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<document id="968319B4FD65838F222753957638A3EA" ID-DOI="10.3897/zookeys.1210.123986" ID-ZooBank="6972216E-2356-4DF7-BD24-2B6B68E639B4" ID-publisher-id="123986" URI-arpha="B6184626-6862-59C4-9047-EF9B7AC24E35" URI-zoobank="http://zoobank.org/6972216E-2356-4DF7-BD24-2B6B68E639B4" article-type="research-article" checkinTime="1723830749198" checkinUser="pensoft" docAuthor="Xu, Xin, Zhan, Yi, Aung, Khin Pyae Pyae, Liu, Fengxiang, Yu, Long &amp; Li, Daiqin" docDate="2024" docId="1148FBC405EA553C96A0DBAFAAE5E4F3" docLanguage="en" docName="ZooKeys 1210: 99-115" docOrigin="ZooKeys 1210" docSource="https://zookeys.pensoft.net/article/123986/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Liphistius nawngau Zhan &amp; Xu 2024, sp. nov." docType="treatment" docVersion="1" dtd-version="3.0" lastPageNumber="115" masterDocId="B6184626686259C49047EF9B7AC24E35" masterDocTitle="Four new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae, Liphistiinae) from Myanmar" masterLastPageNumber="115" masterPageNumber="99" pageNumber="99" updateTime="1723830805509" updateUser="pensoft">
<mods:mods id="6EFFA1AA2E0ADE701BC29332246D46E1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="D924FFB1BE81705ED60C938FE73955F8">Four new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae, Liphistiinae) from Myanmar</mods:title>
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<mods:name id="91339A77146F36FC725C071924D597B9" type="personal">
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<mods:namePart id="CD60D78777F3E5E136C8DF9A52C72A13">Xu, Xin</mods:namePart>
<mods:nameIdentifier id="042CB0522EDA21998C75AA24FCB8C6C6" type="ORCID">0000-0001-5632-6622</mods:nameIdentifier>
<mods:affiliation id="6731F703138990F3D4384D2529126BCC">College of Life Sciences, Hunan Normal University, 36 Lushan Road, Changsha 410081, Hunan Province, China &amp; State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
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<mods:namePart id="7FDDA4F6C557EF6888E971D9D47CE88C">Zhan, Yi</mods:namePart>
<mods:affiliation id="BD6D02B8BC601A680B041B9C308446C7">College of Life Sciences, Hunan Normal University, 36 Lushan Road, Changsha 410081, Hunan Province, China</mods:affiliation>
</mods:name>
<mods:name id="7566F7C5ED904B703CFC546EE0DBD18E" type="personal">
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<mods:namePart id="812DC819A86A89DBA6DC9218F9CE3627">Aung, Khin Pyae Pyae</mods:namePart>
<mods:affiliation id="F44D521F11286E54300B13FD8B91F87F">Department of Zoology, University of Yangon, Kamayut Township, Pyay Road, Yangon, 11041, Myanmar &amp; Department of Biology, Taungoo Education Degree College, Taungoo, 08101, Myanmar</mods:affiliation>
</mods:name>
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<mods:namePart id="2DFCBB11B59B519ED4930DFB549CF1B0">Liu, Fengxiang</mods:namePart>
<mods:affiliation id="361E1B1E5C4D05B95BFE1C8FE9728046">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
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<mods:namePart id="E74177B0F04A968E9DE56A406588DBB5">Yu, Long</mods:namePart>
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<mods:affiliation id="BD60D5CF4706CC1B061C46663A6DDB8E">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
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<mods:namePart id="63CCDF3DABCBC2D960A803387BD52E3C">Li, Daiqin</mods:namePart>
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<mods:affiliation id="F6C7E9F66100EBA3EB9E1CA648E8ECE5">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China &amp; Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, Singapore 117543</mods:affiliation>
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<mods:part id="65300E7F9FBB5459C1E0E4DF75735935">
<mods:date id="15D87897C063B4BF9DCEF0283D786DE3">2024</mods:date>
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<mods:number id="23980C0F5A7EFD7FF4E6C7AFEE8405F0">2024-08-16</mods:number>
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<mods:number id="1ABC3C70807DD9C9D9D4A697C5098F04">1210</mods:number>
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<mods:classification id="F5C0ACDBA108BC1DC116F062679F4EA5">journal article</mods:classification>
<mods:identifier id="5E6ED5A3F7A5F5B80F8457CFF4138EAC" type="DOI">10.3897/zookeys.1210.123986</mods:identifier>
<mods:identifier id="99B556BDAE66423E813BF85A5794D29F" type="ZooBank">6972216E-2356-4DF7-BD24-2B6B68E639B4</mods:identifier>
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<treatment id="1148FBC405EA553C96A0DBAFAAE5E4F3" ID-arpha="1148FBC4-05EA-553C-96A0-DBAFAAE5E4F3" ID-zoobank="https://zoobank.org/1ADFCE8F-66EE-43F9-B61A-927F6B88189F" LSID="urn:lsid:plazi:treatment:1148FBC405EA553C96A0DBAFAAE5E4F3" LSID-ZBK="urn:lsid:zoobank.org:act:1ADFCE8F-66EE-43F9-B61A-927F6B88189F" httpUri="http://treatment.plazi.org/id/1148FBC405EA553C96A0DBAFAAE5E4F3">
<subSubSection id="91EC8776201CB8D09910ABB853F27E75" type="nomenclature">
<paragraph id="15085F835D034EA16CF6B316CD414D3B">
<taxonomicName id="351F29596982D3F6CCCA2BAA2C8513BA" authority="Zhan &amp; Xu" authorityName="Zhan &amp; Xu" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nawngau" status="sp. nov.">
<emphasis id="984167406BB6BA6CBBAF12D4335FF7EF" italics="true">Liphistius nawngau</emphasis>
Zhan &amp; Xu
</taxonomicName>
<taxonomicNameLabel id="8C1F53FB7EC2EA139188E081E7B6A350" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="65E8133EBF945D84B4A72FC1051B2879" type="description">
<paragraph id="B7BDADFCD483D9FBD47B9DC65CCC03D0">
<figureCitation id="5B9D23331179DB5F3A2EAA46ECA6A39A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">Figs 7</figureCitation>
,
<figureCitation id="397A66BF49634B4074C959AB83F5FC88" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Vulva of Liphistius nawngau sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure8" httpUri="https://binary.pensoft.net/fig/1115134">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBXAG" type="type material">
<paragraph id="B2427413558002C8C651BAD42C6317D6">
<heading id="48808AE7FF11DA07348FEBBDF736175D" reason="title">Type material.</heading>
</paragraph>
<paragraph id="C27AC64F39B6B2D8552C857AEF9C94DC">
<materialsCitation id="98B02842AC81616341CF770C7D8F188B" collectingDate="2019-07-26" collectorName="D. Li" country="Myanmar" county="Kyaukme Dist." elevation="1096" latitude="22.26" location="Nawng Au Vill." longLatPrecision="756" longitude="96.83" municipality="Nawnghkio Township" specimenCount="1" specimenCount-male="1" stateProvince="Shan" typeStatus="holotype">
<emphasis id="3967BE0C03BDAFD15E2F720F8C5BD23C" bold="true" italics="true">
<typeStatus id="8331C2623EA9CA57598E737FA09963D6" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="82C41F59AAFBB83F819C51ABA6B3976B" name="Myanmar">Myanmar</collectingCountry>
<specimenCount id="27F1C66CB74841F134A89DB2F205F2DB" count="1" type="male"></specimenCount>
,
<collectingRegion id="7FC831130EB4E3B05E9C83A881787D77" country="Myanmar" name="Shan">Shan State</collectingRegion>
,
<collectingCounty id="3DC684B598ACBFB6F43210EEA13F97DF">Kyaukme Dist.</collectingCounty>
,
<collectingMunicipality id="25DE4FCE47AD34C6DA14F4C7F2A3D788">Nawnghkio Township</collectingMunicipality>
,
<location id="9C115DF75981A5C3823413ACB7C6F46F" LSID="urn:lsid:plazi:treatment:1148FBC405EA553C96A0DBAFAAE5E4F3:9C115DF75981A5C3823413ACB7C6F46F" country="Myanmar" county="Kyaukme Dist." latitude="22.26" longLatPrecision="756" longitude="96.83" municipality="Nawnghkio Township" name="Nawng Au Vill." stateProvince="Shan">Nawng Au Vill.</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPXAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.830000,22.260000]}">
<geoCoordinate id="227AE6416CDD76731B1A80B6F22A2689" degrees="22.26" direction="north" orientation="latitude" precision="555" value="22.26">22.26 ° N</geoCoordinate>
,
<geoCoordinate id="CB3FAF6ED323CD62361EFC8C78E458D9" degrees="96.83" direction="east" orientation="longitude" precision="555" value="96.83">96.83 ° E</geoCoordinate>
</named-content>
, alt.
<quantity id="B79E0E489DE4EE3B6902F0109307F486" metricMagnitude="3" metricUnit="m" metricValue="1.0959999999999999" unit="m" value="1096.0">
<elevation id="300F25DAF517E7FE4B5477CCD530D8B4" metricMagnitude="3" metricUnit="m" metricValue="1.0959999999999999" unit="m" value="1096.0">1096 m</elevation>
</quantity>
;
<date id="D22521C200B8D6B04DDC3D16955576EE" value="2019-07-26">
<collectingDate id="890F7379B699B3BD75B5198C9D009A52" value="2019-07-26">26. 07. 2019</collectingDate>
</date>
;
<collectorName id="4C09A07E9F5C76BD1EDBB0DA053C28B6">D. Li</collectorName>
et al. leg.; XUX- 2019-054 A
</materialsCitation>
.
<emphasis id="87D2C1ECEE3C6D2433D9B2F5630AAB87" bold="true" italics="true">
<typeStatus id="5175CA05864765D837E6BEB4200C6377" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="12782989DF2711F719CDA6182C0E9C6A" count="9" type="male">9 ♂♂</specimenCount>
<specimenCount id="BE40E20D3F6262F735F43D209513ADC8" count="3" type="female">3 ♀♀</specimenCount>
, same data as for
<typeStatus id="89CF1EFA2916D02FDC66CBDB07A51376" type="holotype">holotype</typeStatus>
; XUX- 2019-054 / 054 A / 055 / 055 A / 055 B / 055 D / 056 / 056 A / 057 / 058 / 059 / 060;
<specimenCount id="959E3C847DB5BEA621CC4D9D5064B946" count="5" type="male">5 ♂♂</specimenCount>
<specimenCount id="786D548C5965EFE48F6501A7C8826BCB" count="2" type="female">2 ♀♀</specimenCount>
, same township as for
<typeStatus id="B1E1005FD7BC1D4E2BA57F4A17E5A11D" type="holotype">holotype</typeStatus>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1XAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.730000,22.300000]}">
<geoCoordinate id="C079624369EC4049B17118E4C2D0C485" degrees="22.30" direction="north" orientation="latitude" precision="555" value="22.3">22.30 ° N</geoCoordinate>
,
<geoCoordinate id="1D2DAD04DEC1FB2727F417733318C787" degrees="96.73" direction="east" orientation="longitude" precision="555" value="96.73">96.73 ° E</geoCoordinate>
</named-content>
, alt.
<quantity id="900D0F6B1816747D688313A62529FD47" metricMagnitude="2" metricUnit="m" metricValue="8.45" unit="m" value="845.0">845 m</quantity>
;
<date id="60360393FD5A2E3A868C1276994DE804" value="2019-07-26">26. 07. 2019</date>
and
<date id="8E9A96EF38BE6748B2DDE64053394C26" value="2018-07-14">14. 07. 2018</date>
; XUX- 2019-049 / 051 / 052 / 053, XUX- 2018-116 / 119 / 123.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6XAG" type="diagnosis">
<paragraph id="E622171C10E234893C37A94C1A290FDE">
<heading id="677442942DABB5F6313A398673E134AB" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="CB6CF9E6CA03FADAA9C74D7940C7D77F">
The male of
<taxonomicName id="E48D430159D7DD8EF7C07AF787EB8872" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nawngau" status="sp. nov.">
<emphasis id="AED3D523C2716F33A17BCBA4DC36B3E1" italics="true">L. nawngau</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3247E53B2A878AEBC50484E3555579B1" rank="species">sp. nov.</taxonomicNameLabel>
resembles those of
<taxonomicName id="95E18D322FE590564A4D40AB6528D48D" authorityName="Platnick &amp; Sedgwick" authorityYear="1984" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lordae">
<emphasis id="1BA94450FE7C24C73739B5334C2146AC" italics="true">L. lordae</emphasis>
</taxonomicName>
and
<taxonomicName id="38A60B1481E877F408DD8CA58114A90E" authorityName="Xu, Yu, Aung, Yu, Liu, Lwin, Sang &amp; Li" authorityYear="2021" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="pyinoolwin">
<emphasis id="6C1D4AF212F839CAA5510679372CB83D" italics="true">L. pyinoolwin</emphasis>
</taxonomicName>
in having an adpressed proximal tegular margin (Fig.
<figureCitation id="36C500FC34254F0DC9C68E74987F9DFC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 C, F</figureCitation>
), but it can be distinguished from them by the tegulum (
<collectionCode id="7601C6A74046EE1B7E71F994AAE9318B" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
), which bears a distinct transversal ridge in retrolateral view (Fig.
<figureCitation id="E2F08F526CE359A53490D70DB6621A00" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 C, F</figureCitation>
vs lacking transversal ridge, figs 5,
<quantity id="A69941A57D68D5155096EE39CF535A3A" metricMagnitude="-1" metricUnit="m" metricValue="1.524" unit="in" value="6.0">6 in</quantity>
<bibRefCitation id="D0C44E1564AECCBB7B6D52C0FE3099A1" DOI="10.1111/j.1463-6409.1990.tb00262.x" author="Schwendinger" firstAuthor="Schwendinger" issue="3" journalOrPublisher="Zoologica Scripta" pagination="331-351" refId="B2" refString="Schwendinger PJ (1990) On the spider genus Liphistius (Araneae: Mesothelae) in Thailand and Burma. Zoologica Scripta 19 (3): 331351. https://doi.org/10.1111/j.1463-6409.1990.tb00262.x" title="On the spider genus Liphistius (Araneae: Mesothelae) in Thailand and Burma." volume="19" year="1990">Schwendinger 1990</bibRefCitation>
; fig. 4 F in
<bibRefCitation id="3C7DEF065A7EA75F5F1CACC4B1C030AB" DOI="10.3897/zookeys.1031.59102" author="Xu" etAl="et al." firstAuthor="Xu" journalOrPublisher="ZooKeys" pagination="41-58" refId="B9" refString="Xu X, Yu L, Aung KPP, Yu L, Liu FX, Lwin WW, Sang MZ, Li DQ (2021) A new species of Liphistius from Myanmar and description of the actual male of L. birmanicus Thorell, 1897 (Araneae, Mesothelae, Liphistiidae). ZooKeys 1031: 4158. https://doi.org/10.3897/zookeys.1031.59102" title="A new species of Liphistius from Myanmar and description of the actual male of L. birmanicus Thorell, 1897 (Araneae, Mesothelae, Liphistiidae)." volume="1031" year="2021">Xu et al. 2021</bibRefCitation>
), and by the embolus (E), which has smooth longitudinal ridges reaching tip (Fig.
<figureCitation id="5E06C243BD665C5FFFADA78E7D9D432C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 E</figureCitation>
vs having several denticles along longitudinal ridges, fig. 4 F in
<bibRefCitation id="24DDC3B8F403C959EF46219AE197A54D" DOI="10.3897/zookeys.1031.59102" author="Xu" etAl="et al." firstAuthor="Xu" journalOrPublisher="ZooKeys" pagination="41-58" refId="B9" refString="Xu X, Yu L, Aung KPP, Yu L, Liu FX, Lwin WW, Sang MZ, Li DQ (2021) A new species of Liphistius from Myanmar and description of the actual male of L. birmanicus Thorell, 1897 (Araneae, Mesothelae, Liphistiidae). ZooKeys 1031: 4158. https://doi.org/10.3897/zookeys.1031.59102" title="A new species of Liphistius from Myanmar and description of the actual male of L. birmanicus Thorell, 1897 (Araneae, Mesothelae, Liphistiidae)." volume="1031" year="2021">Xu et al. 2021</bibRefCitation>
). The female of
<taxonomicName id="AC42B1FF463303D449FAAA9E14DF69F6" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nawngau" status="sp. nov.">
<emphasis id="A2EEF69F4593FF8C909CCDE112AFAA60" italics="true">L. nawngau</emphasis>
</taxonomicName>
<taxonomicNameLabel id="220D031B3544CD812EEBCB3788818077" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from that of
<taxonomicName id="C70E2A240CFB13E5123CC40649A301DC" authorityName="Aung, Xu, Lwin, Sang, Yu, Liu, Liu &amp; Li" authorityYear="2019" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="hpruso">
<emphasis id="92D66A463D100963F8DF9C88A7999499" italics="true">L. hpruso</emphasis>
</taxonomicName>
by having the posterior stalk triangular (
<collectionCode id="30DC087311331A2E87FBFE68C86B0ACC">PS</collectionCode>
) (Fig.
<figureCitation id="1CB69AACBC82AEFCFE7D9CD1E0653C07" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Vulva of Liphistius nawngau sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure8" httpUri="https://binary.pensoft.net/fig/1115134">8</figureCitation>
vs nearly oval, fig. 3 B E in
<bibRefCitation id="7572601299EA0288AC6D5DE38590CBD8" DOI="10.3897/zookeys.882.38811" author="Aung" etAl="et al." firstAuthor="Aung" journalOrPublisher="ZooKeys" pagination="29-39" refId="B1" refString="Aung KPP, Xu X, Lwin WW, Sang MZ, Yu L, Liu H, Liu FX, Li DQ (2019) Two new species of the primitively segmented spider genus Liphistius Schiödte, 1849 (Mesothelae, Liphistiidae) from Myanmar. ZooKeys 882: 2939. https://doi.org/10.3897/zookeys.882.38811" title="Two new species of the primitively segmented spider genus Liphistius Schiödte, 1849 (Mesothelae, Liphistiidae) from Myanmar." volume="882" year="2019">Aung et al. 2019</bibRefCitation>
).
</paragraph>
<caption id="55EC15F6C8177BCCB377C0D6AD86E5A4" ID-DOI="10.3897/zookeys.1210.123986.figure7" ID-arpha="167E0A7B-A8FC-5F8B-93C9-EF4CCFD53C17" httpUri="https://binary.pensoft.net/fig/1115133" startId="F7">
<paragraph id="B5EF217E0AC9A38DC1987D435AF81FAB">
<label id="13B70BADA264291994BC80504B5A733D">Figure 7.</label>
</paragraph>
<paragraph id="BBE9FC7E8BEA3D8AA3724CEAC8E7846B">
Male palp of
<taxonomicName id="02DA1C254D20CFA7D986BA823CE081DA" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nawngau" status="sp. nov.">
<emphasis id="F37E9C261CF90F98C7ACD5CFB9DE5733" italics="true">Liphistius nawngau</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CBF0B573733A03D14C8DFB7FB01BDD0B" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="E2E9A079D9465B7B1D81DDE4A5BFCC25" bold="true">A, E</emphasis>
prolateral view
<emphasis id="FAC31AE70C43CA98ECFFF512372EAFF3" bold="true">B</emphasis>
ventral view
<emphasis id="8B867D17AD762AC8515C2344A478E31D" bold="true">C, F</emphasis>
retrolateral view
<emphasis id="F73C6918063F1E5C5F4BB1165B449D04" bold="true">D</emphasis>
dorsal view, red arrows show arched projection and triangular process of
<abbrev id="ABBRID0EX2AG" xlink_title="contrategulum">CT</abbrev>
<emphasis id="96B6E710A9D594DBC9C5E9FE33D7ABFC" bold="true">G</emphasis>
distal view. Abbreviations:
<abbrev id="ABBRID0E42AG" xlink_title="contrategulum">CT</abbrev>
= contrategulum;
<abbrev id="ABBRID0EB3AG" xlink_title="contrategular process">cp</abbrev>
= contrategular process;
<abbrev id="ABBRID0EF3AG" xlink_title="cumulus">Cu</abbrev>
= cumulus; E = embolus;
<abbrev id="ABBRID0EJ3AG" xlink_title="paracymbium">PC</abbrev>
= paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum;
<abbrev id="ABBRID0EN3AG" xlink_title="male palps bearing tibial apophysis">TiA</abbrev>
= tibial apophysis. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="F9500FD4FA1C399A329CE9FB808ADE7B" ID-DOI="10.3897/zookeys.1210.123986.figure8" ID-arpha="B49B9831-E68E-5884-9F01-491C46F8CA97" httpUri="https://binary.pensoft.net/fig/1115134" startId="F8">
<paragraph id="FCBA55ACD07F3BE5B4DF5500AFCE3303">
<label id="D545C0EAE4F0ABE0728345C99AB16C26">Figure 8.</label>
</paragraph>
<paragraph id="7681864D046949A8BE6D09EFE17A4666">
Vulva of
<taxonomicName id="DFAB499436899E303DBA0091F40B57F1" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nawngau" status="sp. nov.">
<emphasis id="204F60E36FF289A6F6F5C7D826D7BDB6" italics="true">Liphistius nawngau</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54FCABF6559B170A5177DBB1F12BD840" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="9D24A29FE825A83C0A3F7C78D1242781" bold="true">A, C</emphasis>
dorsal view
<emphasis id="F7C1B150387C93F12F7F37A7E9058B1C" bold="true">B, D</emphasis>
ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk;
<abbrev id="ABBRID0EN4AG" xlink_title="sclerotized poreplate">PPl</abbrev>
= poreplate;
<abbrev id="ABBRID0ER4AG" xlink_title="receptacular cluster">RC</abbrev>
= receptacular cluster. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E14AG" type="description">
<paragraph id="EC0B07C0424E6B7621A12D815D456854">
<heading id="F0AA6AE3F9BD974655C66A4EC08B06DE" reason="title">Description.</heading>
</paragraph>
<paragraph id="A43F56A996BB9D44E3015B5CD6D01863">
<emphasis id="5071D814A174CD7580E569BC16C6113D" bold="true">
Male
<typeStatus id="536D7EDCE0B7ECE5835A92691638218B" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace brown with few short, scattered bristles; opisthosoma brown, with 12 brown tergites, close to each other, 26
<superScript id="C26C15B5BD5E388F4C454A6E67953ADE">th</superScript>
larger than others, 5
<superScript id="00ED626ECA47D52B3D332A0429DB72EB">th</superScript>
largest; chelicerae robust, promargin of cheliceral groove with 11 denticles of variable size; labium and sternum yellowish brown, sternum with few short setae on anterior tip and many long setae on elongated posterior tip; legs yellowish brown, with strong setae and spines. Measurements:
<abbrev id="ABBRID0EG5AG" xlink_title="body length">BL</abbrev>
15.47,
<abbrev id="ABBRID0EK5AG" xlink_title="carapace length">CL</abbrev>
6.63,
<abbrev id="ABBRID0EO5AG" xlink_title="carapace width">CW</abbrev>
6.57,
<abbrev id="ABBRID0ES5AG" xlink_title="opisthosoma length">OL</abbrev>
7.73,
<abbrev id="ABBRID0EW5AG" xlink_title="opisthosoma width">OW</abbrev>
5.56; eye sizes and interdistances:
<abbrev id="ABBRID0E15AG" xlink_title="anterior median eyes">AME</abbrev>
0.08,
<abbrev id="ABBRID0E55AG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.72,
<abbrev id="ABBRID0EC6AG" xlink_title="posterior median eyes">PME</abbrev>
0.35,
<abbrev id="ABBRID0EG6AG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.49,
<abbrev id="ABBRID0EK6AG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EO6AG" xlink_title="anterior median eyes">AME</abbrev>
0.08,
<abbrev id="ABBRID0ES6AG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EW6AG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.20,
<abbrev id="ABBRID0E16AG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0E56AG" xlink_title="posterior median eyes">PME</abbrev>
0.08,
<abbrev id="ABBRID0ECABG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0EGABG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.07,
<abbrev id="ABBRID0EKABG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EOABG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.81,
<abbrev id="ABBRID0ESABG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EWABG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.11,
<abbrev id="ABBRID0E1ABG" xlink_title="posterior lateral eyes">PLE</abbrev>
<abbrev id="ABBRID0E5ABG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.16,
<abbrev id="ABBRID0ECBBG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EGBBG" xlink_title="posterior median eyes">PME</abbrev>
0.12; labium 0.81 long and 0.58 wide; sternum 2.59 long and 1.01 wide; legs: I 19.58 (5.67, 2.78, 4.03, 4.68, 2.42), II 20.42 (5.66, 2.78, 4.13, 4.96, 2.89), III 22.62 (5.96, 2.63, 4.54, 6.29, 3.20), IV 28.26 (7.16, 2.75, 5.82, 8.55, 3.98).
