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<document id="EC58F103EDF5B60CB6ECB1DCC6482A95" ID-CLB-Dataset="33710" ID-DOI="http://dx.doi.org/10.3897/dez.62.8975" ID-GBIF-Dataset="0c22d54b-65f8-4b75-86ad-1f2e51e873ea" ID-Pensoft-Pub="1860-1324-1-81" ID-ZooBank="CBE1F1254B914A5F8260554A7C453B57" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1860-1324-1-81" ModsDocOrigin="Deutsche Entomologische Zeitschrift 62 (1)" ModsDocTitle="A revision of the genus Chlorocryptus Cameron (Hymenoptera, Ichneumonidae), with the first record of the genus from Japan" checkinTime="1433260402103" checkinUser="pensoft" docAuthor="Yoshida, Takuma &amp; Matsumoto, Rikio" docDate="2015" docId="D0B926F3F891A92BC5CFFC44C5409ECE" docLanguage="en" docName="DeutEntomolZeit 62(1): 81-99" docOrigin="Deutsche Entomologische Zeitschrift 62 (1)" docSource="http://dx.doi.org/10.3897/dez.62.8975" docTitle="Chlorocryptus coreanus" docType="treatment" docVersion="6" lastPageNumber="89" masterDocId="FFC7C411FF8AFF9F6C16FF92FFC7FFFE" masterDocTitle="A revision of the genus Chlorocryptus Cameron (Hymenoptera, Ichneumonidae), with the first record of the genus from Japan" masterLastPageNumber="99" masterPageNumber="81" pageNumber="84" updateTime="1701382121096" updateUser="ExternalLinkService">
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<mods:title id="4CF098DDDD3B1682B2A903022774D142">A revision of the genus Chlorocryptus Cameron (Hymenoptera, Ichneumonidae), with the first record of the genus from Japan</mods:title>
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<treatment id="D0B926F3F891A92BC5CFFC44C5409ECE" ID-GBIF-Taxon="127897300" LSID="urn:lsid:plazi:treatment:D0B926F3F891A92BC5CFFC44C5409ECE" httpUri="http://treatment.plazi.org/id/D0B926F3F891A92BC5CFFC44C5409ECE" lastPageId="8" lastPageNumber="89" pageId="3" pageNumber="84">
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<paragraph id="495B0109EB4974BE7BDD4B33D6541BEB" pageId="3" pageNumber="84">Taxon classification Animalia Hymenoptera Ichneumonidae</paragraph>
</subSubSection>
<subSubSection id="E017E07F8143FCB02A01E85A0FB0E65B" pageId="3" pageNumber="84" type="nomenclature">
<paragraph id="DCC4F70D865AD6658351C0AA3817CBBE" pageId="3" pageNumber="84">
<taxonomicName id="5A43E169655BC2D0F426BA2C697B4DCE" ID-CoL="69Y4P" authority="Szepligeti" class="Insecta" family="Ichneumonidae" genus="Chlorocryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlorocryptus coreanus" order="Hymenoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="coreanus">
Chlorocryptus coreanus (
<normalizedToken id="3EAA472D43A26E6257D351767F7034E0" originalValue="Szépligeti">Szepligeti</normalizedToken>
)
</taxonomicName>
Figs 13-21, 26-27, 36-38, 59
</paragraph>
</subSubSection>
<subSubSection id="C61D63945C79064691A09C61FA03B478" pageId="4" pageNumber="85" type="reference_group">
<paragraph id="1EF11ACF0A238DEBC85DE79559EF76B9" pageId="4" pageNumber="85">
<taxonomicName id="0BC3DD53AA1D3D4E40E9732459699C74" class="Gastropoda" family="Cryptaulacidae" genus="Cryptaulax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptaulax coreanus" pageId="4" pageNumber="85" phylum="Mollusca" rank="species" species="coreanus">
<pageBreakToken id="0A452CFAC20417353F7BDF9F912980E0" pageId="4" pageNumber="85" start="start">Cryptaulax</pageBreakToken>
coreanus
</taxonomicName>
<normalizedToken id="B130506173A18C004FA6980301F00446" originalValue="Szépligeti">Szepligeti</normalizedToken>
, 1916: 287. Holotype: ♀, Korea, Seoul (HNHM) [examined].
</paragraph>
<paragraph id="3D1EC2B7C08D4E3CAA5057A23926CD54" pageId="4" pageNumber="85">
<taxonomicName id="590A566508228742F223FF08299B2D50" class="Insecta" family="Ichneumonidae" genus="Cryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptus trirrhogmaniformis" order="Hymenoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="trirrhogmaniformis">Cryptus trirrhogmaniformis</taxonomicName>
Sonan, 1929: 424. Lectotype: ♂, Formosa, Chikurin (TARI). Synonymized by
<bibRefCitation id="CB746DFAB413E688E18BFD77C0B9BC0D" author="Uchida, T" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido Imperial University" pageId="10" pageNumber="91" pagination="349 - 376" title="Beitrag zur Ichneumoniden-Fauna Japans." url="http://hdl.handle.net/2115/12653" volume="25" year="1930 b">Uchida (1930b)</bibRefCitation>
.
</paragraph>
<paragraph id="EA212393ACD4FAA5A68B8CF829F9D60A" pageId="4" pageNumber="85">
<taxonomicName id="F822B2ED6A536E509117FEE44E99D2B8" class="Insecta" genus="Cochlidionostenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cochlidionostenus coreanus" order="Hymenoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="coreanus">Cochlidionostenus coreanus</taxonomicName>
(
<normalizedToken id="5859CFC7CA39B7C66EAAE5620E4DBE06" originalValue="Szépligeti">Szepligeti</normalizedToken>
):
<bibRefCitation id="0262BEE58A38E9510FA6E6D40C644B87" author="Uchida, T" journalOrPublisher="Insecta Matsumurana" pageId="10" pageNumber="91" pagination="111 - 122" title="Drei neue Gattungen sowie acht neue und fuenf unbeschriebene Arten der Ichneumoniden aus Japan." url="http://hdl.handle.net/2115/9323" volume="10" year="1936">Uchida 1936</bibRefCitation>
: 116.
</paragraph>
<paragraph id="839F49CB5ADE3F087540BDB349F1A8D9" pageId="4" pageNumber="85">
<taxonomicName id="60C95DD077A99D241A102F6C879EE4C0" class="Insecta" family="Ichneumonidae" genus="Chlorocryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlorocryptus coreanus" order="Hymenoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="coreanus">Chlorocryptus coreanus</taxonomicName>
(
<normalizedToken id="C9155ACBAA882027A8B8F4395983F6C0" originalValue="Szépligeti">Szepligeti</normalizedToken>
):
<bibRefCitation id="486567A070CD4F7836264591F503FE95" author="Townes, HK" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="10" pageNumber="91" pagination="1 - 522" title="A catalogue and reclassification of the Indo-Australian Ichneumonidae." volume="1" year="1961">Townes et al. 1961</bibRefCitation>
: 141.
</paragraph>
</subSubSection>
<subSubSection id="28D393ED5A62E8CB18824B13A300E979" pageId="4" pageNumber="85" type="references">
<paragraph id="C3F5B926DE79AB2D94817841FE1D1ECF" pageId="4" pageNumber="85">References.</paragraph>
<paragraph id="42DC77D05111F9105625030DBD519490" pageId="4" pageNumber="85">
<bibRefCitation id="A7CF41B2AFCE1EA510396B640B2423A7" author="Uchida, T" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido Imperial University" pageId="10" pageNumber="91" pagination="299 - 347" title="Fuenfter Beitrag zur Ichneumoniden-Fauna Japans." url="http://hdl.handle.net/2115/12652" volume="25" year="1930 a">Uchida 1930a</bibRefCitation>
: 317 (
<taxonomicName id="4110C96711AC67319BE2AD9C64CC73C1" class="Gastropoda" family="Cryptaulacidae" genus="Cryptaulax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptaulax" pageId="4" pageNumber="85" phylum="Mollusca" rank="genus">Cryptaulax</taxonomicName>
, records, host);
<bibRefCitation id="308453EE7383CE1ED1926CC189D1EC5A" author="Uchida, T" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido Imperial University" pageId="10" pageNumber="91" pagination="349 - 376" title="Beitrag zur Ichneumoniden-Fauna Japans." url="http://hdl.handle.net/2115/12653" volume="25" year="1930 b">Uchida 1930b</bibRefCitation>
: 360 (
<taxonomicName id="D142EB9038E9DB52CDD9A8833B6D1831" class="Gastropoda" family="Cryptaulacidae" genus="Cryptaulax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptaulax" pageId="4" pageNumber="85" phylum="Mollusca" rank="genus">Cryptaulax</taxonomicName>
, hosts);
<bibRefCitation id="16DA6A35881DCFB3F7D83B2CD6986DF0" author="Uchida, T" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido Imperial University" pageId="10" pageNumber="91" pagination="163 - 193" title="Beitrag zur Kenntnis der Cryptinenfauna Formosas." url="http://hdl.handle.net/2115/12678" volume="30" year="1931">Uchida 1931</bibRefCitation>
: 177 (
<taxonomicName id="E6B355B7DF2288EE44469B9122D9EBE2" class="Gastropoda" family="Cryptaulacidae" genus="Cryptaulax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptaulax" pageId="4" pageNumber="85" phylum="Mollusca" rank="genus">Cryptaulax</taxonomicName>
, record, host);
<bibRefCitation id="C8E38C2183757B200EE4C776812711B1" author="Matsumura, S" journalOrPublisher="Toko Shoin, Tokyo" pageId="9" pageNumber="90" year="1931">Matsumura 1931</bibRefCitation>
: 41 (
<taxonomicName id="30D960F4E3F9FB36A11E5CAB4681F932" class="Gastropoda" family="Cryptaulacidae" genus="Cryptaulax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptaulax" pageId="4" pageNumber="85" phylum="Mollusca" rank="genus">Cryptaulax</taxonomicName>
, diagnosis, figure, distribution);
<bibRefCitation id="4981C3ECFBF02E79D67563C6153139B8" author="Uchida, T" journalOrPublisher="Botany and Zoology" pageId="10" pageNumber="91" pagination="259 - 264" volume="8" year="1940">Uchida 1940</bibRefCitation>
: 122 (
<taxonomicName id="C255B79C6C03FFA2A887AC9F06CE2FDA" class="Insecta" genus="Cochlidionostenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cochlidionostenus" order="Hymenoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="genus">Cochlidionostenus</taxonomicName>
, in key);
