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<document id="6572A5DD8BE7BFE7280F9D4BC598CD7E" ENCODING="Cp1252" ID-GBIF-Dataset="1407eb13-b429-4db6-9004-ef57b73c5ccd" ModsDocAuthor="van der Hammen, L. &amp; Strenzke, K." ModsDocDate="1953" ModsDocID="ORI11169" ModsDocOrigin="http://www.repository.naturalis.nl/document/150100" ModsDocTitle="A Partial Revision of the Genus Metabelba Grandjean (Oribatei, Acarina)" checkinTime="1326119974990" checkinUser="thomas" docAuthor="van der Hammen, L. &amp; Strenzke, K." docDate="1953" docId="4CBD140BA9E9EC7E00687B71230394C2" docLanguage="en" docName="vdH_Strenzke_1953_gg1.xml" docOrigin="Zoologische Mededelingen 32" docSource="http://www.repository.naturalis.nl/document/150100" docTitle="Metabelba pulverosa Hammen &amp; Strenzke, 1953, nov. spec." docType="treatment" docVersion="10" lastPageNumber="152" masterDocId="74A8F4D3D959A33D9588A4C422990791" masterDocTitle="A Partial Revision of the Genus Metabelba Grandjean (Oribatei, Acarina)" masterLastPageNumber="154" masterPageNumber="141" pageNumber="148" updateTime="1701313557530" updateUser="plazi">
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<mods:title id="70DA062C7CAAA4C3FA90EA231F6EDFF3">A Partial Revision of the Genus Metabelba Grandjean (Oribatei, Acarina)</mods:title>
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<mods:namePart id="D0043986D338BBCAB712CCF90B8784E3">van der Hammen, L.</mods:namePart>
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<mods:namePart id="5C0185070AC61F8F68D0CE7FEE717278">Strenzke, K.</mods:namePart>
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<mods:title id="61132DD0BA0E1A0FE8985CAF29942ACB">Zoologische Mededelingen</mods:title>
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<mods:part id="DB3919B2C3597892F05EE87430C7EB2F">
<mods:date id="4326EA5AB85D5647114B706C99111752">1953</mods:date>
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<mods:number id="58721BC10242911AEF47931510427CF1">32</mods:number>
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<treatment id="4CBD140BA9E9EC7E00687B71230394C2" ID-DOI="http://doi.org/10.5281/zenodo.6286085" ID-GBIF-Taxon="100112042" ID-Zenodo-Dep="6286085" LSID="urn:lsid:plazi:treatment:4CBD140BA9E9EC7E00687B71230394C2" httpUri="http://treatment.plazi.org/id/4CBD140BA9E9EC7E00687B71230394C2" lastPageId="11" lastPageNumber="152" pageId="7" pageNumber="148">
<subSubSection id="5C27DF28FC46AD91ED60E273C531C92D" pageId="7" pageNumber="148" type="nomenclature">
<paragraph id="E49BDFFF1D692FEDB64DEA9A9AC45CF7" pageId="7" pageNumber="148">
<taxonomicName id="8BA455B2606222451FDB7C04C006EAF3" class="Arachnida" family="Damaeidae" genus="Metabelba" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="pulverosa">Metabelba pulverosa Strenzke</taxonomicName>
<taxonomicNameLabel id="0CBA8633D40021A435F41AA44D074903" pageId="7" pageNumber="148" rank="species">nov. spec.</taxonomicNameLabel>
(figs. 5, 6)
</paragraph>
</subSubSection>
<subSubSection id="6D554A26AE38C810D22ACA120CBC6F55" lastPageId="10" lastPageNumber="151" pageId="7" pageNumber="148" type="description">
<paragraph id="CA467E283D00C83CD39B1FCE2FB9B072" pageId="7" pageNumber="148">
Length of the idiosoma 425-495
<normalizedToken id="F73ED0B02FDABDBD145D757A04A01FAF" originalValue="µ" pageId="7" pageNumber="148">y</normalizedToken>
; length of the propodosoma 190
<normalizedToken id="531C59F8BF428130C9651148EE305E57" originalValue="µ" pageId="7" pageNumber="148">y</normalizedToken>
, of the hysterosoma 305
<normalizedToken id="D2C6A4F03B9C74633A6EC042B24908E8" originalValue="µ" pageId="7" pageNumber="148">y</normalizedToken>
; breadth of the hysterosoma 305
<normalizedToken id="37206A176F9B73CDB7814A4A7DEF0752" originalValue="µ" pageId="7" pageNumber="148">y</normalizedToken>
. Colour glossy brown. Generally the specimens are strongly covered with cerotegument. Often the adults bear the larval and nymphal skins. Rostral hairs, lamellar hairs, interlamellar hairs, and exobothridial hairs slightly coarse; the remaining hairs and the sensillus are smooth.
</paragraph>
<paragraph id="3E995F8E12B4F687C9686F9E1FF52156" pageId="7" pageNumber="148">Propodosoma between the f irst and the second legs with a strong anterior apophysis that is directed to the front, and that is distally truncate or sometimes slightly excavated; the surface of the apophysis is granulate (a.a. in figs. 5, 6a). Anterior parastigmatic apophysis (a.p.a.) slender and conical, directed sideways and upward. A similar, but shorter apophysis in front of acetabulura III (posterior parastigmatic apophysis, a.p.p.) and in front of acetabulum IV (discidium, dis).</paragraph>
<paragraph id="D4C38A3C60189D9D275F005F593D2F1C" pageId="7" pageNumber="148">
Posterior border of the propodosoma without protuberances. In front of the bothridium, at each side, at first a round, light spot, then an area of smaller irregular spots. A similar, unpaired group occurs just between the bothridia. Medially and anally of the bothridia at each side a single light spot, posteriorly of which the chitin is slightly thickened, without, however, forming a protuberance as in related species. The disposition of the propodosomatal hairs is shown in figs. 5a, 6a. The sensillus (length about 120
<normalizedToken id="206DEDB62981FA6C6A3658B761786B13" originalValue="µ" pageId="7" pageNumber="148">y</normalizedToken>
) is flagelliform; it is directed obliquely backward. The interlamellar hairs (length about 80
<normalizedToken id="7D067D579DD2E756C7BCD7B752412D58" originalValue="µ" pageId="7" pageNumber="148">y</normalizedToken>
) are also directed obliquely backward; they stand close to the bothridia. Bothridium cup-shaped. Exobothridial hairs close to the antiaxial side of the bothridia; they are strongly curved, and are directed frontward and upward.
</paragraph>
<paragraph id="62E29DA6CC1639264D38CB19CB15B8FA" lastPageId="8" lastPageNumber="149" pageId="7" pageNumber="148">
Notogastral hairs in two rows, each of 8 curved hairs, which are inserted rather close to each other. The two anterior pairs (c1 and c2) are directed to the front, the remaining hairs 'backward (figs. 5a, 6a). Three pairs of hairs occur at the posterior border of the notogaster (ps1, ps2, ps3 in figs. 5b,
<pageBreakToken id="96895BA83D2A2A3868B9C0D6FD0887FC" pageId="8" pageNumber="149" start="start">6</pageBreakToken>
a). The hairs of the two longitudinal rows possess wing-like expansions that begin just above the base; these are especially striking in the hairs c1 and c2 (fig. 6b). In baisam these features become less distinct. Length of the hairs c1 and c2 80
<normalizedToken id="4FFF88582D48FB90E561C1504215DBDC" originalValue="µ" pageId="8" pageNumber="149">y</normalizedToken>
; the remaining notogastral hairs are about 60
<normalizedToken id="B4C94095C2F3CD483488DEEB214B91A8" originalValue="µ" pageId="8" pageNumber="149">y</normalizedToken>
long. The disposition of the fissures on the notogaster is shown in fig. 6a. There are no Spinae adnatae.
