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<document id="89C621E1A87A610DEFBB1B430E24DC1F" ID-DOI="10.11646/zootaxa.5514.5.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="13914787" ID-ZooBank="36CA84B3-FDA3-4F68-92DA-9294C2E92839" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1728581506313" checkinUser="plazi" docAuthor="Wang, Jian, Lin, Shi-Shi, Gan, Jun-Shen, Chen, Hong-Hui, Yu, Ling-Min, Pan, Zhong, Xiao, Jia-Jie & Zeng, Zhao-Chi" docDate="2024" docId="A43187CDFFA8FFCEFF356D53542193D3" docLanguage="en" docName="zootaxa.5514.5.3.pdf" docOrigin="Zootaxa 5514 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5514.5.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Boulenophrys pepe Wang and Zeng 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="463" masterDocId="5808FFB5FFA0FFC2FFA26A14511C9402" masterDocTitle="A new species of the genus Boulenophrys from South China (Anura, Megophryidae)" masterLastPageNumber="468" masterPageNumber="451" pageNumber="459" updateTime="1728664180886" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="FE3EF53DAD6A8317F92DABF0272586A8">A new species of the genus Boulenophrys from South China (Anura, Megophryidae)</mods:title>
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<mods:namePart id="CA1D0E532CE3E72FCC60CDB7B31DDE2F">Wang, Jian</mods:namePart>
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<mods:affiliation id="6313EFC18F32973F092C0329AD12A6B4">Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
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<mods:nameIdentifier id="0EE37C6DE076C898BF7685A6E480B32A" type="email">wangj-1994@outlook.com</mods:nameIdentifier>
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<mods:namePart id="E2BEB18D94DA5ACF7D9F034E8399C8F7">Lin, Shi-Shi</mods:namePart>
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<mods:affiliation id="F8A4D7FA9DFB70139B037DB1A3E5B639">Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
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<mods:namePart id="0D57BFF9C653BF0291DE79BCA6589069">Gan, Jun-Shen</mods:namePart>
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<mods:affiliation id="C1606DE2B512E27DB036D2FA096F1BA5">Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
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<mods:namePart id="0500ED0D832F5FC5E22ADF76B53D6F10">Chen, Hong-Hui</mods:namePart>
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<mods:namePart id="5D1352BA75FD9AA5C385F99084DE48AA">Yu, Ling-Min</mods:namePart>
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<mods:affiliation id="A83B22810276082BD8716C1A39C189A9">Guangdong Lianshan Bijiashan Provincial Nature Reserve, Qingyuan 513200, China</mods:affiliation>
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<mods:namePart id="45729FC747A5887D423D65B356563130">Pan, Zhong</mods:namePart>
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<mods:namePart id="7150E22948FC251E2C49F0B491092BB5">Xiao, Jia-Jie</mods:namePart>
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<mods:affiliation id="EA167DEE0A9B00D9BB1ADFABDFFCAB51">Guangdong Wildlife Monitoring Rescue and Conservation Center, Guangzhou 510520, China</mods:affiliation>
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<mods:namePart id="C3F3B6DF658C70FBFFAA907952AB4D47">Zeng, Zhao-Chi</mods:namePart>
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<mods:nameIdentifier id="29678C7574BC7EC54207D5BD7D9DF258" type="email">zhaochizeng@outlook.com</mods:nameIdentifier>
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<mods:title id="BEE7EDFCC6E25B503C4AFA9F67A09225">Zootaxa</mods:title>
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<mods:date id="86B2E91758C9B30D77CCD6DA9448030C">2024</mods:date>
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<mods:number id="AB685A421554D78674E84C382DDD6540">2024-10-01</mods:number>
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<mods:classification id="8D814C5950961F5D48EB1F742D96F41A">journal article</mods:classification>
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<treatment id="A43187CDFFA8FFCEFF356D53542193D3" LSID="urn:lsid:plazi:treatment:A43187CDFFA8FFCEFF356D53542193D3" httpUri="http://treatment.plazi.org/id/A43187CDFFA8FFCEFF356D53542193D3" lastPageId="12" lastPageNumber="463" pageId="8" pageNumber="459">
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<subSubSection id="64826550FFA8FFCAFF356D5353AE9360" box="[151,690,1863,1890]" pageId="8" pageNumber="459" type="nomenclature">
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<paragraph id="2C2736DBFFA8FFCAFF356D5353AE9360" blockId="8.[151,1017,1863,1962]" box="[151,690,1863,1890]" pageId="8" pageNumber="459">
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<heading id="776F81B7FFA8FFCAFF356D5353AE9360" bold="true" box="[151,690,1863,1890]" fontSize="11" level="1" pageId="8" pageNumber="459" reason="1">
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<emphasis id="1EECEAC9FFA8FFCAFF356D5353AE9360" bold="true" box="[151,690,1863,1890]" pageId="8" pageNumber="459">
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<taxonomicName id="EB984D58FFA8FFCAFF356D5353529360" authority="Wang and Zeng" authorityName="Wang and Zeng" authorityYear="2024" box="[151,590,1863,1890]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="459" phylum="Chordata" rank="species" species="pepe" status="sp. nov.">
|
||||
<emphasis id="1EECEAC9FFA8FFCAFF356D5350639360" bold="true" box="[151,383,1863,1890]" italics="true" pageId="8" pageNumber="459">Boulenophrys pepe</emphasis>
|
||||
Wang and Zeng
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="05DF57B2FFA8FFCAFDF66D5C53AE9360" box="[596,690,1864,1890]" pageId="8" pageNumber="459" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="64826550FFA8FFCAFF356D7852E59387" box="[151,1017,1899,1926]" pageId="8" pageNumber="459" type="vernacular_names">
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<paragraph id="2C2736DBFFA8FFCAFF356D7852E59387" blockId="8.[151,1017,1863,1962]" box="[151,1017,1899,1926]" pageId="8" pageNumber="459">
|
||||
Pepe Horned Toad (in English) / Lián Shān Jiǎo Chán (
|
||||
<emphasis id="1EECEAC9FFA8FFCAFCA36D78526D9384" bold="true" box="[769,881,1900,1926]" pageId="8" pageNumber="459">ḔƜmDz</emphasis>
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in Chinese)
|
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</paragraph>
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</subSubSection>
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<subSubSection id="64826550FFA8FFCAFF356D9B500093A8" box="[151,284,1935,1962]" pageId="8" pageNumber="459" type="description">
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<paragraph id="2C2736DBFFA8FFCAFF356D9B500093A8" blockId="8.[151,1017,1863,1962]" box="[151,284,1935,1962]" pageId="8" pageNumber="459">
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<figureCitation id="B4A32A5EFFA8FFCAFF356D9B500093A8" box="[151,284,1935,1962]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="10.[152,255,1647,1671]" captionStartId-1="11.[152,255,1904,1928]" captionTargetBox-0="[158,1429,188,1617]" captionTargetBox-1="[157,1429,279,1874]" captionTargetId-0="figure-27@10.[151,1436,181,1623]" captionTargetId-1="figure-62@11.[151,1435,273,1880]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 3. The holotype (GEP a207) of Boulenophrys pepe sp. nov. in life: A. dorsal view; B. dorsolateral view; C. ventral view; D. ventral view of head, arrows showing swollen lips and protrude jaws; E. ventral view of hand; F. ventral view of foot." captionText-1="FIGURE 4. The paratypes SYS a009350 (A–D) and GEP a209 (E–F) of Boulenophrys pepe sp. nov. in life: A. dorsal view; B. dorsolateral view; C. ventral view; D. lateral view of head, with an arrow showing swollen lips and protrude jaws; E. dorsolateral view; F. ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.13914793" figureDoi-1="http://doi.org/10.5281/zenodo.13914795" httpUri-0="https://zenodo.org/record/13914793/files/figure.png" httpUri-1="https://zenodo.org/record/13914795/files/figure.png" pageId="8" pageNumber="459">Figures 3–4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="64826550FFA8FFCBFF356DCC5418958B" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459" type="materials_examined">
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<paragraph id="2C2736DBFFA8FFCBFF356DCC555894D7" blockId="8.[151,1436,2007,2034]" lastBlockId="9.[151,1437,150,1977]" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459">
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<materialsCitation id="9CF03C86FFA8FFCBFF356DCC555894D7" collectingDate="2024-01-17" collectionCode="GEP" collectorName="Jian Wang & Zhao-Chi Zeng & Hong-Hui Chen & Shi-Shi Lin" country="China" county="Yao Autonomous County" elevation="540" lastPageId="9" lastPageNumber="460" latitude="24.216343" location="Lianshan Bijiashan Nature Reserve" longLatPrecision="1" longitude="111.995255" municipality="Qingyuan City" pageId="8" pageNumber="459" specimenCode="GEP a207" specimenCount="1" stateProvince="Guangdong" typeStatus="holotype">
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<emphasis id="1EECEAC9FFA8FFCAFF356DCC501793F0" bold="true" box="[151,267,2008,2034]" pageId="8" pageNumber="459">
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<typeStatus id="F3238879FFA8FFCAFF356DCC501A93F0" box="[151,262,2008,2034]" pageId="8" pageNumber="459" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
|
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<specimenCode id="7C3E9EA0FFA8FFCAFEB06DCC509893F0" box="[274,388,2008,2034]" pageId="8" pageNumber="459">GEP a207</specimenCode>
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(
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<figureCitation id="B4A32A5EFFA8FFCAFE316DC350E893F0" box="[403,500,2007,2034]" captionStart="FIGURE 3" captionStartId="10.[152,255,1647,1671]" captionTargetBox="[158,1429,188,1617]" captionTargetId="figure-27@10.[151,1436,181,1623]" captionTargetPageId="10" captionText="FIGURE 3. The holotype (GEP a207) of Boulenophrys pepe sp. nov. in life: A. dorsal view; B. dorsolateral view; C. ventral view; D. ventral view of head, arrows showing swollen lips and protrude jaws; E. ventral view of hand; F. ventral view of foot." figureDoi="http://doi.org/10.5281/zenodo.13914793" httpUri="https://zenodo.org/record/13914793/files/figure.png" pageId="8" pageNumber="459">Figure 3</figureCitation>
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), adult
|
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<specimenCount id="3A9EFD52FFA8FFCAFDE96DC3539893F3" box="[587,644,2007,2033]" pageId="8" pageNumber="459" type="male">male</specimenCount>
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, collected by
|
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<collectorName id="816D530DFFA8FFCAFCBD6DC3528B93F0" box="[799,919,2007,2034]" pageId="8" pageNumber="459">Jian Wang</collectorName>
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,
|
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<collectorName id="816D530DFFA8FFCAFC006DC3554D93F0" box="[930,1105,2007,2034]" pageId="8" pageNumber="459">Zhao-Chi Zeng</collectorName>
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,
|
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<collectorName id="816D530DFFA8FFCAFBFF6DC3540D93F3" box="[1117,1297,2007,2033]" pageId="8" pageNumber="459">Hong-Hui Chen</collectorName>
|
||||
and
|
||||
<collectorName id="816D530DFFA8FFCBFAEA6DC351A294B3" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459">Shi-Shi Lin</collectorName>
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on
|
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<collectingDate id="4862E9F3FFA9FFCBFF486A8350B994B3" box="[234,421,151,177]" pageId="9" pageNumber="460" value="2024-01-17">17 January 2024</collectingDate>
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</date>
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from
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(
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,
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; ca.