<bibRefCitation id="41B790DBA3B4D651BEC89E7256C48A63" author="Uchida, T" journalOrPublisher="Insecta Matsumurana" pageId="10" pageNumber="91" pagination="107 - 146" title="Ichneumoniden Mandschukuos aus dem Entomologischen Museum der kaiserlichen Hokkaido Universitaet." url="http://hdl.handle.net/2115/9491" volume="16" year="1942">Uchida 1942</bibRefCitation>
: 113 (
<taxonomicName id="E284F84204024C76C9273A1CC0F8BC3D" class="Insecta" genus="Cochlidionostenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cochlidionostenus" order="Hymenoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="genus">Cochlidionostenus</taxonomicName>
, records);
<bibRefCitation id="A7B5C87B20E17DFB7D5F122A661EAA69" author="Sonan, J" journalOrPublisher="Report, Government Research Institute, Department of Agriculture, Formosa" pageId="10" pageNumber="91" pagination="1 - 77" title="A catalogue of parasitic Hymenoptera and their hosts in Taiwan." volume="222" year="1944">Sonan 1944</bibRefCitation>
: 14 (
<taxonomicName id="49C62495A60E5C5B31A996F54350A255" class="Insecta" family="Ichneumonidae" genus="Cryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptus trirrhogmaniformis" order="Hymenoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="trirrhogmaniformis">Cryptus trirrhogmaniformis</taxonomicName>
, listed);
<bibRefCitation id="C773730D8FBB860E527496C5BCFE832A" author="Uchida, T" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido University" pageId="10" pageNumber="91" pagination="95 - 133" title="Die von Dr. K. Tsuneki in Korea gesammelten Ichneumoniden." url="http://hdl.handle.net/2115/12759" volume="50" year="1955">Uchida 1955</bibRefCitation>
: 114 (
<taxonomicName id="974C6D34E2E4B192787A47F68E322672" class="Insecta" genus="Cochlidionostenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cochlidionostenus" order="Hymenoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="genus">Cochlidionostenus</taxonomicName>
, records);
<bibRefCitation id="90905A6F18269723EA94CDDB3835B3E9" pageId="4" pageNumber="85">Kim 1955</bibRefCitation>
: 40 (
<taxonomicName id="8B0391212549A2AD4B1993BA93EF4D30" genus="Cochleonosterus" lsidName="Cochleonosterus" pageId="4" pageNumber="85" rank="genus">Cochleonosterus</taxonomicName>
(lapsus), record);
<bibRefCitation id="B628EE27F50678D8FEDF2782EDD80D81" author="Townes, HK" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="10" pageNumber="91" pagination="1 - 522" title="A catalogue and reclassification of the Indo-Australian Ichneumonidae." volume="1" year="1961">Townes et al. 1961</bibRefCitation>
: 141 (catalogued);
<bibRefCitation id="3FB1FCB8E44406B2B2899E9317DC19BC" author="Chu, Joo-Tso" journalOrPublisher="Shanghai Science and Technology Press, Shanghai" pageId="9" pageNumber="90" year="1963">Chu et al. 1963</bibRefCitation>
: 750 (
<taxonomicName id="C33EF82A532DED6DC188F345DB430EB4" class="Gastropoda" family="Cryptaulacidae" genus="Cryptaulax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptaulax coteanus" pageId="4" pageNumber="85" phylum="Mollusca" rank="species" species="coteanus">Cryptaulax coteanus</taxonomicName>
(lapsus), as parasitoid of
<taxonomicName id="510E1600BD061AD7D0810EC070A12F98" family="Limacodidae" lsidName="" pageId="4" pageNumber="85" rank="family">Limacodidae</taxonomicName>
);
<bibRefCitation id="8900AD3D1D5CA0269555E5C3B76622DF" author="Townes, HK" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="10" pageNumber="91" pagination="1 - 661" title="A catalogue and reclassification of the eastern Palearctic Ichneumonidae." volume="5" year="1965">Townes et al. 1965</bibRefCitation>
: 167 (catalogued);
<bibRefCitation id="2302A9E9F2CDFD071E9BF41C29137C62" author="Yen, DF" journalOrPublisher="Department of Plant Pathology and Entomology, College of Agriculture, National Taiwan University, Taipei" pageId="11" pageNumber="92" title="A natural enemy list of insects of Taiwan." year="1973">Yen 1973</bibRefCitation>
: 86 (
<taxonomicName id="0A9A8959D29A98F2EAE52A4463D835E6" class="Insecta" family="Ichneumonidae" genus="Cryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptus trirrhogmaniformis" order="Hymenoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="trirrhogmaniformis">Cryptus trirrhogmaniformis</taxonomicName>
, listed, host);
<bibRefCitation id="081CA7969F960CA0C44BBC72CEF88769" author="Chao, Hsiu-Fu" journalOrPublisher="Scientific Publisher, Beijing" pageId="9" pageNumber="90" title="An outline of the classification of the Ichneumon-flies of China (Hymenoptera: Ichneumonidae)." year="1976">Chao 1976</bibRefCitation>
: 266 (listed, distribution, hosts);
<bibRefCitation id="8A61D8AB0D9B2FB0721DE404F3AEF905" author="Chu, Joo-Tso" journalOrPublisher="Scientific Publisher, Beijing, China" pageId="9" pageNumber="90" pagination="32 - 49" title="Ichneumonidae." volumeTitle="Atlas of Natural Enemies of Economic Insects" year="1978">Chu et al. 1978</bibRefCitation>
: 40 (diagnosis, distribution, host);
<bibRefCitation id="C00AB1F9A6B3A2AFCFFFC0B6D2848C9A" author="Yang, Xiu-Yuan" journalOrPublisher="Chinese Forestry Publishing House, Beijing" pageId="11" pageNumber="92" title="A check list of the forest insects of China." year="1981">Yang and Wu 1981</bibRefCitation>
: 303 (listed, distribution, host);
<bibRefCitation id="A87A8CAEC134BAED5E2320B794C504DD" author="Sun, Xue-Yi" journalOrPublisher="Liaoning Forestry Science and Technology" pageId="10" pageNumber="91" pagination="46 - 48" volume="1982" year="1982">Sun et al. 1982</bibRefCitation>
: 46 (description, immature stages, host, biology);
<bibRefCitation id="4026DDDC4AECB2747F1CF14A405CBC6B" author="Pan, Lang" journalOrPublisher="Liaoning Agricultural Sciences" pageId="10" pageNumber="91" pagination="50 - 51" volume="1983" year="1983">Pan 1983</bibRefCitation>
: 50 (
<taxonomicName id="A1C791FD5E527850A3D5EB26FA63BA8B" class="Insecta" family="Ichneumonidae" genus="Chlorocryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlorocryptus coevanus" order="Hymenoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="coevanus">coevanus</taxonomicName>
(lapsus), description, figures, host, biology);
<bibRefCitation id="AAF5E8AAA9B29E59B803E41EFEA3AD57" author="Chiu, Shui-Chen" journalOrPublisher="Special Publication" pageId="9" pageNumber="90" pagination="1 - 67" title="A check list of Ichneumonidae (Hymenoptera) of Taiwan. Taiwan Agricultural Research Institute." url="http://ir.tari.gov.tw:8080/handle/345210000/4741" volume="15" year="1984">Chiu et al. 1984</bibRefCitation>
: 17 (listed, distribution, hosts);
<bibRefCitation id="3B6AF809A75DDCFEE94CF76E4451B279" author="Yin, Ping-Fang" journalOrPublisher="Journal of Jilin Forestry Institute" pageId="11" pageNumber="92" pagination="11 - 20" title="A study on the efficiency of some natural enemies to control the community density of oriental moth." volume="3" year="1987">Yin and Zou 1987</bibRefCitation>
: 11 (diagnosis, immature stages, biology, biological control effect on
<taxonomicName id="F0CCFCAF4BF437DECE39C056618C5A85" class="Insecta" family="Limacodidae" genus="Cnidocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cnidocampa flavescens" order="Lepidoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="flavescens">Cnidocampa flavescens</taxonomicName>
);
<bibRefCitation id="3F3D5EB3FBD1F97A120B971FECB0600D" author="He, Jun-Hua" journalOrPublisher="Agricultural Publishing House, Beijing" pageId="9" pageNumber="90" pagination="367 - 400" title="Ichneumonidae." volumeTitle="Agriculture Insects of China" year="1987">He and Wang 1987</bibRefCitation>
: 379 (diagnosis, host, distribution);
<bibRefCitation id="414706BEC3E4E078D3360B009954C0B9" author="Wang, Shu-Fang" editor="Huang, Fu-Sheng" journalOrPublisher="Yunnan Science and Technology Press, Kunming" pageId="11" pageNumber="92" pagination="1317 - 1336" title="Ichneumonidae." volumeTitle="Forest Insects of Yunnan" year="1987">Wang and Li 1987</bibRefCitation>
: 1326 (in key, record, host, distribution);
<bibRefCitation id="DF3B670CA147AA8CEB657BCC9165C72C" author="Gauld, ID" editor="Cook, MJW" journalOrPublisher="CAB International, Wallingford" pageId="9" pageNumber="90" pagination="123 - 138" title="The Ichneumonid parasitoids associated with South-east Asian Limacodidae." volumeTitle="Slug and Nettle Caterpillars: the Biology, Taxonomy and Control of the Limacodidae of Economic Importance on Palms in Southeast Asia" year="1987">Gauld 1987</bibRefCitation>
: 130 (taxonomic remarks);
<bibRefCitation id="7C3AD389D509C7A4BC2B96FB31D7A448" author="Wang, Fu-Wei" journalOrPublisher="Jilin Forestry Science and Technology" pageId="11" pageNumber="92" pagination="30 - 32" volume="1991" year="1991">Wang et al. 1991</bibRefCitation>
: 30 (percentage of parasitism, biological control effect on
<taxonomicName id="B13B94FDE2CFE521F0B8C14EFD55F447" class="Insecta" family="Limacodidae" genus="Cnidocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cnidocampa flavescens" order="Lepidoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="flavescens">Cnidocampa flavescens</taxonomicName>
);
<bibRefCitation id="9FB5816AB152CEA9192102A4D4FEDE88" author="Niu, Yan-Zhang" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="10" pageNumber="91" title="Chlorocryptuscoreanus Szepligeti. In: Xiao Gang-Rou (Ed.). Forest Insects of China" year="1992">Niu and Wang 1992</bibRefCitation>
: 1206 (description, figure, distribution, host, biology);
