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<document id="562D9E6A9E03B9A05AD723F2D8666A69" ENCODING="UTF-8" ID-CLB-Dataset="21556" ID-DOI="10.5281/zenodo.3931989" ID-GBIF-Dataset="751112f9-dacb-4aa7-b6dd-94b531bd95b7" ID-Zenodo-Dep="3931989" checkinTime="1594063508482" checkinUser="admin" docAuthor="Linnaeus, Carolus" docDate="1753" docId="AD40001C44CE4E3AB468B37711C950D4" docLanguage="en" docName="Linnaeus1753Treatments.xml" docOrigin="Stockholm: Laurentius Salvius" docSource="https://doi.org/10.5962/bhl.title.669" docTitle="Papaver hybridum Linnaeus, 1753, spec. nov." docType="treatment" docVersion="6" lastPageNumber="506" masterDocId="B60E6E2598A5F0FB2905D223E4672698" masterDocTitle="Species plantarum: exhibentes plantas rite cognitas, ad genera relatas, cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas" pageNumber="506" updateTime="1710908005168" updateUser="ExternalLinkService">
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<mods:title id="382E42BD78D47D148DBEE27794F9D8E5">Species plantarum: exhibentes plantas rite cognitas, ad genera relatas, cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas</mods:title>
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<mods:namePart id="F10CD2B2FF9FF1C30A4C0A127B1E579A">Linnaeus, Carolus</mods:namePart>
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<mods:dateIssued id="AD78B3EF729C0BCE5AB4E306FB669AD3">1753</mods:dateIssued>
<mods:publisher id="5D3087EA6C277427A5A0C31755204E80">Laurentius Salvius</mods:publisher>
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<mods:identifier id="458A2D7338331834400E634EFBCF3C03" type="DOI">10.5281/zenodo.3931989</mods:identifier>
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<treatment id="AD40001C44CE4E3AB468B37711C950D4" ID-DOI="http://doi.org/10.5281/zenodo.4356826" ID-GBIF-Taxon="164948274" ID-Zenodo-Dep="4356826" LSID="urn:lsid:plazi:treatment:AD40001C44CE4E3AB468B37711C950D4" httpUri="http://treatment.plazi.org/id/AD40001C44CE4E3AB468B37711C950D4" lastPageNumber="506" pageNumber="506">
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<taxonomicName id="272536F055936F27A327C8FC75F34F10" ID-CoL="4CJCN" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Papaveraceae" genus="Papaver" kingdom="Plantae" order="Ranunculales" phylum="Tracheophyta" rank="species" species="hybridum" status="spec. nov.">Papaver hybridum</taxonomicName>
,
<taxonomicNameLabel id="ACDEE0FFF813D8BF1D40CD1D37BBDC4A" rank="species">spec. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="2375F9B57067213C1C8B3CE054D752CE" type="description">
<paragraph id="AEFEDCD44C9735B8610DCBE17DAC4454" pageNumber="506">1. Papaver capsulis subglobosis torosis hispidis, caule folioso multifloro.</paragraph>
<paragraph id="3327EEC239AD2472237CD818FE40C754" pageNumber="506">
Papaver foliis ternato-pinnatifidis, capsulis sulcatis hispidis.
<emphasis id="D29D27A831C9DAE65DD372919A28EB34" italics="true">Hort. cliff. 201.</emphasis>
</paragraph>
<paragraph id="603F1C4B3CD3C6DE829C044EAA2578C4" pageNumber="506">
Papaver caule ramoso, foliis lineari-pinnatifidis, capsulis sulcatis hispidis.
<emphasis id="107122C070DF37225346B500A78CBC78" italics="true">Hort. ups. 136.</emphasis>
</paragraph>
<paragraph id="7F001A4BAF9E83AE7031F694E5F2B6C7" pageNumber="506">
Argemone capitulo breviore.
<emphasis id="6C6D2F05365858992F4F3DCC17DC5696" italics="true">Bauh. pin. 172.</emphasis>
</paragraph>
<paragraph id="6B92AB2C765EFA16407EFD655F4C509D" pageNumber="506">
Argemone capitulo hirsuto rotundo torulis canaliculato.
<emphasis id="16E4892CD3EFFB46A8F96E7F1771BE31" italics="true">Moris. hist. 2. p.278. s.3. t.14. f.9.</emphasis>
<emphasis id="4ADB02BDD9E8ACA6A9888E3B38E5ECDA" italics="true">Lob. ic. 276.</emphasis>
</paragraph>
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<subSubSection id="5143CABA28D97A6D3547FDD53627AF98" type="distribution">
<paragraph id="DF6AF87AE8498E1CB9391FB1724D11DA" pageNumber="506">
<emphasis id="477EECD7C93C4A0516EAE1080016836D" italics="true">Habitat in</emphasis>
Europa
<emphasis id="4394A22EB94CA0A7DEEA1D462BCF73ED" italics="true">australiore. ☉</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.700.11784" ID-GBIF-Dataset="30d1c76a-41e1-4f66-9fb5-460bf381f752" ID-PMC="PMC5711039" ID-Pensoft-Pub="1313-2970-700-1" ID-PubMed="29358897" ID-ZBK="EBA4322720AD4CFFA04E8D2542DDA3D6" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-700-1" ModsDocOrigin="ZooKeys 700" ModsDocTitle="Revision of the ant genus Melophorus (Hymenoptera, Formicidae)" checkinTime="1506009303549" checkinUser="pensoft" docAuthor="Heterick, Brian E., Castalanelli, Mark &amp; Shattuck, Steve O." docDate="2017" docId="AD4049EB3159485B570FB7FBDF36FA48" docLanguage="en" docName="ZooKeys 700: 1-420" docOrigin="ZooKeys 700" docSource="http://dx.doi.org/10.3897/zookeys.700.11784" docTitle="Melophorus longiceps Heterick, Castalanelli &amp; Shattuck, sp. n." docType="treatment" docUuid="9F45E0BF-07F1-4804-B90D-1389936F295B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="177" masterDocId="FF93E60CA711CB7BFF95D521FFFDFF9A" masterDocTitle="Revision of the ant genus Melophorus (Hymenoptera, Formicidae)" masterLastPageNumber="420" masterPageNumber="1" pageNumber="175" updateTime="1668164819442" updateUser="ExternalLinkService">
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<mods:title>Revision of the ant genus Melophorus (Hymenoptera, Formicidae)</mods:title>
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<mods:namePart>Heterick, Brian E.</mods:namePart>
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<mods:namePart>Castalanelli, Mark</mods:namePart>
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<mods:namePart>Shattuck, Steve O.</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2017</mods:date>
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<mods:number>700</mods:number>
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<subSubSection pageId="174" pageNumber="175" type="nomenclature">
<paragraph pageId="174" pageNumber="175">
<taxonomicName LSID="http://zoobank.org/9F45E0BF-07F1-4804-B90D-1389936F295B" authority="Heterick, Castalanelli &amp; Shattuck" class="Insecta" family="Formicidae" genus="Melophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melophorus longiceps" order="Hymenoptera" pageId="174" pageNumber="175" phylum="Arthropoda" rank="species" species="longiceps">Melophorus longiceps Heterick, Castalanelli &amp; Shattuck</taxonomicName>
<taxonomicNameLabel pageId="174" pageNumber="175">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="174" pageNumber="175" type="types">
<paragraph pageId="174" pageNumber="175">Types.</paragraph>
<paragraph pageId="174" pageNumber="175">
Holotype minor worker from CSIRO Lake Mere field stn near Louth, New South Wales, paddock B, January 1995,
<taxonomicName genus="M." lsidName="M." pageId="174" pageNumber="175" rank="genus">M.</taxonomicName>
Bryannah, paddock D (EXC) Interc[ept],
<taxonomicName class="Insecta" family="Formicidae" genus="Melophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melophorus" order="Hymenoptera" pageId="174" pageNumber="175" phylum="Arthropoda" rank="genus">Melophorus</taxonomicName>
sp. K [ANIC32-900104] (ANIC). Paratypes: 2 minor workers from CSIRO Lake Mere field stn near Louth, New South Wales, paddock B, January 1995,
<taxonomicName genus="M." lsidName="M." pageId="174" pageNumber="175" rank="genus">M.</taxonomicName>
Bryannah, paddock E Runoff, '
<taxonomicName class="Insecta" family="Formicidae" genus="Melophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melophorus" order="Hymenoptera" pageId="174" pageNumber="175" phylum="Arthropoda" rank="genus">Melophorus</taxonomicName>
sp.
<normalizedToken originalValue="K">K'</normalizedToken>
[ANIC32-066665] (BMNH); major worker from CSIRO Lake Mere field stn near Louth, New South Wales, paddock B, January 1995,
<taxonomicName genus="M." lsidName="M." pageId="174" pageNumber="175" rank="genus">M.</taxonomicName>
Bryannah, paddock G Runoff, '
<taxonomicName class="Insecta" family="Formicidae" genus="Melophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melophorus" order="Hymenoptera" pageId="174" pageNumber="175" phylum="Arthropoda" rank="genus">Melophorus</taxonomicName>
sp.
<normalizedToken originalValue="K">K'</normalizedToken>
[ANIC32-066665] (MCZ).
</paragraph>
</subSubSection>
<subSubSection pageId="174" pageNumber="175" type="diagnosis">
<paragraph pageId="174" pageNumber="175">Diagnosis.</paragraph>
<paragraph pageId="174" pageNumber="175">
<taxonomicName class="Insecta" family="Formicidae" genus="Melophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melophorus longiceps" order="Hymenoptera" pageId="174" pageNumber="175" phylum="Arthropoda" rank="species" species="longiceps">Melophorus longiceps</taxonomicName>
can be placed in the
<taxonomicName lsidName="M. biroi" pageId="174" pageNumber="175" rank="species" species="biroi">M. biroi</taxonomicName>
species-group on the basis of characters of the clypeus, propodeum, mandible and palps. The species is placed in the
<taxonomicName lsidName="M. biroi" pageId="174" pageNumber="175" rank="species" species="biroi">M. biroi</taxonomicName>
species-complex on the basis of a further suite of characters (viz, metatibia of major worker with only one preapical spur [except rarely in the
<taxonomicName lsidName="mjobergi" pageId="174" pageNumber="175" rank="species" species="mjobergi">mjobergi</taxonomicName>
clade]; clypeal psammophore placed anteriorly at or just above anterior margin of clypeus in the minor worker and often in the major worker; head dorsoventrally compressed to varying degrees in the minor worker of most species with the eyes placed high on the sides; compact legs, and small body size [excluding
<taxonomicName lsidName="mjobergi" pageId="174" pageNumber="175" rank="species" species="mjobergi">mjobergi</taxonomicName>
clade] HW of smallest minor 0.36 mm, average HW of smallest minors 0.46 mm; HW of largest known major 1.29 mm, average HW of largest majors [where known] 1.05 mm). In profile, the head of the minor worker is distinctly dorsoventrally flattened and, in full-face view, extremely narrow (CI ≤ 75). This extreme narrowness is a unique character that defines this species. Some
<taxonomicName lsidName="M. biroi" pageId="174" pageNumber="175" rank="species" species="biroi">M. biroi</taxonomicName>
have rather narrow heads, but in those cases the CI is&gt; 85. The major worker is unknown.
</paragraph>
</subSubSection>
<subSubSection lastPageId="175" lastPageNumber="176" pageId="174" pageNumber="175" type="description">
<paragraph pageId="174" pageNumber="175">Minor worker description.</paragraph>
<paragraph lastPageId="175" lastPageNumber="176" pageId="174" pageNumber="175">
Head. Head rectangular; posterior margin of head planar or weakly concave; frons matt or with weak sheen, shagreenate; frons consisting exclusively or almost exclusively of well-spaced, appressed setae only (small, erect setae, if present, usually confined to ocular triangle or posterior margin of head). Eye moderate (eye length 0.20-0.49 length of side of head capsule); in full-face view, eyes set above midpoint of head capsule; in profile, eye set anteriad of midline of head capsule; eyes elliptical or slightly reniform. In full-face view, frontal carinae distinctly concave; frontal lobes straight in front of antennal insertion. Anteromedial clypeal margin broadly and evenly convex; clypeal psammophore set at or just above anterior clypeal margin; palp formula 6,4. Five mandibular teeth in minor worker; mandibles triangular, weakly incurved; third mandibular tooth distinctly shorter than apical tooth and teeth numbers two and four; masticatory margin of mandibles approximately vertical or weakly
<pageBreakToken pageId="175" pageNumber="176" start="start">oblique</pageBreakToken>
. Mesosoma. Integument of pronotum, mesonotum and mesopleuron with weak to moderate sheen, shagreenate on pronotum and dorsum of mesonotum, otherwise microreticulate; anterior mesosoma in profile weakly elevated anteriad, thereafter gently sinuate, pronotum and mesonotum on same plane; erect pronotal setae absent; in profile, metanotal groove shallow, indicated mainly by an angle; propodeum matt or with a weak sheen and microreticulate; propodeum angulate, propodeal angle blunt, or distinctly angulate, propodeal angle sharp; length ratio of propodeal dorsum to its declivity about 1:1; erect propodeal setae always absent; appressed propodeal setulae short, separated by more than own length and inconspicuous; propodeal spiracle situated on or beside declivitous face of propodeum, and shorter (length &lt;0.50
<normalizedToken originalValue="×">x</normalizedToken>
height of propodeum). Petiole. In profile, petiolar node squamiform; in full-face view, shape of petiolar node uniformly rounded; node shining and distinctly microreticulate. Gaster. Gaster shining, shagreenate ('LP
<normalizedToken originalValue="record">record'</normalizedToken>
appearance); pilosity of first gastral tergite consisting of well-spaced short, inconspicuous, appressed setae only, erect setae always absent. General characters. Colour uniformly chocolate with some tan highlights.
</paragraph>
</subSubSection>
<subSubSection pageId="175" pageNumber="176" type="description">
<paragraph pageId="175" pageNumber="176">Measurements.</paragraph>
<paragraph pageId="175" pageNumber="176">Worker (n = 2): CI 76-81; EI 30-38; EL 0.15-0.16; HL 0.56-0.63; HW 0.43-0.51; ML 0.72-0.82; MTL 0.41-0.43; PpH 0.07-0.10; PpL 0.29-0.34; SI 111-121; SL 0.52-0.56.</paragraph>
</subSubSection>
<subSubSection pageId="176" pageNumber="177" type="comments">
<paragraph pageId="176" pageNumber="177">
<pageBreakToken pageId="176" pageNumber="177" start="start">Comments</pageBreakToken>
.
</paragraph>
<paragraph pageId="176" pageNumber="177">
Currently,
<taxonomicName class="Insecta" family="Formicidae" genus="Melophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melophorus longiceps" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="longiceps">Melophorus longiceps</taxonomicName>
, known only from the minor worker, has been recorded from the Lake Mere Field Station, near Louth, NSW. However, workers morphologically similar to this species were seen in the TERC Collection but time did not permit a detailed examination. This material came from WA (Ningbing Ra) and NT (Kalkarindji, Humbert River and Pigeon Hole Stn) and included a major worker. The different biogeography, however, precludes a too ready association of the two sets of taxa. The elongate, flattened head of this species separates it from the rest of the genus, but in other respects it is identical to
<taxonomicName lsidName="M. biroi" pageId="176" pageNumber="177" rank="species" species="biroi">M. biroi</taxonomicName>
(in which the head capsule varies in length). A worker was collected from a paddock, but there are no other ecological data, and no specimens were available for sequencing.
</paragraph>
</subSubSection>
<subSubSection pageId="176" pageNumber="177" type="etymology">
<paragraph pageId="176" pageNumber="177">Etymology.</paragraph>
<paragraph pageId="176" pageNumber="177">
Latin longus (
<normalizedToken originalValue="long">'long'</normalizedToken>
) plus -ceps (
<normalizedToken originalValue="-headed">'-headed'</normalizedToken>
[from caput]); adjective in the nominative singular.
</paragraph>
<caption pageId="176" pageNumber="177">
<paragraph pageId="176" pageNumber="177">
Figure 40.
<taxonomicName class="Insecta" family="Formicidae" genus="Melophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melophorus longiceps" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="longiceps">Melophorus longiceps</taxonomicName>
sp. n.: minor worker holotype (ANIC32-900104) frons (a), profile (b) and dorsum (c); distribution map for the species (d). Low resolution scale bars: 0.5 mm (b, c); 0.2 mm (a).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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Yamana, Ogawa &amp; Ohtsuka
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(
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,
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<emphasis id="179DA0414132FFD7FF02FBE1FE40FBD0" bold="true" box="[151,386,1051,1077]" pageId="5" pageNumber="6">Material examined.</emphasis>
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<figureCitation id="BDD260D64132FFD7FF02FBC5FF15FBBC" box="[151,215,1087,1113]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
A-a).
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<typeStatus id="FA52C2F14132FFD7FE83FBBAFE4AFBBC" box="[278,392,1088,1113]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
</emphasis>
: WMNH-INV-2015-2497 (ST-
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2015, anesthetized, length
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, width
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,
<figureCitation id="BDD260D64132FFD7FAB5FBC5FAA2FBBF" box="[1312,1376,1087,1114]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
A-b); WMNH-INV-2015-2503 (ST-
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2015, posterior half only, length
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, width
<quantity id="E211D1B64132FFD7FBD9FB99FB50FB9B" box="[1100,1170,1123,1150]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="mm" value="3.0">3 mm</quantity>
,
<figureCitation id="BDD260D64132FFD7FB34FB99FB27FB98" box="[1185,1253,1123,1149]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
A-c);
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E- 12099-1 (ST-
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2015, length
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, width
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,
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accession number
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, 514 bps,
<figureCitation id="BDD260D64132FFD7FB61FB7DFAF7FB47" box="[1268,1333,1159,1186]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
A-d)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD84132FFD7FF52FB51FD29FAB0" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="25567C534132FFD7FF52FB51FD29FAB0" blockId="5.[151,1437,1051,2014]" pageId="5" pageNumber="6">
<emphasis id="179DA0414132FFD7FF52FB51FE80FB20" bold="true" box="[199,322,1195,1221]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
Twenty tentacles arranged in double circles (10 + 10), comprising five pairs of large tentacles in the outer circle and five pairs of small tentacles in the inner circle. Calcareous ring elongate, tubular, with both radial and interradial plates, fragmented into a mosaic of small pieces and prolonged to the posterior, with posterior prolongations distally linked to others forming a loop. Bodywall lacking ossicles. Pedicels only with circular endplate, without supporting plates and rods.
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD84132FFD5FF52FAA5FD8AF8F0" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="description">
<paragraph id="25567C534132FFD7FF52FAA5FE59F97C" blockId="5.[151,1437,1051,2014]" pageId="5" pageNumber="6">
<emphasis id="179DA0414132FFD7FF52FAA5FE9BFA9C" bold="true" box="[199,345,1375,1401]" pageId="5" pageNumber="6">Description.</emphasis>
Body small, up to
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, fusiform, curved, with both ends tapered and turned slightly upwards (
<figureCitation id="BDD260D64132FFD7FF35FA79FF38FA78" box="[160,250,1411,1437]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="5" pageNumber="6">Fig. 2A</figureCitation>
); bodywall thick, soft. Living body color yellowish-brown with dark brown/gray pigmentation, preserved color paler than living color. Twenty tentacles arranged in double circles (10 + 10), comprising five pairs of large tentacles in the outer circle and five pairs of small tentacles in the inner circle. Tentacle bases and first bifurcations dark brown; tentacle trunks and tips transparent. One row of thin, short pharyngeal villi surrounding the oral opening. Oral disk brown. Retractable pedicels covering entire body at constant density, but forming two longitudinal rows along each radius in introvert region. Color of pedicels same or slightly paler than adjacent body. Pedicels becoming gradually smaller anterior and posterior from middle portion of body. Where they are denser. Ten anal papillae and five anal teeth on radii.
</paragraph>
<paragraph id="25567C534132FFD7FF52F959FE8DF8A8" blockId="5.[151,1437,1051,2014]" pageId="5" pageNumber="6">
Calcareous ring elongate, tubular, with both radial and interradial plates fragmented into a mosaic of small pieces and prolonged posteriorly, with the posterior prolongations distally linked to prolongations of neighboring plates (
<figureCitation id="BDD260D64132FFD7FF35F911FF38F8E0" box="[160,250,1771,1797]" captionStart="FIGURE 3" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[152,1426,183,1471]" captionTargetId="figure-18@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 3. Elements of calcareous rings, viewed from the body cavity, anterior upper.A, Massinium toyoshiomaruae sp. nov., (holotype, WMNH-INV-2014-186); B, Thyone liaoi sp. nov., (holotype, WMNH-INV-2014-182, and a paratype, WMNH-INV- 2014-184); C, T. kyushuensis sp. nov., (holotype, WMNH-INV-2015-2437); D, T. toyoshiomaruae sp. nov. (holotype, WMNH- INV-2015-2483). Abbreviations: md, mid-dorsal part; mv, mid-ventral part; -ir, interradial element; -r radial element." figureDoi="http://doi.org/10.5281/zenodo.5225378" httpUri="https://zenodo.org/record/5225378/files/figure.png" pageId="5" pageNumber="6">Fig. 3A</figureCitation>
). Polian vesicles paired, both in medio-ventral position, stone canal single, in mediodorsal. Well-matured gonad situated in mid-body, in two clumps, one on each side of the dorsal mesentery, most tubules branched. Gonad lacking ossicles.
</paragraph>
<paragraph id="25567C534132FFD4FF52F8ADFAB0FE6C" blockId="5.[151,1437,1051,2014]" lastBlockId="6.[151,1437,151,393]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Bodywall lacking ossicles. Tentacle ossicles comprising thin rods in tentacle branches, variously sized, somewhat curved, most with small numbers of distal perforations (
<figureCitation id="BDD260D64132FFD7FC05F881FC28F870" box="[912,1002,1915,1941]" captionStart="FIGURE 4" captionStartId="8.[151,250,1916,1940]" captionTargetBox="[157,1434,384,1893]" captionTargetId="figure-114@8.[151,1436,381,1893]" captionTargetPageId="8" captionText="FIGURE 4. Scanning electron microscope images of ossicles of Massinium toyoshiomaruae sp. nov. (holotype, AE, WMNH- INV-2014-186). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225380" httpUri="https://zenodo.org/record/5225380/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
,
<tableCitation id="686B49E84132FFD7FC63F881FB8EF870" box="[1014,1100,1915,1941]" captionStart="TABLE 3" captionStartId="9.[151,239,480,506]" captionText="TABLE 3. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Massinium toyoshiomaruae sp. nov." pageId="5" pageNumber="6">Table 3</tableCitation>
). Tentacle bases and adjacent area of peri-oral and introvert skin with regular tables with oval discs possessing four large central and numerous small peripheral perforations, and two-pillared low spires mostly with one crossbeam and 26 apical spines (
<figureCitation id="BDD260D64132FFD4FAD5F839FF12FF54" captionStart="FIGURE 4" captionStartId="8.[151,250,1916,1940]" captionTargetBox="[157,1434,384,1893]" captionTargetId="figure-114@8.[151,1436,381,1893]" captionTargetPageId="8" captionText="FIGURE 4. Scanning electron microscope images of ossicles of Massinium toyoshiomaruae sp. nov. (holotype, AE, WMNH- INV-2014-186). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225380" httpUri="https://zenodo.org/record/5225380/files/figure.png" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">Fig. 4A, B, D</figureCitation>
,
<tableCitation id="686B49E84131FFD4FF48FF6DFEF1FF54" box="[221,307,151,177]" captionStart="TABLE 3" captionStartId="9.[151,239,480,506]" captionText="TABLE 3. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Massinium toyoshiomaruae sp. nov." pageId="6" pageNumber="7">Table 3</tableCitation>
). Peri-oral skin ossicles comprising mostly thin-framed, irregular-shaped rods/rosettes, also regular tables in parts adjoining tentacle bases, disc shape of tables in tentacle and peri-oral skin resemble each other, however, mostly have spires higher than those of tentacles and with 23 crossbeams, (
<figureCitation id="BDD260D64131FFD4FBC1FF25FB68FF1C" box="[1108,1194,223,249]" captionStart="FIGURE 4" captionStartId="8.[151,250,1916,1940]" captionTargetBox="[157,1434,384,1893]" captionTargetId="figure-114@8.[151,1436,381,1893]" captionTargetPageId="8" captionText="FIGURE 4. Scanning electron microscope images of ossicles of Massinium toyoshiomaruae sp. nov. (holotype, AE, WMNH- INV-2014-186). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225380" httpUri="https://zenodo.org/record/5225380/files/figure.png" pageId="6" pageNumber="7">Fig. 4B</figureCitation>
,
<tableCitation id="686B49E84131FFD4FB20FF25FAC8FF1C" box="[1205,1290,223,249]" captionStart="TABLE 3" captionStartId="9.[151,239,480,506]" captionText="TABLE 3. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Massinium toyoshiomaruae sp. nov." pageId="6" pageNumber="7">Table 3</tableCitation>
). Pharyngeal villi have many thin-framed rosettes (
<figureCitation id="BDD260D64131FFD4FDA2FEF9FD4FFEF8" box="[567,653,259,285]" captionStart="FIGURE 4" captionStartId="8.[151,250,1916,1940]" captionTargetBox="[157,1434,384,1893]" captionTargetId="figure-114@8.[151,1436,381,1893]" captionTargetPageId="8" captionText="FIGURE 4. Scanning electron microscope images of ossicles of Massinium toyoshiomaruae sp. nov. (holotype, AE, WMNH- INV-2014-186). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225380" httpUri="https://zenodo.org/record/5225380/files/figure.png" pageId="6" pageNumber="7">Fig. 4C</figureCitation>
,
<tableCitation id="686B49E84131FFD4FD02FEF9FD29FEF8" box="[663,747,259,285]" captionStart="TABLE 3" captionStartId="9.[151,239,480,506]" captionText="TABLE 3. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Massinium toyoshiomaruae sp. nov." pageId="6" pageNumber="7">Table 3</tableCitation>
). Introvert possessing tables with the same disc shape of those in tentacles and peri-oral area, however, most have low spires with one or no crossbeams (
<figureCitation id="BDD260D64131FFD4FB36FEDDFB3CFEA4" box="[1187,1278,295,321]" captionStart="FIGURE 4" captionStartId="8.[151,250,1916,1940]" captionTargetBox="[157,1434,384,1893]" captionTargetId="figure-114@8.[151,1436,381,1893]" captionTargetPageId="8" captionText="FIGURE 4. Scanning electron microscope images of ossicles of Massinium toyoshiomaruae sp. nov. (holotype, AE, WMNH- INV-2014-186). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225380" httpUri="https://zenodo.org/record/5225380/files/figure.png" pageId="6" pageNumber="7">Fig. 4D</figureCitation>
,
<tableCitation id="686B49E84131FFD4FA9EFEDDFAA3FEA4" box="[1291,1377,295,321]" captionStart="TABLE 3" captionStartId="9.[151,239,480,506]" captionText="TABLE 3. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Massinium toyoshiomaruae sp. nov." pageId="6" pageNumber="7">Table 3</tableCitation>
). We have several
<typeStatus id="FA52C2F14131FFD4FEBDFEB1FE51FE80" box="[296,403,331,357]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
including smaller individuals, however, we cannot find the evidence supporting a hypothesis the specimens lose/modify their bodywall deposits with age: any individual with nothing of bodywall deposits.
</paragraph>
<caption id="71962CDB4131FFD4FF02F91CFB04F857" ID-DOI="http://doi.org/10.5281/zenodo.5225376" ID-Zenodo-Dep="5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,250,1766,1790]" targetBox="[155,1435,419,1743]" targetPageId="6">
<paragraph id="25567C534131FFD4FF02F91CFB04F857" blockId="6.[151,1437,1766,1970]" pageId="6" pageNumber="7">
<emphasis id="179DA0414131FFD4FF02F91CFED6F91B" bold="true" box="[151,276,1766,1790]" pageId="6" pageNumber="7">FIGURE 2.</emphasis>
Lateral views of the specimens, dorsal upper, anterior left. A,
<taxonomicName id="E2E907D04131FFD4FC0CF91DFB71F91B" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[921,1203,1766,1790]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="7" phylum="Echinodermata" rank="species" species="toyoshiomaruae" status="sp. nov.">
<emphasis id="179DA0414131FFD4FC0CF91DFB71F91B" box="[921,1203,1766,1790]" italics="true" pageId="6" pageNumber="7">Massinium toyoshiomaruae</emphasis>
</taxonomicName>
<emphasis id="179DA0414131FFD4FB2EF91CFAD1F91B" bold="true" box="[1211,1299,1766,1790]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="0CAE1D3A4131FFD4FB2EF91CFACDF91B" box="[1211,1295,1766,1790]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
(a)
<emphasis id="179DA0414131FFD4FAAAF91CFA54F91B" box="[1343,1430,1766,1790]" italics="true" pageId="6" pageNumber="7">holotype</emphasis>
, WMNH-INV-2014-186, (b)
<emphasis id="179DA0414131FFD4FE2FF8F1FDD6F8C7" box="[442,532,1803,1826]" italics="true" pageId="6" pageNumber="7">paratype</emphasis>
, WMNH-INV-2015-2497, (c)
<emphasis id="179DA0414131FFD4FCC5F8F1FC68F8C7" box="[848,938,1803,1826]" italics="true" pageId="6" pageNumber="7">paratype</emphasis>
, WMNH-INV-2015-2503, (d)
<emphasis id="179DA0414131FFD4FB7DF8F1FA80F8C7" box="[1256,1346,1803,1826]" italics="true" pageId="6" pageNumber="7">paratype</emphasis>
, NSMT- E-12099-1; B,
<taxonomicName id="E2E907D04131FFD4FEBAF8D4FE6CF8A3" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[303,430,1838,1862]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="7" phylum="Echinodermata" rank="species" species="liaoi" status="sp. nov.">
<emphasis id="179DA0414131FFD4FEBAF8D4FE6CF8A3" box="[303,430,1838,1862]" italics="true" pageId="6" pageNumber="7">Thyone liaoi</emphasis>
</taxonomicName>
<emphasis id="179DA0414131FFD4FE20F8D4FDC7F8A3" bold="true" box="[437,517,1838,1862]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="0CAE1D3A4131FFD4FE20F8D4FDC7F8A3" box="[437,517,1838,1862]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (e)
<emphasis id="179DA0414131FFD4FDA0F8D4FD4EF8A3" box="[565,652,1838,1862]" italics="true" pageId="6" pageNumber="7">holotype</emphasis>
, WMNH-INV-2014-182, (f)
<emphasis id="179DA0414131FFD4FC20F8D5FBCDF8A3" box="[949,1039,1839,1862]" italics="true" pageId="6" pageNumber="7">paratype</emphasis>
, WMNH-INV-2014-183, (g)
<emphasis id="179DA0414131FFD4FAA9F8D5FA54F8A3" box="[1340,1430,1839,1862]" italics="true" pageId="6" pageNumber="7">paratype</emphasis>
, WMNH-INV-2014-184, (h)
<emphasis id="179DA0414131FFD4FE2BF8A9FDDAF88F" box="[446,536,1875,1898]" italics="true" pageId="6" pageNumber="7">paratype</emphasis>
, WMNH-INV-2014-188; C,
<taxonomicName id="E2E907D04131FFD4FCDCF8A8FBD5F88F" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[841,1047,1874,1898]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="7" phylum="Echinodermata" rank="species" species="kyushuensis" status="sp. nov.">
<emphasis id="179DA0414131FFD4FCDCF8A8FBD5F88F" box="[841,1047,1874,1898]" italics="true" pageId="6" pageNumber="7">Thyone kyushuensis</emphasis>
</taxonomicName>
<emphasis id="179DA0414131FFD4FBB4F8A8FBB7F88F" bold="true" box="[1057,1141,1874,1898]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="0CAE1D3A4131FFD4FBB4F8A8FBB7F88F" box="[1057,1141,1874,1898]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (i)
<emphasis id="179DA0414131FFD4FB32F8A8FB3CF88F" box="[1191,1278,1874,1898]" italics="true" pageId="6" pageNumber="7">holotype</emphasis>
, WMNH-INV- 2015-2437, (j)
<emphasis id="179DA0414131FFD4FEA5F88DFE48F86B" box="[304,394,1911,1934]" italics="true" pageId="6" pageNumber="7">paratype</emphasis>
, NSMT-E-12103; D,
<taxonomicName id="E2E907D04131FFD4FDFDF88DFCE6F86B" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[616,804,1910,1934]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="7" phylum="Echinodermata" rank="species" species="toyoshiomaruae" status="sp. nov.">
<emphasis id="179DA0414131FFD4FDFDF88DFCE6F86B" box="[616,804,1910,1934]" italics="true" pageId="6" pageNumber="7">T. toyoshiomaruae</emphasis>
</taxonomicName>
<emphasis id="179DA0414131FFD4FCBEF88CFCBEF86B" bold="true" box="[811,892,1910,1934]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="0CAE1D3A4131FFD4FCBEF88CFCBEF86B" box="[811,892,1910,1934]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (k)
<emphasis id="179DA0414131FFD4FC38F88DFBC5F86B" box="[941,1031,1911,1934]" italics="true" pageId="6" pageNumber="7">paratype</emphasis>
, WMNH-INV-2015-2438, (l)
<emphasis id="179DA0414131FFD4FAA9F88DFA54F86B" box="[1340,1430,1911,1934]" italics="true" pageId="6" pageNumber="7">paratype</emphasis>
, WMNH-INV-2015-2439, (m)
<emphasis id="179DA0414131FFD4FE5EF860FDE0F857" box="[459,546,1946,1970]" italics="true" pageId="6" pageNumber="7">holotype</emphasis>
, WMNH-INV-2015-2483, (n)
<emphasis id="179DA0414131FFD4FCCEF861FC77F857" box="[859,949,1947,1970]" italics="true" pageId="6" pageNumber="7">paratype</emphasis>
, WMNH-INV-2015-2484.
