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Nixon, 1937
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<paragraph id="32E1A11499177A925A024BC300729102" pageId="0" pageNumber="8151">Distribution</paragraph>
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.108.768" ID-GBIF-Dataset="283587db-b3b0-4b45-9060-0d6c74a4559b" ID-PMC="PMC3119315" ID-Pensoft-Pub="1313-2970-108-49" ID-PubMed="21852927" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-108-49" ModsDocOrigin="ZooKeys 108" ModsDocTitle="Notes on the genus Ismarus Haliday (Hymenoptera, Diapriidae) from China" checkinTime="1451250273262" checkinUser="pensoft" docAuthor="Liu, Jing-xian, Chen, Hua-yan &amp; Xu, Zai-fu" docDate="2011" docId="D0723C4328CE60D26F2DD7277B42BBED" docLanguage="en" docName="ZooKeys 108: 49-60" docOrigin="ZooKeys 108" docSource="http://dx.doi.org/10.3897/zookeys.108.768" docTitle="Ismarus parvicellus Liu, Chen &amp; Xu, sp. n." docType="treatment" docVersion="4" lastPageNumber="55" masterDocId="FFFFD528FFD18755FFECFFBC1E43FFA9" masterDocTitle="Notes on the genus Ismarus Haliday (Hymenoptera, Diapriidae) from China" masterLastPageNumber="60" masterPageNumber="49" pageNumber="54" updateTime="1668151456380" updateUser="ExternalLinkService">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:18E038AE-DB02-4891-95E0-5A7902BAFCE4" authority="Liu, Chen &amp; Xu" class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus parvicellus" order="Hymenoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="parvicellus">Ismarus parvicellus Liu, Chen &amp; Xu</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="54">sp. n.</taxonomicNameLabel>
Figs 9-12
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<paragraph pageId="5" pageNumber="54">Holotype.</paragraph>
<paragraph pageId="5" pageNumber="54">Female. Body length 2.2 mm; fore wing length 1.8 mm.</paragraph>
<paragraph pageId="5" pageNumber="54">Head. Head in dorsal view 2.0 times as wide as long; vertex abruptly sloping behind post ocelli; temple narrowed behind eyes; occipital carina strong and complete, not crenulate; POL as long as OOL; toruli separated from each other; face setose; clypeus evenly convex; epistomal sulcus distinct; eye height 7.0 times length of malar space; malar sulcus absent. Scape cylindrical, with apical rim simple; first to third flagellomeres attenuate basally, and gradually incrassate to apex; ratios of length to width of antennal segments: 20: 5; 8: 5; 11: 4; 11: 4; 9: 4; 8: 5; 8: 5; 7: 6; 7: 6; 7: 6; 6: 6; 6: 6; 6: 6; 6: 5; 10: 5.</paragraph>
<paragraph pageId="5" pageNumber="54">Mesosoma. Mesosoma in dorsal view 0.75 times as long as width of head; pronotum angular in dorsal view, anteriorly punctate and setose; lateral side of pronotum with anterior and upper margins coarsely punctate, lower half rugose punctate, posterior upper part smooth; mesoscutum smooth and convex; notauli present as small pits on anterior face; humeral sulcus strong, 1.4 times length of tegula; anterior scutellar pit transverse, crenulate inside, median longitudinal carina weak; scutellum smooth, posterior rim weakly concave and subtruncate; propodeum rugose and punctate, with transverse carinae and longitudinal carinae strong; mesopleuron mostly smooth and bare, with upper corner below tegula finely punctate and setose; metapleuron rugose and setose.</paragraph>
<paragraph pageId="5" pageNumber="54">Wings. Fore wing with costal, subcostal, basal, marginal, stigmal and postmarginal veins tubular; distal part of median and cubital veins weakly pigmented; radial cell closed, 0.3 times length of marginal vein and 2.3 times as long as its height. Hind wing with a basal cell.</paragraph>
<paragraph pageId="5" pageNumber="54">Legs. Fore and middle legs slender; hind tibia strongly incrassate.</paragraph>
<paragraph pageId="5" pageNumber="54">Metasoma. Petiole transverse, rugose and with irregular longitudinal carinae; second tergite mostly smooth and scattered with a few setae along lower side, median furrow short, extending to 0.2 times length of second tergite; sutures between tergites complete and well impressed.</paragraph>
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<paragraph pageId="5" pageNumber="54">Colour.</paragraph>
<paragraph pageId="5" pageNumber="54">Body black. Antenna yellowish brown but seventh to eighth flagellomeres blackish brown and ninth to thirteenth flagellomeres black. Legs brown. Wings hyaline, veins brown.</paragraph>
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<paragraph pageId="5" pageNumber="54">Male.</paragraph>
<paragraph pageId="5" pageNumber="54">Unknown.</paragraph>
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<paragraph pageId="5" pageNumber="54">Distribution.</paragraph>
<paragraph pageId="5" pageNumber="54">China (Hainan).</paragraph>
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<paragraph pageId="5" pageNumber="54">Material examined.</paragraph>
<paragraph pageId="5" pageNumber="54">
Holotype, female. CHINA: Hainan, Baisha County, Mt. Jiujialing (
<geoCoordinate direction="north" orientation="latitude" precision="555" value="19.21">19.21°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="109.45">109.45°E</geoCoordinate>
), 2010.VII.18, YPT, Hua-yan Chen, No. 20100013.
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<paragraph pageId="5" pageNumber="54">Diagnosis.</paragraph>
<paragraph lastPageId="6" lastPageNumber="55" pageId="5" pageNumber="54">
This species belongs to the
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-group and can be distinguished from
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by having the eye height 7.0 times length of the malar space (4.5 times in
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); the radial cell of the fore wing small and 0.3 times length of marginal
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(radial cell as long as marginal vein); the distal part of the median and cubital veins weakly pigmented (median and cubital veins distinctly pigmented); the posterior rim of the scutellum weakly concave and subtruncate (posterior rim of scutellum round).
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<paragraph pageId="6" pageNumber="55">Etymology.</paragraph>
<paragraph pageId="6" pageNumber="55">
The specific name is derived from the Latin adjective
<normalizedToken originalValue="parvi">'parvi'</normalizedToken>
(small) and
<normalizedToken originalValue="cell">'cell'</normalizedToken>
, referring to the small radial cell of fore wing.
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<paragraph pageId="6" pageNumber="55">
Figures 9-12.
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sp. n. 9 habitus, lateral view 10 head and thorax, dorsal view 11 antenna 12 fore wing.
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<paragraph id="586436FDFFC5FFFCE5FFF9D97963F99A" blockId="18.[151,541,1583,1642]" box="[151,314,1618,1642]" pageId="18" pageNumber="19">
(
<figureCitation id="C0E02A78FFC5FFFCE5F6F9D97968F99A" box="[158,305,1618,1642]" captionStart-0="PLATE 20" captionStart-1="PLATE 21" captionStart-10="PLATE 30" captionStart-2="PLATE 22" captionStart-3="PLATE 23" captionStart-4="PLATE 24" captionStart-5="PLATE 25" captionStart-6="PLATE 26" captionStart-7="PLATE 27" captionStart-8="PLATE 28" captionStart-9="PLATE 29" captionStartId-0="68.[151,233,1655,1678]" captionStartId-1="70.[151,233,1594,1617]" captionStartId-10="88.[151,233,1655,1678]" captionStartId-2="72.[151,233,1625,1648]" captionStartId-3="74.[151,233,1502,1525]" captionStartId-4="76.[151,233,1686,1709]" captionStartId-5="78.[151,233,1594,1617]" captionStartId-6="80.[151,233,1717,1740]" captionStartId-7="82.[151,233,1625,1648]" captionStartId-8="84.[151,233,1655,1678]" captionStartId-9="86.[151,233,1747,1770]" captionText-0="PLATE 20. Protopliomerella kerouaci n. sp., from Section HC6 202T m and 203.5 m, Garden City Formation (lower Floian; Tulean; Psalikilus typicum Zone), west side of Hillyard Canyon, Bear River Range, Franklin County, southeastern Idaho, USA." captionText-1="PLATE 21. Protopliomerella kerouaci n. sp., from Section HC6 202T m, 203.0 m, and 203.5 m, Garden City Formation (lower Floian; Tulean; Psalikilus typicum Zone), west side of Hillyard Canyon, Bear River Range, Franklin County, southeastern Idaho, USA." captionText-10="PLATE 30. Protopliomerella kerouaci n. sp., from Section G 168T m and G 174.0 m, Fillmore Formation (lower Floian; Tulean; Psalikilus typicum Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." captionText-2="PLATE 22. Protopliomerella kerouaci n. sp., from Section HC6 202T m and 203.0 m, Garden City Formation (lower Floian; Tulean; Psalikilus typicum Zone), west side of Hillyard Canyon, Bear River Range, Franklin County, southeastern Idaho, USA." captionText-3="PLATE 23. Protopliomerella kerouaci n. sp., from Section HC6 202T m, 203.0 m, and 203.5 m, Garden City Formation (lower Floian; Tulean; Psalikilus typicum Zone), west side of Hillyard Canyon, Bear River Range, Franklin County, southeastern Idaho, USA." captionText-4="PLATE 24. Protopliomerella kerouaci n. sp., from Section HC6 203.5 m, Garden City Formation (lower Floian; Tulean; Psalikilus typicum Zone), west side of Hillyard Canyon, Bear River Range, Franklin County, southeastern Idaho, USA." captionText-5="PLATE 25. Protopliomerella kerouaci n. sp., from Section HC6 202T m and 203.0 m, Garden City Formation (lower Floian; Tulean; Psalikilus typicum Zone), west side of Hillyard Canyon, Bear River Range, Franklin County, southeastern Idaho, USA." captionText-6="PLATE 26. Protopliomerella kerouaci n. sp., from Section G 168T m, Fillmore Formation (lower Floian; Tulean; Psalikilus typicum Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." captionText-7="PLATE 27. Protopliomerella kerouaci n. sp., from Section G 168T m, Fillmore Formation (lower Floian; Tulean; Psalikilus typicum Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." captionText-8="PLATE 28. Protopliomerella kerouaci n. sp., from Section G 168T m, Fillmore Formation (lower Floian; Tulean; Psalikilus typicum Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." captionText-9="PLATE 29. Protopliomerella kerouaci n. sp., from Section G 168T m, Fillmore Formation (lower Floian; Tulean; Psalikilus typicum Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." pageId="18" pageNumber="19">Plates 2030</figureCitation>
)
</paragraph>
<paragraph id="586436FDFFC5FFFCE5A5F91E7DC5F8F7" blockId="18.[205,1436,1685,1861]" pageId="18" pageNumber="19">
1951
<taxonomicName id="9FDB4D7EFFC5FFFCE471F91E7A3AF95B" authority="Ross" authorityName="Ross" authorityYear="1951" box="[281,611,1685,1707]" class="Trilobita" family="Pliomeridae" genus="Protopliomerops" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="contracta">
<emphasis id="6AAFEAEFFFC5FFFCE471F91E7A73F95B" box="[281,554,1685,1707]" italics="true" pageId="18" pageNumber="19">Protopliomerops contracta</emphasis>
Ross
</taxonomicName>
, p. 136, pl. 33, figs 18, 19, 2332 (only; pl. 33, fig. 15 =
<taxonomicName id="9FDB4D7EFFC5FFFCE187F91E793BF939" authorityName="Adrain" authorityYear="2011" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="seegeri">
<emphasis id="6AAFEAEFFFC5FFFCE187F91E793BF939" italics="true" pageId="18" pageNumber="19">Protopliomerella seegeri</emphasis>
</taxonomicName>
; pl. 33, figs 16, 17, 22 =
<taxonomicName id="9FDB4D7EFFC5FFFCE70AF93F7A85F939" authorityName="Ross" authorityYear="1951" box="[610,732,1716,1737]" class="Trilobita" family="Pliomeridae" genus="Protopliomerops" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="contracta">
<emphasis id="6AAFEAEFFFC5FFFCE70AF93F7A85F939" box="[610,732,1716,1737]" italics="true" pageId="18" pageNumber="19">P. contracta</emphasis>
</taxonomicName>
). 1953
<taxonomicName id="9FDB4D7EFFC5FFFCE471F95979BFF918" box="[281,486,1746,1768]" class="Trilobita" family="Pliomeridae" genus="Protopliomerops" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="6AAFEAEFFFC5FFFCE471F959799AF918" box="[281,451,1746,1768]" italics="true" pageId="18" pageNumber="19">Protopliomerops</emphasis>
sp.
</taxonomicName>
indet.; Hintze, p. 31. 1973 Genus and species undetermined A; Demeter, p. 64, pl. 5, fig. 15 (only; pl. 5, figs 24, 7, 8 =
<taxonomicName id="9FDB4D7EFFC5FFFCE1A3F97A7D63F8F7" authorityName="Poulsen" authorityYear="1927" box="[1227,1338,1777,1799]" class="Trilobita" family="Pliomeridae" genus="Cybelopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="6AAFEAEFFFC5FFFCE1A3F97A7D63F8F7" box="[1227,1338,1777,1799]" italics="true" pageId="18" pageNumber="19">Cybelopsis</emphasis>
</taxonomicName>
? spp.; pl.
</paragraph>
<paragraph id="586436FDFFC5FFFCE471F89B7B29F8B4" blockId="18.[205,1436,1685,1861]" pageId="18" pageNumber="19">
5, fig. 9 =
<taxonomicName id="9FDB4D7EFFC5FFFCE4EBF89B7A26F8D6" authorityName="Adrain" authorityYear="2011" box="[387,639,1808,1830]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="seegeri">
<emphasis id="6AAFEAEFFFC5FFFCE4EBF89B7A26F8D6" box="[387,639,1808,1830]" italics="true" pageId="18" pageNumber="19">Protopliomerella seegeri</emphasis>
</taxonomicName>
; pl. 5, fig. 13 = gen. and sp. indet.). 2009
<taxonomicName id="9FDB4D7EFFC5FFFCE471F8A4799CF8B5" authority="Adrain, 2011" authorityName="Adrain" authorityYear="2011" box="[281,453,1839,1861]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="6AAFEAEFFFC5FFFCE471F8A4799CF8B5" box="[281,453,1839,1861]" italics="true" pageId="18" pageNumber="19">Protopliomerella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="719C5794FFC5FFFCE4A4F8A47A43F8B4" box="[460,538,1839,1860]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
2; Adrain
<emphasis id="6AAFEAEFFFC5FFFCE7EEF8A47AE3F8B5" box="[646,698,1839,1861]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
, p. 561, fig.
<specimenCount id="4EDDFD74FFC5FFFCE653F8A57B29F8B4" box="[827,880,1838,1860]" pageId="18" pageNumber="19" type="generic">11M.</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFC5FFFDE5FFF8E07C52FF07" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="586436FDFFC5FFFDE5FFF8E07C52FF07" blockId="18.[151,1436,1899,2032]" lastBlockId="19.[151,1437,151,2023]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="6AAFEAEFFFC5FFFCE5FFF8E0795FF874" bold="true" box="[151,262,1899,1924]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">Material.</emphasis>
<typeStatus id="8760885FFFC5FFFCE472F8E779DCF874" box="[282,389,1900,1924]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
, cranidium,
<collectionCode id="3ECAAE38FFC5FFFCE77EF8E77A1AF874" box="[534,579,1900,1924]" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="18" pageNumber="19" type="University or college">SUI</collectionCode>
126209, from Section G 168
<collectionCode id="3ECAAE38FFC5FFFCE6F1F8E77BF2F874" box="[921,939,1900,1924]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="19">T</collectionCode>
m, and assigned specimens
<collectionCode id="3ECAAE38FFC5FFFCE190F8E77D7CF874" box="[1272,1317,1900,1924]" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="18" pageNumber="19" type="University or college">SUI</collectionCode>
126206 126208, 126210126246 from Section G 168
<collectionCode id="3ECAAE38FFC5FFFCE7FAF8047AFDF857" box="[658,676,1935,1959]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="19">T</collectionCode>
m and 174.0 m, Fillmore Formation (lower Floian; Tulean;
<taxonomicName id="9FDB4D7EFFC5FFFCE02DF81B793FF83C" authority="Zone" authorityName="Zone" class="Trilobita" family="Hystricuridae" genus="Psalikilus" kingdom="Animalia" order="Aulacopleurida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="typicum">
<emphasis id="6AAFEAEFFFC5FFFCE02DF81B7942F83C" italics="true" pageId="18" pageNumber="19">Psalikilus typicum</emphasis>
Zone
</taxonomicName>
), southern Confusion Range, Ibex area, Millard County, western
<collectingRegion id="9A1FF81FFFC5FFFCE105F83F7CFFF83C" box="[1133,1190,1972,1996]" country="United States of America" name="Utah" pageId="18" pageNumber="19">Utah</collectingRegion>
,
<collectingCountry id="20CC766DFFC5FFFCE1DEF83F7CACF83C" box="[1206,1269,1972,1996]" name="United States of America" pageId="18" pageNumber="19">USA</collectingCountry>
.;
<collectionCode id="3ECAAE38FFC5FFFCE061F83F7D6FF83C" box="[1289,1334,1972,1996]" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="18" pageNumber="19" type="University or college">SUI</collectionCode>
115214, 126146126205 from Section HC6 202
<collectionCode id="3ECAAE38FFC5FFFCE739F8537A3BF800" box="[593,610,2008,2032]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="19">T</collectionCode>
m, 203.0 m, and 203.5 m, Garden City Formation (lower Floian; Tulean;
<taxonomicName id="9FDB4D7EFFC4FFFDE5FFFF1379F4FF5F" authority="Zone" authorityName="Zone" box="[151,429,151,176]" class="Trilobita" family="Hystricuridae" genus="Psalikilus" kingdom="Animalia" order="Aulacopleurida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="typicum">
<emphasis id="6AAFEAEFFFC4FFFDE5FFFF13793EFF5F" box="[151,359,152,176]" italics="true" pageId="19" pageNumber="20">Psalikilus typicum</emphasis>
Zone
</taxonomicName>
), west side of Hillyard Canyon, Bear River Range, Franklin County, southeastern
<collectingRegion id="9A1FF81FFFC4FFFDE03DFF1C7DCEFF5F" box="[1365,1431,151,175]" country="United States of America" name="Idaho" pageId="19" pageNumber="20">Idaho</collectingRegion>
,
<collectingCountry id="20CC766DFFC4FFFDE5FFFF37788AFF24" box="[151,211,188,212]" name="United States of America" pageId="19" pageNumber="20">USA</collectingCountry>
. Also known from Locality YH E, Yellow Hill Limestone (lower Floian; Tulean; high
<taxonomicName id="9FDB4D7EFFC4FFFDE1C3FF377979FF07" authority="Zone" authorityName="Zone" class="Trilobita" family="Bathyuridae" genus="Psalikilopsis" kingdom="Animalia" order="Proetida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="cuspidicauda">
<emphasis id="6AAFEAEFFFC4FFFDE1C3FF377882FF08" italics="true" pageId="19" pageNumber="20">Psalikilopsis cuspidicauda</emphasis>
Zone
</taxonomicName>
), Yellow Hill, near Pioche, Lincoln County, eastern
<collectingRegion id="9A1FF81FFFC4FFFDE60FFF547BE6FF07" box="[871,959,223,247]" country="United States of America" name="Nevada" pageId="19" pageNumber="20">Nevada</collectingRegion>
,
<collectingCountry id="20CC766DFFC4FFFDE6A3FF547C5EFF07" box="[971,1031,223,247]" name="United States of America" pageId="19" pageNumber="20">USA</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFC4FFFDE5AFFE8F7A64FEEB" box="[199,573,260,284]" pageId="19" pageNumber="20" type="etymology">
<paragraph id="586436FDFFC4FFFDE5AFFE8F7A64FEEB" blockId="19.[151,1437,151,2023]" box="[199,573,260,284]" pageId="19" pageNumber="20">
<emphasis id="6AAFEAEFFFC4FFFDE5AFFE8F7914FEEC" bold="true" box="[199,333,260,284]" inLineHeading="true" pageId="19" pageNumber="20" reason="1">Etymology.</emphasis>
After Jack Kerouac.
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFC4FFFDE5AFFEAC7D47FE94" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="586436FDFFC4FFFDE5AFFEAC7D47FE94" blockId="19.[151,1437,151,2023]" pageId="19" pageNumber="20">
<emphasis id="6AAFEAEFFFC4FFFDE5AFFEAC7918FEB0" bold="true" box="[199,321,295,320]" inLineHeading="true" pageId="19" pageNumber="20" reason="1">Diagnosis.</emphasis>
Fixigenae with sparse granules concentrated around edges of pits; thoracic segments dorsally effaced; pygidium nearly effaced, with transverse line of pits on axial rings and down center of pleurae.