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||||
<quantity id="EB609B3EFFA9FFCBFAAA6A82544C94B3" box="[1288,1360,150,177]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="460" unit="m" value="540.0">540 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
),
|
||||
<location id="29476000FFA9FFCBFF356AAF504794D7" LSID="urn:lsid:plazi:treatment:A43187CDFFA8FFCEFF356D53542193D3:29476000FFA9FFCBFF356AAF504794D7" box="[151,347,187,213]" country="China" county="Yao Autonomous County" latitude="24.216343" longLatPrecision="1" longitude="111.995255" municipality="Qingyuan City" name="Lianshan Zhuang" pageId="9" pageNumber="460" stateProvince="Guangdong">Lianshan Zhuang</location>
|
||||
and
|
||||
<collectingCounty id="C5464E57FFA9FFCBFE326AAF53B194D7" box="[400,685,187,213]" pageId="9" pageNumber="460">Yao Autonomous County</collectingCounty>
|
||||
,
|
||||
<collectingMunicipality id="CC43ACA1FFA9FFCBFD156AAF524294D7" box="[695,862,187,213]" pageId="9" pageNumber="460">Qingyuan City</collectingMunicipality>
|
||||
,
|
||||
<collectingRegion id="EE5CF839FFA9FFCBFCCB6AAF52F394D7" box="[873,1007,187,213]" country="China" name="Guangdong" pageId="9" pageNumber="460">Guangdong</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="548F764BFFA9FFCBFC586AAF555D94D7" box="[1018,1089,187,213]" name="China" pageId="9" pageNumber="460">China</collectingCountry>
|
||||
.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="2C2736DBFFA9FFCBFF656ACB5418958B" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="460">
|
||||
<materialsCitation id="9CF03C86FFA9FFCBFF656ACB5502951F" collectingDate="2023-03-18" collectionCode="GEP" collectorName="Jun-Shen Gan & Hu-Jun Pan" elevation="510" latitude="24.222097" location="Lianshan Bijiashan Nature Reserve" longLatPrecision="1" longitude="111.99185" pageId="9" pageNumber="460" specimenCode="GEP a117" specimenCount="1" specimenCount-adult="3" typeStatus="paratype">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFF656ACB505A94FB" bold="true" box="[199,326,223,249]" pageId="9" pageNumber="460">
|
||||
<typeStatus id="F3238879FFA9FFCBFF656ACB505D94FB" box="[199,321,223,249]" pageId="9" pageNumber="460" type="paratype">Paratypes</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
<specimenCount id="3A9EFD52FFA9FFCBFEE86ACB531094FB" box="[330,524,223,249]" pageId="9" pageNumber="460">Three adult males</specimenCount>
|
||||
from
|
||||
<location id="29476000FFA9FFCBFDE96ACB52CE94FB" LSID="urn:lsid:plazi:treatment:A43187CDFFA8FFCEFF356D53542193D3:29476000FFA9FFCBFDE96ACB52CE94FB" box="[587,978,223,249]" latitude="24.222097" longLatPrecision="1" longitude="111.99185" name="Lianshan Bijiashan Nature Reserve" pageId="9" pageNumber="460">Lianshan Bijiashan Nature Reserve</location>
|
||||
.
|
||||
<specimenCode id="7C3E9EA0FFA9FFCBFC7B6ACB555A94FB" box="[985,1094,223,249]" pageId="9" pageNumber="460">GEP a117</specimenCode>
|
||||
(
|
||||
<geoCoordinate id="49AC501CFFA9FFCBFBF06ACB55F194FB" box="[1106,1261,223,249]" degrees="24.222098" direction="north" orientation="latitude" pageId="9" pageNumber="460" precision="1" value="24.222097">24.222098°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="49AC501CFFA9FFCBFB576ACA548494FB" box="[1269,1432,222,249]" degrees="111.991852" direction="east" orientation="longitude" pageId="9" pageNumber="460" precision="1" value="111.99185">111.991852°E</geoCoordinate>
|
||||
; ca.
|
||||
<elevation id="A7B5D1E8FFA9FFCBFF1C6B16505D951F" box="[190,321,258,285]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="9" pageNumber="460" unit="m" value="510.0">
|
||||
<quantity id="EB609B3EFFA9FFCBFF1C6B16501A951F" box="[190,262,258,285]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="9" pageNumber="460" unit="m" value="510.0">510 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
), collected by
|
||||
<collectorName id="816D530DFFA9FFCBFE4A6B17539A951F" box="[488,646,259,285]" pageId="9" pageNumber="460">Jun-Shen Gan</collectorName>
|
||||
and
|
||||
<collectorName id="816D530DFFA9FFCBFD1F6B17525C951F" box="[701,832,259,285]" pageId="9" pageNumber="460">Hu-Jun Pan</collectorName>
|
||||
on
|
||||
<date id="5826101BFFA9FFCBFCC96B175505951F" box="[875,1049,259,285]" pageId="9" pageNumber="460" value="2023-03-18">
|
||||
<collectingDate id="4862E9F3FFA9FFCBFCC96B175505951F" box="[875,1049,259,285]" pageId="9" pageNumber="460" value="2023-03-18">18 March 2023</collectingDate>
|
||||
</date>
|
||||
;
|
||||
</materialsCitation>
|
||||
<materialsCitation id="9CF03C86FFA9FFCBFB876B1753AB9567" collectingDate="2024-01-17" collectionCode="SYS" collectorName="Jian Wang & Zhao-Chi Zeng & Hong-Hui Chen & Shi-Shi Lin" elevation="520" latitude="24.222097" location="Lianshan Bijiashan Nature Reserve" longLatPrecision="1" longitude="111.99185" pageId="9" pageNumber="460" specimenCode="SYS a009350" specimenCount="1" specimenCount-adult="3" typeStatus="paratype">
|
||||
<specimenCode id="7C3E9EA0FFA9FFCBFB876B1755DD951F" box="[1061,1217,258,285]" pageId="9" pageNumber="460">SYS a009350</specimenCode>
|
||||
(field number GEP a208;
|
||||
<figureCitation id="B4A32A5EFFA9FFCBFF7F6B33504B9543" box="[221,343,295,321]" captionStart="FIGURE 4" captionStartId="11.[152,255,1904,1928]" captionTargetBox="[157,1429,279,1874]" captionTargetId="figure-62@11.[151,1435,273,1880]" captionTargetPageId="11" captionText="FIGURE 4. The paratypes SYS a009350 (A–D) and GEP a209 (E–F) of Boulenophrys pepe sp. nov. in life: A. dorsal view; B. dorsolateral view; C. ventral view; D. lateral view of head, with an arrow showing swollen lips and protrude jaws; E. dorsolateral view; F. ventral view." figureDoi="http://doi.org/10.5281/zenodo.13914795" httpUri="https://zenodo.org/record/13914795/files/figure.png" pageId="9" pageNumber="460">Fig. 4A–D</figureCitation>
|
||||
) (
|
||||
<geoCoordinate id="49AC501CFFA9FFCBFECC6B3253159543" box="[366,521,294,321]" degrees="24.219745" direction="north" orientation="latitude" pageId="9" pageNumber="460" precision="1" value="24.219746">24.219745°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="49AC501CFFA9FFCBFDB76B3353AB9543" box="[533,695,295,321]" degrees="111.997946" direction="east" orientation="longitude" pageId="9" pageNumber="460" precision="1" value="111.99795">111.997946°E</geoCoordinate>
|
||||
; ca.
|
||||
<quantity id="EB609B3EFFA9FFCBFD496B32522E9543" box="[747,818,294,321]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="9" pageNumber="460" unit="m" value="520.0">
|
||||
<elevation id="A7B5D1E8FFA9FFCBFD496B32522E9543" box="[747,818,294,321]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="9" pageNumber="460" unit="m" value="520.0">520 m</elevation>
|
||||
</quantity>
|
||||
a.s.l.), collected by
|
||||
<collectorName id="816D530DFFA9FFCBFBB46B3355929543" box="[1046,1166,295,321]" pageId="9" pageNumber="460">Jian Wang</collectorName>
|
||||
,
|
||||
<collectorName id="816D530DFFA9FFCBFB386B3354559543" box="[1178,1353,295,321]" pageId="9" pageNumber="460">Zhao-Chi Zeng</collectorName>
|
||||
,
|
||||
<collectorName id="816D530DFFA9FFCBFAF76B3350199567" pageId="9" pageNumber="460">Hong-Hui Chen</collectorName>
|
||||
and
|
||||
<collectorName id="816D530DFFA9FFCBFEE26B5F50D89567" box="[320,452,331,357]" pageId="9" pageNumber="460">Shi-Shi Lin</collectorName>
|
||||
on
|
||||
<date id="5826101BFFA9FFCBFE506B5F53AF9567" box="[498,691,331,357]" pageId="9" pageNumber="460" value="2024-01-17">
|
||||
<collectingDate id="4862E9F3FFA9FFCBFE506B5F53AF9567" box="[498,691,331,357]" pageId="9" pageNumber="460" value="2024-01-17">17 January 2024</collectingDate>
|
||||
</date>
|
||||
;
|
||||
</materialsCitation>
|
||||
<materialsCitation id="9CF03C86FFA9FFCBFD626B5F541D958B" collectingDate="2024-01-17" collectionCode="GEP" collectorName="Jian Wang & Zhao-Chi Zeng & Hong-Hui Chen & Shi-Shi Lin" elevation="490" latitude="24.222097" location="Lianshan Bijiashan Nature Reserve" longLatPrecision="1" longitude="111.99185" pageId="9" pageNumber="460" specimenCode="GEP a209" specimenCount="1" specimenCount-adult="3" typeStatus="paratype">
|
||||
<specimenCode id="7C3E9EA0FFA9FFCBFD626B5F522F9567" box="[704,819,331,357]" pageId="9" pageNumber="460">GEP a209</specimenCode>
|
||||
(
|
||||
<figureCitation id="B4A32A5EFFA9FFCBFCE76B5F52A69567" box="[837,954,331,357]" captionStart="FIGURE 4" captionStartId="11.[152,255,1904,1928]" captionTargetBox="[157,1429,279,1874]" captionTargetId="figure-62@11.[151,1435,273,1880]" captionTargetPageId="11" captionText="FIGURE 4. The paratypes SYS a009350 (A–D) and GEP a209 (E–F) of Boulenophrys pepe sp. nov. in life: A. dorsal view; B. dorsolateral view; C. ventral view; D. lateral view of head, with an arrow showing swollen lips and protrude jaws; E. dorsolateral view; F. ventral view." figureDoi="http://doi.org/10.5281/zenodo.13914795" httpUri="https://zenodo.org/record/13914795/files/figure.png" pageId="9" pageNumber="460">Fig. 4E–F</figureCitation>
|
||||
) (
|
||||
<geoCoordinate id="49AC501CFFA9FFCBFC706B5E55719567" box="[978,1133,330,357]" degrees="24.219954" direction="north" orientation="latitude" pageId="9" pageNumber="460" precision="1" value="24.219954">24.219954°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="49AC501CFFA9FFCBFBD86B5F54029567" box="[1146,1310,331,357]" degrees="111.997601" direction="east" orientation="longitude" pageId="9" pageNumber="460" precision="1" value="111.997604">111.997601°E</geoCoordinate>
|
||||
; ca.
