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<document id="D07BBB72C616301FE676B0A3B8C2D539" ID-DOI="10.5281/zenodo.185652" ID-GBIF-Dataset="10608bbc-e71a-4a9d-bcfb-4256ea6b0436" ID-ISSN="1175-5326" ID-Zenodo-Dep="185652" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460004875651" checkinUser="plazi" docAuthor="Goin, Francisco J., Zimicz, Natalia, Reyes, Martín De Los &amp; Soibelzon, Leopoldo" docDate="2009" docId="D05D87E3FFCCFFE963FF7D9CFF6CC3AB" docLanguage="en" docName="zt02005p046.pdf" docOrigin="Zootaxa 2005" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Thylophorops lorenzinii Goin, Zimicz, Reyes &amp; Soibelzon, 2009, new species" docType="treatment" docVersion="7" lastPageNumber="38" masterDocId="2C64FF9BFFCFFFEA63687D7EFF87C53C" masterDocTitle="A new large didelphid of the genus Thylophorops (Mammalia: Didelphimorphia: Didelphidae), from the late Tertiary of the Pampean Region (Argentina)" masterLastPageNumber="46" masterPageNumber="35" pageNumber="38" updateTime="1698233198446" updateUser="plazi">
<mods:mods id="0313D8B43B22C155703B8861CC621C45" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D671AD4FE13CA9576016DE46C46362FC">
<mods:title id="7BDF0F0421CD1A83A33388DF5EFA08A8">A new large didelphid of the genus Thylophorops (Mammalia: Didelphimorphia: Didelphidae), from the late Tertiary of the Pampean Region (Argentina)</mods:title>
</mods:titleInfo>
<mods:name id="C9195BDFE2345AF6EAFCE490BDAC7340" type="personal">
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<mods:namePart id="FFA90ED32432DE948F0F32B2C3F78C02">Goin, Francisco J.</mods:namePart>
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<mods:namePart id="4B6E5081C08255AC7AB78B35AF88747F">Zimicz, Natalia</mods:namePart>
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<mods:namePart id="4FD1AD330E53E2AC5D4935763FDAA662">Reyes, Martín De Los</mods:namePart>
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<mods:role id="70E7C52C86536622F8BD90FF8D74EE41">
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<mods:namePart id="94AF99791A7926ABB271E14FE18AB0BF">Soibelzon, Leopoldo</mods:namePart>
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<mods:typeOfResource id="9D490BCD77BE8909152F38EB4E932188">text</mods:typeOfResource>
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<mods:title id="35C20ED7863181D5669EFC13776A3A7D">Zootaxa</mods:title>
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<mods:date id="A9FB013DEC102633911B9D1E09E2A231">2009</mods:date>
<mods:detail id="49FCE2CB7BDC503AF9B549FB537A810C" type="volume">
<mods:number id="3432A65E0348AD4D6841A2472F4E185C">2005</mods:number>
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<mods:identifier id="20DADEFC6E6ED62CB5EDC3B61010FAE4" type="DOI">10.5281/zenodo.185652</mods:identifier>
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<mods:identifier id="05A7B44C3FA860E001FDE025EB3C2BB4" type="Zenodo-Dep">185652</mods:identifier>
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<treatment id="D05D87E3FFCCFFE963FF7D9CFF6CC3AB" ID-DOI="http://doi.org/10.5281/zenodo.6217764" ID-GBIF-Taxon="119380953" ID-Zenodo-Dep="6217764" LSID="urn:lsid:plazi:treatment:D05D87E3FFCCFFE963FF7D9CFF6CC3AB" httpUri="http://treatment.plazi.org/id/D05D87E3FFCCFFE963FF7D9CFF6CC3AB" lastPageNumber="38" pageId="3" pageNumber="38">
<subSubSection id="10EE657EFFCCFFE963FF7D9CFE83C423" pageId="3" pageNumber="38" type="nomenclature">
<paragraph id="584B36F5FFCCFFE963FF7D9CFDDFC5C0" blockId="3.[151,600,226,287]" box="[151,600,226,252]" pageId="3" pageNumber="38">
<heading id="03038199FFCCFFE963FF7D9CFDDFC5C0" bold="true" box="[151,600,226,252]" fontSize="11" level="1" pageId="3" pageNumber="38" reason="1">
<emphasis id="6A80EAE7FFCCFFE963FF7D9CFDDFC5C0" bold="true" box="[151,600,226,252]" pageId="3" pageNumber="38">