</paragraph>
<paragraph id="13DEF346FCB55E63AA0F505965C689DE">
Palp: tibial apophysis (
<abbrev id="ABBRID0EMBBG" xlink_title="male palps bearing tibial apophysis">TiA</abbrev>
) with four setae of similar length (Fig.
<figureCitation id="18F4A67B9B7504651D213AD9E8020364" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 A C</figureCitation>
); paracymbium (
<abbrev id="ABBRID0EUBBG" xlink_title="paracymbium">PC</abbrev>
) with several setae situated on tip (Fig.
<figureCitation id="8D5B01052486702AA3B803E844EE00F1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 A C</figureCitation>
); cumulus (
<abbrev id="ABBRID0E3BBG" xlink_title="cumulus">Cu</abbrev>
) slightly elevated, with 8 tapering spines (Fig.
<figureCitation id="DAB077CD2E370910E8412BD3C22A9874" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 A C</figureCitation>
); subtegulum (ST) with weakly developed apophysis (Fig.
<figureCitation id="AB1EF823FC0F717952355E33E5B94DDF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 B, F</figureCitation>
); contrategulum (
<abbrev id="ABBRID0EICBG" xlink_title="contrategulum">CT</abbrev>
) with an arched projection distally, and triangular process with wide base (Fig.
<figureCitation id="F746B0CE3585B6394BF8A74A91C2ADD5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 D</figureCitation>
); tegulum (
<collectionCode id="D62A7A93BB629BC587F8E76B70679944" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
) with distinct transversal ridge in retrolateral view (Fig.
<figureCitation id="07C3CCC9B4432F82F6C5FFAF95037B28" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 C, F</figureCitation>
); paraembolic plate (PeP) narrow, sclerotized (Fig.
<figureCitation id="17A39763E185FE2B72C2430FA3F76076" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 A, F</figureCitation>
); embolus (E) with several longitudinal ridges reaching tip (Fig.
<figureCitation id="D637E3432CB30856623963395F26BD1F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 E, F</figureCitation>
).
</paragraph>
<paragraph id="F6BF0DB69BC88FEB5B9D6A6430F9FF46">
<emphasis id="4F9B2F36A76E9DE6AE1B7F8DAF7B2521" bold="true">
Female
<typeStatus id="27DCE62D2D9F0711C1B19C8B34FC31ED" type="paratype">paratype</typeStatus>
(XUX- 2019-053).
</emphasis>
Carapace yellowish brown with few short, scattered bristles; opisthosoma brown, with 12 brown tergites, close to each other, with gray patches, 26
<superScript id="BC76F4729343AF25F0626CE05492E997">th</superScript>
larger than others, 5
<superScript id="A806F6080CD91B54E609DD60312A03CF">th</superScript>
largest; chelicerae robust, reddish brown; promargin of cheliceral groove with 12 denticles of variable size; labium and sternum yellowish brown; legs with strong setae and spines. Measurements:
<abbrev id="ABBRID0EEDBG" xlink_title="body length">BL</abbrev>
14.24,
<abbrev id="ABBRID0EIDBG" xlink_title="carapace length">CL</abbrev>
6.10,
<abbrev id="ABBRID0EMDBG" xlink_title="carapace width">CW</abbrev>
5.30,
<abbrev id="ABBRID0EQDBG" xlink_title="opisthosoma length">OL</abbrev>
7.12,
<abbrev id="ABBRID0EUDBG" xlink_title="opisthosoma width">OW</abbrev>
4.96; eye sizes and interdistances:
<abbrev id="ABBRID0EYDBG" xlink_title="anterior median eyes">AME</abbrev>
0.08,
<abbrev id="ABBRID0E3DBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.57,
<abbrev id="ABBRID0EAEBG" xlink_title="posterior median eyes">PME</abbrev>
0.31,
<abbrev id="ABBRID0EEEBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.45,
<abbrev id="ABBRID0EIEBG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EMEBG" xlink_title="anterior median eyes">AME</abbrev>
0.10,
<abbrev id="ABBRID0EQEBG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EUEBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.17,
<abbrev id="ABBRID0EYEBG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0E3EBG" xlink_title="posterior median eyes">PME</abbrev>
0.09,
<abbrev id="ABBRID0EAFBG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0EEFBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.08,
<abbrev id="ABBRID0EIFBG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EMFBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.09,
<abbrev id="ABBRID0EQFBG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EUFBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.10,
<abbrev id="ABBRID0EYFBG" xlink_title="posterior lateral eyes">PLE</abbrev>
<abbrev id="ABBRID0E3FBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.46,
<abbrev id="ABBRID0EAGBG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EEGBG" xlink_title="posterior median eyes">PME</abbrev>
0.06; labium 1.23 long and 0.77 wide; sternum 2.68 long and 1.13 wide; palp 10.42 (3.63, 1.84, 2.77, 2.18), leg I 13.33 (4.16, 2.15, 2.90, 2.58, 1.54), II 16.85 (4.35, 2.26, 2.84, 2.84, 1.72), III 15.04 (4.37, 2.35, 2.90, 3.31, 2.11), IV 20.39 (5.92, 1.92, 4.48, 5.33, 2.74).
</paragraph>
<paragraph id="6FD506E4523891E20ABF022291F4FF64">
Vulva: poreplate (
<abbrev id="ABBRID0EKGBG" xlink_title="sclerotized poreplate">PPl</abbrev>
) with pair of large anterior lobes and pair of relatively small anterolateral lobes; central dorsal opening (
<collectionCode id="E46B961EB81896055B2684E7E78BFDEE">CDO</collectionCode>
) located at center of poreplate (
<abbrev id="ABBRID0EOGBG" xlink_title="sclerotized poreplate">PPl</abbrev>
); receptacular cluster (
<abbrev id="ABBRID0ESGBG" xlink_title="receptacular cluster">
<collectionCode id="87CBA80D545C8628F27374CFF471848D">RC</collectionCode>
</abbrev>
) racemose, protrudes upper edge of poreplate (
<abbrev id="ABBRID0EWGBG" xlink_title="sclerotized poreplate">PPl</abbrev>
); posterior margin of triangular posterior stalk (
<collectionCode id="3A30862C8C429FB335C0B1580DB7DC29">PS</collectionCode>
) almost straight; lateral margins of genital atrium (GA) slightly sclerotized (Fig.
<figureCitation id="C3FC2A0EE0676BD640A01CF5E8CB357C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Vulva of Liphistius nawngau sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure8" httpUri="https://binary.pensoft.net/fig/1115134">8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5GBG" type="variation">
<paragraph id="2B90B85B60376980224C958908C5A36A">
<heading id="068D6CC6F97FE8644086276C31A6E78E" reason="title">Variation.</heading>
</paragraph>
<paragraph id="84ACCB9E393456A9DA61EB3D0812AE94">
Males (
<emphasis id="BE05782BC2912D2287482EB475FA6084" italics="true">N</emphasis>
= 14):
<abbrev id="ABBRID0EGHBG" xlink_title="body length">BL</abbrev>
12.6416.98,
<abbrev id="ABBRID0EKHBG" xlink_title="carapace length">CL</abbrev>
6.277.56,
<abbrev id="ABBRID0EOHBG" xlink_title="carapace width">CW</abbrev>
6.027.56,
<abbrev id="ABBRID0ESHBG" xlink_title="opisthosoma length">OL</abbrev>
6.958.50,
<abbrev id="ABBRID0EWHBG" xlink_title="opisthosoma width">OW</abbrev>
4.606.40; females (
<emphasis id="C6B7BEA17496CFB7EFDBDCC5EFA7F2D2" italics="true">N</emphasis>
= 5):
<abbrev id="ABBRID0E3HBG" xlink_title="body length">BL</abbrev>
10.5514.24,
<abbrev id="ABBRID0EAIBG" xlink_title="carapace length">CL</abbrev>
5.266.37,
<abbrev id="ABBRID0EEIBG" xlink_title="carapace width">CW</abbrev>
4.475.63,
<abbrev id="ABBRID0EIIBG" xlink_title="opisthosoma length">OL</abbrev>
5.247.12,
<abbrev id="ABBRID0EMIBG" xlink_title="opisthosoma width">OW</abbrev>
4.334.96. The number of denticles on the promargin of male cheliceral groove varies from 10 to 13 (
<emphasis id="DA3EF92C33B33DB300E9323658E1FB7A" italics="true">N</emphasis>
= 12); in females, the number of denticles on the promargin of cheliceral groove varies from 11 to 12 (
<emphasis id="1D833BD5A78B28FC37ED503C44B73688" italics="true">N</emphasis>
= 5). The number of setae on tibial apophysis varies from 4 to 6.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUIBG" type="etymology">
<paragraph id="9547AE76BAB6288E82691D4CB49D6211">
<heading id="92856A1013A630F9BAF55F41E45B28F7" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="8B7EA0EAC35CA90B5E231292F4587935">
The species epithet
<taxonomicName id="1B0F9E4BB7AC45DAEC85C08312D6821A" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nawngau">
<emphasis id="8AEB66DE2593A547BBABD28E96214083" italics="true">nawngau</emphasis>
</taxonomicName>
” refers to the
<typeStatus id="DE374C4DA044D02FC1252004E9FA8C71">type</typeStatus>
locality, Nawng Au Village; it is treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDJBG" type="distribution">
<paragraph id="D5267B716836F9C3F677E8EB9BD2C3BD">
<heading id="891C354375E2ED1C6222187E3106511B" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="1D8EC46997F88337EE944622C3BE4074">
<collectingRegion id="CAAAE54240867612AB45B5794A10D990" country="Myanmar" name="Shan">Shan State</collectingRegion>
, (Kyaukme District),
<collectingCountry id="FCD4D0FA5A5AEE5063C962A6CB820501" name="Myanmar">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,396 @@
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<treatment id="2F15F14DFF92FF9F06DEABC1FE2BFA09" LSID="urn:lsid:plazi:treatment:2F15F14DFF92FF9F06DEABC1FE2BFA09" httpUri="http://treatment.plazi.org/id/2F15F14DFF92FF9F06DEABC1FE2BFA09" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
<subSubSection id="EFA613D0FF92FF9806DEABC1FAC8F949" box="[925,1355,1762,1788]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="A703405BFF92FF9806DEABC1FAC8F949" blockId="2.[925,1355,1762,1819]" box="[925,1355,1762,1788]" pageId="2" pageNumber="3">
<heading id="FC4BF737FF92FF9806DEABC1FAC8F949" box="[925,1355,1762,1788]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="3" reason="7">
<taxonomicName id="60BC3BD8FF92FF9806DEABC1FAC8F949" ID-CoL="867NV" authority="Gorjunova, 1970" authorityName="Gorjunova" authorityYear="1970" box="[925,1355,1762,1788]" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="2" pageNumber="3" phylum="Bryozoa" rank="species" species="enodata">
<emphasis id="95C89C49FF92FF9806DEABC1FBFCF949" box="[925,1151,1762,1787]" italics="true" pageId="2" pageNumber="3">Fistulipora enodata</emphasis>
<bibRefCitation id="C32D3DAAFF92FF9801C5ABC1FAC8F949" author="Gorjunova, R. V." box="[1158,1355,1762,1788]" pageId="2" pageNumber="3" pagination="60 - 68" refId="ref27324" refString="Gorjunova, R. V. 1970. New Early Permian bryozoans from the Southeast Darvaz and Pamir, p. 60 - 68. In Astrova, G. G. and Chudinova, I. I. (eds.), New Species of Palaeozoic Bryozoans and Corals. Nauka, Moscow. (In Russian)" type="book chapter" year="1970">Gorjunova, 1970</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EFA613D0FF92FF9806B9AA21FB82F8D3" pageId="2" pageNumber="3" type="description">
<paragraph id="A703405BFF92FF9806B9AA21FB6DF8A9" blockId="2.[925,1355,1762,1819]" box="[1018,1262,1794,1819]" pageId="2" pageNumber="3">
<figureCitation id="3F875CDEFF92FF9806B9AA21FB0EF8A9" box="[1018,1165,1794,1819]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[193,283,1788,1811]" captionStartId-1="7.[200,291,1770,1793]" captionTargetBox-0="[211,1417,164,1758]" captionTargetBox-1="[220,1410,166,1737]" captionTargetId-0="figure-13@4.[211,1417,164,1758]" captionTargetId-1="figure-17@7.[220,1410,166,1738]" captionTargetPageId-0="4" captionTargetPageId-1="7" captionText-0="FIGURE 3. Thin section photographs of Fistulipora enodata Gorjunova, 1970, SMF 23.014 (14); Fistulipora guttata Trizna and Klautzan, 1961, SMF 23.025 (58); and Fistulipora sakagamii n. sp., holotype SMF 23.028 (9 and 10). 1 and 5, longitudinal section showing autozooecial chambers and vesicular skeleton; and 24, 68, 9, and 10, tangential sections showing autozooecial apertures and vesicles." captionText-1="FIGURE 4. Thin section photographs of Fistulipora sakagamii n. sp., holotype SMF 23.028 (1 and 2); Dybowskiella hupehensiformis n. sp., holotype SMF 23.030 and paratype SMF 23.032 (37); and Fistuliramus xianzaensis Liu and Wang, 1987, SMF 23.037 (8). 1, 2, 57, tangential section showing autozooecial apertures and vesicles; 3 and 4, longitudinal section showing autozooecial chambers and vesicular skeleton; and 8, branch transverse section." pageId="2" pageNumber="3">Figure 3.1-4</figureCitation>
;
<tableCitation id="EA3E75E0FF92FF9801DBAA21FB6DF8A9" box="[1176,1262,1794,1819]" captionStart="TABLE 1" captionStartId="5.[190,268,164,187]" captionText="TABLE 1. Measurements of Fistulipora enodata Gorjunova, 1970. Abbreviations: N = number of measurements; X = mean; SD = standard deviation; CV = coefficient of variation; MIN = minimal value; MAX = maximal value." pageId="2" pageNumber="3">Table 1</tableCitation>
</paragraph>
<paragraph id="A703405BFF92FF98060BAA0EFB82F8D3" blockId="2.[840,1440,1836,1889]" pageId="2" pageNumber="3">
1970
<taxonomicName id="60BC3BD8FF92FF980685AA0EFA8EF8F1" authority="Gorjunova" authorityName="Gorjunova" authorityYear="1970" box="[966,1293,1836,1860]" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="2" pageNumber="3" phylum="Bryozoa" rank="species" species="enodata">
<emphasis id="95C89C49FF92FF980685AA0EFB10F8F6" box="[966,1171,1837,1860]" italics="true" pageId="2" pageNumber="3">Fistulipora enodata</emphasis>
Gorjunova
</taxonomicName>
, p. 61, pl. 21, fig. 4.
</paragraph>
</subSubSection>
<caption id="F3C310D3FF93FF990587A8E5FBEFF984" pageId="3" pageNumber="4" startId="3.[196,287,1478,1501]" targetBox="[198,1436,170,1446]" targetPageId="3" targetType="figure">
<paragraph id="A703405BFF93FF990587A8E5FBEFF984" blockId="3.[196,1432,1478,1590]" pageId="3" pageNumber="4">
<emphasis id="95C89C49FF93FF990587A8E5FEBAFA6C" bold="true" box="[196,313,1478,1502]" pageId="3" pageNumber="4">FIGURE 2.</emphasis>
Lithological characteristics of the limestones from the Zhongba Formation. 1, limestone sample with abundant bryozoans (SMF 23.262); 2, rudstone containing crinoids, cystoporate (
<taxonomicName id="60BC3BD8FF93FF990147A8C7FA11FA49" authority="Gorjunova, 1970" authorityName="Gorjunova" authorityYear="1970" box="[1028,1426,1507,1531]" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="enodata">
<emphasis id="95C89C49FF93FF990147A8C7FB51FA49" box="[1028,1234,1508,1531]" italics="true" pageId="3" pageNumber="4">Fistulipora enodata</emphasis>
<bibRefCitation id="C32D3DAAFF93FF99019FA8C0FA11FA49" author="Gorjunova, R. V." box="[1244,1426,1507,1531]" pageId="3" pageNumber="4" pagination="60 - 68" refId="ref27324" refString="Gorjunova, R. V. 1970. New Early Permian bryozoans from the Southeast Darvaz and Pamir, p. 60 - 68. In Astrova, G. G. and Chudinova, I. I. (eds.), New Species of Palaeozoic Bryozoans and Corals. Nauka, Moscow. (In Russian)" type="book chapter" year="1970">Gorjunova, 1970</bibRefCitation>
</taxonomicName>
) and fenestrate bryozoans (SMF 23.263); 3, bindstones with
<taxonomicName id="60BC3BD8FF93FF990621AB22FAD1F9AA" authority="Trizna and Klautzan, 1961" authorityName="Trizna and Klautzan" authorityYear="1961" box="[866,1362,1537,1560]" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="guttata">
<emphasis id="95C89C49FF93FF990621AB22FBA7F9AA" box="[866,1060,1537,1560]" italics="true" pageId="3" pageNumber="4">Fistulipora guttata</emphasis>
<bibRefCitation id="C32D3DAAFF93FF99016CAB22FAD1F9AA" author="Trizna, V. B. &amp; Klautzan, R. A." box="[1071,1362,1537,1560]" pageId="3" pageNumber="4" pagination="331 - 453" refId="ref30392" refString="Trizna, V. B. and Klautzan, R. A. 1961. Bryozoans of the Artinskian of the Ufimian Plateau and their role in the stratigraphy of this stage in western Urals. Trudy Vsesojuznogo Nauchnoissledovatelskogo Geologo-Razvedochnogo Instituta (VNIGRI), 179 th edition, Microfauna of the USSR, 13: 331 - 453. (In Russian)" type="journal article" year="1961">Trizna and Klautzan, 1961</bibRefCitation>
</taxonomicName>
(SMF 23.264); and 4, grain- to packstones with bryozoan and crinoid fragments (SMF 23.265).
</paragraph>
</caption>
<subSubSection id="EFA613D0FF93FF990583AB42FDA5F880" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="A703405BFF93FF990583AB42FDBBF927" blockId="3.[192,792,1632,1879]" pageId="3" pageNumber="4">
1975
<taxonomicName id="60BC3BD8FF93FF99047DAB42FD49F9CA" ID-CoL="867NV" authority="Gorjunova, 1970" authorityName="Gorjunova" authorityYear="1970" box="[318,714,1632,1656]" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="enodata">
<emphasis id="95C89C49FF93FF99047DAB42FD88F9CA" box="[318,523,1633,1656]" italics="true" pageId="3" pageNumber="4">Fistulipora enodata</emphasis>
<bibRefCitation id="C32D3DAAFF93FF990757AB43FD49F9CA" author="Gorjunova, R. V." box="[532,714,1632,1656]" pageId="3" pageNumber="4" pagination="60 - 68" refId="ref27324" refString="Gorjunova, R. V. 1970. New Early Permian bryozoans from the Southeast Darvaz and Pamir, p. 60 - 68. In Astrova, G. G. and Chudinova, I. I. (eds.), New Species of Palaeozoic Bryozoans and Corals. Nauka, Moscow. (In Russian)" type="book chapter" year="1970">Gorjunova, 1970</bibRefCitation>
</taxonomicName>
; Gorjunova, p. 42, pl. 1, fig. 1.
</paragraph>
<paragraph id="A703405BFF93FF990583AB82FE33F940" blockId="3.[192,792,1632,1879]" pageId="3" pageNumber="4">
1987
<taxonomicName id="60BC3BD8FF93FF99047DAB82FD67F90A" ID-CoL="867NV" authority="Gorjunova, 1970" authorityName="Gorjunova" authorityYear="1970" box="[318,740,1696,1720]" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="enodata">
<emphasis id="95C89C49FF93FF99047DAB82FD97F90A" box="[318,532,1697,1720]" italics="true" pageId="3" pageNumber="4">Fistulipora enodata</emphasis>
<bibRefCitation id="C32D3DAAFF93FF990766AB83FD67F90A" author="Gorjunova, R. V." box="[549,740,1696,1720]" pageId="3" pageNumber="4" pagination="60 - 68" refId="ref27324" refString="Gorjunova, R. V. 1970. New Early Permian bryozoans from the Southeast Darvaz and Pamir, p. 60 - 68. In Astrova, G. G. and Chudinova, I. I. (eds.), New Species of Palaeozoic Bryozoans and Corals. Nauka, Moscow. (In Russian)" type="book chapter" year="1970">Gorjunova, 1970</bibRefCitation>
</taxonomicName>
; Liu and Wang, p. 4, pl. 1, figs. 4a-b, pl. 2, figs. 1-a-b.