|
||||
<quantity id="EB609B3EFFA9FFCBFAF16B5F54809567" box="[1363,1436,331,357]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="9" pageNumber="460" unit="m" value="490.0">
|
||||
<elevation id="A7B5D1E8FFA9FFCBFAF16B5F54809567" box="[1363,1436,331,357]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="9" pageNumber="460" unit="m" value="490.0">490 m</elevation>
|
||||
</quantity>
|
||||
a.s.l.), collected by
|
||||
<collectorName id="816D530DFFA9FFCBFED06B7B50F5958B" box="[370,489,367,393]" pageId="9" pageNumber="460">Jian Wang</collectorName>
|
||||
,
|
||||
<collectorName id="816D530DFFA9FFCBFE566B7B53BE958B" box="[500,674,367,393]" pageId="9" pageNumber="460">Zhao-Chi Zeng</collectorName>
|
||||
,
|
||||
<collectorName id="816D530DFFA9FFCBFD0C6B7B527D958B" box="[686,865,367,393]" pageId="9" pageNumber="460">Hong-Hui Chen</collectorName>
|
||||
and
|
||||
<collectorName id="816D530DFFA9FFCBFC3A6B7B5506958B" box="[920,1050,367,393]" pageId="9" pageNumber="460">Shi-Shi Lin</collectorName>
|
||||
on
|
||||
<date id="5826101BFFA9FFCBFBE66B7B541D958B" box="[1092,1281,367,393]" pageId="9" pageNumber="460" value="2024-01-17">
|
||||
<collectingDate id="4862E9F3FFA9FFCBFBE66B7B541D958B" box="[1092,1281,367,393]" pageId="9" pageNumber="460" value="2024-01-17">17 January 2024</collectingDate>
|
||||
</date>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="64826550FFA9FFCBFF656B87551F961B" pageId="9" pageNumber="460" type="etymology">
|
||||
<paragraph id="2C2736DBFFA9FFCBFF656B87551F961B" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="460">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFF656B87505295AF" bold="true" box="[199,334,403,429]" pageId="9" pageNumber="460">Etymology.</emphasis>
|
||||
The specific epithet
|
||||
<taxonomicName id="EB984D58FFA9FFCBFDE56B87539495AE" authorityName="Wang and Zeng" authorityYear="2024" box="[583,648,403,429]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="pepe">
|
||||
“
|
||||
<emphasis id="1EECEAC9FFA9FFCBFDF16B87539495AE" box="[595,648,403,428]" italics="true" pageId="9" pageNumber="460">pepe</emphasis>
|
||||
</taxonomicName>
|
||||
” is named after “Pepe the Frog”, also known as the “Sad Frog”, a webcomic and international. The swollen lips of this new species (
|
||||
<figureCitation id="B4A32A5EFFA9FFCBFCDD6BA352FF95D3" box="[895,995,439,465]" captionStart="FIGURE 3" captionStartId="10.[152,255,1647,1671]" captionTargetBox="[158,1429,188,1617]" captionTargetId="figure-27@10.[151,1436,181,1623]" captionTargetPageId="10" captionText="FIGURE 3. The holotype (GEP a207) of Boulenophrys pepe sp. nov. in life: A. dorsal view; B. dorsolateral view; C. ventral view; D. ventral view of head, arrows showing swollen lips and protrude jaws; E. ventral view of hand; F. ventral view of foot." figureDoi="http://doi.org/10.5281/zenodo.13914793" httpUri="https://zenodo.org/record/13914793/files/figure.png" pageId="9" pageNumber="460">Figs. 3D</figureCitation>
|
||||
,
|
||||
<figureCitation id="B4A32A5EFFA9FFCBFC4D6BA3550E95D3" box="[1007,1042,439,465]" captionStart="FIGURE 4" captionStartId="11.[152,255,1904,1928]" captionTargetBox="[157,1429,279,1874]" captionTargetId="figure-62@11.[151,1435,273,1880]" captionTargetPageId="11" captionText="FIGURE 4. The paratypes SYS a009350 (A–D) and GEP a209 (E–F) of Boulenophrys pepe sp. nov. in life: A. dorsal view; B. dorsolateral view; C. ventral view; D. lateral view of head, with an arrow showing swollen lips and protrude jaws; E. dorsolateral view; F. ventral view." figureDoi="http://doi.org/10.5281/zenodo.13914795" httpUri="https://zenodo.org/record/13914795/files/figure.png" pageId="9" pageNumber="460">4D</figureCitation>
|
||||
) make it look like to has a sad face resembling to Pepe. Moreover, we name the new species after the well-known internet figure in the hope that it could bring attention to the conservation of the micro-endemic genus of frogs.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="64826550FFA9FFCBFF65683650E390C7" pageId="9" pageNumber="460" type="diagnosis">
|
||||
<paragraph id="2C2736DBFFA9FFCBFF656836547C9037" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="460">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFF656836505E963E" bold="true" box="[199,322,546,572]" pageId="9" pageNumber="460">Diagnosis.</emphasis>
|
||||
The new species can be diagnosed from its congeners by a combination of the following characters: (1) body size small and stout, SVL
|
||||
<quantity id="EB609B3EFFA9FFCBFD86685253DB9663" box="[548,711,582,609]" metricMagnitude="-2" metricUnit="m" metricValue="3.5849999999999995" metricValueMax="3.6399999999999997" metricValueMin="3.53" pageId="9" pageNumber="460" unit="mm" value="35.849999999999994" valueMax="36.4" valueMin="35.3">35.3–36.4 mm</quantity>
|
||||
(n = 4) in adult males; (2) head width distinctly larger than head length, (HDW/HDL 1.1–1.2); (3) lips swollen, upper and lower jaws distinctly protrude; tympanic region oblique, tympanum distinct and visible in dorsal view; (4) tongue feebly notched posteriorly; (5) vomerine ridges strong, vomerine teeth weak; (6) dorsal skin rough with dense granules, discontinuous X-shaped ridge on center of dorsum, discontinuous dorsolateral ridges present, dense large tubercles on flanks, dorsal limbs with discontinuous transverse ridges and tubercles, ventral skin smooth; (7) outer margin of upper eyelid with a small horn-like prominent tubercle; supratympanic fold distinct and narrow, curving posteroventrally to above arm; (8) two metacarpal tubercles distinct, inner one observably enlarged; distinct subarticular tubercle at base of each finger; (9) relative finger lengths I <II <IV <III; (10) heels not meeting when hindlimbs folded; tibiotarsal articulation reaching the posterior corner of eye; (11) toes without webbing and lateral fringes; (12) inner metatarsal tubercle long ovoid, outer one absent; (13) dorsal surface dark brown to yellowish brown, dark brown triangular marking between eyes, dorsal surface of forearm with two wide oblique dark brown bands, and dorsal surface of hindlimbs and digits with dark brown transverse bands; (14) dense tiny spines on surface of sacral region and dorsal hindlimbs in males; and (15) dense nuptial spines on dorsal bases of fingers I and II in breeding adult males; subgular vocal sac present in males.
|
||||
</paragraph>
|
||||
<paragraph id="2C2736DBFFA9FFCBFF656E2A50E390C7" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="460">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFF656E2A5071905A" bold="true" box="[199,365,1086,1112]" pageId="9" pageNumber="460">Comparisons.</emphasis>
|
||||
<taxonomicName id="EB984D58FFA9FFCBFEDA6E2B534F905A" authorityName="Wang & Lin & Gan & Chen & Yu & Pan & Xiao & Zeng" authorityYear="2024" box="[376,595,1087,1113]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="pepe" status="sp. nov.">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFEDA6E2B534F905A" box="[376,595,1087,1113]" italics="true" pageId="9" pageNumber="460">Boulenophrys pepe</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="1EECEAC9FFA9FFCBFDFC6E2B53A7905B" bold="true" box="[606,699,1087,1113]" pageId="9" pageNumber="460">
|
||||
<taxonomicNameLabel id="05DF57B2FFA9FFCBFDFC6E2B53A7905B" box="[606,699,1087,1113]" pageId="9" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is the sister species to
|
||||
<taxonomicName id="EB984D58FFA9FFCBFC716E2B552B905B" baseAuthorityName="Wang, Li and Zhao" baseAuthorityYear="2014" box="[979,1079,1087,1113]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="obesa">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFC716E2B552B905B" box="[979,1079,1087,1113]" italics="true" pageId="9" pageNumber="460">B. obesa</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="B4A32A5EFFA9FFCBFBE86E2B55AD905B" box="[1098,1201,1087,1113]" captionStart="FIGURE 2" captionStartId="8.[152,255,1649,1673]" captionTargetBox="[153,1436,741,1623]" captionTargetId="figure-244@8.[151,1436,741,1625]" captionTargetPageId="8" captionText="FIGURE 2. ML tree based on the partial DNA sequences of the mitochondrial 16S and COI genes, with Bootstrap Support (BS) and Posterior Probabilities (BPP) at nodes." figureDoi="http://doi.org/10.5281/zenodo.13914791" httpUri="https://zenodo.org/record/13914791/files/figure.png" pageId="9" pageNumber="460">Figure 2</figureCitation>
|
||||
). However, the new species differs from the later by the vomerine teeth present (vs. absent), tongue notched posteriorly (vs. rounded), lips swollen (vs. normal), upper and lower jaws distinctly protruded (vs. not protruded), and toes without webbing (vs. with weak fleshy webbing).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="64826550FFA9FFCEFF656EDB55BE90A3" lastPageId="12" lastPageNumber="463" pageId="9" pageNumber="460" type="description">
|
||||
<paragraph id="2C2736DBFFA9FFCBFF656EDB53CD9133" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="460">
|
||||
<taxonomicName id="EB984D58FFA9FFCBFF656EDB508090EA" authorityName="Wang & Lin & Gan & Chen & Yu & Pan & Xiao & Zeng" authorityYear="2024" box="[199,412,1231,1257]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="pepe" status="sp. nov.">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFF656EDB508090EA" box="[199,412,1231,1257]" italics="true" pageId="9" pageNumber="460">Boulenophrys pepe</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="1EECEAC9FFA9FFCBFE026EDB50EB90EB" bold="true" box="[416,503,1231,1257]" pageId="9" pageNumber="460">
|
||||
<taxonomicNameLabel id="05DF57B2FFA9FFCBFE026EDB50EB90EB" box="[416,503,1231,1257]" pageId="9" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is further phylogenetically close to
|
||||
<taxonomicName id="EB984D58FFA9FFCBFCDB6EDB550490EB" baseAuthorityName="Messenger and Dahn" baseAuthorityYear="2019" box="[889,1048,1231,1257]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="ombrophila">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFCDB6EDB550490EB" box="[889,1048,1231,1257]" italics="true" pageId="9" pageNumber="460">B. ombrophila</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="B4A32A5EFFA9FFCBFB816EDB559E90EB" box="[1059,1154,1231,1257]" captionStart="FIGURE 2" captionStartId="8.[152,255,1649,1673]" captionTargetBox="[153,1436,741,1623]" captionTargetId="figure-244@8.[151,1436,741,1625]" captionTargetPageId="8" captionText="FIGURE 2. ML tree based on the partial DNA sequences of the mitochondrial 16S and COI genes, with Bootstrap Support (BS) and Posterior Probabilities (BPP) at nodes." figureDoi="http://doi.org/10.5281/zenodo.13914791" httpUri="https://zenodo.org/record/13914791/files/figure.png" pageId="9" pageNumber="460">Figure 2</figureCitation>
|
||||
), but can be distinguished from the latter by the vomerine ridges and vomerine teeth present (vs. both absent), tongue notched posteriorly (vs. rounded posteriorly) and lips swollen (vs. normal).
