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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.570.6095" ID-PMC="PMC4829797" ID-Pensoft-Pub="1313-2970-570-1" ID-Pensoft-UUID="E376FF8EFFF1F22C326D1E0DFF8BFFDF" ID-PubMed="27103878" ID-Zenodo-Dep="579094" ID-ZooBank="CE0DA439F6F64DCF82255700A3C50098" ModsDocID="1313-2970-570-1" checkinTime="1457076378220" checkinUser="pensoft" docAuthor="Dean, David Allen" docDate="2016" docId="ADBB15E36775143A447A4BCD0F0F301D" docLanguage="en" docName="ZooKeys 570: 1-703" docOrigin="ZooKeys 570" docPubDate="2016-03-02" docSource="http://dx.doi.org/10.3897/zookeys.570.6095" docTitle="Anahita punctulata" docType="treatment" docVersion="7" id="E376FF8EFFF1F22C326D1E0DFF8BFFDF" lastPageNumber="80" masterDocId="E376FF8EFFF1F22C326D1E0DFF8BFFDF" masterDocTitle="Catalogue of Texas spiders" masterLastPageNumber="703" masterPageNumber="1" pageNumber="80" updateTime="1668162766282" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Catalogue of Texas spiders</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dean, David Allen</mods:namePart>
<mods:affiliation>Department of Entomology, Texas A &amp; M University, College Station, Texas, United States of America</mods:affiliation>
<mods:nameIdentifier type="email">a-dean-ento@tamu.edu</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2016</mods:date>
<mods:detail type="pubDate">
<mods:number>2016-03-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>570</mods:number>
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<mods:start>1</mods:start>
<mods:end>703</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.570.6095</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.570.6095</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-570-1</mods:identifier>
<mods:identifier type="ZooBank">CE0DA439F6F64DCF82255700A3C50098</mods:identifier>
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<treatment ID-GBIF-Taxon="127856549" LSID="urn:lsid:plazi:treatment:ADBB15E36775143A447A4BCD0F0F301D" httpUri="http://treatment.plazi.org/id/ADBB15E36775143A447A4BCD0F0F301D" lastPageNumber="80" pageId="79" pageNumber="80">
<subSubSection pageId="79" pageNumber="80" type="nomenclature">
<paragraph pageId="79" pageNumber="80">
<taxonomicName LSID="ADBB15E3-6775-143A-447A-4BCD0F0F301D" authority="(Hentz, 1844)" baseAuthorityName="Hentz" baseAuthorityYear="1844" class="Arachnida" family="Ctenidae" genus="Anahita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anahita punctulata" order="Araneae" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="punctulata">Anahita punctulata (Hentz, 1844)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="79" pageNumber="80" type="reference_group">
<paragraph pageId="79" pageNumber="80">
<taxonomicName baseAuthorityName="Hentz" baseAuthorityYear="1844" class="Arachnida" family="Ctenidae" genus="Anahita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anahita punctulata" order="Araneae" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="punctulata">Anahita punctulata</taxonomicName>
<bibRefCitation author="Jackman, JA" pageId="451" pageNumber="452" refId="B378" refString="Jackman, JA, 1997. A field guide to spiders and scorpions of Texas. Texas Monthly Field Guide Series, Gulf Publishing Company, Houston, Texas, 201 pp." title="A field guide to spiders and scorpions of Texas. Texas Monthly Field Guide Series, Gulf Publishing Company, Houston, Texas, 201 pp." year="1997">Jackman 1997</bibRefCitation>
: 162;
<bibRefCitation author="Peck, WB" pageId="461" pageNumber="462" refId="B558" refString="Peck, WB, 1981. The Ctenidae of temperate zone North America. Bulletin of the American Museum of Natural History 170: 157-169." title="The Ctenidae of temperate zone North America. Bulletin of the American Museum of Natural History 170: 157 - 169." year="1981">Peck 1981</bibRefCitation>
: 158 [T], mf, desc. (figs 1-4);
<bibRefCitation author="Roth, VD" pageId="467" pageNumber="468" refId="B685" refString="Roth, VD, 1982. Handbook for spider identification. Published by the author, Portal, Arizona, 62 pp." title="Handbook for spider identification. Published by the author, Portal, Arizona, 62 pp." year="1982">Roth 1982</bibRefCitation>
: 14-1;
