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<document id="CF5A94C5252363BEC13C182C892FE4E1" ID-DOI="10.3897/zookeys.1210.123986" ID-ZooBank="6972216E-2356-4DF7-BD24-2B6B68E639B4" ID-publisher-id="123986" URI-arpha="B6184626-6862-59C4-9047-EF9B7AC24E35" URI-zoobank="http://zoobank.org/6972216E-2356-4DF7-BD24-2B6B68E639B4" article-type="research-article" checkinTime="1723830749198" checkinUser="pensoft" docAuthor="Xu, Xin, Zhan, Yi, Aung, Khin Pyae Pyae, Liu, Fengxiang, Yu, Long &amp; Li, Daiqin" docDate="2024" docId="D0DEE7F143A9512D89942689CD66F8DD" docLanguage="en" docName="ZooKeys 1210: 99-115" docOrigin="ZooKeys 1210" docSource="https://zookeys.pensoft.net/article/123986/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Liphistius kalaw Zhan &amp; Xu 2024, sp. nov." docType="treatment" docVersion="1" dtd-version="3.0" lastPageNumber="115" masterDocId="B6184626686259C49047EF9B7AC24E35" masterDocTitle="Four new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae, Liphistiinae) from Myanmar" masterLastPageNumber="115" masterPageNumber="99" pageNumber="99" updateTime="1723830805509" updateUser="pensoft">
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<mods:title id="D515C10FD4791914DE53061C16006BDA">Four new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae, Liphistiinae) from Myanmar</mods:title>
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<mods:namePart id="AD7EBCDFE08B032E1AA2D7E6F0252D21">Xu, Xin</mods:namePart>
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<mods:affiliation id="1144392E72BBC643158D0A63F29BBE9A">College of Life Sciences, Hunan Normal University, 36 Lushan Road, Changsha 410081, Hunan Province, China &amp; State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
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<mods:namePart id="023D6D00C0F13EDDA671FCD58E6093F5">Zhan, Yi</mods:namePart>
<mods:affiliation id="49CE9E82610F06DAE9198D98471DE474">College of Life Sciences, Hunan Normal University, 36 Lushan Road, Changsha 410081, Hunan Province, China</mods:affiliation>
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<mods:namePart id="FFA7A8783EC347B3F86F72ED7EEF98B4">Aung, Khin Pyae Pyae</mods:namePart>
<mods:affiliation id="FE1DBB220C48910E42987D773916F11D">Department of Zoology, University of Yangon, Kamayut Township, Pyay Road, Yangon, 11041, Myanmar &amp; Department of Biology, Taungoo Education Degree College, Taungoo, 08101, Myanmar</mods:affiliation>
</mods:name>
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<mods:namePart id="F36F0FD22BCCAEF84A8E6AD7379C3DCE">Liu, Fengxiang</mods:namePart>
<mods:affiliation id="A1E883A191335956F5D1BFE6A79194BF">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
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<mods:namePart id="24A7D16C381FCFE97A0C3B1ADE036F30">Yu, Long</mods:namePart>
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<mods:affiliation id="7C57F8CCEF6A0D9D1150C4CBFD6D89A0">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
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<mods:namePart id="1F46B90407CA2981A182DC469994B99E">Li, Daiqin</mods:namePart>
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<mods:affiliation id="686342C7EF0E225CE3303F52AF3F6316">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China &amp; Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, Singapore 117543</mods:affiliation>
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<treatment id="D0DEE7F143A9512D89942689CD66F8DD" ID-arpha="D0DEE7F1-43A9-512D-8994-2689CD66F8DD" ID-zoobank="https://zoobank.org/DE92B21F-6A12-44AA-97C8-18EE985DB623" LSID="urn:lsid:plazi:treatment:D0DEE7F143A9512D89942689CD66F8DD" LSID-ZBK="urn:lsid:zoobank.org:act:DE92B21F-6A12-44AA-97C8-18EE985DB623" httpUri="http://treatment.plazi.org/id/D0DEE7F143A9512D89942689CD66F8DD">