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFC4FFFBE5AFFEE478B3F8D8" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20" type="description">
<paragraph id="586436FDFFC4FFFDE5AFFEE47D61F9CE" blockId="19.[151,1437,151,2023]" pageId="19" pageNumber="20">
<emphasis id="6AAFEAEFFFC4FFFDE5AFFEE47901FE78" bold="true" box="[199,344,367,392]" inLineHeading="true" pageId="19" pageNumber="20" reason="1">Description.</emphasis>
Cranidium with strongly vaulted fixigenae, moderately vaulted axis, narrow anteriorly and very wide posteriorly, with sagittal length 48.4% (44.652.9%) width across genal angles; anterior border narrow, short, subrectangular, very slightly anteriorly curved, moderately inflated, with U-shaped cross-section, densely granulose, more effaced dorsally, with line of pits at mid-length echoing curvature; doublure expressed as anterior face, exposes bottom half of outer face of inner wall of border, scalloped with shallow median arc of rostral suture, only a rim ventrally; anterior border furrow short, very deep, incised, with short anterolateral branches angled at about 30º above horizontal and broad median section with slight anterior convexity; glabella long, narrow, strongly anteriorly tapered, with maximum width across L1 94.9% (81.6106.1%) sagittal length, moderately arched, with convexity higher posteriorly, with four well-defined lateral lobes, and with dense granulose sculpture medially (effaced on larger specimens) and some perforate tubercles or pits, lateral lobes with sculpture extending exsagittally from middle lobe along mid-length and mid-width, but surrounded by short and narrow effaced rim on dorsal face (lobes totally effaced on larger specimens), lobes also outlined by granules on edge extending into lateral and axial furrows; L1 largest, subtriangular, with L2 and L3 slightly shorter and sub-square, L4 short, narrow, triangular, median lobe long, narrow, roughly rectangular, LF just a very short arc behind anterior border furrow; all lateral furrows short, deep, and incised, progressively narrower anteriorly, with S4 about 1/3 width of S1, S1S3 strongly anterolaterally directed, about 30-40º above horizontal, S1 with slightly elongated adaxial end, S4 very strongly anteromedially directed about 50º above horizontal, branching from near exsagittal tip of S3; SO short, slightly longer medially and behind outer halves of L1, deep, shallower medially and deeper laterally, with gently wavy course curved behind L1, lined anteriorly and posteriorly with granules just inside rim; LO wide, about as long as L2, rectangular, with slightly wavy anterior margin, effaced; doublure fairly short, slightly laterally tapered, with a fine transverse ridge at midlength; axial furrows moderately wide, wider at triangular intersections with lateral glabellar furrows, very deep, subparallel along LO, bowed outward around L1, then steadily anteriorly convergent, framed by granulose margins of glabella and fixigenae; palpebro-ocular ridges opposite L3LF, short, narrow, with strongly arcuate outer margin and very narrow, rapidly tapered posterior ridge, highly elevated at about 5060º above horizontal (anterior view), densely granulose; palpebro-ocular furrows narrow, deep, incised, except effaced along extreme anteromedian section of ridge, with gently sigmoid course; interocular fixigenae triangular, narrow, short, a little less than 1/3 maximum length of fixigena; posterior fixigenae long, wide, subrectangular, with broadly curved anterior margin, all fixigena with fairly sparse sculpture of tiny granules, concentrated only on rims of pits in larger specimens, densely scattered small pits, and effaced zone along axial and posterior border furrows; posterior border furrow short, slightly longer toward genal angle, then constricted again, deep, incised, roughly transverse until mid-course, then anteriorly curved (lateral view), rimmed with granules anteriorly and posteriorly; posterior border short medially, expanded laterally to maximum at genal angle, then strongly tapered to point at tip, moderately inflated, mostly effaced, but with scattered granules on posterior edge (concentrated near genal angle), and small nubby spine at genal angle; doublure very short adaxially, turned outward into articulating tongue near axis, longer and wider toward genal angle, cut shallowly by facial suture at anterior corner of genal angle.
</paragraph>
<paragraph id="586436FDFFC4FFFDE5AFF9C27995F991" blockId="19.[151,1437,151,2023]" box="[199,460,1609,1633]" pageId="19" pageNumber="20">Rostral plate unknown.</paragraph>
<paragraph id="586436FDFFC4FFFAE5AFF9E67AEEFE3B" blockId="19.[151,1437,151,2023]" lastBlockId="20.[151,1437,151,2007]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">Hypostome narrow and elongate, with width across shoulders 70.2% (65.873.0%) length; hypostomal suture broad, very slightly anteriorly convex; anterior border vanishingly short, especially medially, nearly overhung by middle body, expanded laterally into long, fairly wide, wedge-shaped anterior wings with small, deep wing process pits located very close to lateral margin, with granulose sculpture and some larger granules medially; anterior bor- der furrow narrow arc, strongly anteriorly bowed, extremely short, especially medially, deep anteriorly, then abruptly shallowed at connection with lateral border furrow; anterior lobe of middle body long, wide, ovoid, strongly ventrally inflated, with inflation decreasing anteriorly and posteriorly from maximum about even with anterior end of lateral notch (lateral view), with sculpture of dense tiny tubercles over anterior half, becoming much less dense posteriorly; middle body furrow ill-defined, moderately long, shallow, strongly posteriorly convex, merged with lateral border furrow at about half length of middle body; posterior lobe of middle body U-shaped, short, tapered anterolaterally, somewhat inflated, with inflation decreasing posteriorly, effaced; lateral border furrows narrow, deep, gently laterally bowed, anteriorly and posteriorly convergent; lateral border very narrow at lateral notch, much expanded at shoulders, and gradually expanded posteriorly to maximum width at posterolateral corners, moderately downturned (lateral view), densely granulose, with small nubby spines at shoulders, small pointed spines between shoulders and posterolateral corners, and larger pointed spines at corners; posterior border furrow moderately long and deep medially, short and shallow laterally, strongly posteriorly convex; posterior bor- der long, narrow, strongly downturned (lateral view), with granulose sculpture and short, triangular median spine; doublure long and wide, reaches slightly inside lateral border furrows and nearly to posterior border furrow, rotated anterolaterally at lateral notch out into anterior wings, posterior section moderately raised and posteroventrally sloped, lateral notch broad and moderately deep.</paragraph>
<paragraph id="586436FDFFC3FFFAE5AFFE5C796DFC17" blockId="20.[151,1437,151,2007]" pageId="20" pageNumber="21">Librigena long and narrow, strongly posteriorly tapered, wedge-shaped with moderate lateral curvature (ventrolateral view); anterior branch of facial suture very short along eye and field, sharply posteriorly angled to make narrow angle with border, then long and steeply anteroventrally sloped to end of anterior projection of border; posterior branch of facial suture very long, steeply sloped from eye to about mid-length of librigenal field, then more shallowly sloped down to border (about 35º and 15º below horizontal, respectively); eye small, elliptical, moderately raised above field on socle (ventrolateral view); librigenal field fairly wide anteriorly, strongly tapered posteriorly, with width of field under midpoint of eye 17.0% (14.519.4%) length of lateral border furrow, moderately ventrolaterally and anteroposteriorly sloped, with dense sculpture of tiny granules and moderately densely distributed small pits; lateral border furrow narrow, deep, deeper anteriorly and slightly shallower near posterior end, very gently laterally bowed and posteriorly sloped in course; lateral border very wide, posteriorly tapered to fine point, with long anterior and posterior projections, anterior anteroventrally rotated, with bluntly squared termination, posterior triangular, rapidly tapered and strongly posterodorsally curved, border well inflated, with dense sculpture of medium granules; doublure wide, reaches about 3/4 to lateral border furrow, slightly tapered anteriorly, then terminated as border rotates to form anterior projection, with inner face exposed along tip of posterior projection in external view.</paragraph>
<paragraph id="586436FDFFC3FFFAE5AFFC787D65F89B" blockId="20.[151,1437,151,2007]" pageId="20" pageNumber="21">Total number of thoracic segments unknown; individual segments highly arched both axially and pleurally, generally short and broad, with width of axis 39.5% (31.380.7%) width across posterior pleural band, posterior segments narrower and more strongly arched, segments nearly effaced dorsally, very finely granulose on other surfaces; articulating half ring short, laterally tapered, somewhat lens-shaped, anterodorsally raised such that anterior margin is roughly equal in elevation with axial ring, anterior rim effaced, posterior half granulose (best seen as change in tone from bright light gray to medium gray in dorsal view); articulating furrow moderately long, very deep laterally in apodemal pits, shallower but still deep medially, transverse medially, with anterolaterally curved ends; axial ring short and broad, slightly wider posteriorly, with small, shallow indent at about midlength of each lateral margin, ring lightly inflated, with inflation stronger near axial furrows (posterior view), with clusters of coarse granules at each corner; doublure short, laterally tapered, semilunate, with several very fine transverse ridges; axial furrows narrow posteriorly, slightly wider at lateral indent of axial ring, and a little more than doubled in width after intersection with pleural furrow, deep, very deep posteriorly, anteriorly convergent in course; inner pleurae and outer pleurae (excluding spine) roughly equally wide in anterior segments, inner pleurae decrease in width posteriorly, joined by steep fulcral angle varying from approximately 45º to approaching subvertical; anterior pleural band as wide as posterior band (excluding spines), very short, with extremely short articulating shelf running along anterior margin and with small hook structure projecting anteriorly from ventral tip, with dense granulose sculpture concentrated posteriorly; pleural furrow very short and very deep, transverse to posterolaterally directed for most of course, anteriorly curved near tips; posterior pleural band slightly longer than axial ring, moderately inflated, higher near axial furrows and just past fulcrum, ventrolateral part of pleurae and pleural spine anterolaterally rotated and somewhat laterally flattened, with fine granules lining all edges and concentrated on anterior face just past fulcrum, and on edges of spine, spine wide (tr.), moderately long, tapered to single fine point on some specimens (Pl. 23, fig. 24; Pl. 28, fig. 21), split by small notch into two blunter tips on others (e.g., Pl. 23, fig. 15; Pl. 29, figs 4, 23), gently anteriorly curved except on far posterior segments (e.g., Pl. 28, fig. 21); doublure wide, covers all of spine, with rounded notch centered over posterior pleural band, posterior rim wider than anterior rim, folded posterodorsally to form very short articulating shelf on inner pleurae of posterior pleural band (posterior view).</paragraph>
<paragraph id="586436FDFFC3FFFBE5AFF8FC7CF6FBA8" blockId="20.[151,1437,151,2007]" lastBlockId="21.[151,1437,151,2012]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">Pygidium made of five segments and terminal piece, strongly pleurally vaulted and moderately axially vaulted, slightly longer than wide, with width measured across tips of anterior pleural band 107.9% (99.7118.7%) sagittal length excluding articulating half ring; articulating half ring short, laterally tapered, semilunate, slightly narrower than first axial ring, with anterior effaced strip and posterior finely granulose strip similar to thoracic articulating half rings; articulating furrow fairly short, deep, somewhat shallower medially; axis conical, broad anteriorly, strongly posteriorly tapered to point at end of terminal piece, moderately vaulted anteriorly, decreasing posteriorly to gently inflated terminal piece; anterior axial rings roughly equal in length, posterior rings slightly shorter, first ring very wide, fifth ring about 2/3 as wide, each ring independently inflated, shape of rings transverse to somewhat anteriorly bowed due to embayment by inter-ring furrows, mostly effaced dorsally, but with some dense granules (mainly on smaller specimens), and tiny pits in transverse line at midlength of rings, also outlined with small granules, especially noticeable at corners; inter-ring furrows lens-shaped, long medially and laterally constricted (anterior furrows) to moderately short (posterior furrows), very deep, especially laterally in apodemal pits; terminal piece triangular, long, wide anteriorly, gradually tapered to point even with or just above base of pleural spines, anterior portion gently inflated, falling along posterior taper, with pits and granules like axial rings, and with complex impression on most specimens varying from two furrows connected to last inter-ring furrow to large, roughly hexagonal pit; axial furrows fairly narrow, wider at intersections with inter-ring furrows, very deep, widely anteriorly divergent, well impressed over fifth segment, then overlapped with fifth interpleural furrows along terminal piece; anterior pleural band expressed on first segment only, extremely short in dorsal view, overhung laterally by posterior pleural band, longer ventrolaterally (lateral view), as wide as posterior pleural bands (without spine), with vanishingly short articulating tongue on anterior margin, and small anteriorly pointed hook at ventrolateral tips; pleural furrow also extremely short in dorsal view and overhung by posterior pleural band, very deep, backturned; posterior pleural bands long, wide, increasingly backturned, with pleurae 13 subparallel and 45 posteriorly convergent, each pleural rib moderately inflated near axis, increasingly inflated posteroventrally (lateral view), ribs somewhat laterally flattened toward base and spine, mostly effaced dorsally, rimmed with granules on anterior, posterior, and adaxial faces, and with line of widely spaced small pits on midline of each rib; spines with strongly posteroventrally curved ventral margin (lateral view), spines fairly short and wide, tapered to blunt point, closely spaced but with free tips, with granules on ventral margin, and small pit close to tip of spine; interpleural furrows short and very deep, posterior furrows shallower on smaller specimens, strongly backturned like pleurae; pygidial border located ventrally, formed from joined spine bases, short and narrow, with raised anterior rim; doublure seen best in anterior view, very long and pointed medially, sharply anterolaterally tapered, smooth.</paragraph>
<paragraph id="586436FDFFC2FFFBE5AFFBE87BDDFAFC" blockId="21.[151,1437,151,2012]" pageId="21" pageNumber="22">
<emphasis id="6AAFEAEFFFC2FFFBE5AFFBE87918FB8C" bold="true" box="[199,321,1123,1148]" inLineHeading="true" pageId="21" pageNumber="22" reason="1">Ontogeny.</emphasis>
Cranidia of
<taxonomicName id="9FDB4D7EFFC2FFFBE4B4FBEF7B5CFB8C" authorityName="Adrain" authorityYear="2011" box="[476,773,1124,1148]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="kerouaci">
<emphasis id="6AAFEAEFFFC2FFFBE4B4FBEF7B5CFB8C" box="[476,773,1124,1148]" italics="true" pageId="21" pageNumber="22">Protopliomerella kerouaci</emphasis>
</taxonomicName>
become wider overall, and all sculpture becomes increasingly effaced; the anterior border narrows and develops pits; the palpebro-ocular ridges move anteromedially, widen, and lose the slight postocular ridge; the glabella widens slightly posteriorly and tapers anteriorly; the lateral furrows lengthen and deepen, and S4 develops; LO lengthens, particularly laterally and becomes more rectangular; the median LO tubercle disappears; and the genal spine is reduced.
</paragraph>
<paragraph id="586436FDFFC2FFFBE5AFFA937C9BFA4F" blockId="21.[151,1437,151,2012]" pageId="21" pageNumber="22">The hypostome lengthens; the anterior wings expand somewhat and rotate ventromedially to better expose the wing process pits in ventral view; the anterior border shortens to near absence medially; the lateral notch deepens (lateral view); the middle body furrow deepens anterolaterally and fades medially; the middle body sculpture reduces in coarseness and becomes restricted to the anterior part of the anterior lobe; the lateral border widens slightly and the posterior border lengthens; and the border spines become shorter and narrower.</paragraph>
<paragraph id="586436FDFFC2FFFBE5AFFA477C78F984" blockId="21.[151,1437,151,2012]" pageId="21" pageNumber="22">Librigenal changes include overall elongation and increase in convexity (ventrolateral view); the anterior branch of the facial suture becomes slightly longer as the anterior part of the field widens and increases the distance between the eye and the lateral border; the librigenal field decreases in posterior taper and its pits enlarge and spread posteriorly; and the lateral border becomes more laterally curved. Thoracic ontogenetic changes cannot be accurately assessed because of variation in size and shape of pleurae by position.</paragraph>
<paragraph id="586436FDFFC2FFFBE5AFF90B78B3F8D8" blockId="21.[151,1437,151,2012]" pageId="21" pageNumber="22">The pygidium becomes longer and laterally rounded; the sculpture changes from coarse granules and small tubercles to nearly complete effacement; the axis narrows slightly relative to the pleurae; the axial rings become shorter medially as the inter-ring furrows lengthen; the axial furrows deepen and widen; the pleurae become longer, wider, and more posteromedially directed; and the tips of the pleural spines become blunter and more closely spaced.</paragraph>
</subSubSection>
<subSubSection id="10C16576FFC2FFF8E5AFF8B87D24FF07" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="discussion">
<paragraph id="586436FDFFC2FFFBE5AFF8B87C6CF82C" blockId="21.[151,1437,151,2012]" pageId="21" pageNumber="22">
<emphasis id="6AAFEAEFFFC2FFFBE5AFF8B87915F8BC" bold="true" box="[199,332,1843,1868]" inLineHeading="true" pageId="21" pageNumber="22" reason="1">Discussion.</emphasis>
The terminal piece of one pygidium (Pl. 30, fig. 1) is marked with a large, deep, roughly hexagonal impression. One librigena (Pl. 28, fig. 1) has a small rounded excursion of the posterior librigenal field interrupting the slope of the posterior branch of the facial suture. Some cranidial specimens, e.g., those of Pl. 28, appear more effaced, and this is likely a combination of ontogenetic and preservational differences. The cranidia in question are both larger and more damaged overall than the more granulose specimens.
</paragraph>
<paragraph id="586436FDFFC1FFF8E5AFFF1C7D24FF07" blockId="22.[151,1436,151,248]" pageId="22" pageNumber="23">
All but three sclerites figured by
<bibRefCitation id="3C4A4B0CFFC1FFF8E728FF137A9AFF5F" author="Ross, R. J., Jr." box="[576,707,151,175]" pageId="22" pageNumber="23" pagination="1 - 161" refId="ref26317" refString="Ross, R. J., Jr. (1951) Stratigraphy of the Garden City Formation in northeastern Utah, and its trilobite faunas. Peabody Museum of Natural History, Yale University, Bulletin, 6, 1 - 161." type="journal article" year="1951">Ross (1951</bibRefCitation>
, pl. 33, figs 1619, 2232) as
<taxonomicName id="9FDB4D7EFFC1FFF8E176FF137D08FF40" baseAuthorityName="Ross" baseAuthorityYear="1951" box="[1054,1361,152,176]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="contracta">
<emphasis id="6AAFEAEFFFC1FFF8E176FF137D08FF40" box="[1054,1361,152,176]" italics="true" pageId="22" pageNumber="23">Protopliomerella contracta</emphasis>
</taxonomicName>
represent
<taxonomicName id="9FDB4D7EFFC1FFF8E5A3FF36791EFF24" authorityName="Adrain" authorityYear="2011" box="[203,327,188,212]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="kerouaci">
<emphasis id="6AAFEAEFFFC1FFF8E5A3FF36791EFF24" box="[203,327,188,212]" italics="true" pageId="22" pageNumber="23">P. kerouaci</emphasis>
</taxonomicName>
, which we have also sampled from the Garden City Formation.