<bibRefCitation author="Roth, VD" pageId="467" pageNumber="468" refId="B686" refString="Roth, VD, 1985. Spider genera of North America. American Arachnological Society, Gainesville, Florida, 171 pp." title="Spider genera of North America. American Arachnological Society, Gainesville, Florida, 171 pp." year="1985">Roth 1985</bibRefCitation>
: B-9-1;
<bibRefCitation author="Roth, VD" pageId="468" pageNumber="469" refId="B688" refString="Roth, VD, 1994. Spider genera of North America, with keys to families and genera, and a guide to literature, third edition, American Arachnological Society, Gainesville, Florida, 203 pp." title="Spider genera of North America, with keys to families and genera, and a guide to literature, third edition, American Arachnological Society, Gainesville, Florida, 203 pp." year="1994">Roth 1994</bibRefCitation>
: 86;
<bibRefCitation author="Sissom, WD" pageId="469" pageNumber="470" refId="B708" refString="Sissom, WD, Peck, WB, Cokendolpher, JC, 1999. New records of wandering spiders from Texas, with a description of the male of Ctenus valverdiensis (Araneae: Ctenidae). Entomological News 110: 260-266." title="New records of wandering spiders from Texas, with a description of the male of Ctenus valverdiensis (Araneae: Ctenidae). Entomological News 110: 260 - 266." year="1999">Sissom et al. 1999</bibRefCitation>
: 260, mf, desc.;
<bibRefCitation author="Ubick, D" pageId="470" pageNumber="471" refId="B739" refString="Ubick, D, Davilla, DS, 2005. Ctenidae. In: Ubick D, Paquin P, Cushing PE, Roth V (Eds) Spiders of North America: an identification manual. American Arachnological Society, 83." title="Ctenidae. In: Ubick D, Paquin P, Cushing PE, Roth V (Eds) Spiders of North America: an identification manual. American Arachnological Society, 83." year="2005">
Ubick and
<normalizedToken originalValue="Dávilla">Davilla</normalizedToken>
2005
</bibRefCitation>
: 84
</paragraph>
<paragraph pageId="79" pageNumber="80">
<taxonomicName class="Arachnida" family="Ctenidae" genus="Ctenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ctenus punctulatus" order="Araneae" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="punctulatus">Ctenus punctulatus</taxonomicName>
Hentz, 1844;
<bibRefCitation author="Marx, G" pageId="459" pageNumber="460" publicationUrl="10.5479/si.00963801.782.497" refId="B506" refString="Marx, G, 1890. Catalogue of the described Araneae of temperate North America. Proceedings of the United States National Museum 12: 497-594. doi: 10.5479/si.00963801.782.497" title="Catalogue of the described Araneae of temperate North America. Proceedings of the United States National Museum 12: 497 - 594. doi:" url="10.5479/si.00963801.782.497" year="1890">Marx 1890</bibRefCitation>
: 567
</paragraph>
</subSubSection>
<subSubSection pageId="79" pageNumber="80" type="distribution">
<paragraph pageId="79" pageNumber="80">Distribution.</paragraph>
<paragraph pageId="79" pageNumber="80">Harris (Houston), Tyler</paragraph>
</subSubSection>
<subSubSection pageId="79" pageNumber="80" type="locality">
<paragraph pageId="79" pageNumber="80">Locality.</paragraph>
<paragraph pageId="79" pageNumber="80">Kirby State Forest</paragraph>
</subSubSection>
<subSubSection pageId="79" pageNumber="80" type="time of activity">
<paragraph pageId="79" pageNumber="80">Time of activity.</paragraph>
<paragraph pageId="79" pageNumber="80">Male (April 27-May 8)</paragraph>
</subSubSection>
<subSubSection pageId="79" pageNumber="80" type="method">
<paragraph pageId="79" pageNumber="80">Method.</paragraph>
<paragraph pageId="79" pageNumber="80">Flight intercept trap on ground [m]</paragraph>
</subSubSection>
<subSubSection pageId="79" pageNumber="80" type="type">
<paragraph pageId="79" pageNumber="80">Type.</paragraph>
<paragraph pageId="79" pageNumber="80">Alabama</paragraph>
</subSubSection>
<subSubSection pageId="79" pageNumber="80" type="etymology">
<paragraph pageId="79" pageNumber="80">Etymology.</paragraph>
<paragraph pageId="79" pageNumber="80">Latin, minute white dots on abdomen</paragraph>
</subSubSection>
<subSubSection pageId="79" pageNumber="80" type="collection">
<paragraph pageId="79" pageNumber="80">Collection.</paragraph>
<paragraph pageId="79" pageNumber="80">TAMU</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="243C569932C1D428BB68CBE65215CD8E">A new species of the genus Nothobranchius (Cyprinodontiformes: Nothobranchiidae) from the coastal area of northeastern Mozambique.</mods:title>