</paragraph>
<paragraph id="A703405BFF93FF990583ABDDFDA5F880" blockId="3.[192,792,1632,1879]" pageId="3" pageNumber="4">
1991
<taxonomicName id="60BC3BD8FF93FF99047DABDDFD4AF8A7" ID-CoL="867NV" authority="Gorjunova, 1970" authorityName="Gorjunova" authorityYear="1970" box="[318,713,1790,1813]" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="enodata">
<emphasis id="95C89C49FF93FF99047DABDDFD88F8A7" box="[318,523,1790,1813]" italics="true" pageId="3" pageNumber="4">Fistulipora enodata</emphasis>
<bibRefCitation id="C32D3DAAFF93FF990750ABDDFD4AF8A7" author="Gorjunova, R. V." box="[531,713,1790,1813]" pageId="3" pageNumber="4" pagination="60 - 68" refId="ref27324" refString="Gorjunova, R. V. 1970. New Early Permian bryozoans from the Southeast Darvaz and Pamir, p. 60 - 68. In Astrova, G. G. and Chudinova, I. I. (eds.), New Species of Palaeozoic Bryozoans and Corals. Nauka, Moscow. (In Russian)" type="book chapter" year="1970">Gorjunova, 1970</bibRefCitation>
</taxonomicName>
; Xia, p. 957, pl. 5, figs. 13-14.
</paragraph>
</subSubSection>
<subSubSection id="EFA613D0FF93FF990583AA1DFDEEF8E5" box="[192,621,1853,1879]" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="A703405BFF93FF990583AA1DFDEEF8E5" blockId="3.[192,792,1632,1879]" box="[192,621,1853,1879]" pageId="3" pageNumber="4">
<emphasis id="95C89C49FF93FF990583AA1DFEAFF8E5" bold="true" box="[192,300,1854,1879]" pageId="3" pageNumber="4">Material.</emphasis>
<collectionCode id="C1ADD89EFF93FF990477AA1EFEEFF8E5" box="[308,364,1853,1879]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="4">SMF</collectionCode>
23.007
<collectionCode id="C1ADD89EFF93FF990490AA1EFD8FF8E5" box="[467,524,1853,1879]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="4">SMF</collectionCode>
23.021.
</paragraph>
</subSubSection>
<subSubSection id="EFA613D0FF93FF9F060BAB43FEB5FBC9" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="description">
<paragraph id="A703405BFF93FF9F060BAB43FEA1FD8C" blockId="3.[840,1441,1632,1913]" lastBlockId="5.[190,290,551,574]" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
<emphasis id="95C89C49FF93FF99060BAB43FC61F9CB" bold="true" box="[840,994,1632,1657]" pageId="3" pageNumber="4">Description.</emphasis>
Encrusting, partly multilayered colony, separate sheets
<quantity id="6044EDBEFF93FF99010AABA3FB55F92B" box="[1097,1238,1664,1689]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.8" metricValueMin="1.8" pageId="3" pageNumber="4" unit="mm" value="2.3" valueMax="2.8" valueMin="1.8">1.82.8 mm</quantity>
thick. Autozooecia growing from thin epitheca, bending at their bases to the colony surface. Autozooecial diaphragms rare to absent, thin. Autozooecial apertures circular to oval. Lunaria well-developed, rounded, disappearing in deeper sections; ends of lunaria not indenting into autozooecia. Vesicles small to medium, high, separating autozooecia in
<heading id="FC4BF737FF95FF9F05FDAF04FEA1FD8C" allCaps="true" bold="true" box="[190,290,551,574]" fontSize="36" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="95C89C49FF95FF9F05FDAF04FEA1FD8C" bold="true" box="[190,290,551,574]" pageId="5" pageNumber="6">TABLE 2</emphasis>
</heading>
</paragraph>
<caption id="F3C310D3FF94FF9E0582ABDFFC80F8DE" pageId="4" pageNumber="5" startId="4.[193,283,1788,1811]" targetBox="[211,1417,164,1758]" targetPageId="4" targetType="figure">
<paragraph id="A703405BFF94FF9E0582ABDFFC80F8DE" blockId="4.[193,1435,1788,1900]" pageId="4" pageNumber="5">
<emphasis id="95C89C49FF94FF9E0582ABDFFEBBF8A6" bold="true" box="[193,312,1788,1812]" pageId="4" pageNumber="5">FIGURE 3.</emphasis>
Thin section photographs of
<taxonomicName id="60BC3BD8FF94FF9E0730ABDEFC7DF8A6" authority="Gorjunova, 1970" authorityName="Gorjunova" authorityYear="1970" box="[627,1022,1788,1812]" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="4" pageNumber="5" phylum="Bryozoa" rank="species" species="enodata">
<emphasis id="95C89C49FF94FF9E0730ABDEFCC3F8A6" box="[627,832,1789,1812]" italics="true" pageId="4" pageNumber="5">Fistulipora enodata</emphasis>
<bibRefCitation id="C32D3DAAFF94FF9E060BABDFFC7DF8A6" author="Gorjunova, R. V." box="[840,1022,1788,1812]" pageId="4" pageNumber="5" pagination="60 - 68" refId="ref27324" refString="Gorjunova, R. V. 1970. New Early Permian bryozoans from the Southeast Darvaz and Pamir, p. 60 - 68. In Astrova, G. G. and Chudinova, I. I. (eds.), New Species of Palaeozoic Bryozoans and Corals. Nauka, Moscow. (In Russian)" type="book chapter" year="1970">Gorjunova, 1970</bibRefCitation>
</taxonomicName>
, SMF 23.014 (14);
<taxonomicName id="60BC3BD8FF94FF9E019FABDEFE5CF883" authority="Trizna and Klautzan, 1961" authorityName="Trizna and Klautzan" authorityYear="1961" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="4" pageNumber="5" phylum="Bryozoa" rank="species" species="guttata">
<emphasis id="95C89C49FF94FF9E019FABDEFA18F8A6" box="[1244,1435,1789,1812]" italics="true" pageId="4" pageNumber="5">Fistulipora guttata</emphasis>
<bibRefCitation id="C32D3DAAFF94FF9E0582AA39FE5CF883" author="Trizna, V. B. &amp; Klautzan, R. A." box="[193,479,1818,1841]" pageId="4" pageNumber="5" pagination="331 - 453" refId="ref30392" refString="Trizna, V. B. and Klautzan, R. A. 1961. Bryozoans of the Artinskian of the Ufimian Plateau and their role in the stratigraphy of this stage in western Urals. Trudy Vsesojuznogo Nauchnoissledovatelskogo Geologo-Razvedochnogo Instituta (VNIGRI), 179 th edition, Microfauna of the USSR, 13: 331 - 453. (In Russian)" type="journal article" year="1961">Trizna and Klautzan, 1961</bibRefCitation>
</taxonomicName>
, SMF 23.025 (58); and
<taxonomicName id="60BC3BD8FF94FF9E07AFAA39FC4CF883" authorityName="Ernst" authorityYear="2016" box="[748,975,1818,1841]" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="4" pageNumber="5" phylum="Bryozoa" rank="species" species="sakagamii" status="sp. nov.">
<emphasis id="95C89C49FF94FF9E07AFAA39FC4CF883" box="[748,975,1818,1841]" italics="true" pageId="4" pageNumber="5">Fistulipora sakagamii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8EFB2132FF94FF9E0694AA39FB95F883" box="[983,1046,1818,1841]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
, holotype SMF 23.028 (9 and 10). 1 and 5, longitudinal section showing autozooecial chambers and vesicular skeleton; and 24, 68, 9, and 10, tangential sections showing autozooecial apertures and vesicles.
</paragraph>
</caption>
<caption id="F3C310D3FF95FF9F05FDAD87FA9EFF6B" pageId="5" pageNumber="6" startId="5.[190,268,164,187]" targetType="table">
<paragraph id="A703405BFF95FF9F05FDAD87FA9EFF6B" blockId="5.[190,1444,164,217]" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F05FDAD87FEAAFF09" bold="true" box="[190,297,164,187]" pageId="5" pageNumber="6">TABLE 1.</emphasis>
Measurements of
<taxonomicName id="60BC3BD8FF95FF9F04B7AD87FCFCFF09" authority="Gorjunova, 1970" authorityName="Gorjunova" authorityYear="1970" box="[500,895,164,187]" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="5" pageNumber="6" phylum="Bryozoa" rank="species" species="enodata">
<emphasis id="95C89C49FF95FF9F04B7AD87FD43FF09" box="[500,704,164,187]" italics="true" pageId="5" pageNumber="6">Fistulipora enodata</emphasis>
<bibRefCitation id="C32D3DAAFF95FF9F078AAD87FCFCFF09" author="Gorjunova, R. V." box="[713,895,164,187]" pageId="5" pageNumber="6" pagination="60 - 68" refId="ref27324" refString="Gorjunova, R. V. 1970. New Early Permian bryozoans from the Southeast Darvaz and Pamir, p. 60 - 68. In Astrova, G. G. and Chudinova, I. I. (eds.), New Species of Palaeozoic Bryozoans and Corals. Nauka, Moscow. (In Russian)" type="book chapter" year="1970">Gorjunova, 1970</bibRefCitation>
</taxonomicName>
. Abbreviations: N = number of measurements; X = mean; SD = standard deviation; CV = coefficient of variation; MIN = minimal value; MAX = maximal value.
</paragraph>
</caption>
<paragraph id="A703405BFF95FF9F078DADCCFAAAFC23" pageId="5" pageNumber="6">
<table id="D5BCB2FBFF9500650461ADCCFAA8FC24" box="[290,1323,239,918]" colsContinueIn="6.[309,1319,206,490]" gridcols="7" gridrows="18" pageId="5" pageNumber="6">
<tr id="198C4219FF9500650461ADCCFAA8FEB1" box="[290,1323,239,259]" gridrow="0" pageId="5" pageNumber="6" rowspan-0="1">
<th id="5A5D2B65FF9500650789ADCCFD6BFEB1" box="[714,744,239,259]" gridcol="1" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F078DADCCFD5EFEB1" bold="true" box="[718,733,239,259]" pageId="5" pageNumber="6">N</emphasis>
</th>
<th id="5A5D2B65FF9500650670ADCCFCE1FEB1" box="[819,866,239,259]" gridcol="2" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F0603ADCCFCCDFEB1" bold="true" box="[832,846,239,259]" pageId="5" pageNumber="6">X</emphasis>
</th>
<th id="5A5D2B65FF95006506DDADCCFC59FEB1" box="[926,986,239,259]" gridcol="3" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F06E9ADCCFC4BFEB1" bold="true" box="[938,968,239,259]" pageId="5" pageNumber="6">SD</emphasis>
</th>
<th id="5A5D2B65FF9500650153ADCCFBC8FEB1" box="[1040,1099,239,259]" gridcol="4" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F0158ADCCFBBAFEB1" bold="true" box="[1051,1081,239,259]" pageId="5" pageNumber="6">CV</emphasis>
</th>
<th id="5A5D2B65FF95006501C4ADCCFB35FEB1" box="[1159,1206,239,259]" gridcol="5" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F01CBADCCFB2CFEB1" bold="true" box="[1160,1199,239,259]" pageId="5" pageNumber="6">MIN</emphasis>
</th>
<th id="5A5D2B65FF95006501B5ADCCFAA8FEB1" box="[1270,1323,239,259]" gridcol="6" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F01B5ADCCFAA6FEB1" bold="true" box="[1270,1317,239,259]" pageId="5" pageNumber="6">MAX</emphasis>
</th>
</tr>
<tr id="198C4219FF9500650461AC31FAA8FE99" box="[290,1323,274,299]" gridrow="1" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461AC31FD05FE99" box="[290,646,274,299]" gridcol="0" gridrow="1" pageId="5" pageNumber="6">Autozooecial aperture width, mm</th>
<td id="5A5D2B65FF9500650789AC31FD6BFE99" box="[714,744,274,299]" gridcol="1" gridrow="1" pageId="5" pageNumber="6">35</td>
<td id="5A5D2B65FF9500650670AC31FCE1FE99" box="[819,866,274,299]" gridcol="2" gridrow="1" pageId="5" pageNumber="6">0.30</td>
<td id="5A5D2B65FF95006506DDAC31FC59FE99" box="[926,986,274,299]" gridcol="3" gridrow="1" pageId="5" pageNumber="6">0.032</td>
<td id="5A5D2B65FF9500650153AC31FBC8FE99" box="[1040,1099,274,299]" gridcol="4" gridrow="1" pageId="5" pageNumber="6">10.87</td>
<td id="5A5D2B65FF95006501C4AC31FB35FE99" box="[1159,1206,274,299]" gridcol="5" gridrow="1" pageId="5" pageNumber="6">0.24</td>
<td id="5A5D2B65FF95006501B5AC31FAA8FE99" box="[1270,1323,274,299]" gridcol="6" gridrow="1" pageId="5" pageNumber="6">0.38</td>
</tr>
<tr id="198C4219FF9500650461AC14FAA8FEE3" box="[290,1323,311,337]" gridrow="2" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461AC14FD05FEE3" box="[290,646,311,337]" gridcol="0" gridrow="2" pageId="5" pageNumber="6">Autozooecial aperture spacing, mm</th>
<td id="5A5D2B65FF9500650789AC14FD6BFEE3" box="[714,744,311,337]" gridcol="1" gridrow="2" pageId="5" pageNumber="6">35</td>
<td id="5A5D2B65FF9500650670AC14FCE1FEE3" box="[819,866,311,337]" gridcol="2" gridrow="2" pageId="5" pageNumber="6">0.58</td>
<td id="5A5D2B65FF95006506DDAC14FC59FEE3" box="[926,986,311,337]" gridcol="3" gridrow="2" pageId="5" pageNumber="6">0.067</td>
<td id="5A5D2B65FF9500650153AC14FBC8FEE3" box="[1040,1099,311,337]" gridcol="4" gridrow="2" pageId="5" pageNumber="6">11.47</td>
<td id="5A5D2B65FF95006501C4AC14FB35FEE3" box="[1159,1206,311,337]" gridcol="5" gridrow="2" pageId="5" pageNumber="6">0.45</td>
<td id="5A5D2B65FF95006501B5AC14FAA8FEE3" box="[1270,1323,311,337]" gridcol="6" gridrow="2" pageId="5" pageNumber="6">0.75</td>
</tr>
<tr id="198C4219FF9500650461AC7EFAA8FEC4" box="[290,1323,349,374]" gridrow="3" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461AC7EFD05FEC4" box="[290,646,349,374]" gridcol="0" gridrow="3" pageId="5" pageNumber="6">Vesicle diameter, mm</th>
<td id="5A5D2B65FF9500650789AC7EFD6BFEC4" box="[714,744,349,374]" gridcol="1" gridrow="3" pageId="5" pageNumber="6">35</td>
<td id="5A5D2B65FF9500650670AC7EFCE1FEC4" box="[819,866,349,374]" gridcol="2" gridrow="3" pageId="5" pageNumber="6">0.13</td>
<td id="5A5D2B65FF95006506DDAC7EFC59FEC4" box="[926,986,349,374]" gridcol="3" gridrow="3" pageId="5" pageNumber="6">0.034</td>
<td id="5A5D2B65FF9500650153AC7EFBC8FEC4" box="[1040,1099,349,374]" gridcol="4" gridrow="3" pageId="5" pageNumber="6">27.24</td>
<td id="5A5D2B65FF95006501C4AC7EFB35FEC4" box="[1159,1206,349,374]" gridcol="5" gridrow="3" pageId="5" pageNumber="6">0.07</td>
<td id="5A5D2B65FF95006501B5AC7EFAA8FEC4" box="[1270,1323,349,374]" gridcol="6" gridrow="3" pageId="5" pageNumber="6">0.21</td>
</tr>
<tr id="198C4219FF9500650461ACA1FAA8FE29" box="[290,1323,386,411]" gridrow="4" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461ACA1FD05FE29" box="[290,646,386,411]" gridcol="0" gridrow="4" pageId="5" pageNumber="6">Vesicle spacing, mm</th>
<td id="5A5D2B65FF9500650789ACA1FD6BFE29" box="[714,744,386,411]" gridcol="1" gridrow="4" pageId="5" pageNumber="6">35</td>
<td id="5A5D2B65FF9500650670ACA1FCE1FE29" box="[819,866,386,411]" gridcol="2" gridrow="4" pageId="5" pageNumber="6">0.11</td>
<td id="5A5D2B65FF95006506DDACA1FC59FE29" box="[926,986,386,411]" gridcol="3" gridrow="4" pageId="5" pageNumber="6">0.025</td>
<td id="5A5D2B65FF9500650153ACA1FBC8FE29" box="[1040,1099,386,411]" gridcol="4" gridrow="4" pageId="5" pageNumber="6">21.80</td>
<td id="5A5D2B65FF95006501C4ACA1FB35FE29" box="[1159,1206,386,411]" gridcol="5" gridrow="4" pageId="5" pageNumber="6">0.08</td>
<td id="5A5D2B65FF95006501B5ACA1FAA8FE29" box="[1270,1323,386,411]" gridcol="6" gridrow="4" pageId="5" pageNumber="6">0.17</td>
</tr>
<tr id="198C4219FF9500650461AC84FAA8FE73" box="[290,1323,423,449]" gridrow="5" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461AC84FD05FE73" box="[290,646,423,449]" gridcol="0" gridrow="5" pageId="5" pageNumber="6">Vesicles per aperture</th>
<td id="5A5D2B65FF9500650789AC84FD6BFE73" box="[714,744,423,449]" gridcol="1" gridrow="5" pageId="5" pageNumber="6">20</td>
<td id="5A5D2B65FF9500650670AC84FCE1FE73" box="[819,866,423,449]" gridcol="2" gridrow="5" pageId="5" pageNumber="6">12.6</td>
<td id="5A5D2B65FF95006506DDAC84FC59FE73" box="[926,986,423,449]" gridcol="3" gridrow="5" pageId="5" pageNumber="6">1.188</td>
<td id="5A5D2B65FF9500650153AC84FBC8FE73" box="[1040,1099,423,449]" gridcol="4" gridrow="5" pageId="5" pageNumber="6">9.43</td>
<td id="5A5D2B65FF95006501C4AC84FB35FE73" box="[1159,1206,423,449]" gridcol="5" gridrow="5" pageId="5" pageNumber="6">11.0</td>
<td id="5A5D2B65FF95006501B5AC84FAA8FE73" box="[1270,1323,423,449]" gridcol="6" gridrow="5" pageId="5" pageNumber="6">15.0</td>
</tr>
<tr id="198C4219FF9500650461ACEEFAA8FE54" box="[290,1323,461,486]" gridrow="6" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461ACEEFD05FE54" box="[290,646,461,486]" gridcol="0" gridrow="6" pageId="5" pageNumber="6">Lunarium length, mm</th>
<td id="5A5D2B65FF9500650789ACEEFD6BFE54" box="[714,744,461,486]" gridcol="1" gridrow="6" pageId="5" pageNumber="6">20</td>
<td id="5A5D2B65FF9500650670ACEEFCE1FE54" box="[819,866,461,486]" gridcol="2" gridrow="6" pageId="5" pageNumber="6">0.14</td>
<td id="5A5D2B65FF95006506DDACEEFC59FE54" box="[926,986,461,486]" gridcol="3" gridrow="6" pageId="5" pageNumber="6">0.024</td>
<td id="5A5D2B65FF9500650153ACEEFBC8FE54" box="[1040,1099,461,486]" gridcol="4" gridrow="6" pageId="5" pageNumber="6">17.26</td>
<td id="5A5D2B65FF95006501C4ACEEFB35FE54" box="[1159,1206,461,486]" gridcol="5" gridrow="6" pageId="5" pageNumber="6">0.10</td>
<td id="5A5D2B65FF95006501B5ACEEFAA8FE54" box="[1270,1323,461,486]" gridcol="6" gridrow="6" pageId="5" pageNumber="6">0.18</td>
</tr>
<tr id="198C4219FF9500650461ACD1FAA8FDB9" box="[290,1323,498,523]" gridrow="7" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461ACD1FD05FDB9" box="[290,646,498,523]" gridcol="0" gridrow="7" pageId="5" pageNumber="6">Lunarium width, mm</th>
<td id="5A5D2B65FF9500650789ACD1FD6BFDB9" box="[714,744,498,523]" gridcol="1" gridrow="7" pageId="5" pageNumber="6">20</td>
<td id="5A5D2B65FF9500650670ACD1FCE1FDB9" box="[819,866,498,523]" gridcol="2" gridrow="7" pageId="5" pageNumber="6">0.25</td>
<td id="5A5D2B65FF95006506DDACD1FC59FDB9" box="[926,986,498,523]" gridcol="3" gridrow="7" pageId="5" pageNumber="6">0.028</td>
<td id="5A5D2B65FF9500650153ACD1FBC8FDB9" box="[1040,1099,498,523]" gridcol="4" gridrow="7" pageId="5" pageNumber="6">11.30</td>
<td id="5A5D2B65FF95006501C4ACD1FB35FDB9" box="[1159,1206,498,523]" gridcol="5" gridrow="7" pageId="5" pageNumber="6">0.20</td>
<td id="5A5D2B65FF95006501B5ACD1FAA8FDB9" box="[1270,1323,498,523]" gridcol="6" gridrow="7" pageId="5" pageNumber="6">0.29</td>
</tr>
<tr id="198C4219FF9500650461AF04FAA8FD8C" box="[290,1323,551,574]" gridrow="8" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461AF04FAA8FD8C" box="[290,1323,551,574]" colspan="7" colspanRight="6" gridcol="0" gridrow="8" pageId="5" pageNumber="6">
. Measurements of
<taxonomicName id="60BC3BD8FF95FF9F04AEAF04FC4FFD8C" authority="Trizna and Klautzan, 1961" authorityName="Trizna and Klautzan" authorityYear="1961" box="[493,972,551,574]" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="5" pageNumber="6" phylum="Bryozoa" rank="species" species="guttata">
<emphasis id="95C89C49FF95FF9F04AEAF04FD28FD8C" box="[493,683,551,574]" italics="true" pageId="5" pageNumber="6">Fistulipora guttata</emphasis>
<bibRefCitation id="C32D3DAAFF95FF9F07F2AF04FC4FFD8C" author="Trizna, V. B. &amp; Klautzan, R. A." box="[689,972,551,574]" pageId="5" pageNumber="6" pagination="331 - 453" refId="ref30392" refString="Trizna, V. B. and Klautzan, R. A. 1961. Bryozoans of the Artinskian of the Ufimian Plateau and their role in the stratigraphy of this stage in western Urals. Trudy Vsesojuznogo Nauchnoissledovatelskogo Geologo-Razvedochnogo Instituta (VNIGRI), 179 th edition, Microfauna of the USSR, 13: 331 - 453. (In Russian)" type="journal article" year="1961">Trizna and Klautzan, 1961</bibRefCitation>
</taxonomicName>
. Abbreviations as for Table 1.
