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<document id="6D0665642C608D47370167DB7A1A7AB5" ID-DOI="10.11646/phytotaxa.668.3.7" ID-ISSN="1179-3163" ID-Zenodo-Dep="14519257" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1734550332556" checkinUser="felipe" docAuthor="López-Floriano, Oscar Mauricio, Claros-Calderón, Brallan, Hágsater, Eric &amp; Santiago, Elizabeth" docDate="2024" docId="4D4887FD0B731E2D95A99DAE6EFA263C" docLanguage="en" docName="phytotaxa.668.3.279-285.pdf" docOrigin="Phytotaxa 668 (3)" docSource="https://doi.org/10.11646/phytotaxa.668.3.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Epidendrum clinidiodontum E. Santiago, Lopez-Flor., Claros &amp; Hagsater 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="282" masterDocId="B171FF850B721E2E95219B5A6A38210C" masterDocTitle="A new species of Epidendrum (Orchidaceae) of the Albertii group of the Eastern Andes in Colombia" masterLastPageNumber="285" masterPageNumber="279" pageNumber="280" updateTime="1740752167110" updateUser="guilherme" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="E70EC997E8925754ECE93E0B995BC506">
<mods:title id="628AC0D14F1E1D86B0332CE0D99BF5D8">A new species of Epidendrum (Orchidaceae) of the Albertii group of the Eastern Andes in Colombia</mods:title>
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<mods:namePart id="9800FF6355E8F259DF20331B9BCF3073">López-Floriano, Oscar Mauricio</mods:namePart>
<mods:nameIdentifier id="B04DC595EA4EB5F7B2F8747F4036683C" type="ORCID">0000-0002-0456-0203</mods:nameIdentifier>
<mods:affiliation id="AAC3A1E28460A35E87423A24F2F298FA">Herbario Enrique Forero HUAZ, Grupo de Investigación en Botánica Uniamazonia, Universidad de la Amazonia, Carrera 11 # 5 - 69, Florencia, Colombia.</mods:affiliation>
<mods:nameIdentifier id="026807AFAA0A4CCAD44FAB1B4AF4421B" type="email">os.lopez@udla.edu.co</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="20245BC0056B5AE38E212C18B7FC5343">Claros-Calderón, Brallan</mods:namePart>
<mods:nameIdentifier id="31624ECD9EAC35681142CFBC48864599" type="ORCID">0009-0009-3588-118X</mods:nameIdentifier>
<mods:affiliation id="4FE2F0E437A4ECC0C2C9177735E063EF">Herbario Enrique Forero HUAZ, Grupo de Investigación en Botánica Uniamazonia, Universidad de la Amazonia, Carrera 11 # 5 - 69, Florencia, Colombia.</mods:affiliation>
<mods:nameIdentifier id="C970E01B3152776D5BA6FE538AF753FD" type="email">brallanclaroscalderon9@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="9B1B59464B58EC6B51704F747E74837B">Hágsater, Eric</mods:namePart>
<mods:nameIdentifier id="31C3A92002482683557C1184F5A5FD6D" type="ORCID">0000-0002-2371-9427</mods:nameIdentifier>
<mods:affiliation id="FF4E36747C97D6FEBB15461F7E29B432">Herbario AMO, Montañas Calizas 490, Lomas de Chapultepec, Miguel Hidalgo, 11000 Ciudad de México, México.</mods:affiliation>
<mods:nameIdentifier id="6C120977DCFCC44DDEB1FFA3FE8689EF" type="email">erichag_1@msn.com</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="87C04C66D9467D4A074DE11A7B7C8A78">Santiago, Elizabeth</mods:namePart>
<mods:nameIdentifier id="434DD360536AACD73FAC796B4E9CC611" type="ORCID">0000-0002-1368-9025</mods:nameIdentifier>
<mods:affiliation id="57AF3D38D2622575E89089F4FCF14222">Herbario AMO, Montañas Calizas 490, Lomas de Chapultepec, Miguel Hidalgo, 11000 Ciudad de México, México.</mods:affiliation>
<mods:nameIdentifier id="088CEC1112483E5888AD4A8A9C86B612" type="email">sayalae2014@gmail.com</mods:nameIdentifier>
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<mods:title id="23964D2BFB2E5E19DE78F967C4E09CCA">Phytotaxa</mods:title>
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<mods:date id="C83861F253FF3892F2C632ACD786EBE7">2024</mods:date>
<mods:detail id="FB911659E9CBDAF9E502B322CB4B2DDF" type="pubDate">
<mods:number id="24F3C2A9B07CBCAD349F10EFEE1BB202">2024-10-16</mods:number>
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<mods:identifier id="8267340F3883565AF107D34357C611CB" type="DOI">10.11646/phytotaxa.668.3.7</mods:identifier>
<mods:identifier id="1CEBE233B78462C829C7FF0C52F64B6A" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="6C655044693EB2C89C06F0F75C2F3DD1" type="Zenodo-Dep">14519257</mods:identifier>
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<treatment id="4D4887FD0B731E2D95A99DAE6EFA263C" LSID="urn:lsid:plazi:treatment:4D4887FD0B731E2D95A99DAE6EFA263C" httpUri="http://treatment.plazi.org/id/4D4887FD0B731E2D95A99DAE6EFA263C" lastPageId="3" lastPageNumber="282" pageId="1" pageNumber="280">
<subSubSection id="8DFB65600B731E2F95A99DAE6EAC2602" box="[136,1172,1780,1806]" pageId="1" pageNumber="280" type="nomenclature">
<paragraph id="C55E36EB0B731E2F95A99DAE6EAC2602" blockId="1.[136,1451,1780,2022]" box="[136,1172,1780,1806]" pageId="1" pageNumber="280">