</paragraph>
<caption id="D9DCA55E4D223383A7656BA35B01A201" pageId="8" pageNumber="149">
<paragraph id="D90461FA21321A4F2DCBBF67B992DD36" pageId="8" pageNumber="149">
Fig. 5.
<taxonomicName id="406D82DA70CDC9B9AEBFCB927D3C977C" class="Arachnida" family="Damaeidae" genus="Metabelba" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="149" phylum="Arthropoda" rank="species" species="pulverosa">Metabelba pulverosa Strenzke</taxonomicName>
<taxonomicNameLabel id="3569ED21FE5590CB80C933E185383009" pageId="8" pageNumber="149" rank="species">nov. spec.</taxonomicNameLabel>
a, dorsal view; b, ventral view. X 240.
</paragraph>
</caption>
<paragraph id="84364A4FAB08BC90FEE1499BE1F04B4B" pageId="8" pageNumber="149">Genital and anal plates separated by a large interval. The chaetotaxy of the ventral surface is shown in fig. 5b.</paragraph>
<paragraph id="8C8BD3E5AE9556D31269E8530E12EA67" pageId="8" pageNumber="149">The shape and the chaetotaxy of the legs is shown in figs. 6c, d. The number of hairs is as follows:</paragraph>
<caption id="A6D509B8DA9D1BFA98A89D5ED76D31C3" ID-Table-UUID="A6D509B8DA9D1BFA98A89D5ED76D31C3" httpUri="http://table.plazi.org/id/A6D509B8DA9D1BFA98A89D5ED76D31C3" pageId="9" pageNumber="150">
<paragraph id="725E4F9AB7035588178C6E2F6D303812" pageId="9" pageNumber="150">
<pageBreakToken id="CB922037DD3DCD1E194C5D74BBF1E7BE" pageId="9" pageNumber="150" start="start">Fig</pageBreakToken>
. 6.
<taxonomicName id="44EDFEC6BC5C6C9B7D6DC51A3962B8AF" class="Arachnida" family="Damaeidae" genus="Metabelba" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="pulverosa">Metabelba pulverosa Strenzke</taxonomicName>
<taxonomicNameLabel id="F139343BD6D79DABE988C2491EC86954" pageId="9" pageNumber="150" rank="species">nov. spec.</taxonomicNameLabel>
a, lateral view; b, hair c2; c, left leg IV from the antiaxial side; d, left leg I from the antiaxial side. a, X 240; b, X 830; c, d, X 335.
</paragraph>
</caption>
<paragraph id="003CCEAB3DB5690FA47FE16BAF1DD42B" pageId="10" pageNumber="151">
<table id="113D7283C3251687FABEBA92DAF6E3A4" pageId="10" pageNumber="151">
<tr id="E0410191575BB0C922DAEE5D0F53E833" pageId="10" pageNumber="151">
<td id="331CB1CE4CB4E4AA462B5223DCA752A6" pageId="10" pageNumber="151">
<pageBreakToken id="5E8336280E350056CB2AC6CA51F749EE" pageId="10" pageNumber="151" start="start">EmTaCe</pageBreakToken>
</td>
<td id="9ECAA3174A07A28B108095D9A97B41BC" pageId="10" pageNumber="151">trochanter</td>
<td id="38AB20BDEA036783A1720C5B5CB398FA" pageId="10" pageNumber="151">femur</td>
<td id="F7C1004B6CB1263A7C8F960D0D12F1EC" pageId="10" pageNumber="151">genu</td>
<td id="4F7A59156B0CDBCDB84F5D91283FB90A" pageId="10" pageNumber="151">tibia</td>
<td id="8367D67ADA7CB9847C388196F28627DD" pageId="10" pageNumber="151">tarsus</td>
</tr>
<tr id="1A36E1122821795EDEF8031E3E761BDE" pageId="10" pageNumber="151">
<td id="666AA74D638D661E75C8D04146DC5763" pageId="10" pageNumber="151">I</td>
<td id="21BA7F03EF2D083CBF41F6BB57F9397C" pageId="10" pageNumber="151">1</td>
<td id="27445FEAA1D5CC93ECB3ECE22B19E858" pageId="10" pageNumber="151">10</td>
<td id="3E5A71EE78B4D2AC3DC5A8CBD2ADC6E5" pageId="10" pageNumber="151">5</td>
<td id="A688562939127388B7815D0261DA973B" pageId="10" pageNumber="151">6</td>
<td id="4D5576BA0F4EE59873A600BAEB6A7003" pageId="10" pageNumber="151">22</td>
</tr>
<tr id="8D3E4214ACA53FE45C5307B88D76BB43" pageId="10" pageNumber="151">
<td id="A09569D2EDBAE8212C82F3A2E864135D" pageId="10" pageNumber="151">IV</td>
<td id="F5BFEDBA372E0346A1E7BD632DCD8D0C" pageId="10" pageNumber="151">3</td>
<td id="FCC1DB34A1DB85C05B3569B49B8D5CCA" pageId="10" pageNumber="151">9</td>
<td id="74DA89FCB80A056A322D384134951F2D" pageId="10" pageNumber="151">4</td>
<td id="C2CC205D9C89334294474EEEB03A2B3F" pageId="10" pageNumber="151">5</td>
<td id="40E5747C543589B98D7B791DFB253F58" pageId="10" pageNumber="151">14</td>
</tr>
</table>
</paragraph>
<paragraph id="AD51CF5236C4CCF89F7BF8BC3E0207D1" pageId="10" pageNumber="151">
On femur I dorsally a strong, serrate hair that is curved to the front close above the base. On leg IV similar hairs on trochanter, femur, and genu. The remaining hairs of the legs partly with similar expansions as the notogastral hairs. Solenidion of tibia IV free, and strongly curved in the middle. The length (in
<normalizedToken id="44C1F01C3F902D546B144E7DE094D8A7" originalValue="µ" pageId="10" pageNumber="151">y</normalizedToken>
) of the joints of the legs is. as follows:
</paragraph>
<paragraph id="CFF2DFED4F8B5EFF740E531ABA475683" pageId="10" pageNumber="151">
<table id="B91F25C17468430A314940538AFCF869" pageId="10" pageNumber="151">
<tr id="12F2AA6E457009035C5EB81221CC4327" pageId="10" pageNumber="151">
<td id="EF50FAF3DF5B4DF1F348EE12624D8CE7" pageId="10" pageNumber="151">EmTaCe</td>
<td id="AEFDDADBFDC864E5E95541350DF0AE84" pageId="10" pageNumber="151">trochanter</td>
<td id="D567C16DA6545692253F9BB70F86DE3F" pageId="10" pageNumber="151">femur</td>
<td id="9657CF1CEFE7B7F2FB2D70D8BC76B138" pageId="10" pageNumber="151">genu</td>
<td id="218AB6976AFF55BFB3840F60CB7A7E2D" pageId="10" pageNumber="151">tibia</td>
<td id="898E731A5A282248D5553811A6BB1D5D" pageId="10" pageNumber="151">tarsus</td>
</tr>
<tr id="0971338358B4B7D96BB90FC0A9C03C97" pageId="10" pageNumber="151">
<td id="1257590CFBC7498223CB0286F0F33244" pageId="10" pageNumber="151">I</td>
<td id="82DD214E6FCB69E820EDC4B48F833687" pageId="10" pageNumber="151">128</td>
<td id="5EBAA1EC56BEDC02709700300F5366E4" pageId="10" pageNumber="151">46</td>
<td id="389776B8B87E7EA777DB10B00211CE68" pageId="10" pageNumber="151">60</td>
<td id="CFC206C6773806CD61C30C4EB8BC3F68" pageId="10" pageNumber="151">122</td>
</tr>