<bibRefCitation id="A289D5D549BC1E6C182594E19362F02E" author="Wang, Fu-Wei" journalOrPublisher="Forest Research" pageId="10" pageNumber="91" pagination="466 - 469" title="Study on the natural enemy complex of oriental moth." volume="6" year="1993">Wang et al. 1993</bibRefCitation>
: 467 (rate of parasitism, life history, biological control effect);
<bibRefCitation id="C2BB416BA29CDE0DAAF61FD167DADCF4" author="Wang, Shug-Fang" editor="Huang, Chun-Mei" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="11" pageNumber="92" pagination="727 - 738" title="Hymenoptera: Ichneumonidae." volumeTitle="Animals of Longqi Mountain" year="1993">Wang and Huang 1993</bibRefCitation>
: 732 (description, figure, host, records, distribution);
<bibRefCitation id="2BA6D045E104029B11D28164C44D97A0" author="He, Jun-Hua" journalOrPublisher="Science Press, Beijing" pageId="9" pageNumber="90" title="Hymenoptera: Ichneumonidae. Economic Insect Fauna of China." year="1996">He et al. 1996</bibRefCitation>
: 510 (description, figure, host, distribution);
<bibRefCitation id="DAAB5AFAFFFAA4D8C8C3AAE5C411FA62" author="Sheng, Mao-Ling" editor="Shen, Xia-Cheng" journalOrPublisher="China Agricultural Scientech Press, Beijing" pageId="10" pageNumber="91" pagination="373 - 379" title="Hymenoptera: Ichneumonidae." volumeTitle="The Fauna and Taxonomy of Insects in Henan Vol. 4. Insects of the Mountains Funiu and Dabie Regions" year="1999">Sheng et al. 1999</bibRefCitation>
: 374 (listed, records, distribution);
<bibRefCitation id="9B17F134B0193654A9D4A3F20C807FF1" author="He, Jun-Hua" editor="Wu, Hong" journalOrPublisher="Science Press, Beijing" pageId="9" pageNumber="90" pagination="710 - 723" title="Hymenoptera: Ichneumonidae." volumeTitle="Insects of Tianmushan National Nature Reserve" year="2001">He et al. 2001</bibRefCitation>
: 721 (listed, hosts, records, distribution);
<bibRefCitation id="0DE26649AC4C83CC419B79E33118BC5A" author="Lin, Kwei-Shui" journalOrPublisher="Special Publication" pageId="9" pageNumber="90" pagination="1 - 2820" title="A list of natural enemies of insect pests. Taiwan Agricultural Research Institute." url="http://ir.tari.gov.tw:8080/handle/345210000/5095" volume="104" year="2003">Lin 2003</bibRefCitation>
: 939 (listed, host, distribution);
<bibRefCitation id="5A4064B8F09E39AE930F20ACA836EA87" author="Wang, Shu-Fang" editor="Huang, Bang-Kan" journalOrPublisher="Fujian Science and Technology Publishing House, Fuzhou" pageId="11" pageNumber="92" pagination="273 - 277" title="Mesosteninae." volumeTitle="Fauna of Insects in Fujian Province of China" year="2003">Wang 2003</bibRefCitation>
: 274 (description, figure, host, records, distribution);
<bibRefCitation id="6405ADAB801BDFD9C7BA8A319D5D2376" author="He, Jun-Hua" journalOrPublisher="Science Press, Beijing, China" pageId="9" pageNumber="90" title="Hymenopteran insect fauna of Zhejiang." year="2004">He et al. 2004</bibRefCitation>
: 514 (description, figures, host, records, distribution);
<bibRefCitation id="A4DD695EA6C38190893A790C2E1A1426" author="He, Jun-Hua" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="9" pageNumber="90" year="2006">He and Chen 2006</bibRefCitation>
: 110 (description, figures, biology, distribution);
<bibRefCitation id="055AAD84E49976A1011AD3C8DE764734" author="Chen, Shu-Pei" journalOrPublisher="Taiwan Agricultural Research Institute Special Publication" pageId="9" pageNumber="90" pagination="1 - 466" title="A list of natural enemies of insect pests in Taiwan." url="http://ir.tari.gov.tw:8080/handle/345210000/2942" volume="137" year="2009">Chen et al. 2009</bibRefCitation>
: 70 (listed, distribution, records);
<bibRefCitation id="52D59E98EEBC02BDBBFA7D9352005735" author="Sheng, Mao-Ling" journalOrPublisher="Science Press, Beijing" pageId="10" pageNumber="91" title="Insect Fauna of Henan. Hymenoptera: Ichneumonidae." year="2009">Sheng and Sun 2009</bibRefCitation>
: 69 (description, photo, hosts, records, distribution);
<bibRefCitation id="2F23D12C739AC007ED11F95B96659EB9" author="Sheng, Mao-Ling" journalOrPublisher="Science Press, Beijing" pageId="10" pageNumber="91" title="Ichneumonid Fauna of Jiangxi (Hymenoptera: Ichneumonidae)." year="2013">Sheng et al. 2013</bibRefCitation>
: 138 (description, photo, hosts, records, distribution).
</paragraph>
</subSubSection>
<subSubSection id="971CD0238F3809D8CEFC5C6F8CA23B0B" pageId="5" pageNumber="86" type="type specimens examined">
<paragraph id="5CF9ECE11C8ADCB41EE855803DC9769E" pageId="5" pageNumber="86">
<pageBreakToken id="3385BE7350E1E075EF0B4D292857F034" pageId="5" pageNumber="86" start="start">Type</pageBreakToken>
specimens examined.
</paragraph>
<paragraph id="2721F4A84E6C91A84D75E4DCD32113B8" pageId="5" pageNumber="86">
Holotype of
<taxonomicName id="A1A0641D9AD0BF217C30594D32D0B9C3" class="Gastropoda" family="Cryptaulacidae" genus="Cryptaulax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptaulax coreanus" pageId="5" pageNumber="86" phylum="Mollusca" rank="species" species="coreanus">Cryptaulax coreanus</taxonomicName>
<normalizedToken id="277C6512D16E75B66E449DB048107C9A" originalValue="Szépligeti">Szepligeti</normalizedToken>
: ♀, &quot;Korea, Sebul&quot; [
<normalizedToken id="477DFC0E4F20CC682D42A537D32F5A43" originalValue="“Soeul”">&quot;Soeul&quot;</normalizedToken>
in the original description], HNHM (ID 115220). Paralectotype of
<taxonomicName id="AA122C03A5D24E36DE73420FFD3E15D6" class="Insecta" family="Ichneumonidae" genus="Cryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptus trirrhogmaniformis" order="Hymenoptera" pageId="5" pageNumber="86" phylum="Arthropoda" rank="species" species="trirrhogmaniformis">Cryptus trirrhogmaniformis</taxonomicName>
Sonan: ♂, &quot;Shinchiku, Chikurin, ft. 3000&quot; [= Chulin, Hsinchu], 10.ix.1928, J. Sonan, reared from a cocoon of
<taxonomicName id="7FB830A65BFC083CB30971CAC61CD42D" class="Insecta" family="Limacodidae" genus="Monema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monema" order="Lepidoptera" pageId="5" pageNumber="86" phylum="Arthropoda" rank="genus">Monema</taxonomicName>
sp. on a cherry tree, SEHU.
</paragraph>
</subSubSection>
<subSubSection id="7CCEDD95ABA0D74B8F007A877BD53D46" pageId="5" pageNumber="86" type="other specimens">
<paragraph id="D6D4108721095BDF24FD16F5AC9606E7" pageId="5" pageNumber="86">Other specimens.</paragraph>
<paragraph id="03463A301269EF04B83879D7015C52DC" pageId="5" pageNumber="86">
South Korea. Poseoksa (N36°03.073
<normalizedToken id="59C8B0C88408FBCDEA50F338886F8033" originalValue="/E127°28.688">'/E127°28.688'</normalizedToken>
), Keumsan, Chungcheong-namdo, P. Tripotin, SEHU: 1 ♂, viii.1998; 1 ♀, 21.ix.2001. Seonunsan (N36°27
<normalizedToken id="189BC44354E3AD81DB9158C1BEF1AFF8" originalValue="50˝/E126°3428˝">'50&quot;/E126°34'28&quot;</normalizedToken>
), Geochang, Jeollabuku-do, 1 ♂, 5.vi.2004, C. L. Young, SEHU. Suigen [= Suwon], Gyeonggi-do, K. Sato, SEHU: 1 ♂, 1925; 1 ♀, 4 ♂, 1.vi.1928; 1 ♀, 14.vi.1928.
<normalizedToken id="2D4704BA6DCD3ABC14F116FA896895CF" originalValue="Shôyo-san">Shoyo-san</normalizedToken>
, Keikido [= Soyosan, Gyeonggi-do], SEHU: 1 ♀, 15.vi.1941; 2 ♂, 10.ix.1943; 2 ♂, 17.ix.1943. Homyeongsan, Cheongpyeong, 1 ♂, 4.v, SEHU. Godaesan, Gyeonggi-do, 1 ♂, 21.vi.1942, SEHU. China. Tetsurei, Manchoukuo, [= Tieling, Liaoning Prov.], 1 ♀, 1 ♂, 11.vi.1938, I. Okada, SEHU. Domonrei, Manchoukuo [= Tumenling, Jiutai, Jilin Prov.], 1 ♀, 1 ♂, 19.viii.1939, H.
<normalizedToken id="0F079743B00284FBC383CB8E05B388AD" originalValue="Kôno">Kono</normalizedToken>
, SEHU. Kaigen, Manchoukuo, [= Kaiyuan, Liaoning Prov.], I. Okada, SEHU: 1 ♀, 15.ix.1935; 1 ♀, 1.vi.1938.
<normalizedToken id="5B4BCC0166AD51210F1EF87F62253B4B" originalValue="Ryôsuiji">Ryosuiji</normalizedToken>
, Dalian, 1 ♀, 23.vi.1929, SEHU. &quot;Nord Pekin&quot;, 1 ♂, A. David, 1865, MNHN.
</paragraph>
</subSubSection>
<subSubSection id="D9236AF888037A73546183E5047D5E95" pageId="5" pageNumber="86" type="diagnosis">
<paragraph id="3584529AC64B06689690E5181F5017C8" pageId="5" pageNumber="86">Diagnosis.</paragraph>
<paragraph id="FA388D7EFFC92417DE03BCF1EA7E40B2" pageId="5" pageNumber="86">
Punctation on head coarser than
<taxonomicName id="0AF861FE44C456304EB6F8D12FCA9165" class="Insecta" family="Ichneumonidae" genus="Chlorocryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlorocryptus purpuratus" order="Hymenoptera" pageId="5" pageNumber="86" phylum="Arthropoda" rank="species" species="purpuratus">Chlorocryptus purpuratus</taxonomicName>
. Temple swollen and wide, occupying 0.3-0.4 of length of head in dorsal view (Fig. 15). Face without transverse rugae (Fig. 13). Clypeus usually with pair of weak tubercles on apical margin (Fig. 14). Mandibular teeth blunt (Fig. 17). Female flagellum with white annulus around mid-length and with apical third ventrally weakly flattened but not widened. Epomia not turned toward mesal line of pronotum (Fig. 19). Prepectus without a short vertical carina opposite lower corner of pronotum. Mesoscutum punctate, without rugae, and glabrous area on lateral lobe narrow (Fig. 18). Propodeum without posterior transverse carina. Nervellar index 1.3-1.8 (Figs 26, 27). Postpetiole long, 0.8-1.0 as long as wide in female, 0.7-1.0 in male (Fig. 21). Median longitudinal carina of 1st tergite not raised at all at beginning of postpetiole (Fig. 20). Subgenital plate (Figs 36, 37) with postero-lateral corner more angulated, with posterior margin weakly notched on median convexity, and with white spot at base of its apodeme.