<taxonomicName id="02E14D680B731E2F95A99DAE698D2602" authority="E.Santiago, Lopez-Flor., Claros &amp; Hagsater" authorityName="E. Santiago, Lopez-Flor., Claros &amp; Hagsater" authorityYear="2024" box="[136,949,1780,1806]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="280" phylum="Tracheophyta" rank="species" species="clinidiodontum" status="sp. nov.">
<emphasis id="F795EAF90B731E2F95A99DAE6BFB2602" box="[136,451,1780,1806]" italics="true" pageId="1" pageNumber="280">Epidendrum clinidiodontum</emphasis>
E.Santiago, López-Flor., Claros &amp; Hágsater
</taxonomicName>
,
<emphasis id="F795EAF90B731E2F96E09DAF6E2E2602" box="[961,1046,1781,1806]" italics="true" pageId="1" pageNumber="280">
<taxonomicNameLabel id="ECA657820B731E2F96E09DAF6E2E2602" box="[961,1046,1781,1806]" pageId="1" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="5DDA2A6E0B731E2F91059DAE6EBA2602" box="[1060,1154,1780,1806]" captionStart="FIGURE 1" captionStartId="2.[136,229,1952,1974]" captionTargetBox="[205,1381,187,1925]" captionTargetId="figure-31@2.[205,1381,187,1925]" captionTargetPageId="2" captionText="FIGURE 1. Epidendrum clinidiodontum A. Plant. B. Flower with closeup of calli and bidentate clinandrium-hood. C. Perianth segments. D. Leaves and Inflorescence. E. Longitudinal section of the column. F. Spathaceous bracts. Based on López and Claros 273. Photographs by Oscar M. López-Floriano; LCDP prepared by A. Cisneros." figureDoi="http://doi.org/10.5281/zenodo.14519259" httpUri="https://zenodo.org/record/14519259/files/figure.png" pageId="1" pageNumber="280">Figure 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8DFB65600B731E2F95A99C4668842659" pageId="1" pageNumber="280" type="materials_examined">
<paragraph id="C55E36EB0B731E2F95A99C4668842659" blockId="1.[136,1451,1780,2022]" pageId="1" pageNumber="280">
<emphasis id="F795EAF90B731E2F95A99C466AFA263D" bold="true" box="[136,194,1820,1841]" pageId="1" pageNumber="280">Type:</emphasis>
—
<materialsCitation id="75893CB60B731E2F95FA9C4668192659" collectingDate="2023-05-09" collectionCode="HUAZ" collectorName="O. Lopez &amp; B. Claros &amp; M. Correa" country="Colombia" elevation="2036" location="Corregimiento el Carano, Vereda la Ruidosa" municipality="Florencia" pageId="1" pageNumber="280" specimenCount="1" stateProvince="Caqueta" typeStatus="holotype">
<collectingCountry id="BDF6767B0B731E2F95FA9C466B65263D" box="[219,349,1820,1841]" name="Colombia" pageId="1" pageNumber="280">COLOMBIA</collectingCountry>
.
<collectingRegion id="0725F8090B731E2F94479C466B80263D" box="[358,440,1820,1841]" country="Colombia" name="Caqueta" pageId="1" pageNumber="280">Caquetá</collectingRegion>
:
<collectingMunicipality id="253AAC910B731E2F94E09C466826263E" box="[449,542,1820,1842]" pageId="1" pageNumber="280">Florencia</collectingMunicipality>
,
<location id="C03E60300B731E2F97069C4669E8263E" LSID="urn:lsid:plazi:treatment:4D4887FD0B731E2D95A99DAE6EFA263C:C03E60300B731E2F97069C4669E8263E" box="[551,976,1820,1842]" country="Colombia" municipality="Florencia" name="Corregimiento el Carano, Vereda la Ruidosa" pageId="1" pageNumber="280" stateProvince="Caqueta">Corregimiento el Caraño, Vereda la Ruidosa</location>
,
<quantity id="02199B0E0B731E2F96FB9C466E1C263D" box="[986,1060,1820,1842]" metricMagnitude="3" metricUnit="m" metricValue="2.036" pageId="1" pageNumber="280" unit="m" value="2036.0">
<elevation id="4ECCD1D80B731E2F96FB9C466E1C263D" box="[986,1060,1820,1842]" metricMagnitude="3" metricUnit="m" metricValue="2.036" pageId="1" pageNumber="280" unit="m" value="2036.0">2036 m</elevation>
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,
<collectingDate id="A11BE9C30B731E2F910F9C466EDD263D" box="[1070,1253,1820,1842]" pageId="1" pageNumber="280" value="2023-05-09">
<date id="B15F102B0B731E2F910F9C466EDD263D" box="[1070,1253,1820,1842]" pageId="1" pageNumber="280" value="2023-05-09">9 de mayo de 2023</date>
</collectingDate>
,
<emphasis id="F795EAF90B731E2F91CE9C466B5D2659" italics="true" pageId="1" pageNumber="280">
<collectorName id="6814533D0B731E2F91CE9C466F70263D" box="[1263,1352,1820,1841]" pageId="1" pageNumber="280">O. López</collectorName>
,
<collectorName id="6814533D0B731E2F90709C466F93263D" box="[1361,1451,1820,1841]" pageId="1" pageNumber="280">B. Claros</collectorName>
&amp;
<collectorName id="6814533D0B731E2F95F79C1A6B022659" box="[214,314,1856,1877]" pageId="1" pageNumber="280">M. Correa</collectorName>
273
</emphasis>
(
<typeStatus id="1A5A88490B731E2F94509C1A6BF2265A" box="[369,458,1856,1878]" pageId="1" pageNumber="280" type="holotype">holotype</typeStatus>
:
<collectionCode id="A3F0AE2E0B731E2F94F29C1A68252659" box="[467,541,1856,1877]" collectionName="HUAZ" pageId="1" pageNumber="280">HUAZ</collectionCode>
!