<subSubSection id="A884F08AD0BECE3E1B07BBA00ED458B7" type="nomenclature">
<paragraph id="7697DD43CA2A31C0E20FBEEA00055E91">
<taxonomicName id="9FAD99FCB5476A2304A84C0549908310" authority="Zhan &amp; Xu" authorityName="Zhan &amp; Xu" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kalaw" status="sp. nov.">
<emphasis id="FFA6E1CA087FE5340DA403329561A2F2" italics="true">Liphistius kalaw</emphasis>
Zhan &amp; Xu
</taxonomicName>
<taxonomicNameLabel id="4B98F382EF4F7E7A9CCD9C62E562B156" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="9A0A9EC0F8B649FD72507E571510472C" type="description">
<paragraph id="49A58B47B4061D201C59CEC137C7AF17">
<figureCitation id="EA99D9B515CCF172F034E5AE643B3A5E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">Figs 3</figureCitation>
,
<figureCitation id="4F4DBB368120D874F923D9C520363B76" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Vulva of Liphistius kalaw sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster; a = length of poreplate transverse; b = width of poreplate. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure4" httpUri="https://binary.pensoft.net/fig/1115130">4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYQAE" type="type material">
<paragraph id="2FC19CB69B34CB7E6282CA1400115C42">
<heading id="1F2D2686356F8F4CF919E53BA54FCD4E" reason="title">Type material.</heading>
</paragraph>
<paragraph id="7AF7B31AEC21A24F6B4B54B483D43A5C">
<materialsCitation id="53F4BCAA58F21C5CC2D7557C53F90B66" collectingDate="2019-07-27" collectorName="D. Li" country="Myanmar" elevation="944" latitude="20.7" location="Kalaw Township" longLatPrecision="760" longitude="96.52" specimenCount="1" specimenCount-male="1" stateProvince="Shan" typeStatus="holotype">
<emphasis id="33C07F1B0EF89C09D7612CEC912C6CAD" bold="true" italics="true">
<typeStatus id="B25E46CF0DEC90DC20451089BC2476F6" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="451162A1B81C14654F3AF5C288883037" name="Myanmar">Myanmar</collectingCountry>
<specimenCount id="21F27CE7B6F92EF9645905833DC0C276" count="1" type="male"></specimenCount>
,
<collectingRegion id="F495C1300AB809EFE4299B1BC2391E80" country="Myanmar" name="Shan">Shan State</collectingRegion>
,
<quantity id="9EE14365F66661529337784230818364" metricMagnitude="4" metricUnit="m" metricValue="1.6" unit="km" value="16.0">16 km</quantity>
W of
<location id="B4ADAFC160019F68D76066695500F4F0" LSID="urn:lsid:plazi:treatment:D0DEE7F143A9512D89942689CD66F8DD:B4ADAFC160019F68D76066695500F4F0" country="Myanmar" latitude="20.7" longLatPrecision="760" longitude="96.52" name="Kalaw Township" stateProvince="Shan">Kalaw Township</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGRAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.520000,20.700000]}">
<geoCoordinate id="00F6DA60F5D2F7AC77867AC94C0233F5" degrees="20.70" direction="north" orientation="latitude" precision="555" value="20.7">20.70 ° N</geoCoordinate>
,
<geoCoordinate id="F963033B304EE9D95E333A95ABC70CFF" degrees="96.52" direction="east" orientation="longitude" precision="555" value="96.52">96.52 ° E</geoCoordinate>
</named-content>
, alt.
<quantity id="A4AE25DAFA7C5387CF6DB229F7C30B9B" metricMagnitude="2" metricUnit="m" metricValue="9.44" unit="m" value="944.0">
<elevation id="80256E94F5B89B8BB8C7CA918AD8AA22" metricMagnitude="2" metricUnit="m" metricValue="9.44" unit="m" value="944.0">944 m</elevation>
</quantity>
;
<date id="7A1B2F1448D5C8ECBBBEB0E89BE0C2F6" value="2019-07-27">
<collectingDate id="0124A5DAAF3CDA8F71C61255838A8BA4" value="2019-07-27">27. 07. 2019</collectingDate>
</date>
;
<collectorName id="69A802D1D8B6DF0AD49DA785D9D1061A">D. Li</collectorName>
et al. leg.; XUX- 2019-061 A
</materialsCitation>
.