<tr id="C3D4E098037FC42D6733C49AC688BD8D" pageId="10" pageNumber="151">
<td id="C2B7EAE01B9FA7256EB59ED18D01B33F" pageId="10" pageNumber="151">IV</td>
<td id="96C08B3B93D48246F0337B66EBCCBB09" pageId="10" pageNumber="151">103</td>
<td id="26E535EB8FFB9A09DD78966B4FACD359" pageId="10" pageNumber="151">82</td>
<td id="D5366BB8496B05E96752DC683E1A0355" pageId="10" pageNumber="151">50</td>
<td id="F846AA5B6D72DDF1CCADBF30141E38F1" pageId="10" pageNumber="151">72</td>
<td id="147D7093E7D374DE4953D2CC77E2E9A7" pageId="10" pageNumber="151">130</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="B96898D34869E8C1F1317D1ED1FDC9D4" pageId="10" pageNumber="151" type="materials_examined">
<paragraph id="BC87BFEBBD4FD811B47FC0B502EE7845" pageId="10" pageNumber="151">
<materialsCitation id="26EF65420B780D7E771A46E46493BD2A" ID-GBIF-Occurrence="923889712" collectingDate="1940-05-18" collectingMethod="Substratprobe" collectorName="K. Strenzke" country="Germany" location="forest meadow at the Keller See" municipality="Sielbeck" stateProvince="Schleswig-Holstein" typeStatus="Holotypus">
<typeStatus id="691BDE307901EE4548602AB9819647A4">Holotype</typeStatus>
: from the soil of a
<location id="8D5E296D49FF3AF868518A1E2416B428" LSID="urn:lsid:plazi:treatment:4CBD140BA9E9EC7E00687B71230394C2:8D5E296D49FF3AF868518A1E2416B428" country="Germany" latitude="54.1827" longLatPrecision="565" longitude="10.6100" municipality="Sielbeck" name="forest meadow at the Keller See" stateProvince="Schleswig-Holstein">forest meadow at the Keller See</location>
,
<collectingRegion id="327E26E062CBC78BF3515B20C5843181">Holstein</collectingRegion>
,
<collectingDate id="B7FC6B94A247813DF5616B806A1577CA" value="1940-05-18">
<date id="49804C349D93E8D5975159AA198BD53C" value="1940-05-18">18.V.1940</date>
</collectingDate>
(sample 239, cf.
<collectorName id="8D6F6EEB58387B8D5A4364B7BA84FFDB">Strenzke</collectorName>
, 1952, p. 48), as slide in Strenzke's collection.
</materialsCitation>
<materialsCitation id="52B1A6C245B5676174FA297A5670376D" ID-GBIF-Occurrence="923889711" collectingDate="1940-05-18" collectingMethod="Substratprobe" collectionCode="RNHL" collectorName="K. Strenzke" country="Germany" location="forest meadow at the Keller See" municipality="Sielbeck" stateProvince="Schleswig-Holstein" typeStatus="Paratypus">
<typeStatus id="A5E8378ED4A569689B91D1AAAC3E46F6">Paratypes</typeStatus>
<location id="BC5E1D63735C84AB9445C9BDE3A33E31" LSID="urn:lsid:plazi:treatment:4CBD140BA9E9EC7E00687B71230394C2:BC5E1D63735C84AB9445C9BDE3A33E31" country="Germany" latitude="54.1827" longLatPrecision="565" longitude="10.6100" municipality="Sielbeck" name="forest meadow at the Keller See" stateProvince="Schleswig-Holstein">from the same sample</location>
, in alcohol, in the collection of the Rijksmuseum van Natuurlijke Historie, Leiden, and in Strenzke's collection.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CB1D67E71DE19FA011CD046AC22EA009" lastPageId="11" lastPageNumber="152" pageId="10" pageNumber="151" type="discussion">
<paragraph id="79167970672676ED7628B18425CD1D94" pageId="10" pageNumber="151">
It is difficult to establish the synonymy of the species because of the absence of reliable figures. An essential part of the records of
<taxonomicName id="211FE38202D2B00B950689169B3C4FE7" class="Arachnida" family="Damaeidae" genus="Metabelba" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="pulverulenta">M. pulverulenta</taxonomicName>
from Central Europe may refer to
<taxonomicName id="B0AC8BCA92B0DBC8D42E6E40B4311A35" class="Arachnida" family="Damaeidae" genus="Metabelba" genus.bestMatchDistance="810" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="pulverosa">M. pulverosa</taxonomicName>
. According to the description and figure reproduced by Oudemans (1937, p. 2595, fig. 1117) it is, anyhow, impossible to identify Koch's species with certainty, because at present it is evident that the genus
<taxonomicName id="73EE2670A115ED5EABF8FDDBC8DBC523" class="Arachnida" family="Damaeidae" genus="Metabelba" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="151" phylum="Arthropoda" rank="genus">Metabelba</taxonomicName>
consists of a rather large number of species that are difficult to distinguish. As Koch's species is characterized by the absence of an anterior apophysis between I and II, and by the occurrence of laterally inserted notogastral hairs that surpass the border of the notogaster (&quot;Hinterleib ... mit einer Reihe kurzer,
<normalizedToken id="43B47100A7133C179D49D83AC7825107" originalValue="gekrümmter" pageId="10" pageNumber="151">gekruemmter</normalizedToken>
, spitzer Borsten
<normalizedToken id="385C74350565AAA4CE1D61E3B4BD3B71" originalValue="über" pageId="10" pageNumber="151">ueber</normalizedToken>
den Seiten und
<normalizedToken id="D5C9848E3F56E54C7AFF0CA71CF5431B" originalValue="über" pageId="10" pageNumber="151">ueber</normalizedToken>
dem Hinterrande&quot;), the identity of
<taxonomicName id="B0308DAB2028B79C5028D708DCE25F8F" class="Arachnida" family="Damaeidae" genus="Metabelba" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="pulverulenta">pulverulenta</taxonomicName>
and
<taxonomicName id="BAA0BF034CA6E841F9780D1CD61361A1" class="Arachnida" family="Damaeidae" genus="Metabelba" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="pulverosa">pulverosa</taxonomicName>
is not probable. As far as concerns the possibility of Koch's species being a
<taxonomicName id="9871CC2C83DE466C147687271784F69D" class="Arachnida" family="Damaeidae" genus="Porobelba" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="151" phylum="Arthropoda" rank="genus">Porobelba</taxonomicName>
, cf. Grandjean (1936, p. 67), van der Hammen (1952, p. 41), Strenzke (1952, p. 101).