</materialsCitation>
,
<materialsCitation id="75893CB60B731E2F970B9C1A688A2659" box="[554,690,1856,1878]" collectingDate="2023-05-09" collectionCode="COL" collectorName="O. Lopez &amp; B. Claros &amp; M. Correa" country="Colombia" elevation="2036" location="Corregimiento el Carano, Vereda la Ruidosa" municipality="Florencia" pageId="1" pageNumber="280" specimenCount="1" stateProvince="Caqueta" typeStatus="isotype">
<typeStatus id="1A5A88490B731E2F970B9C1A6857265A" box="[554,623,1856,1878]" pageId="1" pageNumber="280" type="isotype">isotype</typeStatus>
<collectionCode id="A3F0AE2E0B731E2F97549C1A68952659" box="[629,685,1856,1877]" collectionName="COL" pageId="1" pageNumber="280">COL</collectionCode>
!
</materialsCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8DFB65600B731E2F95A99C3E6B5F26E9" pageId="1" pageNumber="280" type="diagnosis">
<paragraph id="C55E36EB0B731E2F95A99C3E6B5F26E9" blockId="1.[136,1451,1780,2022]" pageId="1" pageNumber="280">
Similar to
<taxonomicName id="02E14D680B731E2F95CC9C3E690B2675" authority="Hagsater &amp; E. Santiago (2004)" authorityName="Hagsater &amp; E. Santiago" authorityYear="2004" box="[237,819,1892,1914]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="280" phylum="Tracheophyta" rank="species" species="hygrohylephilum">
<emphasis id="F795EAF90B731E2F95CC9C3E68332675" box="[237,523,1892,1913]" italics="true" pageId="1" pageNumber="280">Epidendrum hygrohylephilum</emphasis>
Hágsater &amp; E.Santiago (2004)
</taxonomicName>
but the lip entire pentagonal, margin crenate (
<emphasis id="F795EAF90B731E2F91CC9C3E6F302675" box="[1261,1288,1892,1913]" italics="true" pageId="1" pageNumber="280">vs.</emphasis>
entire, basal half semi-elliptic, apical half triangular, margin entire), the apical margin of clinandrium-hood crenulate, apex bidentate (
<emphasis id="F795EAF90B731E2F900C9CD26F7F2691" box="[1325,1351,1928,1949]" italics="true" pageId="1" pageNumber="280">vs.</emphasis>
the apical margin of clinandrium-hood entire, apex truncate), and the lip bicallose, calli laminar, leaning outwards (
<emphasis id="F795EAF90B731E2F91E39CF66EE526CD" box="[1218,1245,1964,1985]" italics="true" pageId="1" pageNumber="280">vs.</emphasis>
lip unicallose, callus triangular, acute).
</paragraph>
</subSubSection>
<caption id="919E66630B701E2C95A99CFA68E126F3" ID-DOI="http://doi.org/10.5281/zenodo.14519259" ID-Zenodo-Dep="14519259" httpUri="https://zenodo.org/record/14519259/files/figure.png" pageId="2" pageNumber="281" startId="2.[136,229,1952,1974]" targetBox="[205,1381,187,1925]" targetPageId="2" targetType="figure">
<paragraph id="C55E36EB0B701E2C95A99CFA68E126F3" blockId="2.[136,1452,1952,2047]" pageId="2" pageNumber="281">
<emphasis id="F795EAF90B701E2C95A99CFA6AC526B9" bold="true" box="[136,253,1952,1974]" pageId="2" pageNumber="281">FIGURE 1.</emphasis>
<taxonomicName id="02E14D680B701E2C94229CFB68F726BB" authority="A. Plant. B. Flower" box="[259,719,1952,1975]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="281" phylum="Tracheophyta" rank="species" species="clinidiodontum">
<emphasis id="F795EAF90B701E2C94229CFB682826BA" box="[259,528,1953,1974]" italics="true" pageId="2" pageNumber="281">Epidendrum clinidiodontum</emphasis>
<emphasis id="F795EAF90B701E2C97379CFA681526B9" bold="true" box="[534,557,1952,1973]" pageId="2" pageNumber="281">A.</emphasis>
Plant.
<emphasis id="F795EAF90B701E2C97519CFB68B826BA" bold="true" box="[624,640,1953,1974]" pageId="2" pageNumber="281">B</emphasis>
. Flower
</taxonomicName>
with closeup of calli and bidentate clinandrium-hood.
<emphasis id="F795EAF90B701E2C91FB9CFA6ED326BA" bold="true" box="[1242,1259,1952,1974]" pageId="2" pageNumber="281">C</emphasis>
. Perianth segments.