<taxonomicName id="9FDB4D7EFFC1FFF8E174FF377D1BFF24" authorityName="Adrain" authorityYear="2011" box="[1052,1346,188,212]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="kerouaci">
<emphasis id="6AAFEAEFFFC1FFF8E174FF377D1BFF24" box="[1052,1346,188,212]" italics="true" pageId="22" pageNumber="23">Protopliomerella kerouaci</emphasis>
</taxonomicName>
is compared to
<taxonomicName id="9FDB4D7EFFC1FFF8E591FF6A7937FF07" authorityName="Adrain" authorityYear="2011" box="[249,366,224,248]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="stegneri">
<emphasis id="6AAFEAEFFFC1FFF8E591FF6A7937FF07" box="[249,366,224,248]" italics="true" pageId="22" pageNumber="23">P. stegneri</emphasis>
</taxonomicName>
,
<taxonomicName id="9FDB4D7EFFC1FFF8E414FF6A79B4FF08" authorityName="Adrain" authorityYear="2011" box="[380,493,224,248]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bowlesi">
<emphasis id="6AAFEAEFFFC1FFF8E414FF6A79B4FF08" box="[380,493,224,248]" italics="true" pageId="22" pageNumber="23">P. bowlesi</emphasis>
</taxonomicName>
,
<emphasis id="6AAFEAEFFFC1FFF8E494FF6A7A37FF07" box="[508,622,224,248]" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="9FDB4D7EFFC1FFF8E494FF6A7A33FF07" authorityName="Adrain" authorityYear="2011" box="[508,618,224,248]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="seegeri">P. seegeri</taxonomicName>
,
</emphasis>
and briefly to
<taxonomicName id="9FDB4D7EFFC1FFF8E67CFF6A7BC3FF08" baseAuthorityName="Ross" baseAuthorityYear="1951" box="[788,922,225,248]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="contracta">
<emphasis id="6AAFEAEFFFC1FFF8E67CFF6A7BC3FF08" box="[788,922,225,248]" italics="true" pageId="22" pageNumber="23">P. contracta</emphasis>
</taxonomicName>
in the discussion sections of those species.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,132 @@
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<treatment id="D07287EBFFCCFFF5E5FFFEE47AE6FAB8" LSID="urn:lsid:plazi:treatment:D07287EBFFCCFFF5E5FFFEE47AE6FAB8" httpUri="http://treatment.plazi.org/id/D07287EBFFCCFFF5E5FFFEE47AE6FAB8" lastPageNumber="28" pageId="27" pageNumber="28">
<subSubSection id="10C16576FFCCFFF5E5FFFEE4799FFE79" box="[151,454,367,393]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="586436FDFFCCFFF5E5FFFEE4799FFE79" blockId="27.[151,454,367,426]" box="[151,454,367,393]" pageId="27" pageNumber="28">
<heading id="032C8191FFCCFFF5E5FFFEE4799FFE79" bold="true" box="[151,454,367,393]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="6AAFEAEFFFCCFFF5E5FFFEE4799FFE79" bold="true" box="[151,454,367,393]" pageId="27" pageNumber="28">
<taxonomicName id="9FDB4D7EFFCCFFF5E5FFFEE4793BFE79" ID-CoL="6XWJ" authority="Adrain, 2011" authorityName="Adrain" authorityYear="2011" box="[151,354,367,393]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="6AAFEAEFFFCCFFF5E5FFFEE4793BFE79" bold="true" box="[151,354,367,393]" italics="true" pageId="27" pageNumber="28">Protopliomerella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="719C5794FFCCFFF5E402FEE479F5FE79" box="[362,428,367,393]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
B
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFCCFFF5E5FFFE1979DDFDFF" pageId="27" pageNumber="28" type="description">
<paragraph id="586436FDFFCCFFF5E5FFFE1979D0FE5A" blockId="27.[151,454,367,426]" box="[151,393,402,426]" pageId="27" pageNumber="28">
(
<figureCitation id="C0E02A78FFCCFFF5E5F6FE1978A7FE5A" box="[158,254,402,426]" captionStart="PLATE 42" captionStartId="112.[151,233,1502,1525]" captionText="PLATE 42. 113. Protopliomerella n. sp. A, from Section YH 129.5 m, Yellow Hill Limestone (lower Floian; Tulean; Heckethornia hyndeae Zone), Yellow Hill, near Pioche, Lincoln County, eastern Nevada, USA." pageId="27" pageNumber="28">Plate 42</figureCitation>
, figs 1425)
</paragraph>
<paragraph id="586436FDFFCCFFF5E5A5FE5179DDFDFF" blockId="27.[205,1436,474,527]" pageId="27" pageNumber="28">1973 Genus and species undetermined (not described); Demeter, pl. 6, figs 10, 12 (only; pl. 6, figs 19, 11 = gen. and sp. indet.).</paragraph>
</subSubSection>
<subSubSection id="10C16576FFCCFFF5E5FFFDB0788FFD6C" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="586436FDFFCCFFF5E5FFFDB0788FFD6C" blockId="27.[151,1437,571,1352]" pageId="27" pageNumber="28">
<emphasis id="6AAFEAEFFFCCFFF5E5FFFDB0795FFDA4" bold="true" box="[151,262,571,596]" inLineHeading="true" pageId="27" pageNumber="28" reason="1">Material.</emphasis>
Assigned specimens
<collectionCode id="3ECAAE38FFCCFFF5E768FDB77A74FDA4" box="[512,557,572,596]" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="27" pageNumber="28" type="University or college">SUI</collectionCode>
126360126364, from Section D 94.4
<collectionCode id="3ECAAE38FFCCFFF5E6B5FDB77BB7FDA4" box="[989,1006,572,596]" name="Tavera, Department of Geology and Geophysics" pageId="27" pageNumber="28">T</collectionCode>
m, Fillmore Formation (lower Floian; Tulean; low
<taxonomicName id="9FDB4D7EFFCCFFF5E44CFDEB7ACFFD87" authority="Zone" authorityName="Zone" box="[292,662,607,632]" class="Trilobita" family="Bathyuridae" genus="Psalikilopsis" kingdom="Animalia" order="Proetida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="cuspidicauda">
<emphasis id="6AAFEAEFFFCCFFF5E44CFDEB7A09FD88" box="[292,592,608,632]" italics="true" pageId="27" pageNumber="28">Psalikilopsis cuspidicauda</emphasis>
Zone
</taxonomicName>
), southern House Range, Tule Valley, Millard County, western
<collectingRegion id="9A1FF81FFFCCFFF5E036FDD47DCEFD87" box="[1374,1431,607,631]" country="United States of America" name="Utah" pageId="27" pageNumber="28">Utah</collectingRegion>
,
<collectingCountry id="20CC766DFFCCFFF5E5FFFD0F788AFD6C" box="[151,211,644,668]" name="United States of America" pageId="27" pageNumber="28">USA</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFCCFFF5E5AFFD2C7AE6FAB8" pageId="27" pageNumber="28" type="discussion">
<paragraph id="586436FDFFCCFFF5E5AFFD2C7A3CFC10" blockId="27.[151,1437,571,1352]" pageId="27" pageNumber="28">
<emphasis id="6AAFEAEFFFCCFFF5E5AFFD2C7915FD30" bold="true" box="[199,332,679,704]" inLineHeading="true" pageId="27" pageNumber="28" reason="1">Discussion.</emphasis>
<taxonomicName id="9FDB4D7EFFCCFFF5E434FD237A42FD30" authority="Adrain, 2011" authorityName="Adrain" authorityYear="2011" box="[348,539,680,704]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="6AAFEAEFFFCCFFF5E434FD237A42FD30" box="[348,539,680,704]" italics="true" pageId="27" pageNumber="28">Protopliomerella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="719C5794FFCCFFF5E74BFD237A39FD4F" box="[547,608,680,703]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
B is known from only a few sclerites (two librigenae and three pygidia), but the unique pygidial morphology of seven segments demonstrates that it is a new species. The pygidium is somewhat similar to that of
<taxonomicName id="9FDB4D7EFFCCFFF5E4A5FD7A7A67FCF8" authorityName="Adrain" authorityYear="2011" box="[461,574,752,776]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bowlesi">
<emphasis id="6AAFEAEFFFCCFFF5E4A5FD7A7A67FCF8" box="[461,574,752,776]" italics="true" pageId="27" pageNumber="28">P. bowlesi</emphasis>
</taxonomicName>
, but in addition to possessing another segment, it is longer and wider overall; the axis is relatively wider anteriorly and narrower posteriorly; the pleurae are more laterally directed, not as tightly backturned, and less strongly vaulted; the inter-ring furrows are shorter; and the sculpture is slightly less effaced, with small tubercles or granules particularly noticeable running along the adaxial side of the pleurae just outside the interpleural furrows. Librigenae differ in that those of
<taxonomicName id="9FDB4D7EFFCCFFF5E64CFC0A7BC1FC68" authorityName="Adrain" authorityYear="2011" box="[804,920,896,920]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bowlesi">
<emphasis id="6AAFEAEFFFCCFFF5E64CFC0A7BC1FC68" box="[804,920,896,920]" italics="true" pageId="27" pageNumber="28">P. bowlesi</emphasis>
</taxonomicName>
are more strongly curved; much wider, particularly across the anterior part of the field; and possess a relatively narrower lateral border, with a shorter posterior projection and longer anterior projection.
</paragraph>
<paragraph id="586436FDFFCCFFF5E5AFFC677BE5FB2C" blockId="27.[151,1437,571,1352]" pageId="27" pageNumber="28">
The librigena of the younger species
<taxonomicName id="9FDB4D7EFFCCFFF5E701FC667ABFFBF4" authorityName="Adrain" authorityYear="2011" box="[617,742,1004,1028]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="kerouaci">
<emphasis id="6AAFEAEFFFCCFFF5E701FC667ABFFBF4" box="[617,742,1004,1028]" italics="true" pageId="27" pageNumber="28">P. kerouaci</emphasis>
</taxonomicName>
more closely resembles that of
<taxonomicName id="9FDB4D7EFFCCFFF5E125FC677D55FBF4" authority="Adrain, 2011" authorityName="Adrain" authorityYear="2011" box="[1101,1292,1004,1028]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="6AAFEAEFFFCCFFF5E125FC677D55FBF4" box="[1101,1292,1004,1028]" italics="true" pageId="27" pageNumber="28">Protopliomerella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="719C5794FFCCFFF5E07BFC677D16FBF3" box="[1299,1359,1004,1027]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
B, but its librigenal field is somewhat wider anteriorly and narrower posteriorly, and it has a slightly narrower lateral bor- der with a much longer and strongly posterodorsally directed posterior projection. The pygidia are not closely similar, as that of
<taxonomicName id="9FDB4D7EFFCCFFF5E45FFBD279EDFB80" authorityName="Adrain" authorityYear="2011" box="[311,436,1112,1136]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="kerouaci">
<emphasis id="6AAFEAEFFFCCFFF5E45FFBD279EDFB80" box="[311,436,1112,1136]" italics="true" pageId="27" pageNumber="28">P. kerouaci</emphasis>
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has only five segments, and a large, complexly impressed terminal piece. The pygidium of the younger species
<taxonomicName id="9FDB4D7EFFCCFFF5E4F0FBF67A5CFB64" authorityName="Adrain" authorityYear="2011" box="[408,517,1149,1172]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="seegeri">
<emphasis id="6AAFEAEFFFCCFFF5E4F0FBF67A5CFB64" box="[408,517,1149,1172]" italics="true" pageId="27" pageNumber="28">P. seegeri</emphasis>
</taxonomicName>
somewhat more closely resembles that of
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<emphasis id="6AAFEAEFFFCCFFF5E688FBF77CC6FB64" box="[992,1183,1148,1172]" italics="true" pageId="27" pageNumber="28">Protopliomerella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="719C5794FFCCFFF5E1CDFBF77CB8FB63" box="[1189,1249,1148,1171]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
B (in that it also resembles that of
<taxonomicName id="9FDB4D7EFFCCFFF5E403FB2A79B8FB48" authorityName="Adrain" authorityYear="2011" box="[363,481,1184,1208]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bowlesi">
<emphasis id="6AAFEAEFFFCCFFF5E403FB2A79B8FB48" box="[363,481,1184,1208]" italics="true" pageId="27" pageNumber="28">P. bowlesi</emphasis>
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), but again, it possesses only six segments, with strongly backturned and highly vaulted pleurae, and its axis is fairly densely covered in small tubercles.
</paragraph>
<paragraph id="586436FDFFCCFFF5E5AFFB637AE6FAB8" blockId="27.[151,1437,571,1352]" pageId="27" pageNumber="28">
<bibRefCitation id="3C4A4B0CFFCCFFF5E5AFFB63792AFAF0" author="Demeter, E. J." box="[199,371,1256,1280]" pageId="27" pageNumber="28" pagination="37 - 65" refId="ref25624" refString="Demeter, E. J. (1973) Lower Ordovician pliomerid trilobites from western Utah. Brigham Young University Geology Studies, 20, 37 - 65." type="journal article" year="1973">Demeter (1973</bibRefCitation>
, pl. 6, figs 10, 12) figured two seven-segmented pygidia likely assignable to
<taxonomicName id="9FDB4D7EFFCCFFF5E1B5FB637DC5FAF0" authority="Adrain, 2011" authorityName="Adrain" authorityYear="2011" box="[1245,1436,1256,1280]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="6AAFEAEFFFCCFFF5E1B5FB637DC5FAF0" box="[1245,1436,1256,1280]" italics="true" pageId="27" pageNumber="28">Protopliomerella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="719C5794FFCCFFF5E5FFFA87788CFAD3" box="[151,213,1292,1315]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
B from Section G at about 150.1 m. We have collected horizons G 148.2, 151T and 151.5T, but have not recovered any material assignable to this species.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="9FDB4D7EFFCDFFF4E5FFF9F4793BF969" ID-CoL="6XWJ" authority="Adrain, 2011" authorityName="Adrain" authorityYear="2011" box="[151,354,1663,1689]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="6AAFEAEFFFCDFFF4E5FFF9F4793BF969" bold="true" box="[151,354,1663,1689]" italics="true" pageId="26" pageNumber="27">Protopliomerella</emphasis>
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A
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<paragraph id="586436FDFFCDFFF4E5FFF92979BDF94A" blockId="26.[151,484,1663,1722]" box="[151,484,1698,1722]" pageId="26" pageNumber="27">
(
<figureCitation id="C0E02A78FFCDFFF4E5F6F92978A4F94A" box="[158,253,1698,1722]" captionStart="PLATE 41" captionStartId="110.[151,233,1655,1678]" captionText="PLATE 41. Protopliomerella n. sp. A, from Section YH 129.5 m, Yellow Hill Limestone (lower Floian; Tulean; Heckethornia hyndeae Zone), Yellow Hill, near Pioche, Lincoln County, eastern Nevada, USA." pageId="26" pageNumber="27">Plate 41</figureCitation>
,
<figureCitation id="C0E02A78FFCDFFF4E461F929793EF94A" box="[265,359,1698,1722]" captionStart="PLATE 42" captionStartId="112.[151,233,1502,1525]" captionText="PLATE 42. 113. Protopliomerella n. sp. A, from Section YH 129.5 m, Yellow Hill Limestone (lower Floian; Tulean; Heckethornia hyndeae Zone), Yellow Hill, near Pioche, Lincoln County, eastern Nevada, USA." pageId="26" pageNumber="27">Plate 42</figureCitation>
, figs 113)
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFCDFFF4E5FFF9617D76F8D6" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="586436FDFFCDFFF4E5FFF9617D76F8D6" blockId="26.[151,1436,1770,2010]" pageId="26" pageNumber="27">
<emphasis id="6AAFEAEFFFCDFFF4E5FFF961795FF8F3" bold="true" box="[151,262,1770,1795]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Material.</emphasis>
Assigned specimens
<collectionCode id="3ECAAE38FFCDFFF4E761F9617A6FF8F2" box="[521,566,1770,1794]" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="26" pageNumber="27" type="University or college">SUI</collectionCode>
126346126359, from Section YH 129.5 m, Yellow Hill Limestone (lower Floian; Tulean;
<taxonomicName id="9FDB4D7EFFCDFFF4E421F8857AD0F8D6" authority="Zone" authorityName="Zone" box="[329,649,1806,1830]" class="Trilobita" family="Dimeropygidae" genus="Heckethornia" kingdom="Animalia" order="Proetida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="hyndeae">
<emphasis id="6AAFEAEFFFCDFFF4E421F8857A1CF8D6" box="[329,581,1806,1830]" italics="true" pageId="26" pageNumber="27">Heckethornia hyndeae</emphasis>
Zone
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), Yellow Hill, near Pioche, Lincoln County, eastern
<collectingRegion id="9A1FF81FFFCDFFF4E1B9F8857D73F8D6" box="[1233,1322,1806,1830]" country="United States of America" name="Nevada" pageId="26" pageNumber="27">Nevada</collectingRegion>
.
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<subSubSection id="10C16576FFCDFFF5E5AFF8B979DBFEEB" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="discussion">
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<emphasis id="6AAFEAEFFFCDFFF4E5AFF8B97915F8BB" bold="true" box="[199,332,1842,1867]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Discussion.</emphasis>
Specimens of
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<emphasis id="6AAFEAEFFFCDFFF4E76FF8B87A9FF8BB" box="[519,710,1843,1867]" italics="true" pageId="26" pageNumber="27">Protopliomerella</emphasis>
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A differ from those of
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<emphasis id="6AAFEAEFFFCDFFF4E144F8B87CECF8BB" box="[1068,1205,1843,1867]" italics="true" pageId="26" pageNumber="27">P. okeeffeae</emphasis>
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in possessing more coarsely granulose cranidial sculpture, and more tuberculate dorsal thoracic and pygidial sculpture. Cranidia have a less anteriorly tapered glabella with a more faintly impressed S3, the interocular fixigenae are longer, and pygidia are slightly narrower and more elongate, with a narrower and more elongate terminal piece and longer, more strongly posteriorly directed pleurae. The librigena (Pl. 42, fig. 4) is not well preserved, but it shows coarser lateral border sculpture, and a possibly wider posterior section of the librigenal field. The species are otherwise very similar, but the sculptural difference between specimens of similar sizes is striking.
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<emphasis id="6AAFEAEFFFCCFFF5E124FF377D26FF24" box="[1100,1407,188,212]" italics="true" pageId="27" pageNumber="28">Protopliomerella okeeffeae</emphasis>
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is known from a single cranidium (
<bibRefCitation id="3C4A4B0CFFCCFFF5E76BFF547A80FF07" author="Adrain, J. M. &amp; McAdams, N. E. B. &amp; Westrop, S. R." box="[515,729,223,248]" pageId="27" pageNumber="28" pagination="541 - 610" refId="ref25562" refString="Adrain, J. M., McAdams, N. E. B. &amp; Westrop, S. R. (2009) Trilobite biostratigraphy and revised bases of the Tulean and Blackhillsian Stages of the Ibexian Series, Lower Ordovician, western United States. Memoirs of the Association of Australasian Palaeontologists, 37, 541 - 610." type="journal article" year="2009">
Adrain
<emphasis id="6AAFEAEFFFCCFFF5E731FF6A7AC8FF08" box="[601,657,224,248]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
, 2009
</bibRefCitation>
, fig. 14L) at Section YH 128.9 m, at which
<taxonomicName id="9FDB4D7EFFCCFFF5E1A9FF6B7DD9FF08" authority="Adrain, 2011" authorityName="Adrain" authorityYear="2011" box="[1217,1408,224,248]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="6AAFEAEFFFCCFFF5E1A9FF6B7DD9FF08" box="[1217,1408,224,248]" italics="true" pageId="27" pageNumber="28">Protopliomerella</emphasis>
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A does not occur.