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<mods:part id="AFF5B6B3C718B538F4A882469A42E73F">
<mods:date id="C8E52AB99C4BCADC5AE5D6E060C3E009">2007</mods:date>
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</mods:location>
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<treatment id="ADBBBAC1FE88AE5EB1F999723C507E22" ID-DOI="http://doi.org/10.5281/zenodo.6248554" ID-GBIF-Taxon="100123204" ID-Zenodo-Dep="6248554" LSID="urn:lsid:plazi:treatment:ADBBBAC1FE88AE5EB1F999723C507E22" httpUri="http://treatment.plazi.org/id/ADBBBAC1FE88AE5EB1F999723C507E22" lastPageNumber="66" pageNumber="62">
<subSubSection id="CB22FD2B3885C81D0BE79EA962A49E9E" type="nomenclature">
<paragraph id="F0DD1ADF2C0351520A97D4643BD6C6D5" pageNumber="62">
<taxonomicName id="3BAFED8893B55EF17655A7E1CC5B70D7" ID-CoL="47SLZ" authorityName="Valdesalici" authorityYear="2007" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="hengstleri" status="new species">Nothobranchius hengstleri</taxonomicName>
,
<taxonomicNameLabel id="A9A622C4D25FE13D26373C3E6A720606" rank="species">new species</taxonomicNameLabel>
</paragraph>
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<paragraph id="17C0BA8BD98302CA2024AF8E56213AD6" pageNumber="62">(Figs. 1-4)</paragraph>
</subSubSection>
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<paragraph id="FA85C032BAA0A648D91AAB8BAF3E0726" pageNumber="62">
<taxonomicName id="C3DBB2B801DD694156D2824302DECCAA" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="62" phylum="Chordata" rank="species" species="nassoro">Nothobranchius species “Nassoro”</taxonomicName>
MZHL-2005-14, Hengstler &amp; Valdesalici (2006)
</paragraph>
</subSubSection>
<subSubSection id="8CFAD0192968BD9CDD3AFBA2EB179205" type="nomenclature">
<paragraph id="32E4D297728DCEFB813D6E4023080951" pageNumber="62">
<materialsCitation id="06D3644DE0EF242B5DBE6971304C1757" ID-GBIF-Occurrence="923890570" country="Mozambique" stateProvince="Cabo Delgado">
<typeStatus id="FA73077A786810092828509A67799F35">Holotype</typeStatus>
.
<collectionCode id="66B7FB8C739228210ACF9853EBE0A13F" collectionName="Germany, Muenchen [= Munich], Zoologische Staatssammlung">ZSM</collectionCode>
34483 (male, 41.3 mm SL);
<collectingCountry id="829F9A2C63065A6A1EA03DF453FBBA7E">Mozambique</collectingCountry>
:
<collectingRegion id="BB8B33A1F40579768FAC787A1DA1976F">Cabo Delgado</collectingRegion>
: about 5 km north of Nassoro village, temporary pools, not connected to any permanent river system,
<geoCoordinate id="AB0AFF30217B34476F259A1E13D076B3" direction="south" orientation="latitude" value="-10.887033">10°53.222’S</geoCoordinate>
,
<geoCoordinate id="603D1B2ADDCBE50E3765C425F3C6ED2A" direction="east" orientation="longitude" value="40.368233">40°22.094’E</geoCoordinate>
, Holger Hengstler and Silverio Vendo,
<date id="09CDF1AE1275DBECAF1461D8CE6F3088" day="18" month="05" value="2005-05-18" year="2005">18 May 2005</date>
. Collected with large hand nets and preserved after seven months of aquarium maintenance.
</materialsCitation>
</paragraph>
<paragraph id="58E035ECB27001DA350114BAA68192F5" pageNumber="62">
<materialsCitation id="4D7C9728128F0A97185F9FB7E3296D5E" ID-GBIF-Occurrence="923890577" country="Mozambique" stateProvince="Cabo Delgado">
<typeStatus id="DA3FF776DD113603D93F53E5F0DCED91">Paratypes</typeStatus>
.
<collectionCode id="01F6E6CDCBF712F209D80CBB3A62144F" collectionName="Germany, Muenchen [= Munich], Zoologische Staatssammlung">ZSM</collectionCode>
34484 (female, 40.5 mm SL), same collection and preservation data as for holotype (upper part of caudal fin removed for DNA extraction).
</materialsCitation>
<materialsCitation id="0F8ED4448284E39BDE57A9EE49DF863F" ID-GBIF-Occurrence="923890574" country="Mozambique" stateProvince="Cabo Delgado">
<collectionCode id="5D287435E4F46E8107C1C263A3A6C1EA" collectionName="Germany, Muenchen [= Munich], Zoologische Staatssammlung">ZSM</collectionCode>
34485 (female, 33.1 mm SL), 34486 (male, 30.9 mm SL), same collection data as for holotype (preserved in field).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="2DD9DA192CFE427D5CC78D8C9A11F03D" type="diagnosis">
<paragraph id="85F1378F535326708A6E42C091070704" pageNumber="62">
Diagnosis.