<emphasis id="F795EAF90B701E2C95A99C9F6AA126D6" bold="true" box="[136,153,1989,2010]" pageId="2" pageNumber="281">D</emphasis>
. Leaves and Inflorescence.
<emphasis id="F795EAF90B701E2C94859C9F6B8C26D6" bold="true" box="[420,436,1989,2010]" pageId="2" pageNumber="281">E</emphasis>
. Longitudinal section of the column.
<emphasis id="F795EAF90B701E2C963A9C9F691226D6" bold="true" box="[795,810,1989,2010]" pageId="2" pageNumber="281">F</emphasis>
. Spathaceous bracts. Based on
<emphasis id="F795EAF90B701E2C91749C9F6F1026D6" box="[1109,1320,1989,2010]" italics="true" pageId="2" pageNumber="281">López and Claros 273</emphasis>
. Photographs by Oscar M. López-Floriano; LCDP prepared by A. Cisneros.
</paragraph>
</caption>
<subSubSection id="8DFB65600B711E2D95A99BC46F6322A1" pageId="3" pageNumber="282" type="description">
<paragraph id="C55E36EB0B711E2D95A99BC46F6322A1" blockId="3.[136,1452,158,1841]" pageId="3" pageNumber="282">
<emphasis id="F795EAF90B711E2D95A99BC46B0021B4" bold="true" box="[136,312,158,184]" pageId="3" pageNumber="282">Description:—</emphasis>
Epiphytic, sympodial, caespitose, erect herb, 34.0 cm tall including inflorescence. Roots 2.0 mm in diameter, basal, scarce, thin. Stems 27.0 ×
<quantity id="02199B0E0B711E2D974A9B9868D321D1" box="[619,747,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="6" metricValueMax="8.0" metricValueMin="4.0" pageId="3" pageNumber="282" unit="cm" value="0.6" valueMax="0.8" valueMin="0.4">0.4–0.8 cm</quantity>
, simple, cane
<emphasis id="F795EAF90B711E2D96A59B9969B521D0" box="[900,909,195,220]" italics="true" pageId="3" pageNumber="282">-</emphasis>
like, laterally compressed at base and ancipitose above, straight. Leaves 5, distributed throughout stems, alternate, articulate, sub-erect; sheaths 3.7–5.2 ×
<quantity id="02199B0E0B711E2D90069BBC6F9F200D" box="[1319,1447,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="6" metricValueMax="8.0" metricValueMin="4.0" pageId="3" pageNumber="282" unit="cm" value="0.6" valueMax="0.8" valueMin="0.4">0.4–0.8 cm</quantity>
, tubular, laterally compressed, ancipitose, minutely striated, green, concolor; blade 7.2–12.5 ×
<quantity id="02199B0E0B711E2D91969A506F012029" box="[1207,1337,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="3" pageNumber="282" unit="cm" value="2.05" valueMax="2.3" valueMin="1.8">1.8–2.3 cm</quantity>
, narrowly elliptic, apex acuminate, sub
<emphasis id="F795EAF90B711E2D94F19A756BE12044" box="[464,473,303,328]" italics="true" pageId="3" pageNumber="282">-</emphasis>
coriaceous, margin entire, spreading. Spathes 2–3, at apex of peduncle, lower spathe 4.6–6.7 × 0.8–1.0 cm (conduplicate), narrowly ovate, apex obtuse, base tubular, ancipitose, green, concolor, second spathe smaller, 2.0 ×
<quantity id="02199B0E0B711E2D94579A2C6BF4209D" box="[374,460,374,401]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="282" unit="mm" value="0.7">0.7 mm</quantity>
(conduplicate), dorsally keeled serrulate at apex, green, concolor, similar to floral bracts but larger. Inflorescence
<quantity id="02199B0E0B711E2D94829AC06BC920B9" box="[419,497,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="282" unit="cm" value="7.5">7.5 cm</quantity>
long, apical (probably with lateral inflorescences in mature stems), short, compact, few
<emphasis id="F795EAF90B711E2D95939AE56A8320D4" box="[178,187,447,472]" italics="true" pageId="3" pageNumber="282">-</emphasis>
flowered, producing new racemes from same inflorescences through time and becoming pluri
<emphasis id="F795EAF90B711E2D91E79AE56EF720D4" box="[1222,1231,447,472]" italics="true" pageId="3" pageNumber="282">-</emphasis>
racemose; peduncle ca. 5.0 cm long, laterally compressed, ancipitose, short, covered at apex by 2 unequal spathes. Floral bracts up to 13.0 mm long (progressively smaller), much longer than ovary, conduplicate, ovate, obtuse, imbricated. Flowers developing one at a time, resupinate, pale green, lip, and column apex cream colored; fragrance not registered. Ovary ca. 9.0–10.0 mm long, completely hidden by floral bract, ventrally inflated for apical half. Sepals 15–16 × 4.0–
<quantity id="02199B0E0B711E2D91F599146F142365" box="[1236,1324,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="282" unit="mm" value="4.3">4.3 mm</quantity>
; spreading, free, convex, narrowly elliptic, sub
<emphasis id="F795EAF90B711E2D972D9929682D2380" box="[524,533,627,652]" italics="true" pageId="3" pageNumber="282">-</emphasis>
acute, apiculate, 5
<emphasis id="F795EAF90B711E2D97FF992968DF2380" box="[734,743,627,652]" italics="true" pageId="3" pageNumber="282">-</emphasis>
veined, margin entire, some revolute; lateral sepals prominently aristate at apex. Petals 14.5 ×
<quantity id="02199B0E0B711E2D94C599CC680723BD" box="[484,575,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="282" unit="mm" value="1.8">1.8 mm</quantity>
, sub-parallel to column, free, linear