</paragraph>
</subSubSection>
</treatment>
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<mods:detail id="52C15C16AE1FFB3C67CFD53FF1DBB6F1" type="pubDate">
<mods:number id="7D9F0D7D47188ABABB511D56770BCCFC">2011-12-23</mods:number>
</mods:detail>
<mods:detail id="2C85C2CD4B5DF5184D275A1828410743" type="volume">
<mods:number id="36E239769104A24305E6109E7E92EF5B">3144</mods:number>
</mods:detail>
<mods:extent id="BD010CBF4AC9AFD90BDCB843944B1BDF" unit="page">
<mods:start id="AC26987465BAC318176ECC8F6E77EE5D">1</mods:start>
<mods:end id="960EB016A33B713FC3B327C7E7657FF4">113</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BE489B63BE0111F71C6621039876F75D">journal article</mods:classification>
<mods:identifier id="47455F1F61E9EF98E5B3A2E5EE9D674C" type="ISSN">1175­5334</mods:identifier>
</mods:mods>
<treatment id="D07287EBFFD8FFFCE5FFFD777956FA13" ID-DOI="http://doi.org/10.5281/zenodo.5247829" ID-Zenodo-Dep="5247829" LSID="urn:lsid:plazi:treatment:D07287EBFFD8FFFCE5FFFD777956FA13" httpUri="http://treatment.plazi.org/id/D07287EBFFD8FFFCE5FFFD777956FA13" lastPageId="18" lastPageNumber="19" pageId="15" pageNumber="16">
<subSubSection id="10C16576FFD8FFE1E5FFFD777A55FCE5" box="[151,524,763,790]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="586436FDFFD8FFE1E5FFFD777A55FCE5" blockId="15.[151,524,763,822]" box="[151,524,763,790]" pageId="15" pageNumber="16">
<heading id="032C8191FFD8FFE1E5FFFD777A55FCE5" bold="true" box="[151,524,763,790]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="6AAFEAEFFFD8FFE1E5FFFD777A55FCE5" bold="true" box="[151,524,763,790]" pageId="15" pageNumber="16">
<taxonomicName id="9FDB4D7EFFD8FFE1E5FFFD77799DFCE6" authority="Adrain, 2011" authorityName="Adrain" authorityYear="2011" box="[151,452,764,790]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bowlesi" status="sp. nov.">
<emphasis id="6AAFEAEFFFD8FFE1E5FFFD77799DFCE6" bold="true" box="[151,452,764,790]" italics="true" pageId="15" pageNumber="16">Protopliomerella bowlesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="719C5794FFD8FFE1E4A2FD707A55FCE5" box="[458,524,763,789]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFD8FFE1E5FFFC957B26FBE5" pageId="15" pageNumber="16" type="description">
<paragraph id="586436FDFFD8FFE1E5FFFC957960FCC6" blockId="15.[151,524,763,822]" box="[151,313,798,822]" pageId="15" pageNumber="16">
(
<figureCitation id="C0E02A78FFD8FFE1E5F6FC957968FCC6" box="[158,305,798,822]" captionStart-0="PLATE 14" captionStart-1="PLATE 15" captionStart-2="PLATE 16" captionStart-3="PLATE 17" captionStart-4="PLATE 18" captionStart-5="PLATE 19" captionStartId-0="56.[151,233,1747,1770]" captionStartId-1="58.[151,233,1717,1740]" captionStartId-2="60.[151,233,1594,1617]" captionStartId-3="62.[151,233,1563,1586]" captionStartId-4="64.[151,233,1686,1709]" captionStartId-5="66.[151,233,1686,1709]" captionText-0="PLATE 14. Protopliomerella bowlesi n. sp., from Section G 162T m, Fillmore Formation (lower Floian; Tulean; low Psalikilopsis cuspidicauda Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." captionText-1="PLATE 15. Protopliomerella bowlesi n. sp., from Section G 162T m, Fillmore Formation (lower Floian; Tulean; low Psalikilopsis cuspidicauda Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." captionText-2="PLATE 16. Protopliomerella bowlesi n. sp., from Section G 162T m, Fillmore Formation (lower Floian; Tulean; low Psalikilopsis cuspidicauda Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." captionText-3="PLATE 17. Protopliomerella bowlesi n. sp., from Section G 162T m, Fillmore Formation (lower Floian; Tulean; low Psalikilopsis cuspidicauda Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." captionText-4="PLATE 18. Protopliomerella bowlesi n. sp., from Section G 162T m, Fillmore Formation (lower Floian; Tulean; low Psalikilopsis cuspidicauda Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." captionText-5="PLATE 19. Protopliomerella bowlesi n. sp., from Section G 162T m, Fillmore Formation (lower Floian; Tulean; low Psalikilopsis cuspidicauda Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." pageId="15" pageNumber="16">Plates 1419</figureCitation>
)
</paragraph>
<paragraph id="586436FDFFD8FFE1E5A5FCED7B0EFC8C" blockId="15.[205,1436,870,1046]" box="[205,855,870,893]" pageId="15" pageNumber="16">
1953
<taxonomicName id="9FDB4D7EFFD8FFE1E471FCEC79A2FC8C" box="[281,507,870,893]" class="Trilobita" family="Pliomeridae" genus="Protopliomerops" kingdom="Animalia" order="Phacopida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="undefined-6">
<emphasis id="6AAFEAEFFFD8FFE1E471FCEC799AFC8D" box="[281,451,871,893]" italics="true" pageId="15" pageNumber="16">Protopliomerops</emphasis>
sp. 6
</taxonomicName>
; Hintze, p. 31, 209, pl. 22, fig. 12.
</paragraph>
<paragraph id="586436FDFFD8FFE1E5A5FC0E7A4FFC49" blockId="15.[205,1436,870,1046]" pageId="15" pageNumber="16">
1953
<taxonomicName id="9FDB4D7EFFD8FFE1E471FC0E799AFC6B" authorityName="Kobayashi" authorityYear="1934" box="[281,451,901,923]" class="Trilobita" family="Pliomeridae" genus="Protopliomerops" kingdom="Animalia" order="Phacopida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="6AAFEAEFFFD8FFE1E471FC0E799AFC6B" box="[281,451,901,923]" italics="true" pageId="15" pageNumber="16">Protopliomerops</emphasis>
</taxonomicName>
aff.
<taxonomicName id="9FDB4D7EFFD8FFE1E499FC0D7AFCFC6B" authority="Ross" authorityName="Ross" box="[497,677,902,923]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="contracta">
<emphasis id="6AAFEAEFFFD8FFE1E499FC0D7A32FC6B" box="[497,619,902,923]" italics="true" pageId="15" pageNumber="16">P. contracta</emphasis>
Ross
</taxonomicName>
; Hintze, p. 31, 207, pl. 22, figs 11, 19 (only; pl. 22, figs 18, 20 =
<taxonomicName id="9FDB4D7EFFD8FFE1E006FC0D7A5FFC4A" authorityName="Adrain" authorityYear="2011" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="okeeffeae">
<emphasis id="6AAFEAEFFFD8FFE1E006FC0D7A5FFC4A" italics="true" pageId="15" pageNumber="16">Protopliomerella okeeffeae</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="586436FDFFD8FFE1E5A5FC49793CFC07" blockId="15.[205,1436,870,1046]" pageId="15" pageNumber="16">
1973
<taxonomicName id="9FDB4D7EFFD8FFE1E471FC497A20FC28" authority="(Ross)" baseAuthorityName="Ross" baseAuthorityYear="1951" box="[281,633,962,984]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="contracta">
<emphasis id="6AAFEAEFFFD8FFE1E471FC497A74FC28" box="[281,557,962,984]" italics="true" pageId="15" pageNumber="16">Protopliomerella contracta</emphasis>
(Ross)
</taxonomicName>
; Demeter, p. 59, pl. 4, fig. 2 (only; pl. 4, figs 6, 11, 12 =
<taxonomicName id="9FDB4D7EFFD8FFE1E1D4FC49790CFC07" authorityName="Adrain" authorityYear="2011" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="okeeffeae">
<emphasis id="6AAFEAEFFFD8FFE1E1D4FC49790CFC07" italics="true" pageId="15" pageNumber="16">Protopliomerella okeeffeae</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="586436FDFFD8FFE1E5A5FB8B7B26FBE5" blockId="15.[205,1436,870,1046]" box="[205,895,1024,1046]" pageId="15" pageNumber="16">
2009
<taxonomicName id="9FDB4D7EFFD8FFE1E471FB8B799CFBE6" authority="Adrain, 2011" authorityName="Adrain" authorityYear="2011" box="[281,453,1024,1046]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="6AAFEAEFFFD8FFE1E471FB8B799CFBE6" box="[281,453,1024,1046]" italics="true" pageId="15" pageNumber="16">Protopliomerella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="719C5794FFD8FFE1E4A4FB8B7A43FBE5" box="[460,538,1024,1045]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
1; Adrain
<emphasis id="6AAFEAEFFFD8FFE1E7EEFB8A7AE3FBE6" box="[646,698,1024,1046]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
, p. 560, fig.10V, Z.
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFD8FFE1E5FFFBC97AA4FB52" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="586436FDFFD8FFE1E5FFFBC97AA4FB52" blockId="15.[151,1436,1090,2015]" pageId="15" pageNumber="16">
<emphasis id="6AAFEAEFFFD8FFE1E5FFFBC9795FFBAB" bold="true" box="[151,262,1090,1115]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">Material.</emphasis>
<materialsCitation id="E8B33CA0FFD8FFE1E47CFBC97AA4FB52" collectionCode="T" collectorName="Section G" country="United States of America" county="Millard County" location="Fillmore Formation" municipality="Ibex area" pageId="15" pageNumber="16" specimenCode="SUI 115195, SUI 115196, 126096-126145" specimenCount="1" stateProvince="Utah" typeStatus="holotype">
<typeStatus id="8760885FFFD8FFE1E47CFBC97926FBAA" box="[276,383,1090,1114]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
, cranidium,
<specimenCode id="087D9E86FFD8FFE1E761FBC97ACDFBAA" box="[521,660,1090,1114]" collectionCode="SUI" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="15" pageNumber="16" type="University or college">SUI 115195</specimenCode>
, and assigned specimens
<specimenCode id="087D9E86FFD8FFE1E6D9FBC97C62FBAA" box="[945,1083,1090,1114]" collectionCode="SUI" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="15" pageNumber="16" type="University or college">SUI 115196</specimenCode>
,
<specimenCode id="087D9E86FFD8FFE1E12EFBC97CA6FBAA" box="[1094,1279,1090,1114]" collectionCode="SUI" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="15" pageNumber="16" type="University or college">126096126145</specimenCode>
, from
<collectorName id="F52E532BFFD8FFE1E020FBC978F2FB8E" pageId="15" pageNumber="16">Section G</collectorName>
162
<collectionCode id="3ECAAE38FFD8FFE1E5B5FBED78B7FB8E" box="[221,238,1126,1150]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
m,
<location id="5D046026FFD8FFE1E473FBED79A0FB8E" LSID="urn:lsid:plazi:treatment:D07287EBFFD8FFFCE5FFFD777956FA13:5D046026FFD8FFE1E473FBED79A0FB8E" box="[283,505,1126,1150]" country="United States of America" county="Millard County" municipality="Ibex area" name="Fillmore Formation" pageId="15" pageNumber="16" stateProvince="Utah">Fillmore Formation</location>
(
<location id="5D046026FFD8FFE1E760FBED7AC4FB8E" LSID="urn:lsid:plazi:treatment:D07287EBFFD8FFFCE5FFFD777956FA13:5D046026FFD8FFE1E760FBED7AC4FB8E" box="[520,669,1126,1150]" country="United States of America" county="Millard County" municipality="Ibex area" name="lower Floian" pageId="15" pageNumber="16" stateProvince="Utah">lower Floian</location>
;
<location id="5D046026FFD8FFE1E7C1FBED7AA0FB8E" LSID="urn:lsid:plazi:treatment:D07287EBFFD8FFFCE5FFFD777956FA13:5D046026FFD8FFE1E7C1FBED7AA0FB8E" box="[681,761,1126,1150]" country="United States of America" county="Millard County" municipality="Ibex area" name="Tulean" pageId="15" pageNumber="16" stateProvince="Utah">Tulean</location>
; low
<taxonomicName id="9FDB4D7EFFD8FFE1E650FBED7CF2FB8E" authority="Zone" authorityName="Zone" box="[824,1195,1126,1150]" class="Trilobita" family="Bathyuridae" genus="Psalikilopsis" kingdom="Animalia" order="Proetida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cuspidicauda">
<emphasis id="6AAFEAEFFFD8FFE1E650FBED7C3DFB8E" box="[824,1124,1126,1150]" italics="true" pageId="15" pageNumber="16">Psalikilopsis cuspidicauda</emphasis>
Zone
</taxonomicName>
),
<location id="5D046026FFD8FFE1E1D7FBED78B8FB53" LSID="urn:lsid:plazi:treatment:D07287EBFFD8FFFCE5FFFD777956FA13:5D046026FFD8FFE1E1D7FBED78B8FB53" country="United States of America" county="Millard County" municipality="Ibex area" name="southern Confusion Range" pageId="15" pageNumber="16" stateProvince="Utah">southern Confusion Range</location>
,
<collectingMunicipality id="B800AC87FFD8FFE1E584FB01790FFB52" box="[236,342,1162,1186]" pageId="15" pageNumber="16">Ibex area</collectingMunicipality>
,
<collectingCounty id="B1054E71FFD8FFE1E40AFB017A49FB52" box="[354,528,1162,1186]" pageId="15" pageNumber="16">Millard County</collectingCounty>
, western
<collectingRegion id="9A1FF81FFFD8FFE1E711FB017AEBFB52" box="[633,690,1162,1186]" country="United States of America" name="Utah" pageId="15" pageNumber="16">Utah</collectingRegion>
,
<collectingCountry id="20CC766DFFD8FFE1E7D6FB017AA3FB52" box="[702,762,1162,1186]" name="United States of America" pageId="15" pageNumber="16">USA</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFD8FFE1E5AFFB247A6BFB36" box="[199,562,1198,1223]" pageId="15" pageNumber="16" type="etymology">
<paragraph id="586436FDFFD8FFE1E5AFFB247A6BFB36" blockId="15.[151,1436,1090,2015]" box="[199,562,1198,1223]" pageId="15" pageNumber="16">
<emphasis id="6AAFEAEFFFD8FFE1E5AFFB247914FB37" bold="true" box="[199,333,1199,1223]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">Etymology.</emphasis>
After Paul Bowles.
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFD8FFE1E5AFFB597B5BFAC2" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="586436FDFFD8FFE1E5AFFB597B5BFAC2" blockId="15.[151,1436,1090,2015]" pageId="15" pageNumber="16">
<emphasis id="6AAFEAEFFFD8FFE1E5AFFB597918FB1B" bold="true" box="[199,321,1234,1259]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">Diagnosis.</emphasis>
Glabella long and narrow; glabellar sculpture very fine and concentrated medially; palpebro-ocular ridges long, with posterior end even with S2; LO with very small median tubercle; effaced six-segmented pygidium, with very long first and second inter-ring furrows.
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFD8FFFCE5AFFAB57C43FE04" lastPageId="18" lastPageNumber="19" pageId="15" pageNumber="16" type="description">
<paragraph id="586436FDFFD8FFFEE5AFFAB5789DFCE4" blockId="15.[151,1436,1090,2015]" lastBlockId="16.[151,1437,151,2012]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="6AAFEAEFFFD8FFE1E5AFFAB57901FAA7" bold="true" box="[199,344,1342,1367]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">Description.</emphasis>
Cranidium long and very wide posteriorly, narrow anteriorly, with sagittal length 53.3% (48.0 57.4%) width across genal angles, strongly vaulted; anterior border short, moderately narrow, slightly anteriorly bowed, moderately inflated, with U-shaped cross-section, densely covered in fine granules; doublure expressed as anterior face of border, exposes roughly bottom 1/3 of outer face of inner wall of border, with broad, very shallow median rostral suture arc (anterior view), and doublure also present ventrally as very short rim; anterior border furrow short, very deep, with large, deep apodemal pits at junctions with axial furrows, somewhat overhung by inflation of anterior border, broad median section gently anteriorly bowed, lateral sections between border and palpebro-ocular ridges anterolaterally directed at about 30º above horizontal; glabella long, narrow, anteriorly tapered, with maximum width across L1 88.3% (79.5100%) sagittal length, moderately arched (sag., tr.) with convexity decreasing anteriorly, formed of four distinct lateral lobes separated by sulci, densely granulose, with sculpture mostly confined to median lobe, but also branched into center of lateral lobes on some specimens (Pl. 14, figs 10, 15), and with a few very small perforate tubercles located at mid-length of each lateral lobe even with ends of sulci, additional tubercles scattered on some specimens, e.g., Pl. 15, fig. 2; Pl. 16, fig. 19; glabellar lobes weakly independently inflated, reach to about 1/3 glabellar width on each side, L1L3 large, roughly equal in area, but L1 subtriangular, L2 and L3 sub-square, L4 less well defined, short, narrow, triangular, and LF present only as very short band echoing curvature of anterior border; S1S3 short, deep, with courses about about 45º, 30º, and 45º above horizontal, with sagittal ends of S2 and especially S1 slightly posteriorly elongated, S4 very narrow, directed opposite other sulci, branches anteromedially from just inside abaxial end of S3, with junction angle of roughly 90º; SO short, moderately deep medially, very slightly longer and much deeper along L1, with course transverse medially and gently posteriorly bowed laterally; LO wide, moderately long, subrectangular, slightly inflated, with sculpture of dense granules concentrated medially, and small median node located slightly posterior from 2/3 length; LO doublure short, hardly laterally tapered, smooth; axial furrows narrow, a little wider at junctions with glabellar sulci, very deep, subparallel along LO, then steadily anteriorly convergent after bulge around L1; palpebro-ocular ridges long and moderately narrow, extend from anterior border to about even with base of L3, with widest part opposite S3, arcuate to gently sigmoid in shape, strongly raised laterally and steeply sloped anteroventrally (anterior view), with finely granulose rims; palpebro-ocular furrows narrow, deep, incised for most of course, effaced anteriorly from S3 forward and far posteriorly, shallowly sigmoid; interocular fixigenae small, triangular, anteriorly tapered, slightly longer than wide, with sculpture identical to posterior fixigenae; posterior fixigenae long and very wide, strongly downturned from fulcrum roughly even with posterolateral end of eye ridge (anterior view), with broadly anterolaterally bowed anterior margin, and with fairly dense sculpture of small pits and tiny granules, except for a border along glabella and in front of posterior border lacking pits, and a rim of granules on all furrows; posterior border furrow short, deep, incised, with broadly anterolateral course nearly transverse adaxially, and strongly anteriorly curved by genal angle; posterior border moderately short adaxially, expanded ventrolaterally to maximum length slightly before genal angle, then tapered anteriorly, with fine granulose sculpture and small, nubby genal spine; doublure fairly short laterally, cut by facial suture at anterior tip, very short medially, until slightly before genal angle, as doublure is posterodorsally rotated outward to form articulating tongue near LO.
