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+0000-0003-4102-9371 +ZTM, Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. +haesa@leibniz-lib.de + + + +Author + +Utevsky, Serge +0000-0003-1290-6742 +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. +serge.utevsky@karazin.ua + + + +Author + +Kristensen, Reinhardt Møbjerg +0000-0001-9549-1188 +Natural History Museum of Denmark, University of Copenhagen, Gothersgade 130, 1123 Copenhagen, Denmark. +rmkristensen@snm.ku.dk + + + +Author + +Kieneke, Alexander +0000-0001-8180-2483 +German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. +alexander.kieneke@senckenberg.de + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +1 +39 + + + + +https://doi.org/10.11646/zootaxa.5543.1.1 + +journal article +10.11646/zootaxa.5543.1.1 +1175-5326 +14385085 +F6372401-5C79-487C-A8C1-DBDEE7C71671 + + + + + + + +Angursa abyssalis +Renaud-Mornant, 1981 + + + + + +N= + +238 specimens +occurrence: ANDEEP I, +II +, +III +, +ANDEEP-SYSTCO +; + +1,088–5,213 m +bsl + + +. + + + + +Fig. 8A–D +; Supplementary +Table 1.5 + + + + \ No newline at end of file diff --git a/data/03/84/87/03848797FF88FFE79B8DE782AD2C1120.xml b/data/03/84/87/03848797FF88FFE79B8DE782AD2C1120.xml new file mode 100644 index 00000000000..16209c14705 --- /dev/null +++ b/data/03/84/87/03848797FF88FFE79B8DE782AD2C1120.xml @@ -0,0 +1,98 @@ + + + +Towards a better understanding of deep-sea tardigrade biogeography: numerous new records from the Southern Ocean + + + +Author + +Trokhymchuk, Roman +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. & German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. + + + +Author + +Schmidt-Rhaesa, Andreas +0000-0003-4102-9371 +ZTM, Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. +haesa@leibniz-lib.de + + + +Author + +Utevsky, Serge +0000-0003-1290-6742 +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. +serge.utevsky@karazin.ua + + + +Author + +Kristensen, Reinhardt Møbjerg +0000-0001-9549-1188 +Natural History Museum of Denmark, University of Copenhagen, Gothersgade 130, 1123 Copenhagen, Denmark. +rmkristensen@snm.ku.dk + + + +Author + +Kieneke, Alexander +0000-0001-8180-2483 +German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. +alexander.kieneke@senckenberg.de + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +1 +39 + + + + +https://doi.org/10.11646/zootaxa.5543.1.1 + +journal article +10.11646/zootaxa.5543.1.1 +1175-5326 +14385085 +F6372401-5C79-487C-A8C1-DBDEE7C71671 + + + + + + +Genus: + +Angursa +Pollock, 1979 + + + + + + +N=1,172 specimens + + + \ No newline at end of file diff --git a/data/03/84/87/03848797FF8AFFE59B8DE081AE3E10E0.xml b/data/03/84/87/03848797FF8AFFE59B8DE081AE3E10E0.xml new file mode 100644 index 00000000000..bccfc363f12 --- /dev/null +++ b/data/03/84/87/03848797FF8AFFE59B8DE081AE3E10E0.xml @@ -0,0 +1,108 @@ + + + +Towards a better understanding of deep-sea tardigrade biogeography: numerous new records from the Southern Ocean + + + +Author + +Trokhymchuk, Roman +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. & German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. + + + +Author + +Schmidt-Rhaesa, Andreas +0000-0003-4102-9371 +ZTM, Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. +haesa@leibniz-lib.de + + + +Author + +Utevsky, Serge +0000-0003-1290-6742 +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. +serge.utevsky@karazin.ua + + + +Author + +Kristensen, Reinhardt Møbjerg +0000-0001-9549-1188 +Natural History Museum of Denmark, University of Copenhagen, Gothersgade 130, 1123 Copenhagen, Denmark. +rmkristensen@snm.ku.dk + + + +Author + +Kieneke, Alexander +0000-0001-8180-2483 +German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. +alexander.kieneke@senckenberg.de + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +1 +39 + + + + +https://doi.org/10.11646/zootaxa.5543.1.1 + +journal article +10.11646/zootaxa.5543.1.1 +1175-5326 +14385085 +F6372401-5C79-487C-A8C1-DBDEE7C71671 + + + + + + + +Angursa antarctica +Víllora-Moreno, 1998 + + + + + +N= +62 specimens +occurrence: ANDEEP I; +2,893 m +bsl. + + + + +Fig. 9A–B +; Supplementary +Table 1.7 + + + + \ No newline at end of file diff --git a/data/03/84/87/03848797FF8AFFE59B8DE3E6AE3F1549.xml b/data/03/84/87/03848797FF8AFFE59B8DE3E6AE3F1549.xml new file mode 100644 index 00000000000..577e7f4f324 --- /dev/null +++ b/data/03/84/87/03848797FF8AFFE59B8DE3E6AE3F1549.xml @@ -0,0 +1,119 @@ + + + +Towards a better understanding of deep-sea tardigrade biogeography: numerous new records from the Southern Ocean + + + +Author + +Trokhymchuk, Roman +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. & German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. + + + +Author + +Schmidt-Rhaesa, Andreas +0000-0003-4102-9371 +ZTM, Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. +haesa@leibniz-lib.de + + + +Author + +Utevsky, Serge +0000-0003-1290-6742 +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. +serge.utevsky@karazin.ua + + + +Author + +Kristensen, Reinhardt Møbjerg +0000-0001-9549-1188 +Natural History Museum of Denmark, University of Copenhagen, Gothersgade 130, 1123 Copenhagen, Denmark. +rmkristensen@snm.ku.dk + + + +Author + +Kieneke, Alexander +0000-0001-8180-2483 +German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. +alexander.kieneke@senckenberg.de + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +1 +39 + + + + +https://doi.org/10.11646/zootaxa.5543.1.1 + +journal article +10.11646/zootaxa.5543.1.1 +1175-5326 +14385085 +F6372401-5C79-487C-A8C1-DBDEE7C71671 + + + + + + + +Angursa capsula +Bussau, 1992 + + + + + +N= + +680 specimens +occurrence: ANDEEP I, +II +, +III +, +ANDEEP-SYSTCO +; + +1,088–5,213 m +bsl + + +. + + + + +Fig. 8 E–F +; Supplementary +Table 1.6 + + + + \ No newline at end of file diff --git a/data/03/84/87/03848797FF8AFFE59B8DE54BAE1312AA.xml b/data/03/84/87/03848797FF8AFFE59B8DE54BAE1312AA.xml new file mode 100644 index 00000000000..8a753968c16 --- /dev/null +++ b/data/03/84/87/03848797FF8AFFE59B8DE54BAE1312AA.xml @@ -0,0 +1,117 @@ + + + +Towards a better understanding of deep-sea tardigrade biogeography: numerous new records from the Southern Ocean + + + +Author + +Trokhymchuk, Roman +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. & German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. + + + +Author + +Schmidt-Rhaesa, Andreas +0000-0003-4102-9371 +ZTM, Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. +haesa@leibniz-lib.de + + + +Author + +Utevsky, Serge +0000-0003-1290-6742 +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. +serge.utevsky@karazin.ua + + + +Author + +Kristensen, Reinhardt Møbjerg +0000-0001-9549-1188 +Natural History Museum of Denmark, University of Copenhagen, Gothersgade 130, 1123 Copenhagen, Denmark. +rmkristensen@snm.ku.dk + + + +Author + +Kieneke, Alexander +0000-0001-8180-2483 +German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. +alexander.kieneke@senckenberg.de + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +1 +39 + + + + +https://doi.org/10.11646/zootaxa.5543.1.1 + +journal article +10.11646/zootaxa.5543.1.1 +1175-5326 +14385085 +F6372401-5C79-487C-A8C1-DBDEE7C71671 + + + + + + + +Angursa lanceolata +Renaud-Mornant, 1981 + + + + + +N= + +36 specimens +occurrence: ANDEEP I, +III +, +ANDEEP-SYSTCO +; + +1,927–5,213 m +bsl + + +. + + + + +Fig. 9 C +; Supplementary +Table 1.8 + + + + \ No newline at end of file diff --git a/data/03/84/87/03848797FF8CFFE39B8DE06EAED817BC.xml b/data/03/84/87/03848797FF8CFFE39B8DE06EAED817BC.xml new file mode 100644 index 00000000000..2f96cc6e978 --- /dev/null +++ b/data/03/84/87/03848797FF8CFFE39B8DE06EAED817BC.xml @@ -0,0 +1,117 @@ + + + +Towards a better understanding of deep-sea tardigrade biogeography: numerous new records from the Southern Ocean + + + +Author + +Trokhymchuk, Roman +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. & German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. + + + +Author + +Schmidt-Rhaesa, Andreas +0000-0003-4102-9371 +ZTM, Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. +haesa@leibniz-lib.de + + + +Author + +Utevsky, Serge +0000-0003-1290-6742 +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. +serge.utevsky@karazin.ua + + + +Author + +Kristensen, Reinhardt Møbjerg +0000-0001-9549-1188 +Natural History Museum of Denmark, University of Copenhagen, Gothersgade 130, 1123 Copenhagen, Denmark. +rmkristensen@snm.ku.dk + + + +Author + +Kieneke, Alexander +0000-0001-8180-2483 +German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. +alexander.kieneke@senckenberg.de + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +1 +39 + + + + +https://doi.org/10.11646/zootaxa.5543.1.1 + +journal article +10.11646/zootaxa.5543.1.1 +1175-5326 +14385085 +F6372401-5C79-487C-A8C1-DBDEE7C71671 + + + + + + + +Styraconyx qivitoq +Kristensen & Higgins, 1984 + + + + + +N= +13 specimens +: + +13 males +occurrence: +ANDEEP-SYSTCO +; + +1,927–1,960 m +bsl + + +. + + + + +Figs 10A–E +; Supplementary +Table 1.10 + + + + \ No newline at end of file diff --git a/data/03/84/87/03848797FF8CFFE39B8DE3E6AE3C1534.xml b/data/03/84/87/03848797FF8CFFE39B8DE3E6AE3C1534.xml new file mode 100644 index 00000000000..70d91dc013c --- /dev/null +++ b/data/03/84/87/03848797FF8CFFE39B8DE3E6AE3C1534.xml @@ -0,0 +1,117 @@ + + + +Towards a better understanding of deep-sea tardigrade biogeography: numerous new records from the Southern Ocean + + + +Author + +Trokhymchuk, Roman +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. & German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. + + + +Author + +Schmidt-Rhaesa, Andreas +0000-0003-4102-9371 +ZTM, Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. +haesa@leibniz-lib.de + + + +Author + +Utevsky, Serge +0000-0003-1290-6742 +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. +serge.utevsky@karazin.ua + + + +Author + +Kristensen, Reinhardt Møbjerg +0000-0001-9549-1188 +Natural History Museum of Denmark, University of Copenhagen, Gothersgade 130, 1123 Copenhagen, Denmark. +rmkristensen@snm.ku.dk + + + +Author + +Kieneke, Alexander +0000-0001-8180-2483 +German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. +alexander.kieneke@senckenberg.de + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +1 +39 + + + + +https://doi.org/10.11646/zootaxa.5543.1.1 + +journal article +10.11646/zootaxa.5543.1.1 +1175-5326 +14385085 +F6372401-5C79-487C-A8C1-DBDEE7C71671 + + + + + + + +Angursa lingua +Bussau, 1992 + + + + + +N= + +65 specimens +occurrence: ANDEEP I, +II +, +ANDEEP-SYSTCO +; + +1,088–3,555 m +bsl + + +. + + + + +Figs 9D, E +; Supplementary +Table 1.9 + + + + \ No newline at end of file diff --git a/data/03/84/87/03848797FF8CFFE39B8DE6F2AE2F1258.xml b/data/03/84/87/03848797FF8CFFE39B8DE6F2AE2F1258.xml new file mode 100644 index 00000000000..0dbf13db458 --- /dev/null +++ b/data/03/84/87/03848797FF8CFFE39B8DE6F2AE2F1258.xml @@ -0,0 +1,115 @@ + + + +Towards a better understanding of deep-sea tardigrade biogeography: numerous new records from the Southern Ocean + + + +Author + +Trokhymchuk, Roman +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. & German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. + + + +Author + +Schmidt-Rhaesa, Andreas +0000-0003-4102-9371 +ZTM, Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. +haesa@leibniz-lib.de + + + +Author + +Utevsky, Serge +0000-0003-1290-6742 +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. +serge.utevsky@karazin.ua + + + +Author + +Kristensen, Reinhardt Møbjerg +0000-0001-9549-1188 +Natural History Museum of Denmark, University of Copenhagen, Gothersgade 130, 1123 Copenhagen, Denmark. +rmkristensen@snm.ku.dk + + + +Author + +Kieneke, Alexander +0000-0001-8180-2483 +German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. +alexander.kieneke@senckenberg.de + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +1 +39 + + + + +https://doi.org/10.11646/zootaxa.5543.1.1 + +journal article +10.11646/zootaxa.5543.1.1 +1175-5326 +14385085 +F6372401-5C79-487C-A8C1-DBDEE7C71671 + + + + + + + +Styraconyx takeshii +Fujimoto, Suzuki, Ito, Tamura & Tsujimoto, 2020 + + + + + +N= + +1 specimen +: female occurrence: ANDEEP +II +; + +1,109 m +bsl + + +. + + + + +Figs 10F–I +; Supplementary +Table 1.11 + + + + \ No newline at end of file diff --git a/data/03/84/87/03848797FF8EFFE19B8DE3AEAE1015FC.xml b/data/03/84/87/03848797FF8EFFE19B8DE3AEAE1015FC.xml new file mode 100644 index 00000000000..c0a2cf9bb6b --- /dev/null +++ b/data/03/84/87/03848797FF8EFFE19B8DE3AEAE1015FC.xml @@ -0,0 +1,119 @@ + + + +Towards a better understanding of deep-sea tardigrade biogeography: numerous new records from the Southern Ocean + + + +Author + +Trokhymchuk, Roman +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. & German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. + + + +Author + +Schmidt-Rhaesa, Andreas +0000-0003-4102-9371 +ZTM, Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. +haesa@leibniz-lib.de + + + +Author + +Utevsky, Serge +0000-0003-1290-6742 +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. +serge.utevsky@karazin.ua + + + +Author + +Kristensen, Reinhardt Møbjerg +0000-0001-9549-1188 +Natural History Museum of Denmark, University of Copenhagen, Gothersgade 130, 1123 Copenhagen, Denmark. +rmkristensen@snm.ku.dk + + + +Author + +Kieneke, Alexander +0000-0001-8180-2483 +German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. +alexander.kieneke@senckenberg.de + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +1 +39 + + + + +https://doi.org/10.11646/zootaxa.5543.1.1 + +journal article +10.11646/zootaxa.5543.1.1 +1175-5326 +14385085 +F6372401-5C79-487C-A8C1-DBDEE7C71671 + + + + + + + +Tholoarctus oleseni +Jørgensen, Boesgaard, Møbjerg, & Kristensen, 2014 + + + + + +N= +8 specimens +: + +1 female +, +4 males +, 3 with undetermined sex occurrence: ANDEEP I, +ANDEEP-SYSTCO +; + +1,927–2,997 m +bsl + + +. + + + + +Fig. 11 +; Supplementary +Table 1.12 + + + + \ No newline at end of file diff --git a/data/03/84/87/03848797FF91FFFD9B8DE58CA9DB11C3.xml b/data/03/84/87/03848797FF91FFFD9B8DE58CA9DB11C3.xml new file mode 100644 index 00000000000..589ed7856d8 --- /dev/null +++ b/data/03/84/87/03848797FF91FFFD9B8DE58CA9DB11C3.xml @@ -0,0 +1,228 @@ + + + +Towards a better understanding of deep-sea tardigrade biogeography: numerous new records from the Southern Ocean + + + +Author + +Trokhymchuk, Roman +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. & German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. + + + +Author + +Schmidt-Rhaesa, Andreas +0000-0003-4102-9371 +ZTM, Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. +haesa@leibniz-lib.de + + + +Author + +Utevsky, Serge +0000-0003-1290-6742 +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. +serge.utevsky@karazin.ua + + + +Author + +Kristensen, Reinhardt Møbjerg +0000-0001-9549-1188 +Natural History Museum of Denmark, University of Copenhagen, Gothersgade 130, 1123 Copenhagen, Denmark. +rmkristensen@snm.ku.dk + + + +Author + +Kieneke, Alexander +0000-0001-8180-2483 +German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. +alexander.kieneke@senckenberg.de + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +1 +39 + + + + +https://doi.org/10.11646/zootaxa.5543.1.1 + +journal article +10.11646/zootaxa.5543.1.1 +1175-5326 +14385085 +F6372401-5C79-487C-A8C1-DBDEE7C71671 + + + + + + + +Batillipes wyedeleinorum +Bartels, Fontoura, Nelson & Kaczmarek, 2024 + + + + + +N= +2 specimens +: +1 female +, +1 juvenile +occurrence: ANDEEP I; +60°38.12′S +, +53°57.67′W +; +2,893 m +bsl (female); ANDEEP 2; +65°20.09′S +, +54°14.72′W +; +1,088 m +bsl (juvenile). + + + + +Both specimens are morphologically similar to the recently described species + +Batillipes wyedeleinorum + +Bartels +et al. +, 2024 + + +. However, we present our detailed description of an adult female since it adds important information to the original description of + +B. wyedeleinorum + +regarding character variation and ecological preferences ( +e.g. +deep sea occurrence). + + +Description of female + + +Figs 4–5 +, Supplementary +Table 1.1 + + +Body. +Small + +Batillipes + +with total body length (from cephalic rim to the tip of caudal appendage) of 147 µm, and maximum body width of 46 µm between leg III and leg IV ( +Figs 4 +, +5 +A-C). Distinct trapezoid head, separated from the body by a distinct neck constriction and with conical lateral projections I of 7 µm length ( +Fig. 5C, G +). Lateral projections between leg I and leg II (2.5 µm) are blunt. Lateral projections between leg II and III (3 µm) are conical. The lateral projections between leg III and leg IV (4 µm) are elongated (up to 20 µm) and rather blunt ( +Fig. 5C, G +). Body cuticle punctuated with internal pillars ( +Fig. 5G–J +). + + +Cephalic region. +Cephalic cirri with cirrophores. Cephalic cirri tips have tufts with additional filaments ( +Fig. 5G, H +). Cirri +A +are 24 µm in length. Unpaired median cirrus is 16 µm long. Internal cirri are 14 µm long and attached dorsally. External cirri are 11 µm long and situated close to the primary clavae. Primary clavae of 9.0 µm length are tube-shaped. ( +Fig. 5A +). Primary clavae, cirrus +A +and external cirri all originate from a common lateral bulge of the head ( +Fig. 4 +). Secondary clavae are indistinct. Eyes not visible. Mouth cone is clearly visible ( +Fig. 5G, H +). Ovoid pharyngeal bulb (15 × 10 µm) connected with the buccal tube with three placoids. + + +Legs +. Sensory organs present on all telescopic legs ( +Fig. 5B +). Sensory organs of legs I and II are about 7 µm long, of legs III to 10 µm. Sensory organs of legs IV with a cirrophore are 18 µm long and have frayed tips ( +Figs 5D, D +2). Bases of legs I and legs II have a small (1.5–2.0 µm) dome-shaped projection. On legs I, digits 2 (4.0 µm) and 4 (5.5 µm) are the shortest (considering digit 1 the most cephalically), digits 3 (10.5 µm) and 5 (11.0 µm) are the longest and digits 1 (6.0 µm) and 6 (7.0 µm) are medium sized. The same pattern of digits on legs II and III ( +Fig. 5A +). On legs IV, digits 3 (6.0 µm) and 4 (8.0 µm) are the shortest, and digits 1 (11.0 µm), 2 (14.5 µm), 5 (14.0 µm), 6 (11.0 µm) are the longest ( +Fig. 5D +). Digits have disc-shaped suction discs ( +Figs 4 +, +5A, D, J +). The specimen has some folded discs on legs II–IV ( +Fig. 4 +, +5I, J +). + + +Caudal region. +The caudal appendage is 8.5 µm long and looks like a single spine attached directly on the body. Rosette-shaped, six-lobed female gonopore and anus present ( +Fig. 5E, J +). Putative smooth area between gonopore and anus is visible on light microscopy figures ( +Fig. 5E +) but hidden by leg IV on SEM figures ( +Fig. 5J +). + + + + +Remarks + + +There are only a few and slight morphological differences between + +B. wyedeleinorum + +and our specimen. We did not see the same medial groove in a smooth area between gonopore and anus. In addition, we were not able to observe the papillae on legs because of legs positioning after preparation for SEM (see +Additional remarks +in + +Bartels +et al. +2024 + +). + +A possible ecological difference is evidenced by our new geographic and bathymetric record of this species. + + + \ No newline at end of file diff --git a/data/03/84/87/03848797FF95FFFA9B8DE0DEAE75102C.xml b/data/03/84/87/03848797FF95FFFA9B8DE0DEAE75102C.xml new file mode 100644 index 00000000000..6e142733123 --- /dev/null +++ b/data/03/84/87/03848797FF95FFFA9B8DE0DEAE75102C.xml @@ -0,0 +1,121 @@ + + + +Towards a better understanding of deep-sea tardigrade biogeography: numerous new records from the Southern Ocean + + + +Author + +Trokhymchuk, Roman +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. & German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. + + + +Author + +Schmidt-Rhaesa, Andreas +0000-0003-4102-9371 +ZTM, Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. +haesa@leibniz-lib.de + + + +Author + +Utevsky, Serge +0000-0003-1290-6742 +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. +serge.utevsky@karazin.ua + + + +Author + +Kristensen, Reinhardt Møbjerg +0000-0001-9549-1188 +Natural History Museum of Denmark, University of Copenhagen, Gothersgade 130, 1123 Copenhagen, Denmark. +rmkristensen@snm.ku.dk + + + +Author + +Kieneke, Alexander +0000-0001-8180-2483 +German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. +alexander.kieneke@senckenberg.de + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +1 +39 + + + + +https://doi.org/10.11646/zootaxa.5543.1.1 + +journal article +10.11646/zootaxa.5543.1.1 +1175-5326 +14385085 +F6372401-5C79-487C-A8C1-DBDEE7C71671 + + + + + + + +Coronarctus tenellus +Renaud-Mornant, 1974 + + + + + +N= +3 specimens +: + +1 female +, +1 male +, +1 juvenile +occurrence: +ANDEEP-SYSTCO +; + +2,891 m +bsl + + +. + + + + +Fig. 6 +; Supplementary +Table 1.3 + + + + \ No newline at end of file diff --git a/data/03/84/87/03848797FF95FFFA9B8DE256AE7515A4.xml b/data/03/84/87/03848797FF95FFFA9B8DE256AE7515A4.xml new file mode 100644 index 00000000000..bf3c8d3d15a --- /dev/null +++ b/data/03/84/87/03848797FF95FFFA9B8DE256AE7515A4.xml @@ -0,0 +1,115 @@ + + + +Towards a better understanding of deep-sea tardigrade biogeography: numerous new records from the Southern Ocean + + + +Author + +Trokhymchuk, Roman +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. & German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. + + + +Author + +Schmidt-Rhaesa, Andreas +0000-0003-4102-9371 +ZTM, Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. +haesa@leibniz-lib.de + + + +Author + +Utevsky, Serge +0000-0003-1290-6742 +VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. +serge.utevsky@karazin.ua + + + +Author + +Kristensen, Reinhardt Møbjerg +0000-0001-9549-1188 +Natural History Museum of Denmark, University of Copenhagen, Gothersgade 130, 1123 Copenhagen, Denmark. +rmkristensen@snm.ku.dk + + + +Author + +Kieneke, Alexander +0000-0001-8180-2483 +German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany. +alexander.kieneke@senckenberg.de + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +1 +39 + + + + +https://doi.org/10.11646/zootaxa.5543.1.1 + +journal article +10.11646/zootaxa.5543.1.1 +1175-5326 +14385085 +F6372401-5C79-487C-A8C1-DBDEE7C71671 + + + + + + + +Coronarctus dissimilis +Gomes-Júnior, Santos, da Rocha, Santos & Fontoura, 2020 + + + + + +N= + +1 specimen +: female occurrence: +ANDEEP-SYSTCO +; + +2,981 m +bsl + + +. + + + + +Fig. 6 +; Supplementary +Table 1.2 + + + + \ No newline at end of file diff --git a/data/03/84/87/038487B7FFA32232FF77B9FF910BFE81.xml b/data/03/84/87/038487B7FFA32232FF77B9FF910BFE81.xml new file mode 100644 index 00000000000..0191c8c3001 --- /dev/null +++ b/data/03/84/87/038487B7FFA32232FF77B9FF910BFE81.xml @@ -0,0 +1,670 @@ + + + +Two remarkable new species of Glaucocharis (Lepidoptera, Crambidae, Crambinae) from the Ogasawara Islands, Japan + + + +Author + +Matsui, Yuki +Entomological Laboratory, Faculty of Agriculture, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, 819 - 0395, Japan. + + + +Author + +Hamaguchi, Junpei +0009-0006-3684-0214 +Entomological Laboratory, Faculty of Agriculture, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, 819 - 0395, Japan. +hamaguchi.jumpei.281@s.kyushu-u.ac.jp + + + +Author + +Yagi, Sadahisa +0000-0002-4261-1219 +Insect DX Laboratory, Faculty of Agriculture, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, 819 - 0395, Japan. +yagi.sadahisa@gmail.com + + + +Author + +Hirowatari, Toshiya +0000-0002-6839-2229 +Entomological Laboratory, Faculty of Agriculture, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, 819 - 0395, Japan. & Entomological Laboratory, Faculty of Agriculture, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, 819 - 0395, Japan. +irowat_t@agr.kyushu-u.ac.jp + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +83 +96 + + + + +https://doi.org/10.11646/zootaxa.5543.1.4 + +journal article +10.11646/zootaxa.5543.1.4 +1175-5326 +14385137 +DA37B0E4-61F6-4C12-958D-91E15DC76B4C + + + + + + + +Glaucocharis triocellaris +Matsui, Yagi & Hirowatari + +, +sp. nov. + + + +(Japanese name: nishiki-eguri-tsutoga) + + + +( +Figs 1A +, +2A, 2B +, +3A +, +4A, 4C +, +5A, 5C, 5E +, +6A, 6C, 6D +, +7A +) + + + + +Diagnosis. +This new species can be easily distinguished from other + +Glaucocharis +species + +by distinctive morphological characters, such as the three black marginal spots, each enclosing a median silvery-white dot in the forewing; the valva with a sturdy hook-shaped projection emerging from the center of the ventral valva margin and the phallus with a ventrally curved hook-shaped apical thorn in the male genitalia; and the circular spinose lamella antevaginalis in the female genitalia. + + + + +Description. Head +( +Fig. 1A +). Frons brown, rounded. Vertex brown, lateral sides ocherous. Maxillary palpus almost as long as compound eye, outer side ocherous basally, pale ocherous apically, inner side white. Labial palpus about 1.6 times longer than compound eye; first palpomere ocherous dorsally, white ventrally; second palpomere with apically expanded scale tuft, ocherous on outer side, white on inner side; third palpomere white basally, brown apically, apex pointed. Maxillary and labial palpi strongly upturned in resting posture ( +Fig. 7A +). Antennae about 3/4 of forewing length, brown, ciliate in male, filiform in female; scape brown dorsally, white ventrally. Proboscis covered with white scales basally. + + +Thorax and legs. +Thorax brown dorsally, white ventrally. Patagium brown, lateral sides ocherous. Tegula ocherous on inner half and whitish silver on outer half. Foreleg femur white; tibia white, dorsal side gray apically; tarsus gray. Midleg femur white; tibia gray dorsally, white ventrally, inner and outer spurs gray, almost of equal length; tarsus gray. Hindleg femur white; tibia gray dorsally, white ventrally, mid and hind spurs gray, almost same length; tarsus gray. + + +Wings +( +Fig. 2A, 2B +). Forewing length +3.2–4.5 mm +(mean +3.9 mm +, +n += 10). Forewing ground color dark brown, tinged with reddish yellow from wing base to medial fascia and apical region; all fasciae and apical marking silvery white with blue luster; basal fascia represented by two oval spots; antemedial and medial fasciae almost straight; postmedial fascia oblique outwardly, ending near termen; submarginal fascia oblique outwardly, disappearing at R +5 +; apical mark represented by a short stripe vertical to costa; three marginal spots black, comma-shaped, connected, each enclosing silvery-white dot medially; marginal spots surrounded by yellow area with irregular black scaling along inner margin; subapical indentation of termen deep with white cilia; rest of cilia dark brown. Hindwing ground color dark brown, medial area white, with a narrow white band along CuA + +2 +in + +male; cilia brown. + + +Wing venation +( +Fig. 3A +) ( +n += 3). Forewing Sc concurrent with R +1 +at 2/3 of length, then diverging as short spur distally; R +2 +separate; R +3 +stalked with R +4 +at 3/5 distance from cell; R +5 +, M +1 +, M +2 +, and M +3 +separate, almost parallel; CuA +1 +close to M +3 +basally; CuA +2 +distant from CuA +1 +; A +1+2 +strong; A +3 +weak, not looped; cell opened. Hind wing Sc+R +1 +stalked with Rs at 4/5 of length; M +1 +, M +2 +, M +3 +, CuA +1 +, and CuA +2 +separate; CuP, A +1+2 +and A +3 +well-marked. Female with three frenular bristles. + + +Abdomen +. Dorsal side dark brown, ventral side pale brown, 1st and 2nd sternites tinged white. Tympanal organs ( +Fig. 4A +): bulla tympani small, bean-shaped, anterior margin convex; fornix tympani narrow, recessed under venula prima; pons tympani absent; venula prima developed, extending to about 2/3 from posterior margin of 2nd sternite. Male 8th tergite with a triangular sclerotization on anterior half ( +Fig. 4C +). Male 8th sternite with a long trapezoid sclerotization, posterior margin concaved medially ( +Fig. 4C +). Female eighth sternite wrinkled laterally ( +Fig. 6A +). + + +Male genitalia +( +Fig. 5A, 5C, 5E +) ( +n += 6). Uncus almost straight, apex pointed. Gnathos beak-shaped, dorsal side sparsely denticulate, apex pointed and curved dorsally. Tegumen long, dorso- and ventrolateral margins forming sclerotized ridges; socii finger-shaped with six petaloid lobes apically. Valva slender, curved ventrally, with sturdy hook-shaped projection emerging from middle of ventral margin; cucullus slightly enlarging distally, truncated apically, with long setae dorsally; costal margin sclerotized, with thumb-like costal arm at base; sacculus triangular, ventral side setose. Juxta oval, dorsal margin strongly concave medially. Saccus short, rounded. Phallus narrowing posteriorly, distal end pointed and slightly curved ventrally; apical thorn large, hook-shaped, strongly sclerotized, curved ventrally; cornuti absent; ductus ejaculatorius connected at anterior end of phallus. + + + +FIGURE 1. + +Glaucocharis +spp. + +head in lateral view. A: + +G. triocellaris + +male, paratype. B: + +G. plumbofascialis + +male, holotype. + + + + +FIGURE 2. + +Glaucocharis +spp. + +adults in dorsal view. A: + +G. triocellaris + +, holotype male. B: ditto, paratype female. C: + +G. plumbofascialis + +, holotype male. D: ditto, paratype female. af: antemedial fascia, am: apical mark, bf: basal fascia, mf: medial fascia, ms: marginal spot, pf: postmedial fascia, sf: submarginal fascia, si: subapical indentation. + + + + +FIGURE 3. + +Glaucocharis +spp. + +, male wing venations. A: + +G. triocellaris + +, paratype (wing slide no. YM-W-10). B: + +G. plumbofascialis + +, paratype (wing slide no. YM-W-8). + + + +Female genitalia +( +Fig. 6A, 6C, 6D +) ( +n += 6). Papillae anales ovate, membranous, weakly fused. Anterior and posterior apophyses thin and straight, almost equal in length. Lamella postvaginalis a square plate. Lamella antevaginalis a circular spinose sclerotization, covering ostium. Ductus bursae membranous, almost as long as corpus bursae. Ductus seminalis located at anterior 2/3 of ductus bursae. Corpus bursae membranous, anterior half slightly swollen, posterior half granulate; signa absent. + + + + +Type material. + + +Holotype +. + +♂ +[JPN: Tokyo-Pref.] +Shigureyama +, +Chichijima Is. +, +Ogasawara-mura +, +N27.0629 +, +E142.2214 +, alt. + +252 m + +, + +9.iii.2023 + +, LT, Shunsuke TOMURA leg., gen. slide no. YM-853, deposited in +ELKU + +. + + +Paratypes +. [ +Chichijima Is. +] + +2♂ +7♀ +, same locality as holotype, + +18.vi.2022 + +, LT, +T +. +Hirowatari +et al. leg., gen. slide no. YM-534, +Wing +slide no. YM-W-10, DNA sample id. M23-037 + +; + +1♂ +, same locality, + +26.vi.2022 + +, LT, +Shunsuke +TOMURA leg + +.; + +1♀ +, same locality, + +13.xi.2022 + +, LT, +T +. +Hirowatari +et al. leg + +.; + +12♂ +1♀ +, same locality, + +9.iii.2023 + +, LT, +T +. +Hirowatari +et al. leg., gen. slide no. YM-533, YM-851, wing slide no. YM-W-7 + +; + +1♂ +1♀ +, same locality, + +13.iii.2023 + +, LT, +T +. +Hirowatari +et al. leg + +.; + +1♂ +1♀ +, +Hatsuneura Observatory +, + +12.iii.2022 + +, +SW +, +T +. +Hirowatari +et al. leg., gen. slide no. YM-735, YM-835 + +; + +1♂ +, +Higashidaira +, + +26.vi.2022 + +, +Shunsuke +TOMURA leg + +.; + +1♂ +, same locality, + +13.vii.2024 + +, +Y. Matsui +leg. + +; + +1♀ +, same locality, + +10.iii.2023 + +, +SW +, +T +. +Hirowatari +et al. leg., gen. slide no. YM-737 + +; + +1♂ +, +Higashidaira +~ +Mt. Hatsune-yama +, + +13.vi.2023 + +, J.