116315AB" author="Matsumura, S." box="[396,618,387,413]" pageId="3" pageNumber="374" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66925FF87FF0DFE42122C15F7" box="[151,293,423,449]" pageId="3" pageNumber="374" type="description">
<paragraph id="8BA8793D6925FF87FF0DFE42122C15F7" blockId="3.[151,618,387,449]" box="[151,293,423,449]" pageId="3" pageNumber="374">
(
<figureCitation id="132C65B86925FF87FF05FE42121715F7" box="[159,286,423,449]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[151,1436,477,1456]" captionTargetId="figure-420@3.[151,1436,502,1460]" captionTargetPageId="3" captionText="FIGURE 1. Lectotype of Toyoides albipennis Matsumura. A. habitus, dorsal view; B. ditto, lateral view; C. frons, ventral view; D. right hind leg with spur; E. lectotype labels. Abbreviations: f, front; b, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239599" httpUri="https://zenodo.org/record/14239599/files/figure.png" pageId="3" pageNumber="374">Figs 1AD</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="DF6829B56925FF87FF0DFA27165911C8" ID-DOI="http://doi.org/10.5281/zenodo.14239599" ID-Zenodo-Dep="14239599" httpUri="https://zenodo.org/record/14239599/files/figure.png" pageId="3" pageNumber="374" startId="3.[151,250,1474,1498]" targetBox="[151,1436,477,1456]" targetPageId="3" targetType="figure">
<paragraph id="8BA8793D6925FF87FF0DFA27165911C8" blockId="3.[151,1436,1474,1534]" pageId="3" pageNumber="374">
<emphasis id="B963A52F6925FF87FF0DFA27121A11EC" bold="true" box="[151,275,1474,1498]" pageId="3" pageNumber="374">FIGURE 1.</emphasis>
Lectotype of
<taxonomicName id="4C1702BE6925FF87FE04FA27100D11EC" authority="Matsumura. A." authorityName="Matsumura. A." box="[414,772,1474,1498]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="albipennis">
<emphasis id="B963A52F6925FF87FE04FA27116E11EC" box="[414,615,1474,1498]" italics="true" pageId="3" pageNumber="374">Toyoides albipennis</emphasis>
Matsumura. A.
</taxonomicName>
habitus, dorsal view; B.
<emphasis id="B963A52F6925FF87FC67FA27172211EC" box="[1021,1067,1474,1498]" italics="true" pageId="3" pageNumber="374">ditto</emphasis>
, lateral view; C. frons, ventral view; D. right hind leg with spur; E. lectotype labels. Abbreviations:
<emphasis id="B963A52F6925FF87FC8EFA03101211C8" box="[788,795,1510,1534]" italics="true" pageId="3" pageNumber="374">f</emphasis>
, front;
<emphasis id="B963A52F6925FF87FCFCFA03107A11C8" box="[870,883,1510,1534]" italics="true" pageId="3" pageNumber="374">b</emphasis>
, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C30D2AB66925FF87FF0DF9AE11E512C3" pageId="3" pageNumber="374" type="diagnosis">
<paragraph id="8BA8793D6925FF87FF0DF9AE11E512C3" blockId="3.[151,1437,1611,2034]" pageId="3" pageNumber="374">
<emphasis id="B963A52F6925FF87FF0DF9AE121B1253" bold="true" box="[151,274,1611,1637]" pageId="3" pageNumber="374">Diagnosis.</emphasis>
Recognized by body light green to yellowish brown; tegmina with anterior and posterior margins slightly infuscate; vertex without any markings; prosternum with one pair of blackish markings parallel (
<figureCitation id="132C65B86925FF87FAA9F98A13A7129B" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[151,1436,477,1456]" captionTargetId="figure-420@3.[151,1436,502,1460]" captionTargetPageId="3" captionText="FIGURE 1. Lectotype of Toyoides albipennis Matsumura. A. habitus, dorsal view; B. ditto, lateral view; C. frons, ventral view; D. right hind leg with spur; E. lectotype labels. Abbreviations: f, front; b, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239599" httpUri="https://zenodo.org/record/14239599/files/figure.png" pageId="3" pageNumber="374">Figs 1A D</figureCitation>
); tegmina almost straight (
<figureCitation id="132C65B86925FF87FE46F976113A129B" box="[476,563,1683,1709]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[151,1436,477,1456]" captionTargetId="figure-420@3.[151,1436,502,1460]" captionTargetPageId="3" captionText="FIGURE 1. Lectotype of Toyoides albipennis Matsumura. A. habitus, dorsal view; B. ditto, lateral view; C. frons, ventral view; D. right hind leg with spur; E. lectotype labels. Abbreviations: f, front; b, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239599" httpUri="https://zenodo.org/record/14239599/files/figure.png" pageId="3" pageNumber="374">Fig. 1B</figureCitation>
); frons with median carinae forked below level of lower margin of eyes (
<figureCitation id="132C65B86925FF87FAF5F97613B012E7" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[151,1436,477,1456]" captionTargetId="figure-420@3.[151,1436,502,1460]" captionTargetPageId="3" captionText="FIGURE 1. Lectotype of Toyoides albipennis Matsumura. A. habitus, dorsal view; B. ditto, lateral view; C. frons, ventral view; D. right hind leg with spur; E. lectotype labels. Abbreviations: f, front; b, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239599" httpUri="https://zenodo.org/record/14239599/files/figure.png" pageId="3" pageNumber="374">Fig. 1C</figureCitation>
); pygofer without medioventral structures; aedeagus branched at medially, division without spinule (modified from
<bibRefCitation id="EF8604CC6925FF87FF4FF939129412C0" author="Matsumura, S." box="[213,413,1755,1782]" pageId="3" pageNumber="374" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935</bibRefCitation>
,
<bibRefCitation id="EF8604CC6925FF87FE3DF939112E12C3" author="Yang, C. T." box="[423,551,1755,1782]" pageId="3" pageNumber="374" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang, 1989</bibRefCitation>
and
<bibRefCitation id="EF8604CC6925FF87FDC7F93E11E912C3" author="Ding, J. H." box="[605,736,1755,1781]" pageId="3" pageNumber="374" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding, 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66925FF87FF5DF91A169413FB" pageId="3" pageNumber="374" type="description">
<paragraph id="8BA8793D6925FF87FF5DF91A1259130B" blockId="3.[151,1437,1611,2034]" pageId="3" pageNumber="374">
<emphasis id="B963A52F6925FF87FF5DF91A1250132F" bold="true" box="[199,345,1791,1817]" pageId="3" pageNumber="374">Description.</emphasis>
See
<bibRefCitation id="EF8604CC6925FF87FE09F8E5116F132C" author="Matsumura, S." box="[403,614,1791,1818]" pageId="3" pageNumber="374" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura (1935)</bibRefCitation>
for original description,
<bibRefCitation id="EF8604CC6925FF87FC1EF8E51718132F" author="Yang, C. T." box="[900,1041,1791,1818]" pageId="3" pageNumber="374" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang (1989)</bibRefCitation>
and
<bibRefCitation id="EF8604CC6925FF87FBD6F91A17D1132F" author="Ding, J. H." box="[1100,1240,1791,1817]" pageId="3" pageNumber="374" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding (2006)</bibRefCitation>
for redescription and figures also.
</paragraph>
<paragraph id="8BA8793D6925FF87FF5DF8A2169413FB" blockId="3.[151,1437,1611,2034]" pageId="3" pageNumber="374">
<emphasis id="B963A52F6925FF87FF5DF8A212D71357" bold="true" box="[199,478,1863,1889]" pageId="3" pageNumber="374">Measurements (in mm).</emphasis>
Female (n=1). Body length (including tegmina): 5.85; body length (excluding tegmina): 3.43; body width: 1.48; head length: 0.54; head width (including eyes): 0.91; first antennal segment length: 0.19; second antennal segment length: 0.40; vertex length: 0.41; vertex width (basal width): 0.38; frons length: 0.80; frons width: 0.38; pronotum length: 0.33; pronotum width: 1.08; mesonotum length: 0.68; mesonotum width: 1.04.
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66925FF87FF5DF832161C13C7" box="[199,1301,2007,2034]" pageId="3" pageNumber="374" type="materials_examined">
<paragraph id="8BA8793D6925FF87FF5DF832161C13C7" blockId="3.[151,1437,1611,2034]" box="[199,1301,2007,2034]" pageId="3" pageNumber="374">
<emphasis id="B963A52F6925FF87FF5DF83212B713C7" bold="true" box="[199,446,2007,2033]" pageId="3" pageNumber="374">Material examined. [</emphasis>
<materialsCitation id="3B7F73606925FF87FE25F832161C13C7" box="[447,1301,2007,2034]" collectingDate="1907-07-13" collectionCode="SEHU" collectorName="S. Matsumura" country="Taiwan" location="Kanshirei" pageId="3" pageNumber="374" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<emphasis id="B963A52F6925FF87FE25F832119413C7" bold="true" box="[447,669,2007,2033]" pageId="3" pageNumber="374">
<collectionCode id="ED06E1F86925FF87FE25F832111913C7" box="[447,528,2007,2033]" collectionName="SEHU" pageId="3" pageNumber="374">SEHU</collectionCode>
]
<typeStatus id="54ACC79F6925FF87FD87F832119113C7" box="[541,664,2007,2033]" pageId="3" pageNumber="374" type="lectotype">Lectotype</typeStatus>
:
</emphasis>
<specimenCount id="9D11B2B46925FF87FD3EF83211CE13C7" box="[676,711,2007,2033]" count="1" pageId="3" pageNumber="374" type="female">1♀</specimenCount>
,
<location id="8EC82FE66925FF87FD4FF832104C13C7" LSID="urn:lsid:plazi:treatment:03BEC82B6925FF80FF0DFE66111815BE:8EC82FE66925FF87FD4FF832104C13C7" box="[725,837,2007,2033]" country="Taiwan" name="Kanshirei" pageId="3" pageNumber="374">Kanshirei</location>
,
<collectingCountry id="F30039AD6925FF87FCD5F83210AC13C7" box="[847,933,2007,2033]" name="Taiwan" pageId="3" pageNumber="374">Taiwan</collectingCountry>
;
<date id="FFA95FFD6925FF87FC2AF832173213C7" box="[944,1083,2007,2033]" pageId="3" pageNumber="374" value="1907-07-13">
<collectingDate id="EFEDA6156925FF87FC2AF832173213C7" box="[944,1083,2007,2033]" pageId="3" pageNumber="374" value="1907-07-13">13.VII.1907</collectingDate>
</date>
,
<collectorName id="26E21CEB6925FF87FBDDF83217ED13C4" box="[1095,1252,2007,2034]" pageId="3" pageNumber="374">S. Matsumura</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66922FF80FF5DFF73169D1487" box="[199,1428,150,177]" pageId="4" pageNumber="375" type="distribution">
<paragraph id="8BA8793D6922FF80FF5DFF73169D1487" blockId="4.[151,1437,150,392]" box="[199,1428,150,177]" pageId="4" pageNumber="375">
<emphasis id="B963A52F6922FF80FF5DFF7312651486" bold="true" box="[199,364,150,176]" pageId="4" pageNumber="375">Distributions.</emphasis>
<collectingCountry id="F30039AD6922FF80FEE8FF7312BC1486" box="[370,437,150,176]" name="China" pageId="4" pageNumber="375">China</collectingCountry>
(
<collectingRegion id="49D3B7DF6922FF80FE5EFF7311071486" box="[452,526,150,176]" country="China" name="Anhui" pageId="4" pageNumber="375">Anhui</collectingRegion>
,
<collectingRegion id="49D3B7DF6922FF80FD80FF73116C1486" box="[538,613,150,176]" country="China" name="Fujian" pageId="4" pageNumber="375">Fujian</collectingRegion>
,
<collectingRegion id="49D3B7DF6922FF80FDF5FF7311DB1486" box="[623,722,150,176]" country="China" name="Guizhou" pageId="4" pageNumber="375">Guizhou</collectingRegion>
,
<collectingRegion id="49D3B7DF6922FF80FD44FF7210221487" box="[734,811,151,177]" country="China" name="Hunan" pageId="4" pageNumber="375">Hunan</collectingRegion>
,
<collectingRegion id="49D3B7DF6922FF80FCACFF7310841486" box="[822,909,150,176]" country="China" name="Jiangsu" pageId="4" pageNumber="375">Jiangsu</collectingRegion>
,
<collectingRegion id="49D3B7DF6922FF80FC02FF7310FC1486" box="[920,1013,150,176]" country="China" name="Sichuan" pageId="4" pageNumber="375">Sichuan</collectingRegion>
,
<collectingRegion id="49D3B7DF6922FF80FB9AFF7317611486" box="[1024,1128,150,176]" country="China" name="Zhejiang" pageId="4" pageNumber="375">Zhejiang</collectingRegion>