</paragraph>
<paragraph id="586436FDFFC7FFFEE5AFFC947995FCC7" blockId="16.[151,1437,151,2012]" box="[199,460,799,823]" pageId="16" pageNumber="17">Rostral plate unknown.</paragraph>
<paragraph id="586436FDFFC7FFFEE5AFFCCF7D58F9DC" blockId="16.[151,1437,151,2012]" pageId="16" pageNumber="17">Hypostome long and narrow, widest across anterior wings, with width across shoulders 68.2% (67.268.6%) sagittal length; hypostomal suture fairly broad, slightly anteriorly bowed; anterior border extremely short medially, flared laterally into short, narrow, triangular anterior wings with relatively large, deep wing process pits; anterior border furrow very short, deep medially, shallowed laterally along posterior half of anterolateral wings, narrow, strongly anteriorly convex; middle body long, narrow, roughly ovoid but with gradual posterior taper; anterior lobe of middle body long (about 3/4 length of middle body), ovoid with rounded posterior margin, strongly ventrally inflated, inflation decreases slightly anteriorly and significantly posteriorly, with fairly dense sculpture of small tubercles concentrated on anterior half, and some scattered small pits (e.g., Pl. 16, fig. 2) on posterior half; anterolateral branches of middle body furrow overlapped with strongly impressed lateral border furrows, furrow illdefined medially, mostly effaced and more present as break in slope between lobes of middle body, strongly posteriorly bowed; posterior lobe of middle body U-shaped, lateral branches long and very narrow, middle short and narrow, gradually posteriorly tapered in width, moderately ventrally inflated, effaced; lateral border furrows narrow, deep and incised, anterior part posteromedially directed, posterior part gently laterally bowed, with anterior portion separated from posterior portion by anterolateral tips of posterior middle body branches; lateral border narrowest anteriorly along lateral notch, gradually expanded posteriorly (as glabella also narrows) to maximum width at posterolateral corners, moderately downturned (lateral view), with dense sculpture of small granules, and with small nubby spines at shoulders, corners, and roughly halfway between those points; posterior border furrow narrow, moderately long and deep, posteriorly curved; posterior border moderately downturned (lateral view) short, but slightly longer than maximum width of lateral border, posteriorly tapered to tip of small, slightly elongated median spine; doublure turned outward into wings anteriorly from moderately long, shallow lateral notch, short and narrow posteriorly, but reaches nearly to lateral and posterior border furrows, gently dorsally raised, smooth.</paragraph>
<paragraph id="586436FDFFC7FFFFE5AFF9B37A84FF5F" blockId="16.[151,1437,151,2012]" lastBlockId="17.[151,1437,151,2012]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">Librigena long and narrow, moderately laterally convex (ventrolateral view); anterior branch of facial suture very short along eye and field, then about double length and anteroventrally directed along anterior projection of border; posterior branch of facial suture very long, posteroventrally sloped, with change from about 27º to about 10º below horizontal at a little less than half length, then upturned along posterior projection; eye small, ovoid, slightly raised above field (ventrolateral view); librigenal field long and narrow, posteriorly tapered, wedge-shaped, with width under midpoint of eye 20.8% (17.023.8%) length along lateral border, with sculpture of fairly densely spaced small pits, and small granules on raised area below eye (Pl. 16, fig. 26); lateral border furrow long, narrow, deep, slightly laterally bowed; lateral border wide, very slightly posteriorly tapered along field, then more strongly tapered to point along posterior projection, strongly inflated, with long, wide, ventrolaterally directed anterior projection with bluntly squared end, posterior projection slightly shorter, strongly tapered, triangular, with strongly upturned end; doublure moderately wide, tapered to sharp point posteriorly, with narrow strip visible behind tip of posterior projection in external view, slightly tapered and terminated anteriorly at anterior edge of librigenal field, then anteroventrally rotated into anterior projection.</paragraph>
<paragraph id="586436FDFFC6FFFFE5AFFF3779C7FC37" blockId="17.[151,1437,151,2012]" pageId="17" pageNumber="18">Total number of thoracic segments unknown; segments short and relatively wide, with width of axis 38.9% (33.150.5%) width across posterior pleural band, highly vaulted in axis and pleurae, with dense sculpture of fine granules all over, including in furrows; articulating half ring short, tapered laterally, relatively wide, raised to about even with axial ring (lateral view); articulating furrow moderately long, deep medially, deeper laterally, very broadly U-shaped in course; axial ring short, approximately same length as posterior pleural band, wide, slightly wider posteriorly than anteriorly, mildly inflated, granulose, with slightly larger granules medially and projecting into axial furrows from corners; doublure very short, very slightly laterally tapered, with extremely fine parallel transverse ridges; axial furrows narrow and very deep along axial ring, slightly wider and shallower after intersection with pleural furrow, anteriorly convergent, more steeply so over anterior pleural band; inner pleurae and outer pleurae approximately same width (excluding pleural spine) on more anterior segments (Pl. 17, fig. 22), outer pleurae longer on more vaulted posterior segments (Pl. 17, fig. 21), pleural angle steep, from about 45º below horizontal to subvertical; anterior pleural band very short medially, slightly expanded ventrolaterally, with extremely short articulating tongue on anterior edge defined by extremely short furrow, and with small anteriorly directed hook structure at tips of band; pleural furrow very short, deep, incised, course posterolateral until just before tips, then gently anteriorly curved; posterior pleural band fairly long, equally broad as anterior band (excluding pleural spine), gently inflated, highest just past fulcrum (anterior view), anterolaterally rotated and somewhat laterally flattened toward abaxial end and through spine, densely granulose, with long, wide, somewhat bladed genal spine, spine with notch separating two bluntly pointed tips in some specimens, and some specimens with concentration of coarser granules lining tip(s) (Pl. 17, figs 17, 27), spine anteriorly curved in most specimens, but posteriorly directed similar to pygidial spines in more posteriorly positioned segments (Pl. 17, fig. 17); doublure very short proximally, turned out posteriorly into very short articulating tongue, long and wide ventrolaterally over spine, with notch at pleural furrow.</paragraph>
<paragraph id="586436FDFFC6FFFFE5AFFC5F7D58F8D8" blockId="17.[151,1437,151,2012]" pageId="17" pageNumber="18">Pygidium of six segments and terminal piece, relatively short and broad, with width across anterior pleural band 120.3% (111.9132.1%) length, highly vaulted, more pleurally than axially, mostly effaced; articulating half ring short, wide, semilunate; articulating furrow short and deep; axis very wide anteriorly, strongly tapered posteriorly to point at tip of terminal piece, moderately vaulted anteriorly, with convexity decreasing toward nearly flat terminal piece; axial rings subrectangular with rounded margins, broad and long anteriorly, tapering to about half width and half length posteriorly, each independently moderately inflated, with paired tiny pits located at about 1/3 and 2/3 width (e.g., Pl. 18, fig. 4; Pl. 19, figs 2, 4), and with additional smaller pits and tiny tubercles on some smaller specimens (Pl. 19, fig. 22); terminal piece triangular, narrow, highly posteriorly tapered, long, very slightly inflated; inter-ring furrows deep, short laterally, longer medially, with first 2-3 furrows much longer; axial furrows narrow, wider at intersections with inter-ring furrows, deep, strongly anteriorly divergent, slightly laterally bowed, discontinuous over sixth pleurae on most but not all specimens (cf. Pl. 18, figs 3, 13), rapidly convergent toward tip of terminal piece, then merged into single sixth interpleural furrow; inner pleurae narrow and increasingly so posteriorly; fulcral angle very steep (anterior view); outer pleurae very wide and strongly downturned; anterior pleural band present only on first segment, broad, very short, with extremely short articulating tongue on inner half set off by extremely short, fairly shallow furrow (lateral view), and with small hook-like articulating structure at lateral tips; pleural furrow short, deep over most of course, but somewhat shallowed near tips of pleurae; posterior pleural bands long, wide, strongly backturned, with fourth pair roughly parallel, and fifth and sixth pairs increasingly posteriorly convergent, moderately inflated like axial rings, with inflation lessening toward tips, ventrolateral portion somewhat laterally compressed into buttress shape (lateral view), effaced except for tiny, widely spaced line of pits centered on ribs (posterior view); pleural spines long, tapered to blunt point, laterally flattened, triangular, gently posteroventrally curved, with very tips slightly recurved dorsally, effaced except for continuation of line of pits on pleurae, and with small tubercles lining ventral margin (lateral view); interpleural furrows short, deep, incised, posterior furrows slightly shallower, backturned to follow pleurae; ventral pygidial border short and narrow, with raised inner rim; doublure moderately long medially, strongly anterolaterally tapered (anterior view).</paragraph>
<paragraph id="586436FDFFC6FFFFE5AFF8B87B44F848" blockId="17.[151,1437,151,2012]" pageId="17" pageNumber="18">
<emphasis id="6AAFEAEFFFC6FFFFE5AFF8B87918F8BC" bold="true" box="[199,321,1843,1868]" inLineHeading="true" pageId="17" pageNumber="18" reason="1">Ontogeny.</emphasis>
Cranidial ontogenetic changes (cf. Pl. 14, figs 3, 15; Pl. 15, figs 10, 23) include overall reduction in coarseness and density of sculpture; narrowing of the anterior border; posterior expansion and anterior tapering of the glabella; lengthening of LF and development of S4; development of perforate glabellar tubercles; reduction of the median LO tubercle; and reduction of the genal spine.
</paragraph>
<paragraph id="586436FDFFC6FFFCE5AFF84F7B3AFF07" blockId="17.[151,1437,151,2012]" lastBlockId="18.[151,1437,151,1508]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">Hypostomal changes (cf. Pl. 16, figs 2, 20) include overall shortening and widening; slight widening of the anterior wings; deepening of the lateral notch; shortening and widening of the anterior lobe of the middle body; shortening of the lateral branches of the posterior lobe of the middle body; reduction in coarseness and extent of middle body sculpture; and increased taper of the lateral border.</paragraph>
<paragraph id="586436FDFFC5FFFCE5AFFE8F7C5DFE94" blockId="18.[151,1437,151,1508]" pageId="18" pageNumber="19">Librigenae (cf. Pl. 16, fig. 28; Pl. 17, fig. 1) elongate and widen anteriorly; the eye develops a slightly raised socle; the librigenal field pits spread and enlarge; and the lateral border curves more (especially posteriorly). Thoracic segments are not comparable due to variation based on thoracic position.</paragraph>
<paragraph id="586436FDFFC5FFFCE5AFFEE47C43FE04" blockId="18.[151,1437,151,1508]" pageId="18" pageNumber="19">Pygidia (cf. Pl. 18, fig. 1; Pl. 19, fig. 22) elongate slightly; the pleurae become more posteromedially directed and create a rounded lateral margin overall; the axis widens and deflates slightly; the axial rings shorten medially as the inter-ring furrows lengthen; the axial furrows become deeper and wider; the pleural tips separate slightly; and the fine granules and tiny pits on the axis and pleurae become more effaced.</paragraph>
</subSubSection>
<subSubSection id="10C16576FFC5FFFCE5AFFE747956FA13" pageId="18" pageNumber="19" type="discussion">
<paragraph id="586436FDFFC5FFFCE5AFFE747B4BFD74" blockId="18.[151,1437,151,1508]" pageId="18" pageNumber="19">
<emphasis id="6AAFEAEFFFC5FFFCE5AFFE747915FDE8" bold="true" box="[199,332,511,536]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">Discussion.</emphasis>
The left S3 of one cranidium (Pl. 14, fig. 1) does not reach the axial furrow. The posterior border of one hypostome (Pl. 16, fig. 14) is irregularly rounded and spineless; it may be a healed injury. One pygidium (Pl. 18, fig. 23) has a very small, irregularly shaped terminal piece. The axial furrow is impressed over the left side only of the sixth pleurae of one pygidium (Pl. 19, fig. 2).
</paragraph>
<paragraph id="586436FDFFC5FFFCE5AFFD1B7A24FCE4" blockId="18.[151,1437,151,1508]" pageId="18" pageNumber="19">
<taxonomicName id="9FDB4D7EFFC5FFFCE5AFFD1B79BDFD58" authorityName="Adrain" authorityYear="2011" box="[199,484,656,680]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bowlesi">
<emphasis id="6AAFEAEFFFC5FFFCE5AFFD1B79BDFD58" box="[199,484,656,680]" italics="true" pageId="18" pageNumber="19">Protopliomerella bowlesi</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="9FDB4D7EFFC5FFFCE792FD1A7B3EFD57" authorityName="Adrain" authorityYear="2011" box="[762,871,656,680]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="seegeri">
<emphasis id="6AAFEAEFFFC5FFFCE792FD1A7B3EFD57" box="[762,871,656,680]" italics="true" pageId="18" pageNumber="19">P. seegeri</emphasis>
</taxonomicName>
, and they are compared in the differential discussion of the latter. The six-segmented pygidium differentiates this species from the older species
<taxonomicName id="9FDB4D7EFFC5FFFCE1A6FD3E7D1AFD3C" authorityName="Adrain" authorityYear="2011" box="[1230,1347,693,716]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="stegneri">
<emphasis id="6AAFEAEFFFC5FFFCE1A6FD3E7D1AFD3C" box="[1230,1347,693,716]" italics="true" pageId="18" pageNumber="19">P. stegneri</emphasis>
</taxonomicName>
and the younger species
<taxonomicName id="9FDB4D7EFFC5FFFCE438FD527995FD00" authorityName="Adrain" authorityYear="2011" box="[336,460,728,752]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="kerouaci">
<emphasis id="6AAFEAEFFFC5FFFCE438FD527995FD00" box="[336,460,728,752]" italics="true" pageId="18" pageNumber="19">P. kerouaci</emphasis>
</taxonomicName>
, but other sclerites are more comparable.
<taxonomicName id="9FDB4D7EFFC5FFFCE6CCFD537CE6FD00" authorityName="Adrain" authorityYear="2011" box="[932,1215,728,752]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bowlesi">
<emphasis id="6AAFEAEFFFC5FFFCE6CCFD537CE6FD00" box="[932,1215,728,752]" italics="true" pageId="18" pageNumber="19">Protopliomerella bowlesi</emphasis>
</taxonomicName>
is also compared to
<taxonomicName id="9FDB4D7EFFC5FFFCE5FFFD767955FCE4" authorityName="Adrain" authorityYear="2011" box="[151,268,765,788]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="stegneri">
<emphasis id="6AAFEAEFFFC5FFFCE5FFFD767955FCE4" box="[151,268,765,788]" italics="true" pageId="18" pageNumber="19">P. stegneri</emphasis>
</taxonomicName>
in the discussion of that species.
</paragraph>
<paragraph id="586436FDFFC5FFFCE5AFFC947C72FBFF" blockId="18.[151,1437,151,1508]" pageId="18" pageNumber="19">
Compared to
<taxonomicName id="9FDB4D7EFFC5FFFCE401FCAB7ACDFCC8" authorityName="Adrain" authorityYear="2011" box="[361,660,800,824]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="kerouaci">
<emphasis id="6AAFEAEFFFC5FFFCE401FCAB7ACDFCC8" box="[361,660,800,824]" italics="true" pageId="18" pageNumber="19">Protopliomerella kerouaci</emphasis>
</taxonomicName>
,
<taxonomicName id="9FDB4D7EFFC5FFFCE7CEFCAA7B45FCC8" authorityName="Adrain" authorityYear="2011" box="[678,796,800,824]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bowlesi">
<emphasis id="6AAFEAEFFFC5FFFCE7CEFCAA7B45FCC8" box="[678,796,800,824]" italics="true" pageId="18" pageNumber="19">P. bowlesi</emphasis>
</taxonomicName>
has an overall narrower, less anteriorly tapered, more highly inflated (especially posteriorly) glabella with coarser granulose sculpture medially, fewer and smaller tubercles outlining the glabellar lobes, and shallower glabellar furrows, longer and narrower (especially anteriorly) palpebro-ocular ridges; a slightly wider hypostome (particularly across the lateral notch) with a longer posterior lobe of the middle body and narrower and shorter lateral and posterior borders; a wider librigena with a wider (anteriorly) and less pitted field; and a relatively wider six-segmented pygidium with a smaller terminal piece lacking a complex impression, shorter pleurae, and a shorter (anterior view) doublure.
</paragraph>
<paragraph id="586436FDFFC5FFFCE5AFFB977D3CFB17" blockId="18.[151,1437,151,1508]" pageId="18" pageNumber="19">
Hintze figured specimens from Section G 158.1 m and G 238.3 m (1953, pl. 22, figs 11, 1820) referred to as
<taxonomicName id="9FDB4D7EFFC5FFFCE5FFFBCB790AFBA8" authorityName="Kobayashi" authorityYear="1934" box="[151,339,1088,1112]" class="Trilobita" family="Pliomeridae" genus="Protopliomerops" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="6AAFEAEFFFC5FFFCE5FFFBCB790AFBA8" box="[151,339,1088,1112]" italics="true" pageId="18" pageNumber="19">Protopliomerops</emphasis>
</taxonomicName>
aff.
<taxonomicName id="9FDB4D7EFFC5FFFCE4EFFBCA7A57FBA8" baseAuthorityName="Ross" baseAuthorityYear="1951" box="[391,526,1089,1112]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="contracta">
<emphasis id="6AAFEAEFFFC5FFFCE4EFFBCA7A57FBA8" box="[391,526,1089,1112]" italics="true" pageId="18" pageNumber="19">P. contracta</emphasis>
</taxonomicName>
. The
<specimenCount id="4EDDFD74FFC5FFFCE73EFBCB7AA4FBA8" box="[598,765,1088,1112]" pageId="18" pageNumber="19" type="generic">two specimens</specimenCount>
from G 158.1 m are specimens of
<taxonomicName id="9FDB4D7EFFC5FFFCE1ECFBCA7CACFBA8" authorityName="Adrain" authorityYear="2011" box="[1156,1269,1088,1112]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bowlesi">
<emphasis id="6AAFEAEFFFC5FFFCE1ECFBCA7CACFBA8" box="[1156,1269,1088,1112]" italics="true" pageId="18" pageNumber="19">P. bowlesi</emphasis>
</taxonomicName>
. Hintze noted that the cranidium (fig. 19) differed from that of
<taxonomicName id="9FDB4D7EFFC5FFFCE7A9FBEF7BAAFB8C" baseAuthorityName="Ross" baseAuthorityYear="1951" box="[705,1011,1124,1148]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="contracta">
<emphasis id="6AAFEAEFFFC5FFFCE7A9FBEF7BAAFB8C" box="[705,1011,1124,1148]" italics="true" pageId="18" pageNumber="19">Protopliomerella contracta</emphasis>
</taxonomicName>
in its less anteriorly tapered glabella, and lower palpebro-ocular ridges. He also stated that the hypostome (fig. 11) had smaller lateral border spines. The two stratigraphically higher specimens represent
<taxonomicName id="9FDB4D7EFFC5FFFCE785FB267B28FB34" authorityName="Adrain" authorityYear="2011" box="[749,881,1196,1220]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="okeeffeae">
<emphasis id="6AAFEAEFFFC5FFFCE785FB267B28FB34" box="[749,881,1196,1220]" italics="true" pageId="18" pageNumber="19">P. okeeffeae</emphasis>
</taxonomicName>
(see discussion). Hintze figured a six-segmented pygidium from G 158.1 m as
<taxonomicName id="9FDB4D7EFFC5FFFCE48BFB5B7A82FB18" box="[483,731,1232,1256]" class="Trilobita" family="Pliomeridae" genus="Protopliomerops" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="undefined-6">
<emphasis id="6AAFEAEFFFC5FFFCE48BFB5B7AC7FB18" box="[483,670,1232,1256]" italics="true" pageId="18" pageNumber="19">Protopliomerops</emphasis>
sp. 6
</taxonomicName>
(1953, pl. 22, fig. 12), and this also belongs to
<taxonomicName id="9FDB4D7EFFC5FFFCE185FB5A7D04FB18" authorityName="Adrain" authorityYear="2011" box="[1261,1373,1232,1256]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bowlesi">
<emphasis id="6AAFEAEFFFC5FFFCE185FB5A7D04FB18" box="[1261,1373,1232,1256]" italics="true" pageId="18" pageNumber="19">P. bowlesi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="586436FDFFC5FFFCE5AFFB7F7956FA13" blockId="18.[151,1437,151,1508]" pageId="18" pageNumber="19">
A single badly damaged pygidium from G 155.6 m likely represents
<taxonomicName id="9FDB4D7EFFC5FFFCE689FB7F7CA6FAFC" authorityName="Adrain" authorityYear="2011" box="[993,1279,1268,1292]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bowlesi">
<emphasis id="6AAFEAEFFFC5FFFCE689FB7F7CA6FAFC" box="[993,1279,1268,1292]" italics="true" pageId="18" pageNumber="19">Protopliomerella bowlesi</emphasis>
</taxonomicName>
. It is broken anteriorly, but evidence from a count of pleurae and damaged axial rings indicates six segments, which rules out assignment to
<taxonomicName id="9FDB4D7EFFC5FFFCE450FAB679F4FAA4" authorityName="Adrain" authorityYear="2011" box="[312,429,1341,1364]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="stegneri">
<emphasis id="6AAFEAEFFFC5FFFCE450FAB679F4FAA4" box="[312,429,1341,1364]" italics="true" pageId="18" pageNumber="19">P. stegneri</emphasis>
</taxonomicName>
.