</paragraph>
<paragraph id="8121D8C4CD30625723C0C6DBB3CDD6D0" pageId="10" pageNumber="151">
It is difficult to identify Kulczynski's
<taxonomicName id="E634D97F8A0F0361F5195119A4A5FF32" class="Arachnida" family="Achipteriidae" genus="Oribata" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="pulverulentus">Oribata pulverulentus</taxonomicName>
(1902, p. 35). Habitus and measurements (legs!), as well as the absence of protuberances on the propodosoma, point to the identity with
<taxonomicName id="780A022E0ABFD44409511B8EB15F2D70" class="Arachnida" family="Damaeidae" genus="Metabelba" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="pulverosa">pulverosa</taxonomicName>
. There are, however, important differences in the chaetotaxy of the legs, in the number of hairs (according to Kulczynski there is only one hair on trochanter IV), as well as in their shape (Kulczynski does not figure strongly serrate dorsal hairs on the proximal joints) (cf. Kulczynski, 1902, pl. 4 figs. 60, 61).
</paragraph>
<paragraph id="AE27EE83449573319D7FFE57E8D80695" lastPageId="11" lastPageNumber="152" pageId="10" pageNumber="151">
Turk's
<taxonomicName id="2D573B1C15E62C5AD475BA27613CB9B4" class="Arachnida" family="Damaeidae" genus="Porobelba" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="pulverulenta">Porobelba pulverulenta</taxonomicName>
(Turk and Turk, 1952) appears to be a species of
<taxonomicName id="D7A00CE4D74168785680C4D43329A12B" class="Arachnida" family="Damaeidae" genus="Metabelba" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="151" phylum="Arthropoda" rank="genus">Metabelba</taxonomicName>
; the slide is, however, too damaged to be identified
<pageBreakToken id="0EDF44A55224E24DD0811FF674E1F980" pageId="11" pageNumber="152" start="start">with</pageBreakToken>
certainty; possibly it concerns a species not dealt with in the present revision 1).
</paragraph>
<paragraph id="542057231121B1A6C22A1B1B0D8C65AA" pageId="11" pageNumber="152">
It is impossible to draw conclusions concerning the geographical distribution of
<taxonomicName id="15D0FD1704AF6D1B4233DB58BBF0EDA7" class="Arachnida" family="Damaeidae" genus="Metabelba" genus.bestMatchDistance="1220" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="152" phylum="Arthropoda" rank="species" species="pulverosa">M. pulverosa</taxonomicName>
. For a certainty the species is up till now only known from Germany (Holstein, Pomerania, Westphalia). Probably Willmann's records (1931, p. 120) also refer to the species. In northern Germany
<taxonomicName id="C6BC7B69E3F6F773DCE649FB04967BE0" class="Arachnida" family="Damaeidae" genus="Metabelba" genus.bestMatchDistance="1264" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="152" phylum="Arthropoda" rank="species" species="pulverosa">M. pulverosa</taxonomicName>
prefers localities with a strongly developed litter layer; the species is not rare in forest meadows, alder marshes, coniferous and deciduous woods, but it also occurs under
<taxonomicName id="2446E7376F582B61AF8AE9610C3ECCCB" family="Ericaceae" pageId="11" pageNumber="152" rank="family">Ericaceae</taxonomicName>
in moorlands; it avoids, however, bogs (Sphagneta) (cf. Strenzke, 1952, p. 102 sub
<taxonomicName id="A9160E67B22F160303B614C4E7B74518" class="Arachnida" family="Damaeidae" genus="Metabelba" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="152" phylum="Arthropoda" rank="species" species="pulverulenta">pulverulenta</taxonomicName>
).
</paragraph>
<paragraph id="514525C53974AD2E8423886D52321263" pageId="11" pageNumber="152">
It is a striking fact that
<taxonomicName id="96D02B58FAEA48BA74169BF81C6E0FA7" class="Arachnida" family="Damaeidae" genus="Metabelba" genus.bestMatchDistance="2714" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="152" phylum="Arthropoda" rank="species" species="papillipes">M. papillipes</taxonomicName>
, evidently not rare in the Netherlands, was not yet discovered in Germany, whilst on the other hand
<taxonomicName id="1CE26871F0148ED13A06C026C7CB2AAD" class="Arachnida" family="Damaeidae" genus="Metabelba" genus.bestMatchDistance="1354" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="152" phylum="Arthropoda" rank="species" species="pulverosa">M. pulverosa</taxonomicName>
was not yet found in the Netherlands.
</paragraph>
<paragraph id="B5EA0D2D1BB0BAC870E524EBB5341074" pageId="11" pageNumber="152">
1) The diagnosis of the specimen is as follows. Length 625 µ; colour light brown; the cerotegument is thin and slightly granulate. Notogastral hairs coarse and rather short; apart from one or two anterior pairs they are strongly curved. The measurements of the legs correspond exactly with those of the
<taxonomicName id="5E0694E236406AFC4EAB66BAA2D30447" class="Arachnida" family="Damaeidae" genus="Metabelba" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="152" phylum="Arthropoda" rank="species" species="montana">montana</taxonomicName>
specimen studied by us. Possibly Turk's specimen is indeed identical with
<taxonomicName id="64F7D4A25518993E118C7E36B453C11D" class="Arachnida" family="Damaeidae" genus="Metabelba" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="152" phylum="Arthropoda" rank="species" species="montana">montana</taxonomicName>
; the light colour and the different cerotegument may be due to the fact that at the date of its capture it had just become adult.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,186 @@
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<mods:title>Yunguirius gen. nov., a new genus of Coelotinae (Araneae, Agelenidae) spiders from southwest China</mods:title>
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<mods:namePart>Li, Bing</mods:namePart>
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<mods:namePart>Zhao, Zhe</mods:namePart>
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<mods:namePart>Okumura, Ken-ichi</mods:namePart>
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<taxonomicName LSID="4CBD2537-DABA-5386-A3D5-AFD473890A31" authority="(Zhang, Zhu &amp; Wang, 2017)" authorityName="Li &amp; Zhao &amp; Okumura &amp; Meng &amp; Li &amp; Chen" authorityYear="2023" baseAuthorityName="Zhang, Zhu &amp; Wang" baseAuthorityYear="2017" family="Agelenidae" genus="Yunguirius" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Yunguirius subterebratus" order="Araneae" pageId="0" pageNumber="51" rank="species" species="subterebratus" status="comb. nov.">Yunguirius subterebratus (Zhang, Zhu &amp; Wang, 2017)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="51">comb. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection pageId="0" pageNumber="51" type="reference_group">
<paragraph pageId="0" pageNumber="51">
<taxonomicName authorityName="Li &amp; Zhao &amp; Okumura &amp; Meng &amp; Li &amp; Chen" authorityYear="2023" class="Arachnida" family="Agelenidae" genus="Draconarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Draconarius subterebratus" order="Araneae" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="subterebratus" status="comb. nov.">Draconarius subterebratus</taxonomicName>
Zhang, Zhu &amp; Wang, 2017 in
<bibRefCitation author="Zhu, M" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="51" refId="B37" refString="Zhu, M, Wang, X, Zhang, Z, 2017. Fauna Sinica: Invertebrata (Vol. 59) Arachnida: Araneae: Agelenidae and Amaurobiidae. Science Press, Beijing" title="Fauna Sinica: Invertebrata (Vol. 59) Arachnida: Araneae: Agelenidae and Amaurobiidae." year="2017">Zhu et al. 2017</bibRefCitation>
: 379, fig. 245A, B.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="type material">
<paragraph pageId="0" pageNumber="51">Type material</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">
(not examined).