),
<collectingCountry id="F30039AD6922FF80FBE1FF7317C41486" box="[1147,1229,150,176]" name="Taiwan" pageId="4" pageNumber="375">Taiwan</collectingCountry>
(
<collectingRegion id="49D3B7DF6922FF80FB41FF7316231486" box="[1243,1322,150,176]" country="Taiwan" name="Tainan" pageId="4" pageNumber="375">Tainan</collectingRegion>
) (
<figureCitation id="132C65B86922FF80FADBFF73168F1487" box="[1345,1414,150,177]" captionStart="FIGURE 5" captionStartId="7.[151,250,1946,1970]" captionTargetBox="[151,1435,1281,1923]" captionTargetId="figure-410@7.[151,1435,1281,1923]" captionTargetPageId="7" captionText="FIGURE 5. Distribution map of the genus Toyoides species from the Korean Peninsula and neighboring countries." figureDoi="http://doi.org/10.5281/zenodo.14239607" httpUri="https://zenodo.org/record/14239607/files/figure.png" pageId="4" pageNumber="375">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66922FF80FF5DFF5F17C914E2" box="[199,1216,186,212]" pageId="4" pageNumber="375" type="biology_ecology">
<paragraph id="8BA8793D6922FF80FF5DFF5F17C914E2" blockId="4.[151,1437,150,392]" box="[199,1216,186,212]" pageId="4" pageNumber="375">
<emphasis id="B963A52F6922FF80FF5DFF5F125F14E2" bold="true" box="[199,342,186,212]" pageId="4" pageNumber="375">Host plants.</emphasis>
<taxonomicName id="4C1702BE6922FF80FEC7FF5E102914E2" authority="(Yang, 1989)" baseAuthorityName="Yang" baseAuthorityYear="1989" box="[349,800,186,212]" class="Liliopsida" family="Poaceae" genus="Pennisetum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="4" pageNumber="375" phylum="Tracheophyta" rank="species" species="alopecuroides">
<emphasis id="B963A52F6922FF80FEC7FF5E118C14E2" box="[349,645,186,212]" italics="true" pageId="4" pageNumber="375">Pennisetum alopecuroides</emphasis>
(
<bibRefCitation id="EF8604CC6922FF80FD09FF5F101014E2" author="Yang, C. T." box="[659,793,186,212]" pageId="4" pageNumber="375" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang, 1989</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C1702BE6922FF80FCB6FF5F17B514E2" authority="(Ding, 2006)" baseAuthorityName="Ding" baseAuthorityYear="2006" box="[812,1212,186,212]" class="Liliopsida" family="Poaceae" genus="Miscanthus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="375" phylum="Tracheophyta" rank="species" species="floridulus">
<emphasis id="B963A52F6922FF80FCB6FF5F172814E2" box="[812,1057,186,212]" italics="true" pageId="4" pageNumber="375">Miscanthus floridulus</emphasis>
(
<bibRefCitation id="EF8604CC6922FF80FBAAFF5F17BD14E2" author="Ding, J. H." box="[1072,1204,186,212]" pageId="4" pageNumber="375" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding, 2006</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66922FF80FF5DFF3B111815BE" pageId="4" pageNumber="375" type="discussion">
<paragraph id="8BA8793D6922FF80FF5DFF3B111815BE" blockId="4.[151,1437,150,392]" pageId="4" pageNumber="375">
<emphasis id="B963A52F6922FF80FF5DFF3B123214CE" bold="true" box="[199,315,222,248]" pageId="4" pageNumber="375">Remarks.</emphasis>
<bibRefCitation id="EF8604CC6922FF80FED9FF3B12E614CE" author="Ishihara, T." box="[323,495,222,248]" pageId="4" pageNumber="375" pagination="1 - 102" refId="ref5544" refString="Ishihara, T. (1949) Revision of the Araeopidae of Japan, Ryukyu Island and Formosa (Hemiptera). Scientific Reports of the Matsuyama Agricultural College, 2, 1 - 102." type="journal article" year="1949">Ishihara (1949)</bibRefCitation>
transferred this species to the genus
<taxonomicName id="4C1702BE6922FF80FC08FF3B164114CE" authority="Puton. However, Yang (1989)" authorityName="Puton. However, Yang" authorityYear="1989" box="[914,1352,222,249]" class="Insecta" family="Delphacidae" genus="Euidella" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6922FF80FC08FF3B10F814CE" box="[914,1009,222,248]" italics="true" pageId="4" pageNumber="375">Euidella</emphasis>
Puton. However,
<bibRefCitation id="EF8604CC6922FF80FB26FF3A164114CE" author="Yang, C. T." box="[1212,1352,222,249]" pageId="4" pageNumber="375" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang (1989)</bibRefCitation>
</taxonomicName>
moved this species to the genus
<taxonomicName id="4C1702BE6922FF80FE24FEE711A8152B" authority="Matsumura" authorityName="Matsumura. Ding" authorityYear="2006" box="[446,673,258,285]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6922FF80FE24FEE7111C152A" box="[446,533,258,284]" italics="true" pageId="4" pageNumber="375">Kakuna</emphasis>
Matsumura
</taxonomicName>
, while
<bibRefCitation id="EF8604CC6922FF80FD63FEE7108F152A" author="Ding, J. H." box="[761,902,258,284]" pageId="4" pageNumber="375" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding (2006)</bibRefCitation>
resisted the genus
<taxonomicName id="4C1702BE6922FF80FBF2FEE717C2152A" authorityName="Park &amp; Jung" authorityYear="2024" baseAuthorityName="Park &amp; Jung" baseAuthorityYear="2024" box="[1128,1227,258,284]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6922FF80FBF2FEE717C2152A" box="[1128,1227,258,284]" italics="true" pageId="4" pageNumber="375">Toyoides</emphasis>
</taxonomicName>
with this species. The type material was not designated in the original description (
<bibRefCitation id="EF8604CC6922FF80FC1BFEC317421577" author="Matsumura, S." box="[897,1099,294,321]" pageId="4" pageNumber="375" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935</bibRefCitation>
) and in the present study, we designated the
<typeStatus id="54ACC79F6922FF80FEA6FEAF12AA1552" box="[316,419,330,356]" pageId="4" pageNumber="375" type="lectotype">lectotype</typeStatus>
which is deposited in SEHU. The current type location is currently Guanziling (= Kanshirei) in Baihe District,
<collectingRegion id="49D3B7DF6922FF80FEC4FE8B12A415BE" box="[350,429,366,392]" country="Taiwan" name="Tainan" pageId="4" pageNumber="375">Tainan</collectingRegion>
,
<collectingCountry id="F30039AD6922FF80FE2DFE8B110415BE" box="[439,525,366,392]" name="Taiwan" pageId="4" pageNumber="375">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,654 @@
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<paragraph id="8BA8793D6927FF85FF0DFB59113410E0" blockId="1.[151,573,1211,1238]" box="[151,573,1211,1238]" pageId="1" pageNumber="372">
<heading id="D0E0CE516927FF85FF0DFB59113410E0" bold="true" box="[151,573,1211,1238]" fontSize="11" level="1" pageId="1" pageNumber="372" reason="1">
<emphasis id="B963A52F6927FF85FF0DFB59113410E0" bold="true" box="[151,573,1211,1238]" pageId="1" pageNumber="372">
Genus
<taxonomicName id="4C1702BE6927FF85FF74FB5E113410E0" ID-CoL="7Y6P" authority="Matsumura, 1935" authorityName="Matsumura" authorityYear="1935" box="[238,573,1211,1238]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6927FF85FF74FB5E125E10E0" bold="true" box="[238,343,1211,1238]" italics="true" pageId="1" pageNumber="372">Toyoides</emphasis>
<bibRefCitation id="EF8604CC6927FF85FEC5FB59113410E0" author="Matsumura, S." box="[351,573,1212,1238]" pageId="1" pageNumber="372" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66927FF85FF0DFB1710AF113C" box="[151,934,1266,1290]" pageId="1" pageNumber="372" type="type_taxon">
<paragraph id="8BA8793D6927FF85FF0DFB1710AF113C" blockId="1.[151,934,1266,1351]" box="[151,934,1266,1290]" pageId="1" pageNumber="372">
<typeStatus id="54ACC79F6927FF85FF0DFB1713C0113C" box="[151,201,1266,1290]" pageId="1" pageNumber="372">Type</typeStatus>
species:
<taxonomicName id="4C1702BE6927FF85FEBCFB1711A2113C" authority="Matsumura, 1935" authorityName="Matsumura" authorityYear="1935" box="[294,683,1266,1290]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="species" species="albipennis">
<emphasis id="B963A52F6927FF85FEBCFB1712F9113C" box="[294,496,1266,1290]" italics="true" pageId="1" pageNumber="372">Toyoides albipennis</emphasis>
<bibRefCitation id="EF8604CC6927FF85FE6CFB1711A2113C" author="Matsumura, S." box="[502,683,1266,1290]" pageId="1" pageNumber="372" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66927FF85FF0DFAF5111E1171" pageId="1" pageNumber="372" type="reference_group">
<paragraph id="8BA8793D6927FF85FF0DFAF51183111F" blockId="1.[151,934,1266,1351]" box="[151,650,1296,1321]" pageId="1" pageNumber="372">
<treatmentCitationGroup id="AB075E136927FF85FF0DFAF51183111F" box="[151,650,1296,1321]" pageId="1" pageNumber="372">
<treatmentCitation id="0AB65F2C6927FF85FF0DFAF512DA111F" author="Matsumura, S." box="[151,467,1296,1321]" page="78" pageId="1" pageNumber="372" year="1935">
<taxonomicName id="4C1702BE6927FF85FF0DFAF512DA111F" ID-CoL="7Y6P" authority="Matsumura, 1935: 78" authorityName="Matsumura" authorityPageNumber="78" authorityYear="1935" box="[151,467,1296,1321]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6927FF85FF0DFAF513F9111E" box="[151,240,1296,1320]" italics="true" pageId="1" pageNumber="372">Toyoides</emphasis>
<bibRefCitation id="EF8604CC6927FF85FF6DFAF412DA111F" author="Matsumura, S." box="[247,467,1296,1321]" pageId="1" pageNumber="372" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935: 78</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AB65F2C6927FF85FE47FAF5118F111F" author="Ding, J. H." box="[477,646,1296,1321]" page="303" pageId="1" pageNumber="372" year="2006">
<bibRefCitation id="EF8604CC6927FF85FE47FAF5118F111F" author="Ding, J. H." box="[477,646,1296,1321]" pageId="1" pageNumber="372" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding, 2006: 303</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BA8793D6927FF85FF0DFACA111E1171" blockId="1.[151,934,1266,1351]" box="[151,535,1327,1351]" pageId="1" pageNumber="372">
<treatmentCitationGroup id="AB075E136927FF85FF0DFACA111E1171" box="[151,535,1327,1351]" pageId="1" pageNumber="372">
<treatmentCitation id="0AB65F2C6927FF85FF0DFACA12CE1171" author="Matsumura, S." box="[151,455,1327,1351]" page="77" pageId="1" pageNumber="372" year="1935">
<taxonomicName id="4C1702BE6927FF85FF0DFACA12CE1171" ID-CoL="4DL6" authority="Matsumura, 1935: 77" authorityName="Matsumura" authorityPageNumber="77" authorityYear="1935" box="[151,455,1327,1351]" class="Insecta" family="Delphacidae" genus="Epunka" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6927FF85FF0DFACA13ED1171" box="[151,228,1327,1351]" italics="true" pageId="1" pageNumber="372">Epunka</emphasis>
<bibRefCitation id="EF8604CC6927FF85FF71FAD512CE1171" author="Matsumura, S." box="[235,455,1327,1351]" pageId="1" pageNumber="372" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935: 77</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis id="B963A52F6927FF85FE4BFACA111E1171" bold="true" box="[465,535,1327,1351]" pageId="1" pageNumber="372">syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66927FF85FF0DFA90167011CD" pageId="1" pageNumber="372" type="diagnosis">
<paragraph id="8BA8793D6927FF85FF0DFA90167011CD" blockId="1.[151,1437,1397,2035]" pageId="1" pageNumber="372">
<emphasis id="B963A52F6927FF85FF0DFA90121B11B9" bold="true" box="[151,274,1397,1423]" pageId="1" pageNumber="372">Diagnosis.</emphasis>
Distinguished by other genera frons with median carina forked above level of upper margin of eyes (
