diff --git a/data/68/B1/2C/68B12CA5515459339D80CC295A1C0FE0.xml b/data/68/B1/2C/68B12CA5515459339D80CC295A1C0FE0.xml
new file mode 100644
index 00000000000..7525cd1882e
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@@ -0,0 +1,1071 @@
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+<mods:title id="76AA5148A9EF14F820BCB8482481E585">The genus Liocanthydrus Guignot, 1957 (Coleoptera, Noteridae) in Argentina: new records and larval morphology</mods:title>
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+<mods:namePart id="016ACB9E88CD7263D3D8DA40F49C701A">Urcola, Juan I.</mods:namePart>
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+<mods:namePart id="FF3D4AF251E8BDFE4EA76959FE969733">Toledo, Mario E.</mods:namePart>
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+<subSubSection id="20D3434C044ACB774BBCEBF3291693EA" type="nomenclature">
+<paragraph id="68F3761E078334CE9D6386EC9F31F762">
+<taxonomicName id="7E99593927CDC1C4DB0734DAC1729E9F" ID-CoL="C3JRD" authority="Baca et al., 2014" authorityName="Baca" authorityYear="2014" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="910019508B62573162B1E21E2CE4AF6E" italics="true">Liocanthydrus nanops</emphasis>
+<bibRefCitation id="69E0F7CA630425D2E0A5E979CD24C8AF" DOI="10.11646/zootaxa.3793.2.3" author="Baca SM &amp; Gustafson GT &amp; Toledo M &amp; Miller KB" journalOrPublisher="Zootaxa" pagination="231-246" refId="B1" refString="Baca SM, Gustafson GT, Toledo M, Miller KB (2014) Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species. Zootaxa 3793: 231-246. https://doi.org/10.11646/zootaxa.3793.2.3" title="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species." volume="3793" year="2014">Baca et al., 2014</bibRefCitation>
+</taxonomicName>
+</paragraph>
+</subSubSection>
+<subSubSection id="587787517289CAE8933BC7185D893D69" type="description">
+<paragraph id="C54EC3B65628F4FC99F6D426196064E1">
+<figureCitation id="54A7A8332F79B8B4E0167284C0297303" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1–8. Habitus (1, 2) and genitalia (3–8) of Liocanthydrus nanops Baca et al., 2014, male from Argentina, Misiones Province (LEBA) 1 dorsal aspect 2 ventral aspect 3 segment IX, right lateral aspect 4 right lateral lobe, right lateral aspect 5 left lateral lobe, right lateral aspect 6 median lobe, left lateral aspect 7 median lobe, dorsal aspect 8 median lobe, right lateral aspect. Scale bars: 1.00 mm (1, 2); 0.25 mm (3–8)." figureDoi="10.3897/zookeys.1231.144746.figures1-8" httpUri="https://binary.pensoft.net/fig/1279877">Figs 1–8</figureCitation>
+, 
+<figureCitation id="76BD7117AC1A766619E0C7250621E4A7" captionStart="Figures 9–12" captionStartId="F2" captionText="Figures 9–12. Male genitalia of Liocanthydrus nanops Baca et al., 2014, modified from drawings in Grosso (1979) 9 segment IX, right lateral aspect 10 right lateral lobe, right lateral aspect 11 left lateral lobe, right lateral aspect 12 median lobe, right lateral aspect. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1231.144746.figures9-12" httpUri="https://binary.pensoft.net/fig/1279878">9–12</figureCitation>
+, 
+<figureCitation id="1C11181988AE2A38D0494A15561A3AC4" captionStart="Figures 13–16" captionStartId="F3" captionText="Figures 13–16. Habitat of Liocanthydrus nanops Baca et al., 2014, Iguazú NP, Misiones Province, Argentina 13 general view of the stream’s flooding site 14 collecting of beetles 15 details of the stream 16 instar III larva of L. nanops. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1231.144746.figures13-16" httpUri="https://binary.pensoft.net/fig/1279879">16</figureCitation>
+, 
+<figureCitation id="E7B969E59D45F03AC3BEB3E99887FA6B" captionStart="Figures 18–20" captionStartId="F5" captionText="Figures 18–20. Liocanthydrus nanops Baca et al., 2014, instar I 18 habitus, dorsal aspect 19 cephalic capsule, dorsal aspect 20 cephalic capsule, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. Solid square indicates additional seta. EB: egg burster, FR: frontoclypeus, PA: parietal, TP: tentorial pit. Scale bars: 0.50 mm (18); 0.10 mm (19, 20)." figureDoi="10.3897/zookeys.1231.144746.figures18-20" httpUri="https://binary.pensoft.net/fig/1279881">18–20</figureCitation>