-H. +PARK +leg + +.; + +1♀ +, +Mt. Mikazuki-yama +, + +13.vi.2023 + +, +SW +, J.-H. +PARK +leg., gen. slide no. YM-855 + +; + +1♀ +, +Kitafukurozawa +~ +Nishikaigan +, + +13.iii.2023 + +; +T +. +Hirowatari +et al. leg., gen. slide no. YM-856, DNA sample id. M23-180 + +; + +1♂ +, +Kuwanokiyama +, + +14.xi.2022 + +, LT, +T +. +Hirowatari +et al. leg + +.; + +1♂ +, Ogaguwa-no-mori, + +12.iii.2023 + +, +SW +, +T +. +Hirowatari +et al. leg + +.; + +1♂ +, same data except LT. + +[ +Hahajima Is. +] + + + +2♂ +2♀ +, +Mt. Funaki-yama +, + +25.ix.2023 + +(larva), + +2-26.xi.2023 + +em. (host: +Bryopsida +spp.), +J. Hamaguchi +leg. + +, gen. slide no. YM-857, YM-836, DNA sample id. M23-181; + +1♂ +, same locality, + +14.vii.2024 + +, +Y. Matsui +leg. + +; + +3♂ +, +Mt. Kuwanoki-yama +, + +8.xi.2022 + +, LT, +T +. +Hirowatari +et al. leg., wing slide no. YM-W-6 + +; + +1♂ +, same locality, + +9.xi.2022 + +, +T +. +Hirowatari +leg. + +; + +2♂ +, +Tamagawadam +, + +25.ix.2023 + +, +light trap +, +Yuki +MATSUI leg., gen. slide no. YM-749, DNA sample id. M23-161 + +; + +1♂ +, +Mt. Chôkiyama +, + +15.vi.2023 + +, LT, +Sadahisa +YAGI +leg + +.; + +1♂ +, +Sakaigatake +, + +22.vi.2022 + +, +SW +, +S. Yagi +leg. + +; + +1♀ +, same locality and collecting date, +Shunsuke +TOMURA leg., gen. slide no. YM-858, DNA sample id. M23-182 + +. + + + + +Biology +( +Fig. 7A, 7D–F +). Although larvae were not found, the adults emerged from various Bryopsida species collected on Hahajima Island. The larvae made a cocoon using the leaf pieces of Bryopsida spp. ( +Fig. 7E +) and then pupated therein. The pupal exuviae were not exposed from the cocoon after emergence ( +Fig. 7F +). Adults fly both during the day and night, but most specimens were collected by light trapping at night. The species is presumed multivoltine. + + + + +Distribution. +Japan +: Ogasawara Islands (Chichijima and Hahajima Islands). + + + + +Etymology. +The name of the new species is derived from the three marginal spots on the forewing. + + + + \ No newline at end of file diff --git a/data/03/84/87/038487B7FFA6223DFF77BE339511F854.xml b/data/03/84/87/038487B7FFA6223DFF77BE339511F854.xml new file mode 100644 index 00000000000..1becfb293da --- /dev/null +++ b/data/03/84/87/038487B7FFA6223DFF77BE339511F854.xml @@ -0,0 +1,809 @@ + + + +Two remarkable new species of Glaucocharis (Lepidoptera, Crambidae, Crambinae) from the Ogasawara Islands, Japan + + + +Author + +Matsui, Yuki +Entomological Laboratory, Faculty of Agriculture, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, 819 - 0395, Japan. + + + +Author + +Hamaguchi, Junpei +0009-0006-3684-0214 +Entomological Laboratory, Faculty of Agriculture, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, 819 - 0395, Japan. +hamaguchi.jumpei.281@s.kyushu-u.ac.jp + + + +Author + +Yagi, Sadahisa +0000-0002-4261-1219 +Insect DX Laboratory, Faculty of Agriculture, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, 819 - 0395, Japan. +yagi.sadahisa@gmail.com + + + +Author + +Hirowatari, Toshiya +0000-0002-6839-2229 +Entomological Laboratory, Faculty of Agriculture, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, 819 - 0395, Japan. & Entomological Laboratory, Faculty of Agriculture, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, 819 - 0395, Japan. +irowat_t@agr.kyushu-u.ac.jp + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +83 +96 + + + + +https://doi.org/10.11646/zootaxa.5543.1.4 + +journal article +10.11646/zootaxa.5543.1.4 +1175-5326 +14385137 +DA37B0E4-61F6-4C12-958D-91E15DC76B4C + + + + + + + +Glaucocharis plumbofascialis +Matsui, Yagi & Hirowatari + +, +sp. nov. + + + + + + +( +Figs 1B +, +2C, 2D +, +3B +, +4B, 4D +, +5B, 5D, 5F +, +6B, 6E +, +7B, 7C +) + +(Japanese name: munin-eguri-tsutoga) + + + +Diagnosis. +This new species can be easily distinguished from other + +Glaucocharis +species + +by its distinctive morphological characters, such as the plumbeous postmedial and submarginal fasciae sharply angled outwards at + + +M + +1 +in + +the forewing; the valva cucullus with a strongly sclerotized prong and the phallus with a sheath-shaped apical thorn in the male genitalia; and the long square plate-like lamellae postvaginalis and antevaginalis in the female genitalia. + + + + +Description. Head +( +Fig. 1B +). Frons pale ocherous, anterior margin white. Vertex brown, lateral sides ocherous. + +Maxillary palpus about 0.7 times longer than compound eye, outer side ocherous basally, pale ocherous apically, + +inner side white. Labial palpus about 1.6 times longer than compound eye; first palpomere ocherous dorsally, white ventrally; second palpomere with apically expanded scale tuft, ocherous on outer side, white on inner side; third palpomere pale ocherous, with apex pointed. Maxillary and labial palpi strongly upturned in resting posture ( +Fig. 7B +). Antennae about 3/4 of forewing length, brown, ciliate in male, filiform in female; scape brown dorsally, white ventrally. Proboscis covered with white scales basally. + + +Thorax and legs. +Thorax brown dorsally, white ventrally. Patagium and tegula brown. Foreleg entirely white. Midleg femur white, ventral side basally gray; tibia gray dorsally, cream white ventrally, inner and outer spurs cream white, inner one slightly longer than outer one; tarsus cream white. Hindleg femur cream white, ventral side black at 2/3 from base; tibia gray dorsally except basal 1/3, cream white ventrally, mid and hind spurs gray, respective inner and outer spurs almost same length; tarsus cream white. + + + +FIGURE 5. + +Glaucocharis +spp. + +male genitalia. A: + +G. triocellaris + +whole genitalia (gen. slide no. YM853). B: + +G. plumbofascialis + +, ditto (gen. slide no. YM837). C: + +G. triocellaris + +uncus, tegumen, and gnathos lateral view (gen. slide no. YM857). D: + +G. plumbofascialis + +, ditto (gen. slide no. YM738). E: + +G. triocellaris + +juxta (gen. slide no. YM857). F: + +G. plumbofascialis + +, ditto (gen. slide no. YM837). at: apical thorn, ca: costal arm, cl: cucullus, gn: gnathos, jx: juxta, ph: phallus, pl: petaloid lobes, sa: saccus, sc: socius, sl: sacculus, tg: tegumen, un: uncus, va: ventral arms of juxta. Scale bars: A, B = 0.5 mm, C–F = 0.1 mm. + + + + +FIGURE 6. + +Glaucocharis +spp. + +female genitalia.A: + +G. triocellaris + +whole genitalia (gen. slide no. YM836). B: + +G. plumbofascialis + +, ditto (gen. slide no. YM838). C: + +G. triocellaris + +, around ostium (lamella antevaginalis opened anteriorly, gen. slide no. YM836). D: ditto, lamella antevaginalis normal position (gen. slide no. YM735). E: + +G. plumbofascialis + +, ditto (gen. slide no. YM750). cb: corpus bursae, db: ductus bursae, la: lamella antevaginalis, lp: lamella postvaginalis, os: ostium. Scale bars: A, B = 0.5 mm, C–E = 0.1 mm. + + + +Wings +( +Fig. 2C, 2D +). Forewing length +3.1–3.9 mm +(mean +3.4 mm +, +n += 10). Forewing ground color dark brown, outer area of medial fascia ocherous; basal, antemedial, medial, postmedial, and submarginal fasciae plumbeous with blue luster, both sides of each fascia margined with black lines; basal, antemedial, and medial fasciae obscure, angled outwards at M +1 +; postmedial and submarginal fasciae distinct, sharply angled outwards at M +1 +, whitish near costa; discal spot ocherous, oval, located at inner angle of medial fascia; two marginal spots irregular, black, located at M +3 +and CuA +2 +on termen; subapical indentation on termen vestigial; cilia plumbeous basally, dark brown distally. Hindwing ground color brown, distally darker, with whitish fascia subapically; in male, basal area covered with black lustered scales, and a whitish gray fascia along CuA +2 +; cilia as in forewing. + + +Wing venation +( +Fig. 3B +) ( +n += 4). Almost as in + +G. triocellaris + +, but hindwing lacking A +3 +, and M +3 +and CuA +1 +occasionally stalked basally (as in +Fig. 3B +). + + +Abdomen. +Dorsally dark brown. Ventrally brown, with posterior margin of each segment white. Tympanal organs ( +Fig. 4B +) as in + +G. triocellaris + +, but ramus tympani present as horizontal bar connected to venula prima laterally. Male 8th tergite with Y-shaped sclerotization medially ( +Fig. 4D +). Male 8th sternite with long square sclerotization medially, posterior margin slightly concave ( +Fig. 4D +). + + +Male genitalia +( +Fig. 5B, 5D, 5F +) ( +n += 9). Uncus long, curved ventrally, apex pointed. Gnathos subtriangular, at right angle from tegumen, dorsal side denticulate apically. Tegumen long, dorso- and ventrolateral margins forming sclerotized ridges; socii finger-shaped with two petaloid lobes apically. Valva slender; cucullus with a strongly sclerotized thorn apically; costal margin with dense long setae, with a short finger-like costal arm at base; sacculus short, triangular. Juxta heart-shaped, lateral sides extended as a pair of ventral arms. Saccus rounded. Phallus tapering posteriorly; apical thorn sheath-shaped, apex truncated; cornuti absent; ductus ejaculatorius connected at anterior end of phallus. + + +Female genitalia +( +Fig. 6B, 6E +) ( +n += 7). Papillae anales, anterior and posterior apophyses as in those of + +G. triocellaris + +. Lamella postvaginalis a long square plate, posterior margin slightly concave medially. Lamella antevaginalis a shorter square plate than lamella postvaginalis, anterior margin folded postero-internally, then strongly constricted and connected to ductus bursae. Ductus bursae membranous, short and thin. Ductus seminalis located at anterior 2/3 of ductus bursae. Corpus bursae ovate, approximately 2–3 times longer than ductus bursae, posterior half lightly sclerotized with smooth surface, anterior half membranous, with weak granules; signa absent. + + + + +Type material. + + +Holotype +. + +♂ +[JPN: Tokyo-Met.] +Mt. Funaki-yama +, +Haha-jima Is. +, +Ogasawara-mura +, + +25.ix.2023 + +(larva), + +26.xi.2023 + +em. (host: Bryopsida spp.), +J. Hamaguchi +leg., gen. slide no. YM-854 + +. + + +Paratypes +. [ +Chichijima Is. +] + +2♀ +, +Asahiyama +, + +11.vi.2023 + +, +T +. +Hirowatari +leg. + +; + +1♀ +, same locality and collector, + +18.vi.2022 + + +; + +2♂ +, +Asahiyamatenbodai +, +Sakaiura +, + +26.vi.2022 + +, +SW +, +S. Yagi +leg. + +, gen. slide no. YM-535, YM-789, wing slide no. YM-W-8; + +1♂ +, +Higashidaira +, + +18.vi.2022 + +, +Shunsuke +TOMURA leg., gen. slide no. YM-861, DNA sample id. M23-185 + +; + +1♀ +, same locality, + +26.vi.2022 + +, +T +. +Hirowatari +leg. + +; + +1♂ +, +Higashi-machi +, + +11.iii.2023 + +, +T +. +Hirowatari +et al. leg + +.; + +1♂ +, +Kuwanokiyama +, + +19.vi.2022 + +, +LT +, +T +. +Hirowatari +et al.leg + +.; + +3♀ +,same locality, + +23.vi.2022 + +, +M.Kimura +leg. + +; + +1♂ +, +Mikazukiyama +, + +13.vi.2023 + +, +T +. +Hirowatari +leg. + +, gen. slide no. YM-792; + +1♀ +, +Mt. Chûô-san +, + +19.vi.2023 + +, +SW +, +J.-H. Park +leg. + +; + +1♀ +, +Ogamiyama +, + +25.vi.2022 + +, +LT +, +Shunsuke +TOMURA leg., gen. slide no. YM-537 + +; + +1♂ +, same locality, + +11.vii.2024 + +, +Y. Matsui +leg. + +; + +1♂ +, +Mt. Akahata-yama +; + +11.vi.2023 + +, +light trap +, +Yuki +MATSUI leg., gen. slide no. YM-738 + +; + +1♂ +1♀ +, +Shigureyama +, + +18.vi.2022 + +, +LT +, +Shunsuke +TOMURA leg., gen. slide no. YM-862, DNA sample id. M23-186 + +; + +1♀ +, +Yoakeyama +, + +12.iii.2023 + +, +T +. +Hirowatari +leg. + +, gen. slide no. YM-736. +[Anijima Is.] + +1♀ +, +Takinoura +, + +20.vi.2022 + +, beating: dead leaf + +Livistona chinensis +var. +boninensis +, Shunsuke TOMURA + +leg., gen. slide no. YM-750, DNA sample id. M23-162. + +[ +Otôtojima Is. +] + + + +5♂ +, + +12-13.vii.2024 + +, +SW +, +S. Yagi +leg. + + + +[ +Mukohjima Is. +] + + +1♂ +, +16.vii.2024 +, Y. Matsui leg. + + +[ +Hahajima Is. +] + + + +1♂ +, same locality as holotype, + +24.vi.2022 + +, +Shunsuke +TOMURA leg + +.; + +1♀ +, same locality, + +17.iii.2023 + +, +LT +, +T +. +Hirowatari +et al. leg., gen. slide no. YM-859, DNA sample id. M23-183 + +; + +2♂ +, same locality, + +14.vii.2024 + +, +Y. Matsui +leg. + +; + +2♂ +1♀ +, +Choukiyama +, + +15.vi.2023 + +, +LT +, +T +. +Hirowatari +et al. leg., gen. slide no. YM-837 + +; + +1♂ +, +Mt. Chibusa-yama +, + +16.vi.2023 + +(larva), + +11.vii.2023 + +em. (host: +Bryopsida +spp.), +J. Hamaguchi +leg. + +; + +♀ +1, same locality and collecting date, +LT +, +Sadahisa +YAGI +leg + +.; + +1♀ +, +Mt. Kuwanoki-yama +, + +8.xi.2022 + +, +LT +, +T +. +Hirowatari +et al. leg + +.; + +1♀ +, +Mt. Sakaigatake +, + +16.iii.2023 + +, +Toshiya Hirowatari +leg. + +; + +1♂ +2♀ +, same locality, + +17.vi.2023 + +, +LT +, J.-H. +Park +& I. KAWASHIMA leg., DNA sample id. M23-024 + +; + +2♀ +, same locality, + +24.ix.2023 + +(larva), + +8.xi.2023 + +em. (host: +Bryopsida +spp.), +J. Hamaguchi +leg. + +, gen. slide no. YM-838; + +1♀ +, +Nishiura +, + +15.iii.2023 + +, +LT +, +T +. +Hirowatari +et al. leg + +.; + +2♂ +, +Higashikô +, + +24.ix.2023 + +, flower of + +Bidens pilosa +, Masaaki Kimura + +leg., gen. slide no. YM-791, YM-790 + +; + +1♂ +, +Shinyûhigaoka +, + +21.ix.2023 + +light trap +; +Yuki +MATSUI leg., gen. slide no. YM-860, DNA sample id. M23-184 + +. + + + + +Biology +( +Fig. 7B–F +). Similar to + +G. triocellaris + +. We observed that some adults visited the flowers of + +Bidens pilosa +var. +radiata +(Sch. Bip.) J.A. Schmidtthe + +during daytime. We also observed many adults swarming around + +Euonymus boninensis +Koidz. + +at twilight: all individuals were males as far as we could confirm. + + + + +Distribution. +Japan +: Ogasawara Islands (Chichijima, Anijima, Otôtojima, Mukohjima, and Hahajima Islands). + + + + +Etymology. +The name of the new species is derived from the plumbeous (dull-gray colored) fasciae of the forewing. + + + + \ No newline at end of file diff --git a/data/03/86/87/03868796FF92FFF8FF1EE056EFDCFB87.xml b/data/03/86/87/03868796FF92FFF8FF1EE056EFDCFB87.xml new file mode 100644 index 00000000000..27b51ec12ad --- /dev/null +++ b/data/03/86/87/03868796FF92FFF8FF1EE056EFDCFB87.xml @@ -0,0 +1,336 @@ + + + +Free-living marine nematodes of the genus Richtersia Steiner, 1916 (Desmodorida: Richtersiidae) from the Sea of Japan + + + +Author + +Revkova, Tatiana N. +0000-0002-3500-3776 +A. O. Kovalevsky Institute of Biology of the Southern Seas of RAS, 2, Nakhimov ave., 299011, Sevastopol, Russia +alinka8314@gmail.com + + + +Author + +Sergeeva, Nelli G. +0000-0002-3984-0918 +A. O. Kovalevsky Institute of Biology of the Southern Seas of RAS, 2, Nakhimov ave., 299011, Sevastopol, Russia +nserg05@mail.ru + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +123 +134 + + + + +https://doi.org/10.11646/zootaxa.5543.1.7 + +journal article +10.11646/zootaxa.5543.1.7 +1175-5326 +093A5AA0-DAEF-4312-B8D9-897F7A6B779C + + + + + + + +Richtersia amurica + +sp. nov. + + + + + + +( +Figures 4–5 +, +Table 1 +) + + + + +Type material. +Six males +and +six females +. +Holotype +male mounted on slide Meib.53.N.h., +five paratype males +mounted on slides: Meib.19.N.p. ( +2 males +), Meib.55.N.p., Meib.57.N.p., Meib.58.N.p. and +six paratype females +: Meib.22.N.p., Meib.23.N.p., Meib.25.N.p., Meib.51.N.p., Meib.52.N.p., Meib.54.N.p. in pure glycerin. + + + + +Type +locality and habitat. + +Amur Bay +, +Sea +of +Japan +, + +20 June 2021 + +, station 11, water depth + +6 m + +, dense silt, finegrained sand, coordinates: +N 42°52.471 +, +E 131°39.202 + +. + + + + +Etymology. +The new species epithet + +“ +amurica + +” refers to the locality of collection, the +Amur +Bay. + + +Descriptions. + + +Males. +Body plump, short, truncated anteriorly and tapering posteriorly. Cuticle annulated about 1 μm in width. Each annule carrying backward-pointing spines about 36–38 rows in the anterior part, 22–24 rows in the mid-body region and 13–15 rows in the cloacal region. Posterior half of the tail lacking spines but with irregular annules; the tail tip is smooth. On the ventral side, longitudinal rows of robust spines located approximately 19–24 µm from the cloaca. Somatic setae 5–6 μm long in eight rows. The six internal labial setae 9–11 μm long, situated on a hyaline labial membrane; the six external labial setae 8–9 μm long and four cephalic setae 4–5 μm long, at almost the same level. Subcephalic setae 5–6 μm long, one at each side of amphidial fovea. Amphidial fovea large with 3.5 turns, surrounded by spines, 46.9–59.7% of +chd +. Stoma large, unarmed and partly surrounded by pharyngeal tissue. Pharynx cylindrical lacking a posterior bulb. Cardia triangular, surrounded by the intestine. The intestinal cells containing dark brown granules, more abundant in the anterior part of the intestine. Nerve ring and secretory-excretory system unclear. + + +Reproductive system monorchic, with outstretched testis, lying partially ventral to the intestine. Two spicules unequal in shape and size: the right spicule very fine 2.8 (3.4–4.6 times cloacal body diameter) and the left spicule wide 1.0 (0.9–2.0 times cloacal body diameter) with a capitulum; right spicule 2.0–2.7 times longer than the left spicule. Gubernaculum surrounding distal ends of spicules. Tail short, 1.1 (1.1–1.7) +cbd +long. Three caudal glands present. + + + +FIGURE 4. + +Richtersia amurica + + +sp. nov. + +A: holotype male, entire body; B: holotype male, amphidial fovea; C: paratype female, amphidial fovea; D: holotype male, spicules; E: paratype female, entire body; F: holotype male, head; G: paratype female, head. Scale bar: A, E = 100 μm; B–D, F, G = 20 μm. + + + + +FIGURE 5. + +Richtersia amurica + + +sp. nov. + +A: holotype male, entire body; B: holotype male, head region; C: paratype female, head region; D: holotype male, tail region; E: holotype male, amphidial fovea; F: paratype female, amphidial fovea; G: paratype female, tail region (Meib.25.N.p.); H: paratype female, vulva; I: paratype female, entire body; J: paratype female, cuticule on vulval region. Scale bar: A, I = 100 μm; B–H, J = 20 μm. + + + +Female. +General morphology similar to that of male. Amphidial fovea smaller, 23.5–29.4 % of chd, with one turn. Cardia invisible. The intestine is abundantly filled with dark brown granules in the anterior part. Reproductive system didelphic, amphidelphic. Ovaries reflexed. The tail is covered with long hairy spines ( +Figure 5G +), longer on the dorsal side than those on the ventral side. + + + + +Diagnosis. + +Richtersia amurica + + +sp. nov. + +is characterised by the number of longitudinal spines rows, located at the level of amphidial fovea; amphidial fovea with 3.5 turns in males and one turn in females; spicules unequal, gubernaculum with anterior apophyses in males; tail conical and short; females tail with long hairy spines. + + +Differential diagnosis. + +Richtersia amurica + + +sp. nov. + +is similar to + +R. inaequalis +, +R. coomansi + +, + +R. beibuwanensis + +and + +R. staresensis + +in having unequal spicules and in amphidial fovea structure in males and females. However, + +R. amurica + + +sp. nov. + +differs from them by the length of left (53–69 μm +vs +. 39–47 μm in + +R. inaequalis + +, 124–185 μm in + +R. coomansi + +, 98.2–108 μm in + +R. beibuwanensis + +, 41–46 μm in + +R. staresensis + +) and right (123–144 μm +vs +. 113–126 μm in + +R. inaequalis + +, 66–85 μm in + +R. coomansi +, + +44–50.9 μm in + +R. beibuwanensis + +, 87–93 μm in + +R. staresensis + +) spicules. In the arrangement of longitudinal spine rows beginning at the level of the amphidial fovea and in long hairy spines on the tail of females, the new species resembles + +R. inaequalis + +and + +R. coomansi + +, but it differs from them by the presence of anterior apophyses on gubernaculum ( +vs +. absent). + +R. amurica + + +sp. nov. + +also differs from + +R. inaequalis + +by having a relatively broader body (a = 8.9–10.7 +vs +. +12–14.4 in +males and 6.6–9.4 +vs. +10.5 in +females) and longer internal labial setae (9–11 μm +vs +. 6.3–7 μm). The new species can be distinguished also from + +R. coomansi + +by the spirally whorled amphidial fovea in males (3.5 turns +vs +. 5.25 turns) and internal labial setae without a prominent base ( +vs +. present). The new species is similar to + +R. beibuwanensis + +and + +R. staresensis + +by the presence of anterior apophyses on the gubernaculums, but differs from them by patterns of longitudinal spine rows (at the level of amphidial fovea +vs +. middle of the pharynx). + +R. amurica + + +sp. nov. + +also differs from + +R. beibuwanensis + +by having a longer body (515–895 μm +vs +. 373.7–405.6 μm), longer internal and external labial setae (9–11 μm and 8–9 μm +vs +. 6 μm), higher index +c +(10.5–24.2 +vs +. 6.6–7.5) and shorter amphidial fovea in females (23.5–29.4% +vs +. 36.3%). Furthermore, the new species differs from + +R. staresensis + +by having a smaller amphidial fovea in males (46.9–59.7% +chd +with 3.5 turns +vs +. 90% +chd +with 4 turns). + + + + \ No newline at end of file diff --git a/data/03/86/87/03868796FF96FFF3FF1EE518EF87FD5B.xml b/data/03/86/87/03868796FF96FFF3FF1EE518EF87FD5B.xml new file mode 100644 index 00000000000..b15411d0765 --- /dev/null +++ b/data/03/86/87/03868796FF96FFF3FF1EE518EF87FD5B.xml @@ -0,0 +1,90 @@ + + + +Free-living marine nematodes of the genus Richtersia Steiner, 1916 (Desmodorida: Richtersiidae) from the Sea of Japan + + + +Author + +Revkova, Tatiana N. +0000-0002-3500-3776 +A. O. Kovalevsky Institute of Biology of the Southern Seas of RAS, 2, Nakhimov ave., 299011, Sevastopol, Russia +alinka8314@gmail.com + + + +Author + +Sergeeva, Nelli G. +0000-0002-3984-0918 +A. O. Kovalevsky Institute of Biology of the Southern Seas of RAS, 2, Nakhimov ave., 299011, Sevastopol, Russia +nserg05@mail.ru + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +123 +134 + + + + +https://doi.org/10.11646/zootaxa.5543.1.7 + +journal article +10.11646/zootaxa.5543.1.7 +1175-5326 +093A5AA0-DAEF-4312-B8D9-897F7A6B779C + + + + + + +Genus + +Richtersia +Steiner, 1916 + + + + + + + +Emended diagnosis +(based in Gharakhani +et al +. 