</paragraph>
</caption>
<caption id="71962CDB4130FFD5FF02FA26FAF8F984" ID-DOI="http://doi.org/10.5281/zenodo.5225378" ID-Zenodo-Dep="5225378" httpUri="https://zenodo.org/record/5225378/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,1500,1524]" targetBox="[152,1426,183,1471]" targetPageId="7">
<paragraph id="25567C534130FFD5FF02FA26FAF8F984" blockId="7.[151,1437,1500,1633]" pageId="7" pageNumber="8">
<emphasis id="179DA0414130FFD5FF02FA26FED0FA11" bold="true" box="[151,274,1500,1524]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
Elements of calcareous rings, viewed from the body cavity, anterior upper. A,
<taxonomicName id="E2E907D04130FFD5FBBDFA27FA82FA10" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[1064,1344,1501,1525]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="toyoshiomaruae" status="sp. nov.">
<emphasis id="179DA0414130FFD5FBBDFA27FA82FA10" box="[1064,1344,1501,1525]" italics="true" pageId="7" pageNumber="8">Massinium toyoshiomaruae</emphasis>
</taxonomicName>
<emphasis id="179DA0414130FFD5FAD3FA27FA57FA10" bold="true" box="[1350,1429,1501,1525]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="0CAE1D3A4130FFD5FAD3FA27FA57FA10" box="[1350,1429,1501,1525]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (
<emphasis id="179DA0414130FFD5FF35F9FBFF35F9FC" box="[160,247,1537,1561]" italics="true" pageId="7" pageNumber="8">holotype</emphasis>
, WMNH-INV-2014-186); B,
<taxonomicName id="E2E907D04130FFD5FDB1F9FBFD61F9FC" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[548,675,1537,1561]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="liaoi" status="sp. nov.">
<emphasis id="179DA0414130FFD5FDB1F9FBFD61F9FC" box="[548,675,1537,1561]" italics="true" pageId="7" pageNumber="8">Thyone liaoi</emphasis>
</taxonomicName>
<emphasis id="179DA0414130FFD5FD3CF9FBFD3BF9FC" bold="true" box="[681,761,1537,1561]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="0CAE1D3A4130FFD5FD3CF9FBFD3BF9FC" box="[681,761,1537,1561]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (
<emphasis id="179DA0414130FFD5FC9BF9FBFCA7F9FC" box="[782,869,1537,1561]" italics="true" pageId="7" pageNumber="8">holotype</emphasis>
, WMNH-INV-2014-182, and a
<emphasis id="179DA0414130FFD5FB3DF9FBFAC0F9FD" box="[1192,1282,1537,1560]" italics="true" pageId="7" pageNumber="8">paratype</emphasis>
, WMNH-INV- 2014-184); C,
<taxonomicName id="E2E907D04130FFD5FEBEF9DFFE7FF9D8" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[299,445,1573,1597]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="kyushuensis" status="sp. nov.">
<emphasis id="179DA0414130FFD5FEBEF9DFFE7FF9D8" box="[299,445,1573,1597]" italics="true" pageId="7" pageNumber="8">T. kyushuensis</emphasis>
</taxonomicName>
<emphasis id="179DA0414130FFD5FE56F9DFFDD1F9D8" bold="true" box="[451,531,1573,1597]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="0CAE1D3A4130FFD5FE56F9DFFDD1F9D8" box="[451,531,1573,1597]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (
<emphasis id="179DA0414130FFD5FDBDF9DFFDBDF9D8" box="[552,639,1573,1597]" italics="true" pageId="7" pageNumber="8">holotype</emphasis>
, WMNH-INV-2015-2437); D,
<taxonomicName id="E2E907D04130FFD5FC29F9DFFBBAF9D8" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[956,1144,1573,1597]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="toyoshiomaruae" status="sp. nov.">
<emphasis id="179DA0414130FFD5FC29F9DFFBBAF9D8" box="[956,1144,1573,1597]" italics="true" pageId="7" pageNumber="8">T. toyoshiomaruae</emphasis>
</taxonomicName>
<emphasis id="179DA0414130FFD5FBEBF9DFFB0DF9D8" bold="true" box="[1150,1231,1573,1597]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="0CAE1D3A4130FFD5FBEBF9DFFB0DF9D8" box="[1150,1231,1573,1597]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<emphasis id="179DA0414130FFD5FB4BF9DFFAF7F9D8" box="[1246,1333,1573,1597]" italics="true" pageId="7" pageNumber="8">holotype</emphasis>
, WMNH- INV-2015-2483). Abbreviations: md, mid-dorsal part; mv, mid-ventral part; -ir, interradial element; -r radial element.
</paragraph>
</caption>
<paragraph id="25567C534130FFD5FF52F975FD8AF8F0" blockId="7.[151,1437,1679,2029]" pageId="7" pageNumber="8">
Pedicels only with circular endplate, without supporting plates/rods, any kinds of supporting rods could not be observed even in the tips of the pedicels (
<figureCitation id="BDD260D64130FFD5FDE0F949FD0FF928" box="[629,717,1715,1741]" captionStart="FIGURE 4" captionStartId="8.[151,250,1916,1940]" captionTargetBox="[157,1434,384,1893]" captionTargetId="figure-114@8.[151,1436,381,1893]" captionTargetPageId="8" captionText="FIGURE 4. Scanning electron microscope images of ossicles of Massinium toyoshiomaruae sp. nov. (holotype, AE, WMNH- INV-2014-186). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225380" httpUri="https://zenodo.org/record/5225380/files/figure.png" pageId="7" pageNumber="8">Fig. 4E</figureCitation>
,
<tableCitation id="686B49E84130FFD5FD4FF949FCF3F928" box="[730,817,1715,1741]" captionStart="TABLE 3" captionStartId="9.[151,239,480,506]" captionText="TABLE 3. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Massinium toyoshiomaruae sp. nov." pageId="7" pageNumber="8">Table 3</tableCitation>
). Anal papillae mostly with supporting plates in distal part, and also with small tables with low spires, and rosettes in basal part (
<figureCitation id="BDD260D64130FFD5FB95F92DFB94F914" box="[1024,1110,1751,1777]" captionStart="FIGURE 4" captionStartId="8.[151,250,1916,1940]" captionTargetBox="[157,1434,384,1893]" captionTargetId="figure-114@8.[151,1436,381,1893]" captionTargetPageId="8" captionText="FIGURE 4. Scanning electron microscope images of ossicles of Massinium toyoshiomaruae sp. nov. (holotype, AE, WMNH- INV-2014-186). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225380" httpUri="https://zenodo.org/record/5225380/files/figure.png" pageId="7" pageNumber="8">Fig. 4F</figureCitation>
,
<tableCitation id="686B49E84130FFD5FBF1F92DFB79F914" box="[1124,1211,1751,1777]" captionStart="TABLE 3" captionStartId="9.[151,239,480,506]" captionText="TABLE 3. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Massinium toyoshiomaruae sp. nov." pageId="7" pageNumber="8">Table 3</tableCitation>
). Skin around anus occasionally possessing regular tables.
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD84130FFD8FF52F8E5FB50FEA4" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8" type="discussion">
<paragraph id="25567C534130FFD8FF52F8E5FB50FEA4" blockId="7.[151,1437,1679,2029]" lastBlockId="10.[151,1437,151,825]" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8">
<emphasis id="179DA0414130FFD5FF52F8E5FEF9F8DC" bold="true" box="[199,315,1823,1849]" pageId="7" pageNumber="8">Remarks.</emphasis>
This species is characterized by the peculiar shape of calcareous ring and possessing 20dendriform tentacles. Presently, these features are the principal features of the genus
<taxonomicName id="E2E907D04130FFD5FCB3F8BEFC59F8B8" authorityName="Samyn &amp; Thandar" authorityYear="2003" box="[806,923,1860,1885]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="genus">
<emphasis id="179DA0414130FFD5FCB3F8BEFC59F8B8" box="[806,923,1860,1885]" italics="true" pageId="7" pageNumber="8">Massinium</emphasis>
</taxonomicName>
and
<emphasis id="179DA0414130FFD5FC58F8B9FBA0F8B8" box="[973,1122,1859,1885]" italics="true" pageId="7" pageNumber="8">Neotyonidium</emphasis>
, former genus is well-defined from the latter by the lacking of bodywall table ossicles and includes 12 nominal species (WoRMS 2021a) (Table 7). In Japanese waters only two species of this genus,
<taxonomicName id="E2E907D04130FFD5FDE8F876FC66F840" authority="(Ludwig, 1882)" baseAuthorityName="Ludwig" baseAuthorityYear="1882" box="[637,932,1931,1957]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="magnum">
<emphasis id="179DA0414130FFD5FDE8F876FD39F840" box="[637,763,1932,1957]" italics="true" pageId="7" pageNumber="8">M. magnum</emphasis>
(
<bibRefCitation id="417801A24130FFD5FC90F871FC5EF840" author="Ludwig, H." box="[773,924,1931,1957]" pageId="7" pageNumber="8" pagination="127 - 137" refId="ref26104" refString="Ludwig, H. (1882) List of the holothurians in the collection of the Leyden Museum. Notes Leyden Museum, 4, 127 - 137." type="journal article" year="1882">Ludwig, 1882</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="E2E907D04130FFD5FC39F876FA5EF840" authority="(Yamana, Ota &amp; Kohtsuka, 2020)" baseAuthorityName="Yamana, Ota &amp; Kohtsuka" baseAuthorityYear="2020" box="[940,1436,1931,1957]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="japonicum">
<emphasis id="179DA0414130FFD5FC39F876FBFCF840" box="[940,1086,1932,1957]" italics="true" pageId="7" pageNumber="8">M.japonicum</emphasis>
(
<bibRefCitation id="417801A24130FFD5FBDDF871FA57F840" author="Yamana, Y. &amp; Ota, Y. &amp; Kohtsuka, H. &amp; Yoshizaki, K." box="[1096,1429,1931,1957]" pageId="7" pageNumber="8" pagination="227 - 236" refId="ref28233" refString="Yamana, Y., Ota, Y., Kohtsuka, H. &amp; Yoshizaki, K. (2020) A New Species of Massinium (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae: Semperiellinae) from western Japan. Species Diversity, 25, 227 - 236. https: // doi. org / 10.12782 / specdiv. 25.227" type="journal article" year="2020">Yamana, Ota &amp; Kohtsuka, 2020</bibRefCitation>
)
</taxonomicName>
has been reported (
<bibRefCitation id="417801A24130FFD5FEF4F855FE3BF82C" author="Nomura, K." box="[353,505,1967,1993]" pageId="7" pageNumber="8" pagination="126 - 128" refId="ref26411" refString="Nomura, K. (2004) New records of the tropical echinoderms in Kushimoto, southern end of the Kii Peninsula. The NANKI SEIBUTU, published by the NANKI Biological Society, 46, 126 - 128." type="journal article" year="2004">Nomura 2004</bibRefCitation>
;
<bibRefCitation id="417801A24130FFD5FD96F855FC9FF82C" author="Yamana, Y. &amp; Ota, Y. &amp; Kohtsuka, H. &amp; Yoshizaki, K." box="[515,861,1967,1993]" pageId="7" pageNumber="8" pagination="227 - 236" refId="ref28233" refString="Yamana, Y., Ota, Y., Kohtsuka, H. &amp; Yoshizaki, K. (2020) A New Species of Massinium (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae: Semperiellinae) from western Japan. Species Diversity, 25, 227 - 236. https: // doi. org / 10.12782 / specdiv. 25.227" type="journal article" year="2020">Yamana, Ota &amp; Kohtsuka, 2020</bibRefCitation>
). The new species
<taxonomicName id="E2E907D04130FFD5FBB3F84AFB37F82C" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[1062,1269,1967,1993]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="toyoshiomaruae">
<emphasis id="179DA0414130FFD5FBB3F84AFB37F82C" box="[1062,1269,1967,1993]" italics="true" pageId="7" pageNumber="8">M. toyoshiomaruae</emphasis>
</taxonomicName>
lacks bodywall ossicles and pedicel supporting plates/rods, and these features readily distinguish it from all congeners (Table 7). Among the 12
<taxonomicName id="E2E907D0413FFFDAFF61FF6DFE10FF54" box="[244,466,151,177]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="179DA041413FFFDAFF61FF6DFEADFF55" box="[244,367,151,176]" italics="true" pageId="8" pageNumber="9">Massinium</emphasis>
species
</taxonomicName>
, the following nine possess bodywall ossicles:
<taxonomicName id="E2E907D0413FFFDAFB8EFF6DFEBCFF30" authority="Samyn, Thandar &amp; Vandenspiegel, 2010" authorityName="Samyn, Thandar &amp; Vandenspiegel" authorityYear="2010" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="albicans">
<emphasis id="179DA041413FFFDAFB8EFF6DFB65FF54" box="[1051,1191,151,177]" italics="true" pageId="8" pageNumber="9">M. albicans</emphasis>
<bibRefCitation id="417801A2413FFFDAFB23FF6DFEBCFF30" author="Samyn, Y. &amp; Thandar, A. S. &amp; Vandenspiegel, D." pageId="8" pageNumber="9" pagination="1 - 19" refId="ref27454" refString="Samyn, Y., Thandar, A. S. &amp; Vandenspiegel, D. (2010) Two new species in the phyllophorid genus Massinium (Echinodermata: Holothuroidea) with redescription of Massinium magnum. Zootaxa, 2399 (1), 1 - 19. https: // doi. org / 10.11646 / zootaxa. 2399.1.1" type="journal article" year="2010">Samyn, Thandar &amp; Vandenspiegel, 2010</bibRefCitation>
</taxonomicName>
having rosettes and pseudobuttons,
<taxonomicName id="E2E907D0413FFFDAFC9CFF41FB62FF30" authority="(Thandar, 1989)" baseAuthorityName="Thandar" baseAuthorityYear="1989" box="[777,1184,187,213]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="arthroprocessum">
<emphasis id="179DA041413FFFDAFC9CFF41FC2BFF30" box="[777,1001,187,213]" italics="true" pageId="8" pageNumber="9">M. arthroprocessum</emphasis>
(
<bibRefCitation id="417801A2413FFFDAFC60FF41FB5AFF30" author="Thandar, A. S." box="[1013,1176,187,213]" pageId="8" pageNumber="9" pagination="637 - 644" refId="ref27692" refString="Thandar, A. S. (1989) A new species of a phyllophorid holothurian from southern Africa. Journal of Zoology, 19, 637 - 644. https: // doi. org / 10.1111 / j. 1469 - 7998.1989. tb 02605. x" type="journal article" year="1989">Thandar, 1989</bibRefCitation>
)
</taxonomicName>
with flattened rods,
<taxonomicName id="E2E907D0413FFFDAFAEBFF41FE69FF1C" authority="O'Loughlin" authorityName="O'Loughlin" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="bonapartum">
<emphasis id="179DA041413FFFDAFAEBFF41FEE2FF1C" italics="true" pageId="8" pageNumber="9">M. bonapartum</emphasis>
OLoughlin
</taxonomicName>
in
<bibRefCitation id="417801A2413FFFDAFE5AFF25FD21FF1C" author="O'Loughlin, P. &amp; Mackenzie, M. &amp; Vandenspiegel, D." box="[463,739,223,249]" pageId="8" pageNumber="9" pagination="5 - 23" refId="ref26740" refString="O'Loughlin, P., Mackenzie, M. &amp; Vandenspiegel, D. (2014) New dendrochirotid sea cucumbers from northern Australia (Echinodermata: Holothuroidea: Dendrochirotida). Memoirs of Museum Victoria, 72, 5 - 23. https: // doi. org / 1024199 / j. mmv. 2014.72.02" type="journal article" year="2014">
OLoughlin
<emphasis id="179DA041413FFFDAFDCFFF25FD50FF1C" box="[602,658,223,249]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
(2014)
</bibRefCitation>
with scales and tables,
<taxonomicName id="E2E907D0413FFFDAFC7FFF25FAA6FF1C" authority="(Cherbonnier, 1988)" baseAuthorityName="Cherbonnier" baseAuthorityYear="1988" box="[1002,1380,223,249]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="dissimilis">
<emphasis id="179DA041413FFFDAFC7FFF25FBB8FF1C" box="[1002,1146,223,249]" italics="true" pageId="8" pageNumber="9">M. dissimilis</emphasis>
(
<bibRefCitation id="417801A2413FFFDAFB1DFF25FA99FF1C" author="Cherbonnier, G." box="[1160,1371,223,249]" pageId="8" pageNumber="9" refId="ref24964" refString="Cherbonnier, G. (1988) Echinodermes: Holothurides. In: de l'ORSTOM (Ed.), Faune de Madagascar. Vol. 70. Museum national d'Histoire naturelle, Paris, pp. 1 - 292." type="book" year="1988">Cherbonnier, 1988</bibRefCitation>
)
</taxonomicName>
with tables and pseudobuttons,
<taxonomicName id="E2E907D0413FFFDAFE56FEF9FB8AFEF8" authority="Samyn, Thandar, &amp; Vandenspiegel, 2010" authorityName="Samyn, Thandar, &amp; Vandenspiegel" authorityYear="2010" box="[451,1096,259,285]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="granulosum">
<emphasis id="179DA041413FFFDAFE56FEF9FDB2FEF8" box="[451,624,259,285]" italics="true" pageId="8" pageNumber="9">M. granulosum</emphasis>
Samyn, Thandar, &amp; Vandenspiegel, 2010
</taxonomicName>
bearing granuliform rods and pseudobuttons,
<taxonomicName id="E2E907D0413FFFDAFED3FEDDFC9DFEA4" authority="(Yamana, Ota &amp; Kohtsuka, 2020)" baseAuthorityName="Yamana, Ota &amp; Kohtsuka" baseAuthorityYear="2020" box="[326,863,295,321]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="japonicum">
<emphasis id="179DA041413FFFDAFED3FEDDFE23FEA5" box="[326,481,295,320]" italics="true" pageId="8" pageNumber="9">M. japonicum</emphasis>
(
<bibRefCitation id="417801A2413FFFDAFE65FEDDFC95FEA4" author="Yamana, Y. &amp; Ota, Y. &amp; Kohtsuka, H. &amp; Yoshizaki, K." box="[496,855,295,321]" pageId="8" pageNumber="9" pagination="227 - 236" refId="ref28233" refString="Yamana, Y., Ota, Y., Kohtsuka, H. &amp; Yoshizaki, K. (2020) A New Species of Massinium (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae: Semperiellinae) from western Japan. Species Diversity, 25, 227 - 236. https: // doi. org / 10.12782 / specdiv. 25.227" type="journal article" year="2020">Yamana, Ota &amp; Kohtsuka, 2020</bibRefCitation>
)
</taxonomicName>
with rosettes (
<bibRefCitation id="417801A2413FFFDAFB94FEDDFAA9FEA4" author="Yamana, Y. &amp; Ota, Y. &amp; Kohtsuka, H. &amp; Yoshizaki, K." box="[1025,1387,295,321]" pageId="8" pageNumber="9" pagination="227 - 236" refId="ref28233" refString="Yamana, Y., Ota, Y., Kohtsuka, H. &amp; Yoshizaki, K. (2020) A New Species of Massinium (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae: Semperiellinae) from western Japan. Species Diversity, 25, 227 - 236. https: // doi. org / 10.12782 / specdiv. 25.227" type="journal article" year="2020">Yamana, Ota &amp; Kohtsuka, 2020</bibRefCitation>
),
<taxonomicName id="E2E907D0413FFFDAFAEBFEDDFEBCFE80" authority="O'Loughlin" authorityName="O'Loughlin" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="keesingi">
<emphasis id="179DA041413FFFDAFAEBFEDDFF31FE80" italics="true" pageId="8" pageNumber="9">M. keesingi</emphasis>
OLoughlin
</taxonomicName>
in
<bibRefCitation id="417801A2413FFFDAFE34FEB1FD77FE80" author="O'Loughlin, P. &amp; Mackenzie, M. &amp; Vandenspiegel, D." box="[417,693,331,357]" pageId="8" pageNumber="9" pagination="5 - 23" refId="ref26740" refString="O'Loughlin, P., Mackenzie, M. &amp; Vandenspiegel, D. (2014) New dendrochirotid sea cucumbers from northern Australia (Echinodermata: Holothuroidea: Dendrochirotida). Memoirs of Museum Victoria, 72, 5 - 23. https: // doi. org / 1024199 / j. mmv. 2014.72.02" type="journal article" year="2014">
OLoughlin
<emphasis id="179DA041413FFFDAFDB9FEB1FDA6FE80" box="[556,612,331,357]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
(2014)
</bibRefCitation>
with tables,
<taxonomicName id="E2E907D0413FFFDAFCD6FEB1FAC3FE80" authority="Samyn &amp; Thandar, 2003" authorityName="Samyn &amp; Thandar" authorityYear="2003" box="[835,1281,331,357]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="maculosum">
<emphasis id="179DA041413FFFDAFCD6FEB1FC2AFE80" box="[835,1000,331,357]" italics="true" pageId="8" pageNumber="9">M. maculosum</emphasis>
<bibRefCitation id="417801A2413FFFDAFC7BFEB1FAC3FE80" author="Samyn, Y. &amp; Thandar, A. S." box="[1006,1281,331,357]" pageId="8" pageNumber="9" pagination="132 - 142" refId="ref27401" refString="Samyn, Y. &amp; Thandar, A. S. (2003) Massinium, a new genus in the family Phyllophoridae (Echinodermata: Holothuroidea: Dendrochirotida) with description of a new south-west Indian Ocean species Massinium maculosum. Belgian Journal of Zoology, 133, 132 - 142." type="journal article" year="2003">Samyn &amp; Thandar, 2003</bibRefCitation>
</taxonomicName>
with rosettes, and
<taxonomicName id="E2E907D0413DFFD8FF53FF6DFDC3FF54" authority="(Ludwig, 1882)" baseAuthorityName="Ludwig" baseAuthorityYear="1882" box="[198,513,151,177]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="magnum">
<emphasis id="179DA041413DFFD8FF53FF6DFE89FF55" box="[198,331,151,176]" italics="true" pageId="10" pageNumber="11">M. magnum</emphasis>
(
<bibRefCitation id="417801A2413DFFD8FECFFF6DFE3BFF54" author="Ludwig, H." box="[346,505,151,177]" pageId="10" pageNumber="11" pagination="127 - 137" refId="ref26104" refString="Ludwig, H. (1882) List of the holothurians in the collection of the Leyden Museum. Notes Leyden Museum, 4, 127 - 137." type="journal article" year="1882">Ludwig, 1882</bibRefCitation>
)
</taxonomicName>
with rosettes and pseudobuttons. However, three Australian species,
<taxonomicName id="E2E907D0413DFFD8FA91FF6DFED9FF30" authority="O'Loughlin" authorityName="O'Loughlin" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="melanieae">
<emphasis id="179DA041413DFFD8FA91FF6DFA5EFF54" box="[1284,1436,151,177]" italics="true" pageId="10" pageNumber="11">M. melanieae</emphasis>
OLoughlin
</taxonomicName>
in
<bibRefCitation id="417801A2413DFFD8FEDDFF41FDACFF30" author="O'Loughlin, P. &amp; Bamos, S. &amp; Vandenspiegel, D." box="[328,622,187,213]" pageId="10" pageNumber="11" pagination="269 - 308" refId="ref26681" refString="O'Loughlin, P., Bamos, S. &amp; Vandenspiegel, D. (2012) The phyllophorid sea cucumbers of southern Australia (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae). Memoirs of Museum Victoria, 69, 269 - 308. https: // doi. org / 10.24199 / j. mmv. 2012.69.05" type="journal article" year="2012">
OLoughlin
<emphasis id="179DA041413DFFD8FE4DFF41FDD7FF30" box="[472,533,187,213]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
(2012)
</bibRefCitation>
,
<taxonomicName id="E2E907D0413DFFD8FDEBFF41FCB0FF30" authority="O'Loughlin" authorityName="O'Loughlin" box="[638,882,187,213]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="vimsi">
<emphasis id="179DA041413DFFD8FDEBFF41FD20FF31" box="[638,738,187,212]" italics="true" pageId="10" pageNumber="11">M. vimsi</emphasis>
OLoughlin
</taxonomicName>
in
<bibRefCitation id="417801A2413DFFD8FC0AFF41FB06FF30" author="O'Loughlin, P. &amp; Bamos, S. &amp; Vandenspiegel, D." box="[927,1220,187,213]" pageId="10" pageNumber="11" pagination="269 - 308" refId="ref26681" refString="O'Loughlin, P., Bamos, S. &amp; Vandenspiegel, D. (2012) The phyllophorid sea cucumbers of southern Australia (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae). Memoirs of Museum Victoria, 69, 269 - 308. https: // doi. org / 10.24199 / j. mmv. 2012.69.05" type="journal article" year="2012">
OLoughlin
<emphasis id="179DA041413DFFD8FBBAFF41FBAEFF30" box="[1071,1132,187,213]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
(2012)
</bibRefCitation>
, and
<taxonomicName id="E2E907D0413DFFD8FA9CFF41FED9FF1C" authority="O'Loughlin" authorityName="O'Loughlin" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="watsonae">
<emphasis id="179DA041413DFFD8FA9CFF41FA5EFF31" box="[1289,1436,187,212]" italics="true" pageId="10" pageNumber="11">M. watsonae</emphasis>
OLoughlin
</taxonomicName>
in
<bibRefCitation id="417801A2413DFFD8FEAAFF25FD97FF1C" author="O'Loughlin, P. &amp; Bamos, S. &amp; Vandenspiegel, D." box="[319,597,223,249]" pageId="10" pageNumber="11" pagination="269 - 308" refId="ref26681" refString="O'Loughlin, P., Bamos, S. &amp; Vandenspiegel, D. (2012) The phyllophorid sea cucumbers of southern Australia (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae). Memoirs of Museum Victoria, 69, 269 - 308. https: // doi. org / 10.24199 / j. mmv. 2012.69.05" type="journal article" year="2012">
OLoughlin
<emphasis id="179DA041413DFFD8FE5FFF25FDC0FF1C" box="[458,514,223,249]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
(2012)
</bibRefCitation>
, are frequently lacking mid-bodywall ossicles. Although the new species is also lacking mid-bodywall ossicles, it differs from these three species by the absence of the pedicel supporting rods and the presence of the supporting rods in the tips of the pedicels (
<bibRefCitation id="417801A2413DFFD8FCEBFEDDFB45FEA4" author="O'Loughlin, P. &amp; Bamos, S. &amp; Vandenspiegel, D." box="[894,1159,295,321]" pageId="10" pageNumber="11" pagination="269 - 308" refId="ref26681" refString="O'Loughlin, P., Bamos, S. &amp; Vandenspiegel, D. (2012) The phyllophorid sea cucumbers of southern Australia (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae). Memoirs of Museum Victoria, 69, 269 - 308. https: // doi. org / 10.24199 / j. mmv. 2012.69.05" type="journal article" year="2012">
OLoughlin
<emphasis id="179DA041413DFFD8FB9EFEDDFBFEFEA4" box="[1035,1084,295,321]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
, 2012
</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="71962CDB413FFFDAFF02F886FB73F839" ID-DOI="http://doi.org/10.5281/zenodo.5225380" ID-Zenodo-Dep="5225380" httpUri="https://zenodo.org/record/5225380/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1916,1940]" targetBox="[157,1434,384,1893]" targetPageId="8">
<paragraph id="25567C53413FFFDAFF02F886FB73F839" blockId="8.[151,1437,1916,2012]" pageId="8" pageNumber="9">
<emphasis id="179DA041413FFFDAFF02F886FED0F871" bold="true" box="[151,274,1916,1940]" pageId="8" pageNumber="9">FIGURE 4.</emphasis>
Scanning electron microscope images of ossicles of
<taxonomicName id="E2E907D0413FFFDAFCB2F887FBFDF871" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[807,1087,1916,1940]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="toyoshiomaruae" status="sp. nov.">
<emphasis id="179DA041413FFFDAFCB2F887FBFDF871" box="[807,1087,1916,1940]" italics="true" pageId="8" pageNumber="9">Massinium toyoshiomaruae</emphasis>
</taxonomicName>
<emphasis id="179DA041413FFFDAFBD3F887FB54F870" bold="true" box="[1094,1174,1917,1941]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="0CAE1D3A413FFFDAFBD3F887FB54F870" box="[1094,1174,1917,1941]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<emphasis id="179DA041413FFFDAFB31F886FB39F871" box="[1188,1275,1916,1940]" italics="true" pageId="8" pageNumber="9">holotype</emphasis>
, AE, WMNH- INV-2014-186). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F, the ventrolateral anal papilla.