<taxonomicName id="9FF44D76FFCCFFE963FF7D9CFE3EC5C0" box="[151,441,226,252]" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="3" pageNumber="38" phylum="Chordata" rank="species" species="lorenzinii" status="sp. nov.">
<emphasis id="6A80EAE7FFCCFFE963FF7D9CFE3EC5C0" bold="true" box="[151,441,226,252]" italics="true" pageId="3" pageNumber="38">Thylophorops lorenzinii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="71B3579CFFCCFFE962A07D9CFDDFC5C0" box="[456,600,226,252]" pageId="3" pageNumber="38" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="584B36F5FFCCFFE963FF7C7BFE83C423" blockId="3.[151,600,226,287]" box="[151,260,261,287]" pageId="3" pageNumber="38">
<figureCitation id="C0CF2A70FFCCFFE963FF7C7BFE83C423" box="[151,260,261,287]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,255,1874,1898]" captionStartId-1="4.[151,255,1867,1891]" captionStartId-2="5.[151,255,1349,1373]" captionStartId-3="6.[151,255,1192,1216]" captionTargetBox-0="[230,1356,194,1848]" captionTargetBox-1="[231,1354,480,1839]" captionTargetBox-2="[151,1436,501,1338]" captionTargetBox-3="[234,1358,384,1169]" captionTargetId-0="figure@2.[230,1356,194,1850]" captionTargetId-1="figure@4.[231,1355,480,1841]" captionTargetId-2="figure@5.[151,1436,499,1341]" captionTargetId-3="figure@6.[234,1359,384,1170]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1: Thylophorops lorenzinii sp. nov. MLP 08 - III- 10 - 1 (Holotype); (A), labial view of the anterior part of the skull and dentary; (B) occlusal view of the dentary; (C) lingual view of the dentary. Scale: 10 mm." captionText-1="FIGURE 2: Thylophorops lorenzinii sp. nov. MLP 08 - III- 10 - 1 (Holotype); detail of the erupting m 3 in lingual (A), and occlusal-lingual views (B). Scale: 2 mm." captionText-2="FIGURE 3: Thylophorops lorenzinii sp. nov. MLP 08 - III- 10 - 1 (Holotype); schematic drawing of the m 3 showing the cusp homologies and measurements. Abbreviations: End., entoconid; Hyd., hypoconid; Hyld., hypoconulid; Med., metaconid; Pad., paraconid; Prod., protoconid. Scale: 2 mm." captionText-3="FIGURE 4: Lower left molars of Thylophorops chapalmalensis (left, detail of specimen MPH 064), T. lorenzinii sp. nov. (centre, detail of the type specimen), and T. perplanus (right, detail of specimen MLP 97 - XI- 15 - 1). Because of the erupting stage of the m 3 of T. lorenzinii in the type specimen, the anterobasal cingulum is not visible. Scale: 2 mm." httpUri-0="https://zenodo.org/record/185653/files/figure.png" httpUri-1="https://zenodo.org/record/185654/files/figure.png" httpUri-2="https://zenodo.org/record/185655/files/figure.png" httpUri-3="https://zenodo.org/record/185656/files/figure.png" pageId="3" pageNumber="38">Figs. 1–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="10EE657EFFCCFFE963FF7C2BFB9AC483" pageId="3" pageNumber="38" type="materials_examined">
<paragraph id="584B36F5FFCCFFE963FF7C2BFB9AC483" blockId="3.[151,1436,341,1687]" pageId="3" pageNumber="38">
<emphasis id="6A80EAE7FFCCFFE963FF7C2BFE97C453" bold="true" box="[151,272,341,367]" pageId="3" pageNumber="38">
<typeStatus id="874F8857FFCCFFE963FF7C2BFE8CC453" box="[151,267,341,367]" pageId="3" pageNumber="38" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="3EE5AE30FFCCFFE962707C2BFED2C453" box="[280,341,341,367]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="3" pageNumber="38">MLP</collectionCode>
<date id="2C4A1035FFCCFFE962357C2BFE64C453" box="[349,483,341,367]" pageId="3" pageNumber="38" value="2008-03-10" valueMax="2008-03-01" valueMin="2008-03-10">08-III-10-1</date>
, a fragment of skull preserving the left maxilla, and a left dentary preserving the second premolar (p2), the deciduous premolar (dp3) and an erupting third molar (m3), all belonging to the same, juvenile specimen. Collected by Leopoldo Soibelzon in January, 2008.