<taxonomicName id="922FBE6FFC5F7A5FB8CF5548C69523C4" authorityName="Valdesalici" authorityYear="2007" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="hengstleri">Nothobranchius hengstleri</taxonomicName>
is distinguished from
<taxonomicName id="5FADABA0C0D362C6CA701A4FCA1D0E5A" baseAuthorityName="Pfeffer" baseAuthorityYear="1896" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="melanospilus">N. melanospilus</taxonomicName>
and
<taxonomicName id="A1AC317082290D0A3B4981E6E70B35D3" baseAuthorityName="Pfeffer" baseAuthorityYear="1896" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="melanospilus">N. cf. melanospilus</taxonomicName>
on the basis of both meristic and morphometric data, male and female coloration, and by a more convex dorsal profile. Male
<taxonomicName id="C7FF0ADD5876783E7C26BFD240719BEC" authorityName="Valdesalici" authorityYear="2007" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="hengstleri">N. hengstleri</taxonomicName>
differs from male
<taxonomicName id="EC8C9FC07B997449CF599452F1CD967B" baseAuthorityName="Pfeffer" baseAuthorityYear="1896" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="melanospilus">N. melanospilus</taxonomicName>
in having a greater body depth (30.4-31.8 vs. 29.3-30.3 % SL), a longer head (33.3-33.4 vs. 26.8-28.3 % SL), a longer prepelvic length (49.6-50.1 vs. 45.6-48.9 % SL); males and females have fewer scales in longitudinal series (25-26 vs. 31-32).
</paragraph>
<paragraph id="E9B707F36E3CE3FFA641BDB10D897264" pageNumber="63">
Male
<taxonomicName id="B545AA179DE9979A8571C3A08170FB8A" authorityName="Valdesalici" authorityYear="2007" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="hengstleri">N. hengstleri</taxonomicName>
differs from male
<taxonomicName id="6F04400F5E45E853D6C77D9A7604598A" baseAuthorityName="Pfeffer" baseAuthorityYear="1896" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="melanospilus">N. cf. melanospilus</taxonomicName>
in having a greater body depth (30.5-31.8 vs. 25.6-28.2 % SL), a longer head (33.3-33.5 vs. 29.5-32.6 % SL); males and females have fewer scales in the longitudinal series (25-26 vs. 29-30).
</paragraph>
<paragraph id="E39A9F1E31F9F0D2F0DE0B628C78EC5B" pageNumber="63">
Males differ from males of the other two species in having a deeper red coloration over the entire body and head, a spotted anal fin (well defined brown spots over entire fin vs. few spots limited at base), a different caudal-fin pattern (deep red, with a well defined complete black margin vs. red, without any particular marking or with thin rudimentary black margin sometimes reduced on upper and lower angle), and a different caudal fin shape (perfectly rounded vs. rounded to subtruncate). Female
<taxonomicName id="CE3B0BF58084B049CDFDF3CD6929D3D5" authorityName="Valdesalici" authorityYear="2007" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="hengstleri">N. hengstleri</taxonomicName>
differ from female
<taxonomicName id="E9CDF3978D1AB3614C80494437E75CE3" baseAuthorityName="Pfeffer" baseAuthorityYear="1896" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="melanospilus">N. melanospilus</taxonomicName>
by the spots on body and fins (absent vs. present) and by the caudal fin shape indicated immediately before. They also differ from female
<taxonomicName id="E6D81AD80FAD26C1977E13F69C3930F3" baseAuthorityName="Pfeffer" baseAuthorityYear="1896" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="melanospilus">N. cf. melanospilus</taxonomicName>
by the spots on body and fins (absent vs. present but rudimentary in analyzed populations; absent in other populations) (Seegers, 1986; 1997; 2003; Rosenstock, 2003).
</paragraph>
<paragraph id="B9ED466A9D3F3CCC88562EA9DC534BB5" pageNumber="63">
Among other closely related species, male
<taxonomicName id="1A5BD0EB42CEC9B7AAB2C0A10EE65A10" authorityName="Valdesalici" authorityYear="2007" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="hengstleri">N. hengstleri</taxonomicName>
differs from male
<taxonomicName id="47A2AD82568D69532794716F9EDE7CCD" authorityName="Ahl" authorityYear="1924" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="vosseleri">N. vosseleri</taxonomicName>
in having a longer head (33.3-33.4 vs. 25.1-31.5 %SL), anal fin pattern (spots forming approximately three or four arch-like stripes in middle part of fin vs. small spots extending on fin rays and irregular stripes), male
<taxonomicName id="61955586C27062BFF3A2D2FAF918B95A" authorityName="Ahl" authorityYear="1924" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="vosseleri">N. vosseleri</taxonomicName>
from “Korogwe” area by dorsal and anal fins coloration (yellow-green vs. yellow-grey to pale blue), males and females with fewer scales in longitudinal series (25-26 vs. 25-31), and fewer anal fin rays (14-16 vs.15-18). Male
<taxonomicName id="073518709ECC4A09CB235FD027BB6FB8" authorityName="Valdesalici" authorityYear="2007" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="hengstleri">N. hengstleri</taxonomicName>
differs form male
<taxonomicName id="3E2FADF48D1F061D9B656E0805795E5D" authorityName="Wildekamp &amp; Berkenkamp" authorityYear="1979" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="interruptus">N. interruptus</taxonomicName>
in having a deeper red colorations over the entire body and head, caudal fin coloration (deep red with complete black margin vs. pale to clear red with irregular blue-grey to almost colorless border), dorsal fin coloration (yellow-green, with red to brown spots vs. bluegrey to olive with dark grey spots), anal fin coloration (yellow-green with red/brown spots vs. olive-yellow to pale blue with some grey spots near fin base), males and females with fewer scales in longitudinal series (25- 26 vs. 27-32), and caudal fin shape (perfectly rounded vs. subtruncate).