|
||||
</paragraph>
|
||||
<paragraph id="2C2736DBFFA9FFCBFF656F2F5095929B" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="460">
|
||||
<taxonomicName id="EB984D58FFA9FFCBFF656F2F50BF9156" authorityName="Wang & Lin & Gan & Chen & Yu & Pan & Xiao & Zeng" authorityYear="2024" box="[199,419,1339,1365]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="pepe" status="sp. nov.">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFF656F2F50BF9156" box="[199,419,1339,1365]" italics="true" pageId="9" pageNumber="460">Boulenophrys pepe</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="1EECEAC9FFA9FFCBFE0C6F2F53109157" bold="true" box="[430,524,1339,1365]" pageId="9" pageNumber="460">
|
||||
<taxonomicNameLabel id="05DF57B2FFA9FFCBFE0C6F2F53109157" box="[430,524,1339,1365]" pageId="9" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
can be distinguished by its heels not meeting when flexed hindlimbs held at right angles to body axis, from the following congeners:
|
||||
<taxonomicName id="EB984D58FFA9FFCBFC906F4B52C6917B" baseAuthorityName="Li, Lu, Liu and Wang" baseAuthorityYear="2020" box="[818,986,1375,1401]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="anlongensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFC906F4B52C6917B" box="[818,986,1375,1401]" italics="true" pageId="9" pageNumber="460">B. anlongensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFC496F4B55A4917B" baseAuthorityName="Wu, Li, Liu, Wang and Wu" baseAuthorityYear="2020" box="[1003,1208,1375,1401]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="baishanzuensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFC496F4B55A4917B" box="[1003,1208,1375,1401]" italics="true" pageId="9" pageNumber="460">B. baishanzuensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFB6B6F4B546E917B" baseAuthorityName="Jiang, Fei and Ye" baseAuthorityYear="2009" box="[1225,1394,1375,1401]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="binlingensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFB6B6F4B546E917B" box="[1225,1394,1375,1401]" italics="true" pageId="9" pageNumber="460">B. binlingensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFA266F4B503F919F" baseAuthorityName="Shen" baseAuthorityYear="1994" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="caudoprocta">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFA266F4B503F919F" italics="true" pageId="9" pageNumber="460">B. caudoprocta</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFE936F975091919F" baseAuthorityName="Wang and Liu" baseAuthorityYear="2014" box="[305,397,1411,1437]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="cheni">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFE936F975091919F" box="[305,397,1411,1437]" italics="true" pageId="9" pageNumber="460">B. cheni</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFE396F97535F919F" baseAuthorityName="Xu, Li, Liu, Wei and Wang" baseAuthorityYear="2020" box="[411,579,1411,1437]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="chishuiensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFE396F97535F919F" box="[411,579,1411,1437]" italics="true" pageId="9" pageNumber="460">B. chishuiensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFDF26F97520B919E" baseAuthorityName="Luo, Wang, Wang, Lu, Wang, Deng and Zhou" baseAuthorityYear="2021" box="[592,791,1411,1436]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="congjiangensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFDF26F97520B919E" box="[592,791,1411,1436]" italics="true" pageId="9" pageNumber="460">B. congjiangensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFC876F9752CE919F" baseAuthorityName="Rao and Yang" baseAuthorityYear="1997" box="[805,978,1411,1437]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="daweimontis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFC876F9752CE919F" box="[805,978,1411,1437]" italics="true" pageId="9" pageNumber="460">B. daweimontis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFC7D6F97557C919F" authorityName="Zeng, Wang, Chen, Xiao, Zhan, Li and Lin" authorityYear="2024" box="[991,1120,1411,1437]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="elongata">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFC7D6F97557C919F" box="[991,1120,1411,1437]" italics="true" pageId="9" pageNumber="460">B. elongata</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFBCC6F97543B919F" baseAuthorityName="Zhang, Liang, Ran and Shen" baseAuthorityYear="2012" box="[1134,1319,1411,1437]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="fanjingmontis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFBCC6F97543B919F" box="[1134,1319,1411,1437]" italics="true" pageId="9" pageNumber="460">B. fanjingmontis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFA976F975489919E" baseAuthorityName="Liu, Li, Wei, Xu, Cheng, Wang and Wu" baseAuthorityYear="2020" box="[1333,1429,1411,1436]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="jiangi">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFA976F975489919E" box="[1333,1429,1411,1436]" italics="true" pageId="9" pageNumber="460">B. jiangi</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFF356FB3504E91C3" baseAuthorityName="Fei and Ye" baseAuthorityYear="1983" box="[151,338,1447,1473]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="jingdongensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFF356FB3504E91C3" box="[151,338,1447,1473]" italics="true" pageId="9" pageNumber="460">B. jingdongensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFEFD6FB3530591C2" baseAuthorityName="Wang" baseAuthorityYear="2012" box="[351,537,1447,1472]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="jinggangensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFEFD6FB3530591C2" box="[351,537,1447,1472]" italics="true" pageId="9" pageNumber="460">B. jinggangensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFD846FB353DA91C3" baseAuthorityName="Wang, Zeng, Lyu and Wang" baseAuthorityYear="2019" box="[550,710,1447,1473]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="jiulianensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFD846FB353DA91C3" box="[550,710,1447,1473]" italics="true" pageId="9" pageNumber="460">B. jiulianensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFD716FB3526791C3" baseAuthorityName="Li, Xu, Liu, Jiang, Wei and Wang" baseAuthorityYear="2018" box="[723,891,1447,1473]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="leishanensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFD716FB3526791C3" box="[723,891,1447,1473]" italics="true" pageId="9" pageNumber="460">B. leishanensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFC2A6FB3551791C3" baseAuthorityName="Zhang, Li, Xiao, Li, Pan, Wang, Zhang and Zhou" baseAuthorityYear="2017" box="[904,1035,1447,1473]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="liboensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFC2A6FB3551791C3" box="[904,1035,1447,1473]" italics="true" pageId="9" pageNumber="460">B. liboensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFBBA6FB3554791C3" baseAuthorityName="Wang and Yang" baseAuthorityYear="2014" box="[1048,1115,1447,1473]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="lini">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFBBA6FB3554791C3" box="[1048,1115,1447,1473]" italics="true" pageId="9" pageNumber="460">B. lini</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFBCB6FB3541891C3" baseAuthorityName="Shi, Li, Zhu, Jiang, Jiang and Wang" baseAuthorityYear="2021" box="[1129,1284,1447,1473]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="lushuiensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFBCB6FB3541891C3" box="[1129,1284,1447,1473]" italics="true" pageId="9" pageNumber="460">B. lushuiensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFAB06FB3548991C3" baseAuthorityName="Lyu, Wang and Zhao" baseAuthorityYear="2020" box="[1298,1429,1447,1473]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="mirabilis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFAB06FB3548991C3" box="[1298,1429,1447,1473]" italics="true" pageId="9" pageNumber="460">B. mirabilis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFF356FDF504B91E7" baseAuthorityName="Wang, Lyu and Wang" baseAuthorityYear="2019" box="[151,343,1483,1509]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="mufumontana">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFF356FDF504B91E7" box="[151,343,1483,1509]" italics="true" pageId="9" pageNumber="460">B. mufumontana</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFEC96FDF530391E7" baseAuthorityName="Lyu, Wang, Liu and Wang" baseAuthorityYear="2019" box="[363,543,1483,1509]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="nanlingensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFEC96FDF530391E7" box="[363,543,1483,1509]" italics="true" pageId="9" pageNumber="460">B. nanlingensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFD916FDF53C591E6" baseAuthorityName="Liu" baseAuthorityYear="1950" box="[563,729,1483,1508]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="omeimontis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFD916FDF53C591E6" box="[563,729,1483,1508]" italics="true" pageId="9" pageNumber="460">B. omeimontis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFD4C6FDF52FE91E7" baseAuthorityName="Bourret" baseAuthorityYear="1937" box="[750,994,1483,1509]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="palpebralespinosa">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFD4C6FDF52FE91E7" box="[750,994,1483,1509]" italics="true" pageId="9" pageNumber="460">B. palpebralespinosa</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFC556FDF55B491E7" box="[1015,1192,1483,1509]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="qianbeiensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFC556FDF55B491E7" box="[1015,1192,1483,1509]" italics="true" pageId="9" pageNumber="460">B. qianbeiensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFB1F6FDF546C91E7" baseAuthorityName="Jiang, Ye and Fei" baseAuthorityYear="2008" box="[1213,1392,1483,1509]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="sangzhiensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFB1F6FDF546C91E7" box="[1213,1392,1483,1509]" italics="true" pageId="9" pageNumber="460">B. sangzhiensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFA266FDF5032920A" baseAuthorityName="Lyu and Wang" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="sanmingensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFA266FDF5032920A" italics="true" pageId="9" pageNumber="460">B. sanmingensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFE996FFB50FF920B" baseAuthorityName="Lyu, Liu and Wang" baseAuthorityYear="2020" box="[315,483,1519,1545]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="shimentaina">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFE996FFB50FF920B" box="[315,483,1519,1545]" italics="true" pageId="9" pageNumber="460">B. shimentaina</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFE526FFB53DD920B" baseAuthorityName="Wang, Deng, Liu, Wu and Liu" baseAuthorityYear="2019" box="[496,705,1519,1545]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="shunhuangensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFE526FFB53DD920B" box="[496,705,1519,1545]" italics="true" pageId="9" pageNumber="460">B. shunhuangensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFD6C6FFB5223920A" baseAuthorityName="Liu and Hu" baseAuthorityYear="1973" box="[718,831,1519,1544]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="spinata">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFD6C6FFB5223920A" box="[718,831,1519,1544]" italics="true" pageId="9" pageNumber="460">B. spinata</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFCEE6FFB551E920A" baseAuthorityName="Lyu and Wang" baseAuthorityYear="2021" box="[844,1026,1519,1544]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="sanmingensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFCEE6FFB551E920A" box="[844,1026,1519,1544]" italics="true" pageId="9" pageNumber="460">B. sanmingensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFBAD6FFB55AF920B" baseAuthorityName="Wang and Lyu" baseAuthorityYear="2021" box="[1039,1203,1519,1545]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="tongboensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFBAD6FFB55AF920B" box="[1039,1203,1519,1545]" italics="true" pageId="9" pageNumber="460">B. tongboensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFB626FFB548A920B" baseAuthorityName="Shen, Mo & Li" baseAuthorityYear="2010" box="[1216,1430,1519,1545]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="tuberogranulata">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFB626FFB548A920B" box="[1216,1430,1519,1545]" italics="true" pageId="9" pageNumber="460">B. tuberogranulata</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFF356C075061922F" baseAuthorityName="Ye and Fei" baseAuthorityYear="1995" box="[151,381,1555,1581]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="wuliangshanensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFF356C075061922F" box="[151,381,1555,1581]" italics="true" pageId="9" pageNumber="460">B. wuliangshanensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFE286C07533A922E" baseAuthorityName="Wang, Wu, Peng, Shi, Lu and Wu" baseAuthorityYear="2020" box="[394,550,1555,1580]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="xianjuensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFE286C07533A922E" box="[394,550,1555,1580]" italics="true" pageId="9" pageNumber="460">B. xianjuensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFD966C0753E4922E" baseAuthorityName="Lyu, Zeng and Wang" baseAuthorityYear="2020" box="[564,760,1555,1580]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="yangmingensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFD966C0753E4922E" box="[564,760,1555,1580]" italics="true" pageId="9" pageNumber="460">B. yangmingensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFCA46C0752A5922F" authorityName="Qi, Mo, Lyu, Wang and Wang" authorityYear="2021" box="[774,953,1555,1581]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="yaoshanensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFCA46C0752A5922F" box="[774,953,1555,1581]" italics="true" pageId="9" pageNumber="460">B. yaoshanensis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="EB984D58FFA9FFCBFC4D6C07558C922F" authorityName="Qi, Lyu, Wang and Wang" authorityYear="2021" box="[1007,1168,1555,1581]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="yingdeensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFC4D6C07558C922F" box="[1007,1168,1555,1581]" italics="true" pageId="9" pageNumber="460">B. yingdeensis</emphasis>
|
||||
</taxonomicName>
|
||||
(vs. heels overlapping); from
|
||||
<taxonomicName id="EB984D58FFA9FFCBFF756C2350849253" baseAuthorityName="Ye and Fei" baseAuthorityYear="1995" box="[215,408,1591,1617]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="binchuanensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFF756C2350849253" box="[215,408,1591,1617]" italics="true" pageId="9" pageNumber="460">B. binchuanensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFE0A6C2353239253" baseAuthorityName="Wang, Liu and Jiang" baseAuthorityYear="2017" box="[424,575,1591,1617]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="lishuiensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFE0A6C2353239253" box="[424,575,1591,1617]" italics="true" pageId="9" pageNumber="460">B. lishuiensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFDED6C2353AF9252" baseAuthorityName="Stejneger" baseAuthorityYear="1926" box="[591,691,1591,1616]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="minor">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFDED6C2353AF9252" box="[591,691,1591,1616]" italics="true" pageId="9" pageNumber="460">B. minor</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFD616C2352989252" baseAuthorityName="Lyu, Zeng and Wang" baseAuthorityYear="2020" box="[707,900,1591,1616]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="xiangnanensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFD616C2352989252" box="[707,900,1591,1616]" italics="true" pageId="9" pageNumber="460">B. xiangnanensis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="EB984D58FFA9FFCBFC1D6C23559E9253" authorityName="Lyu and Wang" authorityYear="2023" box="[959,1154,1591,1617]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="xuefengmontis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFC1D6C23559E9253" box="[959,1154,1591,1617]" italics="true" pageId="9" pageNumber="460">B. xuefengmontis</emphasis>
|
||||
</taxonomicName>
|
||||
(vs. heels just meeting); and from
|
||||
<taxonomicName id="EB984D58FFA9FFCBFEA46C4F50769277" baseAuthorityName="Wu, Suwannapoom, Poyarkov, Pawangkhanant, Xu, Jin, Murphy and Wu et al. 2019 Che" baseAuthorityYear="2019" box="[262,362,1627,1653]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="angka">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFEA46C4F50769277" box="[262,362,1627,1653]" italics="true" pageId="9" pageNumber="460">B. angka</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFEDB6C4F53019277" baseAuthorityName="Lyu, Wang and Wang" baseAuthorityYear="2021" box="[377,541,1627,1653]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="daiyunensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFEDB6C4F53019277" box="[377,541,1627,1653]" italics="true" pageId="9" pageNumber="460">B. daiyunensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="1EECEAC9FFA9FFCBFD8E6C4F52819277" box="[556,925,1627,1653]" italics="true" pageId="9" pageNumber="460">
|
||||
<taxonomicName id="EB984D58FFA9FFCBFD8E6C4F53F89277" baseAuthorityName="Ye, Fei and Xie" baseAuthorityYear="2007" box="[556,740,1627,1653]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="baolongensis">B. baolongensis</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFD4D6C4F52819277" baseAuthorityName="Ye and Fei" baseAuthorityYear="1995" box="[751,925,1627,1653]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="wushanensis">B. wushanensis</taxonomicName>
|
||||
</emphasis>
|
||||
and
|
||||
<taxonomicName id="EB984D58FFA9FFCBFC776C4F55649277" authorityName="Qi, Wang, Lyu and Wang" authorityYear="2021" box="[981,1144,1627,1653]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="yunkaiensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFC776C4F55649277" box="[981,1144,1627,1653]" italics="true" pageId="9" pageNumber="460">B. yunkaiensis</emphasis>
|
||||
</taxonomicName>
|
||||
(vs. heels just meeting or slightly overlapping).