2020). Body short and stout, yellow-brownish, from cylindrical to sacciform. Cuticle finely annulated with longitudinal rows of small spines. Anterior sensilla arranged in 6 + 10 pattern, all anterior sensilla setose. Amphidial fovea ventrally coiled, uni- to multispiral, or of any other derived shapes. Broad mouth opening surrounded by six soft lips and a cuticular collar. Buccal cavity nearly cylindroid, with weakly sclerotised walls. Pharynx short, cylindrical, and evenly muscular. Male reproductive system monorchic, with outstretched testis. No precloacal supplements except for modified cuticular spines. Female reproductive system didelphic-amphidelphic. Strictly marine. + + + + + +Type +species + +: + +Richtersia collaris +Steiner, 1916 + + + + + \ No newline at end of file diff --git a/data/03/86/87/03868796FF96FFF4FF1EE378ECAEF84B.xml b/data/03/86/87/03868796FF96FFF4FF1EE378ECAEF84B.xml new file mode 100644 index 00000000000..11b25895683 --- /dev/null +++ b/data/03/86/87/03868796FF96FFF4FF1EE378ECAEF84B.xml @@ -0,0 +1,347 @@ + + + +Free-living marine nematodes of the genus Richtersia Steiner, 1916 (Desmodorida: Richtersiidae) from the Sea of Japan + + + +Author + +Revkova, Tatiana N. +0000-0002-3500-3776 +A. O. Kovalevsky Institute of Biology of the Southern Seas of RAS, 2, Nakhimov ave., 299011, Sevastopol, Russia +alinka8314@gmail.com + + + +Author + +Sergeeva, Nelli G. +0000-0002-3984-0918 +A. O. Kovalevsky Institute of Biology of the Southern Seas of RAS, 2, Nakhimov ave., 299011, Sevastopol, Russia +nserg05@mail.ru + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +123 +134 + + + + +https://doi.org/10.11646/zootaxa.5543.1.7 + +journal article +10.11646/zootaxa.5543.1.7 +1175-5326 +093A5AA0-DAEF-4312-B8D9-897F7A6B779C + + + + + + + +Richtersia japonica + +sp. nov. + + + + + + +( +Figures 2–3 +, +Table 1 +) + + + + +Type material. +Eight males +and +four females +. +Holotype +male mounted on slide Meib.18.N.h., +four paratype males +mounted on slides: Meib.17.N.p. ( +2 males +), Meib.21.N.p., Meib.60.N.p. and +two paratype females +: Meib.24.N.p., Meib.59.N.p. in pure glycerin; +three paratype males +mounted on slides: Meib.16.N.p. ( +2 males +), Meib.20.N.p. and +two paratype females +: Meib.20.N.p., Meib.56.N.p. in glycerine-gelatin. + + + +FIGURE 2. + +Richtersia japonica + + +sp. nov. + +A: holotype male, entire body; B. holotype male, spicules; C. holotype male, buccal cavity; D. paratype female, amphid level; E. holotype male, amphid level; F. paratype female, buccal cavity; G. paratype female, entire body. Scale bar: A, G = 100 μm; B–F = 20 μm. + + + + +FIGURE 3. + +Richtersia japonica + + +sp. nov. + +A. paratype female, entire body; B. holotype male, entire body; C. paratype female, amphid level; D. holotype male, amphid level; E. paratype female, buccal cavity; F. paratype female, mature egg; G. paratype male, tail region; H. paratype female, tail region and setae (arrows); I. paratype female, cuticula on vulval region; J. paratype female, vulva (arrow). Scale bar: A, B = 100 μm; C–J = 20 μm. + + + + + +Type +locality and habitat. + +Specimens +were collected from the +Sea +of +Japan +in + +June 2021 + +on station 7, water depth + +22.1 m + +, pelitic dense silt with mica plates, fluffy detritus on the surface, coordinates: +N 42°50.5 +, +E 131°26.4 + +. + + + + +Etymology. +The species epithet is derived from the Latin adjective meaning “Japanese”. + + +Descriptions. + + +Males. +Body short, cylindrical, rounded anteriorly, tapering posteriorly. Cuticle annulated, annuli with numerous short and stout spines from posterior to cephalic region to near tail tip. These spines arranged heterogeneously in regular longitudinal rows from about the level of the middle of the pharynx. Each annule carrying spines arranged in groups of longitudinal rows: about 50–53 rows at the level of the middle of the pharynx, about 31–33 rows in the mid-body region, and about 16–19 rows in the cloacal region. Posterior one-third of the tail lacking spines and with a smooth tail tip. Somatic setae 3–9 μm long, irregularly situated in eight rows between longitudinal rows of spines. Longitudinal rows of robust spines on the ventral side that begin at the level of the middle of the pharynx and end about 17–32 µm from the cloaca. Six internal labial setae 6–8 μm long, situated on a hyaline labial membrane; six external labial setae 5–6 μm long and four cephalic setae 4–5 μm long located at almost the same level. Subcephalic setae 4–5 μm long, one at each side of amphidial fovea. Amphidial fovea 40.7–60% of +chd +(corresponding head diameter), round and ventrally spiralled with 3.5 turns, surrounded by cuticular spines. Labial region with 12 lobes, strengthened by 12 cuticular rods, forming a pyramidal invagination of cheilostome wall into the stoma; invagination alternating with 12 triangular prisms. Stoma wide, funnel-shaped, unarmed and partly surrounded by pharyngeal tissue. Pharynx long, cylindrical and lacking a posterior bulb. Cardia not observed. The intestinal cells containing dark brown granules evenly dispersed all along the intestine. The nerve ring and secretory-excretory system indistinguishable. + + +Reproductive system monorchic, with outstretched testis. Two spicules unequal in shape and size: right spicule very fine, 4.3 (3.4–5.0) times cloacal body diameter; left spicule wider and 1.5 (1.2–2.2) times cloacal body diameter. Right spicule 2.8 (2.2–3.0) times longer than left spicule; left spicule with a capitulum. Gubernaculum surrounding distal ends of spicules, with an anterior apophysis on each side. Tail short, 1.2 (0.7–1.3) +cbd +(cloacal body diameter) long, with two caudal glands. + + +Females. +General morphology similar to that of males. Spines arranged in regular longitudinal 49–51 rows beginning close to the buccal cavity, 37–39 rows in the mid-body region, and 22–24 rows in the cloacal region. Somatic setae 3–6 μm long. Amphidial fovea smaller, 26–39.1% of +chd +, with one turn. Reproductive system didelphic, amphidelphic. Ovaries reflexed. Spermatozoa present in uterus. +One female +with a mature egg ( +Figure 3F +). The tail is covered with numerous rows of short hairy spines ( +Figure 3H +), in which substrate particles can accumulate. + + + + +Diagnosis. + +Richtersia japonica + + +sp. nov. + +is characterised by 450–655 μm long body; presence of multispiral amphidial fovea with 3.5 turns in males and one turn in females; longitudinal rows of spines located approximately from the level of the middle of the pharynx in males and behind the buccal cavity in females; two unequal spicules and gubernaculum with anterior apophyses, short conical tail; tail with hairy spines in females. + + +Differential diagnosis. +Based on the amphidial fovea structure and unequal spicules + +R. japonica + + +sp. nov. + +is similar to + +R +. +beibuwanensis +Fu, Cai, Boucher, Cao & Wu, 2013 + +, + +R. staresensis +Soetaert & Vincx, 1987 + +, + +R. coomansi +Soetaert & Vincx, 1987 + +and + +R. inaequalis +Riemann, 1966 + +. However, the new species differs from + +R. coomansi + +and + +R. inaequalis + +by having an anterior apophysis on gubernaculum ( +vs +. absent); length of the left (51–68 μm +vs +. 124– 185 μm in + +R. coomansi + +and 39–47 μm in + +R. inaequalis + +) and right 139–173 μm +vs +. 66–85 μm in + +R. coomansi + +and 113–126 μm in + +R. inaequalis + +) spicules; longitudinal rows of spines begin at the level of the middle of the pharynx in males and behind buccal cavity in females ( +vs. +at the level of amphidial fovea in + +R. coomansi + +and + +R. inaequalis + +). The new species differs from + +R. coomansi + +by the number of amphidial fovea turns in males (3.5 turns +vs +. 5.25 turns) and the shape of internal labial setae (usual +vs +. prominent base). + + +The new species mostly resembles + +R. beibuwanensis + +and + +R. staresensis + +in the arrangement of longitudinal rows of spines, but differs from it in de Man +a +value in males (8–11.9 +vs +. +7.1–7.4 in + +R. beibuwanensis + +and +15.4 in + +R. staresensis + +) and lengths of the spicules: right (139–173 μm +vs +. 44–50.9 μm in + +R. beibuwanensis + +and 87–93 μm in + +R. staresensis + +) and left (51–68 μm +vs +. 98.2–108 μm in + +R. beibuwanensis + +and 41–46 μm in + +R. staresensis + +). + +R. japonica + + +sp. nov. + +also differs from + +R. beibuwanensis + +by having a relatively longer body (450–655 μm +vs. +373.7– 399.6 μm in males, 450–561 μm +vs +. 405.6 μm in females) and subcephalic setae (present +vs +. absent). Furthermore, the new species differs from + +R. staresensis + +by having a smaller amphidial fovea in males (40.7–60% +chd +with 3.5 turns +vs +. 90% +chd +with 4 turns). + + + + \ No newline at end of file diff --git a/data/03/96/43/03964341FFDFFF80F8B8CBEF4D847D27.xml b/data/03/96/43/03964341FFDFFF80F8B8CBEF4D847D27.xml new file mode 100644 index 00000000000..c33a7785e21 --- /dev/null +++ b/data/03/96/43/03964341FFDFFF80F8B8CBEF4D847D27.xml @@ -0,0 +1,187 @@ + + + +A new genus of tribe Dikraneurini (Hemiptera: Cicadellidae: Typhlocybinae) from China based on a new species and genus checklist from China provided + + + +Author + +Jiao, Meng +Institute of Entomology, Guizhou University, Guiyang, 550025, P. R. China. & Guizhou Provincial Key Laboratory for Agricultural Pest Management of the Mountainous Region, Guizhou University, Guiyang, 550025, P. R. China. + + + +Author + +Yang, Maofa +Institute of Entomology, Guizhou University, Guiyang, 550025, P. R. China. & Guizhou Provincial Key Laboratory for Agricultural Pest Management of the Mountainous Region, Guizhou University, Guiyang, 550025, P. R. China. + + + +Author + +Yan, Bin +Institute of Entomology, Guizhou University, Guiyang, 550025, P. R. China. & Guizhou Provincial Key Laboratory for Agricultural Pest Management of the Mountainous Region, Guizhou University, Guiyang, 550025, P. R. China. + + + +Author + +Yu, Xiaofei +Institute of Entomology, Guizhou University, Guiyang, 550025, P. R. China. & Guizhou Provincial Key Laboratory for Agricultural Pest Management of the Mountainous Region, Guizhou University, Guiyang, 550025, P. R. China. + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +135 +140 + + + + +https://doi.org/10.11646/zootaxa.5543.1.8 + +journal article +10.11646/zootaxa.5543.1.8 +1175-5326 +43452907-49AA-4060-9167-7E12EC8015A6 + + + + + + +Genus. + +Sugea + +gen. nov. + + + + + + +Fig. 1 +: A–P + + + + + +Type +species. + + +Sugea obtusangula + + +sp. nov. + + + + + +Description +. Body ( +Fig. 1 +: A, B) relatively large, center of crown ( +Fig. 1 +: C, D, F) with prominent bulge, coronal suture short, head in dorsal view narrower than pronotum. Face ( +Fig. 1 +: E) protuberant in profile, with well-defined angles at dorsal margin of frontoclypeus and midlength of anteclypeus, and clypeal sulcus distinct, anteclypeus strongly swollen and lorum very narrow in male (female unknown); ocelli vestigial, crescent-shaped. Pronotum ( +Fig. 1 +: C, D) with small angulate protrusion at center of anterior margin, with sublateral visible in frontal view. Mesonotum and scutellum depressed, forming single plane narrower than pronotum. Color of forewings relatively uniform, elliptical waxy area located near the middle of the outer margin of the forewing, 3rd apical cell ( +Fig. 1 +: G) small and stalked. Hind wing ( +Fig. 1 +: H) transparent, anal veins and CuA veins forked. Abdominal apodemes ( +Fig. 1 +: I, N) underdeveloped. Pygofer side ( +Fig. 1 +: J, O) broad but irregular in shape, ventral margin with indentations but without appendage. Subgenital plate ( +Fig. 1 +: J, O, P) longer than pygofer, not fused at base, with longitudinal row of sparse macrosetae, apex expanded with protuberances. Style ( +Fig. 1 +: L, M) elongate, without preapical lobe. Connective short and broad, V-shaped. Aedeagus ( +Fig. 1 +: L, M, P) with gonopore-bearing shaft greatly reduced, with pair of elongate basal processes. Gonopore apical. + + + + +Distribution. +China +( +Yunnan +). + + + + +Etymology: +The new genus name is derived from the Chinese “Suge”, which symbolizes vitality and dynamism. Gender: feminine. + + + + +Remarks: +The new genus + +Sugea + +is similar to + +Igutettix +Matsumura (1932) + +, + +Vilbasteana +Anufriev (1970) + +, + +Sweta +Viraktamath et Dietrich (2011) + +, + +Koreoneura +Hossain et Kwon (2021) + +and + +Cornicola +Ohara et Hayashi (2022) + +, in the comparatively large body-size and large pronotum. Some features of the pronotum, including the distinct submarginal furrow present along the anterior and lateral margins, and subgenital plates not fused to each other or to valve, show similarity to those of + +Sweta + +. But it is distinguished from + +Sweta + +by the black coloration, short and smooth pronotum, forewing with 3 +rd +apical cell small and with stalk, and hindwings CuA and anal veins forked. Moreover this new genus is characterized by the face with distinct angular features in profile, the less strongly raised center of the crown, pronotum with a small protruding angle at the center of the anterior margin and end of the subgenital plate with a horn-like protrusion. These traits distinguish it from other genera. + + + + \ No newline at end of file diff --git a/data/03/96/43/03964341FFDFFF83F8B8CE624C637966.xml b/data/03/96/43/03964341FFDFFF83F8B8CE624C637966.xml new file mode 100644 index 00000000000..27ce32c02d1 --- /dev/null +++ b/data/03/96/43/03964341FFDFFF83F8B8CE624C637966.xml @@ -0,0 +1,174 @@ + + + +A new genus of tribe Dikraneurini (Hemiptera: Cicadellidae: Typhlocybinae) from China based on a new species and genus checklist from China provided + + + +Author + +Jiao, Meng +Institute of Entomology, Guizhou University, Guiyang, 550025, P. R. China. & Guizhou Provincial Key Laboratory for Agricultural Pest Management of the Mountainous Region, Guizhou University, Guiyang, 550025, P. R. China. + + + +Author + +Yang, Maofa +Institute of Entomology, Guizhou University, Guiyang, 550025, P. R. China. & Guizhou Provincial Key Laboratory for Agricultural Pest Management of the Mountainous Region, Guizhou University, Guiyang, 550025, P. R. China. + + + +Author + +Yan, Bin +Institute of Entomology, Guizhou University, Guiyang, 550025, P. R. China. & Guizhou Provincial Key Laboratory for Agricultural Pest Management of the Mountainous Region, Guizhou University, Guiyang, 550025, P. R. China. + + + +Author + +Yu, Xiaofei +Institute of Entomology, Guizhou University, Guiyang, 550025, P. R. China. & Guizhou Provincial Key Laboratory for Agricultural Pest Management of the Mountainous Region, Guizhou University, Guiyang, 550025, P. R. China. + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +135 +140 + + + + +https://doi.org/10.11646/zootaxa.5543.1.8 + +journal article +10.11646/zootaxa.5543.1.8 +1175-5326 +43452907-49AA-4060-9167-7E12EC8015A6 + + + + + + + +Sugea obtusangula + +gen. et sp. nov. + + + + + + +Fig. 1 +: A–P + + +Description +. Length: male +4.2–4.5 mm +. + + +Body ( +Fig. 1 +: A, B) black with white spots, four white spots surround the central protrusion of crown ( +Fig. 1 +: F); eyes brown. Face ( +Fig. 1 +: E) black, postclypeus and anteclypeus relatively pale, antennae yellow. Front and postclypeus separated by distinct transverse ridge, most evident in lateral view. Pronotum ( +Fig. 1 +: C, D, F) black with white spots located at depressions near lateral margins. Lateral angles and apical angle of scutellum black, central part yellow, and area near transverse depression black. + + +Forewing ( +Fig. 1 +: G) black, middle of claval area, with two slanted white stripes, and white patch at apex. Area along the R and M veins red. First apical cell large, occupying about half of apical area, third apical cell small with stalk. Abdominal apodemes ( +Fig. 1 +: I, N) extended to 4th abdominal sternite. Pygofer side ( +Fig. 1 +: J, O) covered with truncate setae, heavy sclerotization distributed along notches of ventral margin. + + +Subgenital plate ( +Fig. 1 +: J, O, P) long and upturned with scattered large setae along ventral margin, apex expanded laterad, with sclerotized horn-like protrusion at top of expanded section. Style ( +Fig. 1 +; L, M) longer than aedeagus, slightly sinuate. Connective ( +Fig. 1 +: L, M) shorter than wide. Aedeagus ( +Fig. 1 +: L, M, P) broad at base, atrial complex enclosing shaft, shaft short and tubular, curved, pair of curved short protrusions at base and a pair of long protrusions at the end; long protrusions slender and crossing each other, comprising about two-thirds of total length of aedeagus. + + + + +Type material. + + +Holotype + +, +1♂ +, +CHINA +: +Yunnan Province +, Lvchun. +102.4251°E +22.9432°N +, + +2075m + +, + +8.V.2015 + +, (light-trap), collected by +Wu Yunfei. + + + +Paratype + +, +5♂♂ +, same data as holotype + + + + + +Etymology. +The specific epithet is the Latin word + +obtusangula + +referring to the angulate profile of the face. + + + + +Remarks. +Unusual characteristics of this new species include the sclerotization distributed along the notches of the pygofer ventral margin and the aedeagus with well-developed processes but the shaft short and tubular. + + + + \ No newline at end of file diff --git a/data/03/B3/16/03B3161E234050204B9BFBC0FC7B7A92.xml b/data/03/B3/16/03B3161E234050204B9BFBC0FC7B7A92.xml new file mode 100644 index 00000000000..d2740d9ef7b --- /dev/null +++ b/data/03/B3/16/03B3161E234050204B9BFBC0FC7B7A92.xml @@ -0,0 +1,294 @@ + + + +Description of six new species forming the sumbana species group of the genus Nemophora Hoffmannsegg (Lepidoptera, Adelidae) from the Lesser Sunda Islands in Indonesia + + + +Author + +Kozlov, Mikhail V. + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +111 +122 + + + + +https://doi.org/10.11646/zootaxa.5543.1.6 + +journal article +10.11646/zootaxa.5543.1.6 +1175-5326 +F220768D-2EC2-4ECC-8DE8-7097F1BFA633 + + + + + + + +Nemophora sumbana +Kozlov + +, +sp. nov. + + + + + +urn:lsid:zoobank.org:act: +C3D8CCC3-EFFB-4E0B-A6A2-DDC74B34DED6 + + + + + +( +Figs. 1 +, +9 +, +15 +) + + + + + + +Holotype + +♂ +: +Indonesia +, +Sumba Island +, +Pogobina +( +9º 36’ S +, +119º 25’ E +); labelled: +8 mm +circle with red border, print + +‘ +Holo- +│ type’; 6 × +14 mm +, print ‘W. Sumba, +500 m +│ Pogobina, +ix 1949 +│ Sutter & Wegner’; 6 × +18 mm +, print ‘ + +HOLOTYPE +♂ +│ + +Nemophora +│ +sumbana + +Kozlov’ ( +RMNH +) [examined] + +. + +Paratypes + +. +4 ♂ +, labelled: +8 mm +circle with yellow border, print ‘ +Para- +│ type’; 6 × +14 mm +, print ‘W. Sumba, +500 m +│ Pogobina, +ix 1949 +│ Sutter & Wegner’. +2 ♂ +, labelled: 6 × +14 mm +, print ‘W. Sumba, +500 m +│ Pogobina, +ix 1949 +│ Sutter & Wegner’. +1 ♂ +, labelled: 9 × +13 mm +, print ‘[K. W.] Dammerman [leg.] │ O. Soemba │ +450 M. +│ Mao Marroe │ +V. 1925 +’. All +paratypes +bear the label: 6 × +18 mm +, print ‘ + +PARATYPE +♂ +│ + +Nemophora +│ +sumbana + +Kozlov’ (all in +RMNH +) [examined] + +. + + + + +Diagnosis. + +Nemophora sumbana + +differs from other species in the + +sumbana + +group by its large size, the extremely oblique external border of the basal field on the forewing (reaching the costa at 0.4 and the dorsum at 0.2 × FWL), the spot at the outer wing margin that does not reach the forewing apex, and relatively short labial palpi (0.3‒0.4 × vertical eye diameter). In male genitalia, + +N. sumbana + +is very similar to + +N. umbronitidella + +( +Figs. 3, 4 +), from which it differs by the smaller medial process of the transtilla, the shape of the anellus, and the relatively thick and straight tip of the phallus. + + + + +Description. +Male ( +Fig. 1 +). FWL +6.6‒7.5 mm +, WLR 0.35‒0.40. Vertex bright ochreous; frons glossy golden. PLB 0.3‒0.4 × vertical eye diameter, ochreous brown, with sparse raised ochreous brown piliform scales. Proboscis light brown, base anterolaterally with dense tuft of long and wide dark brown scales. Eyes enlarged, but not touching each other; interocular index 1.3‒1.5; occipital distance 0.05‒0.10. Antenna 3.0‒3.9 × FWL. Scape and basal 5‒8 flagellomeres ochreous brown, dorsally slightly thickened by ochreous scales; then colour of flagellum sharply changes to dark coppery brown, with irregular white marks on lateral sides of some flagellomeres; these marks never form complete rings. At about middle of flagellum its colour steadily changes to bronze, and apical 5‒15 flagellomeres are white in some specimens. Tegulae and thorax brown to bronze; tegulae with brassy green tint. Forewing ( +Fig. 9 +) bronze, with ochreous pattern. Basal field with three glossy bronze longitudinal stripes; costal silver stripe reaches 0.30–0.35 × FWL; external border of basal field oblique, reaching costa at 0.4 and dorsum at 0.2 × FWL. Oblique band next to basal field of about same width at costa and dorsum, of same ochreous colour as forewing base, except for narrow glossy bronze marginal lines. Oblique ochreous band linking costa with outer wing margin bordered by dotted narrow line of dark brown scales; its dorsal part extends along outer wing margin only towards wing base. Bronze background of distal part of forewing suffused with yellow scales, which form diffuse spot along external border of oblique ochreous band. Fringe bronze. Hindwing brown, with coppery tint; costal area yellowish grey; fringe brown. Legs glossy bronze to yellow; apices of tibiae and all tarsomeres brown. Epiphysis at 0.55, not reaching apex of tibia. Abdomen brown to bronze. + + + +FIGURES 1–8. +Adults of + +Nemophora +spp. 1 + +, + +N. sumbana +Kozlov + +, + +sp. nov. + +, male, holotype, from Pogobina, Sumba Island, Indonesia; 2, + +N. timorella +Kozlov + +, + +sp. nov. + +, male, paratype, from Timor Island, Indonesia; 3, + +N. umbronitidella +Kozlov + +, + +sp. nov. + +, male, paratype, from Sumbawa Island, Indonesia; 4, ditto, female, paratype, from the same locality; 5, + +N. wegneri +Kozlov + +, + +sp. nov. + +, male, holotype, from Mao Marroe, Sumba Island, Indonesia; 6, ditto, female, paratype, from Pogobina, Sumba Island, Indonesia; 7, + +N. longipeniculella +Kozlov + +, + +sp. nov. + +, female, paratype, from Bali Island, Indonesia; 8, + +N. brevipeniculella +Kozlov + +, + +sp. nov. + +, female, holotype, from Timor Island, Indonesia. Scale bar: 2 mm. + + +Female unknown. + +Male genitalia ( +Fig. 15 +). Tegumen dome-shaped, with small medial ridge. Socii 1.6 × medial diameter of phallus. Vinculum 2.8 × length of valva, with slightly convex lateral margins and nearly straight distal margin with small medial protuberance. Tips of valvae at about same level as tip of tegumen. Ventral margin of valva with small lobe at 0.5 × total length; dorsal valvar margin almost straight; tip of valva rounded. Valvae fused basally up to 0.3 × total length; internal valvar margins indistinct. Anellus 0.25 × length of valva. Transtilla with small triangular medial process. Juxta 0.55 × length of phallus, anchor-shaped; arrow head extremely wide (WLR 1.3), with rounded tip and pointed lateral arms. Phallus 0.90‒0.95 × length of vinculum, in lateral view gently S-shaped; base of phallus narrowly bell-shaped, tip pointed. + + + + +Distribution. +Indonesia +, Sumba Island. + + + + +Etymology. +The species is named after the +type +locality, Sumba Island. + + + + \ No newline at end of file diff --git a/data/03/B3/16/03B3161E2344502C4B9BF906FE1F7C1E.xml b/data/03/B3/16/03B3161E2344502C4B9BF906FE1F7C1E.xml new file mode 100644 index 00000000000..c30de82e2ca --- /dev/null +++ b/data/03/B3/16/03B3161E2344502C4B9BF906FE1F7C1E.xml @@ -0,0 +1,291 @@ + + + +Description of six new species forming the sumbana species group of the genus Nemophora Hoffmannsegg (Lepidoptera, Adelidae) from the Lesser Sunda Islands in Indonesia + + + +Author + +Kozlov, Mikhail V. + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +111 +122 + + + + +https://doi.org/10.11646/zootaxa.5543.1.6 + +journal article +10.11646/zootaxa.5543.1.6 +1175-5326 +F220768D-2EC2-4ECC-8DE8-7097F1BFA633 + + + + + + + +Nemophora wegneri +Kozlov + +, +sp. nov. + + + + + +urn:lsid:zoobank.org:act: +37C2CA18-18CE-45CF-A632-FF1258E1DEB6 + + + + + +( +Figs. 5, 6 +, +12 +, +18 +) + + + + + + +Holotype + +♂ +: +Indonesia +, +Sumba Island +, +Mao Marroe +(approx. +9º 58’ 30” S +, +120º 30’ 45” E +); labelled: +8 mm +circle with red border, print + +‘ +Holo- +│ type’; 9 × +13 mm +, print ‘[K. W.] Dammerman [leg.]│ O. Soemba │ +450 M. +│ Mao Marroe │ +V. 1925 +’; 6 × +18 mm +, print ‘ + +HOLOTYPE +♂ +│ + +Nemophora +│ +wegneri + +Kozlov’ ( +RMNH +) [examined] + +. + +Paratypes + +. +5 ♂ +2 ♀ +, labelled: +8 mm +circle with yellow border, print ‘ +Para- +│ type’; 6 × +14 mm +, print ‘W. Sumba, +500 m +│ Pogobina, +ix 1949 +│ Sutter & Wegner’. +6 ♂ +, labelled: 6 × +14 mm +, print ‘W. Sumba, +500 m +│ Pogobina, +ix 1949 +│ Sutter & Wegner’. +1 ♂ +, labelled: +8 mm +circle with yellow border, print ‘ +Para- +│ type’; 7 × +14 mm +, print ‘C. Sumba, +4‒500 m +│ Lindi Watju │ +27.ix‒15.x.1949 +│ Sutter & Wegner’. All +paratypes +bear the label: 6 × +18 mm +, print ‘ + +PARATYPE +♂ +[or +♀ +] │ + +Nemophora +│ +wegneri + +Kozlov’ (all in +RMNH +) [examined] + +. + + + + +Diagnosis +. + +Nemophora wegneri + +is most similar to + +N. umbronitidella + +( +Figs. 3, 4 +), from which it differs by the bright yellow posterior margin of the otherwise dark brown metathorax (in males only), shorter male antennae, dark brown scape, and a long phallus (1.1 × length of the vinculum) with a prominent ventral lobe near the apex. + + + + +FIGURES 15–18. +Male genitalia of + +Nemophora +spp. + +15, + +N. sumbana +Kozlov + +, + +sp. nov. + +; 16, + +N. timorella +Kozlov + +, + +sp. nov. + +; 17, + +N. umbronitidella +Kozlov + +, + +sp. nov. + +; 18, + +N. wegneri +Kozlov + +, + +sp. nov. + +; a: genital complex, ventral view (right valva not shown); b: genital complex, lateral view; c: juxta; d: phallus, ventral view; e: phallus, lateral view. Scale bar: 0.2 mm. + + + + +Description +. Male ( +Fig. 5 +). FWL +5.3‒5.9 mm +, WLR 0.36‒0.42. Vertex with dark brown to brown piliform scales along occipital margin and above antennal sockets, and pale yellow scales between compound eyes; frons glossy golden. PLB 0.45‒0.55 × vertical eye diameter, ochreous brown, with sparse raised dark brown scales. Proboscis light brown, base laterally with bronze to coppery bronze scales which, however, do not form a tuft. Eyes enlarged, occipitally closely approaching; interocular index 1.15‒1.35; occipital distance 0.05‒0.15.Antenna 2.9‒3.5 × FWL. Scape dark brown, ventrally in some specimens with a few yellow scales; basal 0.6‒0.7 of flagellum with alternating rings of dark brown and white scales; apical part of flagellum silver to white. Tegulae dark brown with bronze tint; thorax dark brown, except for bright yellow posterior margin of mesothorax. Forewing ( +Fig. 12 +) bright yellow with ochreous pattern. Basal field with two glossy golden longitudinal stripes; costal margin dark brown; external border oblique, reaching costa at 0.35 and dorsum at 0.25 × FWL. Oblique stripe next to basal field at costa narrower than at dorsum (0.13 vs. 0.22 × FWL, respectively), ochreous, clearly contrasting yellow colour of forewing base, except for narrow glossy bronze lines preceding dark brown margins. Transverse band and oblique band linking costa with outer wing margin bright yellow, bordered near wing margins by dotted narrow lines of dark brown scales and in medial zone of forewing by narrow glossy golden lines; yellow spot expands along outer wing margin up to wing apex. Bronze background of distal part of forewing suffused with ochreous scales forming compact spot along external border of oblique ochreous band. Fringe bronze, with few ochreous scales forming dotted marginal line. Hindwing brown, with coppery tint; costal area grey; fringe brown. Legs glossy bronze to yellow; apices of tibiae and all tarsomeres brown. Epiphysis at 0.5, reaching apex of tibia. Abdomen brown to bronze. + + +Female ( +Fig. 6 +). FWL 5.0‒6.0 mm. Antenna 1.5‒1.6 × FWL, basal 0.30‒0.35 of flagellum dorsally with dense cover of raised scales forming very thick brush, which is ochreous yellow except for tip where its colour turns to dark brown; distal part of flagellum dark bronze, with irregular markings of white scales. Tegulae and thorax bright yellow. Otherwise similar to male. + + +Male genitalia ( +Fig. 18 +). Tegumen dome-shaped, with small medial ridge. Socii oval, 1.2 × medial diameter of phallus. Vinculum 2.6‒2.7 × length of valva, with slightly convex lateral margins and gently W-shaped distal margin. Tips of valvae at about same level as tip of tegumen. Ventral margin of valva with small lobe at 0.6 × total length; dorsal valvar margin almost straight; tip of valva pointed. Valvae fused basally up to 0.3 × total length and also fused to vinculum; internal valvar margins indistinct. Anellus 0.3 × length of valva. Transtilla with wide triangular medial process. Juxta 0.55 × length of phallus; arrow head triangular, extremely wide (WLR 1.3), with pointed tip and pointed lateral arms. Phallus 1.1 × length of vinculum, in lateral view gently S-shaped; tip pointed, with prominent ventral lobe; base of phallus narrowly bell-shaped. + + + + +Distribution +. +Indonesia +, Sumba Island. + + + + +Etymology +. The species is named in honour of A. M. R. Wegner, a former director of the Zoological Museum in Bogor, +Indonesia +, a renowned insect collector, and a well-known specialist on the fauna of the Tengger Mountains, who collected most of the specimens of this species. + + + + +Comments. +The coordinates of the +type +locality (Mao Marroe) were identified using a map published by +Dammerman (1926) +. + + + + \ No newline at end of file diff --git a/data/03/B3/16/03B3161E234650214B9BFC97FD8A7AAE.xml b/data/03/B3/16/03B3161E234650214B9BFC97FD8A7AAE.xml new file mode 100644 index 00000000000..e39a66983e2 --- /dev/null +++ b/data/03/B3/16/03B3161E234650214B9BFC97FD8A7AAE.xml @@ -0,0 +1,238 @@ + + + +Description of six new species forming the sumbana species group of the genus Nemophora Hoffmannsegg (Lepidoptera, Adelidae) from the Lesser Sunda Islands in Indonesia + + + +Author + +Kozlov, Mikhail V. + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +111 +122 + + + + +https://doi.org/10.11646/zootaxa.5543.1.6 + +journal article +10.11646/zootaxa.5543.1.6 +1175-5326 +F220768D-2EC2-4ECC-8DE8-7097F1BFA633 + + + + + + + +Nemophora timorella +Kozlov + +, +sp. nov. + + + + + +urn:lsid:zoobank.org:act: +2E572409-7357-4135-B22D-032FDC2E7113 + + + + + +( +Figs. 2 +, +10 +, +16 +, +19 +) + + + + + + +Holotype + +♂ +: +Indonesia +, +Timor Island +(approx. +9º S +, +124º E +); labelled: +8 mm +circle with red border, print + +‘ + +Holo- +│ type’; 9 × +12 mm +, print + black ink ‘S. W. TIMUR [= +Timor Island +] │ + +1500‒3000 ft. + +│ XI.‒XII. │ Doherty, 1891 │ No. 41708’; 8 × +10 mm +, print ‘Walsingham │ Collection │ 1910‒427’; 9 × +16 mm +, print ‘B. M. │ Genitalia slide │ No. 29981’; 6 × +18 mm +, print + +‘ + +HOLOTYPE +♂ +│ + +Nemophora +│ +timorella + +Kozlov’ ( +NHM +) [examined] + +. + +Paratypes + +. +15 ♂ +, labelled: +8 mm +circle with yellow border, print ‘ +Para- +│ type’; 9 × +12 mm +, print + black ink ‘S. W. TIMUR │ +1500‒3000 ft. +│ XI.