<emphasis id="F795EAF90B711E2D96F899CD69DA23BC" box="[985,994,663,688]" italics="true" pageId="3" pageNumber="282">-</emphasis>
oblong, apex rounded, 3
<emphasis id="F795EAF90B711E2D91D999CD6F3923BC" box="[1272,1281,663,688]" italics="true" pageId="3" pageNumber="282">-</emphasis>
veined, margin entire, some revolute. Lip 9.0 ×
<quantity id="02199B0E0B711E2D94D199E0686E23D9" box="[496,598,698,725]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="282" unit="mm" value="10.5">10.5 mm</quantity>
, united to column, entire, slightly convex, pentagonal, base slightly cordate, margin crenate, spreading, somewhat hyaline; bicallose, calli large, laminar, fleshy, sub
<emphasis id="F795EAF90B711E2D914E99856E4023F4" box="[1135,1144,735,760]" italics="true" pageId="3" pageNumber="282">-</emphasis>
erect and leaning outwards meeting sides of apex of clinandrium
<emphasis id="F795EAF90B711E2D9711985968012210" box="[560,569,771,796]" italics="true" pageId="3" pageNumber="282">-</emphasis>
hood; disc fleshy. Column 9.0 mm long, thin at base, thickened in apical 2/3, straight, apex with a pair of teeth each side. Clinandrium
<emphasis id="F795EAF90B711E2D97DD987D693D224C" box="[764,773,807,832]" italics="true" pageId="3" pageNumber="282">-</emphasis>
hood prominent, nearly half length of column, tubular, apical margin crenulate, apex bidentate, teeth triangular, central, adjacent. Anther not seen. Pollinia not seen. Rostellum apical, slit; viscarium semi
<emphasis id="F795EAF90B711E2D949698356BF82284" box="[439,448,879,904]" italics="true" pageId="3" pageNumber="282">-</emphasis>
liquid. Lateral lobes of stigma small, about ¼ length of stigmatic cavity. Cuniculus deep, penetrating ¾ of pedicellate ovary, widened behind perianth and forming prominent vesicle. Capsule not seen.
</paragraph>
</subSubSection>
<subSubSection id="8DFB65600B711E2D959C98EC6A9E2514" pageId="3" pageNumber="282" type="distribution">
<paragraph id="C55E36EB0B711E2D959C98EC6A9E2514" blockId="3.[136,1452,158,1841]" pageId="3" pageNumber="282">
<emphasis id="F795EAF90B711E2D959C98EC6BC522DC" bold="true" box="[189,509,950,976]" pageId="3" pageNumber="282">Distribution and Ecology:</emphasis>
—Presently known from the eastern slope of the Andes in southern and central
<collectingCountry id="BDF6767B0B711E2D95A998806AC122F8" box="[136,249,986,1012]" name="Colombia" pageId="3" pageNumber="282">Colombia</collectingCountry>
, in the departments of
<collectingRegion id="0725F8090B711E2D94D69880686B22F8" box="[503,595,986,1012]" country="Colombia" name="Caqueta" pageId="3" pageNumber="282">Caquetá</collectingRegion>
and
<collectingRegion id="0725F8090B711E2D97A99880691722F8" box="[648,815,986,1012]" country="Colombia" name="Cundinamarca" pageId="3" pageNumber="282">Cundinamarca</collectingRegion>
. Flowering in May, probably throughout the year (
<figureCitation id="5DDA2A6E0B711E2D904198806AA12514" captionStart="FIGURE 2" captionStartId="4.[136,229,1899,1921]" captionTargetBox="[161,1426,200,1865]" captionTargetId="figure-31@4.[151,1436,190,1875]" captionTargetPageId="4" captionText="FIGURE 2. Distribution map of Epidendrum clinidiodontum E.Santiago, López-Flor., Claros &amp; Hágsater in Florencia (Caquetá), Gachalá and Guayabetal (Cundinamarca), Colombia." figureDoi="http://doi.org/10.5281/zenodo.14519261" httpUri="https://zenodo.org/record/14519261/files/figure.png" pageId="3" pageNumber="282">Figure 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8DFB65600B711E2D959C9F78690F2588" pageId="3" pageNumber="282" type="etymology">
<paragraph id="C55E36EB0B711E2D959C9F78690F2588" blockId="3.[136,1452,158,1841]" pageId="3" pageNumber="282">
<emphasis id="F795EAF90B711E2D959C9F786B702530" bold="true" box="[189,328,1058,1084]" pageId="3" pageNumber="282">Etymology:</emphasis>
—From the Greek κλινη bed, the cavity in which the anther cap is situated, at the apex of the column, δυο, two, and οσοντος, tooth. In reference to the two triangular teeth at the apex of the clinandrium
<emphasis id="F795EAF90B711E2D903A9F1D6F1C256C" box="[1307,1316,1095,1120]" italics="true" pageId="3" pageNumber="282">-</emphasis>
hood which distinguish this species from most other
<taxonomicName id="02E14D680B711E2D976C9F30690F2588" box="[589,823,1130,1156]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="282" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="F795EAF90B711E2D976C9F3068EF2588" box="[589,727,1130,1156]" italics="true" pageId="3" pageNumber="282">Epidendrum</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="8DFB65600B711E2D959C9FD4688925A4" box="[189,689,1166,1193]" pageId="3" pageNumber="282" type="conservation">
<paragraph id="C55E36EB0B711E2D959C9FD4688925A4" blockId="3.[136,1452,158,1841]" box="[189,689,1166,1193]" pageId="3" pageNumber="282">
<emphasis id="F795EAF90B711E2D959C9FD46B8125A4" bold="true" box="[189,441,1166,1192]" pageId="3" pageNumber="282">Conservation Status:</emphasis>
—DD. Data deficient.