<emphasis id="8F19F0D7ACDB88497C40541B40E5861E" bold="true" italics="true">
<typeStatus id="2DE4C9F46275B618B90CCA5B88E70815" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="90597EFB0DCD1507B1F445FA2DB9FC61" count="8" type="female">8 ♀♀</specimenCount>
, same data as for
<typeStatus id="8F002A51716E322FDEDCE4AA2132B443" type="holotype">holotype</typeStatus>
;
<date id="1F8DB126501D817DD5F58C9C75CE585A" value="2019-07-27">27. 07. 2019</date>
and
<date id="A087323410CCDE602CFF3B959E76545B" value="2018-07-15">15. 07. 2018</date>
; XUX- 2019-062 / 063 / 064 / 065 / 066 / 067 A / 068; XUX- 2018-124.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EORAE" type="diagnosis">
<paragraph id="BDB6BC3B442498F5D15CF85BB72645CC">
<heading id="80DA1317773417AF600C6709BFEE2C24" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="B6EDD8C75065E76CF562B587720E6232">
The male of
<taxonomicName id="7D55D93C6E3B132B12348BEC02C0FDF3" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kalaw" status="sp. nov.">
<emphasis id="A8A4BCB54A28680C5CCDA84B3F22CA2C" italics="true">L. kalaw</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DB324125438FD76707239F2A2D51ACF" rank="species">sp. nov.</taxonomicNameLabel>
resembles those of
<taxonomicName id="6F0FBF3F651E0DD240759E87A8E1EDAC" authorityName="Thorell" authorityYear="1897" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="birmanicus">
<emphasis id="45FEC0A7F53BB77F4A5FD033FB352D56" italics="true">L. birmanicus</emphasis>
</taxonomicName>
and
<taxonomicName id="D862C29F2C7B1FDF85472CC91929D55F" authorityName="Aung, Xu, Lwin, Sang, Yu, Liu, Liu &amp; Li" authorityYear="2019" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="pinlaung">
<emphasis id="4490ACDA21C90B2C50CB4CFF1E47488E" italics="true">L. pinlaung</emphasis>
</taxonomicName>
in having a distinct contrategular process (
<abbrev id="ABBRID0EVSAE" xlink_title="contrategular process">cp</abbrev>
) (Fig.
<figureCitation id="532C0F54AC3D39A91F3799FAAE080BA8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 B, E</figureCitation>
), but it can be distinguished from
<taxonomicName id="695E7ECAA72C02A841D7654E2A19B49B" authorityName="Thorell" authorityYear="1897" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="birmanicus">
<emphasis id="F68D371600E1F8E1D9E90ADA793B0B30" italics="true">L. birmanicus</emphasis>
</taxonomicName>
by the relatively wider base of the contrategular process (
<abbrev id="ABBRID0EITAE" xlink_title="contrategular process">cp</abbrev>
) (Fig.
<figureCitation id="8FA71DCAF8C402E237D5BE4DCC6AA45A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 E</figureCitation>
vs fig. 13 E in
<bibRefCitation id="D21EAFCB97E687EE8341AEEE7DF1BB82" DOI="10.35929/RSZ.0083" author="Schwendinger" etAl="et al." firstAuthor="Schwendinger" issue="2" journalOrPublisher="Revue Suisse de Zoologie" pagination="375-424" refId="B5" refString="Schwendinger PJ, Huber S, Lehmann-Graber C, Ono H, Aung MM, Hongpadharakiree K (2022) A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species. Revue Suisse de Zoologie 129 (2): 375424. https://doi.org/10.35929/RSZ.0083" title="A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species." volume="129" year="2022">Schwendinger et al. 2022</bibRefCitation>
), and from
<taxonomicName id="28FC45154F217CE46AD73A6CF1E5983E" authorityName="Aung, Xu, Lwin, Sang, Yu, Liu, Liu &amp; Li" authorityYear="2019" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="pinlaung">
<emphasis id="3CAA47B0C9F244FE089922B06199F184" italics="true">L. pinlaung</emphasis>
</taxonomicName>
by the cumulus (
<abbrev id="ABBRID0E6TAE" xlink_title="cumulus">Cu</abbrev>
) which has shorter spines (Fig.