</paragraph>
</caption>
<caption id="71962CDB413EFFDBFF02FE1AFA1EFE1F" box="[151,1500,480,506]" pageId="9" pageNumber="10" startId="9.[151,239,480,506]">
<paragraph id="25567C53413EFFDBFF02FE1AFA1EFE1F" blockId="9.[151,1500,480,506]" box="[151,1500,480,506]" pageId="9" pageNumber="10">
<emphasis id="179DA041413EFFDBFF02FE1AFEC8FE1F" bold="true" box="[151,266,480,506]" pageId="9" pageNumber="10">TABLE 3.</emphasis>
Counts and measurements (Mean ± sd μm) of ossicles from four specimens of
<taxonomicName id="E2E907D0413EFFDBFBCCFE1AFA42FE1F" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[1113,1408,480,506]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="toyoshiomaruae" status="sp. nov.">
<emphasis id="179DA041413EFFDBFBCCFE1AFA42FE1F" box="[1113,1408,480,506]" italics="true" pageId="9" pageNumber="10">Massinium toyoshiomaruae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CAE1D3A413EFFDBFA12FE1AFA1EFE1F" box="[1415,1500,480,506]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="25567C53413EFFDBFE28FDFDFD91F955" pageId="9" pageNumber="10">
<table id="57E98EF3413E002DFF30FDFDF809F954" box="[165,1995,519,1713]" gridcols="17" gridrows="35" pageId="9" pageNumber="10">
<tr id="9BD97E11413E002DFF30FDFDF809FDFB" box="[165,1995,519,542]" gridrow="0" pageId="9" pageNumber="10" rowspan-0="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-15="1" rowspan-16="1" rowspan-4="1" rowspan-5="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413E002DFE08FDFDFDBCFDFB" box="[413,638,519,542]" colspan="3" colspanRight="2" gridcol="1" gridrow="0" pageId="9" pageNumber="10">Plate and rosette</th>
<th id="D808176D413E002DFCC4FDFDFBDEFDFB" box="[849,1052,519,542]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="9" pageNumber="10">Endplate</th>
<th id="D808176D413E002DFB68FDFDFA92FDFB" box="[1277,1360,519,542]" gridcol="10" gridrow="0" pageId="9" pageNumber="10">Rod</th>
<th id="D808176D413E002DF900FDFDF93EFDFB" box="[1685,1788,519,542]" gridcol="14" gridrow="0" pageId="9" pageNumber="10">Table</th>
</tr>
<tr id="9BD97E11413E002DFF30FDD2F809FDA4" box="[165,1995,552,577]" gridrow="1" pageId="9" pageNumber="10" rowspan-1="1" rowspan-12="1" rowspan-4="1" rowspan-9="1">
<th id="D808176D413E002DFF30FDD2FEA0FDA4" box="[165,354,552,577]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">Samplea</th>
<td id="D808176D413E002DFE43FDD2FDBCFDA4" box="[470,638,552,577]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="9" pageNumber="10">Length</td>
<td id="D808176D413E002DFD76FDD2FC5AFDA4" box="[739,920,552,577]" colspan="2" colspanRight="1" gridcol="5" gridrow="1" pageId="9" pageNumber="10">Diameter</td>
<td id="D808176D413E002DFC4CFDD2FB45FDA4" box="[985,1159,552,577]" colspan="2" colspanRight="1" gridcol="7" gridrow="1" pageId="9" pageNumber="10">Length</td>
<td id="D808176D413E002DFB68FDD2FA77FDA4" box="[1277,1461,552,577]" colspan="2" colspanRight="1" gridcol="10" gridrow="1" pageId="9" pageNumber="10">Length</td>
<td id="D808176D413E002DF9BEFDD2F93EFDA4" box="[1579,1788,552,577]" colspan="2" colspanRight="1" gridcol="13" gridrow="1" pageId="9" pageNumber="10">Disc length</td>
<td id="D808176D413E002DF888FDD2F809FDA4" box="[1821,1995,552,577]" colspan="2" colspanRight="1" gridcol="15" gridrow="1" pageId="9" pageNumber="10">No. disc hole</td>
</tr>
<tr id="9BD97E11413E002DFF30FDB1F809FD84" box="[165,1995,587,609]" gridrow="2" pageId="9" pageNumber="10" rowspan-0="1">
<td id="D808176D413E002DFE08FDB1FE74FD84" box="[413,438,587,609]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">
<emphasis id="179DA041413EFFDBFE36FDB1FE6DFD84" box="[419,431,587,609]" italics="true" pageId="9" pageNumber="10">n</emphasis>
</td>
<td id="D808176D413E002DFE43FDB1FDD8FD84" box="[470,538,587,609]" gridcol="2" gridrow="2" pageId="9" pageNumber="10">Mean</td>
<td id="D808176D413E002DFDAFFDB1FDBCFD84" box="[570,638,587,609]" gridcol="3" gridrow="2" pageId="9" pageNumber="10">Range</td>
<td id="D808176D413E002DFD24FDB1FD7CFD84" box="[689,702,587,609]" gridcol="4" gridrow="2" pageId="9" pageNumber="10">
<emphasis id="179DA041413EFFDBFD27FDB1FD7CFD84" box="[690,702,587,609]" italics="true" pageId="9" pageNumber="10">n</emphasis>
</td>
<td id="D808176D413E002DFD76FDB1FCF2FD84" box="[739,816,587,609]" gridcol="5" gridrow="2" pageId="9" pageNumber="10">Mean</td>
<td id="D808176D413E002DFCC4FDB1FC5AFD84" box="[849,920,587,609]" gridcol="6" gridrow="2" pageId="9" pageNumber="10">Range</td>
<td id="D808176D413E002DFC4CFDB1FBDEFD84" box="[985,1052,587,609]" gridcol="7" gridrow="2" pageId="9" pageNumber="10">Mean</td>
<td id="D808176D413E002DFBD6FDB1FB45FD84" box="[1091,1159,587,609]" gridcol="8" gridrow="2" pageId="9" pageNumber="10">Range</td>
<td id="D808176D413E002DFB56FDB1FB18FD84" box="[1219,1242,587,609]" gridcol="9" gridrow="2" pageId="9" pageNumber="10">
<emphasis id="179DA041413EFFDBFB5DFDB1FB16FD84" box="[1224,1236,587,609]" italics="true" pageId="9" pageNumber="10">n</emphasis>
</td>
<td id="D808176D413E002DFB68FDB1FA92FD84" box="[1277,1360,587,609]" gridcol="10" gridrow="2" pageId="9" pageNumber="10">Mean</td>
<td id="D808176D413E002DFAE0FDB1FA77FD84" box="[1397,1461,587,609]" gridcol="11" gridrow="2" pageId="9" pageNumber="10">Range</td>
<td id="D808176D413E002DFA79FDB1F9C3FD84" box="[1516,1537,587,609]" gridcol="12" gridrow="2" pageId="9" pageNumber="10">
<emphasis id="179DA041413EFFDBFA64FDB1FA3FFD84" box="[1521,1533,587,609]" italics="true" pageId="9" pageNumber="10">n</emphasis>
</td>
<td id="D808176D413E002DF9BEFDB1F9ACFD84" box="[1579,1646,587,609]" gridcol="13" gridrow="2" pageId="9" pageNumber="10">Mean</td>
<td id="D808176D413E002DF900FDB1F93EFD84" box="[1685,1788,587,609]" gridcol="14" gridrow="2" pageId="9" pageNumber="10">Range</td>
<td id="D808176D413E002DF888FDB1F89AFD84" box="[1821,1880,587,609]" gridcol="15" gridrow="2" pageId="9" pageNumber="10">Mean</td>
<td id="D808176D413E002DF805FDB1F809FD84" box="[1936,1995,587,609]" gridcol="16" gridrow="2" pageId="9" pageNumber="10">Range</td>
</tr>
<tr id="9BD97E11413E002DFF30FD94F809FD60" box="[165,1995,622,645]" gridrow="3" pageId="9" pageNumber="10">
<th id="D808176D413E002DFF30FD94F809FD60" box="[165,1995,622,645]" colspan="17" colspanRight="16" gridcol="0" gridrow="3" pageId="9" pageNumber="10">WMNH-INV-2014-186</th>
</tr>
<tr id="9BD97E11413E002DFF30FD6AF809FD42" box="[165,1995,656,679]" gridrow="4" pageId="9" pageNumber="10" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30FD6AFEA0FD42" box="[165,354,656,679]" gridcol="0" gridrow="4" pageId="9" pageNumber="10">Tentacle</th>
<td id="D808176D413E002DFE08FD6AFE74FD42" box="[413,438,656,679]" gridcol="1" gridrow="4" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFD24FD6AFD7CFD42" box="[689,702,656,679]" gridcol="4" gridrow="4" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56FD6AFB18FD42" box="[1219,1242,656,679]" gridcol="9" gridrow="4" pageId="9" pageNumber="10">17</td>
<td id="D808176D413E002DFB68FD6AFA77FD42" box="[1277,1461,656,679]" colspan="2" colspanRight="1" gridcol="10" gridrow="4" pageId="9" pageNumber="10">100 ± 29 47153</td>
<td id="D808176D413E002DFA79FD6AF9C3FD42" box="[1516,1537,656,679]" gridcol="12" gridrow="4" pageId="9" pageNumber="10">10</td>
<td id="D808176D413E002DF9BEFD6AF9ACFD42" box="[1579,1646,656,679]" gridcol="13" gridrow="4" pageId="9" pageNumber="10">80 ± 10</td>
<td id="D808176D413E002DF900FD6AF93EFD42" box="[1685,1788,656,679]" gridcol="14" gridrow="4" pageId="9" pageNumber="10">6292</td>
</tr>
<tr id="9BD97E11413E002DFF30FD4EF809FD2E" box="[165,1995,692,715]" gridrow="5" pageId="9" pageNumber="10" rowspan-15="1" rowspan-16="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30FD4EFEA0FD2E" box="[165,354,692,715]" gridcol="0" gridrow="5" pageId="9" pageNumber="10">Peri-oral skin</th>
<td id="D808176D413E002DFE08FD4EFE74FD2E" box="[413,438,692,715]" gridcol="1" gridrow="5" pageId="9" pageNumber="10">4</td>
<td id="D808176D413E002DFE43FD4EFDD8FD2E" box="[470,538,692,715]" gridcol="2" gridrow="5" pageId="9" pageNumber="10">39 ± 8</td>
<td id="D808176D413E002DFDAFFD4EFDBCFD2E" box="[570,638,692,715]" gridcol="3" gridrow="5" pageId="9" pageNumber="10">3249</td>
<td id="D808176D413E002DFD24FD4EFD7CFD2E" box="[689,702,692,715]" gridcol="4" gridrow="5" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56FD4EFB18FD2E" box="[1219,1242,692,715]" gridcol="9" gridrow="5" pageId="9" pageNumber="10">17</td>
<td id="D808176D413E002DFB68FD4EFA92FD2E" box="[1277,1360,692,715]" gridcol="10" gridrow="5" pageId="9" pageNumber="10">54 ±17</td>
<td id="D808176D413E002DFAE0FD4EFA77FD2E" box="[1397,1461,692,715]" gridcol="11" gridrow="5" pageId="9" pageNumber="10">2585</td>
<td id="D808176D413E002DFA79FD4EF9C3FD2E" box="[1516,1537,692,715]" gridcol="12" gridrow="5" pageId="9" pageNumber="10">5</td>
<td id="D808176D413E002DF9BEFD4EF9ACFD2E" box="[1579,1646,692,715]" gridcol="13" gridrow="5" pageId="9" pageNumber="10">84 ± 20</td>
<td id="D808176D413E002DF900FD4EF93EFD2E" box="[1685,1788,692,715]" gridcol="14" gridrow="5" pageId="9" pageNumber="10">61105</td>
</tr>
<tr id="9BD97E11413E002DFF30FD2CF809FD08" box="[165,1995,726,749]" gridrow="6" pageId="9" pageNumber="10" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30FD2CFEA0FD08" box="[165,354,726,749]" gridcol="0" gridrow="6" pageId="9" pageNumber="10">Oesophagus skin</th>
<td id="D808176D413E002DFE08FD2CFE74FD08" box="[413,438,726,749]" gridcol="1" gridrow="6" pageId="9" pageNumber="10">16</td>
<td id="D808176D413E002DFE43FD2CFDD8FD08" box="[470,538,726,749]" gridcol="2" gridrow="6" pageId="9" pageNumber="10">29 ± 4</td>
<td id="D808176D413E002DFDAFFD2CFDBCFD08" box="[570,638,726,749]" gridcol="3" gridrow="6" pageId="9" pageNumber="10">2337</td>
<td id="D808176D413E002DFD24FD2CFD7CFD08" box="[689,702,726,749]" gridcol="4" gridrow="6" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56FD2CFB18FD08" box="[1219,1242,726,749]" gridcol="9" gridrow="6" pageId="9" pageNumber="10">2</td>
<td id="D808176D413E002DFB68FD2CFA92FD08" box="[1277,1360,726,749]" gridcol="10" gridrow="6" pageId="9" pageNumber="10">48</td>
<td id="D808176D413E002DFAE0FD2CFA77FD08" box="[1397,1461,726,749]" gridcol="11" gridrow="6" pageId="9" pageNumber="10">33, 63</td>
<td id="D808176D413E002DFA79FD2CF9C3FD08" box="[1516,1537,726,749]" gridcol="12" gridrow="6" pageId="9" pageNumber="10">0</td>
</tr>
<tr id="9BD97E11413E002DFF30FD02F809FCEA" box="[165,1995,760,783]" gridrow="7" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-15="1" rowspan-16="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30FD02FEA0FCEA" box="[165,354,760,783]" gridcol="0" gridrow="7" pageId="9" pageNumber="10">Introvert skin</th>
<td id="D808176D413E002DFE08FD02FE74FCEA" box="[413,438,760,783]" gridcol="1" gridrow="7" pageId="9" pageNumber="10">1</td>
<td id="D808176D413E002DFE43FD02FDD8FCEA" box="[470,538,760,783]" gridcol="2" gridrow="7" pageId="9" pageNumber="10">53</td>
<td id="D808176D413E002DFDAFFD02FDBCFCEA" box="[570,638,760,783]" gridcol="3" gridrow="7" pageId="9" pageNumber="10">53</td>
<td id="D808176D413E002DFD24FD02FD7CFCEA" box="[689,702,760,783]" gridcol="4" gridrow="7" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56FD02FB18FCEA" box="[1219,1242,760,783]" gridcol="9" gridrow="7" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFA79FD02F9C3FCEA" box="[1516,1537,760,783]" gridcol="12" gridrow="7" pageId="9" pageNumber="10">11</td>
<td id="D808176D413E002DF9BEFD02F9ACFCEA" box="[1579,1646,760,783]" gridcol="13" gridrow="7" pageId="9" pageNumber="10">69 ± 8</td>
<td id="D808176D413E002DF900FD02F93EFCEA" box="[1685,1788,760,783]" gridcol="14" gridrow="7" pageId="9" pageNumber="10">5878</td>
</tr>
<tr id="9BD97E11413E002DFF30FCE0F809FCD4" box="[165,1995,794,817]" gridrow="8" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1">
<th id="D808176D413E002DFF30FCE0FEA0FCD4" box="[165,354,794,817]" gridcol="0" gridrow="8" pageId="9" pageNumber="10">Pedicels (ventral)</th>
<td id="D808176D413E002DFE08FCE0FE74FCD4" box="[413,438,794,817]" gridcol="1" gridrow="8" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFD24FCE0FD7CFCD4" box="[689,702,794,817]" gridcol="4" gridrow="8" pageId="9" pageNumber="10">4</td>
<td id="D808176D413E002DFD76FCE0FCF2FCD4" box="[739,816,794,817]" gridcol="5" gridrow="8" pageId="9" pageNumber="10">145 ± 23</td>
<td id="D808176D413E002DFCC4FCE0FC5AFCD4" box="[849,920,794,817]" gridcol="6" gridrow="8" pageId="9" pageNumber="10">111164</td>
<td id="D808176D413E002DFC4CFCE0FBDEFCD4" box="[985,1052,794,817]" gridcol="7" gridrow="8" pageId="9" pageNumber="10">83 ± 32</td>
<td id="D808176D413E002DFBD6FCE0FB45FCD4" box="[1091,1159,794,817]" gridcol="8" gridrow="8" pageId="9" pageNumber="10">51112</td>
<td id="D808176D413E002DFB56FCE0FB18FCD4" box="[1219,1242,794,817]" gridcol="9" gridrow="8" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFA79FCE0F9C3FCD4" box="[1516,1537,794,817]" gridcol="12" gridrow="8" pageId="9" pageNumber="10">0</td>
</tr>
<tr id="9BD97E11413E002DFF30FCC6F809FCB6" box="[165,1995,828,851]" gridrow="9" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30FCC6FEA0FCB6" box="[165,354,828,851]" gridcol="0" gridrow="9" pageId="9" pageNumber="10">Bodywall (ventral)</th>
<td id="D808176D413E002DFE08FCC6FE74FCB6" box="[413,438,828,851]" gridcol="1" gridrow="9" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFD24FCC6FD7CFCB6" box="[689,702,828,851]" gridcol="4" gridrow="9" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56FCC6FB18FCB6" box="[1219,1242,828,851]" gridcol="9" gridrow="9" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFA79FCC6F9C3FCB6" box="[1516,1537,828,851]" gridcol="12" gridrow="9" pageId="9" pageNumber="10">0</td>
</tr>
<tr id="9BD97E11413E002DFF30FCA4F809FC93" box="[165,1995,862,886]" gridrow="10" pageId="9" pageNumber="10" rowspan-15="1" rowspan-16="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30FCA4FEA0FC93" box="[165,354,862,886]" gridcol="0" gridrow="10" pageId="9" pageNumber="10">Anal papilla</th>
<td id="D808176D413E002DFE08FCA4FE74FC93" box="[413,438,862,886]" gridcol="1" gridrow="10" pageId="9" pageNumber="10">16</td>
<td id="D808176D413E002DFE43FCA4FDD8FC93" box="[470,538,862,886]" gridcol="2" gridrow="10" pageId="9" pageNumber="10">47 ± 19</td>
<td id="D808176D413E002DFDAFFCA4FDBCFC93" box="[570,638,862,886]" gridcol="3" gridrow="10" pageId="9" pageNumber="10">3190</td>
<td id="D808176D413E002DFD24FCA4FD7CFC93" box="[689,702,862,886]" gridcol="4" gridrow="10" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56FCA4FB18FC93" box="[1219,1242,862,886]" gridcol="9" gridrow="10" pageId="9" pageNumber="10">3</td>
<td id="D808176D413E002DFB68FCA4FA92FC93" box="[1277,1360,862,886]" gridcol="10" gridrow="10" pageId="9" pageNumber="10">59 ± 17</td>
<td id="D808176D413E002DFAE0FCA4FA77FC93" box="[1397,1461,862,886]" gridcol="11" gridrow="10" pageId="9" pageNumber="10">4277</td>
<td id="D808176D413E002DFA79FCA4F9C3FC93" box="[1516,1537,862,886]" gridcol="12" gridrow="10" pageId="9" pageNumber="10">2</td>
<td id="D808176D413E002DF9BEFCA4F9ACFC93" box="[1579,1646,862,886]" gridcol="13" gridrow="10" pageId="9" pageNumber="10">50.5</td>
<td id="D808176D413E002DF900FCA4F93EFC93" box="[1685,1788,862,886]" gridcol="14" gridrow="10" pageId="9" pageNumber="10">48, 53</td>
</tr>
<tr id="9BD97E11413E002DFF30FC7BF809FC7D" box="[165,1995,897,920]" gridrow="11" pageId="9" pageNumber="10">
<th id="D808176D413E002DFF30FC7BF809FC7D" box="[165,1995,897,920]" colspan="17" colspanRight="16" gridcol="0" gridrow="11" pageId="9" pageNumber="10">WMNH-INV-2015-2497</th>
</tr>
<tr id="9BD97E11413E002DFF30FC59F809FC5F" box="[165,1995,931,954]" gridrow="12" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30FC59FEA0FC5F" box="[165,354,931,954]" gridcol="0" gridrow="12" pageId="9" pageNumber="10">Tentacle</th>
<td id="D808176D413E002DFE08FC59FE74FC5F" box="[413,438,931,954]" gridcol="1" gridrow="12" pageId="9" pageNumber="10">20</td>
<td id="D808176D413E002DFE43FC59FDD8FC5F" box="[470,538,931,954]" gridcol="2" gridrow="12" pageId="9" pageNumber="10">80 ± 10</td>
<td id="D808176D413E002DFDAFFC59FDBCFC5F" box="[570,638,931,954]" gridcol="3" gridrow="12" pageId="9" pageNumber="10">59101</td>
<td id="D808176D413E002DFD24FC59FD7CFC5F" box="[689,702,931,954]" gridcol="4" gridrow="12" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56FC59FB18FC5F" box="[1219,1242,931,954]" gridcol="9" gridrow="12" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFA79FC59F9C3FC5F" box="[1516,1537,931,954]" gridcol="12" gridrow="12" pageId="9" pageNumber="10">0</td>
</tr>
<tr id="9BD97E11413E002DFF30FC3FF809FC39" box="[165,1995,965,988]" gridrow="13" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-15="1" rowspan-16="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30FC3FFEA0FC39" box="[165,354,965,988]" gridcol="0" gridrow="13" pageId="9" pageNumber="10">Peri-oral skin</th>
<td id="D808176D413E002DFE08FC3FFE74FC39" box="[413,438,965,988]" gridcol="1" gridrow="13" pageId="9" pageNumber="10">20</td>
<td id="D808176D413E002DFE43FC3FFDD8FC39" box="[470,538,965,988]" gridcol="2" gridrow="13" pageId="9" pageNumber="10">42 ± 10</td>
<td id="D808176D413E002DFDAFFC3FFDBCFC39" box="[570,638,965,988]" gridcol="3" gridrow="13" pageId="9" pageNumber="10">2367</td>
<td id="D808176D413E002DFD24FC3FFD7CFC39" box="[689,702,965,988]" gridcol="4" gridrow="13" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56FC3FFB18FC39" box="[1219,1242,965,988]" gridcol="9" gridrow="13" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFA79FC3FF9C3FC39" box="[1516,1537,965,988]" gridcol="12" gridrow="13" pageId="9" pageNumber="10">8</td>
<td id="D808176D413E002DF9BEFC3FF9ACFC39" box="[1579,1646,965,988]" gridcol="13" gridrow="13" pageId="9" pageNumber="10">73 ± 14</td>
<td id="D808176D413E002DF900FC3FF93EFC39" box="[1685,1788,965,988]" gridcol="14" gridrow="13" pageId="9" pageNumber="10">5495</td>
</tr>
<tr id="9BD97E11413E002DFF30FC1DF809FC1B" box="[165,1995,999,1022]" gridrow="14" pageId="9" pageNumber="10" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30FC1DFEA0FC1B" box="[165,354,999,1022]" gridcol="0" gridrow="14" pageId="9" pageNumber="10">Oesophagus skin</th>
<td id="D808176D413E002DFE08FC1DFE74FC1B" box="[413,438,999,1022]" gridcol="1" gridrow="14" pageId="9" pageNumber="10">20</td>
<td id="D808176D413E002DFE43FC1DFDD8FC1B" box="[470,538,999,1022]" gridcol="2" gridrow="14" pageId="9" pageNumber="10">30 ± 10</td>
<td id="D808176D413E002DFDAFFC1DFDBCFC1B" box="[570,638,999,1022]" gridcol="3" gridrow="14" pageId="9" pageNumber="10">1547</td>
<td id="D808176D413E002DFD24FC1DFD7CFC1B" box="[689,702,999,1022]" gridcol="4" gridrow="14" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56FC1DFB18FC1B" box="[1219,1242,999,1022]" gridcol="9" gridrow="14" pageId="9" pageNumber="10">4</td>
<td id="D808176D413E002DFB68FC1DFA92FC1B" box="[1277,1360,999,1022]" gridcol="10" gridrow="14" pageId="9" pageNumber="10">31 ± 12</td>
<td id="D808176D413E002DFAE0FC1DFA77FC1B" box="[1397,1461,999,1022]" gridcol="11" gridrow="14" pageId="9" pageNumber="10">1743</td>
<td id="D808176D413E002DFA79FC1DF9C3FC1B" box="[1516,1537,999,1022]" gridcol="12" gridrow="14" pageId="9" pageNumber="10">0</td>
</tr>
<tr id="9BD97E11413E002DFF30FBF3F809FBC5" box="[165,1995,1033,1056]" gridrow="15" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30FBF3FEA0FBC5" box="[165,354,1033,1056]" gridcol="0" gridrow="15" pageId="9" pageNumber="10">Introvert skin</th>
<td id="D808176D413E002DFE08FBF3FE74FBC5" box="[413,438,1033,1056]" gridcol="1" gridrow="15" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFD24FBF3FD7CFBC5" box="[689,702,1033,1056]" gridcol="4" gridrow="15" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56FBF3FB18FBC5" box="[1219,1242,1033,1056]" gridcol="9" gridrow="15" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFA79FBF3F9C3FBC5" box="[1516,1537,1033,1056]" gridcol="12" gridrow="15" pageId="9" pageNumber="10">14</td>
<td id="D808176D413E002DF9BEFBF3F9ACFBC5" box="[1579,1646,1033,1056]" gridcol="13" gridrow="15" pageId="9" pageNumber="10">68 ± 6</td>
<td id="D808176D413E002DF900FBF3F93EFBC5" box="[1685,1788,1033,1056]" gridcol="14" gridrow="15" pageId="9" pageNumber="10">5278</td>
</tr>
<tr id="9BD97E11413E002DFF30FBD1F809FBA7" box="[165,1995,1067,1090]" gridrow="16" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1">
<th id="D808176D413E002DFF30FBD1FEA0FBA7" box="[165,354,1067,1090]" gridcol="0" gridrow="16" pageId="9" pageNumber="10">Pedicels (ventral)</th>
<td id="D808176D413E002DFE08FBD1FE74FBA7" box="[413,438,1067,1090]" gridcol="1" gridrow="16" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFD24FBD1FD7CFBA7" box="[689,702,1067,1090]" gridcol="4" gridrow="16" pageId="9" pageNumber="10">6</td>
<td id="D808176D413E002DFD76FBD1FCF2FBA7" box="[739,816,1067,1090]" gridcol="5" gridrow="16" pageId="9" pageNumber="10">98 ± 7</td>
<td id="D808176D413E002DFCC4FBD1FC5AFBA7" box="[849,920,1067,1090]" gridcol="6" gridrow="16" pageId="9" pageNumber="10">89110</td>
<td id="D808176D413E002DFC4CFBD1FBDEFBA7" box="[985,1052,1067,1090]" gridcol="7" gridrow="16" pageId="9" pageNumber="10">66 ± 7</td>
<td id="D808176D413E002DFBD6FBD1FB45FBA7" box="[1091,1159,1067,1090]" gridcol="8" gridrow="16" pageId="9" pageNumber="10">5573</td>
<td id="D808176D413E002DFB56FBD1FB18FBA7" box="[1219,1242,1067,1090]" gridcol="9" gridrow="16" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFA79FBD1F9C3FBA7" box="[1516,1537,1067,1090]" gridcol="12" gridrow="16" pageId="9" pageNumber="10">0</td>
</tr>
<tr id="9BD97E11413E002DFF30FBB7F809FB81" box="[165,1995,1101,1124]" gridrow="17" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30FBB7FEA0FB81" box="[165,354,1101,1124]" gridcol="0" gridrow="17" pageId="9" pageNumber="10">Bodywall (ventral)</th>
<td id="D808176D413E002DFE08FBB7FE74FB81" box="[413,438,1101,1124]" gridcol="1" gridrow="17" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFD24FBB7FD7CFB81" box="[689,702,1101,1124]" gridcol="4" gridrow="17" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56FBB7FB18FB81" box="[1219,1242,1101,1124]" gridcol="9" gridrow="17" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFA79FBB7F9C3FB81" box="[1516,1537,1101,1124]" gridcol="12" gridrow="17" pageId="9" pageNumber="10">0</td>
</tr>
<tr id="9BD97E11413E002DFF30FB95F809FB63" box="[165,1995,1135,1158]" gridrow="18" pageId="9" pageNumber="10" rowspan-15="1" rowspan-16="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30FB95FEA0FB63" box="[165,354,1135,1158]" gridcol="0" gridrow="18" pageId="9" pageNumber="10">Anal papilla</th>
<td id="D808176D413E002DFE08FB95FE74FB63" box="[413,438,1135,1158]" gridcol="1" gridrow="18" pageId="9" pageNumber="10">13</td>
<td id="D808176D413E002DFE43FB95FDD8FB63" box="[470,538,1135,1158]" gridcol="2" gridrow="18" pageId="9" pageNumber="10">42 ± 6</td>
<td id="D808176D413E002DFDAFFB95FDBCFB63" box="[570,638,1135,1158]" gridcol="3" gridrow="18" pageId="9" pageNumber="10">3150</td>
<td id="D808176D413E002DFD24FB95FD7CFB63" box="[689,702,1135,1158]" gridcol="4" gridrow="18" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56FB95FB18FB63" box="[1219,1242,1135,1158]" gridcol="9" gridrow="18" pageId="9" pageNumber="10">1</td>
<td id="D808176D413E002DFB68FB95FA92FB63" box="[1277,1360,1135,1158]" gridcol="10" gridrow="18" pageId="9" pageNumber="10">22</td>
<td id="D808176D413E002DFAE0FB95FA77FB63" box="[1397,1461,1135,1158]" gridcol="11" gridrow="18" pageId="9" pageNumber="10">22</td>
<td id="D808176D413E002DFA79FB95F9C3FB63" box="[1516,1537,1135,1158]" gridcol="12" gridrow="18" pageId="9" pageNumber="10">10</td>
<td id="D808176D413E002DF9BEFB95F9ACFB63" box="[1579,1646,1135,1158]" gridcol="13" gridrow="18" pageId="9" pageNumber="10">57 ± 7</td>
<td id="D808176D413E002DF900FB95F93EFB63" box="[1685,1788,1135,1158]" gridcol="14" gridrow="18" pageId="9" pageNumber="10">4367</td>
</tr>
<tr id="9BD97E11413E002DFF30FB6BF809FB4D" box="[165,1995,1169,1192]" gridrow="19" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413E002DFF30FB6BFE74FB4D" box="[165,438,1169,1192]" colspan="2" colspanRight="1" gridcol="0" gridrow="19" pageId="9" pageNumber="10">WMNH-INV-2015-2503</th>
</tr>
<tr id="9BD97E11413E002DFF30FB48F809FB2F" box="[165,1995,1202,1226]" gridrow="20" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413E002DFF30FB48FEA0FB2F" box="[165,354,1202,1226]" gridcol="0" gridrow="20" pageId="9" pageNumber="10">Tentacle</th>
<td id="D808176D413E002DFE08FB48FDD8FB2F" box="[413,538,1202,1226]" colspan="2" colspanRight="1" gridcol="1" gridrow="20" pageId="9" pageNumber="10">Amputated</td>
</tr>
<tr id="9BD97E11413E002DFF30FB2FF809FB0B" box="[165,1995,1237,1262]" gridrow="21" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413E002DFF30FB2FFEA0FB0B" box="[165,354,1237,1262]" gridcol="0" gridrow="21" pageId="9" pageNumber="10">Peri-oral skin</th>
<td id="D808176D413E002DFE08FB2FFDD8FB0B" box="[413,538,1237,1262]" colspan="2" colspanRight="1" gridcol="1" gridrow="21" pageId="9" pageNumber="10">Amputated</td>
</tr>
<tr id="9BD97E11413E002DFF30FB02F809FAF4" box="[165,1995,1272,1297]" gridrow="22" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413E002DFF30FB02FEA0FAF4" box="[165,354,1272,1297]" gridcol="0" gridrow="22" pageId="9" pageNumber="10">Oesophagus skin</th>
<td id="D808176D413E002DFE08FB02FDD8FAF4" box="[413,538,1272,1297]" colspan="2" colspanRight="1" gridcol="1" gridrow="22" pageId="9" pageNumber="10">Amputated</td>
</tr>
<tr id="9BD97E11413E002DFF30FAE1F809FAD1" box="[165,1995,1307,1332]" gridrow="23" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413E002DFF30FAE1FEA0FAD1" box="[165,354,1307,1332]" gridcol="0" gridrow="23" pageId="9" pageNumber="10">Introvert skin</th>
<td id="D808176D413E002DFE08FAE1FDD8FAD1" box="[413,538,1307,1332]" colspan="2" colspanRight="1" gridcol="1" gridrow="23" pageId="9" pageNumber="10">Amputated</td>
</tr>
<tr id="9BD97E11413E002DFF30FAC4F809FAB2" box="[165,1995,1342,1367]" gridrow="24" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1">
<th id="D808176D413E002DFF30FAC4FEA0FAB2" box="[165,354,1342,1367]" gridcol="0" gridrow="24" pageId="9" pageNumber="10">Pedicels (ventral)</th>
<td id="D808176D413E002DFE08FAC4FE74FAB2" box="[413,438,1342,1367]" gridcol="1" gridrow="24" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFD24FAC4FD7CFAB2" box="[689,702,1342,1367]" gridcol="4" gridrow="24" pageId="9" pageNumber="10">4</td>
<td id="D808176D413E002DFD76FAC4FCF2FAB2" box="[739,816,1342,1367]" gridcol="5" gridrow="24" pageId="9" pageNumber="10">80 ± 11</td>
<td id="D808176D413E002DFCC4FAC4FC5AFAB2" box="[849,920,1342,1367]" gridcol="6" gridrow="24" pageId="9" pageNumber="10">6488</td>
<td id="D808176D413E002DFC4CFAC4FBDEFAB2" box="[985,1052,1342,1367]" gridcol="7" gridrow="24" pageId="9" pageNumber="10">51 ± 12</td>
<td id="D808176D413E002DFBD6FAC4FB45FAB2" box="[1091,1159,1342,1367]" gridcol="8" gridrow="24" pageId="9" pageNumber="10">3662</td>
<td id="D808176D413E002DFB56FAC4FB18FAB2" box="[1219,1242,1342,1367]" gridcol="9" gridrow="24" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFA79FAC4F9C3FAB2" box="[1516,1537,1342,1367]" gridcol="12" gridrow="24" pageId="9" pageNumber="10">0</td>
</tr>
<tr id="9BD97E11413E002DFF30FA9BF809FA9C" box="[165,1995,1377,1401]" gridrow="25" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30FA9BFEA0FA9C" box="[165,354,1377,1401]" gridcol="0" gridrow="25" pageId="9" pageNumber="10">Bodywall (ventral)</th>
<td id="D808176D413E002DFE08FA9BFE74FA9C" box="[413,438,1377,1401]" gridcol="1" gridrow="25" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFD24FA9BFD7CFA9C" box="[689,702,1377,1401]" gridcol="4" gridrow="25" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56FA9BFB18FA9C" box="[1219,1242,1377,1401]" gridcol="9" gridrow="25" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFA79FA9BF9C3FA9C" box="[1516,1537,1377,1401]" gridcol="12" gridrow="25" pageId="9" pageNumber="10">0</td>
</tr>
<tr id="9BD97E11413E002DFF30FA79F809FA7E" box="[165,1995,1411,1435]" gridrow="26" pageId="9" pageNumber="10" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30FA79FEA0FA7E" box="[165,354,1411,1435]" gridcol="0" gridrow="26" pageId="9" pageNumber="10">Anal papilla</th>
<td id="D808176D413E002DFE08FA79FE74FA7E" box="[413,438,1411,1435]" gridcol="1" gridrow="26" pageId="9" pageNumber="10">20</td>
<td id="D808176D413E002DFE43FA79FDD8FA7E" box="[470,538,1411,1435]" gridcol="2" gridrow="26" pageId="9" pageNumber="10">46 ± 5</td>
<td id="D808176D413E002DFDAFFA79FDBCFA7E" box="[570,638,1411,1435]" gridcol="3" gridrow="26" pageId="9" pageNumber="10">4151</td>
<td id="D808176D413E002DFD24FA79FD7CFA7E" box="[689,702,1411,1435]" gridcol="4" gridrow="26" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56FA79FB18FA7E" box="[1219,1242,1411,1435]" gridcol="9" gridrow="26" pageId="9" pageNumber="10">5</td>
<td id="D808176D413E002DFB68FA79FA92FA7E" box="[1277,1360,1411,1435]" gridcol="10" gridrow="26" pageId="9" pageNumber="10">63 ± 13</td>
<td id="D808176D413E002DFAE0FA79FA77FA7E" box="[1397,1461,1411,1435]" gridcol="11" gridrow="26" pageId="9" pageNumber="10">4277</td>
<td id="D808176D413E002DFA79FA79F9C3FA7E" box="[1516,1537,1411,1435]" gridcol="12" gridrow="26" pageId="9" pageNumber="10">3</td>
<td id="D808176D413E002DF9BEFA79F9ACFA7E" box="[1579,1646,1411,1435]" gridcol="13" gridrow="26" pageId="9" pageNumber="10">46 ± 5</td>
<td id="D808176D413E002DF900FA79F93EFA7E" box="[1685,1788,1411,1435]" gridcol="14" gridrow="26" pageId="9" pageNumber="10">4151</td>
<td id="D808176D413E002DF888FA79F89AFA7E" box="[1821,1880,1411,1435]" gridcol="15" gridrow="26" pageId="9" pageNumber="10">12 ± 7</td>
<td id="D808176D413E002DF805FA79F809FA7E" box="[1936,1995,1411,1435]" gridcol="16" gridrow="26" pageId="9" pageNumber="10">820</td>
</tr>
<tr id="9BD97E11413E002DFF30FA52F809FA5A" box="[165,1995,1448,1471]" gridrow="27" pageId="9" pageNumber="10" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413E002DFF30FA52FEA0FA5A" box="[165,354,1448,1471]" gridcol="0" gridrow="27" pageId="9" pageNumber="10">NMST-E-12099-1</th>
</tr>
<tr id="9BD97E11413E002DFF30FA33F809FA07" box="[165,1995,1481,1506]" gridrow="28" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413E002DFF30FA33FEA0FA07" box="[165,354,1481,1506]" gridcol="0" gridrow="28" pageId="9" pageNumber="10">Tentacle</th>
<td id="D808176D413E002DFE08FA33FDBCFA07" box="[413,638,1481,1506]" colspan="3" colspanRight="2" gridcol="1" gridrow="28" pageId="9" pageNumber="10">Not measured</td>
</tr>
<tr id="9BD97E11413E002DFF30FA16F809F9E0" box="[165,1995,1516,1541]" gridrow="29" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413E002DFF30FA16FEA0F9E0" box="[165,354,1516,1541]" gridcol="0" gridrow="29" pageId="9" pageNumber="10">Peri-oral skin</th>
<td id="D808176D413E002DFE08FA16FDBCF9E0" box="[413,638,1516,1541]" colspan="3" colspanRight="2" gridcol="1" gridrow="29" pageId="9" pageNumber="10">Not measured</td>
</tr>
<tr id="9BD97E11413E002DFF30F9EAF809F9CC" box="[165,1995,1552,1577]" gridrow="30" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413E002DFF30F9EAFEA0F9CC" box="[165,354,1552,1577]" gridcol="0" gridrow="30" pageId="9" pageNumber="10">Oesophagus skin</th>
<td id="D808176D413E002DFE08F9EAFDBCF9CC" box="[413,638,1552,1577]" colspan="3" colspanRight="2" gridcol="1" gridrow="30" pageId="9" pageNumber="10">Not measured</td>
</tr>
<tr id="9BD97E11413E002DFF30F9C9F809F9AF" box="[165,1995,1587,1610]" gridrow="31" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30F9C9FEA0F9AF" box="[165,354,1587,1610]" gridcol="0" gridrow="31" pageId="9" pageNumber="10">Introvert skin</th>
<td id="D808176D413E002DFE08F9C9FE74F9AF" box="[413,438,1587,1610]" gridcol="1" gridrow="31" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFD24F9C9FD7CF9AF" box="[689,702,1587,1610]" gridcol="4" gridrow="31" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56F9C9FB18F9AF" box="[1219,1242,1587,1610]" gridcol="9" gridrow="31" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFA79F9C9F9C3F9AF" box="[1516,1537,1587,1610]" gridcol="12" gridrow="31" pageId="9" pageNumber="10">11</td>
<td id="D808176D413E002DF9BEF9C9F9ACF9AF" box="[1579,1646,1587,1610]" gridcol="13" gridrow="31" pageId="9" pageNumber="10">76 ± 5</td>
<td id="D808176D413E002DF900F9C9F93EF9AF" box="[1685,1788,1587,1610]" gridcol="14" gridrow="31" pageId="9" pageNumber="10">6983</td>
<td id="D808176D413E002DF888F9C9F89AF9AF" box="[1821,1880,1587,1610]" gridcol="15" gridrow="31" pageId="9" pageNumber="10">39 ± 11</td>
<td id="D808176D413E002DF805F9C9F809F9AF" box="[1936,1995,1587,1610]" gridcol="16" gridrow="31" pageId="9" pageNumber="10">2666</td>
</tr>
<tr id="9BD97E11413E002DFF30F9AFF809F989" box="[165,1995,1621,1644]" gridrow="32" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1">
<th id="D808176D413E002DFF30F9AFFEA0F989" box="[165,354,1621,1644]" gridcol="0" gridrow="32" pageId="9" pageNumber="10">Pedicels (ventral)</th>
<td id="D808176D413E002DFE08F9AFFE74F989" box="[413,438,1621,1644]" gridcol="1" gridrow="32" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFD24F9AFFD7CF989" box="[689,702,1621,1644]" gridcol="4" gridrow="32" pageId="9" pageNumber="10">3</td>
<td id="D808176D413E002DFD76F9AFFCF2F989" box="[739,816,1621,1644]" gridcol="5" gridrow="32" pageId="9" pageNumber="10">103 ± 2</td>
<td id="D808176D413E002DFCC4F9AFFC5AF989" box="[849,920,1621,1644]" gridcol="6" gridrow="32" pageId="9" pageNumber="10">101105</td>
<td id="D808176D413E002DFC4CF9AFFBDEF989" box="[985,1052,1621,1644]" gridcol="7" gridrow="32" pageId="9" pageNumber="10">57 ± 21</td>
<td id="D808176D413E002DFBD6F9AFFB45F989" box="[1091,1159,1621,1644]" gridcol="8" gridrow="32" pageId="9" pageNumber="10">3270</td>
<td id="D808176D413E002DFB56F9AFFB18F989" box="[1219,1242,1621,1644]" gridcol="9" gridrow="32" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFA79F9AFF9C3F989" box="[1516,1537,1621,1644]" gridcol="12" gridrow="32" pageId="9" pageNumber="10">0</td>
</tr>
<tr id="9BD97E11413E002DFF30F98DF809F96B" box="[165,1995,1655,1678]" gridrow="33" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413E002DFF30F98DFEA0F96B" box="[165,354,1655,1678]" gridcol="0" gridrow="33" pageId="9" pageNumber="10">Bodywall (ventral)</th>
<td id="D808176D413E002DFE08F98DFE74F96B" box="[413,438,1655,1678]" gridcol="1" gridrow="33" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFD24F98DFD7CF96B" box="[689,702,1655,1678]" gridcol="4" gridrow="33" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFB56F98DFB18F96B" box="[1219,1242,1655,1678]" gridcol="9" gridrow="33" pageId="9" pageNumber="10">0</td>
<td id="D808176D413E002DFA79F98DF9C3F96B" box="[1516,1537,1655,1678]" gridcol="12" gridrow="33" pageId="9" pageNumber="10">0</td>
</tr>
<tr id="9BD97E11413E002DFF30F963F809F954" box="[165,1995,1689,1713]" gridrow="34" pageId="9" pageNumber="10" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413E002DFF30F963FEA0F954" box="[165,354,1689,1713]" gridcol="0" gridrow="34" pageId="9" pageNumber="10">Anal papilla</th>
<td id="D808176D413E002DFE08F963FDBCF954" box="[413,638,1689,1713]" colspan="3" colspanRight="2" gridcol="1" gridrow="34" pageId="9" pageNumber="10">Not measured</td>
</tr>
</table>
</paragraph>
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WMNH-INV: Invertebrate collection of the Wakayama Prefectural Museum of Natural History, NMST-E: Echinoderms collection of National Museum of Nature and Science.
</tableNote>
</paragraph>
<subSubSection id="6DF32FD8413DFFD8FF52FEB0FD71FE48" pageId="10" pageNumber="11" type="distribution">
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<emphasis id="179DA041413DFFD8FF52FEB0FEA3FE81" bold="true" box="[199,353,330,356]" pageId="10" pageNumber="11">Distribution.</emphasis>
In the Seto Inland Sea and western part of the Sea of
<collectingCountry id="5DFE3CC3413DFFD8FC3CFEB1FC2EFE80" box="[937,1004,331,357]" name="Japan" pageId="10" pageNumber="11">Japan</collectingCountry>
, depth
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12
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</quantity>
. The Bisan-Seto Strait, eastern part of the Seto Inland Sea, sand bottom, depth
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(ST-
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2014); western part of the Sea of
<collectingCountry id="5DFE3CC3413DFFD8FAC3FE95FA5BFE6C" box="[1366,1433,367,393]" name="Japan" pageId="10" pageNumber="11">Japan</collectingCountry>
, sand bottom, depth
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(ST-9, 10 in 2015).