<bibRefCitation id="3C4A4B0CFFC5FFFCE4A9FAB77A35FAA4" author="Demeter, E. J." box="[449,620,1340,1364]" pageId="18" pageNumber="19" pagination="37 - 65" refId="ref25624" refString="Demeter, E. J. (1973) Lower Ordovician pliomerid trilobites from western Utah. Brigham Young University Geology Studies, 20, 37 - 65." type="journal article" year="1973">Demeter (1973</bibRefCitation>
, pl. 6, figs 10, 12) figured pygidia from approximately G 150.1 m, which have seven segments and are likely
<taxonomicName id="9FDB4D7EFFC5FFFCE75BFAEB7AABFA88" authority="Adrain, 2011" authorityName="Adrain" authorityYear="2011" box="[563,754,1376,1400]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="6AAFEAEFFFC5FFFCE75BFAEB7AABFA88" box="[563,754,1376,1400]" italics="true" pageId="18" pageNumber="19">Protopliomerella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="719C5794FFC5FFFCE792FAEB7B61FA87" box="[762,824,1376,1399]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
B (see discussion of that taxon). However, in lateral view (Pl. 19, fig. 23) the pygidium from G 155.6 m appears to include the ventrolateral portion of the anterior pleural band, suggesting that the damage has not removed additional anterior segments, and that the pygidium is that of
<taxonomicName id="9FDB4D7EFFC5FFFCE5FFFA46795EFA14" authorityName="Adrain" authorityYear="2011" box="[151,263,1484,1508]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bowlesi">
<emphasis id="6AAFEAEFFFC5FFFCE5FFFA46795EFA14" box="[151,263,1484,1508]" italics="true" pageId="18" pageNumber="19">P. bowlesi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="56A8907C4BB76C7B4A3A67F775BFE1AF">Revision of the Lower Ordovician (lower Floian; Tulean) pliomerid trilobite Protopliomerella, with new species from the Great Basin, western USA 3144</mods:title>
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<mods:name id="7A2A1E46E1441A1A8BB49BD39B3A01EB" type="personal">
<mods:role id="94CEC490970F9659BF23730E754BBC70">
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<mods:namePart id="D657CB6837A7C903E24D1BDE69D1590A">Adrain, Jonathan M.</mods:namePart>
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<mods:typeOfResource id="67E5B92A28C632F56F6B2839A8B6F3D5">text</mods:typeOfResource>
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<mods:titleInfo id="8CCA8EF1A07FF306BDD4E802F213EF6F">
<mods:title id="334E88022E588577F7B6DCEC18A81A42">Zootaxa</mods:title>
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<mods:part id="AA68CFC3D1AAA771830A3C543217F34F">
<mods:date id="AF95F3C71F818137396F9AFA4F8D1D56">2011</mods:date>
<mods:detail id="206A988CCC7E5BAD630905047F8C898A" type="pubDate">
<mods:number id="E95648ED6B92534F1E6A8A5C52CFEA05">2011-12-23</mods:number>
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<mods:number id="D19746A0A5E4BE3CC6C21F2FE8CEA9CC">3144</mods:number>
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<mods:start id="19055C133F837961AAFD7321A7570BB9">1</mods:start>
<mods:end id="527A39BCCB37D45847B661D0B1FF252C">113</mods:end>
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<mods:identifier id="A98DC0306CB57FE000310A4365C7594F" type="ISSN">1175­5334</mods:identifier>
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<treatment id="D07287EBFFDBFFE1E5FFFDAF7CF6FD57" LSID="urn:lsid:plazi:treatment:D07287EBFFDBFFE1E5FFFDAF7CF6FD57" httpUri="http://treatment.plazi.org/id/D07287EBFFDBFFE1E5FFFDAF7CF6FD57" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13">
<subSubSection id="10C16576FFDBFFE2E5FFFDAF7A4AFDCD" box="[151,531,547,574]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="586436FDFFDBFFE2E5FFFDAF7A4AFDCD" blockId="12.[151,531,547,606]" box="[151,531,547,574]" pageId="12" pageNumber="13">
<heading id="032C8191FFDBFFE2E5FFFDAF7A4AFDCD" bold="true" box="[151,531,547,574]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="6AAFEAEFFFDBFFE2E5FFFDAF7A4AFDCD" bold="true" box="[151,531,547,574]" pageId="12" pageNumber="13">
<taxonomicName id="9FDB4D7EFFDBFFE2E5FFFDAF7993FDCE" authority="Adrain, 2011" authorityName="Adrain" authorityYear="2011" box="[151,458,548,574]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="stegneri" status="sp. nov.">
<emphasis id="6AAFEAEFFFDBFFE2E5FFFDAF7993FDCE" bold="true" box="[151,458,548,574]" italics="true" pageId="12" pageNumber="13">Protopliomerella stegneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="719C5794FFDBFFE2E4B9FDA87A4AFDCD" box="[465,531,547,573]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFDBFFE2E5FFFDCD7D3DFD54" pageId="12" pageNumber="13" type="description">
<paragraph id="586436FDFFDBFFE2E5FFFDCD7972FDAE" blockId="12.[151,531,547,606]" box="[151,299,582,606]" pageId="12" pageNumber="13">
(
<figureCitation id="C0E02A78FFDBFFE2E5F6FDCD797AFDAE" box="[158,291,582,606]" captionStart-0="PLATE 9" captionStart-1="PLATE 10" captionStart-2="PLATE 11" captionStart-3="PLATE 12" captionStart-4="PLATE 13" captionStartId-0="46.[151,233,1625,1648]" captionStartId-1="48.[151,233,1625,1648]" captionStartId-2="50.[151,233,1625,1648]" captionStartId-3="52.[151,233,1655,1678]" captionStartId-4="54.[151,233,1594,1617]" captionText-0="PLATE 9. Protopliomerella stegneri n. sp., from Section G 138144.5T m, Fillmore Formation (lower Floian; Tulean; unzoned strata beneath Psalikilopsis cuspidicauda Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." captionText-1="PLATE 10. Protopliomerella stegneri n. sp., from Section G 138144.5T m, Fillmore Formation (lower Floian; Tulean; unzoned strata beneath Psalikilopsis cuspidicauda Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." captionText-2="PLATE 11. Protopliomerella stegneri n. sp., from Section G 138144.5T m, Fillmore Formation (Tulean; unzoned strata beneath Psalikilopsis cuspidicauda Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." captionText-3="PLATE 12. Protopliomerella stegneri n. sp., from Section G 138144.5T m, Fillmore Formation (lower Floian; Tulean; unzoned strata beneath Psalikilopsis cuspidicauda Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." captionText-4="PLATE 13. Protopliomerella stegneri n. sp., from Section G 142144.5T m, Fillmore Formation (lower Floian; Tulean; unzoned strata beneath Psalikilopsis cuspidicauda Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." pageId="12" pageNumber="13">Plates 913</figureCitation>
)
</paragraph>
<paragraph id="586436FDFFDBFFE2E5A5FD057D3DFD54" blockId="12.[205,1380,654,677]" box="[205,1380,654,677]" pageId="12" pageNumber="13">
1973
<taxonomicName id="9FDB4D7EFFDBFFE2E477FD047A44FD54" authority="(Ross)" baseAuthorityName="Ross" baseAuthorityYear="1951" box="[287,541,654,677]" class="Trilobita" family="Pliomeridae" genus="Hintzeia" kingdom="Animalia" order="Phacopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="celsaora">
<emphasis id="6AAFEAEFFFDBFFE2E477FD04798BFD55" box="[287,466,655,677]" italics="true" pageId="12" pageNumber="13">Hintzeia celsaora</emphasis>
(Ross)
</taxonomicName>
; Demeter, p. 58, pl. 3, figs 9, 10, 14 (only; pl. 3, figs 1113 =?
<taxonomicName id="9FDB4D7EFFDBFFE2E1CBFD047D0CFD55" baseAuthorityName="Ross" baseAuthorityYear="1951" box="[1187,1365,655,677]" class="Trilobita" family="Pliomeridae" genus="Hintzeia" kingdom="Animalia" order="Phacopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="celsaora">
<emphasis id="6AAFEAEFFFDBFFE2E1CBFD047D0CFD55" box="[1187,1365,655,677]" italics="true" pageId="12" pageNumber="13">Hintzeia celsaora</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFDBFFE2E5FFFD5A7981FCA5" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="586436FDFFDBFFE2E5FFFD5A7981FCA5" blockId="12.[151,1436,721,2006]" pageId="12" pageNumber="13">
<emphasis id="6AAFEAEFFFDBFFE2E5FFFD5A795FFD1A" bold="true" box="[151,262,721,746]" inLineHeading="true" pageId="12" pageNumber="13" reason="1">Material.</emphasis>
<materialsCitation id="E8B33CA0FFDBFFE2E47DFD5A7981FCA5" collectionCode="T" collectorName="Section G" country="United States of America" county="Millard County" location="Fillmore Formation" municipality="Ibex area" pageId="12" pageNumber="13" specimenCode="SUI 126054, SUI 126052, 126053, 126055-126095" specimenCount="1" stateProvince="Utah" typeStatus="holotype">
<typeStatus id="8760885FFFDBFFE2E47DFD5A79D9FD19" box="[277,384,721,745]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
, cranidium,
<specimenCode id="087D9E86FFDBFFE2E764FD5A7AC1FD19" box="[524,664,721,745]" collectionCode="SUI" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="12" pageNumber="13" type="University or college">SUI 126054</specimenCode>
, from
<collectorName id="F52E532BFFDBFFE2E789FD5A7B08FD19" box="[737,849,721,745]" pageId="12" pageNumber="13">Section G</collectorName>
138139
<collectionCode id="3ECAAE38FFDBFFE2E6D2FD5A7B95FD19" box="[954,972,721,745]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="13">T</collectionCode>
m, and assigned specimens
<specimenCode id="087D9E86FFDBFFE2E064FD5A7DCEFD19" box="[1292,1431,721,745]" collectionCode="SUI" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="12" pageNumber="13" type="University or college">SUI 126052</specimenCode>
,
<specimenCode id="087D9E86FFDBFFE2E5FFFD7E78B7FCFD" box="[151,238,757,781]" collectionCode="SUI" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="12" pageNumber="13" type="University or college">126053</specimenCode>
,
<specimenCode id="087D9E86FFDBFFE2E594FD7E79ECFCFD" box="[252,437,757,781]" collectionCode="SUI" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="12" pageNumber="13" type="University or college">126055126095</specimenCode>
, from
<collectorName id="F52E532BFFDBFFE2E76DFD7E7A21FCFD" box="[517,632,757,781]" pageId="12" pageNumber="13">Section G</collectorName>
138139
<collectionCode id="3ECAAE38FFDBFFE2E78CFD7E7AAFFCFD" box="[740,758,757,781]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="13">T</collectionCode>
m and 142144.5
<collectionCode id="3ECAAE38FFDBFFE2E6A3FD7E7B85FCFD" box="[971,988,757,781]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="13">T</collectionCode>
m,
<location id="5D046026FFDBFFE2E166FD7E7CB7FCFD" LSID="urn:lsid:plazi:treatment:D07287EBFFDBFFE1E5FFFDAF7CF6FD57:5D046026FFDBFFE2E166FD7E7CB7FCFD" box="[1038,1262,757,781]" country="United States of America" county="Millard County" municipality="Ibex area" name="Fillmore Formation" pageId="12" pageNumber="13" stateProvince="Utah">Fillmore Formation</location>
(
<location id="5D046026FFDBFFE2E068FD7E7DCEFCFD" LSID="urn:lsid:plazi:treatment:D07287EBFFDBFFE1E5FFFDAF7CF6FD57:5D046026FFDBFFE2E068FD7E7DCEFCFD" box="[1280,1431,757,781]" country="United States of America" county="Millard County" municipality="Ibex area" name="lower Floian" pageId="12" pageNumber="13" stateProvince="Utah">lower Floian</location>
;
<location id="5D046026FFDBFFE2E5FFFC9278BEFCC1" LSID="urn:lsid:plazi:treatment:D07287EBFFDBFFE1E5FFFDAF7CF6FD57:5D046026FFDBFFE2E5FFFC9278BEFCC1" box="[151,231,793,817]" country="United States of America" county="Millard County" municipality="Ibex area" name="Tulean" pageId="12" pageNumber="13" stateProvince="Utah">Tulean</location>
; unzoned strata beneath
<taxonomicName id="9FDB4D7EFFDBFFE2E76CFC927B20FCC1" authority="Zone" authorityName="Zone" box="[516,889,793,817]" class="Trilobita" family="Bathyuridae" genus="Psalikilopsis" kingdom="Animalia" order="Proetida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cuspidicauda">
<emphasis id="6AAFEAEFFFDBFFE2E76CFC927B68FCC1" box="[516,817,793,817]" italics="true" pageId="12" pageNumber="13">Psalikilopsis cuspidicauda</emphasis>
Zone
</taxonomicName>
),
<location id="5D046026FFDBFFE2E6E6FC927C99FCC2" LSID="urn:lsid:plazi:treatment:D07287EBFFDBFFE1E5FFFDAF7CF6FD57:5D046026FFDBFFE2E6E6FC927C99FCC2" box="[910,1216,793,818]" country="United States of America" county="Millard County" municipality="Ibex area" name="southern Confusion Range" pageId="12" pageNumber="13" stateProvince="Utah">southern Confusion Range</location>
,
<collectingMunicipality id="B800AC87FFDBFFE2E1A6FC927D63FCC1" box="[1230,1338,793,817]" pageId="12" pageNumber="13">Ibex area</collectingMunicipality>
,
<collectingCounty id="B1054E71FFDBFFE2E020FC9278B2FCA5" pageId="12" pageNumber="13">Millard County</collectingCounty>
, western
<collectingRegion id="9A1FF81FFFDBFFE2E43CFCB679D4FCA5" box="[340,397,829,853]" country="United States of America" name="Utah" pageId="12" pageNumber="13">Utah</collectingRegion>
,
<collectingCountry id="20CC766DFFDBFFE2E4F0FCB6798DFCA5" box="[408,468,829,853]" name="United States of America" pageId="12" pageNumber="13">USA</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFDBFFE2E5AFFCEA7A00FC89" box="[199,601,865,889]" pageId="12" pageNumber="13" type="etymology">
<paragraph id="586436FDFFDBFFE2E5AFFCEA7A00FC89" blockId="12.[151,1436,721,2006]" box="[199,601,865,889]" pageId="12" pageNumber="13">
<emphasis id="6AAFEAEFFFDBFFE2E5AFFCEA7914FC89" bold="true" box="[199,333,865,889]" inLineHeading="true" pageId="12" pageNumber="13" reason="1">Etymology.</emphasis>
After Wallace Stegner.
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFDBFFE2E5AFFC0E7966FC15" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="586436FDFFDBFFE2E5AFFC0E7966FC15" blockId="12.[151,1436,721,2006]" pageId="12" pageNumber="13">
<emphasis id="6AAFEAEFFFDBFFE2E5AFFC0E7918FC6E" bold="true" box="[199,321,901,926]" inLineHeading="true" pageId="12" pageNumber="13" reason="1">Diagnosis.</emphasis>
Anterior border moderately broad; S4 barely impressed; hypostomal suture broad, with anterior wings long and much wider than shoulders; posterior section of librigenal field fairly wide; pygidial pleural spines well separated.
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFDBFFE0E5AFFC7A79D6F880" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
<paragraph id="586436FDFFDBFFE3E5AFFC7A7A39FE5C" blockId="12.[151,1436,721,2006]" lastBlockId="13.[151,1437,151,2012]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="6AAFEAEFFFDBFFE2E5AFFC7A7901FBFA" bold="true" box="[199,344,1009,1034]" inLineHeading="true" pageId="12" pageNumber="13" reason="1">Description.</emphasis>
Cranidium relatively short and wide, with sagittal length 48.4% (45.650.6%) maximum width across genal angles, highly vaulted (tr.) mainly because of strongly downturned fixigenae, axis moderately vaulted, cranidium covered in dense, finely granulose sculpture; anterior border moderately broad, short, gently anteriorly bowed, moderately inflated, densely granulose; doublure expressed as anterior face of border, exposes ventral half of inner wall of border, with very shallow median rostral suture arc of about 1/3 total border width; anterior border furrow very short, deep, incised, somewhat anteroventrally directed underneath posterior edge of anterior border, course gently anteriorly bowed over broad glabellar section, roughly transverse on narrow lateral sections in front of palpebro-ocular ridges, with deeper, longer apodemal pits at junctions with axial furrows; glabella relatively short and wide, gradually anteriorly tapered, with maximum width across L1 100.4% (94.7106.5%) sagittal length, mildly vaulted (sag., tr.), with convexity highest posteriorly and falling anteriorly, with three well defined independently inflated lateral lobes, and with sculpture of fine granules, somewhat effaced in centers of L1L3, denser medially, with lobes outlined by slightly larger granules on edges of furrows; L2 and L3 very similar, subrectangular, anterolaterally oriented at about 20º above horizontal, with L2 slightly longer and wider, L1 smaller, narrower, subtriangular, LF short, narrow, wedge-shaped, with very narrow, faint trace of S4 diverging anteromedially from S3 suggesting L4 (better visible on larger specimens, e.g., Pl. 9, figs 3, 21), S1 and S2 moderately long, very deep, with L1 slightly longer with inner end slightly posteriorly elongated, S3 shorter and shallower; SO short and deep medially, longer and deeper laterally in apodemal pits behind mid-L1, then constricted toward axial furrow, abaxial portions lined by small granules on margins of L1 and LO; LO wide, somewhat longer medially and tapered laterally, with very shallowly W-shaped anterior margin indented by apodemal pits in SO, moderately inflated, highest just posterior from mid-length (lateral view), densely granulose, with small median node and slightly larger granules on lateral margins; doublure fairly short, laterally tapered, lens-shaped; axial furrows deep, fairly narrow, wider at triangular junctions with glabellar furrows, anteriorly convergent along LO, then moderately laterally bowed around base of L1, then gradually anteriorly convergent; palpebro-ocular ridges fairly long, reaching from anterior border to anterior of L2, narrow, arcuate, tapered anteriorly and posteriorly, with widest point even with mid-L3, dorsolaterally raised (anterior view); palpebro-ocular furrows arcuate to gently sigmoid, very short, deep over most of course, shallow anteriorly and far posteriorly, anteromedially directed at about 45º above horizontal; interocular fixigenae short, fairly narrow, tapered anteriorly to point at junction of glabella and palpebro-ocular ridge, roughly equilaterally triangular; posterior fixigenae slightly longer, very wide, with broadly anteriorly curved anterior margin, very strongly downturned from fulcrum, all areas of fixigenae with dense granulose sculpture and slightly less densely concentrated small pits; posterior border furrow roughly transverse until genal angle, then anteriorly curved (lateral view), deep, short, constricted adaxially, longer over most of course, then tapered anteriorly after genal angle; posterior border moderately long, expanded laterally toward maximum length at genal angle of not quite double adaxial length, then abruptly tapered to end, moderately inflated, granulose, with small nubby spine at genal angle; doublure very short adaxially, rotated posterodorsally into articulating device near axis, gradually expanded laterally to maximum just before genal angle, and terminated by cut of facial suture at and slightly anterior from genal angle.
</paragraph>
<paragraph id="586436FDFFDAFFE3E5AFFE3C7995FE3F" blockId="13.[151,1437,151,2012]" box="[199,460,439,463]" pageId="13" pageNumber="14">Rostral plate unknown.</paragraph>
<paragraph id="586436FDFFDAFFE3E5AFFE577973FAFC" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="14">Hypostome long and narrow, widest across anterior wings, with width across shoulders 68.6% (65.9, 71.2%) sagittal length; hypostomal suture broad, very shallowly arcuate to nearly transverse; anterior border extremely short medially, abruptly expanded laterally into large triangular anterior wings of approximately equal width and length with small, deep wing process pits, densely covered in granules; anterior border furrow strongly anteriorly bowed, short, deep, incised medially, nearly effaced posterolaterally toward meeting with lateral border furrows; middle body long, narrow, ovoid; anterior lobe large, elongate, ovoid, about 80% total length of middle body, strongly ventrally inflated, with inflation decreasing slightly posteriorly, with dense sculpture of small tubercles over at least anterior half of lobe (entire lobe in some specimens); lateral branches of middle body furrow narrow, deep, incised, posteromedially sloped and gently laterally bowed in course, intersect lateral border furrows even with lateral notch, poorly impressed medially (longer and much shallower) on most specimens, but strongly posteriorly curved; posterior lobe of middle body U-shaped, with long, slender, anteriorly tapered arms and short, narrow midsection, lobe moderately inflated, effaced; lateral border furrows narrow, deep except over tips of lateral branches of posterior lobe of middle body and along anterior wings, incised, somewhat laterally bowed posterolaterally, subparallel for most of course and then anteriorly convergent with anterior border furrow; lateral border strongly downturned and inflated, narrow, gradually expanded posteriorly to maximum width at posterolateral corners, with dense sculpture of coarse granules and some anastomosing ridges, and with small nubby spines (just rounded angles in some specimens) at shoulders and slightly posterior from shoulders, and a longer, thicker, more pointed spine at posterolateral corners; posterior border furrow long, shallow, ill-defined, strongly posteriorly convex in course; posterior border long, fairly wide, moderately steeply ventrally angled (lateral view), with background sculpture like that of lateral border, and with small, broad-based, triangular median spine; doublure short and narrow, but reaches to lateral border furrow and about halfway to posterior border furrow, with anterior parts turned outward at lateral notch to form anterior wings, doublure moderately strongly upturned, with long, shallow lateral notch.</paragraph>
<paragraph id="586436FDFFDAFFE3E5AFFA937924F880" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="14">Librigena long and moderately wide, laterally convex but not well curved; anterior branch of facial suture short along eye and field, meets lateral border at slightly greater than 90º, branch along anterior projection of lateral bor- der longer and strongly downturned; posterior branch of facial suture very long, with two major changes of slope, the first at about half length of field, changing from angle of about 45º to very gentle slope less than 10º, and the second at junction with lateral border, changing to posterodorsal curvature along posterior projection of lateral bor- der; eye elliptical, narrow and elongate, elevated above librigenal field on socle about same height as eye (ventrolateral view); librigenal field roughly wedge-shaped, fairly broad anteriorly, strongly tapered after about half field length, with width of field under midpoint of eye 22.9% (20.424.9%) length along lateral border furrow, moderately steeply ventrolaterally sloped, with sculpture of dense granules and fairly dense small pits concentrated under eye and near facial sutures; lateral border furrow narrow, deep, with short deeper segment at anterior end and slightly shallower toward posterior tip, curved, with slope steeper anteriorly; lateral border highly inflated, with inflation greater ventrolaterally, broad, gradually posteriorly tapered, rotated anteroventrally to form very long anterior projection with blunt termination, posterior projection short, triangular, strongly posteriorly tapered to point, with inner face of doublure protruding, border with dense sculpture of granules slightly coarser than those of field, but effaced far ventrolaterally; doublure wide, posteriorly tapered, with slight protrusion above tip in external view, bluntly terminated anteriorly even with edge of librigenal field, rotated anteroventrally outward into anterior projection of border.</paragraph>
<paragraph id="586436FDFFDAFFE0E5AFF8F77B25FD74" blockId="13.[151,1437,151,2012]" lastBlockId="14.[151,1437,151,2012]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">Total number of thoracic segments unknown; segments highly arched axially and pleurally (varies somewhat with thoracic position), short and broad, with width of axis 34.4% (33.5, 35.2%) width across posterior pleural band, with dense finely granulose sculpture all over, including in furrows; articulating half ring short, laterally tapered, semilunate, anteriorly upturned; articulating furrow deep, long medially, sharply constricted and deeper ventrolaterally, broadly arcuate and posteriorly convex in course; axial ring narrow, about same width as inner pleurae (may change depending on thoracic position), short, transverse medially with shallowly anterolaterally curved extremities, well inflated, slightly higher posteriorly; doublure short, laterally tapered, with fine transverse ridge at midlength; axial furrows fairly narrow, widest at junction with pleural furrow, deep, anteriorly convergent; inner pleurae and outer pleurae of roughly equal widths (may vary with thoracic position), inner pleurae nearly horizontal (anterior view), joined to steeply sloped outer pleurae by fulcral angle of about 115º; anterior pleural band very short, broad, uninflated, with extremely short articulating tongue on anterior margin set off posteriorly by incised furrow; pleural furrow very short and deep, slightly shallower near adaxial and abaxial ends, transverse for most of course, anteriorly curved at ventrolateral tip; posterior pleural band about twice length of anterior band dorsally, expanded ventrolaterally toward spine, broad, well inflated, higher near axis, laterally flattened and anterolaterally rotated to form broad-based spine, spine separated into two prongs by sharp notch in some specimens (Pl. 11, fig. 22); doublure on inner pleurae posterodorsally twisted into articulating ledge on posterior margin, long and wide ventrolaterally over spine, with deep notch at pleural furrow.</paragraph>
<paragraph id="586436FDFFD9FFE0E5AFFD047D29F9F8" blockId="14.[151,1437,151,2012]" pageId="14" pageNumber="15">Pygidium highly vaulted pleurally and axially, composed of five segments and terminal piece, relatively short and wide, with width across tips of anterior pleural band of first segment 124.2% (117.7133.1%) sagittal length excluding articulating half ring and free tips of spines, and with dense finely granulose sculpture; articulating half ring broad, short, laterally tapered; articulating furrow long and deep; axis conical, broad anteriorly, strongly posteriorly tapered to point at tip of terminal piece, highly vaulted anteriorly, decreasing posteriorly to barely inflated terminal piece; axial rings short and broad, with fifth ring a little shorter and slightly greater than half width of first, rings highly independently inflated, lined on all margins with small tubercles and granules, with small tubercles or pitted tubercles medially; inter-ring furrows long medially, shorter ventrolaterally into apodemal pits, deep, very deep in pits; terminal piece triangular, broad anteriorly, strongly tapered, long, with narrow posterior section extending almost to base of pygidial spines on some specimens (posterior view), mildly inflated, with impression of two short furrows projecting from anterior margin; axial furrows fairly narrow, wider at intersections with interpleural furrows, deep, widely anteriorly divergent, strong over fifth segment, then merged with fifth interpleural furrows along terminal piece; anterior pleural band present only on first segment, short, as wide as posterior band excluding its spine, strongly posterolaterally curved, with slightly flared tips (dorsal view), and with extremely short articulating tongue on anterior edge (best seen in lateral view, e.g., Pl. 12, fig. 7; Pl. 13, fig. 5), and small anteriorly directed hook structure at adaxial tip; pleural furrow short, deep, incised; posterior pleural bands long, broad, strongly independently inflated, increasingly backturned, with pleurae 14 approaching parallel and fifth pleurae medially convergent, granulose, with larger granules concentrated on margins of pleurae, and with small pits (some with slightly raised rims) on center of pleurae, and with large pleural spine; spines fairly short (posterior view), broad, tapered to blunt point, somewhat laterally flattened and blade-like, well separated at tips, projecting posteroventrally with gently curved ventral margin (lateral view) and slightly raised ridge at margin; interpleural furrows moderately long, anterior furrows very deep, furrows 45 relatively shallow but still well impressed, strongly backturned like pleurae, with 5th furrows posteriorly convergent into single furrow behind terminal piece; pygidial border located ventrally, formed of fused spine bases, with slightly raised rim anterolaterally, grading into doublure medially; doublure roughly triangular, moderately long medially and strongly anterolaterally tapered.</paragraph>
<paragraph id="586436FDFFD9FFE0E5AFF99879D6F880" blockId="14.[151,1437,151,2012]" pageId="14" pageNumber="15">
<emphasis id="6AAFEAEFFFD9FFE0E5AFF9987918F9DC" bold="true" box="[199,321,1555,1580]" inLineHeading="true" pageId="14" pageNumber="15" reason="1">Ontogeny.</emphasis>
<taxonomicName id="9FDB4D7EFFD9FFE0E426F99F7A34F9DC" authorityName="Adrain" authorityYear="2011" box="[334,621,1556,1580]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="stegneri">
<emphasis id="6AAFEAEFFFD9FFE0E426F99F7A34F9DC" box="[334,621,1556,1580]" italics="true" pageId="14" pageNumber="15">Protopliomerella stegneri</emphasis>
</taxonomicName>
is not as well known as most other species of the genus. The small sample size and similar sizes of individual specimens hinder discussion of ontogenetic changes. However, it is possible to see that the glabella narrows and elongates slightly, all the cranidial furrows deepen, the palpebro-ocular ridges increase in anterolateral curvature, the coarseness of the pits and granules increases, and the genal angle widens. The hypostomes are very close in size, but it appears that larger hypostomes have coarser middle body tubercles and shorter lateral and posterior border spines. Librigenal ontogenetic changes include widening of the field, with a relative decrease in width of the lateral border, and with a less marked change in slope of the posterior branch of the facial suture. Thoracic segments are too poorly known to be informative. The pygidium elongates slightly, the axial ring sculpture is reduced, the inter-ring and interpleural furrows lengthen and deepen, and the terminal piece develops impressions.