</paragraph>
</subSubSection>
<subSubSection id="10EE657EFFCCFFE963AE7CB3FD80C733" pageId="3" pageNumber="38" type="etymology">
<paragraph id="584B36F5FFCCFFE963AE7CB3FD80C733" blockId="3.[151,1436,341,1687]" pageId="3" pageNumber="38">
<emphasis id="6A80EAE7FFCCFFE963AE7CB3FED4C4DB" bold="true" box="[198,339,461,487]" pageId="3" pageNumber="38">Etymology.</emphasis>
Honouring Mr. Silvio Lorenzini, outstanding amateur collector of fossil specimens in Southeastern Buenos Aires Province.
</paragraph>
</subSubSection>
<subSubSection id="10EE657EFFCCFFE963AE7F63FDCFC793" pageId="3" pageNumber="38" type="description">
<paragraph id="584B36F5FFCCFFE963AE7F63FDCFC793" blockId="3.[151,1436,341,1687]" pageId="3" pageNumber="38">
<emphasis id="6A80EAE7FFCCFFE963AE7F63FE02C70B" bold="true" box="[198,389,541,567]" pageId="3" pageNumber="38">Measurements.</emphasis>
Extra-alveolar length of upper canine (C): 10.15; CL: 4.33; CW: 3.41; height of dentary at m3: 11.87; width of dentary at same point: 8.08; length of the p2-m3 series:
<quantity id="9F0C9B10FFCCFFE967257F3BFB4EC763" box="[1101,1225,581,607]" metricMagnitude="-2" metricUnit="m" metricValue="4.002" pageId="3" pageNumber="38" unit="mm" value="40.02">40.02 mm</quantity>
; p2L: 7.64; p2W: 3.22; dp3L: 5.76; dp3W: 2.83; m3L (total): 9.69; trigonid m3L: 4.22; talonid m3L: 5.47; talonid m3W: 4.31; trigonid m3W: 4.38 (see also
<figureCitation id="C0CF2A70FFCCFFE9629A7FEBFDBFC793" box="[498,568,661,687]" captionStart="FIGURE 3" captionStartId="5.[151,255,1349,1373]" captionTargetBox="[151,1436,501,1338]" captionTargetId="figure@5.[151,1436,499,1341]" captionTargetPageId="5" captionText="FIGURE 3: Thylophorops lorenzinii sp. nov. MLP 08 - III- 10 - 1 (Holotype); schematic drawing of the m 3 showing the cusp homologies and measurements. Abbreviations: End., entoconid; Hyd., hypoconid; Hyld., hypoconulid; Med., metaconid; Pad., paraconid; Prod., protoconid. Scale: 2 mm." httpUri="https://zenodo.org/record/185655/files/figure.png" pageId="3" pageNumber="38">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="10EE657EFFCCFFE963AE7FC3FF6BC673" pageId="3" pageNumber="38" type="diagnosis">
<paragraph id="584B36F5FFCCFFE963AE7FC3FF6BC673" blockId="3.[151,1436,341,1687]" pageId="3" pageNumber="38">
<emphasis id="6A80EAE7FFCCFFE963AE7FC3FEC1C7EB" bold="true" box="[198,326,701,727]" pageId="3" pageNumber="38">Diagnosis.</emphasis>
Largest known didelphid; differs from the other species of
<taxonomicName id="9FF44D76FFCCFFE967797FC3FB34C7EA" box="[1041,1203,701,726]" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="3" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="6A80EAE7FFCCFFE967797FC3FB34C7EA" box="[1041,1203,701,726]" italics="true" pageId="3" pageNumber="38">Thylophorops</emphasis>
</taxonomicName>
in having its lower molars with proportionally large hypoconulids which are not antero-posteriorly compressed; labially salient hypoconids; p2 is proportionally larger, higher, and posteriorly wider than in