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</subSubSection>
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<emphasis id="179DA041413DFFD8FF52FE4DFE8CFE34" bold="true" box="[199,334,439,465]" pageId="10" pageNumber="11">Etymology.</emphasis>
The specific name
<taxonomicName id="E2E907D0413DFFD8FDBEFE4DFD1DFE34" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[555,735,439,465]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="toyoshiomaruae">
<emphasis id="179DA041413DFFD8FDBEFE4DFD1DFE34" box="[555,735,439,465]" italics="true" pageId="10" pageNumber="11">toyoshiomaruae</emphasis>
</taxonomicName>
alludes to the name (TR/V)
<emphasis id="179DA041413DFFD8FBB6FE4DFAC2FE34" box="[1059,1280,439,465]" italics="true" pageId="10" pageNumber="11">TOYOSHIO MARU</emphasis>
of
<collectingRegion id="E72DB2B1413DFFD8FAB3FE4DFA5EFE34" box="[1318,1436,439,465]" country="Japan" name="Hiroshima" pageId="10" pageNumber="11">Hiroshima</collectingRegion>
University.
</paragraph>
<paragraph id="25567C53413DFFD8FF52FE05FB5DFCDD" blockId="10.[151,1437,151,825]" pageId="10" pageNumber="11">
<emphasis id="179DA041413DFFD8FF52FE05FDD7FDFD" bold="true" box="[199,533,510,537]" pageId="10" pageNumber="11">Molecular data comparison.</emphasis>
In BLAST searches, the three closest hits obtained for the partial COI gene sequence were those of
<taxonomicName id="E2E907D0413DFFD8FEA4FDD9FE3EFDD8" box="[305,508,547,573]" class="Holothuroidea" family="Phyllophoridae" genus="Neothyonidium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="179DA041413DFFD8FEA4FDD9FE15FDD8" box="[305,471,547,573]" italics="true" pageId="10" pageNumber="11">Neothyonidium</emphasis>
sp.
</taxonomicName>
(INSD accession number
<accessionNumber id="3ABAE1B0413DFFD8FC89FDD9FC52FDD8" box="[796,912,547,573]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF142179" pageId="10" pageNumber="11">KF142179</accessionNumber>
and
<accessionNumber id="3ABAE1B0413DFFD8FC50FDD9FBFFFDD8" box="[965,1085,547,573]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF142176" pageId="10" pageNumber="11">KF142176</accessionNumber>
, 86.5% and 86.2% respectively) and
<taxonomicName id="E2E907D0413DFFD8FF5FFDBDFE69FD85" baseAuthorityName="Ludwig" baseAuthorityYear="1882" box="[202,427,583,608]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="magnum">
<emphasis id="179DA041413DFFD8FF5FFDBDFE69FD85" box="[202,427,583,608]" italics="true" pageId="10" pageNumber="11">Massinium magnum</emphasis>
</taxonomicName>
(
<accessionNumber id="3ABAE1B0413DFFD8FE2BFDBDFDF4FD84" box="[446,566,583,609]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF142162" pageId="10" pageNumber="11">KF142162</accessionNumber>
, 84.6%). The results of BLAST searches confirmed that the present species is genetically related to
<emphasis id="179DA041413DFFD8FE31FD91FDF2FD61" box="[420,560,619,644]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="E2E907D0413DFFD8FE31FD91FDEFFD61" baseAuthorityName="Ludwig" baseAuthorityYear="1882" box="[420,557,619,644]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="magnum">M. magnum</taxonomicName>
,
</emphasis>
and supported that the present species belongs to the genus
<taxonomicName id="E2E907D0413DFFD8FB4EFD91FA91FD61" authorityName="Samyn &amp; Thandar" authorityYear="2003" box="[1243,1363,619,644]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="179DA041413DFFD8FB4EFD91FA91FD61" box="[1243,1363,619,644]" italics="true" pageId="10" pageNumber="11">Massinium</emphasis>
</taxonomicName>
and it and is distinct from
<taxonomicName id="E2E907D0413DFFD8FEEEFD75FDC2FD4D" baseAuthorityName="Ludwig" baseAuthorityYear="1882" box="[379,512,655,680]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="magnum">
<emphasis id="179DA041413DFFD8FEEEFD75FDC2FD4D" box="[379,512,655,680]" italics="true" pageId="10" pageNumber="11">M. magnum</emphasis>
</taxonomicName>
. The results of BLAST searches also showed that the present species is similar to
<taxonomicName id="E2E907D0413DFFD8FF02FD49FEA6FD28" box="[151,356,691,717]" class="Holothuroidea" family="Phyllophoridae" genus="Neothyonidium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="179DA041413DFFD8FF02FD49FEFFFD28" box="[151,317,691,717]" italics="true" pageId="10" pageNumber="11">Neothyonidium</emphasis>
sp.
</taxonomicName>
However, the detailed morphologies of voucher specimens of
<taxonomicName id="E2E907D0413DFFD8FB8FFD49FB25FD28" box="[1050,1255,691,717]" class="Holothuroidea" family="Phyllophoridae" genus="Neothyonidium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="179DA041413DFFD8FB8FFD49FB02FD28" box="[1050,1216,691,717]" italics="true" pageId="10" pageNumber="11">Neothyonidium</emphasis>
sp.
</taxonomicName>
(
<accessionNumber id="3ABAE1B0413DFFD8FB63FD49FAAEFD28" box="[1270,1388,691,717]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF142179" pageId="10" pageNumber="11">KF142179</accessionNumber>
and
<accessionNumber id="3ABAE1B0413DFFD8FF02FD2DFECFFD17" box="[151,269,727,754]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF142176" pageId="10" pageNumber="11">KF142176</accessionNumber>
) are unavailable, and INSD sometimes includes misidentification; thus, morphological studies to confirm the relatedness of these species are required. However, this result is quite acceptable because the previous nominated species of
<taxonomicName id="E2E907D0413DFFD8FF2EFCE5FEF1FCDD" authorityName="Samyn &amp; Thandar" authorityYear="2003" box="[187,307,799,824]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="179DA041413DFFD8FF2EFCE5FEF1FCDD" box="[187,307,799,824]" italics="true" pageId="10" pageNumber="11">Massinium</emphasis>
</taxonomicName>
were put in
<taxonomicName id="E2E907D0413DFFD8FE2DFCE5FD9CFCDC" authorityName="Deichmann" authorityYear="1938" box="[440,606,799,825]" class="Holothuroidea" family="Phyllophoridae" genus="Neothyonidium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="179DA041413DFFD8FE2DFCE5FD9CFCDC" box="[440,606,799,825]" italics="true" pageId="10" pageNumber="11">Neothyonidium</emphasis>
</taxonomicName>
until
<bibRefCitation id="417801A2413DFFD8FD09FCE5FC07FCDC" author="Samyn, Y. &amp; Thandar, A. S." box="[668,965,799,825]" pageId="10" pageNumber="11" pagination="132 - 142" refId="ref27401" refString="Samyn, Y. &amp; Thandar, A. S. (2003) Massinium, a new genus in the family Phyllophoridae (Echinodermata: Holothuroidea: Dendrochirotida) with description of a new south-west Indian Ocean species Massinium maculosum. Belgian Journal of Zoology, 133, 132 - 142." type="journal article" year="2003">Samyn and Thandar (2003)</bibRefCitation>
erected
<emphasis id="179DA041413DFFD8FBB5FCE5FB5DFCDD" box="[1056,1183,799,824]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="E2E907D0413DFFD8FBB5FCE5FB59FCDD" authorityName="Samyn &amp; Thandar" authorityYear="2003" box="[1056,1179,799,824]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">Massinium</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,557 @@
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<mods:title id="79D15AC2C18CC9E22C49A2DE21C3382C">Four new dendrochirotid holothurians collected from the Seto Inland Sea and the western part of the Sea of Japan, western Japan</mods:title>
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<mods:classification id="25A4CA79E012993E3E2DC66F48F7BE2D">journal article</mods:classification>
<mods:identifier id="6D0CA26834D5A82B5871E5EF9CA4A2DF" type="DOI">10.11646/zootaxa.5023.1.1</mods:identifier>
<mods:identifier id="170CF8978E3A6F63263D35DCB573D5B4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="35B2E0D884BB629F0D805E463E86BEEC" type="Zenodo-Dep">5225372</mods:identifier>
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</mods:mods>
<treatment id="AD40CD454139FFC3FF02FDA0FDB4FAE8" ID-DOI="http://doi.org/10.5281/zenodo.5225388" ID-Zenodo-Dep="5225388" LSID="urn:lsid:zoobank.org:act:2D90339A-1315-48D9-ACC9-8A0DFDC5C01F" httpUri="http://treatment.plazi.org/id/AD40CD454139FFC3FF02FDA0FDB4FAE8" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
<subSubSection id="6DF32FD84139FFDCFF02FDA0FD6FFD7C" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="25567C534139FFDCFF02FDA0FC98FD90" blockId="14.[151,858,602,702]" box="[151,858,602,629]" pageId="14" pageNumber="15">
<heading id="7E1ECB3F4139FFDCFF02FDA0FC98FD90" bold="true" box="[151,858,602,629]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="179DA0414139FFDCFF02FDA0FC98FD90" bold="true" box="[151,858,602,629]" pageId="14" pageNumber="15">
<taxonomicName id="E2E907D04139FFDCFF02FDA0FD34FD90" authority="Yamana, Ogawa &amp; Ohtsuka" authorityName="Yamana, Ogawa &amp; Ohtsuka" authorityYear="2021" box="[151,758,602,629]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="kyushuensis" status="sp. nov.">
<emphasis id="179DA0414139FFDCFF02FDA0FE4CFD90" bold="true" box="[151,398,602,629]" italics="true" pageId="14" pageNumber="15">Thyone kyushuensis</emphasis>
Yamana, Ogawa &amp; Ohtsuka
</taxonomicName>
<taxonomicNameLabel id="0CAE1D3A4139FFDCFD68FDA1FC98FD90" box="[765,858,603,629]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="25567C534139FFDCFF02FD85FD6FFD7C" blockId="14.[151,858,602,702]" box="[151,685,638,665]" pageId="14" pageNumber="15">[New Japanese name: Kyushu-kehada-namako]</paragraph>
</subSubSection>
<subSubSection id="6DF32FD84139FFDCFF02FD59FE41FD5B" box="[151,387,675,702]" pageId="14" pageNumber="15" type="description">
<paragraph id="25567C534139FFDCFF02FD59FE41FD5B" blockId="14.[151,858,602,702]" box="[151,387,675,702]" pageId="14" pageNumber="15">
(
<figureCitation id="BDD260D64139FFDCFF0AFD59FF39FD58" box="[159,251,675,701]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="14" pageNumber="15">Figs 2C</figureCitation>
,
<figureCitation id="BDD260D64139FFDCFE93FD59FEEBFD58" box="[262,297,675,701]" captionStart="FIGURE 3" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[152,1426,183,1471]" captionTargetId="figure-18@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 3. Elements of calcareous rings, viewed from the body cavity, anterior upper.A, Massinium toyoshiomaruae sp. nov., (holotype, WMNH-INV-2014-186); B, Thyone liaoi sp. nov., (holotype, WMNH-INV-2014-182, and a paratype, WMNH-INV- 2014-184); C, T. kyushuensis sp. nov., (holotype, WMNH-INV-2015-2437); D, T. toyoshiomaruae sp. nov. (holotype, WMNH- INV-2015-2483). Abbreviations: md, mid-dorsal part; mv, mid-ventral part; -ir, interradial element; -r radial element." figureDoi="http://doi.org/10.5281/zenodo.5225378" httpUri="https://zenodo.org/record/5225378/files/figure.png" pageId="14" pageNumber="15">3C</figureCitation>
,
<figureCitation id="BDD260D64139FFDCFEA0FD59FEB9FD5B" box="[309,379,675,702]" captionStart="FIGURE 6" captionStartId="15.[151,250,1910,1934]" captionTargetBox="[175,1412,182,1886]" captionTargetId="figure-18@15.[174,1413,181,1887]" captionTargetPageId="15" captionText="FIGURE 6. Scanning electron microscope images of ossicles of Thyone kyushuensis sp. nov., (holotype, AH, WMNH-INV- 2015-2437). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal and dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225384" httpUri="https://zenodo.org/record/5225384/files/figure.png" pageId="14" pageNumber="15">6AH</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD84139FFDCFF02FD11FBC8FCAB" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="25567C534139FFDCFF02FD11FBC8FCAB" blockId="14.[151,1437,747,2034]" pageId="14" pageNumber="15">
<emphasis id="179DA0414139FFDCFF02FD11FE43FCE0" bold="true" box="[151,385,747,773]" pageId="14" pageNumber="15">Material examined.</emphasis>
<materialsCitation id="9581760E4139FFDCFE1CFD11FCD1FCCF" accessionNumber="LC528688" collectionCode="INSD" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<emphasis id="179DA0414139FFDCFE1CFD11FE32FCE0" box="[393,496,747,773]" italics="true" pageId="14" pageNumber="15">
<typeStatus id="FA52C2F14139FFDCFE1CFD11FE32FCE0" box="[393,496,747,773]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
, WMNH-INV-2015-2437 (ST-
<quantity id="E211D1B64139FFDCFCDAFD11FCBBFCE0" box="[847,889,747,774]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="14" pageNumber="15" unit="in" value="5.0">5 in</quantity>
2015, anesthetized, length
<quantity id="E211D1B64139FFDCFB3BFD11FAC2FCE0" box="[1198,1280,747,773]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="15" unit="mm" value="13.0">13 mm</quantity>
, width
<quantity id="E211D1B64139FFDCFAC1FD11FA5AFCE0" box="[1364,1432,747,773]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="15" unit="mm" value="4.0">4 mm</quantity>
,
<collectionCode id="43F8E4964139FFDCFF02FCF5FF1AFCCC" box="[151,216,783,809]" pageId="14" pageNumber="15">INSD</collectionCode>
accession number
<accessionNumber id="3ABAE1B04139FFDCFE2CFCF5FDF6FCCF" box="[441,564,783,810]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC528688" pageId="14" pageNumber="15">LC528688</accessionNumber>
, 616bps,
<figureCitation id="BDD260D64139FFDCFD30FCF5FD21FCCF" box="[677,739,783,810]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="14" pageNumber="15">Fig 2</figureCitation>
C-i)
</materialsCitation>
.
<materialsCitation id="9581760E4139FFDCFCB7FCEAFBC4FCAB" collectionCode="NSMT" pageId="14" pageNumber="15" specimenCount="1" typeStatus="paratype">
<emphasis id="179DA0414139FFDCFCB7FCEAFC4BFCCC" box="[802,905,784,809]" italics="true" pageId="14" pageNumber="15">
<typeStatus id="FA52C2F14139FFDCFCB7FCEAFC4BFCCC" box="[802,905,784,809]" pageId="14" pageNumber="15" type="paratype">Paratype</typeStatus>
</emphasis>
,
<collectionCode id="43F8E4964139FFDCFC0FFCF5FC25FCCC" box="[922,999,783,809]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="14" pageNumber="15">NSMT</collectionCode>
E-12103 (ST-
<quantity id="E211D1B64139FFDCFB1AFCF5FB08FCCC" box="[1167,1226,783,809]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="15" unit="in" value="10.0">10 in</quantity>
2015, eviscerated calcareous ring, length
<quantity id="E211D1B64139FFDCFE09FCC9FE1DFCA8" box="[412,479,819,845]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="15" unit="mm" value="6.0">6 mm</quantity>
, width
<quantity id="E211D1B64139FFDCFDA7FCC9FDB7FCA8" box="[562,629,819,845]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="mm" value="2.0">2 mm</quantity>
, eviscerated organs only,
<figureCitation id="BDD260D64139FFDCFC01FCC9FC14FCAB" box="[916,982,819,846]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="14" pageNumber="15">Fig. 2</figureCitation>
C-j)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD84139FFDCFF52FCADFC54FBE4" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="25567C534139FFDCFF52FCADFC54FBE4" blockId="14.[151,1437,747,2034]" pageId="14" pageNumber="15">
<emphasis id="179DA0414139FFDCFF52FCADFE80FC94" bold="true" box="[199,322,855,881]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
Bodywall table ossicles with regular oval or rhomboidal disc with undulant margin, high spire without crossbeam, with two pillars united distally, ending in x-shaped crown with 46 minute apical teeth on each tip. Pedicel supporting plate ossicles with narrow curved disc with two large central-most perforations and two by standing minute perforations, and 26 small distal perforations, without spire. Tentacle ossicles comprising mostly unbranching rods and/or rod-like rosettes, varying in size and shape.
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD84139FFC3FF52FBF1FBFCFEA4" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15" type="description">
<paragraph id="25567C534139FFDCFF52FBF1FD9DFB18" blockId="14.[151,1437,747,2034]" pageId="14" pageNumber="15">
<emphasis id="179DA0414139FFDCFF52FBF1FE9BFBC0" bold="true" box="[199,345,1035,1061]" pageId="14" pageNumber="15">Description.</emphasis>
Bodywall thin, soft. Living body color whitish-yellow, not altered by fixation/preservation. Tentacles 10, arranged in a single circle, medioventral two tentacles smaller than other eight. Tentacles and introvert mostly transparent in life, partially with light-brown pigmentation, but contracted and yellowish in
<typeStatus id="FA52C2F14139FFDCFB6BFBA9FAA0FB88" box="[1278,1378,1107,1133]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
. One row of short, villi surrounding oral opening. Color of oral periphery, dark grayish brown. Pedicels long, nonretractile, covering entire body at constant density, lacking in introvert region. Color of pedicels same or slightly deeper than adjacent body. Pedicels becoming gradually smaller anteriorly and posteriorly to middle of body. Ten anal papillae and five anal teeth on radii.
</paragraph>
<paragraph id="25567C534139FFDCFF52FAFDFADFFA54" blockId="14.[151,1437,747,2034]" pageId="14" pageNumber="15">
Calcareous ring long, radial and interradial main plates weakly fragmented (
<figureCitation id="BDD260D64139FFDCFB9BFAFDFBA7FAC4" box="[1038,1125,1287,1313]" captionStart="FIGURE 3" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[152,1426,183,1471]" captionTargetId="figure-18@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 3. Elements of calcareous rings, viewed from the body cavity, anterior upper.A, Massinium toyoshiomaruae sp. nov., (holotype, WMNH-INV-2014-186); B, Thyone liaoi sp. nov., (holotype, WMNH-INV-2014-182, and a paratype, WMNH-INV- 2014-184); C, T. kyushuensis sp. nov., (holotype, WMNH-INV-2015-2437); D, T. toyoshiomaruae sp. nov. (holotype, WMNH- INV-2015-2483). Abbreviations: md, mid-dorsal part; mv, mid-ventral part; -ir, interradial element; -r radial element." figureDoi="http://doi.org/10.5281/zenodo.5225378" httpUri="https://zenodo.org/record/5225378/files/figure.png" pageId="14" pageNumber="15">Fig. 3C</figureCitation>
). Radial elements elongated with an anterior projection and anterior notch, and with paired posterior, weakly fragmented prolongations. Interradial elements also long, without posterior prolongations, with anterior projection, shortest in ventro-lateral elements. One Polian vesicle in medioventral, single stone canal in mediodorsal position. Well-matured gonad situated in posterior mid-body, in single clump on dorsal mesentery, tubules not branched. Gonad lacking ossicles.
</paragraph>
<paragraph id="25567C534139FFDCFF52FA41FE89F9A4" blockId="14.[151,1437,747,2034]" pageId="14" pageNumber="15">
Bodywall with only table ossicles (
<figureCitation id="BDD260D64139FFDCFDCDFA41FD12FA30" box="[600,720,1467,1493]" captionStart="FIGURE 6" captionStartId="15.[151,250,1910,1934]" captionTargetBox="[175,1412,182,1886]" captionTargetId="figure-18@15.[174,1413,181,1887]" captionTargetPageId="15" captionText="FIGURE 6. Scanning electron microscope images of ossicles of Thyone kyushuensis sp. nov., (holotype, AH, WMNH-INV- 2015-2437). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal and dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225384" httpUri="https://zenodo.org/record/5225384/files/figure.png" pageId="14" pageNumber="15">Fig. 6F, G</figureCitation>
,
<tableCitation id="686B49E84139FFDCFD49FA41FCF1FA33" box="[732,819,1467,1494]" captionStart="TABLE 5" captionStartId="16.[151,239,703,729]" captionText="TABLE 5. Counts and measurements (Mean ± sd μm) of ossicles from one specimen of Thyone kyushuensis sp. nov." pageId="14" pageNumber="15">Table 5</tableCitation>
); disc oval or rhomboidal with undulant margin, spire high, without crossbeam, with two pillars united distally, ending in x-shaped top with 46 minute apical spines on each tip. Table disc with 517 perforations, most frequently 912, four central perforations slightly larger than peripheral ones.
</paragraph>
<paragraph id="25567C534139FFDCFF52F9B1FD3FF817" blockId="14.[151,1437,747,2034]" pageId="14" pageNumber="15">
Tentacle ossicles mostly unbranching rods and/or rod-like rosettes, clustered in small white spots of trunk and branches, varying in size and shape, also large table ossicles in basal part of tentacle of
<typeStatus id="FA52C2F14139FFDCFBF0F995FB05F96C" box="[1125,1223,1647,1673]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(
<figureCitation id="BDD260D64139FFDCFB43F995FAEDF96C" box="[1238,1327,1647,1673]" captionStart="FIGURE 6" captionStartId="15.[151,250,1910,1934]" captionTargetBox="[175,1412,182,1886]" captionTargetId="figure-18@15.[174,1413,181,1887]" captionTargetPageId="15" captionText="FIGURE 6. Scanning electron microscope images of ossicles of Thyone kyushuensis sp. nov., (holotype, AH, WMNH-INV- 2015-2437). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal and dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225384" httpUri="https://zenodo.org/record/5225384/files/figure.png" pageId="14" pageNumber="15">Fig. 6A</figureCitation>
,
<tableCitation id="686B49E84139FFDCFAACF995FA4CF96F" box="[1337,1422,1647,1674]" captionStart="TABLE 5" captionStartId="16.[151,239,703,729]" captionText="TABLE 5. Counts and measurements (Mean ± sd μm) of ossicles from one specimen of Thyone kyushuensis sp. nov." pageId="14" pageNumber="15">Table 5</tableCitation>
). Large tables with circler or oval disc, with two pillars united distally mostly with 12 crossbeams and 36 apical spines. Table disc with four large central holes and up to 94 smaller, peripheral ones (
<figureCitation id="BDD260D64139FFDCFBEEF94DFB15F934" box="[1147,1239,1719,1745]" captionStart="FIGURE 6" captionStartId="15.[151,250,1910,1934]" captionTargetBox="[175,1412,182,1886]" captionTargetId="figure-18@15.[174,1413,181,1887]" captionTargetPageId="15" captionText="FIGURE 6. Scanning electron microscope images of ossicles of Thyone kyushuensis sp. nov., (holotype, AH, WMNH-INV- 2015-2437). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal and dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225384" httpUri="https://zenodo.org/record/5225384/files/figure.png" pageId="14" pageNumber="15">Fig. 6A</figureCitation>
,
<tableCitation id="686B49E84139FFDCFB70F94DFAFEF937" box="[1253,1340,1719,1746]" captionStart="TABLE 5" captionStartId="16.[151,239,703,729]" captionText="TABLE 5. Counts and measurements (Mean ± sd μm) of ossicles from one specimen of Thyone kyushuensis sp. nov." pageId="14" pageNumber="15">Table 5</tableCitation>
). Tables not detected in the
<typeStatus id="FA52C2F14139FFDCFEE4F921FE13F910" box="[369,465,1755,1781]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
(
<tableCitation id="686B49E84139FFDCFE74F921FDFBF913" box="[481,569,1755,1782]" captionStart="TABLE 5" captionStartId="16.[151,239,703,729]" captionText="TABLE 5. Counts and measurements (Mean ± sd μm) of ossicles from one specimen of Thyone kyushuensis sp. nov." pageId="14" pageNumber="15">Table 5</tableCitation>
). Rosettes and rods with numerous short branches, with intermediate shapes between rosettes and rods common (
<figureCitation id="BDD260D64139FFDCFDA6F905FD4FF8FC" box="[563,653,1791,1817]" captionStart="FIGURE 6" captionStartId="15.[151,250,1910,1934]" captionTargetBox="[175,1412,182,1886]" captionTargetId="figure-18@15.[174,1413,181,1887]" captionTargetPageId="15" captionText="FIGURE 6. Scanning electron microscope images of ossicles of Thyone kyushuensis sp. nov., (holotype, AH, WMNH-INV- 2015-2437). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal and dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225384" httpUri="https://zenodo.org/record/5225384/files/figure.png" pageId="14" pageNumber="15">Fig. 6A</figureCitation>
,
<tableCitation id="686B49E84139FFDCFD0DF905FD2FF8FF" box="[664,749,1791,1818]" captionStart="TABLE 5" captionStartId="16.[151,239,703,729]" captionText="TABLE 5. Counts and measurements (Mean ± sd μm) of ossicles from one specimen of Thyone kyushuensis sp. nov." pageId="14" pageNumber="15">Table 5</tableCitation>
). Peri-oral skin ossicles comprising mostly rosettes, and also large tables in parts adjoining tentacle base. Rosettes, with numerous short, occasionally mulberry-like, granulated, branches. Tables with circuler or oval disc, with two pillars united distally (
<figureCitation id="BDD260D64139FFDCFC71F8BDFBFEF884" box="[996,1084,1863,1889]" captionStart="FIGURE 6" captionStartId="15.[151,250,1910,1934]" captionTargetBox="[175,1412,182,1886]" captionTargetId="figure-18@15.[174,1413,181,1887]" captionTargetPageId="15" captionText="FIGURE 6. Scanning electron microscope images of ossicles of Thyone kyushuensis sp. nov., (holotype, AH, WMNH-INV- 2015-2437). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal and dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225384" httpUri="https://zenodo.org/record/5225384/files/figure.png" pageId="14" pageNumber="15">Fig. 6B</figureCitation>
,
<tableCitation id="686B49E84139FFDCFBD2F8BDFB5EF887" box="[1095,1180,1863,1890]" captionStart="TABLE 5" captionStartId="16.[151,239,703,729]" captionText="TABLE 5. Counts and measurements (Mean ± sd μm) of ossicles from one specimen of Thyone kyushuensis sp. nov." pageId="14" pageNumber="15">Table 5</tableCitation>
). Pharyngeal villi with small rosettes, with granulated, grape-like branches (
<figureCitation id="BDD260D64139FFDCFD73F891FCFFF860" box="[742,829,1899,1925]" captionStart="FIGURE 6" captionStartId="15.[151,250,1910,1934]" captionTargetBox="[175,1412,182,1886]" captionTargetId="figure-18@15.[174,1413,181,1887]" captionTargetPageId="15" captionText="FIGURE 6. Scanning electron microscope images of ossicles of Thyone kyushuensis sp. nov., (holotype, AH, WMNH-INV- 2015-2437). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal and dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225384" httpUri="https://zenodo.org/record/5225384/files/figure.png" pageId="14" pageNumber="15">Fig. 6C</figureCitation>
,
<tableCitation id="686B49E84139FFDCFCDCF891FC5FF863" box="[841,925,1899,1926]" captionStart="TABLE 5" captionStartId="16.[151,239,703,729]" captionText="TABLE 5. Counts and measurements (Mean ± sd μm) of ossicles from one specimen of Thyone kyushuensis sp. nov." pageId="14" pageNumber="15">Table 5</tableCitation>
). Introvert with large tables and rosettes (
<figureCitation id="BDD260D64139FFDCFAFAF891FF7EF84C" captionStart="FIGURE 6" captionStartId="15.[151,250,1910,1934]" captionTargetBox="[175,1412,182,1886]" captionTargetId="figure-18@15.[174,1413,181,1887]" captionTargetPageId="15" captionText="FIGURE 6. Scanning electron microscope images of ossicles of Thyone kyushuensis sp. nov., (holotype, AH, WMNH-INV- 2015-2437). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal and dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225384" httpUri="https://zenodo.org/record/5225384/files/figure.png" pageId="14" pageNumber="15">Fig. 6D</figureCitation>
,
<tableCitation id="686B49E84139FFDCFF5DF875FEDCF84F" box="[200,286,1935,1962]" captionStart="TABLE 5" captionStartId="16.[151,239,703,729]" captionText="TABLE 5. Counts and measurements (Mean ± sd μm) of ossicles from one specimen of Thyone kyushuensis sp. nov." pageId="14" pageNumber="15">Table 5</tableCitation>
). Large table ossicles with distorted oval or rhomboidal disc, with four large central perforations and 3075 smaller peripheral ones. Rosettes with numerous short branches. These tables have smaller disc length and fewer perforations in
<typeStatus id="FA52C2F14139FFDCFE1CF82DFE2BF814" box="[393,489,2007,2033]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
than
<typeStatus id="FA52C2F14139FFDCFDBDF82DFD48F814" box="[552,650,2007,2033]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(
<tableCitation id="686B49E84139FFDCFD0DF82DFD2DF817" box="[664,751,2007,2034]" captionStart="TABLE 5" captionStartId="16.[151,239,703,729]" captionText="TABLE 5. Counts and measurements (Mean ± sd μm) of ossicles from one specimen of Thyone kyushuensis sp. nov." pageId="14" pageNumber="15">Table 5</tableCitation>
).
</paragraph>
<caption id="71962CDB4138FFDDFF02F88CFCBEF81E" ID-DOI="http://doi.org/10.5281/zenodo.5225384" ID-Zenodo-Dep="5225384" httpUri="https://zenodo.org/record/5225384/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1910,1934]" targetBox="[175,1412,182,1886]" targetPageId="15">
<paragraph id="25567C534138FFDDFF02F88CFCBEF81E" blockId="15.[151,1437,1910,2043]" pageId="15" pageNumber="16">
<emphasis id="179DA0414138FFDDFF02F88CFED6F86B" bold="true" box="[151,276,1910,1934]" pageId="15" pageNumber="16">FIGURE 6.</emphasis>
Scanning electron microscope images of ossicles of
<taxonomicName id="E2E907D04138FFDDFCA7F88DFC3FF86A" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[818,1021,1911,1935]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="kyushuensis" status="sp. nov.">
<emphasis id="179DA0414138FFDDFCA7F88DFC3FF86A" box="[818,1021,1911,1935]" italics="true" pageId="15" pageNumber="16">Thyone kyushuensis</emphasis>
</taxonomicName>
<emphasis id="179DA0414138FFDDFB91F88DFB94F86A" bold="true" box="[1028,1110,1911,1935]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="0CAE1D3A4138FFDDFB91F88DFB94F86A" box="[1028,1110,1911,1935]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (
<emphasis id="179DA0414138FFDDFBF9F88DFB01F86A" box="[1132,1219,1911,1935]" italics="true" pageId="15" pageNumber="16">holotype</emphasis>
, AH, WMNH-INV- 2015-2437). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal and dorsal side, respectively; H, the ventrolateral anal papilla.