+, 
+<figureCitation id="69C400DDB0490AA5EF66FC26041AD1DC" captionStart="Figures 21–27" captionStartId="F6" captionText="Figures 21–27. Liocanthydrus nanops Baca et al., 2014, instar I 21 left antenna, dorsal aspect 22 right antenna, ventral aspect 23 left mandible, dorsal aspect 24 right maxilla, dorsal aspect 25 left maxilla, ventral aspect 26 labium, dorsal aspect 27 labium, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AN: antenna, LA: labium, MN: mandible, MX: maxilla. Scale bars: 0.04 mm." figureDoi="10.3897/zookeys.1231.144746.figures21-27" httpUri="https://binary.pensoft.net/fig/1279882">21–27</figureCitation>
+, 
+<figureCitation id="D8A84CFFF69DD406D9C4DD9B641D70E5" captionStart="Figures 28–29" captionStartId="F7" captionText="Figures 28–29. Liocanthydrus nanops Baca et al., 2014, instar I 28 left metathoracic leg, anterior aspect 29 right metathoracic leg, posterior aspect. Numbers and lowercase letters indicate primary setae and pores respectively. Solid square indicates additional seta. CO: coxa, FE: femur, PT: pretarsus, TA: tarsus, TI: tibia, TR: trochanter. Scale bar: 0.10 mm." figureDoi="10.3897/zookeys.1231.144746.figures28-29" httpUri="https://binary.pensoft.net/fig/1279883">28–29</figureCitation>
+, 
+<figureCitation id="6D2499A019A2D2B0FAED724BD85099DD" captionStart="Figures 30–33" captionStartId="F8" captionText="Figures 30–33. Liocanthydrus nanops Baca et al., 2014, instar I 30 abdominal segment VIII, dorsal aspect 31 abdominal segment VIII and urogomphi, ventral aspect 32 urogomphi, dorsal aspect 33 urogomphi, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AB: abdominal segment VIII, UR: urogomphus. Scale bars: 0.10 mm (30, 31); 0.02 mm (32, 33)." figureDoi="10.3897/zookeys.1231.144746.figures30-33" httpUri="https://binary.pensoft.net/fig/1279884">30–33</figureCitation>
+, 
+<figureCitation id="FBCC772FC2ABDB51CB69D1F2E889E522" captionStart="Figures 34–37" captionStartId="F9" captionText="Figures 34–37. Liocanthydrus nanops Baca et al., 2014, instar III 34 head, dorsal aspect 35 left prothoracic leg, anterior aspect 36 right prothoracic leg, posterior aspect 37 abdominal segment VIII, dorsal aspect. Numbers and lowercase letters indicate primary setae and pores, respectively (some setae on abdominal segment VIII could not be identified). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1231.144746.figures34-37" httpUri="https://binary.pensoft.net/fig/1279885">34–37</figureCitation>
+, 
+<tableCitation id="BACDC271464360D9BA9AC371AEB5631A" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements and ratios for the larval instars of Liocanthydrus nanops Baca et al. 2014 (n = 1)." httpUri="http://table.plazi.org/id/FA6E299B8663F3E9739748C1C56820F3" tableUuid="FA6E299B8663F3E9739748C1C56820F3">Tables 1</tableCitation>
+, 
+<tableCitation id="6505AF5990C12DB4F0038BBE88A4F2D7" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Number and position of secondary setae on the legs of larvae of Liocanthydrus nanops Baca et al. 2014. Numbers between slash marks refer to pro-, meso-, and metathoracic leg, respectively. A = anterior, PD = posterodorsal, PV = posteroventral; Total = total number of secondary setae on the article (i. e., excluding primary setae) (n = 1)." httpUri="http://table.plazi.org/id/40AE87F8CC844DB08E7B09B2CD312719" tableUuid="40AE87F8CC844DB08E7B09B2CD312719">2</tableCitation>
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0EEPAC" type="material">
+<paragraph id="C9C0A4710210C69BFC110131AB34140A">
+<heading id="A6AB4FDE3CC0E7A6C0A28E5E5D217FDA" reason="title">Material examined.</heading>
+</paragraph>
+<paragraph id="8EC8814B5935D962880103509A911152">