<emphasis italics="true" pageId="0" pageNumber="51">Holotype</emphasis>
</emphasis>
♀ (MHBU): China: Guizhou Province: Zunyi City: Daozhen County, Dashahegou Nature Reserve, Xieshiyan Cave to Dashahe River, 18.VIII.2004, Z. Zhang leg.
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Paratypes</emphasis>
: 3♀♀ (MHBU): same data as the holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="diagnosis">
<paragraph pageId="0" pageNumber="51">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName authorityName="Li &amp; Zhao &amp; Okumura &amp; Meng &amp; Li &amp; Chen" authorityYear="2023" baseAuthorityName="Zhang, Zhu &amp; Wang" baseAuthorityYear="2017" family="Agelenidae" genus="Yunguirius" kingdom="Animalia" lsidName="Yunguirius subterebratus" order="Araneae" pageId="0" pageNumber="51" rank="species" species="subterebratus">
<emphasis italics="true" pageId="0" pageNumber="51">Yunguirius subterebratus</emphasis>
</taxonomicName>
can be distinguished from other species of this genus as follows: 1) atrium cordiform (fig. 245A in
<bibRefCitation author="Zhu, M" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="51" refId="B37" refString="Zhu, M, Wang, X, Zhang, Z, 2017. Fauna Sinica: Invertebrata (Vol. 59) Arachnida: Araneae: Agelenidae and Amaurobiidae. Science Press, Beijing" title="Fauna Sinica: Invertebrata (Vol. 59) Arachnida: Araneae: Agelenidae and Amaurobiidae." year="2017">Zhu et al. 2017</bibRefCitation>
) in
<taxonomicName lsidName="Y. subterebratus" pageId="0" pageNumber="51" rank="species" species="subterebratus">
<emphasis italics="true" pageId="0" pageNumber="51">Y. subterebratus</emphasis>
</taxonomicName>
, but inverted trapezoid (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Epigyne and female habitus of Yunguirius ornatus comb. nov. A epigyne, ventral B vulva, dorsal C female habitus, dorsal D same, ventral E same, lateral. Scale bar equal for C-E. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; F = fold; FD = fertilization duct; H = hood; MS = median septum; PES = posterior epigynal sclerite; S = spermatheca; SB = spermathecal base; SH = spermathecal head; St = stalk." figureDoi="10.3897/zookeys.1159.100786.figure3" httpUri="https://binary.pensoft.net/fig/842425" pageId="0" pageNumber="51">3A</figureCitation>
; fig. 53 in
<bibRefCitation author="Wang, J" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="0" pageNumber="51" refId="B26" refString="Wang, J, Yin, C, Peng, X, Xie, L, 1990. New species of the spiders of the genus Coelotes from China (Araneae: Agelenidae). Spiders in China: One Hundred New and Newly Recorded Species of the Families Araneidae and Agelenidae. Hunan Normal University Press, 172-253." title="New species of the spiders of the genus Coelotes from China (Araneae: Agelenidae). Spiders in China: One Hundred New and Newly Recorded Species of the Families Araneidae and Agelenidae. Hunan Normal University Press, 172 - 253." year="1990">Wang et al. 1990</bibRefCitation>
) in
<taxonomicName lsidName="Y. ornatus" pageId="0" pageNumber="51" rank="species" species="ornatus">
<emphasis italics="true" pageId="0" pageNumber="51">Y. ornatus</emphasis>
</taxonomicName>
, or subrounded (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epigyne and female habitus of Yunguirius duoge sp. nov. A epigyne, ventral B vulva, dorsal C female habitus, dorsal D same, ventral E same, lateral. Scale bar equal for C-E. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; F = fold; FD = fertilization duct; H = hood; PES = posterior epigynal sclerite; S = spermatheca; SB = spermathecal base; SH = spermathecal head; St = stalk." figureDoi="10.3897/zookeys.1159.100786.figure2" httpUri="https://binary.pensoft.net/fig/842424" pageId="0" pageNumber="51">2A</figureCitation>
) in
<taxonomicName lsidName="Y. duoge" pageId="0" pageNumber="51" rank="species" species="duoge">
<emphasis italics="true" pageId="0" pageNumber="51">Y. duoge</emphasis>
</taxonomicName>
sp. nov. and (fig. 28 in
<bibRefCitation author="Peng, X" journalOrPublisher="Bulletin - British Arachnological Society" pageId="0" pageNumber="51" pagination="327 - 333" refId="B19" refString="Peng, X, Wang, J, 1997. Seven new species of the genus Coelotes (Araneae: Agelenidae) from China. Bulletin - British Arachnological Society 10: 327 - 333" title="Seven new species of the genus Coelotes (Araneae: Agelenidae) from China." volume="10" year="1997">Peng and Wang 1997</bibRefCitation>
)
<taxonomicName lsidName="Y. terebratus" pageId="0" pageNumber="51" rank="species" species="terebratus">
<emphasis italics="true" pageId="0" pageNumber="51">Y. terebratus</emphasis>
</taxonomicName>
; and 2) posterior epigynal sclerite longer than wide, waist-drum-shaped (fig. 245A in
<bibRefCitation author="Zhu, M" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="51" refId="B37" refString="Zhu, M, Wang, X, Zhang, Z, 2017. Fauna Sinica: Invertebrata (Vol. 59) Arachnida: Araneae: Agelenidae and Amaurobiidae. Science Press, Beijing" title="Fauna Sinica: Invertebrata (Vol. 59) Arachnida: Araneae: Agelenidae and Amaurobiidae." year="2017">Zhu et al. 2017</bibRefCitation>
) in
<taxonomicName lsidName="Y. subterebratus" pageId="0" pageNumber="51" rank="species" species="subterebratus">
<emphasis italics="true" pageId="0" pageNumber="51">Y. subterebratus</emphasis>
</taxonomicName>
, but vase-shaped (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epigyne and female habitus of Yunguirius xiangding sp. nov. A epigyne, ventral B vulva, dorsal C female habitus, dorsal D same, ventral E same, lateral. Scale bar equal for C-E. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; F = fold; FD = fertilization duct; H = hood; PES = posterior epigynal sclerite; S = spermatheca; SB = spermathecal base; SH = spermathecal head; St = stalk." figureDoi="10.3897/zookeys.1159.100786.figure4" httpUri="https://binary.pensoft.net/fig/842426" pageId="0" pageNumber="51">4A</figureCitation>
) in
<taxonomicName lsidName="Y. xiangding" pageId="0" pageNumber="51" rank="species" species="xiangding">
<emphasis italics="true" pageId="0" pageNumber="51">Y. xiangding</emphasis>
</taxonomicName>
sp. nov., or rectangular (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epigyne and female habitus of Yunguirius duoge sp. nov. A epigyne, ventral B vulva, dorsal C female habitus, dorsal D same, ventral E same, lateral. Scale bar equal for C-E. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; F = fold; FD = fertilization duct; H = hood; PES = posterior epigynal sclerite; S = spermatheca; SB = spermathecal base; SH = spermathecal head; St = stalk." figureDoi="10.3897/zookeys.1159.100786.figure2" httpUri="https://binary.pensoft.net/fig/842424" pageId="0" pageNumber="51">2A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Epigyne and female habitus of Yunguirius ornatus comb. nov. A epigyne, ventral B vulva, dorsal C female habitus, dorsal D same, ventral E same, lateral. Scale bar equal for C-E. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; F = fold; FD = fertilization duct; H = hood; MS = median septum; PES = posterior epigynal sclerite; S = spermatheca; SB = spermathecal base; SH = spermathecal head; St = stalk." figureDoi="10.3897/zookeys.1159.100786.figure3" httpUri="https://binary.pensoft.net/fig/842425" pageId="0" pageNumber="51">3A</figureCitation>
; fig. 53 in
<bibRefCitation author="Wang, J" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="0" pageNumber="51" refId="B26" refString="Wang, J, Yin, C, Peng, X, Xie, L, 1990. New species of the spiders of the genus Coelotes from China (Araneae: Agelenidae). Spiders in China: One Hundred New and Newly Recorded Species of the Families Araneidae and Agelenidae. Hunan Normal University Press, 172-253." title="New species of the spiders of the genus Coelotes from China (Araneae: Agelenidae). Spiders in China: One Hundred New and Newly Recorded Species of the Families Araneidae and Agelenidae. Hunan Normal University Press, 172 - 253." year="1990">Wang et al. 1990</bibRefCitation>
; fig. 28 in
<bibRefCitation author="Peng, X" journalOrPublisher="Bulletin - British Arachnological Society" pageId="0" pageNumber="51" pagination="327 - 333" refId="B19" refString="Peng, X, Wang, J, 1997. Seven new species of the genus Coelotes (Araneae: Agelenidae) from China. Bulletin - British Arachnological Society 10: 327 - 333" title="Seven new species of the genus Coelotes (Araneae: Agelenidae) from China." volume="10" year="1997">Peng and Wang 1997</bibRefCitation>
) in others.