</paragraph>
<paragraph id="2DE0810EFD896922B9E8211023B24B12" pageNumber="63">
Male
<taxonomicName id="73A2198FC2F21B7919C0331302ADD125" authorityName="Valdesalici" authorityYear="2007" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="hengstleri">N. hengstleri</taxonomicName>
differs from male
<taxonomicName id="05DEBF212DB737DB0393188923803141" authorityName="Wildekamp &amp; Berkenkamp" authorityYear="1979" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="jubbi">N. jubbi</taxonomicName>
in having a deeper red colorations over the entire body and head, dorsal fin coloration (yellowish with red/brown spots vs. blue-grey to olive with red brown spots), anal fin coloration (yellow-green, with red-brown spots vs. olive-yellow to pale blue with some grey spots near fin base), caudal fin coloration (deep red with complete black margin vs. blue-grey with a pattern of red- brown spots and a white margin in blue morph and clear red followed by a wide blue-grey band and white border in red morph), female
<taxonomicName id="D72828782FB07EFA874EE204B707762E" authorityName="Valdesalici" authorityYear="2007" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="hengstleri">N. hengstleri</taxonomicName>
differs from female
<taxonomicName id="6978FBE675D1182F566A23E8476CF7A2" authorityName="Wildekamp &amp; Berkenkamp" authorityYear="1979" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="jubbi">N. jubbi</taxonomicName>
in body and unpaired fins pattern (not marked vs.dark border on body scales and small dark gray marking at base of fins), males and females with fewer scales in longitudinal series (25-26 vs. 29-32), fewer dorsal (13-16 vs.15-20) and anal fin rays (14-16 vs.15- 19), caudal fin shape (perfectly rounded vs. subtruncate).
</paragraph>
<paragraph id="46B7F87E1BAD5F41AD69C0F9070680D6" pageNumber="63">
Male
<taxonomicName id="0342A3CF669C7D30E0758B3F43072082" authorityName="Valdesalici" authorityYear="2007" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="hengstleri">N. hengstleri</taxonomicName>
differs from male
<taxonomicName id="5D312740AA0B09BB370D7682DE8782D1" authorityName="Wildekamp" authorityYear="1982" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="elongatus">N. elongatus</taxonomicName>
in having a deeper red colorations over the entire body and head, anal fin coloration (yellow-green, with red/brown spots vs. yellow-grey with few red spots near fin base); males and females have fewer scales in longitudinal series (25-26 vs.30-32), fewer anal fin rays (14- 16 vs.16-18), and caudal fin shape (perfectly rounded vs. subtruncate).
</paragraph>
</subSubSection>
<subSubSection id="2942E074C7FA23254D84AA7F6A4089E8" type="description">
<paragraph id="E4473DA9D7E1CDA7D60D7C8B33F4FF9F" pageNumber="63">Description. Morphometric data appear in Table 1.</paragraph>
<paragraph id="C58C2D9BEC44C0B417A723FD04A112E4" pageNumber="63">
A robust, moderately deep-bodied
<taxonomicName id="8D10E0277AA2D73862AEAB0A99F04A96" authorityName="Peters" authorityYear="1868" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="genus">Nothobranchius</taxonomicName>
with pointed snout, terminal mouth directed upward. Dorsal fin rays 13-16; anal fin rays 14-16. Longitudinal series of scales 25-26 + 2-4 on caudal fin base. Cephalic squamation pattern variable, with one specimen (ZSM 34486) showing G-type. Anterior neuromast series on head ‘open’ type; central series in two shallow grooves, lined with low lobes; posterior cephalic neuromast series with curved shallow pits on each side. Preopercular neuromast system in open groove, distal ridge slightly overlapping opercle. One neuromast on each scale of mid-lateral line.