</paragraph>
</caption>
<caption id="71962CDB4127FFC2FF02FD45FABBFD3C" box="[151,1401,703,729]" pageId="16" pageNumber="17" startId="16.[151,239,703,729]">
<paragraph id="25567C534127FFC2FF02FD45FABBFD3C" blockId="16.[151,1401,703,729]" box="[151,1401,703,729]" pageId="16" pageNumber="17">
<emphasis id="179DA0414127FFC2FF02FD45FEC8FD3C" bold="true" box="[151,266,703,729]" pageId="16" pageNumber="17">TABLE 5.</emphasis>
Counts and measurements (Mean ± sd μm) of ossicles from one specimen of
<taxonomicName id="E2E907D04127FFC2FBDDFD45FADEFD3C" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[1096,1308,703,729]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="kyushuensis" status="sp. nov.">
<emphasis id="179DA0414127FFC2FBDDFD45FADEFD3C" box="[1096,1308,703,729]" italics="true" pageId="16" pageNumber="17">Thyone kyushuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CAE1D3A4127FFC2FAB6FD45FABBFD3C" box="[1315,1401,703,729]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="25567C534127FFC2FE35FD12FE3CFA20" pageId="16" pageNumber="17">
<table id="57E98EF34127002DFF30FD12F81AFA20" box="[165,2008,744,1477]" gridcols="17" gridrows="19" pageId="16" pageNumber="17">
<tr id="9BD97E114127002DFF30FD12F81AFD1A" box="[165,2008,744,767]" gridrow="0" pageId="16" pageNumber="17" rowspan-0="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-15="1" rowspan-16="1" rowspan-4="1" rowspan-5="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D4127002DFE03FD12FD44FD1A" box="[406,646,744,767]" colspan="3" colspanRight="2" gridcol="1" gridrow="0" pageId="16" pageNumber="17">Rod and plate (or rosette)</th>
<th id="D808176D4127002DFCC1FD12FBCDFD1A" box="[852,1039,744,767]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="16" pageNumber="17">Endplate</th>
<th id="D808176D4127002DFB63FD12FA8CFD1A" box="[1270,1358,744,767]" gridcol="10" gridrow="0" pageId="16" pageNumber="17">Rosette</th>
<th id="D808176D4127002DF935FD12F93FFD1A" box="[1696,1789,744,767]" gridcol="14" gridrow="0" pageId="16" pageNumber="17">Table</th>
</tr>
<tr id="9BD97E114127002DFF30FCF5F81AFCCD" box="[165,2008,783,808]" gridrow="1" pageId="16" pageNumber="17" rowspan-1="1" rowspan-11="1" rowspan-12="1" rowspan-4="1" rowspan-9="1">
<th id="D808176D4127002DFF30FCF5FE97FCCD" box="[165,341,783,808]" gridcol="0" gridrow="1" pageId="16" pageNumber="17">Samplea</th>
<td id="D808176D4127002DFE4DFCF5FD44FCCD" box="[472,646,783,808]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="16" pageNumber="17">Length</td>
<td id="D808176D4127002DFD72FCF5FC52FCCD" box="[743,912,783,808]" colspan="2" colspanRight="1" gridcol="5" gridrow="1" pageId="16" pageNumber="17">Diameter</td>
<td id="D808176D4127002DFC46FCF5FBB4FCCD" box="[979,1142,783,808]" colspan="2" colspanRight="1" gridcol="7" gridrow="1" pageId="16" pageNumber="17">No. plate hole</td>
<td id="D808176D4127002DFB63FCF5FA8CFCCD" box="[1270,1358,783,808]" gridcol="10" gridrow="1" pageId="16" pageNumber="17">Length</td>
<td id="D808176D4127002DF9B1FCF5F93FFCCD" box="[1572,1789,783,808]" colspan="2" colspanRight="1" gridcol="13" gridrow="1" pageId="16" pageNumber="17">Disc length</td>
<td id="D808176D4127002DF8B5FCF5F81AFCCD" box="[1824,2008,783,808]" colspan="2" colspanRight="1" gridcol="15" gridrow="1" pageId="16" pageNumber="17">No.disc hole</td>
</tr>
<tr id="9BD97E114127002DFF30FCCDF81AFCAB" box="[165,2008,823,846]" gridrow="2" pageId="16" pageNumber="17" rowspan-0="1">
<td id="D808176D4127002DFE03FCCDFE6FFCAB" box="[406,429,823,846]" gridcol="1" gridrow="2" pageId="16" pageNumber="17">
<emphasis id="179DA0414127FFC2FE09FCCDFE6AFCAB" box="[412,424,823,846]" italics="true" pageId="16" pageNumber="17">n</emphasis>
</td>
<td id="D808176D4127002DFE4DFCCDFDD8FCAB" box="[472,538,823,846]" gridcol="2" gridrow="2" pageId="16" pageNumber="17">Mean</td>
<td id="D808176D4127002DFDD1FCCDFD44FCAB" box="[580,646,823,846]" gridcol="3" gridrow="2" pageId="16" pageNumber="17">Range</td>
<td id="D808176D4127002DFD26FCCDFD7DFCAB" box="[691,703,823,846]" gridcol="4" gridrow="2" pageId="16" pageNumber="17">
<emphasis id="179DA0414127FFC2FD26FCCDFD7DFCAB" box="[691,703,823,846]" italics="true" pageId="16" pageNumber="17">n</emphasis>
</td>
<td id="D808176D4127002DFD72FCCDFCE1FCAB" box="[743,803,823,846]" gridcol="5" gridrow="2" pageId="16" pageNumber="17">Mean</td>
<td id="D808176D4127002DFCC1FCCDFC52FCAB" box="[852,912,823,846]" gridcol="6" gridrow="2" pageId="16" pageNumber="17">Range</td>
<td id="D808176D4127002DFC46FCCDFBCDFCAB" box="[979,1039,823,846]" gridcol="7" gridrow="2" pageId="16" pageNumber="17">Mean</td>
<td id="D808176D4127002DFBAFFCCDFBB4FCAB" box="[1082,1142,823,846]" gridcol="8" gridrow="2" pageId="16" pageNumber="17">Range</td>
<td id="D808176D4127002DFB3BFCCDFB00FCAB" box="[1198,1218,823,846]" gridcol="9" gridrow="2" pageId="16" pageNumber="17">
<emphasis id="179DA0414127FFC2FB26FCCDFB7DFCAB" box="[1203,1215,823,846]" italics="true" pageId="16" pageNumber="17">n</emphasis>
</td>
<td id="D808176D4127002DFB63FCCDFA8CFCAB" box="[1270,1358,823,846]" gridcol="10" gridrow="2" pageId="16" pageNumber="17">Mean</td>
<td id="D808176D4127002DFAF8FCCDFA68FCAB" box="[1389,1450,823,846]" gridcol="11" gridrow="2" pageId="16" pageNumber="17">Range</td>
<td id="D808176D4127002DFA76FCCDFA35FCAB" box="[1507,1527,823,846]" gridcol="12" gridrow="2" pageId="16" pageNumber="17">
<emphasis id="179DA0414127FFC2FA72FCCDFA31FCAB" box="[1511,1523,823,846]" italics="true" pageId="16" pageNumber="17">n</emphasis>
</td>
<td id="D808176D4127002DF9B1FCCDF9BBFCAB" box="[1572,1657,823,846]" gridcol="13" gridrow="2" pageId="16" pageNumber="17">Mean</td>
<td id="D808176D4127002DF935FCCDF93FFCAB" box="[1696,1789,823,846]" gridcol="14" gridrow="2" pageId="16" pageNumber="17">Range</td>
<td id="D808176D4127002DF8B5FCCDF8ABFCAB" box="[1824,1897,823,846]" gridcol="15" gridrow="2" pageId="16" pageNumber="17">Mean</td>
<td id="D808176D4127002DF809FCCDF81AFCAB" box="[1948,2008,823,846]" gridcol="16" gridrow="2" pageId="16" pageNumber="17">Range</td>
</tr>
<tr id="9BD97E114127002DFF30FC9AF81AFC93" box="[165,2008,864,886]" gridrow="3" pageId="16" pageNumber="17">
<th id="D808176D4127002DFF30FC9AF81AFC93" box="[165,2008,864,886]" colspan="17" colspanRight="16" gridcol="0" gridrow="3" pageId="16" pageNumber="17">WMNH-INV-2015-2437</th>
</tr>
<tr id="9BD97E114127002DFF30FC72F81AFC7A" box="[165,2008,904,927]" gridrow="4" pageId="16" pageNumber="17" rowspan-10="1" rowspan-11="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D4127002DFF30FC72FE97FC7A" box="[165,341,904,927]" gridcol="0" gridrow="4" pageId="16" pageNumber="17">Tentacle</th>
<td id="D808176D4127002DFE03FC72FE6FFC7A" box="[406,429,904,927]" gridcol="1" gridrow="4" pageId="16" pageNumber="17">23</td>
<td id="D808176D4127002DFE4DFC72FD44FC7A" box="[472,646,904,927]" colspan="2" colspanRight="1" gridcol="2" gridrow="4" pageId="16" pageNumber="17">61±14 46109</td>
<td id="D808176D4127002DFD26FC72FD7DFC7A" box="[691,703,904,927]" gridcol="4" gridrow="4" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFB3BFC72FB00FC7A" box="[1198,1218,904,927]" gridcol="9" gridrow="4" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFA76FC72FA35FC7A" box="[1507,1527,904,927]" gridcol="12" gridrow="4" pageId="16" pageNumber="17">4</td>
<td id="D808176D4127002DF9B1FC72F9BBFC7A" box="[1572,1657,904,927]" gridcol="13" gridrow="4" pageId="16" pageNumber="17">117 ± 11</td>
<td id="D808176D4127002DF935FC72F93FFC7A" box="[1696,1789,904,927]" gridcol="14" gridrow="4" pageId="16" pageNumber="17">106127</td>
<td id="D808176D4127002DF8B5FC72F8ABFC7A" box="[1824,1897,904,927]" gridcol="15" gridrow="4" pageId="16" pageNumber="17">87 ± 11</td>
<td id="D808176D4127002DF809FC72F81AFC7A" box="[1948,2008,904,927]" gridcol="16" gridrow="4" pageId="16" pageNumber="17">7298</td>
</tr>
<tr id="9BD97E114127002DFF30FC55F81AFC23" box="[165,2008,943,966]" gridrow="5" pageId="16" pageNumber="17" rowspan-14="1" rowspan-16="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D4127002DFF30FC55FE97FC23" box="[165,341,943,966]" gridcol="0" gridrow="5" pageId="16" pageNumber="17">Peri-oral skin</th>
<td id="D808176D4127002DFE03FC55FE6FFC23" box="[406,429,943,966]" gridcol="1" gridrow="5" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFD26FC55FD7DFC23" box="[691,703,943,966]" gridcol="4" gridrow="5" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFB3BFC55FB00FC23" box="[1198,1218,943,966]" gridcol="9" gridrow="5" pageId="16" pageNumber="17">5</td>
<td id="D808176D4127002DFB63FC55FA8CFC23" box="[1270,1358,943,966]" gridcol="10" gridrow="5" pageId="16" pageNumber="17">31 ±15</td>
<td id="D808176D4127002DFAF8FC55FA68FC23" box="[1389,1450,943,966]" gridcol="11" gridrow="5" pageId="16" pageNumber="17">2357</td>
<td id="D808176D4127002DFA76FC55FA35FC23" box="[1507,1527,943,966]" gridcol="12" gridrow="5" pageId="16" pageNumber="17">2</td>
<td id="D808176D4127002DF9B1FC55F9BBFC23" box="[1572,1657,943,966]" gridcol="13" gridrow="5" pageId="16" pageNumber="17">87, 170</td>
<td id="D808176D4127002DF8B5FC55F8ABFC23" box="[1824,1897,943,966]" gridcol="15" gridrow="5" pageId="16" pageNumber="17">39, 112</td>
</tr>
<tr id="9BD97E114127002DFF30FC2CF81AFC08" box="[165,2008,982,1005]" gridrow="6" pageId="16" pageNumber="17" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D4127002DFF30FC2CFE97FC08" box="[165,341,982,1005]" gridcol="0" gridrow="6" pageId="16" pageNumber="17">Oesophagus skin</th>
<td id="D808176D4127002DFE03FC2CFE6FFC08" box="[406,429,982,1005]" gridcol="1" gridrow="6" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFD26FC2CFD7DFC08" box="[691,703,982,1005]" gridcol="4" gridrow="6" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFB3BFC2CFB00FC08" box="[1198,1218,982,1005]" gridcol="9" gridrow="6" pageId="16" pageNumber="17">16</td>
<td id="D808176D4127002DFB63FC2CFA8CFC08" box="[1270,1358,982,1005]" gridcol="10" gridrow="6" pageId="16" pageNumber="17">20 ± 3</td>
<td id="D808176D4127002DFAF8FC2CFA68FC08" box="[1389,1450,982,1005]" gridcol="11" gridrow="6" pageId="16" pageNumber="17">1725</td>
<td id="D808176D4127002DFA76FC2CFA35FC08" box="[1507,1527,982,1005]" gridcol="12" gridrow="6" pageId="16" pageNumber="17">0</td>
</tr>
<tr id="9BD97E114127002DFF30FC04F81AFBF0" box="[165,2008,1022,1045]" gridrow="7" pageId="16" pageNumber="17" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D4127002DFF30FC04FE97FBF0" box="[165,341,1022,1045]" gridcol="0" gridrow="7" pageId="16" pageNumber="17">Introvert skin</th>
<td id="D808176D4127002DFE03FC04FE6FFBF0" box="[406,429,1022,1045]" gridcol="1" gridrow="7" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFD26FC04FD7DFBF0" box="[691,703,1022,1045]" gridcol="4" gridrow="7" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFB3BFC04FB00FBF0" box="[1198,1218,1022,1045]" gridcol="9" gridrow="7" pageId="16" pageNumber="17">9</td>
<td id="D808176D4127002DFB63FC04FA8CFBF0" box="[1270,1358,1022,1045]" gridcol="10" gridrow="7" pageId="16" pageNumber="17">24 ± 4</td>
<td id="D808176D4127002DFAF8FC04FA68FBF0" box="[1389,1450,1022,1045]" gridcol="11" gridrow="7" pageId="16" pageNumber="17">2535</td>
<td id="D808176D4127002DFA76FC04FA35FBF0" box="[1507,1527,1022,1045]" gridcol="12" gridrow="7" pageId="16" pageNumber="17">4</td>
<td id="D808176D4127002DF9B1FC04F9BBFBF0" box="[1572,1657,1022,1045]" gridcol="13" gridrow="7" pageId="16" pageNumber="17">120 ± 27</td>
<td id="D808176D4127002DF935FC04F93FFBF0" box="[1696,1789,1022,1045]" gridcol="14" gridrow="7" pageId="16" pageNumber="17">93157</td>
<td id="D808176D4127002DF8B5FC04F8ABFBF0" box="[1824,1897,1022,1045]" gridcol="15" gridrow="7" pageId="16" pageNumber="17">63 ± 22</td>
<td id="D808176D4127002DF809FC04F81AFBF0" box="[1948,2008,1022,1045]" gridcol="16" gridrow="7" pageId="16" pageNumber="17">3485</td>
</tr>
<tr id="9BD97E114127002DFF30FBDCF81AFBD8" box="[165,2008,1062,1085]" gridrow="8" pageId="16" pageNumber="17" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1">
<th id="D808176D4127002DFF30FBDCFE97FBD8" box="[165,341,1062,1085]" gridcol="0" gridrow="8" pageId="16" pageNumber="17">Pedicels (ventral)</th>
<td id="D808176D4127002DFE03FBDCFE6FFBD8" box="[406,429,1062,1085]" gridcol="1" gridrow="8" pageId="16" pageNumber="17">25</td>
<td id="D808176D4127002DFE4DFBDCFDD8FBD8" box="[472,538,1062,1085]" gridcol="2" gridrow="8" pageId="16" pageNumber="17">70 ± 9</td>
<td id="D808176D4127002DFDD1FBDCFD44FBD8" box="[580,646,1062,1085]" gridcol="3" gridrow="8" pageId="16" pageNumber="17">5291</td>
<td id="D808176D4127002DFD26FBDCFD7DFBD8" box="[691,703,1062,1085]" gridcol="4" gridrow="8" pageId="16" pageNumber="17">5</td>
<td id="D808176D4127002DFD72FBDCFCE1FBD8" box="[743,803,1062,1085]" gridcol="5" gridrow="8" pageId="16" pageNumber="17">63 ± 5</td>
<td id="D808176D4127002DFCC1FBDCFC52FBD8" box="[852,912,1062,1085]" gridcol="6" gridrow="8" pageId="16" pageNumber="17">5568</td>
<td id="D808176D4127002DFC46FBDCFBCDFBD8" box="[979,1039,1062,1085]" gridcol="7" gridrow="8" pageId="16" pageNumber="17">60 ± 9</td>
<td id="D808176D4127002DFBAFFBDCFBB4FBD8" box="[1082,1142,1062,1085]" gridcol="8" gridrow="8" pageId="16" pageNumber="17">5070</td>
<td id="D808176D4127002DFB3BFBDCFB00FBD8" box="[1198,1218,1062,1085]" gridcol="9" gridrow="8" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFA76FBDCFA35FBD8" box="[1507,1527,1062,1085]" gridcol="12" gridrow="8" pageId="16" pageNumber="17">0</td>
</tr>
<tr id="9BD97E114127002DFF30FBB7F81AFB81" box="[165,2008,1101,1124]" gridrow="9" pageId="16" pageNumber="17" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D4127002DFF30FBB7FE97FB81" box="[165,341,1101,1124]" gridcol="0" gridrow="9" pageId="16" pageNumber="17">Bodywall (ventral)</th>
<td id="D808176D4127002DFE03FBB7FE6FFB81" box="[406,429,1101,1124]" gridcol="1" gridrow="9" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFD26FBB7FD7DFB81" box="[691,703,1101,1124]" gridcol="4" gridrow="9" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFB3BFBB7FB00FB81" box="[1198,1218,1101,1124]" gridcol="9" gridrow="9" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFA76FBB7FA35FB81" box="[1507,1527,1101,1124]" gridcol="12" gridrow="9" pageId="16" pageNumber="17">13</td>
<td id="D808176D4127002DF9B1FBB7F9BBFB81" box="[1572,1657,1101,1124]" gridcol="13" gridrow="9" pageId="16" pageNumber="17">57 ± 4</td>
<td id="D808176D4127002DF935FBB7F93FFB81" box="[1696,1789,1101,1124]" gridcol="14" gridrow="9" pageId="16" pageNumber="17">5163</td>
<td id="D808176D4127002DF8B5FBB7F8ABFB81" box="[1824,1897,1101,1124]" gridcol="15" gridrow="9" pageId="16" pageNumber="17">16 ± 1</td>
<td id="D808176D4127002DF809FBB7F81AFB81" box="[1948,2008,1101,1124]" gridcol="16" gridrow="9" pageId="16" pageNumber="17">517</td>
</tr>
<tr id="9BD97E114127002DFF30FB8EF81AFB6E" box="[165,2008,1140,1163]" gridrow="10" pageId="16" pageNumber="17" rowspan-10="1" rowspan-11="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D4127002DFF30FB8EFE97FB6E" box="[165,341,1140,1163]" gridcol="0" gridrow="10" pageId="16" pageNumber="17">Anal papilla</th>
<td id="D808176D4127002DFE03FB8EFE6FFB6E" box="[406,429,1140,1163]" gridcol="1" gridrow="10" pageId="16" pageNumber="17">7</td>
<td id="D808176D4127002DFE4DFB8EFDD8FB6E" box="[472,538,1140,1163]" gridcol="2" gridrow="10" pageId="16" pageNumber="17">74 ±10</td>
<td id="D808176D4127002DFDD1FB8EFD44FB6E" box="[580,646,1140,1163]" gridcol="3" gridrow="10" pageId="16" pageNumber="17">6291</td>
<td id="D808176D4127002DFD26FB8EFD7DFB6E" box="[691,703,1140,1163]" gridcol="4" gridrow="10" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFB3BFB8EFB00FB6E" box="[1198,1218,1140,1163]" gridcol="9" gridrow="10" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFA76FB8EFA35FB6E" box="[1507,1527,1140,1163]" gridcol="12" gridrow="10" pageId="16" pageNumber="17">3</td>
<td id="D808176D4127002DF9B1FB8EF9BBFB6E" box="[1572,1657,1140,1163]" gridcol="13" gridrow="10" pageId="16" pageNumber="17">61 ± 2</td>
<td id="D808176D4127002DF935FB8EF93FFB6E" box="[1696,1789,1140,1163]" gridcol="14" gridrow="10" pageId="16" pageNumber="17">5963</td>
<td id="D808176D4127002DF8B5FB8EF8ABFB6E" box="[1824,1897,1140,1163]" gridcol="15" gridrow="10" pageId="16" pageNumber="17">16 ± 3</td>
<td id="D808176D4127002DF809FB8EF81AFB6E" box="[1948,2008,1140,1163]" gridcol="16" gridrow="10" pageId="16" pageNumber="17">1320</td>
</tr>
<tr id="9BD97E114127002DFF30FB61F81AFB54" box="[165,2008,1179,1201]" gridrow="11" pageId="16" pageNumber="17">
<th id="D808176D4127002DFF30FB61F81AFB54" box="[165,2008,1179,1201]" colspan="17" colspanRight="16" gridcol="0" gridrow="11" pageId="16" pageNumber="17">NMST-E-12103 (eviscerated organs only)</th>
</tr>
<tr id="9BD97E114127002DFF30FB38F81AFB3C" box="[165,2008,1218,1241]" gridrow="12" pageId="16" pageNumber="17" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D4127002DFF30FB38FE97FB3C" box="[165,341,1218,1241]" gridcol="0" gridrow="12" pageId="16" pageNumber="17">Tentacle</th>
<td id="D808176D4127002DFE03FB38FE6FFB3C" box="[406,429,1218,1241]" gridcol="1" gridrow="12" pageId="16" pageNumber="17">35</td>
<td id="D808176D4127002DFE4DFB38FDD8FB3C" box="[472,538,1218,1241]" gridcol="2" gridrow="12" pageId="16" pageNumber="17">60 ±15</td>
<td id="D808176D4127002DFDD1FB38FD44FB3C" box="[580,646,1218,1241]" gridcol="3" gridrow="12" pageId="16" pageNumber="17">3394</td>
<td id="D808176D4127002DFD26FB38FD7DFB3C" box="[691,703,1218,1241]" gridcol="4" gridrow="12" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFB3BFB38FB00FB3C" box="[1198,1218,1218,1241]" gridcol="9" gridrow="12" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFA76FB38FA35FB3C" box="[1507,1527,1218,1241]" gridcol="12" gridrow="12" pageId="16" pageNumber="17">0</td>
</tr>
<tr id="9BD97E114127002DFF30FB13F81AFAE5" box="[165,2008,1257,1280]" gridrow="13" pageId="16" pageNumber="17" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D4127002DFF30FB13FE97FAE5" box="[165,341,1257,1280]" gridcol="0" gridrow="13" pageId="16" pageNumber="17">Peri-oral skin</th>
<td id="D808176D4127002DFE03FB13FE6FFAE5" box="[406,429,1257,1280]" gridcol="1" gridrow="13" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFD26FB13FD7DFAE5" box="[691,703,1257,1280]" gridcol="4" gridrow="13" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFB3BFB13FB00FAE5" box="[1198,1218,1257,1280]" gridcol="9" gridrow="13" pageId="16" pageNumber="17">11</td>
<td id="D808176D4127002DFB63FB13FA8CFAE5" box="[1270,1358,1257,1280]" gridcol="10" gridrow="13" pageId="16" pageNumber="17">30 ± 10</td>
<td id="D808176D4127002DFAF8FB13FA68FAE5" box="[1389,1450,1257,1280]" gridcol="11" gridrow="13" pageId="16" pageNumber="17">1949</td>
<td id="D808176D4127002DFA76FB13FA35FAE5" box="[1507,1527,1257,1280]" gridcol="12" gridrow="13" pageId="16" pageNumber="17">6</td>
<td id="D808176D4127002DF9B1FB13F9BBFAE5" box="[1572,1657,1257,1280]" gridcol="13" gridrow="13" pageId="16" pageNumber="17">68 ± 10</td>
<td id="D808176D4127002DF935FB13F93FFAE5" box="[1696,1789,1257,1280]" gridcol="14" gridrow="13" pageId="16" pageNumber="17">5182</td>
<td id="D808176D4127002DF8B5FB13F8ABFAE5" box="[1824,1897,1257,1280]" gridcol="15" gridrow="13" pageId="16" pageNumber="17">18 ± 5</td>
<td id="D808176D4127002DF809FB13F81AFAE5" box="[1948,2008,1257,1280]" gridcol="16" gridrow="13" pageId="16" pageNumber="17">925</td>
</tr>
<tr id="9BD97E114127002DFF30FAEBF81AFACD" box="[165,2008,1297,1320]" gridrow="14" pageId="16" pageNumber="17" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D4127002DFF30FAEBFE97FACD" box="[165,341,1297,1320]" gridcol="0" gridrow="14" pageId="16" pageNumber="17">Oesophagus skin</th>
<td id="D808176D4127002DFE03FAEBFE6FFACD" box="[406,429,1297,1320]" gridcol="1" gridrow="14" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFD26FAEBFD7DFACD" box="[691,703,1297,1320]" gridcol="4" gridrow="14" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFB3BFAEBFB00FACD" box="[1198,1218,1297,1320]" gridcol="9" gridrow="14" pageId="16" pageNumber="17">17</td>
<td id="D808176D4127002DFB63FAEBFA8CFACD" box="[1270,1358,1297,1320]" gridcol="10" gridrow="14" pageId="16" pageNumber="17">23 ± 3</td>
<td id="D808176D4127002DFAF8FAEBFA68FACD" box="[1389,1450,1297,1320]" gridcol="11" gridrow="14" pageId="16" pageNumber="17">1829</td>
<td id="D808176D4127002DFA76FAEBFA35FACD" box="[1507,1527,1297,1320]" gridcol="12" gridrow="14" pageId="16" pageNumber="17">0</td>
</tr>
<tr id="9BD97E114127002DFF30FAC2F81AFAAA" box="[165,2008,1336,1359]" gridrow="15" pageId="16" pageNumber="17" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D4127002DFF30FAC2FE97FAAA" box="[165,341,1336,1359]" gridcol="0" gridrow="15" pageId="16" pageNumber="17">Introvert skin</th>
<td id="D808176D4127002DFE03FAC2FE6FFAAA" box="[406,429,1336,1359]" gridcol="1" gridrow="15" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFD26FAC2FD7DFAAA" box="[691,703,1336,1359]" gridcol="4" gridrow="15" pageId="16" pageNumber="17">0</td>
<td id="D808176D4127002DFB3BFAC2FB00FAAA" box="[1198,1218,1336,1359]" gridcol="9" gridrow="15" pageId="16" pageNumber="17">8</td>
<td id="D808176D4127002DFB63FAC2FA8CFAAA" box="[1270,1358,1336,1359]" gridcol="10" gridrow="15" pageId="16" pageNumber="17">33 ± 6</td>
<td id="D808176D4127002DFAF8FAC2FA68FAAA" box="[1389,1450,1336,1359]" gridcol="11" gridrow="15" pageId="16" pageNumber="17">2845</td>
<td id="D808176D4127002DFA76FAC2FA35FAAA" box="[1507,1527,1336,1359]" gridcol="12" gridrow="15" pageId="16" pageNumber="17">8</td>
<td id="D808176D4127002DF9B1FAC2F9BBFAAA" box="[1572,1657,1336,1359]" gridcol="13" gridrow="15" pageId="16" pageNumber="17">78 ± 11</td>
<td id="D808176D4127002DF935FAC2F93FFAAA" box="[1696,1789,1336,1359]" gridcol="14" gridrow="15" pageId="16" pageNumber="17">6192</td>
<td id="D808176D4127002DF8B5FAC2F8ABFAAA" box="[1824,1897,1336,1359]" gridcol="15" gridrow="15" pageId="16" pageNumber="17">23 ± 7</td>
<td id="D808176D4127002DF809FAC2F81AFAAA" box="[1948,2008,1336,1359]" gridcol="16" gridrow="15" pageId="16" pageNumber="17">1535</td>
</tr>
<tr id="9BD97E114127002DFF30FAA5F81AFA93" box="[165,2008,1375,1398]" gridrow="16" pageId="16" pageNumber="17" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D4127002DFF30FAA5FE97FA93" box="[165,341,1375,1398]" gridcol="0" gridrow="16" pageId="16" pageNumber="17">Pedicels (ventral)</th>
<td id="D808176D4127002DFE03FAA5FDD8FA93" box="[406,538,1375,1398]" colspan="2" colspanRight="1" gridcol="1" gridrow="16" pageId="16" pageNumber="17">Lacking</td>
</tr>
<tr id="9BD97E114127002DFF30FA7DF81AFA7B" box="[165,2008,1415,1438]" gridrow="17" pageId="16" pageNumber="17" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D4127002DFF30FA7DFE97FA7B" box="[165,341,1415,1438]" gridcol="0" gridrow="17" pageId="16" pageNumber="17">Bodywall (ventral)</th>
<td id="D808176D4127002DFE03FA7DFDD8FA7B" box="[406,538,1415,1438]" colspan="2" colspanRight="1" gridcol="1" gridrow="17" pageId="16" pageNumber="17">Lacking</td>
</tr>
<tr id="9BD97E114127002DFF30FA54F81AFA20" box="[165,2008,1454,1477]" gridrow="18" pageId="16" pageNumber="17" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D4127002DFF30FA54FE97FA20" box="[165,341,1454,1477]" gridcol="0" gridrow="18" pageId="16" pageNumber="17">Anal papilla</th>
<td id="D808176D4127002DFE03FA54FDD8FA20" box="[406,538,1454,1477]" colspan="2" colspanRight="1" gridcol="1" gridrow="18" pageId="16" pageNumber="17">Lacking</td>
</tr>
</table>
</paragraph>
<paragraph id="25567C534127FFC2FF02FA28F8F4FA0F" blockId="16.[151,1846,1490,1514]" box="[151,1846,1490,1514]" pageId="16" pageNumber="17">
<tableNote id="D80F7DDD4127FFC2FF02FA28F8F4FA0F" box="[151,1846,1490,1514]" pageId="16" pageNumber="17" targetBox="[165,2008,744,1477]" targetPageId="16">
<superScript id="D29CD11B4127FFC2FF02FA28FEE0FA0F" attach="right" box="[151,290,1490,1514]" fontSize="5" pageId="16" pageNumber="17">aWMNH-INV</superScript>
: Invertebrate collection of the Wakayama Prefectural Museum of Natural History, NMST-E: Echinoderms collection of National Museum of Nature and Science.
</tableNote>
</paragraph>
<paragraph id="25567C534126FFC3FF52FF6DFBFCFEA4" blockId="17.[151,1437,150,1293]" pageId="17" pageNumber="18">
Pedicels with endplate and supporting plates (
<figureCitation id="BDD260D64126FFC3FD43FF6DFCECFF54" box="[726,814,151,177]" captionStart="FIGURE 6" captionStartId="15.[151,250,1910,1934]" captionTargetBox="[175,1412,182,1886]" captionTargetId="figure-18@15.[174,1413,181,1887]" captionTargetPageId="15" captionText="FIGURE 6. Scanning electron microscope images of ossicles of Thyone kyushuensis sp. nov., (holotype, AH, WMNH-INV- 2015-2437). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal and dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225384" httpUri="https://zenodo.org/record/5225384/files/figure.png" pageId="17" pageNumber="18">Fig. 6E</figureCitation>
,
<tableCitation id="686B49E84126FFC3FCAEFF6DFC50FF54" box="[827,914,150,177]" captionStart="TABLE 5" captionStartId="16.[151,239,703,729]" captionText="TABLE 5. Counts and measurements (Mean ± sd μm) of ossicles from one specimen of Thyone kyushuensis sp. nov." pageId="17" pageNumber="18">Table 5</tableCitation>
). Supporting plates with narrow, curved disc, with two large central perforations, two minute, adjacent ones, and 26 small distal ones. Endplate small, with a few, small peripheral perforations in outer rim, followed by a few large perforations in middle that separated by small clusters or pores, and with a central area of medium-sized perforations. (
<figureCitation id="BDD260D64126FFC3FB98FEF9FBA1FEF8" box="[1037,1123,259,285]" captionStart="FIGURE 6" captionStartId="15.[151,250,1910,1934]" captionTargetBox="[175,1412,182,1886]" captionTargetId="figure-18@15.[174,1413,181,1887]" captionTargetPageId="15" captionText="FIGURE 6. Scanning electron microscope images of ossicles of Thyone kyushuensis sp. nov., (holotype, AH, WMNH-INV- 2015-2437). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal and dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225384" httpUri="https://zenodo.org/record/5225384/files/figure.png" pageId="17" pageNumber="18">Fig. 6E</figureCitation>
,
<tableCitation id="686B49E84126FFC3FBFAFEF9FB06FEF8" box="[1135,1220,258,285]" captionStart="TABLE 5" captionStartId="16.[151,239,703,729]" captionText="TABLE 5. Counts and measurements (Mean ± sd μm) of ossicles from one specimen of Thyone kyushuensis sp. nov." pageId="17" pageNumber="18">Table 5</tableCitation>
). Anal papilla with irregular supporting plates (
<figureCitation id="BDD260D64126FFC3FE5BFEDDFDE5FEA4" box="[462,551,295,321]" captionStart="FIGURE 6" captionStartId="15.[151,250,1910,1934]" captionTargetBox="[175,1412,182,1886]" captionTargetId="figure-18@15.[174,1413,181,1887]" captionTargetPageId="15" captionText="FIGURE 6. Scanning electron microscope images of ossicles of Thyone kyushuensis sp. nov., (holotype, AH, WMNH-INV- 2015-2437). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal and dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225384" httpUri="https://zenodo.org/record/5225384/files/figure.png" pageId="17" pageNumber="18">Fig. 6H</figureCitation>
,
<tableCitation id="686B49E84126FFC3FDA7FEDDFD45FEA4" box="[562,647,294,321]" captionStart="TABLE 5" captionStartId="16.[151,239,703,729]" captionText="TABLE 5. Counts and measurements (Mean ± sd μm) of ossicles from one specimen of Thyone kyushuensis sp. nov." pageId="17" pageNumber="18">Table 5</tableCitation>
). Skin around anus with regular tables.
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD84126FFC3FF52FEB1FE05FBF4" pageId="17" pageNumber="18" type="discussion">
<paragraph id="25567C534126FFC3FF52FEB1FE05FBF4" blockId="17.[151,1437,150,1293]" pageId="17" pageNumber="18">
<emphasis id="179DA0414126FFC3FF52FEB1FEF9FE80" bold="true" box="[199,315,331,357]" pageId="17" pageNumber="18">Remarks.</emphasis>
This new species is based mostly on one complete specimen; however, it is readily distinguished from most previously described species of
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<emphasis id="179DA0414126FFC3FDE8FE95FD0DFE6C" box="[637,719,367,393]" italics="true" pageId="17" pageNumber="18">Thyone</emphasis>
</taxonomicName>
by the peculiar bodywall ossicles: regular tables with x-shaped crowns with 46 minute apical spines on each tip. The presence of such table with x-shaped top was also reported in
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<emphasis id="179DA0414126FFC3FF02FE4DFEABFE34" box="[151,361,439,465]" italics="true" pageId="17" pageNumber="18">Thyone pohaiensis</emphasis>
<bibRefCitation id="417801A24126FFC3FEE5FE4DFE28FE34" author="Liao, Y." box="[368,490,439,465]" pageId="17" pageNumber="18" pagination="313 - 316" refId="ref25985" refString="Liao, Y. (1986) Thyone pohaiensis, a new sea cucumber from the Bohai Sea, China. Chinese Journal of Oceanology and Limnology, 4, 313 - 316. https: // doi. org / 10.1007 / BF 02850417" type="journal article" year="1986">Liao, 1986</bibRefCitation>
</taxonomicName>
from the Bohai Sea (Bokkai) of
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, which the new species resembles both in its ossicles and anatomy. However,
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<emphasis id="179DA0414126FFC3FD94FE21FD51FE10" box="[513,659,475,501]" italics="true" pageId="17" pageNumber="18">T. pohaiensis</emphasis>
</taxonomicName>
usually has supporting tables in the pedicels, whereas the new species only has supporting plates lacking a spire. Furthermore, the tentacle ossicles of
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<emphasis id="179DA0414126FFC3FB85FE05FB61FDFC" box="[1040,1187,511,537]" italics="true" pageId="17" pageNumber="18">T. pohaiensis</emphasis>
</taxonomicName>
are comprised of rods with flat surfaces, whereas in the new species they are mostly comprised of rods as well as rod-like rosettes. From the feature of tentacle ossicles,
<typeStatus id="FA52C2F14126FFC3FE65FDBDFD92FD84" box="[496,592,583,609]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
(NSMT E-12103) is detected as this new species, although the table ossicles of peri-oral skin and introvert skin are smaller in disc length and lesser in disc perforation number than those of the
<typeStatus id="FA52C2F14126FFC3FF57FD75FEE6FD4C" box="[194,292,655,681]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
(
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),
<emphasis id="179DA0414126FFC3FE31FD75FE08FD4D" box="[420,458,655,680]" italics="true" pageId="17" pageNumber="18">viz.</emphasis>
, both species have polyporous large tables in the introvert and tentacles base, but the table disc in
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<emphasis id="179DA0414126FFC3FEBDFD49FE7FFD28" box="[296,445,691,717]" italics="true" pageId="17" pageNumber="18">T. pohaiensis</emphasis>
</taxonomicName>
is up to approximately 80 µm long and has approximately 25 perforations, whereas the table disc in the present new species is up to approximately 100150 µm long and has up to approximately 35100 perforations. Although both species resemble one another in body size, they differ in living body color:
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<emphasis id="179DA0414126FFC3FA9FFD01FA5EFCF0" box="[1290,1436,763,789]" italics="true" pageId="17" pageNumber="18">T. pohaiensis</emphasis>
</taxonomicName>
is translucent light red with brownish tentacles with body length
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and width
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, whereas the present new species is whitish-yellow with body length
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and width
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. However, our complete specimen is only one individual so that the two species can not be differentiated by these external morphology. This new species and
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<emphasis id="179DA0414126FFC3FF02FC71FEEBFC40" box="[151,297,907,933]" italics="true" pageId="17" pageNumber="18">T. pohaiensis</emphasis>
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both live in sandy bottoms; however, they differ from each other in bathymetrical distribution; because of small sample numbers also it must not be decisive, the present new species were collected from the continental shelf,
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deep in ST-5 and
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deep in ST-10, whereas
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<emphasis id="179DA0414126FFC3FCCAFC29FC31FC08" box="[863,1011,979,1005]" italics="true" pageId="17" pageNumber="18">T. pohaiensis</emphasis>
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was found in the sublittoral zone, the low-tide mark to
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deep.
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD84126FFC3FF52FBE0FC74FB98" pageId="17" pageNumber="18" type="distribution">
<paragraph id="25567C534126FFC3FF52FBE0FC74FB98" blockId="17.[151,1437,150,1293]" pageId="17" pageNumber="18">
<emphasis id="179DA0414126FFC3FF52FBE0FEA3FBD1" bold="true" box="[199,353,1050,1076]" pageId="17" pageNumber="18">Distribution.</emphasis>
Known only from two sampling localities of northern islands of off Kyushu Island of western
<collectingCountry id="5DFE3CC34126FFC3FF02FBC5FF19FBBC" box="[151,219,1087,1113]" name="Japan" pageId="17" pageNumber="18">Japan</collectingCountry>
: off Hisaka-shima Island, Goto Islands,
<collectingRegion id="E72DB2B14126FFC3FD0CFBC5FC82FBBC" box="[665,832,1087,1113]" country="Japan" name="Nagasaki" pageId="17" pageNumber="18">Nagasaki Pref.</collectingRegion>
, sand bottom, depth
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(ST-
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2015), and Genkainada Sea,
<collectingRegion id="E72DB2B14126FFC3FE9CFB99FE69FB98" box="[265,427,1123,1149]" country="Japan" name="Fukuoka" pageId="17" pageNumber="18">Fukuoka Pref.</collectingRegion>
, sand bottom, depth
<quantity id="E211D1B64126FFC3FD06FB99FD34FB98" box="[659,758,1123,1149]" metricMagnitude="1" metricUnit="m" metricValue="7.25" metricValueMax="7.3" metricValueMin="7.2" pageId="17" pageNumber="18" unit="m" value="72.5" valueMax="73.0" valueMin="72.0">7273 m</quantity>
(ST-
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2015).