</th>
</tr>
<tr id="198C4219FF9500650461AF77FAA8FDDB" box="[290,1323,596,617]" gridrow="9" pageId="5" pageNumber="6" rowspan-0="1">
<td id="5A5D2B65FF9500650789AF77FD6BFDDB" box="[714,744,596,617]" gridcol="1" gridrow="9" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F0796AF76FD67FDDB" bold="true" box="[725,740,597,617]" pageId="5" pageNumber="6">N</emphasis>
</td>
<td id="5A5D2B65FF9500650670AF77FCE1FDDB" box="[819,866,596,617]" gridcol="2" gridrow="9" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F0604AF76FCD6FDDB" bold="true" box="[839,853,597,617]" pageId="5" pageNumber="6">X</emphasis>
</td>
<td id="5A5D2B65FF95006506DDAF77FC59FDDB" box="[926,986,596,617]" gridcol="3" gridrow="9" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F06F3AF77FC4DFDDA" bold="true" box="[944,974,596,616]" pageId="5" pageNumber="6">SD</emphasis>
</td>
<td id="5A5D2B65FF9500650153AF77FBC8FDDB" box="[1040,1099,596,617]" gridcol="4" gridrow="9" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F0161AF77FBC3FDDA" bold="true" box="[1058,1088,596,616]" pageId="5" pageNumber="6">CV</emphasis>
</td>
<td id="5A5D2B65FF95006501C4AF77FB35FDDB" box="[1159,1206,596,617]" gridcol="5" gridrow="9" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F01CCAF76FB35FDDB" bold="true" box="[1167,1206,597,617]" pageId="5" pageNumber="6">MIN</emphasis>
</td>
<td id="5A5D2B65FF95006501B5AF77FAA8FDDB" box="[1270,1323,596,617]" gridcol="6" gridrow="9" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F01BFAF76FAA8FDDB" bold="true" box="[1276,1323,597,617]" pageId="5" pageNumber="6">MAX</emphasis>
</td>
</tr>
<tr id="198C4219FF9500650461AF54FAA8FD23" box="[290,1323,631,657]" gridrow="10" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461AF54FD05FD23" box="[290,646,631,657]" gridcol="0" gridrow="10" pageId="5" pageNumber="6">Autozooecial aperture width, mm</th>
<td id="5A5D2B65FF9500650789AF54FD6BFD23" box="[714,744,631,657]" gridcol="1" gridrow="10" pageId="5" pageNumber="6">30</td>
<td id="5A5D2B65FF9500650670AF54FCE1FD23" box="[819,866,631,657]" gridcol="2" gridrow="10" pageId="5" pageNumber="6">0.28</td>
<td id="5A5D2B65FF95006506DDAF54FC59FD23" box="[926,986,631,657]" gridcol="3" gridrow="10" pageId="5" pageNumber="6">0.038</td>
<td id="5A5D2B65FF9500650153AF54FBC8FD23" box="[1040,1099,631,657]" gridcol="4" gridrow="10" pageId="5" pageNumber="6">13.89</td>
<td id="5A5D2B65FF95006501C4AF54FB35FD23" box="[1159,1206,631,657]" gridcol="5" gridrow="10" pageId="5" pageNumber="6">0.23</td>
<td id="5A5D2B65FF95006501B5AF54FAA8FD23" box="[1270,1323,631,657]" gridcol="6" gridrow="10" pageId="5" pageNumber="6">0.38</td>
</tr>
<tr id="198C4219FF9500650461AFBEFAA8FD04" box="[290,1323,669,694]" gridrow="11" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461AFBEFD05FD04" box="[290,646,669,694]" gridcol="0" gridrow="11" pageId="5" pageNumber="6">Autozooecial aperture spacing, mm</th>
<td id="5A5D2B65FF9500650789AFBEFD6BFD04" box="[714,744,669,694]" gridcol="1" gridrow="11" pageId="5" pageNumber="6">30</td>
<td id="5A5D2B65FF9500650670AFBEFCE1FD04" box="[819,866,669,694]" gridcol="2" gridrow="11" pageId="5" pageNumber="6">0.66</td>
<td id="5A5D2B65FF95006506DDAFBEFC59FD04" box="[926,986,669,694]" gridcol="3" gridrow="11" pageId="5" pageNumber="6">0.073</td>
<td id="5A5D2B65FF9500650153AFBEFBC8FD04" box="[1040,1099,669,694]" gridcol="4" gridrow="11" pageId="5" pageNumber="6">11.04</td>
<td id="5A5D2B65FF95006501C4AFBEFB35FD04" box="[1159,1206,669,694]" gridcol="5" gridrow="11" pageId="5" pageNumber="6">0.55</td>
<td id="5A5D2B65FF95006501B5AFBEFAA8FD04" box="[1270,1323,669,694]" gridcol="6" gridrow="11" pageId="5" pageNumber="6">0.80</td>
</tr>
<tr id="198C4219FF9500650461AFE1FAA8FD69" box="[290,1323,706,731]" gridrow="12" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461AFE1FD05FD69" box="[290,646,706,731]" gridcol="0" gridrow="12" pageId="5" pageNumber="6">Vesicle diameter, mm</th>
<td id="5A5D2B65FF9500650789AFE1FD6BFD69" box="[714,744,706,731]" gridcol="1" gridrow="12" pageId="5" pageNumber="6">30</td>
<td id="5A5D2B65FF9500650670AFE1FCE1FD69" box="[819,866,706,731]" gridcol="2" gridrow="12" pageId="5" pageNumber="6">0.11</td>
<td id="5A5D2B65FF95006506DDAFE1FC59FD69" box="[926,986,706,731]" gridcol="3" gridrow="12" pageId="5" pageNumber="6">0.030</td>
<td id="5A5D2B65FF9500650153AFE1FBC8FD69" box="[1040,1099,706,731]" gridcol="4" gridrow="12" pageId="5" pageNumber="6">28.08</td>
<td id="5A5D2B65FF95006501C4AFE1FB35FD69" box="[1159,1206,706,731]" gridcol="5" gridrow="12" pageId="5" pageNumber="6">0.05</td>
<td id="5A5D2B65FF95006501B5AFE1FAA8FD69" box="[1270,1323,706,731]" gridcol="6" gridrow="12" pageId="5" pageNumber="6">0.19</td>
</tr>
<tr id="198C4219FF9500650461AFC4FAA8FCB3" box="[290,1323,743,769]" gridrow="13" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461AFC4FD05FCB3" box="[290,646,743,769]" gridcol="0" gridrow="13" pageId="5" pageNumber="6">Vesicle spacing, mm</th>
<td id="5A5D2B65FF9500650789AFC4FD6BFCB3" box="[714,744,743,769]" gridcol="1" gridrow="13" pageId="5" pageNumber="6">25</td>
<td id="5A5D2B65FF9500650670AFC4FCE1FCB3" box="[819,866,743,769]" gridcol="2" gridrow="13" pageId="5" pageNumber="6">0.10</td>
<td id="5A5D2B65FF95006506DDAFC4FC59FCB3" box="[926,986,743,769]" gridcol="3" gridrow="13" pageId="5" pageNumber="6">0.022</td>
<td id="5A5D2B65FF9500650153AFC4FBC8FCB3" box="[1040,1099,743,769]" gridcol="4" gridrow="13" pageId="5" pageNumber="6">21.78</td>
<td id="5A5D2B65FF95006501C4AFC4FB35FCB3" box="[1159,1206,743,769]" gridcol="5" gridrow="13" pageId="5" pageNumber="6">0.05</td>
<td id="5A5D2B65FF95006501B5AFC4FAA8FCB3" box="[1270,1323,743,769]" gridcol="6" gridrow="13" pageId="5" pageNumber="6">0.13</td>
</tr>
<tr id="198C4219FF9500650461AE2EFAA8FC94" box="[290,1323,781,806]" gridrow="14" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461AE2EFD05FC94" box="[290,646,781,806]" gridcol="0" gridrow="14" pageId="5" pageNumber="6">Vesicles per aperture</th>
<td id="5A5D2B65FF9500650789AE2EFD6BFC94" box="[714,744,781,806]" gridcol="1" gridrow="14" pageId="5" pageNumber="6">30</td>
<td id="5A5D2B65FF9500650670AE2EFCE1FC94" box="[819,866,781,806]" gridcol="2" gridrow="14" pageId="5" pageNumber="6">12.0</td>
<td id="5A5D2B65FF95006506DDAE2EFC59FC94" box="[926,986,781,806]" gridcol="3" gridrow="14" pageId="5" pageNumber="6">1.564</td>
<td id="5A5D2B65FF9500650153AE2EFBC8FC94" box="[1040,1099,781,806]" gridcol="4" gridrow="14" pageId="5" pageNumber="6">13.07</td>
<td id="5A5D2B65FF95006501C4AE2EFB35FC94" box="[1159,1206,781,806]" gridcol="5" gridrow="14" pageId="5" pageNumber="6">9.0</td>
<td id="5A5D2B65FF95006501B5AE2EFAA8FC94" box="[1270,1323,781,806]" gridcol="6" gridrow="14" pageId="5" pageNumber="6">16.0</td>
</tr>
<tr id="198C4219FF9500650461AE11FAA8FCF9" box="[290,1323,818,843]" gridrow="15" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461AE11FD05FCF9" box="[290,646,818,843]" gridcol="0" gridrow="15" pageId="5" pageNumber="6">Lunarium length, mm</th>
<td id="5A5D2B65FF9500650789AE11FD6BFCF9" box="[714,744,818,843]" gridcol="1" gridrow="15" pageId="5" pageNumber="6">15</td>
<td id="5A5D2B65FF9500650670AE11FCE1FCF9" box="[819,866,818,843]" gridcol="2" gridrow="15" pageId="5" pageNumber="6">0.14</td>
<td id="5A5D2B65FF95006506DDAE11FC59FCF9" box="[926,986,818,843]" gridcol="3" gridrow="15" pageId="5" pageNumber="6">0.037</td>
<td id="5A5D2B65FF9500650153AE11FBC8FCF9" box="[1040,1099,818,843]" gridcol="4" gridrow="15" pageId="5" pageNumber="6">26.91</td>
<td id="5A5D2B65FF95006501C4AE11FB35FCF9" box="[1159,1206,818,843]" gridcol="5" gridrow="15" pageId="5" pageNumber="6">0.08</td>
<td id="5A5D2B65FF95006501B5AE11FAA8FCF9" box="[1270,1323,818,843]" gridcol="6" gridrow="15" pageId="5" pageNumber="6">0.23</td>
</tr>
<tr id="198C4219FF9500650461AE74FAA8FCC3" box="[290,1323,855,881]" gridrow="16" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461AE74FD05FCC3" box="[290,646,855,881]" gridcol="0" gridrow="16" pageId="5" pageNumber="6">Lunarium width, mm</th>
<td id="5A5D2B65FF9500650789AE74FD6BFCC3" box="[714,744,855,881]" gridcol="1" gridrow="16" pageId="5" pageNumber="6">15</td>
<td id="5A5D2B65FF9500650670AE74FCE1FCC3" box="[819,866,855,881]" gridcol="2" gridrow="16" pageId="5" pageNumber="6">0.21</td>
<td id="5A5D2B65FF95006506DDAE74FC59FCC3" box="[926,986,855,881]" gridcol="3" gridrow="16" pageId="5" pageNumber="6">0.030</td>
<td id="5A5D2B65FF9500650153AE74FBC8FCC3" box="[1040,1099,855,881]" gridcol="4" gridrow="16" pageId="5" pageNumber="6">14.18</td>
<td id="5A5D2B65FF95006501C4AE74FB35FCC3" box="[1159,1206,855,881]" gridcol="5" gridrow="16" pageId="5" pageNumber="6">0.17</td>
<td id="5A5D2B65FF95006501B5AE74FAA8FCC3" box="[1270,1323,855,881]" gridcol="6" gridrow="16" pageId="5" pageNumber="6">0.29</td>
</tr>
<tr id="198C4219FF9500650461AE5EFAA8FC24" box="[290,1323,893,918]" gridrow="17" pageId="5" pageNumber="6">
<th id="5A5D2B65FF9500650461AE5EFD05FC24" box="[290,646,893,918]" gridcol="0" gridrow="17" pageId="5" pageNumber="6">Lunarium thickness, mm</th>
<td id="5A5D2B65FF9500650789AE5EFD6BFC24" box="[714,744,893,918]" gridcol="1" gridrow="17" pageId="5" pageNumber="6">15</td>
<td id="5A5D2B65FF9500650670AE5EFCE1FC24" box="[819,866,893,918]" gridcol="2" gridrow="17" pageId="5" pageNumber="6">0.08</td>
<td id="5A5D2B65FF95006506DDAE5EFC59FC24" box="[926,986,893,918]" gridcol="3" gridrow="17" pageId="5" pageNumber="6">0.031</td>
<td id="5A5D2B65FF9500650153AE5EFBC8FC24" box="[1040,1099,893,918]" gridcol="4" gridrow="17" pageId="5" pageNumber="6">40.31</td>
<td id="5A5D2B65FF95006501C4AE5EFB35FC24" box="[1159,1206,893,918]" gridcol="5" gridrow="17" pageId="5" pageNumber="6">0.03</td>
<td id="5A5D2B65FF95006501B5AE5EFAA8FC24" box="[1270,1323,893,918]" gridcol="6" gridrow="17" pageId="5" pageNumber="6">0.13</td>
</tr>
</table>
</paragraph>
<paragraph id="A703405BFF95FF9F0583AEC1FEB5FBC9" blockId="5.[192,793,994,1467]" pageId="5" pageNumber="6">
12 rows, 1115 surrounding each autozooecia aperture, with rounded to flat roofs, polygonal in tangential section. Autozooecial walls granular prismatic,
<quantity id="6044EDBEFF95FF9F0406A961FD9BFBE9" box="[325,536,1090,1115]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="5" pageNumber="6" unit="mm" value="0.01" valueMax="0.015" valueMin="0.005">
<geoCoordinate id="C288269CFF95FF9F0406A961FE0AFBE9" box="[325,393,1090,1115]" degrees="0.005" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="55" value="0.005">0.005</geoCoordinate>
<geoCoordinate id="C288269CFF95FF9F04CAA961FE5AFBE9" box="[393,473,1090,1115]" degrees="0.015" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="55" value="-0.015">0.015</geoCoordinate>
mm
</quantity>
thick. Maculae not observed.
</paragraph>
</subSubSection>
<subSubSection id="EFA613D0FF95FF9F0583A9A1FE2BFA09" pageId="5" pageNumber="6" type="discussion">
<paragraph id="A703405BFF95FF9F0583A9A1FDDFFB49" blockId="5.[192,793,994,1467]" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F0583A9A1FEBBFB29" bold="true" box="[192,312,1154,1179]" pageId="5" pageNumber="6">Remarks.</emphasis>
<taxonomicName id="60BC3BD8FF95FF9F047CA9A1FD6FFB29" authority="Gorjunova, 1970" authorityName="Gorjunova" authorityYear="1970" box="[319,748,1154,1180]" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="5" pageNumber="6" phylum="Bryozoa" rank="species" species="enodata">
<emphasis id="95C89C49FF95FF9F047CA9A1FDA2FB29" box="[319,545,1154,1179]" italics="true" pageId="5" pageNumber="6">Fistulipora enodata</emphasis>
<bibRefCitation id="C32D3DAAFF95FF9F076BA9A1FD6FFB29" author="Gorjunova, R. V." box="[552,748,1154,1180]" pageId="5" pageNumber="6" pagination="60 - 68" refId="ref27324" refString="Gorjunova, R. V. 1970. New Early Permian bryozoans from the Southeast Darvaz and Pamir, p. 60 - 68. In Astrova, G. G. and Chudinova, I. I. (eds.), New Species of Palaeozoic Bryozoans and Corals. Nauka, Moscow. (In Russian)" type="book chapter" year="1970">Gorjunova, 1970</bibRefCitation>
</taxonomicName>
differs from
<taxonomicName id="60BC3BD8FF95FF9F0473A981FDEEFB09" authority="Bassler, 1929" authorityName="Bassler" authorityYear="1929" box="[304,621,1186,1211]" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="5" pageNumber="6" phylum="Bryozoa" rank="species" species="timorensis">
<emphasis id="95C89C49FF95FF9F0473A981FE47FB09" box="[304,452,1186,1211]" italics="true" pageId="5" pageNumber="6">F. timorensis</emphasis>
<bibRefCitation id="C32D3DAAFF95FF9F048FA981FDEEFB09" author="Bassler, R. S." box="[460,621,1186,1211]" pageId="5" pageNumber="6" pagination="37 - 90" refId="ref26181" refString="Bassler, R. S. 1929. The Permian Bryozoa of Timor. Palaontologie von Timor, 16: 37 - 90." type="journal article" year="1929">
Bassler,
<quantity id="6044EDBEFF95FF9F0771A981FDEEFB09" box="[562,621,1186,1211]" metricMagnitude="1" metricUnit="m" metricValue="4.89966" pageId="5" pageNumber="6" unit="in" value="1929.0">1929</quantity>
</bibRefCitation>
</taxonomicName>
in larger autozooecial apertures (aperture width
<quantity id="6044EDBEFF95FF9F0729A9E1FC9BFB69" box="[618,792,1218,1243]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" metricValueMax="3.8" metricValueMin="2.4" pageId="5" pageNumber="6" unit="mm" value="0.31" valueMax="0.38" valueMin="0.24">0.240.38 mm</quantity>
vs.
<quantity id="6044EDBEFF95FF9F05AAA9C1FE10FB49" box="[233,403,1250,1275]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="5" pageNumber="6" unit="mm" value="0.25" valueMax="0.28" valueMin="0.22">0.220.28 mm</quantity>
in
<taxonomicName id="60BC3BD8FF95FF9F04F4A9C1FDCFFB49" authorityName="Bassler" authorityYear="1929" box="[439,588,1250,1275]" class="Stenolaemata" family="Fistuliporidae" genus="Fistulipora" kingdom="Animalia" order="Cystoporida" pageId="5" pageNumber="6" phylum="Bryozoa" rank="species" species="timorensis">
<emphasis id="95C89C49FF95FF9F04F4A9C1FDCFFB49" box="[439,588,1250,1275]" italics="true" pageId="5" pageNumber="6">F. timorensis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="A703405BFF95FF9F0583A821FE2BFA09" blockId="5.[192,793,994,1467]" pageId="5" pageNumber="6">
<emphasis id="95C89C49FF95FF9F0583A821FED8FAAE" bold="true" box="[192,347,1282,1308]" pageId="5" pageNumber="6">Occurrence.</emphasis>
Lower Permian, Artinskian;
<collectingCountry id="DFAB00CBFF95FF9F07E8A821FC9BFAA9" box="[683,792,1282,1307]" name="Tajikistan" pageId="5" pageNumber="6">Tajikistan</collectingCountry>
(Pamir). Lower Permian; Rutog, southwestern Tibet. Xiala Formation,?Middle Permian; Xainza, southwestern Tibet. Zhongba Formation, Permian (late CisuralianGuadalupian); Zhongba area of southwestern Tibet.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,465 @@
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<mods:title id="2E8F82371292AFEB829A711AB864C5F0">Four new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae, Liphistiinae) from Myanmar</mods:title>
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<mods:namePart id="74B1F3056EF13C35F4C499F81BF2475D">Zhan, Yi</mods:namePart>
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</mods:role>
<mods:namePart id="DCDDA27E7DDA46A6FF14935C66625140">Yu, Long</mods:namePart>
<mods:nameIdentifier id="FD6301E26664B6FBDA36981B14A4E587" type="ORCID">0000-0001-5675-1864</mods:nameIdentifier>
<mods:affiliation id="7B3A75734E88E1A2A7C9FFCCC00E12F0">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="09D7BE9B641A3819F2D97D977B84DD38">Li, Daiqin</mods:namePart>
<mods:nameIdentifier id="06A1A3C1DCD7DDCC7CD0D204632751F7" type="ORCID">0000-0001-8269-7734</mods:nameIdentifier>
<mods:affiliation id="D358AC90ED783EF5A33AC18B964734B4">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China &amp; Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, Singapore 117543</mods:affiliation>
</mods:name>
<mods:typeOfResource id="F3159DA38D9885B76CF4B4073972F71B">text</mods:typeOfResource>
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<mods:part id="76FD6C130C03BBF842F5B328F231696F">
<mods:date id="74EDC2068AE832D89460FCECC9F4760D">2024</mods:date>
<mods:detail id="0149568B476132D361FE5613F15D62ED" type="pubDate">
<mods:number id="DDB1C231311B4DD6F58DBD29807262AB">2024-08-16</mods:number>
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<mods:number id="07FF866DF1434D912E2ECAC8AE1A7CAB">1210</mods:number>
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<mods:classification id="BDEA820559F0D8DAA60FBFC64E328172">journal article</mods:classification>
<mods:identifier id="D95D9F787DFCAB0CD74E287F2FB40B02" type="DOI">10.3897/zookeys.1210.123986</mods:identifier>
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<treatment id="A5B909E355755A208FD79858B83ECEA8" ID-arpha="A5B909E3-5575-5A20-8FD7-9858B83ECEA8" ID-zoobank="https://zoobank.org/8982C984-06DB-4636-9C22-92E2C69A62CB" LSID="urn:lsid:plazi:treatment:A5B909E355755A208FD79858B83ECEA8" LSID-ZBK="urn:lsid:zoobank.org:act:8982C984-06DB-4636-9C22-92E2C69A62CB" httpUri="http://treatment.plazi.org/id/A5B909E355755A208FD79858B83ECEA8">
<subSubSection id="8C2504C15D0E5BE0BC65D0FD9A165F5F" type="nomenclature">
<paragraph id="0B05D507883012FB9CAFC33F60112DB9">
<taxonomicName id="2415FCD747AA9E74D1E4F102285390FE" authority="Zhan &amp; Xu" authorityName="Zhan &amp; Xu" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kanpetlet" status="sp. nov.">
<emphasis id="58B62507CB93046124DB3EFD109C3A4D" italics="true">Liphistius kanpetlet</emphasis>
Zhan &amp; Xu
</taxonomicName>
<taxonomicNameLabel id="11EFF3D6B633B3E0250AEC443345A790" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="F093B20C8C30B5D11B7745674F16B08F" type="description">
<paragraph id="DEC67AE839CDD879430CDBB247F6E68D">
<figureCitation id="AD2898434AF1F831CA0F9597D083A038" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Liphistius kanpetlet sp. nov. A, E prolateral view B ventral view; C, F retrolateral view, red arrow shows distal process of PC D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure5" httpUri="https://binary.pensoft.net/fig/1115131">Figs 5</figureCitation>
,
<figureCitation id="AF618D723389B1CA2CB8E630D26F302C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Vulva of Liphistius kanpetlet sp. nov. A C dorsal view D F ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure6" httpUri="https://binary.pensoft.net/fig/1115132">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIEAG" type="type material">
<paragraph id="2C24314CCC6BDF6675474C6FD623D569">
<heading id="C2EBB46F7B8C1BF42E9F7C919DD5B05B" reason="title">Type material.</heading>
</paragraph>
<paragraph id="2292FA9CDBBC785CA7B0004860B5EA58">
<materialsCitation id="ACE9C03F8D65E35792CDA87AFC2C95EC" collectingDate="2019-07-17" collectorName="D. Li" country="Myanmar" elevation="1469" latitude="21.19" location="Kanpetlet Matupi Rd" longLatPrecision="758" longitude="94.05" municipality="Kanpetlet Township" specimenCount="1" specimenCount-male="1" stateProvince="Chin" typeStatus="holotype">
<emphasis id="29EECEDC618D56275B845BDDC3546CDD" bold="true" italics="true">
<typeStatus id="257E87C718168E4B3ECD52B3E55F9C75" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="CCFF3E816159A4C08400D77223086B36" name="Myanmar">Myanmar</collectingCountry>
<specimenCount id="0D6D5E5F4805C712EAF89EA6449FA181" count="1" type="male"></specimenCount>
,
<collectingRegion id="25BFBC75B084B104261F1A62B3201A5F" country="Myanmar" name="Chin">Chin State</collectingRegion>
,
<collectingMunicipality id="751270E397E8DCE0483F5AB9C2CE1202">Kanpetlet Township</collectingMunicipality>
,
<location id="D7E4757585A6E4064114A1DBF60B3702" LSID="urn:lsid:plazi:treatment:A5B909E355755A208FD79858B83ECEA8:D7E4757585A6E4064114A1DBF60B3702" country="Myanmar" latitude="21.19" longLatPrecision="758" longitude="94.05" municipality="Kanpetlet Township" name="Kanpetlet Matupi Rd" stateProvince="Chin">Kanpetlet Matupi Rd</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWEAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[94.050000,21.190000]}">
<geoCoordinate id="6E9F16F4F060E55DF4B7C482D27CE806" degrees="21.19" direction="north" orientation="latitude" precision="555" value="21.19">21.19 ° N</geoCoordinate>
,
<geoCoordinate id="1C58B0FADECE53137D9C6005041DB050" degrees="94.05" direction="east" orientation="longitude" precision="555" value="94.05">94.05 ° E</geoCoordinate>
</named-content>
, alt.