<figureCitation id="1430FFA7133F3D9027E5713E841A3F7A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 A C</figureCitation>
vs fig. 4 C in
<bibRefCitation id="81544AB47DBD03953F3A7F785740772B" DOI="10.3897/zookeys.882.38811" author="Aung" etAl="et al." firstAuthor="Aung" journalOrPublisher="ZooKeys" pagination="29-39" refId="B1" refString="Aung KPP, Xu X, Lwin WW, Sang MZ, Yu L, Liu H, Liu FX, Li DQ (2019) Two new species of the primitively segmented spider genus Liphistius Schiödte, 1849 (Mesothelae, Liphistiidae) from Myanmar. ZooKeys 882: 2939. https://doi.org/10.3897/zookeys.882.38811" title="Two new species of the primitively segmented spider genus Liphistius Schiödte, 1849 (Mesothelae, Liphistiidae) from Myanmar." volume="882" year="2019">Aung et al. 2019</bibRefCitation>
). The female of
<taxonomicName id="E19C7BB574185BBABFAC9B84DD5F054E" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kalaw" status="sp. nov.">
<emphasis id="576E90D97DFAEA4C0486602F9B2667D5" italics="true">L. kalaw</emphasis>
</taxonomicName>
<taxonomicNameLabel id="931BBF0F0BF8269EAD486516DCEA8CF6" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from that of
<taxonomicName id="BC2B77502C1CB1BD12A0DC2B2EFD01F2" authorityName="Aung, Xu, Lwin, Sang, Yu, Liu, Liu &amp; Li" authorityYear="2019" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="hpruso">
<emphasis id="537B2011B4AB10CE0B93F661A803AC43" italics="true">L. hpruso</emphasis>
</taxonomicName>
by the relatively wider posterior stalk (
<collectionCode id="867F3C6C4452B7042B22B740955FA135">PS</collectionCode>
) (Fig.
<figureCitation id="CD4EABC3E90DAF5C2C327AFA6D301EF2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Vulva of Liphistius kalaw sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster; a = length of poreplate transverse; b = width of poreplate. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure4" httpUri="https://binary.pensoft.net/fig/1115130">4 C</figureCitation>
vs fig. 3 B E in
<bibRefCitation id="734CD7994C9D6E7710355B987BEAA366" DOI="10.3897/zookeys.882.38811" author="Aung" etAl="et al." firstAuthor="Aung" journalOrPublisher="ZooKeys" pagination="29-39" refId="B1" refString="Aung KPP, Xu X, Lwin WW, Sang MZ, Yu L, Liu H, Liu FX, Li DQ (2019) Two new species of the primitively segmented spider genus Liphistius Schiödte, 1849 (Mesothelae, Liphistiidae) from Myanmar. ZooKeys 882: 2939. https://doi.org/10.3897/zookeys.882.38811" title="Two new species of the primitively segmented spider genus Liphistius Schiödte, 1849 (Mesothelae, Liphistiidae) from Myanmar." volume="882" year="2019">Aung et al. 2019</bibRefCitation>
).