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD84126FFC3FF52FB7DFDB4FAE8" pageId="17" pageNumber="18" type="etymology">
<paragraph id="25567C534126FFC3FF52FB7DFAC2FB44" blockId="17.[151,1437,150,1293]" box="[199,1280,1159,1185]" pageId="17" pageNumber="18">
<emphasis id="179DA0414126FFC3FF52FB7DFE8CFB44" bold="true" box="[199,334,1159,1185]" pageId="17" pageNumber="18">Etymology.</emphasis>
The specific name is an adjective, derived from the locality name of Kyushu Island.
</paragraph>
<paragraph id="25567C534126FFC3FF52FB51FDB4FAE8" blockId="17.[151,1437,150,1293]" pageId="17" pageNumber="18">
<emphasis id="179DA0414126FFC3FF52FB51FDD9FB21" bold="true" box="[199,539,1194,1221]" pageId="17" pageNumber="18">Molecular data comparison.</emphasis>
In BLAST searches, the closest hit to the COI sequence is from
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<emphasis id="179DA0414126FFC3FA91FB51FA5FFB21" box="[1284,1437,1195,1221]" italics="true" pageId="17" pageNumber="18">Thyone nigra</emphasis>
</taxonomicName>
(INSD accession number
<accessionNumber id="3ABAE1B04126FFC3FE26FB35FDEFFB0C" box="[435,557,1231,1257]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF142163" pageId="17" pageNumber="18">KF142163</accessionNumber>
, 82.7% similarity with 93% coverage). The results of BLAST searches supports that the present species belongs to
<emphasis id="179DA0414126FFC3FD89FB09FDB4FAE8" box="[540,630,1267,1293]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="E2E907D04126FFC3FD89FB09FDB0FAE8" authorityName="Oken" authorityYear="1815" box="[540,626,1267,1293]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">Thyone</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="179DA041413DFFD8FF02FC28FE2FFC09" bold="true" box="[151,493,978,1005]" pageId="10" pageNumber="11">
Genus
<taxonomicName id="E2E907D0413DFFD8FF7BFC28FE23FC09" ID-CoL="7Q4FF" authority="Oken, 1815" authorityName="Oken" authorityYear="1815" box="[238,481,978,1005]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="179DA041413DFFD8FF7BFC28FE8BFC08" bold="true" box="[238,329,978,1005]" italics="true" pageId="10" pageNumber="11">Thyone</emphasis>
<bibRefCitation id="417801A2413DFFD8FEC4FC28FE23FC09" author="Oken, L." box="[337,481,978,1004]" pageId="10" pageNumber="11" refId="ref26651" refString="Oken, L. (1815 - 1816) Lehrbuch der Naturgeschichte. Vol 3. Zoologie. A. Schmid &amp; Co., Jena, 841 pp." type="book" year="1815">Oken, 1815</bibRefCitation>
</taxonomicName>
*
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<paragraph id="25567C53413DFFD8FF02FC0DFD51FBF4" blockId="10.[151,659,978,1041]" box="[151,659,1014,1041]" pageId="10" pageNumber="11">[New Japanese name: Kehada-namako-zoku]</paragraph>
</subSubSection>
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*
<emphasis id="179DA041413DFFD8FF30FBC5FF26FBBC" bold="true" box="[165,228,1087,1113]" pageId="10" pageNumber="11">Note.</emphasis>
Names established in
<bibRefCitation id="417801A2413DFFD8FE77FBC5FDA5FBBC" author="Oken, L." box="[482,615,1086,1113]" pageId="10" pageNumber="11" refId="ref26651" refString="Oken, L. (1815 - 1816) Lehrbuch der Naturgeschichte. Vol 3. Zoologie. A. Schmid &amp; Co., Jena, 841 pp." type="book" year="1815">Oken (1815</bibRefCitation>
1816), including
<taxonomicName id="E2E907D0413DFFD8FCA3FBC5FC4AFBBC" authorityName="Oken" authorityYear="1815" box="[822,904,1087,1113]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="179DA041413DFFD8FCA3FBC5FC4AFBBC" box="[822,904,1087,1113]" italics="true" pageId="10" pageNumber="11">Thyone</emphasis>
</taxonomicName>
, were once ruled to be unavailable because the works did not strictly adapt the Principle of Binominal Nomenclature (International Commission on Zoological Nomenclature [ICZN] 1956). However, recently,
<bibRefCitation id="417801A2413DFFD8FD44FB7DFB85FB44" author="Paulay, G. &amp; O'Loughlin, M." box="[721,1095,1159,1185]" pageId="10" pageNumber="11" pagination="276 - 278" refId="ref26898" refString="Paulay, G. &amp; O'Loughlin, M. (2012) Case 3598: Thyone Oken, 1815 and Psolus Oken, 1815 (Echinodermata, Holothuroidea): proposed reinstatement as available names. Bulletin of Zoological Nomenclature, 69, 276 - 278. https: // doi. org / 10.21805 / bzn. v 69 i 4. a 7" type="journal article" year="2012">Paulay and OLoughlins (2012)</bibRefCitation>
Case 3598, in which
<taxonomicName id="E2E907D0413DFFD8FADFFB7DFEDCFB20" authority="Oken, 1815" authorityName="Oken" authorityYear="1815" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="179DA041413DFFD8FADFFB7DFA5EFB44" box="[1354,1436,1159,1185]" italics="true" pageId="10" pageNumber="11">Thyone</emphasis>
<bibRefCitation id="417801A2413DFFD8FF02FB51FEDCFB20" author="Oken, L." box="[151,286,1194,1221]" pageId="10" pageNumber="11" refId="ref26651" refString="Oken, L. (1815 - 1816) Lehrbuch der Naturgeschichte. Vol 3. Zoologie. A. Schmid &amp; Co., Jena, 841 pp." type="book" year="1815">Oken, 1815</bibRefCitation>
</taxonomicName>
is treated as an available and valid name within the
<taxonomicName id="E2E907D0413DFFD8FC01FB51FB8EFB20" authorityName="Grube" authorityYear="1840" box="[916,1100,1195,1221]" class="Holothuroidea" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="order">Dendrochirotida</taxonomicName>
has been accepted by ICZN (2015).
<bibRefCitation id="417801A2413DFFD8FF67FB35FE4CFB0C" author="Jaeger, G. F." box="[242,398,1231,1257]" pageId="10" pageNumber="11" refId="ref25711" refString="Jaeger, G. F. (1833) De Holothuriis. typis Gessnerianis, Zurich, 40 pp." type="book" year="1833">Jaeger (1833)</bibRefCitation>
made the dendrochirotid
<taxonomicName id="E2E907D0413DFFD8FD22FB35FCCBFB0C" authorityName="Oken" authorityYear="1815" box="[695,777,1231,1257]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="179DA041413DFFD8FD22FB35FCCBFB0C" box="[695,777,1231,1257]" italics="true" pageId="10" pageNumber="11">Thyone</emphasis>
</taxonomicName>
available; however,
<bibRefCitation id="417801A2413DFFD8FC6DFB35FB5EFB0C" author="Lesson, R. P." box="[1016,1180,1231,1257]" pageId="10" pageNumber="11" refId="ref25942" refString="Lesson, R. P. (1830) Centurie Zoologique, ou Choix d'Animaux Rares, Nouveaux ou Imparfaitement Connus. Levrault, Paris, 254 pp. https: // doi. org / 10.5962 / bhl. title. 41400" type="book" year="1830">Lesson (1830)</bibRefCitation>
had earlier erected an aspidochirotid holothurian genus with the same name:
<emphasis id="179DA041413DFFD8FC81FB09FCF4FAE9" box="[788,822,1267,1292]" italics="true" pageId="10" pageNumber="11">i.e.</emphasis>
, the aspidochirotid
<taxonomicName id="E2E907D0413DFFD8FB8BFB09FAD5FAE8" authority="Lesson, 1830" authorityName="Lesson" authorityYear="1830" box="[1054,1303,1267,1293]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="179DA041413DFFD8FB8BFB09FBB2FAE8" box="[1054,1136,1267,1293]" italics="true" pageId="10" pageNumber="11">Thyone</emphasis>
<bibRefCitation id="417801A2413DFFD8FBEEFB09FAD5FAE8" author="Lesson, R. P." box="[1147,1303,1267,1293]" pageId="10" pageNumber="11" refId="ref25942" refString="Lesson, R. P. (1830) Centurie Zoologique, ou Choix d'Animaux Rares, Nouveaux ou Imparfaitement Connus. Levrault, Paris, 254 pp. https: // doi. org / 10.5962 / bhl. title. 41400" type="book" year="1830">Lesson, 1830</bibRefCitation>
</taxonomicName>
, which had precedence over the dendrochirotid
<taxonomicName id="E2E907D0413DFFD8FDA3FAEDFCE7FAD4" authority="Jaeger, 1833" authorityName="Jaeger" authorityYear="1833" box="[566,805,1303,1329]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="179DA041413DFFD8FDA3FAEDFD4AFAD4" box="[566,648,1303,1329]" italics="true" pageId="10" pageNumber="11">Thyone</emphasis>
<bibRefCitation id="417801A2413DFFD8FD07FAEDFCE7FAD4" author="Jaeger, G. F." box="[658,805,1303,1329]" pageId="10" pageNumber="11" refId="ref25711" refString="Jaeger, G. F. (1833) De Holothuriis. typis Gessnerianis, Zurich, 40 pp." type="book" year="1833">Jaeger, 1833</bibRefCitation>
</taxonomicName>
. Despite the above ICZN ruling, however,
<taxonomicName id="E2E907D0413DFFD8FA8FFAEDFAAEFAD4" authorityName="Jaeger" authorityYear="1833" box="[1306,1388,1303,1329]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="179DA041413DFFD8FA8FFAEDFAAEFAD4" box="[1306,1388,1303,1329]" italics="true" pageId="10" pageNumber="11">Thyone</emphasis>
</taxonomicName>
has been in prevailing usage in the sense of
<taxonomicName id="E2E907D0413DFFD8FDF1FAC1FCDDFAB0" authorityName="Grube" authorityYear="1840" box="[612,799,1339,1365]" class="Holothuroidea" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="order">Dendrochirotida</taxonomicName>
, rather than Aspidochirotida. This prompted
<bibRefCitation id="417801A2413DFFD8FA8AFAC1FEAFFA9C" author="Paulay, G. &amp; O'Loughlin, M." pageId="10" pageNumber="11" pagination="276 - 278" refId="ref26898" refString="Paulay, G. &amp; O'Loughlin, M. (2012) Case 3598: Thyone Oken, 1815 and Psolus Oken, 1815 (Echinodermata, Holothuroidea): proposed reinstatement as available names. Bulletin of Zoological Nomenclature, 69, 276 - 278. https: // doi. org / 10.21805 / bzn. v 69 i 4. a 7" type="journal article" year="2012">Paulay and OLoughlin (2012)</bibRefCitation>
to ask the ICZN to make
<taxonomicName id="E2E907D0413DFFD8FD1AFAA5FCA9FA9C" authority="Oken, 1815" authorityName="Oken" authorityYear="1815" box="[655,875,1374,1401]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="179DA041413DFFD8FD1AFAA5FD23FA9C" box="[655,737,1375,1401]" italics="true" pageId="10" pageNumber="11">Thyone</emphasis>
<bibRefCitation id="417801A2413DFFD8FD7CFAA5FCA9FA9C" author="Oken, L." box="[745,875,1374,1401]" pageId="10" pageNumber="11" refId="ref26651" refString="Oken, L. (1815 - 1816) Lehrbuch der Naturgeschichte. Vol 3. Zoologie. A. Schmid &amp; Co., Jena, 841 pp." type="book" year="1815">Oken, 1815</bibRefCitation>
</taxonomicName>
available under its plenary power.
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<subSubSection id="6DF32FD8413DFFD8FF02FA30FD09F9EC" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="25567C53413DFFD8FF02FA30FCC1FA00" blockId="10.[151,771,1482,1581]" box="[151,771,1482,1509]" pageId="10" pageNumber="11">
<heading id="7E1ECB3F413DFFD8FF02FA30FCC1FA00" bold="true" box="[151,771,1482,1509]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="179DA041413DFFD8FF02FA30FCC1FA00" bold="true" box="[151,771,1482,1509]" pageId="10" pageNumber="11">
<taxonomicName id="E2E907D0413DFFD8FF02FA30FD5DFA01" ID-CoL="8RW7R" authority="Yamana, Ogawa &amp; Ohtsuka" authorityName="Yamana, Ogawa &amp; Ohtsuka" authorityYear="2021" box="[151,671,1482,1509]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="liaoi" status="sp. nov.">
<emphasis id="179DA041413DFFD8FF02FA30FEF2FA00" bold="true" box="[151,304,1482,1509]" italics="true" pageId="10" pageNumber="11">Thyone liaoi</emphasis>
Yamana, Ogawa &amp; Ohtsuka
</taxonomicName>
<taxonomicNameLabel id="0CAE1D3A413DFFD8FD33FA31FCC1FA00" box="[678,771,1483,1509]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="25567C53413DFFD8FF02FA15FD09F9EC" blockId="10.[151,771,1482,1581]" box="[151,715,1518,1545]" pageId="10" pageNumber="11">
[Japanese name: Kehada-namako (
<bibRefCitation id="417801A2413DFFD8FD89FA15FD7EF9EC" author="Yamana, Y." box="[540,700,1518,1545]" pageId="10" pageNumber="11" pagination="6 - 8" refId="ref28119" refString="Yamana, Y. (2016) Kii-suido shuhen no namako bunnrui no genjou [The present situation of the studies on Holothurioidea in the Kii Strait], The Umiushi Tsushin (ISSN 1341 - 5972), published by the Research Institute of Marine Invertebrates, Tokyo, 92, 6 - 8. [only in Japanese]" type="journal article" year="2016">Yamana 2016</bibRefCitation>
)]
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD8413DFFD8FF02F9E9FE41F9C8" box="[151,387,1554,1581]" pageId="10" pageNumber="11" type="description">
<paragraph id="25567C53413DFFD8FF02F9E9FE41F9C8" blockId="10.[151,771,1482,1581]" box="[151,387,1554,1581]" pageId="10" pageNumber="11">
(
<figureCitation id="BDD260D6413DFFD8FF0AF9E9FF39F9C8" box="[159,251,1555,1581]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="10" pageNumber="11">Figs 2B</figureCitation>
,
<figureCitation id="BDD260D6413DFFD8FE93F9E9FEEBF9C8" box="[262,297,1555,1581]" captionStart="FIGURE 3" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[152,1426,183,1471]" captionTargetId="figure-18@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 3. Elements of calcareous rings, viewed from the body cavity, anterior upper.A, Massinium toyoshiomaruae sp. nov., (holotype, WMNH-INV-2014-186); B, Thyone liaoi sp. nov., (holotype, WMNH-INV-2014-182, and a paratype, WMNH-INV- 2014-184); C, T. kyushuensis sp. nov., (holotype, WMNH-INV-2015-2437); D, T. toyoshiomaruae sp. nov. (holotype, WMNH- INV-2015-2483). Abbreviations: md, mid-dorsal part; mv, mid-ventral part; -ir, interradial element; -r radial element." figureDoi="http://doi.org/10.5281/zenodo.5225378" httpUri="https://zenodo.org/record/5225378/files/figure.png" pageId="10" pageNumber="11">3B</figureCitation>
,
<figureCitation id="BDD260D6413DFFD8FEA0F9E8FEB9F9C8" box="[309,379,1554,1581]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="10" pageNumber="11">5AH</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD8413DFFD8FF02F9ADFD26F96B" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="25567C53413DFFD8FF02F9ADFC29F98A" blockId="10.[151,1003,1623,1678]" box="[151,1003,1623,1647]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="05F95B7D413DFFD8FF02F9ADFC29F98A" box="[151,1003,1623,1647]" pageId="10" pageNumber="11">
<treatmentCitation id="A4485A42413DFFD8FF02F9ADFDA1F98A" author="Liao, Y. &amp; Clark, A. M." box="[151,611,1623,1647]" page="506" pageId="10" pageNumber="11" year="1995">
<taxonomicName id="E2E907D0413DFFD8FF02F9ADFDA1F98A" ID-CoL="56RN8" authority=": Liao &amp; Clark 1995: 505 - 506" authorityName="Liao &amp; Clark" authorityPageNumber="505 - 506" authorityYear="1995" box="[151,611,1623,1647]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="pedata">
<emphasis id="179DA041413DFFD8FF02F9ADFEEFF98A" box="[151,301,1623,1647]" italics="true" pageId="10" pageNumber="11">Thyone pedata</emphasis>
:
<bibRefCitation id="417801A2413DFFD8FEAFF9ADFDA1F98A" author="Liao, Y. &amp; Clark, A. M." box="[314,611,1623,1647]" pageId="10" pageNumber="11" refId="ref26030" refString="Liao, Y. &amp; Clark, A. M. (1995) The Echinoderms of Southern China. Science Press Beijing, 614 pp." type="book" year="1995">Liao &amp; Clark 1995: 505506</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, Fig. 307 a, b;
<treatmentCitation id="A4485A42413DFFD8FD6CF9A2FC5DF98A" author="Yamana, Y." box="[761,927,1623,1647]" page="7" pageId="10" pageNumber="11" year="2016">
<bibRefCitation id="417801A2413DFFD8FD6CF9A2FC5DF98A" author="Yamana, Y." box="[761,927,1623,1647]" pageId="10" pageNumber="11" pagination="6 - 8" refId="ref28119" refString="Yamana, Y. (2016) Kii-suido shuhen no namako bunnrui no genjou [The present situation of the studies on Holothurioidea in the Kii Strait], The Umiushi Tsushin (ISSN 1341 - 5972), published by the Research Institute of Marine Invertebrates, Tokyo, 92, 6 - 8. [only in Japanese]" type="journal article" year="2016">Yamana 2016: 7</bibRefCitation>
</treatmentCitation>
, Fig. a.
</treatmentCitationGroup>
</paragraph>
<paragraph id="25567C53413DFFD8FF02F98DFD26F96B" blockId="10.[151,1003,1623,1678]" box="[151,740,1654,1678]" pageId="10" pageNumber="11">
<emphasis id="179DA041413DFFD8FF02F98DFF7FF96B" box="[151,189,1655,1678]" italics="true" pageId="10" pageNumber="11">non</emphasis>
:
<taxonomicName id="E2E907D0413DFFD8FF5EF98CFE34F96B" ID-CoL="56RN8" authority="Semper, 1867" authorityName="Semper" authorityYear="1867" box="[203,502,1654,1678]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="pedata">
<emphasis id="179DA041413DFFD8FF5EF98CFEA2F96B" box="[203,352,1654,1678]" italics="true" pageId="10" pageNumber="11">Thyone pedata</emphasis>
<bibRefCitation id="417801A2413DFFD8FEF2F98CFE34F96B" author="Semper, C." box="[359,502,1654,1678]" pageId="10" pageNumber="11" refId="ref27545" refString="Semper, C. (1867 - 1868) Holothurien. Reisen im Archipel der Philippinen. Teil 2. Wissenschaftliche Resultate. C. W. Kreidel, Wiesbaden, 288 pp. https: // doi. org / 10.5962 / bhl. title. 11687" type="book" year="1867">Semper, 1867</bibRefCitation>
</taxonomicName>
8: p 67. (see Remarks)
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD8413DFFD8FF02F946FC26F882" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="25567C53413DFFD8FF02F946FC26F882" blockId="10.[151,1436,1724,2039]" pageId="10" pageNumber="11">
<emphasis id="179DA041413DFFD8FF02F946FE40F933" bold="true" box="[151,386,1724,1750]" pageId="10" pageNumber="11">Material examined.</emphasis>
<materialsCitation id="9581760E413DFFD8FE1EF946FBC6F91F" accessionNumber="LC547978" collectingDate="2014-11-06" collectionCode="INSD" pageId="10" pageNumber="11" specimenCount="1" typeStatus="holotype">
<emphasis id="179DA041413DFFD8FE1EF946FE30F933" box="[395,498,1724,1750]" italics="true" pageId="10" pageNumber="11">
<typeStatus id="FA52C2F1413DFFD8FE1EF946FE30F933" box="[395,498,1724,1750]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
, WMNH-INV-2014-182 (ST-
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2014,
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<collectingDate id="4113A37B413DFFD8FC54F946FB53F933" box="[961,1169,1724,1750]" pageId="10" pageNumber="11" value="2014-11-06">6 November 2014</collectingDate>
</date>
, anesthetized length
<quantity id="E211D1B6413DFFD8FA15F946FF07F91E" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="10" pageNumber="11" unit="mm" value="36.0">36 mm</quantity>
, width
<quantity id="E211D1B6413DFFD8FE89F91AFEADF91E" box="[284,367,1760,1787]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="mm" value="10.0">10 mm</quantity>
,
<collectionCode id="43F8E496413DFFD8FEE8F91AFE7CF91F" box="[381,446,1760,1786]" pageId="10" pageNumber="11">INSD</collectionCode>
accession number
<accessionNumber id="3ABAE1B0413DFFD8FD0FF91AFCD7F91E" box="[666,789,1760,1787]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC547978" pageId="10" pageNumber="11">LC547978</accessionNumber>
, 622 bps,
<figureCitation id="BDD260D6413DFFD8FC19F91AFC0DF91F" box="[908,975,1760,1786]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
B-e)
</materialsCitation>
.
<emphasis id="179DA041413DFFD8FB84F91BFB41F91F" box="[1041,1155,1761,1786]" italics="true" pageId="10" pageNumber="11">
<typeStatus id="FA52C2F1413DFFD8FB84F91BFB41F91F" box="[1041,1155,1761,1786]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
</emphasis>
: WMNH-INV-2014-183 (ST-
<quantity id="E211D1B6413DFFD8FF5CF8FEFF34F8FB" box="[201,246,1796,1822]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="10" pageNumber="11" unit="in" value="7.0">7 in</quantity>
2014,
<date id="51575A93413DFFD8FEDFF8FEFDDCF8FB" box="[330,542,1796,1822]" pageId="10" pageNumber="11" value="2014-11-06">6 November 2014</date>
, length
<quantity id="E211D1B6413DFFD8FDE9F8FEFD13F8FA" box="[636,721,1796,1823]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="10" pageNumber="11" unit="mm" value="39.0">39 mm</quantity>
, width
<quantity id="E211D1B6413DFFD8FCBFF8FEFCB2F8FA" box="[810,880,1796,1823]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="mm" value="9.0">9 mm</quantity>
, eviscerated,
<figureCitation id="BDD260D6413DFFD8FB9BF8FEFB91F8FA" box="[1038,1107,1796,1823]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
B-f); WMNH-INV-2014-184 (ST-
<quantity id="E211D1B6413DFFD8FF5CF8D2FF31F8A7" box="[201,243,1832,1858]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="10" pageNumber="11" unit="in" value="7.0">7 in</quantity>
2014,
<date id="51575A93413DFFD8FEAAF8D2FDCEF8A7" box="[319,524,1832,1858]" pageId="10" pageNumber="11" value="2014-11-06">6 November 2014</date>
, length
<quantity id="E211D1B6413DFFD8FDF6F8D2FD76F8A6" box="[611,692,1832,1859]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="10" pageNumber="11" unit="mm" value="34.0">34 mm</quantity>
, width
<quantity id="E211D1B6413DFFD8FC93F8D2FC8BF8A6" box="[774,841,1832,1859]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="mm" value="9.0">9 mm</quantity>
,
<figureCitation id="BDD260D6413DFFD8FCC1F8D2FC57F8A7" box="[852,917,1832,1858]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
B-g); WMNH-INV-2014-188 (ST-
<quantity id="E211D1B6413DFFD8FA82F8D2FA80F8A7" box="[1303,1346,1832,1858]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="10" pageNumber="11" unit="in" value="4.0">4 in</quantity>
2014,
<date id="51575A93413DFFD8FA1BF8D2FE8DF883" pageId="10" pageNumber="11" value="2014-11-05">5 November 2014</date>
, anterior half only, length
<quantity id="E211D1B6413DFFD8FDE3F8B6FD05F882" box="[630,711,1868,1895]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="11" unit="mm" value="13.0">13 mm</quantity>
, width
<quantity id="E211D1B6413DFFD8FC8CF8B6FC9EF882" box="[793,860,1868,1895]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="11" unit="mm" value="5.0">5 mm</quantity>
,
<figureCitation id="BDD260D6413DFFD8FCFDF8B6FC6BF882" box="[872,937,1868,1895]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
B-h).
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD8413DFFD8FF52F88AFC1FF812" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="25567C53413DFFD8FF52F88AFC1FF812" blockId="10.[151,1436,1724,2039]" pageId="10" pageNumber="11">
<emphasis id="179DA041413DFFD8FF52F88AFE80F86F" bold="true" box="[199,322,1904,1930]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
Bodywall table ossicles with distorted oval or rhomboidal disc and delicate spire, mostly with one pillar formed by fusion of two pillars and spire with 46 minute apical spines. Pedicel supporting table ossicles with narrow curved disc with 24 large central perforations and 28 small distal perforations; spire high, with two (occasionally three) pillars united distally, ending in 24 minute processes.
</paragraph>
</subSubSection>
<caption id="71962CDB413CFFD9FF02F89AFC92F801" ID-DOI="http://doi.org/10.5281/zenodo.5225382" ID-Zenodo-Dep="5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1888,1912]" targetBox="[182,1404,182,1863]" targetPageId="11">
<paragraph id="25567C53413CFFD9FF02F89AFC92F801" blockId="11.[151,1436,1888,2020]" pageId="11" pageNumber="12">
<emphasis id="179DA041413CFFD9FF02F89AFED7F89D" bold="true" box="[151,277,1888,1912]" pageId="11" pageNumber="12">FIGURE 5.</emphasis>
Scanning electron microscope images of ossicles of
<taxonomicName id="E2E907D0413CFFD9FCA9F89AFC7EF89D" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[828,956,1888,1912]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="11" pageNumber="12" phylum="Echinodermata" rank="species" species="liaoi" status="sp. nov.">
<emphasis id="179DA041413CFFD9FCA9F89AFC7EF89D" box="[828,956,1888,1912]" italics="true" pageId="11" pageNumber="12">Thyone liaoi</emphasis>
</taxonomicName>
<emphasis id="179DA041413CFFD9FC50F89AFBD5F89D" bold="true" box="[965,1047,1888,1912]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="0CAE1D3A413CFFD9FC50F89AFBD5F89D" box="[965,1047,1888,1912]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (
<emphasis id="179DA041413CFFD9FBBAF89AFB44F89D" box="[1071,1158,1888,1912]" italics="true" pageId="11" pageNumber="12">holotype</emphasis>
, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla.