<quantity id="E7735ED85FBB46CE8BE6A102C0752AEC" metricMagnitude="3" metricUnit="m" metricValue="1.469" unit="m" value="1469.0">
<elevation id="04268C023A90010FF8FFB36755744255" metricMagnitude="3" metricUnit="m" metricValue="1.469" unit="m" value="1469.0">1469 m</elevation>
</quantity>
;
<date id="A1B6854C6E7FB3442DB1FB75D5E52F93" value="2019-07-17">
<collectingDate id="672A02F50D968238A7F155CB136821CE" value="2019-07-17">17. 07. 2019</collectingDate>
</date>
;
<collectorName id="E0346B079194789D2FADF0E1CD982932">D. Li</collectorName>
et al. leg.; XUX- 2019-026
</materialsCitation>
.
<emphasis id="8E6E5DBC04CDECADD036971EC350EB5C" bold="true" italics="true">
<typeStatus id="017E1E13C54FCC0BEA30F0A7D669F07A" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="E2DDFA95CF0CB5D018EC161D11A3B46C" count="5" type="female">5 ♀♀</specimenCount>
, same locality as for
<typeStatus id="3BD1861D7E6A5E35A7AC998A3EF1707B" type="holotype">holotype</typeStatus>
;
<date id="931ECB5017594BDFC2027D1894664A88" value="2019-07-17" valueMax="2019-07-18" valueMin="2019-07-17">1718.07. 2019</date>
; XUX- 2019-020 / 021 / 024 / 031 / 033.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5EAG" type="diagnosis">
<paragraph id="7416784097A544D295A557D56C19DDC6">
<heading id="E5EF24C24B796621947A44C8F57DC5FE" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="A1F6CFF054F37DF540D317989D1FED07">
The male of
<taxonomicName id="0B515ECE2B6B41F7D17BEFE3BDB55540" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kanpetlet" status="sp. nov.">
<emphasis id="E295122E33A7893D295B5499089B709F" italics="true">L. kanpetlet</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6D92AEDE167168C510886A29E2E9A164" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from those of
<taxonomicName id="F316F7CC100A94E389C0860819E52324" authorityName="Thorell" authorityYear="1897" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="birmanicus">
<emphasis id="7495CFF8D49587F4BFD57625B993A93D" italics="true">L. birmanicus</emphasis>
</taxonomicName>
by having the paracymbium (
<abbrev id="ABBRID0E1FAG" xlink_title="paracymbium">PC</abbrev>
) bearing a distal process (Fig.
<figureCitation id="31964DDD8312FB433E949AF9BDFD9509" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Liphistius kanpetlet sp. nov. A, E prolateral view B ventral view; C, F retrolateral view, red arrow shows distal process of PC D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure5" httpUri="https://binary.pensoft.net/fig/1115131">5 C</figureCitation>
vs lacking a distal process, fig. 13 H in
<bibRefCitation id="39499AB4A7AD2F2724650FB1E4182CAD" DOI="10.35929/RSZ.0083" author="Schwendinger" etAl="et al." firstAuthor="Schwendinger" issue="2" journalOrPublisher="Revue Suisse de Zoologie" pagination="375-424" refId="B5" refString="Schwendinger PJ, Huber S, Lehmann-Graber C, Ono H, Aung MM, Hongpadharakiree K (2022) A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species. Revue Suisse de Zoologie 129 (2): 375424. https://doi.org/10.35929/RSZ.0083" title="A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species." volume="129" year="2022">Schwendinger et al. 2022</bibRefCitation>
), and from
<taxonomicName id="121D65E44420C28274CB389B0E864415" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kalaw" status="sp. nov.">
<emphasis id="97C51A2F665E9E36B0AF18AEB6D198A3" italics="true">L. kalaw</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDEF5FBA510C4B32F66B5E49EC26DC51" rank="species">sp. nov.</taxonomicNameLabel>
by the contrategulum (
<abbrev id="ABBRID0ERGAG" xlink_title="contrategulum">CT</abbrev>
) which has a blunt process (Fig.
<figureCitation id="6FFFEFDF87A925187E72996CC84A9900" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Liphistius kanpetlet sp. nov. A, E prolateral view B ventral view; C, F retrolateral view, red arrow shows distal process of PC D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure5" httpUri="https://binary.pensoft.net/fig/1115131">5 D</figureCitation>
vs slightly sharp process, Fig.
<figureCitation id="D01D5A09FE2139A3DAF77550B7A08C26" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 E</figureCitation>
). The female of
<taxonomicName id="DFC01E95FCECC84375006F0536DECED5" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kanpetlet" status="sp. nov.">
<emphasis id="41BCDBCDE2CAFACCC2A0CA87E7942318" italics="true">L. kanpetlet</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3531FDCB6A7BD251888A5C1D77A5B73A" rank="species">sp. nov.</taxonomicNameLabel>
differs from that of
<taxonomicName id="D275B7A0BBA7F7C99AE87C0F964290FC" authorityName="Aung, Xu, Lwin, Sang, Yu, Liu, Liu &amp; Li" authorityYear="2019" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="hpruso">
<emphasis id="FB3A2844636FCB65292D3384A14BFD92" italics="true">L. hpruso</emphasis>
</taxonomicName>
by having two relatively larger anterior lobes (Fig.
<figureCitation id="35F11033F5CAE1ED5F47DA65C07C30E2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Vulva of Liphistius kanpetlet sp. nov. A C dorsal view D F ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure6" httpUri="https://binary.pensoft.net/fig/1115132">6</figureCitation>
vs fig. 3 B E in
<bibRefCitation id="9CB56E0A49FBBBC4D60133047FC515C2" DOI="10.3897/zookeys.882.38811" author="Aung" etAl="et al." firstAuthor="Aung" journalOrPublisher="ZooKeys" pagination="29-39" refId="B1" refString="Aung KPP, Xu X, Lwin WW, Sang MZ, Yu L, Liu H, Liu FX, Li DQ (2019) Two new species of the primitively segmented spider genus Liphistius Schiödte, 1849 (Mesothelae, Liphistiidae) from Myanmar. ZooKeys 882: 2939. https://doi.org/10.3897/zookeys.882.38811" title="Two new species of the primitively segmented spider genus Liphistius Schiödte, 1849 (Mesothelae, Liphistiidae) from Myanmar." volume="882" year="2019">Aung et al. 2019</bibRefCitation>
).
</paragraph>
<caption id="9157607886C444CF8ADEC6BF7785CFB5" ID-DOI="10.3897/zookeys.1210.123986.figure5" ID-arpha="DB0BCA64-6973-58C4-B5C2-9229E32A9D76" httpUri="https://binary.pensoft.net/fig/1115131" startId="F5">
<paragraph id="A7B4EB0530E25F158B866124D2D7BF56">
<label id="E3B15A2334AAB343D862B219DC91D63C">Figure 5.</label>
</paragraph>
<paragraph id="26BEF0B4C4F6A5DD3C0BA105C26C061D">
Male palp of
<taxonomicName id="C5D40C7C87B56C3DD0F84925E326DCE3" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kanpetlet" status="sp. nov.">
<emphasis id="14D2DAE887F1C4C4A075328A70A47B93" italics="true">Liphistius kanpetlet</emphasis>
</taxonomicName>
<taxonomicNameLabel id="03C8DD383C638174ACF5FCEA1036F4B7" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="2938BC4505F10113B97199A34B424CE5" bold="true">A, E</emphasis>
prolateral view
<emphasis id="FCE4E305F5B69343B19AB1023D93F44C" bold="true">B</emphasis>
ventral view;
<emphasis id="093F0CD96326B51B096A12D6DE8D383E" bold="true">C, F</emphasis>
retrolateral view, red arrow shows distal process of
<abbrev id="ABBRID0EUIAG" xlink_title="paracymbium">PC</abbrev>
<emphasis id="3FEC9AB80D493E3086E96DE9BFE97F17" bold="true">D</emphasis>
dorsal view, red arrows show arched projection and triangular process of
<abbrev id="ABBRID0E1IAG" xlink_title="contrategulum">CT</abbrev>
<emphasis id="C21AA54D39C8DB1ED1E3A2F8D0F00C10" bold="true">G</emphasis>
distal view. Abbreviations:
<abbrev id="ABBRID0EAJAG" xlink_title="contrategulum">CT</abbrev>
= contrategulum;
<abbrev id="ABBRID0EEJAG" xlink_title="contrategular process">cp</abbrev>
= contrategular process;
<abbrev id="ABBRID0EIJAG" xlink_title="cumulus">Cu</abbrev>
= cumulus; E = embolus;
<abbrev id="ABBRID0EMJAG" xlink_title="paracymbium">PC</abbrev>
= paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum;
<abbrev id="ABBRID0EQJAG" xlink_title="male palps bearing tibial apophysis">TiA</abbrev>
= tibial apophysis. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="5C83FF0B1DAEC166FA09741542C21B30" ID-DOI="10.3897/zookeys.1210.123986.figure6" ID-arpha="610A59C0-AAB6-5786-A074-A23EE9C7E9D1" httpUri="https://binary.pensoft.net/fig/1115132" startId="F6">
<paragraph id="DA2E6CF65C163774A12850608C3822E8">
<label id="36B241DDC2EE287C848FF8A4E1095CEC">Figure 6.</label>
</paragraph>
<paragraph id="649D6944130AF23D5184130778CDC1A5">
Vulva of
<taxonomicName id="93A572A3D0C33AD078D5C28D5563EB1A" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kanpetlet" status="sp. nov.">
<emphasis id="F7E444C964450AA1F179E26800B4043E" italics="true">Liphistius kanpetlet</emphasis>
</taxonomicName>
<taxonomicNameLabel id="286D6E3F88575EDD3559C960CF3C739C" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="82CF0EBA4F144043C9964085319EF98D" bold="true">A C</emphasis>
dorsal view
<emphasis id="177014E0D3BEF90BF8BF7DFFC6C0CF18" bold="true">D F</emphasis>
ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk;
<abbrev id="ABBRID0EQKAG" xlink_title="sclerotized poreplate">PPl</abbrev>
= poreplate;
<abbrev id="ABBRID0EUKAG" xlink_title="receptacular cluster">RC</abbrev>
= receptacular cluster. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E4KAG" type="description">
<paragraph id="62F9EA58A96FAD5DBF2DFF03FC608BFB">
<heading id="FA1F02E36FB39A3AAA09EC269B6173BB" reason="title">Description.</heading>
</paragraph>
<paragraph id="62969D8C8F1C1C5D96AF5ED54E3B72CE">
<emphasis id="A26E8E6D63331555DE2135AF30713DE3" bold="true">
Male
<typeStatus id="790423CA56E53315D730DED447B7EDF1" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace yellowish brown with few short, scattered bristles; opisthosoma brown, with 12 brown tergites, close to each other, 26
<superScript id="E91D0C2083BB858187D5BFF31F49CD61">th</superScript>
larger than others, 4
<superScript id="55C5E5F21C973D05915DED53F96D8E43">th</superScript>
largest; chelicerae robust, promargin of cheliceral groove with 13 denticles of variable size; labium yellow, sternum yellow, with few short setae on anterior tip and many long setae on elongated posterior tip; legs reddish brown, with strong setae and spines; 8 spinnerets. Measurements:
<abbrev id="ABBRID0EJLAG" xlink_title="body length">BL</abbrev>
12.41,
<abbrev id="ABBRID0ENLAG" xlink_title="carapace length">CL</abbrev>
6.01,
<abbrev id="ABBRID0ERLAG" xlink_title="carapace width">CW</abbrev>
6.06,
<abbrev id="ABBRID0EVLAG" xlink_title="opisthosoma length">OL</abbrev>
6.30,
<abbrev id="ABBRID0EZLAG" xlink_title="opisthosoma width">OW</abbrev>
4.19; eye sizes and interdistances:
<abbrev id="ABBRID0E4LAG" xlink_title="anterior median eyes">AME</abbrev>
0.06,
<abbrev id="ABBRID0EBMAG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.56,
<abbrev id="ABBRID0EFMAG" xlink_title="posterior median eyes">PME</abbrev>
0.32,
<abbrev id="ABBRID0EJMAG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.42,
<abbrev id="ABBRID0ENMAG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0ERMAG" xlink_title="anterior median eyes">AME</abbrev>
0.06,
<abbrev id="ABBRID0EVMAG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EZMAG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.16,
<abbrev id="ABBRID0E4MAG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0EBNAG" xlink_title="posterior median eyes">PME</abbrev>
0.10,
<abbrev id="ABBRID0EFNAG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0EJNAG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.11,
<abbrev id="ABBRID0ENNAG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0ERNAG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.14,
<abbrev id="ABBRID0EVNAG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EZNAG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.14,
<abbrev id="ABBRID0E4NAG" xlink_title="posterior lateral eyes">PLE</abbrev>
<abbrev id="ABBRID0EBOAG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.34,
<abbrev id="ABBRID0EFOAG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EJOAG" xlink_title="posterior median eyes">PME</abbrev>
0.08; labium 0.96 long and 0.50 wide; sternum 2.63 long and 1.00 wide; legs: I 17.51 (4.82, 2.52, 3.79, 4.14, 2.24), II 18.36 (5.20, 2.19, 3.95, 4.72, 2.3), III 20.63 (5.46, 2.20, 4.41, 5.86, 2.70), IV 26.62 (7.18, 2.64, 5.40, 7.71, 3.69).
</paragraph>
<paragraph id="0A78FBEB549E2AC28420C4A78DEBE941">
Palp: tibial apophysis (
<abbrev id="ABBRID0EPOAG" xlink_title="male palps bearing tibial apophysis">TiA</abbrev>
) with 5 setae of different lengths (Fig.
<figureCitation id="D00106DBA9D918C19E39BA7AA846FEF5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Liphistius kanpetlet sp. nov. A, E prolateral view B ventral view; C, F retrolateral view, red arrow shows distal process of PC D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure5" httpUri="https://binary.pensoft.net/fig/1115131">5 A C</figureCitation>
); paracymbium (
<abbrev id="ABBRID0EXOAG" xlink_title="paracymbium">PC</abbrev>
) with some setae situated on tip, and with pointed lateral process (Fig.
<figureCitation id="625E46180D796CFD7F452D6BF2FC1FD4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Liphistius kanpetlet sp. nov. A, E prolateral view B ventral view; C, F retrolateral view, red arrow shows distal process of PC D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure5" httpUri="https://binary.pensoft.net/fig/1115131">5 C</figureCitation>
); and several tapering spines on slightly elevated cumulus (
<abbrev id="ABBRID0E6OAG" xlink_title="cumulus">Cu</abbrev>
) (Fig.
<figureCitation id="FD4EDCDD3F252BDD683222DB12B2E84A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Liphistius kanpetlet sp. nov. A, E prolateral view B ventral view; C, F retrolateral view, red arrow shows distal process of PC D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure5" httpUri="https://binary.pensoft.net/fig/1115131">5 B</figureCitation>
); subtegular apophysis weakly developed (Fig.
<figureCitation id="1CD9866FA4AEA5F48CB253ADB9E4FE90" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Liphistius kanpetlet sp. nov. A, E prolateral view B ventral view; C, F retrolateral view, red arrow shows distal process of PC D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure5" httpUri="https://binary.pensoft.net/fig/1115131">5 B, F</figureCitation>
); contrategulum (
<abbrev id="ABBRID0ELPAG" xlink_title="contrategulum">CT</abbrev>
) with arched projection distally, and wide triangular process in prolateral view (Fig.
<figureCitation id="7B744B3EC703220770C29298A2FD46BD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Liphistius kanpetlet sp. nov. A, E prolateral view B ventral view; C, F retrolateral view, red arrow shows distal process of PC D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure5" httpUri="https://binary.pensoft.net/fig/1115131">5 D</figureCitation>
); fan-shaped tegulum (
<collectionCode id="86803715476F6EEEF9EC582CFE85779A" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
) with 6 longitudinal ridges (Fig.
<figureCitation id="11F8E1D3064ACB9591BB177001FE7C48" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Liphistius kanpetlet sp. nov. A, E prolateral view B ventral view; C, F retrolateral view, red arrow shows distal process of PC D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure5" httpUri="https://binary.pensoft.net/fig/1115131">5 C</figureCitation>
); paraembolic plate (PeP) widely rounded, short, with curved margin in ventral view (Fig.
<figureCitation id="08790CC9D49FC80B8AA3791A31156B83" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Liphistius kanpetlet sp. nov. A, E prolateral view B ventral view; C, F retrolateral view, red arrow shows distal process of PC D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure5" httpUri="https://binary.pensoft.net/fig/1115131">5 A, F</figureCitation>
); embolus (E) sclerotized basally, with several denticles along longitudinal ridges reaching tip (Fig.
<figureCitation id="C5BBB125CA29EA998254B2F5BCD5051F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Liphistius kanpetlet sp. nov. A, E prolateral view B ventral view; C, F retrolateral view, red arrow shows distal process of PC D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure5" httpUri="https://binary.pensoft.net/fig/1115131">5 C G</figureCitation>
).
</paragraph>
<paragraph id="BBE80F60AF8436A33C26F0483FD554AF">
<emphasis id="BC1116C812F24D6CA53880DCC30773D9" bold="true">
Female
<typeStatus id="C31E262A02D0A839BA551C503DCF2CF8" type="paratype">paratype</typeStatus>
(XUX- 2019-024).
</emphasis>
Carapace brown with few short, scattered bristles; opisthosoma gray, with 12 brown tergites, close to each other, with gray patches, 26
<superScript id="546F971F8ACCCB9EA598299FFB608A3F">th</superScript>
larger than others, 5
<superScript id="31BBDE7AC7C6C27094EB116A2C6263A4">th</superScript>
largest; chelicerae robust, reddish brown; promargin of cheliceral groove with 10 denticles of variable size; labium yellow, sternum yellow with several setae; legs with strong setae and spines, without distinct annulations. Measurements:
<abbrev id="ABBRID0EHQAG" xlink_title="body length">BL</abbrev>
11.71,
<abbrev id="ABBRID0ELQAG" xlink_title="carapace length">CL</abbrev>
5.26,
<abbrev id="ABBRID0EPQAG" xlink_title="carapace width">CW</abbrev>
5.15,
<abbrev id="ABBRID0ETQAG" xlink_title="opisthosoma length">OL</abbrev>
6.36,
<abbrev id="ABBRID0EXQAG" xlink_title="opisthosoma width">OW</abbrev>
4.70; eye sizes and interdistances:
<abbrev id="ABBRID0E2QAG" xlink_title="anterior median eyes">AME</abbrev>
0.09,
<abbrev id="ABBRID0E6QAG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.50,
<abbrev id="ABBRID0EDRAG" xlink_title="posterior median eyes">PME</abbrev>
0.34,
<abbrev id="ABBRID0EHRAG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.40,
<abbrev id="ABBRID0ELRAG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EPRAG" xlink_title="anterior median eyes">AME</abbrev>
0.06,
<abbrev id="ABBRID0ETRAG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EXRAG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.19,
<abbrev id="ABBRID0E2RAG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0E6RAG" xlink_title="posterior median eyes">PME</abbrev>
0.07,
<abbrev id="ABBRID0EDSAG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0EHSAG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.10,
<abbrev id="ABBRID0ELSAG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EPSAG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.09,
<abbrev id="ABBRID0ETSAG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EXSAG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.14,
<abbrev id="ABBRID0E2SAG" xlink_title="posterior lateral eyes">PLE</abbrev>
<abbrev id="ABBRID0E6SAG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.35,
<abbrev id="ABBRID0EDTAG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EHTAG" xlink_title="posterior median eyes">PME</abbrev>
0.06; Labium 1.38 long and 0.59 wide; sternum 2.31 long and 1.19 wide; palp 8.05 (2.76, 1.36, 2.15, 1.78), leg I 10.24 (3.21, 1.60, 2.30, 2.01, 1.12), II 10.82 (3.43, 1.68, 2.27, 2.14, 1.30), III 11.14 (3.49, 1.51, 2.16, 2.51, 1.47), IV 15.54 (4.45, 1.94, 3.13, 3.71, 2.31).
</paragraph>
<paragraph id="BF8D823E26DA2AF3D581E1D15AE21456">
Vulva: approximately rectangular poreplate (
<abbrev id="ABBRID0ENTAG" xlink_title="sclerotized poreplate">PPl</abbrev>
) wider than long with smoothly curved posterior margin; with pair of large anterior lobes and pair of small anterolateral lobes, 2 anterior lobes separated from each other, but close to anterolateral lobes; central dorsal opening (
<collectionCode id="4EA96EF85FB56FAB005C111659F703F2">CDO</collectionCode>
) small, situated in center of poreplate (
<abbrev id="ABBRID0ERTAG" xlink_title="sclerotized poreplate">PPl</abbrev>
); racemose receptacular cluster (
<abbrev id="ABBRID0EVTAG" xlink_title="receptacular cluster">
<collectionCode id="D4611EFDD86DFA4E137BC076C80A3157">RC</collectionCode>
</abbrev>
) long and narrow; posterior stalk (
<collectionCode id="5FBDD2BF7A423D70BB5609E709DC50A9">PS</collectionCode>
) axe-shaped; genital atrium (GA) with slightly sclerotized lateral margins (Fig.