</paragraph>
</subSubSection>
<subSubSection id="10C16576FFD9FFE1E5AFF8F07CF6FD57" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="discussion">
<paragraph id="586436FDFFD9FFE0E5AFF8F07AF0F82B" blockId="14.[151,1437,151,2012]" pageId="14" pageNumber="15">
<emphasis id="6AAFEAEFFFD9FFE0E5AFF8F07915F864" bold="true" box="[199,332,1915,1940]" inLineHeading="true" pageId="14" pageNumber="15" reason="1">Discussion.</emphasis>
Two damaged hypostomes (Pl. 11, figs 11, 15) appear to have malformed lateral and posterior borders, although the deformation could be breakage.
<bibRefCitation id="3C4A4B0CFFD9FFE0E79EF82A7BF9F848" author="Demeter, E. J." box="[758,928,1952,1976]" pageId="14" pageNumber="15" pagination="37 - 65" refId="ref25624" refString="Demeter, E. J. (1973) Lower Ordovician pliomerid trilobites from western Utah. Brigham Young University Geology Studies, 20, 37 - 65." type="journal article" year="1973">Demeter (1973</bibRefCitation>
, pl. 3, figs. 9, 10, 14) figured two pygidia and a cranidium of
<taxonomicName id="9FDB4D7EFFD9FFE0E429F84E79EFF82C" authorityName="Adrain" authorityYear="2011" box="[321,438,1989,2012]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="stegneri">
<emphasis id="6AAFEAEFFFD9FFE0E429F84E79EFF82C" box="[321,438,1989,2012]" italics="true" pageId="14" pageNumber="15">P. stegneri</emphasis>
</taxonomicName>
as
<taxonomicName id="9FDB4D7EFFD9FFE0E4B4F84E7AFBF82C" baseAuthorityName="Ross" baseAuthorityYear="1951" box="[476,674,1988,2012]" class="Trilobita" family="Pliomeridae" genus="Hintzeia" kingdom="Animalia" order="Phacopida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="celsaora">
<emphasis id="6AAFEAEFFFD9FFE0E4B4F84E7AFBF82C" box="[476,674,1988,2012]" italics="true" pageId="14" pageNumber="15">Hintzeia celsaora</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="586436FDFFD8FFE1E5AFFF137ADBFE5C" blockId="15.[151,1436,151,680]" pageId="15" pageNumber="16">
<taxonomicName id="9FDB4D7EFFD8FFE1E5AFFF1379BFFF5F" authorityName="Adrain" authorityYear="2011" box="[199,486,152,176]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="stegneri">
<emphasis id="6AAFEAEFFFD8FFE1E5AFFF1379BFFF5F" box="[199,486,152,176]" italics="true" pageId="15" pageNumber="16">Protopliomerella stegneri</emphasis>
</taxonomicName>
is most morphologically similar to
<taxonomicName id="9FDB4D7EFFD8FFE1E61FFF127BAAFF40" authorityName="Adrain" authorityYear="2011" box="[887,1011,152,176]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="kerouaci">
<emphasis id="6AAFEAEFFFD8FFE1E61FFF127BAAFF40" box="[887,1011,152,176]" italics="true" pageId="15" pageNumber="16">P. kerouaci</emphasis>
</taxonomicName>
, but can be distinguished by its wider anterior border, shorter and wider glabella, weakly developed S4, longer and more posterolaterally located palpebro-ocular ridges; a hypostome with much wider anterior wings, shorter border spines, shorter posterior lobe of the middle body, and coarser, more sparsely spaced tubercles on the anterior lobe of the middle body; a slightly shorter and much wider librigena with a lower eye and less raised eye socle area, longer anterior branch of the facial suture along the field, and less pitted field; and a relatively shorter and wider pygidium with a less tapered posterior margin, less posteromedially directed pleurae, wider and slightly more separated pleural spine tips, a less complex terminal piece impression, and finer sculpture.
</paragraph>
<paragraph id="586436FDFFD8FFE1E5AFFE337B25FD74" blockId="15.[151,1436,151,680]" pageId="15" pageNumber="16">
<taxonomicName id="9FDB4D7EFFD8FFE1E5AFFE3379B0FE3F" authorityName="Adrain" authorityYear="2011" box="[199,489,440,464]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="stegneri">
<emphasis id="6AAFEAEFFFD8FFE1E5AFFE3379B0FE3F" box="[199,489,440,464]" italics="true" pageId="15" pageNumber="16">Protopliomerella stegneri</emphasis>
</taxonomicName>
is stratigraphically closest to
<taxonomicName id="9FDB4D7EFFD8FFE1E62CFE327BEEFE20" authorityName="Adrain" authorityYear="2011" box="[836,951,440,464]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bowlesi">
<emphasis id="6AAFEAEFFFD8FFE1E62CFE327BEEFE20" box="[836,951,440,464]" italics="true" pageId="15" pageNumber="16">P. bowlesi</emphasis>
</taxonomicName>
, but the latter taxon possesses a narrower anterior border, a considerably longer and narrower glabella with a much more clearly impressed S4, narrower interocular fixigenae and more anteriorly located palpebro-ocular ridges, more steeply sloped fixigenae; a librigena with a more evenly sloped posterior branch of the facial suture and a narrower librigenal field, particularly posteriorly; and a distinctive six-segmented pygidium with less inflated axial rings and pleurae, a smaller, narrower terminal piece, and narrower, much more closely spaced pleural spines.
</paragraph>
<paragraph id="586436FDFFD8FFE1E5AFFD1B7CF6FD57" blockId="15.[151,1436,151,680]" box="[199,1199,655,680]" pageId="15" pageNumber="16">
<taxonomicName id="9FDB4D7EFFD8FFE1E5AFFD1B79BFFD57" authorityName="Adrain" authorityYear="2011" box="[199,486,656,680]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="stegneri">
<emphasis id="6AAFEAEFFFD8FFE1E5AFFD1B79BFFD57" box="[199,486,656,680]" italics="true" pageId="15" pageNumber="16">Protopliomerella stegneri</emphasis>
</taxonomicName>
is compared to
<taxonomicName id="9FDB4D7EFFD8FFE1E7F2FD1A7B5FFD57" authorityName="Adrain" authorityYear="2011" box="[666,774,656,680]" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" kingdom="Animalia" order="Phacopida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="seegeri">
<emphasis id="6AAFEAEFFFD8FFE1E7F2FD1A7B5FFD57" box="[666,774,656,680]" italics="true" pageId="15" pageNumber="16">P. seegeri</emphasis>
</taxonomicName>
in the discussion of the latter species.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.31.6251" ID-PMC="PMC3881344" ID-Pensoft-Pub="1314-2003-31-21" ID-Pensoft-UUID="3273FFA3FF812C50FFE0FF98024A9177" ID-PubMed="24399902" ID-Zenodo-Dep="576190" ModsDocID="1314-2003-31-21" checkinTime="1451251795732" checkinUser="pensoft" docAuthor="Ickert-Bond, Stefanie M. &amp; Wen, Jun" docDate="2013" docId="D07290A615D852A1BB3E5448F7942605" docLanguage="en" docName="PhytoKeys 31: 21-61" docOrigin="PhytoKeys 31" docPubDate="2013-12-17" docSource="http://dx.doi.org/10.3897/phytokeys.31.6251" docTitle="Liquidambar gracilipes Ickert-Bond &amp; J. Wen 2013, comb. nov." docType="treatment" docVersion="3" id="3273FFA3FF812C50FFE0FF98024A9177" lastPageNumber="33" masterDocId="3273FFA3FF812C50FFE0FF98024A9177" masterDocTitle="A taxonomic synopsis of Altingiaceae with nine new combinations" masterLastPageNumber="61" masterPageNumber="21" pageNumber="33" updateTime="1668140986105" updateUser="ExternalLinkService">
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<mods:title>A taxonomic synopsis of Altingiaceae with nine new combinations</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ickert-Bond, Stefanie M.</mods:namePart>
<mods:affiliation>UA Museum of the North Herbarium (ALA), Department of Biology and Wildlife, University of Alaska Fairbanks, 907 Yukon Drive, Fairbanks, Alaska 99775 - 6590 USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wen, Jun</mods:namePart>
<mods:affiliation>Department of Botany, MRC- 166, Smithsonian Institution, P. O. Box 37012, Washington, D. C. 20013 - 7012 USA</mods:affiliation>
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<treatment ID-GBIF-Taxon="182228863" LSID="urn:lsid:plazi:treatment:D07290A615D852A1BB3E5448F7942605" httpUri="http://treatment.plazi.org/id/D07290A615D852A1BB3E5448F7942605" lastPageId="13" lastPageNumber="33" pageId="12" pageNumber="33">
<subSubSection pageId="12" pageNumber="33" type="nomenclature">
<paragraph pageId="12" pageNumber="33">
8.
<taxonomicName LSID="D07290A6-15D8-52A1-BB3E-5448F7942605" authority="(Hemsl.) Ickert-Bond &amp; J. Wen" authorityName="Ickert-Bond &amp; J. Wen" authorityYear="2013" baseAuthorityName="Ickert-Bond &amp; J. Wen" baseAuthorityYear="2013" class="Dicotyledoneae" family="Hamamelidaceae" genus="Liquidambar" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Liquidambar gracilipes" order="Rosales" pageId="12" pageNumber="33" phylum="Angiospermae" rank="species" species="gracilipes" status="comb. nov.">Liquidambar gracilipes (Hemsl.) Ickert-Bond &amp; J. Wen</taxonomicName>
<taxonomicNameLabel pageId="12" pageNumber="33">comb. nov.</taxonomicNameLabel>
Basionym:
<taxonomicName authorityName="Ickert-Bond &amp; Wen" authorityYear="2013" baseAuthorityName="Ickert-Bond &amp; J. Wen" baseAuthorityYear="2013" class="Dicotyledoneae" family="Hamamelidaceae" genus="Altingia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Rosales" pageId="12" pageNumber="33" phylum="Angiospermae" rank="species" species="gracilipes" status="comb. nov.">
<emphasis italics="true" pageId="12" pageNumber="33">Altingia gracilipes</emphasis>
</taxonomicName>
Hemsl., in Hook. Ic. Pl. t. 2837, 1907. TYPE. CHINA: Fujian,
<emphasis italics="true" pageId="12" pageNumber="33">S.T. Dunn</emphasis>
Hb. Hongk. 2682. (Lectotype here designated HK [HK 10948!]; isolectotypes: A [A 00043390!], IBSC [IBSC 001001!])
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="33" type="reference_group">
<paragraph pageId="12" pageNumber="33">
<taxonomicName authorityName="Ickert-Bond &amp; Wen" authorityYear="2013" genus="Altingi" lsidName="" pageId="12" pageNumber="33" rank="species" species="gracilipes" status="comb. nov.">Altingi gracilipes</taxonomicName>
Hemsl. var.
<emphasis italics="true" pageId="12" pageNumber="33">serrulata</emphasis>
Tutcher, Hong Kong Administrative Report 1914: M31. TYPE. CHINA, Hong Kong, Ukautang,
<emphasis italics="true" pageId="12" pageNumber="33">W.J. Tutcher</emphasis>
Herb. Hongkong No. 10947) (holotype: HK [HK 10949!, collected 25/5/1914]; isotypes: HK [Garden Department Hong Kong 1711!, collected 25/5/1914], HK [HK10950!])
</paragraph>
<paragraph pageId="12" pageNumber="33">
<taxonomicName authorityName="Ickert-Bond &amp; Wen" authorityYear="2013" genus="Altinga" lsidName="" pageId="12" pageNumber="33" rank="species" species="gracilipes" status="comb. nov.">Altinga gracilipes</taxonomicName>
Hemsl. f.
<emphasis italics="true" pageId="12" pageNumber="33">uniflora</emphasis>
H. T. Chang, Sunyatsenia 7: 74: 1948. TYPE. CHINA, Fujian,
<emphasis italics="true" pageId="12" pageNumber="33">Y. Ling 2295</emphasis>
(holotype: IBSC 001003!)
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="34" pageId="12" pageNumber="33" type="distribution">
<paragraph pageId="12" pageNumber="33">Distribution.</paragraph>
<paragraph lastPageId="13" lastPageNumber="34" pageId="12" pageNumber="33">
China (Fujian, Guangdong, Hong Kong, and Zhejiang).