</paragraph>
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<paragraph pageId="0" pageNumber="51">Description.</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Female</emphasis>
: See
<bibRefCitation author="Zhu, M" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="51" refId="B37" refString="Zhu, M, Wang, X, Zhang, Z, 2017. Fauna Sinica: Invertebrata (Vol. 59) Arachnida: Araneae: Agelenidae and Amaurobiidae. Science Press, Beijing" title="Fauna Sinica: Invertebrata (Vol. 59) Arachnida: Araneae: Agelenidae and Amaurobiidae." year="2017">Zhu et al. (2017)</bibRefCitation>
for complete description (fig. 245A, B).
</paragraph>
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<emphasis bold="true" pageId="0" pageNumber="51">Male.</emphasis>
Unknown.
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<paragraph pageId="0" pageNumber="51">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="51">
Guizhou Province, China (Fig.
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).
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<mods:titleInfo>
<mods:title>Nine genera of Eucnemidae (Coleoptera) new to Peru, with a key to Peruvian genera</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vahtera, Varpu</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Muona, Jyrki</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Linna, Ari</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Saeaeksjaervi, Ilari E.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>4493</mods:start>
<mods:end>4493</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/BDJ.3.e4493</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.3.e4493</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828-3-4493</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:4CBDC80859D8652DE690AB431C13BF5C" httpUri="http://treatment.plazi.org/id/4CBDC80859D8652DE690AB431C13BF5C" lastPageNumber="4493" pageId="0" pageNumber="4493">
<subSubSection pageId="0" pageNumber="4493" type="nomenclature">
<paragraph pageId="0" pageNumber="4493">
<taxonomicName authority="Bonvouloir, 1871" authorityName="Bonvouloir" authorityYear="1871" class="Insecta" family="Curculionidae" genus="Macraulacus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macraulacus" order="Coleoptera" pageId="0" pageNumber="4493" phylum="Arthropoda" rank="genus">Macraulacus Bonvouloir, 1871</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="4493" type="materials_examined">
<paragraph pageId="0" pageNumber="4493">Materials</paragraph>
<paragraph pageId="0" pageNumber="4493">
Type status:
<typeStatus pageId="0" pageNumber="4493">Other material</typeStatus>
. Taxon: higherClassification: Coleoptera; Eucnemidae; Macraulacinae; Macraulacini; genus: Macraulacus Bonvouloir, 1871
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="4493" type="notes">
<paragraph pageId="0" pageNumber="4493">Notes</paragraph>
<paragraph pageId="0" pageNumber="4493">Not found in our study, but an undescribed species is previously known from Peru (JM collection).</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.899.47070" ID-GBIF-Dataset="2c7e9ce7-8a11-4d64-9a73-40bb2eb487fb" ID-PMC="PMC6926428" ID-Pensoft-Pub="1313-2970-899-109" ID-Pensoft-UUID="E0FA12EBCCD55E3B8BC61796697C69C3" ID-PubMed="31875090" ID-ZooBank="9AA5216D3150475DA165B36EABCB61E2" ModsDocID="1313-2970-899-109" checkinTime="1576248706674" checkinUser="pensoft" docAuthor="Taylor, John D. &amp; Glover, Emily A." docDate="2019" docId="4CBDD7281E8250DC96E8AC1BAD8C281D" docLanguage="en" docName="ZooKeys 899: 109-140" docOrigin="ZooKeys 899" docSource="http://dx.doi.org/10.3897/zookeys.899.47070" docTitle="Pusillolucina arabica Taylor &amp; Glover, 2019, sp. nov." docType="treatment" docVersion="4" id="E0FA12EBCCD55E3B8BC61796697C69C3" lastPageNumber="109" masterDocId="E0FA12EBCCD55E3B8BC61796697C69C3" masterDocTitle="Unloved, paraphyletic or misplaced: new genera and species of small to minute lucinid bivalves and their relationships (Bivalvia, Lucinidae)" masterLastPageNumber="140" masterPageNumber="109" pageNumber="109" updateTime="1668168230035" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Unloved, paraphyletic or misplaced: new genera and species of small to minute lucinid bivalves and their relationships (Bivalvia, Lucinidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Taylor, John D.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Glover, Emily A.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>899</mods:number>
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<mods:start>109</mods:start>
<mods:end>140</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.899.47070</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.899.47070</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-899-109</mods:identifier>
<mods:identifier type="ZooBank">9AA5216D3150475DA165B36EABCB61E2</mods:identifier>
<mods:identifier type="Pensoft-UUID">E0FA12EBCCD55E3B8BC61796697C69C3</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="160702939" LSID="urn:lsid:plazi:treatment:4CBDD7281E8250DC96E8AC1BAD8C281D" httpUri="http://treatment.plazi.org/id/4CBDD7281E8250DC96E8AC1BAD8C281D" lastPageNumber="109" pageId="0" pageNumber="109">
<subSubSection pageId="0" pageNumber="109" type="nomenclature">
<paragraph pageId="0" pageNumber="109">
<taxonomicName LSID="4CBDD728-1E82-50DC-96E8-AC1BAD8C281D" class="Bivalvia" family="Lucinidae" genus="Pusillolucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pusillolucina arabica" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="arabica">Pusillolucina arabica</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="109">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Internal drawings of left valves of Rugalucina and Pusillolucina species. Not to scale." figureDoi="10.3897/zookeys.899.47070.figure3" httpUri="https://binary.pensoft.net/fig/364610" pageId="0" pageNumber="109" tableDoi="10.3897/zookeys.899.47070.figure3">Figs 3</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Pusillolucina arabica sp. nov. Tarut Bay, Saudi Arabia, Arabian Gulf. A-C Holotype (NHMUK 20191077) exterior of right valve and interior of left and right valves, L 2.4 mm D, E holotype, interior of left and right valves SEMs F-H paratype (NHMUK 20191078) exterior of right valve and interior of left and right valves, L 2.1 mm I, J paratype F-H (NHMUK 20191078) interior of left and right valves SEM K, L paratype (NHMUK 20191078) exterior of left and right valves, L 2.0 mm M paratype (NHMUK 20191078) dorsal view, L 1.9 mm N protoconch of holotype, scale bar 50 µm O, P paratype I-J detail of multi-cuspate posterior lateral hinge teeth. Scale bar: 200 µm." figureDoi="10.3897/zookeys.899.47070.figure9" httpUri="https://binary.pensoft.net/fig/364616" pageId="0" pageNumber="109" tableDoi="10.3897/zookeys.899.47070.figure9">9</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Pusillolucina