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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1138.90580" ID-Pensoft-Pub="1313-2970-1138-89" ID-Pensoft-UUID="49C7E0FDF49955939BFFF3C31BC8326B" ID-ZooBank="AD364502AA604A3BA7A8CA3716D9B23F" ModsDocID="1313-2970-1138-89" checkinTime="1673085713316" checkinUser="pensoft" docAuthor="Novikov, Aleksandr, Sharafutdinova, Dayana &amp; Chertoprud, Elena" docDate="2023" docId="ADC42B81DD2652E38A4AB967ABA4BA5D" docLanguage="en" docName="ZooKeys 1138: 89-141" docOrigin="ZooKeys 1138" docPubDate="2023-01-05" docSource="http://dx.doi.org/10.3897/zookeys.1138.90580" docTitle="Bryocamptus (Bryocamptus) minutus subsp. Subgenus Chappuis 1929" docType="treatment" docVersion="1" id="49C7E0FDF49955939BFFF3C31BC8326B" lastPageNumber="89" masterDocId="49C7E0FDF49955939BFFF3C31BC8326B" masterDocTitle="Two new species of Bryocamptus (Copepoda, Harpacticoida, Canthocamptidae) from the Russian Arctic and comparison with Bryocamptus minutus (Claus, 1863)" masterLastPageNumber="141" masterPageNumber="89" pageNumber="89" updateTime="1673085713316" updateUser="pensoft">
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<mods:title>Two new species of Bryocamptus (Copepoda, Harpacticoida, Canthocamptidae) from the Russian Arctic and comparison with Bryocamptus minutus (Claus, 1863)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Novikov, Aleksandr</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2026-2510</mods:nameIdentifier>
<mods:affiliation>Kazan Federal University, Kremlyovskaya St. 18, 420008 Kazan, Russia</mods:affiliation>
<mods:nameIdentifier type="email">aleksandr-novikov-2011@list.ru</mods:nameIdentifier>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sharafutdinova, Dayana</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6533-5686</mods:nameIdentifier>
<mods:affiliation>Kazan Federal University, Kremlyovskaya St. 18, 420008 Kazan, Russia</mods:affiliation>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Chertoprud, Elena</mods:namePart>
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<mods:affiliation>Department of Hydrobiology, Biological Faculty, M. V. Lomonosov Moscow State University, Leninskie Gory, Moscow 119991, Russia &amp; A. N. Severtsov Institute of Ecology and Evolution, Leninsky Prospect, 33, Moscow 119071, Russia</mods:affiliation>
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<taxonomicName LSID="ADC42B81-DD26-52E3-8A4A-B967ABA4BA5D" authority="Bryocamptus Bryocamptus Chappuis, 1929" authorityName="Chappuis" authorityYear="1929" class="Hexanauplia" family="Canthocamptidae" genus="Bryocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryocamptus (Bryocamptus) subsp. Subgenus" order="Harpacticoida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="subGenus" subGenus="Bryocamptus">Subgenus Bryocamptus Bryocamptus Chappuis, 1929</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="89" type="remarks">
<paragraph pageId="0" pageNumber="89">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="89">
<taxonomicName authorityName="Novikov &amp; Sharafutdinova &amp; Chertoprud" authorityYear="2023" class="Hexanauplia" family="Canthocamptidae" genus="Bryocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryocamptus" order="Harpacticoida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="genus">Bryocamptus</taxonomicName>
is a very large genus with ~ 135 species and subspecies in four subgenera:
<taxonomicName authority="" family="Canthocamptidae" kingdom="Animalia" lsidName="B. (Arcticocamptus)" order="Siphonostomatoida" pageId="0" pageNumber="89" rank="subGenus" subGenus="Arcticocamptus">B. (Arcticocamptus)</taxonomicName>
Chappuis, 1929,
<taxonomicName family="Canthocamptidae" genus="B." kingdom="Animalia" lsidName="B." order="Siphonostomatoida" pageId="0" pageNumber="89" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="89">B.</emphasis>