</paragraph>
</subSubSection>
<subSubSection id="8DFB65600B711E2D959C9FE86F022434" pageId="3" pageNumber="282" type="materials_examined">
<paragraph id="C55E36EB0B711E2D959C9FE86F022434" blockId="3.[136,1452,158,1841]" pageId="3" pageNumber="282">
<materialsCitation id="75893CB60B711E2D959C9FE869D225FC" collectingDate="2023-05-07" collectingDateMax="2023-05-07" collectingDateMin="2019-09-24" collectionCode="HUAZ, AMO" collectorName="Arias" country="Colombia" county="Doncello" elevation="2570" location="Vda. Las Acacias: camino al centro poblado Berlin" municipality="Doncello" pageId="3" pageNumber="282" specimenCount="1" stateProvince="Caqueta" typeStatus="paratype">
<emphasis id="F795EAF90B711E2D959C9FE868F025C0" bold="true" box="[189,712,1202,1228]" pageId="3" pageNumber="282">
Additional specimens examined (
<typeStatus id="1A5A88490B711E2D97689FE8688325C0" box="[585,699,1202,1228]" pageId="3" pageNumber="282" type="paratype">paratype</typeStatus>
):
</emphasis>
—
<collectingCountry id="BDF6767B0B711E2D97C49FE8694525C0" box="[741,893,1202,1228]" name="Colombia" pageId="3" pageNumber="282">COLOMBIA</collectingCountry>
.
<collectingRegion id="0725F8090B711E2D96A99FE869D025C0" box="[904,1000,1202,1228]" country="Colombia" name="Caqueta" pageId="3" pageNumber="282">Caquetá</collectingRegion>
:
<collectingMunicipality id="253AAC910B711E2D96D29FE86E6625C0" box="[1011,1118,1202,1228]" pageId="3" pageNumber="282">Doncello</collectingMunicipality>
:
<location id="C03E60300B711E2D91489FE86B4525FC" LSID="urn:lsid:plazi:treatment:4D4887FD0B731E2D95A99DAE6EFA263C:C03E60300B711E2D91489FE86B4525FC" country="Colombia" county="Doncello" municipality="Doncello" name="Vda. Las Acacias: camino al centro poblado Berlin" pageId="3" pageNumber="282" stateProvince="Caqueta">Vda. Las Acacias: camino al centro poblado Berlín</location>
,
<date id="B15F102B0B711E2D94A69F8D683725FC" box="[391,527,1238,1265]" pageId="3" pageNumber="282" value="2023-05-07">
<collectingDate id="A11BE9C30B711E2D94A69F8D683725FC" box="[391,527,1238,1265]" pageId="3" pageNumber="282" value="2023-05-07">7 May 2023</collectingDate>
</date>
,
<emphasis id="F795EAF90B711E2D97389F8D68BE25FC" box="[537,646,1238,1265]" italics="true" pageId="3" pageNumber="282">
<collectorName id="6814533D0B711E2D97389F8D686E25FD" box="[537,598,1239,1265]" pageId="3" pageNumber="282">Arias</collectorName>
960
</emphasis>
,
<collectionCode id="A3F0AE2E0B711E2D97B29F8C68DF25FC" box="[659,743,1238,1264]" collectionName="HUAZ" pageId="3" pageNumber="282">HUAZ</collectionCode>
! digital images,
<collectionCode id="A3F0AE2E0B711E2D96B99F8C69DB25FC" box="[920,995,1238,1264]" pageId="3" pageNumber="282">AMO</collectionCode>
!.
