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<document ID-CLB-Dataset="297060" ID-DOI="10.5281/zenodo.7316519" ID-GBIF-Dataset="4ec5bc66-0ad8-4214-a6a1-9f3a733ea999" ID-ISBN="0-8018-8221-4" checkinTime="1670905277960" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="D02C2F3669FCF4EFA6FA9C0BF27BCB4A" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Vespertilionidae.451-529.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1, Baltimore: The Johns Hopkins University Press" docTitle="Hypsugo macrotis" docType="treatment" docVersion="4" id="1612F121776AEBF203A8EB376B6B87C6" lastPageNumber="491" masterDocId="1612F121776AEBF203A8EB376B6B87C6" masterDocTitle="Order Chiroptera - Family Vespertilionidae" masterLastPageNumber="529" masterPageNumber="451" pageNumber="491" updateTime="1716775078212" updateUser="ExternalLinkService">
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<mods:title id="E3CC774B267FA1DBA0405B12459015CB">Order Chiroptera - Family Vespertilionidae</mods:title>
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<paragraph id="BFDE6A98F034A627663516E559F9CA2B" pageNumber="491">
<taxonomicName id="399A61360A239CBB32DEC04ECCA1797C" ID-CoL="3P4S2" authority="(Temminck 1840)" baseAuthorityName="Temminck" baseAuthorityYear="1840" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageNumber="491" phylum="Chordata" rank="species" species="macrotis">
<emphasis id="143F8247913024170574F11AA2B2825D" italics="true" pageNumber="491">Hypsugo macrotis</emphasis>
(Temminck 1840)
</taxonomicName>
</paragraph>
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<subSubSection id="7502F9F8611037F375DD2E4626517975" pageNumber="491" type="reference_group">
<paragraph id="C76B7CC6212F8EB3056F3786BA9C7C81" pageNumber="491">
<treatmentCitation id="237062A5351211DB78F1DC024835B2D6" author="Temminck" authority="Temminck 1840" authorityName="Temminck" authorityYear="1840" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" page="218" pageNumber="491" phylum="Chordata" rank="species" species="macrotis" title="Vespertilio macrotis" volumeTitle="Monogr. Mamm., Vol. 2" year="1840">
<taxonomicName id="6F48A7FF30B2A1CBD6884D293DF0C0DE" ID-CoL="7G2S9" authority="Temminck 1840" authorityName="Temminck" authorityYear="1840" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageNumber="491" phylum="Chordata" rank="species" species="macrotis">
<emphasis id="7AFD7D6ABE5627669DABF22BC764A078" italics="true" pageNumber="491">[Vespertilio] macrotis</emphasis>
Temminck 1840
</taxonomicName>
,
<bibCitation id="AEC51042FA36AACD29AF0DEF866B3502" author="Temminck" pageNumber="491" pagination="218" title="Vespertilio macrotis" volumeTitle="Monogr. Mamm., Vol. 2" year="1840">Monogr. Mamm., Vol. 2: 218</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="323DB48ECFCD831226393CEB46846F9E" pageNumber="491" type="materials_examined">
<paragraph id="C14C92DFBC7A1E3A93692DD616E641A1" pageNumber="491">
<emphasis id="E4FBD8D2842702BD19FDEF19C9DDFC35" inLineHeading="true" pageNumber="491">Type Locality:</emphasis>
<materialsCitation id="9E319DDD0F7256B10CE37E60E479A591" ID-GBIF-Occurrence="4867287479" country="Indonesia" pageNumber="491" stateProvince="Sumatera Utara">
<collectingCountry id="00537BE55822886DD7DC980DE5081984" name="Indonesia">Indonesia</collectingCountry>
,
<collectingRegion id="1CFC8EE01D56B82BDF9C47C5ADB9351E" country="Indonesia" name="Sumatera Utara">Sumatra</collectingRegion>
, Padang.
</materialsCitation>
</paragraph>
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<subSubSection id="CE51626472B5CFD7C076F59F7F42C67B" pageNumber="491" type="vernacular_names">
<paragraph id="A861154667C98DAE591D9A68DDF6CB01" pageNumber="491">
<emphasis id="757F27551EAE8C2CF7392389828ED26D" inLineHeading="true" pageNumber="491">Vernacular Names:</emphasis>
<vernacularName id="BFB7F755EB6AF2B0329A404A060C73DD" language="eng" pageNumber="491">Big-eared Pipistrelle</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="8690CC7701D880402BE7CBD00FD78E15" pageNumber="491" type="synonym_list">
<paragraph id="78E0A8752235BB4516B41E675AA5B260" pageNumber="491">
<emphasis id="4FB775B7D396583D03279DD4D4361BA5" inLineHeading="true" pageNumber="491">Synonyms:</emphasis>
<taxonomicName id="B4E0CAA603CA0DA0AC3520D297BC46DF" authority="Miller 1911" authorityName="Miller" authorityYear="1911" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageNumber="491" phylum="Chordata" rank="species" species="curtatus">
<emphasis id="9C777AE9ED583C78F5A7D9904E3D9C66" italics="true" pageNumber="491">Hypsugo curtatus</emphasis>
Miller 1911
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A485CB271A55544C7215315D9047A38D" pageNumber="491" type="distribution">
<paragraph id="E78E2606ED6F52886C1F307B6F095981" pageNumber="491">
<emphasis id="B70679C3D657436493ACAA77717707F9" inLineHeading="true" pageNumber="491">Distribution:</emphasis>
W
<collectingCountry id="CDDA4C47D23DF399A39690603D3C0F4E" name="Malaysia">Malaysia</collectingCountry>
, Sumatra, Bali, adjacent small islands.
</paragraph>
</subSubSection>
<subSubSection id="1CA5BFDDF94AF3F22359E822ECFFE794" pageNumber="491" type="conservation">
<paragraph id="CCFF6C2A279423F11ECBDEBCD12B8401" pageNumber="491">
<emphasis id="62C07041400D276BE13C1372F92BA221" inLineHeading="true" pageNumber="491">Conservation:</emphasis>
<collectionCode id="3FE8F7BFDA2998917ECF48C73E7E8191">IUCN</collectionCode>
2003 and
<collectionCode id="8D316B305A5038411BDA2D94B792BAA7">IUCN</collectionCode>
/
<collectionCode id="32D32E0BFE4702E0E7C2AF99982C6088" country="USA" httpUri="http://grbio.org/cool/9wqd-8bvt" name="Sacramento State University" type="">SSC</collectionCode>
Action Plan (2001) Lower Risk (nt) as
<taxonomicName id="C978790A841891CDAC4EEA9828D8449A" baseAuthorityName="Temminck" baseAuthorityYear="1840" class="Mammalia" family="Vespertilionidae" genus="Pipistrellus" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="species" species="macrotis">
<emphasis id="48A274EF88EDEAD47154FD6008BDCC82" italics="true" pageNumber="491">Pipistrellus macrotis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A875B69D9E9929A8C3DDF7BD483E1BCD" pageNumber="491" type="discussion">
<paragraph id="6DC4E4C8A554ECBE12EFBCE9E9DC29FB" pageNumber="491">
<emphasis id="EAE9800785EB12C9B390F269A0376076" inLineHeading="true" pageNumber="491">Discussion:</emphasis>
Listed as a subspecies of
<taxonomicName id="7071D129370C80A73A6DC8DE9BA811C0" baseAuthorityName="Horsfield" baseAuthorityYear="1824" class="Mammalia" family="Vespertilionidae" genus="Pipistrellus" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="species" species="imbricatus">
<emphasis id="F7A375381FC3BAC85FFE852374E7FA70" italics="true" pageNumber="491">imbricatus</emphasis>
</taxonomicName>
by
<bibRefCitation id="34CDDF4FCAE4E72FEEF513F08073BA25" author="Medway, L." refId="ref235372" refString="Medway, L. 1969. The wild mammals of Malaya and offshore islands including Singapore. Oxford University Press, London, 118 pp." year="1969">
<collectingRegion id="EDFA61CD51E1EED13809F96DF14602AA" country="United Kingdom" name="Medway">Medway</collectingRegion>
(1969)
</bibRefCitation>
, but see
<bibRefCitation id="160F9E738E9BD1CDB9B32626771DE6F8" author="Tate, G. H. H." refId="ref347428" refString="Tate, G. H. H. 1942 a. Review of the vespertilionine bats, with special attention to genera and species of the Archbold collections. Bulletin of the American Museum of Natural History, 80: 221 - 297." year="1942">
Tate (1942
<emphasis id="708B1FC484EAE554C103745C5BAF261D" italics="true" pageNumber="491">a</emphasis>
)
</bibRefCitation>
and
<bibRefCitation id="A15F98F8D84E43B626C6CF70D0A132D6" author="Corbet, G. B. &amp; J. E. Hill" refId="ref67843" refString="Corbet, G. B., and J. E. Hill. 1980. A world list of mammalian species. British Museum (Natural History), London, 226 pp." year="1980">Corbet and Hill (1980)</bibRefCitation>
. May include
<taxonomicName id="1DBC6E0761419A589D484DFDC41B2FF2" authorityName="Jentink" authorityYear="1890" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="species" species="vordermanni">
<emphasis id="D6DF702D871287131D1C31421564A89A" italics="true" pageNumber="491">vordermanni</emphasis>
</taxonomicName>
; see
<bibRefCitation id="EFCAEDB1F40CE6932897C509C2497E33" author="Corbet, G. B. &amp; J. E. Hill" refId="ref67955" refString="Corbet, G. B., and J. E. Hill. 1992. Mammals of the Indomalayan region. A systematic review. Oxford University Press, Oxford, 488 pp." year="1992">Corbet and Hill (1992)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="6AF1EABC1F70FF9676330AFAFD5AA9E5" bold="true" box="[263,648,1750,1773]" pageId="8" pageNumber="525">
<emphasis id="6AF1EABC1F70FF9676330AFAFE08A9E5" bold="true" box="[263,474,1750,1773]" italics="true" pageId="8" pageNumber="525">Neoalardus typicus</emphasis>
(
<bibRefCitation id="3C144B5F1F70FF9676DC0AFAFD53A9E5" author="DISTANT W. L." box="[488,641,1750,1773]" pageId="8" pageNumber="525" pagination="219 - 274" refId="ref7875" refString="DISTANT W. L. 1903: Report on the Rhynchota. Part I. Heteroptera. Fascicula Malayenses Zoology 2: 219 - 274." type="journal article" year="1903">Distant, 1903</bibRefCitation>
)
</emphasis>
</taxonomicName>
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(
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)
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<emphasis id="6AF1EABC1F70FF9677A30B32FE9BA839" bold="true" box="[151,329,1822,1841]" pageId="8" pageNumber="525">Material examined</emphasis>
(
<collectionCode id="3E94AE6B1F70FF96766C0B32FE4BA839" box="[344,409,1822,1841]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="525">NMPC</collectionCode>
,
<collectionCode id="3E94AE6B1F70FF9676900B32FE3DA839" box="[420,495,1822,1841]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="525" type="Museum">NHMW</collectionCode>
). 1 J (apterous), labelled “
<geoCoordinate id="3DB150691F70FF9675D30B32FF28A844" degrees="27" direction="north" minutes="08" orientation="latitude" pageId="8" pageNumber="525" precision="15" seconds="22" value="27.139444">N 27° 08 22″</geoCoordinate>
<geoCoordinate id="3DB150691F70FF9676340B15FE53A844" box="[256,385,1849,1868]" degrees="076" direction="east" minutes="20" orientation="longitude" pageId="8" pageNumber="525" precision="15" seconds="38" value="76.343895">E 076° 20 38″</geoCoordinate>
\
<collectingCountry id="2092763E1F70FF9676A70B15FE1CA844" box="[403,462,1849,1868]" name="India" pageId="8" pageNumber="525">INDIA</collectingCountry>
BOR. OCC.
<collectingRegion id="9A41F84C1F70FF9675770B15FD45A844" box="[579,663,1849,1868]" country="India" name="Rajasthan" pageId="8" pageNumber="525">Rajasthan</collectingRegion>
st. \ Alwar distr., Naranimata env., \
<date id="2C3B106E1F70FF9676400B7FFE08A86E" box="[372,474,1875,1894]" pageId="8" pageNumber="525" value="2002-08-16">16. 8. 2002</date>
, lgt. P. Šrámek”; 1 J (apterous), labelled “
<geoCoordinate id="3DB150691F70FF9677DD0B42FEB2A889" box="[233,352,1902,1921]" degrees="27" direction="north" minutes="08" orientation="latitude" pageId="8" pageNumber="525" precision="15" seconds="22" value="27.139444">N 27° 08 22″</geoCoordinate>
<geoCoordinate id="3DB150691F70FF9676510B42FE36A889" box="[357,484,1902,1921]" degrees="076" direction="east" minutes="20" orientation="longitude" pageId="8" pageNumber="525" precision="15" seconds="38" value="76.343895">E 076° 20 38″</geoCoordinate>
\
<collectingCountry id="2092763E1F70FF9676C10B42FDE2A889" box="[501,560,1902,1921]" name="India" pageId="8" pageNumber="525">INDIA</collectingCountry>
BOR. OCC.
<collectingRegion id="9A41F84C1F70FF9675970B42FD25A889" box="[675,759,1902,1921]" country="India" name="Rajasthan" pageId="8" pageNumber="525">Rajasthan</collectingRegion>
st. \ Alwar distr., Naranimata env., \
<date id="2C3B106E1F70FF9676F80BA5FDF6A894" box="[460,548,1929,1948]" pageId="8" pageNumber="525" value="2002-08-01">1. 8. 2002</date>
, lgt. P. Šrámek”; 2 JJ
<specimenCount id="4E83FD271F70FF9675D90BA5FF75A8BE" pageId="8" pageNumber="525" type="female">1 ♀</specimenCount>
(all apterous), labelled “
<geoCoordinate id="3DB150691F70FF96764E0B8FFE23A8BE" box="[378,497,1955,1974]" degrees="27" direction="north" minutes="08" orientation="latitude" pageId="8" pageNumber="525" precision="15" seconds="22" value="27.139444">N 27° 08 22″</geoCoordinate>
<geoCoordinate id="3DB150691F70FF9676C20B8FFDA7A8BE" box="[502,629,1955,1974]" degrees="076" direction="east" minutes="20" orientation="longitude" pageId="8" pageNumber="525" precision="15" seconds="38" value="76.343895">E 076° 20 38″</geoCoordinate>
\
<collectingCountry id="2092763E1F70FF9675B20B8FFD13A8BE" box="[646,705,1955,1974]" name="India" pageId="8" pageNumber="525">INDIA</collectingCountry>
BOR. OCC.
<collectingRegion id="9A41F84C1F70FF9677F90B92FE9DA8D9" box="[205,335,1982,2001]" country="India" name="Rajasthan" pageId="8" pageNumber="525">RAJASTHAN</collectingRegion>
st., Dausa distr. \ Golakabas env., Bhangar \
<date id="2C3B106E1F70FF9675FC0B92FF16A8E4" pageId="8" pageNumber="525" value="2002-08-16">16. 8. 2002</date>
, lgt. P. Šrámek”;
<specimenCount id="4E83FD271F70FF9676600BF5FEA0A8E3" box="[340,370,2002,2028]" pageId="8" pageNumber="525" type="female">1 ♀</specimenCount>
(apterous), labelled “
<geoCoordinate id="3DB150691F70FF96751C0BF5FD49A8E4" box="[552,667,2009,2028]" degrees="27" direction="north" minutes="05" orientation="latitude" pageId="8" pageNumber="525" precision="15" seconds="46" value="27.096111">N 27° 05 46″</geoCoordinate>
<geoCoordinate id="3DB150691F70FF9675AB0BF5FF67A70E" degrees="076" direction="east" minutes="20" orientation="longitude" pageId="8" pageNumber="525" precision="15" seconds="38" value="76.343895">E 076° 20 38″</geoCoordinate>
\
<collectingCountry id="2092763E1F70FF9677F30BDFFED0A70E" box="[199,258,2035,2054]" name="India" pageId="8" pageNumber="525">INDIA</collectingCountry>
BOR. OCC.
<collectingRegion id="9A41F84C1F70FF9676410BDFFE1BA70E" box="[373,457,2035,2054]" country="India" name="Rajasthan" pageId="8" pageNumber="525">Rajasthan</collectingRegion>
st. \ Dausa distr., Golakabas env., \ “Sarsadewi lake”,
<quantity id="9F7D9B4B1F70FF9676010422FEB4A729" box="[309,358,2062,2081]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="525" unit="m" value="400.0">400m</quantity>
a.s.l. \
<date id="2C3B106E1F70FF9676900422FDD4A729" box="[420,518,2062,2081]" pageId="8" pageNumber="525" value="2002-08-14">14. 8. 2002</date>
, lgt. P. Šrámek”.
</paragraph>
</subSubSection>
<subSubSection id="109F65251F70FF9674080F8BFA4FABD6" pageId="8" pageNumber="525" type="distribution">
<paragraph id="583A36AE1F70FF9674080F8BFA4FABD6" blockId="8.[828,1437,935,1246]" pageId="8" pageNumber="525">
<emphasis id="6AF1EABC1F70FF9674080F8BFC18ACB6" bold="true" box="[828,970,935,958]" pageId="8" pageNumber="525">Distribution.</emphasis>
<taxonomicName id="9F854D2D1F70FF9674FB0F8BFB45ACB6" baseAuthorityName="Distant" baseAuthorityYear="1903" box="[975,1175,935,958]" class="Insecta" family="Veliidae" genus="Neoalardus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="typicus">
<emphasis id="6AF1EABC1F70FF9674FB0F8BFB45ACB6" box="[975,1175,935,958]" italics="true" pageId="8" pageNumber="525">Neoalardus typicus</emphasis>
</taxonomicName>
was described from West
<collectingCountry id="2092763E1F70FF9674080FEBFC4CACD6" box="[828,926,967,990]" name="Malaysia" pageId="8" pageNumber="525">Malaysia</collectingCountry>
(Dඌඍൺඇඍ 1903). YൺඇG et al. (1997) recorded it from
<collectingCountry id="2092763E1F70FF96744D0FCBFC3BACF6" box="[889,1001,999,1022]" name="Singapore" pageId="8" pageNumber="525">Singapore</collectingCountry>
. Zൾඍඍൾඅ (1998) gave a redescription of
<taxonomicName id="9F854D2D1F70FF967408082BFC7BAB16" baseAuthorityName="Distant" baseAuthorityYear="1903" box="[828,937,1031,1054]" class="Insecta" family="Veliidae" genus="Neoalardus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="typicus">
<emphasis id="6AF1EABC1F70FF967408082BFC7BAB16" box="[828,937,1031,1054]" italics="true" pageId="8" pageNumber="525">N. typicus</emphasis>
</taxonomicName>
and some new country records (
<collectingCountry id="2092763E1F70FF967225082BFA87AB16" box="[1297,1365,1031,1054]" name="China" pageId="8" pageNumber="525">China</collectingCountry>
,
<collectingCountry id="2092763E1F70FF967256082BFA4AAB16" box="[1378,1432,1031,1054]" name="Laos" pageId="8" pageNumber="525">Laos</collectingCountry>
,
<collectingCountry id="2092763E1F70FF967408080BFC7BAB36" box="[828,937,1063,1086]" name="Indonesia" pageId="8" pageNumber="525">Indonesia</collectingCountry>
). THඋඎආൺඅൺ (2002) listed
<taxonomicName id="9F854D2D1F70FF9673E0080BFA92AB36" baseAuthorityName="Distant" baseAuthorityYear="1903" box="[1236,1344,1063,1086]" class="Insecta" family="Veliidae" genus="Neoalardus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="typicus">
<emphasis id="6AF1EABC1F70FF9673E0080BFA92AB36" box="[1236,1344,1063,1086]" italics="true" pageId="8" pageNumber="525">N. typicus</emphasis>
</taxonomicName>
only for the “Andaman &amp;
<collectingCountry id="2092763E1F70FF9674CC086BFB76AB56" box="[1016,1188,1095,1118]" name="India" pageId="8" pageNumber="525">Nicobar Islands</collectingCountry>
”, but Aඇൽൾඋඌൾඇ (1982: fig. 615: map) already indicated that this species occurs in continental
<collectingCountry id="2092763E1F70FF96748D08ABFC21AB96" box="[953,1011,1159,1182]" name="India" pageId="8" pageNumber="525">India</collectingCountry>
. The species has recently been recorded from
<collectingRegion id="9A41F84C1F70FF96744F088BFC21ABB6" box="[891,1011,1191,1214]" country="India" name="Meghalaya" pageId="8" pageNumber="525">Meghalaya</collectingRegion>
(JൾHൺආൺඅൺඋ &amp;
<collectingCountry id="2092763E1F70FF967399088BFB18ABB6" box="[1197,1226,1191,1214]" name="Switzerland" pageId="8" pageNumber="525">CH</collectingCountry>
ൺඇൽඋൺ 2020). The new record extends the known distribution far westwards.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,274 @@
<document id="BE3A244CB85AAC4CAF30923ABC629C16" ID-DOI="10.37520/aemnp.2021.029" ID-ISSN="1804-6487" ID-Zenodo-Dep="5821215" ID-ZooBank="70C7FD52-9D69-4401-ADE8-0D043D39BB2C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641390478763" checkinUser="plazi" docAuthor="Zettel, Herbert &amp; Laciny, Alice" docDate="2021" docId="D02C87B81F79FF9D74FD0AF0FADCAA3D" docLanguage="en" docName="ActaEntMusNatPra.61.2.517-527.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 61 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Thirumalaia ocularis Zettel &amp; Laciny 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="520" masterDocId="2C15FFC01F78FF9E77340C2CFFD2AF08" masterDocTitle="The taxonomy of some unusual Microveliinae (Hemiptera: Heteroptera: Veliidae) from India" masterLastPageNumber="527" masterPageNumber="517" pageNumber="518" updateTime="1699285328636" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="4BA879CBBF494CD2B7B0431B5BBBFA9E">The taxonomy of some unusual Microveliinae (Hemiptera: Heteroptera: Veliidae) from India</mods:title>
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<mods:namePart id="E57F3AA9809DD4A4E7ACCC75371ADBA4">Laciny, Alice</mods:namePart>
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<mods:date id="7820736C0DDFDC09DC90FFD659D4FDC6">2021</mods:date>
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<treatment id="D02C87B81F79FF9D74FD0AF0FADCAA3D" LSID="urn:lsid:plazi:treatment:D02C87B81F79FF9D74FD0AF0FADCAA3D" httpUri="http://treatment.plazi.org/id/D02C87B81F79FF9D74FD0AF0FADCAA3D" lastPageId="3" lastPageNumber="520" pageId="1" pageNumber="518">
<subSubSection id="109F65251F79FF9F74FD0AF0FAC2A9FB" box="[969,1296,1756,1779]" pageId="1" pageNumber="518" type="nomenclature">
<paragraph id="583A36AE1F79FF9F74FD0AF0FAC2A9FB" blockId="1.[969,1296,1756,1779]" box="[969,1296,1756,1779]" pageId="1" pageNumber="518">
<heading id="037281C21F79FF9F74FD0AF0FAC2A9FB" box="[969,1296,1756,1779]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="518" reason="3">
<emphasis id="6AF1EABC1F79FF9F74FD0AF0FAC2A9FB" bold="true" box="[969,1296,1756,1779]" pageId="1" pageNumber="518">
<taxonomicName id="9F854D2D1F79FF9F74FD0AF0FB61A9FB" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[969,1203,1756,1779]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="species" species="ocularis" status="sp. nov.">
<emphasis id="6AF1EABC1F79FF9F74FD0AF0FB61A9FB" bold="true" box="[969,1203,1756,1779]" italics="true" pageId="1" pageNumber="518">Thirumalaia ocularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F79FF9F738E0AF0FAC2A9FB" box="[1210,1296,1756,1779]" pageId="1" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="109F65251F79FF9F73110AD6FB60A805" box="[1061,1202,1786,1805]" pageId="1" pageNumber="518" type="description">
<paragraph id="583A36AE1F79FF9F73110AD6FB60A805" blockId="1.[828,1437,1786,2080]" box="[1061,1202,1786,1805]" pageId="1" pageNumber="518">
(
<figureCitation id="C0BE2A2B1F79FF9F731F0AD6FBAAA805" box="[1067,1144,1786,1805]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="1" pageNumber="518">Figs 1, 2</figureCitation>
,
<figureCitation id="C0BE2A2B1F79FF9F73B50AD6FB7EA805" box="[1153,1196,1786,1805]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">915</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="109F65251F79FF9F74080B31FA4FA728" pageId="1" pageNumber="518" type="materials_examined">
<paragraph id="583A36AE1F79FF9F74080B31FA4FA728" blockId="1.[828,1437,1786,2080]" pageId="1" pageNumber="518">
<emphasis id="6AF1EABC1F79FF9F74080B31FC10A838" bold="true" box="[828,962,1821,1840]" pageId="1" pageNumber="518">Type material.</emphasis>
Hඈඅඈඍඒඉൾ: J (apterous) (
<collectionCode id="3E94AE6B1F79FF9F739C0B31FB20A838" box="[1192,1266,1821,1840]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="518" type="Museum">NHMW</collectionCode>
), labelled “
<collectingCountry id="2092763E1F79FF9F72620B31FA4AA838" box="[1366,1432,1821,1840]" name="India" pageId="1" pageNumber="518">INDIA</collectingCountry>
:
<collectingRegion id="9A41F84C1F79FF9F74080B1BFC73A842" box="[828,929,1847,1866]" country="India" name="Tamil Nadu" pageId="1" pageNumber="518">Tamil Nadu</collectingRegion>
,
<date id="2C3B106E1F79FF9F749D0B1BFC2CA842" box="[937,1022,1847,1866]" pageId="1" pageNumber="518" value="1999-01-23">23.1.1999</date>
\ Nilgiri Hills,
<quantity id="9F7D9B4B1F79FF9F734F0B1BFB73A842" box="[1147,1185,1847,1866]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="518" unit="km" value="5.0">5km</quantity>
E Kundah \
<geoCoordinate id="3DB150691F79FF9F723C0B1BFA9CA842" box="[1288,1358,1847,1866]" degrees="11" direction="north" minutes="18" orientation="latitude" pageId="1" pageNumber="518" precision="925" value="11.3">11°18N</geoCoordinate>
<geoCoordinate id="3DB150691F79FF9F72660B1BFA4BA842" box="[1362,1433,1847,1866]" degrees="76" direction="east" minutes="40" orientation="longitude" pageId="1" pageNumber="518" precision="925" value="76.666664">76°40E</geoCoordinate>
,
<quantity id="9F7D9B4B1F79FF9F74080B7EFCA5A86D" box="[828,887,1874,1893]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="518" unit="m" value="1900.0">1900m</quantity>
\ leg. D. Boukal (80)”. Pൺඋൺඍඒඉൾඌ (
<collectionCode id="3E94AE6B1F79FF9F73AA0B7EFB34A86D" box="[1182,1254,1874,1893]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="518" type="Museum">NHMW</collectionCode>
,
<collectionCode id="3E94AE6B1F79FF9F73D90B7EFAFCA86D" box="[1261,1326,1874,1893]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="518">NMPC</collectionCode>
): 2 JJ
<specimenCount id="4E83FD271F79FF9F725A0B7EFA49A86D" box="[1390,1435,1868,1893]" pageId="1" pageNumber="518" type="female">5 ♀♀</specimenCount>
(all apterous) (include
<typeStatus id="873E880C1F79FF9F73370B41FB9BA888" box="[1027,1097,1901,1920]" pageId="1" pageNumber="518" type="allotype">allotype</typeStatus>
) with same label data; 1 J (apterous), labelled “
<collectingCountry id="2092763E1F79FF9F74B90BABFC6EA892" box="[909,956,1927,1946]" name="India" pageId="1" pageNumber="518">India</collectingCountry>
:
<collectingRegion id="9A41F84C1F79FF9F74F00BABFBF8A892" box="[964,1066,1927,1946]" country="India" name="Tamil Nadu" pageId="1" pageNumber="518">Tamil Nadu</collectingRegion>
, Nilgiri Hills \ Kotagiri env., Kadashola \
<geoCoordinate id="3DB150691F79FF9F72B90BABFCA1A8BD" degrees="11" direction="north" minutes="25" orientation="latitude" pageId="1" pageNumber="518" precision="925" value="11.416667">N 11°25</geoCoordinate>
<geoCoordinate id="3DB150691F79FF9F74430B8EFC13A8BD" box="[887,961,1954,1973]" degrees="76" direction="east" minutes="55" orientation="longitude" pageId="1" pageNumber="518" precision="925" value="76.916664">E 76°55</geoCoordinate>
,
<quantity id="9F7D9B4B1F79FF9F74FF0B8EFBD9A8BD" box="[971,1035,1954,1973]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="518" unit="m" value="1500.0">1500 m</quantity>
\ a.s.l.,
<date id="2C3B106E1F79FF9F737F0B8EFB75A8BD" box="[1099,1191,1954,1973]" pageId="1" pageNumber="518" value="1999-01-07" valueMax="1999-01-08" valueMin="1999-01-07">78.I.1999</date>
\ leg. D. Boukal (74)”; 3 JJ
<specimenCount id="4E83FD271F79FF9F74080B91FC88A8C7" box="[828,858,1974,2000]" pageId="1" pageNumber="518" type="female">1 ♀</specimenCount>
(all apterous), labelled “
<collectingCountry id="2092763E1F79FF9F731D0B91FBB9A8D8" box="[1065,1131,1981,2000]" name="India" pageId="1" pageNumber="518">INDIA</collectingCountry>
:
<collectingRegion id="9A41F84C1F79FF9F73460B91FB0AA8D8" box="[1138,1240,1981,2000]" country="India" name="Tamil Nadu" pageId="1" pageNumber="518">Tamil Nadu</collectingRegion>
,
<date id="2C3B106E1F79FF9F73EB0B91FAE7A8D8" box="[1247,1333,1981,2000]" pageId="1" pageNumber="518" value="1999-12-01">12.1.1999</date>
\ Palni Hills
<geoCoordinate id="3DB150691F79FF9F74080BFBFC51A8E2" box="[828,899,2007,2026]" degrees="10" direction="north" minutes="16" orientation="latitude" pageId="1" pageNumber="518" precision="925" value="10.266666">10°16N</geoCoordinate>
<geoCoordinate id="3DB150691F79FF9F74BD0BFBFC1CA8E2" box="[905,974,2007,2026]" degrees="77" direction="east" minutes="33" orientation="longitude" pageId="1" pageNumber="518" precision="925" value="77.55">77°33E</geoCoordinate>
\ Perumalmalai,
<quantity id="9F7D9B4B1F79FF9F73560BFBFB4FA8E2" box="[1122,1181,2007,2026]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="518" unit="m" value="1600.0">1600m</quantity>
\ leg. D. Boukal (60)”;
<specimenCount id="4E83FD271F79FF9F72580BFBFA4FA8E2" box="[1388,1437,2001,2026]" pageId="1" pageNumber="518" type="female">2 ♀♀</specimenCount>
(apterous), labelled “
<collectingCountry id="2092763E1F79FF9F74DC0BDEFBFBA70D" box="[1000,1065,2034,2053]" name="India" pageId="1" pageNumber="518">INDIA</collectingCountry>
:
<collectingRegion id="9A41F84C1F79FF9F73040BDEFB47A70D" box="[1072,1173,2034,2053]" country="India" name="Tamil Nadu" pageId="1" pageNumber="518">Tamil Nadu</collectingRegion>
,
<date id="2C3B106E1F79FF9F73AF0BDEFB3CA70D" box="[1179,1262,2034,2053]" pageId="1" pageNumber="518" value="1999-11-01">11.1.1999</date>
\ Palni Hills
<geoCoordinate id="3DB150691F79FF9F72620BDEFA4EA70D" box="[1366,1436,2034,2053]" degrees="10" direction="north" minutes="14" orientation="latitude" pageId="1" pageNumber="518" precision="925" value="10.233334">10°14N</geoCoordinate>
<geoCoordinate id="3DB150691F79FF9F74080421FC53A728" box="[828,897,2061,2080]" degrees="77" direction="east" minutes="29" orientation="longitude" pageId="1" pageNumber="518" precision="925" value="77.48333">77°29E</geoCoordinate>
\ Kodaikanal, Bear Shola Falls \
<quantity id="9F7D9B4B1F79FF9F73A80421FB05A728" box="[1180,1239,2061,2080]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="518" unit="m" value="2100.0">2100m</quantity>
\ leg. D. Boukal (57)”.