+<materialsCitation id="05F247FE813575E9C2A89F7F2D807C5C" collectingDate="1997-09-27" collectionCode="LEBA" collectorName="Lopez Ruf" country="Argentina" latitude="-25.666666" location="Iguazu National Park" longLatPrecision="1245" longitude="-54.45" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Misiones Province">
+<collectingCountry id="CDFC682C5EB07CAC26E2D0002CD1F3DF" name="Argentina">Argentina</collectingCountry>
+– 
+<collectingRegion id="99BFEE018DFE6569E726D6E37369E2D5">Misiones Province</collectingRegion>
+: 
+<specimenCount id="4B9D39991FF00AF564FD3C3B94B705B1" count="2" type="male">2 ♂</specimenCount>
+and 
+<specimenCount id="496A15DD7416B4F0397F1C3B3A436912" count="4" type="female">4 ♀</specimenCount>
+, 
+<location id="1DB4DFBF662602E3DE5A629BD34FD0A1" LSID="urn:lsid:plazi:treatment:68B12CA5515459339D80CC295A1C0FE0:1DB4DFBF662602E3DE5A629BD34FD0A1" country="Argentina" latitude="-25.666666" longLatPrecision="1245" longitude="-54.45" municipality="Iguazu National Park" name="Iguazu National Park" stateProvince="Misiones Province">Iguazú National Park</location>
+, 
+<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENPAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-54.450000,-25.666667]}">
+<geoCoordinate id="4DD27B228D94BD6FC62F63D08C8F51AE" degrees="25" direction="south" minutes="40" orientation="latitude" precision="925" value="-25.666666">25°40'S</geoCoordinate>
+, 
+<geoCoordinate id="126A8118D474385F8772D8E974833ACA" degrees="54" direction="west" minutes="27" orientation="longitude" precision="925" value="-54.45">54°27'W</geoCoordinate>
+</named-content>
+, 
+<date id="EBB6801E870EA4A21F6CDBC8A040EDFE" value="1997-09-27">
+<collectingDate id="BE4D43BAB8D1DB5D6DBDFFA905BEF967" value="1997-09-27">27. IX. 1997</collectingDate>
+</date>
+, 
+<collectorName id="FF0FDD94125B78EA5C3083C49422584B">López Ruf</collectorName>
+leg. (
+<abbrev id="ABBRID0ESPAC" xlink_title="Laboratory of Entomology, Buenos Aires, Argentina">
+<collectionCode id="AAD156FFA27BBA2BDB63B486CDF286BC">LEBA</collectionCode>
+</abbrev>
+)
+</materialsCitation>
+• 
+<materialsCitation id="DE81CE4D4E4FE473C0E70622703A4D0F" collectingDate="2024-01-09" collectionCode="LEBA" collectorName="Urcola" country="Argentina" county="Somay" elevation="197" latitude="-25.715" location="Daniel &quot; Pupi &quot; Somay bird observatory" longLatPrecision="21" longitude="-54.448334" municipality="Iguazu National Park" specimenCount="20" specimenCount-adult="19" specimenCount-larva="1" stateProvince="Misiones Province">
+<specimenCount id="BD904A6BD699E50973730958F0B9423D" count="1" type="larva">1 larva</specimenCount>
+of instar III and 
+<specimenCount id="365F8ECC37C0A51CCB43F108E201C527" count="19" type="adult">19 adults</specimenCount>
+, 
+<collectingMunicipality id="4C60221B3FD3CE7914EA6619FBFDF4FD">Iguazú National Park</collectingMunicipality>
+, 
+<location id="FDED8E6094C72FD4CA89CD750AA4D53F" LSID="urn:lsid:plazi:treatment:68B12CA5515459339D80CC295A1C0FE0:FDED8E6094C72FD4CA89CD750AA4D53F" country="Argentina" county="Somay" latitude="-25.715" longLatPrecision="21" longitude="-54.448334" municipality="Iguazu National Park" name="Daniel &quot; Pupi &quot; Somay bird observatory">Daniel “Pupi” Somay bird observatory</location>
+, 
+<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZPAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-54.448333,-25.715000]}">
+<geoCoordinate id="5E996FF9D34BEFD9C4DAC84CDDE5E772" degrees="25" direction="south" minutes="42" orientation="latitude" precision="15" seconds="54" value="-25.715">25°42'54&quot;S</geoCoordinate>
+, 
+<geoCoordinate id="17B18D2D01386CFAA644835FA64C5981" degrees="54" direction="west" minutes="26" orientation="longitude" precision="15" seconds="54" value="-54.448334">54°26'54&quot;W</geoCoordinate>
+</named-content>
+, alt. 
+<elevation id="32993FDDBE49F5A0142171D22C547461" metricMagnitude="2" metricUnit="m" metricValue="1.97" unit="m" value="197.0">
+<quantity id="B6F64E781639D96A130B1FDCF1F26FE1" metricMagnitude="2" metricUnit="m" metricValue="1.97" unit="m" value="197.0">197 m</quantity>
+a. s. l.