<taxonomicName id="9FF44D76FFCCFFE967517E73FA88C61A" box="[1081,1295,781,806]" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="3" pageNumber="38" phylum="Chordata" rank="species" species="chapalmalensis">
<emphasis id="6A80EAE7FFCCFFE967517E73FA88C61A" box="[1081,1295,781,806]" italics="true" pageId="3" pageNumber="38">T. chapalmalensis</emphasis>
</taxonomicName>
and
<taxonomicName id="9FF44D76FFCCFFE966247E73FF62C672" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="3" pageNumber="38" phylum="Chordata" rank="species" species="perplanus">
<emphasis id="6A80EAE7FFCCFFE966247E73FF62C672" italics="true" pageId="3" pageNumber="38">T. perplanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="10EE657EFFCCFFE963AE7E23FF6CC3AB" pageId="3" pageNumber="38" type="description">
<paragraph id="584B36F5FFCCFFE963AE7E23FACBC103" blockId="3.[151,1436,341,1687]" pageId="3" pageNumber="38">
<emphasis id="6A80EAE7FFCCFFE963AE7E23FDA3C64B" bold="true" box="[198,548,861,887]" pageId="3" pageNumber="38">
Description of the
<typeStatus id="874F8857FFCCFFE962D97E23FD98C64B" box="[433,543,861,887]" pageId="3" pageNumber="38" type="holotype">holotype</typeStatus>
.
</emphasis>
Specimen MLP
<date id="2C4A1035FFCCFFE961877E23FCF4C64B" box="[751,883,861,887]" pageId="3" pageNumber="38" value="2008-03-10" valueMax="2008-03-01" valueMin="2008-03-10">08-III-10-1</date>
belongs to a juvenile individual, as evidenced by the persistence of a dp3 and an erupting m3 which is placed immediately anterior to the masseteric crest (i.e. m4 is not observable). Only the anterior part of the snout is preserved (
<figureCitation id="C0CF2A70FFCCFFE9674F7ED3FBECC6FB" box="[1063,1131,941,967]" captionStart="FIGURE 1" captionStartId="2.[151,255,1874,1898]" captionTargetBox="[230,1356,194,1848]" captionTargetId="figure@2.[230,1356,194,1850]" captionTargetPageId="2" captionText="FIGURE 1: Thylophorops lorenzinii sp. nov. MLP 08 - III- 10 - 1 (Holotype); (A), labial view of the anterior part of the skull and dentary; (B) occlusal view of the dentary; (C) lingual view of the dentary. Scale: 10 mm." httpUri="https://zenodo.org/record/185653/files/figure.png" pageId="3" pageNumber="38">Fig. 1</figureCitation>
A), which is broken. The dentary (
<figureCitation id="C0CF2A70FFCCFFE963977EABFEC7C6D3" box="[255,320,981,1007]" captionStart="FIGURE 1" captionStartId="2.[151,255,1874,1898]" captionTargetBox="[230,1356,194,1848]" captionTargetId="figure@2.[230,1356,194,1850]" captionTargetPageId="2" captionText="FIGURE 1: Thylophorops lorenzinii sp. nov. MLP 08 - III- 10 - 1 (Holotype); (A), labial view of the anterior part of the skull and dentary; (B) occlusal view of the dentary; (C) lingual view of the dentary. Scale: 10 mm." httpUri="https://zenodo.org/record/185653/files/figure.png" pageId="3" pageNumber="38">Fig. 1</figureCitation>
B–C) is partially broken and lacks the incisor region and the angular process; it is quite robust, especially under m3. The posterior end of the symphysis reaches the distal margin of p2. The mental foramen is in line with the mesial root of m1. The distal root of p1, and complete p2, dp3 and m3 are preserved.