arabica sp. nov. Arabian Gulf NHMUK consultancy samples- see text for details. A Lateral view of right valve stn 41 C (NHMUK 20191079) stained with Rose Bengal, L 1.3 mm B lateral view of left valve stn 41 C (NHMUK 20191079), L 1.2 mm C, D exterior of left (L 1.5 mm) and right (L 1.3 mm) valves stn 39 FC (NHMUK 20191081) E, F Interior of right (L 1.4 mm) and left (L 1.5 mm) valves stn 36 FC (NHMUK 20191080) G dorsal view, stn 39 FC (NHMUK 20191081), L 1.3 mm. H juvenile shell, stn 48 FC (NHMUK 20191082), L 1.1 mm I juvenile shell, stn 36 FC (NHMUK 20191080), L 0.9 mm J-L hinge teeth of left and right valves stn 36 FC (NHMUK 20191080) M detail of posterior lateral tooth of right valve stn 36 FC (NHMUK 20191080) N section through a ctenidial demibranch with thickened bacteriocyte zone, critical point dried preparation O symbiotic bacteria. Scale bars: 300 µm (J-L); 100 µm (M); 50 µm (N); 4 µm (O)." figureDoi="10.3897/zookeys.899.47070.figure10" httpUri="https://binary.pensoft.net/fig/364617" pageId="0" pageNumber="109" tableDoi="10.3897/zookeys.899.47070.figure10">10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="type material">
<paragraph pageId="0" pageNumber="109">Type material.</paragraph>
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" italics="true" pageId="0" pageNumber="109">Holotype</emphasis>
: NHMUK 20191077 L 2.4 mm.
<emphasis bold="true" italics="true" pageId="0" pageNumber="109">Paratypes</emphasis>
: figured L 2.1 mm, L 2.0 mm, L 1.9 mm, unfigured 11 sh, 2 v (NHMUK 20191078).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="type locality">
<paragraph pageId="0" pageNumber="109">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="109">Arabian Gulf, Tarut Bay, Saudi Arabia, dredged (17.5.1971) K. Smythe collection.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="materials_examined">
<paragraph pageId="0" pageNumber="109">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="109">
NHMUK consultancy report ECM 5027C/06. Arabian Gulf: station 36 FC,
<geoCoordinate degrees="27" direction="north" minutes="42" orientation="latitude" precision="15" seconds="38" value="27.710556">27°42'38&quot;N</geoCoordinate>
,
<geoCoordinate degrees="52" direction="east" minutes="11" orientation="longitude" precision="15" seconds="16" value="52.18778">52°11'16&quot;E</geoCoordinate>
25 m (NHMUK20191080), stn 39 FC,
<geoCoordinate degrees="27" direction="north" minutes="42" orientation="latitude" precision="15" seconds="05" value="27.70139">27°42'05&quot;N</geoCoordinate>
,
<geoCoordinate degrees="52" direction="east" minutes="10" orientation="longitude" precision="15" seconds="50" value="52.180557">52°10'50&quot;E</geoCoordinate>
31 m, NHMUK 20191081), stn 41 FC, 27°42'31&quot;N, 52°10'02&quot;E 32 m (NHMUK 20191079), stn 48 FC,
<geoCoordinate degrees="27" direction="north" minutes="45" orientation="latitude" precision="1" seconds="0.27" value="27.750074">27°45'0.27&quot;N</geoCoordinate>
,
<geoCoordinate degrees="52" direction="east" minutes="07" orientation="longitude" precision="15" seconds="46" value="52.129444">52°07'46&quot;E</geoCoordinate>
23 m (NHMUK 20191082).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="etymology">
<paragraph pageId="0" pageNumber="109">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName lsidName="arabica" pageId="0" pageNumber="109" rank="species" species="arabica">
<emphasis italics="true" pageId="0" pageNumber="109">arabica</emphasis>
</taxonomicName>
from Latin
<emphasis italics="true" pageId="0" pageNumber="109">arabicus</emphasis>
. Used as an adjective.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="diagnosis">
<paragraph pageId="0" pageNumber="109">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Pusillolucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pusillolucina" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Pusillolucina</emphasis>
</taxonomicName>
with posterior lateral teeth divided into four or five cusps and sockets.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="description">
<paragraph pageId="0" pageNumber="109">Description.</paragraph>
<paragraph pageId="0" pageNumber="109">
Shell very small, L to 2.4 mm, ovate, umbones prominent, sculpture of closely spaced, narrow, commarginal lamellae, sometimes elevated at posterior and anterior dorsal margins, crossed at anterior and posterior by low radial ribs, juvenile shells with commarginal lamellae only. Colour: white, translucent when wet. Protoconch: P1 84
<normalizedToken originalValue="µm">µm</normalizedToken>
, P1 + P2 = 155
<normalizedToken originalValue="µm">µm</normalizedToken>
, P2 with numerous growth increments. Lunule broadly lanceolate, smooth. Ligament internal, short, set alongside cardinal teeth. Hinge: right valve with single cardinal tooth, anterior lateral tooth located above anterior adductor muscle. Posterior lateral tooth long, divided into four or five cusps, left valve with two cardinal teeth, the anterior larger, anterior lateral tooth small, posterior lateral tooth divided into four or five sockets for cusps of right valve. Anterior adductor muscle scar short, barely detached from pallial line, posterior scar ovoid. Inner shell margin crenulate, more strongly to anterior and posterior.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="remarks">
<paragraph pageId="0" pageNumber="109">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Pusillolucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pusillolucina arabica" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="arabica">
<emphasis italics="true" pageId="0" pageNumber="109">Pusillolucina arabica</emphasis>
</taxonomicName>
differs from the Philippine
<taxonomicName lsidName="P. pusilla" pageId="0" pageNumber="109" rank="species" species="pusilla">
<emphasis italics="true" pageId="0" pageNumber="109">P. pusilla</emphasis>
</taxonomicName>
by the more ovate outline, and the less prominent commarginal and radial sculpture. By comparison,
<taxonomicName lsidName="P. africana" pageId="0" pageNumber="109" rank="species" species="africana">
<emphasis italics="true" pageId="0" pageNumber="109">P. africana</emphasis>
</taxonomicName>
has finer more closely spaced commarginal sculpture and more cusps (7-8) on the posterior lateral tooth. From South Africa
<taxonomicName lsidName="P. denticula" pageId="0" pageNumber="109" rank="species" species="denticula">
<emphasis italics="true" pageId="0" pageNumber="109">P. denticula</emphasis>
</taxonomicName>
has much finer commarginal sculpture and up to 10 smaller cusps on the posterior lateral tooth. From Madagascar
<taxonomicName lsidName="P. biritika" pageId="0" pageNumber="109" rank="species" species="biritika">
<emphasis italics="true" pageId="0" pageNumber="109">P. biritika</emphasis>
</taxonomicName>
sp. nov. has only three cusps on the posterior lateral tooth.