</paragraph>
<paragraph id="9873401376E70917E7B56AC45350D504" lastPageNumber="64" pageNumber="63">Males. Body profile convex on dorsal and ventral margins, slightly concave on caudal peduncle posterior to dorsal and anal fins. Greatest body depth observed at level of pelvic fins base. Dorsal and anal fins rounded, their tips with rudimentary filamentous rays. Caudal fin perfectly rounded. Membranes of pelvic fins fusing near the base, tips reaching urogenital papilla. Dorsal and anal fins with small papillate contact organs. Opercular membrane projecting from opercle, distal edge slightly wrinkled.</paragraph>
<paragraph id="67AAAADD4B67570601F96AAC362F13A8" pageNumber="65">Females. Body less compressed and deep compared to males. Dorsal fin rounded; anal fin triangular, with rounded tip; caudal fin perfecly rounded. Anal fin positioned more posteriorly compared to males. Membranes of pelvic fins fusing near the base, tips reaching urogenital papilla. Opercular membrane not projecting from opercle.</paragraph>
<paragraph id="61A0C4892322A6458CBE8A52F5B699D7" pageNumber="65">Maximum observed size of males 41.3 mm SL. Maximum observed size of females 40.5 mm SL.</paragraph>
<paragraph id="1608047FBA20E4B085F070D30A1036DA" pageNumber="65">Color in life. Male (Fig. 3). Scales on body and head light blue, with distinct red margins, resulting in a reticulated pattern. Scale margins in ventral area orange. Snout, throat, frontal and upper/dorsal part of head deep red. Scales on middle part of caudal peduncle and on caudal fin base red. Dorsal fin yellowish-green, with pattern of red/brown spots forming arch-like stripes and narrow lines; spots on distal part of fin smaller and more dense, with white to light blue border and with thin submarginal blue band. Anal fin yellow-green, with pattern of red/brown spots forming approximately three or four arch-like stripes in middle part of fin; spots near base distally fused in narrow lines extending onto fin rays; fin with white to light blue border. Caudal fin deep red, with complete black margin. Pelvic fins light yellow-green, with some brown spots parallel to fin rays, light blue border and brown submarginal band. Pectoral fins hyaline, with light blue margin. Iris golden, with light blue iridescences and faint dark vertical bar.</paragraph>
<paragraph id="D2C2720D4334C8209E48B336E4CF37F6" pageNumber="65">Female (Fig. 4). Body pale olive-brown, darker grey-brown dorsally; light brown to almost silvery-white ventrally. Flank scales with light blue iridescence. Unpaired and paired fins hyaline. Iris golden, with a faint dark vertical bar.</paragraph>
<paragraph id="76BFFA5B24AE5EB58822C7E0AF3AA4EA" pageNumber="65">Color in alcohol. Male (Fig.1). Body scales light brown to whitish, almost all scales with distinct dark red margin. Dorsal fin light brown with a pattern dark brown spots. Anal fin light brown. Caudal fin light brown with dark red lines exending on fin rays. Pelvic and pectoral fins light brown. Iris bluish.</paragraph>
<paragraph id="021868AE918A92B8F61212EA7F98CE78" pageNumber="65">Female (Fig. 2). Body light brown to whitish, scales with a narrow brown margin. Opercular and ventral area yellowish to light orange. Unpaired and paired fins pale light brown to whitish. Iris blueish.</paragraph>
</subSubSection>
<subSubSection id="985A3E060411AF1C9E27E844AA62DC0C" type="distribution">
<paragraph id="E9FEE50DDEE7317ED6E2A0AD1476456E" lastPageNumber="66" pageNumber="65">
Distribution and habitat. (Figs. 5-6).
<taxonomicName id="58ECB37D4A6B4C4C1AE55F70B013CCA3" authorityName="Valdesalici" authorityYear="2007" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="hengstleri">N. hengstleri</taxonomicName>
is only known from the type locality, a temporary pool in a small coastal plateau, about 10 km long by 20 km wide, altitude 67 m, in northeastern Mozambique, approximately 5 km north Nassoro village, Cabo Delgado province. The species probably has a wider range, but this can only be ascertained through additional collecting.