</materialsCitation>
<materialsCitation id="75893CB60B711E2D96D19F8C6F2E2418" collectingDate="2019-09-24" collectingDateMax="2023-05-07" collectingDateMin="2019-09-24" collectionCode="COL" collectorName="Hernandez-Rodriguez" country="Colombia" county="Doncello" elevation="2570" location="Vda. Tendidos del rio Negro: sector del Alto de Siberia" municipality="Gachala" pageId="3" pageNumber="282" specimenCount="1" stateProvince="Cundinamarca">
<collectingRegion id="0725F8090B711E2D96D19F8C6EA025FC" box="[1008,1176,1238,1264]" country="Colombia" name="Cundinamarca" pageId="3" pageNumber="282">Cundinamarca</collectingRegion>
:
<collectingMunicipality id="253AAC910B711E2D91809F8C6F3925FC" box="[1185,1281,1238,1264]" pageId="3" pageNumber="282">Gachalá</collectingMunicipality>
:
<location id="C03E60300B711E2D902B9F8C687E2418" LSID="urn:lsid:plazi:treatment:4D4887FD0B731E2D95A99DAE6EFA263C:C03E60300B711E2D902B9F8C687E2418" country="Colombia" county="Doncello" municipality="Gachala" name="Vda. Tendidos del rio Negro: sector del Alto de Siberia" pageId="3" pageNumber="282" stateProvince="Cundinamarca">Vda. Tendidos del río Negro: sector del Alto de Siberia</location>
,
<quantity id="02199B0E0B711E2D97709FA068922419" box="[593,682,1274,1301]" metricMagnitude="3" metricUnit="m" metricValue="2.57" pageId="3" pageNumber="282" unit="m" value="2570.0">
<elevation id="4ECCD1D80B711E2D97709FA068922419" box="[593,682,1274,1301]" metricMagnitude="3" metricUnit="m" metricValue="2.57" pageId="3" pageNumber="282" unit="m" value="2570.0">2570 m</elevation>
</quantity>
.,
<date id="B15F102B0B711E2D97989FA069AD2418" box="[697,917,1274,1300]" pageId="3" pageNumber="282" value="2019-09-24">
<collectingDate id="A11BE9C30B711E2D97989FA069AD2418" box="[697,917,1274,1300]" pageId="3" pageNumber="282" value="2019-09-24">24 September 2019</collectingDate>
</date>
,
<emphasis id="F795EAF90B711E2D96BE9FA06EFF2418" box="[927,1223,1274,1300]" italics="true" pageId="3" pageNumber="282">
<collectorName id="6814533D0B711E2D96BE9FA06EAE2418" box="[927,1174,1274,1300]" pageId="3" pageNumber="282">Hernández-Rodríguez</collectorName>
233
</emphasis>
,
<collectionCode id="A3F0AE2E0B711E2D91F49FA06F2E2418" box="[1237,1302,1274,1300]" collectionName="COL" pageId="3" pageNumber="282">COL</collectionCode>
</materialsCitation>
!;
<materialsCitation id="75893CB60B711E2D90059FA06F0E2434" collectingDate="2019-01-28" collectionCode="AMO" collectorName="Parra" country="Colombia" county="Guayabetal" elevation="2387" location="Vereda Limoncitos, Finca Casa Roja" municipality="Guayabetal" pageId="3" pageNumber="282" specimenCount="1">
<collectingMunicipality id="253AAC910B711E2D90059FA06F902418" box="[1316,1448,1274,1300]" pageId="3" pageNumber="282">Guayabetal</collectingMunicipality>
:
<location id="C03E60300B711E2D95A99E44681D2434" LSID="urn:lsid:plazi:treatment:4D4887FD0B731E2D95A99DAE6EFA263C:C03E60300B711E2D95A99E44681D2434" box="[136,549,1310,1336]" country="Colombia" county="Guayabetal" municipality="Guayabetal" name="Vereda Limoncitos, Finca Casa Roja" pageId="3" pageNumber="282">Vereda Limoncitos, Finca Casa Roja</location>
,
<quantity id="02199B0E0B711E2D97119E4468BF2435" box="[560,647,1310,1337]" metricMagnitude="3" metricUnit="m" metricValue="2.3869999999999996" pageId="3" pageNumber="282" unit="m" value="2387.0">
<elevation id="4ECCD1D80B711E2D97119E4468BF2435" box="[560,647,1310,1337]" metricMagnitude="3" metricUnit="m" metricValue="2.3869999999999996" pageId="3" pageNumber="282" unit="m" value="2387.0">2387 m</elevation>
</quantity>
, received
<date id="B15F102B0B711E2D97D89E44698D2434" box="[761,949,1310,1337]" pageId="3" pageNumber="282" value="2019-01-28">
<collectingDate id="A11BE9C30B711E2D97D89E44698D2434" box="[761,949,1310,1337]" pageId="3" pageNumber="282" value="2019-01-28">28 January 2019</collectingDate>
</date>
,
<emphasis id="F795EAF90B711E2D96E19E456E082434" box="[960,1072,1311,1336]" italics="true" pageId="3" pageNumber="282">
<collectorName id="6814533D0B711E2D96E19E456E3B2434" box="[960,1027,1311,1336]" pageId="3" pageNumber="282">Parra</collectorName>
s.n.
</emphasis>
, Digital images,
<collectionCode id="A3F0AE2E0B711E2D91CE9E446F0E2434" box="[1263,1334,1310,1336]" pageId="3" pageNumber="282">AMO</collectionCode>
</materialsCitation>
!