</paragraph>
</subSubSection>
<subSubSection id="2AF589AC6DD0DE70AC482FFA8ACB69C7" lastPageId="8" lastPageNumber="89" pageId="5" pageNumber="86" type="description">
<paragraph id="09F8A25E9B036E115C73B3D443E93D83" pageId="5" pageNumber="86">Description.</paragraph>
<paragraph id="A01BEDB272748A2A2B0C2E463179FAF0" lastPageId="6" lastPageNumber="87" pageId="5" pageNumber="86">
Adult. ♀. Head (Fig. 15) 1.7-1.8 times as wide as long in dorsal view. Temple swollen, occupying 0.3-0.4 of length of head in dorsal view (Fig. 15). POL/OOL=0.6-0.8. Vertex and gena densely with coarse punctures; punctures smaller and shallower on gena than
<pageBreakToken id="CA286E4E97FEDA508E3DD2D14C16B0D3" pageId="6" pageNumber="87" start="start">on</pageBreakToken>
vertex. Face 1.4-1.5 times as wide as high, distinctly convex medially, punctate-reticulate (Figs 5, 13). Clypeus with pair of weak tubercles on apical margin, punctate-reticulate, punctures larger than on face (Fig. 14). Malar space 0.8-1.0 times as wide as mandible width. Mandible stout, with short and blunt teeth; upper tooth broad, a little longer than lower one (Fig. 17). Antenna with 32-34 flagellomeres. Apical third of flagellum weakly flattened ventrally but not thickened, weakly tapered to apex. First flagellomere 4.9-5.8 times as long as apical width, a little longer than 2nd flagellomere. Fifth flagellomere 2.3-2.5 times as long as apical width.
</paragraph>
<paragraph id="D362B3EC6BF6149075DEEC8E6C3B164C" pageId="6" pageNumber="87">Epomia ends at midway to upper edge of pronotum without turning toward mesal line of pronotum (Fig. 19). Prepectus without a short vertical carina opposite lower corner of pronotum. Mesoscutum and scutellum lacunose, without rugae, and with only narrow glabrous stripe on lateral lobe (Fig. 18). Mesopleuron punctate to areolate-rugose on lower part to transversely strigate on upper frontal part. Mesosternum densely punctate. Metapleuron areolate-rugose. Upper division of metapleuron punctulate. Propodeum areolate rugose; only anterior transverse carina present, other carinae absent.</paragraph>
<paragraph id="79409A429FA27BE79F746F76A9A66748" pageId="6" pageNumber="87">Legs slender. Hind femur 6.0-7.7 times as long as maximum width. Hind tibia 8.5-9.5 times as long as apical width.</paragraph>
<paragraph id="8EF9DDE612C31FE5308BAA1D924E749D" pageId="6" pageNumber="87">Fore wing about 11-14 mm long. Nervellar index 1.3-1.8 (Figs 26, 27).</paragraph>
<paragraph id="DDE3EC2567309288D45E1CE545E53CE1" lastPageId="7" lastPageNumber="88" pageId="6" pageNumber="87">
First metasomal tergite 1.9-2.3 times as long as apical width, with postpetiole 0.7-1.0 times as long as apical
<pageBreakToken id="1FAB92C7F78B094AD9D8BB9AC2F8A2F0" pageId="7" pageNumber="88" start="start">width</pageBreakToken>
; dorsal face not distinctly raised; median longitudinal carina absent or very weak on its entire length (Figs 20, 21). Ovipositor sheath 0.9-1.0 times as long as hind tibia.
</paragraph>
<paragraph id="2F5B74D11E061E650EE81E91F29F172B" pageId="7" pageNumber="88">
Body with metallic luster in blue, purple or green. Flagellum black with white band on 6
<normalizedToken id="545EE1EE3A3C42BF0234A9716B886626" originalValue="th">th-</normalizedToken>
9th flagellomeres. Fore and middle tibiae and all tarsi are dark brown to black. Ovipositor sheath and ovipositor black. Wings narrowly infumate apically (Fig. 26).
</paragraph>
<paragraph id="CE396934F83AA7495621682E3FBF4866" lastPageId="8" lastPageNumber="89" pageId="7" pageNumber="88">
♂. Similar to female except as follows: face 1.0-1.2 times as wide as long; flagellum with 37-39 flagellomeres, with 5 or 6 tyloids which can be on 14
<normalizedToken id="A6F9BB1A066CDD624418FA2E9CC477DC" originalValue="th">th-</normalizedToken>
20th flagellomeres; 1st flagellomere 2.8-3.3 times as long as
<pageBreakToken id="99762875151A554333C0E3E6C6E3318C" pageId="8" pageNumber="89" start="start">apical</pageBreakToken>
width; 1st metasomal segment 2.0-2.4 times as long as apical width; hind femur 7.0-8.5 times as long as median depth; hind tibia 9.3-10.0 times as long as apical width; stripe on anterior face of fore femur, fore tibia and dorsal stripe of middle tibia light brown. Subgenital plate with postero-lateral corner more angulated than in
<taxonomicName id="0DB4EE41B9EF155FCEEEA3B16BA618FB" class="Insecta" family="Ichneumonidae" genus="Chlorocryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlorocryptus purpuratus" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="purpuratus">Chlorocryptus purpuratus</taxonomicName>
, with posterior margin convex, with small concavity on middle of convexity (Figs 36, 37), but one aberrant specimen from Korea without concavity (Fig. 38); usually with white spot near base of apodeme. Wings narrowly infumate in specimens from Korea and North China, but Taiwanese specimens with extensively brown wings (Fig. 27).
</paragraph>
</subSubSection>
<subSubSection id="7774CF90FFA40009CE7384138BCC861F" pageId="8" pageNumber="89" type="biology">
<paragraph id="E85CDD1AF7CA48C246B65E48829AB78A" pageId="8" pageNumber="89">Biology.</paragraph>
<paragraph id="12284DD2E6483E96CF5B089ACD4D88F0" pageId="8" pageNumber="89">
<taxonomicName id="C462455F32037BA17513C5ECDA1F2CEC" class="Insecta" family="Limacodidae" genus="Monema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monema flavescens" order="Lepidoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="flavescens">Monema flavescens</taxonomicName>
Walker (
<bibRefCitation id="D59F498FF5432FE7A7EF1EFBD0930543" author="Uchida, T" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido Imperial University" pageId="10" pageNumber="91" pagination="349 - 376" title="Beitrag zur Ichneumoniden-Fauna Japans." url="http://hdl.handle.net/2115/12653" volume="25" year="1930 b">Uchida 1930b</bibRefCitation>
,
<bibRefCitation id="6048DF000094E1A756AFA61B33763C64" author="Sun, Xue-Yi" journalOrPublisher="Liaoning Forestry Science and Technology" pageId="10" pageNumber="91" pagination="46 - 48" volume="1982" year="1982">Sun et al. 1982</bibRefCitation>
,
<bibRefCitation id="F942D20AC535EFB015ECA649AB481F46" author="Pan, Lang" journalOrPublisher="Liaoning Agricultural Sciences" pageId="10" pageNumber="91" pagination="50 - 51" volume="1983" year="1983">Pan 1983</bibRefCitation>
,
<bibRefCitation id="B35D690A5C9980A1FEA22EB9F6C06F61" author="Yin, Ping-Fang" journalOrPublisher="Journal of Jilin Forestry Institute" pageId="11" pageNumber="92" pagination="11 - 20" title="A study on the efficiency of some natural enemies to control the community density of oriental moth." volume="3" year="1987">Yin and Zou 1987</bibRefCitation>
,
<bibRefCitation id="63F59D0E46C03F9A629F2A3B28B9EF1B" author="Wang, Fu-Wei" journalOrPublisher="Jilin Forestry Science and Technology" pageId="11" pageNumber="92" pagination="30 - 32" volume="1991" year="1991">Wang et al. 1991</bibRefCitation>
,
<bibRefCitation id="0D0DBFC5E74FADF96F663404E0BBB5ED" author="Wang, Fu-Wei" journalOrPublisher="Forest Research" pageId="10" pageNumber="91" pagination="466 - 469" title="Study on the natural enemy complex of oriental moth." volume="6" year="1993">1993</bibRefCitation>
) and
<taxonomicName id="4843C067BCC18F3E986975EFEC0E3153" class="Insecta" family="Limacodidae" genus="Miresa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Miresa inornata" order="Lepidoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="inornata">Miresa inornata</taxonomicName>
Walker (
<taxonomicName id="799006547286D386B07BC291705759EE" family="Limacodidae" lsidName="" pageId="8" pageNumber="89" rank="family">Limacodidae</taxonomicName>
) (
<bibRefCitation id="E693CAB4B0D92FA5237119D82FCB17B6" author="Uchida, T" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido Imperial University" pageId="10" pageNumber="91" pagination="299 - 347" title="Fuenfter Beitrag zur Ichneumoniden-Fauna Japans." url="http://hdl.handle.net/2115/12652" volume="25" year="1930 a">Uchida 1930a</bibRefCitation>
) have been recorded as hosts.