+</elevation>
+, 
+<date id="F212C777A9F53E2468470CFC8D80E5E6" value="2024-01-09">
+<collectingDate id="32EB3A9AC91A401DB33BC346078E63C3" value="2024-01-09">9. I. 2024</collectingDate>
+</date>
+, 
+<collectorName id="7F843333834E56342D089B295BEEDBA2">Urcola</collectorName>
+leg. (
+<abbrev id="ABBRID0E5PAC" xlink_title="Laboratory of Entomology, Buenos Aires, Argentina">
+<collectionCode id="2EE267C6CEA5D0AB0429B00A97D81529">LEBA</collectionCode>
+</abbrev>
+)
+</materialsCitation>
+• 
+<materialsCitation id="E3887A097488E025A848482DD88137BD" collectingDate="2024-01-10" collectionCode="LEBA" collectorName="Urcola" country="Argentina" county="Somay" elevation="197" latitude="-25.715" location="Iguazu National Park" longLatPrecision="21" longitude="-54.448334" municipality="Daniel" specimenCount="19" specimenCount-adult="18" specimenCount-larva="1" stateProvince="Misiones Province">
+<specimenCount id="25320B861BDDA35AC5F89CFB94F022F3" count="1" type="larva">1 larva</specimenCount>
+of instar I and 
+<specimenCount id="5DEE587CEBB5914E0B8838354CD86938" count="18" type="adult">18 adults</specimenCount>
+, same data except 
+<date id="9F77A97E2FCAF35E1E66055242A652CF" value="2024-01-10">
+<collectingDate id="8F6058C9D97C06EB85B9A98980383745" value="2024-01-10">10. I. 2024</collectingDate>
+</date>
+(
+<abbrev id="ABBRID0EDAAE" xlink_title="Laboratory of Entomology, Buenos Aires, Argentina">
+<collectionCode id="934C9CE0A8A07577AA9936506549B34D">LEBA</collectionCode>
+</abbrev>
+)
+</materialsCitation>
+.
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0EHAAE" type="description">
+<paragraph id="0AAE55F1F1E019F7B6F4CB0AB835B030">
+<heading id="705504AE998B19E2890A35B1006A8512" reason="title">Measurements.</heading>
+</paragraph>
+<paragraph id="A2771B1E38A51503B351D9F6D7CD9473">
+<abbrev id="ABBRID0ENAAE" xlink_title="total length">TL</abbrev>
+= 
+<quantity id="397449F521072B183B0FCFE3B597C55B" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" unit="mm" value="2.95" valueMax="3.1" valueMin="2.8">2.8–3.10 mm</quantity>
+, mean = 
+<quantity id="FD35EB1C949A04E40D2C1EAAFDEB3E69" metricMagnitude="-3" metricUnit="m" metricValue="2.95" unit="mm" value="2.95">2.95 mm</quantity>
+; 
+<abbrev id="ABBRID0ERAAE" xlink_title="greatest width">GW</abbrev>
+= 
+<quantity id="F2E1613F4D7DD474D8E8D111B2729399" metricMagnitude="-3" metricUnit="m" metricValue="1.425" metricValueMax="1.5" metricValueMin="1.35" unit="mm" value="1.425" valueMax="1.5" valueMin="1.35">1.35–1.50 mm</quantity>
+, mean = 
+<quantity id="693A1696D90395733259E86BB19D053C" metricMagnitude="-3" metricUnit="m" metricValue="1.4" unit="mm" value="1.4">1.4 mm</quantity>
+; 
+<abbrev id="ABBRID0EVAAE" xlink_title="total length">TL</abbrev>
+/ 
+<abbrev id="ABBRID0EZAAE" xlink_title="greatest width">GW</abbrev>
+= 2.0– 
+<quantity id="2F1B091D2F5CA89EF067E18FA9C6A8B2" metricMagnitude="-3" metricUnit="m" metricValue="2.2" unit="mm" value="2.2">2.2 mm</quantity>
+, mean = 
+<quantity id="E3862D559734C9F029E4D2E14FD07C80" metricMagnitude="-3" metricUnit="m" metricValue="2.1" unit="mm" value="2.1">2.1 mm</quantity>
+; 
+<abbrev id="ABBRID0E4AAE" xlink_title="greatest width of head">HW</abbrev>
+= 
+<quantity id="89AF9BEC01949AA94AC47C05FC07C23C" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" unit="mm" value="0.8999999999999999" valueMax="0.95" valueMin="0.85">0.85–0.95 mm</quantity>
+, mean = 