</paragraph>
</subSubSection>
<caption id="0CFA66261F7AFF9C77A30A99FE24A9EB" ID-DOI="http://doi.org/10.5281/zenodo.5821217" ID-Zenodo-Dep="5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="2" pageNumber="519" startId="2.[151,188,1717,1736]" targetBox="[151,1436,189,1698]" targetPageId="2">
<paragraph id="583A36AE1F7AFF9C77A30A99FE24A9EB" blockId="2.[151,1438,1717,1763]" pageId="2" pageNumber="519">
Figs 14. Habitus of
<taxonomicName id="9F854D2D1F7AFF9C767E0A99FDD2A9C0" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[330,512,1717,1736]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="519" phylum="Arthropoda" rank="species" species="ocularis" status="gen. nov.">
<emphasis id="6AF1EABC1F7AFF9C767E0A99FDD2A9C0" box="[330,512,1717,1736]" italics="true" pageId="2" pageNumber="519">Thirumalaia ocularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7AFF9C75310A99FD81A9C0" box="[517,595,1717,1736]" pageId="2" pageNumber="519" rank="genus">gen. nov.</taxonomicNameLabel>
&amp; sp. nov. (1 apterous male, 2 apterous female), and of
<taxonomicName id="9F854D2D1F7AFF9C737A0A99FB3FA9C0" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[1102,1261,1717,1736]" class="Insecta" family="Veliidae" genus="Eyarinella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="519" phylum="Arthropoda" rank="species" species="robusta" status="gen. nov.">
<emphasis id="6AF1EABC1F7AFF9C737A0A99FB3FA9C0" box="[1102,1261,1717,1736]" italics="true" pageId="2" pageNumber="519">Eyarinella robusta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7AFF9C73C70A99FA93A9C0" box="[1267,1345,1717,1736]" pageId="2" pageNumber="519" rank="genus">gen. nov.</taxonomicNameLabel>
&amp; sp. nov. (3 apterous male, 4 apterous female).
</paragraph>
</caption>
<subSubSection id="109F65251F7AFF9D77A30B24FC35ABFD" lastPageId="3" lastPageNumber="520" pageId="2" pageNumber="519" type="description">
<paragraph id="583A36AE1F7AFF9C77A30B24FC22A8B7" blockId="2.[151,761,1800,2079]" lastBlockId="2.[828,1437,1800,2079]" pageId="2" pageNumber="519">
<emphasis id="6AF1EABC1F7AFF9C77A30B24FE0BA817" bold="true" box="[151,473,1800,1823]" pageId="2" pageNumber="519">Description of apterous male.</emphasis>
Measurements of
<typeStatus id="873E880C1F7AFF9C75A10B24FD27A817" box="[661,757,1800,1823]" pageId="2" pageNumber="519" type="holotype">holotype</typeStatus>
: Body length 2.62; maximum body width (at metapleura) 0.96. Head length 0.46, width 0.64. Synthlipsis 0.28. Pronotum length 0.53, width 0.86. Lengths of antennomeres, I 0.49, II 0.30, III 0.54, IV 0.54. Lengths of leg segments: profemur 0.81, protibia 0.72, protarsus 0.31, mesofemur 0.99, mesotibia 0.93, mesotarsus 0.20 + 0.31, metafemur 1.09, metatibia 1.36, metatarsus 0.26 + 0.31; protibial comb length 0.25; comb index 0.35. Measurements of
<typeStatus id="873E880C1F7AFF9C75A50424FD25A717" box="[657,759,2056,2079]" pageId="2" pageNumber="519" type="paratype">paratypes</typeStatus>
(n = 6): Body length 2.552.64; maximum body width (at metapleura or abdominal segment 4) 0.920.98. Head length 0.440.46, width 0.640.66. Synthlipsis 0.290.31. Pronotum length 0.500.54, width 0.880.92. Protibia length 0.700.76; protibial comb length 0.210.27; comb index 0.300.36.
</paragraph>
<paragraph id="583A36AE1F7AFF9D746E0BE4FE08AE1D" blockId="2.[828,1437,1800,2079]" lastBlockId="3.[151,761,190,2069]" lastPageId="3" lastPageNumber="520" pageId="2" pageNumber="519">
Colour (
<figureCitation id="C0BE2A2B1F7AFF9C74860BE4FC22A8D7" box="[946,1008,1992,2015]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="2" pageNumber="519">Fig. 1</figureCitation>
): Black. Sides of head yellowish brown. Pronotum anteromedially with a narrow transverse yellow stripe, narrowly interrupted at midline. Acetabula orange-brown to black. Rostrum yellow, terminal segment black.Antenna dark brown. Legs yellow, apices of femora, protibiae, and tarsi infuscated.
</paragraph>
<paragraph id="583A36AE1F7BFF9D77810D32FE0AADBD" blockId="3.[151,761,190,2069]" pageId="3" pageNumber="520">
Pilosity: Body covered by a short appressed greyish brown pilosity; pilosity on legs yellowish to whitish. Antenna with two long setae on antennomere I, and one on antennomere II; additionally, with numerous shorter, oblique setae on antennomeres IIIV. Most of body with brown, relatively short to moderately long oblique standing setae (longest on mediotergites). Legs with long whitish standing setae on flexor sides, and with more oblique, shorter, mostly brownish setae on extensor side. Silverish pilosity (
<figureCitation id="C0BE2A2B1F7BFF9D77C00E12FEE2AD5D" box="[244,304,574,597]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="3" pageNumber="520">Fig. 1</figureCitation>
) extended on anterior part of pronotum and sides of mediotergite 1; small spots often developed at hind margins of laterotergites and (minute and indistinct) on sides of mediotergites 57.
</paragraph>
<paragraph id="583A36AE1F7BFF9D77810E92FD3FA99D" blockId="3.[151,761,190,2069]" pageId="3" pageNumber="520">
Structures: Body moderately stout (
<figureCitation id="C0BE2A2B1F7BFF9D75770E92FD57ADDD" box="[579,645,702,725]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="3" pageNumber="520">Fig. 1</figureCitation>
); sides of abdomen subparallel. Head wider than long, anteriorly blunt, posteriorly weakly protruded, in a shallow concavity of the anterior margin of pronotum. Anteclypeus and labrum shiny. Eyes prominent; eye width larger than half of synthlipsis (
<figureCitation id="C0BE2A2B1F7BFF9D76090F72FE51AC7D" box="[317,387,862,885]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="3" pageNumber="520">Fig. 9</figureCitation>
). Antenna very long and slender; antennomere II slightly slenderer and shorter than I; antennomeres III and IV similar in length and width, or IV slightly longer than III. Rostrum stout, reaching base of mesosternum. Pronotum long (
<figureCitation id="C0BE2A2B1F7BFF9D76DE0FF2FDFDACFD" box="[490,559,990,1013]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="3" pageNumber="520">Fig. 9</figureCitation>
), reaching mediotergite 1 posteriorly; sides of pronotum slightly protruded anteriad; disc with a shallow transverse groove bearing a row of sensory pores and separating anterior part from posterior lobe; anterior margin with another row of pores; numerous pores scattered over pronotal lobe. Legs long, hardly modified. Profemur (
<figureCitation id="C0BE2A2B1F7BFF9D76F708B2FDC3ABBD" box="[451,529,1182,1205]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="3" pageNumber="520">Fig. 10</figureCitation>
) slightly thicker than meso- and metafemur, not curved. Tibiae and tarsi slender. Protibial comb about one third as long as protibia (
<figureCitation id="C0BE2A2B1F7BFF9D75AB08F2FD39ABFD" box="[671,747,1246,1269]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="3" pageNumber="520">Fig. 10</figureCitation>
). Second tarsomeres on middle and hind leg longer than first. Abdomen moderately wide. All mediotergites transverse (
<figureCitation id="C0BE2A2B1F7BFF9D77AB0912FF08AA5D" box="[159,218,1342,1365]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="3" pageNumber="520">Fig. 1</figureCitation>
); mediotergite 7 about 1.6 times as wide as long. Abdominal sterna hardly modified. Sterna 57 very shallowly impressed along midline. Hind margin of sternum 7 broadly concave. Segment 8 (
<figureCitation id="C0BE2A2B1F7BFF9D76B309B2FE05AABD" box="[391,471,1438,1461]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="3" pageNumber="520">Fig. 12</figureCitation>
) subcylindrical, 1.5 times as long as wide; ventrally, posterior and anterior margin strongly concave, dorsally posterior margin with concavity in medial third. Pygophore (
<figureCitation id="C0BE2A2B1F7BFF9D768909D2FDDAA91D" box="[445,520,1534,1557]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="3" pageNumber="520">Fig. 13</figureCitation>
) pear-shaped, but with truncated posterior margin, widest at paramere insertion, about 1.6 times as long as wide. Proctiger (
<figureCitation id="C0BE2A2B1F7BFF9D75560A12FD7DA95D" box="[610,687,1598,1621]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="3" pageNumber="520">Fig. 15</figureCitation>
) small and slender, posterior part almost round. Parameres (
<figureCitation id="C0BE2A2B1F7BFF9D75F80A72FF60A99D" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="3" pageNumber="520">Fig. 14</figureCitation>
) symmetrical, medium-sized, of characteristic shape.
</paragraph>
<paragraph id="583A36AE1F7BFF9D77A30AB2FC7CAFDD" blockId="3.[151,761,190,2069]" lastBlockId="3.[828,1437,190,1333]" pageId="3" pageNumber="520">
<emphasis id="6AF1EABC1F7BFF9D77A30AB2FE2CA9BD" bold="true" box="[151,510,1694,1717]" pageId="3" pageNumber="520">Description of apterous female.</emphasis>
Measurements of
<typeStatus id="873E880C1F7BFF9D75F30AB2FF1BA9DD" pageId="3" pageNumber="520" type="allotype">allotype</typeStatus>
: Body length 2.88; maximum body width (at abdominal segment 4) 1.38. Head length 0.51, width 0.68. Synthlipsis 0.34. Pronotum length 0.50, width 0.98. Lengths of antennomeres, I 0.44, II 0.28, III 0.47, IV 0.51. Lengths of leg segments: profemur 0.81, protibia 0.70, protarsus 0.33, mesofemur 0.91, mesotibia 0.95, mesotarsus 0.19 + 0.32, metafemur 1.10, metatibia 1.38, metatarsus 0.27 + 0.34. Measurements of
<typeStatus id="873E880C1F7BFF9D76980BB2FDC0A8BD" box="[428,530,1950,1973]" pageId="3" pageNumber="520" type="paratype">paratypes</typeStatus>
(n = 7): Body length 2.803.00; maximum body width (at abdominal segment 3 or 4) 1.181.38. Head length 0.470.52, width 0.670.70. Synthlipsis 0.310.34. Pronotum length 0.500.54, width 0.930.99.
</paragraph>
<paragraph id="583A36AE1F7BFF9D746E0CF2FB03AFFD" blockId="3.[828,1437,190,1333]" box="[858,1233,222,245]" pageId="3" pageNumber="520">
Colour (
<figureCitation id="C0BE2A2B1F7BFF9D74870CF2FC26AFFD" box="[947,1012,222,245]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="3" pageNumber="520">Fig. 2</figureCitation>
): Similar as in male.
</paragraph>
<paragraph id="583A36AE1F7BFF9D746E0CD2FA9CAE7D" blockId="3.[828,1437,190,1333]" pageId="3" pageNumber="520">Pilosity: Similar as in male. Silvery pilosity on mediotergites and laterotergites on average more extended. Setae on mediotergite 8 similar as on mediotergite 7. Gonocoxa with moderately long, oblique pilosity.</paragraph>
<paragraph id="583A36AE1F7BFF9D746E0D52FC1FADDD" blockId="3.[828,1437,190,1333]" pageId="3" pageNumber="520">
Structures: Body stout (
<figureCitation id="C0BE2A2B1F7BFF9D73580D52FB61AE9D" box="[1132,1203,382,405]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="3" pageNumber="520">Fig. 2</figureCitation>
); shape of abdomen ovate. Most characters similar as in male. Antennomere IV slightly longer than 3. Legs unmodified; all femora of similar width; protibia without grasping comb. Mediotergites strongly transverse (
<figureCitation id="C0BE2A2B1F7BFF9D735C0DD2FB78AD1D" box="[1128,1194,510,533]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="3" pageNumber="520">Fig. 2</figureCitation>
). Mediotergite 7 about 1.25 times as long as 6, at anterior margin 2.4 times as wide as long, posteriorly strongly narrowed; posterior margin straight. Mediotergite 8 (
<figureCitation id="C0BE2A2B1F7BFF9D737C0E72FB46AD7D" box="[1096,1172,606,629]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="3" pageNumber="520">Fig. 11</figureCitation>
) almost semi-circular in shape, directed straight posteriorly. Proctiger knob-shaped, directed posteriorly. Gonocoxa plate-like, not concealed by sternum 7.
</paragraph>
<paragraph id="583A36AE1F7BFF9D74080EF2FC35ABFD" blockId="3.[828,1437,190,1333]" pageId="3" pageNumber="520">
<emphasis id="6AF1EABC1F7BFF9D74080EF2FBC8ADFD" bold="true" box="[828,1050,734,757]" pageId="3" pageNumber="520">Comparative notes.</emphasis>
The generic diagnosis distinguishes
<taxonomicName id="9F854D2D1F7BFF9D74080ED2FBCEAC1D" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[828,1052,766,789]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="species" species="ocularis" status="sp. nov.">
<emphasis id="6AF1EABC1F7BFF9D74080ED2FBCEAC1D" box="[828,1052,766,789]" italics="true" pageId="3" pageNumber="520">Thirumalaia ocularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7BFF9D73150ED2FBA0AC1D" box="[1057,1138,766,789]" pageId="3" pageNumber="520" rank="species">sp. nov.</taxonomicNameLabel>
from all other Indian Microveliinae except
<taxonomicName id="9F854D2D1F7BFF9D74D40F32FA4BAC3D" authority="Hafiz &amp; Ribeiro, 1939" authorityName="Hafiz &amp; Ribeiro" authorityYear="1939" box="[992,1433,798,821]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="species" species="santala">
<emphasis id="6AF1EABC1F7BFF9D74D40F32FB7AAC3D" box="[992,1192,798,821]" italics="true" pageId="3" pageNumber="520">Microvelia santala</emphasis>
<bibRefCitation id="3C144B5F1F7BFF9D73980F32FA4BAC3D" author="HAFIZ H. A. &amp; RIBEIRO K. S." box="[1196,1433,798,821]" pageId="3" pageNumber="520" pagination="423 - 436" refId="ref8192" refString="HAFIZ H. A. &amp; RIBEIRO K. S. 1939: On a further collection of aquatic Rhynchota from the Rajmahal Hills, Santal Parganas, Bihar, with a description of Microvelia santala, sp. nov. Records of the Indian Museums 41 (4): 423 - 436." type="journal article" year="1939">Hafiz &amp; Ribeiro, 1939</bibRefCitation>
</taxonomicName>
. This species was described from apterous males collected in
<collectingRegion id="9A41F84C1F7BFF9D74630F72FC17AC7D" box="[855,965,862,885]" country="India" name="Jharkhand" pageId="3" pageNumber="520">Jharkhand</collectingRegion>
(formerly
<collectingRegion id="9A41F84C1F7BFF9D730C0F72FBA5AC7D" box="[1080,1143,862,885]" country="India" name="Bihar" pageId="3" pageNumber="520">Bihar</collectingRegion>
; Hൺൿ&amp; Rൻൾඋඈ 1939). It has several similarities with
<taxonomicName id="9F854D2D1F7BFF9D73510F52FA94AC9D" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[1125,1350,894,917]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="species" species="ocularis" status="sp. nov.">
<emphasis id="6AF1EABC1F7BFF9D73510F52FA94AC9D" box="[1125,1350,894,917]" italics="true" pageId="3" pageNumber="520">Thirumalaia ocularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7BFF9D72780F52FA4FAC9D" box="[1356,1437,894,917]" pageId="3" pageNumber="520" rank="species">sp. nov.</taxonomicNameLabel>
as it is also a slender species with large eyes, very long appendages, and a long protibial grasping comb (cf. Hൺൿ&amp; Rൻൾඋඈ 1939), a character combination that does not fit
<taxonomicName id="9F854D2D1F7BFF9D74080FD2FC62AB1D" authorityName="Westwood" authorityYear="1834" box="[828,944,1022,1045]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F7BFF9D74080FD2FC62AB1D" box="[828,944,1022,1045]" italics="true" pageId="3" pageNumber="520">Microvelia</emphasis>
</taxonomicName>
. Therefore, it may represent a second species of
<taxonomicName id="9F854D2D1F7BFF9D74680832FC30AB3D" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[860,994,1054,1077]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F7BFF9D74680832FC30AB3D" box="[860,994,1054,1077]" italics="true" pageId="3" pageNumber="520">Thirumalaia</emphasis>
</taxonomicName>
. However, genitalia were not treated in the original description, and therefore we do not transfer
<taxonomicName id="9F854D2D1F7BFF9D74080872FC62AB7D" authorityName="Hafiz &amp; Ribeiro" authorityYear="1939" box="[828,944,1118,1141]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="species" species="santala">
<emphasis id="6AF1EABC1F7BFF9D74080872FC62AB7D" box="[828,944,1118,1141]" italics="true" pageId="3" pageNumber="520">M. santala</emphasis>
</taxonomicName>
to the new genus.
<taxonomicName id="9F854D2D1F7BFF9D73B40872FA9EAB7D" authorityName="Hafiz &amp; Ribeiro" authorityYear="1939" box="[1152,1356,1118,1141]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="species" species="santala">
<emphasis id="6AF1EABC1F7BFF9D73B40872FA9EAB7D" box="[1152,1356,1118,1141]" italics="true" pageId="3" pageNumber="520">Microvelia santala</emphasis>
</taxonomicName>
clearly differs from
<taxonomicName id="9F854D2D1F7BFF9D74F60852FB74AB9D" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[962,1190,1150,1173]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="species" species="ocularis" status="sp. nov.">
<emphasis id="6AF1EABC1F7BFF9D74F60852FB74AB9D" box="[962,1190,1150,1173]" italics="true" pageId="3" pageNumber="520">Thirumalaia ocularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7BFF9D739A0852FAD3AB9D" box="[1198,1281,1150,1173]" pageId="3" pageNumber="520" rank="species">sp. nov.</taxonomicNameLabel>
by the greater slenderness of its body and a mediotergite 7 of the male that is longer than broad (cf. Hൺൿ&amp; Rൻൾඋඈ 1939: p. 426, text-fig. a).