</paragraph>
</caption>
<caption id="71962CDB413BFFDEFF02FE1FFAF9FE1A" box="[151,1339,485,511]" pageId="12" pageNumber="13" startId="12.[151,239,485,511]">
<paragraph id="25567C53413BFFDEFF02FE1FFAF9FE1A" blockId="12.[151,1339,485,511]" box="[151,1339,485,511]" pageId="12" pageNumber="13">
<emphasis id="179DA041413BFFDEFF02FE1FFEC8FE1A" bold="true" box="[151,266,485,511]" pageId="12" pageNumber="13">TABLE 4.</emphasis>
Counts and measurements (Mean ± sd μm) of ossicles from four specimens of
<taxonomicName id="E2E907D0413BFFDEFBCCFE1FFB1DFE1A" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[1113,1247,485,511]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="12" pageNumber="13" phylum="Echinodermata" rank="species" species="liaoi" status="sp. nov.">
<emphasis id="179DA041413BFFDEFBCCFE1FFB1DFE1A" box="[1113,1247,485,511]" italics="true" pageId="12" pageNumber="13">Thyone liaoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CAE1D3A413BFFDEFB70FE1FFAF9FE1A" box="[1253,1339,485,511]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="25567C53413BFFDEFE2AFDF7FDF3F976" pageId="12" pageNumber="13">
<table id="57E98EF3413B002DFF37FDF7F829F976" box="[162,2027,525,1683]" gridcols="19" gridrows="35" pageId="12" pageNumber="13">
<tr id="9BD97E11413B002DFF37FDF7F829FDC1" box="[162,2027,525,548]" gridrow="0" pageId="12" pageNumber="13" rowspan-0="1" rowspan-1="1" rowspan-12="1" rowspan-18="1" rowspan-4="1" rowspan-5="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413B002DFE28FDF7FDB7FDC1" box="[445,629,525,548]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="12" pageNumber="13">Plate and rod</th>
<th id="D808176D413B002DFCA9FDF7FC37FDC1" box="[828,1013,525,548]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="12" pageNumber="13">Endplate</th>
<th id="D808176D413B002DFB2CFDF7FA8DFDC1" box="[1209,1359,525,548]" colspan="2" colspanRight="1" gridcol="10" gridrow="0" pageId="12" pageNumber="13">Rosette</th>
<th id="D808176D413B002DFA2CFDF7F852FDC1" box="[1465,1936,525,548]" colspan="5" colspanRight="4" gridcol="13" gridrow="0" pageId="12" pageNumber="13">Table (T) and supporting table (ST)</th>
</tr>
<tr id="9BD97E11413B002DFF37FDD6F829FDA0" box="[162,2027,556,581]" gridrow="1" pageId="12" pageNumber="13" rowspan-1="1" rowspan-12="1" rowspan-4="1" rowspan-9="1">
<th id="D808176D413B002DFF37FDD6FE9BFDA0" box="[162,345,556,581]" gridcol="0" gridrow="1" pageId="12" pageNumber="13">Samplea</th>
<td id="D808176D413B002DFE28FDD6FDB7FDA0" box="[445,629,556,581]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="12" pageNumber="13">Length</td>
<td id="D808176D413B002DFD46FDD6FCBCFDA0" box="[723,894,556,581]" colspan="2" colspanRight="1" gridcol="5" gridrow="1" pageId="12" pageNumber="13">Diameter</td>
<td id="D808176D413B002DFC24FDD6FB98FDA0" box="[945,1114,556,581]" colspan="2" colspanRight="1" gridcol="7" gridrow="1" pageId="12" pageNumber="13">No. plate hole</td>
<td id="D808176D413B002DFB2CFDD6FA8DFDA0" box="[1209,1359,556,581]" colspan="2" colspanRight="1" gridcol="10" gridrow="1" pageId="12" pageNumber="13">Length</td>
<td id="D808176D413B002DFA2CFDD6F9AAFDA0" box="[1465,1640,556,581]" colspan="2" colspanRight="1" gridcol="13" gridrow="1" pageId="12" pageNumber="13">Disc length</td>
<td id="D808176D413B002DF905FDD6F8EDFDA0" box="[1680,1839,556,581]" colspan="2" colspanRight="1" gridcol="15" gridrow="1" pageId="12" pageNumber="13">No. disc hole (T)</td>
<td id="D808176D413B002DF8CDFDD6F829FDA0" box="[1880,2027,556,581]" colspan="2" colspanRight="1" gridcol="17" gridrow="1" pageId="12" pageNumber="13">Spire (ST)</td>
</tr>
<tr id="9BD97E11413B002DFF37FDB5F829FD80" box="[162,2027,591,613]" gridrow="2" pageId="12" pageNumber="13" rowspan-0="1">
<td id="D808176D413B002DFE1CFDB5FE5DFD80" box="[393,415,591,613]" gridcol="1" gridrow="2" pageId="12" pageNumber="13">
<emphasis id="179DA041413BFFDEFE1BFDB5FE58FD80" box="[398,410,591,613]" italics="true" pageId="12" pageNumber="13">n</emphasis>
</td>
<td id="D808176D413B002DFE28FDB5FDD7FD80" box="[445,533,591,613]" gridcol="2" gridrow="2" pageId="12" pageNumber="13">Mean</td>
<td id="D808176D413B002DFDA4FDB5FDB7FD80" box="[561,629,591,613]" gridcol="3" gridrow="2" pageId="12" pageNumber="13">Range</td>
<td id="D808176D413B002DFD31FDB5FD72FD80" box="[676,688,591,613]" gridcol="4" gridrow="2" pageId="12" pageNumber="13">
<emphasis id="179DA041413BFFDEFD31FDB5FD72FD80" box="[676,688,591,613]" italics="true" pageId="12" pageNumber="13">n</emphasis>
</td>
<td id="D808176D413B002DFD46FDB5FCD4FD80" box="[723,790,591,613]" gridcol="5" gridrow="2" pageId="12" pageNumber="13">Mean</td>
<td id="D808176D413B002DFCA9FDB5FCBCFD80" box="[828,894,591,613]" gridcol="6" gridrow="2" pageId="12" pageNumber="13">Range</td>
<td id="D808176D413B002DFC24FDB5FC37FD80" box="[945,1013,591,613]" gridcol="7" gridrow="2" pageId="12" pageNumber="13">Mean</td>
<td id="D808176D413B002DFB8DFDB5FB98FD80" box="[1048,1114,591,613]" gridcol="8" gridrow="2" pageId="12" pageNumber="13">Range</td>
<td id="D808176D413B002DFB11FDB5FB59FD80" box="[1156,1179,591,613]" gridcol="9" gridrow="2" pageId="12" pageNumber="13">
<emphasis id="179DA041413BFFDEFB1CFDB5FB57FD80" box="[1161,1173,591,613]" italics="true" pageId="12" pageNumber="13">n</emphasis>
</td>
<td id="D808176D413B002DFB2CFDB5FB36FD80" box="[1209,1268,591,613]" gridcol="10" gridrow="2" pageId="12" pageNumber="13">Mean</td>
<td id="D808176D413B002DFA87FDB5FA8DFD80" box="[1298,1359,591,613]" gridcol="11" gridrow="2" pageId="12" pageNumber="13">Range</td>
<td id="D808176D413B002DFAEBFDB5FA56FD80" box="[1406,1428,591,613]" gridcol="12" gridrow="2" pageId="12" pageNumber="13">
<emphasis id="179DA041413BFFDEFA16FDB5FA4DFD80" box="[1411,1423,591,613]" italics="true" pageId="12" pageNumber="13">n</emphasis>
</td>
<td id="D808176D413B002DFA2CFDB5F9C6FD80" box="[1465,1540,591,613]" gridcol="13" gridrow="2" pageId="12" pageNumber="13">Mean</td>
<td id="D808176D413B002DF9B6FDB5F9AAFD80" box="[1571,1640,591,613]" gridcol="14" gridrow="2" pageId="12" pageNumber="13">Range</td>
<td id="D808176D413B002DF905FDB5F909FD80" box="[1680,1739,591,613]" gridcol="15" gridrow="2" pageId="12" pageNumber="13">Mean</td>
<td id="D808176D413B002DF978FDB5F8EDFD80" box="[1773,1839,591,613]" gridcol="16" gridrow="2" pageId="12" pageNumber="13">Range</td>
<td id="D808176D413B002DF8CDFDB5F852FD80" box="[1880,1936,591,613]" gridcol="17" gridrow="2" pageId="12" pageNumber="13">Mean</td>
<td id="D808176D413B002DF825FDB5F829FD80" box="[1968,2027,591,613]" gridcol="18" gridrow="2" pageId="12" pageNumber="13">Range</td>
</tr>
<tr id="9BD97E11413B002DFF37FD8AF829FD62" box="[162,2027,624,647]" gridrow="3" pageId="12" pageNumber="13">
<th id="D808176D413B002DFF37FD8AF829FD62" box="[162,2027,624,647]" colspan="19" colspanRight="18" gridcol="0" gridrow="3" pageId="12" pageNumber="13">WMNH-INV-2014-182</th>
</tr>
<tr id="9BD97E11413B002DFF37FD6BF829FD4D" box="[162,2027,657,680]" gridrow="4" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FD6BFE9BFD4D" box="[162,345,657,680]" gridcol="0" gridrow="4" pageId="12" pageNumber="13">Tentacle</th>
<td id="D808176D413B002DFE1CFD6BFE5DFD4D" box="[393,415,657,680]" gridcol="1" gridrow="4" pageId="12" pageNumber="13">30</td>
<td id="D808176D413B002DFE28FD6BFDD7FD4D" box="[445,533,657,680]" gridcol="2" gridrow="4" pageId="12" pageNumber="13">139 ± 108</td>
<td id="D808176D413B002DFDA4FD6BFDB7FD4D" box="[561,629,657,680]" gridcol="3" gridrow="4" pageId="12" pageNumber="13">36432</td>
<td id="D808176D413B002DFD31FD6BFD72FD4D" box="[676,688,657,680]" gridcol="4" gridrow="4" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FD6BFB59FD4D" box="[1156,1179,657,680]" gridcol="9" gridrow="4" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFD6BFA56FD4D" box="[1406,1428,657,680]" gridcol="12" gridrow="4" pageId="12" pageNumber="13">4</td>
<td id="D808176D413B002DFA2CFD6BF9C6FD4D" box="[1465,1540,657,680]" gridcol="13" gridrow="4" pageId="12" pageNumber="13">147 ± 100</td>
<td id="D808176D413B002DF9B6FD6BF9AAFD4D" box="[1571,1640,657,680]" gridcol="14" gridrow="4" pageId="12" pageNumber="13">77294</td>
<td id="D808176D413B002DF905FD6BF909FD4D" box="[1680,1739,657,680]" gridcol="15" gridrow="4" pageId="12" pageNumber="13">19 ± 11</td>
<td id="D808176D413B002DF978FD6BF8EDFD4D" box="[1773,1839,657,680]" gridcol="16" gridrow="4" pageId="12" pageNumber="13">1035</td>
</tr>
<tr id="9BD97E11413B002DFF37FD4EF829FD2E" box="[162,2027,692,715]" gridrow="5" pageId="12" pageNumber="13" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FD4EFE9BFD2E" box="[162,345,692,715]" gridcol="0" gridrow="5" pageId="12" pageNumber="13">Peri-oral skin</th>
<td id="D808176D413B002DFE1CFD4EFE5DFD2E" box="[393,415,692,715]" gridcol="1" gridrow="5" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FD4EFD72FD2E" box="[676,688,692,715]" gridcol="4" gridrow="5" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FD4EFB59FD2E" box="[1156,1179,692,715]" gridcol="9" gridrow="5" pageId="12" pageNumber="13">25</td>
<td id="D808176D413B002DFB2CFD4EFB36FD2E" box="[1209,1268,692,715]" gridcol="10" gridrow="5" pageId="12" pageNumber="13">37±11</td>
<td id="D808176D413B002DFA87FD4EFA8DFD2E" box="[1298,1359,692,715]" gridcol="11" gridrow="5" pageId="12" pageNumber="13">2369</td>
<td id="D808176D413B002DFAEBFD4EFA56FD2E" box="[1406,1428,692,715]" gridcol="12" gridrow="5" pageId="12" pageNumber="13">6</td>
<td id="D808176D413B002DFA2CFD4EF9C6FD2E" box="[1465,1540,692,715]" gridcol="13" gridrow="5" pageId="12" pageNumber="13">104 ± 11</td>
<td id="D808176D413B002DF9B6FD4EF9AAFD2E" box="[1571,1640,692,715]" gridcol="14" gridrow="5" pageId="12" pageNumber="13">82112</td>
<td id="D808176D413B002DF905FD4EF909FD2E" box="[1680,1739,692,715]" gridcol="15" gridrow="5" pageId="12" pageNumber="13">20 ± 4</td>
<td id="D808176D413B002DF978FD4EF8EDFD2E" box="[1773,1839,692,715]" gridcol="16" gridrow="5" pageId="12" pageNumber="13">1524</td>
</tr>
<tr id="9BD97E11413B002DFF37FD2DF829FD0B" box="[162,2027,727,750]" gridrow="6" pageId="12" pageNumber="13" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FD2DFE9BFD0B" box="[162,345,727,750]" gridcol="0" gridrow="6" pageId="12" pageNumber="13">Oesophagus skin</th>
<td id="D808176D413B002DFE1CFD2DFE5DFD0B" box="[393,415,727,750]" gridcol="1" gridrow="6" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FD2DFD72FD0B" box="[676,688,727,750]" gridcol="4" gridrow="6" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FD2DFB59FD0B" box="[1156,1179,727,750]" gridcol="9" gridrow="6" pageId="12" pageNumber="13">10</td>
<td id="D808176D413B002DFB2CFD2DFB36FD0B" box="[1209,1268,727,750]" gridcol="10" gridrow="6" pageId="12" pageNumber="13">36 ± 5</td>
<td id="D808176D413B002DFA87FD2DFA8DFD0B" box="[1298,1359,727,750]" gridcol="11" gridrow="6" pageId="12" pageNumber="13">2743</td>
<td id="D808176D413B002DFAEBFD2DFA56FD0B" box="[1406,1428,727,750]" gridcol="12" gridrow="6" pageId="12" pageNumber="13">0</td>
</tr>
<tr id="9BD97E11413B002DFF37FD03F829FCF5" box="[162,2027,761,784]" gridrow="7" pageId="12" pageNumber="13" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FD03FE9BFCF5" box="[162,345,761,784]" gridcol="0" gridrow="7" pageId="12" pageNumber="13">Introvert skin</th>
<td id="D808176D413B002DFE1CFD03FE5DFCF5" box="[393,415,761,784]" gridcol="1" gridrow="7" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FD03FD72FCF5" box="[676,688,761,784]" gridcol="4" gridrow="7" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FD03FB59FCF5" box="[1156,1179,761,784]" gridcol="9" gridrow="7" pageId="12" pageNumber="13">4</td>
<td id="D808176D413B002DFB2CFD03FB36FCF5" box="[1209,1268,761,784]" gridcol="10" gridrow="7" pageId="12" pageNumber="13">47 ± 13</td>
<td id="D808176D413B002DFA87FD03FA8DFCF5" box="[1298,1359,761,784]" gridcol="11" gridrow="7" pageId="12" pageNumber="13">3362</td>
<td id="D808176D413B002DFAEBFD03FA56FCF5" box="[1406,1428,761,784]" gridcol="12" gridrow="7" pageId="12" pageNumber="13">12</td>
<td id="D808176D413B002DFA2CFD03F9C6FCF5" box="[1465,1540,761,784]" gridcol="13" gridrow="7" pageId="12" pageNumber="13">88 ± 11</td>
<td id="D808176D413B002DF9B6FD03F9AAFCF5" box="[1571,1640,761,784]" gridcol="14" gridrow="7" pageId="12" pageNumber="13">68108</td>
<td id="D808176D413B002DF905FD03F909FCF5" box="[1680,1739,761,784]" gridcol="15" gridrow="7" pageId="12" pageNumber="13">17 ± 4</td>
<td id="D808176D413B002DF978FD03F8EDFCF5" box="[1773,1839,761,784]" gridcol="16" gridrow="7" pageId="12" pageNumber="13">1023</td>
</tr>
<tr id="9BD97E11413B002DFF37FCE0F829FCD4" box="[162,2027,794,817]" gridrow="8" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1">
<th id="D808176D413B002DFF37FCE0FE9BFCD4" box="[162,345,794,817]" gridcol="0" gridrow="8" pageId="12" pageNumber="13">Pedicels (ventral)</th>
<td id="D808176D413B002DFE1CFCE0FE5DFCD4" box="[393,415,794,817]" gridcol="1" gridrow="8" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FCE0FD72FCD4" box="[676,688,794,817]" gridcol="4" gridrow="8" pageId="12" pageNumber="13">5</td>
<td id="D808176D413B002DFD46FCE0FCD4FCD4" box="[723,790,794,817]" gridcol="5" gridrow="8" pageId="12" pageNumber="13">176 ± 3</td>
<td id="D808176D413B002DFCA9FCE0FCBCFCD4" box="[828,894,794,817]" gridcol="6" gridrow="8" pageId="12" pageNumber="13">172179</td>
<td id="D808176D413B002DFC24FCE0FC37FCD4" box="[945,1013,794,817]" gridcol="7" gridrow="8" pageId="12" pageNumber="13">193 ± 14</td>
<td id="D808176D413B002DFB8DFCE0FB98FCD4" box="[1048,1114,794,817]" gridcol="8" gridrow="8" pageId="12" pageNumber="13">171209</td>
<td id="D808176D413B002DFB11FCE0FB59FCD4" box="[1156,1179,794,817]" gridcol="9" gridrow="8" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFCE0FA56FCD4" box="[1406,1428,794,817]" gridcol="12" gridrow="8" pageId="12" pageNumber="13">24</td>
<td id="D808176D413B002DFA2CFCE0F9C6FCD4" box="[1465,1540,794,817]" gridcol="13" gridrow="8" pageId="12" pageNumber="13">90 ± 6</td>
<td id="D808176D413B002DF9B6FCE0F9AAFCD4" box="[1571,1640,794,817]" gridcol="14" gridrow="8" pageId="12" pageNumber="13">78100</td>
<td id="D808176D413B002DF8CDFCE0F852FCD4" box="[1880,1936,794,817]" gridcol="17" gridrow="8" pageId="12" pageNumber="13">37 ± 4</td>
<td id="D808176D413B002DF825FCE0F829FCD4" box="[1968,2027,794,817]" gridcol="18" gridrow="8" pageId="12" pageNumber="13">2945</td>
</tr>
<tr id="9BD97E11413B002DFF37FCC1F829FCB7" box="[162,2027,827,850]" gridrow="9" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FCC1FE9BFCB7" box="[162,345,827,850]" gridcol="0" gridrow="9" pageId="12" pageNumber="13">Bodywall (ventral)</th>
<td id="D808176D413B002DFE1CFCC1FE5DFCB7" box="[393,415,827,850]" gridcol="1" gridrow="9" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FCC1FD72FCB7" box="[676,688,827,850]" gridcol="4" gridrow="9" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FCC1FB59FCB7" box="[1156,1179,827,850]" gridcol="9" gridrow="9" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFCC1FA56FCB7" box="[1406,1428,827,850]" gridcol="12" gridrow="9" pageId="12" pageNumber="13">16</td>
<td id="D808176D413B002DFA2CFCC1F9C6FCB7" box="[1465,1540,827,850]" gridcol="13" gridrow="9" pageId="12" pageNumber="13">86 ± 12</td>
<td id="D808176D413B002DF9B6FCC1F9AAFCB7" box="[1571,1640,827,850]" gridcol="14" gridrow="9" pageId="12" pageNumber="13">68109</td>
<td id="D808176D413B002DF905FCC1F909FCB7" box="[1680,1739,827,850]" gridcol="15" gridrow="9" pageId="12" pageNumber="13">9 ± 2</td>
<td id="D808176D413B002DF978FCC1F8EDFCB7" box="[1773,1839,827,850]" gridcol="16" gridrow="9" pageId="12" pageNumber="13">513</td>
</tr>
<tr id="9BD97E11413B002DFF37FCA7F829FC91" box="[162,2027,861,884]" gridrow="10" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FCA7FE9BFC91" box="[162,345,861,884]" gridcol="0" gridrow="10" pageId="12" pageNumber="13">Anal papilla</th>
<td id="D808176D413B002DFE1CFCA7FE5DFC91" box="[393,415,861,884]" gridcol="1" gridrow="10" pageId="12" pageNumber="13">7</td>
<td id="D808176D413B002DFE28FCA7FDD7FC91" box="[445,533,861,884]" gridcol="2" gridrow="10" pageId="12" pageNumber="13">74 ± 10</td>
<td id="D808176D413B002DFDA4FCA7FDB7FC91" box="[561,629,861,884]" gridcol="3" gridrow="10" pageId="12" pageNumber="13">6291</td>
<td id="D808176D413B002DFD31FCA7FD72FC91" box="[676,688,861,884]" gridcol="4" gridrow="10" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FCA7FB59FC91" box="[1156,1179,861,884]" gridcol="9" gridrow="10" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFCA7FA56FC91" box="[1406,1428,861,884]" gridcol="12" gridrow="10" pageId="12" pageNumber="13">4</td>
<td id="D808176D413B002DFA2CFCA7F9C6FC91" box="[1465,1540,861,884]" gridcol="13" gridrow="10" pageId="12" pageNumber="13">63 ± 3</td>
<td id="D808176D413B002DF9B6FCA7F9AAFC91" box="[1571,1640,861,884]" gridcol="14" gridrow="10" pageId="12" pageNumber="13">6067</td>
<td id="D808176D413B002DF905FCA7F909FC91" box="[1680,1739,861,884]" gridcol="15" gridrow="10" pageId="12" pageNumber="13">16 ± 3</td>
<td id="D808176D413B002DF978FCA7F8EDFC91" box="[1773,1839,861,884]" gridcol="16" gridrow="10" pageId="12" pageNumber="13">1320</td>
</tr>
<tr id="9BD97E11413B002DFF37FC84F829FC70" box="[162,2027,894,917]" gridrow="11" pageId="12" pageNumber="13">
<th id="D808176D413B002DFF37FC84F829FC70" box="[162,2027,894,917]" colspan="19" colspanRight="18" gridcol="0" gridrow="11" pageId="12" pageNumber="13">WMNH-INV-2014-183</th>
</tr>
<tr id="9BD97E11413B002DFF37FC5AF829FC52" box="[162,2027,928,951]" gridrow="12" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FC5AFE9BFC52" box="[162,345,928,951]" gridcol="0" gridrow="12" pageId="12" pageNumber="13">Tentacle</th>
<td id="D808176D413B002DFE1CFC5AFE5DFC52" box="[393,415,928,951]" gridcol="1" gridrow="12" pageId="12" pageNumber="13">29</td>
<td id="D808176D413B002DFE28FC5AFDD7FC52" box="[445,533,928,951]" gridcol="2" gridrow="12" pageId="12" pageNumber="13">103 ± 72</td>
<td id="D808176D413B002DFDA4FC5AFDB7FC52" box="[561,629,928,951]" gridcol="3" gridrow="12" pageId="12" pageNumber="13">38295</td>
<td id="D808176D413B002DFD31FC5AFD72FC52" box="[676,688,928,951]" gridcol="4" gridrow="12" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FC5AFB59FC52" box="[1156,1179,928,951]" gridcol="9" gridrow="12" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFC5AFA56FC52" box="[1406,1428,928,951]" gridcol="12" gridrow="12" pageId="12" pageNumber="13">0</td>
</tr>
<tr id="9BD97E11413B002DFF37FC3BF829FC3D" box="[162,2027,961,984]" gridrow="13" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FC3BFE9BFC3D" box="[162,345,961,984]" gridcol="0" gridrow="13" pageId="12" pageNumber="13">Peri-oral skin</th>
<td id="D808176D413B002DFE1CFC3BFE5DFC3D" box="[393,415,961,984]" gridcol="1" gridrow="13" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FC3BFD72FC3D" box="[676,688,961,984]" gridcol="4" gridrow="13" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FC3BFB59FC3D" box="[1156,1179,961,984]" gridcol="9" gridrow="13" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFC3BFA56FC3D" box="[1406,1428,961,984]" gridcol="12" gridrow="13" pageId="12" pageNumber="13">17</td>
<td id="D808176D413B002DFA2CFC3BF9C6FC3D" box="[1465,1540,961,984]" gridcol="13" gridrow="13" pageId="12" pageNumber="13">75 ± 9</td>
<td id="D808176D413B002DF9B6FC3BF9AAFC3D" box="[1571,1640,961,984]" gridcol="14" gridrow="13" pageId="12" pageNumber="13">6492</td>
<td id="D808176D413B002DF905FC3BF909FC3D" box="[1680,1739,961,984]" gridcol="15" gridrow="13" pageId="12" pageNumber="13">18 ± 5</td>
<td id="D808176D413B002DF978FC3BF8EDFC3D" box="[1773,1839,961,984]" gridcol="16" gridrow="13" pageId="12" pageNumber="13">1225</td>
</tr>
<tr id="9BD97E11413B002DFF37FC18F829FC1C" box="[162,2027,994,1017]" gridrow="14" pageId="12" pageNumber="13" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FC18FE9BFC1C" box="[162,345,994,1017]" gridcol="0" gridrow="14" pageId="12" pageNumber="13">Oesophagus skin</th>
<td id="D808176D413B002DFE1CFC18FE5DFC1C" box="[393,415,994,1017]" gridcol="1" gridrow="14" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FC18FD72FC1C" box="[676,688,994,1017]" gridcol="4" gridrow="14" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FC18FB59FC1C" box="[1156,1179,994,1017]" gridcol="9" gridrow="14" pageId="12" pageNumber="13">26</td>
<td id="D808176D413B002DFB2CFC18FB36FC1C" box="[1209,1268,994,1017]" gridcol="10" gridrow="14" pageId="12" pageNumber="13">27 ± 7</td>
<td id="D808176D413B002DFA87FC18FA8DFC1C" box="[1298,1359,994,1017]" gridcol="11" gridrow="14" pageId="12" pageNumber="13">1744</td>
<td id="D808176D413B002DFAEBFC18FA56FC1C" box="[1406,1428,994,1017]" gridcol="12" gridrow="14" pageId="12" pageNumber="13">0</td>
</tr>
<tr id="9BD97E11413B002DFF37FBF9F829FBFF" box="[162,2027,1027,1050]" gridrow="15" pageId="12" pageNumber="13" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FBF9FE9BFBFF" box="[162,345,1027,1050]" gridcol="0" gridrow="15" pageId="12" pageNumber="13">Introvert skin</th>
<td id="D808176D413B002DFE1CFBF9FE5DFBFF" box="[393,415,1027,1050]" gridcol="1" gridrow="15" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FBF9FD72FBFF" box="[676,688,1027,1050]" gridcol="4" gridrow="15" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FBF9FB59FBFF" box="[1156,1179,1027,1050]" gridcol="9" gridrow="15" pageId="12" pageNumber="13">13</td>
<td id="D808176D413B002DFB2CFBF9FB36FBFF" box="[1209,1268,1027,1050]" gridcol="10" gridrow="15" pageId="12" pageNumber="13">40 ± 16</td>
<td id="D808176D413B002DFA87FBF9FA8DFBFF" box="[1298,1359,1027,1050]" gridcol="11" gridrow="15" pageId="12" pageNumber="13">2283</td>
<td id="D808176D413B002DFAEBFBF9FA56FBFF" box="[1406,1428,1027,1050]" gridcol="12" gridrow="15" pageId="12" pageNumber="13">5</td>
<td id="D808176D413B002DFA2CFBF9F9C6FBFF" box="[1465,1540,1027,1050]" gridcol="13" gridrow="15" pageId="12" pageNumber="13">74 ± 18</td>
<td id="D808176D413B002DF9B6FBF9F9AAFBFF" box="[1571,1640,1027,1050]" gridcol="14" gridrow="15" pageId="12" pageNumber="13">4895</td>
<td id="D808176D413B002DF905FBF9F909FBFF" box="[1680,1739,1027,1050]" gridcol="15" gridrow="15" pageId="12" pageNumber="13">15 ± 5</td>
<td id="D808176D413B002DF978FBF9F8EDFBFF" box="[1773,1839,1027,1050]" gridcol="16" gridrow="15" pageId="12" pageNumber="13">1122</td>
</tr>
<tr id="9BD97E11413B002DFF37FBDFF829FBD9" box="[162,2027,1061,1084]" gridrow="16" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1">
<th id="D808176D413B002DFF37FBDFFE9BFBD9" box="[162,345,1061,1084]" gridcol="0" gridrow="16" pageId="12" pageNumber="13">Pedicels (ventral)</th>
<td id="D808176D413B002DFE1CFBDFFE5DFBD9" box="[393,415,1061,1084]" gridcol="1" gridrow="16" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FBDFFD72FBD9" box="[676,688,1061,1084]" gridcol="4" gridrow="16" pageId="12" pageNumber="13">3</td>
<td id="D808176D413B002DFD46FBDFFCD4FBD9" box="[723,790,1061,1084]" gridcol="5" gridrow="16" pageId="12" pageNumber="13">179 ± 12</td>
<td id="D808176D413B002DFCA9FBDFFCBCFBD9" box="[828,894,1061,1084]" gridcol="6" gridrow="16" pageId="12" pageNumber="13">166187</td>
<td id="D808176D413B002DFC24FBDFFC37FBD9" box="[945,1013,1061,1084]" gridcol="7" gridrow="16" pageId="12" pageNumber="13">161 ± 36</td>
<td id="D808176D413B002DFB8DFBDFFB98FBD9" box="[1048,1114,1061,1084]" gridcol="8" gridrow="16" pageId="12" pageNumber="13">127198</td>
<td id="D808176D413B002DFB11FBDFFB59FBD9" box="[1156,1179,1061,1084]" gridcol="9" gridrow="16" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFBDFFA56FBD9" box="[1406,1428,1061,1084]" gridcol="12" gridrow="16" pageId="12" pageNumber="13">19</td>
<td id="D808176D413B002DFA2CFBDFF9C6FBD9" box="[1465,1540,1061,1084]" gridcol="13" gridrow="16" pageId="12" pageNumber="13">90 ± 10</td>
<td id="D808176D413B002DF9B6FBDFF9AAFBD9" box="[1571,1640,1061,1084]" gridcol="14" gridrow="16" pageId="12" pageNumber="13">74103</td>
<td id="D808176D413B002DF8CDFBDFF852FBD9" box="[1880,1936,1061,1084]" gridcol="17" gridrow="16" pageId="12" pageNumber="13">34 ± 6</td>
<td id="D808176D413B002DF825FBDFF829FBD9" box="[1968,2027,1061,1084]" gridcol="18" gridrow="16" pageId="12" pageNumber="13">1941</td>
</tr>
<tr id="9BD97E11413B002DFF37FBBCF829FBB8" box="[162,2027,1094,1117]" gridrow="17" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FBBCFE9BFBB8" box="[162,345,1094,1117]" gridcol="0" gridrow="17" pageId="12" pageNumber="13">Bodywall (ventral)</th>
<td id="D808176D413B002DFE1CFBBCFE5DFBB8" box="[393,415,1094,1117]" gridcol="1" gridrow="17" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FBBCFD72FBB8" box="[676,688,1094,1117]" gridcol="4" gridrow="17" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FBBCFB59FBB8" box="[1156,1179,1094,1117]" gridcol="9" gridrow="17" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFBBCFA56FBB8" box="[1406,1428,1094,1117]" gridcol="12" gridrow="17" pageId="12" pageNumber="13">28</td>
<td id="D808176D413B002DFA2CFBBCF9C6FBB8" box="[1465,1540,1094,1117]" gridcol="13" gridrow="17" pageId="12" pageNumber="13">100 ± 11</td>
<td id="D808176D413B002DF9B6FBBCF9AAFBB8" box="[1571,1640,1094,1117]" gridcol="14" gridrow="17" pageId="12" pageNumber="13">81120</td>
<td id="D808176D413B002DF905FBBCF909FBB8" box="[1680,1739,1094,1117]" gridcol="15" gridrow="17" pageId="12" pageNumber="13">9 ± 2</td>
<td id="D808176D413B002DF978FBBCF8EDFBB8" box="[1773,1839,1094,1117]" gridcol="16" gridrow="17" pageId="12" pageNumber="13">514</td>
</tr>
<tr id="9BD97E11413B002DFF37FB92F829FB9A" box="[162,2027,1128,1151]" gridrow="18" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FB92FE9BFB9A" box="[162,345,1128,1151]" gridcol="0" gridrow="18" pageId="12" pageNumber="13">Anal papilla</th>
<td id="D808176D413B002DFE1CFB92FE5DFB9A" box="[393,415,1128,1151]" gridcol="1" gridrow="18" pageId="12" pageNumber="13">12</td>
<td id="D808176D413B002DFE28FB92FDD7FB9A" box="[445,533,1128,1151]" gridcol="2" gridrow="18" pageId="12" pageNumber="13">67 ± 6</td>
<td id="D808176D413B002DFDA4FB92FDB7FB9A" box="[561,629,1128,1151]" gridcol="3" gridrow="18" pageId="12" pageNumber="13">5977</td>
<td id="D808176D413B002DFD31FB92FD72FB9A" box="[676,688,1128,1151]" gridcol="4" gridrow="18" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FB92FB59FB9A" box="[1156,1179,1128,1151]" gridcol="9" gridrow="18" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFB92FA56FB9A" box="[1406,1428,1128,1151]" gridcol="12" gridrow="18" pageId="12" pageNumber="13">14</td>
<td id="D808176D413B002DFA2CFB92F9C6FB9A" box="[1465,1540,1128,1151]" gridcol="13" gridrow="18" pageId="12" pageNumber="13">55 ± 7</td>
<td id="D808176D413B002DF9B6FB92F9AAFB9A" box="[1571,1640,1128,1151]" gridcol="14" gridrow="18" pageId="12" pageNumber="13">4569</td>
<td id="D808176D413B002DF905FB92F909FB9A" box="[1680,1739,1128,1151]" gridcol="15" gridrow="18" pageId="12" pageNumber="13">11 ± 2</td>
<td id="D808176D413B002DF978FB92F8EDFB9A" box="[1773,1839,1128,1151]" gridcol="16" gridrow="18" pageId="12" pageNumber="13">816</td>
</tr>
<tr id="9BD97E11413B002DFF37FB73F829FB45" box="[162,2027,1161,1184]" gridrow="19" pageId="12" pageNumber="13">
<th id="D808176D413B002DFF37FB73F829FB45" box="[162,2027,1161,1184]" colspan="19" colspanRight="18" gridcol="0" gridrow="19" pageId="12" pageNumber="13">WMNH-INV-2014-184</th>
</tr>
<tr id="9BD97E11413B002DFF37FB50F829FB24" box="[162,2027,1194,1217]" gridrow="20" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FB50FE9BFB24" box="[162,345,1194,1217]" gridcol="0" gridrow="20" pageId="12" pageNumber="13">Tentacle</th>
<td id="D808176D413B002DFE1CFB50FE5DFB24" box="[393,415,1194,1217]" gridcol="1" gridrow="20" pageId="12" pageNumber="13">30</td>
<td id="D808176D413B002DFE28FB50FDD7FB24" box="[445,533,1194,1217]" gridcol="2" gridrow="20" pageId="12" pageNumber="13">108 ± 69</td>
<td id="D808176D413B002DFDA4FB50FDB7FB24" box="[561,629,1194,1217]" gridcol="3" gridrow="20" pageId="12" pageNumber="13">33268</td>
<td id="D808176D413B002DFD31FB50FD72FB24" box="[676,688,1194,1217]" gridcol="4" gridrow="20" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FB50FB59FB24" box="[1156,1179,1194,1217]" gridcol="9" gridrow="20" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFB50FA56FB24" box="[1406,1428,1194,1217]" gridcol="12" gridrow="20" pageId="12" pageNumber="13">0</td>
</tr>
<tr id="9BD97E11413B002DFF37FB31F829FB06" box="[162,2027,1227,1251]" gridrow="21" pageId="12" pageNumber="13" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FB31FE9BFB06" box="[162,345,1227,1251]" gridcol="0" gridrow="21" pageId="12" pageNumber="13">Peri-oral skin</th>
<td id="D808176D413B002DFE1CFB31FE5DFB06" box="[393,415,1227,1251]" gridcol="1" gridrow="21" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FB31FD72FB06" box="[676,688,1227,1251]" gridcol="4" gridrow="21" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FB31FB59FB06" box="[1156,1179,1227,1251]" gridcol="9" gridrow="21" pageId="12" pageNumber="13">27</td>
<td id="D808176D413B002DFB2CFB31FB36FB06" box="[1209,1268,1227,1251]" gridcol="10" gridrow="21" pageId="12" pageNumber="13">38 ±1 0</td>
<td id="D808176D413B002DFA87FB31FA8DFB06" box="[1298,1359,1227,1251]" gridcol="11" gridrow="21" pageId="12" pageNumber="13">2459</td>
<td id="D808176D413B002DFAEBFB31FA56FB06" box="[1406,1428,1227,1251]" gridcol="12" gridrow="21" pageId="12" pageNumber="13">8</td>
<td id="D808176D413B002DFA2CFB31F9C6FB06" box="[1465,1540,1227,1251]" gridcol="13" gridrow="21" pageId="12" pageNumber="13">89 ± 13</td>
<td id="D808176D413B002DF9B6FB31F9AAFB06" box="[1571,1640,1227,1251]" gridcol="14" gridrow="21" pageId="12" pageNumber="13">70104</td>
<td id="D808176D413B002DF905FB31F909FB06" box="[1680,1739,1227,1251]" gridcol="15" gridrow="21" pageId="12" pageNumber="13">15 ± 6</td>
<td id="D808176D413B002DF978FB31F8EDFB06" box="[1773,1839,1227,1251]" gridcol="16" gridrow="21" pageId="12" pageNumber="13">623</td>
</tr>
<tr id="9BD97E11413B002DFF37FB17F829FAE1" box="[162,2027,1261,1284]" gridrow="22" pageId="12" pageNumber="13" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FB17FE9BFAE1" box="[162,345,1261,1284]" gridcol="0" gridrow="22" pageId="12" pageNumber="13">Oesophagus skin</th>
<td id="D808176D413B002DFE1CFB17FE5DFAE1" box="[393,415,1261,1284]" gridcol="1" gridrow="22" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FB17FD72FAE1" box="[676,688,1261,1284]" gridcol="4" gridrow="22" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FB17FB59FAE1" box="[1156,1179,1261,1284]" gridcol="9" gridrow="22" pageId="12" pageNumber="13">26</td>
<td id="D808176D413B002DFB2CFB17FB36FAE1" box="[1209,1268,1261,1284]" gridcol="10" gridrow="22" pageId="12" pageNumber="13">36 ±10</td>
<td id="D808176D413B002DFA87FB17FA8DFAE1" box="[1298,1359,1261,1284]" gridcol="11" gridrow="22" pageId="12" pageNumber="13">1457</td>
<td id="D808176D413B002DFAEBFB17FA56FAE1" box="[1406,1428,1261,1284]" gridcol="12" gridrow="22" pageId="12" pageNumber="13">0</td>
</tr>
<tr id="9BD97E11413B002DFF37FAF4F829FAC0" box="[162,2027,1294,1317]" gridrow="23" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FAF4FE9BFAC0" box="[162,345,1294,1317]" gridcol="0" gridrow="23" pageId="12" pageNumber="13">Introvert skin</th>
<td id="D808176D413B002DFE1CFAF4FE5DFAC0" box="[393,415,1294,1317]" gridcol="1" gridrow="23" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FAF4FD72FAC0" box="[676,688,1294,1317]" gridcol="4" gridrow="23" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FAF4FB59FAC0" box="[1156,1179,1294,1317]" gridcol="9" gridrow="23" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFAF4FA56FAC0" box="[1406,1428,1294,1317]" gridcol="12" gridrow="23" pageId="12" pageNumber="13">16</td>
<td id="D808176D413B002DFA2CFAF4F9C6FAC0" box="[1465,1540,1294,1317]" gridcol="13" gridrow="23" pageId="12" pageNumber="13">92 ±13</td>
<td id="D808176D413B002DF9B6FAF4F9AAFAC0" box="[1571,1640,1294,1317]" gridcol="14" gridrow="23" pageId="12" pageNumber="13">71125</td>
<td id="D808176D413B002DF905FAF4F909FAC0" box="[1680,1739,1294,1317]" gridcol="15" gridrow="23" pageId="12" pageNumber="13">15 ± 4</td>
<td id="D808176D413B002DF978FAF4F8EDFAC0" box="[1773,1839,1294,1317]" gridcol="16" gridrow="23" pageId="12" pageNumber="13">1022</td>
</tr>
<tr id="9BD97E11413B002DFF37FAD5F829FAA3" box="[162,2027,1327,1350]" gridrow="24" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1">
<th id="D808176D413B002DFF37FAD5FE9BFAA3" box="[162,345,1327,1350]" gridcol="0" gridrow="24" pageId="12" pageNumber="13">Pedicels (ventral)</th>
<td id="D808176D413B002DFE1CFAD5FE5DFAA3" box="[393,415,1327,1350]" gridcol="1" gridrow="24" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FAD5FD72FAA3" box="[676,688,1327,1350]" gridcol="4" gridrow="24" pageId="12" pageNumber="13">7</td>
<td id="D808176D413B002DFD46FAD5FCD4FAA3" box="[723,790,1327,1350]" gridcol="5" gridrow="24" pageId="12" pageNumber="13">176 ± 6</td>
<td id="D808176D413B002DFCA9FAD5FCBCFAA3" box="[828,894,1327,1350]" gridcol="6" gridrow="24" pageId="12" pageNumber="13">164183</td>
<td id="D808176D413B002DFC24FAD5FC37FAA3" box="[945,1013,1327,1350]" gridcol="7" gridrow="24" pageId="12" pageNumber="13">119 ± 20</td>
<td id="D808176D413B002DFB8DFAD5FB98FAA3" box="[1048,1114,1327,1350]" gridcol="8" gridrow="24" pageId="12" pageNumber="13">85141</td>
<td id="D808176D413B002DFB11FAD5FB59FAA3" box="[1156,1179,1327,1350]" gridcol="9" gridrow="24" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFAD5FA56FAA3" box="[1406,1428,1327,1350]" gridcol="12" gridrow="24" pageId="12" pageNumber="13">23</td>
<td id="D808176D413B002DFA2CFAD5F9C6FAA3" box="[1465,1540,1327,1350]" gridcol="13" gridrow="24" pageId="12" pageNumber="13">99 ± 8</td>
<td id="D808176D413B002DF9B6FAD5F9AAFAA3" box="[1571,1640,1327,1350]" gridcol="14" gridrow="24" pageId="12" pageNumber="13">79112</td>
<td id="D808176D413B002DF8CDFAD5F852FAA3" box="[1880,1936,1327,1350]" gridcol="17" gridrow="24" pageId="12" pageNumber="13">32 ± 6</td>