<figureCitation id="ACAC12FD8DE5F8CC744F34F832D9F198" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Vulva of Liphistius kanpetlet sp. nov. A C dorsal view D F ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure6" httpUri="https://binary.pensoft.net/fig/1115132">6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4TAG" type="variation">
<paragraph id="030BD4DE8844222FBAC564FD7E56F19B">
<heading id="A5B91399210A99C713393B5E5268ED69" reason="title">Variation.</heading>
</paragraph>
<paragraph id="4AC05AD8BFE343CFCCAB4F5FC0ED3F09">
Females vary in body size. The range of measurements of females (
<emphasis id="DE5EAC5F0D8FB59E745FE2D5FCEBD3C7" italics="true">N</emphasis>
= 5):
<abbrev id="ABBRID0EFUAG" xlink_title="body length">BL</abbrev>
9.7014.85,
<abbrev id="ABBRID0EJUAG" xlink_title="carapace length">CL</abbrev>
4.866.55,
<abbrev id="ABBRID0ENUAG" xlink_title="carapace width">CW</abbrev>
4.208.63,
<abbrev id="ABBRID0ERUAG" xlink_title="opisthosoma length">OL</abbrev>
3.828.63,
<abbrev id="ABBRID0EVUAG" xlink_title="opisthosoma width">OW</abbrev>
3.827.09. The number of denticles on the promargin of cheliceral groove varies from 1013 (
<emphasis id="94455FE6B02EA093065F602F785B6063" italics="true">N</emphasis>
= 5).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2UAG" type="etymology">
<paragraph id="574181A739BEE8E110A9339FDAAB9EC0">
<heading id="04C11C654CE0FCBBB046133FFBF2EB37" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="FF9EB9E2D67A4E2E602F820FE7FA5686">
The species epithet
<taxonomicName id="EE6F16981D32808F42C6914A3FA473E9" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kanpetlet">
<emphasis id="666AAD992A1A6D9FBE4F9203B020ACAF" italics="true">kanpetlet</emphasis>
</taxonomicName>
” refers to the
<typeStatus id="A01591096180335ADBC0604682512FB3">type</typeStatus>
locality, Kanpetlet Township; it is treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKVAG" type="distribution">
<paragraph id="18B77FD5B879D2C109EE242D6D8F0E9A">
<heading id="12DD3D3480BA1F41B73F616F5932FDA1" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="C1206B508660ACA7DB65F3E38AD607D5">
<collectingRegion id="17DE8A5B2E5BFF4816064279B01D20AF" country="Myanmar" name="Chin">Chin State</collectingRegion>
(Kanpetlet Township),
<collectingCountry id="085F8C15E70DB19B817003DEE4342FC1" name="Myanmar">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,472 @@
<document id="04BE13E67393E186F0BEA8020CB72AB2" ID-DOI="10.3897/zookeys.1210.123986" ID-ZooBank="6972216E-2356-4DF7-BD24-2B6B68E639B4" ID-publisher-id="123986" URI-arpha="B6184626-6862-59C4-9047-EF9B7AC24E35" URI-zoobank="http://zoobank.org/6972216E-2356-4DF7-BD24-2B6B68E639B4" article-type="research-article" checkinTime="1723830749198" checkinUser="pensoft" docAuthor="Xu, Xin, Zhan, Yi, Aung, Khin Pyae Pyae, Liu, Fengxiang, Yu, Long &amp; Li, Daiqin" docDate="2024" docId="BE9AF2F05BBA5A87B548EEDB4256870E" docLanguage="en" docName="ZooKeys 1210: 99-115" docOrigin="ZooKeys 1210" docSource="https://zookeys.pensoft.net/article/123986/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Liphistius rostratus Zhan &amp; Xu 2024, sp. nov." docType="treatment" docVersion="1" dtd-version="3.0" lastPageNumber="115" masterDocId="B6184626686259C49047EF9B7AC24E35" masterDocTitle="Four new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae, Liphistiinae) from Myanmar" masterLastPageNumber="115" masterPageNumber="99" pageNumber="99" updateTime="1723830805509" updateUser="pensoft">
<mods:mods id="26ED0726C24CC570EB2B0140AA5B3CEC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="14DAFEF81F25D7AE15EC85BDA31BFFDD">
<mods:title id="68CB80660B468754C6BC8C9C117DFBDA">Four new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae, Liphistiinae) from Myanmar</mods:title>
</mods:titleInfo>
<mods:name id="8A3185180DE7A71783CD181F1D7D3833" type="personal">
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<mods:namePart id="80EB64DE67ABFF6CE91AB6A3065AF4AE">Xu, Xin</mods:namePart>
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<mods:affiliation id="1E351DE5C427B29E7FDEB501ED5EE1AF">College of Life Sciences, Hunan Normal University, 36 Lushan Road, Changsha 410081, Hunan Province, China &amp; State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
</mods:name>
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<mods:namePart id="91C8940CBD05A1C1F9589715A5A71811">Zhan, Yi</mods:namePart>
<mods:affiliation id="7F28C9640A09FBDD6AACB1295A73E92E">College of Life Sciences, Hunan Normal University, 36 Lushan Road, Changsha 410081, Hunan Province, China</mods:affiliation>
</mods:name>
<mods:name id="88318A2CF614C93ED8A25289A17552B3" type="personal">
<mods:role id="89FC81F64E7EE298E2152E415B49D657">
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<mods:namePart id="83A2FBE51C538699F555EE8D67584443">Aung, Khin Pyae Pyae</mods:namePart>
<mods:affiliation id="00183F93FA2D24BC7C07E9A28BDA37D8">Department of Zoology, University of Yangon, Kamayut Township, Pyay Road, Yangon, 11041, Myanmar &amp; Department of Biology, Taungoo Education Degree College, Taungoo, 08101, Myanmar</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="6AB556D52C7B9A5A15D733043543AFDE">Liu, Fengxiang</mods:namePart>
<mods:affiliation id="B6464542F1D4A97785C7AAAE80B0E368">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
</mods:name>
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<mods:namePart id="DDBA0CCD27121CBD671E13B6766045CB">Yu, Long</mods:namePart>
<mods:nameIdentifier id="ECE779E0095BAC5408AF7F3290EFF859" type="ORCID">0000-0001-5675-1864</mods:nameIdentifier>
<mods:affiliation id="14F41A69F68562694BD888CD9CC9D448">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
</mods:name>
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<mods:role id="DBEF83E89A9ABB8E9679A30A6267A4E2">
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<mods:namePart id="F0B863EF8575FFBE800417B4983E9409">Li, Daiqin</mods:namePart>
<mods:nameIdentifier id="2DEB6E4089B7AD985E7D310BFEAD59F7" type="ORCID">0000-0001-8269-7734</mods:nameIdentifier>
<mods:affiliation id="07EFB7E5937BB80A691C17B67E140C47">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China &amp; Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, Singapore 117543</mods:affiliation>
</mods:name>
<mods:typeOfResource id="4E70E2B7E68E3598276B0451B3486EEF">text</mods:typeOfResource>
<mods:relatedItem id="8C6B324DEC5DDAF000DEE4E85D887453" type="host">
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<mods:title id="C8AE9461942DAB341ED99EEC60658A4A">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="53512BB31739FB336B3B91A3900BC5BB">
<mods:date id="972CE7F291513A9B3D0E3085D7203D0A">2024</mods:date>
<mods:detail id="B72885404E46523CDE3461483D386F28" type="pubDate">
<mods:number id="F668328ECC7A518CBF0F86736491D4B3">2024-08-16</mods:number>
</mods:detail>
<mods:detail id="3B2ECE42F32B6EE6ACE41A271B3CC8B0" type="volume">
<mods:number id="1DE87AE90F89BC90732778D5C3DD7B4B">1210</mods:number>
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<mods:classification id="EC3B8DB474D2B7F4061B0E4320FDA3AC">journal article</mods:classification>
<mods:identifier id="1CDFB1E1160C364B7B4F6FBF8DBB78E6" type="DOI">10.3897/zookeys.1210.123986</mods:identifier>
<mods:identifier id="EB2BCFAADC1F162A92F0E61456AF06BA" type="ZooBank">6972216E-2356-4DF7-BD24-2B6B68E639B4</mods:identifier>
</mods:mods>
<treatment id="BE9AF2F05BBA5A87B548EEDB4256870E" ID-arpha="BE9AF2F0-5BBA-5A87-B548-EEDB4256870E" ID-zoobank="https://zoobank.org/35E36D61-84CB-4C01-830F-BCFE53FBAA6E" LSID="urn:lsid:plazi:treatment:BE9AF2F05BBA5A87B548EEDB4256870E" LSID-ZBK="urn:lsid:zoobank.org:act:35E36D61-84CB-4C01-830F-BCFE53FBAA6E" httpUri="http://treatment.plazi.org/id/BE9AF2F05BBA5A87B548EEDB4256870E">
<subSubSection id="0145F8F88E6DA1E1558D672278BAC01D" type="nomenclature">
<paragraph id="262AFC0357A6F4368FBCEE1F125FDBB7">
<taxonomicName id="E8597CBC1292B2409900E2E3EE7C47D5" authority="Zhan &amp; Xu" authorityName="Zhan &amp; Xu" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rostratus" status="sp. nov.">
<emphasis id="09EC6147A3D2562E225F29B9BD346948" italics="true">Liphistius rostratus</emphasis>
Zhan &amp; Xu
</taxonomicName>
<taxonomicNameLabel id="CB335354974F3FA422691FBCB2719DF2" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="311A172FCD5516F7D55B47E8AD657749" type="description">
<paragraph id="BD270E97EA6B21E53BDAF7D12AD3C2E3">
<figureCitation id="77611BDFE971920F33CD9C4238A7BA83" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">Figs 9</figureCitation>
,
<figureCitation id="EA2FE05A41A0AF18E590CCF09DE2F640" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Vulva of Liphistius rostratus sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure10" httpUri="https://binary.pensoft.net/fig/1115136">10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1KBG" type="type material">
<paragraph id="5559C4DD5D7B5F47CF46371F79E07F58">
<heading id="5DBECA201F1CEBFA7709FD2BC42EC9DE" reason="title">Type material.</heading>
</paragraph>
<paragraph id="815DA4A2B33723FD4D0CDA7AB43B8172">
<materialsCitation id="42AAFE8EA0C379F3CE00162A028F1D6B" collectingDate="2019-07-25" collectorName="D. Li" country="Myanmar" elevation="553" latitude="22.88" location="Mandalay Region" longLatPrecision="754" longitude="96.12" specimenCount="1" specimenCount-male="1" stateProvince="Mandalay" typeStatus="holotype">
<emphasis id="CBF8BB9F608FC32E55E57B086EC5EF5D" bold="true" italics="true">
<typeStatus id="A7FBEB8069606D6FFA4390DB953A83B9" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="18643CDD17069BD6AEA450A8CACF5459" name="Myanmar">Myanmar</collectingCountry>
<specimenCount id="E4D36A634ABA1F69646DCBCE707584E6" count="1" type="male"></specimenCount>
,
<collectingRegion id="F0B7438BB556196C9ADE58665E480187" country="Myanmar" name="Mandalay">Mandalay Region</collectingRegion>
, War Phyu Taung Vill.;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EILBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.120000,22.880000]}">
<geoCoordinate id="CB0FE2FF1669ABDFCB357CAE878797A4" degrees="22.88" direction="north" orientation="latitude" precision="555" value="22.88">22.88 ° N</geoCoordinate>
,
<geoCoordinate id="D37014C72A308503B5608047F3A79C48" degrees="96.12" direction="east" orientation="longitude" precision="555" value="96.12">96.12 ° E</geoCoordinate>
</named-content>
, alt.
<quantity id="E28B7169FE62F39A92FC18A21F689EFC" metricMagnitude="2" metricUnit="m" metricValue="5.529999999999999" unit="m" value="553.0">
<elevation id="002D392D7BB1ED6E1EEE1F20705BDDEE" metricMagnitude="2" metricUnit="m" metricValue="5.529999999999999" unit="m" value="553.0">553 m</elevation>
</quantity>
;
<date id="F82CAB061B07ADAC4009B99B6F14EA09" value="2019-07-25">
<collectingDate id="BD2D02078B30BA8E4E3BF1B07B33480A" value="2019-07-25">25. 07. 2019</collectingDate>
</date>
;
<collectorName id="9134E72FA719992C3F02B0B1F9B1EEB6">D. Li</collectorName>
et al. leg.; XUX- 2019-038
</materialsCitation>
.
<emphasis id="3FA207C1E0A8C04308C09D253E0A8691" bold="true" italics="true">
<typeStatus id="3C4CA49C29155610F512511819E9A11B" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="ECF665EA5B1D05C6C7A96123315342CD" count="2" type="male">2 ♂♂</specimenCount>
<specimenCount id="8232C0E4DDB0F1CD5EF13D05C65B2B48" count="3" type="female">3 ♀♀</specimenCount>
, same data as for
<typeStatus id="45FBA5322C7DE95E163B0C4F03DFF000" type="holotype">holotype</typeStatus>
; XUX- 2019-034 037 / 041.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQLBG" type="diagnosis">
<paragraph id="91CDBF6F21FAD11643F8D6145A2A45A4">
<heading id="20B55ABE211E805F1644DEA0F5B7E9C6" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="FA021153CBB28AC9FF888FE0396F38C8">
The male of
<taxonomicName id="5EE438C63650F296A788D8EBE97DB57A" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rostratus" status="sp. nov.">
<emphasis id="326FDEED752920213DC44168082F3F31" italics="true">L. rostratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EF3FD97276FA1CA5B2C609688B4EFAD3" rank="species">sp. nov.</taxonomicNameLabel>
resembles those of
<taxonomicName id="806EFDE3EF784101464218C6B884297E" authorityName="Schwendinger &amp; Huber" authorityYear="2022" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="8B745EC089BF21D40A0788EC3620A681" italics="true">L. cupreus</emphasis>
</taxonomicName>
,
<taxonomicName id="B17090B61F98652838B2D3D50D6BE3DB" authorityName="Yu, Zhang &amp; Zhang" authorityYear="2021" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nabang">
<emphasis id="2298C0E783E79AE07CE56EDF535BD6FC" italics="true">L. nabang</emphasis>
</taxonomicName>
, and
<taxonomicName id="D783036F8103731304717CE8FF57FCF4" authorityName="Schwendinger &amp; Huber" authorityYear="2022" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="platnicki">
<emphasis id="C2A8F1308FA34F6242B05AAB1F04640C" italics="true">L. platnicki</emphasis>
</taxonomicName>
in having distinctly elevated cumulus (
<abbrev id="ABBRID0ECNBG" xlink_title="cumulus">Cu</abbrev>
) (Fig.
<figureCitation id="4E1A1B43485D9D404C97FAF8B75ACD57" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 C</figureCitation>
), but it can be distinguished from them by the relatively larger paraembolic plate (PeP) (Fig.
<figureCitation id="B76081F6297AD279FCACCC0CE1589A36" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 A, E</figureCitation>
vs figs 21 A, 23 K in
<bibRefCitation id="E2C75F6D6D0679E2CB4BE737950037DF" DOI="10.35929/RSZ.0083" author="Schwendinger" etAl="et al." firstAuthor="Schwendinger" issue="2" journalOrPublisher="Revue Suisse de Zoologie" pagination="375-424" refId="B5" refString="Schwendinger PJ, Huber S, Lehmann-Graber C, Ono H, Aung MM, Hongpadharakiree K (2022) A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species. Revue Suisse de Zoologie 129 (2): 375424. https://doi.org/10.35929/RSZ.0083" title="A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species." volume="129" year="2022">Schwendinger et al. 2022</bibRefCitation>
; fig. 3 A in
<bibRefCitation id="CEB0280A19B688A0583B0B98F79BE71A" DOI="10.3969/j.issn.1005-9628.2021.01.006" author="Yu" etAl="et al." firstAuthor="Yu" refId="B10" refString="Yu K, Zhang F, Zhang JX (2021) First new species of the genus Liphistius Schiödte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30: 3640. https://doi.org/10.3969/j.issn.1005-9628.2021.01.006" year="2021">Yu et al. 2021</bibRefCitation>
), and an indistinct contrategular process (
<abbrev id="ABBRID0EWNBG" xlink_title="contrategular process">cp</abbrev>
) (Fig.
<figureCitation id="CCBD1A1031F5CBF1A1B8A09A88D036A0" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 B</figureCitation>
vs relatively large contrategular process, figs 21 E, 23 E in
<bibRefCitation id="D0F3F1DFD57EDEA751B6F1F6E5A81FA6" DOI="10.35929/RSZ.0083" author="Schwendinger" etAl="et al." firstAuthor="Schwendinger" issue="2" journalOrPublisher="Revue Suisse de Zoologie" pagination="375-424" refId="B5" refString="Schwendinger PJ, Huber S, Lehmann-Graber C, Ono H, Aung MM, Hongpadharakiree K (2022) A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species. Revue Suisse de Zoologie 129 (2): 375424. https://doi.org/10.35929/RSZ.0083" title="A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species." volume="129" year="2022">Schwendinger et al. 2022</bibRefCitation>
; fig. 3 B in
<bibRefCitation id="0DE6AD9633D08F12299E02222821F12F" DOI="10.3969/j.issn.1005-9628.2021.01.006" author="Yu" etAl="et al." firstAuthor="Yu" refId="B10" refString="Yu K, Zhang F, Zhang JX (2021) First new species of the genus Liphistius Schiödte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30: 3640. https://doi.org/10.3969/j.issn.1005-9628.2021.01.006" year="2021">Yu et al. 2021</bibRefCitation>
). The female of
<taxonomicName id="F0318219C914F1BCB24E470947000AED" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rostratus" status="sp. nov.">
<emphasis id="7FA8BC96D2354B8BFFF62E13B3BCACB4" italics="true">L. rostratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0758C94BA8C2283E146ADBA70F364A58" rank="species">sp. nov.</taxonomicNameLabel>
resembles to that of
<taxonomicName id="C41C835EBC262ACDB92DE9F47F619683" authorityName="Schwendinger &amp; Huber" authorityYear="2022" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="391714BD7BEC57302065ED389C3A5C35" italics="true">L. cupreus</emphasis>
</taxonomicName>
in having vesicle clusters along the anterior margin of the poreplate (
<abbrev id="ABBRID0E3OBG" xlink_title="sclerotized poreplate">PPl</abbrev>
), but it can be distinguished by the relatively longer posterior stalk (
<collectionCode id="450895A77A2275F4574CB105711F65BC">PS</collectionCode>
), with a length / width ratio 0.40.6 (Fig.
<figureCitation id="2C11D1F39D5BC7932BDF22C55DAB05F1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Vulva of Liphistius rostratus sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure10" httpUri="https://binary.pensoft.net/fig/1115136">10</figureCitation>
vs length / width ratio about 0.20.4, fig.
<quantity id="DCBEA25F36C8526B3EBA95F6C94D7995" metricMagnitude="-1" metricUnit="m" metricValue="5.588" unit="in" value="22.0">22 in</quantity>
<bibRefCitation id="17F5A1ADA3061878F531AEAB8F61D438" DOI="10.35929/RSZ.0083" author="Schwendinger" etAl="et al." firstAuthor="Schwendinger" issue="2" journalOrPublisher="Revue Suisse de Zoologie" pagination="375-424" refId="B5" refString="Schwendinger PJ, Huber S, Lehmann-Graber C, Ono H, Aung MM, Hongpadharakiree K (2022) A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species. Revue Suisse de Zoologie 129 (2): 375424. https://doi.org/10.35929/RSZ.0083" title="A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species." volume="129" year="2022">Schwendinger et al. 2022</bibRefCitation>
).
</paragraph>
<caption id="C997D08E1B159160B1111B4367F6CB80" ID-DOI="10.3897/zookeys.1210.123986.figure9" ID-arpha="EE41BB75-A320-5356-97B2-C4FE897FAE94" httpUri="https://binary.pensoft.net/fig/1115135" startId="F9">
<paragraph id="0B8918B9F376B296EA199C5BA95B021C">
<label id="A74677FAF2EE11C49ED98FC00B948041">Figure 9.</label>
</paragraph>
<paragraph id="E4E7A17E2483CB868FB73476506F0EDB">
Male palp of
<taxonomicName id="C2BD0368F33DAB0CC5DCB577EA022CC2" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rostratus" status="sp. nov.">
<emphasis id="5A93A17998703565FA74E33CAB344E46" italics="true">Liphistius rostratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EEF583394BB7FDEF2CE6D8F072D19351" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="3A9CA78A8EA9C297E4DB53F04F4B1926" bold="true">A, E</emphasis>
prolateral view
<emphasis id="9B6B17F6047FF04203177E70EC9D8295" bold="true">B</emphasis>
ventral view
<emphasis id="568E02C705575F2866D9051E81F0BB98" bold="true">C, F</emphasis>
retrolateral view
<emphasis id="1EEB3159C4B6601AD0628B05921C0E49" bold="true">D</emphasis>
dorsal view
<emphasis id="E9B8CECB46EEBB67947497C325673E2F" bold="true">G</emphasis>
distal view, red arrow shows beak-like contrategular projection. Abbreviations:
<abbrev id="ABBRID0EFQBG" xlink_title="contrategulum">CT</abbrev>
= contrategulum;
<abbrev id="ABBRID0EJQBG" xlink_title="contrategular process">cp</abbrev>
= contrategular process;
<abbrev id="ABBRID0ENQBG" xlink_title="cumulus">Cu</abbrev>
= cumulus; E = embolus;
<abbrev id="ABBRID0ERQBG" xlink_title="paracymbium">PC</abbrev>
= paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum;
<abbrev id="ABBRID0EVQBG" xlink_title="male palps bearing tibial apophysis">TiA</abbrev>
= tibial apophysis. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="5874242F0D3072C948265B27F6789200" ID-DOI="10.3897/zookeys.1210.123986.figure10" ID-arpha="CC0BDFD5-5D2E-5F2C-8AD4-7292A822B05C" httpUri="https://binary.pensoft.net/fig/1115136" startId="F10">
<paragraph id="C208B9D425864655700D5C5F10C19B72">
<label id="4811C0E7F56A72BED1E4D5CA07417F31">Figure 10.</label>
</paragraph>
<paragraph id="8331AD0BBE8B9D2E9AFE25EDABE9A38C">
Vulva of
<taxonomicName id="8F21898EC6B18F97129EFC8C00B880C6" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rostratus" status="sp. nov.">
<emphasis id="78ACCF574389A7EA04C50AC7E663E281" italics="true">Liphistius rostratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0898230BBC5D53A55CD93726DA6A1F75" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="6187C14FC0B43D9414700CAF988C3C98" bold="true">A, C</emphasis>
dorsal view
<emphasis id="9D80BAE9E565F11389839EB7C457D5A8" bold="true">B, D</emphasis>
ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk;
<abbrev id="ABBRID0EVRBG" xlink_title="sclerotized poreplate">PPl</abbrev>
= poreplate;
<abbrev id="ABBRID0EZRBG" xlink_title="receptacular cluster">RC</abbrev>
= receptacular cluster. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ECSBG" type="description">
<paragraph id="F9882118EB337685EFEA08690C49D4F2">
<heading id="883A1ACE69E9534BEE4740E1852B23D1" reason="title">Description.</heading>
</paragraph>
<paragraph id="CE23517DE7E8169960F48501E2F16111">
<emphasis id="3A3AEF8E20E8D44AAC607CE574F7DFD0" bold="true">
Male
<typeStatus id="D7BF5644204985CFDF002223F9135C5F" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace yellowish brown, with few short, scattered bristles; opisthosoma brown, with 12 brown tergites, close to each other, 26
<superScript id="D36AD013E6027708BEE4BA99B83B2CFD">th</superScript>
larger than others, 4
<superScript id="248A3F61E9222084A6B2E905A609018B">th</superScript>
largest; chelicerae robust, promargin of cheliceral groove with 12 denticles of variable size; labium and sternum yellowish brown, sternum with few short setae on anterior tip and many long setae on elongated posterior tip; legs yellowish brown, with strong setae and spines. Measurements:
<abbrev id="ABBRID0EOSBG" xlink_title="body length">BL</abbrev>
9.34,
<abbrev id="ABBRID0ESSBG" xlink_title="carapace length">CL</abbrev>
4.57,
<abbrev id="ABBRID0EWSBG" xlink_title="carapace width">CW</abbrev>
4.26,
<abbrev id="ABBRID0E1SBG" xlink_title="opisthosoma length">OL</abbrev>
5.02,
<abbrev id="ABBRID0E5SBG" xlink_title="opisthosoma width">OW</abbrev>
3.59; eye sizes and interdistances:
<abbrev id="ABBRID0ECTBG" xlink_title="anterior median eyes">AME</abbrev>
reduced,
<abbrev id="ABBRID0EGTBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.40,
<abbrev id="ABBRID0EKTBG" xlink_title="posterior median eyes">PME</abbrev>
reduced,
<abbrev id="ABBRID0EOTBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.24,
<abbrev id="ABBRID0ESTBG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EWTBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.07,
<abbrev id="ABBRID0E1TBG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0E5TBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.05,
<abbrev id="ABBRID0ECUBG" xlink_title="posterior lateral eyes">PLE</abbrev>
<abbrev id="ABBRID0EGUBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.30; labium 0.98 long and 0.48 wide; sternum 2.09 long and 0.83 wide; legs: I 12.63 (3.80, 1.32, 2.60, 3.07, 1.84), II 13.19 (3.80, 1.32, 2.60, 3.07, 1.84), III 14.35 (3.79, 1.58, 3.33, 3.44, 2.21), IV 19.76 (4.76, 1.66, 4.13, 6.01, 3.20).