</paragraph>
</subSubSection>
<subSubSection id="109F65251F7BFF9D740808D2FADCAA3D" pageId="3" pageNumber="520" type="etymology">
<paragraph id="583A36AE1F7BFF9D740808D2FADCAA3D" blockId="3.[828,1437,190,1333]" pageId="3" pageNumber="520">
<emphasis id="6AF1EABC1F7BFF9D740808D2FC6EAA1D" bold="true" box="[828,956,1278,1301]" pageId="3" pageNumber="520">Etymology.</emphasis>
The species epithet, Latin adjective
<taxonomicName id="9F854D2D1F7BFF9D727108D2FA4EAA1D" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[1349,1436,1278,1301]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="species" species="ocularis">
<emphasis id="6AF1EABC1F7BFF9D727108D2FA4EAA1D" box="[1349,1436,1278,1301]" italics="true" pageId="3" pageNumber="520">ocularis</emphasis>
</taxonomicName>
(-
<emphasis id="6AF1EABC1F7BFF9D74790932FC8DAA3D" box="[845,863,1310,1333]" italics="true" pageId="3" pageNumber="520">is</emphasis>
, -
<emphasis id="6AF1EABC1F7BFF9D74410932FC53AA3D" box="[885,897,1310,1333]" italics="true" pageId="3" pageNumber="520">e</emphasis>
), refers to the large, protruding eyes.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="583A36AE1F79FF9F74C50CC1FB35AE0C" blockId="1.[1009,1255,237,260]" box="[1009,1255,237,260]" pageId="1" pageNumber="518">
<heading id="037281C21F79FF9F74C50CC1FB35AE0C" box="[1009,1255,237,260]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="518" reason="3">
<emphasis id="6AF1EABC1F79FF9F74C50CC1FB35AE0C" bold="true" box="[1009,1255,237,260]" pageId="1" pageNumber="518">
<taxonomicName id="9F854D2D1F79FF9F74C50CC1FBAEAE0C" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[1009,1148,237,260]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F79FF9F74C50CC1FBAEAE0C" bold="true" box="[1009,1148,237,260]" italics="true" pageId="1" pageNumber="518">Thirumalaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F79FF9F73B70CC1FB35AE0C" box="[1155,1255,237,260]" pageId="1" pageNumber="518" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="109F65251F79FF9F73110D27FB60AE16" box="[1061,1202,267,286]" pageId="1" pageNumber="518" type="description">
<paragraph id="583A36AE1F79FF9F73110D27FB60AE16" blockId="1.[1061,1202,267,286]" box="[1061,1202,267,286]" pageId="1" pageNumber="518">
(
<figureCitation id="C0BE2A2B1F79FF9F731F0D27FBAAAE16" box="[1067,1144,267,286]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="1" pageNumber="518">Figs 1, 2</figureCitation>
,
<figureCitation id="C0BE2A2B1F79FF9F73B50D27FB7EAE16" box="[1153,1196,267,286]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">915</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="109F65251F79FF9F74080D1AFAC1AE45" box="[828,1299,310,333]" pageId="1" pageNumber="518" type="type_taxon">
<paragraph id="583A36AE1F79FF9F74080D1AFAC1AE45" blockId="1.[828,1299,310,333]" box="[828,1299,310,333]" pageId="1" pageNumber="518">
<emphasis id="6AF1EABC1F79FF9F74080D1AFC02AE45" bold="true" box="[828,976,310,333]" pageId="1" pageNumber="518">
<typeStatus id="873E880C1F79FF9F74080D1AFCA6AE45" box="[828,884,310,333]" pageId="1" pageNumber="518">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="9F854D2D1F79FF9F74E20D1AFB6BAE45" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[982,1209,310,333]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="species" species="ocularis" status="sp. nov.">
<emphasis id="6AF1EABC1F79FF9F74E20D1AFB6BAE45" box="[982,1209,310,333]" italics="true" pageId="1" pageNumber="518">Thirumalaia ocularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F79FF9F73F40D1AFAC1AE45" box="[1216,1299,310,333]" pageId="1" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="109F65251F79FF9F74080D4AFB52A9B5" pageId="1" pageNumber="518" type="diagnosis">
<paragraph id="583A36AE1F79FF9F74080D4AFBFBACF5" blockId="1.[828,1437,358,1725]" pageId="1" pageNumber="518">
<emphasis id="6AF1EABC1F79FF9F74080D4AFC63AE75" bold="true" box="[828,945,358,381]" pageId="1" pageNumber="518">Diagnosis.</emphasis>
Body of male moderately stout (
<figureCitation id="C0BE2A2B1F79FF9F721D0D4AFAB8AE75" box="[1321,1386,358,381]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="1" pageNumber="518">Fig. 1</figureCitation>
), of female much stouter (
<figureCitation id="C0BE2A2B1F79FF9F731F0DAAFBBDAE95" box="[1067,1135,390,413]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="1" pageNumber="518">Fig. 2</figureCitation>
). Postocular region of head deeply inserted in anterior part of pronotum (
<figureCitation id="C0BE2A2B1F79FF9F72730D8AFA5DAEB5" box="[1351,1423,422,445]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">Fig. 9</figureCitation>
). Eyes very large, synthlipsis 0.280.34 (
<figureCitation id="C0BE2A2B1F79FF9F73DE0DEAFAFCAED5" box="[1258,1326,454,477]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">Fig. 9</figureCitation>
). Antenna very long and slender, about two thirds of body length (
<figureCitation id="C0BE2A2B1F79FF9F74770E2AFC76AD15" box="[835,932,518,541]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="1" pageNumber="518">Figs 1, 2</figureCitation>
); antennomere I approximately as long as head length (in male longer than in female); relative lengths of antennomeres: IV ≧ III&gt; I&gt; II. Pronotum convexly rounded posteriorly (
<figureCitation id="C0BE2A2B1F79FF9F732A0E4AFBB3AD75" box="[1054,1121,614,637]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">Fig. 9</figureCitation>
). Lateral evaporatorium situated approximately in middle of metapleuron. Legs long and slender. Claws preterminal. Male: Propleuron unmodified. Femora unmodified. Protibia unmodified; protibial grasping comb long, about one third of tibia length (
<figureCitation id="C0BE2A2B1F79FF9F74770F2AFC42AC15" box="[835,912,774,797]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">Fig. 10</figureCitation>
). Mesotibia without grasping comb. Claws simple, minimally surpassing apex of tarsus. Abdomen without projections. Segment 8 elongated. Pygophore pear-shaped, narrow at base, with truncated posterior margin (
<figureCitation id="C0BE2A2B1F79FF9F72750F4AFA5DAC75" box="[1345,1423,870,893]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">Fig. 13</figureCitation>
). Proctiger very slender (
<figureCitation id="C0BE2A2B1F79FF9F73030FAAFB55AC95" box="[1079,1159,902,925]" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">Fig. 15</figureCitation>
). Parameres symmetrical, medium-sized, of characteristic shape as shown in
<figureCitation id="C0BE2A2B1F79FF9F72630F8AFC88ACD5" captionStart="Figs 915" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 915. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 head, first antennomere, and pronotum, dorsal; 10 foreleg of male (arrow points at proximal end of grasping comb); 11 abdominal segments 7 and 8, and proctiger of female, dorsal; 12 abdominal segment 8 of male, ventral; 13 pygophore of male, ventral; 14 left paramere, lateral; 15 proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">Figure 14</figureCitation>
. Female: Genitalia situated in a strictly posterior opening of segment 7.
</paragraph>
<paragraph id="583A36AE1F79FF9F7408082AFB52A9B5" blockId="1.[828,1437,358,1725]" pageId="1" pageNumber="518">
<emphasis id="6AF1EABC1F79FF9F7408082AFBCDAB15" bold="true" box="[828,1055,1030,1053]" pageId="1" pageNumber="518">Comparative notes.</emphasis>
<taxonomicName id="9F854D2D1F79FF9F731D082AFB60AB15" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[1065,1202,1030,1053]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F79FF9F731D082AFB60AB15" box="[1065,1202,1030,1053]" italics="true" pageId="1" pageNumber="518">Thirumalaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F79FF9F7389082AFAF7AB15" box="[1213,1317,1030,1053]" pageId="1" pageNumber="518" rank="genus">gen. nov.</taxonomicNameLabel>
resembles some species of
<taxonomicName id="9F854D2D1F79FF9F74CC080AFB21AB35" authority="Distant, 1903" authorityName="Distant" authorityYear="1903" box="[1016,1267,1062,1085]" class="Insecta" family="Veliidae" genus="Baptista" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F79FF9F74CC080AFB81AB35" box="[1016,1107,1062,1085]" italics="true" pageId="1" pageNumber="518">Baptista</emphasis>
<bibRefCitation id="3C144B5F1F79FF9F7369080AFB21AB35" author="DISTANT W. L." box="[1117,1267,1062,1085]" pageId="1" pageNumber="518" pagination="219 - 274" refId="ref7875" refString="DISTANT W. L. 1903: Report on the Rhynchota. Part I. Heteroptera. Fascicula Malayenses Zoology 2: 219 - 274." type="journal article" year="1903">
Distant,
<quantity id="9F7D9B4B1F79FF9F7389080AFB21AB35" box="[1213,1267,1062,1085]" metricMagnitude="1" metricUnit="m" metricValue="4.83362" pageId="1" pageNumber="518" unit="in" value="1903.0">1903</quantity>
</bibRefCitation>
</taxonomicName>
in habitus and shares with this genus some other characters like the long protibial grasping comb, structure of antenna, and a concave anterior margin of the pronotum. Most species of
<taxonomicName id="9F854D2D1F79FF9F74B9088AFC3AABB5" authorityName="Distant" authorityYear="1903" box="[909,1000,1190,1213]" class="Insecta" family="Veliidae" genus="Baptista" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F79FF9F74B9088AFC3AABB5" box="[909,1000,1190,1213]" italics="true" pageId="1" pageNumber="518">Baptista</emphasis>
</taxonomicName>
, including those hitherto reported from
<collectingCountry id="2092763E1F79FF9F740808EAFCA5ABD5" box="[828,887,1222,1245]" name="India" pageId="1" pageNumber="518">India</collectingCountry>
, differ from
<taxonomicName id="9F854D2D1F79FF9F74CA08EAFB54ABD5" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[1022,1158,1222,1245]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F79FF9F74CA08EAFB54ABD5" box="[1022,1158,1222,1245]" italics="true" pageId="1" pageNumber="518">Thirumalaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F79FF9F73B908EAFB22ABD5" box="[1165,1264,1222,1245]" pageId="1" pageNumber="518" rank="genus">gen. nov.</taxonomicNameLabel>
by strong modifications of foreleg and abdominal segments 67 of the males (=
<taxonomicName id="9F854D2D1F79FF9F74AB092AFBB5AA15" authorityName="Andersen" authorityYear="1989" box="[927,1127,1286,1309]" class="Insecta" family="Veliidae" genus="Baptista" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="6AF1EABC1F79FF9F74AB092AFBB5AA15" box="[927,1127,1286,1309]" italics="true" pageId="1" pageNumber="518">Baptista femoralis</emphasis>
</taxonomicName>
group sensu Kඈඏൺർ &amp; YൺඇG 2000). Four species of
<taxonomicName id="9F854D2D1F79FF9F7307090AFB5CAA35" authorityName="Distant" authorityYear="1903" box="[1075,1166,1318,1341]" class="Insecta" family="Veliidae" genus="Baptista" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F79FF9F7307090AFB5CAA35" box="[1075,1166,1318,1341]" italics="true" pageId="1" pageNumber="518">Baptista</emphasis>
</taxonomicName>
from Southeast Asia and Southern
<collectingCountry id="2092763E1F79FF9F7497096AFC31AA55" box="[931,995,1350,1373]" name="China" pageId="1" pageNumber="518">China</collectingCountry>
form the
<taxonomicName id="9F854D2D1F79FF9F737C096AFB29AA55" baseAuthorityName="Andersen" baseAuthorityYear="1989" box="[1096,1275,1350,1373]" class="Insecta" family="Veliidae" genus="Baptista" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="species" species="collaris">
<emphasis id="6AF1EABC1F79FF9F737C096AFB29AA55" box="[1096,1275,1350,1373]" italics="true" pageId="1" pageNumber="518">Baptista collaris</emphasis>
</taxonomicName>
group and also lack such modifications (Zൾඍඍൾඅ 2004, Yൾ et al. 2014b). However,
<taxonomicName id="9F854D2D1F79FF9F749A09AAFBE4AA95" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[942,1078,1414,1437]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F79FF9F749A09AAFBE4AA95" box="[942,1078,1414,1437]" italics="true" pageId="1" pageNumber="518">Thirumalaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F79FF9F730B09AAFB76AA95" box="[1087,1188,1414,1437]" pageId="1" pageNumber="518" rank="genus">gen. nov.</taxonomicNameLabel>
possesses none of the characters defining this group either (described in detail by Zൾඍඍൾඅ 2004). The sexually dimorphic body shape, the enlarged compound eyes, and the characteristic parameres are the most important characteristics of
<taxonomicName id="9F854D2D1F79FF9F72270A2AFA4EA915" authority="For" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[1299,1436,1542,1565]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F79FF9F72270A2AFA4EA915" box="[1299,1436,1542,1565]" italics="true" pageId="1" pageNumber="518">Thirumalaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F79FF9F74080A0AFC4DA935" box="[828,927,1574,1597]" pageId="1" pageNumber="518" rank="genus">gen. nov.</taxonomicNameLabel>
For distinction from other genera see the key.
<emphasis id="6AF1EABC1F79FF9F74080A6AFC6BA955" bold="true" box="[828,953,1606,1629]" pageId="1" pageNumber="518">Etymology.</emphasis>
Dedicated to the memory of the Scientist of the Zoological Survey of
<collectingCountry id="2092763E1F79FF9F73110A4AFB8EA975" box="[1061,1116,1638,1661]" name="India" pageId="1" pageNumber="518">India</collectingCountry>
Dr. G. Thirumalai, honouring his significant contribution to the knowledge of Indian water bugs. Gender: feminine.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,197 @@
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<paragraph id="583A36AE1F7BFF9D74C90972FB09AA7D" blockId="3.[1021,1243,1374,1397]" box="[1021,1243,1374,1397]" pageId="3" pageNumber="520">
<heading id="037281C21F7BFF9D74C90972FB09AA7D" box="[1021,1243,1374,1397]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="520" reason="3">
<emphasis id="6AF1EABC1F7BFF9D74C90972FB09AA7D" bold="true" box="[1021,1243,1374,1397]" pageId="3" pageNumber="520">
<taxonomicName id="9F854D2D1F7BFF9D74C90972FBA3AA7D" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[1021,1137,1374,1397]" class="Insecta" family="Veliidae" genus="Eyarinella" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F7BFF9D74C90972FBA3AA7D" bold="true" box="[1021,1137,1374,1397]" italics="true" pageId="3" pageNumber="520">Eyarinella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7BFF9D73430972FB09AA7D" box="[1143,1243,1374,1397]" pageId="3" pageNumber="520" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="109F65251F7BFF9D73140950FB6AAA87" box="[1056,1208,1404,1423]" pageId="3" pageNumber="520" type="description">
<paragraph id="583A36AE1F7BFF9D73140950FB6AAA87" blockId="3.[1056,1208,1404,1423]" box="[1056,1208,1404,1423]" pageId="3" pageNumber="520">
(
<figureCitation id="C0BE2A2B1F7BFF9D73120950FBA0AA87" box="[1062,1138,1404,1423]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="3" pageNumber="520">Figs 3, 4</figureCitation>
,
<figureCitation id="C0BE2A2B1F7BFF9D73480950FB60AA87" box="[1148,1202,1404,1423]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="3" pageNumber="520">1624</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="109F65251F7BFF9D74080984FB25AAB7" box="[828,1271,1448,1471]" pageId="3" pageNumber="520" type="type_taxon">
<paragraph id="583A36AE1F7BFF9D74080984FB25AAB7" blockId="3.[828,1437,1448,2079]" box="[828,1271,1448,1471]" pageId="3" pageNumber="520">
<emphasis id="6AF1EABC1F7BFF9D74080984FC02AAB7" bold="true" box="[828,976,1448,1471]" pageId="3" pageNumber="520">
<typeStatus id="873E880C1F7BFF9D74080984FCA6AAB7" box="[828,884,1448,1471]" pageId="3" pageNumber="520">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="9F854D2D1F7BFF9D74E20984FB4CAAB7" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[982,1182,1448,1471]" class="Insecta" family="Veliidae" genus="Eyarinella" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="6AF1EABC1F7BFF9D74E20984FB4CAAB7" box="[982,1182,1448,1471]" italics="true" pageId="3" pageNumber="520">Eyarinella robusta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7BFF9D73900984FB25AAB7" box="[1188,1271,1448,1471]" pageId="3" pageNumber="520" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="109F65251F7BFF9B740809E4FE88AA26" lastPageId="5" lastPageNumber="522" pageId="3" pageNumber="520" type="diagnosis">
<paragraph id="583A36AE1F7BFF9A740809E4FE56A874" blockId="3.[828,1437,1448,2079]" lastBlockId="4.[151,760,1861,2076]" lastPageId="4" lastPageNumber="521" pageId="3" pageNumber="520">
<emphasis id="6AF1EABC1F7BFF9D740809E4FC62AAD7" bold="true" box="[828,944,1480,1503]" pageId="3" pageNumber="520">Diagnosis.</emphasis>
Body stout, ovate (
<figureCitation id="C0BE2A2B1F7BFF9D734E09E4FB05AAD7" box="[1146,1239,1480,1503]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="3" pageNumber="520">Figs 3, 4</figureCitation>
). Posterior of head deeply inserted in anterior part of pronotum (
<figureCitation id="C0BE2A2B1F7BFF9D720F09C4FA42AAF7" box="[1339,1424,1512,1535]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="3" pageNumber="520">Fig. 16</figureCitation>
). Eyes moderately large (
<figureCitation id="C0BE2A2B1F7BFF9D73750A24FB11A917" box="[1089,1219,1544,1567]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="3" pageNumber="520">Figs 16, 17</figureCitation>
). Antenna long and slender, about two thirds of body length; antennomere I (
<figureCitation id="C0BE2A2B1F7BFF9D74700A64FC42A957" box="[836,912,1608,1631]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="3" pageNumber="520">Fig. 16</figureCitation>
) 0.9 times as long as head length; relative lengths of antennomeres: IV&gt; III ≧ I&gt; II. Pronotum convexly rounded posteriorly (
<figureCitation id="C0BE2A2B1F7BFF9D73200AA4FBB2A997" box="[1044,1120,1672,1695]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="3" pageNumber="520">Fig. 16</figureCitation>
). Lateral evaporatorium small, situated in ventral half of metapleuron. Legs stout. Claws preterminal, their apices not reaching apex of tarsus. Male: Propleuron with a sharp ridge behind eye (
<figureCitation id="C0BE2A2B1F7BFF9D72750AC4FA5DA9F7" box="[1345,1423,1768,1791]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="3" pageNumber="520">Fig. 17</figureCitation>
), in dorsal view appearing tooth-like. Femora stout (
<figureCitation id="C0BE2A2B1F7BFF9D72730B24FC8BA837" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="3" pageNumber="520">Figs 18, 19</figureCitation>
), otherwise unmodified. Protibia unmodified; protibial grasping comb extremely short (
<figureCitation id="C0BE2A2B1F7BFF9D73AB0B64FB22A857" box="[1183,1264,1864,1887]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="3" pageNumber="520">Fig. 18</figureCitation>
), restricted to a short ear-shaped spur at apex of tibia. Mesotibia without grasping comb. Pregenital abdomen without projections. Segment 8 moderately elongated (
<figureCitation id="C0BE2A2B1F7BFF9D73AC0B84FB36A8B7" box="[1176,1252,1960,1983]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="3" pageNumber="520">Fig. 21</figureCitation>
). Pygophore (
<figureCitation id="C0BE2A2B1F7BFF9D72460B84FC8AA8D7" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="3" pageNumber="520">Fig. 22</figureCitation>
) distally upcurved and with concave posterior margin. Proctiger (
<figureCitation id="C0BE2A2B1F7BFF9D749F0BC4FC2AA8F7" box="[939,1016,2024,2047]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="3" pageNumber="520">Fig. 23</figureCitation>
) narrow, tongue-shaped; apex narrowly rounded. Parameres (
<figureCitation id="C0BE2A2B1F7BFF9D73160424FBA0A717" box="[1058,1138,2056,2079]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="3" pageNumber="520">Fig. 24</figureCitation>
) symmetrical, curved, very long and slender. Female: Genitalia situated in a posterior opening of segment 7.
</paragraph>
<caption id="0CFA66261F7CFF9A77A30AC5FDE9A81E" ID-DOI="http://doi.org/10.5281/zenodo.5821219" ID-Zenodo-Dep="5821219" httpUri="https://zenodo.org/record/5821219/files/figure.png" pageId="4" pageNumber="521" startId="4.[151,188,1769,1788]" targetBox="[151,1436,189,1749]" targetPageId="4">
<paragraph id="583A36AE1F7CFF9A77A30AC5FDE9A81E" blockId="4.[151,1437,1769,1814]" pageId="4" pageNumber="521">
Figs 58. Habitus of
<taxonomicName id="9F854D2D1F7CFF9A767B0AC5FE31A9F4" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[335,483,1769,1788]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="521" phylum="Arthropoda" rank="species" species="fikaceki" status="sp. nov.">
<emphasis id="6AF1EABC1F7CFF9A767B0AC5FE31A9F4" box="[335,483,1769,1788]" italics="true" pageId="4" pageNumber="521">Geovelia fikaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7CFF9A76DD0AC5FDFFA9F4" box="[489,557,1769,1788]" pageId="4" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
(5 apterous male, 6 apterous female), and
<taxonomicName id="9F854D2D1F7CFF9A74F50AC5FB3DA9F4" authority="(Distant, 1903)" baseAuthorityName="Distant" baseAuthorityYear="1903" box="[961,1263,1769,1788]" class="Insecta" family="Veliidae" genus="Neoalardus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="521" phylum="Arthropoda" rank="species" species="typicus">
<emphasis id="6AF1EABC1F7CFF9A74F50AC5FBB5A9F4" box="[961,1127,1769,1788]" italics="true" pageId="4" pageNumber="521">Neoalardus typicus</emphasis>
(
<bibRefCitation id="3C144B5F1F7CFF9A73470AC5FB38A9F4" author="DISTANT W. L." box="[1139,1258,1769,1788]" pageId="4" pageNumber="521" pagination="219 - 274" refId="ref7875" refString="DISTANT W. L. 1903: Report on the Rhynchota. Part I. Heteroptera. Fascicula Malayenses Zoology 2: 219 - 274." type="journal article" year="1903">Distant, 1903</bibRefCitation>
)
</taxonomicName>
(7 apterous male, 8 apterous female), both specimens from India.
</paragraph>
</caption>
<paragraph id="583A36AE1F7CFF9B77A30BA9FE88AA26" blockId="4.[151,760,1861,2076]" lastBlockId="5.[151,760,1079,1326]" lastPageId="5" lastPageNumber="522" pageId="4" pageNumber="521">
<emphasis id="6AF1EABC1F7CFF9A77A30BA9FEA7A894" bold="true" box="[151,373,1925,1948]" pageId="4" pageNumber="521">Comparative notes.</emphasis>
The stout body distinguishes
<taxonomicName id="9F854D2D1F7CFF9A75870BA9FF1EA8B4" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" class="Insecta" family="Veliidae" genus="Eyarinella" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="521" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F7CFF9A75870BA9FF1EA8B4" italics="true" pageId="4" pageNumber="521">Eyarinella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7CFF9A77E50B89FEFDA8B4" box="[209,303,1957,1980]" pageId="4" pageNumber="521" rank="genus">gen. nov.</taxonomicNameLabel>
from all other Oriental genera of Microveliinae, except from
<taxonomicName id="9F854D2D1F7CFF9A767B0BE9FE7CA8D4" authorityName="Zimmermann" authorityYear="1984" box="[335,430,1989,2012]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F7CFF9A767B0BE9FE7CA8D4" box="[335,430,1989,2012]" italics="true" pageId="4" pageNumber="521">Geovelia</emphasis>
</taxonomicName>
and a few species of
<taxonomicName id="9F854D2D1F7CFF9A75B10BE9FD25A8D4" authorityName="Westwood" authorityYear="1834" box="[645,759,1989,2012]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F7CFF9A75B10BE9FD25A8D4" box="[645,759,1989,2012]" italics="true" pageId="4" pageNumber="521">Microvelia</emphasis>
</taxonomicName>
(
<taxonomicName id="9F854D2D1F7CFF9A77940BC9FEC6A8F4" authorityName="Westwood" authorityYear="1834" box="[160,276,2021,2044]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F7CFF9A77940BC9FEC6A8F4" box="[160,276,2021,2044]" italics="true" pageId="4" pageNumber="521">Microvelia</emphasis>
</taxonomicName>
shows a great variation in body shape). However, in
<taxonomicName id="9F854D2D1F7CFF9A77DC0429FE89A714" authorityName="Westwood" authorityYear="1834" box="[232,347,2053,2076]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F7CFF9A77DC0429FE89A714" box="[232,347,2053,2076]" italics="true" pageId="4" pageNumber="521">Microvelia</emphasis>
</taxonomicName>
the antenna is much shorter and stouter, and the genitalia of females are less protruding.