‒XII. │ Doherty, 1891 │ No. 417**’ [** = 03, 04, 06, 07, 09‒16, 18, 19; specimen no. 41718 has additional label: 11 × +32 mm +, pencil ‘timurensis’]; 8 × +10 mm +, print ‘Walsingham │ Collection │ 1910‒427’; 6 × +18 mm +, print ‘ + +PARATYPE +♂ +│ + +Nemophora +│ +timorella + +Kozlov’ (all in +NHM +) [examined] + +. + + + + +Diagnosis +. + +Nemophora timorella + +is most similar to + +N. umbronitidella + +( +Figs. 3, 4 +), from which differs by its smaller size (FWL 4.3‒5.0 mm), the pale yellowish brown basal field of the forewing, the nearly perpendicular external border of the basal field relative to the wing margins, a longer vinculum, the tips of the valvae slightly extending beyond the tip of the tegumen, a medial indentation in the fused basal parts of the valvae, and the triangular shape of the arrow head of the juxta. It differs from + +N. longipeniculella + +( +Fig. 7 +) and + +N. brevipeniculella + +( +Fig. 8 +) by the presence of two longitudinal silver stripes with brilliant iridescence in the dark ochreous brown basal field of the forewing. + + + + +Description +. Male ( +Fig. 2 +). FWL 4.3‒5.0 mm, WLR 0.37‒0.39. Vertex light brown; frons with iridescent silver to light golden scales, and with row of pale ochreous brown piliform scales below antennal sockets. PLB 0.5‒0.6 × vertical eye diameter (1.3‒1.6 × length of scape), brown. Proboscis brown, base covered with dark coppery brown semi-erect elongated scales, forming small tuft. Eyes enlarged, occipitally closely approaching, with rounded dorsal margins; interocular index 1.3‒1.6, occipital distance 0.05‒0.07. Antenna 3.4‒3.9 × FWL. Scape ochreous brown, flagellum bronze, basal 20‒30 flagellomeres with rings of white scales, which gradually disappear towards apex. Tegulae and thorax light ochreous brown, with slight coppery tint on external margins of tegulae. Forewing ( +Fig. 10 +) bronze, with pale yellow basal field reaching costa at 0.35 and dorsum at 0.40 × FWL, ochreous yellow medial band, and oblique yellow band linking costa with outer wing margin. Basal field with two longitudinal glossy silver stripes with slight brilliant iridescence; lower stripe very narrow.Yellow medial band with sinuate margins, medially suffused with ochreous scales; on both sides bordered by narrow lines of dark brown scales. Bronze background of distal part of forewing suffused with ochreous scales, except for costal area near apex that is entirely bronze. Oblique yellow band, linking costa with outer wing margin, bordered by narrow line of dark brown scales; at outer wing margin it expands between veins M1 and CuA1 and is suffused with dark brown scales. Some of dark brown scales all over forewing show brilliant iridescence. Fringe bronze. Hindwing brown with coppery iridescence; costal area grey; anal field semitranslucent; fringe brown to light brown. Legs bronze; bases of all tarsomeres light yellow. Epiphysis at 0.6, reaching apex of tibia. Abdomen coppery brown dorsally, brown with bronze iridescence ventrally. + +Female unknown. + +Male genitalia ( +Figs. 16 +, +19 +). Tegumen onion-shaped. Socii 1.6‒1.8 × medial diameter of phallus. Vinculum 2.7 × length of valva, with straight lateral margins and slightly convex distal margin. Tips of valvae slightly extend beyond tip of tegumen. Ventral margin of valva medially with small lobe at 0.4 × total length; dorsal margin straight; tip of valva narrowly rounded. Valvae fused basally up to 0.2 × total length; internal margins not visible. Fused parts of valvae between ventral margins with distinct medial indentation. Anellus 0.35 × length of valva. Transtilla with wide triangular medial process. Juxta 0.6 × length of phallus; arrow head triangular, extremely wide (WLR 1.3), with pointed tip and short pointed lateral arms. Phallus 0.9 × length of vinculum, in lateral view shallowly C-shaped; tip narrow, hook-shaped; base narrowly funnel-shaped. + + + + +Distribution +. +Indonesia +, Timor Island. + + + + +Etymology +. The species is named after the +type +locality, Timor Island. + + + + +Comments. +Although Walsingham clearly labelled +one male +specimen as the +type +of a proposed new species, he never published its description. + + + + \ No newline at end of file diff --git a/data/03/B3/16/03B3161E234750224B9BF967FD8A7EE2.xml b/data/03/B3/16/03B3161E234750224B9BF967FD8A7EE2.xml new file mode 100644 index 00000000000..57f3e25f5e5 --- /dev/null +++ b/data/03/B3/16/03B3161E234750224B9BF967FD8A7EE2.xml @@ -0,0 +1,282 @@ + + + +Description of six new species forming the sumbana species group of the genus Nemophora Hoffmannsegg (Lepidoptera, Adelidae) from the Lesser Sunda Islands in Indonesia + + + +Author + +Kozlov, Mikhail V. + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +111 +122 + + + + +https://doi.org/10.11646/zootaxa.5543.1.6 + +journal article +10.11646/zootaxa.5543.1.6 +1175-5326 +F220768D-2EC2-4ECC-8DE8-7097F1BFA633 + + + + + + + +Nemophora umbronitidella +Kozlov + +, +sp. nov. + + + + + +urn:lsid:zoobank.org:act: +9FAB7629-54B3-4DA0-AEA5-ED6B83500DB4 + + + + + +( +Figs. 3, 4 +, +11 +, +17 +, +20 +) + + + + + + +Holotype + +♂ +: +Indonesia +, +Sumbawa Island +(approx. +8º 20’ S +, +118º E +); labelled: +8 mm +circle with red border, print + +‘ + +Holo- +│ type’; 9 × +13 mm +, print + black ink ‘SAMBAWA [=Sumbawa Island], │ + +3000‒5500 ft. + +, │ (virgin forest), │ +Doherty +, 1891. │ +No +40729’; 8 × +10 mm +, print ‘ +Walsingham +│ +Collection +│ 1910‒427’; 7 × +17 mm +, black frame, black ink ‘ +Typ +│ umbreonitida +♂ +│ W │ Named by Wlsm. ’; 9 × +16 mm +, print ‘B. M. │ Genitalia slide │ No. 29492’; 6 × +18 mm +, print + +‘ + +HOLOTYPE +♂ +│ + +Nemophora +│ +umbronitidella + +Kozlov’ ( +NHM +) [examined] + +. + +Paratypes + +. +2 ♂ +, labelled: +8 mm +circle with yellow border, print ‘ +Para- +│ type’; 9 × +13 mm +, print + black ink ‘SAMBAWA, │ +3000‒5500 ft. +, │ (virgin forest), │ Doherty, 1891. │ No 40730 [or 40731]’; 8 × +10 mm +, print ‘Walsingham │ Collection │ 1910‒427.’; 6 × +18 mm +, print ‘ +PARATYPE +♂ +│ + +Nemophora +│ +umbronitidella + +Kozlov’. +1 ♀ +, labelled: +8 mm +circle with yellow border, print ‘ +Para- +│ type’; 9 × +13 mm +, print + black ink ‘SAMBAWA, │ +3000‒5500 ft. +, │ (virgin forest), │ Doherty, 1891. │ No 40732’; 8 × +10 mm +, print ‘Walsingham │ Collection │ 1910‒427.’; 7 × +17 mm +, black frame, black ink ‘Typ │ umbreonitida +♀ +│ W. │ Named by Wlsm.’; 6 × +18 mm +, print ‘ + +PARATYPE +♀ +│ + +Nemophora +│ +umbronitidella + +Kozlov’ (all in +NHM +) [examined] + +. + + + + +Diagnosis. + +Nemophora umbronitidella + +is most similar to + +N. timorella + +, from which it differs by larger size (FWL +5.2‒5.7 mm +), an oblique external border of the basal field of forewing, a shorter vinculum, the tip of the tegumen slightly extending beyond the tips of the valvae, a medial protuberance in the fused basal parts of the valvae and an anchor-shaped juxta. It differs from + +N. longipeniculella + +( +Fig. 7 +) and + +N. brevipeniculella + +( +Fig. 8 +) by the presence of two longitudinal silver stripes with brilliant iridescence in the dark ochreous brown basal field of the forewing. + + + + +Description +. Male ( +Fig. 3 +). FWL +5.2‒5.7 mm +, WLR 0.35‒0.37. Vertex pale yellow above antennal sockets to ochreous brown along occipital margin; frons with iridescent silver to light golden scales through which brown colour of head capsule may be visible, and with row of pale ochreous yellow piliform scales below antennal sockets. PLB 0.5 × vertical eye diameter (0.8‒0.9 × length of scape), light brown. Proboscis brown, base densely covered with dark coppery brown semi-erect elongated scales, forming characteristic tuft. Eyes enlarged, occipitally closely approaching, with rounded dorsal margins; interocular index 1.3‒1.4, occipital distance 0.05‒0.07. Antenna 3.7‒4.1 × FWL. Scape yellow, flagellum coppery brown, basal 27‒32 flagellomeres with rings of white scales, which gradually disappear towards apex. Tegulae and thorax dark ochreous brown. Forewing ( +Fig. 11 +) bronze, with dark ochreous brown basal field reaching costa at 0.35 and dorsum at 0.30 × FWL, ochreous yellow medial band, and oblique yellow band linking costa with outer wing margin. Basal field with two longitudinal glossy silver stripes with slight brilliant iridescence; lower stripe very narrow. Ochreous yellow medial band with sinuate margins, on both sides bordered by narrow lines of dark brown scales. Bronze background of distal part of forewing suffused with yellow scales. Oblique yellow band linking costa with outer wing margin bordered by narrow line of dark brown scales; at outer margin it expands between veins RS4 and CuA2 and is suffused with brown scales. Some of dark brown scales all over forewing show brilliant iridescence. Fringe bronze. Hindwing brown with coppery iridescence; costal area grey; anal field semitranslucent; fringe brown to light brown. Legs bronze; bases of all tarsomeres white to light yellow. Epiphysis at 0.65, reaching apex of tibia. Abdomen brown with bronze iridescence. + + +Female ( +Fig. 4 +). FWL +5.6 mm +. Base of antenna thickened by appressed dark brown scales (distal part of antenna missing). Otherwise similar to male. + + +Male genitalia ( +Figs. 17 +, +20 +). Tegumen onion-shaped. Socii 1.3 × medial diameter of phallus. Vinculum 2.2 × length of valva, with straight lateral margins and slightly convex distal margin. Tip of tegumen slightly extends beyond tips of valvae. Ventral margin of valva with small lobe at 0.55 × total length; dorsal margin straight; tip of valva narrowly rounded. Valvae fused basally up to 0.4 × total length; internal margins not visible. Fused parts of valvae between ventral margins with distinct triangular protuberance. Anellus 0.2 × length of valva. Transtilla with wide triangular medial process. Juxta 0.55 × length of phallus; arrow head anchor-shaped, extremely wide (WLR 1.1), with widely rounded tip and long pointed lateral arms. Phallus 0.95 × length of vinculum, in lateral view shallowly C-shaped; tip narrow, base widely funnel-shaped. + + + + +Distribution +. +Indonesia +, Sumbawa Island. + + + + +Etymology +. The specific epithet is derived from umbro (Latin: shade) and nitidus (Latin: bright, shining) and refers to the diagnostic trait of this species. + + + + +Comments +. Although Walsingham clearly labelled +one male +and +one female +as +types +of a proposed new species, he never published its description. + + + + \ No newline at end of file diff --git a/data/03/B3/16/03B3161E234A502D4B9BFB1BFD887996.xml b/data/03/B3/16/03B3161E234A502D4B9BFB1BFD887996.xml new file mode 100644 index 00000000000..554ee3f5cfb --- /dev/null +++ b/data/03/B3/16/03B3161E234A502D4B9BFB1BFD887996.xml @@ -0,0 +1,224 @@ + + + +Description of six new species forming the sumbana species group of the genus Nemophora Hoffmannsegg (Lepidoptera, Adelidae) from the Lesser Sunda Islands in Indonesia + + + +Author + +Kozlov, Mikhail V. + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +111 +122 + + + + +https://doi.org/10.11646/zootaxa.5543.1.6 + +journal article +10.11646/zootaxa.5543.1.6 +1175-5326 +F220768D-2EC2-4ECC-8DE8-7097F1BFA633 + + + + + + + +Nemophora longipeniculella +Kozlov + +, +sp. nov. + + + + + +urn:lsid:zoobank.org:act: +6D766E8A-4D2B-4D65-8D20-3C542866281C + + + + + +( +Figs. 7 +, +13 +) + + + + + + +Holotype + +♀ +: +Indonesia +, +Bali +Island +(approx. +8º 20’ S +, +115º E +); labelled: +8 mm +circle with red border, print + +‘ + +Holo- +│ type’; 7 × +15 mm +, print ‘ +BALI +│ Doherty. 1896’; 8 × +10 mm +, print ‘ +Walsingham +│ +Collection +│ 1910‒427’; 6 × +18 mm +, print + +‘ + +HOLOTYPE +♀ +│ + +Nemophora +longi- + +│ +peniculella +Kozlov’ ( +NHM +) [examined] + +. + +Paratype + +. +1 ♀ +, labelled: +8 mm +circle with yellow border, print ‘ + +Para- +│ type’; 7 × +15 mm +, print ‘ +BALI +│ Doherty. 1896’; 8 × +10 mm +, print ‘ +Walsingham +│ +Collection +│ 1910‒427.’; 6 × +18 mm +, print + +‘ + +PARATYPE +♀ +│ + +Nemophora +longi- + +│ +peniculella +Kozlov’ ( +NHM +) [examined] + +. + + + + +Diagnosis +. + +Nemophora longipeniculella + +is most similar to + +N. brevipeniculella + +( +Fig. 8 +), from which it differs by the longer (almost equal to the forewing length) part of the female antenna thickened with dark brown scales, the ochreous yellow scales suffusing the bronze band adjacent to the forewing base, the ochreous scales at the forewing apex, and the unusually long epiphysis articulated at 0.25 of the total length of tibia and reaching its apex. It differs from females of + +N. umbronitidella + +( +Fig. 4 +) by the yellow forewing base (females of + +N. timorella + +are unknown). + + + + +Description +. Male unknown. + + +Female ( +Fig. 7 +). FWL +4.9‒5.1 mm +, WLR 0.35. Vertex and frons brown. PLB 1.0‒1.1 × vertical eye diameter (1.0‒1.1 × length of scape), brown. Proboscis brown, base covered with dark coppery brown semi-erect elongated scales forming small tuft.Antenna>1.3 × FWL. Scape and basal part of flagellum dark coppery brown, dorsolaterally thickened by semi-erect dark brown scales; brush formed by these scales reaches 0.95 × FWL; flagellum grey beyond this brush. Tegulae and thorax ochreous yellow to yellow. Forewing ( +Fig. 13 +) ochreous brown, with pale yellow basal field expanding to 0.4 × FWL at both costa and dorsum, pale yellow medial band, and oblique yellow band linking costa with outer wing margin. Basal spot reduced to dark line along costa; forewing base separated from bronze background by interrupted line of coppery brown scales. Pale yellow medial band with straight internal margin and sinuate external margin, on both sides bordered by interrupted lines of coppery brown scales. Oblique yellow band linking costa with outer wing margin bordered by sparse coppery brown scales; this band expands into wide spot along outer margin. Costal part of forewing bronze near apex. Fringe brown. Hindwing dark brown with coppery iridescence; costal area yellowish grey; fringe brown to light brown. Legs coppery brown to bronze; bases of all tarsomeres light yellow. Epiphysis at 0.25, unusually long, reaching apex of tibia. Abdomen dorsally ochreous brown, ventrally bronze at base to coppery brown at apex. + + + + +Distribution. +Indonesia +, +Bali +Island. + + + + +Etymology +. The specific epithet is derived from longus (Latin: long) and peniculus (Latin: brush) and refers to the diagnostic trait of this species. + + + + \ No newline at end of file diff --git a/data/03/B3/16/03B3161E234B502E4B9BFD93FECE7D52.xml b/data/03/B3/16/03B3161E234B502E4B9BFD93FECE7D52.xml new file mode 100644 index 00000000000..76ebc930108 --- /dev/null +++ b/data/03/B3/16/03B3161E234B502E4B9BFD93FECE7D52.xml @@ -0,0 +1,298 @@ + + + +Description of six new species forming the sumbana species group of the genus Nemophora Hoffmannsegg (Lepidoptera, Adelidae) from the Lesser Sunda Islands in Indonesia + + + +Author + +Kozlov, Mikhail V. + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +111 +122 + + + + +https://doi.org/10.11646/zootaxa.5543.1.6 + +journal article +10.11646/zootaxa.5543.1.6 +1175-5326 +F220768D-2EC2-4ECC-8DE8-7097F1BFA633 + + + + + + + +Nemophora brevipeniculella +Kozlov + +, +sp. nov. + + + + + +urn:lsid:zoobank.org:act: +CDB9D2FC-5604-4372-95E0-9D0245FCD4F3 + + + + + +( +Figs. 8 +, +14 +) + + + + + + +Holotype + +♀ +: +Indonesia +, +Timor Island +(approx. +9º S +, +124º E +); labelled: +8 mm +circle with red border, print + +‘ + +Holo- +│ type’; 9 × +12 mm +, print + black ink ‘S. W. TIMUR [=Timor Island] │ + +1500‒3000 ft. + +│ XI.‒XII. │ +Doherty +, 1891 │ No. 41721’; 8 × +10 mm +, print ‘ +Walsingham +│ +Collection +│ 1910‒427.’; 9 × +22 mm +, black ink ‘didesma │ +♀ +type’; 6 × +18 mm +, print + +‘ + +HOLOTYPE +♀ +│ + +Nemophora +brevi- + +│ +peniculella +Kozlov’ ( +NHM +) [examined] + +. + +Paratypes + +. +2 ♀ +, labelled: +8 mm +circle with yellow border, print ‘ +Para- +│ type’; 9 × +12 mm +, print + black ink ‘S. W. TIMUR │ +1500‒3000 ft. +│ XI.‒XII. │ Doherty, 1891 │ No. 41717 or 41720]’; 8 × +10 mm +, print ‘Walsingham │ Collection │ 1910‒427.’; 6 × +18 mm +, print ‘ + +PARATYPE +♀ +│ + +Nemophora +brevi- + +│ +peniculella +Kozlov’ (both in +NHM +) [examined] + +. + + + + +FIGURES 19–20. +Male genitalia of + +Nemophora +spp. + +19, + +N. timorella +Kozlov + +, + +sp. nov. + +, holotype, from Timor Island, Indonesia, genitalia preparation 29981 (NHM); 20, + +N. umbronitidella +Kozlov + +, + +sp. nov. + +, holotype, from Sumbawa Island, Indonesia, genitalia preparation 29492 (NHM); a: genital complex, ventral view; b: phallus; c: apex of phallus; d: juxta. Scale bar: 0.2 mm (valid for a, b and d only). + + + + +Diagnosis +. + +Nemophora brevipeniculella + +is most similar to + +N. longipeniculella + +, from which it differs by the shorter (0.8 × forewing length) part of the female antenna thickened with dark brown scales, the dark brown scales densely suffusing the bronze band adjacent to the forewing base, the brown scales at the forewing apex, and the typical size of the epiphysis, which is articulated at 0.5 of the total length of tibia and does not reach its apex. It differs from females of + +N. umbronitidella + +( +Fig. 4 +) by the yellow forewing base (females of + +N. timorella + +are unknown). + + + + +Description +. Male unknown. + + +Female ( +Fig. 8 +). FWL 4.7‒5.0 mm, WLR 0.33‒0.38. Vertex brown; frons with iridescent silver to light golden scales, and with row of brown piliform scales below antennal sockets. PLB 0.9‒1.0 × vertical eye diameter (1.0 × length of scape), brown. Proboscis brown, base covered with dark coppery brown semi-erect elongated scales, forming small tuft. Antenna 2.0 × FWL. Scape and basal part of flagellum dark brown, dorsolaterally thickened by semi-erect dark brown scales; brush formed by these scales reaches 0.8 × FWL. Flagellum bronze beyond this brush; 10‒15 flagellomeres next to brush with white rings. Tegulae and thorax yellow. Forewing ( +Fig. 14 +) bronze, with pale yellow basal field expanding to 0.3 × FWL at both costa and dorsum, pale yellow medial band, and oblique yellow band linking costa with outer wing margin. Basal spot dark brown, connected with dark brown band separating yellow forewing base from bronze background of distal part of forewing. Pale yellow medial band almost straight, on both sides bordered by dark brown bands. Bronze field between basal field and fascia medially densely suffused with dark brown scales. Oblique yellow band linking costa with outer wing margin bordered by dark brown scales; irregular spot along outer wing margin dark brown, sparsely suffused with ochreous brown scales. Costal part of forewing bronze near apex. Fringe brown. Hindwing dark brown with coppery iridescence; costal area light yellow; fringe brown to light brown. Legs coppery brown to bronze; bases of all tarsomeres light yellow. Epiphysis at 0.5, not reaching apex of tibia. Abdomen dorsally brown with coppery iridescence, ventrally bronze except terminal sternite which is coppery brown; lateral parts of tergites with yellow scales. + + + + +Distribution. +Indonesia +, Timor Island. + + + + +Etymology +. The specific epithet is derived from brevis (Latin: short) and peniculus (Latin: brush) and refers to the diagnostic trait of this species. + + + + +Comments +. Although Walsingham clearly labelled the +holotype +of a proposed new species, he never published its description. + + +The females of + +N. brevipeniculella + +were collected from the same locality as the males of + +N. timorella + +. Although the males of + +N. brevipeniculella + +and the females of + +N. timorella + +remain unknown, I have opted to describe + +N. brevipeniculella + +as a separate species rather than consider these specimens as females of + +N. timorella + +. This decision is primarily based on the lack of sexual dimorphism in wing coloration, as well as in the position and length of the epiphysis in both + +N. umbronitidella + +and + +N. wegneri + +, species closely related to both + +N. timorella + +and + +N. brevipeniculella + +. + + + + \ No newline at end of file diff --git a/data/61/25/87/612587B8C96AD659FE14FCEBFB1E644E.xml b/data/61/25/87/612587B8C96AD659FE14FCEBFB1E644E.xml index dea4c749ac5..755cba48bb4 100644 --- a/data/61/25/87/612587B8C96AD659FE14FCEBFB1E644E.xml +++ b/data/61/25/87/612587B8C96AD659FE14FCEBFB1E644E.xml @@ -1,71 +1,91 @@ - - - -New species and new records of terrestrial isopods (Crustacea, Isopoda, Oniscidea) of the families Philosciidae and Scleropactidae from Brazilian caves + + + +New species and new records of terrestrial isopods (Crustacea, Isopoda, Oniscidea) of the families Philosciidae and Scleropactidae from Brazilian caves - - -Author + + +Author -Campos-Filho, Ivanklin Soares +Campos-Filho, Ivanklin Soares +C752F864-3C84-4AF4-9EDA-4D1AF464D615 +Universidade Federal de Campina Grande, Programa de Pós-Graduação em Engenharia e Gestão de Recursos Naturais, Av. Aprígio Veloso, 882, Bairro Universitário, 58429 - 140 Campina Grande, Paraíba, Brazil. +ivanklin.filho@gmail.com - - -Author + + +Author -Fernandes, Camile Sorbo +Fernandes, Camile Sorbo +C8246067-8235-4981-87D8-56FD9C43FDC2 +Universidade Federal de São Carlos, Departamento de Ecologia e Biologia Evolutiva, Rodovia Washington Luis, Km 235, 13565 - 905 São Carlos, São Paulo, Brazil. +camilesorbofernandes@yahoo.com.br - - -Author + + +Author -Cardoso, Giovanna Monticelli +Cardoso, Giovanna Monticelli +C829E5B7-B87E-4C4A-B88E-D5FE377AE60B +Universidade Federal do Rio Grande do Sul, Programa de Pós-Graduação em Biologia Animal, Departamento de Zoologia, Laboratório de Carcinologia, Av. Bento Gonçalves, 9500, Agronomia, 91510 - 979 Porto Alegre, Rio Grande do Sul, Brazil. +jojomonticelli@hotmail.com - - -Author + + +Author -Bichuette, Maria Elina +Bichuette, Maria Elina +7B740115-BCA0-4711-8C9D-CD014FD3122B +Universidade Federal de São Carlos, Departamento de Ecologia e Biologia Evolutiva, Rodovia Washington Luis, Km 235, 13565 - 905 São Carlos, São Paulo, Brazil. +lina.cave@gmail.com - - -Author + + +Author -Aguiar, José Otávio +Aguiar, José Otávio +8D91D781-EF49-42CC-B7AE-12789CB1CD71 +Universidade Federal de Campina Grande, Programa de Pós-Graduação em Engenharia e Gestão de Recursos Naturais, Av. Aprígio Veloso, 882, Bairro Universitário, 58429 - 140 Campina Grande, Paraíba, Brazil. +otavio.j.aguiar@gmail.com - - -Author + + +Author -Taiti, Stefano +Taiti, Stefano +62E97059-6AE5-4984-9ABB-7FB6F7358BD6 +Istituto di Ricerca sugli Ecosistemi Terrestri, Consiglio Nazionale delle Ricerche, Via Madonna del Piano 10, 50019 Sesto Fiorentino (Florence), Italy. & Museo di Storia Naturale, Sezione di Zoologia “ La Specola ”, Via Romana 17, 50125 Florence, Italy. +stefano.taiti@cnr.it -text - - -European Journal of Taxonomy +text + + +European Journal of Taxonomy - -2020 - -2020-02-20 + +2020 + +2020-02-20 - -606 + +606 - -1 -38 + +1 +38 -journal article -10.5852/ejt.2020.606 -e75c913b-d6ce-48b6-b5c4-dd09034b4547 -3676984 -95D497A6-2022-406A-989A-2DA7F04223B0 +journal article +23893 +10.5852/ejt.2020.606 +e75c913b-d6ce-48b6-b5c4-dd09034b4547 +2118-9773 +3676984 +95D497A6-2022-406A-989A-2DA7F04223B0 - - + + @@ -75,6 +95,8 @@ Campos-Filho, Cardoso & Taiti sp. nov. + + urn:lsid:zoobank.org:act: @@ -109,15 +131,13 @@ urn:lsid:zoobank.org:act: Material examined + - Holotype - - BRAZIL – @@ -149,191 +169,130 @@ urn:lsid:zoobank.org:act: leg.; MZUSP 39670 . - - + + + - Paratypes - - - BRAZIL – - -Pará -State - -, Parauapebas +Pará State +, +Parauapebas • 6 ♀♀ -, 2 juvs; -Gruta -N4E77 -Cave -; +, +2 juvs +; Gruta N4E77 Cave; 6°01′57″ S , 50°09′02″ W ; + 13– 30 Jan. 2010 + ; -R -. -Andrade leg -.; - -MZUSP -39671 • -1 ♀ -; -Gruta -N4E78 -Cave -; +R. Andrade +leg.; + + +MZUSP 39671 +• +1 ♀ +; Gruta N4E78 Cave; 6°01′57″ S , 50°09′04″ W ; - - -19 Nov. -– -4 - -Dec. - -2010 +19 Nov.–4 Dec. 2010 +; -; - -MZUSP -39672 • -1 ♀ -; -Gruta -S11D-79 -Cave -; + +MZUSP 39672 +• +1 ♀ +; Gruta S11D-79 Cave; 6°23′32″ S , 50°18′57″ W ; -1–14 Jul. 2010 + +1–14 Jul. 2010 + ; leg. -R -. -Andrade leg -. +R. Andrade +leg. ; - -MZUSP -39673 • -1 ♀ -; -Gruta -S11D-33 -Cave -; + +MZUSP 39673 +• +1 ♀ +; Gruta S11D-33 Cave; 6°24′39″ S , 50°20′37″ W ; - -13– -30 Jan. - -2010 +13–30 Jan. 2010 ; -R -. -Andrade leg -. +R. Andrade +leg. ; - -MZUSP -39674 • -1 ♀ -; -Gruta -S11D-37 -Cave -; + +MZUSP 39674 +• +1 ♀ +; Gruta S11D-37 Cave; 6°24′46″ S , 50°21′30″ W ; - -3– - -19 -Aug. - - -2010 +3–19 Aug. 2010 ; -R -. -Andrade leg -. +R. Andrade +leg. ; - -MZUSP -39675 – + +MZUSP 39675 +– Pará State -, Canaãdos Carajás +, +Canaãdos Carajás • - -1 -♀ - -; -Gruta -S11- 23 -Cave -; +1 ♀ +; Gruta S11- 23 Cave; 6°25′21″ S , 50°17′57″ W ; - - -24 Feb. -– -4 - -Mar. - -2010 +24 Feb.–4 Mar. 2010 ; -R -. -Andrade leg -.; -MZUSP -39676. +R. Andrade +leg.; +MZUSP 39676 +. diff --git a/data/61/25/87/612587B8C970D647FDCFFC96FA996663.xml b/data/61/25/87/612587B8C970D647FDCFFC96FA996663.xml new file mode 100644 index 00000000000..e87aea10fd7 --- /dev/null +++ b/data/61/25/87/612587B8C970D647FDCFFC96FA996663.xml @@ -0,0 +1,188 @@ + + + +New species and new records of terrestrial isopods (Crustacea, Isopoda, Oniscidea) of the families Philosciidae and Scleropactidae from Brazilian caves + + + +Author + +Campos-Filho, Ivanklin Soares +C752F864-3C84-4AF4-9EDA-4D1AF464D615 +Universidade Federal de Campina Grande, Programa de Pós-Graduação em Engenharia e Gestão de Recursos Naturais, Av. Aprígio Veloso, 882, Bairro Universitário, 58429 - 140 Campina Grande, Paraíba, Brazil. +ivanklin.filho@gmail.com + + + +Author + +Fernandes, Camile Sorbo +C8246067-8235-4981-87D8-56FD9C43FDC2 +Universidade Federal de São Carlos, Departamento de Ecologia e Biologia Evolutiva, Rodovia Washington Luis, Km 235, 13565 - 905 São Carlos, São Paulo, Brazil. +camilesorbofernandes@yahoo.com.br + + + +Author + +Cardoso, Giovanna Monticelli +C829E5B7-B87E-4C4A-B88E-D5FE377AE60B +Universidade Federal do Rio Grande do Sul, Programa de Pós-Graduação em Biologia Animal, Departamento de Zoologia, Laboratório de Carcinologia, Av. Bento Gonçalves, 9500, Agronomia, 91510 - 979 Porto Alegre, Rio Grande do Sul, Brazil. +jojomonticelli@hotmail.com + + + +Author + +Bichuette, Maria Elina +7B740115-BCA0-4711-8C9D-CD014FD3122B +Universidade Federal de São Carlos, Departamento de Ecologia e Biologia Evolutiva, Rodovia Washington Luis, Km 235, 13565 - 905 São Carlos, São Paulo, Brazil. +lina.cave@gmail.com + + + +Author + +Aguiar, José Otávio +8D91D781-EF49-42CC-B7AE-12789CB1CD71 +Universidade Federal de Campina Grande, Programa de Pós-Graduação em Engenharia e Gestão de Recursos Naturais, Av. Aprígio Veloso, 882, Bairro Universitário, 58429 - 140 Campina Grande, Paraíba, Brazil. +otavio.j.aguiar@gmail.com + + + +Author + +Taiti, Stefano +62E97059-6AE5-4984-9ABB-7FB6F7358BD6 +Istituto di Ricerca sugli Ecosistemi Terrestri, Consiglio Nazionale delle Ricerche, Via Madonna del Piano 10, 50019 Sesto Fiorentino (Florence), Italy. & Museo di Storia Naturale, Sezione di Zoologia “ La Specola ”, Via Romana 17, 50125 Florence, Italy. +stefano.taiti@cnr.it + +text + + +European Journal of Taxonomy + + +2020 + +2020-02-20 + + +606 + + +1 +38 + + + +journal article +23893 +10.5852/ejt.2020.606 +e75c913b-d6ce-48b6-b5c4-dd09034b4547 +2118-9773 +3676984 +95D497A6-2022-406A-989A-2DA7F04223B0 + + + + + + +Benthana olfersii +( +Brandt, 1833 +) + + + + + + +Fig. 14 + + + + + + + +Philoscia Olfersii + + +Brandt, 1833: 183 + + +. + + + + + +Benthana olfersii + +– + + +Campos-Filho +et al +. 2018a: 11 + + +(for previous records). + + + + + + +Material examined + + + + +BRAZIL +– + +Minas Gerais State +, Itabirito + +• +1 ♂ +; Caverna VL-33; +20°19’54″ S +, +43°56′15″ W +; +29 Mar.