<figureCitation id="132C65B86927FF85FAF0FA9013B01185" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[151,1436,477,1456]" captionTargetId="figure-420@3.[151,1436,502,1460]" captionTargetPageId="3" captionText="FIGURE 1. Lectotype of Toyoides albipennis Matsumura. A. habitus, dorsal view; B. ditto, lateral view; C. frons, ventral view; D. right hind leg with spur; E. lectotype labels. Abbreviations: f, front; b, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239599" httpUri="https://zenodo.org/record/14239599/files/figure.png" pageId="1" pageNumber="372">Figs 1C</figureCitation>
,
<figureCitation id="132C65B86927FF85FF5CFA7C13E21185" box="[198,235,1433,1459]" captionStart="FIGURE 2" captionStartId="4.[151,250,1653,1677]" captionTargetBox="[151,1431,678,1626]" captionTargetId="figure-252@4.[151,1436,669,1635]" captionTargetPageId="4" captionText="FIGURE 2. Lectotype and paralectotype of Toyoides bilineata Matsumura comb. n. A. habitus, dorsal view; B. lectotype, habitus, lateral view; C. ditto, frons, ventral view; D. ditto, right hind leg with spur; E. lectotype and paralectotype labels. Abbreviations: f, front; b, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239601" httpUri="https://zenodo.org/record/14239601/files/figure.png" pageId="1" pageNumber="372">2C</figureCitation>
); anal segment with one pair of spinosus processes protrudes caudally (
<figureCitation id="132C65B86927FF85FB8BFA7C17981185" box="[1041,1169,1433,1459]" captionStart="FIGURE 4" captionStartId="6.[151,250,1530,1554]" captionTargetBox="[158,1429,188,1499]" captionTargetId="figure-20@6.[151,1436,181,1506]" captionTargetPageId="6" captionText="FIGURE 4. Male genitalia of Toyoides bilineata Matsumura comb. n. A. genitalia, caudal view; B. ditto, lateral view; C. anal segment, caudal view; D. ditto, lateral view; E. pygofer, caudal view; F. ditto, lateral view; G. genital styles, caudoventral view; H. ditto, lateral view; I. aedeagus with suspensorium, left-lateral view; J. aedeagus without suspensorium, dorsal view; K. suspensorium. Scale bar: AB, EF: 0.5 mm; CD, GH: 0.3 mm; IK: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239605" httpUri="https://zenodo.org/record/14239605/files/figure.png" pageId="1" pageNumber="372">Figs 4CD</figureCitation>
); genital styles forceps-shaped, converging apically (
<figureCitation id="132C65B86927FF85FE41FA58115311E1" box="[475,602,1469,1495]" captionStart="FIGURE 4" captionStartId="6.[151,250,1530,1554]" captionTargetBox="[158,1429,188,1499]" captionTargetId="figure-20@6.[151,1436,181,1506]" captionTargetPageId="6" captionText="FIGURE 4. Male genitalia of Toyoides bilineata Matsumura comb. n. A. genitalia, caudal view; B. ditto, lateral view; C. anal segment, caudal view; D. ditto, lateral view; E. pygofer, caudal view; F. ditto, lateral view; G. genital styles, caudoventral view; H. ditto, lateral view; I. aedeagus with suspensorium, left-lateral view; J. aedeagus without suspensorium, dorsal view; K. suspensorium. Scale bar: AB, EF: 0.5 mm; CD, GH: 0.3 mm; IK: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239605" httpUri="https://zenodo.org/record/14239605/files/figure.png" pageId="1" pageNumber="372">Figs 4A, G</figureCitation>
); aedeagus hook-shaped, upper process longer than lower distinctly, curved ventrally; suspensorium ring-shaped, length longer than width visibly, fused at middle of aedeagus (
<figureCitation id="132C65B86927FF85FB6CFA04166511CD" box="[1270,1388,1505,1531]" captionStart="FIGURE 4" captionStartId="6.[151,250,1530,1554]" captionTargetBox="[158,1429,188,1499]" captionTargetId="figure-20@6.[151,1436,181,1506]" captionTargetPageId="6" captionText="FIGURE 4. Male genitalia of Toyoides bilineata Matsumura comb. n. A. genitalia, caudal view; B. ditto, lateral view; C. anal segment, caudal view; D. ditto, lateral view; E. pygofer, caudal view; F. ditto, lateral view; G. genital styles, caudoventral view; H. ditto, lateral view; I. aedeagus with suspensorium, left-lateral view; J. aedeagus without suspensorium, dorsal view; K. suspensorium. Scale bar: AB, EF: 0.5 mm; CD, GH: 0.3 mm; IK: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239605" httpUri="https://zenodo.org/record/14239605/files/figure.png" pageId="1" pageNumber="372">Figs 4I, K</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66927FF85FF5DF9E011E713B1" pageId="1" pageNumber="372" type="description">
<paragraph id="8BA8793D6927FF85FF5DF9E011E713B1" blockId="1.[151,1437,1397,2035]" pageId="1" pageNumber="372">
<emphasis id="B963A52F6927FF85FF5DF9E0127C1229" bold="true" box="[199,373,1541,1567]" pageId="1" pageNumber="372">Redescription.</emphasis>
Recognized by body length
<quantity id="4CEFD4D86927FF85FD2FF9E010061216" box="[693,783,1541,1568]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="1" pageNumber="372" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5-7 mm</quantity>
, large and macropterous taxon; general coloration of body light green to brown; wings hyaline or brown.Head narrower than pronotum, in lateral view head anteriorly projecting, appearing slightly upcurved; carinae of vertex other vertex carinae visible. Frons parallel, narrower between below level of eyes; median carina forked above level of upper margin of eyes; vertex much longer than wide; antennae cylindrical; segment II nearly two times longer than segment I; segment II with several sensory pits; lateral ocelli not visible, near anteroventral margin of compound eye. Lateral carinae of pronotum not reaching posterior margin. Hind tibiae with one lateral spine; spur long and triangular, black-tipped teeth. Male pygofer longer than wide in lateral and caudal view; opening of pygofer semicircular. Genital styles forceps-shaped, converging and narrowing towards the apex. Aedeagus hook-shaped, overlapped with suspensorium, dividing medially and upper appendage much longer than lower. Suspensorium oval, longer than wide. Anal segment short, protruded caudally and curved to ventrally (modified from
<bibRefCitation id="EF8604CC6927FF85FE4AF88B115A13B1" author="Yang, C. T." box="[464,595,1901,1928]" pageId="1" pageNumber="372" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang, 1989</bibRefCitation>
;
<bibRefCitation id="EF8604CC6927FF85FDC5F88811EB13B1" author="Ding, J. H." box="[607,738,1901,1927]" pageId="1" pageNumber="372" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding, 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66927FF85FF5DF874116F13C5" pageId="1" pageNumber="372" type="distribution">
<paragraph id="8BA8793D6927FF85FF5DF874116F13C5" blockId="1.[151,1437,1397,2035]" pageId="1" pageNumber="372">
<emphasis id="B963A52F6927FF85FF5DF8741268139D" bold="true" box="[199,353,1937,1963]" pageId="1" pageNumber="372">Distribution.</emphasis>
<collectingCountry id="F30039AD6927FF85FEF2F87412FC139A" box="[360,501,1937,1964]" name="South Korea" pageId="1" pageNumber="372">South Korea</collectingCountry>
(
<collectingRegion id="49D3B7DF6927FF85FD9FF874117E139D" box="[517,631,1937,1963]" country="South Korea" name="Gyeonggi" pageId="1" pageNumber="372">Gyeonggi</collectingRegion>
)
<superScript id="7C62D4756927FF85FDE4F874118F13A9" attach="none" box="[638,646,1937,1951]" fontSize="6" pageId="1" pageNumber="372">*</superScript>
,
<collectingCountry id="F30039AD6927FF85FD0FF87411D1139D" box="[661,728,1937,1963]" name="China" pageId="1" pageNumber="372">China</collectingCountry>
(
<collectingRegion id="49D3B7DF6927FF85FD72F874103B139D" box="[744,818,1937,1963]" country="China" name="Anhui" pageId="1" pageNumber="372">Anhui</collectingRegion>
,
<collectingRegion id="49D3B7DF6927FF85FCA4F8741080139D" box="[830,905,1937,1963]" country="China" name="Fujian" pageId="1" pageNumber="372">Fujian</collectingRegion>
,
<collectingRegion id="49D3B7DF6927FF85FC0EF87410FE139D" box="[916,1015,1937,1963]" country="China" name="Guizhou" pageId="1" pageNumber="372">Guizhou</collectingRegion>
,
<collectingRegion id="49D3B7DF6927FF85FB99F8771759139A" box="[1027,1104,1938,1964]" country="China" name="Hunan" pageId="1" pageNumber="372">Hunan</collectingRegion>
,
<collectingRegion id="49D3B7DF6927FF85FBC7F87417BD139D" box="[1117,1204,1937,1963]" country="China" name="Jiangsu" pageId="1" pageNumber="372">Jiangsu</collectingRegion>
,
<collectingRegion id="49D3B7DF6927FF85FB25F8741615139D" box="[1215,1308,1937,1963]" country="China" name="Sichuan" pageId="1" pageNumber="372">Sichuan</collectingRegion>
,
<collectingRegion id="49D3B7DF6927FF85FAB2F8741699139D" box="[1320,1424,1937,1963]" country="China" name="Zhejiang" pageId="1" pageNumber="372">Zhejiang</collectingRegion>
),
<collectingCountry id="F30039AD6927FF85FF0DF85313DE13E6" box="[151,215,1974,2000]" name="Japan" pageId="1" pageNumber="372">Japan</collectingCountry>
(
<collectingRegion id="49D3B7DF6927FF85FF7FF850125113F9" box="[229,344,1973,1999]" country="Japan" name="Hokkaido" pageId="1" pageNumber="372">Hokkaido</collectingRegion>
, Honshu, Kyushu, Tsushima Island),
<collectingCountry id="F30039AD6927FF85FD6DF850104013F9" box="[759,841,1973,1999]" name="Taiwan" pageId="1" pageNumber="372">Taiwan</collectingCountry>
(
<collectingRegion id="49D3B7DF6927FF85FCCDF85010DC13F9" box="[855,981,1973,1999]" country="Taiwan" name="Kaohsiung" pageId="1" pageNumber="372">Kaohsiung</collectingRegion>
,
<collectingRegion id="49D3B7DF6927FF85FC44F850172513F9" box="[990,1068,1973,1999]" country="Taiwan" name="Tainan" pageId="1" pageNumber="372">Tainan</collectingRegion>
) (
<bibRefCitation id="EF8604CC6927FF85FBDBF850160213E6" author="Matsumura, S." box="[1089,1291,1973,2000]" pageId="1" pageNumber="372" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935</bibRefCitation>
;
<bibRefCitation id="EF8604CC6927FF85FA8FF853169E13F9" author="Yang, C. T." box="[1301,1431,1973,2000]" pageId="1" pageNumber="372" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang, 1989</bibRefCitation>
;
<bibRefCitation id="EF8604CC6927FF85FF0DF83C121013C5" author="Ding, J. H." box="[151,281,2009,2035]" pageId="1" pageNumber="372" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding, 2006</bibRefCitation>
;
<bibRefCitation id="EF8604CC6927FF85FEBEF83C115313C5" author="Hayashi, M. &amp; Fujinuma, S." box="[292,602,2009,2035]" pageId="1" pageNumber="372" pagination="323 - 355" refId="ref5480" refString="Hayashi, M. &amp; Fujinuma, S. (2016) Part Fulgoromorpha. In: Entomological Society of Japan (2016) Catalogue of the insects of Japan. Vol. 4. Paraneoptera (Psocodea, Thysanoptera, Hemiptera). 4. Editorial Committee of Catalogue of the Insects of Japan, 4, pp. 323 - 355." type="journal article" year="2016">Hayashi &amp; Fujinuma, 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66924FF86FF5DFF72177214CF" pageId="2" pageNumber="373" type="etymology">
<paragraph id="8BA8793D6924FF86FF5DFF72177214CF" blockId="2.[151,1437,150,1041]" pageId="2" pageNumber="373">
<emphasis id="B963A52F6924FF86FF5DFF7212471487" bold="true" box="[199,334,151,177]" pageId="2" pageNumber="373">Etymology.</emphasis>
The original description does not provide the etymology (
<bibRefCitation id="EF8604CC6924FF86FC64FF7217C41487" author="Matsumura, S." box="[1022,1229,150,177]" pageId="2" pageNumber="373" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935</bibRefCitation>
); it simply states, Closely allied to
<taxonomicName id="4C1702BE6924FF86FEF9FF5E12EE14E3" authority="DIST." authorityName="DIST." box="[355,487,187,213]" class="Insecta" family="Delphacidae" genus="Toya" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FEF9FF5E129114E2" box="[355,408,187,212]" italics="true" pageId="2" pageNumber="373">Toya</emphasis>
DIST.