<td id="D808176D413B002DF825FAD5F829FAA3" box="[1968,2027,1327,1350]" gridcol="18" gridrow="24" pageId="12" pageNumber="13">2245</td>
</tr>
<tr id="9BD97E11413B002DFF37FAABF829FA8D" box="[162,2027,1361,1384]" gridrow="25" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FAABFE9BFA8D" box="[162,345,1361,1384]" gridcol="0" gridrow="25" pageId="12" pageNumber="13">Bodywall (ventral)</th>
<td id="D808176D413B002DFE1CFAABFE5DFA8D" box="[393,415,1361,1384]" gridcol="1" gridrow="25" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FAABFD72FA8D" box="[676,688,1361,1384]" gridcol="4" gridrow="25" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FAABFB59FA8D" box="[1156,1179,1361,1384]" gridcol="9" gridrow="25" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFAABFA56FA8D" box="[1406,1428,1361,1384]" gridcol="12" gridrow="25" pageId="12" pageNumber="13">29</td>
<td id="D808176D413B002DFA2CFAABF9C6FA8D" box="[1465,1540,1361,1384]" gridcol="13" gridrow="25" pageId="12" pageNumber="13">112 ± 20</td>
<td id="D808176D413B002DF9B6FAABF9AAFA8D" box="[1571,1640,1361,1384]" gridcol="14" gridrow="25" pageId="12" pageNumber="13">81161</td>
<td id="D808176D413B002DF905FAABF909FA8D" box="[1680,1739,1361,1384]" gridcol="15" gridrow="25" pageId="12" pageNumber="13">9 ± 6</td>
<td id="D808176D413B002DF978FAABF8EDFA8D" box="[1773,1839,1361,1384]" gridcol="16" gridrow="25" pageId="12" pageNumber="13">431</td>
</tr>
<tr id="9BD97E11413B002DFF37FA88F829FA6C" box="[162,2027,1394,1417]" gridrow="26" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FA88FE9BFA6C" box="[162,345,1394,1417]" gridcol="0" gridrow="26" pageId="12" pageNumber="13">Anal papilla</th>
<td id="D808176D413B002DFE1CFA88FE5DFA6C" box="[393,415,1394,1417]" gridcol="1" gridrow="26" pageId="12" pageNumber="13">19</td>
<td id="D808176D413B002DFE28FA88FDD7FA6C" box="[445,533,1394,1417]" gridcol="2" gridrow="26" pageId="12" pageNumber="13">71 ± 15</td>
<td id="D808176D413B002DFDA4FA88FDB7FA6C" box="[561,629,1394,1417]" gridcol="3" gridrow="26" pageId="12" pageNumber="13">51107</td>
<td id="D808176D413B002DFD31FA88FD72FA6C" box="[676,688,1394,1417]" gridcol="4" gridrow="26" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FA88FB59FA6C" box="[1156,1179,1394,1417]" gridcol="9" gridrow="26" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFA88FA56FA6C" box="[1406,1428,1394,1417]" gridcol="12" gridrow="26" pageId="12" pageNumber="13">7</td>
<td id="D808176D413B002DFA2CFA88F9C6FA6C" box="[1465,1540,1394,1417]" gridcol="13" gridrow="26" pageId="12" pageNumber="13">76 ± 6</td>
<td id="D808176D413B002DF9B6FA88F9AAFA6C" box="[1571,1640,1394,1417]" gridcol="14" gridrow="26" pageId="12" pageNumber="13">6886</td>
<td id="D808176D413B002DF905FA88F909FA6C" box="[1680,1739,1394,1417]" gridcol="15" gridrow="26" pageId="12" pageNumber="13">6 ± 2</td>
<td id="D808176D413B002DF978FA88F8EDFA6C" box="[1773,1839,1394,1417]" gridcol="16" gridrow="26" pageId="12" pageNumber="13">48</td>
</tr>
<tr id="9BD97E11413B002DFF37FA69F829FA4F" box="[162,2027,1427,1450]" gridrow="27" pageId="12" pageNumber="13">
<th id="D808176D413B002DFF37FA69F829FA4F" box="[162,2027,1427,1450]" colspan="19" colspanRight="18" gridcol="0" gridrow="27" pageId="12" pageNumber="13">WMNH-INV-2014-188</th>
</tr>
<tr id="9BD97E11413B002DFF37FA4EF829FA2E" box="[162,2027,1460,1483]" gridrow="28" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FA4EFE9BFA2E" box="[162,345,1460,1483]" gridcol="0" gridrow="28" pageId="12" pageNumber="13">Tentacle</th>
<td id="D808176D413B002DFE1CFA4EFE5DFA2E" box="[393,415,1460,1483]" gridcol="1" gridrow="28" pageId="12" pageNumber="13">30</td>
<td id="D808176D413B002DFE28FA4EFDD7FA2E" box="[445,533,1460,1483]" gridcol="2" gridrow="28" pageId="12" pageNumber="13">111 ± 76</td>
<td id="D808176D413B002DFDA4FA4EFDB7FA2E" box="[561,629,1460,1483]" gridcol="3" gridrow="28" pageId="12" pageNumber="13">35321</td>
<td id="D808176D413B002DFD31FA4EFD72FA2E" box="[676,688,1460,1483]" gridcol="4" gridrow="28" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FA4EFB59FA2E" box="[1156,1179,1460,1483]" gridcol="9" gridrow="28" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFA4EFA56FA2E" box="[1406,1428,1460,1483]" gridcol="12" gridrow="28" pageId="12" pageNumber="13">0</td>
</tr>
<tr id="9BD97E11413B002DFF37FA2CF829FA08" box="[162,2027,1494,1517]" gridrow="29" pageId="12" pageNumber="13" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FA2CFE9BFA08" box="[162,345,1494,1517]" gridcol="0" gridrow="29" pageId="12" pageNumber="13">Peri-oral skin</th>
<td id="D808176D413B002DFE1CFA2CFE5DFA08" box="[393,415,1494,1517]" gridcol="1" gridrow="29" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FA2CFD72FA08" box="[676,688,1494,1517]" gridcol="4" gridrow="29" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FA2CFB59FA08" box="[1156,1179,1494,1517]" gridcol="9" gridrow="29" pageId="12" pageNumber="13">29</td>
<td id="D808176D413B002DFB2CFA2CFB36FA08" box="[1209,1268,1494,1517]" gridcol="10" gridrow="29" pageId="12" pageNumber="13">38 ± 11</td>
<td id="D808176D413B002DFA87FA2CFA8DFA08" box="[1298,1359,1494,1517]" gridcol="11" gridrow="29" pageId="12" pageNumber="13">2574</td>
<td id="D808176D413B002DFAEBFA2CFA56FA08" box="[1406,1428,1494,1517]" gridcol="12" gridrow="29" pageId="12" pageNumber="13">4</td>
<td id="D808176D413B002DFA2CFA2CF9C6FA08" box="[1465,1540,1494,1517]" gridcol="13" gridrow="29" pageId="12" pageNumber="13">86 ± 25</td>
<td id="D808176D413B002DF9B6FA2CF9AAFA08" box="[1571,1640,1494,1517]" gridcol="14" gridrow="29" pageId="12" pageNumber="13">65117</td>
<td id="D808176D413B002DF905FA2CF909FA08" box="[1680,1739,1494,1517]" gridcol="15" gridrow="29" pageId="12" pageNumber="13">21 ±1 0</td>
<td id="D808176D413B002DF978FA2CF8EDFA08" box="[1773,1839,1494,1517]" gridcol="16" gridrow="29" pageId="12" pageNumber="13">933</td>
</tr>
<tr id="9BD97E11413B002DFF37FA0DF829F9EB" box="[162,2027,1527,1550]" gridrow="30" pageId="12" pageNumber="13" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FA0DFE9BF9EB" box="[162,345,1527,1550]" gridcol="0" gridrow="30" pageId="12" pageNumber="13">Oesophagus skin</th>
<td id="D808176D413B002DFE1CFA0DFE5DF9EB" box="[393,415,1527,1550]" gridcol="1" gridrow="30" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FA0DFD72F9EB" box="[676,688,1527,1550]" gridcol="4" gridrow="30" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FA0DFB59F9EB" box="[1156,1179,1527,1550]" gridcol="9" gridrow="30" pageId="12" pageNumber="13">25</td>
<td id="D808176D413B002DFB2CFA0DFB36F9EB" box="[1209,1268,1527,1550]" gridcol="10" gridrow="30" pageId="12" pageNumber="13">34 ± 5</td>
<td id="D808176D413B002DFA87FA0DFA8DF9EB" box="[1298,1359,1527,1550]" gridcol="11" gridrow="30" pageId="12" pageNumber="13">2443</td>
<td id="D808176D413B002DFAEBFA0DFA56F9EB" box="[1406,1428,1527,1550]" gridcol="12" gridrow="30" pageId="12" pageNumber="13">0</td>
</tr>
<tr id="9BD97E11413B002DFF37F9E2F829F9CA" box="[162,2027,1560,1583]" gridrow="31" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37F9E2FE9BF9CA" box="[162,345,1560,1583]" gridcol="0" gridrow="31" pageId="12" pageNumber="13">Introvert skin</th>
<td id="D808176D413B002DFE1CF9E2FE5DF9CA" box="[393,415,1560,1583]" gridcol="1" gridrow="31" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31F9E2FD72F9CA" box="[676,688,1560,1583]" gridcol="4" gridrow="31" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11F9E2FB59F9CA" box="[1156,1179,1560,1583]" gridcol="9" gridrow="31" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBF9E2FA56F9CA" box="[1406,1428,1560,1583]" gridcol="12" gridrow="31" pageId="12" pageNumber="13">24</td>
<td id="D808176D413B002DFA2CF9E2F9C6F9CA" box="[1465,1540,1560,1583]" gridcol="13" gridrow="31" pageId="12" pageNumber="13">85 ± 14</td>
<td id="D808176D413B002DF9B6F9E2F9AAF9CA" box="[1571,1640,1560,1583]" gridcol="14" gridrow="31" pageId="12" pageNumber="13">56113</td>
<td id="D808176D413B002DF905F9E2F909F9CA" box="[1680,1739,1560,1583]" gridcol="15" gridrow="31" pageId="12" pageNumber="13">19 ± 8</td>
<td id="D808176D413B002DF978F9E2F8EDF9CA" box="[1773,1839,1560,1583]" gridcol="16" gridrow="31" pageId="12" pageNumber="13">1035</td>
</tr>
<tr id="9BD97E11413B002DFF37F9C3F829F9B4" box="[162,2027,1593,1617]" gridrow="32" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1">
<th id="D808176D413B002DFF37F9C3FE9BF9B4" box="[162,345,1593,1617]" gridcol="0" gridrow="32" pageId="12" pageNumber="13">Pedicels (ventral)</th>
<td id="D808176D413B002DFE1CF9C3FE5DF9B4" box="[393,415,1593,1617]" gridcol="1" gridrow="32" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31F9C3FD72F9B4" box="[676,688,1593,1617]" gridcol="4" gridrow="32" pageId="12" pageNumber="13">3</td>
<td id="D808176D413B002DFD46F9C3FCD4F9B4" box="[723,790,1593,1617]" gridcol="5" gridrow="32" pageId="12" pageNumber="13">170 ± 12</td>
<td id="D808176D413B002DFCA9F9C3FCBCF9B4" box="[828,894,1593,1617]" gridcol="6" gridrow="32" pageId="12" pageNumber="13">160184</td>
<td id="D808176D413B002DFC24F9C3FC37F9B4" box="[945,1013,1593,1617]" gridcol="7" gridrow="32" pageId="12" pageNumber="13">143 ± 42</td>
<td id="D808176D413B002DFB8DF9C3FB98F9B4" box="[1048,1114,1593,1617]" gridcol="8" gridrow="32" pageId="12" pageNumber="13">96176</td>
<td id="D808176D413B002DFB11F9C3FB59F9B4" box="[1156,1179,1593,1617]" gridcol="9" gridrow="32" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBF9C3FA56F9B4" box="[1406,1428,1593,1617]" gridcol="12" gridrow="32" pageId="12" pageNumber="13">15</td>
<td id="D808176D413B002DFA2CF9C3F9C6F9B4" box="[1465,1540,1593,1617]" gridcol="13" gridrow="32" pageId="12" pageNumber="13">91 ± 8</td>
<td id="D808176D413B002DF9B6F9C3F9AAF9B4" box="[1571,1640,1593,1617]" gridcol="14" gridrow="32" pageId="12" pageNumber="13">78106</td>
<td id="D808176D413B002DF8CDF9C3F852F9B4" box="[1880,1936,1593,1617]" gridcol="17" gridrow="32" pageId="12" pageNumber="13">38 ± 7</td>
<td id="D808176D413B002DF825F9C3F829F9B4" box="[1968,2027,1593,1617]" gridcol="18" gridrow="32" pageId="12" pageNumber="13">2651</td>
</tr>
<tr id="9BD97E11413B002DFF37F9A1F829F997" box="[162,2027,1627,1650]" gridrow="33" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37F9A1FE9BF997" box="[162,345,1627,1650]" gridcol="0" gridrow="33" pageId="12" pageNumber="13">Bodywall (ventral)</th>
<td id="D808176D413B002DFE1CF9A1FE5DF997" box="[393,415,1627,1650]" gridcol="1" gridrow="33" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31F9A1FD72F997" box="[676,688,1627,1650]" gridcol="4" gridrow="33" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11F9A1FB59F997" box="[1156,1179,1627,1650]" gridcol="9" gridrow="33" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBF9A1FA56F997" box="[1406,1428,1627,1650]" gridcol="12" gridrow="33" pageId="12" pageNumber="13">30</td>
<td id="D808176D413B002DFA2CF9A1F9C6F997" box="[1465,1540,1627,1650]" gridcol="13" gridrow="33" pageId="12" pageNumber="13">107 ± 12</td>
<td id="D808176D413B002DF9B6F9A1F9AAF997" box="[1571,1640,1627,1650]" gridcol="14" gridrow="33" pageId="12" pageNumber="13">88135</td>
<td id="D808176D413B002DF905F9A1F909F997" box="[1680,1739,1627,1650]" gridcol="15" gridrow="33" pageId="12" pageNumber="13">10 ± 2</td>
<td id="D808176D413B002DF978F9A1F8EDF997" box="[1773,1839,1627,1650]" gridcol="16" gridrow="33" pageId="12" pageNumber="13">717</td>
</tr>
<tr id="9BD97E11413B002DFF37F986F829F976" box="[162,2027,1660,1683]" gridrow="34" pageId="12" pageNumber="13" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413B002DFF37F986FE9BF976" box="[162,345,1660,1683]" gridcol="0" gridrow="34" pageId="12" pageNumber="13">Anal papilla</th>
<td id="D808176D413B002DFE28F986FDB7F976" box="[445,629,1660,1683]" colspan="2" colspanRight="1" gridcol="2" gridrow="34" pageId="12" pageNumber="13">Amputated</td>
</tr>
</table>
</paragraph>
<paragraph id="25567C53413BFFDEFF36F966FB81F950" blockId="12.[163,1091,1692,1717]" box="[163,1091,1692,1717]" pageId="12" pageNumber="13">
<tableNote id="D80F7DDD413BFFDEFF36F966FB81F950" box="[163,1091,1692,1717]" pageId="12" pageNumber="13" targetBox="[162,2027,525,1683]" targetPageId="12">
<superScript id="D29CD11B413BFFDEFF36F966FF6BF94F" attach="left" box="[163,169,1692,1706]" fontSize="5" pageId="12" pageNumber="13">a</superScript>
WMNH-INV: Invertebrate collection of the Wakayama Prefectural Museum of Natural History.
</tableNote>
</paragraph>
<subSubSection id="6DF32FD8413AFFDFFF52FF6CFD1FFA3E" pageId="13" pageNumber="14" type="description">
<paragraph id="25567C53413AFFDFFF52FF6CFBDEFE2A" blockId="13.[151,1437,150,2036]" pageId="13" pageNumber="14">
<emphasis id="179DA041413AFFDFFF52FF6CFE9BFF55" bold="true" box="[199,345,150,176]" pageId="13" pageNumber="14">Description.</emphasis>
Body medium, up to
<quantity id="E211D1B6413AFFDFFDC0FF6DFD64FF54" box="[597,678,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="13" pageNumber="14" unit="mm" value="39.0">39 mm</quantity>
long, fusiform, curved, with both ends tapered and turned slightly upwards (
<figureCitation id="BDD260D6413AFFDFFE99FF41FEA9FF30" box="[268,363,187,213]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="13" pageNumber="14">Fig. 2B</figureCitation>
); bodywall thin, soft. Color fading upon preservation. Living body color whitish orange. Tentacles 10, arranged in a single circle, two medioventral tentacles smaller than the other eight. Tentacles and introvert transparent in most parts in living animals, and with some dark brown pigmentation. Several rows of villi surrounding oral opening. Oral disk and pharynx of the preserved specimens dark grayish brown and dark purplish brown, respectively. Pedicels long, non-retractile, covering entire body at constant density, lacking in introvert region. Color of pedicels same and slightly deeper than adjacent body. Pedicels becoming gradually smaller toward anterior and posterior from middle portion of body. Ten anal papillae and five anal teeth in radii. After preservation, color denoted above turned more yellowish and whitish than that of living color.
</paragraph>
<paragraph id="25567C53413AFFDFFF52FE23FB42FD2C" blockId="13.[151,1437,150,2036]" pageId="13" pageNumber="14">
Calcareous ring long, cracking of mosaic patterns mostly caused by the damage of material; main radial plates and interradial plates not fragmented in original condition in the several well preserved specimens (
<figureCitation id="BDD260D6413AFFDFFB7BFE06FAFDFDF2" box="[1262,1343,508,535]" captionStart="FIGURE 3" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[152,1426,183,1471]" captionTargetId="figure-18@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 3. Elements of calcareous rings, viewed from the body cavity, anterior upper.A, Massinium toyoshiomaruae sp. nov., (holotype, WMNH-INV-2014-186); B, Thyone liaoi sp. nov., (holotype, WMNH-INV-2014-182, and a paratype, WMNH-INV- 2014-184); C, T. kyushuensis sp. nov., (holotype, WMNH-INV-2015-2437); D, T. toyoshiomaruae sp. nov. (holotype, WMNH- INV-2015-2483). Abbreviations: md, mid-dorsal part; mv, mid-ventral part; -ir, interradial element; -r radial element." figureDoi="http://doi.org/10.5281/zenodo.5225378" httpUri="https://zenodo.org/record/5225378/files/figure.png" pageId="13" pageNumber="14">Fig 3B</figureCitation>
). Radial elements elongated, with paired, posterior, weakly fragmented prolongations; each with an anterior projection and notch. Interradial elements also long but without posterior prolongations with slight anterior projections; shortest in dorso-lateral elements. Radial plates and interradial plates loosely adherent. One (or rarely two) Polian vesicles in medioventral, single stone canal in mediodorsal position. Well-matured gonad situated in mid-body, in two clumps, one on each side of dorsal mesentery, most tubules not branched. Gonad lacking ossicles.
</paragraph>
<paragraph id="25567C53413AFFDFFF52FD29FACFFCD1" blockId="13.[151,1437,150,2036]" pageId="13" pageNumber="14">
Bodywall ossicles comprising abundant tables (
<figureCitation id="BDD260D6413AFFDFFD4DFD29FC8EFD08" box="[728,844,723,750]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5F, G</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFCC0FD29FC6BFD08" box="[853,937,723,749]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
), with distorted, oval or rhomboidal disc and delicate spire, mostly with two pillars fusing into one distally, ending in 46 minute apical spines. Table disc with 816 perforations, most frequently 810; central two perforations large, peripheral perforations small.
</paragraph>
<paragraph id="25567C53413AFFDFFF52FCC4FF0BFAE0" blockId="13.[151,1437,150,2036]" pageId="13" pageNumber="14">
Tentacle ossicles comprising mostly plates of varying size and shape in trunk and branches, also table ossicles in basal parts (
<figureCitation id="BDD260D6413AFFDFFEABFC98FE5BFC98" box="[318,409,866,893]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5A</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFE30FC98FE3EFC99" box="[421,508,866,892]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
). Table ossicles with distorted oval or rhomboidal disc, and high spires with twopillars with mostly 23 crossbeams, and 48 apical spines. Some table ossicles have oval discs, possessing two larges central perforations and small distal perforations. Plates elongated, with two, large, central perforations and small distal perforations, occasionally developing a rudimentary spire and/or extra layer in central part. Peri-oral skin ossicles comprising mostly rosettes, with tables in part adjoining tentacle bases (
<figureCitation id="BDD260D6413AFFDFFBE5FC0BFB02FBE9" box="[1136,1216,1009,1036]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig.5B</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFB5BFC0BFAE5FBEE" box="[1230,1319,1009,1035]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
). Rosettes have granulated, mulberry-like, densely branched in more than one plane branches. Tables with distorted oval or rhomboidal discs, high spires with two-pillars, mostly with 13 crossbeams and 46 apical spines. Occasionally plates also exist, some with rudimentary spires and/or extra layer in central part. Pharyngeal villi have rosettes with granulated, mulberry-like (
<figureCitation id="BDD260D6413AFFDFFE5CFB7AFDE1FB7F" box="[457,547,1151,1178]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5C</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFDA4FB7AFD4BFB7F" box="[561,649,1152,1178]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
). Introvert with tables and mulberry-like rosettes (
<figureCitation id="BDD260D6413AFFDFFB5BFB7AFAEBFB7F" box="[1230,1321,1151,1178]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5D</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFAA2FB7AFA4DFB7F" box="[1335,1423,1152,1178]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
). Table ossicles with distorted, oval or rhomboidal disc, and low spires of two-pillars, with mostly 12 crossbeams, and 48 minute apical spines, occasionally with an extra layer in central part. Rosettes with granulated, mulberrylike.
</paragraph>
<paragraph id="25567C53413AFFDFFF52FAF5FD1FFA3E" blockId="13.[151,1437,150,2036]" pageId="13" pageNumber="14">
Most pedicels with endplate and supporting tables (
<figureCitation id="BDD260D6413AFFDFFC80FAF5FCAFFACC" box="[789,877,1294,1321]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5E</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFCEFFAF5FC10FACC" box="[890,978,1295,1321]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
). Supporting tables with narrow, curved disc, with 24 large, central perforations and 28 small, distal perforations; spire high, with two (occasionally three) pillars united distally, ending in 24 minute processes (
<figureCitation id="BDD260D6413AFFDFFC84FAACFCABFA94" box="[785,873,1366,1393]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5E</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFCE3FAACFC0CFA95" box="[886,974,1366,1392]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
). Endplates with holes arranged in three concentric areas,
<emphasis id="179DA041413AFFDFFECEFA81FE43FA71" box="[347,385,1403,1428]" italics="true" pageId="13" pageNumber="14">viz.</emphasis>
, commarginally-elongated holes near rim, radially-elongated holes in mid-region, and almost circular holes in central area. Anal papillae with small, irregular supporting plates and tables (
<figureCitation id="BDD260D6413AFFDFFB0AFA67FB35FA5D" box="[1183,1271,1437,1464]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5H</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFA94FA67FA94FA5D" box="[1281,1366,1437,1464]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
). Skin around anus with relatively small and simple tables.
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD8413AFFDCFF52FA1FFBC4FF1D" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="discussion">
<paragraph id="25567C53413AFFDCFF52FA1FFBC4FF1D" blockId="13.[151,1437,150,2036]" lastBlockId="14.[151,1437,150,535]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="179DA041413AFFDFFF52FA1FFEF9FA1A" bold="true" box="[199,315,1509,1535]" pageId="13" pageNumber="14">Remarks.</emphasis>
<taxonomicName id="E2E907D0413AFFDFFEDFFA1FFE1DFA1A" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[330,479,1509,1535]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="liaoi" status="sp. nov.">
<emphasis id="179DA041413AFFDFFEDFFA1FFE1DFA1A" box="[330,479,1509,1535]" italics="true" pageId="13" pageNumber="14">Thyone liaoi</emphasis>
</taxonomicName>
<emphasis id="179DA041413AFFDFFE7DFA1FFD85FA1A" bold="true" box="[488,583,1509,1535]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="0CAE1D3A413AFFDFFE7DFA1FFD85FA1A" box="[488,583,1509,1535]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was previously reported as a regional form of “
<taxonomicName id="E2E907D0413AFFDFFB10FA1FFF15F9C6" authority="Semper, 1867" authorityName="Semper" authorityYear="1867" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="pedata">
<emphasis id="179DA041413AFFDFFB10FA1FFAF6FA1A" box="[1157,1332,1509,1535]" italics="true" pageId="13" pageNumber="14">Thyone pedata</emphasis>
<bibRefCitation id="417801A2413AFFDFFAA8FA1FFF15F9C6" author="Semper, C." pageId="13" pageNumber="14" refId="ref27545" refString="Semper, C. (1867 - 1868) Holothurien. Reisen im Archipel der Philippinen. Teil 2. Wissenschaftliche Resultate. C. W. Kreidel, Wiesbaden, 288 pp. https: // doi. org / 10.5962 / bhl. title. 11687" type="book" year="1867">Semper, 1867</bibRefCitation>
</taxonomicName>
,” from the Chinese and Japanese waters (
<emphasis id="179DA041413AFFDFFD55F9F0FD28F9C6" box="[704,746,1546,1571]" italics="true" pageId="13" pageNumber="14">e.g.</emphasis>
,
<bibRefCitation id="417801A2413AFFDFFD6FF9F3FC19F9C1" author="Liao, Y. &amp; Clark, A. M." box="[762,987,1545,1572]" pageId="13" pageNumber="14" refId="ref26030" refString="Liao, Y. &amp; Clark, A. M. (1995) The Echinoderms of Southern China. Science Press Beijing, 614 pp." type="book" year="1995">Liao &amp; Clark 1995</bibRefCitation>
,
<bibRefCitation id="417801A2413AFFDFFC73F9F3FB4EF9C1" author="Yamana, Y." box="[998,1164,1545,1572]" pageId="13" pageNumber="14" pagination="6 - 8" refId="ref28119" refString="Yamana, Y. (2016) Kii-suido shuhen no namako bunnrui no genjou [The present situation of the studies on Holothurioidea in the Kii Strait], The Umiushi Tsushin (ISSN 1341 - 5972), published by the Research Institute of Marine Invertebrates, Tokyo, 92, 6 - 8. [only in Japanese]" type="journal article" year="2016">Yamana 2016</bibRefCitation>
), because the bodywall ossicle morphologies (reported in
<bibRefCitation id="417801A2413AFFDFFDA6F9D6FCEAF9A2" author="Clark, A. M. &amp; Rowe, F. W. E." box="[563,808,1580,1607]" pageId="13" pageNumber="14" refId="ref25186" refString="Clark, A. M. &amp; Rowe, F. W. E. (1971) Monograph of shallow-water Indo-West Pacific echinoderms. British Museum of Natural History, London, 238 pp." type="book" year="1971">Clark &amp; Rowe 1971</bibRefCitation>
;
<bibRefCitation id="417801A2413AFFDFFCA3F9D6FBDCF9A2" author="Liao, Y. &amp; Clark, A. M." box="[822,1054,1580,1607]" pageId="13" pageNumber="14" refId="ref26030" refString="Liao, Y. &amp; Clark, A. M. (1995) The Echinoderms of Southern China. Science Press Beijing, 614 pp." type="book" year="1995">Liao &amp; Clark 1995</bibRefCitation>
; and the present study) of these species resemble each other. However,
<bibRefCitation id="417801A2413AFFDFFDF7F9AAFCCAF98E" author="Semper, C." box="[610,776,1616,1643]" pageId="13" pageNumber="14" refId="ref27545" refString="Semper, C. (1867 - 1868) Holothurien. Reisen im Archipel der Philippinen. Teil 2. Wissenschaftliche Resultate. C. W. Kreidel, Wiesbaden, 288 pp. https: // doi. org / 10.5962 / bhl. title. 11687" type="book" year="1867">Semper (1867</bibRefCitation>
8) plainly stated that the gonadal tubules of this species have needle-shaped ossicles and that the bodywall possesses quite large ossicles. (These ossicle morphologies were not mentioned in
<bibRefCitation id="417801A2413AFFDFFE32F962FD86F957" author="Semper, C." box="[423,580,1688,1714]" pageId="13" pageNumber="14" refId="ref27545" refString="Semper, C. (1867 - 1868) Holothurien. Reisen im Archipel der Philippinen. Teil 2. Wissenschaftliche Resultate. C. W. Kreidel, Wiesbaden, 288 pp. https: // doi. org / 10.5962 / bhl. title. 11687" type="book" year="1867">Semper 1867</bibRefCitation>
8, p 67). The presence of such ossicles were not observed in the present new species. From seas worldwide, there are more than 60 nominal species and subspecies in the genus
<taxonomicName id="E2E907D0413AFFDFFAD3F941FA5EF930" authorityName="Oken" authorityYear="1815" box="[1350,1436,1723,1749]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="genus">
<emphasis id="179DA041413AFFDFFAD3F941FA5EF930" box="[1350,1436,1723,1749]" italics="true" pageId="13" pageNumber="14">Thyone</emphasis>
</taxonomicName>
(65 or 68 species/subspecies nominated in WoRMS 2021b), in which 14 species are presently classified into the Group III (comprising species with both rosettes and tables in the introvert) congeners which had been implied by
<bibRefCitation id="417801A2413AFFDFFEB4F8DDFDC6F8A4" author="Deichmann, E." box="[289,516,1831,1857]" pageId="13" pageNumber="14" pagination="43 - 236" refId="ref25222" refString="Deichmann, E. (1930) The Holothurians of the Western Part of the Atlantic Ocean. Bulletin of the Museum of Comparative Zooology at Harvard College, 71, 43 - 236." type="journal article" year="1930">Deichmann (1930)</bibRefCitation>
,
<bibRefCitation id="417801A2413AFFDFFD87F8DDFD09F8A4" author="Panning, A." box="[530,715,1831,1857]" pageId="13" pageNumber="14" pagination="404 - 470" refId="ref26870" refString="Panning, A. (1949) Versuch einer Neuordnung der Familie Cucumariidae (Holothurioidea, Dendrochirota). Zoologischen Jahrbuchern, 78, 404 - 470." type="journal article" year="1949">Panning (1949)</bibRefCitation>
and Pawson &amp; Millar (1981). The present new species also should be involved into this group as 15th species (Table 8). Among the 15 species of Group III, this species have the four identical characters: 1) granulated, mulberry-like small rosettes (
<figureCitation id="BDD260D6413AFFDFFBABF894FB7FF86C" box="[1086,1213,1902,1929]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5BD</figureCitation>
) are detected from the peri-oral, pharynx, and introvert skin; 2) length (approx. 70160 µm) of the bodywall table disc (
<figureCitation id="BDD260D6413AFFDFFAF8F868FF1EF835" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5F, G</figureCitation>
) is exceedingly larger than that of the other 14 species (Table 8-2); 3) length (approx. 30430 µm) of the tentacle ossicles (
<figureCitation id="BDD260D6413AFFDFFE29F823FDD8F811" box="[444,538,2009,2036]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5A</figureCitation>
) is exceedingly larger than that of the other 14 species (Table 8-2); 4) the two-pillared spire of the bodywall table is united one-pillar with 46 minute apical spines (
<figureCitation id="BDD260D64139FFDCFB74FF6DFAA2FF54" box="[1249,1376,150,177]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="14" pageNumber="15">Fig. 5F, G</figureCitation>
), but not forming a crown. From these features, this new species is easily distinguished from the 14 species of the Group III and other 24 species of undetectable group of
<taxonomicName id="E2E907D04139FFDCFCA1FF24FC48FF1D" authorityName="Oken" authorityYear="1815" box="[820,906,222,248]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="genus">
<emphasis id="179DA0414139FFDCFCA1FF24FC48FF1D" box="[820,906,222,248]" italics="true" pageId="14" pageNumber="15">Thyone</emphasis>
</taxonomicName>
(Table 8).
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD84139FFDCFF52FEF8FA52FE81" pageId="14" pageNumber="15" type="distribution">
<paragraph id="25567C534139FFDCFF52FEF8FA52FE81" blockId="14.[151,1437,150,535]" pageId="14" pageNumber="15">
<emphasis id="179DA0414139FFDCFF52FEF8FEA3FEF9" bold="true" box="[199,353,258,284]" pageId="14" pageNumber="15">Distribution.</emphasis>
Known from the Gulf of
<collectingCountry id="5DFE3CC34139FFDCFDE5FEF8FD7DFEF9" box="[624,703,258,284]" name="Vietnam" pageId="14" pageNumber="15">Tonkin</collectingCountry>
,
<collectingCountry id="5DFE3CC34139FFDCFD5CFEF8FCCEFEF9" box="[713,780,258,284]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
, depth
<quantity id="E211D1B64139FFDCFCC2FEF8FC4FFEF9" box="[855,909,258,285]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="14" pageNumber="15" unit="m" value="55.0">55 m</quantity>
(
<bibRefCitation id="417801A24139FFDCFC0FFEF8FBABFEF8" author="Liao, Y. &amp; Clark, A. M." box="[922,1129,258,285]" pageId="14" pageNumber="15" refId="ref26030" refString="Liao, Y. &amp; Clark, A. M. (1995) The Echinoderms of Southern China. Science Press Beijing, 614 pp." type="book" year="1995">Liao &amp; Clark 1995</bibRefCitation>
), and several localities in the Seto Inland Sea,
<collectingCountry id="5DFE3CC34139FFDCFEDAFEDCFE53FEA5" box="[335,401,294,320]" name="Japan" pageId="14" pageNumber="15">Japan</collectingCountry>
: middle and eastern parts of the Seto Inland Sea, sand bottom, depth
<quantity id="E211D1B64139FFDCFBEAFEDFFB74FEA5" box="[1151,1206,293,320]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="14" pageNumber="15" unit="m" value="26.0">26 m</quantity>
(ST-
<quantity id="E211D1B64139FFDCFB78FEDCFAD4FEDA" box="[1261,1302,293,320]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="14" pageNumber="15" unit="in" value="4.0">4 in</quantity>
2014); mud bottom, depth
<quantity id="E211D1B64139FFDCFEA1FEB3FE51FE81" box="[308,403,329,356]" metricMagnitude="1" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="14" pageNumber="15" unit="m" value="26.0" valueMax="27.0" valueMin="25.0">2527 m</quantity>
(ST-
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2014), and northen part of the Kii Strait, sand bottom, shallow water (
<bibRefCitation id="417801A24139FFDCFB7CFEB3FA46FE81" author="Yamana, Y." box="[1257,1412,329,356]" pageId="14" pageNumber="15" pagination="6 - 8" refId="ref28119" refString="Yamana, Y. (2016) Kii-suido shuhen no namako bunnrui no genjou [The present situation of the studies on Holothurioidea in the Kii Strait], The Umiushi Tsushin (ISSN 1341 - 5972), published by the Research Institute of Marine Invertebrates, Tokyo, 92, 6 - 8. [only in Japanese]" type="journal article" year="2016">Yamana 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD84139FFDCFF52FE97FBEFFDF2" pageId="14" pageNumber="15" type="etymology">
<paragraph id="25567C534139FFDCFF52FE97FB58FE4E" blockId="14.[151,1437,150,535]" pageId="14" pageNumber="15">
<emphasis id="179DA0414139FFDCFF52FE97FE8CFE62" bold="true" box="[199,334,365,391]" pageId="14" pageNumber="15">Etymology.</emphasis>
The species named in honor of the famous researcher Dr. Yulin Liao of the Institute of Oceanology, Academia Sinica (IOAS) Qingdao,
<collectingCountry id="5DFE3CC34139FFDCFDB3FE6BFDAEFE4E" box="[550,620,401,427]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
, who first reported this species as
<emphasis id="179DA0414139FFDCFC79FE6BFB58FE4E" box="[1004,1178,401,427]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="E2E907D04139FFDCFC79FE6BFB54FE4E" authorityName="Semper" authorityYear="1867" box="[1004,1174,401,427]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="pedata">Thyone pedata</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="25567C534139FFDCFF52FE4FFBEFFDF2" blockId="14.[151,1437,150,535]" pageId="14" pageNumber="15">
<emphasis id="179DA0414139FFDCFF52FE4FFDD5FE2B" bold="true" box="[199,535,436,463]" pageId="14" pageNumber="15">Molecular data comparison.</emphasis>
In BLAST searches, the obtained partial COI gene sequence is deeply divergent from all other sequences. A sequence identified as
<taxonomicName id="E2E907D04139FFDCFD48FE22FC5BFE16" box="[733,921,472,499]" class="Holothuroidea" family="Synallactidae" genus="Paelopatides" kingdom="Animalia" order="Synallactida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="179DA0414139FFDCFD48FE22FCADFE17" box="[733,879,472,498]" italics="true" pageId="14" pageNumber="15">Paelopatides</emphasis>
sp.
</taxonomicName>
(INSD accession number
<accessionNumber id="3ABAE1B04139FFDCFB5DFE22FA85FE16" box="[1224,1351,472,499]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX874356" pageId="14" pageNumber="15">KX874356</accessionNumber>
, 80.6% similarity with 100% coverage) was closest (meaningless at generic/family level).
</paragraph>
</subSubSection>
</treatment>
</document>