+<quantity id="64457BBB4CE243E8201D5C7F3B7B7BC5" metricMagnitude="-4" metricUnit="m" metricValue="9.0" unit="mm" value="0.9">0.9 mm</quantity>
+; 
+<abbrev id="ABBRID0EBBAE" xlink_title="distance between eyes">EW</abbrev>
+= 
+<quantity id="72CF3950A06C3EE063E7B0FDECC01970" metricMagnitude="-4" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" unit="mm" value="0.575" valueMax="0.65" valueMin="0.5">0.5–0.65 mm</quantity>
+, mean = 
+<quantity id="26A9DD7408928BB955DB953574F4DCA8" metricMagnitude="-4" metricUnit="m" metricValue="5.5" unit="mm" value="0.55">0.55 mm</quantity>
+; 
+<abbrev id="ABBRID0EFBAE" xlink_title="greatest width of head">HW</abbrev>
+/ 
+<abbrev id="ABBRID0EJBAE" xlink_title="distance between eyes">EW</abbrev>
+= 
+<quantity id="6FE95168ADB262C1F2CDC2E979CFC0C8" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
+, mean = 
+<quantity id="E44BB4B660B6582CA1F80F8280D6ED8D" metricMagnitude="-3" metricUnit="m" metricValue="1.55" unit="mm" value="1.55">1.55 mm</quantity>
+; PNWant = 0.85–1.0 mm, mean = 
+<quantity id="D35F38D3F51BBF51A871DDF24DF80F28" metricMagnitude="-4" metricUnit="m" metricValue="9.5" unit="mm" value="0.95">0.95 mm</quantity>
+; PNWpost = 
+<quantity id="6C80574BFC574C232ABF576F7BBFAF50" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
+, mean = 
+<quantity id="3546B016F8823538E25AEF5BA9097AE7" metricMagnitude="-3" metricUnit="m" metricValue="1.35" unit="mm" value="1.35">1.35 mm</quantity>
+; PNWpost / PNWant = 
+<quantity id="80572355E5576BC65EEF8C993821763B" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
+, mean = 
+<quantity id="83A7BC44F1319177CBC1EF55347F0755" metricMagnitude="-3" metricUnit="m" metricValue="1.45" unit="mm" value="1.45">1.45 mm</quantity>
+; 
+<abbrev id="ABBRID0ENBAE" xlink_title="total length of prosternum plus noterid platform at midline">
+<collectionCode id="19518A0CC7EEC843172F5C85777A5BC8">TLVP</collectionCode>
+</abbrev>
+= 
+<quantity id="7DE72085FFD6219E1685BF12651A385C" metricMagnitude="-3" metricUnit="m" metricValue="1.225" metricValueMax="1.3" metricValueMin="1.15" unit="mm" value="1.225" valueMax="1.3" valueMin="1.15">1.15–1.3 mm</quantity>
+, mean = 
+<quantity id="3AECB094223D8C12B1D96D57F9C67A98" metricMagnitude="-3" metricUnit="m" metricValue="1.2" unit="mm" value="1.2">1.2 mm</quantity>
+.
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0ERBAE" type="variation">
+<paragraph id="219EA9B480DDC25B425749D74D4E6685">
+<heading id="BC955E4A547A6B973BF2C7ABF24E0AC1" reason="title">Variation in adult morphology.</heading>
+</paragraph>
+<paragraph id="000D88213109D80139C274263D28AE82">
+Similar to what was reported by 
+<bibRefCitation id="19488892B4A47C25B90E6EBFE42FC0FB" DOI="10.11646/zootaxa.3793.2.3" author="Baca SM &amp; Gustafson GT &amp; Toledo M &amp; Miller KB" etAl="et al." firstAuthor="Baca" journalOrPublisher="Zootaxa" pagination="231-246" refId="B1" refString="Baca SM, Gustafson GT, Toledo M, Miller KB (2014) Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species. Zootaxa 3793: 231–246. https://doi.org/10.11646/zootaxa.3793.2.3" title="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species." volume="3793" year="2014">Baca et al. (2014)</bibRefCitation>
+, the specimens examined here show considerable variation in color. Many of the specimens exhibit a yellow head and pronotum, whereas in others these parts are reddish brown. The dark macula on the anteromedial region of the pronotum (Fig. 