</paragraph>
<caption id="716D9CA6E2CC7F89ED84E1B48F9C5D3B" ID-DOI="10.3897/zookeys.1210.123986.figure3" ID-arpha="CAB41D52-B773-5360-B4B5-E82229AE1B2B" httpUri="https://binary.pensoft.net/fig/1115129" startId="F3">
<paragraph id="FEDAA9C605721BD9C81AA96023EB5C1A">
<label id="9E1E807F39D3FFFE611B96010363CBC5">Figure 3.</label>
</paragraph>
<paragraph id="06EB19FB0A5B2107C75A2F5F3D472380">
Male palp of
<taxonomicName id="A64F30F01376BF074BDF8B5F190530A2" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kalaw" status="sp. nov.">
<emphasis id="803471CCE6C39ED0B6C1677828D3CEA0" italics="true">Liphistius kalaw</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0660BAAFEB09D0520E9B02756D401417" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="3AE3EAF7200FB35B3B82E9C6B5D7E083" bold="true">A, E</emphasis>
prolateral view
<emphasis id="074CACA811D02B032D62FCC513D96A4C" bold="true">B</emphasis>
ventral view
<emphasis id="E60E5B15F3B10B713FDF0E36FB512DFD" bold="true">C, F</emphasis>
retrolateral view
<emphasis id="EF0424CE16C88FB3968F8E3F2EAFF6A6" bold="true">D</emphasis>
dorsal view, red arrows show arched projection and triangular process of
<abbrev id="ABBRID0EEWAE" xlink_title="contrategulum">CT</abbrev>
<emphasis id="55D80DD0D7A89193ACB6410621E9EC9C" bold="true">G</emphasis>
distal view. Abbreviations:
<abbrev id="ABBRID0EKWAE" xlink_title="contrategulum">CT</abbrev>
= contrategulum;
<abbrev id="ABBRID0EOWAE" xlink_title="contrategular process">cp</abbrev>
= contrategular process;
<abbrev id="ABBRID0ESWAE" xlink_title="cumulus">Cu</abbrev>
= cumulus; E = embolus;
<abbrev id="ABBRID0EWWAE" xlink_title="paracymbium">PC</abbrev>
= paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum;
<abbrev id="ABBRID0E1WAE" xlink_title="male palps bearing tibial apophysis">TiA</abbrev>
= tibial apophysis. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="D24C36FAB214EEC5F71D0BDA2D0C0FAB" ID-DOI="10.3897/zookeys.1210.123986.figure4" ID-arpha="9C5A999A-9CB0-5F28-B4B1-27C4BBCBC908" httpUri="https://binary.pensoft.net/fig/1115130" startId="F4">
<paragraph id="B9BB8AB9254FBFAFBA1406B0739C9B73">
<label id="D33A8680BD432C7EF8F71FB2013FA702">Figure 4.</label>
</paragraph>
<paragraph id="45B303E90176297F0D30C2A5AFE812FD">
Vulva of
<taxonomicName id="8AC642C7F71F2D2449FC5712FCE26911" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kalaw" status="sp. nov.">
<emphasis id="C00778387F2CBF39D7FC4DA10BE194E7" italics="true">Liphistius kalaw</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A4F3CB83A96D267F7E08FCA399391E05" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="73649E32C01D87E82FA8CAD4AD08DC60" bold="true">A, C</emphasis>
dorsal view
<emphasis id="84C92196A550ED904F9920C7C18EAD5D" bold="true">B, D</emphasis>
ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk;
<abbrev id="ABBRID0E1XAE" xlink_title="sclerotized poreplate">PPl</abbrev>
= poreplate;
<abbrev id="ABBRID0E5XAE" xlink_title="receptacular cluster">RC</abbrev>
= receptacular cluster; a = length of poreplate transverse; b = width of poreplate. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EHYAE" type="description">
<paragraph id="68379F02305BB81DCF08325BD74FD61D">
<heading id="A3F034C61FEE7332CFD3D31B3BE8268D" reason="title">Description.</heading>
</paragraph>
<paragraph id="BA3D7639B62701E883BE0F2E64179897">
<emphasis id="3EC9C52795447FF00FA8315352203D8E" bold="true">
Male
<typeStatus id="8D783900399CE8F712314C2C043C4FC3" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace yellowish brown, with few short, scattered bristles; opisthosoma yellowish brown, with 12 brown tergites, close to each other, 26
<superScript id="F42D3169641C538756673A519AD8C2D9">th</superScript>