<taxonomicName id="9F854D2D1F7CFF9A72080B69FA4EA854" authorityName="Zimmermann" authorityYear="1984" box="[1340,1436,1861,1884]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F7CFF9A72080B69FA4EA854" box="[1340,1436,1861,1884]" italics="true" pageId="4" pageNumber="521">Geovelia</emphasis>
</taxonomicName>
differs from
<taxonomicName id="9F854D2D1F7CFF9A748F0B49FBF8A874" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[955,1066,1893,1916]" class="Insecta" family="Veliidae" genus="Eyarinella" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="521" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F7CFF9A748F0B49FBF8A874" box="[955,1066,1893,1916]" italics="true" pageId="4" pageNumber="521">Eyarinella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7CFF9A731B0B49FB5CA874" box="[1071,1166,1893,1916]" pageId="4" pageNumber="521" rank="genus">gen. nov.</taxonomicNameLabel>
in several characters, e.g., by a short head that is not deeply embedded in a concavity of the pronotum, by large eyes (e.g.,
<figureCitation id="C0BE2A2B1F7CFF9A73E30B89FAE9A8B4" box="[1239,1339,1957,1980]" captionStart="Figs 58" captionStartId="4.[151,188,1769,1788]" captionTargetBox="[151,1436,189,1749]" captionTargetId="figure-142@4.[151,1436,189,1749]" captionTargetPageId="4" captionText="Figs 58. Habitus of Geovelia fikaceki sp. nov. (5 apterous male, 6 apterous female), and Neoalardus typicus (Distant, 1903) (7 apterous male, 8 apterous female), both specimens from India." figureDoi="http://doi.org/10.5281/zenodo.5821219" httpUri="https://zenodo.org/record/5821219/files/figure.png" pageId="4" pageNumber="521">Figs 5, 6</figureCitation>
), and by subterminal claws. The long antenna of
<taxonomicName id="9F854D2D1F7CFF9A73C10BE9FAB4A8D4" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[1269,1382,1989,2012]" class="Insecta" family="Veliidae" genus="Eyarinella" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="521" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F7CFF9A73C10BE9FAB4A8D4" box="[1269,1382,1989,2012]" italics="true" pageId="4" pageNumber="521">Eyarinella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7CFF9A725B0BE9FCBBA8F4" pageId="4" pageNumber="521" rank="genus">gen. nov.</taxonomicNameLabel>
resembles several genera of Microveliinae with a hygropetric lifestyle, e.g.,
<taxonomicName id="9F854D2D1F7CFF9A73190429FAC4A714" authority="Distant, 1903" authorityName="Distant" authorityYear="1903" box="[1069,1302,2053,2076]" class="Insecta" family="Veliidae" genus="Baptista" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F7CFF9A73190429FB56A714" box="[1069,1156,2053,2076]" italics="true" pageId="4" pageNumber="521">Baptista</emphasis>
<bibRefCitation id="3C144B5F1F7CFF9A73BD0429FAC4A714" author="DISTANT W. L." box="[1161,1302,2053,2076]" pageId="4" pageNumber="521" pagination="219 - 274" refId="ref7875" refString="DISTANT W. L. 1903: Report on the Rhynchota. Part I. Heteroptera. Fascicula Malayenses Zoology 2: 219 - 274." type="journal article" year="1903">Distant, 1903</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="9F854D2D1F7CFF9B722A0429FE80AB46" authority="Thirumalai, 1999" authorityName="Thirumalai" authorityYear="1999" class="Insecta" family="Veliidae" genus="Aquulavelia" kingdom="Animalia" lastPageId="5" lastPageNumber="522" order="Hemiptera" pageId="4" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F7CFF9A722A0429FA4EA714" box="[1310,1436,2053,2076]" italics="true" pageId="4" pageNumber="521">Aquulavelia</emphasis>
<bibRefCitation id="3C144B5F1F7DFF9B77A3081BFE80AB46" author="THIRUMALAI G." box="[151,338,1079,1102]" pageId="5" pageNumber="522" pagination="205 - 211" refId="ref8607" refString="THIRUMALAI G. 1999: A new genus of Microveliinae (Hemiptera: Heteroptera: Veliidae) from North Eastern India with a checklist of the Indian species of the subfamily. Records of the Zoological Survey of India 97 (3): 205 - 211." type="journal article" year="1999">Thirumalai, 1999</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="9F854D2D1F7DFF9B76BC081BFDDFAB46" authority="The" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[392,525,1079,1102]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="522" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F7DFF9B76BC081BFDDFAB46" box="[392,525,1079,1102]" italics="true" pageId="5" pageNumber="522">Thirumalaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7DFF9B7527081BFDA6AB46" box="[531,628,1079,1102]" pageId="5" pageNumber="522" rank="genus">gen. nov.</taxonomicNameLabel>
The male of
<taxonomicName id="9F854D2D1F7DFF9B77A3087BFEDAAB66" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[151,264,1111,1134]" class="Insecta" family="Veliidae" genus="Eyarinella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="522" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F7DFF9B77A3087BFEDAAB66" box="[151,264,1111,1134]" italics="true" pageId="5" pageNumber="522">Eyarinella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7DFF9B763B087BFEA0AB66" box="[271,370,1111,1134]" pageId="5" pageNumber="522" rank="genus">gen. nov.</taxonomicNameLabel>
possesses the following two unique characters: the sharp ridge anteriorly on propleuron (
<figureCitation id="C0BE2A2B1F7DFF9B75F8085BFF61ABA6" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="5" pageNumber="522">Fig. 17</figureCitation>
), and the rudimentary protibial grasping comb (
<figureCitation id="C0BE2A2B1F7DFF9B75AB08BBFD39ABA6" box="[671,747,1175,1198]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="5" pageNumber="522">Fig. 18</figureCitation>
).
<emphasis id="6AF1EABC1F7DFF9B77A3089BFECEABC6" bold="true" box="[151,284,1207,1230]" pageId="5" pageNumber="522">Etymology.</emphasis>
The generic name is dedicated to Dr. E. Eyarin Jehamalar of the Zoological Survey of
<collectingCountry id="2092763E1F7DFF9B75AC08FBFD1DABE6" box="[664,719,1239,1262]" name="India" pageId="5" pageNumber="522">India</collectingCountry>
for her numerous valuable publications on Indian water bugs. Gender: feminine.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,235 @@
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<mods:title id="C74953593F7CFDA9F4E8EDC40E263725">The taxonomy of some unusual Microveliinae (Hemiptera: Heteroptera: Veliidae) from India</mods:title>
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<paragraph id="583A36AE1F7DFF9B7607097BFD8EAA66" blockId="5.[307,604,1367,1390]" box="[307,604,1367,1390]" pageId="5" pageNumber="522">
<heading id="037281C21F7DFF9B7607097BFD8EAA66" box="[307,604,1367,1390]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="522" reason="3">
<emphasis id="6AF1EABC1F7DFF9B7607097BFD8EAA66" bold="true" box="[307,604,1367,1390]" pageId="5" pageNumber="522">
<taxonomicName id="9F854D2D1F7DFF9B7607097BFDD2AA66" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[307,512,1367,1390]" class="Insecta" family="Veliidae" genus="Eyarinella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="522" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="6AF1EABC1F7DFF9B7607097BFDD2AA66" bold="true" box="[307,512,1367,1390]" italics="true" pageId="5" pageNumber="522">Eyarinella robusta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7DFF9B7533097BFD8EAA66" box="[519,604,1367,1390]" pageId="5" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="109F65251F7DFF9B764F095AFDC1AA81" box="[379,531,1398,1417]" pageId="5" pageNumber="522" type="description">
<paragraph id="583A36AE1F7DFF9B764F095AFDC1AA81" blockId="5.[151,761,1398,2072]" box="[379,531,1398,1417]" pageId="5" pageNumber="522">
(
<figureCitation id="C0BE2A2B1F7DFF9B76B5095AFE1CAA81" box="[385,462,1398,1417]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="5" pageNumber="522">Figs 3, 4</figureCitation>
,
<figureCitation id="C0BE2A2B1F7DFF9B76E3095AFDDFAA81" box="[471,525,1398,1417]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="5" pageNumber="522">1624</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="109F65251F7DFF9877A309B3FDA5A704" lastPageId="6" lastPageNumber="523" pageId="5" pageNumber="522" type="materials_examined">
<paragraph id="583A36AE1F7DFF9B77A309B3FBE0ABA6" blockId="5.[151,761,1398,2072]" lastBlockId="5.[828,1437,1079,2062]" pageId="5" pageNumber="522">
<emphasis id="6AF1EABC1F7DFF9B77A309B3FECDAABA" bold="true" box="[151,287,1439,1458]" pageId="5" pageNumber="522">Type material:</emphasis>
Hඈඅඈඍඒඉൾ: J (apterous) (
<collectionCode id="3E94AE6B1F7DFF9B753009B3FD9CAABA" box="[516,590,1439,1458]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="522" type="Museum">NHMW</collectionCode>
), labelled “
<collectingCountry id="2092763E1F7DFF9B758609B3FD26AABA" box="[690,756,1439,1458]" name="India" pageId="5" pageNumber="522">INDIA</collectingCountry>
:
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,
<date id="2C3B106E1F7DFF9B77EE0996FEF6AAC5" box="[218,292,1466,1485]" pageId="5" pageNumber="522" value="1999-01-08">8.1.1999</date>
\
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WSW Munnar,
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\ Kallar Valley,
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<geoCoordinate id="3DB150691F7DFF9B77A309F9FF0EAAE0" box="[151,220,1493,1512]" degrees="76" direction="east" minutes="59" orientation="longitude" pageId="5" pageNumber="522" precision="925" value="76.98333">76°59E</geoCoordinate>
\ leg.D. Boukal (52)”. Pൺඋൺඍඒඉൾඌ (
<collectionCode id="3E94AE6B1F7DFF9B76CA09F9FD9BAAE0" box="[510,585,1493,1512]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="522" type="Museum">NHMW</collectionCode>
): 1J (apterous) with same label data;
<specimenCount id="4E83FD271F7DFF9B762909C3FEEBA90A" box="[285,313,1513,1538]" pageId="5" pageNumber="522" type="female">1♀</specimenCount>
(apterous),labelled “
<collectingCountry id="2092763E1F7DFF9B76DE09C3FDF9A90A" box="[490,555,1519,1538]" name="India" pageId="5" pageNumber="522">INDIA</collectingCountry>
:
<collectingRegion id="9A41F84C1F7DFF9B750609C3FDBEA90A" box="[562,620,1519,1538]" country="India" name="Kerala" pageId="5" pageNumber="522">Kerala</collectingRegion>
,
<date id="2C3B106E1F7DFF9B754609C3FD69A90A" box="[626,699,1519,1538]" pageId="5" pageNumber="522" value="1999-01-07">7.1.1999</date>
\
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WSW Munnar,
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\ Kallar Valley,
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<geoCoordinate id="3DB150691F7DFF9B751D0A26FDBCA915" box="[553,622,1546,1565]" degrees="76" direction="east" minutes="58" orientation="longitude" pageId="5" pageNumber="522" precision="925" value="76.96667">76°58E</geoCoordinate>
\ leg. D. Boukal (50)”;
<specimenCount id="4E83FD271F7DFF9B77FB0A09FF3DA93F" box="[207,239,1566,1592]" pageId="5" pageNumber="522" type="female">1 ♀</specimenCount>
(apterous;
<typeStatus id="873E880C1F7DFF9B767B0A09FE45A930" box="[335,407,1573,1592]" pageId="5" pageNumber="522" type="allotype">allotype</typeStatus>
), labelled “
<collectingCountry id="2092763E1F7DFF9B76CC0A09FDE8A930" box="[504,570,1573,1592]" name="India" pageId="5" pageNumber="522">INDIA</collectingCountry>
:
<collectingRegion id="9A41F84C1F7DFF9B75770A09FDACA930" box="[579,638,1573,1592]" country="India" name="Kerala" pageId="5" pageNumber="522">Kerala</collectingRegion>
, Kallar Valley \
<quantity id="9F7D9B4B1F7DFF9B77900A13FF07A95A" box="[164,213,1599,1618]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="522" unit="km" value="10.0">10km</quantity>
WSW Munnar, \
<geoCoordinate id="3DB150691F7DFF9B76450A13FE6AA95A" box="[369,440,1599,1618]" degrees="10" direction="north" minutes="03" orientation="latitude" pageId="5" pageNumber="522" precision="925" value="10.05">10°03N</geoCoordinate>
76°5859E \ 1100- \,
<quantity id="9F7D9B4B1F7DFF9B75B60A13FD16A95A" box="[642,708,1599,1618]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="522" unit="m" value="1200.0">1200 m</quantity>
a.s.l., 7
<date id="2C3B106E1F7DFF9B77980A76FF25A965" box="[172,247,1626,1645]" pageId="5" pageNumber="522" value="1999-01-08">8.1.1999</date>
\ leg. D. Boukal (49/52)”;
<specimenCount id="4E83FD271F7DFF9B76EF0A76FE28A965" box="[475,506,1620,1645]" pageId="5" pageNumber="522" type="female">1 ♀</specimenCount>
(apterous), labelled “
<collectingCountry id="2092763E1F7DFF9B75860A76FD26A965" box="[690,756,1626,1645]" name="India" pageId="5" pageNumber="522">INDIA</collectingCountry>
:
<collectingRegion id="9A41F84C1F7DFF9B77A30A59FF2DA980" box="[151,255,1653,1672]" country="India" name="Tamil Nadu" pageId="5" pageNumber="522">Tamil Nadu</collectingRegion>
,
<date id="2C3B106E1F7DFF9B763D0A59FE8CA980" box="[265,350,1653,1672]" pageId="5" pageNumber="522" value="1999-01-25">25.1.1999</date>
\ Shevaroy Hills, Yercaud \ Kiliyur Falls env. \
<geoCoordinate id="3DB150691F7DFF9B77A30AA3FF0FA9AA" box="[151,221,1679,1698]" degrees="11" direction="north" minutes="47" orientation="latitude" pageId="5" pageNumber="522" precision="925" value="11.783334">11°47N</geoCoordinate>
<geoCoordinate id="3DB150691F7DFF9B77D60AA3FEFBA9AA" box="[226,297,1679,1698]" degrees="78" direction="east" minutes="12" orientation="longitude" pageId="5" pageNumber="522" precision="925" value="78.2">78°12E</geoCoordinate>
,
<quantity id="9F7D9B4B1F7DFF9B76050AA3FEBEA9AA" box="[305,364,1679,1698]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="522" unit="m" value="1400.0">1400m</quantity>
\ leg. D. Boukal (85)”;
<specimenCount id="4E83FD271F7DFF9B75050AA3FD9DA9AA" box="[561,591,1673,1698]" pageId="5" pageNumber="522" type="female">1 ♀</specimenCount>
(apterous), labelled “S.
<collectingCountry id="2092763E1F7DFF9B77820A86FF27A9B5" box="[182,245,1706,1725]" name="India" pageId="5" pageNumber="522">INDIA</collectingCountry>
,
<collectingRegion id="9A41F84C1F7DFF9B77CF0A86FE86A9B5" box="[251,340,1706,1725]" country="India" name="Kerala" pageId="5" pageNumber="522">KERALA</collectingRegion>
, Thekkady \ Periyar Lake,
<geoCoordinate id="3DB150691F7DFF9B75070A86FDA8A9B5" box="[563,634,1706,1725]" degrees="09" direction="north" minutes="34" orientation="latitude" pageId="5" pageNumber="522" precision="925" value="9.566667">09°34N</geoCoordinate>
<geoCoordinate id="3DB150691F7DFF9B754A0A86FD11A9B5" box="[638,707,1706,1725]" degrees="77" direction="east" minutes="10" orientation="longitude" pageId="5" pageNumber="522" precision="925" value="77.166664">77°10E</geoCoordinate>
\
<quantity id="9F7D9B4B1F7DFF9B75E50A86FF07A9D0" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="5" pageNumber="522" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">900- 1000m</quantity>
,
<date id="2C3B106E1F7DFF9B77E90AE9FE8FA9D0" box="[221,349,1733,1752]" pageId="5" pageNumber="522" value="1997-04-19" valueMax="1997-04-27" valueMin="1997-04-19">19.-27.IV.1997</date>
\ leg. Dembicky &amp; Pacholátko leg.”
<emphasis id="6AF1EABC1F7DFF9B77A30ACDFE0BA9F0" bold="true" box="[151,473,1761,1784]" pageId="5" pageNumber="522">Description of apterous male.</emphasis>
Measurements of
<typeStatus id="873E880C1F7DFF9B75A10ACDFD27A9F0" box="[661,757,1761,1784]" pageId="5" pageNumber="522" type="holotype">holotype</typeStatus>
: Body length 2.52; maximum body width (at abdominal segment 4) 1.05. Head length 0.44, width 0.65. Synthlipsis 0.36. Pronotum length 0.38, width 0.91. Lengths of antennomeres, I 0.44, II 0.34, III 0.48, IV 0.60. Lengths of leg segments: profemur 0.76, protibia 0.72, protarsus 0.34, mesofemur 0.88, mesotibia 0.82, mesotarsus 0.14 + 0.29, metafemur 1.00, metatibia 1.02, metatarsus 0.12 + 0.28; protibial comb length 0.02; comb index 0.03. Measurements of
<typeStatus id="873E880C1F7DFF9B77C8042DFEB0A710" box="[252,354,2049,2072]" pageId="5" pageNumber="522" type="paratype">paratypes</typeStatus>
(n = 1): Body length 2.57; maximum body width (at abdominal segment 4) 1.08. Head length 0.44, width 0.66. Synthlipsis 0.35. Pronotum length 0.34, width 0.89. Protibia length 0.70; protibial comb length 0.03; comb index 0.03.
</paragraph>
<paragraph id="583A36AE1F7DFF9B746E089BFC71A926" blockId="5.[828,1437,1079,2062]" pageId="5" pageNumber="522">
Colour (
<figureCitation id="C0BE2A2B1F7DFF9B7486089BFC23ABC6" box="[946,1009,1207,1230]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="5" pageNumber="522">Fig. 3</figureCitation>
): Dark brown to black. Posterior margin of head and stripes along medial eye margin more or less dark orange; only
<typeStatus id="873E880C1F7DFF9B732408DBFBA2AA06" box="[1040,1136,1271,1294]" pageId="5" pageNumber="522" type="holotype">holotype</typeStatus>
provided posteriorly with a pair of yellow marks. Pronotum anteromedially with a transverse, broad, yellow stripe, sides and a narrow hind margin more or less orange-brown; sharp edge on propleuron below eye yellow. Mediotergite 1 at middle orange-brown. Narrow lateral margins of laterotergites and sterna pale orange. Sides of segment 8 yellowish. Antenna pale brown. Legs basally pale yellow, apices of femora, bases and apices of tibiae, and apices of tarsi indistinctly infuscate.
</paragraph>
<paragraph id="583A36AE1F7DFF9B746E0A1BFB5FA706" blockId="5.[828,1437,1079,2062]" pageId="5" pageNumber="522">
Pilosity: A short grey appressed pilosity covering body except disc of pronotum. In addition, almost entire body (except sides and venter of head and thorax) with scattered long setae, longest on pronotum and dorsalmost part of metapleuron; those on head and pronotum standing, those on abdomen oblique. Metapleuron densely hirsute. Silverish hairs (
<figureCitation id="C0BE2A2B1F7DFF9B74AA0ADBFC09A806" box="[926,987,1783,1806]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="5" pageNumber="522">Fig. 3</figureCitation>
) arranged in a pair of large patches laterally on mediotergite 1 and small patches at hind margins of laterotergites 26. Antenna with two long setae on antennomere I, and one on antennomere II; all antennomeres with several shorter setae. Legs densely beset with decumbent pilosity; profemur with row of standing setae on flexor side; meso- and metafemur with some longer oblique setae on flexor side subapically; all tibiae with moderately long, oblique setae on extensor sides.
</paragraph>
<caption id="0CFA66261F7EFF9877A3083FFD8BAB7E" ID-DOI="http://doi.org/10.5281/zenodo.5821223" ID-Zenodo-Dep="5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="6" pageNumber="523" startId="6.[151,188,1043,1062]" targetBox="[151,1432,190,1020]" targetPageId="6">
<paragraph id="583A36AE1F7EFF9877A3083FFD8BAB7E" blockId="6.[151,1436,1043,1142]" pageId="6" pageNumber="523">
Figs 1624. Structural details of
<taxonomicName id="9F854D2D1F7EFF987687083FFD86AB2E" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[435,596,1043,1062]" class="Insecta" family="Veliidae" genus="Eyarinella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="523" phylum="Arthropoda" rank="species" species="robusta" status="gen. nov.">
<emphasis id="6AF1EABC1F7EFF987687083FFD86AB2E" box="[435,596,1043,1062]" italics="true" pageId="6" pageNumber="523">Eyarinella robusta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7EFF98756E083FFD78AB2E" box="[602,682,1043,1062]" pageId="6" pageNumber="523" rank="genus">gen. nov.</taxonomicNameLabel>
&amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24.
</paragraph>
</caption>
<paragraph id="583A36AE1F7EFF9877810899FDA5A704" blockId="6.[151,760,1205,2060]" pageId="6" pageNumber="523">
Structures: Head (
<figureCitation id="C0BE2A2B1F7EFF98764B0899FE03ABC4" box="[383,465,1205,1228]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="6" pageNumber="523">Fig. 16</figureCitation>
) short, with pointed apex, posterodorsally deeply inserted in strongly concave anterior margin of pronotum. Eye moderately large (
<figureCitation id="C0BE2A2B1F7EFF9875AB08D9FF66AA24" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="6" pageNumber="523">Figs 16, 17</figureCitation>
), posteriorly touching pronotum. Antenna very long and slender; antennomere I surpassing apex of head by about three fourths of its length (
<figureCitation id="C0BE2A2B1F7EFF9876D80979FDE8AA64" box="[492,570,1365,1388]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="6" pageNumber="523">Fig. 16</figureCitation>
); antennomeres II more slender, III and IV very slender, filiform. Pronotum (
<figureCitation id="C0BE2A2B1F7EFF9877AB09B9FF39AAA4" box="[159,235,1429,1452]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="6" pageNumber="523">Fig. 16</figureCitation>
) long, anterior margin broadly concave, posterior margin broadly convex, but with a shallow concavity in approximately medial fourth; only there metanotum visible as a narrow transverse line. Pronotum with transverse rows of punctures in front of and behind yellow stripe, and with scattered punctures on disc except on midline. Propleuron with short ridge at anterior margin behind eye (
<figureCitation id="C0BE2A2B1F7EFF9875AE0A79FD38A964" box="[666,746,1621,1644]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="6" pageNumber="523">Fig. 17</figureCitation>
). Legs moderately long, stout; profemur slightly narrower than mesofemur and much narrower than metafemur (
<figureCitation id="C0BE2A2B1F7EFF9875F90AB9FF66A9C4" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="6" pageNumber="523">Fig. 19</figureCitation>
). Protibial grasping comb very stout, situated on a small apical spur (
<figureCitation id="C0BE2A2B1F7EFF9876100AF9FEABA9E4" box="[292,377,1749,1772]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="6" pageNumber="523">Fig. 18</figureCitation>
). Pregenital abdomen very broad; mediotergites about three times wider than laterotergites; mediotergite 7 ca. 1.5 times longer than 6, about 2.2 times wider than long. Segment 8 (
<figureCitation id="C0BE2A2B1F7EFF9876EE0B19FDF8A844" box="[474,554,1845,1868]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="6" pageNumber="523">Fig. 21</figureCitation>
) moderately wide, subcylindrical. Pygophore (
<figureCitation id="C0BE2A2B1F7EFF9876860B79FE2EA864" box="[434,508,1877,1900]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="6" pageNumber="523">Fig. 22</figureCitation>
) slender, ca. 1.9 times as long as wide, distally upcurved, and with concave posterior margin. Proctiger (
<figureCitation id="C0BE2A2B1F7EFF9876540BB9FE7CA8A4" box="[352,430,1941,1964]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="6" pageNumber="523">Fig. 23</figureCitation>
) narrow, tongue-shaped, about twice as long as wide, apex narrowly rounded. Parameres (
<figureCitation id="C0BE2A2B1F7EFF9877AB0BF9FF3DA8E4" box="[159,239,2005,2028]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="6" pageNumber="523">Fig. 24</figureCitation>
) symmetrical, very long and slender, curved, in resting position surpassing apex of proctiger.
</paragraph>
</subSubSection>
<subSubSection id="109F65251F7EFF9974080899FDB8AE3D" lastPageId="7" lastPageNumber="524" pageId="6" pageNumber="523" type="description">
<paragraph id="583A36AE1F7EFF9874080899FAF0A924" blockId="6.[828,1437,1205,2060]" pageId="6" pageNumber="523">
<emphasis id="6AF1EABC1F7EFF9874080899FB71ABC4" bold="true" box="[828,1187,1205,1228]" pageId="6" pageNumber="523">Description of apterous female.</emphasis>
Measurements of
<typeStatus id="873E880C1F7EFF98725F0899FCBCABE4" pageId="6" pageNumber="523" type="allotype">allotype</typeStatus>
: Body length 2.69; maximum body width (at abdominal segment 4) 1.34. Head length 0.50, width 0.68. Synthlipsis 0.39. Pronotum length 0.34, width 0.92. Lengths of antennomeres, I 0.47, II 0.35, III 0.47, IV 0.57. Lengths of leg segments: profemur 0.80, protibia 0.74, protarsus 0.38, mesofemur 0.90, mesotibia 0.88, mesotarsus 0.14 + 0.31, metafemur 1.10, metatibia 1.15, metatarsus 0.16 + 0.31. Measurements of
<typeStatus id="873E880C1F7EFF9874C70999FB85AAC4" box="[1011,1111,1461,1484]" pageId="6" pageNumber="523" type="paratype">paratypes</typeStatus>
(n = 3): Body length 2.532.59; maximum body width (at metapleura) 1.261.29. Head length 0.440.47, width 0.630.66. Synthlipsis 0.350.36. Pronotum length 0.310.33, width 0.890.90.
</paragraph>
<paragraph id="583A36AE1F7EFF98746E0A19FC0FA964" blockId="6.[828,1437,1205,2060]" pageId="6" pageNumber="523">
Colour (
<figureCitation id="C0BE2A2B1F7EFF9874820A19FC28A944" box="[950,1018,1589,1612]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="6" pageNumber="523">Fig. 4</figureCitation>
): Similar to male, in some specimens slightly darker.
</paragraph>
<paragraph id="583A36AE1F7EFF98746E0A59FB10A864" blockId="6.[828,1437,1205,2060]" pageId="6" pageNumber="523">
Pilosity: Similar to male, but short pilosity also present on pronotal disk, and very long, erect setae all over dorsum of head and thorax, mediotergites, laterotergites, and sides of sterna. Long pilosity partially rubbed off in some specimens. Patches of silverish pilosity (
<figureCitation id="C0BE2A2B1F7EFF9873C70AD9FAE7A804" box="[1267,1333,1781,1804]" captionStart="Figs 14" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 apterous male, 2 apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 apterous male, 4 apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="6" pageNumber="523">Fig. 4</figureCitation>
) laterally on mediotergite 1 more extended than in male; in addition, a larger central part of mediotergites 6 and 7 silverish. Gonocoxa 1 only with short pilosity.