</taxonomicName>
(
<emphasis italics="true" pageId="0" pageNumber="89">
<taxonomicName authorityName="Novikov &amp; Sharafutdinova &amp; Chertoprud" authorityYear="2023" class="Hexanauplia" family="Canthocamptidae" genus="Bryocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryocamptus" order="Harpacticoida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="genus">Bryocamptus</taxonomicName>
)
</emphasis>
Chappuis, 1928,
<taxonomicName authority="" family="Canthocamptidae" kingdom="Animalia" lsidName="B. (Echinocamptus)" order="Siphonostomatoida" pageId="0" pageNumber="89" rank="subGenus" subGenus="Echinocamptus">B. (Echinocamptus)</taxonomicName>
Chappuis, 1929 and
<taxonomicName authority="" family="Canthocamptidae" kingdom="Animalia" lsidName="B. (Rheocamptus)" order="Siphonostomatoida" pageId="0" pageNumber="89" rank="subGenus" subGenus="Rheocamptus">B. (Rheocamptus)</taxonomicName>
Borutzky, 1952. Additionally, two subgenera were earlier designated as not valid
<taxonomicName authority="" family="Canthocamptidae" kingdom="Animalia" lsidName="B. (Limocamptus)" order="Siphonostomatoida" pageId="0" pageNumber="89" rank="subGenus" subGenus="Limocamptus">B. (Limocamptus)</taxonomicName>
Chappuis, 1929 and
<taxonomicName authority="" family="Canthocamptidae" kingdom="Animalia" lsidName="B. (Pentacamptus)" order="Siphonostomatoida" pageId="0" pageNumber="89" rank="subGenus" subGenus="Pentacamptus">B. (Pentacamptus)</taxonomicName>
Wiley, 1934.
</paragraph>
<paragraph pageId="0" pageNumber="89">
In our opinion, this is one of the genera of the family most in need of revision. The first reason is that there are no clear diagnostic characters for the entire genus. Previously, this character was the two-segment exopod A2; however, this character is plesiomorphic for the entire family
<taxonomicName authorityName="Brady" authorityYear="1880" class="Hexanauplia" family="Canthocamptidae" kingdom="Animalia" lsidName="" order="Harpacticoida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="family">Canthocamptidae</taxonomicName>
, so it may be an adequate solution to separate at least part of the subgenera into separate genera. The second reason is the blurred line between
<taxonomicName authority="" family="Canthocamptidae" kingdom="Animalia" lsidName="B. (Bryocamptus)" order="Siphonostomatoida" pageId="0" pageNumber="89" rank="subGenus" subGenus="Bryocamptus">B. (Bryocamptus)</taxonomicName>
and
<taxonomicName authority="" family="Canthocamptidae" kingdom="Animalia" lsidName="B. (Rheocamptus)" order="Siphonostomatoida" pageId="0" pageNumber="89" rank="subGenus" subGenus="Rheocamptus">B. (Rheocamptus)</taxonomicName>
.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.46313" author="Borutzky, EV" journalOrPublisher="Fauna SSSR, Rakoobraznye [Fauna of the USSR, Crustacea]" pageId="0" pageNumber="89" pagination="1 - 424" refId="B5" refString="Borutzky, EV, 1952. Harpacticoida presnykh vod [Harpacticoida of fresh waters]. Fauna SSSR, Rakoobraznye [Fauna of the USSR, Crustacea] 3: 1 - 424, DOI: https://doi.org/10.5962/bhl.title.46313" title="Harpacticoida presnykh vod [Harpacticoida of fresh waters]." url="https://doi.org/10.5962/bhl.title.46313" volume="3" year="1952">Borutzky (1952)</bibRefCitation>
in the differences between these subgenera indicates the difference in segmentation of the endopods P1-P4, which again contrasts plesiomorphic and apomorphic characters. In our opinion, an essential part of the
<taxonomicName authority="" family="Canthocamptidae" kingdom="Animalia" lsidName="B. (Rheocamptus)" order="Siphonostomatoida" pageId="0" pageNumber="89" rank="subGenus" subGenus="Rheocamptus">B. (Rheocamptus)</taxonomicName>
species should in fact be transferred to the type subgenus.