</paragraph>
<paragraph id="D191393659F1496FACD56540BD510CEC">
Palp: tibial apophysis (
<abbrev id="ABBRID0EMUBG" xlink_title="male palps bearing tibial apophysis">TiA</abbrev>
) with 4 setae of similar length (Fig.
<figureCitation id="FF1E6D481C3B87FC8A6303E15D731FAC" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 A C</figureCitation>
); paracymbium (
<abbrev id="ABBRID0EUUBG" xlink_title="paracymbium">PC</abbrev>
) with several setae situated on tip (Fig.
<figureCitation id="7C097F7273A1C825966B474F60897A40" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 B, C</figureCitation>
); cumulus (
<abbrev id="ABBRID0E3UBG" xlink_title="cumulus">Cu</abbrev>
) distinct elevated with 5 tapering spines (Fig.
<figureCitation id="714D27869EFE0A63388481EF36FC6C71" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 C</figureCitation>
); subtegulum (ST) without apophysis (Fig.
<figureCitation id="1A5986685701DDF1168BD26D037F6AE0" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 F</figureCitation>
); contrategulum (
<abbrev id="ABBRID0EIVBG" xlink_title="contrategulum">CT</abbrev>
) with beak-like projection distally in distal view and small process in ventral view (Fig.
<figureCitation id="08E6A40F6B8C646A7FD7F2373472D156" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 B, G</figureCitation>
); tegulum (
<collectionCode id="F59DBCE55A561ABEE3F32EEAE4A84996" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
) semicircular with flat surface and serrated edges, and proximal tegular margin with an elevated transverse ridge (Fig.
<figureCitation id="D1C454F4665E54D898464D1BA7F320B7" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 B, C, F</figureCitation>
); paraembolic plate (PeP) sclerotized, narrow (Fig.
<figureCitation id="5F3882DE8B0D654A0D33C9B87653B029" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 A, E</figureCitation>
); embolus (E) with 3 distinct longitudinal ridges reaching tip retrolaterally, and several denticles along these longitudinal ridges (Fig.
<figureCitation id="A4ABB38C28D297CA1E131C2A3ED6AC01" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 B, F</figureCitation>
).
</paragraph>
<paragraph id="293BE238132A85EFC7869AD75DB4DC08">
<emphasis id="A9033D247D702D04B5FE4497DD818650" bold="true">
Female
<typeStatus id="1F980AEAE0707D0397515DE3B5AE9A91" type="paratype">paratype</typeStatus>
(XUX- 2019-034).
</emphasis>
Carapace brown with few short, scattered bristles; opisthosoma brown, with 12 brown tergites, close to each other, 26
<superScript id="A2E8FFE5CC3DE05F5CA69360B180BE24">th</superScript>
larger than others, 5
<superScript id="A6E39A9204A4DD25FDA26DF5C2CD5511">th</superScript>
largest; chelicerae robust, reddish brown; promargin of cheliceral groove with 12 denticles of variable size; labium yellowish brown, sternum yellowish brown with several setae; legs with strong setae and spines. Measurements:
<abbrev id="ABBRID0EEWBG" xlink_title="body length">BL</abbrev>
10.54,
<abbrev id="ABBRID0EIWBG" xlink_title="carapace length">CL</abbrev>
4.5,
<abbrev id="ABBRID0EMWBG" xlink_title="carapace width">CW</abbrev>
3.93,
<abbrev id="ABBRID0EQWBG" xlink_title="opisthosoma length">OL</abbrev>
6.47,
<abbrev id="ABBRID0EUWBG" xlink_title="opisthosoma width">OW</abbrev>
4.74; eye sizes and interdistances:
<abbrev id="ABBRID0EYWBG" xlink_title="anterior median eyes">AME</abbrev>
0.05,
<abbrev id="ABBRID0E3WBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.37,
<abbrev id="ABBRID0EAXBG" xlink_title="posterior median eyes">PME</abbrev>
0.19,
<abbrev id="ABBRID0EEXBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.29,
<abbrev id="ABBRID0EIXBG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EMXBG" xlink_title="anterior median eyes">AME</abbrev>
0.07,
<abbrev id="ABBRID0EQXBG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EUXBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.12,
<abbrev id="ABBRID0EYXBG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0E3XBG" xlink_title="posterior median eyes">PME</abbrev>
0.06,
<abbrev id="ABBRID0EAYBG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0EEYBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.06,
<abbrev id="ABBRID0EIYBG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EMYBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.09,
<abbrev id="ABBRID0EQYBG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EUYBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.10,
<abbrev id="ABBRID0EYYBG" xlink_title="posterior lateral eyes">PLE</abbrev>
<abbrev id="ABBRID0E3YBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.32,
<abbrev id="ABBRID0EAZBG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EEZBG" xlink_title="posterior median eyes">PME</abbrev>
0.08; labium 0.95 long and 0.38 wide; sternum 2.41 long and 0.99 wide; palp 7.10 (2.44, 0.98, 1.86, 1.82), leg I 8.89 (2.94, 1.34, 2.00, 1.62, 0.99), II 9.69 (3.40, 1.09, 2.02, 1.98, 1.20), III 9.48 (2.29, 1.05, 2.59, 2.38, 1.17), IV 13.71 (3.21, 1.44, 3.11, 3.80, 2.21).
</paragraph>
<paragraph id="33580FF4BFA1C1C1D14C8E77933699A4">
Vulva: about rectangular poreplate (
<abbrev id="ABBRID0EKZBG" xlink_title="sclerotized poreplate">PPl</abbrev>
) with pair of anterior lobes; central dorsal opening (
<collectionCode id="905D1A264670DAA4400755019B26DA0F">CDO</collectionCode>
) located below center of poreplate (
<abbrev id="ABBRID0EOZBG" xlink_title="sclerotized poreplate">PPl</abbrev>
); racemose receptacular cluster (
<abbrev id="ABBRID0ESZBG" xlink_title="receptacular cluster">
<collectionCode id="235FC8924BD81EFE2297B3C9243BB828">RC</collectionCode>
</abbrev>
) small; posterior stalk (
<collectionCode id="03B36BD368534646B7262D7B23756475">PS</collectionCode>
) as wide as poreplate (
<abbrev id="ABBRID0EWZBG" xlink_title="sclerotized poreplate">PPl</abbrev>
), lateral margins of genital atrium (GA) slightly sclerotized (Fig.
<figureCitation id="A94D011792C7BE6AB12129E711F2D17D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Vulva of Liphistius rostratus sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure10" httpUri="https://binary.pensoft.net/fig/1115136">10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5ZBG" type="variation">
<paragraph id="B6590AE953A64F095D8115141E43C726">
<heading id="912DC78384B05B2CDE02C031DA46A89C" reason="title">Variation.</heading>
</paragraph>
<paragraph id="7FF2073944B35C35DC7D414C8FBCE720">
Males (
<emphasis id="817B360FBD7EF3EF703834D6433B6453" italics="true">N</emphasis>
= 3):
<abbrev id="ABBRID0EG1BG" xlink_title="body length">BL</abbrev>
9.3410.31,
<abbrev id="ABBRID0EK1BG" xlink_title="carapace length">CL</abbrev>
4.575.24,
<abbrev id="ABBRID0EO1BG" xlink_title="carapace width">CW</abbrev>
4.265.83,
<abbrev id="ABBRID0ES1BG" xlink_title="opisthosoma length">OL</abbrev>
4.735.10,
<abbrev id="ABBRID0EW1BG" xlink_title="opisthosoma width">OW</abbrev>
3.403.40; females (
<emphasis id="52702B85236FB06A351F605BE28BF7E4" italics="true">N</emphasis>
= 4):
<abbrev id="ABBRID0E31BG" xlink_title="body length">BL</abbrev>
10.5412.35,
<abbrev id="ABBRID0EA2BG" xlink_title="carapace length">CL</abbrev>
4.506.31,
<abbrev id="ABBRID0EE2BG" xlink_title="carapace width">CW</abbrev>
3.935.47,
<abbrev id="ABBRID0EI2BG" xlink_title="opisthosoma length">OL</abbrev>
6.476.87,
<abbrev id="ABBRID0EM2BG" xlink_title="opisthosoma width">OW</abbrev>
4.644.79. The number of denticles on the cheliceral promargin varies from 12 to 13 (
<emphasis id="089289E164CCBD9ED51C231FD76CB1E3" italics="true">N</emphasis>
= 6). In addition, the male lacking an
<abbrev id="ABBRID0ES2BG" xlink_title="anterior median eyes">AME</abbrev>
and
<abbrev id="ABBRID0EW2BG" xlink_title="posterior median eyes">PME</abbrev>
, which may be degenerate during molting, is chosen as the
<typeStatus id="005E9B890AE75A15F04DAAF85D5DE9CC" type="holotype">holotype</typeStatus>
, because all
<typeStatus id="1684F557AB335E52DF1FE705B4814689" type="paratype">paratype</typeStatus>
male palps are deformed.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E12BG" type="etymology">
<paragraph id="295AED6F79390A21DAA2A02E5F4D8876">
<heading id="3778F46B2963AC25B42089628846C180" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="3968C4DF81B826F56791C1ECA0647ECD">
The species name is derived from the Latin word
<taxonomicName id="79903805B446A136631194A4653ADB56" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rostratus">
<emphasis id="4C8DD574F58FCD9534F44EE8B3CA1A62" italics="true">rostratus</emphasis>
</taxonomicName>
”, referring to the beak-like contrategulum of the male palp in distal view.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EK3BG" type="distribution">
<paragraph id="C4859CBF17DBAD7600019C4D24EF553D">
<heading id="3C6513F111C92CF0310A9F28FA73C9C9" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="DEAD99E9BC2F569366116E4CA1C1B3F8">
<collectingRegion id="B90F85A46ABE6D42BD1BC3BD739E8E33" country="Myanmar" name="Mandalay">Mandalay Region</collectingRegion>
(War Hpyu Taung),
<collectingCountry id="5F0FBAF34B9B2ADBA5B93D55D286CDFD" name="Myanmar">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,477 @@
<document id="CF5A94C5252363BEC13C182C892FE4E1" ID-DOI="10.3897/zookeys.1210.123986" ID-ZooBank="6972216E-2356-4DF7-BD24-2B6B68E639B4" ID-publisher-id="123986" URI-arpha="B6184626-6862-59C4-9047-EF9B7AC24E35" URI-zoobank="http://zoobank.org/6972216E-2356-4DF7-BD24-2B6B68E639B4" article-type="research-article" checkinTime="1723830749198" checkinUser="pensoft" docAuthor="Xu, Xin, Zhan, Yi, Aung, Khin Pyae Pyae, Liu, Fengxiang, Yu, Long &amp; Li, Daiqin" docDate="2024" docId="D0DEE7F143A9512D89942689CD66F8DD" docLanguage="en" docName="ZooKeys 1210: 99-115" docOrigin="ZooKeys 1210" docSource="https://zookeys.pensoft.net/article/123986/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Liphistius kalaw Zhan &amp; Xu 2024, sp. nov." docType="treatment" docVersion="1" dtd-version="3.0" lastPageNumber="115" masterDocId="B6184626686259C49047EF9B7AC24E35" masterDocTitle="Four new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae, Liphistiinae) from Myanmar" masterLastPageNumber="115" masterPageNumber="99" pageNumber="99" updateTime="1723830805509" updateUser="pensoft">
<mods:mods id="A9051C9765FF156746688128D3819540" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EDF83D5F07BDDD2BB4DD3A0C3A975C7F">
<mods:title id="D515C10FD4791914DE53061C16006BDA">Four new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae, Liphistiinae) from Myanmar</mods:title>
</mods:titleInfo>
<mods:name id="D37589D4127E0E21DD5B0694C93911B2" type="personal">
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<mods:namePart id="AD7EBCDFE08B032E1AA2D7E6F0252D21">Xu, Xin</mods:namePart>
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<mods:affiliation id="1144392E72BBC643158D0A63F29BBE9A">College of Life Sciences, Hunan Normal University, 36 Lushan Road, Changsha 410081, Hunan Province, China &amp; State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
</mods:name>
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<mods:namePart id="023D6D00C0F13EDDA671FCD58E6093F5">Zhan, Yi</mods:namePart>
<mods:affiliation id="49CE9E82610F06DAE9198D98471DE474">College of Life Sciences, Hunan Normal University, 36 Lushan Road, Changsha 410081, Hunan Province, China</mods:affiliation>
</mods:name>
<mods:name id="7E699F479334976F1E810851470DA102" type="personal">
<mods:role id="B745884588F63852A455ACF13B1F7F1C">
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<mods:namePart id="FFA7A8783EC347B3F86F72ED7EEF98B4">Aung, Khin Pyae Pyae</mods:namePart>
<mods:affiliation id="FE1DBB220C48910E42987D773916F11D">Department of Zoology, University of Yangon, Kamayut Township, Pyay Road, Yangon, 11041, Myanmar &amp; Department of Biology, Taungoo Education Degree College, Taungoo, 08101, Myanmar</mods:affiliation>
</mods:name>
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<mods:role id="243E04F608DF573782E440F83ADA96F1">
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</mods:role>
<mods:namePart id="F36F0FD22BCCAEF84A8E6AD7379C3DCE">Liu, Fengxiang</mods:namePart>
<mods:affiliation id="A1E883A191335956F5D1BFE6A79194BF">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
</mods:name>
<mods:name id="E14BDDC985A8FD03D133D846B2742600" type="personal">
<mods:role id="6F964ACDDBFDAB931F7B88E33D0BFB1A">
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</mods:role>
<mods:namePart id="24A7D16C381FCFE97A0C3B1ADE036F30">Yu, Long</mods:namePart>
<mods:nameIdentifier id="3BCD715C23CB0CB267F8EE0EC7891B4F" type="ORCID">0000-0001-5675-1864</mods:nameIdentifier>
<mods:affiliation id="7C57F8CCEF6A0D9D1150C4CBFD6D89A0">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
</mods:name>
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<mods:roleTerm id="CF6CA11B815498E92566652A3ED21699">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1F46B90407CA2981A182DC469994B99E">Li, Daiqin</mods:namePart>
<mods:nameIdentifier id="54CCE02D29D6613F473AAA17C01E94CF" type="ORCID">0000-0001-8269-7734</mods:nameIdentifier>
<mods:affiliation id="686342C7EF0E225CE3303F52AF3F6316">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China &amp; Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, Singapore 117543</mods:affiliation>
</mods:name>
<mods:typeOfResource id="B2E7567224C49144442B18112C83F62C">text</mods:typeOfResource>
<mods:relatedItem id="A66A4D7C1C8637BB44949B2A0876834B" type="host">
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<mods:title id="599B95CD32A9A86ED12E0D2FEEE93137">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="DF6F602564F8BD009E6FF94B30D599A8">
<mods:date id="7F5ECA8E37E4D8F56F08D5A2A5391948">2024</mods:date>
<mods:detail id="2DD6B86E57004FE4EC34C244641961E2" type="pubDate">
<mods:number id="3EB2AD1C7A8A2244B4FED91518951969">2024-08-16</mods:number>
</mods:detail>
<mods:detail id="4D0160988E16AE98F5C954E1C18E2DC4" type="volume">
<mods:number id="C4E63D768F371BEC2297A5E25FFB2AA6">1210</mods:number>
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</mods:extent>
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<mods:classification id="A18803E4F8051C4BA0696AD31BA04DE7">journal article</mods:classification>
<mods:identifier id="1D4D117E8DECB6464614BFACE1FE18A5" type="DOI">10.3897/zookeys.1210.123986</mods:identifier>
<mods:identifier id="5D7ED8FF5CA33A1F4BE7AE0AAC5AE3B2" type="ZooBank">6972216E-2356-4DF7-BD24-2B6B68E639B4</mods:identifier>
</mods:mods>
<treatment id="D0DEE7F143A9512D89942689CD66F8DD" ID-arpha="D0DEE7F1-43A9-512D-8994-2689CD66F8DD" ID-zoobank="https://zoobank.org/DE92B21F-6A12-44AA-97C8-18EE985DB623" LSID="urn:lsid:plazi:treatment:D0DEE7F143A9512D89942689CD66F8DD" LSID-ZBK="urn:lsid:zoobank.org:act:DE92B21F-6A12-44AA-97C8-18EE985DB623" httpUri="http://treatment.plazi.org/id/D0DEE7F143A9512D89942689CD66F8DD">
<subSubSection id="A884F08AD0BECE3E1B07BBA00ED458B7" type="nomenclature">
<paragraph id="7697DD43CA2A31C0E20FBEEA00055E91">
<taxonomicName id="9FAD99FCB5476A2304A84C0549908310" authority="Zhan &amp; Xu" authorityName="Zhan &amp; Xu" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kalaw" status="sp. nov.">
<emphasis id="FFA6E1CA087FE5340DA403329561A2F2" italics="true">Liphistius kalaw</emphasis>
Zhan &amp; Xu
</taxonomicName>
<taxonomicNameLabel id="4B98F382EF4F7E7A9CCD9C62E562B156" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="9A0A9EC0F8B649FD72507E571510472C" type="description">
<paragraph id="49A58B47B4061D201C59CEC137C7AF17">
<figureCitation id="EA99D9B515CCF172F034E5AE643B3A5E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">Figs 3</figureCitation>
,
<figureCitation id="4F4DBB368120D874F923D9C520363B76" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Vulva of Liphistius kalaw sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster; a = length of poreplate transverse; b = width of poreplate. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure4" httpUri="https://binary.pensoft.net/fig/1115130">4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYQAE" type="type material">
<paragraph id="2FC19CB69B34CB7E6282CA1400115C42">
<heading id="1F2D2686356F8F4CF919E53BA54FCD4E" reason="title">Type material.</heading>
</paragraph>
<paragraph id="7AF7B31AEC21A24F6B4B54B483D43A5C">
<materialsCitation id="53F4BCAA58F21C5CC2D7557C53F90B66" collectingDate="2019-07-27" collectorName="D. Li" country="Myanmar" elevation="944" latitude="20.7" location="Kalaw Township" longLatPrecision="760" longitude="96.52" specimenCount="1" specimenCount-male="1" stateProvince="Shan" typeStatus="holotype">
<emphasis id="33C07F1B0EF89C09D7612CEC912C6CAD" bold="true" italics="true">
<typeStatus id="B25E46CF0DEC90DC20451089BC2476F6" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="451162A1B81C14654F3AF5C288883037" name="Myanmar">Myanmar</collectingCountry>
<specimenCount id="21F27CE7B6F92EF9645905833DC0C276" count="1" type="male"></specimenCount>
,
<collectingRegion id="F495C1300AB809EFE4299B1BC2391E80" country="Myanmar" name="Shan">Shan State</collectingRegion>
,
<quantity id="9EE14365F66661529337784230818364" metricMagnitude="4" metricUnit="m" metricValue="1.6" unit="km" value="16.0">16 km</quantity>
W of
<location id="B4ADAFC160019F68D76066695500F4F0" LSID="urn:lsid:plazi:treatment:D0DEE7F143A9512D89942689CD66F8DD:B4ADAFC160019F68D76066695500F4F0" country="Myanmar" latitude="20.7" longLatPrecision="760" longitude="96.52" name="Kalaw Township" stateProvince="Shan">Kalaw Township</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGRAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.520000,20.700000]}">
<geoCoordinate id="00F6DA60F5D2F7AC77867AC94C0233F5" degrees="20.70" direction="north" orientation="latitude" precision="555" value="20.7">20.70 ° N</geoCoordinate>
,
<geoCoordinate id="F963033B304EE9D95E333A95ABC70CFF" degrees="96.52" direction="east" orientation="longitude" precision="555" value="96.52">96.52 ° E</geoCoordinate>
</named-content>
, alt.
<quantity id="A4AE25DAFA7C5387CF6DB229F7C30B9B" metricMagnitude="2" metricUnit="m" metricValue="9.44" unit="m" value="944.0">
<elevation id="80256E94F5B89B8BB8C7CA918AD8AA22" metricMagnitude="2" metricUnit="m" metricValue="9.44" unit="m" value="944.0">944 m</elevation>
</quantity>
;
<date id="7A1B2F1448D5C8ECBBBEB0E89BE0C2F6" value="2019-07-27">
<collectingDate id="0124A5DAAF3CDA8F71C61255838A8BA4" value="2019-07-27">27. 07. 2019</collectingDate>
</date>
;
<collectorName id="69A802D1D8B6DF0AD49DA785D9D1061A">D. Li</collectorName>
et al. leg.; XUX- 2019-061 A
</materialsCitation>
.