larger than others, 5
<superScript id="2C2E14C13BFBD0AA9FA3D6E9A78D0DBE">th</superScript>
largest; chelicerae robust, promargin with 13 denticles of variable size; labium and sternum yellowish brown, with few short setae on anterior margin and many long setae on elongated posterior margin; legs yellowish brown, with strong setae and spines. Measurements:
<abbrev id="ABBRID0ETYAE" xlink_title="body length">BL</abbrev>
13.65,
<abbrev id="ABBRID0EXYAE" xlink_title="carapace length">CL</abbrev>
5.94,
<abbrev id="ABBRID0E2YAE" xlink_title="carapace width">CW</abbrev>
6.07,
<abbrev id="ABBRID0E6YAE" xlink_title="opisthosoma length">OL</abbrev>
8.00,
<abbrev id="ABBRID0EDZAE" xlink_title="opisthosoma width">OW</abbrev>
5.36; eye sizes and interdistances:
<abbrev id="ABBRID0EHZAE" xlink_title="anterior median eyes">AME</abbrev>
0.08,
<abbrev id="ABBRID0ELZAE" xlink_title="anterior lateral eyes">ALE</abbrev>
0.54,
<abbrev id="ABBRID0EPZAE" xlink_title="posterior median eyes">PME</abbrev>
0.27,
<abbrev id="ABBRID0ETZAE" xlink_title="posterior lateral eyes">PLE</abbrev>
0.43,
<abbrev id="ABBRID0EXZAE" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0E2ZAE" xlink_title="anterior median eyes">AME</abbrev>
0.11,
<abbrev id="ABBRID0E6ZAE" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0ED1AE" xlink_title="anterior lateral eyes">ALE</abbrev>
0.15,
<abbrev id="ABBRID0EH1AE" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0EL1AE" xlink_title="posterior median eyes">PME</abbrev>
0.08,
<abbrev id="ABBRID0EP1AE" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0ET1AE" xlink_title="posterior lateral eyes">PLE</abbrev>
0.10,
<abbrev id="ABBRID0EX1AE" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0E21AE" xlink_title="posterior lateral eyes">PLE</abbrev>
0.17,
<abbrev id="ABBRID0E61AE" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0ED2AE" xlink_title="anterior lateral eyes">ALE</abbrev>
0.12,
<abbrev id="ABBRID0EH2AE" xlink_title="posterior lateral eyes">PLE</abbrev>
<abbrev id="ABBRID0EL2AE" xlink_title="posterior lateral eyes">PLE</abbrev>
0.34,
<abbrev id="ABBRID0EP2AE" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0ET2AE" xlink_title="posterior median eyes">PME</abbrev>
0.10; labium 1.06 long and 0.98 wide; sternum 3.08 long and 0.96 wide; legs: I 18.36 (4.81, 2.57, 3.87, 4.60, 2.51), II 20.24 (5.64, 2.56, 4.23, 5.07, 2.74), III 20.46 (5.64, 2.37, 4.42, 6.00, 3.03), IV 27.56 (6.97, 2.84, 5.57, 8.17, 4.01).
</paragraph>
<paragraph id="151E46E3B3E43F5AD90E2A46648C2363">
Palp: tibial apophysis (
<abbrev id="ABBRID0EZ2AE" xlink_title="male palps bearing tibial apophysis">TiA</abbrev>
) with 4 tapering setae of similar length (Fig.
<figureCitation id="317D03CBDE0ADF7DB0952CADB441043F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 C</figureCitation>
); paracymbium (
<abbrev id="ABBRID0EB3AE" xlink_title="paracymbium">PC</abbrev>
) with many setae situated on tip (Fig.
<figureCitation id="0A14F7F7BB52FB1B5F72EDE1FBBBB6C7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 A C</figureCitation>
); cumulus (
<abbrev id="ABBRID0EJ3AE" xlink_title="cumulus">Cu</abbrev>
) slightly elevated, with 8 tapering spines (Fig.
<figureCitation id="F269D10666944891FD481D554480A434" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 A C</figureCitation>
); subtegulum with weakly developed apophysis (Fig.
<figureCitation id="0EC8FE3CAC6CAD203C1C88C77A1776AE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 B</figureCitation>
); contrategulum (
<abbrev id="ABBRID0EV3AE" xlink_title="contrategulum">CT</abbrev>
) with arched projection distally, and large triangular process in prolateral view (Fig.