</taxonomicName>
, but differs from the latter as follows: …’. It is speculated that the genus name was formed by combining
<taxonomicName id="4C1702BE6924FF86FE00FF3A12C614CE" box="[410,463,223,248]" class="Insecta" family="Delphacidae" genus="Toya" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FE00FF3A12C614CE" box="[410,463,223,248]" italics="true" pageId="2" pageNumber="373">Toya</emphasis>
</taxonomicName>
with the suffix -oides (likeness), along with a Latin suffix.
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66924FF86FF5DFEE612501553" pageId="2" pageNumber="373" type="biology_ecology">
<paragraph id="8BA8793D6924FF86FF5DFEE612501553" blockId="2.[151,1437,150,1041]" pageId="2" pageNumber="373">
<emphasis id="B963A52F6924FF86FF5DFEE61259152B" bold="true" box="[199,336,259,285]" pageId="2" pageNumber="373">Host plants.</emphasis>
<taxonomicName id="4C1702BE6924FF86FECCFEE61167152B" baseAuthorityName="Yang" baseAuthorityYear="1989" box="[342,622,259,285]" class="Liliopsida" family="Poaceae" genus="Pennisetum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="2" pageNumber="373" phylum="Tracheophyta" rank="species" species="alopecuroides">
<emphasis id="B963A52F6924FF86FECCFEE61167152B" box="[342,622,259,285]" italics="true" pageId="2" pageNumber="373">Pennisetum alopecuroides</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1702BE6924FF86FD38FEE61083152B" baseAuthorityName="Ding" baseAuthorityYear="2006" box="[674,906,259,285]" class="Liliopsida" family="Poaceae" genus="Miscanthus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="373" phylum="Tracheophyta" rank="species" species="floridulus">
<emphasis id="B963A52F6924FF86FD38FEE61083152B" box="[674,906,259,285]" italics="true" pageId="2" pageNumber="373">Miscanthus floridulus</emphasis>
</taxonomicName>
have been recorded as hosts of
<taxonomicName id="4C1702BE6924FF86FB44FEE61661152B" box="[1246,1384,259,285]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="albipennis">
<emphasis id="B963A52F6924FF86FB44FEE617E4152A" box="[1246,1261,259,284]" italics="true" pageId="2" pageNumber="373">T</emphasis>
.
<emphasis id="B963A52F6924FF86FB60FEE61661152B" box="[1274,1384,259,285]" italics="true" pageId="2" pageNumber="373">albipennis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C1702BE6924FF86FF0DFEC2127C1577" box="[151,373,295,321]" class="Liliopsida" family="Poaceae" genus="Phacelurus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="373" phylum="Tracheophyta" rank="species" species="latifolius">
<emphasis id="B963A52F6924FF86FF0DFEC2127C1577" box="[151,373,295,321]" italics="true" pageId="2" pageNumber="373">Phacelurus latifolius</emphasis>
</taxonomicName>
is newly recorded as a host of
<taxonomicName id="4C1702BE6924FF86FD5FFEC210491577" authorityName="Park &amp; Jung" authorityYear="2024" baseAuthorityName="Matsumura" baseAuthorityYear="1935" box="[709,832,295,321]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="bilineata" status="comb. n.">
<emphasis id="B963A52F6924FF86FD5FFEC211DD1576" box="[709,724,295,320]" italics="true" pageId="2" pageNumber="373">T</emphasis>
.
<emphasis id="B963A52F6924FF86FD78FEC210491577" box="[738,832,295,321]" italics="true" pageId="2" pageNumber="373">bilineata</emphasis>
</taxonomicName>
<emphasis id="B963A52F6924FF86FCDDFEC210A01577" bold="true" box="[839,937,295,321]" pageId="2" pageNumber="373">
<taxonomicNameLabel id="A25018546924FF86FCDDFEC210A01577" box="[839,937,295,321]" pageId="2" pageNumber="373" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
All hosts belong to the subfamily
<taxonomicName id="4C1702BE6924FF86FA86FEC216951577" box="[1308,1436,295,321]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="2" pageNumber="373" phylum="Tracheophyta" rank="subFamily" subFamily="Panicoideae">Panicoideae</taxonomicName>
(
<taxonomicName id="4C1702BE6924FF86FF05FEAE13E01553" box="[159,233,331,357]" class="Liliopsida" kingdom="Plantae" order="Poales" pageId="2" pageNumber="373" phylum="Tracheophyta" rank="order">Poales</taxonomicName>
:
<taxonomicName id="4C1702BE6924FF86FF68FEAE12471553" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[242,334,331,357]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="2" pageNumber="373" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66924FF86FF5DFE8A16191027" pageId="2" pageNumber="373" type="discussion">
<paragraph id="8BA8793D6924FF86FF5DFE8A1683160B" blockId="2.[151,1437,150,1041]" pageId="2" pageNumber="373">
<emphasis id="B963A52F6924FF86FF5DFE8A123215BF" bold="true" box="[199,315,367,393]" pageId="2" pageNumber="373">Remarks.</emphasis>
<emphasis id="B963A52F6924FF86FEDCFE8A11B915BF" box="[326,688,367,393]" italics="true" pageId="2" pageNumber="373">
Synopsis of the genus
<taxonomicName id="4C1702BE6924FF86FDD7FE8A11B915BF" authorityName="Matsumura" authorityYear="1935" box="[589,688,367,393]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">Toyoides</taxonomicName>
</emphasis>
. After
<bibRefCitation id="EF8604CC6924FF86FC9CFE8A10D215BF" author="Matsumura, S." box="[774,987,366,393]" pageId="2" pageNumber="373" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura (1935)</bibRefCitation>
initially described this taxon,
<bibRefCitation id="EF8604CC6924FF86FAD8FE8A13EB159B" author="Ishihara, T." pageId="2" pageNumber="373" pagination="1 - 102" refId="ref5544" refString="Ishihara, T. (1949) Revision of the Araeopidae of Japan, Ryukyu Island and Formosa (Hemiptera). Scientific Reports of the Matsuyama Agricultural College, 2, 1 - 102." type="journal article" year="1949">Ishihara (1949)</bibRefCitation>
transferred it to the genus
<taxonomicName id="4C1702BE6924FF86FD83FE761099159B" authority="Puton, and Yang (1989)" authorityName="Puton, and Yang" authorityYear="1989" box="[537,912,403,429]" class="Insecta" family="Delphacidae" genus="Euidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FD83FE761171159B" box="[537,632,403,429]" italics="true" pageId="2" pageNumber="373">Euidella</emphasis>
Puton, and
<bibRefCitation id="EF8604CC6924FF86FC99FE761099159B" author="Yang, C. T." box="[771,912,403,429]" pageId="2" pageNumber="373" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang (1989)</bibRefCitation>
</taxonomicName>
reclassified to the genus
<taxonomicName id="4C1702BE6924FF86FB2CFE76122D15E7" authority="Matsumura. Ding (2006)" authorityName="Matsumura. Ding" authorityYear="2006" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FB2CFE761604159B" box="[1206,1293,403,429]" italics="true" pageId="2" pageNumber="373">Kakuna</emphasis>
Matsumura.
<bibRefCitation id="EF8604CC6924FF86FF0DFE52122D15E7" author="Ding, J. H." box="[151,292,439,465]" pageId="2" pageNumber="373" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding (2006)</bibRefCitation>
</taxonomicName>
resisted this genus and state that this genus differs from the genus
<taxonomicName id="4C1702BE6924FF86FBA9FE52178315E7" authorityName="Matsumura. Ding" authorityYear="2006" box="[1075,1162,439,465]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FBA9FE52178315E7" box="[1075,1162,439,465]" italics="true" pageId="2" pageNumber="373">Kakuna</emphasis>
</taxonomicName>
based on the hind tibia spines and its genital characteristics are similar to those of the genus
<taxonomicName id="4C1702BE6924FF86FC02FE3E10ED15C3" box="[920,996,475,501]" class="Insecta" family="Delphacidae" genus="Euides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FC02FE3E10ED15C3" box="[920,996,475,501]" italics="true" pageId="2" pageNumber="373">Euides</emphasis>
</taxonomicName>
. In summary,
<taxonomicName id="4C1702BE6924FF86FB1EFE3E17EE15C3" authorityName="Matsumura" authorityYear="1935" box="[1156,1255,475,501]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FB1EFE3E17EE15C3" box="[1156,1255,475,501]" italics="true" pageId="2" pageNumber="373">Toyoides</emphasis>
</taxonomicName>
is distinguished from other related genera based on the forked portion of the frons and hook-shaped aedeagus. In this study, we have redescribed the generic characteristics (detailed taxonomic histories are provided in the remarks of each species).
</paragraph>
<paragraph id="8BA8793D6924FF86FF5DFDA216191027" blockId="2.[151,1437,150,1041]" pageId="2" pageNumber="373">
<emphasis id="B963A52F6924FF86FF5DFDA211131657" box="[199,538,583,609]" italics="true" pageId="2" pageNumber="373">
Synonymy of the genus
<taxonomicName id="4C1702BE6924FF86FE5EFDA211131657" authorityName="Matsumura" authorityYear="1935" box="[452,538,583,609]" class="Insecta" family="Delphacidae" genus="Epunka" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">Epunka</taxonomicName>
</emphasis>
. The genus
<taxonomicName id="4C1702BE6924FF86FD01FDA2107C1657" authority="Matsumura" authorityName="Matsumura" authorityYear="1935" box="[667,885,583,609]" class="Insecta" family="Delphacidae" genus="Epunka" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FD01FDA211F81657" box="[667,753,583,609]" italics="true" pageId="2" pageNumber="373">Epunka</emphasis>
Matsumura
</taxonomicName>
is endemic to
<collectingCountry id="F30039AD6924FF86FB94FDA217581657" box="[1038,1105,583,609]" name="Japan" pageId="2" pageNumber="373">Japan</collectingCountry>
, distributed only in
<collectingRegion id="49D3B7DF6924FF86FABDFDA216911657" box="[1319,1432,583,609]" country="Japan" name="Hokkaido" pageId="2" pageNumber="373">Hokkaido</collectingRegion>
, Honshu, Kyushu and Tsushima Island (
<bibRefCitation id="EF8604CC6924FF86FDFBFD8E102616B3" author="Matsumura, S." box="[609,815,618,645]" pageId="2" pageNumber="373" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935</bibRefCitation>
;
<bibRefCitation id="EF8604CC6924FF86FCA5FD8E178916B3" author="Hayashi, M. &amp; Fujinuma, S." box="[831,1152,619,645]" pageId="2" pageNumber="373" pagination="323 - 355" refId="ref5480" refString="Hayashi, M. &amp; Fujinuma, S. (2016) Part Fulgoromorpha. In: Entomological Society of Japan (2016) Catalogue of the insects of Japan. Vol. 4. Paraneoptera (Psocodea, Thysanoptera, Hemiptera). 4. Editorial Committee of Catalogue of the Insects of Japan, 4, pp. 323 - 355." type="journal article" year="2016">Hayashi &amp; Fujinuma, 2016</bibRefCitation>
). Historically, this taxon comprised only one species,
<taxonomicName id="4C1702BE6924FF86FE4CFD6A1151169F" authorityName="Park &amp; Jung" authorityYear="2024" baseAuthorityName="Matsumura" baseAuthorityYear="1935" box="[470,600,655,681]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="bilineata">
<emphasis id="B963A52F6924FF86FE4CFD6A12EE169E" box="[470,487,655,680]" italics="true" pageId="2" pageNumber="373">E</emphasis>
.