</paragraph>
<paragraph id="584B36F5FFCCFFE963AF7933FB36C1B3" blockId="3.[151,1436,341,1687]" pageId="3" pageNumber="38">The p2 is large and high. The crown is relatively narrow over the mesial root but is much wider and more robust over the distal root. The posterior cingulum ends near the labial side of the talonid.</paragraph>
<paragraph id="584B36F5FFCCFFE963AF79E3FDD0C013" blockId="3.[151,1436,341,1687]" pageId="3" pageNumber="38">The dp3 is the smallest tooth of this specimen; the trigonid and talonid are similar in length, but the talonid is wider. The metaconid is slightly smaller than the protoconid and is located on the posterolabial margin of the trigonid. The small paraconid is positioned anteromedially. The entoconid is the highest cusp of the talonid; the hypoconid is labially salient.</paragraph>
<paragraph id="584B36F5FFCCFFE963AF7843FF6CC3AB" blockId="3.[151,1436,341,1687]" pageId="3" pageNumber="38">
The erupting m3 (
<figureCitation id="C0CF2A70FFCCFFE962F27843FE66C06B" box="[410,481,1341,1367]" captionStart="FIGURE 2" captionStartId="4.[151,255,1867,1891]" captionTargetBox="[231,1354,480,1839]" captionTargetId="figure@4.[231,1355,480,1841]" captionTargetPageId="4" captionText="FIGURE 2: Thylophorops lorenzinii sp. nov. MLP 08 - III- 10 - 1 (Holotype); detail of the erupting m 3 in lingual (A), and occlusal-lingual views (B). Scale: 2 mm." httpUri="https://zenodo.org/record/185654/files/figure.png" pageId="3" pageNumber="38">Fig. 2</figureCitation>
) is very large, with its trigonid slightly longer than the talonid. All cusps are well preserved. The paraconid is triangular in occlusal view; its postero-lingual side is flat. In contrast, in
<taxonomicName id="9FF44D76FFCCFFE966EE781BFECAC09A" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="3" pageNumber="38" phylum="Chordata" rank="species" species="chapalmalensis">
<emphasis id="6A80EAE7FFCCFFE966EE781BFECAC09A" italics="true" pageId="3" pageNumber="38">T. chapalmalensis</emphasis>
</taxonomicName>
(
<figureCitation id="C0CF2A70FFCCFFE9623678F3FE20C09B" box="[350,423,1421,1447]" captionStart="FIGURE 4" captionStartId="6.[151,255,1192,1216]" captionTargetBox="[234,1358,384,1169]" captionTargetId="figure@6.[234,1359,384,1170]" captionTargetPageId="6" captionText="FIGURE 4: Lower left molars of Thylophorops chapalmalensis (left, detail of specimen MPH 064), T. lorenzinii sp. nov. (centre, detail of the type specimen), and T. perplanus (right, detail of specimen MLP 97 - XI- 15 - 1). Because of the erupting stage of the m 3 of T. lorenzinii in the type specimen, the anterobasal cingulum is not visible. Scale: 2 mm." httpUri="https://zenodo.org/record/185656/files/figure.png" pageId="3" pageNumber="38">Fig. 4</figureCitation>
, left), this side is rounded and the cusp is conical in occlusal view. The protoconid is the highest and largest cusp of the trigonid; it lies on the lingual side and occupies almost the entire trigonid surface. The metaconid is proportionally larger than that of
<taxonomicName id="9FF44D76FFCCFFE9603278A3FBAAC0CA" box="[858,1069,1501,1526]" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="3" pageNumber="38" phylum="Chordata" rank="species" species="chapalmalensis">
<emphasis id="6A80EAE7FFCCFFE9603278A3FBAAC0CA" box="[858,1069,1501,1526]" italics="true" pageId="3" pageNumber="38">T. chapalmalensis</emphasis>
</taxonomicName>
. The talonid is well developed in all dimensions. The entoconid is tall and prominent. The hypoconulid is large and well developed anteroposteriorly. The entoconid and hypoconulid are similar in size, in sharp contrast with