+<figureCitation id="A7CBEABFDC479EBDF22633D00E685C41" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1–8. Habitus (1, 2) and genitalia (3–8) of Liocanthydrus nanops Baca et al., 2014, male from Argentina, Misiones Province (LEBA) 1 dorsal aspect 2 ventral aspect 3 segment IX, right lateral aspect 4 right lateral lobe, right lateral aspect 5 left lateral lobe, right lateral aspect 6 median lobe, left lateral aspect 7 median lobe, dorsal aspect 8 median lobe, right lateral aspect. Scale bars: 1.00 mm (1, 2); 0.25 mm (3–8)." figureDoi="10.3897/zookeys.1231.144746.figures1-8" httpUri="https://binary.pensoft.net/fig/1279877">1</figureCitation>
+) is absent in some individuals. Most of the specimens have dark elytra with clearly visible yellow spots (Fig. 
+<figureCitation id="90ECE0122023C5FB8DF6CA64FA75E836" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1–8. Habitus (1, 2) and genitalia (3–8) of Liocanthydrus nanops Baca et al., 2014, male from Argentina, Misiones Province (LEBA) 1 dorsal aspect 2 ventral aspect 3 segment IX, right lateral aspect 4 right lateral lobe, right lateral aspect 5 left lateral lobe, right lateral aspect 6 median lobe, left lateral aspect 7 median lobe, dorsal aspect 8 median lobe, right lateral aspect. Scale bars: 1.00 mm (1, 2); 0.25 mm (3–8)." figureDoi="10.3897/zookeys.1231.144746.figures1-8" httpUri="https://binary.pensoft.net/fig/1279877">1</figureCitation>
+), while in others the elytra are dark reddish brown and the spots are only slightly lighter. The specimens collected in 1997 have a reddish-brown head and pronotum, the latter lacking the dark anteromedial macula, and the elytra are slightly darker with the spots barely visible (similar in color to the head and pronotum). However, examination of the aedeagus revealed the same diagnostic characters as reported by 
+<bibRefCitation id="D9A8A8973964B2821D53369DABE62303" DOI="10.11646/zootaxa.3793.2.3" author="Baca SM &amp; Gustafson GT &amp; Toledo M &amp; Miller KB" etAl="et al." firstAuthor="Baca" journalOrPublisher="Zootaxa" pagination="231-246" refId="B1" refString="Baca SM, Gustafson GT, Toledo M, Miller KB (2014) Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species. Zootaxa 3793: 231–246. https://doi.org/10.11646/zootaxa.3793.2.3" title="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species." volume="3793" year="2014">Baca et al. (2014)</bibRefCitation>
+and the other specimens collected here. Finally, the 
+<specimenCount id="6622FFA0F03567E903A7E8058CF3DC02" count="2" type="male">two males</specimenCount>
+collected in 1997 have the anterodorsal margin of sternite IX widened, similar to that reported by 
+<bibRefCitation id="589443864FE3B7709EF84C511EEA7550" author="Grosso LE" firstAuthor="Grosso" refId="B5" refString="Grosso LE (1979) Contribución al conocimiento biológico y sistemático de las especies argentinas de Noteridae (Coleoptera, Adephaga). Doctoral thesis N ° 374, Universidad Nacional de La Plata, La Plata, Argentina." year="1979">Grosso (1979)</bibRefCitation>
+for specimens identified as 
+<taxonomicName id="2BC6503BFF6789526B207AF6E531F7DA" authorityName="Zimmermann" authorityYear="1921" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="octoguttatus">
+<emphasis id="E9B8AD6C349E49C8E9141389EC807709" italics="true">L. octoguttatus</emphasis>
+</taxonomicName>
+. However, we observed that this character is variable in the rest of the material examined, with this widening absent in most specimens (Fig. 
+<figureCitation id="36FAB50658750439FB366BB6AE19C5A9" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1–8. Habitus (1, 2) and genitalia (3–8) of Liocanthydrus nanops Baca et al., 2014, male from Argentina, Misiones Province (LEBA) 1 dorsal aspect 2 ventral aspect 3 segment IX, right lateral aspect 4 right lateral lobe, right lateral aspect 5 left lateral lobe, right lateral aspect 6 median lobe, left lateral aspect 7 median lobe, dorsal aspect 8 median lobe, right lateral aspect. Scale bars: 1.00 mm (1, 2); 0.25 mm (3–8)." figureDoi="10.3897/zookeys.1231.144746.figures1-8" httpUri="https://binary.pensoft.net/fig/1279877">3</figureCitation>
+).