</paragraph>
<paragraph id="583A36AE1F7EFF99746E0B59FDB8AE3D" blockId="6.[828,1437,1205,2060]" lastBlockId="7.[151,760,190,309]" lastPageId="7" lastPageNumber="524" pageId="6" pageNumber="523">
Structures: Head and thorax similar as in male. Pronotum without concavity of hind margin. Propleuron without ridge behind eye. Abdomen wider than in male, but posteriorly more narrowed. Mediotergite 7 (
<figureCitation id="C0BE2A2B1F7EFF9873CA0BF9FA9DA8E4" box="[1278,1359,2005,2028]" captionStart="Figs 1624" captionStartId="6.[151,188,1043,1062]" captionTargetBox="[151,1432,190,1020]" captionTargetId="figure-472@6.[151,1436,190,1023]" captionTargetPageId="6" captionText="Figs 1624. Structural details of Eyarinella robusta gen. nov. &amp; sp. nov.: 16 head, first antennomere, and pronotum, dorsal; 17 head and anterior margin of pronotum, lateral; 18 foreleg of male (arrow points at proximal end of grasping comb); 19 metafemur of male; 20 abdominal segments 68, and proctiger of female, dorsal; 21 abdominal segment 8 of male, ventral; 22 pygophore of male, ventral; 23 proctiger of male, dorsal; 24 left paramere, lateral. Pilosity omitted, except in Fig. 24." figureDoi="http://doi.org/10.5281/zenodo.5821223" httpUri="https://zenodo.org/record/5821223/files/figure.png" pageId="6" pageNumber="523">Fig. 20</figureCitation>
) about 2.1 times as long as wide. Mediotergite 8 flat, wider than long, directed posteriorly. Proctiger narrow, posteriorly protruded. Gonocoxa 1 flat, base concealed by sternum 7.
<emphasis id="6AF1EABC1F7FFF9977A30CD2FEC5AE1D" bold="true" box="[151,279,254,277]" pageId="7" pageNumber="524">Etymology.</emphasis>
The species epithet, the Latin adjective
<emphasis id="6AF1EABC1F7FFF9975870CD2FF64AE3D" italics="true" pageId="7" pageNumber="524">robustus</emphasis>
(-
<emphasis id="6AF1EABC1F7FFF9977FB0D32FF0EAE3D" box="[207,220,286,309]" italics="true" pageId="7" pageNumber="524">a</emphasis>
, -
<emphasis id="6AF1EABC1F7FFF9977C60D32FEC1AE3D" box="[242,275,286,309]" italics="true" pageId="7" pageNumber="524">um</emphasis>
), refers to the stout body shape.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,303 @@
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<mods:title id="16623D3CEE6A5CA61335DA0178654853">The taxonomy of some unusual Microveliinae (Hemiptera: Heteroptera: Veliidae) from India</mods:title>
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<subSubSection id="109F65251F7FFF99760F0D72FD86AE7D" box="[315,596,349,373]" pageId="7" pageNumber="524" type="nomenclature">
<paragraph id="583A36AE1F7FFF99760F0D72FD86AE7D" blockId="7.[315,596,349,373]" box="[315,596,349,373]" pageId="7" pageNumber="524">
<heading id="037281C21F7FFF99760F0D72FD86AE7D" box="[315,596,349,373]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="524" reason="3">
<emphasis id="6AF1EABC1F7FFF99760F0D72FD86AE7D" bold="true" box="[315,596,349,373]" pageId="7" pageNumber="524">
<taxonomicName id="9F854D2D1F7FFF99760F0D72FE2AAE7D" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[315,504,349,373]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="524" phylum="Arthropoda" rank="species" species="fikaceki" status="sp. nov.">
<emphasis id="6AF1EABC1F7FFF99760F0D72FE2AAE7D" bold="true" box="[315,504,349,373]" italics="true" pageId="7" pageNumber="524">Geovelia fikaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7FFF9976CA0D72FD86AE7D" box="[510,596,350,373]" pageId="7" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="109F65251F7FFF99764F0D50FDC1AE87" box="[379,531,380,399]" pageId="7" pageNumber="524" type="description">
<paragraph id="583A36AE1F7FFF99764F0D50FDC1AE87" blockId="7.[151,761,380,2041]" box="[379,531,380,399]" pageId="7" pageNumber="524">
(
<figureCitation id="C0BE2A2B1F7FFF9976B50D50FE1CAE87" box="[385,462,380,399]" captionStart="Figs 58" captionStartId="4.[151,188,1769,1788]" captionTargetBox="[151,1436,189,1749]" captionTargetId="figure-142@4.[151,1436,189,1749]" captionTargetPageId="4" captionText="Figs 58. Habitus of Geovelia fikaceki sp. nov. (5 apterous male, 6 apterous female), and Neoalardus typicus (Distant, 1903) (7 apterous male, 8 apterous female), both specimens from India." figureDoi="http://doi.org/10.5281/zenodo.5821219" httpUri="https://zenodo.org/record/5821219/files/figure.png" pageId="7" pageNumber="524">Figs 5, 6</figureCitation>
,
<figureCitation id="C0BE2A2B1F7FFF9976E30D50FDDFAE87" box="[471,525,380,399]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="7" pageNumber="524">2531</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="109F65251F7FFF9977A30D8AFDCCAD51" pageId="7" pageNumber="524" type="materials_examined">
<paragraph id="583A36AE1F7FFF9977A30D8AFDCCAD51" blockId="7.[151,761,380,2041]" pageId="7" pageNumber="524">
<emphasis id="6AF1EABC1F7FFF9977A30D8AFEF2AEB1" bold="true" box="[151,288,422,441]" pageId="7" pageNumber="524">Type material:</emphasis>
Hඈඅඈඍඒඉൾ: J (apterous) (
<collectionCode id="3E94AE6B1F7FFF99753F0D8AFD9EAEB1" box="[523,588,422,441]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="7" pageNumber="524">NMPC</collectionCode>
), labelled “
<collectingCountry id="2092763E1F7FFF9975860D8AFD21AEB1" box="[690,755,422,441]" name="India" pageId="7" pageNumber="524">INDIA</collectingCountry>
,
<collectingRegion id="9A41F84C1F7FFF9977A30DEDFEE8AEDC" box="[151,314,449,468]" country="India" name="Arunachal Pradesh" pageId="7" pageNumber="524">Arunachal Pradesh</collectingRegion>
(15) \
<quantity id="9F7D9B4B1F7FFF9976470DEDFE7AAEDC" box="[371,424,449,468]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="524" unit="km" value="11.0">11 km</quantity>
SSE of Tenga, Eagles \ Nest WL sanctuary,
<date id="2C3B106E1F7FFF9977F80DF7FEC7AEE6" box="[204,277,475,494]" pageId="7" pageNumber="524" value="2008-05-04">4.v.2008</date>
\
<geoCoordinate id="3DB150691F7FFF9976120DF7FE59AEE6" box="[294,395,475,494]" degrees="27" direction="north" minutes="06" orientation="latitude" pageId="7" pageNumber="524" precision="15" seconds="23" value="27.10639">27°0623″N</geoCoordinate>
<geoCoordinate id="3DB150691F7FFF9976A40DF7FE21AEE6" box="[400,499,475,494]" degrees="92" direction="east" minutes="26" orientation="longitude" pageId="7" pageNumber="524" precision="15" seconds="02" value="92.43389">92°2602″E</geoCoordinate>
<quantity id="9F7D9B4B1F7FFF9976CD0DF7FDE6AEE6" box="[505,564,475,494]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="7" pageNumber="524" unit="m" value="2510.0">2510m</quantity>
\ Fikáček, Podskalská, Šípek lgt.”, “primary evergreen mountain \ cloud forest with very sparse \ understory vegetation \ sifting: wet leaf litter”. Pൺඋൺඍඒඉൾඌ (including
<typeStatus id="873E880C1F7FFF9977A30E07FF0EAD36" box="[151,220,555,574]" pageId="7" pageNumber="524" type="allotype">allotype</typeStatus>
female;
<collectionCode id="3E94AE6B1F7FFF9976110E07FEB6AD36" box="[293,356,555,574]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="7" pageNumber="524">NMPC</collectionCode>
,
<collectionCode id="3E94AE6B1F7FFF99765F0E07FE63AD36" box="[363,433,555,574]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="524" type="Museum">NHMW</collectionCode>
,
<collectionCode id="3E94AE6B1F7FFF99768D0E07FE29AD36" box="[441,507,555,574]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="7" pageNumber="524">ZMUC</collectionCode>
): 10 JJ
<specimenCount id="4E83FD271F7FFF9975710E07FDAEAD36" box="[581,636,549,574]" pageId="7" pageNumber="524" type="female">11 ♀♀</specimenCount>
(all apterous), same label data (5 JJ
<specimenCount id="4E83FD271F7FFF9976680E6AFE59AD51" box="[348,395,576,601]" pageId="7" pageNumber="524" type="female">4 ♀♀</specimenCount>
in pure ethanol).
</paragraph>
</subSubSection>
<subSubSection id="109F65251F7FFF9977A30E4EFE2AAC91" pageId="7" pageNumber="524" type="diagnosis">
<paragraph id="583A36AE1F7FFF9977A30E4EFE2AAC91" blockId="7.[151,761,380,2041]" pageId="7" pageNumber="524">
<emphasis id="6AF1EABC1F7FFF9977A30E4EFEDEAD71" bold="true" box="[151,268,610,633]" pageId="7" pageNumber="524">Diagnosis.</emphasis>
Body small and stout, length 1.882.04. Head (
<figureCitation id="C0BE2A2B1F7FFF9977AB0EAEFF3FAD91" box="[159,237,642,665]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="7" pageNumber="524">Fig. 25</figureCitation>
) wide, with globular eyes. Pronotum of apterous morph (
<figureCitation id="C0BE2A2B1F7FFF9977DF0E8EFEEAADB1" box="[235,312,674,697]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="7" pageNumber="524">Fig. 25</figureCitation>
) moderately short (width / length ca. 3.4), reaching abdominal mediotergite 1; middle of anterior margin moderately concave; middle of hind margin straight or shallowly concave. Grasping comb of male shorter than subapical tibia width (
<figureCitation id="C0BE2A2B1F7FFF9976B60F0EFE03AC31" box="[386,465,802,825]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="7" pageNumber="524">Fig. 26</figureCitation>
), less than one tenth of tibia length. Apterous morph with shiny spots anteromedially on mediotergites 7 and 8 (in male spot on segment 8 not visible in undissected specimen).
</paragraph>
</subSubSection>
<subSubSection id="109F65251F7FFF9677A30F8EFD13AA95" lastPageId="8" lastPageNumber="525" pageId="7" pageNumber="524" type="description">
<paragraph id="583A36AE1F7FFF9977A30F8EFE99AA71" blockId="7.[151,761,380,2041]" pageId="7" pageNumber="524">
<emphasis id="6AF1EABC1F7FFF9977A30F8EFE0BACB1" bold="true" box="[151,473,930,953]" pageId="7" pageNumber="524">Description of apterous male.</emphasis>
Measurements of
<typeStatus id="873E880C1F7FFF9975A10F8EFD27ACB1" box="[661,757,930,953]" pageId="7" pageNumber="524" type="holotype">holotype</typeStatus>
: Body length 1.88; maximum body width (at laterotergites 3) 0.94. Head length 0.41, width 0.72. Synthlipsis 0.33. Pronotum length 0.22, width 0.74. Lengths of antennomeres, I 0.25, II 0.23, III 0.42, IV 0.49. Lengths of leg segments: profemur 0.62, protibia 0.74, protarsus 0.21, mesofemur 0.66, mesotibia 0.62, mesotarsus 0.09 + 0.19, metafemur 0.72, metatibia 0.83, metatarsus 0.13 + 0.21; protibial comb length 0.07; comb index 0.09. Measurements of
<typeStatus id="873E880C1F7FFF99761F08EEFE43ABD1" box="[299,401,1218,1241]" pageId="7" pageNumber="524" type="paratype">paratypes</typeStatus>
(n = 5): Body length 1.851.93; maximum body width (at laterotergites 3) 0.920.96. Head length 0.390.43, width 0.710.74. Synthlipsis 0.330.36. Pronotum length 0.200.22, width 0.740.78. Protibia length 0.740.76; protibial comb length 0.060.07; comb index 0.080.09.
</paragraph>
<paragraph id="583A36AE1F7FFF99778109AEFD14A951" blockId="7.[151,761,380,2041]" pageId="7" pageNumber="524">
Colour (
<figureCitation id="C0BE2A2B1F7FFF99762409AEFE86AA91" box="[272,340,1410,1433]" captionStart="Figs 58" captionStartId="4.[151,188,1769,1788]" captionTargetBox="[151,1436,189,1749]" captionTargetId="figure-142@4.[151,1436,189,1749]" captionTargetPageId="4" captionText="Figs 58. Habitus of Geovelia fikaceki sp. nov. (5 apterous male, 6 apterous female), and Neoalardus typicus (Distant, 1903) (7 apterous male, 8 apterous female), both specimens from India." figureDoi="http://doi.org/10.5281/zenodo.5821219" httpUri="https://zenodo.org/record/5821219/files/figure.png" pageId="7" pageNumber="524">Fig. 5</figureCitation>
): Ground colour pale brown, ventrally more yellowish brown. Dorsum of head with paired blackish markings. Pronotum with a transverse black stripe in middle of length. Sutural zones between mediotergites 17 black. Sides of thorax with black spots around grooves. Sides of abdomen with elongate blackish stripe. Rostrum yellow with black tip. Antenna and legs pale yellow.
</paragraph>
<paragraph id="583A36AE1F7FFF9977810A4EFEAAA811" blockId="7.[151,761,380,2041]" pageId="7" pageNumber="524">
Pilosity: A sparse pale yellow appressed pilosity inconspicuously developed all over body. Stripes along medial eye margins, and at hind margins of pronotum, mediotergites 16 and laterotergites 26 with variously developed shiny white pilosity (
<figureCitation id="C0BE2A2B1F7FFF9976D00ACEFDF1A9F1" box="[484,547,1762,1785]" captionStart="Figs 58" captionStartId="4.[151,188,1769,1788]" captionTargetBox="[151,1436,189,1749]" captionTargetId="figure-142@4.[151,1436,189,1749]" captionTargetPageId="4" captionText="Figs 58. Habitus of Geovelia fikaceki sp. nov. (5 apterous male, 6 apterous female), and Neoalardus typicus (Distant, 1903) (7 apterous male, 8 apterous female), both specimens from India." figureDoi="http://doi.org/10.5281/zenodo.5821219" httpUri="https://zenodo.org/record/5821219/files/figure.png" pageId="7" pageNumber="524">Fig. 5</figureCitation>
). Erect setae absent, except from antenna.
</paragraph>
<paragraph id="583A36AE1F7FFF9977810B0EFD25A8F1" blockId="7.[151,761,380,2041]" pageId="7" pageNumber="524">
Structures: Head (
<figureCitation id="C0BE2A2B1F7FFF9976410B0EFE11A831" box="[373,451,1826,1849]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="7" pageNumber="524">Fig. 25</figureCitation>
) including eyes almost twice as wide as long, with large globular eyes; apex of head rectangularly protruded. Posterior of head only slightly protruded into a shallow concavity of pronotum. Antenna long; antennomere I short, but surpassing apex of head by more than half of its length (
<figureCitation id="C0BE2A2B1F7FFF9975360BEEFD84A8D1" box="[514,598,1986,2009]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="7" pageNumber="524">Fig. 25</figureCitation>
); antennomere II slightly shorter and more slender than antennomere
</paragraph>
<paragraph id="583A36AE1F7FFF9974080C92FAE0AC1D" blockId="7.[828,1437,190,2037]" pageId="7" pageNumber="524">
I, but clearly thicker than filiform antennomeres III and IV; antennomere IV longest. Rostrum reaching middle of mesosternum. Pronotum (
<figureCitation id="C0BE2A2B1F7FFF9973650CD2FB72AE1D" box="[1105,1184,254,277]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="7" pageNumber="524">Fig. 25</figureCitation>
) moderately short, medially touching mediotergite 1; anterior margin shallowly concave in middle; posterior margin laterally convex, medially straight to very shallowly concave; at midlength with a deep transverse groove beset with numerous sensory pores; another transverse row of pores just behind anterior margin. Legs relatively short; femora moderately stout. Protibial grasping comb very short, occupying less than one tenth of tibia length (
<figureCitation id="C0BE2A2B1F7FFF9973700DD2FB41AD1D" box="[1092,1171,510,533]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="7" pageNumber="524">Fig. 26</figureCitation>
). All mediotergites much wider than long, mediotergite 7 more than twice as long as 6, anteromedially with a large shiny area. Sutural zones between mediotergites 17 with transverse rows of pores. Laterotergites evenly sloping. All sterna simple, convex. Segment 8 (
<figureCitation id="C0BE2A2B1F7FFF9974F70EB2FBC6ADBD" box="[963,1044,670,693]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="7" pageNumber="524">Fig. 28</figureCitation>
) about one-and-a-half times longer than wide, dorsally shiny. Genitalia very small. Pygophore unmodified (
<figureCitation id="C0BE2A2B1F7FFF99748B0EF2FBDEADFD" box="[959,1036,734,757]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="7" pageNumber="524">Fig. 29</figureCitation>
). Proctiger with pointed apex (
<figureCitation id="C0BE2A2B1F7FFF9972700EF2FA43ADFD" box="[1348,1425,734,757]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="7" pageNumber="524">Fig. 30</figureCitation>
). Paramere very slender, sickle-shaped (
<figureCitation id="C0BE2A2B1F7FFF9973E20ED2FAF7AC1D" box="[1238,1317,766,789]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="7" pageNumber="524">Fig. 31</figureCitation>
).
</paragraph>
<paragraph id="583A36AE1F7FFF9974080F32FC7CABBD" blockId="7.[828,1437,190,2037]" pageId="7" pageNumber="524">
<emphasis id="6AF1EABC1F7FFF9974080F32FB15AC3D" bold="true" box="[828,1223,798,821]" pageId="7" pageNumber="524">Description of apterous female.</emphasis>
Measurements of
<typeStatus id="873E880C1F7FFF9974080F12FC4BAC5D" box="[828,921,830,853]" pageId="7" pageNumber="524" type="allotype">allotype</typeStatus>
: Body length 1.90; maximum body width (at laterotergites 3) 0.98. Head length 0.42, width 0.73. Synthlipsis 0.35. Pronotum length 0.20, width 0.76. Lengths of antennomeres, I 0.25, II 0.23, III 0.36, IV 0.47. Lengths of leg segments: profemur 0.55, protibia 0.50, protarsus 0.21, mesofemur 0.62, mesotibia 0.61, mesotarsus 0.09 + 0.15, metafemur 0.77, metatibia 0.83, metatarsus 0.12 + 0.21. Measurements of
<typeStatus id="873E880C1F7FFF9973640832FB64AB3D" box="[1104,1206,1054,1077]" pageId="7" pageNumber="524" type="paratype">paratypes</typeStatus>
(n = 6): Body length 1.982.04; maximum body width (at laterotergites 3 or 4) 1.021.06. Head length 0.410.43, width 0.740.78. Synthlipsis 0.340.38. Pronotum length 0.200.22, width 0.730.78.
</paragraph>
<paragraph id="583A36AE1F7FFF99746E0892FBE2AA9D" blockId="7.[828,1437,190,2037]" pageId="7" pageNumber="524">
Colour (
<figureCitation id="C0BE2A2B1F7FFF9974820892FC28ABDD" box="[950,1018,1214,1237]" captionStart="Figs 58" captionStartId="4.[151,188,1769,1788]" captionTargetBox="[151,1436,189,1749]" captionTargetId="figure-142@4.[151,1436,189,1749]" captionTargetPageId="4" captionText="Figs 58. Habitus of Geovelia fikaceki sp. nov. (5 apterous male, 6 apterous female), and Neoalardus typicus (Distant, 1903) (7 apterous male, 8 apterous female), both specimens from India." figureDoi="http://doi.org/10.5281/zenodo.5821219" httpUri="https://zenodo.org/record/5821219/files/figure.png" pageId="7" pageNumber="524">Fig. 6</figureCitation>
), pilosity, and structures almost same as in male. Body slightly broader. Abdomen unmodified. Mediotergite 7 about one third longer than 6 and 8, respectively. Mediotergite 8 directed straight caudad (
<figureCitation id="C0BE2A2B1F7FFF99720B0932FA42AA3D" box="[1343,1424,1310,1333]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="7" pageNumber="524">Fig. 27</figureCitation>
), with moderately convex hind margin. Gonocoxa plate-like, unmodified, not concealed by sternum 7. Proctiger with pointed apex (
<figureCitation id="C0BE2A2B1F7FFF9974E70952FBF1AA9D" box="[979,1059,1406,1429]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="7" pageNumber="524">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="583A36AE1F7FFF99740809B2FBF2A83D" blockId="7.[828,1437,190,2037]" pageId="7" pageNumber="524">
<emphasis id="6AF1EABC1F7FFF99740809B2FACBAABD" bold="true" box="[828,1305,1438,1461]" pageId="7" pageNumber="524">
Notes on
<taxonomicName id="9F854D2D1F7FFF99748609B2FAC6AABD" authority="Zimmermann, 1984" authorityName="Zimmermann" authorityYear="1984" box="[946,1300,1438,1461]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F7FFF99748609B2FBC5AABD" bold="true" box="[946,1047,1438,1461]" italics="true" pageId="7" pageNumber="524">Geovelia</emphasis>
<bibRefCitation id="3C144B5F1F7FFF99731609B2FAC6AABD" author="ZIMMERMANN G." box="[1058,1300,1438,1461]" pageId="7" pageNumber="524" pagination="65 - 74" refId="ref9005" refString="ZIMMERMANN G. 1984: Heteroptera aus dem Nepal-Himalaya. Geovelia. n. gen., eine Gattung terrestrischer Microveliinae (Insecta: Gerromorpha: Veliidae). Senckenbergiana Biologica 65 (1 - 2): 65 - 74." type="journal article" year="1984">Zimmermann, 1984</bibRefCitation>
</taxonomicName>
.
</emphasis>
Species of
<taxonomicName id="9F854D2D1F7FFF9974080992FC4EAADD" authorityName="Zimmermann" authorityYear="1984" box="[828,924,1470,1493]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F7FFF9974080992FC4EAADD" box="[828,924,1470,1493]" italics="true" pageId="7" pageNumber="524">Geovelia</emphasis>
</taxonomicName>
inhabit moist leaf litter in forests (Zආආൾඋආൺඇඇ 1984, 2014; Zൾඍඍൾඅ 2011). They were described from
<collectingCountry id="2092763E1F7FFF99740809D2FCAFA91D" box="[828,893,1534,1557]" name="Nepal" pageId="7" pageNumber="524">Nepal</collectingCountry>
(five species, all known only from the apterous morph; Zආආൾඋආൺඇඇ 1984, 2014) and
<collectingCountry id="2092763E1F7FFF9973E40A32FAE5A93D" box="[1232,1335,1566,1589]" name="Myanmar" pageId="7" pageNumber="524">Myanmar</collectingCountry>
(one species, known only in macropterous morph; Zൾඍඍൾඅ 2011).
<taxonomicName id="9F854D2D1F7FFF9974080A72FB27A97D" authority="Ye, Chen &amp; Bu, 2014" authorityName="Ye, Chen &amp; Bu" authorityYear="2014" box="[828,1269,1630,1653]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="524" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="6AF1EABC1F7FFF9974080A72FC27A97D" box="[828,1013,1630,1653]" italics="true" pageId="7" pageNumber="524">Geovelia hirsuta</emphasis>
<bibRefCitation id="3C144B5F1F7FFF9974CA0A72FB27A97D" author="YE ZH &amp; CHEN P. &amp; BU W." box="[1022,1269,1630,1653]" pageId="7" pageNumber="524" pagination="16 - 22" refId="ref8744" refString="YE ZH., CHEN P. &amp; BU W. 2014 a: First record of the genus Geovelia Zimmermann 1984 (Hemiptera: Heteroptera: Veliidae) from China, with description of a new species. Pan-Pacific Entomologist 90: 16 - 22." type="journal article" year="2014">Ye, Chen &amp; Bu, 2014</bibRefCitation>
</taxonomicName>
was described from Southwestern
<collectingCountry id="2092763E1F7FFF99733A0A52FB9CA99D" box="[1038,1102,1662,1685]" name="China" pageId="7" pageNumber="524">China</collectingCountry>
(Yൾ et al. 2014a), but has been transferred to a new genus (Zൾඍඍൾඅ et al. 2021).
<taxonomicName id="9F854D2D1F7FFF9972080AB2FC5FA9DD" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="524" phylum="Arthropoda" rank="species" species="fikaceki" status="sp. nov.">
<emphasis id="6AF1EABC1F7FFF9972080AB2FC5FA9DD" italics="true" pageId="7" pageNumber="524">Geovelia fikaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F7FFF9974A60A92FC31A9DD" box="[914,995,1726,1749]" pageId="7" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
is the first record of this genus from
<collectingCountry id="2092763E1F7FFF99726B0A92FA4BA9DD" box="[1375,1433,1726,1749]" name="India" pageId="7" pageNumber="524">India</collectingCountry>
. The altitudinal range of species extends from
<quantity id="9F7D9B4B1F7FFF9972150AF2FAB7A9FD" box="[1313,1381,1758,1781]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="524" unit="m" value="350.0">350 m</quantity>
a.s.l. (
<taxonomicName id="9F854D2D1F7FFF9974710AD2FB84A81D" authority="Zettel, 2011" authorityName="Zettel" authorityYear="2011" box="[837,1110,1790,1813]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="524" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="6AF1EABC1F7FFF9974710AD2FC1EA81D" box="[837,972,1790,1813]" italics="true" pageId="7" pageNumber="524">G. orientalis</emphasis>
<bibRefCitation id="3C144B5F1F7FFF9974E00AD2FB84A81D" author="ZETTEL H." box="[980,1110,1790,1813]" pageId="7" pageNumber="524" pagination="89 - 114" refId="ref8916" refString="ZETTEL H. 2011: A contribution to the knowledge of Gerromorpha (Insecta: Hemiptera) of Myanmar, with seven new species, eight new records, and a catalogue. Annalen des Naturhistorischen Museums Wien, Series B 112: 89 - 114." type="journal article" year="2011">Zettel, 2011</bibRefCitation>
</taxonomicName>
) to
<quantity id="9F7D9B4B1F7FFF9973B60AD2FB06A81D" box="[1154,1236,1790,1813]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="524" unit="m" value="3200.0">3200 m</quantity>
a.s.l. (
<taxonomicName id="9F854D2D1F7FFF9972290AD2FBC6A83D" authority="Zimmermann, 1984" authorityName="Zimmermann" authorityYear="1984" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="524" phylum="Arthropoda" rank="species" species="martensi">
<emphasis id="6AF1EABC1F7FFF9972290AD2FA4EA81D" box="[1309,1436,1790,1813]" italics="true" pageId="7" pageNumber="524">G. martensi</emphasis>
<bibRefCitation id="3C144B5F1F7FFF9974080B32FBC6A83D" author="ZIMMERMANN G." box="[828,1044,1822,1845]" pageId="7" pageNumber="524" pagination="65 - 74" refId="ref9005" refString="ZIMMERMANN G. 1984: Heteroptera aus dem Nepal-Himalaya. Geovelia. n. gen., eine Gattung terrestrischer Microveliinae (Insecta: Gerromorpha: Veliidae). Senckenbergiana Biologica 65 (1 - 2): 65 - 74." type="journal article" year="1984">Zimmermann, 1984</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="583A36AE1F7FFF99746E0B12FB13A8FD" blockId="7.[828,1437,190,2037]" pageId="7" pageNumber="524">
Species of
<taxonomicName id="9F854D2D1F7FFF9974E20B12FBE5A85D" authorityName="Zimmermann" authorityYear="1984" box="[982,1079,1854,1877]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F7FFF9974E20B12FBE5A85D" box="[982,1079,1854,1877]" italics="true" pageId="7" pageNumber="524">Geovelia</emphasis>
</taxonomicName>
can be recognized primarily by the subterminal claws that clearly surpass the tarsal apex; this character is not found in other Oriental
<taxonomicName id="9F854D2D1F7FFF9973C90B52FA85A89D" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[1277,1367,1918,1941]" class="Insecta" family="Veliidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="524" phylum="Arthropoda" rank="family">Veliidae</taxonomicName>
. Other important characters are a very small and stout body, a short head with very large eyes and long antennae, and a short protibial grasping comb of the male.