</paragraph>
</subSubSection>
<subSubSection id="1AEBBAE70C3AD25F509C06F5BA02706B" pageId="8" pageNumber="89" type="distribution">
<paragraph id="C44893AF76760A0CDCB39F2C510A4046" pageId="8" pageNumber="89">Distribution.</paragraph>
<paragraph id="084A951AC27972A56C53A2CA9EFA4B24" pageId="8" pageNumber="89">
(Fig. 58). Korea (
<bibRefCitation id="7CE5D2B18523B9A7C0E1D35CA8879789" author="Szepligeti, G" journalOrPublisher="Annales Musei Nationalis Hungarici" pageId="10" pageNumber="91" pagination="225 - 380" title="Ichneumoniden aus der Sammlung des ungarischen National-Museums. II." volume="14" year="1916">
<normalizedToken id="68C1DD5CF884F278DF9364C8F8FC259D" originalValue="Szépligeti">Szepligeti</normalizedToken>
1916
</bibRefCitation>
,
<bibRefCitation id="569B60E6FC3FF3070708437B4EE07192" author="Uchida, T" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido Imperial University" pageId="10" pageNumber="91" pagination="299 - 347" title="Fuenfter Beitrag zur Ichneumoniden-Fauna Japans." url="http://hdl.handle.net/2115/12652" volume="25" year="1930 a">Uchida 1930a</bibRefCitation>
,
<bibRefCitation id="D685F873DE8C4F580B9667064926418A" author="Uchida, T" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido University" pageId="10" pageNumber="91" pagination="95 - 133" title="Die von Dr. K. Tsuneki in Korea gesammelten Ichneumoniden." url="http://hdl.handle.net/2115/12759" volume="50" year="1955">1955</bibRefCitation>
,
<bibRefCitation id="953974B281DAE82593A1E8CFA2738B4B" pageId="8" pageNumber="89">Kim 1955</bibRefCitation>
). China: Fujian (
<bibRefCitation id="877BD1E5043E9780550A8AC20973AB3D" author="Wang, Shu-Fang" editor="Huang, Bang-Kan" journalOrPublisher="Fujian Science and Technology Publishing House, Fuzhou" pageId="11" pageNumber="92" pagination="273 - 277" title="Mesosteninae." volumeTitle="Fauna of Insects in Fujian Province of China" year="2003">Wang 2003</bibRefCitation>
,
<bibRefCitation id="D9C8907265550FC34C4D957B7E962615" author="Wang, Shug-Fang" editor="Huang, Chun-Mei" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="11" pageNumber="92" pagination="727 - 738" title="Hymenoptera: Ichneumonidae." volumeTitle="Animals of Longqi Mountain" year="1993">Wang and Huang 1993</bibRefCitation>
), Guangxi (
<bibRefCitation id="FFB1A218BCA1A8876CA99C4F2DD7A048" author="Niu, Yan-Zhang" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="10" pageNumber="91" title="Chlorocryptuscoreanus Szepligeti. In: Xiao Gang-Rou (Ed.). Forest Insects of China" year="1992">Niu and Wang 1992</bibRefCitation>
), Heilongjiang (
<bibRefCitation id="940F4611F9F395D1E79C9ACF810CD22C" author="Niu, Yan-Zhang" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="10" pageNumber="91" title="Chlorocryptuscoreanus Szepligeti. In: Xiao Gang-Rou (Ed.). Forest Insects of China" year="1992">Niu and Wang 1992</bibRefCitation>
), Henan (
<bibRefCitation id="E702792CDCF17E6C88ADB0B8B7DA0DD2" author="Sheng, Mao-Ling" editor="Shen, Xia-Cheng" journalOrPublisher="China Agricultural Scientech Press, Beijing" pageId="10" pageNumber="91" pagination="373 - 379" title="Hymenoptera: Ichneumonidae." volumeTitle="The Fauna and Taxonomy of Insects in Henan Vol. 4. Insects of the Mountains Funiu and Dabie Regions" year="1999">Sheng et al. 1999</bibRefCitation>
,
<bibRefCitation id="56D1DEDB10A9A67BE297015C92302B90" author="Sheng, Mao-Ling" journalOrPublisher="Science Press, Beijing" pageId="10" pageNumber="91" title="Insect Fauna of Henan. Hymenoptera: Ichneumonidae." year="2009">Sheng and Sun 2009</bibRefCitation>
), Jiangxi (
<bibRefCitation id="D0904833983B23464AB77B7FDA59F245" author="Sheng, Mao-Ling" journalOrPublisher="Science Press, Beijing" pageId="10" pageNumber="91" title="Ichneumonid Fauna of Jiangxi (Hymenoptera: Ichneumonidae)." year="2013">Sheng et al. 2013</bibRefCitation>
), Jilin (
<bibRefCitation id="15A73DE4F998CD2BDFDE8EA2DBAC504B" author="Yin, Ping-Fang" journalOrPublisher="Journal of Jilin Forestry Institute" pageId="11" pageNumber="92" pagination="11 - 20" title="A study on the efficiency of some natural enemies to control the community density of oriental moth." volume="3" year="1987">Yin and Zou 1987</bibRefCitation>
), Liaoning (
<bibRefCitation id="A3233220D352E4D6196B12CB12C0FA1F" author="Uchida, T" journalOrPublisher="Insecta Matsumurana" pageId="10" pageNumber="91" pagination="107 - 146" title="Ichneumoniden Mandschukuos aus dem Entomologischen Museum der kaiserlichen Hokkaido Universitaet." url="http://hdl.handle.net/2115/9491" volume="16" year="1942">Uchida 1942</bibRefCitation>
,
<bibRefCitation id="57DD3F79CA85A09A9BC0E7B8A43C43C0" author="Sun, Xue-Yi" journalOrPublisher="Liaoning Forestry Science and Technology" pageId="10" pageNumber="91" pagination="46 - 48" volume="1982" year="1982">Sun et al. 1982</bibRefCitation>
,
<bibRefCitation id="367E99B1FDCE170E4F09108D674249BA" author="Pan, Lang" journalOrPublisher="Liaoning Agricultural Sciences" pageId="10" pageNumber="91" pagination="50 - 51" volume="1983" year="1983">Pan 1983</bibRefCitation>
), Neimenggu (
<bibRefCitation id="E04715826A583BD5F55550F4905EBEF1" author="Niu, Yan-Zhang" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="10" pageNumber="91" title="Chlorocryptuscoreanus Szepligeti. In: Xiao Gang-Rou (Ed.). Forest Insects of China" year="1992">Niu and Wang 1992</bibRefCitation>
), Shaanxi (
<bibRefCitation id="A5BE6C56F002E39ABD0790AE4A18F415" author="He, Jun-Hua" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="9" pageNumber="90" year="2006">He and Chen 2006</bibRefCitation>
), Sichuan (
<bibRefCitation id="4A1F20F2E19104488C978A9BC812EDCD" author="Wang, Shug-Fang" editor="Huang, Chun-Mei" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="11" pageNumber="92" pagination="727 - 738" title="Hymenoptera: Ichneumonidae." volumeTitle="Animals of Longqi Mountain" year="1993">Wang and Huang 1993</bibRefCitation>
), Yunnan (
<bibRefCitation id="D6B02A1D4DFAA66B04AB8C36A1B99E9C" author="Wang, Shu-Fang" editor="Huang, Fu-Sheng" journalOrPublisher="Yunnan Science and Technology Press, Kunming" pageId="11" pageNumber="92" pagination="1317 - 1336" title="Ichneumonidae." volumeTitle="Forest Insects of Yunnan" year="1987">Wang and Li 1987</bibRefCitation>
), Zhejiang (
<bibRefCitation id="1ADDC82E6FC33CCD757E4A8481929973" author="He, Jun-Hua" journalOrPublisher="Agricultural Publishing House, Beijing" pageId="9" pageNumber="90" pagination="367 - 400" title="Ichneumonidae." volumeTitle="Agriculture Insects of China" year="1987">He and Wang 1987</bibRefCitation>
,
<bibRefCitation id="B62E2A9DEF71574244A411777611D1F9" author="He, Jun-Hua" editor="Wu, Hong" journalOrPublisher="Science Press, Beijing" pageId="9" pageNumber="90" pagination="710 - 723" title="Hymenoptera: Ichneumonidae." volumeTitle="Insects of Tianmushan National Nature Reserve" year="2001">He et al. 2001</bibRefCitation>
). Taiwan (
<bibRefCitation id="C911CA78E611E5AD8019D2C6710C3F85" author="Sonan, J" journalOrPublisher="Transactions of the Natural History Society of Formosa" pageId="10" pageNumber="91" pagination="415 - 425" title="A few host-known Ichneumonidae found in Formosa (Hym.)." volume="19" year="1929">Sonan 1929</bibRefCitation>
,
<bibRefCitation id="A9769EDCCCCE80379BA8D938C2BD77D5" author="Uchida, T" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido Imperial University" pageId="10" pageNumber="91" pagination="163 - 193" title="Beitrag zur Kenntnis der Cryptinenfauna Formosas." url="http://hdl.handle.net/2115/12678" volume="30" year="1931">Uchida 1931</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="383F9057C6A469B395A825639D5D966B" pageId="8" pageNumber="89" type="remarks">
<paragraph id="B487D040F11C41460008B8DFEE4905A9" pageId="8" pageNumber="89">Remarks.</paragraph>
<paragraph id="29B7F905A9279830AAB654EE19CAFE9F" pageId="8" pageNumber="89">
Similar to the preceding species, this species shows considerable intraspecific morphological variation. Only the Taiwanese specimens, namely the type series of
<taxonomicName id="D5781C70F105E0D9EC5071976662D485" class="Insecta" family="Ichneumonidae" genus="Cryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptus trirrhogmaniformis" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="trirrhogmaniformis">Cryptus trirrhogmaniformis</taxonomicName>
, have the fore wings entirely tinged with brown (Fig. 27) (according to the original description of
<taxonomicName id="45DEBCA17C141FB89D8E96F82B71F9F5" class="Insecta" family="Ichneumonidae" genus="Cryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptus trirrhogmaniformis" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="trirrhogmaniformis">Cryptus trirrhogmaniformis</taxonomicName>
, the lectotype has also brown wings), and all the other specimens examined have the fore wings darkened only apically (Fig. 26). One specimen from Korea (Fig. 38) has a differently shaped subgenital plate, which we considered to be aberrant.
</paragraph>
<paragraph id="9A5ED0AE43DA3E77DDE0391E2AC8B75A" pageId="8" pageNumber="89">
In the description of
<taxonomicName id="875B959EAF72E1EA79B4953F24EDA2CE" class="Gastropoda" family="Cryptaulacidae" genus="Cryptaulax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptaulax coreanus" pageId="8" pageNumber="89" phylum="Mollusca" rank="species" species="coreanus">Cryptaulax coreanus</taxonomicName>
,
<bibRefCitation id="52E18BB86C69F8821644E7D76CDB364A" author="Szepligeti, G" journalOrPublisher="Annales Musei Nationalis Hungarici" pageId="10" pageNumber="91" pagination="225 - 380" title="Ichneumoniden aus der Sammlung des ungarischen National-Museums. II." volume="14" year="1916">
<normalizedToken id="BFA0E2DF8BB76418D56DD73FD4798B61" originalValue="Szépligeti">Szepligeti</normalizedToken>
(1916)
</bibRefCitation>
did not originally designate the holotype nor mentioned the number of specimens he examined. The
<normalizedToken id="945CBDB3DACD6FFF12B608F42F427EF4" originalValue="Szépligetis">Szepligeti's</normalizedToken>
collection at HNHM contains only one specimen of the species, which
<bibRefCitation id="3350D1D9527D976A636B87D0022EE613" author="Townes, HK" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="10" pageNumber="91" pagination="1 - 522" title="A catalogue and reclassification of the Indo-Australian Ichneumonidae." volume="1" year="1961">Townes et al. (1961)</bibRefCitation>
treated as the holotype fixed by monotypy.
</paragraph>
<paragraph id="A6EF9FFF59B5ED8BCB5EA62790562430" pageId="8" pageNumber="89">
<taxonomicName id="597B215BDA052ADC681EBBC77DBBC79F" class="Insecta" family="Ichneumonidae" genus="Cryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptus trirrhogmaniformis" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="trirrhogmaniformis">Cryptus trirrhogmaniformis</taxonomicName>
Sonan was described based on two male syntypes. One was deposited at TARI and the other was probably donated to Uchida at that time and now is located at SEHU. The TARI specimen was treated as
<normalizedToken id="194E77C57B1482347BA3F5B362A82643" originalValue="“type”">&quot;type&quot;</normalizedToken>
by
<bibRefCitation id="53298BC9821300940C908911E2AACE1E" author="Townes, HK" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="10" pageNumber="91" pagination="1 - 522" title="A catalogue and reclassification of the Indo-Australian Ichneumonidae." volume="1" year="1961">Townes et al. (1961)</bibRefCitation>
, which is considered a lectotype designation as it is evident that the authors cited the specimen to serve as the name-bearing type (ICZN 74.5).