<figureCitation id="FE99AB1F8A6F2CC4104B52ED87172A07" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 A, B, D</figureCitation>
); tegulum (
<collectionCode id="B515C6B28621EE251CE0FB31D4C9DF10" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
) fan-shaped with serrated edge and longitudinal ridges (Fig.
<figureCitation id="CA68FDA3C5CE175A6DABCF73FF8DEED7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 C, F</figureCitation>
); paraembolic plate (PeP) narrow, sclerotized (Fig.
<figureCitation id="3AE4579A0CAE3C377A8738494513BF1F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 A, F</figureCitation>
); embolus (E) with 4 distinct longitudinal ridges reaching tip in retrolateral view, and with several denticles along longitudinal ridges (Fig.
<figureCitation id="B98613C76790664835D1D7614D05BBF3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 B F</figureCitation>
).
</paragraph>
<paragraph id="5322CF90A434395C0C62B7BAB5DCE00D">
<emphasis id="2C213BCDE17F12FD0DEDBEC2B2E5A626" bold="true">
Female
<typeStatus id="44BBB99C39D30E80741EAC3F16F2A807" type="paratype">paratype</typeStatus>
(XUX- 2019-062).
</emphasis>
Carapace yellowish brown, with few short, scattered bristles; opisthosoma brown, with 12 brown tergites, close to each other, with gray patches, 26
<superScript id="77F1A73F37BF694F336DD811FDC3DB07">th</superScript>
larger than others, 5
<superScript id="E05C2DA8482D7CB75BD0A3BBAA458E01">th</superScript>
largest; chelicerae robust, reddish brown; promargin of cheliceral groove with 12 denticles of variable size; labium yellowish brown, sternum with several setae; legs with strong setae and spines. Measurements:
<abbrev id="ABBRID0ER4AE" xlink_title="body length">BL</abbrev>
11.56,
<abbrev id="ABBRID0EV4AE" xlink_title="carapace length">CL</abbrev>
5.82,
<abbrev id="ABBRID0EZ4AE" xlink_title="carapace width">CW</abbrev>
5.22,
<abbrev id="ABBRID0E44AE" xlink_title="opisthosoma length">OL</abbrev>
6.23,
<abbrev id="ABBRID0EB5AE" xlink_title="opisthosoma width">OW</abbrev>
4.62; eye sizes and interdistances:
<abbrev id="ABBRID0EF5AE" xlink_title="anterior median eyes">AME</abbrev>
0.07,
<abbrev id="ABBRID0EJ5AE" xlink_title="anterior lateral eyes">ALE</abbrev>
0.56,
<abbrev id="ABBRID0EN5AE" xlink_title="posterior median eyes">PME</abbrev>
0.29,
<abbrev id="ABBRID0ER5AE" xlink_title="posterior lateral eyes">PLE</abbrev>
0.42,
<abbrev id="ABBRID0EV5AE" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EZ5AE" xlink_title="anterior median eyes">AME</abbrev>
0.09,
<abbrev id="ABBRID0E45AE" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EB6AE" xlink_title="anterior lateral eyes">ALE</abbrev>
0.15,
<abbrev id="ABBRID0EF6AE" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0EJ6AE" xlink_title="posterior median eyes">PME</abbrev>
0.06,
<abbrev id="ABBRID0EN6AE" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0ER6AE" xlink_title="posterior lateral eyes">PLE</abbrev>
0.09,
<abbrev id="ABBRID0EV6AE" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EZ6AE" xlink_title="posterior lateral eyes">PLE</abbrev>
0.11,
<abbrev id="ABBRID0E46AE" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0ECAAG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.07,
<abbrev id="ABBRID0EGAAG" xlink_title="posterior lateral eyes">PLE</abbrev>
<abbrev id="ABBRID0EKAAG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.35,
<abbrev id="ABBRID0EOAAG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0ESAAG" xlink_title="posterior median eyes">PME</abbrev>
0.08; labium 3.00 long and 0.61 wide; sternum 1.19 long and 1.13 wide; palp 10.25 (3.62, 1.74, 2.36, 2.53), legs: I 12.78 (4.29, 2.03, 2.63, 2.37, 1.46), II 12.89 (4.17, 2.03, 2.56, 2.59, 1.54), III 13.42 (4.13, 1.84, 2.75, 2.78, 1.92), IV 19.33 (5.57, 2.32, 3.81, 4.91, 2.72).