</paragraph>
<paragraph pageId="0" pageNumber="89">
Unfortunately, at the moment we do not have enough data and material to revise the subgenera, so in this work we adhere to the classification given by
<bibRefCitation author="Dussart, B" journalOrPublisher="Brill, Leiden" pageId="0" pageNumber="89" refId="B10" refString="Dussart, B, Defaye, D, 1990. Repertoire mondial des Crustaces Copepodes des eaux interieures: III. Harpacticoides. E. J. Brill, Leiden" title="Repertoire mondial des Crustaces Copepodes des eaux interieures: III. Harpacticoides. E. J." year="1990">Dussart and Defaye (1990)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="FF0BD074C7788B30B38D3DD888446019" ID-CoL="7JLYR" LSID="ADC4D560-9E18-56FE-933D-C33D65120117" authority="Santschi, 1933" authorityName="Santschi" authorityYear="1933" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName="Crematogaster brunneipennis subsp. yorubosa" order="Hymenoptera" pageId="0" pageNumber="99555" phylum="Arthropoda" rank="subSpecies" species="brunneipennis" subSpecies="yorubosa">Crematogaster brunneipennis yorubosa Santschi, 1933</taxonomicName>
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<paragraph id="4FDCF1D52C88109DF42A85BE42B96719" pageId="0" pageNumber="99555">Notes</paragraph>
<paragraph id="92E17DEA14594326648262CA7484ACAC" pageId="0" pageNumber="99555">
(
<bibRefCitation id="5157922C58CE187BA1446BFC519EEA49" author="Santschi, F." journalOrPublisher="Bulletin et Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="99555" pagination="95 - 108" refId="B7740379" refString="Santschi, F., 1933. Contribution a l'etude des fourmis de l'Afrique tropicale. Bulletin et Annales de la Societe Entomologique de Belgique 73: 95 - 108" title="Contribution a l'etude des fourmis de l'Afrique tropicale" volume="73" year="1933">Santschi 1933</bibRefCitation>
)
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<paragraph id="4A4C87C4E7D65C9D06617D8F493E12AA" pageId="6" pageNumber="43">Bestimmung nach SELLNICK &amp; FORSSLUND (1955:486), Grandjean (1936:51)</paragraph>
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810-910
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,
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:Breite 1,80-2,2 (7 Ex.); mit dickem Cerotegument und darin eingelagerten Substratpartikeln bedeckt.
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:
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<collectingRegion id="DBE134F084B6F66582C706B3A0103FDA">Stadtwald Ettlingen</collectingRegion>
, Moderbuchenwald, Streu an
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,
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, 1 Ex.,
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;
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Moos von
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,
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leg., 3 Ex.,
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.
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<paragraph id="74B3907B3698192172FE98EE30460565" pageId="7" pageNumber="44">Diskussion</paragraph>
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Zur Darstellung bei Grandjean (1936) besteht ein deutlicher Unterschied: Der Hinterrand des Notogaster ist dort (Abb. 3A) nicht durch eine klare Kante zwischen den beiden k1-Haaren abgeschlossen; die Kanten laufen dort ziemlich weit rostrad und enden unverbunden. Im Gegensatz dazu ist bei unseren Tieren an dieser Stelle eine deutlich Kante ausgebildet, die in schwachem Bogen rostrad eingebogen ist. Die Gestalt
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daher in Dorsalansicht eher der Darstellung von
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(Thorell, 1872) bei Sellnick &amp; Forsslund (1955). Diese Art
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aber 2 Paar Notogasterhaare (pn2, f1) auf dem
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caudalen Notogasterfeld.
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