– 3 Apr. 2012 +; +Andrade leg +.; +LES +18860 + +. + + + + + +Distribution + + + +Typical in Atlantic forest areas in the states of +Rio de Janeiro +and +São Paulo +( + +Campos-Filho +et al. +2015a + +). This is the first record of this species in cave environments, suggesting a troglophilic status. + + + + \ No newline at end of file diff --git a/data/61/25/87/612587B8C970D647FDCFFC96FC8A60F9.xml b/data/61/25/87/612587B8C970D647FDCFFC96FC8A60F9.xml deleted file mode 100644 index c5916e53877..00000000000 --- a/data/61/25/87/612587B8C970D647FDCFFC96FC8A60F9.xml +++ /dev/null @@ -1,86 +0,0 @@ - - - -New species and new records of terrestrial isopods (Crustacea, Isopoda, Oniscidea) of the families Philosciidae and Scleropactidae from Brazilian caves - - - -Author - -Campos-Filho, Ivanklin Soares - - - -Author - -Fernandes, Camile Sorbo - - - -Author - -Cardoso, Giovanna Monticelli - - - -Author - -Bichuette, Maria Elina - - - -Author - -Aguiar, José Otávio - - - -Author - -Taiti, Stefano - -text - - -European Journal of Taxonomy - - -2020 - -2020-02-20 - - -606 - - -1 -38 - - - -journal article -10.5852/ejt.2020.606 -e75c913b-d6ce-48b6-b5c4-dd09034b4547 -3676984 -95D497A6-2022-406A-989A-2DA7F04223B0 - - - - - - -Benthana olfersii -( -Brandt, 1833 -) - - - - - - -Fig. 14 - - - - \ No newline at end of file diff --git a/data/61/25/87/612587B8C970D647FF24FC34FA996663.xml b/data/61/25/87/612587B8C970D647FF24FC34FA996663.xml deleted file mode 100644 index 4e8ec8f25df..00000000000 --- a/data/61/25/87/612587B8C970D647FF24FC34FA996663.xml +++ /dev/null @@ -1,153 +0,0 @@ - - - -New species and new records of terrestrial isopods (Crustacea, Isopoda, Oniscidea) of the families Philosciidae and Scleropactidae from Brazilian caves - - - -Author - -Campos-Filho, Ivanklin Soares - - - -Author - -Fernandes, Camile Sorbo - - - -Author - -Cardoso, Giovanna Monticelli - - - -Author - -Bichuette, Maria Elina - - - -Author - -Aguiar, José Otávio - - - -Author - -Taiti, Stefano - -text - - -European Journal of Taxonomy - - -2020 - -2020-02-20 - - -606 - - -1 -38 - - - -journal article -10.5852/ejt.2020.606 -e75c913b-d6ce-48b6-b5c4-dd09034b4547 -3676984 -95D497A6-2022-406A-989A-2DA7F04223B0 - - - - - - - -Philoscia Olfersii - - -Brandt, 1833: 183 - - -. - - - - - - - -Benthana olfersii - -– - - -Campos-Filho -et al -. 2018a: 11 - - -(for previous records). - - - - - - -Material examined - - - - -BRAZIL -– - -Minas Gerais State -, Itabirito - -• -1 ♂ -; Caverna VL-33; -20°19’54″ S -, -43°56′15″ W -; -29 Mar.– 3 Apr. 2012 -; -Andrade leg -.; -LES -18860 - -. - - - - - -Distribution - - - -Typical in Atlantic forest areas in the states of -Rio de Janeiro -and -São Paulo -( - -Campos-Filho -et al. -2015a - -). This is the first record of this species in cave environments, suggesting a troglophilic status. - - - - \ No newline at end of file diff --git a/data/61/25/87/612587B8C971D646FD80FD58FDA0666D.xml b/data/61/25/87/612587B8C971D646FD80FD58FDA0666D.xml index 3b952d25eed..6c31c8e9757 100644 --- a/data/61/25/87/612587B8C971D646FD80FD58FDA0666D.xml +++ b/data/61/25/87/612587B8C971D646FD80FD58FDA0666D.xml @@ -1,70 +1,90 @@ - - - -New species and new records of terrestrial isopods (Crustacea, Isopoda, Oniscidea) of the families Philosciidae and Scleropactidae from Brazilian caves + + + +New species and new records of terrestrial isopods (Crustacea, Isopoda, Oniscidea) of the families Philosciidae and Scleropactidae from Brazilian caves - - -Author + + +Author -Campos-Filho, Ivanklin Soares +Campos-Filho, Ivanklin Soares +C752F864-3C84-4AF4-9EDA-4D1AF464D615 +Universidade Federal de Campina Grande, Programa de Pós-Graduação em Engenharia e Gestão de Recursos Naturais, Av. Aprígio Veloso, 882, Bairro Universitário, 58429 - 140 Campina Grande, Paraíba, Brazil. +ivanklin.filho@gmail.com - - -Author + + +Author -Fernandes, Camile Sorbo +Fernandes, Camile Sorbo +C8246067-8235-4981-87D8-56FD9C43FDC2 +Universidade Federal de São Carlos, Departamento de Ecologia e Biologia Evolutiva, Rodovia Washington Luis, Km 235, 13565 - 905 São Carlos, São Paulo, Brazil. +camilesorbofernandes@yahoo.com.br - - -Author + + +Author -Cardoso, Giovanna Monticelli +Cardoso, Giovanna Monticelli +C829E5B7-B87E-4C4A-B88E-D5FE377AE60B +Universidade Federal do Rio Grande do Sul, Programa de Pós-Graduação em Biologia Animal, Departamento de Zoologia, Laboratório de Carcinologia, Av. Bento Gonçalves, 9500, Agronomia, 91510 - 979 Porto Alegre, Rio Grande do Sul, Brazil. +jojomonticelli@hotmail.com - - -Author + + +Author -Bichuette, Maria Elina +Bichuette, Maria Elina +7B740115-BCA0-4711-8C9D-CD014FD3122B +Universidade Federal de São Carlos, Departamento de Ecologia e Biologia Evolutiva, Rodovia Washington Luis, Km 235, 13565 - 905 São Carlos, São Paulo, Brazil. +lina.cave@gmail.com - - -Author + + +Author -Aguiar, José Otávio +Aguiar, José Otávio +8D91D781-EF49-42CC-B7AE-12789CB1CD71 +Universidade Federal de Campina Grande, Programa de Pós-Graduação em Engenharia e Gestão de Recursos Naturais, Av. Aprígio Veloso, 882, Bairro Universitário, 58429 - 140 Campina Grande, Paraíba, Brazil. +otavio.j.aguiar@gmail.com - - -Author + + +Author -Taiti, Stefano +Taiti, Stefano +62E97059-6AE5-4984-9ABB-7FB6F7358BD6 +Istituto di Ricerca sugli Ecosistemi Terrestri, Consiglio Nazionale delle Ricerche, Via Madonna del Piano 10, 50019 Sesto Fiorentino (Florence), Italy. & Museo di Storia Naturale, Sezione di Zoologia “ La Specola ”, Via Romana 17, 50125 Florence, Italy. +stefano.taiti@cnr.it -text - - -European Journal of Taxonomy +text + + +European Journal of Taxonomy - -2020 - -2020-02-20 + +2020 + +2020-02-20 - -606 + +606 - -1 -38 + +1 +38 -journal article -10.5852/ejt.2020.606 -e75c913b-d6ce-48b6-b5c4-dd09034b4547 -3676984 -95D497A6-2022-406A-989A-2DA7F04223B0 +journal article +23893 +10.5852/ejt.2020.606 +e75c913b-d6ce-48b6-b5c4-dd09034b4547 +2118-9773 +3676984 +95D497A6-2022-406A-989A-2DA7F04223B0 - + @@ -83,7 +103,7 @@ - + Bentheana taeiata Araujo & Buckup, 1994: 269 @@ -117,7 +137,7 @@ Campos-Filho - + BRAZIL – @@ -137,27 +157,28 @@ Campos-Filho Bichuette leg .; LES -4419 • +4419 + + +• 2 ♂♂ , 1 ♀ - -; - -same collection data as for preceding; +; same collection data as for preceding; LES -4421 • +4421 + + +• 1 ♀ - -; - -same collection data as for preceding; +; same collection data as for preceding; LES -18861 • -1 ♂ +18861 + +• +1 ♂ ; - Gruta do Capinzal; 24°19′55″ S diff --git a/data/61/25/87/612587B8C971D646FDA5FAA8FB436687.xml b/data/61/25/87/612587B8C971D646FDA5FAA8FB436687.xml index 494212a6072..5f13ff51907 100644 --- a/data/61/25/87/612587B8C971D646FDA5FAA8FB436687.xml +++ b/data/61/25/87/612587B8C971D646FDA5FAA8FB436687.xml @@ -1,70 +1,90 @@ - - - -New species and new records of terrestrial isopods (Crustacea, Isopoda, Oniscidea) of the families Philosciidae and Scleropactidae from Brazilian caves + + + +New species and new records of terrestrial isopods (Crustacea, Isopoda, Oniscidea) of the families Philosciidae and Scleropactidae from Brazilian caves - - -Author + + +Author -Campos-Filho, Ivanklin Soares +Campos-Filho, Ivanklin Soares +C752F864-3C84-4AF4-9EDA-4D1AF464D615 +Universidade Federal de Campina Grande, Programa de Pós-Graduação em Engenharia e Gestão de Recursos Naturais, Av. Aprígio Veloso, 882, Bairro Universitário, 58429 - 140 Campina Grande, Paraíba, Brazil. +ivanklin.filho@gmail.com - - -Author + + +Author -Fernandes, Camile Sorbo +Fernandes, Camile Sorbo +C8246067-8235-4981-87D8-56FD9C43FDC2 +Universidade Federal de São Carlos, Departamento de Ecologia e Biologia Evolutiva, Rodovia Washington Luis, Km 235, 13565 - 905 São Carlos, São Paulo, Brazil. +camilesorbofernandes@yahoo.com.br - - -Author + + +Author -Cardoso, Giovanna Monticelli +Cardoso, Giovanna Monticelli +C829E5B7-B87E-4C4A-B88E-D5FE377AE60B +Universidade Federal do Rio Grande do Sul, Programa de Pós-Graduação em Biologia Animal, Departamento de Zoologia, Laboratório de Carcinologia, Av. Bento Gonçalves, 9500, Agronomia, 91510 - 979 Porto Alegre, Rio Grande do Sul, Brazil. +jojomonticelli@hotmail.com - - -Author + + +Author -Bichuette, Maria Elina +Bichuette, Maria Elina +7B740115-BCA0-4711-8C9D-CD014FD3122B +Universidade Federal de São Carlos, Departamento de Ecologia e Biologia Evolutiva, Rodovia Washington Luis, Km 235, 13565 - 905 São Carlos, São Paulo, Brazil. +lina.cave@gmail.com - - -Author + + +Author -Aguiar, José Otávio +Aguiar, José Otávio +8D91D781-EF49-42CC-B7AE-12789CB1CD71 +Universidade Federal de Campina Grande, Programa de Pós-Graduação em Engenharia e Gestão de Recursos Naturais, Av. Aprígio Veloso, 882, Bairro Universitário, 58429 - 140 Campina Grande, Paraíba, Brazil. +otavio.j.aguiar@gmail.com - - -Author + + +Author -Taiti, Stefano +Taiti, Stefano +62E97059-6AE5-4984-9ABB-7FB6F7358BD6 +Istituto di Ricerca sugli Ecosistemi Terrestri, Consiglio Nazionale delle Ricerche, Via Madonna del Piano 10, 50019 Sesto Fiorentino (Florence), Italy. & Museo di Storia Naturale, Sezione di Zoologia “ La Specola ”, Via Romana 17, 50125 Florence, Italy. +stefano.taiti@cnr.it -text - - -European Journal of Taxonomy +text + + +European Journal of Taxonomy - -2020 - -2020-02-20 + +2020 + +2020-02-20 - -606 + +606 - -1 -38 + +1 +38 -journal article -10.5852/ejt.2020.606 -e75c913b-d6ce-48b6-b5c4-dd09034b4547 -3676984 -95D497A6-2022-406A-989A-2DA7F04223B0 +journal article +23893 +10.5852/ejt.2020.606 +e75c913b-d6ce-48b6-b5c4-dd09034b4547 +2118-9773 +3676984 +95D497A6-2022-406A-989A-2DA7F04223B0 - + @@ -76,7 +96,7 @@ Genus - + @@ -85,8 +105,6 @@ species - - Metaprosekia nodilinearis diff --git a/data/61/25/87/612587B8C976D641FE2FFEF9FC3067F9.xml b/data/61/25/87/612587B8C976D641FE2FFEF9FB406738.xml similarity index 75% rename from data/61/25/87/612587B8C976D641FE2FFEF9FC3067F9.xml rename to data/61/25/87/612587B8C976D641FE2FFEF9FB406738.xml index b547f04f20f..ba037643b5c 100644 --- a/data/61/25/87/612587B8C976D641FE2FFEF9FC3067F9.xml +++ b/data/61/25/87/612587B8C976D641FE2FFEF9FB406738.xml @@ -1,70 +1,90 @@ - - - -New species and new records of terrestrial isopods (Crustacea, Isopoda, Oniscidea) of the families Philosciidae and Scleropactidae from Brazilian caves + + + +New species and new records of terrestrial isopods (Crustacea, Isopoda, Oniscidea) of the families Philosciidae and Scleropactidae from Brazilian caves - - -Author + + +Author -Campos-Filho, Ivanklin Soares +Campos-Filho, Ivanklin Soares +C752F864-3C84-4AF4-9EDA-4D1AF464D615 +Universidade Federal de Campina Grande, Programa de Pós-Graduação em Engenharia e Gestão de Recursos Naturais, Av. Aprígio Veloso, 882, Bairro Universitário, 58429 - 140 Campina Grande, Paraíba, Brazil. +ivanklin.filho@gmail.com - - -Author + + +Author -Fernandes, Camile Sorbo +Fernandes, Camile Sorbo +C8246067-8235-4981-87D8-56FD9C43FDC2 +Universidade Federal de São Carlos, Departamento de Ecologia e Biologia Evolutiva, Rodovia Washington Luis, Km 235, 13565 - 905 São Carlos, São Paulo, Brazil. +camilesorbofernandes@yahoo.com.br - - -Author + + +Author -Cardoso, Giovanna Monticelli +Cardoso, Giovanna Monticelli +C829E5B7-B87E-4C4A-B88E-D5FE377AE60B +Universidade Federal do Rio Grande do Sul, Programa de Pós-Graduação em Biologia Animal, Departamento de Zoologia, Laboratório de Carcinologia, Av. Bento Gonçalves, 9500, Agronomia, 91510 - 979 Porto Alegre, Rio Grande do Sul, Brazil. +jojomonticelli@hotmail.com - - -Author + + +Author -Bichuette, Maria Elina +Bichuette, Maria Elina +7B740115-BCA0-4711-8C9D-CD014FD3122B +Universidade Federal de São Carlos, Departamento de Ecologia e Biologia Evolutiva, Rodovia Washington Luis, Km 235, 13565 - 905 São Carlos, São Paulo, Brazil. +lina.cave@gmail.com - - -Author + + +Author -Aguiar, José Otávio +Aguiar, José Otávio +8D91D781-EF49-42CC-B7AE-12789CB1CD71 +Universidade Federal de Campina Grande, Programa de Pós-Graduação em Engenharia e Gestão de Recursos Naturais, Av. Aprígio Veloso, 882, Bairro Universitário, 58429 - 140 Campina Grande, Paraíba, Brazil. +otavio.j.aguiar@gmail.com - - -Author + + +Author -Taiti, Stefano +Taiti, Stefano +62E97059-6AE5-4984-9ABB-7FB6F7358BD6 +Istituto di Ricerca sugli Ecosistemi Terrestri, Consiglio Nazionale delle Ricerche, Via Madonna del Piano 10, 50019 Sesto Fiorentino (Florence), Italy. & Museo di Storia Naturale, Sezione di Zoologia “ La Specola ”, Via Romana 17, 50125 Florence, Italy. +stefano.taiti@cnr.it -text - - -European Journal of Taxonomy +text + + +European Journal of Taxonomy - -2020 - -2020-02-20 + +2020 + +2020-02-20 - -606 + +606 - -1 -38 + +1 +38 -journal article -10.5852/ejt.2020.606 -e75c913b-d6ce-48b6-b5c4-dd09034b4547 -3676984 -95D497A6-2022-406A-989A-2DA7F04223B0 +journal article +23893 +10.5852/ejt.2020.606 +e75c913b-d6ce-48b6-b5c4-dd09034b4547 +2118-9773 +3676984 +95D497A6-2022-406A-989A-2DA7F04223B0 - + @@ -116,7 +136,7 @@ Campos-Filho ; - + 2013: 466 , fig. 12b. — @@ -170,7 +190,7 @@ Campos-Filho - + BRAZIL – @@ -193,12 +213,12 @@ Campos-Filho Moreira leg .; LES -10818 • -3 ♀♀ +10818 -; - -Laranjeiras, + +• +3 ♀♀ +; Laranjeiras, Caverna dos Aventureiros; 10°48′11.6″ S @@ -213,12 +233,12 @@ dos Aventureiros; Moreira leg .; LES -10819 • -1 ♂ +10819 -; - -Japaratuba, + +• +1 ♂ +; Japaratuba, Caverna Casa do Caboclo; 10°37′57.16″ S @@ -233,11 +253,10 @@ Casa do Caboclo; Moreira leg .; LES -10820 • 4 juvs +10820 -; - -Murici, Fazenda São José, + +• 4 juvs; Murici, Fazenda São José, Caverna Toca da Raposa; 9°13′35.95″ S @@ -287,14 +306,5 @@ Campos-Filho is considered here as trogloxene. - - -Family - -Scleropactidae -Verhoeff, 1938 - - - \ No newline at end of file diff --git a/data/7E/13/87/7E1387F6FFC3FF94E987FD96AE0AFAAE.xml b/data/7E/13/87/7E1387F6FFC3FF94E987FD96AE0AFAAE.xml new file mode 100644 index 00000000000..8c6459ee291 --- /dev/null +++ b/data/7E/13/87/7E1387F6FFC3FF94E987FD96AE0AFAAE.xml @@ -0,0 +1,345 @@ + + + +Revisionary notes on the genera Symplanella Fennah, 1987 and Pseudosymplanella Che, Zhang et Webb, 2009 (Hemiptera: Fulgoromorpha: Caliscelidae, Ommatidiotinae), with description of new subgenera and new species + + + +Author + +Gnezdilov, Vladimir M. + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +40 +56 + + + + +https://doi.org/10.11646/zootaxa.5543.1.2 + +journal article +10.11646/zootaxa.5543.1.2 +1175-5326 +14384924 +98F81CC2-4AF3-46A8-A9A0-81008EF4EFE9 + + + + + + + +Symplanella +( +Bambunella +) +cattiensis + +sp. nov. + + + + + + +( +Figs 14–32 +) + + +Description. +Structure. +Coryphe wide, 1.3 times as wide between the eyes as long medially, with convex anterior margin, deeply concave posterior margin, and lateral margin weakly convex ( +Figs 14 +, +18 +). Eyes large, each is 0.7 as wide as coryphe (in dorsal view) ( +Fig. 18 +). Metope 1.5 times as long medially as wide between eyes, slightly enlarged above clypeus, with median carina running from its upper margin to basal part of anteclypeus and with sublaleral carinae running from its upper margin, but not reaching metopoclypeal suture ( +Fig. 16 +). Median and sublateral carinae not joint at upper margin of metope. Metopoclypeal suture weak, slightly convex. Rostrum reaching hind coxae, with very short not narrowing apically third segment. Pronotum with acutely angulated anterior margin and deeply concave posterior margin, without carinae. Pronotum slightly shorter than coryphe medially. Paradiscal fields of pronotum not narrow. Paranotal lobes of pronotum elongate. Mesonotum large, nearly 2.5 times as long as pronotum, without carinae. Fore wings with elongate basal cell. Clavus long, 4/5 of whole wing length, reaching abdominal apex ( +Figs 15 +, +19 +). Clavus closed, Pcu + A +1 +running to CuP before claval apex. Radius and median running by short common stem from the basal cell. Forewing vein sequence: R 3, firstly furcating before nodal line; r-m 2; M 3–4, furcating after nodal line; m-cua 2–3; CuA 2–3, furcating after nodal line; cua-cup 1–2 ( +Figs 19, 20 +). Hind wings as long as fore wings. Hind tibia with a single lateral spine below its middle and with six spines apically. First metatarsomere nearly three times longer than second one, both without spines. + + +Coloration. +General coloration brownish light yellow ( +Figs 14, 15 +). Coryphe, pro- and mesonotum with red median line ( +Fig. 14 +). Upper half of metope between eyes light brown to brown. Lower half of metope and postclypeus and anteclypeus frontally dark brown to black ( +Fig. 16 +). Ocelli surrounded by red. Pedicel with large black spot outside and with small brown to dark brown spot on inner side. Male paranotal lobes of pronotum and episternae and epimerae sometimes reddish. Fore and hind wings slightly opaque, with light brown veins. Hind coxae with black hind margins. Claws dark brown. Tips of leg spines black. Abdominal sternites with dark brown to black stripes anteriorly and laterally, including the lateral furrows ( +Fig. 17 +, +lf +). Abdominal tergites with wide dark brown lateral stripes. Male abdominal sternites and pygofer sometimes greenish. Female sternite VII sometimes reddish yellow. + + + +Male terminalia ( +Figs 21–30 +). + +Abdominal sternite II with wide and short apodemes ( +Fig. 22 +). Pygofer elongate vertically, with vertical lobe-shaped processes on concaved hind margins below anal tube; posterior margins sharply concave; lower margin widely concave below styles ( +Figs 23, 24 +). Anal tube large, protruding downwards (in lateral view), elongate, with obtusely angulate apex (in dorsal view) ( +Figs 24, 25 +). Anal column wide. Penis massive, with aedeagal shaft strongly curved, U-shaped ( +Fig. 28 +, +ash +). Aedeagal shaft narrowing apically ( +Fig. 29 +). Gonopore apical. Penis with pair of large processes ventrally ( +Figs 27, 28 +, +vpp +). Penis fused with connective which is fused with styles ( +Figs 26, 27 +, +s +). Style rounded, without processes ( +Figs 27, 30 +). + + + +FIGURES 14–17. + +Symplanella +( +Bambunella +) +cattiensis + + +sp. nov. + +14—male paratype, dorsal view; 15—same, lateral view; 16—same, frontal view; 17—female paratype, abdomen ventrally. Abbrevitation: lf—lateral furrow. Scale bar—1 mm. + + + + +FIGURES 18–20. + +Symplanella +( +Bambunella +) +cattiensis + + +sp. nov. + +, holotype. 18—head, dorsal view; 19—left forewing; 20— apex of right forewing. Not to scale. + + + + +FIGURES 21–30. + +Symplanella +( +Bambunella +) +cattiensis + + +sp. nov. + +, male terminalia (21, 23–30—holotype, 22—paratype). 21— abdominal sternites IV–VI; 22—left apodeme of sternite II; 23—pygofer, penis, and styles, ventral view; 24—pygofer and anal tube, lateral view; 25—anal tube, dorsal view; 26— penis, connective, and style, lateral view; 27—penis and styles, ventral view; 28—penis and connective, lateral view; 29—apex of aedeagal shaft, dorsal view; 30—style, dorsal view. Abbrevitations: ash—aedeagal shaft; lf—lateral furrow; vpp—ventral processes of penis; s—style. Not to scale. + + + + +Female terminalia ( +Figs 17 +, +31, 32 +). + +Sternite VII with very weak concavity of hind margin ( +Fig. 31 +). Endogonocoxal lobes fused in shape of large conical sclerotization well visible behind hind margin of sternite VII ( +Figs 31, 32 +, +GxL +). + + +Total length +(from apex of coryphe to the apices of fore wings). Males—5.5–6.0 mm. Females—6.0–6.2 mm. + + + + +Type material. + +Holotype +, +♂ +, +Vietnam +, +Dong Nai Province +, +Cat Tien National Park +, + +1.VI.2012 + +, +A.F. Emeljanov +leg.( +ZIN +) + +. + +Paratypes +: +1♂ +, +Vietnam +, +Dong Nai Province +, +Cat Tien National Park +, +10°25´N +107°25´E +, + +15.XI.2012 + +, +V + +. +M. Gnezdilov +leg. ( +ZIN +); + +2♂ +, +Vietnam +, +Dong Nai Province +, +Cat Tien National Park +, +10°25´N +107°25´E +, + +24.XI.2012 + +, +V + +. +M. Gnezdilov +leg. ( +ZIN +and +MMBC +); + +1♀ +, +Vietnam +, +Dong Nai Province +, +Cat Tien National Park +, + +4. +VI + + +.2012, +A.F. Emeljanov +leg. ( +ZIN +); + +1♀ +, +Vietnam +, +Dong Nai Province +, +Cat Tien National Park +, +10°25´N +107°25´E +, + +18.XI.2012 + +, +V + +. +M. Gnezdilov +leg. ( +ZIN +). + + + + +FIGURES 31–32. + +Symplanella +( +Bambunella +) +cattiensis + + +sp. nov. + +, paratype, female terminalia. 31—sternite VII, fused endogonocoxal lobes, and gonocoxae VIII; 32—fused endogonocoxal lobes. Abbrevitations: VII—sternite VII; Gx VIII— gonocoxa VIII; GxL—fused endogonocoxal lobes. Not to scale. + + + + +Etymology. +The species is named after the +type +locality—Cat Tien National Park in southern +Vietnam +. + + + + \ No newline at end of file diff --git a/data/7E/13/87/7E1387F6FFC3FF9BE987FF19AF7EFE0B.xml b/data/7E/13/87/7E1387F6FFC3FF9BE987FF19AF7EFE0B.xml new file mode 100644 index 00000000000..ea917ebdbb9 --- /dev/null +++ b/data/7E/13/87/7E1387F6FFC3FF9BE987FF19AF7EFE0B.xml @@ -0,0 +1,89 @@ + + + +Revisionary notes on the genera Symplanella Fennah, 1987 and Pseudosymplanella Che, Zhang et Webb, 2009 (Hemiptera: Fulgoromorpha: Caliscelidae, Ommatidiotinae), with description of new subgenera and new species + + + +Author + +Gnezdilov, Vladimir M. + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +40 +56 + + + + +https://doi.org/10.11646/zootaxa.5543.1.2 + +journal article +10.11646/zootaxa.5543.1.2 +1175-5326 +14384924 +98F81CC2-4AF3-46A8-A9A0-81008EF4EFE9 + + + + + + +Subgenus + +Bambunella + +subgen. nov. + + + + + + +Type +species: + +Symplanella zhongtua +Yang et Chen, 2014 + +. + + + + +Diagnosis. +Coryphe almost square or slightly elongate. Metope 1.5 times as long medially as wide between eyes. Pronotum with wide medial part and weakly acutely angulated anterior margin. Penis rather narrow, not flattened laterally. Hind margins of male pygofer often with processes variable in shape among the species. + + + + +Etymology. +Subgeneric name is an amalgamation of “bamboo” and “nella”. + + +Composition and distribution. +Nine species known from +China +and +Vietnam +. + + + + \ No newline at end of file diff --git a/data/7E/13/87/7E1387F6FFC7FF98E987F991AAE2F93E.xml b/data/7E/13/87/7E1387F6FFC7FF98E987F991AAE2F93E.xml new file mode 100644 index 00000000000..96de3a469f9 --- /dev/null +++ b/data/7E/13/87/7E1387F6FFC7FF98E987F991AAE2F93E.xml @@ -0,0 +1,254 @@ + + + +Revisionary notes on the genera Symplanella Fennah, 1987 and Pseudosymplanella Che, Zhang et Webb, 2009 (Hemiptera: Fulgoromorpha: Caliscelidae, Ommatidiotinae), with description of new subgenera and new species + + + +Author + +Gnezdilov, Vladimir M. + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +40 +56 + + + + +https://doi.org/10.11646/zootaxa.5543.1.2 + +journal article +10.11646/zootaxa.5543.1.2 +1175-5326 +14384924 +98F81CC2-4AF3-46A8-A9A0-81008EF4EFE9 + + + + + + + +Symplanella +( +Symplanella +) +breviceps +Fennah, 1987 + + + + + + + +( +Figs 1–13 +) + + + + + + + +Symplanella breviceps +Fennah, 1987: 245 + + +, figs 1–9. + + + +Supplementary description. +Structure. +Coryphe elongate, 1.5 times as long medially as wide between eyes, with obtusely angulate anterior margin and sharply concave posterior margin ( +Fig. 1 +). Eyes large, each is as wide as coryphe (in dorsal view). Metope narrow, twice longer than wide between eyes, slightly enlarged above clypeus, with distinct median carina running throughout postclypeus and sublateral carinae not reaching metopoclypeal suture ( +Fig. 3 +). Median and sublateral carinae joint at one point at upper margin of metope. Ocelli present. Rostrum with short and thick apical segment. Pronotum with narrow medial part and strongly acutely angulated anterior margin. Paradiscal fields of pronotum not narrow. Paranotal lobes nearly oval. Fore wings long and narrow, not narrowing apically, with a nodal line ( +Fig. 2 +). Fore wings with elongate basal cell. Clavus long, 4/5 of whole wing length, reaching abdominal apex. Clavus closed, Pcu + A +1 +running to CuP before claval apex. Radius and median running by short common stem from the basal cell. Forewing vein sequence: R 3, firstly furcating before nodal line; + + + +r-m 2; M 2–4, furcating after nodal line; m-cua 2–3; CuA 1–2; cua-cup 1. Hind wings as long as fore wings. Hind tibia with a single lateral spine medially and with two lateral and four (3 + 1) intermediate spines apically. First metatarsomere 3–4 times as long as second one. First and second metatarsomeres without spines. + + +FIGURES 1–3. + +Symplanella +( +Symplanella +) +breviceps +Fennah, 1987 + +, male (Thailand, Chumphon Province). 1—dorsal view; 2—lateral view; 3—frontal view. Scale bar—1 mm. + + + +Coloration +( +Figs 1–3 +). Coryphe, pro- and mesonotum with wide red stripe medially. Metope pale, sometimes reddish between eyes. Pro- and mesonotum red laterally. Clypeus, fore and middle coxae, femora, apices of third tarsomeres, and claws dark brown to black. Pedicel with black spot. Fore and hind wings slightly opaque. Males with reddish veins of fore wings up to nodal line and black laterally abdominal sternites. Tips of leg spines black. Females with reddish abdominal tergites. + + + +Male terminalia ( +Figs 4–11 +). + +Abdominal sternite II with wide and short apodemes each bearing three spine-shaped processes on inner margins ( +Fig. 11 +). Anal tube short, with deeply concave hind margin (in dorsal view) ( +Fig. 6 +). Anal column wide. Pygofer elongate vertically, with weak processes of concaved hind margins below anal tube; lower margin without processes ( +Figs 4, 5 +). Penis fused with connective, wide and flattened laterally, bottleshaped ventrally, with a short tubular apex ( +Figs 7, 8 +). Gonopore apical. Style with long and narrow caudo-dorsal angle of the plate and long neck, with large capitulum and finger-shaped apical part ( +Figs 9, 10 +). + + + +FIGURES 4–11. + +Symplanella +( +Symplanella +) +breviceps +Fennah, 1987 + +, male terminalia (Thailand, Chumphon Province). 4— pygofer and anal tube, lateral view; 5—pygofer, caudal view; 6—anal tube, dorsal view; 7—penis and connective, lateral view; 8—penis and connective, ventral view; 9—style, lateral view; 10—style, dorsal view; 11—left apodeme of sternite II. Not to scale. + + + + +Female terminalia ( +Figs 12, 13 +). + +Hind margin of sternite VII widely concave, with endogonocoxal lobes fused in shape of large rounded sclerotization well visible behind its margin ( +Fig. 13 +, +GxL +). Anal tube short and wide, with widely concave hind margin (in ventral view) ( +Fig. 12 +). Gonoplacs shortly triangular. + + + +FIGURES 12–13. + +Symplanella +( +Symplanella +) +breviceps +Fennah, 1987 + +, female terminalia (Thailand, Chumphon Province). 12—anal tube, ventral view; 13—sternite VII, gonocoxae VIII, and fused endogonocoxal lobes, ventral view. Abbrevitations: VII—sternite VII; Gx VIII—gonocoxa VIII; GxL—fused endogonocoxal lobes. Not to scale. + + + +Total length. +Males and females—5.0 mm (from apex of coryphe to the apices of fore wings). + + + + +Material examined. + +1♂ +, +1♀ +, “ +Thailand +, +Chumphon +/ prov., + +27.iii.–14.iv.1996 + +/ +Pha To +env. +9°48´N +98°47´E +/ +P. Průdek +leg.” // “Collectio / Moravské museum / Brno” ( +MMBC +) + +; + +1♂ +, “S. +Thailand +, 18–23.iv., / +Chumphon +pr., + + +200 m + +. + +,/ +Phato +, +9°48´N +98°48´E +, / +Jiří Kolibáč +leg., 1997” // “Collectio / Moravské museum / Brno” ( +ZIN +) + +; + +1♀ +, “ +Thailand +, + +22–26.iii.1996 + +/ +Ranong prov. +9°34´N +98°42´E +/ Ban Na env. / +P. Průdek +leg.” // “Collectio / Moravské museum / Brno” ( +ZIN +) + +. + + + + \ No newline at end of file diff --git a/data/7E/13/87/7E1387F6FFCFFF91E987F95AAA7BF95D.xml b/data/7E/13/87/7E1387F6FFCFFF91E987F95AAA7BF95D.xml new file mode 100644 index 00000000000..a847cd5b15d --- /dev/null +++ b/data/7E/13/87/7E1387F6FFCFFF91E987F95AAA7BF95D.xml @@ -0,0 +1,272 @@ + + + +Revisionary notes on the genera Symplanella Fennah, 1987 and Pseudosymplanella Che, Zhang et Webb, 2009 (Hemiptera: Fulgoromorpha: Caliscelidae, Ommatidiotinae), with description of new subgenera and new species + + + +Author + +Gnezdilov, Vladimir M. + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +40 +56 + + + + +https://doi.org/10.11646/zootaxa.5543.1.2 + +journal article +10.11646/zootaxa.5543.1.2 +1175-5326 +14384924 +98F81CC2-4AF3-46A8-A9A0-81008EF4EFE9 + + + + + + + +Pseudosymplanella +( +Nelobamba +) +kubani + +sp. nov. + + + + + + +( +Figs 33–51 +) + + +Description. +Structure. +Eyes large, each is 0.6 as wide as coryphe (in dorsal view) ( +Figs 33 +, +37 +). Coryphe almost square, 1.3 times as wide between eyes as long at midline, with convex anterior margin, deeply concave posterior margin, and lateral margins nearly straight.. Coryphe with short median carina developed only basally ( +Fig. 37 +). Metope as wide between eyes as long medially, enlarged above clypeus, with median and sublateral carinae running from its upper margin separately ( +Fig. 35 +). Median carina running throughout postclypeus; sublateral carinae not reaching metopoclypeal suture. Metopoclypeal suture strongly convex. Rostrum reaching hind coxae, 3 +rd +segment short and wide. Metope and coryphe joint at acute angle in lateral view ( +Fig. 34 +). Pronotum with convex anterior margin and deeply concave posterior margin, without carinae. Paradiscal fields of pronotum not narrow. Paranotal lobes of pronotum elongately oval. Mesonotum twice longer than pronotum medially, without carinae. Fore wings long and narrow, with narrowly rounded apices, surpassing beyond abdominal apex at 1/5 of its length; without nodal line. Clavus long, 2/3 of whole wing length, reaching abdominal apex. Clavus opened, Pcu + A +1 +running to its apex. Basal cell long and narrow. R and M running by one short stem from the basal cell ( +Fig. 38 +). Forewing vein sequence: R 2, furcating apically; r-m 2; M 3–5, furcating apically; m-cua 2; CuA 1–2; cua-cup 1 ( +Figs 38, 39 +). Hind tibia with a single lateral spine medially and two lateral and four (3 + 1) intermediate spines apically. First metatarsomere nearly three times as long as second one. First and second metatarsomeres without spines. Hind margin of arolium of pretarsus surpassing beyond claw apices in dorsal view. + + +Coloration. +General coloration straw-yellow ( +Figs 33–36 +). Metope whitish above clypeus ( +Fig. 35 +). Postclypeus light brown basally, sometimes with brown stripes. Pedicel with a large black spot. Ocelli surrounded by red. Fore and hind wings slightly opaque. Claws and tips of leg spine black. + + + +FIGURES 33–36. + +Pseudosymplanella +( +Nelobamba +) +kubani + + +sp. nov. + +, female, paratype. 