</paragraph>
</subSubSection>
</treatment>
</document>

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Type status:
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. Occurrence: recordNumber: 21 L; recordedBy:
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; Location: county: NOR-Sogn og Fjordane; locality:
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; eventDate:
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; habitat: Grass tussocks on stony beach, humic soil, herbaceous plants, moss
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Sweden (
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) and new for Norway.
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<treatment LSID="urn:lsid:zoobank.org:act:15602B59-C063-4007-82C0-9141BD06D2BE" httpUri="http://treatment.plazi.org/id/D0B9A23F51045CDEA8EE3DE5A93BF24E" lastPageNumber="215" pageId="0" pageNumber="215">
<subSubSection pageId="0" pageNumber="215" type="nomenclature">
<paragraph pageId="0" pageNumber="215">
<taxonomicName LSID="http://zoobank.org/15602B59-C063-4007-82C0-9141BD06D2BE" authority="Cavalcante-Silva &amp; Pereira &amp; Calor, 2022" authorityName="Cavalcante-Silva &amp; Pereira &amp; Calor" authorityYear="2022" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Helicopsyche (Feropsyche) ralphi" order="Trichoptera" pageId="0" pageNumber="215" phylum="Mollusca" rank="species" species="ralphi" status="sp. nov." subGenus="Feropsyche">Helicopsyche (Feropsyche) ralphi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="215">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="215" type="description">
<paragraph pageId="0" pageNumber="215">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">Figs 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Helicopsyche ralphi sp. nov., larvae and case A larval case, dorsal view B larval case, ventral view C larval lateral habitus." figureDoi="10.3897/zookeys.1111.77541.figure6" httpUri="https://binary.pensoft.net/fig/712454" pageId="0" pageNumber="215">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Helicopsyche ralphi sp. nov., pupa and case A pupa front B abdominal segment IX and anal processes, dorsal view C pupa lateral habitus D-H abdominal segments I-V, dorsal, with details of dorsal hook plates I pupa case, ventral view J pupa case with sieve membrane highlighted, ventral view." figureDoi="10.3897/zookeys.1111.77541.figure7" httpUri="https://binary.pensoft.net/fig/712455" pageId="0" pageNumber="215">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="215" type="materials_examined">
<paragraph pageId="0" pageNumber="215">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="215">
<materialsCitation collectingDate="2017-09-21" collectingMethod="UV light pan trap" collectorName="Cavalcante-Silva, A." country="Brazil" elevation="890" latitude="-7.8675003" location="Pinga" longLatPrecision="21" longitude="-38.120277" municipality="Triunfo" specimenCount="1" specimenCount-male="1" stateProvince="Pernambuco" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="215">
<emphasis italics="true" pageId="0" pageNumber="215">
<typeStatus>Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry name="Brazil">Brazil</collectingCountry>
,
<specimenCount type="male">1 male</specimenCount>
;
<collectingRegion country="Brazil" name="Pernambuco">Pernambuco</collectingRegion>
,
<collectingMunicipality>Triunfo</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:D0B9A23F51045CDEA8EE3DE5A93BF24E:A3CD7196BE49228CAEEFBC93D5D638F8" country="Brazil" latitude="-7.8675003" longLatPrecision="21" longitude="-38.120277" municipality="Triunfo" name="Pinga" stateProvince="Pernambuco">Pinga</location>
stream;
<geoCoordinate degrees="7" direction="south" minutes="52" orientation="latitude" precision="15" seconds="3" value="-7.8675003">7°52'3&quot;S</geoCoordinate>
,
<geoCoordinate degrees="38" direction="west" minutes="7" orientation="longitude" precision="15" seconds="13" value="-38.120277">38°7'13&quot;W</geoCoordinate>
, el.
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">890 m</elevation>
</quantity>
;
<collectingDate value="2017-09-21">21.ix.2017</collectingDate>
;
<collectorName>Cavalcante-Silva, A.</collectorName>
leg.;
<collectingMethod>UV light pan trap</collectingMethod>
(MZSP)
</materialsCitation>
.
<materialsCitation collectingDate="2017-09-21" collectingDateMax="2019-05-01" collectingDateMin="2017-09-21" collectorName="Cavalcante-Silva, A, Pereira, R." country="Brazil" elevation="740" latitude="-7.8681946" location="Pinga" longLatPrecision="1" longitude="-38.121025" municipality="Triunfo" specimenCount="44" specimenCount-female="24" specimenCount-male="20" stateProvince="Pernambuco" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="215">
<emphasis italics="true" pageId="0" pageNumber="215">
<typeStatus>Paratypes</typeStatus>
</emphasis>
.
</emphasis>
Same data as holotype, except
<specimenCount type="male">5 males</specimenCount>
; Grito stream;
<geoCoordinate degrees="7" direction="south" minutes="51" orientation="latitude" precision="15" seconds="41" value="-7.8613887">7°51'41&quot;S</geoCoordinate>
,
<geoCoordinate degrees="38" direction="west" minutes="5" orientation="longitude" precision="15" seconds="25" value="-38.090275">38°5'25&quot;W</geoCoordinate>
, el.
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.4" unit="m" value="740.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.4" unit="m" value="740.0">740 m</elevation>
</quantity>
;
<collectingDate value="2018-08-09">09.viii.2018</collectingDate>
(UFBA); same except
<specimenCount type="male">3 males</specimenCount>
; Laje stream;
<geoCoordinate degrees="7" direction="south" minutes="52" orientation="latitude" precision="15" seconds="13" value="-7.870278">7°52'13&quot;S</geoCoordinate>
,
<geoCoordinate degrees="38" direction="west" minutes="5" orientation="longitude" precision="15" seconds="18" value="-38.088333">38°5'18&quot;W</geoCoordinate>
, el.
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.8" unit="m" value="580.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.8" unit="m" value="580.0">580 m</elevation>
</quantity>
;
<collectingDate value="2018-08-07">07.viii.2018</collectingDate>
(DZRJ); same except
<specimenCount type="male">6 males</specimenCount>
; Pinga stream;
<geoCoordinate degrees="7" direction="south" minutes="52" orientation="latitude" precision="15" seconds="3" value="-7.8675003">7°52'3&quot;S</geoCoordinate>
,
<geoCoordinate degrees="38" direction="west" minutes="7" orientation="longitude" precision="15" seconds="13" value="-38.120277">38°7'13&quot;W</geoCoordinate>
, el.
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">890 m</elevation>
</quantity>
;
<collectingDate value="2017-09-21">21.ix.2017</collectingDate>
(MZSP); same except
<specimenCount type="male">6 males</specimenCount>
(INPA); same except
<specimenCount type="female">1 female</specimenCount>
;
<geoCoordinate degrees="7" direction="south" minutes="52" orientation="latitude" precision="15" seconds="3" value="-7.8675003">7°52'3&quot;S</geoCoordinate>
,
<geoCoordinate degrees="38" direction="west" minutes="7" orientation="longitude" precision="15" seconds="13" value="-38.120277">38°7'13&quot;W</geoCoordinate>
, el.
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">890 m</elevation>
</quantity>
;
<collectingDate value="2019-02-03">03.ii.2019</collectingDate>
(MZSP); same except
<specimenCount type="female">5 females</specimenCount>
;
<geoCoordinate degrees="7" direction="south" minutes="52" orientation="latitude" precision="1" seconds="5,5" value="-7.8681946">7°52'5,5&quot;S</geoCoordinate>
,
<geoCoordinate degrees="38" direction="west" minutes="7" orientation="longitude" precision="1" seconds="15,6" value="-38.121">38°7'15,6&quot;W</geoCoordinate>
, el.
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.7" unit="m" value="870.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.7" unit="m" value="870.0">870 m</elevation>
</quantity>
;
<collectingDate value="2019-05-01">01.v.2019</collectingDate>
;
<collectorName>Cavalcante-Silva, A</collectorName>
,
<collectorName>Pereira, R.</collectorName>
leg. (MZSP); same except
<specimenCount type="female">6 females</specimenCount>
;
<geoCoordinate degrees="7" direction="south" minutes="52" orientation="latitude" precision="1" seconds="5,5" value="-7.8681946">7°52'5,5&quot;S</geoCoordinate>
,
<geoCoordinate degrees="38" direction="west" minutes="7" orientation="longitude" precision="1" seconds="15,7" value="-38.121025">38°7'15,7&quot;W</geoCoordinate>
, el.
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.65" unit="m" value="865.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.65" unit="m" value="865.0">865 m</elevation>
</quantity>
;
<collectingDate value="2019-05-01">01.v.2019</collectingDate>
(UFBA); same except
<specimenCount type="female">6 females</specimenCount>
(DZRJ); same except
<specimenCount type="female">6 females</specimenCount>
(INPA)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">Figure 2.</emphasis>
<taxonomicName authorityName="Cavalcante-Silva &amp; Pereira &amp; Calor" authorityYear="2022" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="Helicopsyche ralphi" order="Trichoptera" pageId="0" pageNumber="215" phylum="Mollusca" rank="species" species="ralphi">
<emphasis italics="true" pageId="0" pageNumber="215">Helicopsyche ralphi</emphasis>
</taxonomicName>
sp. nov., male
<emphasis bold="true" pageId="0" pageNumber="215">A</emphasis>
genitalia, lateral view
<emphasis bold="true" pageId="0" pageNumber="215">B</emphasis>
segments IX and X and inferior appendages, dorsal view
<emphasis bold="true" pageId="0" pageNumber="215">C</emphasis>
inferior appendage, ventral view
<emphasis bold="true" pageId="0" pageNumber="215">D</emphasis>
phallus, lateral view
<emphasis bold="true" pageId="0" pageNumber="215">E</emphasis>
phallus, ventral view
<emphasis bold="true" pageId="0" pageNumber="215">F</emphasis>
sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a.seg. IX = apodeme of abdominal segment IX; sup.a = superior appendage; seg. X = abdominal segment X; bas.l = basomesal lobe; inf.a = inferior appendage; pha.b = phallobase; pha.scl. = phallotremal sclerite.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="215" type="diagnosis">
<paragraph pageId="0" pageNumber="215">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="215">
The new species is distinguished from all other congeners by the following characters of the male genitalia: inferior appendage subtriangular, acuminated in posterior region, basomesal lobe subtriangular ~ 1/2 the length of the inferior appendage, in lateral view, trapezoid, with spine-like setae in posterior margin, in ventral view; abdominal segment X slender, slightly cleft at the apex, in dorsal view. The characters of the genitalia of new species are morphologically similar to
<taxonomicName authorityName="Johanson" authorityYear="1999" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="Helicopsyche flinti" order="Trichoptera" pageId="0" pageNumber="215" phylum="Mollusca" rank="species" species="flinti">
<emphasis italics="true" pageId="0" pageNumber="215">Helicopsyche flinti</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.1076/aqin.21.2.127.4535" author="Johanson, KA" journalOrPublisher="Aquatic Insects" pageId="0" pageNumber="215" pagination="127 - 132" refId="B36" refString="Johanson, KA, 1999. Description of two new Neotropical Helicopsyche (Trichoptera: Helicopsychidae). Aquatic Insects 21 (2): 127 - 132, DOI: https://doi.org/10.1076/aqin.21.2.127.4535" title="Description of two new Neotropical Helicopsyche (Trichoptera: Helicopsychidae)." url="https://doi.org/10.1076/aqin.21.2.127.4535" volume="21" year="1999">Johanson (1999)</bibRefCitation>
. The new species presents abdominal segment X with a rounded apex, and a medial row of spine-like setae, in dorsal view (while
<taxonomicName genus="H." lsidName="H. flinti" pageId="0" pageNumber="215" rank="species" species="flinti">
<emphasis italics="true" pageId="0" pageNumber="215">H. flinti</emphasis>
</taxonomicName>
presents abdominal segment X with apex nearly straight, side row of spine-like setae), and inferior appendage with strongly projecting mesal margin, forming a large, rounded lobe, in dorsal view (while
<taxonomicName genus="H." lsidName="H. flinti" pageId="0" pageNumber="215" rank="species" species="flinti">
<emphasis italics="true" pageId="0" pageNumber="215">H. flinti</emphasis>
</taxonomicName>
presents an inferior appendage without a large mesal lobe).