<emphasis bold="true" pageId="12" pageNumber="33">Representative specimens examined.</emphasis>
<emphasis bold="true" pageId="12" pageNumber="33">CHINA:Fujian</emphasis>
: Jian Ou, Wang Mu Lin,
<emphasis italics="true" pageId="12" pageNumber="33">Fujian Forestry Coll. 41</emphasis>
(MO); Long Yan, Jiang Shan, Shuang Che,
<emphasis italics="true" pageId="12" pageNumber="33">H. Chen 1272</emphasis>
(F, MO); Nan Ping, Mang Tang Mountains,
<emphasis italics="true" pageId="12" pageNumber="33">G. He 4690</emphasis>
(MO),
<emphasis italics="true" pageId="12" pageNumber="33">G. He 5479</emphasis>
(MO),
<emphasis italics="true" pageId="12" pageNumber="33">G. He 6155</emphasis>
(MO); Yong An, Hong Tian,
<emphasis italics="true" pageId="12" pageNumber="33">Fujian Team 5567</emphasis>
(MO); Yungyung Mts., Fujian, central,
<emphasis italics="true" pageId="12" pageNumber="33">S.T. Dunn 2682</emphasis>
(GH);
<emphasis bold="true" pageId="12" pageNumber="33">Guangdong</emphasis>
: Chaochow district,
<emphasis italics="true" pageId="12" pageNumber="33">N.K. Chun 42718</emphasis>
(MO); Da Pu, Yan Shang, Min Ying,
<emphasis italics="true" pageId="12" pageNumber="33">L. Deng 5438</emphasis>
(MO); Da Pu, Feng Xi,
<emphasis italics="true" pageId="12" pageNumber="33">X. Wang 153</emphasis>
(MO); Da Pu, Ying Jian,
<emphasis italics="true" pageId="12" pageNumber="33">Z. Li 963</emphasis>
(MO); Da Pu, Pu Cheng,
<emphasis italics="true" pageId="12" pageNumber="33">X. Li 202545</emphasis>
(MO); Feng Xun, Da Tian, Bei Xi,
<emphasis italics="true" pageId="12" pageNumber="33">X. Li 200921</emphasis>
(MO); He Yuan, Xin Feng Jian,
<emphasis italics="true" pageId="12" pageNumber="33">B. Yu 103106</emphasis>
(MO); He Ping, Qing Zhou,
<emphasis italics="true" pageId="12" pageNumber="33">Z. Wei 120579</emphasis>
(MO); Jiu Lian Xiang, Xiao Shui,
<emphasis italics="true" pageId="12" pageNumber="33">Z. Wei 120217</emphasis>
(MO); Lian Ping, Zhong xin, Da Shui,
<emphasis italics="true" pageId="12" pageNumber="33">Y. Liu 242</emphasis>
(MO); Ping Yuan,
<emphasis italics="true" pageId="12" pageNumber="33">L. Dong 4331</emphasis>
(MO); Qiao Lin, Shi Hu,
<emphasis italics="true" pageId="12" pageNumber="33">L. Dong 4685</emphasis>
(MO); Raoping Xian,
<emphasis italics="true" pageId="12" pageNumber="33">N.K. Chun 42718</emphasis>
(2 sheets, MO); Rao Ping, Feng Huang,
<emphasis italics="true" pageId="12" pageNumber="33">X. Li 200701</emphasis>
(MO); Rao Ping, Feng Huang Yu,
<emphasis italics="true" pageId="12" pageNumber="33">H. Zeng 72101</emphasis>
(MO); Tung Koo Shan, Tapu district, Tan Shue,
<emphasis italics="true" pageId="12" pageNumber="33">W.T. Tsang 21697</emphasis>
(GH, HK, P); Wu Hua, Shuang
<pageBreakToken pageId="13" pageNumber="34" start="start">Hua</pageBreakToken>
,
<emphasis italics="true" pageId="13" pageNumber="34">Z. Li 1162</emphasis>
(MO); Wu Hua, Qi Mu Zhang,
<emphasis italics="true" pageId="13" pageNumber="34">Y.-J. Wang 56484</emphasis>
(MO); Xin Yi, Da Wu Lin,
<emphasis italics="true" pageId="13" pageNumber="34">Z. Feng 36293</emphasis>
(MO); Yam Na Shan [Yit Nga Shan] Mei [Kaying] District,
<emphasis italics="true" pageId="13" pageNumber="34">W.T. Tsang 21514</emphasis>
(GH), Yunfu Xian,
<emphasis italics="true" pageId="13" pageNumber="34">C. Wang 37057</emphasis>
(MO);
<emphasis bold="true" pageId="13" pageNumber="34">Hong Kong</emphasis>
:
<emphasis italics="true" pageId="13" pageNumber="34">M.T. Sin &amp; W.K. Woo s.n</emphasis>
. (HK); Pat Sin Leng Country Park, Nam Chung trail,
<emphasis italics="true" pageId="13" pageNumber="34">S. Ickert-Bond 1272</emphasis>
(F); Nam Chung,
<emphasis italics="true" pageId="13" pageNumber="34">D. Lau 43</emphasis>
(GH, MO),
<emphasis italics="true" pageId="13" pageNumber="34">T.W. Lau 98</emphasis>
(GH, MO),
<emphasis italics="true" pageId="13" pageNumber="34">Y.-W. Lam 1429</emphasis>
(HK),
<emphasis italics="true" pageId="13" pageNumber="34">K.-L. Yip 4082</emphasis>
(MO); Pat Sin Leng, Plover Cove Country Park, Plover Cove Reservoir,
<emphasis italics="true" pageId="13" pageNumber="34">S. Ickert-Bond 1266</emphasis>
(F); Pat Sing Leng Nature Trails,
<emphasis italics="true" pageId="13" pageNumber="34">L.T. Lo 629</emphasis>
(HK); Sha Tau Kok-luk Keng,
<emphasis italics="true" pageId="13" pageNumber="34">S.Y. Hu 9984</emphasis>
(GH); Wu Kau Tan N.T.,
<emphasis italics="true" pageId="13" pageNumber="34">S.Y. Hu &amp; Y.C. Kong 175</emphasis>
(MO),
<emphasis italics="true" pageId="13" pageNumber="34">Y.S. Lau 2634</emphasis>
(HK);
<emphasis bold="true" pageId="13" pageNumber="34">Zhejiang</emphasis>
: Between Ping Yung and Tai Suan,
<emphasis italics="true" pageId="13" pageNumber="34">R.-C. Ching 2199</emphasis>
(E); Feng Yang Mtn.,
<emphasis italics="true" pageId="13" pageNumber="34">H.-Y. Zou 79</emphasis>
(MO),
<emphasis italics="true" pageId="13" pageNumber="34">H.-Y. Zou 151</emphasis>
(GH),
<emphasis italics="true" pageId="13" pageNumber="34">H.-Y. Zou 762</emphasis>
(GH); Long Qian, Ju Shui,
<emphasis italics="true" pageId="13" pageNumber="34">P.L. Chiu 1072</emphasis>
(MO); Qing Yuan, Wu Du Mountain,
<emphasis italics="true" pageId="13" pageNumber="34">Z.G. Mao 10231</emphasis>
(MO); Shui Chang, Pu Nang,
<emphasis italics="true" pageId="13" pageNumber="34">P.L. Chiu 1470</emphasis>
(MO); Taishun Hsien,
<emphasis italics="true" pageId="13" pageNumber="34">Y.L. Keng 316</emphasis>
(GH).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,140 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1093.72339" ID-GBIF-Dataset="8bb856a5-26cb-4abc-8e4a-04d3716144b1" ID-PMC="PMC9010403" ID-Pensoft-Pub="1313-2970-1093-1" ID-Pensoft-UUID="D3A8D50FF61A5B61B8776D63EB0D3F4C" ID-PubMed="35586542" ID-ZooBank="23B7070849A94681AC20494D06F98CCE" ModsDocID="1313-2970-1093-1" checkinTime="1649449468463" checkinUser="pensoft" docAuthor="Benda, Daniel, Pohl, Hans, Nakase, Yuta, Beutel, Rolf &amp; Straka, Jakub" docDate="2022" docId="D07299E98BCD5F5E86E7B8243B4C0215" docLanguage="en" docName="ZooKeys 1093: 1-134" docOrigin="ZooKeys 1093" docPubDate="2022-04-07" docSource="http://dx.doi.org/10.3897/zookeys.1093.72339" docTitle="Eupathocera mendozae Benda &amp; Pohl &amp; Nakase &amp; Beutel &amp; Straka 2022, comb. nov." docType="treatment" docVersion="4" id="D3A8D50FF61A5B61B8776D63EB0D3F4C" lastPageNumber="1" masterDocId="D3A8D50FF61A5B61B8776D63EB0D3F4C" masterDocTitle="A generic classification of Xenidae (Strepsiptera) based on the morphology of the female cephalothorax and male cephalotheca with a preliminary checklist of species" masterLastPageNumber="134" masterPageNumber="1" pageNumber="1" updateTime="1668151620981" updateUser="ExternalLinkService">
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<mods:title>A generic classification of Xenidae (Strepsiptera) based on the morphology of the female cephalothorax and male cephalotheca with a preliminary checklist of species</mods:title>
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<mods:namePart>Benda, Daniel</mods:namePart>
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<mods:affiliation>Department of Zoology, Faculty of Science, Charles University, Prague, Czech Republic &amp; Department of Entomology, National Museum, Prague, Czech Republic</mods:affiliation>
<mods:nameIdentifier type="email">benda.daniel@email.cz</mods:nameIdentifier>
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<mods:namePart>Pohl, Hans</mods:namePart>
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<mods:affiliation>Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet, Jena, Germany</mods:affiliation>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nakase, Yuta</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Shinshu University, Matsumoto, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Beutel, Rolf</mods:namePart>
<mods:affiliation>Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet, Jena, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Straka, Jakub</mods:namePart>
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<mods:affiliation>Department of Zoology, Faculty of Science, Charles University, Prague, Czech Republic</mods:affiliation>
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<mods:date>2022</mods:date>
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<mods:number>2022-04-07</mods:number>
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<treatment ID-GBIF-Taxon="194520467" LSID="urn:lsid:plazi:treatment:D07299E98BCD5F5E86E7B8243B4C0215" httpUri="http://treatment.plazi.org/id/D07299E98BCD5F5E86E7B8243B4C0215" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="D07299E9-8BCD-5F5E-86E7-B8243B4C0215" authority="(Brethes, 1923)" authorityName="Benda &amp; Pohl &amp; Nakase &amp; Beutel &amp; Straka" authorityYear="2022" baseAuthorityName="Brethes" baseAuthorityYear="1923" class="Insecta" family="Xenidae" genus="Eupathocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupathocera mendozae" order="Strepsiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mendozae" status="comb. nov.">
Eupathocera mendozae (
<normalizedToken originalValue="Brèthes">Brethes</normalizedToken>
, 1923)
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">comb. nov.</taxonomicNameLabel>
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</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Benda &amp; Pohl &amp; Nakase &amp; Beutel &amp; Straka" authorityYear="2022" class="Insecta" family="Xenidae" genus="Ophthalmochlus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophthalmochlus (Homilops) mendozae" order="Strepsiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mendozae" status="comb. nov." subGenus="Homilops">Ophthalmochlus (Homilops) mendozae</taxonomicName>
<normalizedToken originalValue="Brèthes">Brethes</normalizedToken>
, 1923: 51.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Benda &amp; Pohl &amp; Nakase &amp; Beutel &amp; Straka" authorityYear="2022" class="Insecta" family="Xenidae" genus="Pseudoxenos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudoxenos mendozae" order="Strepsiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mendozae" status="comb. nov.">Pseudoxenos mendozae</taxonomicName>
(
<normalizedToken originalValue="Brèthes">Brethes</normalizedToken>
, 1923) (new combination by
<bibRefCitation DOI="https://doi.org/10.1155/1937/28072" author="Bohart, RM" journalOrPublisher="Psyche: A Journal of Entomology" pageId="0" pageNumber="1" pagination="132 - 137" refId="B7" refString="Bohart, RM, 1937. Synonymy of the Genus Pseudoxenos Saunders (Strepsiptera, Xenidae) and records of stylopized Hymenoptera from North Carolina. Psyche: A Journal of Entomology 44: 132 - 137, DOI: https://doi.org/10.1155/1937/28072" title="Synonymy of the Genus Pseudoxenos Saunders (Strepsiptera, Xenidae) and records of stylopized Hymenoptera from North Carolina." url="https://doi.org/10.1155/1937/28072" volume="44" year="1937">Bohart 1937</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Benda &amp; Pohl &amp; Nakase &amp; Beutel &amp; Straka" authorityYear="2022" class="Insecta" family="Xenidae" genus="Paraxenos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraxenos mendozae" order="Strepsiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mendozae" status="comb. nov.">Paraxenos mendozae</taxonomicName>
(
<normalizedToken originalValue="Brèthes">Brethes</normalizedToken>
, 1923) (new combination by
<bibRefCitation author="Kinzelbach, R" journalOrPublisher="Schweizerbart'sche Verlagsbuchhandlung, Stuttgart" pageId="0" pageNumber="1" refId="B54" refString="Kinzelbach, R, 1971b. Morphologische Befunde an Faecherflueglern und ihre phylogenetische Bedeutung (Insecta: Strepsiptera). Schweizerbart'sche Verlagsbuchhandlung, Stuttgart" title="Morphologische Befunde an Faecherflueglern und ihre phylogenetische Bedeutung (Insecta: Strepsiptera)." year="1971 b">Kinzelbach 1971b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="host">
<paragraph pageId="0" pageNumber="1">Host.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Kohl" baseAuthorityYear="1890" class="Insecta" family="Sphecidae" genus="Prionyx" kingdom="Animalia" lsidName="Prionyx neoxenus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="neoxenus">
<emphasis italics="true" pageId="0" pageNumber="1">Prionyx neoxenus</emphasis>
</taxonomicName>
(Kohl, 1890) (as
<taxonomicName baseAuthorityName="Kohl" baseAuthorityYear="1890" class="Insecta" family="Sphecidae" genus="Priononyx" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Priononyx neoxenus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="neoxenus">
<emphasis italics="true" pageId="0" pageNumber="1">Priononyx neoxenus</emphasis>
</taxonomicName>
, var.
<taxonomicName class="Insecta" family="Sphecidae" genus="Priononyx" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Priononyx neoxenus var. melanogaster" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="variety" species="neoxenus" variety="melanogaster">Priononyx neoxenus melanogaster</taxonomicName>
<normalizedToken originalValue="Brèthes">Brethes</normalizedToken>
, 1910) (
<bibRefCitation author="Brethes, J" journalOrPublisher="Revista de la Facultad de Agronomia de la Plata" pageId="0" pageNumber="1" pagination="41 - 56" refId="B10" refString="Brethes, J, 1923. Primera contribucion para el conocimiento de los &quot; Strepsiptera &quot; argentinos. Revista de la Facultad de Agronomia de la Plata 15: 41 - 56" title="Primera contribucion para el conocimiento de los &quot; Strepsiptera &quot; argentinos." volume="15" year="1923">
<normalizedToken originalValue="Brèthes">Brethes</normalizedToken>
1923
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Argentina: Mendoza (
<bibRefCitation author="Brethes, J" journalOrPublisher="Revista de la Facultad de Agronomia de la Plata" pageId="0" pageNumber="1" pagination="41 - 56" refId="B10" refString="Brethes, J, 1923. Primera contribucion para el conocimiento de los &quot; Strepsiptera &quot; argentinos. Revista de la Facultad de Agronomia de la Plata 15: 41 - 56" title="Primera contribucion para el conocimiento de los &quot; Strepsiptera &quot; argentinos." volume="15" year="1923">
<normalizedToken originalValue="Brèthes">Brethes</normalizedToken>
1923
</bibRefCitation>
).
</paragraph>
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View file

@ -0,0 +1,123 @@
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<treatment ID-GBIF-Taxon="127902641" LSID="urn:lsid:plazi:treatment:D072A4E6F71F314658D2236C70A7038D" httpUri="http://treatment.plazi.org/id/D072A4E6F71F314658D2236C70A7038D" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44">
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<paragraph pageId="1" pageNumber="44">Taxon classification Animalia Lepidoptera Coleophoridae</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="44" type="nomenclature">
<paragraph pageId="1" pageNumber="44">
<taxonomicName authority="Toll, 1956" authorityName="Toll" authorityYear="1956" class="Insecta" family="Coleophoridae" genus="Coleophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coleophora micronotella" order="Lepidoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="micronotella">Coleophora micronotella Toll, 1956</taxonomicName>
Figs 1-4
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="44" type="material">
<paragraph pageId="1" pageNumber="44">Material.</paragraph>
<paragraph pageId="1" pageNumber="44">
2 ♂ (one dissected, slide MHNG ENTO 5970), ♀ [Italy] 'SARDEGNA, Orien. | La Caletta | 2.VIII.81-lux | leg.
<normalizedToken originalValue="Baldizzone">Baldizzone'</normalizedToken>
; 1 ♂ (head prepared for SEM), same data except
<normalizedToken originalValue="29.VII.81">'29.VII.81'</normalizedToken>
; 2 ♂ (one dissected, slide MHNG ENTO 7501), 'SARDEGNA, Merid. | Porto Botte | 23.VII.81-lux | leg.
<normalizedToken originalValue="Baldizzone">Baldizzone'</normalizedToken>
, deposited in MHNG. 1 ♂ (dissected, slide MHNG ENTO 7503) [Tunisia] 'Mauretania | Tunesia-Sud | Oase Tozeur | 28.4.-11.5.1981 | leg. M. u. W.
<normalizedToken originalValue="Glaser">Glaser'</normalizedToken>
, deposited in MHNG. Additional material, undissected: 194 specimens from Sardinia (leg. Baldizzone); 30 from Tunisia (24 from Tozeur, 1921, leg. Dumont; 7 from Tozeur e Gabes, 1980, leg. Glaser); deposited in collection Baldizzone.
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44" type="description">
<paragraph pageId="1" pageNumber="44">Description of head.</paragraph>
<paragraph pageId="1" pageNumber="44">
Frontal aspect with appressed scales, with short, apparent scaled palpi, without proboscis. Ventral edge of fronto-clypeus slightly sinuate. Labrum not apparent. Maxillary palpi unsegmented, very short, with few scale sockets and setae, with apical sensillum styloconicum. Labial palpi slightly longer than maxillary palpi, reaching apices of subgenal apophyses, with two segments visible, apical segment about half as long as basal segment and lacking vom
<normalizedToken originalValue="Raths">Rath's</normalizedToken>
organ. With sac-like rounded structure laterally from labial palpi.
</paragraph>
<caption pageId="2" pageNumber="45">
<paragraph pageId="2" pageNumber="45">
<pageBreakToken pageId="2" pageNumber="45" start="start">Figures</pageBreakToken>
1-4. Head of
<taxonomicName class="Insecta" family="Coleophoridae" genus="Coleophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coleophora micronotella" order="Lepidoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="micronotella">Coleophora micronotella</taxonomicName>
Toll. 1. Frontal view. 2. Scanning electron micrograph (fc: fronto-clypeus; lp: labial palp, partly collapsed; mp; maxillary palpus; se: sensillum styloconicum). 3. Ventral view showing labial palpi. 4. Dorsal view showing maxillary palpi.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="2" pageNumber="45" type="notes">
<paragraph pageId="2" pageNumber="45">Notes.</paragraph>
<paragraph pageId="2" pageNumber="45">
The halophilous biotope of La Caletta, where three of the specimens of
<taxonomicName class="Insecta" family="Coleophoridae" genus="Coleophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coleophora micronotella" order="Lepidoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="micronotella">Coleophora micronotella</taxonomicName>
studied were collected, is about one kilometer away from the Mediterranean, on the sides of a small canal where tamarix (
<taxonomicName class="Magnoliopsida" family="Tamaricaceae" genus="Tamarix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tamarix" order="Caryophyllales" pageId="2" pageNumber="45" phylum="Tracheophyta" rank="genus">Tamarix</taxonomicName>
sp.,
<taxonomicName family="Tamaricaceae" genus="Tamaricaceae" lsidName="Tamaricaceae" pageId="2" pageNumber="45" rank="family">Tamaricaceae</taxonomicName>
) grows. The specimens were collected with a black light in a salt marsh where
<taxonomicName class="Magnoliopsida" family="Amaranthaceae" genus="Salicornia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salicornia" order="Caryophyllales" pageId="2" pageNumber="45" phylum="Tracheophyta" rank="genus">Salicornia</taxonomicName>
(
<taxonomicName family="Amaranthaceae" genus="Amaranthaceae" lsidName="Amaranthaceae" pageId="2" pageNumber="45" rank="family">Amaranthaceae</taxonomicName>
) and other halophilous herbaceous plants were growing. Hundreds of specimens came to light, but could not come to rest on the sheet because of the presence of thousands of staphylinid beetles and flies.
<bibRefCitation author="Falkovitsh, MI" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="4" pageNumber="47" pagination="167 - 176" title="New taxa of casebearers (Lepidoptera, Coleophoridae) from the desert zone of the Palaearctic." volume="84" year="2005">Falkovitsh (2005)</bibRefCitation>
recorded that the larva of
<taxonomicName class="Insecta" family="Coleophoridae" genus="Coleophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coleophora micronotella" order="Lepidoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="micronotella">Coleophora micronotella</taxonomicName>
feeds as a borer, without making a case, in 'green branches (assimilating shoots)' of
<taxonomicName class="Magnoliopsida" family="Amaranthaceae" genus="Halostachys" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Halostachys caspica" order="Caryophyllales" pageId="2" pageNumber="45" phylum="Tracheophyta" rank="species" species="caspica">Halostachys caspica</taxonomicName>
C.A. Mey. and
<taxonomicName class="Magnoliopsida" family="Amaranthaceae" genus="Halocnemum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Halocnemum strobilaceum" order="Caryophyllales" pageId="2" pageNumber="45" phylum="Tracheophyta" rank="species" species="strobilaceum">Halocnemum strobilaceum</taxonomicName>
(Pallas) Bieb. (
<taxonomicName family="Amaranthaceae" genus="Amaranthaceae" lsidName="Amaranthaceae" pageId="2" pageNumber="45" rank="family">Amaranthaceae</taxonomicName>
).
</paragraph>
<paragraph pageId="2" pageNumber="45">
Described from Tozeur, Tunisia, the species is also found in Italy (mainland, Sardinia, Sicily) and Spain. It has also been reported from Iran (
<bibRefCitation author="Baldizzone, G" journalOrPublisher="Apollo Books, Stenstrup" pageId="4" pageNumber="47" title="Contibuzioni alla conoscenza dei Coleophoridae. LXXV. Coleophoridae dell'Area Irano-Anatolica e regioni limitrofe (Lepidoptera). Associazone Naturalistica Piemontese, Memorie" year="1994">Baldizzone 1994</bibRefCitation>
), and Algeria, Kazakhstan, and Tadjikistan (
<bibRefCitation author="Falkovitsh, MI" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="4" pageNumber="47" pagination="167 - 176" title="New taxa of casebearers (Lepidoptera, Coleophoridae) from the desert zone of the Palaearctic." volume="84" year="2005">Falkovitsh 2005</bibRefCitation>
). However, a specimen from the Iranian series mentioned in
<bibRefCitation author="Baldizzone, G" journalOrPublisher="Apollo Books, Stenstrup" pageId="4" pageNumber="47" title="Contibuzioni alla conoscenza dei Coleophoridae. LXXV. Coleophoridae dell'Area Irano-Anatolica e regioni limitrofe (Lepidoptera). Associazone Naturalistica Piemontese, Memorie" year="1994">Baldizzone (1994)</bibRefCitation>
was re-examined and found to represent another, probably undescribed species. Its mouth parts are more developed than in
<taxonomicName class="Insecta" family="Coleophoridae" genus="Coleophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coleophora micronotella" order="Lepidoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="micronotella">Coleophora micronotella</taxonomicName>
, with the labial palpi about as long as 3/4 the widest diameter of the compound eye, but without apparent proboscis. Also, the male valva differs slightly in the distally larger and more evidently separated sacculus from the basally narrower valvula, and the phallus is narrow, needle-like, instead of spatulate.
</paragraph>
</subSubSection>
</treatment>
</document>