33—dorsal view; 34—lateral view; 35—frontal view; 36—abdomen ventrally. Scale bar—1 mm. + + + + +FIGURES 37–39. + +Pseudosymplanella +( +Nelobamba +) +kubani + + +sp. nov. + +37—holotype, head, dorsal view; 38—male paratype, left forewing; 39—holotype, apex of left forewing. Not to scale. + + + + +Male terminalia ( +Figs 40–49 +). + +Abdominal sternite II with long and narrow apodemes each bearing spine-shaped process on its inner side ( +Fig. 49 +). Pygofer wide; hind margins concave below anal tube, with long undulating filiform processes ( +Fig. 40 +). Lower margin of pygofer weakly convex medially ( +Fig. 41 +). Anal tube large (in lateral view), slightly longer than wide medially (in dorsal view), with pair of short processes ventro-apically ( +Figs 40, 42, 43 +). Anal column long, 1/3 of anal tube length. Penis fused with connective, with tubular-shaped and curved aedeagal shaft (in lateral view) ( +Fig. 44 +). Aedeagal shaft wide, with denticles laterally below apex (in ventral view) ( +Fig. 45 +). Gonopore apical ( +Figs 44–46 +). Style with wide plate and wide capitulum bearing denticles ( +Fig. 47 +). Styles fused medially with a large elongate lobe between them ( +Fig. 48 +, +ls +). + + + +Female terminalia ( +Figs 36 +, +50, 51 +). + +Hind margin of female sternite VII sharply concave medially, with angularly protruding margins of this concavity ( +Figs 36 +, +51 +). Endogonocoxal lobes fused in shape of large sclerotizaton, with a median carina ( +Fig. 51 +, GxL). Anal tube short, cylindrical, weakly concave apically (in ventral view) ( +Fig. 50 +). Gonoplacs shortly triangular. + + +Total length (from apex of coryphe to the apices of forewings). +Males—4.5 mm. Females—5.0 mm. + + + + +Type material. + +Holotype +, +♂ +, “ +Laos +—N, + +21.IV.1999 + +/ +Louang Phrabang prov. +, / +19°53´N +102°69´E, / Khan riv., + +300 m + +/ +Vit. Kubáň +leg.” // “Collectio / Moravské museum / Brno” ( +MMBC +) + +. + +Paratypes +: +2♀ +, “ +Laos +—N, + +21.IV.1999 + +/ +Louang Phrabang prov. +, / +19°53´N +102°69´E, / Khan riv., + +300 m + +/ +Vit. Kubáň +leg.” // “Collectio / Moravské museum / Brno” ( +MMBC +and +ZIN +) + +; + +1♂ +, “ +Laos +—N, + +21.IV.1999 + +/ +Louang Phrabang prov. +, / +20°43´N +102°41´E +, / Muang Ngoy, + +500 m + +/ +Vit. Kubáň +leg.” // “Collectio / Moravské museum / Brno” ( +ZIN +) + +. + + + + +Etymology. +The species is named in honour of Czech coleopterologist Vítězslav Kubáň who collected the +type +series in +Laos +. + + + + \ No newline at end of file diff --git a/data/7E/13/87/7E1387F6FFCFFF97E987FA85AAA0F94E.xml b/data/7E/13/87/7E1387F6FFCFFF97E987FA85AAA0F94E.xml new file mode 100644 index 00000000000..019ef11c42a --- /dev/null +++ b/data/7E/13/87/7E1387F6FFCFFF97E987FA85AAA0F94E.xml @@ -0,0 +1,93 @@ + + + +Revisionary notes on the genera Symplanella Fennah, 1987 and Pseudosymplanella Che, Zhang et Webb, 2009 (Hemiptera: Fulgoromorpha: Caliscelidae, Ommatidiotinae), with description of new subgenera and new species + + + +Author + +Gnezdilov, Vladimir M. + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +40 +56 + + + + +https://doi.org/10.11646/zootaxa.5543.1.2 + +journal article +10.11646/zootaxa.5543.1.2 +1175-5326 +14384924 +98F81CC2-4AF3-46A8-A9A0-81008EF4EFE9 + + + + + + +Subgenus + +Nelobamba + +subgen. nov. + + + + + + +Type +species: + +Pseudosymplanella maxima +Gong, Chen et Yang, 2023 + +. + + + + +Diagnosis. +Generally green or straw-yellow except pedicel with a black spot and postclypeus sometimes brown. Style wide, not curved apically. Hind margin of female sternite VII sharply concave medially. Endogonocoxal lobes fused in shape of large sclerotizaton. + + + + +Etymology. +Subgeneric name is an amalgamation of “nella +” +and “bamboo”. + + +Composition and distribution. +Two species from southern +China +( +Yunnan Province +) and northern +Laos +(Louang Phrabang Province). + + + + \ No newline at end of file diff --git a/data/7E/13/87/7E1387F6FFCFFF97E987FB97AE11FB75.xml b/data/7E/13/87/7E1387F6FFCFFF97E987FB97AE11FB75.xml new file mode 100644 index 00000000000..353d3fa1c2f --- /dev/null +++ b/data/7E/13/87/7E1387F6FFCFFF97E987FB97AE11FB75.xml @@ -0,0 +1,103 @@ + + + +Revisionary notes on the genera Symplanella Fennah, 1987 and Pseudosymplanella Che, Zhang et Webb, 2009 (Hemiptera: Fulgoromorpha: Caliscelidae, Ommatidiotinae), with description of new subgenera and new species + + + +Author + +Gnezdilov, Vladimir M. + +text + + +Zootaxa + + +2024 + +2024-12-02 + + +5543 + + +1 + + +40 +56 + + + + +https://doi.org/10.11646/zootaxa.5543.1.2 + +journal article +10.11646/zootaxa.5543.1.2 +1175-5326 +14384924 +98F81CC2-4AF3-46A8-A9A0-81008EF4EFE9 + + + + + + + +Pseudosymplanella +( +Pseudosymplanella +) +nigrifasciata +Che, Zhang et Webb, 2009 + + + + + + + + + + +Pseudosymplanella nigrifasciata +Che, Zhang et Webb, 2009: 50 + + +, figs 1–3, 7–21. + + + + + +Material examined. + +1♂ +, +1♀ +, +China +, +Yunnan Province +, +Menghai county +, + +25. +VI + + + +.2019, +Nian Gong +leg. ( +ZIN +) + +. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FF0039D092DA912D37A8DAAD.xml b/data/98/1B/87/981B8798FF0039D092DA912D37A8DAAD.xml new file mode 100644 index 00000000000..41a94e346f0 --- /dev/null +++ b/data/98/1B/87/981B8798FF0039D092DA912D37A8DAAD.xml @@ -0,0 +1,178 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Porphyrophora victoriae +Jashenko + + + + + + + +( +Fig. 77 +, distribution map +Fig. 92G +) + + + + + + + +Porphyrophora victoriae +Jashenko, 1994: 36 + + +. + + + +Field characters: +Appearance of live specimens not recorded. + + + + +Microscopic diagnosis: +Body of slide-mounted adult female broadly oval, with segments on both surfaces each with a dense band of long setae, except for abdominal segment I on dorsum and segment III on venter each with only a single transverse row of setae; also with long setae in belts on sides of dorsal abdomen and other segments. Antennae each with 7 or 8 segments; basal segment with a few setae; sensilla placodeum on segment IV usually present, circular, containing 5 or 6 small sensilla, and usually with 1 very long fleshy seta present next to sensilla placodeum; other intermediate segments naked; apical segment only slightly swollen, bearing some flagellate setae, several fleshy setae and sensilla. Prothoracic legs each with tarsus fused with claw, enlarged claw without a denticle. Thoracic spiracles each with an apodeme; with 4 or 5 spiracular disc-pores, each pore with a central loculus and 1 ring of peripheral loculi; with 2‒4 perispriracular sensilla present near each thoracic spiracle. Abdominal spiracles absent. Multilocular disc-pores each with a central loculus and 1 or 2 rings of peripheral loculi; pores on head, thorax and abdominal segments I‒VI each with second ring of loculi incomplete; abdominal segments VII and VIII with pores each containing 2 complete rings of loculi. Anal opening with sclerotized rim on dorsal margin, surrounded by an area of derm without setae or pores. Vulva membranous. + + + + +FIGURE 76. +Adult female of + +Porphyrophora tritici +(Bodenheimer) + +, modified from +Vahedi & Hodgson (2007) +, page 106, fig. 34. + + + + +FIGURE 77. +Adult female of + +Porphyrophora victoriae +Jashenko + +, modified from +Jashenko (1994) +, page 37, fig. 18. + + + + +Distribution: + +Porphyrophora victoriae + +is known from +Kazakhstan +( +Jashenko 1994 +) and +Iran +, +Markazi province +( +Moghaddam 2013 +). + + +Host-plants: +The species has been found on + +Acanthophyllum pungens + +( +Caryophyllaceae +) ( +Jashenko 1994 +) and + +Lepidium draba + +( +Brassicaceae +) ( +Moghaddam 2013 +). + + +Economic importance: + +Porphyrophora victoriae + +is of no economic importance in +Iran +. + + +Natural enemies: +None recorded in +Iran +. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FF0739D292DA916537A8DCEE.xml b/data/98/1B/87/981B8798FF0739D292DA916537A8DCEE.xml new file mode 100644 index 00000000000..ea790c50008 --- /dev/null +++ b/data/98/1B/87/981B8798FF0739D292DA916537A8DCEE.xml @@ -0,0 +1,200 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Porphyrophora medicaginis +Jashenko + + + + + + + +( +Fig. 75 +, distribution map +Fig. 92E +) + + + + + + + +Porphyrophora medicaginis +Jashenko, 1994: 22 + + +. + + + +Field characteristics: +Body more-or-less egg-shaped; derm flexible and soft; long hair setae rather stiff; body light to dark red ( +Jashenko 1994 +). + + + + +Microscopic diagnosis: +Slide-mounted adult female with both surfaces of body quite densely covered by long hair-like setae and small setae.Antennae each with 7–9 segments; sensilla placodeum on segment IV usually present, circular or elliptical, containing about 5–7 small sensilla; other intermediate segments naked; apical segment slightly swollen, bearing some flagellate setae, several fleshy setae and sensilla. Prothoracic legs each with tarsus fused with claw, enlarged claw without a denticle. Each meso- and metathoracic leg with long pretarsus, 1/4 length of claw; meso- and metathoracic claws generally each constricted between claw and pretarsus. Thoracic spiracles each with peritreme containing 4–7 spiracular disc-pores, and texture of membranous peritreme with a mosaic pattern of diamond shapes; perispiracular sensilla situated far from margin of peritreme in a group of 2–7, occasionally with about 2 pores fusing to form a twin pore. Abdominal spiracles apparently absent. Anterior multilocular disc-pores each with 1 even or uneven ring of loculi, each ring mostly with 9–20 loculi; pores mainly present in transverse segmental bands 2 or 3 pores wide; some (not all) posterior multilocular disc-pores tending to be larger, particularly around genital opening, each of these generally with 2 or 3 rings of loculi. Anal opening with narrow, semicircular sclerotized rim along dorsal margin, surrounded by an area of derm without setae or pores. + + + + +FIGURE 74 +. Adult female of + +Porphyrophora jashenkoi +Vahedi + +, modified from +Vahedi & Hodgson (2007) +, page 69, fig. 17. + + + + +FIGURE 75. +Adult female of + +Porphyrophora medicaginis +Jashenko + +, modified from +Vahedi & Hodgson (2007) +, page 82, fig. 23. + + + + +Distribution: + +Porphyrophora medicaginis + +is only known from +Kazakhstan +(Vostochno +Kazakhstan +Oblast) and +Iran +, where it has been found in Azarbaijan -e Sharghi and +Kerman +provinces ( +Jashenko 1994 +; +Moghaddam, 2013 +). + + +Host-plants: +The species has been recorded on host-plants belonging to the families +Fabaceae +and + +Poaceae ( + +García Morales +et al +. 2016 + +) + +. In +Iran +, the host-plants + +Medicago vulgare + +( +Fabaceae +), + +Cynodon +sp. + +and + +Phragmites +sp. + +( +Poaceae +) were recorded ( +Vahedi & Hodgson 2007 +). + + +Economic importance: + +Porphyrophora medicaginis + +is of no economic importance in +Iran +. + + +Natural enemies: +None recorded in +Iran +. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FF0739D592DA95123758D8A3.xml b/data/98/1B/87/981B8798FF0739D592DA95123758D8A3.xml new file mode 100644 index 00000000000..7695c74ebb5 --- /dev/null +++ b/data/98/1B/87/981B8798FF0739D592DA95123758D8A3.xml @@ -0,0 +1,143 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Porphyrophora jashenkoi +Vahedi + + + + + + + +( +Fig. 74 +, distribution map +Fig. 92D +) + + + + + + +Porphyrophora jashenkoi + +Vahedi +2007 + + +in + +Vahedi & Hodgson, 2007: 68 + +. + + + +Field characteristics: +Appearance of live specimens not recorded. + + + + +Microscopic diagnosis: +Slide-mounted adult female oval; dorsal and ventral surfaces of body covered quite sparsely by long hair-like setae and small setae, although setae on margins of head and prothorax more numerous. Antennae each with 7 segments; sensillum placodeum on segment IV usually present, containing a dense group of 3–5 small sensilla, also with a short seta (not illustrated); segment V with a minute seta; other intermediate antennal segments naked; apical segment as wide as, or slightly wider than long, bearing some flagellate setae, several fleshy setae and sensilla. Prothoracic legs each with tarsus fused with claw; enlarged claw at least 2.0x longer than basal width, without a denticle. Thoracic spiracles each with an apodeme; each peritreme with 4–6 spiracular disc-pores; each pore with 1 ring of about 8 evenly-spaced loculi; each disc-pore with a bright central zone; with a group of 2 perispiracular sensilla on lateral margin of peritreme. Abdominal spiracles apparently absent. Multilocular disc-pores very sparse, each pore with a single ring of 8–10 loculi; mainly present in segmental transverse bands 1 or 2 pores wide; posterior multilocular disc-pores tending to be larger, these almost always with 1 or 2 complex rings of loculi. Microducts abundant over dorsum of abdomen (not illustrated). Anal opening with thin, semicircular, poorly sclerotized rim on dorsal margin, surrounded by an area of derm without setae or pores. + + + + +Distribution: + +Porphyrophora jashenkoi + +is known only from +Iran +, Hamadan-Kermanshah province ( +Vahedi & Hodgson 2007 +). + + +Host-plants: +The species was recorded feeding on wheat, + +Triticum aestivum + +( +Poaceae +) ( +Vahedi & Hodgson 2007 +). + + +Economic importance: + +Porphyrophora jashenkoi + +is no economic importance in +Iran +, but might become a problem on wheat if accidentally introduced to other parts of the world. + + +Natural enemies: +None recorded in +Iran +. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FF0939DB92DA93D337A8DA2E.xml b/data/98/1B/87/981B8798FF0939DB92DA93D337A8DA2E.xml new file mode 100644 index 00000000000..80ea8d41b46 --- /dev/null +++ b/data/98/1B/87/981B8798FF0939DB92DA93D337A8DA2E.xml @@ -0,0 +1,137 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Porphyrophora chelodonta +Vahedi + + + + + + + +( +Fig. 71 +, distribution map +Fig. 92A +) + + + + + + +Porphyrophora chelodonta + +Vahedi +2007 + + +in + +Vahedi & Hodgson, 2007: 40 + +. + + + +Field characteristics: +Appearance of live specimens not recorded. + + + + +Microscopic diagnosis: +Slide-mounted adult female broadly oval; both surfaces of body quite sparsely covered with long hair-like setae and small setae. Antennae each with 7 segments; sensillum placodeum on antennal segment IV usually present, small and circular, containing 1–3 or 4 sensilla; segments I‒III, V and VI naked; apical segment enlarged, bearing some flagellate setae, several fleshy setae and sensilla. Prothoracic legs each with a wide femur, deeply curved posteriorly at an angle of about 80˚; tarsus fused with claw; enlarged claw very short (claw length less than 1.7x basal width) and pointed, with a serrate edge. Thoracic spiracles each with a long apodeme; each peritreme with about 6 spiracular disc-pores, and a group of 2–4 perispiracular sensilla on lateral margin. Abdominal spiracles apparently absent. Multilocular disc-pores each normally with 1 or 2 rings of loculi; outer ring with about 8–14 loculi and inner ring with up to 10 loculi; pores present across each segment in transverse bands 2 pores wide. Anal opening situated in a naked area surrounded by numerous setae. + + + + +Distribution: + +Porphyrophora chelodonta + +is known only from +Iran +, +Kermanshah province +( +Vahedi & Hodgson 2007 +). + + +Host-plants: +On an unidentified plant. + + +Economic importance: + +Porphyrophora chelodonta + +is of no economic importance in +Iran +. + + +Natural enemies: +None recorded in +Iran +. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FF0A39D892DA97C335D1D9A7.xml b/data/98/1B/87/981B8798FF0A39D892DA97C335D1D9A7.xml new file mode 100644 index 00000000000..8ffd09cea19 --- /dev/null +++ b/data/98/1B/87/981B8798FF0A39D892DA97C335D1D9A7.xml @@ -0,0 +1,191 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Porphyrophora cynodontis +(Archangelskaya) + + + + + + + +( +Fig. 72 +, distribution map +Fig. 92B +) + + + + + + + +Margarodes cynodontis +Archangelskaya, 1935: 15 + + +. + + + +Field characteristics: +Body more-or-less egg-shaped and hairy; derm flexible and soft; body densely pubescent ( +Jakubski 1965 +); colour ranging from bright violet to dark red. + + + + +Microscopic diagnosis: +Slide-mounted adult female broadly oval; dorsum of body densely covered by long hair-like setae and small setae; ventral surface densely covered by long hair-like setae but without small setae. Antennae each with 9–12 or 13 segments; sensillum placodeum on segment IV usually present, elongate oval to almond shaped, containing 7–12 sensilla plus a short fleshy seta; most other intermediate segments naked but a few bearing 1 or 2 setae; apical segment enlarged, bearing some flagellate setae, several fleshy setae and sensilla. Prothoracic legs each with tarsus fused with claw; enlarged claw at least 2.0x longer than basal width, and pointed, normally without a denticle (rarely, a small one present). Thoracic spiracles larger than those in most other + +Porphyrophora +species + +, each with an apodeme; peritreme with about 10 spiracular disc-pores and with a group of 3–15 perispiracular sensilla on lateral margin. Abdominal spiracles usually absent, but some geographical races from +Iran +with 1 pair of simple abdominal spiracles, very poorly developed, on segment I. Multilocular disc-pores each normally with 1 ring of loculi, each ring mostly with about 12 loculi; pores mainly present in segmental bands 5 or 6 pores wide across each segment; some posterior multilocular disc-pores tending to be larger. Anal opening with thin, semicircular, poorly sclerotized rim along dorsal margin, surrounded by an area of derm without setae or pores. + + + + +Distribution: + +Porphyrophora cynodontis + +is known from +Uzbekistan +and +Iran +( +Jakubski 1965 +; +Vahedi & Hodgson 2007 +), where it has been recorded from Azarbaijan-Sharghi and +Kermanshah +provinces ( +Vahedi & Hodgson 2007 +). + + +Host-plants: +The species has been recorded on host-plants belonging to three grass genera in the family +Poaceae +: + +Aeluropus +, +Cynodon + +and + +Phragmites +( + +García Morales +et al +. 2016 + +) + +. In +Iran +, it has been found on + +Phragmites +sp. + +( +Vahedi 2002 +). + + +Economic importance: + +Porphyrophora cynodontis + +is of no economic importance in +Iran +. + + +Natural enemies: +None recorded in +Iran +. + + + + +Comments: + +Porphyrophora cynodontis + +is probably a synonym of + +P. hamelii +Brandt ( +Vahedi & Hodgson 2007 +) + +. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FF0D39DF92DA94AE37A8DAB6.xml b/data/98/1B/87/981B8798FF0D39DF92DA94AE37A8DAB6.xml new file mode 100644 index 00000000000..032e6334341 --- /dev/null +++ b/data/98/1B/87/981B8798FF0D39DF92DA94AE37A8DAB6.xml @@ -0,0 +1,177 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Nidularia balachowskii +Bodenheimer + + + + + + + +( +Fig. 68 +, +Plate 4G +, distribution map +Fig. 91E +) + + + + + + + +Nidularia balachowskii +Bodenheimer, 1941: 78–80 + + +. + + + +Field characters: +Pre-reproductive adult female oval, soft and flat; dorsum brownish and venter yellowish-white; dorsal surface covered with 10 longitudinal rows of rectangular wax plates. Post-reproductive adult female oval, moderately convex and sclerotized, each side with 5 longitudinal rows of almost fused dark brown wax plates; with lighter brown wax in between rows of plates. A waxy ovisac is produced. + + + + +Microscopic diagnosis: +Slide-mounted adult female fusiform. Marginal setae spinose. Antennae each with 1 segment, bearing 2–6 fleshy setae. Labium triangular, with 3 segments, apical segment with 4 pairs of setae. + + +Dorsum +: Margin with a single row of slender setae; setae absent from rest of dorsum. Microtubular ducts and small, circular simple pores present throughout. + + +Venter +: Setae of 2 sizes: longer setae forming a single row across medial area of each abdominal segment, accompanied by spinules; and shorter setae sparsely present on medial areas of thorax and head. Quinquelocular pores numbering 8–11 between each anterior spiracle and submarginal band of tubular ducts, but only 2‒4 laterad to each posterior spiracle; quinquelocular pores also present in a broad band of 10–13 pores medially between posterior spiracles and in a single, complete submarginal band 1 pore wide, totaling about 50–70 pores on each side. Submarginal tubular duct band also containing dispersed bilocular pores, numbering 103–135 per side. Multilocular pores, each with 9 or 10 loculi, forming group of 3 or 4 pores between anterior and posterior spiracles on each side, and present in transverse bands across abdominal segments. Tubular ducts forming complete submarginal band 2 or 3 ducts wide, and sparsely present medially on thorax. Anal ring ventral, composed of 2 sclerotized semicircles, each half bearing 3 spinose setae and 10–12 pores. + + + + +Distribution: + +Nidularia balachowskii + +is known from +Israel +, +Türkiye +( +Turkey +) and +Iran +( +Bodenheimer 1941 +; + +Spodek +et al +. 2014 + +), where it has been recorded from +Kermanshah +and +Lorestan +provinces ( +Moghaddam 2013 +). + + +Host-plants: +The species is oligophagous, having been recorded on + +Quercus +ithaburensis + +and + +Quercus + +spp. ( +Fagaceae +) ( +Bodenheimer 1941 +; + +Spodek +et al +. 2014 + +). + + +Economic importance: + +Nidularia balachowskii + +is of no economic importance in +Iran +. + + +Natural enemies: +None recorded in +Iran +. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FF1D39CD92DA92B437A8DC02.xml b/data/98/1B/87/981B8798FF1D39CD92DA92B437A8DC02.xml new file mode 100644 index 00000000000..c2f57ac9a4a --- /dev/null +++ b/data/98/1B/87/981B8798FF1D39CD92DA92B437A8DC02.xml @@ -0,0 +1,151 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Rhizococcus asperulae +Moghaddam + + + + + + + +( +Fig. 59 +, +Plate 4B +, distribution map +Fig. 93C +) + + + + + +Rhizococcus asperulae +Moghaddam, 2019: 454 + +. + + +Field characteristics: +Found on upper leaf surfaces. Live adult female oval, body contents dark green; ovisac white, rounded, completely covering body. + + + + +Microscopic diagnosis: +Slide-mounted adult female oval. Antennae each with 7 segments. Frontal lobes and frontal tubercles absent. Legs well developed; hind coxa with fewer than 5 translucent pores, but with scattered spinulae; tibiae each with 5 setae, with median seta present; tarsal and claw digitules slightly knobbed and longer than claw; claw with a denticle. Anal lobes well developed, dorsum of each lobe with 3 enlarged setae and 3 or 4 microtubular ducts. Anal ring with a partial double row of pores and bearing 8 robust setae. Cauda absent. + + +Dorsum +with enlarged setae all spine-like with pointed apices, forming transverse rows across each segment, reaching to margins. Margin of abdominal segment VII with 2 lateral enlarged setae on each side. Macrotubular ducts all of 1 size, present throughout. Microducts distributed throughout. + + + +FIGURE 59. +Adult female of Rhizo +coccus + +asperulae +Moghaddam + +, from Moghaddam (2019), page 456, fig. 3. © Magnolia Press, www.mapress.com/j/zt, reproduced with permission from the author and the copyright holder. + + + +Venter +with normal hair-like setae of various sizes, scattered on median and submedian areas of segments; enlarged setae, same size as those on dorsum, present in a broad band on margins and submargins. Quinquelocular pores sparsely distributed on thorax, and forming rows across abdominal segments I–VIII+IX. Disc-pores absent. Trilocular pores numbering 2 or 3, present singly near each anterior spiracle and hind coxa. Macrotubular ducts similar to those on dorsum, forming a submarginal band and present in medial areas. Microducts present in a regular arrangement throughout. Cruciform pores absent. + + + + +Distribution: + +Rhizococcus asperulae + +has been recorded only from +Iran +, +Markazi province +(Moghaddam 2019). + + +Host-plants: +The scale was found on + +Asperula glomerata + +( +Rubiaceae +) (Moghaddam 2019). + + +Economic importance: +Not known as a pest in +Iran +. + + +Natural enemies: +None recorded in +Iran +. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FF1F39CD92DA96C637A8D8D4.xml b/data/98/1B/87/981B8798FF1F39CD92DA96C637A8D8D4.xml new file mode 100644 index 00000000000..0f73ea77895 --- /dev/null +++ b/data/98/1B/87/981B8798FF1F39CD92DA96C637A8D8D4.xml @@ -0,0 +1,149 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Rhizococcus avicennae +Moghaddam + + + + + + + +( +Fig. 60 +, +Plate 4C +, distribution map +Fig. 93D +) + + + + + + + +Rhizococcus avicennae +Moghaddam, 2018: 520 + + +. + + + +Field characteristics: +Live adult female oval, with dark green body contents; ovisac white, rounded, and completely covering body. + + + + +Microscopic diagnosis: +Slide-mounted adult female body elongate oval, tapering posteriorly. Antennae each with 7 segments. Frontal lobes and frontal tubercles absent. Eyes situated on margins. Legs well developed; hind coxae each with translucent pores; tarsal and claw digitules knobbed apically and longer than claw; claw with a denticle. Anal lobes strongly developed, dorsum of each with 4 enlarged setae and 1 or 2 microtubular ducts. Anal ring with a partial double row of pores and bearing 6 setae. Cauda absent. Large setae on margins few or absent. + + +Dorsum +with enlarged conical setae of +2 types +, both with pointed tips: (i) longer +type +situated on margin, few; and (ii) smaller +type +forming transverse rows across each segment, and irregular rows across head. Margin of abdominal segment VII with 3 lateral enlarged setae on each side. Macrotubular ducts of 3 sizes: (i) large ducts present on head, thorax and abdominal segments; (ii) intermediate-sized ducts present on all segments but much fewer than large ducts; and (iii) small ducts forming transverse rows across each abdominal segment. + + +Venter +with normal flagellate setae of various sizes, present from median area to margin. Labium basal segment bearing 2 unequal pairs of setae. Disc-pores each with 7 loculi in a single ring, forming transverse rows across abdominal segments III–VIII+X, plus a few near each anterior spiracle. Hind femur without translucent pores. Quinquelocular pores present on abdominal segments I–VI and near anterior and posterior spiracles. Macrotubular ducts of 3 sizes, similar to those on dorsum: large ducts forming transverse bands in marginal areas; intermediate-sized ducts scattered but absent from margins; and small ducts forming transverse rows across abdominal segments IV–VI. Cruciform pores few, present on submargins of prothorax and head. + + + + +Distribution: + +Rhizococcus avicennae + +has been recorded only from +Iran +, +Hamadan province +( +Moghaddam 2018 +). + + +Host-plants: +The scale was found on an unidentified grass (P +oaceae) +( +Moghaddam 2018 +). + + +Economic importance: +Not known as a pest in +Iran +. + + +Natural enemies: +None recorded in +Iran +. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FF3339E192DA944D37A8DB60.xml b/data/98/1B/87/981B8798FF3339E192DA944D37A8DB60.xml new file mode 100644 index 00000000000..d1a44879b05 --- /dev/null +++ b/data/98/1B/87/981B8798FF3339E192DA944D37A8DB60.xml @@ -0,0 +1,160 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Ortheziola marottai +Kaydan & Szita + + + + + + + +( +Fig. 86 +, distribution map +Fig. 91G +) + + + + + + +Ortheziola marottai +Kaydan & Szita + +in + + +Kaydan +et al +., 2014: 72–74 + + +. + + + +Field characters: +Intact adult female not seen. + + + + +Microscopic diagnosis: +Slide-mounted adult female body oval. Antennae each with 3 segments, third (apical) antennal segment with long slender apical seta. Legs each with rows of robust spine-like setae. Wax plates (represented by groups of spines) absent from ventral marginal areas of head and thorax, except for a small spine cluster next to antenna and a normal wax plate between antennae; each thoracic spiracle surrounded by marginal wax band. Setae few, scattered; and associated with anterior and posterior multilocular pore rows, several more associated with posterior multilocular pores surrounding vulva. Abdominal spiracles present with 3 pairs on each side of body anterior to ovisac band and 1 pair situated inside ovisac band, near anterolateral angle; each abdominal spiracle with a sclerotized vestibule. Anal ring with incomplete triple row of circular pores. A sclerotized tergite present on abdominal segment VII, anterior to anal ring. Each multilocular pore cluster laterad of anal ring with an abdominal spiracle present in centre. + + + + +Distribution: + +Ortheziola marottai + +has been recorded from +Croatia +, +Greece +(including +Cyprus +), +Romania +, +Türkiye +( +Turkey +) and +Iran +( + +García Morales +et al +. 2016 + +), where it was found in +Gilan province +( + +Kaydan +et al +. 2014 + +). + + +Host-plants: +Unknown. + + +Economic importance: + +Ortheziola marottai + +is of no economic importance in +Iran +. + + +Natural enemies: +None recorded in +Iran +. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FF3539E592DA911637A6DF7E.xml b/data/98/1B/87/981B8798FF3539E592DA911637A6DF7E.xml new file mode 100644 index 00000000000..cafa8f058c7 --- /dev/null +++ b/data/98/1B/87/981B8798FF3539E592DA911637A6DF7E.xml @@ -0,0 +1,174 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Pseudaspidoproctus gramineus +Jashenko & Danzig + + + + + + + +( +Fig. 83 +) + + + + + + + +Pseudaspidoproctus gramineus +Jashenko & Danzig, 1992: 89 + + +. + + + +Field characteristics: +Live adult female exposed on host-plant during growing period; body elongate ovoid, with dorsum strongly convex and venter flattened; eggs deposited within an internal ventral marsupium with a U- or Vshaped opening ( +Jashenko & Danzig 1992 +). + + + + +Microscopic diagnosis: +Slide-mounted adult female oval, about +4–5 mm +long. Antennae each with 8‒10 segments, each sparsely covered with setae. Legs well developed, covered with long setae; inner margin of tibia and tarsus with 1 or 2 rows of short, thick spines; tarsus apparently partly divided, with a small second pseudosegment present near claw, almost completely merged with long first segment. Cicatrices situated posterior to vulva, numbering 3, each usually elongate oval, median one much larger than lateral ones. Abdominal spiracles probably numbering 7 pairs, but difficult to see. Multilocular pores of +2 types +: (i) with 1 central circular or oval loculus and 9‒12 peripheral loculi; (ii) with 3 central loculi and 7‒9 peripheral loculi; multilocular disc-pores few on dorsum, occurring singly; more numerous on venter, in groups on thoracic segments; on abdomen, concentrated around vulva. + + + + +FIGURE 83. +Adult female of + +Pseudaspidoproctus gramineus +Jashenko & Danzig + +, reproduced from +Jashenko & Danzig (1992) +, page 88, fig. 4, from +Entomologicheskoe Obozrenie +. + + + + +Distribution: + +Pseudaspidoproctus gramineus + +is known only from +Yemen +(Afrotropical region), +Afghanistan +and +Iran +(Palaearctic region) ( + +García Morales +et al +. 2016 + +), where + +Kozár +et al +. (1996) + +recorded it, without specifying in which province it was found. + + +Host-plants: +The species has been recorded on host-plants in the family +Poaceae +in +Afghanistan +( +Jashenko & Danzig 1992 +), and on + +Odyssea mucronata + +( +Poaceae +) in +Yemen +, but its host in +Iran +was not determined. + + +Economic importance: +None recorded. + + +Natural enemies +: None recorded in +Iran +. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FFA0397192DA966E37A8DE8A.xml b/data/98/1B/87/981B8798FFA0397192DA966E37A8DE8A.xml new file mode 100644 index 00000000000..1300aa671ce --- /dev/null +++ b/data/98/1B/87/981B8798FFA0397192DA966E37A8DE8A.xml @@ -0,0 +1,201 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Eulecanium ficiphilum +Borchsenius + + + + + + + +( +Fig. 23 +, distribution map +Fig. 90D +) + + + + + + + +Eulecanium ficiphilum +Borchsenius, 1955: 293 + + +. + + + +Field characteristics: +Live adult female body irregularly subspherical, wider than long, with rear end perpendicular to substrate; dead adult female light or dark yellow, with brown spots, wrinkled and with numerous concavities in surface. + + + + +Microscopic diagnosis: +Slide-mounted adult female body more-or-less circular; stigmatic clefts absent; anal cleft quite well developed. + + + +FIGURE 23. +Adult female of + +Eulecanium ficiphilum +Borchsenius. + + + + +Dorsum +. Derm membranous in young specimens but becoming sclerotized at maturity, particularly in area around anal plates. Setae spinose, each with a well-developed basal socket, scattered throughout. Pores of +1 type +present throughout. Preopercular pores absent. Tubular ducts of +1 type +present throughout, each with fairly long outer ductule, and slightly shorter, narrower inner filament ending in a terminal gland.Anal plates together quadrate, each with 3 or 4 apical or subapical setae and 1 or 2 setae along inner margin. Anal ring bearing probably 6 setae. + + +Margin +. Marginal setae of +2 types +: (i) some long flagellate setae present at anterior end, and a few at posterior end of abdomen, extending onto distal anal cleft margins; and (ii) most marginal setae thick, elongate-conical, blunt and spinelike, mainly on lateral margins. Stigmatic areas each with 0–3 spiracular setae, not differentiated from marginal setae. + + +Venter +. Derm completely membranous. Pregenital disc-pores each with 10 loculi, present around anogenital fold and forming large medial and submedial segmental clusters on abdominal and thoracic segments. Spiracular disc-pores each with 5 loculi, present in a narrow band between each spiracle and margin. Ventral tubular ducts of +2 types +: (i) larger ducts, present in a wide submarginal band around entire submargin; and (ii) smaller ducts, present around mouthparts. Legs well developed, each without a tibio-tarsal articulatory sclerosis; claw lacking a denticle; claws very thick, rough, each held at 90º to tarsus; claw digitules absent or shorter than claw. Tarsal digitules slender, short, and approximately as long as claw. Antennae each with 8 segments. + + + + +Distribution: + +Eulecanium ficiphilum + +is only known from +Afghanistan +, +Türkiye +( +Turkey +), +Turkmenistan +and +Iran +( + +García Morales +et al +. 2016 + +); in +Iran +, it is found in +Kerman +and Sistan & Balouchestan provinces ( +Moghaddam 2013 +). + + +Host-plants: +The species has only been recorded on + +Ficus +spp. + +including + +F. carica + +( +Moraceae +) ( + +García Morales +et al +. 2016 + +). In +Iran +, it has been recorded on edible fig, + +F. carica +( +Bodenheimer 1944b +) + +. + + +Economic importance: +There are no records of economic damage in +Iran +. + + +Natural enemies: +None recorded in +Iran +. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FFAC397E92DA963A304FDBFB.xml b/data/98/1B/87/981B8798FFAC397E92DA963A304FDBFB.xml new file mode 100644 index 00000000000..5141c4c3272 --- /dev/null +++ b/data/98/1B/87/981B8798FFAC397E92DA963A304FDBFB.xml @@ -0,0 +1,219 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Ceroplastes sinensis +Del Guercio + + + + + + + +( +Fig. 17 +, distribution map +Fig. 89H +) + + + + + + + +Ceroplastes sinensis +Del Guercio, 1900: 3 + + +. + + + +Field characteristics: +Live adult female body almost circular with an irregular margin, convex, covered with thick wax. Wax of young females white, divided into 8 plates, 5 near margins with a blurred brownish patch at the centre of each; in ventral view, with narrow white strip of secretion between each spiracle and margin. Old females with reddish brown wax, with plate divisions and darker brown patches indistinct. + + + + +Microscopic diagnosis: +Slide-mounted adult female body oval, convex, with shallow stigmatic clefts. Caudal process short and stout. + + +Dorsum +. Derm membranous in young females, except for sclerotized caudal process. Derm with 8 clear areas. Setae each cylindrical with a rounded or pointed apex. Pores mostly trilocular, of 3 main +types +: (i) oval trilocular pores, (ii) triangular trilocular pores, and (iii) bilocular pores; a few 4-locular pores also present, particularly at anterior end. Preopercular pores present in a transverse band, numbering about 8–20 pores. Tubular ducts and duct tubercles absent. Anal plates each with 3 large dorsal setae on posterior quarter of each plate, plus a smaller apical seta (not shown in +Fig. 17 +). + + +Margin +. Marginal setae flagellate; with 2−4 marginal setae between anterior and posterior stigmatic clefts on each side; anal lobe with 3 or 4 marginal setae; submarginal setae more frequent than marginal setae. Stigmatic clefts shallow, each with 18−54 hemispherical, bullet-shaped or lanceolate spiracular setae in 3 or 4 irregular rows in each cleft. + + +Venter +. Derm membranous. Pregenital disc-pores restricted to around anogenital fold and across 2 preceding abdominal segments; much less frequent medially and occasionally present mediolaterally on segment V; generally absent from segment IV and more anteriorly. Spiracular disc-pore bands at narrowest point each only 3 or 4 pores wide, but about 5 or 6 pores wide near margin. Cruciform pores scattered (not illustrated). Simple microducts (filamentous ducts) present around margins, not always visible. Ventral tubular ducts present in groups on head and around anogenital fold. Antennae each usually with 7 segments (but +8 in +Iranian specimens). Legs well-developed, each with a tibio-tarsal articulatory sclerosis; claw digitules both broad; each claw with an almost imperceptible denticle. + + + + +Distribution: + +Ceroplastes sinensis + +is a widespread species in warm temperate to subtropical countries; in +Iran +, it is found in +Gilan +and +Mazandaran +provinces ( +Moghaddam 2017 +). + + +Host-plants: +The scale is polyphagous, mostly on woody broad-leafed hosts. In +Iran +, it has been found on + +Punica granatum + +( +Lythraceae +), + +Rosa +sp. + +( +Rosaceae +) ( +Moghaddam 2017 +), + +Hedera helix + +( +Araliaceae +) ( +Feli Kohikheili & Damavandian 2015 +), and + +Citrus +spp. + +( +Rutaceae +). + + +Economic importance: +The species is a serious pest of + +Citrus + +trees in the northern provinces of +Iran +. + + +Natural enemies: +The following parasitoid wasps ( +Hymenoptera +) have been observed attacking + +C. sinensis + +in +Iran +: + +Aphytis hispanicus +(Mercet) + +and + +Coccophagus lycimnia +(Walker) + +( +Aphelinidae +); + +Microterys nietneri +(Motschulsky) + +( +Encyrtidae +), and + +Alaptus priesneri +Soyka + +( +Mymaridae +) ( +Feli Kohikheili & Damavandian 2015 +). + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FFB0396092DA91FA3788DFEA.xml b/data/98/1B/87/981B8798FFB0396092DA91FA3788DFEA.xml new file mode 100644 index 00000000000..730e44f291d --- /dev/null +++ b/data/98/1B/87/981B8798FFB0396092DA91FA3788DFEA.xml @@ -0,0 +1,184 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Ceroplastes rubens +Maskell + + + + + + + +( +Fig. 15 +, +Plate 2A +, distribution map +Fig. 89F +) + + + + + + + +Ceroplastes rubens +Maskell, 1893: 214 + + +. + + + +Field characteristics: +Live adult female body convex, up to +3.1 mm +long, covered with thick pinkish brown wax not divided into plates; dorsum with short, raised, narrow whiteish ridge leading to each stigmatic cleft. In dorsal view, the waxy test appears sub-rectangular or oval. The pink eggs are laid under the female body, protected by the waxy test. + + + + +Microscopic diagnosis: +Slide-mounted adult female body broadly oval and convex. Stigmatic clefts quite wide and deep. + + +Dorsum +. Derm membranous on young females, but with a rather flat, heavily sclerotized caudal process; with 1 cephalic, 1 mediodorsal, and 6 lateral clear areas devoid of pores and setae. Setae short, cylindrical, and with truncate apex. Pores of +2 types +: bilocular, and irregularly oval trilocular. Tubular ducts present throughout. Duct tubercles absent. Preopercular pores present in a group of about 9−15 pores. Anal plates each with 3 long flagellate dorsal setae. Anal ring with 8 setae. + + +Margin +. Marginal setae slightly longer than dorsal setae, except each anal lobe with 3−5 longer, sharply pointed setae. Stigmatic clefts quite deep, each with about 15–18 stigmatic setae along margin of each stigmatic cleft, each seta bollard-like, plus, more dorsally, 2 significantly larger bollard-like setae and with a much larger, bluntly pointed, spinose seta at apex of each group. + + + +FIGURE 15. +Adult female of + +Ceroplastes rubens +Maskell + +, reproduced from +Moghaddam & Nematian (2021) +page 179, fig. 2, with kind permission from the Editor of +Journal of Entomological Society of Iran +. + + + +Venter +. Derm entirely membranous. Pregenital disc-pores each with 10 loculi, restricted to around anogenital fold (segment VII). Spiracular disc-pores each with 5 loculi, present in broad bands between margin and each spiracle, and with a few extending medially past spiracular apodeme towards prothoracic coxa. Cruciform pores present at margins. Tubular ducts absent. Minute setae present; interantennal setae numbering at most 3 pairs; setae on pregenital segments small. Antennae quite short, each with 6 segments. Legs much reduced in size, with trochanter and femur occasionally appearing fused, lacking tibio-tarsal articulatory sclerosis; tibia and tarsus fused; claw digitules dissimilar, 1 slightly thicker and longer than other; tarsal digitules both narrow. + + + + +Distribution: + +Ceroplastes rubens + +is a cosmopolitan species of African origin; in +Iran +, it is found in +Tehran province +( +Moghaddam & Nematian 2021 +). + + +Host-plants: +The scale highly polyphagous, usually feeding on woody hosts and some +Cycadaceae +and +Pteridaceae +; in +Iran +, it has been recorded on + +Aglaonema commutatum + +( +Araceae +) ( +Moghaddam & Nematian 2021 +) and + +Ficus carica + +( +Moraceae +). + + +Economic importance: +It is highly likely that this species was introduced to +Iran +on plants that had not been properly inspected. At present, there are some reports of it causing serious damage to fig trees in +Iran +. + + +Natural enemies: +None recorded from +Iran +. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FFCE391A92DA910F35D3DF5A.xml b/data/98/1B/87/981B8798FFCE391A92DA910F35D3DF5A.xml new file mode 100644 index 00000000000..876ab4e8e21 --- /dev/null +++ b/data/98/1B/87/981B8798FFCE391A92DA910F35D3DF5A.xml @@ -0,0 +1,196 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Pulvinaria urbicola +Cockerell + + + + + + + +( +Fig. 35 +, distribution map +Fig. 93A +) + + + + + + + +Pulvinaria urbicola +Cockerell, 1893b: 160 + + +. + + + +Field characteristics: +Adult female elongate oval, yellow to yellowish green. Mature female convex. + + + + +Microscopic diagnosis: +Slide-mounted adult female body oval to broadly oval. Stigmatic clefts rather shallow. Anal cleft relatively short. + + +Dorsum +. Derm entirely membranous. Setae relatively long and spiniform, scattered throughout. Pores absent. Preopercular pores oval to circular, in group of 4−12 present anterior to anal plates. Submarginal duct tubercles usually absent but occasionally present in small numbers ( +Williams & Watson 1990 +). Tubular ducts scattered throughout. Anal plates together quadrate, each with 3 or 4 apical setae. Anal ring usually bearing 8 setae. + + +Margin +. Marginal setae slender, with parallel sides or curved, some bluntly pointed, some with fimbriate or bifid tips; with about 9 setae present between anterior and posterior stigmatic clefts on each side. Stigmatic clefts rather shallow, each containing 3 setae, with median seta about 2 or 3 times longer than lateral setae. + + + +FIGURE 35. +Adult female of + +Pulvinaria urbicola +Cockerell + +: tubular ducts of types (i), (ii) and (iii) shown; for details of their respective distributions, see also the diagnosis. + + + +Venter +. + +Derm membranous. Pregenital disc-pores mostly each with 7 loculi, present around anogenital fold, and on preceding 3 or 4 abdominal segments; a small group also present lateral to each metacoxa. Spiracular disc-pores each with 5 loculi, present in single to triple rows between each spiracle and margin. Microducts present in a submarginal band and scattered throughout. Tubular ducts of +3 types +: (i) a short duct, with short filamentous inner ductule and minute terminal gland, present in a broad submarginal band extending from anal cleft to posterior spiracular disc-pore band and with a few ducts also present on marginal area of head and between spiracular disc-pore band; (ii) a large duct, with fairly long and slender inner ductule and a well-developed terminal gland, present in submarginal areas; and (iii) a duct with a shallow cup-shaped invagination, very long slender inner ductule and a well-developed terminal gland, present medially on posterior abdominal segments. Legs well developed, each with freely articulated tibia and tarsus and with articulatory sclerosis; claw without a denticle; both claw digitules broad and shorter than thin tarsal digitules. Antennae each with 8 segments, with segments +II +and +III +subequal in length + +. + + + + +Distribution: + +Pulvinaria urbicola + +is a cosmopolitan species ( + +García Morales +et al +. 2016 + +); in +Iran +, it is found in Sistan & Balouchestan province. + + +Host-plants: +The species is polyphagous, having been recorded on host-plants belonging to 45 families ( + +García Morales +et al +. 2016 + +). In +Iran +, it has been found on + +Psidium guajava + +( +Myrtaceae +). + + +Economic importance: +No damage caused by this species has been recorded in +Iran +. + + +Natural enemies: +None recorded in +Iran +. + + + + +Comments: +This is the first record of + +P. urbicola + +from +Iran +. Please see the comments under + +P. camelicola + +above. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FFD1390192DA95E334A5DDBA.xml b/data/98/1B/87/981B8798FFD1390192DA95E334A5DDBA.xml new file mode 100644 index 00000000000..f4cae0947f9 --- /dev/null +++ b/data/98/1B/87/981B8798FFD1390192DA95E334A5DDBA.xml @@ -0,0 +1,196 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Pulvinaria aligarhensis +Avasthi & Shafee + + + + + + + +( +Fig. 31 +, distribution map +Fig. 92J +) + + + + + + + +Pulvinaria aligarhensis +Avasthi & Shafee, 1985: 1289 + + +. + + + +Field characteristics: +Live adult female oval, flat, grey with a longitudinal dark brown median patch. Mature adult female becomes enclosed in ovisac material. + + + + +Microscopic diagnosis: +Slide-mounted adult female body broadly oval. Stigmatic clefts relatively deep. Anal cleft of moderate depth. + + +Dorsum +. Slide mounted adult female broadly oval. Derm membranous; lacking dermal areolations. Setae small and lanceolate, scattered throughout. Preopercular pores each oval to circular, present anterior to anal plates and extending to level of mouthparts. Tubular ducts and duct tubercles absent. Anal plates together quadrate; each plate with 3 apical and 1 subdiscal setae dorsally. Anal ring with 6 setae present. + + +Margin +. Marginal setae slender, with apices knobbed or rounded, with about 15‒22 on each side between anterior and posterior stigmatic areas; setae on either side of anal cleft longer. Stigmatic cleft well developed; each spiracular cleft containing group of 3 or rarely 4 stigmatic setae; median stigmatic seta 2 or 3 times longer than lateral setae. Marginal setae slender with clubbed apices, except for a few long, simple setae on apex of each anal lobe (in Iranian specimens). + + +Venter +. + +Derm completely membranous. Multilocular disc-pores each with 7 or 10 loculi, numerous around anogenital fold and extending medially and submedially as far forward as head, where they also occur on the submargin. Quinquelocular spiracular pores present in a band between each spiracle and stigmatic cleft. Microducts scattered throughout (not illustrated). Tubular ducts of +3 types +: (i) a duct with fairly short outer ductule and short, filamentous inner ductule with a minute terminal gland, present in a broad submarginal band extending from anal cleft to near antennae; (ii) a duct rather similar to +type +(i) but with inner ductule broader and often longer than outer ductule, and with a large terminal gland, present medially in more posterior abdominal segments and laterally interspersed with +type +(i) ducts; and (iii) a larger duct, with a deeper cup-shaped invagination and inner ductule even broader than that in +type +(ii) and with a large terminal gland; this is the only duct +type +present medially on head, thorax and more anterior abdominal segments; they also extend laterally to meet submarginal band of +type +(i) and (ii) ducts, where they become infrequent or absent. Venter with setae of variable lengths, sparsely distributed; all abdominal segments each with a median pair of long setae. With 4 pairs of setae between antennal bases. Legs well developed, each with an articulation between tibia and tarsus, with a tibio-tarsal sclerosis; claw without a denticle; both claw digitules broad and shorter than thin tarsal digitules.Antennae each with mostly 8 segments, with segment +III +longest + +. + + + + +Distribution: + +Pulvinaria aligarhensis + +has only been recorded from +India +before; this is the first record from +Iran +, where it is found in Sistan & Balouchestan province. + + +Host-plants: + +Pulvinaria aligarhensis + +has been recorded only on + +Azadirachta indica + +( +Meliaceae +) ( + +García Morales +et al +. 2016 + +), including in +Iran +. + + + +FIGURE 31. +Adult female of + +Pulvinaria aligarhensis +Avasthi & Shafee + +, based on an Iranian specimen, with enlargements of tubular ducts of types (i), (ii) and (iii) shown; for details of their respective distributions, see the diagnosis. + + + +Economic importance: +Not of any economic importance in +Iran +. + + +Natural enemies: +None recorded in +Iran +. + + + + +Comments: + +Pulvinaria aligarhensis + +is recorded here for the first time in +Iran +. However, there are some differences between Indian and Iranian specimens (character states of Indian specimens in parenthesis): (i) pregenital disc-pores exceptionally widespread, present around anogenital fold and medially and submedially as far forward as head, where they also occur on the submargin (numerous anterior to anal fold and extending forward medially as far as mouthparts); (ii) marginal setae blunt, some with knobbed apices (all with knobbed apices); and (iii) long marginal setae at apex of each anal lobe numbering 3 or 4 (numbering only 1). These differences are regarded as intraspecific variation. + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FFD2391D92DA91A631A2DD96.xml b/data/98/1B/87/981B8798FFD2391D92DA91A631A2DD96.xml new file mode 100644 index 00000000000..35ffa252f33 --- /dev/null +++ b/data/98/1B/87/981B8798FFD2391D92DA91A631A2DD96.xml @@ -0,0 +1,303 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Pulvinaria camelicola +Signoret + + + + + + + +( +Fig. 33 +) + + + + + + + +Pulvinaria camelicola +Signoret, 1873b: 32 + + +. + + + +Field characteristics: +Live adult female elongate oval, slightly convex; body yellow with small reddish spots, quite similar to + +P. floccifera +( +Tanaka & Kamitani 2022 +) + +. Mature adult female produces a relatively long ovisac. + + + + +FIGURE 33. +Adult female of + +Pulvinaria camelicola +Signoret + +, from +Tanaka & Kamitani (2022) +page 441, fig. 1. © Magnolia Press, www.mapress.com/j/zt, reproduced with permission from the authors and the copyright holder. Distributions of tubular duct types (i), (ii) and (iii) shown; for details of their respective distributions, see also the diagnosis. + + + + +Microscopic diagnosis: +Slide-mounted adult female body elongate oval. Stigmatic clefts rather shallow. Anal cleft of moderate depth. + + +Dorsum +. Derm entirely membranous; dermal areolations well developed. Setae spiniform, scattered throughout. Preopercular pores oval to circular, often difficult to see, present anterior to anal plates. Submarginal dorsal tubercles present, numbering 0−4 on each side. Tubular ducts frequent throughout, each situated within an areolation. Anal plates together quadrate, each plate with 3 or 4 apical setae. Anal ring usually bearing 5−7 setae. + + +Margin +. Marginal setae with well-developed basal sockets and with tips fimbriate, frayed, spatulate, split, or simply pointed, those on either side of anal cleft tending to be longer; each side with 12–25 setae between anterior and posterior stigmatic clefts. Stigmatic clefts distinct but shallow, each containing 3 setae, median seta about 2−4 times longer than lateral setae. + + +Venter +. + +Derm membranous. Pregenital disc-pores mostly each with 5−8 loculi, present around genital opening, and on medial areas of preceding 3 to 5 abdominal segments; a small group also present lateral to each metacoxa and occasionally to each mesocoxa also. Spiracular disc-pores each with 5 loculi, present in a narrow band between each spiracle and margin. Microducts scattered throughout venter. Tubular ducts of +3 types +: +Type +I with a large outer ductule and a broad inner ductule ending in a well-developed flower-shaped terminal gland: present in posterior medial area of head, medial area of all thoracic segments and anterior abdominal segments, also in inner submarginal area extending from near anterior abdominal segments to prothoracic segments; +Type +II +with a rather narrower outer ductule ending in a shallow cup-shaped invagination, and a long, slender inner ductule with a well-developed terminal gland, present medially on posterior abdominal segments; and submarginal band of tubular ducts but reaching only as far forward as posterior spiracular pore band; and +Type +III +with a short, filamentous inner ductule and minute terminal gland, present in a broad submarginal band on thoracic and abdominal segments, but mostly concentrated on posterior area, intermixed with +Type +I or +Type +II +ducts. Posterior 3 abdominal segments each with 1 pair of long ventral setae on medial area. With 2–6 pairs of long setae present between antennal bases; other setae short and fine, distributed over entire venter. Legs well developed, each with an articulated tibio-tarsal joint and an articulatory sclerosis; claw without a denticle; both claw digitules rather broad and slightly shorter than thin tarsal digitules. Antennae each 6–8 segmented ( +Tanaka & Kamitani 2022 +) + +. + + + + +Distribution: + +Pulvinaria camelicola + +has been reported from 54 countries in five zoogeograpical regions ( + +García Morales +et al +. 2016 + +), despite it not having been well described until very recently ( +Tanaka & Kamitani, 2022 +). + + +Host-plants: + +Pulvinaria camelicola + +has been recorded on host-plants in 62 genera belonging to 44 families ( + +García Morales +et al +. 