</paragraph>
<caption doi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">Figure 3.</emphasis>
<taxonomicName authorityName="Cavalcante-Silva &amp; Pereira &amp; Calor" authorityYear="2022" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="Helicopsyche ralphi" order="Trichoptera" pageId="0" pageNumber="215" phylum="Mollusca" rank="species" species="ralphi">
<emphasis italics="true" pageId="0" pageNumber="215">Helicopsyche ralphi</emphasis>
</taxonomicName>
sp. nov., female
<emphasis bold="true" pageId="0" pageNumber="215">A</emphasis>
genitalia, lateral view
<emphasis bold="true" pageId="0" pageNumber="215">B</emphasis>
genitalia, ventral view
<emphasis bold="true" pageId="0" pageNumber="215">C</emphasis>
genitalia, dorsal view
<emphasis bold="true" pageId="0" pageNumber="215">D</emphasis>
sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e.seg. IX = external part of abdominal segment IX; pre.a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="215" type="description">
<paragraph pageId="0" pageNumber="215">Description.</paragraph>
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">Adults</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4</figureCitation>
): length of forewing 4.1-5.2 mm (
<emphasis italics="true" pageId="0" pageNumber="215">n</emphasis>
= 20). Wings: forewing without discoidal cell, without medial cell, with thyridial cells; hind wing without discoidal cell, without thyridial cell. Head: brownish; antennae yellowish, shorter than forewing length, scape yellowish, shorter than head length, covered with long setae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4E-G</figureCitation>
). Thorax: pronotum brownish, with warts, filiform, covered with small and ferruginous setae; mesoscutum brownish, with mesoscutal warts spherical and not covered with setae; mesoscutellum brownish, with mesoscutellar warts spherical and not covered with setae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4G</figureCitation>
); legs yellowish, tibial spur formula 2,2,4.
</paragraph>
<caption doi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">Figure 4.</emphasis>
<taxonomicName authorityName="Cavalcante-Silva &amp; Pereira &amp; Calor" authorityYear="2022" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="Helicopsyche ralphi" order="Trichoptera" pageId="0" pageNumber="215" phylum="Mollusca" rank="species" species="ralphi">
<emphasis italics="true" pageId="0" pageNumber="215">Helicopsyche ralphi</emphasis>
</taxonomicName>
sp. nov., adult
<emphasis bold="true" pageId="0" pageNumber="215">A</emphasis>
male head, dorsal view
<emphasis bold="true" pageId="0" pageNumber="215">B</emphasis>
male head, frontal view
<emphasis bold="true" pageId="0" pageNumber="215">C</emphasis>
male habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="215">D</emphasis>
female head, dorsal view
<emphasis bold="true" pageId="0" pageNumber="215">E</emphasis>
male head, frontal view
<emphasis bold="true" pageId="0" pageNumber="215">F</emphasis>
female habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="215">G</emphasis>
female habitus, lateral view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">Male</emphasis>
(Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4A-C</figureCitation>
): body length ~ 3.3-4.6 mm (
<emphasis italics="true" pageId="0" pageNumber="215">n</emphasis>
= 20).
</paragraph>
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" italics="true" pageId="0" pageNumber="215">Head</emphasis>
: interantennal warts present, brownish, spherical, covered with small setae; posteroantennal warts present, brownish, club shaped, covered with long setae; cephalic warts present, brownish, subtriangular, covered with long setae; postocular warts present, filiform, brownish, covered with long setae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4A-C</figureCitation>
); maxillary palps yellowish, with two segments, covered with long ferruginous setae; labial palps yellowish, with three segments, covered with long ferruginous setae. Abdomen: abdominal sternum VI process present, almost same length as segment, tubular, apically rounded, covered with small microtrichiae (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2F</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">
<emphasis italics="true" pageId="0" pageNumber="215">Genitalia</emphasis>
.
</emphasis>
Abdominal segment IX with slightly concave anteroventral margin in ventral half; apodeme well developed laterally, located midlaterally on segment; posterior margin nearly straight, in lateral view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2A</figureCitation>
), anterior margin strongly concave, in dorsal view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2B</figureCitation>
); preanal appendages setose, rounded in lateral view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2A</figureCitation>
), clavate in dorsal view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2B</figureCitation>
). Abdominal segment X tubular, dorsal margin slightly curved, in lateral view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2A</figureCitation>
); slender, mesodorsal borders inverted Y-shaped, bearing two rows of short setae, near the center, in dorsal view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2B</figureCitation>
). Inferior appendage subtriangular, acuminated in posterior region, in lateral view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2A</figureCitation>
); anterior margin slightly convex, posterior margin undulated and tapered apex, in dorsal view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2B</figureCitation>
); basomesal lobe of inferior appendage, in lateral view well developed, with ventral margin covered with long setae and dorsal margin with spine-like setae, in ventral view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2C</figureCitation>
). Phallus tubular, phallobase rounded, ventral view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2E</figureCitation>
), acuminate at anteroventral border, in lateral view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2D</figureCitation>
), slightly down curved; phallotremal sclerite conspicuous, moon shaped in lateral view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2D</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">Female</emphasis>
(Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4D-G</figureCitation>
): body length ~ 3.9-5.4 mm (
<emphasis italics="true" pageId="0" pageNumber="215">n</emphasis>
= 20).
</paragraph>
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" italics="true" pageId="0" pageNumber="215">Head</emphasis>
: interantennal warts present, brownish, spherical, covered with small setae; postero-antennal warts present, brownish, covered with long setae; cephalic warts present, brownish, subtriangular, covered with long setae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4D-F</figureCitation>
); postocular warts present, filiform, brownish, covered with long setae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4F</figureCitation>
); maxillary palps yellowish, with 5-segments, covered with long and yellowish setae; labial palps yellowish, with 3-segments, covered with long yellowish setae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4G</figureCitation>
). Abdomen: abdominal sternum VI process present, ~ 1/3 segment length, tubular and apically rounded, covered with small microtrichiae (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3D</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">
<emphasis italics="true" pageId="0" pageNumber="215">Genitalia</emphasis>
.
</emphasis>
Abdominal segment IX is well separated from abdominal segment VIII and indistinctly separated from abdominal segment X, anterior margin convex, in lateral view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3A</figureCitation>
); external part of abdominal segment IX apically incised, in ventral view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3B</figureCitation>
). Preanal appendage long and filiform, in lateral view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3A</figureCitation>
). Abdominal segment X with two branches; dorsal branch narrow, base with apex broad, rounded and covered with long setae, in lateral view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3A</figureCitation>
), bilobed with U-shaped with apical incision, in dorsal view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3C</figureCitation>
); ventral branch with sinuous margin, in lateral view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3A</figureCitation>
), and apex obtuse in ventral and dorsal view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3B, C</figureCitation>
). Vagina with thick anterior margin, in ventral view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3B</figureCitation>
); vaginal sclerite slender along its length, in lateral view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3A</figureCitation>
), finger-shaped projection on the anterior margin, internal sclerite long, with sclerotized lateral margins, in ventral view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">Larva (5th instar)</emphasis>
(Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5A-L</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Helicopsyche ralphi sp. nov., larvae and case A larval case, dorsal view B larval case, ventral view C larval lateral habitus." figureDoi="10.3897/zookeys.1111.77541.figure6" httpUri="https://binary.pensoft.net/fig/712454" pageId="0" pageNumber="215">6C</figureCitation>
): Body total length 2.9-3.6 mm (
<emphasis italics="true" pageId="0" pageNumber="215">n</emphasis>
= 10).