</paragraph>
<paragraph pageId="0" pageNumber="109">
Despite its small size,
<taxonomicName lsidName="P. arabica" pageId="0" pageNumber="109" rank="species" species="arabica">
<emphasis italics="true" pageId="0" pageNumber="109">P. arabica</emphasis>
</taxonomicName>
has the characteristic, thick inner ctenidial demibranchs with a well-developed bacteriocyte zone packed with symbiotic bacteria ca 3-4
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Pusillolucina arabica sp. nov. Arabian Gulf NHMUK consultancy samples- see text for details. A Lateral view of right valve stn 41 C (NHMUK 20191079) stained with Rose Bengal, L 1.3 mm B lateral view of left valve stn 41 C (NHMUK 20191079), L 1.2 mm C, D exterior of left (L 1.5 mm) and right (L 1.3 mm) valves stn 39 FC (NHMUK 20191081) E, F Interior of right (L 1.4 mm) and left (L 1.5 mm) valves stn 36 FC (NHMUK 20191080) G dorsal view, stn 39 FC (NHMUK 20191081), L 1.3 mm. H juvenile shell, stn 48 FC (NHMUK 20191082), L 1.1 mm I juvenile shell, stn 36 FC (NHMUK 20191080), L 0.9 mm J-L hinge teeth of left and right valves stn 36 FC (NHMUK 20191080) M detail of posterior lateral tooth of right valve stn 36 FC (NHMUK 20191080) N section through a ctenidial demibranch with thickened bacteriocyte zone, critical point dried preparation O symbiotic bacteria. Scale bars: 300 µm (J-L); 100 µm (M); 50 µm (N); 4 µm (O)." figureDoi="10.3897/zookeys.899.47070.figure10" httpUri="https://binary.pensoft.net/fig/364617" pageId="0" pageNumber="109" tableDoi="10.3897/zookeys.899.47070.figure10">Fig. 10N, O</figureCitation>
) similar to other lucinids.
</paragraph>
<caption doi="10.3897/zookeys.899.47070.figure9" httpUri="https://binary.pensoft.net/fig/364616" pageId="0" pageNumber="109" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figure 9.</emphasis>
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Pusillolucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pusillolucina arabica" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="arabica">
<emphasis italics="true" pageId="0" pageNumber="109">Pusillolucina arabica</emphasis>
</taxonomicName>
sp. nov. Tarut Bay, Saudi Arabia, Arabian Gulf.
<emphasis bold="true" pageId="0" pageNumber="109">
<normalizedToken originalValue="AC">A-C</normalizedToken>
</emphasis>
Holotype (NHMUK 20191077) exterior of right valve and interior of left and right valves, L 2.4 mm
<emphasis bold="true" pageId="0" pageNumber="109">D, E</emphasis>
holotype, interior of left and right valves SEMs
<emphasis bold="true" pageId="0" pageNumber="109">
<normalizedToken originalValue="FH">F-H</normalizedToken>
</emphasis>
paratype (NHMUK 20191078) exterior of right valve and interior of left and right valves, L 2.1 mm
<emphasis bold="true" pageId="0" pageNumber="109">I, J</emphasis>
paratype F-H (NHMUK 20191078) interior of left and right valves SEM
<emphasis bold="true" pageId="0" pageNumber="109">K, L</emphasis>
paratype (NHMUK 20191078) exterior of left and right valves, L 2.0 mm
<emphasis bold="true" pageId="0" pageNumber="109">M</emphasis>
paratype (NHMUK 20191078) dorsal view, L 1.9 mm
<emphasis bold="true" pageId="0" pageNumber="109">N</emphasis>
protoconch of holotype, scale bar 50
<normalizedToken originalValue="µm">µm</normalizedToken>
<emphasis bold="true" pageId="0" pageNumber="109">O, P</emphasis>
paratype
<emphasis bold="true" pageId="0" pageNumber="109">
<normalizedToken originalValue="IJ">I-J</normalizedToken>
</emphasis>
detail of multi-cuspate posterior lateral hinge teeth. Scale bar: 200
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.899.47070.figure10" httpUri="https://binary.pensoft.net/fig/364617" pageId="0" pageNumber="109" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figure 10.</emphasis>
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Pusillolucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pusillolucina arabica" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="arabica">
<emphasis italics="true" pageId="0" pageNumber="109">Pusillolucina arabica</emphasis>
</taxonomicName>
sp. nov. Arabian Gulf NHMUK consultancy samples- see text for details.
<emphasis bold="true" pageId="0" pageNumber="109">A</emphasis>
Lateral view of right valve stn 41C (NHMUK 20191079) stained with Rose Bengal, L 1.3 mm
<emphasis bold="true" pageId="0" pageNumber="109">B</emphasis>
lateral view of left valve stn 41C (NHMUK 20191079), L 1.2 mm
<emphasis bold="true" pageId="0" pageNumber="109">C, D</emphasis>
exterior of left (L 1.5 mm) and right (L 1.3 mm) valves stn 39FC (NHMUK 20191081)
<emphasis bold="true" pageId="0" pageNumber="109">E, F</emphasis>
Interior of right (L 1.4 mm) and left (L 1.5 mm) valves stn 36FC (NHMUK 20191080)
<emphasis bold="true" pageId="0" pageNumber="109">G</emphasis>
dorsal view, stn 39FC (NHMUK 20191081), L 1.3 mm.
<emphasis bold="true" pageId="0" pageNumber="109">H</emphasis>
juvenile shell, stn 48FC (NHMUK 20191082), L 1.1 mm
<emphasis bold="true" pageId="0" pageNumber="109">I</emphasis>
juvenile shell, stn 36FC (NHMUK 20191080), L 0.9 mm
<emphasis bold="true" pageId="0" pageNumber="109">
<normalizedToken originalValue="JL">J-L</normalizedToken>
</emphasis>
hinge teeth of left and right valves stn 36FC (NHMUK 20191080)
<emphasis bold="true" pageId="0" pageNumber="109">M</emphasis>
detail of posterior lateral tooth of right valve stn 36FC (NHMUK 20191080)
<emphasis bold="true" pageId="0" pageNumber="109">N</emphasis>
section through a ctenidial demibranch with thickened bacteriocyte zone, critical point dried preparation
<emphasis bold="true" pageId="0" pageNumber="109">O</emphasis>
symbiotic bacteria. Scale bars: 300
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="109">
<normalizedToken originalValue="JL">J-L</normalizedToken>
</emphasis>
); 100
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="109">M</emphasis>
); 50
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="109">N</emphasis>
); 4
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="109">O</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>