<taxonomicName id="9FF44D76FFCCFFE967F77B53FAF3C37A" box="[1183,1396,1581,1606]" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="3" pageNumber="38" phylum="Chordata" rank="species" species="chapalmalensis">
<emphasis id="6A80EAE7FFCCFFE967F77B53FAF3C37A" box="[1183,1396,1581,1606]" italics="true" pageId="3" pageNumber="38">T. chapalmalensis</emphasis>
</taxonomicName>
, in which the entoconid is much larger. The hypoconid is located on the labial side of the talonid and is noticeably salient.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="C8D6F8E210D90CC847E2A7777D827827" pageId="8" pageNumber="129">Taxon classification Animalia Araneae Liphistiidae</paragraph>
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<paragraph id="201CAF5FE80F2C67D94CDEE7E9929C9D" pageId="8" pageNumber="129">
Genus
<taxonomicName id="AEB64E8BE3741BBD5C3EE9DA29F65F21" ID-CoL="76B3" ID-ENA="1649364" LSID="http://zoobank.org/BA103085-F5FB-4DF1-81C7-CA5CDE26E65E" authority="Xu &amp; Kuntner" class="Arachnida" family="Heptathelidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela" order="Araneae" pageId="8" pageNumber="129" phylum="Arthropoda" rank="genus">Qiongthela Xu &amp; Kuntner</taxonomicName>
<taxonomicNameLabel id="06E0087A4502ACA3E6A97EDC5075FB96" pageId="8" pageNumber="129">gen. n.</taxonomicNameLabel>
Figures 22-27
</paragraph>
</subSubSection>
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<paragraph id="73DA8D2251C8EBF2EB7797746D1E4A37" pageId="8" pageNumber="129">Type species.</paragraph>
<paragraph id="6E2C754B785FD94632CF8015A68FD3B5" pageId="8" pageNumber="129">
<taxonomicName id="7F296016A50B1960E36DFCDC70371F0F" class="Arachnida" family="Heptathelidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela baishensis" order="Araneae" pageId="8" pageNumber="129" phylum="Arthropoda" rank="species" species="baishensis">Qiongthela baishensis</taxonomicName>
sp. n.
</paragraph>
</subSubSection>
<subSubSection id="0538D4FCBDDC983B2479475D3E20E113" pageId="8" pageNumber="129" type="etymology">
<paragraph id="800EE998C4EF0731B65776D7AD72E239" pageId="8" pageNumber="129">Etymology.</paragraph>
<paragraph id="65F230436ED443796567869042B044F8" pageId="8" pageNumber="129">
The genera of heptathelines contain in their name the word
<normalizedToken id="5C0390307BC4320C0A4A217A3B1A3ED6" originalValue="‘thela’">'thela'</normalizedToken>
referring to spinnerets as the Greek word thele means nipple-like protuberance (
<bibRefCitation id="59BA4C45C94F3901BC963786BF34549C" author="Ono, H" journalOrPublisher="Memoirs of the National Science Museum" pageId="20" pageNumber="141" pagination="145 - 151" title="Zoogeographic and taxonomic notes on spiders of the subfamily Heptathelinae (Araneae, Mesothelae, Liphistiidae)." volume="33" year="2000">Ono 2000</bibRefCitation>
). We continue this tradition, but name the genus to start with Qiong-, referring to Hainan Province, China.
</paragraph>
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<paragraph id="867C262E257EE3C577A7506FEEF887CB" pageId="8" pageNumber="129">Diagnosis.</paragraph>
<paragraph id="947E2A2B802F485F57E635DA7B09CE7D" pageId="8" pageNumber="129">
<taxonomicName id="D7E13F7961053FFE9B603754E3CCC0BB" class="Arachnida" family="Heptathelidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela" order="Araneae" pageId="8" pageNumber="129" phylum="Arthropoda" rank="genus">Qiongthela</taxonomicName>
males differ from all other
<taxonomicName id="5E79F01028D4752A225B6637230BE140" lsidName="" pageId="8" pageNumber="129" rank="subFamily" subFamily="Heptathelinae">Heptathelinae</taxonomicName>
genera by the conductor with a narrow, blade-like, slightly hooked apex (Figures 22, 25-26), and by tegulum with two apophyses (Figures 23, 25-26).