</paragraph>
</subSubSection>
<subSubSection id="8DFB65600B711E2D959C9E186EFA263C" pageId="3" pageNumber="282" type="discussion">
<paragraph id="C55E36EB0B711E2D959C9E186EFA263C" blockId="3.[136,1452,158,1841]" pageId="3" pageNumber="282">
<emphasis id="F795EAF90B711E2D959C9E186BF52450" bold="true" box="[189,461,1346,1372]" pageId="3" pageNumber="282">Taxonomic discussion:</emphasis>
—
<taxonomicName id="02E14D680B711E2D94C89E18691F2450" box="[489,807,1346,1372]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="282" phylum="Tracheophyta" rank="species" species="clinidiodontum">
<emphasis id="F795EAF90B711E2D94C89E18691F2450" box="[489,807,1346,1372]" italics="true" pageId="3" pageNumber="282">Epidendrum clinidiodontum</emphasis>
</taxonomicName>
belongs to the Albertii group which is characterized by the sympodial habit, the laterally compressed to ancipitose or somewhat fusiform
<emphasis id="F795EAF90B711E2D91009E3D6E12248C" box="[1057,1066,1383,1408]" italics="true" pageId="3" pageNumber="282">-</emphasis>
thickened stems, the apical and/or lateral, distichous inflorescences, more or less with imbricating bracts on the peduncle, producing one flower at a time, and the Ellipsophyllum subgroup, which has generally apical inflorescences, the apex of leaves acuminate, and the clinandrium-hood elongate, margin entire to dentate. The new species is recognized by the short apical inflorescences with a two
<emphasis id="F795EAF90B711E2D95DE9EAD6B30271C" box="[255,264,1527,1552]" italics="true" pageId="3" pageNumber="282">-</emphasis>
winged, short peduncle, the two prominent spathes at the apex of peduncle, the leaves 7.2–12.5 ×
<quantity id="02199B0E0B711E2D90769EAC6A952739" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="3" pageNumber="282" unit="cm" value="2.05" valueMax="2.3" valueMin="1.8">1.8–2.3 cm</quantity>
, narrowly elliptic, apex acuminate, the flowers pale green and lip cream colored, the sepals
<quantity id="02199B0E0B711E2D91809D406F202739" box="[1185,1304,1562,1589]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="3" pageNumber="282" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">15–16 mm</quantity>
long, margin revolute; the petals 14.5 ×
<quantity id="02199B0E0B711E2D94E09D6468242755" box="[449,540,1598,1625]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="282" unit="mm" value="1.8">1.8 mm</quantity>
, sub-parallel to column, linear-oblong, the lip 9.0 ×
<quantity id="02199B0E0B711E2D915F9D646EDF2755" box="[1150,1255,1598,1625]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="282" unit="mm" value="10.5">10.5 mm</quantity>
, slightly convex, pentagonal with hyaline margin, bicallose, calli laminar,, leaning outwards and the column with a bidentate apex. The flowers are similar to
<taxonomicName id="02E14D680B711E2D94A99DDC68E127AC" authorityName="Hagsater &amp; E. Santiago" authorityYear="2004" box="[392,729,1670,1696]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="282" phylum="Tracheophyta" rank="species" species="hygrohylephilum">
<emphasis id="F795EAF90B711E2D94A99DDC68E127AC" box="[392,729,1670,1696]" italics="true" pageId="3" pageNumber="282">Epidendrum hygrohylephilum</emphasis>
</taxonomicName>
but that species has the callus triangular, acute, that continues into a thin, long rib reaching apical sinus, the lip 7.0–10.0 × 7.0–
<quantity id="02199B0E0B711E2D96509DF069F227C9" box="[881,970,1706,1733]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="3" pageNumber="282" unit="mm" value="7.3">7.3 mm</quantity>
, united to column, entire, basal half semi-elliptic, apical half triangular, acute, margin entire and clinandrium-hood with entire margin.
<taxonomicName id="02E14D680B711E2D91E59D946AE92600" authority="Schltr." authorityName="Schltr." class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="282" phylum="Tracheophyta" rank="species" species="albertii">
<emphasis id="F795EAF90B711E2D91E59D946F9327E4" box="[1220,1451,1742,1768]" italics="true" pageId="3" pageNumber="282">Epidendrum albertii</emphasis>
Schltr.
</taxonomicName>
(Schlechter 1922: 208;
<bibRefCitation id="A1704B1A0B711E2D94C19DA869352600" author="Santiago, E. &amp; Hagsater, E." box="[480,781,1778,1804]" pageId="3" pageNumber="282" refId="ref3333" refString="Santiago, E. &amp; Hagsater, E. (2007) In: Hagsater, E. &amp; Sanchez, S. L. (Eds.) The Genus Epidendrum, Part 6. Icones Orchidacearum 9: t. 901 - 1000." type="journal volume" year="2007">Santiago &amp; Hágsater 2007</bibRefCitation>
) has elliptic
<emphasis id="F795EAF90B711E2D96B69DA969982600" box="[919,928,1779,1804]" italics="true" pageId="3" pageNumber="282">-</emphasis>
oblanceolate leaves 2.8–7.7 × 2.0–
<quantity id="02199B0E0B711E2D90099DA86F4F2601" box="[1320,1399,1778,1805]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="282" unit="cm" value="3.9">3.9 cm</quantity>
, and the lip is ovate
<emphasis id="F795EAF90B711E2D940D9C4D6B0D263C" box="[300,309,1815,1840]" italics="true" pageId="3" pageNumber="282">-</emphasis>
oblong with an acute apex and the column strongly arched at the apex (
<figureCitation id="5DDA2A6E0B711E2D91739C4C6E8C263C" box="[1106,1204,1814,1840]" captionStart="FIGURE 3" captionStartId="5.[136,229,1952,1974]" captionTargetBox="[215,1373,202,1916]" captionTargetId="figure-22@5.[202,1386,190,1928]" captionTargetPageId="5" captionText="FIGURE 3. A–B. Epidendrum clinidiodontum, C–D. Epidendrum hygrohylephilum and E–F. Epidendrum albertii. Photographs by Oscar M. López-Floriano, Guillermo A. Reina-Rodríguez and Hugo Medina." figureDoi="http://doi.org/10.5281/zenodo.14519263" httpUri="https://zenodo.org/record/14519263/files/figure.png" pageId="3" pageNumber="282">Figure 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>