</paragraph>
<caption id="0CFA66261F70FF9677A30F26FBA7AC5A" ID-DOI="http://doi.org/10.5281/zenodo.5821225" ID-Zenodo-Dep="5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="8" pageNumber="525" startId="8.[151,188,778,797]" targetBox="[152,1431,189,754]" targetPageId="8">
<paragraph id="583A36AE1F70FF9677A30F26FBA7AC5A" blockId="8.[151,1436,778,850]" pageId="8" pageNumber="525">
Figs 2531. Structural details of
<taxonomicName id="9F854D2D1F70FF9676860F26FD94AC15" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[434,582,778,797]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="fikaceki" status="sp. nov.">
<emphasis id="6AF1EABC1F70FF9676860F26FD94AC15" box="[434,582,778,797]" italics="true" pageId="8" pageNumber="525">Geovelia fikaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F70FF9675780F26FD41AC15" box="[588,659,778,797]" pageId="8" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31.
</paragraph>
</caption>
<paragraph id="583A36AE1F70FF9677A30F8AFD13AA95" blockId="8.[151,760,934,1661]" pageId="8" pageNumber="525">
<emphasis id="6AF1EABC1F70FF9677A30F8AFEBDACB5" bold="true" box="[151,367,934,957]" pageId="8" pageNumber="525">Comparative notes.</emphasis>
<taxonomicName id="9F854D2D1F70FF9676400F8AFDF4ACB5" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[372,550,934,957]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="fikaceki" status="sp. nov.">
<emphasis id="6AF1EABC1F70FF9676400F8AFDF4ACB5" box="[372,550,934,957]" italics="true" pageId="8" pageNumber="525">Geovelia fikaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F70FF96751F0F8AFDA8ACB5" box="[555,634,934,957]" pageId="8" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
differs from most species (except
<taxonomicName id="9F854D2D1F70FF9676450FEAFD7EACD5" authority="Zimmermann, 2014" authorityName="Zimmermann" authorityYear="2014" box="[369,684,966,989]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="riegeri">
<emphasis id="6AF1EABC1F70FF9676450FEAFE05ACD5" box="[369,471,966,989]" italics="true" pageId="8" pageNumber="525">G. riegeri</emphasis>
<bibRefCitation id="3C144B5F1F70FF9676E80FEAFD7EACD5" author="ZIMMERMANN G." box="[476,684,966,989]" pageId="8" pageNumber="525" pagination="251 - 256" refId="ref9047" refString="ZIMMERMANN G. 2014: Neue Arten der Gattung Geovelia (Insecta, Gerromorpha, Veliidae) aus Nepal. Andrias 20: 251 - 256." type="journal article" year="2014">Zimmermann, 2014</bibRefCitation>
</taxonomicName>
) by the protibial grasping comb being shorter than the subapical protibia width (
<figureCitation id="C0BE2A2B1F70FF967677082AFE44AB15" box="[323,406,1030,1053]" captionStart="Figs 2531" captionStartId="8.[151,188,778,797]" captionTargetBox="[152,1431,189,754]" captionTargetId="figure-698@8.[151,1436,189,756]" captionTargetPageId="8" captionText="Figs 2531. Structural details of Geovelia fikaceki sp. nov.: 25 head, first antennomere, and pronotum, dorsal; 26 foreleg of male (arrow points at proximal end of grasping comb); 27 abdominal segments 78, and proctiger of female, dorsal; 28 abdominal segment 8 of male, ventral; 29 pygophore of male, ventral; 30 proctiger of male, dorsal; 31 left paramere, lateral. Pilosity omitted, except in Fig. 31." figureDoi="http://doi.org/10.5281/zenodo.5821225" httpUri="https://zenodo.org/record/5821225/files/figure.png" pageId="8" pageNumber="525">Fig. 26</figureCitation>
).
<taxonomicName id="9F854D2D1F70FF967698082AFDBAAB15" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[428,616,1030,1053]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="fikaceki" status="sp. nov.">
<emphasis id="6AF1EABC1F70FF967698082AFDBAAB15" box="[428,616,1030,1053]" italics="true" pageId="8" pageNumber="525">Geovelia fikaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F70FF967546082AFD1AAB15" box="[626,712,1030,1053]" pageId="8" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="9F854D2D1F70FF9677A3080AFED0AB35" authorityName="Zimmermann" authorityYear="2014" box="[151,258,1062,1085]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="riegeri">
<emphasis id="6AF1EABC1F70FF9677A3080AFED0AB35" box="[151,258,1062,1085]" italics="true" pageId="8" pageNumber="525">G. riegeri</emphasis>
</taxonomicName>
are very similar in appearance by their stocky body, wide head, and globular eyes (these characters are also present in
<taxonomicName id="9F854D2D1F70FF967602084AFD56AB75" authority="Zimmermann, 1984" authorityName="Zimmermann" authorityYear="1984" box="[310,644,1126,1149]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="ilamica">
<emphasis id="6AF1EABC1F70FF967602084AFE75AB75" box="[310,423,1126,1149]" italics="true" pageId="8" pageNumber="525">G. ilamica</emphasis>
<bibRefCitation id="3C144B5F1F70FF967699084AFD56AB75" author="ZIMMERMANN G." box="[429,644,1126,1149]" pageId="8" pageNumber="525" pagination="65 - 74" refId="ref9005" refString="ZIMMERMANN G. 1984: Heteroptera aus dem Nepal-Himalaya. Geovelia. n. gen., eine Gattung terrestrischer Microveliinae (Insecta: Gerromorpha: Veliidae). Senckenbergiana Biologica 65 (1 - 2): 65 - 74." type="journal article" year="1984">Zimmermann, 1984</bibRefCitation>
</taxonomicName>
), but differ strongly in the length of the pronotum: In
<taxonomicName id="9F854D2D1F70FF96756B08AAFD00AB95" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[607,722,1158,1181]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="fikaceki" status="sp. nov.">
<emphasis id="6AF1EABC1F70FF96756B08AAFD00AB95" box="[607,722,1158,1181]" italics="true" pageId="8" pageNumber="525">G. fikaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F70FF9675ED08AAFF17ABB5" pageId="8" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
the pronotum is longer (pronotum width: pronotum length = 3.4) and reaches mediotergite
<quantity id="9F7D9B4B1F70FF96751708EAFD9BABD5" box="[547,585,1222,1245]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="525" unit="in" value="1.0">1 in</quantity>
middle, whereas in
<taxonomicName id="9F854D2D1F70FF96778108CAFEF3ABF5" authorityName="Zimmermann" authorityYear="2014" box="[181,289,1254,1277]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="riegeri">
<emphasis id="6AF1EABC1F70FF96778108CAFEF3ABF5" box="[181,289,1254,1277]" italics="true" pageId="8" pageNumber="525">G. riegeri</emphasis>
</taxonomicName>
the pronotum is shorter (pronotum width: pronotum length = 3.6) and leaves the metanotum uncovered in middle (compare measurements and drawings in Zආආൾඋආൺඇඇ 2014). The females of
<taxonomicName id="9F854D2D1F70FF967528096AFD01AA55" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[540,723,1350,1373]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="fikaceki" status="sp. nov.">
<emphasis id="6AF1EABC1F70FF967528096AFD01AA55" box="[540,723,1350,1373]" italics="true" pageId="8" pageNumber="525">Geovelia fikaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F70FF9675ED096AFF17AA75" pageId="8" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
are smaller than those of
<taxonomicName id="9F854D2D1F70FF9676D3094AFD86AA75" authorityName="Zimmermann" authorityYear="2014" box="[487,596,1382,1405]" class="Insecta" family="Veliidae" genus="Geovelia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="riegeri">
<emphasis id="6AF1EABC1F70FF9676D3094AFD86AA75" box="[487,596,1382,1405]" italics="true" pageId="8" pageNumber="525">G. riegeri</emphasis>
</taxonomicName>
(1.902.04 vs.
<quantity id="9F7D9B4B1F70FF9677A309AAFEE7AA95" box="[151,309,1414,1437]" metricMagnitude="-3" metricUnit="m" metricValue="2.225" metricValueMax="2.25" metricValueMin="2.2" pageId="8" pageNumber="525" unit="mm" value="2.225" valueMax="2.25" valueMin="2.2">2.202.25 mm</quantity>
), whereas the size of males overlaps.
</paragraph>
</subSubSection>
<subSubSection id="109F65251F70FF9677A3098AFD30A975" pageId="8" pageNumber="525" type="etymology">
<paragraph id="583A36AE1F70FF9677A3098AFE2CAAF5" blockId="8.[151,760,934,1661]" pageId="8" pageNumber="525">
<emphasis id="6AF1EABC1F70FF9677A3098AFEC6AAB5" bold="true" box="[151,276,1446,1469]" pageId="8" pageNumber="525">Etymology.</emphasis>
We dedicate this species to Dr. Martin Fikáček, coleopterist at the National Museum,
<collectingRegion id="9A41F84C1F70FF96751909EAFDA8AAD5" box="[557,634,1478,1501]" country="Czech Republic" name="Praha" pageId="8" pageNumber="525">Prague</collectingRegion>
, and one of the collectors of this new species.
</paragraph>
<paragraph id="583A36AE1F70FF9677A30A2AFD30A975" blockId="8.[151,760,934,1661]" pageId="8" pageNumber="525">
<emphasis id="6AF1EABC1F70FF9677A30A2AFE6AA915" bold="true" box="[151,440,1542,1565]" pageId="8" pageNumber="525">Collecting circumstances.</emphasis>
All examined specimens were collected by sifting accumulations of leaf litter in primary evergreen cloud forest at ca.
<quantity id="9F7D9B4B1F70FF9676F00A6AFDC1A955" box="[452,531,1606,1629]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="525" unit="m" value="2500.0">2500 m</quantity>
a.s.l., on the southern slope of the Himalaya Mts. (M. Fikáček, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,189 @@
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<paragraph id="2A2AB7D66FA0339380167D1CFC9E86AE" pageNumber="645">
<taxonomicName id="39F4278E717EB550D39205C5B4FE6311" ID-CoL="Q9XD" authority="Linnaeus 1758" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" pageNumber="645" phylum="Chordata" rank="species" species="dromedarius">
<emphasis id="5639D6807CA8733C7F336FBD3639675E" italics="true" pageNumber="645">Camelus dromedarius</emphasis>
Linnaeus 1758
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="7FE7C44610E2E65C31695591331A26EC" pageNumber="645" type="reference_group">
<paragraph id="1A4D050B1C96522A68A6633678D56CA4" pageNumber="645">
<treatmentCitation id="56C224FF59400CF5D0262D6CACAC1360" ID-DOI="http://doi.org/10.5281/zenodo.3921045" ID-Zenodo-Dep="3921045" LSID="urn:lsid:zoobank.org:act:39FF5CE5-F340-42AD-B9AA-FF84032A6E55" author="Linnaeus" authority="Linnaeus 1758" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Camelidae" genus="Camelus" httpUri="http://treatment.plazi.org/id/F6AF9490E2A640F29A2F854F7F9DED61" kingdom="Animalia" order="Artiodactyla" page="65" pageNumber="645" phylum="Chordata" rank="species" species="dromedarius" title="Camelus dromedarius" volumeTitle="Syst. Nat., 10th ed., Vol. 1" year="1758">
<taxonomicName id="C515E73FCFFF4DB45A29657B9A15BE4C" ID-CoL="Q9XD" authority="Linnaeus 1758" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" pageNumber="645" phylum="Chordata" rank="species" species="dromedarius">
<emphasis id="50E30C2C1E7195873D235C49D23DB825" italics="true" pageNumber="645">Camelus dromedarius</emphasis>
<bibRefCitation id="92574AD8905F65134CBEA7D15C948CFD" author="Linnaeus, C." refId="ref213949" refString="Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classis, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tenth ed. Vol. 1. Laurentii Salvii, Stockholm, 824 pp." year="1758">Linnaeus 1758</bibRefCitation>
</taxonomicName>
,
<bibCitation id="240329EA4967D095D16AA29C9131DE04" author="Linnaeus" pageNumber="645" pagination="65" title="Camelus dromedarius" volumeTitle="Syst. Nat., 10th ed., Vol. 1" year="1758">Syst. Nat., 10th ed., Vol. 1: 65</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="ED41AAEAEAD0C48B8256E03303662805" pageNumber="645" type="materials_examined">
<paragraph id="64A2593B167D453B9A7ED7E20629036D" pageNumber="645">
<emphasis id="7064240E539E75D2EEA91EA9F2883E5F" inLineHeading="true" pageNumber="645">Type Locality:</emphasis>
<materialsCitation id="05545E74CB22E617AF6DE6C761180CC4" ID-GBIF-Occurrence="4867329323" country="Libya" pageNumber="645">
&quot;Habitat in Africae desertis arenosis siticulosis&quot;, identified as &quot;deserts of
<collectingCountry id="A3C5BF01D84FE48C454F9AAA333E7CF0" name="Libya">Libya</collectingCountry>
and Arabia&quot; by
<bibRefCitation id="AFA0378D8F7DAE4F7F5C2D80E6EE01FF" author="Thomas, O." refId="ref353417" refString="Thomas, O. 1911 a. The mammals of the tenth edition of Linnaeus; an attempt to fix the types of the genera and the exact bases and localities of the species. Proceedings of the Zoological Society of London, 1911: 120 - 158." year="1911">
Thomas (1911
<emphasis id="559C857E21E5DF43E35C583FDB133A31" italics="true" pageNumber="645">a</emphasis>
:150)
</bibRefCitation>
; based on domesticated stock
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E355CDA0714F3B69E23A8B4433945946" pageNumber="645" type="vernacular_names">
<paragraph id="A46D0A83C34BC7B305EF807219F47A7E" pageNumber="645">
<emphasis id="C5B9FFBE70BB3EE1EA1C61C5D4819575" inLineHeading="true" pageNumber="645">Vernacular Names:</emphasis>
<vernacularName id="C342CC05C3B95ECEFF92EEAAF9235906" language="eng" pageNumber="645">One-humped Camel</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="7C1EEBD9496278F3ED1334AA679587DB" pageNumber="645" type="synonym_list">
<paragraph id="A06843C350F3231833169D79BF58604C" pageNumber="645">
<emphasis id="582BBD9A306B5AC1CC7AEA7D097CCF9B" inLineHeading="true" pageNumber="645">Synonyms:</emphasis>
<taxonomicName id="F02CD7F443B1A46CC7A26B3D5E3539BC" authority="Kolenati 1847" authorityName="Kolenati" authorityYear="1847" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" pageNumber="645" phylum="Chordata" rank="species" species="aegyptiacus">
<emphasis id="3ACD6E48F06B2192FD9BA9D691A5B1DE" italics="true" pageNumber="645">Camelus aegyptiacus</emphasis>
Kolenati 1847
</taxonomicName>
;
<taxonomicName id="F495BB53A2C22A169275831620E53383" authority="(Gloger 1841)" baseAuthorityName="Gloger" baseAuthorityYear="1841" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" pageNumber="645" phylum="Chordata" rank="species" species="africanus">
<emphasis id="AE39E542AE17D0E5F75107FD9AAE5309" italics="true" pageNumber="645">Camelus africanus</emphasis>
(Gloger 1841)
</taxonomicName>
;
<taxonomicName id="B626317B6D78FBBE57C326BEF998C622" authority="Desmoulins 1823" authorityName="Desmoulins" authorityYear="1823" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" pageNumber="645" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="C5B164E8725410147DA25736C276F8D8" italics="true" pageNumber="645">Camelus arabicus</emphasis>
Desmoulins 1823
</taxonomicName>
;
<taxonomicName id="DD7144840BCBD9AB9DAD926AF06CECBB" authority="Pallas 1811" authorityName="Pallas" authorityYear="1811" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" pageNumber="645" phylum="Chordata" rank="species" species="dromas">
<emphasis id="BC1448FF9AADF88F546769C652D53238" italics="true" pageNumber="645">Camelus dromas</emphasis>
<bibRefCitation id="0738E8715EE39EA6EDF9CB58ADFFAB3D" author="Pallas, P. S." refId="ref271664" refString="Pallas, P. S. 1811 [1831]. Zoographia Rosso-Asiatica, sistens omnium Animalium in extenso Imperio Rossico et adjacentibus maribus observatorum recensionem, domicillia, mores et descriptiones, anatomen atque icones plurimorum. [ICZN Opinion 212 -- dates of volumes: 1 &amp; 2: 1811; 3: 1814]. Petropoli, in officina Caes. acadamiae scientiarum. 3 vol. [official date of publ for vol 1 - 1811 impress], 1: 1 - 568." year="1811">Pallas 1811</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="AA3A681C04DE50A2511BB66AD06BEAC0" authority="Kerr 1792" authorityName="Kerr" authorityYear="1792" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" pageNumber="645" phylum="Chordata" rank="species" species="dromos">
<emphasis id="8A265B7D912821A021ED5EADC3F40209" italics="true" pageNumber="645">Camelus dromos</emphasis>
<bibRefCitation id="2829FACC93B3162CA29D9BEB1BE9BB52" author="Kerr, R." refId="ref183257" refString="Kerr, R. 1792. The animal kingdom, or zoological system, of the celebrated Sir Charles Linnaeus; class I: Mammalia. London, J. Murray &amp; R. Faulder, 664 pp." year="1792">Kerr 1792</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="D7903C80A12831D61F8D3D7230465CE3" authority="Falk 1786" authorityName="Falk" authorityYear="1786" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" pageNumber="645" phylum="Chordata" rank="species" species="ferus">
<emphasis id="EDA844D76C8976D38AAB4D11C94C69A4" italics="true" pageNumber="645">Camelus ferus</emphasis>
Falk 1786
</taxonomicName>
;
<taxonomicName id="CB805F2CE27602FF7209EE7BECD97A65" authority="Kolenati 1847" authorityName="Kolenati" authorityYear="1847" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" pageNumber="645" phylum="Chordata" rank="species" species="lukius">
<emphasis id="9A29DCB959EFA5E5CEE6E35EDA48F03F" italics="true" pageNumber="645">Camelus lukius</emphasis>
Kolenati 1847
</taxonomicName>
;
<taxonomicName id="F45CB6F03F250557BA31C8AAB6E71720" authority="Kolenati 1847" authorityName="Kolenati" authorityYear="1847" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" pageNumber="645" phylum="Chordata" rank="species" species="polytrichus">
<emphasis id="8CA86035F0D8314E736C4F1B678A0422" italics="true" pageNumber="645">Camelus polytrichus</emphasis>
Kolenati 1847
</taxonomicName>
;
<taxonomicName id="A5E9F459183ACF73F715F6A243E1E7D8" authority="J. Fischer 1829" authorityName="J. Fischer" authorityYear="1829" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" pageNumber="645" phylum="Chordata" rank="species" species="turcomanicus">
<emphasis id="BAEAD26AD28E745DBE4A02742DD952B0" italics="true" pageNumber="645">Camelus turcomanicus</emphasis>
J. Fischer 1829
</taxonomicName>
;
<taxonomicName id="011A728F74393B2DD3641E2A27B263C1" authority="Kolenati 1847" authorityName="Kolenati" authorityYear="1847" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" pageNumber="645" phylum="Chordata" rank="species" species="vulgaris">
<emphasis id="97A40C9435F425E410882C7257A87441" italics="true" pageNumber="645">Camelus vulgaris</emphasis>
Kolenati 1847
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="900C6800F5882FED5F297F28D2CB0BDC" pageNumber="645" type="distribution">
<paragraph id="245C42A040B1268065E4F8F05E4E88D5" pageNumber="645">
<emphasis id="1EC5E9CBD8A8B505B9A13ADE93A7F2C6" inLineHeading="true" pageNumber="645">Distribution:</emphasis>
Extinct in the wild; domesticated from wild populations which presumably had become restricted to the S Arabian Peninsula; domesticated in
<collectingCountry id="02479F1C24EA8E1A0B6296A174D81083" name="Senegal">Senegal</collectingCountry>
and
<collectingCountry id="791ADD0D113C401CAB9DA2CAA1666E96" name="Mauritania">Mauritania</collectingCountry>
to
<collectingCountry id="6B03AE7AB4C4C4C9990EC93EBCEB499E" name="Somalia">Somalia</collectingCountry>
and
<collectingCountry id="0B0DEB02BE56D22F0BF9AE50E8767FCB" name="Kenya">Kenya</collectingCountry>
, throughout N Africa, the Middle East, Arabia, and
<collectingCountry id="94F3A9AD2CFAD32EC139964BF5966128" name="Iran">Iran</collectingCountry>
to NW
<collectingCountry id="AA52EC9561FBF1FD3E830D670A8D7298" name="India">India</collectingCountry>
; feral populations in
<collectingCountry id="D60796BD4ACF1B1C201D02A21E019A2A" name="Australia">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="8A1801254ECDA772839EED3BDBDCA29A" pageNumber="645" type="discussion">
<paragraph id="3C190B1C7B2A720E4D4265FBF01733ED" pageNumber="645">
<emphasis id="71CAE9DC89A5F7069F37F66EBE921B43" inLineHeading="true" pageNumber="645">Discussion:</emphasis>
Produces viable hybrids with
<taxonomicName id="F75332F2D5BE029005DE7A693D7E7FE1" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" phylum="Chordata" rank="species" species="bactrianus">
<emphasis id="335DD407CAC9B20FB075A3E88CEA12D8" italics="true" pageNumber="645">bactrianus</emphasis>
</taxonomicName>
(see comments therein).