</paragraph>
</subSubSection>
<subSubSection id="7574C8FD1D8F0F7E404D5F5B67A1935C" type="biology_ecology">
<paragraph id="0F37CF83B886C84E90D1F78681536DC7" pageNumber="66">The very small pool was at the time of collection about 6 m long, 1.5 m wide and shallow (about 0.10 m deep). No aquatic vegetation was present, with only dry grass along the shore. The water was brown and turbid and the pH 7.5. No other fish species were present.</paragraph>
</subSubSection>
<subSubSection id="99A9791B496EB4A1EF80944AB68D47B5" type="etymology">
<paragraph id="4E7A1935CCBEC7734944E206CE20E123" pageNumber="66">Etymology. The new species is named in honor of the collector, Mr. Holger Hengstler of Munich, Germany.</paragraph>
</subSubSection>
<subSubSection id="FC848F68AD45C7DB8D0F7321B855DA53" type="discussion">
<paragraph id="0FB6ADAAFE69AE22C8BE84D3F3C58E0D" pageNumber="66">Discussion</paragraph>
<paragraph id="A831E2E204D5B67087B6BEB61B253FF3" pageNumber="66">
Additional and modified synapomorphies important in defining the
<taxonomicName id="40D95696EE4497CEB1C695E283671E11" baseAuthorityName="Pfeffer" baseAuthorityYear="1896" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="melanospilus">N. melanospilus</taxonomicName>
species group are: maximum observed body size (over 30 mm SL), a relatively elongate to moderately deep body (less than 35% of SL), variable cephalic squamation, pointed triangular head, dorsal profile gently convex from snout to end of dorsal-fin base, male coloration consisting of uniform red reticulation over blue scales and a dorsal fin with white to light blue rudimentary filamentous rays. The last character was not mentioned in the redescription of
<taxonomicName id="10DE6AF6AB37564B6592CB95E6FC4028" authorityName="Ahl" authorityYear="1924" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="vosseleri">N. vosseleri</taxonomicName>
(Wildekamp et al, 1998), but is present in some specimens; see illustration in Seegers (1997: 82).
</paragraph>
<paragraph id="A4CCDE41A934675ACB45A05D5A53272E" pageNumber="66">
<taxonomicName id="9E5DBFDA330B13B9ECB14B1708D18A78" authorityName="Valdesalici" authorityYear="2007" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="hengstleri">Nothobranchius hengstleri</taxonomicName>
is a member of this group, presenting all of the diagnostic features of this complex. Aspects of the colour pattern and morphology suggest that
<taxonomicName id="22B2FD98B9AABE8D050BA25F327D0D23" authorityName="Valdesalici" authorityYear="2007" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="hengstleri">N. hengstleri</taxonomicName>
is more closely related to
<taxonomicName id="4BCE24B2A19E6E45325DE09A9673334A" authorityName="Ahl" authorityYear="1924" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="vosseleri">N. vosseleri</taxonomicName>
than to the others members of the group.
</paragraph>
<paragraph id="608C0B611E360ADB31843F904E959B08" pageNumber="66">The distribution of the species group is almost entirely restricted to the area from southern Somalia to Mozambique (north of the Zambezi River), a pattern congruent to the paleo-ichthyofaunal boundaries proposed by Stewart (2001).</paragraph>
</subSubSection>
</treatment>
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Linnaeus, 1758
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<paragraph id="A5ED1117CB5603E1B44CE71E57687CCB" pageId="0" pageNumber="8050">England, Scotland, Wales, Ireland, Isle of Man</paragraph>
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<mods:date id="63494FC2E58B81D2D9757A889282177C">2019</mods:date>
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<treatment id="ADBBDB3ED0BF41D954C9ACC42BF0D9FD" ID-GBIF-Taxon="156200493" LSID="urn:lsid:plazi:treatment:ADBBDB3ED0BF41D954C9ACC42BF0D9FD" httpUri="http://treatment.plazi.org/id/ADBBDB3ED0BF41D954C9ACC42BF0D9FD" lastPageNumber="1" pageId="0" pageNumber="1">
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<paragraph id="D069293581E8ABCB9C0BC86F4F4D16F7" pageId="0" pageNumber="1">
<taxonomicName id="87ECA6AADEAE8A906FC6645E6B19ECEC" ID-CoL="66FPZ" authority="Pfeiffer, 1861" authorityName="Pfeiffer" authorityYear="1861" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphidromus flavus" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="flavus">Amphidromus flavus (Pfeiffer, 1861)</taxonomicName>
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<taxonomicName id="7CF5DFD71526D2D6DBA15330E729956D" class="Gastropoda" family="Camaenidae" genus="Bulimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimus flavus" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="flavus">Bulimus flavus</taxonomicName>
Pfeiffer, 1861a: 194. Type locality: Siam [Thailand].
<bibRefCitation id="324119353B67C5C254A8176E506E1451" pageId="0" pageNumber="1">Pfeiffer 1861b</bibRefCitation>
: 171, 172, pl. 46, figs 7, 8.
</paragraph>
<paragraph id="D951666FF34B177878ECCB90A3976F75" pageId="0" pageNumber="1">
<taxonomicName id="317816BE92C6F888E6B201EFAF2CE853" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphidromus (Syndromus) flavus" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="flavus" subGenus="Syndromus">Amphidromus (Syndromus) flavus</taxonomicName>
:
<bibRefCitation id="7F81EF29BF90E6286B116BE42C85A225" author="Laidlaw, FF" journalOrPublisher="Fieldiana: Zoology" pageId="0" pageNumber="1" pagination="507 - 677" title="The land snail genus Amphidromus: A synoptic catalogue." volume="41" year="1961">Laidlaw and Solem 1961</bibRefCitation>
: 563, 564, 619.