<emphasis id="B963A52F6924FF86FE6EFD6A1151169F" box="[500,600,655,681]" italics="true" pageId="2" pageNumber="373">bilineata</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF8604CC6924FF86FDFFFD6A1019169F" author="Ishihara, T." box="[613,784,655,681]" pageId="2" pageNumber="373" pagination="1 - 102" refId="ref5544" refString="Ishihara, T. (1949) Revision of the Araeopidae of Japan, Ryukyu Island and Formosa (Hemiptera). Scientific Reports of the Matsuyama Agricultural College, 2, 1 - 102." type="journal article" year="1949">Ishihara (1949)</bibRefCitation>
synonymized this genus with the genus
<taxonomicName id="4C1702BE6924FF86FB54FD6A1624169F" authorityName="Puton, and Yang" authorityYear="1989" box="[1230,1325,655,681]" class="Insecta" family="Delphacidae" genus="Euidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FB54FD6A1624169F" box="[1230,1325,655,681]" italics="true" pageId="2" pageNumber="373">Euidella</emphasis>
</taxonomicName>
, although this classification was not accepted in more recent publications (
<bibRefCitation id="EF8604CC6924FF86FC13FD56170216FB" author="Nast, J." box="[905,1035,691,717]" pageId="2" pageNumber="373" refId="ref5641" refString="Nast, J. (1972) Palaearctic Auchenorrhyncha (Homoptera). Polish Scientific publishers, Warszawa, 550 pp." type="book" year="1972">Nast, 1972</bibRefCitation>
;
<bibRefCitation id="EF8604CC6924FF86FB80FD56165216FB" author="Hayashi, M. &amp; Fujinuma, S." box="[1050,1371,691,717]" pageId="2" pageNumber="373" pagination="323 - 355" refId="ref5480" refString="Hayashi, M. &amp; Fujinuma, S. (2016) Part Fulgoromorpha. In: Entomological Society of Japan (2016) Catalogue of the insects of Japan. Vol. 4. Paraneoptera (Psocodea, Thysanoptera, Hemiptera). 4. Editorial Committee of Catalogue of the Insects of Japan, 4, pp. 323 - 355." type="journal article" year="2016">Hayashi &amp; Fujinuma, 2016</bibRefCitation>
). The genera
<taxonomicName id="4C1702BE6924FF86FF70FD32124916C7" authorityName="Matsumura" authorityYear="1935" box="[234,320,727,753]" class="Insecta" family="Delphacidae" genus="Epunka" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FF70FD32124916C7" box="[234,320,727,753]" italics="true" pageId="2" pageNumber="373">Epunka</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1702BE6924FF86FEE2FD3212D216C7" authorityName="Matsumura" authorityYear="1935" box="[376,475,727,753]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FEE2FD3212D216C7" box="[376,475,727,753]" italics="true" pageId="2" pageNumber="373">Toyoides</emphasis>
</taxonomicName>
were first established in a paper by
<bibRefCitation id="EF8604CC6924FF86FCE2FD32174716C7" author="Matsumura, S." box="[888,1102,726,753]" pageId="2" pageNumber="373" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura (1935)</bibRefCitation>
. However, the morphological differences between the two genera were not presented; only the differences between each genus and their closely related genera (
<emphasis id="B963A52F6924FF86FEDFFCFA12FC170F" box="[325,501,799,825]" italics="true" pageId="2" pageNumber="373">
<taxonomicName id="4C1702BE6924FF86FEDFFCFA1292170F" authorityName="Matsumura" authorityYear="1935" box="[325,411,799,825]" class="Insecta" family="Delphacidae" genus="Epunka" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">Epunka</taxonomicName>
<taxonomicName id="4C1702BE6924FF86FE33FCFA12FC170F" box="[425,501,799,825]" class="Insecta" family="Delphacidae" genus="Euides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">Euides</taxonomicName>
</emphasis>
and
<emphasis id="B963A52F6924FF86FDB1FCFA11D8170E" box="[555,721,799,825]" italics="true" pageId="2" pageNumber="373">
<taxonomicName id="4C1702BE6924FF86FDB1FCFA1187170F" authorityName="Matsumura" authorityYear="1935" box="[555,654,799,825]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">Toyoides</taxonomicName>
<taxonomicName id="4C1702BE6924FF86FD06FCFA11D8170E" box="[668,721,799,824]" class="Insecta" family="Delphacidae" genus="Toya" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">Toya</taxonomicName>
</emphasis>
) were provided. The protologue indicates that the genus
<taxonomicName id="4C1702BE6924FF86FACAFCFA1695170F" box="[1360,1436,799,825]" class="Insecta" family="Delphacidae" genus="Euides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FACAFCFA1695170F" box="[1360,1436,799,825]" italics="true" pageId="2" pageNumber="373">Euides</emphasis>
</taxonomicName>
can be distinguished by the presence of a frontal fork that is much longer and does not become narrow between the eyes; the middle keel in the pro-and mesonotum is not conspicuous. Morphological comparison of the
<typeStatus id="54ACC79F6924FF86FAF6FC82169517B7" box="[1388,1436,871,897]" pageId="2" pageNumber="373">type</typeStatus>
specimens for the two genera in this study revealed that, the male genitalia of the genus
<taxonomicName id="4C1702BE6924FF86FBF4FC6E17CD1793" authorityName="Matsumura" authorityYear="1935" box="[1134,1220,907,933]" class="Insecta" family="Delphacidae" genus="Epunka" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FBF4FC6E17CD1793" box="[1134,1220,907,933]" italics="true" pageId="2" pageNumber="373">Epunka</emphasis>
</taxonomicName>
could be described in detail for the first time.
<bibRefCitation id="EF8604CC6924FF86FE5BFC4A101817FF" author="Matsumura, S. &amp; Ishihara, T." box="[449,785,942,969]" pageId="2" pageNumber="373" pagination="59 - 76" refId="ref5610" refString="Matsumura, S. &amp; Ishihara, T. (1945) Species novae vel cognitae Araeopidarum Imperii Japonici (Hemiptera). Mushi, 16, 59 - 76." type="journal article" year="1945">Matsumura &amp; Ishihara (1945)</bibRefCitation>
only offered a brief illustration of the male anal segment, the habitus (forked portion on frons of median carina) and genital characters (hook-shaped aedeagus) are accorded with the genus
<taxonomicName id="4C1702BE6924FF86FEDBFC1212AD1027" authorityName="Matsumura" authorityYear="1935" box="[321,420,1015,1041]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FEDBFC1212AD1027" box="[321,420,1015,1041]" italics="true" pageId="2" pageNumber="373">Toyoides</emphasis>
</taxonomicName>
, here we treat the genus
<taxonomicName id="4C1702BE6924FF86FD21FC1210181027" authorityName="Matsumura" authorityYear="1935" box="[699,785,1015,1041]" class="Insecta" family="Delphacidae" genus="Epunka" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FD21FC1210181027" box="[699,785,1015,1041]" italics="true" pageId="2" pageNumber="373">Epunka</emphasis>
</taxonomicName>
as the junior synonym of the genus
<taxonomicName id="4C1702BE6924FF86FB3DFC1216031027" authorityName="Matsumura" authorityYear="1935" box="[1191,1290,1015,1041]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FB3DFC1216031027" box="[1191,1290,1015,1041]" italics="true" pageId="2" pageNumber="373">Toyoides</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66924FF86FF0DFB861743104A" box="[151,1098,1122,1149]" pageId="2" pageNumber="373" type="reference_group">
<paragraph id="8BA8793D6924FF86FF0DFB861743104A" blockId="2.[151,1098,1122,1149]" box="[151,1098,1122,1149]" pageId="2" pageNumber="373">
<emphasis id="B963A52F6924FF86FF0DFB861743104A" bold="true" box="[151,1098,1122,1149]" pageId="2" pageNumber="373">
Key to the genus
<taxonomicName id="4C1702BE6924FF86FEF4FB8712DE104B" ID-CoL="7Y6P" authorityName="Matsumura" authorityYear="1935" box="[366,471,1122,1149]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FEF4FB8712DE104B" bold="true" box="[366,471,1122,1149]" italics="true" pageId="2" pageNumber="373">Toyoides</emphasis>
</taxonomicName>
and its related groups (modified from
<bibRefCitation id="EF8604CC6924FF86FC20FB871748104A" author="Ding, J. H." box="[954,1089,1122,1148]" pageId="2" pageNumber="373" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding, 2006</bibRefCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66924FF86FF0DFB4416951138" pageId="2" pageNumber="373" type="key">
<key id="FF862E0C6924FF86FF0DFB4416951138" pageId="2" pageNumber="373">
<keyStep id="30E361986924FF86FF0DFB44169510E2" pageId="2" pageNumber="373">
<paragraph id="8BA8793D6924FF86FF0DFB4416951080" blockId="2.[151,1436,1185,1295]" box="[151,1436,1185,1207]" pageId="2" pageNumber="373">
<keyLead id="30E6DA086924FF86FF0DFB4416951080" box="[151,1436,1185,1207]" pageId="2" pageNumber="373">
1. Frons with median carina forked above level of upper margin of eyes (
<figureCitation id="132C65B86924FF86FC1DFB4410DC1080" box="[903,981,1185,1207]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[151,1436,477,1456]" captionTargetId="figure-420@3.[151,1436,502,1460]" captionTargetPageId="3" captionText="FIGURE 1. Lectotype of Toyoides albipennis Matsumura. A. habitus, dorsal view; B. ditto, lateral view; C. frons, ventral view; D. right hind leg with spur; E. lectotype labels. Abbreviations: f, front; b, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239599" httpUri="https://zenodo.org/record/14239599/files/figure.png" pageId="2" pageNumber="373">Figs 1C</figureCitation>
,
<figureCitation id="132C65B86924FF86FC45FB4410F51080" box="[991,1020,1185,1206]" captionStart="FIGURE 2" captionStartId="4.[151,250,1653,1677]" captionTargetBox="[151,1431,678,1626]" captionTargetId="figure-252@4.[151,1436,669,1635]" captionTargetPageId="4" captionText="FIGURE 2. Lectotype and paralectotype of Toyoides bilineata Matsumura comb. n. A. habitus, dorsal view; B. lectotype, habitus, lateral view; C. ditto, frons, ventral view; D. ditto, right hind leg with spur; E. lectotype and paralectotype labels. Abbreviations: f, front; b, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239601" httpUri="https://zenodo.org/record/14239601/files/figure.png" pageId="2" pageNumber="373">2C</figureCitation>
)................................ 2
</keyLead>
</paragraph>
<paragraph id="8BA8793D6924FF86FF0DFB5A169510E2" blockId="2.[151,1436,1185,1295]" box="[151,1436,1214,1236]" pageId="2" pageNumber="373">
<keyLead id="30E6DA086924FF86FF0DFB5A169510E2" box="[151,1436,1214,1236]" pageId="2" pageNumber="373">
- Frons with median carina forked below level of upper margin of eyes............................
<taxonomicName id="4C1702BE6924FF86FB44FB5B169510E2" ID-CoL="5777" authority="Matsumura" authorityName="Matsumura. Ding" authorityYear="2006" box="[1246,1436,1214,1236]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FB44FB5B162010E5" box="[1246,1321,1214,1235]" italics="true" pageId="2" pageNumber="373">Kakuna</emphasis>
Matsumura
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E361986924FF86FF0DFB3916951138" pageId="2" pageNumber="373">
<paragraph id="8BA8793D6924FF86FF0DFB39169510C4" blockId="2.[151,1436,1185,1295]" box="[151,1436,1244,1266]" pageId="2" pageNumber="373">
<keyLead id="30E6DA086924FF86FF0DFB39169510C4" box="[151,1436,1244,1266]" pageId="2" pageNumber="373">
2. Pygofer with medioventral process.............................................................
<taxonomicName id="4C1702BE6924FF86FA82FB39169510C4" ID-CoL="87FYD" authority="Fieber" authorityName="Fieber" box="[1304,1436,1244,1266]" class="Insecta" family="Delphacidae" genus="Euides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FA82FB39165010C7" box="[1304,1369,1244,1265]" italics="true" pageId="2" pageNumber="373">Euides</emphasis>
Fieber
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA8793D6924FF86FF0DFB1C16951138" blockId="2.[151,1436,1185,1295]" box="[151,1436,1273,1295]" pageId="2" pageNumber="373">
<keyLead id="30E6DA086924FF86FF0DFB1C16951138" box="[151,1436,1273,1295]" pageId="2" pageNumber="373">
- Pygofer without medioventral process (
<figureCitation id="132C65B86924FF86FDC6FB1C11CA1138" box="[604,707,1273,1295]" captionStart="FIGURE 4" captionStartId="6.[151,250,1530,1554]" captionTargetBox="[158,1429,188,1499]" captionTargetId="figure-20@6.[151,1436,181,1506]" captionTargetPageId="6" captionText="FIGURE 4. Male genitalia of Toyoides bilineata Matsumura comb. n. A. genitalia, caudal view; B. ditto, lateral view; C. anal segment, caudal view; D. ditto, lateral view; E. pygofer, caudal view; F. ditto, lateral view; G. genital styles, caudoventral view; H. ditto, lateral view; I. aedeagus with suspensorium, left-lateral view; J. aedeagus without suspensorium, dorsal view; K. suspensorium. Scale bar: AB, EF: 0.5 mm; CD, GH: 0.3 mm; IK: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239605" httpUri="https://zenodo.org/record/14239605/files/figure.png" pageId="2" pageNumber="373">Figs 4EF</figureCitation>
)..........................................