|
||||
</paragraph>
|
||||
<paragraph id="2C2736DBFFA9FFCBFF656CB755049307" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="460">
|
||||
For the other congeners,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFE526CB753D092BE" authorityName="Wang & Lin & Gan & Chen & Yu & Pan & Xiao & Zeng" authorityYear="2024" box="[496,716,1699,1725]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="pepe" status="sp. nov.">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFE526CB753D092BE" box="[496,716,1699,1725]" italics="true" pageId="9" pageNumber="460">Boulenophrys pepe</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="1EECEAC9FFA9FFCBFD756CB7522992BF" bold="true" box="[727,821,1699,1725]" pageId="9" pageNumber="460">
|
||||
<taxonomicNameLabel id="05DF57B2FFA9FFCBFD756CB7522992BF" box="[727,821,1699,1725]" pageId="9" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
can be distinguished by the vomerine teeth present, from the following congeners:
|
||||
<taxonomicName id="EB984D58FFA9FFCBFE556CD3534A92E2" baseAuthorityName="Wang, Li and Jin" baseAuthorityYear="2014" box="[503,598,1735,1760]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="acuta">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFE556CD3534A92E2" box="[503,598,1735,1760]" italics="true" pageId="9" pageNumber="460">B. acuta</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFDC76CD353F792E3" baseAuthorityName="Boulenger" baseAuthorityYear="1899" box="[613,747,1735,1761]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="boettgeri">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFDC76CD353F792E3" box="[613,747,1735,1761]" italics="true" pageId="9" pageNumber="460">B. boettgeri</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFD586CD352AF92E3" baseAuthorityName="Nguyen, Pham, Nguyen, Luong and Ziegler" baseAuthorityYear="2020" box="[762,947,1735,1761]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="caobangensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFD586CD352AF92E3" box="[762,947,1735,1761]" italics="true" pageId="9" pageNumber="460">B. caobangensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFC606CD3550192E3" baseAuthorityName="Lyu, Zeng, Wang and Wang" baseAuthorityYear="2021" box="[962,1053,1735,1761]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="daoji">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFC606CD3550192E3" box="[962,1053,1735,1761]" italics="true" pageId="9" pageNumber="460">B. daoji</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFB8E6CD355BE92E3" authorityName="Wang, Zeng, Lyu, Xiao and Wang" authorityYear="2022" box="[1068,1186,1735,1761]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="hungtai">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFB8E6CD355BE92E3" box="[1068,1186,1735,1761]" italics="true" pageId="9" pageNumber="460">B. hungtai</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFB136CD3546892E3" authorityName="Qian, Hu, Mo, Gao, Zhang and Yang" authorityYear="2023" box="[1201,1396,1735,1761]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="hengshanensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFB136CD3546892E3" box="[1201,1396,1735,1761]" italics="true" pageId="9" pageNumber="460">B. hengshanensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFA266CD350079307" baseAuthorityName="Pope" baseAuthorityYear="1929" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="kuatunensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFA266CD350079307" italics="true" pageId="9" pageNumber="460">B. kuatunensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFE8B6CFF50E89307" baseAuthorityName="Tian and Sun" baseAuthorityYear="1995" box="[297,500,1771,1797]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="shuichengensis">B. shuichengensis</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="EB984D58FFA9FFCBFD886CFF53C69306" baseAuthorityName="Wang, Lyu and Wang" baseAuthorityYear="2019" box="[554,730,1771,1796]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="wugongensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFD886CFF53C69306" box="[554,730,1771,1796]" italics="true" pageId="9" pageNumber="460">B. wugongensis</emphasis>
|
||||
</taxonomicName>
|
||||
(vs. vomerine teeth absent).
|
||||
</paragraph>
|
||||
<paragraph id="2C2736DBFFA9FFCBFF656D1B54739373" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="460">
|
||||
<taxonomicName id="EB984D58FFA9FFCBFF656D1B5082932A" authorityName="Wang & Lin & Gan & Chen & Yu & Pan & Xiao & Zeng" authorityYear="2024" box="[199,414,1807,1833]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="pepe" status="sp. nov.">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFF656D1B5082932A" box="[199,414,1807,1833]" italics="true" pageId="9" pageNumber="460">Boulenophrys pepe</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="1EECEAC9FFA9FFCBFE076D1B50E2932B" bold="true" box="[421,510,1807,1833]" pageId="9" pageNumber="460">
|
||||
<taxonomicNameLabel id="05DF57B2FFA9FFCBFE076D1B50E2932B" box="[421,510,1807,1833]" pageId="9" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
can be further distinguished by its lateral fringes and webbing on toes absent, from the following congeners:
|
||||
<taxonomicName id="EB984D58FFA9FFCBFE156D27537C934F" baseAuthorityName="Inger and Romer" baseAuthorityYear="1961" box="[439,608,1843,1869]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="brachykolos">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFE156D27537C934F" box="[439,608,1843,1869]" italics="true" pageId="9" pageNumber="460">B. brachykolos</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFDCD6D27522A934F" baseAuthorityName="Wang and Wang" baseAuthorityYear="2019" box="[623,822,1843,1869]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="dongguanensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFDCD6D27522A934F" box="[623,822,1843,1869]" italics="true" pageId="9" pageNumber="460">B. dongguanensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFCE76D27551E934F" authorityName="Wang, Zeng, Lyu and Wang" authorityYear="2022" box="[837,1026,1843,1869]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="fengshunensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFCE76D27551E934F" box="[837,1026,1843,1869]" italics="true" pageId="9" pageNumber="460">B. fengshunensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="1EECEAC9FFA9FFCBFBB36D27558F934F" box="[1041,1171,1843,1869]" italics="true" pageId="9" pageNumber="460">
|
||||
B,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFB936D27558F934F" baseAuthorityName="Wang, Liu, Lyu, Zeng and Wang" baseAuthorityYear="2017" box="[1073,1171,1843,1869]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="insularis">insularis</taxonomicName>
|
||||
</emphasis>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFB006D275450934F" baseAuthorityName="Wang, Zeng and Wang" baseAuthorityYear="2019" box="[1186,1356,1843,1869]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="nankunensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFB006D275450934F" box="[1186,1356,1843,1869]" italics="true" pageId="9" pageNumber="460">B. nankunensis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="EB984D58FFA9FFCBFA266D2750019372" authorityName="Wang, Zeng, Lyu, Xiao and Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="puningensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFA266D2750019372" italics="true" pageId="9" pageNumber="460">B. puningensis</emphasis>
|
||||
</taxonomicName>
|
||||
(vs. toes with rudimentary webbing); and from
|
||||
<taxonomicName id="EB984D58FFA9FFCBFC946D4352D49373" baseAuthorityName="Tapley, Cutajar, Mahony, Chung, Dau, Nguyen, Luong and Rowley" baseAuthorityYear="2017" box="[822,968,1879,1905]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="rubrimera">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFC946D4352D49373" box="[822,968,1879,1905]" italics="true" pageId="9" pageNumber="460">B. rubrimera</emphasis>
|
||||
</taxonomicName>
|
||||
(vs. toes with narrow lateral fringes).
|
||||
</paragraph>
|
||||
<paragraph id="2C2736DBFFA9FFCBFF656D6F552193BB" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="460">
|
||||
<taxonomicName id="EB984D58FFA9FFCBFF656D6F50819396" authorityName="Wang & Lin & Gan & Chen & Yu & Pan & Xiao & Zeng" authorityYear="2024" box="[199,413,1915,1941]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="pepe" status="sp. nov.">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFF656D6F50819396" box="[199,413,1915,1941]" italics="true" pageId="9" pageNumber="460">Boulenophrys pepe</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="1EECEAC9FFA9FFCBFE016D6F50E79397" bold="true" box="[419,507,1915,1941]" pageId="9" pageNumber="460">
|
||||
<taxonomicNameLabel id="05DF57B2FFA9FFCBFE016D6F50E79397" box="[419,507,1915,1941]" pageId="9" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
can be distinguished by its tongue notched posterior, from the remaining congeners:
|
||||
<taxonomicName id="EB984D58FFA9FFCBFF356D8B504293BB" baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong and Rowley" baseAuthorityYear="2018" box="[151,350,1951,1977]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="hoanglienensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFF356D8B504293BB" box="[151,350,1951,1977]" italics="true" pageId="9" pageNumber="460">B. hoanglienensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EB984D58FFA9FFCBFECE6D8B533F93BB" baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong and Rowley" baseAuthorityYear="2018" box="[364,547,1951,1977]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="fansipanensis">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFECE6D8B533F93BB" box="[364,547,1951,1977]" italics="true" pageId="9" pageNumber="460">B. fansipanensis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="EB984D58FFA9FFCBFDF86D8B53D993BB" baseAuthorityName="Tapley, Cutaja, Nguyen, Portway, Mahony, Nguyen, Harding, Luong and Rowley" baseAuthorityYear="2021" box="[602,709,1951,1977]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="frigida">
|
||||
<emphasis id="1EECEAC9FFA9FFCBFDF86D8B53D993BB" box="[602,709,1951,1977]" italics="true" pageId="9" pageNumber="460">B. frigida</emphasis>
|
||||
</taxonomicName>
|
||||
(vs. tongue rounded posteriorly).