+</paragraph>
+<caption id="B065D1A81AEDF78E29B9770DABEB3CBD" ID-DOI="10.3897/zookeys.1231.144746.figures1-8" ID-arpha="AAAE4736-27C8-555D-BF3C-E48EBE93BD49" httpUri="https://binary.pensoft.net/fig/1279877" startId="F1">
+<paragraph id="622564AC9D0B0478C5ABA5D46DD0B9FB">
+<label id="98CB19BF5B7664A1B1125D528597992F">Figures 1–8.</label>
+</paragraph>
+<paragraph id="DE2797C62398E073DCB93BA57AA49344">
+Habitus (
+<emphasis id="9176EC202EE647B08A69D64DA4113026" bold="true">1, 2</emphasis>
+) and genitalia (
+<emphasis id="145C10E670D7999016825610EBE8DA66" bold="true">3–8</emphasis>
+) of 
+<taxonomicName id="F2DEA346CCFC104246583E79D5A3352E" authority="Baca et al., 2014" authorityName="Baca" authorityYear="2014" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="BCFA695BB919B3D88279CA6D3648067F" italics="true">Liocanthydrus nanops</emphasis>
+<bibRefCitation id="BA57BE1EAAE87CD3766EBE52D8141399" DOI="10.11646/zootaxa.3793.2.3" author="Baca" etAl="et al." firstAuthor="Baca" journalOrPublisher="Zootaxa" pagination="231-246" refId="B1" refString="Baca SM, Gustafson GT, Toledo M, Miller KB (2014) Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species. Zootaxa 3793: 231–246. https://doi.org/10.11646/zootaxa.3793.2.3" title="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species." volume="3793" year="2014">Baca et al., 2014</bibRefCitation>
+</taxonomicName>
+, male from Argentina, Misiones Province (
+<abbrev id="ABBRID0EVDAE" xlink_title="Laboratory of Entomology, Buenos Aires, Argentina">LEBA</abbrev>
+) 
+<emphasis id="29BDFF23FE4B6138E84CFAF0BEFF02D7" bold="true">1</emphasis>
+dorsal aspect 
+<emphasis id="817EF608705FF47003BCA076F53303A1" bold="true">2</emphasis>
+ventral aspect 
+<emphasis id="CB5BA7079496A676C73D8BAE9EB362D8" bold="true">3</emphasis>
+segment IX, right lateral aspect 
+<emphasis id="05DEDF1F89C51B3BDB2677F7462912C3" bold="true">4</emphasis>
+right lateral lobe, right lateral aspect 
+<emphasis id="01FC998425D7CAD9887EF03DF79DD8B0" bold="true">5</emphasis>
+left lateral lobe, right lateral aspect 
+<emphasis id="D680B6486704AE811375A445137A33C9" bold="true">6</emphasis>
+median lobe, left lateral aspect 
+<emphasis id="562CBFECF0474EBC2791237765575CD8" bold="true">7</emphasis>
+median lobe, dorsal aspect 
+<emphasis id="978912F8DE6A7A423E22B4DE33464ADC" bold="true">8</emphasis>
+median lobe, right lateral aspect. Scale bars: 1.00 mm (
+<emphasis id="BE1C6A0A828D053FE1E6D2ED76B65BD1" bold="true">1, 2</emphasis>
+); 0.25 mm (
+<emphasis id="A0ABC3ACE76071872F2A59B75DE8A279" bold="true">3–8</emphasis>
+).