</paragraph>
<caption doi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">Figure 5.</emphasis>
<taxonomicName authorityName="Cavalcante-Silva &amp; Pereira &amp; Calor" authorityYear="2022" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="Helicopsyche ralphi" order="Trichoptera" pageId="0" pageNumber="215" phylum="Mollusca" rank="species" species="ralphi">
<emphasis italics="true" pageId="0" pageNumber="215">Helicopsyche ralphi</emphasis>
</taxonomicName>
sp. nov., larvae
<emphasis bold="true" pageId="0" pageNumber="215">A</emphasis>
head, dorsal view
<emphasis bold="true" pageId="0" pageNumber="215">B</emphasis>
head, ventral view
<emphasis bold="true" pageId="0" pageNumber="215">C</emphasis>
head, left lateral view
<emphasis bold="true" pageId="0" pageNumber="215">D</emphasis>
mandibles, dorsal view
<emphasis bold="true" pageId="0" pageNumber="215">E</emphasis>
labrum, dorsal view
<emphasis bold="true" pageId="0" pageNumber="215">F</emphasis>
notos, dorsal view
<emphasis bold="true" pageId="0" pageNumber="215">G</emphasis>
proleg, lateral view
<emphasis bold="true" pageId="0" pageNumber="215">H</emphasis>
mesoleg, lateral view
<emphasis bold="true" pageId="0" pageNumber="215">I</emphasis>
metaleg, lateral view
<emphasis bold="true" pageId="0" pageNumber="215">J</emphasis>
lateral hump
<emphasis bold="true" pageId="0" pageNumber="215">L</emphasis>
anal legs.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" italics="true" pageId="0" pageNumber="215">Head</emphasis>
: oval, with anterior margin 1.3
<normalizedToken originalValue="×">x</normalizedToken>
broader than posterior margin, in dorsal view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5A</figureCitation>
), mostly light brown, with pale region on anterolateral margin of the head capsule until antenna region, around stemmata, medial region of frontoclypeus margin, and posteromedial region of head capsule, in dorsal view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5A</figureCitation>
), with lateral region light brown, in ventral view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5B</figureCitation>
), cardo and anterior ventral apotome dark brown, six and eight brown muscle scars in left and right, respectively, in dorsal view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5A</figureCitation>
), head capsule with muscle scar light brown in posterolateral region, in dorsal view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5A</figureCitation>
), with frontal area flattened, muscle scars in basal region, cardo and ventral apotome sclerotized, ventral view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5B</figureCitation>
); with muscle scar light brown in posterior region, in lateral view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5C</figureCitation>
); frontoclypeus and adjacent areas nearly flat and margined with semicircular carina, frontoclypeal suture with strongly delimited margin, frontoclypeal with one muscle scar brown in medial region, and three in posterior region, in dorsal view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5A</figureCitation>
); labrum translucent, with short setae covering the anterior margin, in ventral view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5B</figureCitation>
); mandibles asymmetrical, each internal margin with pale, long, thin setae; left mandible with three teeth, the apical tooth trilobed, mesal and basal tooth acute; right mandible with three teeth, the apical tooth trilobed, mesal and basal tooth obtuse, in dorsal view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5D-E</figureCitation>
); chaetotaxy of head as in Figure
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5A-E</figureCitation>
. Thorax: pronotum brown with dark muscle scars, anterior region with row of long setae at margin, covered long setae to near medial region, posterior margin sinuous and lighter with few and scattered setae, in dorsal view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5F</figureCitation>
), trochantin almost as long as foreleg coxae, finger shaped with one spinelike setae in apex (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5G</figureCitation>
); mesonotum lighter than pronotum, with pale regions in medial region, muscle scars in dark brown shades, four pairs in anterior region and one pair in posterior region, posterior margins angulate, in dorsal view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5F</figureCitation>
); metanotum with three pairs of sclerites, two pairs of anteromesal (sa1) sclerites small, one pairs anterior subtriangular, and one pairs irregular bearing one setae and one pair of posterior subtriangular sclerites (fused sa2 and sa3 sclerites), each bearing single seta posteromesally (sa2) and several setae anterolateral (sa3), in dorsal view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5F</figureCitation>
); lateral hump oval, apical region mostly dark, one short setae in anteromedial region (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5J</figureCitation>
); thoracic legs with chaetotaxy as in Figure
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5G-I</figureCitation>
; The foreleg has a length equivalent to 2/3 of the midleg and 1/2 of the hindleg, foreleg segments robust and short, mid and hind leg segments filiform and long (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5I</figureCitation>
). Abdomen: anal prolegs each with lateral sclerites curved, anal claw elongates, with accessory parallel teeth pectinate, arranged like comb (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5L</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" italics="true" pageId="0" pageNumber="215">Larval case</emphasis>
(Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Helicopsyche ralphi sp. nov., larvae and case A larval case, dorsal view B larval case, ventral view C larval lateral habitus." figureDoi="10.3897/zookeys.1111.77541.figure6" httpUri="https://binary.pensoft.net/fig/712454" pageId="0" pageNumber="215">6A-B</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Helicopsyche ralphi sp. nov., pupa and case A pupa front B abdominal segment IX and anal processes, dorsal view C pupa lateral habitus D-H abdominal segments I-V, dorsal, with details of dorsal hook plates I pupa case, ventral view J pupa case with sieve membrane highlighted, ventral view." figureDoi="10.3897/zookeys.1111.77541.figure7" httpUri="https://binary.pensoft.net/fig/712455" pageId="0" pageNumber="215">7I-J</figureCitation>
): length 2.9-3.6 mm (
<emphasis italics="true" pageId="0" pageNumber="215">n</emphasis>
= 10). Made with cemented sand grains, forming a snail-like, helical case, case with two 1/2 whorls at the end of the phase, with umbilicus open and deep.
</paragraph>
<caption doi="10.3897/zookeys.1111.77541.figure6" httpUri="https://binary.pensoft.net/fig/712454" pageId="0" pageNumber="215" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">Figure 6.</emphasis>
<taxonomicName authorityName="Cavalcante-Silva &amp; Pereira &amp; Calor" authorityYear="2022" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="Helicopsyche ralphi" order="Trichoptera" pageId="0" pageNumber="215" phylum="Mollusca" rank="species" species="ralphi">
<emphasis italics="true" pageId="0" pageNumber="215">Helicopsyche ralphi</emphasis>
</taxonomicName>
sp. nov., larvae and case
<emphasis bold="true" pageId="0" pageNumber="215">A</emphasis>
larval case, dorsal view
<emphasis bold="true" pageId="0" pageNumber="215">B</emphasis>
larval case, ventral view
<emphasis bold="true" pageId="0" pageNumber="215">C</emphasis>
larval lateral habitus.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1111.77541.figure7" httpUri="https://binary.pensoft.net/fig/712455" pageId="0" pageNumber="215" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">Figure 7.</emphasis>
<taxonomicName authorityName="Cavalcante-Silva &amp; Pereira &amp; Calor" authorityYear="2022" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="Helicopsyche ralphi" order="Trichoptera" pageId="0" pageNumber="215" phylum="Mollusca" rank="species" species="ralphi">
<emphasis italics="true" pageId="0" pageNumber="215">Helicopsyche ralphi</emphasis>
</taxonomicName>
sp. nov., pupa and case
<emphasis bold="true" pageId="0" pageNumber="215">A</emphasis>
pupa front
<emphasis bold="true" pageId="0" pageNumber="215">B</emphasis>
abdominal segment IX and anal processes, dorsal view
<emphasis bold="true" pageId="0" pageNumber="215">C</emphasis>
pupa lateral habitus
<emphasis bold="true" pageId="0" pageNumber="215">D-H</emphasis>
abdominal segments I-V, dorsal, with details of dorsal hook plates
<emphasis bold="true" pageId="0" pageNumber="215">I</emphasis>
pupa case, ventral view
<emphasis bold="true" pageId="0" pageNumber="215">J</emphasis>
pupa case with sieve membrane highlighted, ventral view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" italics="true" pageId="0" pageNumber="215">Pupa</emphasis>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Helicopsyche ralphi sp. nov., pupa and case A pupa front B abdominal segment IX and anal processes, dorsal view C pupa lateral habitus D-H abdominal segments I-V, dorsal, with details of dorsal hook plates I pupa case, ventral view J pupa case with sieve membrane highlighted, ventral view." figureDoi="10.3897/zookeys.1111.77541.figure7" httpUri="https://binary.pensoft.net/fig/712455" pageId="0" pageNumber="215">7A-J</figureCitation>
): body length 3.3-4.1 mm (
<emphasis italics="true" pageId="0" pageNumber="215">n</emphasis>
= 10). Generally dark brown, almost black, with yellowish abdomen; Head: mandibles curved with wide bases, each with length 2.5
<normalizedToken originalValue="×">x</normalizedToken>
basal width, apex pointed and internal margin smooth (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Helicopsyche ralphi sp. nov., pupa and case A pupa front B abdominal segment IX and anal processes, dorsal view C pupa lateral habitus D-H abdominal segments I-V, dorsal, with details of dorsal hook plates I pupa case, ventral view J pupa case with sieve membrane highlighted, ventral view." figureDoi="10.3897/zookeys.1111.77541.figure7" httpUri="https://binary.pensoft.net/fig/712455" pageId="0" pageNumber="215">7A</figureCitation>
). Abdomen: paired anterior dorsal hook plates on segments II-V, pair of posterior dorsal hook plates on segment IV asymmetrical; general morphology of dorsal hook plates as in (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5C-H</figureCitation>
); terminal abdominal segment rounded, with two divergent digitate processes, each process bearing one subapical and three apical setae (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Helicopsyche ralphi sp. nov., pupa and case A pupa front B abdominal segment IX and anal processes, dorsal view C pupa lateral habitus D-H abdominal segments I-V, dorsal, with details of dorsal hook plates I pupa case, ventral view J pupa case with sieve membrane highlighted, ventral view." figureDoi="10.3897/zookeys.1111.77541.figure7" httpUri="https://binary.pensoft.net/fig/712455" pageId="0" pageNumber="215">7B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="215" type="etymology">
<paragraph pageId="0" pageNumber="215">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="215">This species is named in honor of Dr. Ralph W. Holzenthal, for his outstanding contribution to the study of Neotropical caddisflies, and also as an acknowledgment for collaboration and his contributing to the training of young researchers.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="215" type="materials_examined">
<paragraph pageId="0" pageNumber="215">Material additional.</paragraph>
<paragraph pageId="0" pageNumber="215">
A1 (
<specimenCount type="female">1 female</specimenCount>
), C6 (
<specimenCount type="male">643 males</specimenCount>
); C7 (
<specimenCount type="larva">10 larvae</specimenCount>
,
<specimenCount type="pupa">10 pupae</specimenCount>
); D2 (
<specimenCount type="male">34 males</specimenCount>
); D7 (
<specimenCount type="male">1 male</specimenCount>
); D17 (
<specimenCount type="male">14 males</specimenCount>
); D18 (
<specimenCount type="male">379 males</specimenCount>
); D18 (
<specimenCount type="female">49 females</specimenCount>
); D20 (
<specimenCount type="female">32 females</specimenCount>
); D22 (
<specimenCount type="larva">1 larvae</specimenCount>
); E1 (
<specimenCount type="male">1 male</specimenCount>
); F2 (
<specimenCount type="male">30 males</specimenCount>
); F3 (
<specimenCount type="male">17 males</specimenCount>
) (UFBA) (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Collection data from Brejo de Altitude de Triunfo, with the sample of each collection site, geographic coordinates, elevation, date, traps (LPT = UV Light Pan Trap, MAL = Malaise and WSA = White Sheet Attraction), and acronyms of collectors." httpUri="http://table.plazi.org/id/6FB051B14B382AEF1FDDD28788D65E4B" pageId="0" pageNumber="215" tableUuid="6FB051B14B382AEF1FDDD28788D65E4B">1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="215" type="distribution">
<paragraph pageId="0" pageNumber="215">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="215">Brazil (Pernambuco state).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="215" type="caddisflies from brejo de altitude de triunfo">
<paragraph pageId="0" pageNumber="215">Caddisflies from Brejo de Altitude de Triunfo</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="215" type="hydropsychidae">
<paragraph pageId="0" pageNumber="215">
<taxonomicName authorityName="J Curtis" authorityYear="1835" class="Insecta" family="HYDROPSYCHIDAE" kingdom="Animalia" lsidName="" order="Trichoptera" pageId="0" pageNumber="215" phylum="Arthropoda" rank="family">HYDROPSYCHIDAE</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>