<bibRefCitation id="9D70F11D02659D2B6425832594D9076B" author="Inkhavilay, K" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="1" pagination="1 - 40" title="Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species." url="https://doi.org/10.5852/ejt.2017.330" volume="330" year="2017">Inkhavilay et al. 2017</bibRefCitation>
: 24, figs 9b, 10
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, 11
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, 12
<normalizedToken id="3EF68001AAF9A2312F944CC5831DB7CD" originalValue="a–c">a-c</normalizedToken>
.
</paragraph>
<paragraph id="7D62E4C2650CB7F65A1D32A80BDCBC9E" pageId="0" pageNumber="1">
<taxonomicName id="2328CDE7D57F6F20344B6E8EBEB7FAF8" class="Gastropoda" family="Camaenidae" genus="Syndromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syndromus flavus" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="flavus">Syndromus flavus</taxonomicName>
:
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: 51.
</paragraph>
<paragraph id="912A5F96A0E66666A7CD98F727F948EA" pageId="0" pageNumber="1">
<taxonomicName id="1B767B7EF0E819C988C0F282461FB32A" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphidromus flavus" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="flavus">Amphidromus flavus</taxonomicName>
:
<bibRefCitation id="C1D798E83EAD685812DECB8974A684E1" author="Sutcharit, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="49 - 105" title="Illustrated type catalogue of Amphidromus Albers, 1850 in Natural History Museum, London, and descriptions of two new species." url="https://doi.org/10.3897/zookeys.492.8641" volume="492" year="2015">Sutcharit et al. 2015</bibRefCitation>
: 70, fig. 7i, j.
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<paragraph id="D3ACCF04234C2A8F4627352E53ED6582" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="036C8507C8138800F65DD2759DA76F74" pageId="0" pageNumber="1">
Lectotype NHMUK 19601436 and paralectotype NHMUK 19601437 (1 shell) figured in
<bibRefCitation id="F536D358C7CEC1C319ABFA11404723D9" author="Sutcharit, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="49 - 105" title="Illustrated type catalogue of Amphidromus Albers, 1850 in Natural History Museum, London, and descriptions of two new species." url="https://doi.org/10.3897/zookeys.492.8641" volume="492" year="2015">Sutcharit et al. (2015</bibRefCitation>
: fig. 7i, j). Specimens CUMZ 7029 from Tam Pou Kham, Vangvieng District, Vientiane (Fig. 42E).
</paragraph>
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<paragraph id="8854366563EEDC23D8B72AFC3A200AAC" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="9BDA0026F62653080E3D06A6CDF332FF" pageId="0" pageNumber="1">
Laos, Malaysia, Thailand and Vietnam (
<bibRefCitation id="8B420E0F183969C8729EED92B824E52B" author="Laidlaw, FF" journalOrPublisher="Fieldiana: Zoology" pageId="0" pageNumber="1" pagination="507 - 677" title="The land snail genus Amphidromus: A synoptic catalogue." volume="41" year="1961">Laidlaw and Solem 1961</bibRefCitation>
,
<bibRefCitation id="0D4608C159252D37CD2CC608DB789E5A" author="Inkhavilay, K" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="1" pagination="1 - 40" title="Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species." url="https://doi.org/10.5852/ejt.2017.330" volume="330" year="2017">Inkhavilay et al. 2017</bibRefCitation>
).
</paragraph>
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<paragraph id="35538554EDFEB7DFB1440F7D497A0797" pageId="0" pageNumber="1">
Figure 42. A, B
<taxonomicName id="B228E62521183E181C9C0486F58656C7" class="Gastropoda" family="Camaenidae" genus="Bradybaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bradybaena jourdyi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jourdyi">Bradybaena jourdyi</taxonomicName>
A syntype MNHN-IM-2000-1944 and B CUMZ collection C
<taxonomicName id="8986F58415273B2F06CF8C3754404769" class="Gastropoda" family="Camaenidae" genus="Bradybaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bradybaena similaris" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="similaris">Bradybaena similaris</taxonomicName>
, syntype MNHN-IM-2000-31776 D
<taxonomicName id="2BF9C9D9FA59AE4BA013A985900946DD" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphidromus areolatus" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="areolatus">Amphidromus areolatus</taxonomicName>
, specimen CUMZ 7022 E
<taxonomicName id="52644CC69E2FE2502E4CD95B13AAEF1F" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphidromus flavus" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="flavus">Amphidromus flavus</taxonomicName>
, specimen CUMZ 7029 F
<taxonomicName id="BA1EDFC0F793E19CA85F560586E54D0F" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphidromus fuscolabris" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fuscolabris">Amphidromus fuscolabris</taxonomicName>
, holotype of
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MNHN-IM-2000-33216.
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