<emphasis id="8F19F0D7ACDB88497C40541B40E5861E" bold="true" italics="true">
<typeStatus id="2DE4C9F46275B618B90CCA5B88E70815" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="90597EFB0DCD1507B1F445FA2DB9FC61" count="8" type="female">8 ♀♀</specimenCount>
, same data as for
<typeStatus id="8F002A51716E322FDEDCE4AA2132B443" type="holotype">holotype</typeStatus>
;
<date id="1F8DB126501D817DD5F58C9C75CE585A" value="2019-07-27">27. 07. 2019</date>
and
<date id="A087323410CCDE602CFF3B959E76545B" value="2018-07-15">15. 07. 2018</date>
; XUX- 2019-062 / 063 / 064 / 065 / 066 / 067 A / 068; XUX- 2018-124.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EORAE" type="diagnosis">
<paragraph id="BDB6BC3B442498F5D15CF85BB72645CC">
<heading id="80DA1317773417AF600C6709BFEE2C24" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="B6EDD8C75065E76CF562B587720E6232">
The male of
<taxonomicName id="7D55D93C6E3B132B12348BEC02C0FDF3" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kalaw" status="sp. nov.">
<emphasis id="A8A4BCB54A28680C5CCDA84B3F22CA2C" italics="true">L. kalaw</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DB324125438FD76707239F2A2D51ACF" rank="species">sp. nov.</taxonomicNameLabel>
resembles those of
<taxonomicName id="6F0FBF3F651E0DD240759E87A8E1EDAC" authorityName="Thorell" authorityYear="1897" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="birmanicus">
<emphasis id="45FEC0A7F53BB77F4A5FD033FB352D56" italics="true">L. birmanicus</emphasis>
</taxonomicName>
and
<taxonomicName id="D862C29F2C7B1FDF85472CC91929D55F" authorityName="Aung, Xu, Lwin, Sang, Yu, Liu, Liu &amp; Li" authorityYear="2019" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="pinlaung">
<emphasis id="4490ACDA21C90B2C50CB4CFF1E47488E" italics="true">L. pinlaung</emphasis>
</taxonomicName>
in having a distinct contrategular process (
<abbrev id="ABBRID0EVSAE" xlink_title="contrategular process">cp</abbrev>
) (Fig.
<figureCitation id="532C0F54AC3D39A91F3799FAAE080BA8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 B, E</figureCitation>
), but it can be distinguished from
<taxonomicName id="695E7ECAA72C02A841D7654E2A19B49B" authorityName="Thorell" authorityYear="1897" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="birmanicus">
<emphasis id="F68D371600E1F8E1D9E90ADA793B0B30" italics="true">L. birmanicus</emphasis>
</taxonomicName>
by the relatively wider base of the contrategular process (
<abbrev id="ABBRID0EITAE" xlink_title="contrategular process">cp</abbrev>
) (Fig.
<figureCitation id="8FA71DCAF8C402E237D5BE4DCC6AA45A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 E</figureCitation>
vs fig. 13 E in
<bibRefCitation id="D21EAFCB97E687EE8341AEEE7DF1BB82" DOI="10.35929/RSZ.0083" author="Schwendinger" etAl="et al." firstAuthor="Schwendinger" issue="2" journalOrPublisher="Revue Suisse de Zoologie" pagination="375-424" refId="B5" refString="Schwendinger PJ, Huber S, Lehmann-Graber C, Ono H, Aung MM, Hongpadharakiree K (2022) A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species. Revue Suisse de Zoologie 129 (2): 375424. https://doi.org/10.35929/RSZ.0083" title="A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species." volume="129" year="2022">Schwendinger et al. 2022</bibRefCitation>
), and from
<taxonomicName id="28FC45154F217CE46AD73A6CF1E5983E" authorityName="Aung, Xu, Lwin, Sang, Yu, Liu, Liu &amp; Li" authorityYear="2019" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="pinlaung">
<emphasis id="3CAA47B0C9F244FE089922B06199F184" italics="true">L. pinlaung</emphasis>
</taxonomicName>
by the cumulus (
<abbrev id="ABBRID0E6TAE" xlink_title="cumulus">Cu</abbrev>
) which has shorter spines (Fig.
<figureCitation id="1430FFA7133F3D9027E5713E841A3F7A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 A C</figureCitation>
vs fig. 4 C in
<bibRefCitation id="81544AB47DBD03953F3A7F785740772B" DOI="10.3897/zookeys.882.38811" author="Aung" etAl="et al." firstAuthor="Aung" journalOrPublisher="ZooKeys" pagination="29-39" refId="B1" refString="Aung KPP, Xu X, Lwin WW, Sang MZ, Yu L, Liu H, Liu FX, Li DQ (2019) Two new species of the primitively segmented spider genus Liphistius Schiödte, 1849 (Mesothelae, Liphistiidae) from Myanmar. ZooKeys 882: 2939. https://doi.org/10.3897/zookeys.882.38811" title="Two new species of the primitively segmented spider genus Liphistius Schiödte, 1849 (Mesothelae, Liphistiidae) from Myanmar." volume="882" year="2019">Aung et al. 2019</bibRefCitation>
). The female of
<taxonomicName id="E19C7BB574185BBABFAC9B84DD5F054E" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kalaw" status="sp. nov.">
<emphasis id="576E90D97DFAEA4C0486602F9B2667D5" italics="true">L. kalaw</emphasis>
</taxonomicName>
<taxonomicNameLabel id="931BBF0F0BF8269EAD486516DCEA8CF6" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from that of
<taxonomicName id="BC2B77502C1CB1BD12A0DC2B2EFD01F2" authorityName="Aung, Xu, Lwin, Sang, Yu, Liu, Liu &amp; Li" authorityYear="2019" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="hpruso">
<emphasis id="537B2011B4AB10CE0B93F661A803AC43" italics="true">L. hpruso</emphasis>
</taxonomicName>
by the relatively wider posterior stalk (
<collectionCode id="867F3C6C4452B7042B22B740955FA135">PS</collectionCode>
) (Fig.
<figureCitation id="CD4EABC3E90DAF5C2C327AFA6D301EF2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Vulva of Liphistius kalaw sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster; a = length of poreplate transverse; b = width of poreplate. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure4" httpUri="https://binary.pensoft.net/fig/1115130">4 C</figureCitation>
vs fig. 3 B E in
<bibRefCitation id="734CD7994C9D6E7710355B987BEAA366" DOI="10.3897/zookeys.882.38811" author="Aung" etAl="et al." firstAuthor="Aung" journalOrPublisher="ZooKeys" pagination="29-39" refId="B1" refString="Aung KPP, Xu X, Lwin WW, Sang MZ, Yu L, Liu H, Liu FX, Li DQ (2019) Two new species of the primitively segmented spider genus Liphistius Schiödte, 1849 (Mesothelae, Liphistiidae) from Myanmar. ZooKeys 882: 2939. https://doi.org/10.3897/zookeys.882.38811" title="Two new species of the primitively segmented spider genus Liphistius Schiödte, 1849 (Mesothelae, Liphistiidae) from Myanmar." volume="882" year="2019">Aung et al. 2019</bibRefCitation>
).
</paragraph>
<caption id="716D9CA6E2CC7F89ED84E1B48F9C5D3B" ID-DOI="10.3897/zookeys.1210.123986.figure3" ID-arpha="CAB41D52-B773-5360-B4B5-E82229AE1B2B" httpUri="https://binary.pensoft.net/fig/1115129" startId="F3">
<paragraph id="FEDAA9C605721BD9C81AA96023EB5C1A">
<label id="9E1E807F39D3FFFE611B96010363CBC5">Figure 3.</label>
</paragraph>
<paragraph id="06EB19FB0A5B2107C75A2F5F3D472380">
Male palp of
<taxonomicName id="A64F30F01376BF074BDF8B5F190530A2" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kalaw" status="sp. nov.">
<emphasis id="803471CCE6C39ED0B6C1677828D3CEA0" italics="true">Liphistius kalaw</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0660BAAFEB09D0520E9B02756D401417" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="3AE3EAF7200FB35B3B82E9C6B5D7E083" bold="true">A, E</emphasis>
prolateral view
<emphasis id="074CACA811D02B032D62FCC513D96A4C" bold="true">B</emphasis>
ventral view
<emphasis id="E60E5B15F3B10B713FDF0E36FB512DFD" bold="true">C, F</emphasis>
retrolateral view
<emphasis id="EF0424CE16C88FB3968F8E3F2EAFF6A6" bold="true">D</emphasis>
dorsal view, red arrows show arched projection and triangular process of
<abbrev id="ABBRID0EEWAE" xlink_title="contrategulum">CT</abbrev>
<emphasis id="55D80DD0D7A89193ACB6410621E9EC9C" bold="true">G</emphasis>
distal view. Abbreviations:
<abbrev id="ABBRID0EKWAE" xlink_title="contrategulum">CT</abbrev>
= contrategulum;
<abbrev id="ABBRID0EOWAE" xlink_title="contrategular process">cp</abbrev>
= contrategular process;
<abbrev id="ABBRID0ESWAE" xlink_title="cumulus">Cu</abbrev>
= cumulus; E = embolus;
<abbrev id="ABBRID0EWWAE" xlink_title="paracymbium">PC</abbrev>
= paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum;
<abbrev id="ABBRID0E1WAE" xlink_title="male palps bearing tibial apophysis">TiA</abbrev>
= tibial apophysis. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="D24C36FAB214EEC5F71D0BDA2D0C0FAB" ID-DOI="10.3897/zookeys.1210.123986.figure4" ID-arpha="9C5A999A-9CB0-5F28-B4B1-27C4BBCBC908" httpUri="https://binary.pensoft.net/fig/1115130" startId="F4">
<paragraph id="B9BB8AB9254FBFAFBA1406B0739C9B73">
<label id="D33A8680BD432C7EF8F71FB2013FA702">Figure 4.</label>
</paragraph>
<paragraph id="45B303E90176297F0D30C2A5AFE812FD">
Vulva of
<taxonomicName id="8AC642C7F71F2D2449FC5712FCE26911" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kalaw" status="sp. nov.">
<emphasis id="C00778387F2CBF39D7FC4DA10BE194E7" italics="true">Liphistius kalaw</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A4F3CB83A96D267F7E08FCA399391E05" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="73649E32C01D87E82FA8CAD4AD08DC60" bold="true">A, C</emphasis>
dorsal view
<emphasis id="84C92196A550ED904F9920C7C18EAD5D" bold="true">B, D</emphasis>
ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk;
<abbrev id="ABBRID0E1XAE" xlink_title="sclerotized poreplate">PPl</abbrev>
= poreplate;
<abbrev id="ABBRID0E5XAE" xlink_title="receptacular cluster">RC</abbrev>
= receptacular cluster; a = length of poreplate transverse; b = width of poreplate. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EHYAE" type="description">
<paragraph id="68379F02305BB81DCF08325BD74FD61D">
<heading id="A3F034C61FEE7332CFD3D31B3BE8268D" reason="title">Description.</heading>
</paragraph>
<paragraph id="BA3D7639B62701E883BE0F2E64179897">
<emphasis id="3EC9C52795447FF00FA8315352203D8E" bold="true">
Male
<typeStatus id="8D783900399CE8F712314C2C043C4FC3" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace yellowish brown, with few short, scattered bristles; opisthosoma yellowish brown, with 12 brown tergites, close to each other, 26
<superScript id="F42D3169641C538756673A519AD8C2D9">th</superScript>
larger than others, 5
<superScript id="2C2E14C13BFBD0AA9FA3D6E9A78D0DBE">th</superScript>
largest; chelicerae robust, promargin with 13 denticles of variable size; labium and sternum yellowish brown, with few short setae on anterior margin and many long setae on elongated posterior margin; legs yellowish brown, with strong setae and spines. Measurements:
<abbrev id="ABBRID0ETYAE" xlink_title="body length">BL</abbrev>
13.65,
<abbrev id="ABBRID0EXYAE" xlink_title="carapace length">CL</abbrev>
5.94,
<abbrev id="ABBRID0E2YAE" xlink_title="carapace width">CW</abbrev>
6.07,
<abbrev id="ABBRID0E6YAE" xlink_title="opisthosoma length">OL</abbrev>
8.00,
<abbrev id="ABBRID0EDZAE" xlink_title="opisthosoma width">OW</abbrev>
5.36; eye sizes and interdistances:
<abbrev id="ABBRID0EHZAE" xlink_title="anterior median eyes">AME</abbrev>
0.08,
<abbrev id="ABBRID0ELZAE" xlink_title="anterior lateral eyes">ALE</abbrev>
0.54,
<abbrev id="ABBRID0EPZAE" xlink_title="posterior median eyes">PME</abbrev>
0.27,
<abbrev id="ABBRID0ETZAE" xlink_title="posterior lateral eyes">PLE</abbrev>
0.43,
<abbrev id="ABBRID0EXZAE" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0E2ZAE" xlink_title="anterior median eyes">AME</abbrev>
0.11,
<abbrev id="ABBRID0E6ZAE" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0ED1AE" xlink_title="anterior lateral eyes">ALE</abbrev>
0.15,
<abbrev id="ABBRID0EH1AE" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0EL1AE" xlink_title="posterior median eyes">PME</abbrev>
0.08,
<abbrev id="ABBRID0EP1AE" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0ET1AE" xlink_title="posterior lateral eyes">PLE</abbrev>
0.10,
<abbrev id="ABBRID0EX1AE" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0E21AE" xlink_title="posterior lateral eyes">PLE</abbrev>
0.17,
<abbrev id="ABBRID0E61AE" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0ED2AE" xlink_title="anterior lateral eyes">ALE</abbrev>
0.12,
<abbrev id="ABBRID0EH2AE" xlink_title="posterior lateral eyes">PLE</abbrev>
<abbrev id="ABBRID0EL2AE" xlink_title="posterior lateral eyes">PLE</abbrev>
0.34,
<abbrev id="ABBRID0EP2AE" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0ET2AE" xlink_title="posterior median eyes">PME</abbrev>
0.10; labium 1.06 long and 0.98 wide; sternum 3.08 long and 0.96 wide; legs: I 18.36 (4.81, 2.57, 3.87, 4.60, 2.51), II 20.24 (5.64, 2.56, 4.23, 5.07, 2.74), III 20.46 (5.64, 2.37, 4.42, 6.00, 3.03), IV 27.56 (6.97, 2.84, 5.57, 8.17, 4.01).
</paragraph>
<paragraph id="151E46E3B3E43F5AD90E2A46648C2363">
Palp: tibial apophysis (
<abbrev id="ABBRID0EZ2AE" xlink_title="male palps bearing tibial apophysis">TiA</abbrev>
) with 4 tapering setae of similar length (Fig.
<figureCitation id="317D03CBDE0ADF7DB0952CADB441043F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 C</figureCitation>
); paracymbium (
<abbrev id="ABBRID0EB3AE" xlink_title="paracymbium">PC</abbrev>
) with many setae situated on tip (Fig.
<figureCitation id="0A14F7F7BB52FB1B5F72EDE1FBBBB6C7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 A C</figureCitation>
); cumulus (
<abbrev id="ABBRID0EJ3AE" xlink_title="cumulus">Cu</abbrev>
) slightly elevated, with 8 tapering spines (Fig.
<figureCitation id="F269D10666944891FD481D554480A434" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 A C</figureCitation>
); subtegulum with weakly developed apophysis (Fig.
<figureCitation id="0EC8FE3CAC6CAD203C1C88C77A1776AE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 B</figureCitation>
); contrategulum (
<abbrev id="ABBRID0EV3AE" xlink_title="contrategulum">CT</abbrev>
) with arched projection distally, and large triangular process in prolateral view (Fig.
<figureCitation id="FE99AB1F8A6F2CC4104B52ED87172A07" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 A, B, D</figureCitation>
); tegulum (
<collectionCode id="B515C6B28621EE251CE0FB31D4C9DF10" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
) fan-shaped with serrated edge and longitudinal ridges (Fig.
<figureCitation id="CA68FDA3C5CE175A6DABCF73FF8DEED7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 C, F</figureCitation>
); paraembolic plate (PeP) narrow, sclerotized (Fig.
<figureCitation id="3AE4579A0CAE3C377A8738494513BF1F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 A, F</figureCitation>
); embolus (E) with 4 distinct longitudinal ridges reaching tip in retrolateral view, and with several denticles along longitudinal ridges (Fig.
<figureCitation id="B98613C76790664835D1D7614D05BBF3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Liphistius kalaw sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure3" httpUri="https://binary.pensoft.net/fig/1115129">3 B F</figureCitation>
).
</paragraph>
<paragraph id="5322CF90A434395C0C62B7BAB5DCE00D">
<emphasis id="2C213BCDE17F12FD0DEDBEC2B2E5A626" bold="true">
Female
<typeStatus id="44BBB99C39D30E80741EAC3F16F2A807" type="paratype">paratype</typeStatus>
(XUX- 2019-062).
</emphasis>
Carapace yellowish brown, with few short, scattered bristles; opisthosoma brown, with 12 brown tergites, close to each other, with gray patches, 26
<superScript id="77F1A73F37BF694F336DD811FDC3DB07">th</superScript>
larger than others, 5
<superScript id="E05C2DA8482D7CB75BD0A3BBAA458E01">th</superScript>
largest; chelicerae robust, reddish brown; promargin of cheliceral groove with 12 denticles of variable size; labium yellowish brown, sternum with several setae; legs with strong setae and spines. Measurements:
<abbrev id="ABBRID0ER4AE" xlink_title="body length">BL</abbrev>
11.56,
<abbrev id="ABBRID0EV4AE" xlink_title="carapace length">CL</abbrev>
5.82,
<abbrev id="ABBRID0EZ4AE" xlink_title="carapace width">CW</abbrev>
5.22,
<abbrev id="ABBRID0E44AE" xlink_title="opisthosoma length">OL</abbrev>
6.23,
<abbrev id="ABBRID0EB5AE" xlink_title="opisthosoma width">OW</abbrev>
4.62; eye sizes and interdistances:
<abbrev id="ABBRID0EF5AE" xlink_title="anterior median eyes">AME</abbrev>
0.07,
<abbrev id="ABBRID0EJ5AE" xlink_title="anterior lateral eyes">ALE</abbrev>
0.56,
<abbrev id="ABBRID0EN5AE" xlink_title="posterior median eyes">PME</abbrev>
0.29,
<abbrev id="ABBRID0ER5AE" xlink_title="posterior lateral eyes">PLE</abbrev>
0.42,
<abbrev id="ABBRID0EV5AE" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EZ5AE" xlink_title="anterior median eyes">AME</abbrev>
0.09,
<abbrev id="ABBRID0E45AE" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EB6AE" xlink_title="anterior lateral eyes">ALE</abbrev>
0.15,
<abbrev id="ABBRID0EF6AE" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0EJ6AE" xlink_title="posterior median eyes">PME</abbrev>
0.06,
<abbrev id="ABBRID0EN6AE" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0ER6AE" xlink_title="posterior lateral eyes">PLE</abbrev>
0.09,
<abbrev id="ABBRID0EV6AE" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EZ6AE" xlink_title="posterior lateral eyes">PLE</abbrev>
0.11,
<abbrev id="ABBRID0E46AE" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0ECAAG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.07,
<abbrev id="ABBRID0EGAAG" xlink_title="posterior lateral eyes">PLE</abbrev>
<abbrev id="ABBRID0EKAAG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.35,
<abbrev id="ABBRID0EOAAG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0ESAAG" xlink_title="posterior median eyes">PME</abbrev>
0.08; labium 3.00 long and 0.61 wide; sternum 1.19 long and 1.13 wide; palp 10.25 (3.62, 1.74, 2.36, 2.53), legs: I 12.78 (4.29, 2.03, 2.63, 2.37, 1.46), II 12.89 (4.17, 2.03, 2.56, 2.59, 1.54), III 13.42 (4.13, 1.84, 2.75, 2.78, 1.92), IV 19.33 (5.57, 2.32, 3.81, 4.91, 2.72).
</paragraph>
<paragraph id="938FCCC7F0A38AC548919A1E45238441">
Vulva: poreplate (
<abbrev id="ABBRID0EYAAG" xlink_title="sclerotized poreplate">PPl</abbrev>
) slightly wider than long, with pair of large anterior lobes and pair of small anterolateral lobes; central dorsal opening (
<collectionCode id="9226ACD7B8675A600D5572086888F07B">CDO</collectionCode>
) circular, racemose receptacular cluster (
<abbrev id="ABBRID0E3AAG" xlink_title="receptacular cluster">
<collectionCode id="2E6977B2735A391E9D5475AEA0918F8D">RC</collectionCode>
</abbrev>
) slightly long; posterior stalk (
<collectionCode id="E619BC1CD17003EFD198FE8513DADDA2">PS</collectionCode>
) axe-shaped; lateral margins of genital atrium (GA) membranous (Fig.
<figureCitation id="CCCB716771F9FD14919F77421759A4BE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Vulva of Liphistius kalaw sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster; a = length of poreplate transverse; b = width of poreplate. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure4" httpUri="https://binary.pensoft.net/fig/1115130">4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEBAG" type="variation">
<paragraph id="03528DA065840EEB59FBA8989CF1FDB3">
<heading id="EF694DC53E75499C6FD51376A931A75F" reason="title">Variation.</heading>
</paragraph>
<paragraph id="9839E25EF1369BE62AA7BBFA54D98D70">
Females vary in body size. The range measurements of females (
<emphasis id="6667B023D4E697D68C852E0A46F4DD6C" italics="true">N</emphasis>
= 8):
<abbrev id="ABBRID0EMBAG" xlink_title="body length">BL</abbrev>
8.5516.11,
<abbrev id="ABBRID0EQBAG" xlink_title="carapace length">CL</abbrev>
4.487.20,
<abbrev id="ABBRID0EUBAG" xlink_title="carapace width">CW</abbrev>
3.906.43,
<abbrev id="ABBRID0EYBAG" xlink_title="opisthosoma length">OL</abbrev>
4.188.56,
<abbrev id="ABBRID0E3BAG" xlink_title="opisthosoma width">OW</abbrev>
3.007.36. The number of denticles on the promargin of cheliceral groove varies from 11 to 14 (
<emphasis id="30AF1ADADBE80742CA2BD35A78B5446B" italics="true">N</emphasis>
= 8).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECCAG" type="etymology">
<paragraph id="6706ECFB1287FA2076CEC9813261683A">
<heading id="10F64A3BA0A101433248B22E7D6A9C86" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="A6B9EF402EBCC0240F7D4DEB88733B89">
The species epithet
<taxonomicName id="2956E68DACCFDF0ED9FBB93A5CDE31B9" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kalaw">
<emphasis id="13D831DC53F6E23B2BE39E30D220197C" italics="true">kalaw</emphasis>
</taxonomicName>
” refers to the
<typeStatus id="8906A6F0378FA69FBCA31F8906F70424">type</typeStatus>
locality, Kalaw Township; it is treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERCAG" type="distribution">
<paragraph id="3804DC6EE60363984BD9D8FFACD9CD7C">
<heading id="6C11D500CA8CA99C378810AAD38737CE" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="B8C8A001B7EF17D62B3E9C15153461E4">
<collectingRegion id="75AB6F8757624AAF38FA42EDCD2E80F6" country="Myanmar" name="Mandalay">Mandalay Region</collectingRegion>
(Kalaw Township),
<collectingCountry id="81FECEB190C4DFBB79684CF80AF2D2A9" name="Myanmar">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>