<bibRefCitation id="B41D03A3868D7ADC03D864E33DE2A659" author="Bohlken, H." refId="ref34927" refString="Bohlken, H. 1961. Haustier und zoologische Systematik. Zeitschrift fur Tierzuchtung und Zuchtungbiologie, 75: 107 - 113." year="1961">Bohlken (1961)</bibRefCitation>
considered
<taxonomicName id="0BBDF9F3D659EEB9127E90EE10696A93" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" phylum="Chordata" rank="species" species="dromedarius">
<emphasis id="9582B3816F497F948642FE7C8F043135" italics="true" pageNumber="645">dromedarius</emphasis>
</taxonomicName>
a synonym of
<taxonomicName id="6DA1B9132407F39BB39212783CD55942" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" phylum="Chordata" rank="species" species="bactrianus">
<emphasis id="163887F95797AA2352466000102105D5" italics="true" pageNumber="645">bactrianus</emphasis>
</taxonomicName>
. Reviewed by Köhler-Rollefson (1991, Mammalian Species, 375). Biology reviewed by
<bibRefCitation id="C47BDC3D77E60F0C94D30FEAEB3DB115" author="Gauthier-Pilters, H. &amp; A. Innis Dagg" refId="ref115665" refString="Gauthier-Pilters, H., and A. Innis Dagg. 1981. The camel: Its evolution, ecology, behavior and relationships to man. University of Chicago Press, Chicago, IL, 208 pp." year="1981">Gauthier-Pilters and Innis Dagg (1981)</bibRefCitation>
. For history of domestication, see
<collectionCode id="E4322F61C460F717A20336592CA24277" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<collectionCode id="132847A592506D3A3DE927E77C6873B0" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
. Wilson (1984).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,345 @@
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<mods:title id="330D032DCD720B41CE773336BA95A2EB">Description of the mature larva of Gastrophysa janthina (Suffrian, 1851) [= G. unicolor auct.] (Coleoptera: Chrysomelidae: Chrysomelinae) and key to the larvae of the European Gastrophysa</mods:title>
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<treatment id="D02CFF38FF85C71D91B5535EFB9F1D2A" ID-DOI="http://doi.org/10.5281/zenodo.6244699" ID-GBIF-Taxon="119373228" ID-Zenodo-Dep="6244699" LSID="urn:lsid:plazi:treatment:D02CFF38FF85C71D91B5535EFB9F1D2A" httpUri="http://treatment.plazi.org/id/D02CFF38FF85C71D91B5535EFB9F1D2A" lastPageId="6" lastPageNumber="67" pageId="1" pageNumber="62">
<subSubSection id="109F1DA5FF85C71A91B5535EFEF4192F" pageId="1" pageNumber="62" type="nomenclature">
<paragraph id="583A4E2EFF85C71A91B5535EFCDB190C" blockId="1.[151,794,1586,1647]" box="[151,794,1586,1612]" pageId="1" pageNumber="62">
<heading id="0372F942FF85C71A91B5535EFCDB190C" bold="true" box="[151,794,1586,1612]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
<emphasis id="6AF1923CFF85C71A91B5535EFCDB190C" bold="true" box="[151,794,1586,1612]" pageId="1" pageNumber="62">
<taxonomicName id="9F8535ADFF85C71A91B5535EFDA8190C" authority="Suffrian, 1851" authorityName="Suffrian" authorityYear="1851" box="[151,617,1586,1612]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="janthina">
<emphasis id="6AF1923CFF85C71A91B5535EFE5D190C" bold="true" box="[151,412,1586,1612]" italics="true" pageId="1" pageNumber="62">Gastrophysa janthina</emphasis>
(Suffrian, 1851)
</taxonomicName>
, mature larva
</emphasis>
</heading>
</paragraph>
<paragraph id="583A4E2EFF85C71A91B55339FEF4192F" blockId="1.[151,794,1586,1647]" box="[151,309,1621,1647]" pageId="1" pageNumber="62">
(
<figureCitation id="C0BE52ABFF85C71A91BD5339FECE192F" box="[159,271,1621,1647]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2 7" captionStartId-0="3.[151,255,1223,1247]" captionStartId-1="4.[151,255,1786,1810]" captionTargetBox-0="[153,1436,595,1198]" captionTargetBox-1="[151,1436,195,1761]" captionTargetId-0="figure@3.[153,1438,594,1199]" captionTargetId-1="figure@4.[151,1436,194,1762]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Habitus of Gastrophysa unicolor, mature larva from Chandrexa de Queixa. Length = 6.4 mm." captionText-1="FIGURE 2 7. Gastrophysa spp. (2) Head of G. janthina, frontal view. (3) Antenna of G. janthina, frontal view. (4) Labrum of G. janthina, dorsal view. (5) Maxillae and labium of G. janthina, ventral view. (6) Leg of G. janthina, lateral view. (7) Labrum of G. v i r i d u l a, dorsal view. Scale bars = 0.1 mm." httpUri-0="https://zenodo.org/record/178651/files/figure.png" httpUri-1="https://zenodo.org/record/178652/files/figure.png" pageId="1" pageNumber="62">Figs. 16</figureCitation>
,
<figureCitation id="C0BE52ABFF85C71A903E5339FEEA192F" box="[284,299,1621,1647]" captionStart="FIGURE 8" captionStartId="5.[151,255,1930,1954]" captionTargetBox="[243,1344,195,1906]" captionTargetId="figure@5.[243,1345,194,1906]" captionTargetPageId="5" captionText="FIGURE 8. Gastrophysa viridula: location of tubercles and body chaetotaxy, right side. Pr, prothorax; Ms, mesothorax; Mt, metathorax; A 1 A 10, abdominal segments 1 10. See text for definition of tubercles abbreviations. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/178653/files/figure.png" pageId="1" pageNumber="62">8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="109F1DA5FF85C71A91B553C9FB33184F" pageId="1" pageNumber="62" type="materials_examined">
<paragraph id="583A4E2EFF85C71A91B553C9FB33184F" blockId="1.[151,1437,1701,2007]" pageId="1" pageNumber="62">
<emphasis id="6AF1923CFF85C71A91B553C9FE4A19FF" bold="true" box="[151,395,1701,1727]" pageId="1" pageNumber="62">Material examined.</emphasis>
<collectingCountry id="20920EBEFF85C71A90B653C9FE2B19FF" box="[404,490,1701,1727]" name="Spain" pageId="1" pageNumber="62">SPAIN</collectingCountry>
. A Coruña, A Capela, Caaveiro, (UTM 29TNJ7407),
<quantity id="9F7DE3CBFF85C71A954753C9FB6019FF" box="[1125,1185,1701,1727]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="62" unit="m" value="50.0">50 m</quantity>
asl,
<date id="2C3B68EEFF85C71A95FB53C9FA4119FF" box="[1241,1408,1701,1727]" pageId="1" pageNumber="62" value="1999-04-11">11 April 1999</date>
, 5 mature larvae; Ourense, Chandrexa de Queixa (UTM 29TPG3478),
<quantity id="9F7DE3CBFF85C71A929653A1FC3F19A7" box="[948,1022,1741,1767]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="62" unit="m" value="600.0">600 m</quantity>
asl,
<date id="2C3B68EEFF85C71A951653A1FB0519A7" box="[1076,1220,1741,1767]" pageId="1" pageNumber="62" value="1999-05-08">8 May 1999</date>
, 14 mature larvae. Voucher specimens are deposited in the Museo Nacional de Ciencias Naturales, Madrid,
<collectingCountry id="20920EBEFF85C71A958A5399FB2E184F" box="[1192,1263,1781,1807]" name="Spain" pageId="1" pageNumber="62">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="109F1DA5FF85C71891E45271FD5D1FF2" lastPageId="3" lastPageNumber="64" pageId="1" pageNumber="62" type="description">
<paragraph id="583A4E2EFF85C71A91E45271FA5D1897" blockId="1.[151,1437,1701,2007]" pageId="1" pageNumber="62">
<emphasis id="6AF1923CFF85C71A91E45271FE9F1877" bold="true" box="[198,350,1821,1847]" pageId="1" pageNumber="62">Description.</emphasis>
Habitus as in
<figureCitation id="C0BE52ABFF85C71A932B5271FD911877" box="[521,592,1821,1847]" captionStart="FIGURE 1" captionStartId="3.[151,255,1223,1247]" captionTargetBox="[153,1436,595,1198]" captionTargetId="figure@3.[153,1438,594,1199]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gastrophysa unicolor, mature larva from Chandrexa de Queixa. Length = 6.4 mm." httpUri="https://zenodo.org/record/178651/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
. Length: 5.57.1 mm. Body eruciform, moderately convex and slightly arched in preserved specimens. Inter-tubercular plates and tubercles concolor, brown in dorsal region, light brown in ventral region.
<emphasis id="6AF1923CFF85C71A90E05201FDCD18C7" bold="true" box="[450,524,1901,1927]" pageId="1" pageNumber="62">Head.</emphasis>
Hypognathous, well sclerotized. Color brown. Epicranial suture well developed and long, frontal arms distinct, V-shaped and almost straight (
<figureCitation id="C0BE52ABFF85C71A929452F9FC3E18EF" box="[950,1023,1941,1967]" captionStart="FIGURE 2 7" captionStartId="4.[151,255,1786,1810]" captionTargetBox="[151,1436,195,1761]" captionTargetId="figure@4.[151,1436,194,1762]" captionTargetPageId="4" captionText="FIGURE 2 7. Gastrophysa spp. (2) Head of G. janthina, frontal view. (3) Antenna of G. janthina, frontal view. (4) Labrum of G. janthina, dorsal view. (5) Maxillae and labium of G. janthina, ventral view. (6) Leg of G. janthina, lateral view. (7) Labrum of G. v i r i d u l a, dorsal view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/178652/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
). Endocarina present, extending to the middle of the frons and thus not reaching the clypeus. Vertex bearing 4 pairs of setae (
<date id="2C3B68EEFF85C71A95C052D1FAC01897" box="[1250,1281,1981,2007]" pageId="1" pageNumber="62">v1</date>
,
<date id="2C3B68EEFF85C71A943252D1FAEE1897" box="[1296,1327,1981,2007]" pageId="1" pageNumber="62">v3</date>
,
<date id="2C3B68EEFF85C71A941D52D1FA9E1897" box="[1343,1375,1981,2007]" pageId="1" pageNumber="62">v4</date>
,
<date id="2C3B68EEFF85C71A944F52D1FA4F1897" box="[1389,1422,1981,2007]" pageId="1" pageNumber="62">v6</date>
).
</paragraph>
<paragraph id="583A4E2EFF86C71891B555F4FD5D1FF2" blockId="2.[151,1437,152,2018]" lastBlockId="3.[151,1437,152,538]" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63">
Frons with 4 pairs of setae (f1, f3, f4, f6). Antennae short and well sclerotized, three-segmented (
<figureCitation id="C0BE52ABFF86C719942F55F4FA941FF2" box="[1293,1365,152,178]" captionStart="FIGURE 2 7" captionStartId="4.[151,255,1786,1810]" captionTargetBox="[151,1436,195,1761]" captionTargetId="figure@4.[151,1436,194,1762]" captionTargetPageId="4" captionText="FIGURE 2 7. Gastrophysa spp. (2) Head of G. janthina, frontal view. (3) Antenna of G. janthina, frontal view. (4) Labrum of G. janthina, dorsal view. (5) Maxillae and labium of G. janthina, ventral view. (6) Leg of G. janthina, lateral view. (7) Labrum of G. v i r i d u l a, dorsal view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/178652/files/figure.png" pageId="2" pageNumber="63">Fig. 3</figureCitation>
): first joint highly transverse, bearing 2 placoid sensilla (only 1 visible in frontal view); second joint 1.5 times longer than wide, bearing a large conical membranous sensillum and 5 minute setae; distal joint narrow, bottleshaped, constricted anteapically, with membranous apex bearing 6 highly minute setae similar in size (no central one larger than the others) and 1 placoid sensillum slightly produced and situated before the anteapical constriction. Stemmata arranged in two groups, 1 pair located below the base of antenna and 2 pairs behind the antenna. Clypeus with 2 pairs of setae.
<emphasis id="6AF1923CFF86C71993AB54EBFCE71EE1" bold="true" box="[649,806,391,417]" pageId="2" pageNumber="63">Mouthparts.</emphasis>
Labrum (
<figureCitation id="C0BE52ABFF86C71992BB54E4FC1E1EE2" box="[921,991,392,418]" captionStart="FIGURE 2 7" captionStartId="4.[151,255,1786,1810]" captionTargetBox="[151,1436,195,1761]" captionTargetId="figure@4.[151,1436,194,1762]" captionTargetPageId="4" captionText="FIGURE 2 7. Gastrophysa spp. (2) Head of G. janthina, frontal view. (3) Antenna of G. janthina, frontal view. (4) Labrum of G. janthina, dorsal view. (5) Maxillae and labium of G. janthina, ventral view. (6) Leg of G. janthina, lateral view. (7) Labrum of G. v i r i d u l a, dorsal view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/178652/files/figure.png" pageId="2" pageNumber="63">Fig. 4</figureCitation>
) bearing 1 pair of placoid sensilla and 2 pairs of setae on dorsal surface, one near middle, the other near lateral sides; anterior border with a median wide V-shaped notch; ventral surface with 2 pairs of short setae not reaching the anterior border. Mandibles symmetrical, 5-toothed, bearing 2 setae on external face and 1 placoid sensillum on dorsal side. Maxillae (
<figureCitation id="C0BE52ABFF86C719944E576CFF691D02" captionStart="FIGURE 2 7" captionStartId="4.[151,255,1786,1810]" captionTargetBox="[151,1436,195,1761]" captionTargetId="figure@4.[151,1436,194,1762]" captionTargetPageId="4" captionText="FIGURE 2 7. Gastrophysa spp. (2) Head of G. janthina, frontal view. (3) Antenna of G. janthina, frontal view. (4) Labrum of G. janthina, dorsal view. (5) Maxillae and labium of G. janthina, ventral view. (6) Leg of G. janthina, lateral view. (7) Labrum of G. v i r i d u l a, dorsal view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/178652/files/figure.png" pageId="2" pageNumber="63">Fig. 5</figureCitation>
): cardo transverse, without setae; stipes elongate, with 2 large setae on external face and 1 short seta on the internal one; mala bearing 10 setae on internal margin and apex, basal setae longer than apical ones; maxillary palpi 4-segmented, first joint almost as long as wide, bearing 2 long setae basally in ventral side, second joint transverse with 1 minute seta on external side, third joint longer than second, but still transverse, with 1 seta on internal face, 1 placoid sensillum on ventral face and two setae on external side, and fourth joint conical with 1 minute seta on internal face and membranous apex bearing 911 highly minute setae. Labium (
<figureCitation id="C0BE52ABFF86C719946E579CFA531C4A" box="[1356,1426,752,778]" captionStart="FIGURE 2 7" captionStartId="4.[151,255,1786,1810]" captionTargetBox="[151,1436,195,1761]" captionTargetId="figure@4.[151,1436,194,1762]" captionTargetPageId="4" captionText="FIGURE 2 7. Gastrophysa spp. (2) Head of G. janthina, frontal view. (3) Antenna of G. janthina, frontal view. (4) Labrum of G. janthina, dorsal view. (5) Maxillae and labium of G. janthina, ventral view. (6) Leg of G. janthina, lateral view. (7) Labrum of G. v i r i d u l a, dorsal view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/178652/files/figure.png" pageId="2" pageNumber="63">Fig. 5</figureCitation>
) with postmentum membranous, bearing 3 pairs of setae, anterolateral one shorter than the others; prementum with 4 pairs, 2 posterior and 2 anterior to labial palpi (only one visible in
<figureCitation id="C0BE52ABFF86C7199538562CFBA21C1A" box="[1050,1123,832,858]" captionStart="FIGURE 2 7" captionStartId="4.[151,255,1786,1810]" captionTargetBox="[151,1436,195,1761]" captionTargetId="figure@4.[151,1436,194,1762]" captionTargetPageId="4" captionText="FIGURE 2 7. Gastrophysa spp. (2) Head of G. janthina, frontal view. (3) Antenna of G. janthina, frontal view. (4) Labrum of G. janthina, dorsal view. (5) Maxillae and labium of G. janthina, ventral view. (6) Leg of G. janthina, lateral view. (7) Labrum of G. v i r i d u l a, dorsal view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/178652/files/figure.png" pageId="2" pageNumber="63">Fig. 5</figureCitation>
), one of the former much longer than the others; labial palpi 2-segmented, first joint transverse, distal joint conical with membranous apex bearing 9 highly minute setae.
<emphasis id="6AF1923CFF86C719936756E3FD681CE9" bold="true" box="[581,681,911,937]" pageId="2" pageNumber="63">Thorax.</emphasis>
Prothorax with tubercles D (dorsal), DL (dorsolateral) and EPa (epipleural) fused together in a pronotal sclerite (
<figureCitation id="C0BE52ABFF86C71993FD56D4FCE61C92" box="[735,807,952,978]" captionStart="FIGURE 8" captionStartId="5.[151,255,1930,1954]" captionTargetBox="[243,1344,195,1906]" captionTargetId="figure@5.[243,1345,194,1906]" captionTargetPageId="5" captionText="FIGURE 8. Gastrophysa viridula: location of tubercles and body chaetotaxy, right side. Pr, prothorax; Ms, mesothorax; Mt, metathorax; A 1 A 10, abdominal segments 1 10. See text for definition of tubercles abbreviations. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/178653/files/figure.png" pageId="2" pageNumber="63">Fig. 8</figureCitation>
), pronotum (D-DL-EPa) bearing 10 pairs of primary setae along with 1 pair of extremely minute ones; tubercle EPp bearing 1 setae; tubercle P (pleural) with 1 setae; ventral region with midventral tubercle ES (eusternal) fused with tubercle SS (sternellar), tubercle SS- ES bearing 2 pairs of setae. Meso- and metathorax with 5 tubercles on each side of dorsal region: Dai (dorsal anterior interior, bearing 1 seta), Dae (dorsal anterior exterior, 1 seta), Dpi-Dpe (dorsal posterior interior and dorsal posterior exterior fused together, 1 seta), DLai-DLpi (dorsolateral anterior interior and dorsolateral posterior interior fused together, 1 long seta and 2 smaller ones), and DLae-DLpe (dorsolateral anterior exterior and dorsolateral posterior exterior fused together, 2 large setae along with 12 minute ones) enclosing the eversible gland; epipleural region with 2 tubercles, EPa (epipleural anterior, 23 setae, variable among individuals and even lateral sides of a specimen) and EPp (epipleural posterior, 1 seta); mesothoracic spiracle (st) very close to EPa tubercle; P tubercle bearing 1 seta; SS-ES bearing 2 pairs of setae.
<emphasis id="6AF1923CFF86C7199555502BFB781A21" bold="true" box="[1143,1209,1351,1377]" pageId="2" pageNumber="63">Legs.</emphasis>
All pairs similar in size; trochantin (
<collectingCountry id="20920EBEFF86C719907E501CFEBB1ACA" box="[348,378,1392,1418]" name="Chile" pageId="2" pageNumber="63">Tr</collectingCountry>
) located in front of P tubercle (
<figureCitation id="C0BE52ABFF86C71993D3501CFCF81ACA" box="[753,825,1392,1418]" captionStart="FIGURE 8" captionStartId="5.[151,255,1930,1954]" captionTargetBox="[243,1344,195,1906]" captionTargetId="figure@5.[243,1345,194,1906]" captionTargetPageId="5" captionText="FIGURE 8. Gastrophysa viridula: location of tubercles and body chaetotaxy, right side. Pr, prothorax; Ms, mesothorax; Mt, metathorax; A 1 A 10, abdominal segments 1 10. See text for definition of tubercles abbreviations. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/178653/files/figure.png" pageId="2" pageNumber="63">Fig. 8</figureCitation>
), bearing 2 extremely minute setae in anterior half; prothoracic trochantin also bearing a large seta near posterior angle; coxa 1.5 times longer than wide in lateral view (
<figureCitation id="C0BE52ABFF86C71991C050ACFEED1A9A" box="[226,300,1472,1498]" captionStart="FIGURE 2 7" captionStartId="4.[151,255,1786,1810]" captionTargetBox="[151,1436,195,1761]" captionTargetId="figure@4.[151,1436,194,1762]" captionTargetPageId="4" captionText="FIGURE 2 7. Gastrophysa spp. (2) Head of G. janthina, frontal view. (3) Antenna of G. janthina, frontal view. (4) Labrum of G. janthina, dorsal view. (5) Maxillae and labium of G. janthina, ventral view. (6) Leg of G. janthina, lateral view. (7) Labrum of G. v i r i d u l a, dorsal view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/178652/files/figure.png" pageId="2" pageNumber="63">Fig. 6</figureCitation>
), with 4 large setae on dorsal half of anterior side and 5 shorter ones in lateral declivities; trochanter triangular in lateral view, with 2 setae on each side, 5 placoid sensilla near coxal articulation on anterior side and 2 on posterior side; femur trapezoidal in lateral view, with 3 setae dorsally (1 pair near apex and 1 single minute one near base, 3 ones on each lateral declivity and 1 placoid sensillum on anterior side; tibiotarsus twice longer than wide, bearing 4 setae dorsally, 1 on each side and 2 ventrally; unguis wide basally, curved apically, bearing 1 seta on lower side.
<emphasis id="6AF1923CFF86C719939053EBFCF219E1" bold="true" box="[690,819,1671,1697]" pageId="2" pageNumber="63">Abdomen.</emphasis>
Segments 16 with only 3 tubercles on each side of dorsal region (
<figureCitation id="C0BE52ABFF86C719907653DCFE5E198A" box="[340,415,1712,1738]" captionStart="FIGURE 8" captionStartId="5.[151,255,1930,1954]" captionTargetBox="[243,1344,195,1906]" captionTargetId="figure@5.[243,1345,194,1906]" captionTargetPageId="5" captionText="FIGURE 8. Gastrophysa viridula: location of tubercles and body chaetotaxy, right side. Pr, prothorax; Ms, mesothorax; Mt, metathorax; A 1 A 10, abdominal segments 1 10. See text for definition of tubercles abbreviations. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/178653/files/figure.png" pageId="2" pageNumber="63">Fig. 8</figureCitation>
): Dpi-Dpe (2 setae), Dae (1 seta) and DLae-DLpe (2 setae) enclosing the eversible gland; Dai and DLai-DLpi lacking; epipleural region with tubercle EP bearing 2 setae; spiracle (st) located between EP and DLae-DLpe tubercles, closer to EP; P tubercle with 2 setae, one of them shorter (lacking in segment 1); sternal region presents the following tubercles: PS (parasternal, 2 setae, excepting segment 1 with only 1 seta), SS (1 seta), ES (1 pair of setae); secondary tubercle as1 (with 1 minute seta) present. Segment 7 with tubercles Dpi-Dpe fused together in a single unpaired dorsal sclerite, and with tubercles as1, PS, SS and ES fused together in a ventral sclerite; segment 8 and 9 with dorsal and dorsolateral tubercles fused together, as well as pleural and ventral ones; segment 9 with tubercle EP lacking; segment 10 forming anal pseudopod, without dorsal tubercles, ventral ones fused.
</paragraph>
</subSubSection>
<subSubSection id="109F1DA5FF87C71891E455D3FB8F1D5A" pageId="3" pageNumber="64" type="distribution">
<paragraph id="583A4E2EFF87C71891E455D3FB8F1D5A" blockId="3.[151,1437,152,538]" pageId="3" pageNumber="64">
<emphasis id="6AF1923CFF87C71891E455D3FE3E1F99" bold="true" box="[198,511,191,217]" pageId="3" pageNumber="64">Distribution and ecology.</emphasis>
Although
<taxonomicName id="9F8535ADFF87C718935955ACFD3C1F99" box="[635,765,192,217]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="janthina">
<emphasis id="6AF1923CFF87C718935955ACFD3C1F99" box="[635,765,192,217]" italics="true" pageId="3" pageNumber="64">G. janthina</emphasis>
</taxonomicName>
was cited by
<bibRefCitation id="3C1433DFFF87C71892BE55ACFB901F9A" author="Oliveira" box="[924,1105,192,218]" pageId="3" pageNumber="68" refString="Oliveira, M. P. (1893) Catalogue des Insectes du Coleptres Universidade de Coimbra, Coimbra, 393 pp." type="book" year="1893">Oliveira (1893)</bibRefCitation>
from “
<emphasis id="6AF1923CFF87C718958755ACFA9C1F99" box="[1189,1373,192,217]" italics="true" pageId="3" pageNumber="64">
tout le
<collectingCountry id="20920EBEFF87C71895D755ACFA9C1F99" box="[1269,1373,192,217]" name="Portugal" pageId="3" pageNumber="64">Portugal</collectingCountry>
</emphasis>
” and after
<bibRefCitation id="3C1433DFFF87C71891F65584FEB81E42" author="Jolivet" box="[212,377,232,258]" pageId="3" pageNumber="68" refString="Jolivet, P. (1951) Contribution a l'Etude de genre Gastrophysa Chevrolat (Coleoptera: Chrysomelidae) (3 eme note). Bulletin de l'Institut royal des sciences naturelles de Belgique, 27, 1 - 47." type="journal article" year="1951">Jolivet (1951)</bibRefCitation>
this species is distributed throughout the Iberian peninsula, considering the available data the taxon is restricted to the northwest quadrant of the Iberian peninsula: it is very common in Galicia (
<bibRefCitation id="3C1433DFFF87C71891BD5454FE141E12" author="Baselga" box="[159,469,312,338]" pageId="3" pageNumber="67" refString="Baselga, A. &amp; Novoa, F. (2000 a) Los Chrysomelidae (Coleoptera) de la Sierra de Ancares, Noroeste de la Espana (Coleoptera). Nouvelle Revue d'Entomologie (N. S.), 17, 165 - 180." type="journal article" year="2000" yearSuffix="a">Baselga &amp; Novoa, 2000a</bibRefCitation>
;
<bibRefCitation id="3C1433DFFF87C71890C75454FDF31E12" author="Baselga" box="[485,562,312,338]" pageId="3" pageNumber="67" refString="Baselga, A. &amp; Novoa, F. (2000 b) Los Chrysomelidae (Coleoptera) de los sistemas dunares de Galicia (Noroeste de Espana). Boletin de la Real Sociedad Espanola de Historia Natural (Sec. Biol.), 96, 113 - 124." type="journal article" year="2000" yearSuffix="b">2000b</bibRefCitation>
;
<bibRefCitation id="3C1433DFFF87C71893635454FDBE1E12" author="Baselga" box="[577,639,312,338]" pageId="3" pageNumber="67" refString="Baselga, A. &amp; Novoa, F. (2002) Los Chrysomelidae (Coleoptera) de las sierras orientales de Ourense (Galicia, noroeste de la PenRnsula Iberica). Boletin de la Asociacion espanola de Entomologia, 26, 57 - 73." type="journal article" year="2002">2002</bibRefCitation>
;
<bibRefCitation id="3C1433DFFF87C71893AC5454FD1B1E12" author="Baselga" box="[654,730,312,338]" pageId="3" pageNumber="67" refString="Baselga, A. &amp; Novoa, F. (2004 a) Coleopteros del Parque Natural de las Fragas del Eume (Galicia, noroeste de la PenRnsula Iberica), II: Scarabaeoidea, Buprestoidea, Byrrhoidea, Elateroidea, Bostrichoidea, Lymexyloidea, Cleroidea, Cucujoidea, Tenebrionoidea, Chrysomeloidea y Curculionoidea. Boletin de la Asociacion espanola de Entomologia, 28, 121 - 143." type="journal article" year="2004" yearSuffix="a">2004a</bibRefCitation>
;
<bibRefCitation id="3C1433DFFF87C71893CB5454FCE41E12" author="Baselga" box="[745,805,312,338]" pageId="3" pageNumber="68" refString="Baselga, A. &amp; Novoa, F. (2006) Diversity of Chrysomelidae (Coleoptera) in Galicia, Northwest Spain: estimating the completeness of the regional inventory. Biodiversity and Conservation, 15, 205 - 230." type="journal article" year="2006">2006</bibRefCitation>
) and it was also recorded from León (
<bibRefCitation id="3C1433DFFF87C71895DC5454FE481E3A" author="Petitpierre" pageId="3" pageNumber="68" refString="Petitpierre, E. &amp; Gomez-Zurita, J. (1998) Los Chrysomelidae de Leon; NO de Espana (Coleoptera). Nouvelle Revue d'Entomologie (N. S.), 15, 13 - 26." type="journal article" year="1998">Petitpierre &amp; Gómez-Zurita, 1998</bibRefCitation>
), North
<collectingCountry id="20920EBEFF87C71890CB540CFD8D1E3A" box="[489,588,352,378]" name="Portugal" pageId="3" pageNumber="64">Portugal</collectingCountry>
(
<bibRefCitation id="3C1433DFFF87C718937E540CFC481E3A" author="Baselga" box="[604,905,352,378]" pageId="3" pageNumber="67" refString="Baselga, A. &amp; Novoa, F. (2000 b) Los Chrysomelidae (Coleoptera) de los sistemas dunares de Galicia (Noroeste de Espana). Boletin de la Real Sociedad Espanola de Historia Natural (Sec. Biol.), 96, 113 - 124." type="journal article" year="2000" yearSuffix="b">Baselga &amp; Novoa, 2000b</bibRefCitation>
), Zamora (
<bibRefCitation id="3C1433DFFF87C7189525540CFAE71E3A" author="Baselga" box="[1031,1318,352,378]" pageId="3" pageNumber="67" refString="Baselga, A. &amp; Novoa, F. (2003) Los Chrysomelidae de los Arribes del Duero, noroeste de la PenRnsula Iberica (Coleoptera). Nouvelle Revue d'Entomologie (N. S.), 20, 117 - 131." type="journal article" year="2003">Baselga &amp; Novoa, 2003</bibRefCitation>
), Asturias and Cantabria (
<bibRefCitation id="3C1433DFFF87C718906854E4FDD51EE2" author="Petitpierre" box="[330,532,392,418]" pageId="3" pageNumber="68" refString="Petitpierre, E. (2005) Listado de Chrysomelidae (Coleoptera) de Asturias y Cantabria. Boletin de la Asociacion espanola de espanla Entomologia, 29, 51 - 72." type="journal article" year="2005">Petitpierre, 2005</bibRefCitation>
). In all these citations the species is referred as
<taxonomicName id="9F8535ADFF87C718956254E4FB031EE1" box="[1088,1218,392,417]" class="Insecta" family="Chrysomelidae" genus="Chrysomela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="6AF1923CFF87C718956254E4FB031EE1" box="[1088,1218,392,417]" italics="true" pageId="3" pageNumber="64">G. unicolor</emphasis>
</taxonomicName>
.