2016 + +). The host-plant in +Iran +was not recorded by +Bodenheimer (1944b) +. + + +Economic importance: +Not recorded in +Iran +. + + +Natural enemies: +None recorded in +Iran + + + + +Comments: +Bodenheimer (1944b) +recorded + +P. camelicola + +from +Iran +, but neither his material nor any other specimens from +Iran +have been seen in the present study so it has not been possible to confirm its presence in the country. It is possible that +Bodenheimer’s (1944b) +record may have been a misidentification of + +P. floccifera + +. + + +In the past, several little-studied species (including + +P. camelicola + +) have been misidentified as the well-known polyphagous pest species, + +P. floccifera + +, because its original description is too brief and subsequent redescriptions were based on misidentified material. This situation subjectively invalidates the identity of + +P. floccifera + +because several species have been studied under the same (inappropriate) scientific name in the absence of +type +fixation, leading to taxonomic confusion ( + +García Morales +et al. +2016 + +). + + + +Tanaka & +Amano +(2007) + +found that the morphology of +five adult +female +syntypes +of + +P. floccifera + +did not correspond with the descriptions of several of the species that were considered synonyms. To confer taxonomic stability to + +P. floccifera + +, in accordance with the provisions of Article 23.3.5 of the Fourth edition of the International Code of Zoological Nomenclature ( +ICZN 1999 +), its oldest synonym ( + +P. camelicola + +) was revived and redescribed by Tanaka & Kanitani (2022) based on Japanese specimens of a distinct species hitherto misidentified as + +P. floccifera + +. However, they could not determine which other species had been misidentified as + +P. floccifera + +because of the difficulty of obtaining the relevant type specimens for study. Aspects of this taxonomic confusion therefore have not yet been resolved and molecular studies are needed to further elucidate the species boundaries. + + +As presently understood, + +P. camelicola + +is close to + +P. floccifera + +and + +P. urbicola + +in having multilocular disc-pores each with 6‒8 loculi and similarly distributed ventral tubular ducts, each with a filamentous inner ductule and lacking a terminal gland ( +Tanaka & Kamitani 2022 +). + +Pulvinaria camelicola + +differs from these species by having (contrasting character states of + +P. floccifera + +and + +P. urbicola + +given in parenthesis): (1) large, well-developed dorsal areolations (areolations rarely present, small and restricted to submarginal areas); and (2) dorsum with a small number of tubular ducts (dorsum mostly lacking tubular ducts) ( +Tanaka & Kamitani 2022 +). + + + + \ No newline at end of file diff --git a/data/98/1B/87/981B8798FFF0392292DA944E37A8DA63.xml b/data/98/1B/87/981B8798FFF0392292DA944E37A8DA63.xml new file mode 100644 index 00000000000..1f0670d349b --- /dev/null +++ b/data/98/1B/87/981B8798FFF0392292DA944E37A8DA63.xml @@ -0,0 +1,168 @@ + + + +The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families + + + +Author + +Moghaddam, Masumeh +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. + + + +Author + +Watson, Gillian W. +0000-0001-9914-0094 +Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. +gillian.watson@nhm.ac.uk + +text + + +Zootaxa + + +2024 + +2024-11-29 + + +5542 + + +1 + + +1 +202 + + + + +https://doi.org/10.11646/zootaxa.5542.1.1 + +journal article +10.11646/zootaxa.5542.1.1 +1175-5334 +2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16 + + + + + + + +Acanthococcus aceris +Signoret + + + + + + + +( +Fig. 48 +, distribution map +Fig. 88A +) + + + + + + + +Acanthococcus aceris +Signoret, 1875: 35‒36 + + +. + + + +Field characteristics: +Body of live adult female oval, chestnut brown, with surface covered by compact, greyish felted ovisac. + + + + +Microscopic diagnosis: +Slide-mounted adult female oval, tapering posteriorly. Antennae each with 6 or 7 segments. Frontal lobes well developed, and frontal tubercles present. Legs well developed; meso- and metathoracic coxae each with spinulae on ventral surface; translucent pores absent; claw and tarsal digitules all with apical knobs; claws each with denticle. Anal lobes strongly developed and sclerotized, frequently inner margins with tooth-like projections; each lobe with 3 enlarged setae and 3–5 microtubular ducts on dorsal surface, and ventrally with 2 flagellate setae. Anal ring strongly sclerotized, bearing 8 setae and pores. Cauda triangular, slightly sclerotized. + + +Dorsum +with spinose setae of 2 sizes: (i) marginal enlarged setae conical, long (mostly more than 2x as long as a dorsal enlarged seta) and blunt; and (ii) other dorsal setae shorter, conical and blunt, present in transverse segmental rows; medial area of abdominal segment VIII with 6–8 enlarged setae. Enlarged tubular ducts absent. Macrotubular ducts present throughout. Microtubular ducts long, scattered. + + +Venter +with flagellate setae in median areas, lateral areas with spine-like setae. Apical labial segment with 5 pairs of hair-like setae, median (apical) pair somewhat shorter than others. Disc-pores present in sparse bands across all abdominal segments, also scattered on thorax and head. Macrotubular ducts of 3 sizes: (i) smallest ducts situated on median areas of abdominal segments; (ii) medium-sized ducts present on median areas of abdominal segments; and (iii) largest ducts present in a submarginal band on abdomen, thorax and head. Microtubular ducts absent. Cruciform pores generally forming a submarginal band on thorax and abdominal segments, and a few present on head. + + + + +Distribution: + +Acanthococcus aceris + +is a Palaearctic species, recorded from 24 countries including +Iran +, +Khuzestan province +( + +García Morales +et al +. 2016 + +, + +Rozedar +et al +. 2013 + +). + + +Host-plants: +The scale is quite polyphagous, having been recorded on host-plants belonging to 13 genera in 10 families ( + +García Morales +et al +. 2016 + +); in +Iran +, it was found on + +Citrus +spp. + +( +Rutaceae +) ( + +Rozedar +et al +. 2013 + +). + + +Economic importance: +Not known as a pest in +Iran +. + + +Natural enemies: +None recorded in +Iran +. + + + + \ No newline at end of file diff --git a/data/9F/07/87/9F07878AA865CD7C19AADC599BAE5B55.xml b/data/9F/07/87/9F07878AA865CD7C19AADC599BAE5B55.xml new file mode 100644 index 00000000000..444058461fa --- /dev/null +++ b/data/9F/07/87/9F07878AA865CD7C19AADC599BAE5B55.xml @@ -0,0 +1,714 @@ + + + +Comparative phylogeography of bamboo bats of the genus Tylonycteris (Chiroptera, Vespertilionidae) in Southeast Asia + + + +Author + +Tu, Vuong Tan + + + +Author + +Csorba, Gábor + + + +Author + +Ruedi, Manuel + + + +Author + +Furey, Neil M. + + + +Author + +Son, Nguyen Truong + + + +Author + +Thong, Vu Dinh + + + +Author + +Bonillo, Céline + + + +Author + +Hassanin, Alexandre + +text + + +European Journal of Taxonomy + + +2017 + +2017-02-09 + + +274 + + +274 + + +1 +38 + + + + +https://doi.org/10.5852/ejt.2017.274 + +journal article +10.5852/ejt.2017.274 +d7506d3e-5d6b-4914-9de0-208a44712770 +2118-9773 +291919 +urn:lsid:zoobank.org:pub:DEFAD552-9C2E-497B-83CA-1E04E3353EA4 + + + + + + +Cryptic species diversity in the genus + +Tylonycteris + + + + + + + +Previous studies have detected high levels of genetic and karyological variation among specimens identified as either + +T. pachypus + +or + +T. robustula + +collected from different geographic locations in mainland Southeast Asia ( + +Francis +et al. +2010 + +; + +Huang +et al. +2014 + +). Our +COI +analyses further revealed the existence of three divergent geographic haplogroups for both + +T. pachypus + +and + +T. robustula + +, for each corresponding to Sumatra, northern Indochina and southern Indochina (but also northwestern India and Peninsular Malaysia for + +T. robustula + +). Our +Cytb +dataset confirmed the existence of these geographic haplogroups in mainland Southeast Asia. In addition, a specimen of + +T. pachypus + +collected from Borneo and originally described as + +T. robustula + +was found to be highly divergent from the two other Indochinese haplogroups ( +Fig. 2 +). In the absence of +Cytb +data for Sumatran specimens, it is impossible to know whether they share the same mitochondrial lineage as those from Borneo. + + +Taken together, our mtDNA analyses show that all haplotypes sequenced for insular + +Tylonycteris + +are very divergent from those identified in mainland Southeast Asia. However, genetic inferences based on the maternally inherited mitochondrial genes are prone to be discordant with the true evolutionary history of the taxa, owing to various evolutionary processes, such as mtDNA introgression, incomplete lineage sorting or female philopatry ( +Avise 2000 +; +Ballard & Whitlock 2004 +; +Hassanin & Ropiquet 2007 +; + +Nesi +et al. +2011 + +; + +Rivers +et al. +2005 + +). Here, the geographic pattern of mtDNA diversity observed for the two species of + +Tylonycteris + +could be the consequence of female philopatry, i.e., the behavior of remaining in, or returning to the natal territory. Indeed, bat species with philopatric females generally display high geographic structure when relationships are examined with maternally inherited markers, such as the mitochondrial DNA. This pattern can disappear with biparentally inherited markers when adult males are able to disperse over long distances, allowing gene flow between otherwise isolated populations ( + +Castella +et al. +2001 + +; + +Hassanin +et al. +2015 + +; + +Hulva +et al. +2010 + +; + +Pereira +et al. +2009 + +; + +Rivers +et al. +2005 + +). Behavioral and population genetic studies in southern China have shown that + +T. pachypus + +bats are philopatric to their natal area and that philopatry is especially pronounced in females ( + +Hua +et al. +2011 + +, +2013 +). Although no data are available for + +T. robustula + +, female philopatry can also be predicted for this species, because it shares similar morphological, behavioural and ecological traits with + +T. pachypus + +( +Medway 1972 +; +Medway & Marshall 1970 +, +1972 +; + +Zhang +et al. +2007 + +). The social organization of + +T. pachypus + +and + +T. robustula + +, combined with their fragmented habitats, is therefore expected to result in limited gene flow between populations, especially among matrilines from distant geographic localities. For both species, this prediction is corroborated by the analyses of mtDNA markers, with the identification of three divergent geographically non-overlapping haplogroups. For + +T. robustula + +, this phylogeographic pattern is also supported by the nuclear sequence data, as the two Indochinese clades were recovered monophyletic with all the three introns containing enough nucleotide variation at the intra-specific level, i.e., +CHPF2 +, +HDAC1 +and +TUFM +( +Appendix 8 +). By contrast, our nuclear analyses ( +Fig. 3 +; Appendices 4, 8) do not support the reciprocal monophyly of the two Indochinese clades of + +T. pachypus + +, suggesting that gene flow was maintained by male dispersal or, alternatively, that their separation was too recent to be detected with our nuclear genes. + + +Nucleotide distances estimated from mtDNA genes between northern and southern Indochinese populations of + +T. robustula + +were more than twice those of + +T. pachypus + +(6.5% vs 2.8% in +COI +; 9.5% vs 2.8% in +Cytb +), indeed supporting a more recent divergence for the latter taxon, if we assume equal evolutionary rates. Similarly, the nuclear distances between the two Indochinese clades of + +T. robustula + +were between 0.41 and 0.56%, which is more than twice those calculated between Indochinese individuals of + +T. pachypus + +(0–0.2%; +Appendix 5 +) and in the range of interspecific distances found in other groups of Laurasiatheria, such as fruit bats of the tribes +Myonycterini +( + +Nesi +et al +. 2013 + +) and Scotonycterini ( + +Hassanin +et al +. 2015 + +), or cattle and buffalo of the tribe Bovini ( + +Hassanin +et al +. 2013 + +). Although none of the nuclear markers could be sequenced for Sumatran and Bornean + +Tylonycteris + +, their high mtDNA divergence from Indochinese populations (> 5.7 % in both +COI +and +Cytb +sequences; +Appendix 5 +) suggests they might represent distinct lineages based on nuclear markers as well. In agreement with this view, our multivariate morphological analyses revealed that Indochinese bats of the + +T. pachypus + +complex constitute a distinct group separated from those collected on Sumatra. For the + +T. robustula + +complex, morphological overlap between haplogroups is more extensive, but pairwise comparisons of their PC mean scores support the distinctness of adjacent geographical taxa, such as Tr2 and Tr3 on the Southeast Asian mainland. + + +The close morphological similarity among taxa of + +Tylonycteris + +suggests that they have evolved under the influences of similar and specialized habitats, i.e., woody bamboo vegetation. Molecular evidence indicates, however, that + +T. pachypus + +should be split into at least two distinct species, + +T. pachypus + +on the Sunda islands (Sumatra and/or Borneo) and + +T. fulvida + +in mainland Southeast Asia, and that + +T. robustula + +should be divided into at least three species, with + +T. robustula + +on Sumatra, + +T. malayana + +in southern and western mainland Southeast Asia, and + +T. tonkinensis + +sp. nov. +in northern Indochina. + + + +The evolution of +Tylonyteris +spp. in Southeast Asia during the Pleistocene + + + +Given that both species complexes, here named + +T. pachypus + +s. lat. +and + +T. robustula + +s. lat. +, are usually found in sympatry across most of their geographic ranges in Southeast Asia, they are expected to share a common phylogeographical history. Our estimates of divergence times based on mtDNA sequences suggest that the genus + +Tylonycteris + +diversified during the Pliocene epoch ( +Cytb +: 5.92 ± 0.65 Mya; +COI +: 4.56 ± 0.72 Mya) ( +Table 1 +). During the Miocene and until the early Pliocene, Southeast Asia was generally covered by large tracks of rain forests as a consequence of warm and humid climatic conditions ( +Meijaard & Groves 2006 +; +Morley 2000 +). Thus, ancestors of both + +Tylonycteris + +species complexes were presumably widely distributed across Southeast Asia during the Pliocene. + + +Our molecular dating estimates indicate that the basal geographic splits within the two species complexes, i.e., between mainland Southeast Asia and Sumatra, took place approximately at the same time during the Early Pleistocene (between 2.70 and 1.96 Mya for + +T. pachypus + +, between 3.07 and 2.22 Mya for + +T. robustula + +; +Table 1 +). The Pleistocene epoch is characterized by the onset of repeated cycles of cold glacial and warm interglacial periods as the results of the glaciations/deglaciations of the Northern Hemisphere, which implied contraction and expansion of rain forests in Asia ( + +An +et al. +2001 + +; +Meijaard & Groves 2006 +; +Morley 2000 +). As bats of the genus + +Tylonycteris + +are highly dependent on woody bamboo vegetation for roosting, foraging and mating ( +Kunz 1982 +; +Medway 1972 +; +Medway & Marshall 1970 +, +1972 +), their Pleistocene biogeographic history was firmly constrained by the distribution of such bamboo habitats. The current distribution of woody bamboo species in Asia ( + +Bystriakova +et al. +2003b + +; +Fig. 6 +) indicates that eight disjunctive biogeographic regions have higher species richness (> 5 species) for some bamboo genera: southern India ( + +Ochlandra +Thwaites + +), northern Myanmar ( + +Cephalostachyum +Munro + +), southern China ( + +Dendrocalamus +Nees + +and + +Bambusa +Schreb. + +), Hainan Island ( + +Bambusa + +), northwestern Thailand ( + +Dendrocalamus + +and + +Gigantochloa +Kurz ex Munro + +), Peninsular Malaysia ( + +Gigantochloa + +), Sumatra and Borneo ( + +Gigantochloa + +and + +Schizostachyum +Nees + +). All these regions may therefore have acted as distinct bamboo forest refugia during the glacial periods of the Pleistocene ( +Fig. 6 +). Evidence for a number of these postulated glacial refugia has been reported in previous studies for many organisms, including bats ( + +Flanders +et al. +2011 + +; + +Khan +et al. +2010 + +; + +Lin +et al. +2014 + +; + +Mao +et al. +2013 + +). Accordingly, we propose that the contraction of woody bamboo forests into different glacial refugia had fragmented the distribution of the Pliocene ancestors of both + +T. pachypus + +s. lat. +and + +T. robustula + +s. lat. +In addition, we can assume that Pleistocene glacial periods resulted in higher interspecific competition between co-distributed species of + +Tylonycteris + +, because the supply of most suitable resources was more limited in glacial bamboo forest refugia ( +Medway & Marshall 1970 +). As noted in previous studies, + +T. pachypus + +s. lat. +has a more manoeuvrable flight in cluttered habitats and forages on smaller insects than + +T. robustula + +s. lat. +( + +Zhang +et al. +2005 + +, +2007 +). Moreover, +Medway & Marshall (1970) +found that the smaller + +T. pachypus + +s. lat. +can roost in the internodes with small entrance holes, which the larger + +T. robustula + +s. lat. +is unable to enter. These differences suggest, therefore, that the smaller + +T. pachypus + +s. lat. +have greater advantages than the larger + +T. robustula + +s. lat. +in interspecific competition when natural resources are limited. Hence, isolated populations of + +T. robustula + +s. lat. +may have been more exposed to bottlenecks and therefore more vulnerable to local extinction than those of co-distributed + +T. pachypus + +s. lat. + + +During interglacial periods of the Early Pleistocene, warmer and humid conditions resulted in the expansion of woody bamboo forests, which in turn may have favored the restoration of connectivity between isolated populations of both complexes. However, the isolated populations may have been connected or not, depending on their dispersal capacity and the distances between refugia. In + +T. robustula + +s. lat. +, these processes may have taken longer, because of its lower population abundance ( +Lande & Barrowclough 1987 +; +Shaffer 1981 +), and may have been prevented in cases of extinction of transitional populations ( + +Huang +et al. +2014 + +and references therein). This scenario is supported by the fact that + +T. pachypus + +s. lat. +is usually found to be more abundant than + +T. robustula + +s. lat. +in bamboo forests ( + +Zhang +et al. +2004 + +; +Medway & Marshall 1972 +) and by the wider geographic range of + +T. pachypus + +s. lat. +( + +Bates +et al. +2008a + +, +2008b +; +Fig. 1 +). Knowing this, the body size differences between the two species complexes may be the key factor explaining why the basal divergence of northern Indochinese populations occurred earlier in + +T. robustula + +s. lat. +(i.e., + +T. tonkinensis + +sp. nov +) than in + +T. pachypus + +s. lat. +, i.e., 2.97–1.70 vs 1.35–0.79 Mya ( +Table 1 +). During Pleistocene interglacials, exchanges of + +Tylonycteris + +spp. between the continent and the islands of Sundaland were probably prevented because of the long distances between the glacial forest refugia, as well as the higher sea levels ( +Fig. 1 +). Our molecular dating estimates corroborate this scenario, as continental populations of + +Tylonycteris + +spp. from Indochina and Peninsular Malaysia diverged from insular populations (Sumatra and Borneo) in the Early Pleistocene ( +Table 1 +). + + + + + +Implications for conservation + + + +Previous studies considered + +T. pachypus + +and + +T. robustula + +to be common species and thus classified them as +Least Concern +in the IUCN Red List ( + +Bates +et al. +2008a + +, +2008b +). Since our study reveals that both species in fact represent several species with more restricted distributions, the IUCN status of the different taxa should be reassessed urgently, including that of the new species, + +T. tonkinensis + +sp. nov. +In addition, our study suggests that several biogeographic regions have acted as Pleistocene glacial refugia. This information is very important for developing more effective conservation strategies, particularly given the high current rates of deforestation affecting most natural habitats in Southeast Asia ( +Kingston 2010 +; + +Sodhi +et al. +2010 + +; + +Tordoff +et al. +2012 + +). + + + + \ No newline at end of file diff --git a/data/9F/07/87/9F07878AA866CD601888DE649C175E02.xml b/data/9F/07/87/9F07878AA866CD601888DE649C175E02.xml new file mode 100644 index 00000000000..b2da7a3c1a3 --- /dev/null +++ b/data/9F/07/87/9F07878AA866CD601888DE649C175E02.xml @@ -0,0 +1,602 @@ + + + +Comparative phylogeography of bamboo bats of the genus Tylonycteris (Chiroptera, Vespertilionidae) in Southeast Asia + + + +Author + +Tu, Vuong Tan + + + +Author + +Csorba, Gábor + + + +Author + +Ruedi, Manuel + + + +Author + +Furey, Neil M. + + + +Author + +Son, Nguyen Truong + + + +Author + +Thong, Vu Dinh + + + +Author + +Bonillo, Céline + + + +Author + +Hassanin, Alexandre + +text + + +European Journal of Taxonomy + + +2017 + +2017-02-09 + + +274 + + +274 + + +1 +38 + + + + +https://doi.org/10.5852/ejt.2017.274 + +journal article +10.5852/ejt.2017.274 +d7506d3e-5d6b-4914-9de0-208a44712770 +2118-9773 +291919 +urn:lsid:zoobank.org:pub:DEFAD552-9C2E-497B-83CA-1E04E3353EA4 + + + + + +Tylonycteris tonkinensis +Tu, Csorba, Ruedi & Hassanin + +sp. nov. + + + +urn:lsid:zoobank.org:act:C59B0774-79D6-4A84-9489-CF04BE35FC49 + +Fig. 5 +B + + + + + +Tylonycteris robustula +Thomas, 1915 + +(partim): 227. + + + +Tylonycteris robustula + +– +Osgood 1932: 236 +. — Tate 1942: 268. — + +Hendrichsen +et al. +2001: 90 + +. — +Kruskop 2013: 221 +. — + +Thomas +et al. +2013: 229 + +. + + + + + +Etymology + + + +The specific epithet refers to the current restricted occurrence of the new species in north-eastern Laos and northern Vietnam ( +Fig. 1 +). The Vietnamese portion of this region was previously called “Tonkin” during the Nguyễn dynasty and French colonial era (from the 19th to the mid-20th centuries) to separate it from the country’s centre (Annam) and southern regions (Cochinchina). The proposed English name is “Tonkin’s greater bamboo bat” and the proposed Vietnamese name is ‘Dơi ống tre Bắc Bộ’. + + + + +Table 3. +Factor loadings of craniodental characters for the PCs obtained from principle component analyses (PCAs) of specimens of + +Tylonycteris + +. Acronyms and definitions for measurements are given in Material and methods. The first PCA is based on raw data (PC*1), while the second PCA is based on the log-transformed, standardized data (PC1 and PC2). Values in bold indicate the most significant loadings. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
+Raw data +Standardized +data +
+Characters + +PC*1 + +PC1 + +PC2 +
GLS +0.25 +0.140.04
CCL +0.23 +0.22-0.02
CC +0.36 +-0.180.25
UCI +0.41 + +-0.62 + +-0.38 +
M3M3 +0.32 +-0.020.30
IC +0.34 +-0.15 +-0.62 +
MB0.17 +0.47 +-0.10
BW0.19 +0.43 +-0.28
CM3 +0.35 +-0.220.29
ML +0.27 +0.090.16
CM3 +0.34 +-0.17 +0.35 +
Eigenvalue0.00820.00060.0003
% variance86.937.219.3
+ + + + +Type material + + + + + + +Holotype + + + + +VIETNAM +: +♂ +, +Copia Nature Reserve +, +Co Ma commune +, +Thuan Chau District +, +Son La Province +, +21°21.727′ N +, +103°30.562′ E +, + +1286 m + +a.s.l., + +9 May 2011 + +, +Vuong Tan Tu +leg. ( +IEBR-VN11-0055 +; field number Tu.090511.3; tissue code VN11-0055). Body in alcohol, skull removed. Mass: +4 g +. Measurements (in mm): FA: 27.0; HB: 41.0; Tail: 33.0; Ear: 11.0; GLS: 12.60; CCL: 11.59; UCI: 1.98; CC: 4.06; M3M3: 5.46; IC: 3.97; MB: 7.42; BW: 7.07; CM3: 3.96; ML: 8.64; CM3: 4.22. Accession numbers of mitochondrial and nuclear sequences: +KX496422–KX496429 +. + + + + + + +Paratypes + + + + +LAOS +: +3 adult +♂♂ +( +MHNG 1926.059 +, +MNHN 2006-90 +, +MNHN 2006-93 +), + +1 adult +♀ +( +MHNG 1926.057 +), +Hat Hin, Nam Sing River +, +Phongsaly Province +, +21°40.14′ N +, +102°13.26′ E +, 2004, +Manuel Ruedi +leg., body in alcohol, skull removed. + + + + + +VIETNAM +: +1 ♂ +( +IEBR-VN11-1804 +), +Hang Kia, Pa Co Nature Reserve +, +Hoa Binh Province +, +20°44.910′ N +, +104°54.900′ E +, + +1080 m + +a.s.l., 2012, +Vuong Tan Tu +leg., body in alcohol, skull removed. + + + +Accession numbers of DNA sequences for paratypes are given in +Appendix 1 +. + + + +Referred material + + + +Specimens identified as + +T. robustula + +collected from Na Don, Phuong Vien, Cho Don (Bac Kan Province) and Na Hang Nature Reserve (Tuyen Quang Province) ( +Appendix 1 +) are also referred to + +T. tonkinensis + +sp. nov. + + + + + +Description + + + +A member of the + +T. robustula + +species complex comprising representatives of the Tr3 haplogroup found in northern Indochina. Externally, individuals are small, with a forearm length of 25.1–27.8 mm ( +Table 2 +). The head is dorsoventrally very flattened. Pelage coloration is relatively variable, more or less golden red at the base of the dorsal fur, to dark brown near the tips of the dorsal hairs, and lighter golden brown on the underparts ( +Fig. 5 +B). The ears have a triangular shape, with broadly rounded tips. The tragus is short and blunt. The wing membranes are dark brown. The base of thumbs and soles of hind feet have fleshy pads ( +Fig. 5 +B). + + +The skull is small (GLS: 11.91–12.60 mm), lightly built and very flat ( +Fig. 5 +B). The rostrum is short. The sagittal crest is absent. The lambdoid crests are well developed. The dental formula is I2/3 C1/1, P1/2, M3/3 = 32. The first upper incisor (I2) is bicuspidate, with small cusps on cingulum. I3 is unicuspidate, about half the height and crown area of I2. The upper canine has a posterior supplementary cusp. A diastema between I3 and the upper canine is clearly visible. The protocones of M1 and M2 are welldeveloped. M2 appreciably exceeds M1 in width, and its width clearly exceeds its length. M3 is relatively smaller and without a metastyle. The three lower incisors are tricuspidate. The first (PM 2) and second (PM4) premolars are approximately equal in height and crown area ( +Fig. 5 +B). + + + + +Fig. 5. +Morphological characteristics of the two nominal species of the genus + +Tylonycteris +Peters, 1872 + +. +A +. + +T. pachypus +(Temminck, 1840) + +(corrected taxon name is + +T. fulvida +(Blyth, 1859)) + +, IEBR-VN11- 0 0 15. +B +. + +T. robustula +Thomas, 1915 + +(corrected taxon name is + +T. tonkinensis +Tu, Csorba, Ruedi & Hassanin + +sp. nov. +), holotype, IEBR-VN11-0055. Head profiles, ventral and dorsal views, fleshy pads at the base of the thumb and on the sole of the foot, and different views of the skull (dorsal, ventral and lateral). Scale = 10 mm. + + + + + +Remarks + + + +In northern Indochina, + +T. tonkinensis + +sp. nov. +can be distinguished from + +T. fulvida + +and + +T. pygmaea + +by its significantly larger body and skull size ( +Table 2 +; +Figs 4–5 +; see + +Feng +et al +. 2008 + +for comparisons with + +T. pygmaea + +), by K2P distances of at least 12% for +Cytb +and +COI +sequences and by K2P distances of at least 1.5% for the concatenation of the seven nuclear genes (5604 nt) ( +Appendix 5 +). Within the + +T. robustula + +complex, + +T. tonkinensis + +sp. nov. +is morphologically overlapping with + +T. robustula + +, found in Sumatra, and + +T. malayana + +(= Tr2 haplogroup), collected from the Southeast Asian mainland, but differs from the first taxon by K2P distances of at least 5.2% in +COI +sequences ( +Appendix 5 +) and from the latter by K2P distances of at least 5.5%, 8.6% and 0.4% calculated from +COI +sequences (657 nt), +Cytb +sequences (1140 nt) and the concatenation of seven nuclear DNA sequences (5604 nt), respectively ( +Appendix 5 +). + + + + + +Ecology and habitat + + + +Like other species of + +Tylonycteris + +, + +T. tonkinensis + +sp. nov. +is associated with woody bamboo groves. The new species is usually found in sympatry with the smaller species + +T. fulvida + +( +Fig. 1 +). In northwestern Vietnam, bats of the new species were captured in mist-nests set near bamboo groves in forest edges adjacent to rural-residential areas at relatively high elevations, e.g., at 1010 m a.s.l in Hang Kia, Pa Co Nature Reserve (Hoa Binh Province) or at 1286 m a.s.l in its type locality. In Laos, the known localities are found at lower elevations, between 500 and 800 m a.s.l. + + + + + +Distribution + + + +Currently, the new species is known to occur in north-eastern Laos and northern Vietnam only ( +Fig. 1 +). + + + + \ No newline at end of file