<taxonomicName id="BDA6617CD5AFDF3990FA3E7A3A0322C9" class="Arachnida" family="Heptathelidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela" order="Araneae" pageId="8" pageNumber="129" phylum="Arthropoda" rank="genus">Qiongthela</taxonomicName>
females can be distinguished from all other
<taxonomicName id="2C61C5BE6F96990B857280F304A2B619" lsidName="" pageId="8" pageNumber="129" rank="subFamily" subFamily="Heptathelinae">Heptathelinae</taxonomicName>
by two paired receptacular clusters located at the anterior margin of the bursa copulatrix (Figure 27).
</paragraph>
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<paragraph id="7D4F12D8A6AB049BD8F03A5B8CC5565F" pageId="8" pageNumber="129">Description.</paragraph>
<paragraph id="B6285541763743EF752808D920A76B1D" pageId="8" pageNumber="129">Total length (excluding chelicerae) = 13-31 mm (N = 14); male palp with a distally free conductor, narrow, blade-like with slightly hook-like apex, (Figures 22, 25-26); tegulum with two margins, spinose paracymbium (Figures 23, 25-26); female genitalia with two paired receptacular clusters, all situated at the anterior margin of the bursa copulatrix with more or less distinct tubular stems (Figure 27).</paragraph>
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<paragraph id="9E42628A53DBC9AF74500C32C4FFE686" pageId="8" pageNumber="129">Species composition.</paragraph>
<paragraph id="0F8CDDC51E27BEA3BCB6EDB71C9A19E5" pageId="8" pageNumber="129">
<taxonomicName id="EC99476D7234C07E3C3AF8CF1B37E996" class="Arachnida" family="Heptathelidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela australis" order="Araneae" pageId="8" pageNumber="129" phylum="Arthropoda" rank="species" species="australis">Qiongthela australis</taxonomicName>
(Ono, 2002), comb. n.,
<taxonomicName id="B3F111CA8D5C0558FB022BE07FA93B66" class="Arachnida" family="Heptathelidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela nui" order="Araneae" pageId="8" pageNumber="129" phylum="Arthropoda" rank="species" species="nui">Qiongthela nui</taxonomicName>
(Schwendinger &amp; Ono, 2011), comb. n.,
<taxonomicName id="DDA5B7CA477DCB4ED03D11A410CF5FEF" class="Arachnida" family="Heptathelidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela baishensis" order="Araneae" pageId="8" pageNumber="129" phylum="Arthropoda" rank="species" species="baishensis">Qiongthela baishensis</taxonomicName>
sp. n. (3♂2♀), and three undescribed species (6♂8♀, 1♀ and 1♂1♀, respectively) from Hainan, China.
</paragraph>
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<paragraph id="7893D0C47B6CB8326FF786D36E98B4CF" pageId="8" pageNumber="129">Distribution.</paragraph>
<paragraph id="AB8F9DE9BF393EFF1B8BE7FB9E7D2B65" pageId="8" pageNumber="129">Hainan (China) and Vietnam.</paragraph>
</subSubSection>
<subSubSection id="C8B264313B1C468BC4A892BAB587F6C8" pageId="9" pageNumber="130" type="remarks">
<paragraph id="0FC2D5D3E74E3FEA2EA92C76D2C3A0A1" pageId="9" pageNumber="130">
<pageBreakToken id="695C042782F086D4945738DA3F50C573" pageId="9" pageNumber="130" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph id="D3EA9AFA39F88B6F01127CEE4A1CD8D2" pageId="9" pageNumber="130">
Based on morphological descriptions, but not on phylogenetic analyses, we include two species from Vietnam in this genus, originally described as
<taxonomicName id="3649DAB9D80C543C9236A78EDB23B034" class="Arachnida" family="Heptathelidae" genus="Songthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Songthela australis" order="Araneae" pageId="9" pageNumber="130" phylum="Arthropoda" rank="species" species="australis">Songthela australis</taxonomicName>
Ono, 2002 and
<taxonomicName id="6B2F818F771F3605CA2C5133D32C1093" class="Arachnida" family="Heptathelidae" genus="Heptathela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptathela nui" order="Araneae" pageId="9" pageNumber="130" phylum="Arthropoda" rank="species" species="nui">Heptathela nui</taxonomicName>
Schwendinger &amp; Ono, 2011.