|
||||
</paragraph>
|
||||
<caption id="78E76653FFAAFFC8FF3A6C7B51DB92CD" ID-DOI="http://doi.org/10.5281/zenodo.13914793" ID-Zenodo-Dep="13914793" httpUri="https://zenodo.org/record/13914793/files/figure.png" pageId="10" pageNumber="461" startId="10.[152,255,1647,1671]" targetBox="[158,1429,188,1617]" targetPageId="10" targetType="figure">
|
||||
<paragraph id="2C2736DBFFAAFFC8FF3A6C7B51DB92CD" blockId="10.[151,1436,1647,1743]" pageId="10" pageNumber="461">
|
||||
<emphasis id="1EECEAC9FFAAFFC8FF3A6C7B50059285" bold="true" box="[152,281,1647,1671]" pageId="10" pageNumber="461">FIGURE 3.</emphasis>
|
||||
The holotype (GEP a207) of
|
||||
<taxonomicName id="EB984D58FFAAFFC8FDEB6C7B52119285" authorityName="Wang & Lin & Gan & Chen & Yu & Pan & Xiao & Zeng" authorityYear="2024" box="[585,781,1647,1671]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="461" phylum="Chordata" rank="species" species="pepe" status="sp. nov.">
|
||||
<emphasis id="1EECEAC9FFAAFFC8FDEB6C7B52119285" box="[585,781,1647,1671]" italics="true" pageId="10" pageNumber="461">Boulenophrys pepe</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="1EECEAC9FFAAFFC8FCB66C645279928A" bold="true" box="[788,869,1648,1672]" pageId="10" pageNumber="461">
|
||||
<taxonomicNameLabel id="05DF57B2FFAAFFC8FCB66C645279928A" box="[788,869,1648,1672]" pageId="10" pageNumber="461" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
in life:
|
||||
<emphasis id="1EECEAC9FFAAFFC8FC156C7B52D39285" bold="true" box="[951,975,1647,1671]" pageId="10" pageNumber="461">A.</emphasis>
|
||||
dorsal view;
|
||||
<emphasis id="1EECEAC9FFAAFFC8FBF86C64556D928A" bold="true" box="[1114,1137,1648,1672]" pageId="10" pageNumber="461">B.</emphasis>
|
||||
dorsolateral view;
|
||||
<emphasis id="1EECEAC9FFAAFFC8FA946C7B54539285" bold="true" box="[1334,1359,1647,1671]" pageId="10" pageNumber="461">C.</emphasis>
|
||||
ventral view;
|
||||
<emphasis id="1EECEAC9FFAAFFC8FF7A6C8051ED92AE" bold="true" box="[216,241,1684,1708]" pageId="10" pageNumber="461">D.</emphasis>
|
||||
ventral view of head, arrows showing swollen lips and protrude jaws;
|
||||
<emphasis id="1EECEAC9FFAAFFC8FC786C8052ED92AE" bold="true" box="[986,1009,1684,1708]" pageId="10" pageNumber="461">E.</emphasis>
|
||||
ventral view of hand;
|
||||
<emphasis id="1EECEAC9FFAAFFC8FB406C8055E992AE" bold="true" box="[1250,1269,1684,1708]" pageId="10" pageNumber="461">F.</emphasis>
|
||||
ventral view of foot.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="2C2736DBFFAAFFC9FF656D35523A94FB" blockId="10.[151,1437,1825,2032]" lastBlockId="11.[151,1436,150,249]" lastPageId="11" lastPageNumber="462" pageId="10" pageNumber="461">
|
||||
<emphasis id="1EECEAC9FFAAFFC8FF656D3550FF933E" bold="true" box="[199,483,1825,1852]" pageId="10" pageNumber="461">
|
||||
Description of
|
||||
<typeStatus id="F3238879FFAAFFC8FED46D3650C3933E" box="[374,479,1826,1852]" pageId="10" pageNumber="461" type="holotype">holotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Adult male, body stout, SVL
|
||||
<quantity id="EB609B3EFFAAFFC8FC926D36528A933E" box="[816,918,1826,1852]" metricMagnitude="-2" metricUnit="m" metricValue="3.6399999999999997" pageId="10" pageNumber="461" unit="mm" value="36.4">36.4 mm</quantity>
|
||||
. Head width distinctly larger than head length, HWD/HDL 1.12; snout rounded in dorsal view, projecting, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; top of head flat; eyes moderate in size, ED 0.31 of HDL, pupil vertical, near diamondshaped; nostril obliquely ovoid; canthus rostralis well developed; loreal region slightly oblique; internasal distance slightly larger than interorbital distance; lips swollen, upper and lower jaws distinctly protrude, tympanic region oblique, tympanum distinct and visible in dorsal view; tympanum moderate in size, margin clear, upper margin in contact with supratympanic fold, lower margin in contact with upper lip, TD/ED 0.54; large ovoid choanae at base of maxilla; strong vomerine ridge and weak vomerine teeth present, maxillary teeth present; margin of tongue feebly notched posteriorly; presence of single subgular vocal sac.
|
||||
</paragraph>
|
||||
<caption id="78E76653FFABFFC9FF3A6D6450EC93D2" ID-DOI="http://doi.org/10.5281/zenodo.13914795" ID-Zenodo-Dep="13914795" httpUri="https://zenodo.org/record/13914795/files/figure.png" pageId="11" pageNumber="462" startId="11.[152,255,1904,1928]" targetBox="[157,1429,279,1874]" targetPageId="11" targetType="figure">
|
||||
<paragraph id="2C2736DBFFABFFC9FF3A6D6450EC93D2" blockId="11.[151,1437,1904,2000]" pageId="11" pageNumber="462">
|
||||
<emphasis id="1EECEAC9FFABFFC9FF3A6D645005938A" bold="true" box="[152,281,1904,1928]" pageId="11" pageNumber="462">FIGURE 4.</emphasis>
|
||||
The paratypes SYS a009350 (A–D) and GEP a209 (E–F) of
|
||||
<taxonomicName id="EB984D58FFABFFC9FC326D645548938A" authorityName="Wang & Lin & Gan & Chen & Yu & Pan & Xiao & Zeng" authorityYear="2024" box="[912,1108,1904,1928]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="462" phylum="Chordata" rank="species" species="pepe" status="sp. nov.">
|
||||
<emphasis id="1EECEAC9FFABFFC9FC326D645548938A" box="[912,1108,1904,1928]" italics="true" pageId="11" pageNumber="462">Boulenophrys pepe</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="1EECEAC9FFABFFC9FBF96D6455B1938A" bold="true" box="[1115,1197,1904,1928]" pageId="11" pageNumber="462">
|
||||
<taxonomicNameLabel id="05DF57B2FFABFFC9FBF96D6455B1938A" box="[1115,1197,1904,1928]" pageId="11" pageNumber="462" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
in life: A. dorsal view; B. dorsolateral view; C. ventral view; D. lateral view of head, with an arrow showing swollen lips and protrude jaws; E. dorsolateral view; F. ventral view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="2C2736DBFFACFFCEFF656A8352C495AF" blockId="12.[151,1437,151,1185]" pageId="12" pageNumber="463">Forearm length 0.23 of SVL, hand 0.24 of SVL; webbing absent between fingers, lateral fringes absent, relative finger length I <II <IV <III; tips of fingers slightly dilated, round; subarticular tubercles present, distinct; inner metacarpal tubercle observably enlarged, outer one slightly smaller; single nuptial pad bearing nuptial spines present on dorsal surface of first and second fingers, respectively. Hindlimbs short, tibio-tarsal articulation reaching forward to posterior conner of eye when hindlimb stretched along body; heels not meeting when flexed hindlimbs held at right angles to body axis; shank length 0.40 of SVL and foot length 0.55 of SVL; relative toe length I <II <V <III <IV; tips of toes round and slightly dilated; toes without lateral fringes and webbing; subarticular present and distinct; inner metatarsal tubercle long ovoid and lacking outer metatarsal tubercle.</paragraph>
|
||||
<paragraph id="2C2736DBFFACFFCEFF656BA3529E96AB" blockId="12.[151,1437,151,1185]" pageId="12" pageNumber="463">Dorsal skin rough and highly granular; dense large tubercles on flanks; single horn-like prominent tubercle on edge of upper eyelid; obvious supratympanic fold curving posteroventrally from posterior corner of eye to level above insertion of arm; upper lip, mandibular articulation, loreal, temporal region excluding tympanum, upper eyelid and surface around cloaca with conical tubercles; discontinuous X-shaped ridge on center of dorsum, discontinuous dorsolateral ridges present; dense large tubercles on flanks; dense tiny spines on surface of sacral region and dorsal hindlimbs; ventral surface relatively smooth; small pectoral gland closer to axilla; single femoral gland positioned on posterior surface of thigh at midpoint between knee and cloaca.</paragraph>
|
||||
<paragraph id="2C2736DBFFACFFCEFF6568A6540E97A7" blockId="12.[151,1437,151,1185]" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF6568A6533E96CE" bold="true" box="[199,546,690,717]" pageId="12" pageNumber="463">
|
||||
Coloration of
|
||||
<typeStatus id="F3238879FFACFFCEFEC968A750CE96CF" box="[363,466,691,717]" pageId="12" pageNumber="463" type="holotype">holotype</typeStatus>
|
||||
in life.
|
||||
</emphasis>
|
||||
Dorsal surface of body yellowish brown, with incomplete dark brown triangular marking between eyes. Two wide oblique dark bands present on forearm. Dorsal surface of digits and hindlimbs with dark brown transverse bands. Tubercles on edge of upper eyelids orange. Supratympanic fold light brown. Spines on surface of sacral region and dorsal hindlimbs gray brown. Ventral surface dark brown, with black longitudinal band on surface of throat; surface of chest mottled with orange dots and patches; surface of ventral limbs and abdomen with grayish white mottling. Digits gray white; subarticular tubercles, inner and outer metacarpal tubercles and inner metatarsal tubercle orange. Pectoral glands and femoral glands gray white. Iris yellowish brown.
|
||||
</paragraph>
|
||||
<paragraph id="2C2736DBFFACFFCEFF6569BA53179037" blockId="12.[151,1437,151,1185]" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF6569BA53B497CA" bold="true" box="[199,680,942,969]" pageId="12" pageNumber="463">
|
||||
Coloration of
|
||||
<typeStatus id="F3238879FFACFFCEFEDA69BB50C397CB" box="[376,479,943,969]" pageId="12" pageNumber="463" type="holotype">holotype</typeStatus>
|
||||
in preservative.
|
||||
</emphasis>
|
||||
Yellowish brown fades to grayish brown dorsally. Color of the triangular marking between eyes, oblique bands on forearms, patterns on ventral surface more distinct. Orange patches and dots on surface of chest absent, white mottling on ventral limbs and abdomen faded; color of pectoral glands and femoral glands faded.