<taxonomicName id="9F8535ADFF87C71895F354E4FF151E89" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="janthina">
<emphasis id="6AF1923CFF87C71895F354E4FF151E89" italics="true" pageId="3" pageNumber="64">Gastrophysa janthina</emphasis>
</taxonomicName>
lives on
<taxonomicName id="9F8535ADFF87C718906654DCFE261E8A" box="[324,487,432,458]" class="Magnoliopsida" family="Polygonaceae" kingdom="Plantae" order="Polygonales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="family">Polygonaceae</taxonomicName>
of the genera
<taxonomicName id="9F8535ADFF87C71893BA54DCFCDF1E89" box="[664,798,432,457]" class="Magnoliopsida" family="Polygonaceae" genus="Polygonum" kingdom="Plantae" order="Polygonales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="genus">
<emphasis id="6AF1923CFF87C71893BA54DCFCDF1E89" box="[664,798,432,457]" italics="true" pageId="3" pageNumber="64">Polygonum</emphasis>
</taxonomicName>
L. (Petitpierre &amp; G³mez-Zurita, 1998) and
<taxonomicName id="9F8535ADFF87C718940B54DCFAB81E89" box="[1321,1401,432,457]" class="Magnoliopsida" family="Polygonaceae" genus="Rumex" kingdom="Plantae" order="Polygonales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="genus">
<emphasis id="6AF1923CFF87C718940B54DCFAB81E89" box="[1321,1401,432,457]" italics="true" pageId="3" pageNumber="64">Rumex</emphasis>
</taxonomicName>
L. (
<bibRefCitation id="3C1433DFFF87C71891BD54B4FE0B1EB2" author="Baselga" box="[159,458,472,498]" pageId="3" pageNumber="67" refString="Baselga, A. &amp; Novoa, F. (2000 a) Los Chrysomelidae (Coleoptera) de la Sierra de Ancares, Noroeste de la Espana (Coleoptera). Nouvelle Revue d'Entomologie (N. S.), 17, 165 - 180." type="journal article" year="2000" yearSuffix="a">Baselga &amp; Novoa, 2000a</bibRefCitation>
;
<bibRefCitation id="3C1433DFFF87C71890F454B4FDE21EB2" author="Baselga" box="[470,547,472,498]" pageId="3" pageNumber="67" refString="Baselga, A. &amp; Novoa, F. (2000 b) Los Chrysomelidae (Coleoptera) de los sistemas dunares de Galicia (Noroeste de Espana). Boletin de la Real Sociedad Espanola de Historia Natural (Sec. Biol.), 96, 113 - 124." type="journal article" year="2000" yearSuffix="b">2000b</bibRefCitation>
;
<bibRefCitation id="3C1433DFFF87C718930C54B4FDAD1EB2" author="Baselga" box="[558,620,472,498]" pageId="3" pageNumber="67" refString="Baselga, A. &amp; Novoa, F. (2002) Los Chrysomelidae (Coleoptera) de las sierras orientales de Ourense (Galicia, noroeste de la PenRnsula Iberica). Boletin de la Asociacion espanola de Entomologia, 26, 57 - 73." type="journal article" year="2002">2002</bibRefCitation>
;
<bibRefCitation id="3C1433DFFF87C718935A54B4FD761EB2" author="Baselga" box="[632,695,472,498]" pageId="3" pageNumber="67" refString="Baselga, A. &amp; Novoa, F. (2003) Los Chrysomelidae de los Arribes del Duero, noroeste de la PenRnsula Iberica (Coleoptera). Nouvelle Revue d'Entomologie (N. S.), 20, 117 - 131." type="journal article" year="2003">2003</bibRefCitation>
), but no host plant was ever identified to the specific level. The larvae described in this paper were collected on
<taxonomicName id="9F8535ADFF87C71893EE576CFCDD1D59" box="[716,796,512,537]" class="Magnoliopsida" family="Polygonaceae" genus="Rumex" kingdom="Plantae" order="Polygonales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="genus">
<emphasis id="6AF1923CFF87C71893EE576CFCDD1D59" box="[716,796,512,537]" italics="true" pageId="3" pageNumber="64">Rumex</emphasis>
</taxonomicName>
spp. (field identification).
</paragraph>
</subSubSection>
<caption id="0CFA1EA6FF87C71891B551ABFAC61B9E" box="[151,1287,1223,1247]" httpUri="https://zenodo.org/record/178651/files/figure.png" pageId="3" pageNumber="64" targetBox="[153,1436,595,1198]" targetPageId="3">
<paragraph id="583A4E2EFF87C71891B551ABFAC61B9E" blockId="3.[151,1287,1223,1247]" box="[151,1287,1223,1247]" pageId="3" pageNumber="64">
<emphasis id="6AF1923CFF87C71891B551ABFED81B9F" bold="true" box="[151,281,1223,1247]" pageId="3" pageNumber="64">FIGURE 1.</emphasis>
Habitus of
<taxonomicName id="9F8535ADFF87C71890B451ABFDBC1B9E" box="[406,637,1223,1246]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="6AF1923CFF87C71890B451ABFDBC1B9E" box="[406,637,1223,1246]" italics="true" pageId="3" pageNumber="64">Gastrophysa unicolor</emphasis>
</taxonomicName>
, mature larva from Chandrexa de Queixa. Length = 6.4 mm.
</paragraph>
</caption>
<subSubSection id="109F1DA5FF87C71D91E45060FB9F1D2A" lastPageId="6" lastPageNumber="67" pageId="3" pageNumber="64" type="discussion">
<paragraph id="583A4E2EFF87C71891E45060FD7318E6" blockId="3.[151,1437,1292,1958]" pageId="3" pageNumber="64">
<emphasis id="6AF1923CFF87C71891E45060FE901A66" bold="true" box="[198,337,1292,1318]" pageId="3" pageNumber="64">Discussion.</emphasis>
Four species of
<taxonomicName id="9F8535ADFF87C71893315061FD691A66" box="[531,680,1293,1318]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1923CFF87C71893315061FD691A66" box="[531,680,1293,1318]" italics="true" pageId="3" pageNumber="64">Gastrophysa</emphasis>
</taxonomicName>
are distributed in the western Palaearctic region (
<bibRefCitation id="3C1433DFFF87C71895C95060FF121A0E" author="Warchalowski" pageId="3" pageNumber="68" refString="Warchalowski, A. (2003) Chrysomelidae. The leaf-beetles of Europe and the Mediterranean area. Natura optima dux Foundation, Warszawa, 600 pp." type="book" year="2003">Warchalowski, 2003</bibRefCitation>
) but the larval stages are known only for
<taxonomicName id="9F8535ADFF87C71893975059FCFD1A0E" box="[693,828,1333,1358]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="polygoni">
<emphasis id="6AF1923CFF87C71893975059FCFD1A0E" box="[693,828,1333,1358]" italics="true" pageId="3" pageNumber="64">G. polygoni</emphasis>
</taxonomicName>
and
<emphasis id="6AF1923CFF87C71892565059FC2D1A0E" box="[884,1004,1333,1358]" italics="true" pageId="3" pageNumber="64">G. v i r i d u l a</emphasis>
(
<bibRefCitation id="3C1433DFFF87C71895245058FB6B1A0E" author="Hennig" box="[1030,1194,1332,1358]" pageId="3" pageNumber="68" refString="Hennig, W. (1938) U bersicht uber die Larven der wichtigsten deutschen Chrysomelinen (Coleoptera). Arbeiten uber physiologische und angewandte Entomologie aus Berlin-Dahlem, 5, 85 - 136." type="journal article" year="1938">Hennig, 1938</bibRefCitation>
;
<bibRefCitation id="3C1433DFFF87C71895975058FA4E1A0E" author="Steinhausen" box="[1205,1423,1332,1358]" pageId="3" pageNumber="68" refString="Steinhausen, W. R. (1994) Familie Chrysomelidae. In: Klausnitzer, B. (Ed.). Goecke &amp; Evers, Krefeld, Germany, pp. 231 - 314." type="book chapter" year="1994">Steinhausen, 1994</bibRefCitation>
). The larvae of the genus
<taxonomicName id="9F8535ADFF87C71890925031FD851A36" box="[432,580,1373,1398]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1923CFF87C71890925031FD851A36" box="[432,580,1373,1398]" italics="true" pageId="3" pageNumber="64">Gastrophysa</emphasis>
</taxonomicName>
are well defined by the following combination of characters (
<bibRefCitation id="3C1433DFFF87C718943A5030FA561A36" author="Cox" box="[1304,1431,1372,1398]" pageId="3" pageNumber="68" refString="Cox, M. L. (1982) Larvae of the British genera of chrysomeline beetles (Coleoptera Chrysomelidae). Systematic Entomology, 7, 297 - 310." type="journal article" year="1982">Cox, 1982</bibRefCitation>
;
<bibRefCitation id="3C1433DFFF87C71891B550E8FEB81ADE" author="Steinhausen" box="[151,377,1412,1438]" pageId="3" pageNumber="68" refString="Steinhausen, W. R. (1994) Familie Chrysomelidae. In: Klausnitzer, B. (Ed.). Goecke &amp; Evers, Krefeld, Germany, pp. 231 - 314." type="book chapter" year="1994">Steinhausen, 1994</bibRefCitation>
): body dorsally not strongly convex, epicraneal suture long, labrum with an anterior median notch and 2 pairs of setae on upper surface, pronotum bearing a low number of setae on disc, dorsolateral tubercles of meso-, metathorax and abdominal segments 17 with eversible glands, abdominal segments 16 with a dorsal anterior tubercle, tarsal claws not toothed on lower side. The mature larva of
<taxonomicName id="9F8535ADFF87C718943A5091FA5D1956" box="[1304,1436,1533,1558]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="janthina">
<emphasis id="6AF1923CFF87C718943A5091FA5D1956" box="[1304,1436,1533,1558]" italics="true" pageId="3" pageNumber="64">G. janthina</emphasis>
</taxonomicName>
agrees all these diagnostic characters, as described in this paper, and keys out with
<taxonomicName id="9F8535ADFF87C71895525349FAAB197E" box="[1136,1386,1573,1598]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="viridula">
<emphasis id="6AF1923CFF87C71895525349FAAB197E" box="[1136,1386,1573,1598]" italics="true" pageId="3" pageNumber="64">Gastrophysa viridula</emphasis>
</taxonomicName>
following both
<bibRefCitation id="3C1433DFFF87C71890135320FE211926" author="Hennig" box="[305,480,1612,1638]" pageId="3" pageNumber="68" refString="Hennig, W. (1938) U bersicht uber die Larven der wichtigsten deutschen Chrysomelinen (Coleoptera). Arbeiten uber physiologische und angewandte Entomologie aus Berlin-Dahlem, 5, 85 - 136." type="journal article" year="1938">Hennig (1938)</bibRefCitation>
and
<bibRefCitation id="3C1433DFFF87C718933C5320FCC21926" author="Steinhausen" box="[542,771,1612,1638]" pageId="3" pageNumber="68" refString="Steinhausen, W. R. (1994) Familie Chrysomelidae. In: Klausnitzer, B. (Ed.). Goecke &amp; Evers, Krefeld, Germany, pp. 231 - 314." type="book chapter" year="1994">Steinhausen (1994)</bibRefCitation>
works, due to its brown dorsum with tubercles clearly delimited from intertubercular tegument. No characters are available in the cited papers to separate
<taxonomicName id="9F8535ADFF87C71894395319FA5D19CE" box="[1307,1436,1653,1678]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="janthina">
<emphasis id="6AF1923CFF87C71894395319FA5D19CE" box="[1307,1436,1653,1678]" italics="true" pageId="3" pageNumber="64">G. janthina</emphasis>
</taxonomicName>
from
<taxonomicName id="9F8535ADFF87C71891FE53F1FE9D19F6" box="[220,348,1693,1718]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="viridula">
<emphasis id="6AF1923CFF87C71891FE53F1FE9D19F6" box="[220,348,1693,1718]" italics="true" pageId="3" pageNumber="64">G. viridula</emphasis>
</taxonomicName>
, but larvae of the latter species were collected [
<collectingCountry id="20920EBEFF87C718929353F0FBCA19F6" box="[945,1035,1692,1718]" name="Spain" pageId="3" pageNumber="64">SPAIN</collectingCountry>
: Lleida, Valle de Ar&lt;n, Circo de Colomers (31TCH2922),
<date id="2C3B68EEFF87C71890E653A8FDA0199E" box="[452,609,1732,1758]" pageId="3" pageNumber="64" value="2006-07-23">23 July 2006</date>
, 19 mature larvae and
<specimenCount id="4E8385A7FF87C718925253A8FC12199E" box="[880,979,1732,1758]" pageId="3" pageNumber="64" type="adult">5 adults</specimenCount>
], and the comparison of the larvae has showed that they are almost identical in both species. No differences were observed in the number and configuration of tubercles or setae and even the same variation in the number of setae of meso- and metathoracical tubercles DLae-DLpe and EPa is observed in both species. The only diagnostic character that could be detected is the outline of the labrum which has a wide V-shaped median notch in
<taxonomicName id="9F8535ADFF87C71895725209FB13183E" box="[1104,1234,1893,1918]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="janthina">
<emphasis id="6AF1923CFF87C71895725209FB13183E" box="[1104,1234,1893,1918]" italics="true" pageId="3" pageNumber="64">G. janthina</emphasis>
</taxonomicName>
, whereas it has a narrow U-shaped notch in G.
<taxonomicName id="9F8535ADFF87C71890CD52E1FD8D18E6" box="[495,588,1933,1958]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="viridula">
<emphasis id="6AF1923CFF87C71890CD52E1FD8D18E6" box="[495,588,1933,1958]" italics="true" pageId="3" pageNumber="64">viridula</emphasis>
</taxonomicName>
(
<figureCitation id="C0BE52ABFF87C718937E52E0FD6518E6" box="[604,676,1932,1958]" captionStart="FIGURE 2 7" captionStartId="4.[151,255,1786,1810]" captionTargetBox="[151,1436,195,1761]" captionTargetId="figure@4.[151,1436,194,1762]" captionTargetPageId="4" captionText="FIGURE 2 7. Gastrophysa spp. (2) Head of G. janthina, frontal view. (3) Antenna of G. janthina, frontal view. (4) Labrum of G. janthina, dorsal view. (5) Maxillae and labium of G. janthina, ventral view. (6) Leg of G. janthina, lateral view. (7) Labrum of G. v i r i d u l a, dorsal view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/178652/files/figure.png" pageId="3" pageNumber="64">Fig. 7</figureCitation>
).
</paragraph>
<caption id="0CFA1EA6FF80C71F91B55396FC9D1816" httpUri="https://zenodo.org/record/178652/files/figure.png" pageId="4" pageNumber="65" targetBox="[151,1436,195,1761]" targetPageId="4">
<paragraph id="583A4E2EFF80C71F91B55396FC9D1816" blockId="4.[151,1436,1786,1879]" pageId="4" pageNumber="65">
<emphasis id="6AF1923CFF80C71F91B55396FEF61852" bold="true" box="[151,311,1786,1810]" pageId="4" pageNumber="65">FIGURE 27.</emphasis>
<taxonomicName id="9F8535ADFF80C71F90635396FE091851" box="[321,456,1786,1809]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1923CFF80C71F90635396FE091851" box="[321,456,1786,1809]" italics="true" pageId="4" pageNumber="65">Gastrophysa</emphasis>
</taxonomicName>
spp. (2) Head of
<taxonomicName id="9F8535ADFF80C71F93B65396FCCC1851" box="[660,781,1786,1809]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="janthina">
<emphasis id="6AF1923CFF80C71F93B65396FCCC1851" box="[660,781,1786,1809]" italics="true" pageId="4" pageNumber="65">G. janthina</emphasis>
</taxonomicName>
, frontal view. (3) Antenna of
<taxonomicName id="9F8535ADFF80C71F957D5396FB191851" box="[1119,1240,1786,1809]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="janthina">
<emphasis id="6AF1923CFF80C71F957D5396FB191851" box="[1119,1240,1786,1809]" italics="true" pageId="4" pageNumber="65">G. janthina</emphasis>
</taxonomicName>
, frontal view. (4) Labrum of
<taxonomicName id="9F8535ADFF80C71F90325271FE461874" box="[272,391,1821,1844]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="janthina">
<emphasis id="6AF1923CFF80C71F90325271FE461874" box="[272,391,1821,1844]" italics="true" pageId="4" pageNumber="65">G. janthina</emphasis>
</taxonomicName>
, dorsal view. (5) Maxillae and labium of
<taxonomicName id="9F8535ADFF80C71F926A5271FC7E1874" box="[840,959,1821,1844]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="janthina">
<emphasis id="6AF1923CFF80C71F926A5271FC7E1874" box="[840,959,1821,1844]" italics="true" pageId="4" pageNumber="65">G. janthina</emphasis>
</taxonomicName>
, ventral view. (6) Leg of
<taxonomicName id="9F8535ADFF80C71F95F65271FA8A1874" box="[1236,1355,1821,1844]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="janthina">
<emphasis id="6AF1923CFF80C71F95F65271FA8A1874" box="[1236,1355,1821,1844]" italics="true" pageId="4" pageNumber="65">G. janthina</emphasis>
</taxonomicName>
, lateral view. (7) Labrum of
<emphasis id="6AF1923CFF80C71F9056522CFE231817" box="[372,482,1856,1879]" italics="true" pageId="4" pageNumber="65">G. v i r i d u l a</emphasis>
, dorsal view. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="0CFA1EA6FF81C71E91B552E6FF0918A6" httpUri="https://zenodo.org/record/178653/files/figure.png" pageId="5" pageNumber="66" targetBox="[243,1344,195,1906]" targetPageId="5">
<paragraph id="583A4E2EFF81C71E91B552E6FF0918A6" blockId="5.[151,1436,1930,2022]" pageId="5" pageNumber="66">
<emphasis id="6AF1923CFF81C71E91B552E6FED818E2" bold="true" box="[151,281,1930,1954]" pageId="5" pageNumber="66">FIGURE 8.</emphasis>
<taxonomicName id="9F8535ADFF81C71E900252E6FDC518E1" box="[288,516,1930,1953]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="viridula">
<emphasis id="6AF1923CFF81C71E900252E6FDC518E1" box="[288,516,1930,1953]" italics="true" pageId="5" pageNumber="66">Gastrophysa viridula</emphasis>
</taxonomicName>
: location of tubercles and body chaetotaxy, right side. Pr, prothorax; Ms, mesothorax; Mt, metathorax; A1A10, abdominal segments 110. See text for definition of tubercles abbreviations. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="583A4E2EFF82C71D91E455F4FB9F1D2A" blockId="6.[151,1436,152,618]" pageId="6" pageNumber="67">
The description of the larva of
<taxonomicName id="9F8535ADFF82C71D931055F4FD751FF1" box="[562,692,152,177]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="janthina">
<emphasis id="6AF1923CFF82C71D931055F4FD751FF1" box="[562,692,152,177]" italics="true" pageId="6" pageNumber="67">G. janthina</emphasis>
</taxonomicName>
is the first one within the subgenus
<taxonomicName id="9F8535ADFF82C71D957E55F4FA891FF2" authority="Jolivet" authorityName="Jolivet" box="[1116,1352,152,178]" class="Insecta" family="Chrysomelidae" genus="Exiguipenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1923CFF82C71D957E55F4FB311FF1" box="[1116,1264,152,177]" italics="true" pageId="6" pageNumber="67">Exiguipenna</emphasis>
Jolivet
</taxonomicName>
, which includes only this species and
<taxonomicName id="9F8535ADFF82C71D90D455ACFD9F1F99" box="[502,606,192,217]" class="Insecta" family="Chrysomelidae" genus="Gastrophysa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="6AF1923CFF82C71D90D455ACFD9F1F99" box="[502,606,192,217]" italics="true" pageId="6" pageNumber="67">G. analis</emphasis>
</taxonomicName>
from the Balkan peninsula. The subgenus was established based on the reduction of wings in adults of both species. No other characters support the splitting of the genus, and the larval morphology does not supply any other diagnostic feature, providing that the two species are virtually identical at this stage, except for the shape of the labrum, which is highly variable among close species of Chrysomelinae (
<bibRefCitation id="3C1433DFFF82C71D907D540CFD831E3A" author="Steinhausen" box="[351,578,352,378]" pageId="6" pageNumber="68" refString="Steinhausen, W. R. (1994) Familie Chrysomelidae. In: Klausnitzer, B. (Ed.). Goecke &amp; Evers, Krefeld, Germany, pp. 231 - 314." type="book chapter" year="1994">Steinhausen, 1994</bibRefCitation>
;
<bibRefCitation id="3C1433DFFF82C71D9377540CFC571E3A" author="Baselga" box="[597,918,352,378]" lastPageId="6" lastPageNumber="68" pageId="6" pageNumber="67" refString="Baselga, A. &amp; Novoa, F. (2004 b) Larvae of the Gonioctena subgenus Spartoxena: description of mature larvae of G. l e p - rieuri and G. aegrota (Coleoptera: Chrysomelidae: Chrysomelinae). Canadian Entomologist, 136, 313 - 321." type="journal article" year="2004" yearSuffix="b">Baselga &amp; Novoa, 2004b</bibRefCitation>
) and thus not appropriate for supporting supraspecific taxa. On the other hand, wing reduction or loss is a common evolutionary event frequently produced in independent lineages due to environmental conditions (see
<bibRefCitation id="3C1433DFFF82C71D929B54DCFB261E8A" author="Wagner" box="[953,1255,432,458]" pageId="6" pageNumber="68" refString="Wagner, D. L. &amp; Liebherr, J. K. (1992) Flightlessness in insects. Trends in Ecology &amp; Evolution, 7, 216 - 220." type="journal article" year="1992">Wagner &amp; Liebherr, 1992</bibRefCitation>
for a review of the causes leading to apterism and its frequent independence of phylogeny;
<bibRefCitation id="3C1433DFFF82C71D950354B4FB7F1EB2" author="Philips" box="[1057,1214,472,498]" pageId="6" pageNumber="68" refString="Philips, T. K. (2000) Phylogenetic analysis of the New World Ptininae (Coleoptera: Bostrichoidea). Systematic Entomology, 25, 235 - 262." type="journal article" year="2000">Philips, 2000</bibRefCitation>
for
<bibRefCitation id="3C1433DFFF82C71D95D054B4FB3E1EB2" author="Philips" box="[1266,1279,472,498]" pageId="6" pageNumber="68" refString="Philips, T. K. (2000) Phylogenetic analysis of the New World Ptininae (Coleoptera: Bostrichoidea). Systematic Entomology, 25, 235 - 262." type="journal article" year="2000" yearSuffix="a">a</bibRefCitation>
report of the independent origin of flightlessness in New World
<taxonomicName id="9F8535ADFF82C71D93CF576CFC901D5A" box="[749,849,512,538]" class="Insecta" family="Ptinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="subFamily" subFamily="Ptininae">Ptininae</taxonomicName>
; and
<bibRefCitation id="3C1433DFFF82C71D92AC576CFBF61D5A" author="Flowers" box="[910,1079,512,538]" pageId="6" pageNumber="68" refString="Flowers, R. W. (2004) New flightless Eumolpinae of the genera Apterodina Bechyne and Brachypterodina n. gen. (Coleoptera: Chrysomelidae) from the Neotropics. Zootaxa, 549, 1 - 18." type="journal article" year="2004">Flowers, 2004</bibRefCitation>
for an example in other
<taxonomicName id="9F8535ADFF82C71D9475576CFED51D02" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="family">Chrysomelidae</taxonomicName>
). In my opinion it is questionable that brachypterism justify the subgenus
<taxonomicName id="9F8535ADFF82C71D955F5744FAD01D01" box="[1149,1297,552,577]" class="Insecta" family="Chrysomelidae" genus="Exiguipenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1923CFF82C71D955F5744FAD01D01" box="[1149,1297,552,577]" italics="true" pageId="6" pageNumber="67">Exiguipenna</emphasis>
</taxonomicName>
, in absence of any other evidence, and further phylogenetic research should clarify its validity.
</paragraph>
</subSubSection>
</treatment>
</document>