</paragraph>
<paragraph id="938FCCC7F0A38AC548919A1E45238441">
Vulva: poreplate (
<abbrev id="ABBRID0EYAAG" xlink_title="sclerotized poreplate">PPl</abbrev>
) slightly wider than long, with pair of large anterior lobes and pair of small anterolateral lobes; central dorsal opening (
<collectionCode id="9226ACD7B8675A600D5572086888F07B">CDO</collectionCode>
) circular, racemose receptacular cluster (
<abbrev id="ABBRID0E3AAG" xlink_title="receptacular cluster">
<collectionCode id="2E6977B2735A391E9D5475AEA0918F8D">RC</collectionCode>
</abbrev>
) slightly long; posterior stalk (
<collectionCode id="E619BC1CD17003EFD198FE8513DADDA2">PS</collectionCode>
) axe-shaped; lateral margins of genital atrium (GA) membranous (Fig.
<figureCitation id="CCCB716771F9FD14919F77421759A4BE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Vulva of Liphistius kalaw sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster; a = length of poreplate transverse; b = width of poreplate. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure4" httpUri="https://binary.pensoft.net/fig/1115130">4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEBAG" type="variation">
<paragraph id="03528DA065840EEB59FBA8989CF1FDB3">
<heading id="EF694DC53E75499C6FD51376A931A75F" reason="title">Variation.</heading>
</paragraph>
<paragraph id="9839E25EF1369BE62AA7BBFA54D98D70">
Females vary in body size. The range measurements of females (
<emphasis id="6667B023D4E697D68C852E0A46F4DD6C" italics="true">N</emphasis>
= 8):
<abbrev id="ABBRID0EMBAG" xlink_title="body length">BL</abbrev>
8.5516.11,
<abbrev id="ABBRID0EQBAG" xlink_title="carapace length">CL</abbrev>
4.487.20,
<abbrev id="ABBRID0EUBAG" xlink_title="carapace width">CW</abbrev>
3.906.43,
<abbrev id="ABBRID0EYBAG" xlink_title="opisthosoma length">OL</abbrev>
4.188.56,
<abbrev id="ABBRID0E3BAG" xlink_title="opisthosoma width">OW</abbrev>
3.007.36. The number of denticles on the promargin of cheliceral groove varies from 11 to 14 (
<emphasis id="30AF1ADADBE80742CA2BD35A78B5446B" italics="true">N</emphasis>
= 8).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECCAG" type="etymology">
<paragraph id="6706ECFB1287FA2076CEC9813261683A">
<heading id="10F64A3BA0A101433248B22E7D6A9C86" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="A6B9EF402EBCC0240F7D4DEB88733B89">
The species epithet
<taxonomicName id="2956E68DACCFDF0ED9FBB93A5CDE31B9" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kalaw">
<emphasis id="13D831DC53F6E23B2BE39E30D220197C" italics="true">kalaw</emphasis>
</taxonomicName>
” refers to the
<typeStatus id="8906A6F0378FA69FBCA31F8906F70424">type</typeStatus>
locality, Kalaw Township; it is treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERCAG" type="distribution">
<paragraph id="3804DC6EE60363984BD9D8FFACD9CD7C">
<heading id="6C11D500CA8CA99C378810AAD38737CE" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="B8C8A001B7EF17D62B3E9C15153461E4">
<collectingRegion id="75AB6F8757624AAF38FA42EDCD2E80F6" country="Myanmar" name="Mandalay">Mandalay Region</collectingRegion>
(Kalaw Township),
<collectingCountry id="81FECEB190C4DFBB79684CF80AF2D2A9" name="Myanmar">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>