|
||||
</paragraph>
|
||||
<paragraph id="2C2736DBFFACFFCEFF656E2A55BE90A3" blockId="12.[151,1437,151,1185]" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF656E2A5023905A" bold="true" box="[199,319,1086,1112]" pageId="12" pageNumber="463">Variation.</emphasis>
|
||||
Morphometric variation is listed in
|
||||
<tableCitation id="611A0360FFACFFCEFD406E2B5225905B" box="[738,825,1087,1113]" captionStart="TABLE 3" captionStartId="12.[152,244,1232,1256]" captionTargetPageId="12" captionText="TABLE 3. Measurements (in mm) of the type series of Boulenophrys pepe sp. nov., * indicates the holotype." pageId="12" pageNumber="463">Table 3</tableCitation>
|
||||
. Most of the
|
||||
<typeStatus id="F3238879FFACFFCEFC7A6E2B555F905B" box="[984,1091,1087,1113]" pageId="12" pageNumber="463" type="paratype">paratypes</typeStatus>
|
||||
are similar to the
|
||||
<typeStatus id="F3238879FFACFFCEFAB96E2B5461905B" box="[1307,1405,1087,1113]" pageId="12" pageNumber="463" type="holotype">holotype</typeStatus>
|
||||
in morphology and color pattern, except for the following: dorsal surface of body dark brown in the
|
||||
<typeStatus id="F3238879FFACFFCEFB5D6E775443907F" box="[1279,1375,1123,1149]" pageId="12" pageNumber="463" type="paratype">paratype</typeStatus>
|
||||
GEP a209; orange patches and dots denser in the
|
||||
<typeStatus id="F3238879FFACFFCEFD246E9353FA90A3" box="[646,742,1159,1185]" pageId="12" pageNumber="463" type="paratype">paratype</typeStatus>
|
||||
SYS a009350 and absent in GEP a209.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="78E76653FFACFFCEFF3A6EC4543790EA" box="[152,1323,1232,1257]" pageId="12" pageNumber="463" startId="12.[152,244,1232,1256]" targetBox="[239,1357,1272,1820]" targetIsTable="true" targetPageId="12" targetType="table">
|
||||
<paragraph id="2C2736DBFFACFFCEFF3A6EC4543790EA" blockId="12.[152,1323,1232,1257]" box="[152,1323,1232,1257]" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF3A6EC4501390EA" bold="true" box="[152,271,1232,1256]" pageId="12" pageNumber="463">TABLE 3.</emphasis>
|
||||
Measurements (in mm) of the type series of
|
||||
<taxonomicName id="EB984D58FFACFFCEFD4C6EC452A790EA" authorityName="Wang & Lin & Gan & Chen & Yu & Pan & Xiao & Zeng" authorityYear="2024" box="[750,955,1232,1256]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="463" phylum="Chordata" rank="species" species="pepe" status="sp. nov.">
|
||||
<emphasis id="1EECEAC9FFACFFCEFD4C6EC452A790EA" box="[750,955,1232,1256]" italics="true" pageId="12" pageNumber="463">Boulenophrys pepe</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="05DF57B2FFACFFCEFC606EC5550490EA" box="[962,1048,1233,1257]" pageId="12" pageNumber="463" rank="species">
|
||||
<emphasis id="1EECEAC9FFACFFCEFC606EC5550D90EB" bold="true" box="[962,1041,1233,1257]" pageId="12" pageNumber="463">sp. nov</emphasis>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
, * indicates the holotype.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="2C2736DBFFACFFCEFE6E6EED5432931E" pageId="12" pageNumber="463">
|
||||
<table id="5E98C47BFFAC003DFF4D6EEC5451931E" box="[239,1357,1272,1820]" gridcols="5" gridrows="14" pageId="12" pageNumber="463">
|
||||
<tr id="92A83499FFAC003DFF4D6EEC54519113" box="[239,1357,1272,1297]" gridrow="0" isHeader="true" pageId="12" pageNumber="463" rowspan-0="1">
|
||||
<th id="D1795DE5FFAC003DFE6E6EEC535C9113" box="[460,576,1272,1297]" gridcol="1" gridrow="0" pageId="12" pageNumber="463">GEP a207*</th>
|
||||
<th id="D1795DE5FFAC003DFD616EEC52539113" box="[707,847,1272,1297]" gridcol="2" gridrow="0" pageId="12" pageNumber="463">SYS a009350</th>
|
||||
<th id="D1795DE5FFAC003DFC7E6EEC555F9113" box="[988,1091,1272,1297]" gridcol="3" gridrow="0" pageId="12" pageNumber="463">GEP a209</th>
|
||||
<th id="D1795DE5FFAC003DFB476EEC54519113" box="[1253,1357,1272,1297]" gridcol="4" gridrow="0" pageId="12" pageNumber="463">GEP a117</th>
|
||||
</tr>
|
||||
<tr id="92A83499FFAC003DFF4D6F345451913A" box="[239,1357,1312,1336]" gridrow="1" pageId="12" pageNumber="463">
|
||||
<th id="D1795DE5FFAC003DFF4D6F345032913A" box="[239,302,1312,1336]" gridcol="0" gridrow="1" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF596F34503D913A" bold="true" box="[251,289,1312,1336]" pageId="12" pageNumber="463">Sex</emphasis>
|
||||
</th>
|
||||
<td id="D1795DE5FFAC003DFE6E6F34535C913A" box="[460,576,1312,1336]" gridcol="1" gridrow="1" pageId="12" pageNumber="463">♂</td>
|
||||
<td id="D1795DE5FFAC003DFD616F345253913A" box="[707,847,1312,1336]" gridcol="2" gridrow="1" pageId="12" pageNumber="463">♂</td>
|
||||
<td id="D1795DE5FFAC003DFC7E6F34555F913A" box="[988,1091,1312,1336]" gridcol="3" gridrow="1" pageId="12" pageNumber="463">♂</td>
|
||||
<td id="D1795DE5FFAC003DFB476F345451913A" box="[1253,1357,1312,1336]" gridcol="4" gridrow="1" pageId="12" pageNumber="463">♂</td>
|
||||
</tr>
|
||||
<tr id="92A83499FFAC003DFF4D6F5C54519163" box="[239,1357,1352,1377]" gridrow="2" pageId="12" pageNumber="463">
|
||||
<th id="D1795DE5FFAC003DFF4D6F5C50329163" box="[239,302,1352,1377]" gridcol="0" gridrow="2" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF546F5C503B9162" bold="true" box="[246,295,1352,1376]" pageId="12" pageNumber="463">SVL</emphasis>
|
||||
</th>
|
||||
<td id="D1795DE5FFAC003DFE6E6F5C535C9163" box="[460,576,1352,1377]" gridcol="1" gridrow="2" pageId="12" pageNumber="463">36.4</td>
|
||||
<td id="D1795DE5FFAC003DFD616F5C52539163" box="[707,847,1352,1377]" gridcol="2" gridrow="2" pageId="12" pageNumber="463">35.7</td>
|
||||
<td id="D1795DE5FFAC003DFC7E6F5C555F9163" box="[988,1091,1352,1377]" gridcol="3" gridrow="2" pageId="12" pageNumber="463">35.3</td>
|
||||
<td id="D1795DE5FFAC003DFB476F5C54519163" box="[1253,1357,1352,1377]" gridcol="4" gridrow="2" pageId="12" pageNumber="463">36.0</td>
|
||||
</tr>
|
||||
<tr id="92A83499FFAC003DFF4D6F6554519188" box="[239,1357,1393,1418]" gridrow="3" pageId="12" pageNumber="463">
|
||||
<th id="D1795DE5FFAC003DFF4D6F6550329188" box="[239,302,1393,1418]" gridcol="0" gridrow="3" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF516F655036918B" bold="true" box="[243,298,1393,1417]" pageId="12" pageNumber="463">HDL</emphasis>
|
||||
</th>
|
||||
<td id="D1795DE5FFAC003DFE6E6F65535C9188" box="[460,576,1393,1418]" gridcol="1" gridrow="3" pageId="12" pageNumber="463">12.6</td>
|
||||
<td id="D1795DE5FFAC003DFD616F6552539188" box="[707,847,1393,1418]" gridcol="2" gridrow="3" pageId="12" pageNumber="463">12.5</td>
|
||||
<td id="D1795DE5FFAC003DFC7E6F65555F9188" box="[988,1091,1393,1418]" gridcol="3" gridrow="3" pageId="12" pageNumber="463">12.7</td>
|
||||
<td id="D1795DE5FFAC003DFB476F6554519188" box="[1253,1357,1393,1418]" gridcol="4" gridrow="3" pageId="12" pageNumber="463">12.5</td>
|
||||
</tr>
|
||||
<tr id="92A83499FFAC003DFF4D6F8D545191B0" box="[239,1357,1433,1458]" gridrow="4" pageId="12" pageNumber="463">
|
||||
<th id="D1795DE5FFAC003DFF4D6F8D503291B0" box="[239,302,1433,1458]" gridcol="0" gridrow="4" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF4D6F8D503291B3" bold="true" box="[239,302,1433,1457]" pageId="12" pageNumber="463">HDW</emphasis>
|
||||
</th>
|
||||
<td id="D1795DE5FFAC003DFE6E6F8D535C91B0" box="[460,576,1433,1458]" gridcol="1" gridrow="4" pageId="12" pageNumber="463">14.1</td>
|
||||
<td id="D1795DE5FFAC003DFD616F8D525391B0" box="[707,847,1433,1458]" gridcol="2" gridrow="4" pageId="12" pageNumber="463">14.4</td>
|
||||
<td id="D1795DE5FFAC003DFC7E6F8D555F91B0" box="[988,1091,1433,1458]" gridcol="3" gridrow="4" pageId="12" pageNumber="463">14.2</td>
|
||||
<td id="D1795DE5FFAC003DFB476F8D545191B0" box="[1253,1357,1433,1458]" gridcol="4" gridrow="4" pageId="12" pageNumber="463">13.7</td>
|
||||
</tr>
|
||||
<tr id="92A83499FFAC003DFF4D6FD5545191D8" box="[239,1357,1473,1498]" gridrow="5" pageId="12" pageNumber="463">
|
||||
<th id="D1795DE5FFAC003DFF4D6FD5503291D8" box="[239,302,1473,1498]" gridcol="0" gridrow="5" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF546FD5503B91DB" bold="true" box="[246,295,1473,1497]" pageId="12" pageNumber="463">SNT</emphasis>
|
||||
</th>
|
||||
<td id="D1795DE5FFAC003DFE6E6FD5535C91D8" box="[460,576,1473,1498]" gridcol="1" gridrow="5" pageId="12" pageNumber="463">4.5</td>
|
||||
<td id="D1795DE5FFAC003DFD616FD5525391D8" box="[707,847,1473,1498]" gridcol="2" gridrow="5" pageId="12" pageNumber="463">4.1</td>
|
||||
<td id="D1795DE5FFAC003DFC7E6FD5555F91D8" box="[988,1091,1473,1498]" gridcol="3" gridrow="5" pageId="12" pageNumber="463">4.5</td>
|
||||
<td id="D1795DE5FFAC003DFB476FD5545191D8" box="[1253,1357,1473,1498]" gridcol="4" gridrow="5" pageId="12" pageNumber="463">4.4</td>
|
||||
</tr>
|
||||
<tr id="92A83499FFAC003DFF4D6FFE54519200" box="[239,1357,1514,1538]" gridrow="6" pageId="12" pageNumber="463">
|
||||
<th id="D1795DE5FFAC003DFF4D6FFE50329200" box="[239,302,1514,1538]" gridcol="0" gridrow="6" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF556FFE50399200" bold="true" box="[247,293,1514,1538]" pageId="12" pageNumber="463">IND</emphasis>
|
||||
</th>
|
||||
<td id="D1795DE5FFAC003DFE6E6FFE535C9200" box="[460,576,1514,1538]" gridcol="1" gridrow="6" pageId="12" pageNumber="463">4.3</td>
|
||||
<td id="D1795DE5FFAC003DFD616FFE52539200" box="[707,847,1514,1538]" gridcol="2" gridrow="6" pageId="12" pageNumber="463">4.9</td>
|
||||