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<paragraph id="2552BBA57733B8F313D16E68788CF00A" pageNumber="49">9. Scirpus culmo tereti nudo, spicis subglobosis glomeratis pedunculatis, involucro diphyllo inaequali mucronato.</paragraph>
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Scirpus panicula solum foliosa, spicis globosis pedunculatis lateralibus.
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Juncus acutus maritimus, capitulis rotundis.
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Scirpoides maritimum, capitulis sparsis glomeratis.
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Holoschoenus.
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Europa
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Lendemer, J. North Carolina Acad. Sci. 129(3): 74 (2013). Basionym.
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lebhaft azur- bis violettblau, Durchmesser
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lang
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Zeigerwerte nach
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<td id="F8AEE320F787AC03081C085B76FBC1FA" pageNumber="812">Salzzeichen</td>
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<td id="44F00686DD46736C64E144BC3DD1B768" pageNumber="812">Reaktionszahl R</td>
<td id="6403150D3B27083CE7013008CB5B5B16" pageNumber="812" style="text-align:left">neutral bis basisch (pH 5.5-8.5)</td>
<td id="50B1AE7CF82E39D0BEE7CF20017AB382" pageNumber="812">Temperaturzahl T</td>
<td id="5969FA1410989E8FEACB0A136E4527F0" pageNumber="812" style="text-align:left">
sehr warm-kollin (nur an
<normalizedToken id="3EFCB60CDF016BE1C580C85575C071D6" originalValue="wärmsten">waermsten</normalizedToken>
Stellen, Hauptverbreitung in
<normalizedToken id="27F9F7BC1B135C3F40FD9D576C3361A5" originalValue="Südeuropa">Suedeuropa</normalizedToken>
)
</td>
</tr>
<tr id="41C94B39AD551DFC05E37AE7ED6F4798" pageNumber="812">
<td id="55FD3E45A74F706103813B2D60E0A702" pageNumber="812">
<normalizedToken id="9BFAD2CCA7E839D98E83CD207F69FFEB" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="8ACA244A080DC89E01E462F0710CABCA" pageNumber="812" style="text-align:left">
<normalizedToken id="D8F81928301627B66A947E54CED1633E" originalValue="mässig">maessig</normalizedToken>
<normalizedToken id="769D614E70DE14CC3A1DE7A0C42592F6" originalValue="nährstoffarm">naehrstoffarm</normalizedToken>
bis
<normalizedToken id="2E61A2724F4587CBEBD356A529CF8C65" originalValue="mässig">maessig</normalizedToken>
<normalizedToken id="65BC66868BDB8A5D117C23023319A588" originalValue="nährstoffreich">naehrstoffreich</normalizedToken>
</td>
<td id="4430824CCCC3A293435965D4AFAA3639" pageNumber="812">
<normalizedToken id="D5E6BA3FB7B57DE49923D5CC863E8438" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="EFCF9AE515FFCCCCE35E916A7091B8A0" pageNumber="812" style="text-align:left">subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter)</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="5B7733713B6B286D0CE48DEE8DF1B480" pageNumber="812" type="vernacular_names">
<paragraph id="938E0823F06746854DFBE8F61627C5CA" pageNumber="812">
Volksname Deutscher Name:
<vernacularName id="2036ACA7C369FE43245B5FE092DE5CCC" language="deu" pageNumber="812">Italienische Ochsenzunge</vernacularName>
Nom
<normalizedToken id="EFB6257B22572FD4C58ED3EC81C6F12B" originalValue="français">francais</normalizedToken>
:
<vernacularName id="AB5CE92BC76553B505B0D5E8323E07AD" language="fra" pageNumber="812">Buglosse d'Italie</vernacularName>
Nome italiano:
<vernacularName id="751EFE0223BA8B13F5A5A0439C5DB465" language="ita" pageNumber="812">Buglossa azzurra</vernacularName>
</paragraph>
</subSubSection>
</treatment>
</document>