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<document id="9C48A3BE890F0FBF28AEFD5417A4A842" ENCODING="UTF-8" ID-GBIF-Dataset="77d07a5c-cc8e-45f7-84bd-5587ecf0d3d5" ModsDocID="8111" checkinTime="1243377608815" checkinUser="christiana" docAuthor="Santschi, F." docDate="1914" docId="4CE6DB43841A62E71CA56F3463B8956A" docLanguage="fr" docName="8111" docOrigin="Voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale (1911 - 1912). Résultats scientifiques. Hyménoptères 2" docSource="http://antbase.org/ants/publications/8111/8111.pdf" docTitle="Dorylus (Alaopone) buyssoni" docType="treatment" docVersion="11" lastPageNumber="65" masterDocId="FE06B16C2BBDFCD084D4F6343B802CB0" masterDocTitle="Formicidae." masterLastPageNumber="148" masterPageNumber="41" pageNumber="65" updateTime="1701312626052" updateUser="plazi">
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<mods:title id="97E73CDDAAB6513529EF0A4A764316AC">Formicidae.</mods:title>
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<mods:namePart id="256E06B655586B20659FFCE1DB307088">Santschi, F.</mods:namePart>
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<mods:title id="4F23866629923B55D13FBE155B64E117">Voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale (1911 - 1912). Résultats scientifiques. Hyménoptères</mods:title>
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<mods:date id="AB4B07CEBD50213821272E29863A849B">1914</mods:date>
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<mods:number id="7D7EB503E6574C89493D4F1770654F4B">2</mods:number>
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<mods:start id="5D0277A009D2B2598F2B789980507F85">41</mods:start>
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<treatment id="4CE6DB43841A62E71CA56F3463B8956A" ID-DOI="http://doi.org/10.5281/zenodo.5063540" ID-GBIF-Taxon="100126795" ID-Zenodo-Dep="5063540" LSID="urn:lsid:plazi:treatment:4CE6DB43841A62E71CA56F3463B8956A" httpUri="http://treatment.plazi.org/id/4CE6DB43841A62E71CA56F3463B8956A" lastPageNumber="65" pageNumber="65">
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<taxonomicName id="B71228E165DDD5CB0F1688C8D32010C2" ID-CoL="37FBV" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:137837" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" lsidName="Dorylus (Alaopone) buyssoni Santschi" lsidName-HNS="Dorylus (Alaopone) buyssoni Santschi" order="Hymenoptera" pageNumber="65" phylum="Arthropoda" rank="species" species="buyssoni" subGenus="Alaopone">Dorylus (Alaopone) Buyssoni</taxonomicName>
Santschi.
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<paragraph id="47370FA2CA516AE677B8A0525CCDDE6B" pageNumber="65">Rev. Suisse Zool., vol. 18, p. 748, fig. 7 (1910), [[male]].</paragraph>
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<subSubSection id="29A7A42A8AE48A4F0C8CCA4DA521C624" type="materials_examined">
<paragraph id="C3FB834E6B03845A4B89EEF1A8EB194F" pageNumber="65">Afrique orientale anglaise: Nairobi (1904), 4 [[male]] [types]; - Nairobi (1909), 3 [[male]]; - region cotiere: Cheteni (st. n° 4, nov. 1911), 1 [[male]].</paragraph>
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<paragraph pageId="76" pageNumber="77">
† Tribe
<taxonomicName LSID="4CE6FD9D-F500-5F11-A9E3-45E745B1D0BB" authority="Kolibáč &amp; Huang, 2008" authorityName="Handlirsch" authorityYear="1906" class="Insecta" family="Trogossitidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="tribe" tribe="Lithostomatini">
Lithostomatini
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
&amp; Huang, 2008
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<paragraph pageId="76" pageNumber="77">
<taxonomicName authorityName="Kolibac &amp; Huang" authorityYear="2008" lsidName="" pageId="76" pageNumber="77" rank="tribe" tribe="Lithostomini">Lithostomini</taxonomicName>
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
, J. &amp; Huang, D.-Y. 2008: 142 (as). Bouchard, P. et al. 2011: 57 (emendation to
<taxonomicName authorityName="Kolibac &amp; Huang" authorityYear="2008" lsidName="" pageId="76" pageNumber="77" rank="tribe" tribe="Lithostomatini">Lithostomatini</taxonomicName>
)
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<paragraph pageId="76" pageNumber="77">Type genus.</paragraph>
<paragraph pageId="76" pageNumber="77">
<taxonomicName authorityName="Martynov" authorityYear="1926" class="Insecta" family="Trogossitidae" genus="Lithostoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="76" pageNumber="77">Lithostoma</emphasis>
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Martynov, 1926 [by monotypy and
<normalizedToken originalValue="authors">author's</normalizedToken>
designation]
</paragraph>
<paragraph pageId="76" pageNumber="77">
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
, J. &amp; Huang, D.-Y. 2008: 142. Yu, Y. et al. 2012: 250
</paragraph>
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<subSubSection lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77" type="remarks">
<paragraph pageId="76" pageNumber="77">Remarks.</paragraph>
<paragraph pageId="76" pageNumber="77">
This fossil differs from all other Mesozoic