<taxonomicName id="4C1702BE6924FF86FB4EFB1C16951138" ID-CoL="7Y6P" authority="Matsumura" authorityName="Matsumura" authorityYear="1935" box="[1236,1436,1273,1294]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FB4EFB1C16211138" box="[1236,1320,1273,1294]" italics="true" pageId="2" pageNumber="373">Toyoides</emphasis>
Matsumura
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="C30D2AB66924FF86FF0DFAB11122130D" pageId="2" pageNumber="373" type="taxon_list">
<paragraph id="8BA8793D6924FF86FF0DFAB111BC1158" blockId="2.[151,693,1363,1390]" box="[151,693,1363,1390]" pageId="2" pageNumber="373">
<emphasis id="B963A52F6924FF86FF0DFAB111BC1158" bold="true" box="[151,693,1363,1390]" pageId="2" pageNumber="373">
Checklist of the genus
<taxonomicName id="4C1702BE6924FF86FE2BFAB611BC1158" ID-CoL="7Y6P" authority="Matsumura" authorityName="Matsumura" authorityYear="1935" box="[433,693,1363,1390]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6924FF86FE2BFAB61113115B" bold="true" box="[433,538,1363,1389]" italics="true" pageId="2" pageNumber="373">Toyoides</emphasis>
Matsumura
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA8793D6924FF86FF0DFA70117B1186" blockId="2.[151,1328,1429,1887]" box="[151,626,1429,1456]" pageId="2" pageNumber="373">
1.
<taxonomicName id="4C1702BE6924FF86FF5DFA70117B1186" authority="Matsumura, 1935" authorityName="Matsumura" authorityYear="1935" box="[199,626,1429,1456]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="albipennis">
<emphasis id="B963A52F6924FF86FF5DFA7012AF1199" box="[199,422,1429,1455]" italics="true" pageId="2" pageNumber="373">Toyoides albipennis</emphasis>
<bibRefCitation id="EF8604CC6924FF86FE36FA73117B1186" author="Matsumura, S." box="[428,626,1429,1456]" pageId="2" pageNumber="373" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA8793D6924FF86FF5DFA5C106311E5" blockId="2.[151,1328,1429,1887]" box="[199,874,1465,1492]" pageId="2" pageNumber="373">
<treatmentCitationGroup id="AB075E136924FF86FF5DFA5C106311E5" box="[199,874,1465,1492]" pageId="2" pageNumber="373">
<treatmentCitation id="0AB65F2C6924FF86FF5DFA5C11A911E5" author="Matsumura, S." box="[199,672,1465,1492]" page="78" pageId="2" pageNumber="373" year="1935">
<taxonomicName id="4C1702BE6924FF86FF5DFA5C11A911E5" ID-CoL="57N2L" authority="Matsumura, 1935: 78" authorityName="Matsumura" authorityPageNumber="78" authorityYear="1935" box="[199,672,1465,1492]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="albipennis">
<emphasis id="B963A52F6924FF86FF5DFA5C12AF11E5" box="[199,422,1465,1491]" italics="true" pageId="2" pageNumber="373">Toyoides albipennis</emphasis>
<bibRefCitation id="EF8604CC6924FF86FE36FA5F11A911E5" author="Matsumura, S." box="[428,672,1465,1492]" pageId="2" pageNumber="373" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935: 78</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="0AB65F2C6924FF86FD31FA5C106C11E5" author="Ding, J. H." box="[683,869,1465,1491]" page="304" pageId="2" pageNumber="373" year="2006">
<bibRefCitation id="EF8604CC6924FF86FD31FA5C106C11E5" author="Ding, J. H." box="[683,869,1465,1491]" pageId="2" pageNumber="373" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding, 2006: 304</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BA8793D6924FF86FF5DFA38117211C1" blockId="2.[151,1328,1429,1887]" box="[199,635,1501,1527]" pageId="2" pageNumber="373">
<treatmentCitationGroup id="AB075E136924FF86FF5DFA38117211C1" box="[199,635,1501,1527]" pageId="2" pageNumber="373">
<treatmentCitation id="0AB65F2C6924FF86FF5DFA38117E11C1" author="Ishihara, T." box="[199,631,1501,1527]" page="34" pageId="2" pageNumber="373" year="1949">
<taxonomicName id="4C1702BE6924FF86FF5DFA38117E11C1" authority="Ishihara, 1949: 34" authorityName="Ishihara" authorityPageNumber="34" authorityYear="1949" box="[199,631,1501,1527]" class="Insecta" family="Delphacidae" genus="Euidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="albipennis">
<emphasis id="B963A52F6924FF86FF5DFA3812AB11C1" box="[199,418,1501,1527]" italics="true" pageId="2" pageNumber="373">Euidella albipennis</emphasis>
<bibRefCitation id="EF8604CC6924FF86FE33FA38117E11C1" author="Ishihara, T." box="[425,631,1501,1527]" pageId="2" pageNumber="373" pagination="1 - 102" refId="ref5544" refString="Ishihara, T. (1949) Revision of the Araeopidae of Japan, Ryukyu Island and Formosa (Hemiptera). Scientific Reports of the Matsuyama Agricultural College, 2, 1 - 102." type="journal article" year="1949">Ishihara, 1949: 34</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA8793D6924FF86FF5DF9E4115B122D" blockId="2.[151,1328,1429,1887]" box="[199,594,1537,1564]" pageId="2" pageNumber="373">
<treatmentCitationGroup id="AB075E136924FF86FF5DF9E4115B122D" box="[199,594,1537,1564]" pageId="2" pageNumber="373">
<treatmentCitation id="0AB65F2C6924FF86FF5DF9E41147122D" author="Yang, C. T." box="[199,590,1537,1564]" page="62" pageId="2" pageNumber="373" year="1989">
<taxonomicName id="4C1702BE6924FF86FF5DF9E41147122D" authority="Yang, 1989: 62" authorityName="Yang" authorityPageNumber="62" authorityYear="1989" box="[199,590,1537,1564]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="albipennis">
<emphasis id="B963A52F6924FF86FF5DF9E41293122D" box="[199,410,1537,1563]" italics="true" pageId="2" pageNumber="373">Kakuna albipennis</emphasis>
<bibRefCitation id="EF8604CC6924FF86FE3AF9E71147122D" author="Yang, C. T." box="[416,590,1537,1564]" pageId="2" pageNumber="373" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang, 1989: 62</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA8793D6924FF86FF5DF9C016391209" blockId="2.[151,1328,1429,1887]" box="[199,1328,1573,1600]" pageId="2" pageNumber="373">
Distributions:
<collectingCountry id="F30039AD6924FF86FEF2F9C012A21209" box="[360,427,1573,1599]" name="China" pageId="2" pageNumber="373">China</collectingCountry>
(
<collectingRegion id="49D3B7DF6924FF86FE20F9C0110D1209" box="[442,516,1573,1599]" country="China" name="Anhui" pageId="2" pageNumber="373">Anhui</collectingRegion>
,
<collectingRegion id="49D3B7DF6924FF86FD95F9C011531209" box="[527,602,1573,1599]" country="China" name="Fujian" pageId="2" pageNumber="373">Fujian</collectingRegion>
,
<collectingRegion id="49D3B7DF6924FF86FDFFF9C011C11209" box="[613,712,1573,1599]" country="China" name="Guizhou" pageId="2" pageNumber="373">Guizhou</collectingRegion>
,
<collectingRegion id="49D3B7DF6924FF86FD49F9C310291276" box="[723,800,1574,1600]" country="China" name="Hunan" pageId="2" pageNumber="373">Hunan</collectingRegion>
,
<collectingRegion id="49D3B7DF6924FF86FCB6F9C0108A1209" box="[812,899,1573,1599]" country="China" name="Jiangsu" pageId="2" pageNumber="373">Jiangsu</collectingRegion>
,
<collectingRegion id="49D3B7DF6924FF86FC14F9C010E21209" box="[910,1003,1573,1599]" country="China" name="Sichuan" pageId="2" pageNumber="373">Sichuan</collectingRegion>
,
<collectingRegion id="49D3B7DF6924FF86FC6CF9C017571209" box="[1014,1118,1573,1599]" country="China" name="Zhejiang" pageId="2" pageNumber="373">Zhejiang</collectingRegion>
),
<collectingCountry id="F30039AD6924FF86FBEBF9C017CA1209" box="[1137,1219,1573,1599]" name="Taiwan" pageId="2" pageNumber="373">Taiwan</collectingCountry>
(
<collectingRegion id="49D3B7DF6924FF86FB4BF9C0162A1209" box="[1233,1315,1573,1599]" country="Taiwan" name="Tainan" pageId="2" pageNumber="373">Tainan</collectingRegion>
).
</paragraph>
<paragraph id="8BA8793D6924FF86FF0DF9AC11E81255" blockId="2.[151,1328,1429,1887]" box="[151,737,1609,1636]" pageId="2" pageNumber="373">
2.
<taxonomicName id="4C1702BE6924FF86FF5DF9AC117A1252" authority="(Matsumura, 1935)" authorityName="Park &amp; Jung" authorityYear="2024" baseAuthorityName="Matsumura" baseAuthorityYear="1935" box="[199,627,1609,1636]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="bilineata" status="comb. nov.">
<emphasis id="B963A52F6924FF86FF5DF9AC129C1255" box="[199,405,1609,1635]" italics="true" pageId="2" pageNumber="373">Toyoides bilineata</emphasis>
(
<bibRefCitation id="EF8604CC6924FF86FE39F9AC11621252" author="Matsumura, S." box="[419,619,1609,1636]" pageId="2" pageNumber="373" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935</bibRefCitation>
)
</taxonomicName>
<emphasis id="B963A52F6924FF86FDE0F9AC11E81255" bold="true" box="[634,737,1609,1635]" pageId="2" pageNumber="373">
<taxonomicNameLabel id="A25018546924FF86FDE0F9AC11E81255" box="[634,737,1609,1635]" pageId="2" pageNumber="373" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BA8793D6924FF86FF5DF988118F12BE" blockId="2.[151,1328,1429,1887]" box="[199,646,1645,1672]" pageId="2" pageNumber="373">
<treatmentCitationGroup id="AB075E136924FF86FF5DF988118F12BE" box="[199,646,1645,1672]" pageId="2" pageNumber="373">
<treatmentCitation id="0AB65F2C6924FF86FF5DF988118B12BE" author="Matsumura, S." box="[199,642,1645,1672]" page="77" pageId="2" pageNumber="373" year="1935">
<taxonomicName id="4C1702BE6924FF86FF5DF988118B12BE" ID-CoL="6FYTL" authority="Matsumura, 1935: 77" authorityName="Matsumura" authorityPageNumber="77" authorityYear="1935" box="[199,642,1645,1672]" class="Insecta" family="Delphacidae" genus="Epunka" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="bilineata">
<emphasis id="B963A52F6924FF86FF5DF988128112B1" box="[199,392,1645,1671]" italics="true" pageId="2" pageNumber="373">Epunka bilineata</emphasis>
<bibRefCitation id="EF8604CC6924FF86FE14F98B118B12BE" author="Matsumura, S." box="[398,642,1645,1672]" pageId="2" pageNumber="373" pagination="71 - 78" refId="ref5580" refString="Matsumura, S. (1935) Supplementary note to the revision of Stenocranus and its allied species in Japan-Empire. Insecta Matsumurana, 10, 71 - 78." type="journal article" year="1935">Matsumura, 1935: 77</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA8793D6924FF86FF5DF9741163129D" blockId="2.[151,1328,1429,1887]" box="[199,618,1681,1707]" pageId="2" pageNumber="373">
<treatmentCitationGroup id="AB075E136924FF86FF5DF9741163129D" box="[199,618,1681,1707]" pageId="2" pageNumber="373">
<treatmentCitation id="0AB65F2C6924FF86FF5DF974116F129D" author="Ishihara, T." box="[199,614,1681,1707]" page="34" pageId="2" pageNumber="373" year="1949">
<taxonomicName id="4C1702BE6924FF86FF5DF974116F129D" authority="Ishihara, 1949: 34" authorityName="Ishihara" authorityPageNumber="34" authorityYear="1949" box="[199,614,1681,1707]" class="Insecta" family="Delphacidae" genus="Euidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="bilineata">
<emphasis id="B963A52F6924FF86FF5DF9741298129D" box="[199,401,1681,1707]" italics="true" pageId="2" pageNumber="373">Euidella bilineata</emphasis>
<bibRefCitation id="EF8604CC6924FF86FE02F974116F129D" author="Ishihara, T." box="[408,614,1681,1707]" pageId="2" pageNumber="373" pagination="1 - 102" refId="ref5544" refString="Ishihara, T. (1949) Revision of the Araeopidae of Japan, Ryukyu Island and Formosa (Hemiptera). Scientific Reports of the Matsuyama Agricultural College, 2, 1 - 102." type="journal article" year="1949">Ishihara, 1949: 34</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA8793D6924FF86FF5DF95017F812F9" blockId="2.[151,1328,1429,1887]" box="[199,1265,1717,1744]" pageId="2" pageNumber="373">
Distributions:
<collectingCountry id="F30039AD6924FF86FEF2F95012FC12E6" box="[360,501,1717,1744]" name="South Korea" pageId="2" pageNumber="373">South Korea</collectingCountry>
(
<collectingRegion id="49D3B7DF6924FF86FD9EF950117F12F9" box="[516,630,1717,1743]" country="South Korea" name="Gyeonggi" pageId="2" pageNumber="373">Gyeonggi</collectingRegion>
)
<superScript id="7C62D4756924FF86FDE7F950118C12F5" attach="none" box="[637,645,1717,1731]" fontSize="6" pageId="2" pageNumber="373">*</superScript>
,
<collectingCountry id="F30039AD6924FF86FD09F95311DA12E6" box="[659,723,1718,1744]" name="Japan" pageId="2" pageNumber="373">Japan</collectingCountry>
(
<collectingRegion id="49D3B7DF6924FF86FD7BF950105D12F9" box="[737,852,1717,1743]" country="Japan" name="Hokkaido" pageId="2" pageNumber="373">Hokkaido</collectingRegion>
, Honshu, Kyushu, Tsushima Island).