<td id="D1795DE5FFAC003DFC7E6FFE555F9200" box="[988,1091,1514,1538]" gridcol="3" gridrow="6" pageId="12" pageNumber="463">4.6</td>
|
||||
<td id="D1795DE5FFAC003DFB476FFE54519200" box="[1253,1357,1514,1538]" gridcol="4" gridrow="6" pageId="12" pageNumber="463">4.3</td>
|
||||
</tr>
|
||||
<tr id="92A83499FFAC003DFF4D6C0654519228" box="[239,1357,1554,1578]" gridrow="7" pageId="12" pageNumber="463">
|
||||
<th id="D1795DE5FFAC003DFF4D6C0650329228" box="[239,302,1554,1578]" gridcol="0" gridrow="7" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF546C06503A9228" bold="true" box="[246,294,1554,1578]" pageId="12" pageNumber="463">IOD</emphasis>
|
||||
</th>
|
||||
<td id="D1795DE5FFAC003DFE6E6C06535C9228" box="[460,576,1554,1578]" gridcol="1" gridrow="7" pageId="12" pageNumber="463">3.9</td>
|
||||
<td id="D1795DE5FFAC003DFD616C0652539228" box="[707,847,1554,1578]" gridcol="2" gridrow="7" pageId="12" pageNumber="463">4.1</td>
|
||||
<td id="D1795DE5FFAC003DFC7E6C06555F9228" box="[988,1091,1554,1578]" gridcol="3" gridrow="7" pageId="12" pageNumber="463">4.1</td>
|
||||
<td id="D1795DE5FFAC003DFB476C0654519228" box="[1253,1357,1554,1578]" gridcol="4" gridrow="7" pageId="12" pageNumber="463">4.0</td>
|
||||
</tr>
|
||||
<tr id="92A83499FFAC003DFF4D6C2E54519251" box="[239,1357,1594,1619]" gridrow="8" pageId="12" pageNumber="463">
|
||||
<th id="D1795DE5FFAC003DFF4D6C2E50329251" box="[239,302,1594,1619]" gridcol="0" gridrow="8" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF5F6C2E503C9250" bold="true" box="[253,288,1594,1618]" pageId="12" pageNumber="463">ED</emphasis>
|
||||
</th>
|
||||
<td id="D1795DE5FFAC003DFE6E6C2E535C9251" box="[460,576,1594,1619]" gridcol="1" gridrow="8" pageId="12" pageNumber="463">3.9</td>
|
||||
<td id="D1795DE5FFAC003DFD616C2E52539251" box="[707,847,1594,1619]" gridcol="2" gridrow="8" pageId="12" pageNumber="463">4.0</td>
|
||||
<td id="D1795DE5FFAC003DFC7E6C2E555F9251" box="[988,1091,1594,1619]" gridcol="3" gridrow="8" pageId="12" pageNumber="463">4.1</td>
|
||||
<td id="D1795DE5FFAC003DFB476C2E54519251" box="[1253,1357,1594,1619]" gridcol="4" gridrow="8" pageId="12" pageNumber="463">4.1</td>
|
||||
</tr>
|
||||
<tr id="92A83499FFAC003DFF4D6C7754519279" box="[239,1357,1635,1659]" gridrow="9" pageId="12" pageNumber="463">
|
||||
<th id="D1795DE5FFAC003DFF4D6C7750329279" box="[239,302,1635,1659]" gridcol="0" gridrow="9" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF5F6C77503C9279" bold="true" box="[253,288,1635,1659]" pageId="12" pageNumber="463">TD</emphasis>
|
||||
</th>
|
||||
<td id="D1795DE5FFAC003DFE6E6C77535C9279" box="[460,576,1635,1659]" gridcol="1" gridrow="9" pageId="12" pageNumber="463">2.1</td>
|
||||
<td id="D1795DE5FFAC003DFD616C7752539279" box="[707,847,1635,1659]" gridcol="2" gridrow="9" pageId="12" pageNumber="463">2.5</td>
|
||||
<td id="D1795DE5FFAC003DFC7E6C77555F9279" box="[988,1091,1635,1659]" gridcol="3" gridrow="9" pageId="12" pageNumber="463">2.5</td>
|
||||
<td id="D1795DE5FFAC003DFB476C7754519279" box="[1253,1357,1635,1659]" gridcol="4" gridrow="9" pageId="12" pageNumber="463">2.4</td>
|
||||
</tr>
|
||||
<tr id="92A83499FFAC003DFF4D6C9F545192A1" box="[239,1357,1675,1699]" gridrow="10" pageId="12" pageNumber="463">
|
||||
<th id="D1795DE5FFAC003DFF4D6C9F503292A1" box="[239,302,1675,1699]" gridcol="0" gridrow="10" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF506C9F503692A1" bold="true" box="[242,298,1675,1699]" pageId="12" pageNumber="463">HND</emphasis>
|
||||
</th>
|
||||
<td id="D1795DE5FFAC003DFE6E6C9F535C92A1" box="[460,576,1675,1699]" gridcol="1" gridrow="10" pageId="12" pageNumber="463">8.9</td>
|
||||
<td id="D1795DE5FFAC003DFD616C9F525392A1" box="[707,847,1675,1699]" gridcol="2" gridrow="10" pageId="12" pageNumber="463">9.0</td>
|
||||
<td id="D1795DE5FFAC003DFC7E6C9F555F92A1" box="[988,1091,1675,1699]" gridcol="3" gridrow="10" pageId="12" pageNumber="463">8.6</td>
|
||||
<td id="D1795DE5FFAC003DFB476C9F545192A1" box="[1253,1357,1675,1699]" gridcol="4" gridrow="10" pageId="12" pageNumber="463">8.4</td>
|
||||
</tr>
|
||||
<tr id="92A83499FFAC003DFF4D6CA7545192C9" box="[239,1357,1715,1739]" gridrow="11" pageId="12" pageNumber="463">
|
||||
<th id="D1795DE5FFAC003DFF4D6CA7503292C9" box="[239,302,1715,1739]" gridcol="0" gridrow="11" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF516CA7503692C9" bold="true" box="[243,298,1715,1739]" pageId="12" pageNumber="463">RAD</emphasis>
|
||||
</th>
|
||||
<td id="D1795DE5FFAC003DFE6E6CA7535C92C9" box="[460,576,1715,1739]" gridcol="1" gridrow="11" pageId="12" pageNumber="463">8.2</td>
|
||||
<td id="D1795DE5FFAC003DFD616CA7525392C9" box="[707,847,1715,1739]" gridcol="2" gridrow="11" pageId="12" pageNumber="463">7.5</td>
|
||||
<td id="D1795DE5FFAC003DFC7E6CA7555F92C9" box="[988,1091,1715,1739]" gridcol="3" gridrow="11" pageId="12" pageNumber="463">8.1</td>
|
||||
<td id="D1795DE5FFAC003DFB476CA7545192C9" box="[1253,1357,1715,1739]" gridcol="4" gridrow="11" pageId="12" pageNumber="463">7.9</td>
|
||||
</tr>
|
||||
<tr id="92A83499FFAC003DFF4D6CCF545192F6" box="[239,1357,1755,1780]" gridrow="12" pageId="12" pageNumber="463">
|
||||
<th id="D1795DE5FFAC003DFF4D6CCF503292F6" box="[239,302,1755,1780]" gridcol="0" gridrow="12" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF5A6CCF503892F1" bold="true" box="[248,292,1755,1779]" pageId="12" pageNumber="463">TIB</emphasis>
|
||||
</th>
|
||||
<td id="D1795DE5FFAC003DFE6E6CCF535C92F6" box="[460,576,1755,1780]" gridcol="1" gridrow="12" pageId="12" pageNumber="463">14.5</td>
|
||||
<td id="D1795DE5FFAC003DFD616CCF525392F6" box="[707,847,1755,1780]" gridcol="2" gridrow="12" pageId="12" pageNumber="463">14.6</td>
|
||||
<td id="D1795DE5FFAC003DFC7E6CCF555F92F6" box="[988,1091,1755,1780]" gridcol="3" gridrow="12" pageId="12" pageNumber="463">14.6</td>
|
||||
<td id="D1795DE5FFAC003DFB476CCF545192F6" box="[1253,1357,1755,1780]" gridcol="4" gridrow="12" pageId="12" pageNumber="463">14.7</td>
|
||||
</tr>
|
||||
<tr id="92A83499FFAC003DFF4D6D105451931E" box="[239,1357,1796,1820]" gridrow="13" pageId="12" pageNumber="463">
|
||||
<th id="D1795DE5FFAC003DFF4D6D105032931E" box="[239,302,1796,1820]" gridcol="0" gridrow="13" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF546D10503B931E" bold="true" box="[246,295,1796,1820]" pageId="12" pageNumber="463">FTL</emphasis>
|
||||
</th>
|
||||
<td id="D1795DE5FFAC003DFE6E6D10535C931E" box="[460,576,1796,1820]" gridcol="1" gridrow="13" pageId="12" pageNumber="463">20.2</td>
|
||||
<td id="D1795DE5FFAC003DFD616D105253931E" box="[707,847,1796,1820]" gridcol="2" gridrow="13" pageId="12" pageNumber="463">22.0</td>
|
||||
<td id="D1795DE5FFAC003DFC7E6D10555F931E" box="[988,1091,1796,1820]" gridcol="3" gridrow="13" pageId="12" pageNumber="463">21.0</td>
|
||||
<td id="D1795DE5FFAC003DFB476D105451931E" box="[1253,1357,1796,1820]" gridcol="4" gridrow="13" pageId="12" pageNumber="463">20.5</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<subSubSection id="64826550FFACFFCEFF656D7B542193D3" pageId="12" pageNumber="463" type="distribution">
|
||||
<paragraph id="2C2736DBFFACFFCEFF656D7B542193D3" blockId="12.[151,1437,1903,2002]" pageId="12" pageNumber="463">
|
||||
<emphasis id="1EECEAC9FFACFFCEFF656D7B5351938B" bold="true" box="[199,589,1903,1929]" pageId="12" pageNumber="463">Distribution and natural history.</emphasis>
|
||||
Currently,
|
||||
<taxonomicName id="EB984D58FFACFFCEFD696D7B52BE938B" authorityName="Wang & Lin & Gan & Chen & Yu & Pan & Xiao & Zeng" authorityYear="2024" box="[715,930,1903,1929]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="463" phylum="Chordata" rank="species" species="pepe" status="sp. nov.">
|
||||
<emphasis id="1EECEAC9FFACFFCEFD696D7B52BE938B" box="[715,930,1903,1929]" italics="true" pageId="12" pageNumber="463">Boulenophrys pepe</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="1EECEAC9FFACFFCEFC0B6D7B551E938B" bold="true" box="[937,1026,1903,1929]" pageId="12" pageNumber="463">
|
||||
<taxonomicNameLabel id="05DF57B2FFACFFCEFC0B6D7B551E938B" box="[937,1026,1903,1929]" pageId="12" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is only known from its
|
||||
<typeStatus id="F3238879FFACFFCEFAAE6D6454209388" box="[1292,1340,1904,1930]" pageId="12" pageNumber="463">type</typeStatus>
|
||||
locality. It inhabits flowing montane seeps and the nearby forest floor and leaf litter at elevations between
|
||||
<quantity id="EB609B3EFFACFFCEFB656D87545893AC" box="[1223,1348,1939,1966]" metricMagnitude="2" metricUnit="m" metricValue="5.15" metricValueMax="5.4" metricValueMin="4.9" pageId="12" pageNumber="463" unit="m" value="515.0" valueMax="540.0" valueMin="490.0">490–540 m</quantity>
|
||||
(
|
||||
<figureCitation id="B4A32A5EFFACFFCEFAF36D8751BB93D0" captionStart="FIGURE 5" captionStartId="13.[152,255,1061,1085]" captionTargetBox="[151,1436,181,1038]" captionTargetId="figure-19@13.[151,1436,181,1038]" captionTargetPageId="13" captionText="FIGURE 5. Habitat (A) and microhabitat (B) of Boulenophrys pepe sp. nov. at the type locality." figureDoi="http://doi.org/10.5281/zenodo.13914797" httpUri="https://zenodo.org/record/13914797/files/figure.png" pageId="12" pageNumber="463">Figure 5</figureCitation>
|
||||
). On January, males were found calling in rock crevices or under leaf litters and tadpoles were observed.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
Reference in a new issue