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Hexapoda" family="Trogossitidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="family">Trogossitidae</taxonomicName>
described to date. If
<bibRefCitation author="Martynov, AV" journalOrPublisher="Ann. Soc. Paleont. Russie" pageId="157" pageNumber="158" pagination="1 - 39" refId="B200" refString="Martynov, AV, 1926. KPoznaniyu Iskopaemykh Nasekomykh Yurskikh Slantsev Turkestana. 5. O Nekotorykh Formakh Zhukov (Coleoptera). Ann. Soc. Paleont. Russie 5: 1 - 39" title="KPoznaniyu Iskopaemykh Nasekomykh Yurskikh Slantsev Turkestana. 5. O Nekotorykh Formakh Zhukov (Coleoptera)." volume="5" year="1926">Martynov (1926)</bibRefCitation>
interpreted the shapes of the head and antennal segments well, it is the first known member of
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Hexapoda" family="Trogossitidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="family">Trogossitidae</taxonomicName>
without a distinct antennal club and with the head narrowed towards its base. The following features of
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Hexapoda" family="Trogossitidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="family">Trogossitidae</taxonomicName>
appear in the fossil: (1) general shape and size of body, (2) distinctly flattened sides of pronotum and elytra, (3) double rows of punctures/tubercles among elytral carinae, (4) robust bidentate mandibles, (5) extremely large scapus, and (6) dilated antennal segments with what are perhaps sensorial fields in the enlarged parts of each segment. The classification within
<taxonomicName authorityName="Latreille" authorityYear="1802" lsidName="" pageId="76" pageNumber="77" rank="subFamily" subFamily="Trogossitinae">Trogossitinae</taxonomicName>
is based on the presence of the sensorial fields in the enlarged parts of the antennomeres alone. Small tubercles occurring in pronotum and elytra are known in trogossitine genera
<taxonomicName authorityName="Thomson" authorityYear="1859" class="Hexapoda" family="Trogossitidae" genus="Calitys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="76" pageNumber="77">Calitys</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Reitter" authorityYear="1876" class="Hexapoda" family="Trogossitidae" genus="Phanodesta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="76" pageNumber="77">Phanodesta</emphasis>
</taxonomicName>
only; no peltine representative possesses such structures. The tribe differs from the recent and fossil members of
<taxonomicName authorityName="Latreille" authorityYear="1802" lsidName="" pageId="76" pageNumber="77" rank="subFamily" subFamily="Trogossitinae">Trogossitinae</taxonomicName>
in broadly oval body (this occurs in some
<taxonomicName authorityName="Lacordaire" authorityYear="1854" lsidName="" pageId="76" pageNumber="77" rank="tribe" tribe="Gymnochilini">Gymnochilini</taxonomicName>
only), pronotum narrowed anteriad, antennae without conspicuous club and asymmetrical segments in flagellum, head narrowed towards base. The shape of antennal segments 10 and 11 is unknown because they are missing (only a trace of segment 10 is visible). The antennae may be only 10-segmented with the last segment enlarged (as in e.g.
<taxonomicName authorityName="Lacordaire" authorityYear="1854" lsidName="" pageId="76" pageNumber="77" rank="tribe" tribe="Egoliini">Egoliini</taxonomicName>
). The new tribe is probably isolated from other tribes of
<taxonomicName authorityName="Latreille" authorityYear="1802" lsidName="" pageId="76" pageNumber="77" rank="subFamily" subFamily="Trogossitinae">Trogossitinae</taxonomicName>
and may be considered a sister group to them. The ventral part of the fossil is unfortunately unknown, so a classification of
<taxonomicName authorityName="Martynov" authorityYear="1926" class="Insecta" family="Trogossitidae" genus="Lithostoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="76" pageNumber="77">Lithostoma</emphasis>
</taxonomicName>
remains uncertain, chiefly based on the distinctly enlarged scapus.
</paragraph>
<paragraph pageId="77" pageNumber="78">
<pageBreakToken pageId="77" pageNumber="78" start="start">On</pageBreakToken>
the other hand, the concept of
<taxonomicName authorityName="Kolibac &amp; Huang" authorityYear="2008" lsidName="" pageId="77" pageNumber="78" rank="tribe" tribe="Lithostomatini">Lithostomatini</taxonomicName>
was justifiably called into question by
<bibRefCitation author="Yu, Y" journalOrPublisher="Annales Zoologici (Warszawa)" pageId="164" pageNumber="165" pagination="245 - 252" publicationUrl="10.3161/000345412X652765" refId="B322" refString="Yu, Y, Leschen, RA B, Ślipinski, A, Ren, D, Pang, H, 2012. The first fossil bark-gnawing beetle from the Middle Jurassic of Inner Mongolia, China (Coleoptera: Trogossitidae). Annales Zoologici (Warszawa) 62: 245 - 252, 10.3161/000345412X652765" title="The first fossil bark-gnawing beetle from the Middle Jurassic of Inner Mongolia, China (Coleoptera: Trogossitidae)." url="10.3161/000345412X652765" volume="62" year="2012">Yu et al. (2012)</bibRefCitation>
who argued that insufficient morphological information existed for the establishment of a higher taxon.
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