</paragraph>
<paragraph id="8BA8793D6924FF86FF0DF93C12F612C5" blockId="2.[151,1328,1429,1887]" box="[151,511,1753,1779]" pageId="2" pageNumber="373">
3.
<taxonomicName id="4C1702BE6924FF86FF5DF93C12F612C5" authority="(Yang, 1989)" authorityName="Ding" authorityYear="2006" baseAuthorityName="Yang" baseAuthorityYear="1989" box="[199,511,1753,1779]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B963A52F6924FF86FF5DF93C126F12C5" box="[199,358,1753,1779]" italics="true" pageId="2" pageNumber="373">Toyoides mira</emphasis>
(
<bibRefCitation id="EF8604CC6924FF86FEEEF93C12FE12C5" author="Yang, C. T." box="[372,503,1753,1779]" pageId="2" pageNumber="373" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang, 1989</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BA8793D6924FF86FF5DF918111B1321" blockId="2.[151,1328,1429,1887]" box="[199,530,1789,1816]" pageId="2" pageNumber="373">
<treatmentCitationGroup id="AB075E136924FF86FF5DF918111B1321" box="[199,530,1789,1816]" pageId="2" pageNumber="373">
<treatmentCitation id="0AB65F2C6924FF86FF5DF91811071321" author="Yang, C. T." box="[199,526,1789,1816]" page="64" pageId="2" pageNumber="373" year="1989">
<taxonomicName id="4C1702BE6924FF86FF5DF91811071321" ID-CoL="6NJZ5" authority="Yang, 1989: 64" authorityName="Yang" authorityPageNumber="64" authorityYear="1989" box="[199,526,1789,1816]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B963A52F6924FF86FF5DF91812531321" box="[199,346,1789,1815]" italics="true" pageId="2" pageNumber="373">Kakuna mira</emphasis>
<bibRefCitation id="EF8604CC6924FF86FEFAF91B11071321" author="Yang, C. T." box="[352,526,1789,1816]" pageId="2" pageNumber="373" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang, 1989: 64</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA8793D6924FF86FF5DF8C41122130D" blockId="2.[151,1328,1429,1887]" box="[199,555,1825,1851]" pageId="2" pageNumber="373">
<treatmentCitationGroup id="AB075E136924FF86FF5DF8C41122130D" box="[199,555,1825,1851]" pageId="2" pageNumber="373">
<treatmentCitation id="0AB65F2C6924FF86FF5DF8C4112F130D" author="Ding, J. H." box="[199,550,1825,1851]" page="306" pageId="2" pageNumber="373" year="2006">
<taxonomicName id="4C1702BE6924FF86FF5DF8C4112F130D" ID-CoL="57N2M" authority="Ding, 2006: 306" authorityName="Ding" authorityPageNumber="306" authorityYear="2006" baseAuthorityName="Yang" baseAuthorityYear="1989" box="[199,550,1825,1851]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B963A52F6924FF86FF5DF8C4126F130D" box="[199,358,1825,1851]" italics="true" pageId="2" pageNumber="373">Toyoides mira</emphasis>
<bibRefCitation id="EF8604CC6924FF86FEF7F8C4112F130D" author="Ding, J. H." box="[365,550,1825,1851]" pageId="2" pageNumber="373" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding, 2006: 306</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66924FF86FF5DF8A011401369" box="[199,585,1861,1887]" pageId="2" pageNumber="373" type="distribution">
<paragraph id="8BA8793D6924FF86FF5DF8A011401369" blockId="2.[151,1328,1429,1887]" box="[199,585,1861,1887]" pageId="2" pageNumber="373">
Distribution:
<collectingCountry id="F30039AD6924FF86FEC7F8A012A61369" box="[349,431,1861,1887]" name="Taiwan" pageId="2" pageNumber="373">Taiwan</collectingCountry>
(
<collectingRegion id="49D3B7DF6924FF86FE24F8A011371369" box="[446,574,1861,1887]" country="Taiwan" name="Kaohsiung" pageId="2" pageNumber="373">Kaohsiung</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66925FF87FF0DFF7210C21487" box="[151,971,151,178]" pageId="3" pageNumber="374" type="multiple">
<paragraph id="8BA8793D6925FF87FF0DFF7210C21487" blockId="3.[151,971,151,178]" box="[151,971,151,178]" pageId="3" pageNumber="374">
<heading id="D0E0CE516925FF87FF0DFF7210C21487" bold="true" box="[151,971,151,178]" fontSize="11" level="1" pageId="3" pageNumber="374" reason="1">
<emphasis id="B963A52F6925FF87FF0DFF7210C21487" bold="true" box="[151,971,151,178]" pageId="3" pageNumber="374">
Key to species of
<taxonomicName id="4C1702BE6925FF87FEF4FF7212DE1484" ID-CoL="7Y6P" authorityName="Park &amp; Jung" authorityYear="2024" baseAuthorityName="Park &amp; Jung" baseAuthorityYear="2024" box="[366,471,151,178]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F6925FF87FEF4FF7212DE1484" bold="true" box="[366,471,151,178]" italics="true" pageId="3" pageNumber="374">Toyoides</emphasis>
</taxonomicName>
(modified from
<bibRefCitation id="EF8604CC6925FF87FD38FF7210271487" author="Yang, C. T." box="[674,814,151,177]" pageId="3" pageNumber="374" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang, 1989</bibRefCitation>
;
<bibRefCitation id="EF8604CC6925FF87FCA6FF7210CA1487" author="Ding, J. H." box="[828,963,151,177]" pageId="3" pageNumber="374" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding, 2006</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB66925FF87FF0DFF2B1695150D" pageId="3" pageNumber="374" type="key">
<keyStep id="30E361986925FF87FF0DFF2B16951536" pageId="3" pageNumber="374">
<paragraph id="8BA8793D6925FF87FF0DFF2B169514D5" blockId="3.[151,1436,205,315]" box="[151,1436,205,227]" pageId="3" pageNumber="374">
<keyLead id="30E6DA086925FF87FF0DFF2B169514D5" box="[151,1436,205,227]" pageId="3" pageNumber="374">
1. Frons with median carina forked above level of lower margin of eyes (
<figureCitation id="132C65B86925FF87FC1DFF2810DC14D5" box="[903,981,205,227]" captionStart="FIGURE 2" captionStartId="4.[151,250,1653,1677]" captionTargetBox="[151,1431,678,1626]" captionTargetId="figure-252@4.[151,1436,669,1635]" captionTargetPageId="4" captionText="FIGURE 2. Lectotype and paralectotype of Toyoides bilineata Matsumura comb. n. A. habitus, dorsal view; B. lectotype, habitus, lateral view; C. ditto, frons, ventral view; D. ditto, right hind leg with spur; E. lectotype and paralectotype labels. Abbreviations: f, front; b, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239601" httpUri="https://zenodo.org/record/14239601/files/figure.png" pageId="3" pageNumber="374">Figs 2C</figureCitation>
,
<figureCitation id="132C65B86925FF87FC45FF2B10F414D5" box="[991,1021,206,227]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="3" pageNumber="374">3D</figureCitation>
)................................ 2
</keyLead>
</paragraph>
<paragraph id="8BA8793D6925FF87FF0DFF0E16951536" blockId="3.[151,1436,205,315]" box="[151,1436,235,257]" pageId="3" pageNumber="374">
<keyLead id="30E6DA086925FF87FF0DFF0E16951536" box="[151,1436,235,257]" pageId="3" pageNumber="374">
- Frons with median carina forked below level of lower margin of eyes (
<figureCitation id="132C65B86925FF87FC12FF0E10D81536" box="[904,977,235,257]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[151,1436,477,1456]" captionTargetId="figure-420@3.[151,1436,502,1460]" captionTargetPageId="3" captionText="FIGURE 1. Lectotype of Toyoides albipennis Matsumura. A. habitus, dorsal view; B. ditto, lateral view; C. frons, ventral view; D. right hind leg with spur; E. lectotype labels. Abbreviations: f, front; b, back. Scale bar: AB: 1.0 mm; CD: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239599" httpUri="https://zenodo.org/record/14239599/files/figure.png" pageId="3" pageNumber="374">Fig. 1C</figureCitation>
).................
<taxonomicName id="4C1702BE6925FF87FB36FF0E16951536" authority="Matsumura" authorityName="Matsumura" box="[1196,1436,235,256]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="albipennis">
<emphasis id="B963A52F6925FF87FB36FF0E17B01536" box="[1196,1209,235,256]" italics="true" pageId="3" pageNumber="374">T</emphasis>
.
<emphasis id="B963A52F6925FF87FB5FFF0E16201536" box="[1221,1321,235,256]" italics="true" pageId="3" pageNumber="374">albipennis</emphasis>
Matsumura
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E361986925FF87FF0DFEED1695150D" pageId="3" pageNumber="374">
<paragraph id="8BA8793D6925FF87FF0DFEED1695152B" blockId="3.[151,1436,205,315]" box="[151,1436,264,286]" pageId="3" pageNumber="374">
<keyLead id="30E6DA086925FF87FF0DFEED1695152B" box="[151,1436,264,286]" pageId="3" pageNumber="374">
2. Tegmina distinctly curved (
<figureCitation id="132C65B86925FF87FE73FEED113F152B" box="[489,566,264,286]" captionStart="FIGURE 3" captionStartId="5.[151,250,1356,1380]" captionTargetBox="[156,1429,188,1331]" captionTargetId="figure-17@5.[151,1436,181,1332]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of Toyoides bilineata Matsumura comb. n. A. female habitus, dorsal view; B. male habitus, dorsal view; C. ditto, lateral view; D. left tegmina, in alcohol; E. frons, ventral view; F. head and thorax, dorsal view; G. ditto, lateral view. Scale bar: AD: 1.0 mm; EG: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239603" httpUri="https://zenodo.org/record/14239603/files/figure.png" pageId="3" pageNumber="374">Fig. 3D</figureCitation>
); division of aedeagus with spinule (
<figureCitation id="132C65B86925FF87FC0DFEED10D1152B" box="[919,984,264,286]" captionStart="FIGURE 4" captionStartId="6.[151,250,1530,1554]" captionTargetBox="[158,1429,188,1499]" captionTargetId="figure-20@6.[151,1436,181,1506]" captionTargetPageId="6" captionText="FIGURE 4. Male genitalia of Toyoides bilineata Matsumura comb. n. A. genitalia, caudal view; B. ditto, lateral view; C. anal segment, caudal view; D. ditto, lateral view; E. pygofer, caudal view; F. ditto, lateral view; G. genital styles, caudoventral view; H. ditto, lateral view; I. aedeagus with suspensorium, left-lateral view; J. aedeagus without suspensorium, dorsal view; K. suspensorium. Scale bar: AB, EF: 0.5 mm; CD, GH: 0.3 mm; IK: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239605" httpUri="https://zenodo.org/record/14239605/files/figure.png" pageId="3" pageNumber="374">Fig. 4I</figureCitation>
)........
<taxonomicName id="4C1702BE6925FF87FBD7FEEC1637152B" authority="(Matsumura)" authorityName="Park &amp; Jung" authorityYear="2024" baseAuthorityName="Matsumura" box="[1101,1342,264,286]" class="Insecta" family="Delphacidae" genus="Euidella" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="bilineata" status="comb. n.">
<emphasis id="B963A52F6925FF87FBD7FEEC17531528" box="[1101,1114,265,286]" italics="true" pageId="3" pageNumber="374">T</emphasis>
.
<emphasis id="B963A52F6925FF87FBFCFEED17B2152B" box="[1126,1211,264,285]" italics="true" pageId="3" pageNumber="374">bilineata</emphasis>
(Matsumura)
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<emphasis id="B963A52F6925FF87FADFFEED1695152B" bold="true" box="[1349,1436,264,285]" pageId="3" pageNumber="374">
<taxonomicNameLabel id="A25018546925FF87FADFFEED1695152B" box="[1349,1436,264,285]" pageId="3" pageNumber="374" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BA8793D6925FF87FF0DFEC31695150D" blockId="3.[151,1436,205,315]" box="[151,1436,293,315]" pageId="3" pageNumber="374">
<keyLead id="30E6DA086925FF87FF0DFEC31695150D" box="[151,1436,293,315]" pageId="3" pageNumber="374">
- Tegmina almost straight; division of aedeagus without spinule......................................
<taxonomicName id="4C1702BE6925FF87FA94FEC31695150D" ID-CoL="57N2M" authority="(Yang)" authorityName="Ding" authorityYear="2006" baseAuthorityName="Yang" baseAuthorityYear="1989" box="[1294,1436,294,315]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B963A52F6925FF87FA94FEC31612150D" box="[1294,1307,294,315]" italics="true" pageId="3" pageNumber="374">T</emphasis>
.
<emphasis id="B963A52F6925FF87FABDFEC3165D150D" box="[1319,1364,294,315]" italics="true" pageId="3" pageNumber="374">mira</emphasis>
(Yang)
</taxonomicName>
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</paragraph>
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</treatment>
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View file

@ -0,0 +1,152 @@
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<subSubSection id="C30D2AB6692EFF8CFF0DFF7212ED1487" box="[151,484,151,178]" pageId="8" pageNumber="379" type="nomenclature">
<paragraph id="8BA8793D692EFF8CFF0DFF7212ED1487" blockId="8.[151,484,151,178]" box="[151,484,151,178]" pageId="8" pageNumber="379">
<taxonomicName id="4C1702BE692EFF8CFF0DFF7212ED1487" ID-CoL="57N2M" authority="(Yang, 1989)" authorityName="Ding" authorityYear="2006" baseAuthorityName="Yang" baseAuthorityYear="1989" box="[151,484,151,178]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B963A52F692EFF8CFF0DFF7212ED1487" bold="true" box="[151,484,151,178]" pageId="8" pageNumber="379">
<emphasis id="B963A52F692EFF8CFF0DFF7212491487" bold="true" box="[151,320,151,178]" italics="true" pageId="8" pageNumber="379">Toyoides mira</emphasis>
(
<bibRefCitation id="EF8604CC692EFF8CFECAFF7212D51487" author="Yang, C. T." box="[336,476,151,177]" pageId="8" pageNumber="379" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang, 1989</bibRefCitation>
)
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB6692EFF8CFF0DFF3912D01524" pageId="8" pageNumber="379" type="taxon_list">
<paragraph id="8BA8793D692EFF8CFF0DFF3912CA14C5" blockId="8.[151,473,219,274]" box="[151,451,219,244]" pageId="8" pageNumber="379">
<treatmentCitationGroup id="AB075E13692EFF8CFF0DFF3912CA14C5" box="[151,451,219,244]" pageId="8" pageNumber="379">
<treatmentCitation id="0AB65F2C692EFF8CFF0DFF3912B614C5" author="Yang, C. T." box="[151,447,219,244]" page="64" pageId="8" pageNumber="379" year="1989">
<taxonomicName id="4C1702BE692EFF8CFF0DFF3912B614C5" ID-CoL="6NJZ5" authority="Yang, 1989: 64" authorityName="Yang" authorityPageNumber="64" authorityYear="1989" box="[151,447,219,244]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B963A52F692EFF8CFF0DFF39121514C5" box="[151,284,220,244]" italics="true" pageId="8" pageNumber="379">Kakuna mira</emphasis>
<bibRefCitation id="EF8604CC692EFF8CFEB8FF3912B614C5" author="Yang, C. T." box="[290,447,219,244]" pageId="8" pageNumber="379" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang, 1989: 64</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="8BA8793D692EFF8CFF0DFF1F12D01524" blockId="8.[151,473,219,274]" box="[151,473,250,274]" pageId="8" pageNumber="379">
<treatmentCitationGroup id="AB075E13692EFF8CFF0DFF1F12D01524" box="[151,473,250,274]" pageId="8" pageNumber="379">
<treatmentCitation id="0AB65F2C692EFF8CFF0DFF1F12DC1524" author="Ding, J. H." box="[151,469,250,274]" page="306" pageId="8" pageNumber="379" year="2006">
<taxonomicName id="4C1702BE692EFF8CFF0DFF1F12DC1524" ID-CoL="57N2M" authority="Ding, 2006: 306" authorityName="Ding" authorityPageNumber="306" authorityYear="2006" baseAuthorityName="Yang" baseAuthorityYear="1989" box="[151,469,250,274]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B963A52F692EFF8CFF0DFF1F122E1524" box="[151,295,250,274]" italics="true" pageId="8" pageNumber="379">Toyoides mira</emphasis>
<bibRefCitation id="EF8604CC692EFF8CFEB7FF1F12DC1524" author="Ding, J. H." box="[301,469,250,274]" pageId="8" pageNumber="379" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding, 2006: 306</bibRefCitation>
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.
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<paragraph id="8BA8793D692EFF8CFF0DFEA512BC15F0" blockId="8.[151,1437,320,671]" pageId="8" pageNumber="379">
<emphasis id="B963A52F692EFF8CFF0DFEA5121B156C" bold="true" box="[151,274,320,346]" pageId="8" pageNumber="379">Diagnosis.</emphasis>
Recognized by body yellowish white; tegmina with anterior and posterior margins black; vertex without any markings; tegmina almost straight; frons with median carinae forked above level of lower margin of eyes; pygofer with medioventral bump rounded; aedeagus branched at medially, division without spinule (modified from
<bibRefCitation id="EF8604CC692EFF8CFF0DFE48121315F1" author="Yang, C. T." box="[151,282,429,455]" pageId="8" pageNumber="379" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang, 1989</bibRefCitation>
;
<bibRefCitation id="EF8604CC692EFF8CFEBCFE4912A015F0" author="Ding, J. H." box="[294,425,428,454]" pageId="8" pageNumber="379" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding, 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB6692EFF8CFF5DFE35166315DC" box="[199,1386,464,491]" pageId="8" pageNumber="379" type="description">
<paragraph id="8BA8793D692EFF8CFF5DFE35166315DC" blockId="8.[151,1437,320,671]" box="[199,1386,464,491]" pageId="8" pageNumber="379">
<emphasis id="B963A52F692EFF8CFF5DFE35125015DC" bold="true" box="[199,345,464,490]" pageId="8" pageNumber="379">Description.</emphasis>
See
<bibRefCitation id="EF8604CC692EFF8CFE14FE34111015DD" author="Yang, C. T." box="[398,537,465,491]" pageId="8" pageNumber="379" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang (1989)</bibRefCitation>
for original description, and
<bibRefCitation id="EF8604CC692EFF8CFCFAFE3510E315DC" author="Ding, J. H." box="[864,1002,464,490]" pageId="8" pageNumber="379" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding (2006)</bibRefCitation>
for redescription and figures also.
</paragraph>
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<paragraph id="8BA8793D692EFF8CFF5DFE1112F51639" blockId="8.[151,1437,320,671]" box="[199,508,500,527]" pageId="8" pageNumber="379">
<emphasis id="B963A52F692EFF8CFF5DFE1112B81638" bold="true" box="[199,433,500,526]" pageId="8" pageNumber="379">Material examined.</emphasis>
None.
</paragraph>
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<subSubSection id="C30D2AB6692EFF8CFF5DFDFD11A71605" box="[199,686,536,563]" pageId="8" pageNumber="379" type="distribution">
<paragraph id="8BA8793D692EFF8CFF5DFDFD11A71605" blockId="8.[151,1437,320,671]" box="[199,686,536,563]" pageId="8" pageNumber="379">
<emphasis id="B963A52F692EFF8CFF5DFDFD12681604" bold="true" box="[199,353,536,562]" pageId="8" pageNumber="379">Distribution.</emphasis>
<collectingCountry id="F30039AD692EFF8CFEFDFDFD12B01604" box="[359,441,536,562]" name="Taiwan" pageId="8" pageNumber="379">Taiwan</collectingCountry>
(
<collectingRegion id="49D3B7DF692EFF8CFE52FDFD114C1604" box="[456,581,536,562]" country="Taiwan" name="Kaohsiung" pageId="8" pageNumber="379">Kaohsiung</collectingRegion>
) (
<figureCitation id="132C65B8692EFF8CFDC1FDFD11A91605" box="[603,672,536,563]" captionStart="FIGURE 5" captionStartId="7.[151,250,1946,1970]" captionTargetBox="[151,1435,1281,1923]" captionTargetId="figure-410@7.[151,1435,1281,1923]" captionTargetPageId="7" captionText="FIGURE 5. Distribution map of the genus Toyoides species from the Korean Peninsula and neighboring countries." figureDoi="http://doi.org/10.5281/zenodo.14239607" httpUri="https://zenodo.org/record/14239607/files/figure.png" pageId="8" pageNumber="379">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB6692EFF8CFF5DFDD812C01660" box="[199,457,572,599]" pageId="8" pageNumber="379" type="biology_ecology">
<paragraph id="8BA8793D692EFF8CFF5DFDD812C01660" blockId="8.[151,1437,320,671]" box="[199,457,572,599]" pageId="8" pageNumber="379">
<emphasis id="B963A52F692EFF8CFF5DFDD812421661" bold="true" box="[199,331,573,599]" pageId="8" pageNumber="379">Host plant.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30D2AB6692EFF8CFF5DFD84111F16A9" pageId="8" pageNumber="379" type="discussion">
<paragraph id="8BA8793D692EFF8CFF5DFD84111F16A9" blockId="8.[151,1437,320,671]" pageId="8" pageNumber="379">
<emphasis id="B963A52F692EFF8CFF5DFD841232164D" bold="true" box="[199,315,609,635]" pageId="8" pageNumber="379">Remarks.</emphasis>
<bibRefCitation id="EF8604CC692EFF8CFEDBFD8412C4164D" author="Yang, C. T." box="[321,461,609,635]" pageId="8" pageNumber="379" refId="ref5902" refString="Yang, C. T. (1989) Delphacidae of Taiwan (II) (Homoptera: Fulgoroidea). National Science Council Special Publication No 6. National Science Council, Taipei, 334 pp." type="book" year="1989">Yang (1989)</bibRefCitation>
described this species in the genus
<taxonomicName id="4C1702BE692EFF8CFCC5FD851749164D" authority="Matsumura" authorityName="Matsumura. Ding" authorityYear="2006" box="[863,1088,608,635]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F692EFF8CFCC5FD8510BF164C" box="[863,950,608,634]" italics="true" pageId="8" pageNumber="379">Kakuna</emphasis>
Matsumura
</taxonomicName>
, while
<bibRefCitation id="EF8604CC692EFF8CFB0BFD851615164C" author="Ding, J. H." box="[1169,1308,608,634]" pageId="8" pageNumber="379" refId="ref5446" refString="Ding, J. H. (2006) Fauna Sinica. Insecta. Vol. 45. Homoptera Delphacidae. Fauna Sinica. Science Press, Beijing, China, 776 pp." type="book" year="2006">Ding (2006)</bibRefCitation>
transferred this species to the genus
<taxonomicName id="4C1702BE692EFF8CFE37FD61111916A8" authorityName="Park &amp; Jung" authorityYear="2024" baseAuthorityName="Park &amp; Jung" baseAuthorityYear="2024" box="[429,528,644,670]" class="Insecta" family="Delphacidae" genus="Toyoides" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="genus">
<emphasis id="B963A52F692EFF8CFE37FD61111916A8" box="[429,528,644,670]" italics="true" pageId="8" pageNumber="379">Toyoides</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>