+</paragraph>
+</caption>
+<caption id="AAB167CF077C6AC70AB491B743775B81" ID-DOI="10.3897/zookeys.1231.144746.figures9-12" ID-arpha="5FF18595-B32D-5FE0-9527-C1DFA6160C31" httpUri="https://binary.pensoft.net/fig/1279878" startId="F2">
+<paragraph id="575D6A086FE949D505E9783FBAC7101D">
+<label id="18C8C4E3FDD2F33930B138463913382B">Figures 9–12.</label>
+</paragraph>
+<paragraph id="6A4936264B857A83FB5CA0789FBDDE33">
+Male genitalia of 
+<taxonomicName id="FC0A359F69640FEBF24C8CDF27E5E83B" authority="Baca et al., 2014" authorityName="Baca" authorityYear="2014" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="4EE1861AFF9F8A661A41C212B8DE46DE" italics="true">Liocanthydrus nanops</emphasis>
+<bibRefCitation id="196BD0D2B7043F2E18BAE49766A3C792" DOI="10.11646/zootaxa.3793.2.3" author="Baca" etAl="et al." firstAuthor="Baca" journalOrPublisher="Zootaxa" pagination="231-246" refId="B1" refString="Baca SM, Gustafson GT, Toledo M, Miller KB (2014) Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species. Zootaxa 3793: 231–246. https://doi.org/10.11646/zootaxa.3793.2.3" title="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species." volume="3793" year="2014">Baca et al., 2014</bibRefCitation>
+</taxonomicName>
+, modified from drawings in 
+<bibRefCitation id="E55600C33869DB5358D7360E96ABB1F9" author="Grosso LE" firstAuthor="Grosso" refId="B5" refString="Grosso LE (1979) Contribución al conocimiento biológico y sistemático de las especies argentinas de Noteridae (Coleoptera, Adephaga). Doctoral thesis N ° 374, Universidad Nacional de La Plata, La Plata, Argentina." year="1979">Grosso (1979)</bibRefCitation>
+<emphasis id="CEFCFA4D81467F6476A957B35548FD05" bold="true">9</emphasis>
+segment IX, right lateral aspect 
+<emphasis id="9D209993FD3A37EE4C23E803B59E37C1" bold="true">10</emphasis>
+right lateral lobe, right lateral aspect 
+<emphasis id="32CEAD40101D3FCBAEDAA3DB3E8B8F45" bold="true">11</emphasis>
+left lateral lobe, right lateral aspect 
+<emphasis id="5E2287321F2D76D7CA657A3CC34298BE" bold="true">12</emphasis>
+median lobe, right lateral aspect. Scale bar: 0.5 mm.
+</paragraph>
+</caption>
+<caption id="A3327379BB958AC525DB6BF4E6AB61D0" ID-DOI="10.3897/zookeys.1231.144746.figures13-16" ID-arpha="7DF973BB-D9EB-5938-9844-6AC1D1682126" httpUri="https://binary.pensoft.net/fig/1279879" startId="F3">
+<paragraph id="8BDEA69243F548E1648D30AECD0840C1">
+<label id="B35C77C1A63781D4F4358F706440A77D">Figures 13–16.</label>
+</paragraph>
+<paragraph id="2D2C9CF0177DEFB7D28677DD1177285B">
+Habitat of 
+<taxonomicName id="A431089127C0AA0DE7169C9B6AD2F140" authority="Baca et al., 2014" authorityName="Baca" authorityYear="2014" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="C0D96F10919DA0F2A98CAB402988EC0D" italics="true">Liocanthydrus nanops</emphasis>
+<bibRefCitation id="4ED80F08AB4B58D010EED9A2EBA4D91B" DOI="10.11646/zootaxa.3793.2.3" author="Baca" etAl="et al." firstAuthor="Baca" journalOrPublisher="Zootaxa" pagination="231-246" refId="B1" refString="Baca SM, Gustafson GT, Toledo M, Miller KB (2014) Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species. Zootaxa 3793: 231–246. https://doi.org/10.11646/zootaxa.3793.2.3" title="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species." volume="3793" year="2014">Baca et al., 2014</bibRefCitation>
+</taxonomicName>
+, Iguazú NP, Misiones Province, Argentina 
+<emphasis id="5A5796C00BD05A263DCF84D18D698768" bold="true">13</emphasis>
+general view of the stream’s flooding site 
+<emphasis id="C371FCC41DB74B4D5BE7A5A5C17FC588" bold="true">14</emphasis>
+collecting of beetles 
+<emphasis id="B5C53A6D50B09BBDBAD58472FA76266F" bold="true">15</emphasis>
+details of the stream 
+<emphasis id="56752133A300427AC7F15F15BB10D267" bold="true">16</emphasis>
+instar III larva of 
+<taxonomicName id="07102BF26ED77D17ABA1E71F7AE8D8D6" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="7F1216CCFEDA6B102732B9E45D008738" italics="true">L. nanops</emphasis>
+</taxonomicName>
+. Scale bar: 0.5 mm.
+</paragraph>
+</caption>
+</subSubSection>
+<subSubSection id="SECID0EJHAE" type="remarks">
+<paragraph id="DFD5973D0F7031AF1759027AB51E9FD7">
+<heading id="77CF13EC7DF4E0FC11F4A493CED578BA" reason="title">Remarks.</heading>
+</paragraph>
+<paragraph id="0B0BFE331BF25F14FC1DD1274B2CB20E">
+Based on 
+<bibRefCitation id="196998EEB809CD41ECC12202BB80B61F" author="Grosso LE" firstAuthor="Grosso's" refId="B5" refString="Grosso LE (1979) Contribución al conocimiento biológico y sistemático de las especies argentinas de Noteridae (Coleoptera, Adephaga). Doctoral thesis N ° 374, Universidad Nacional de La Plata, La Plata, Argentina." year="1979">Grosso’s (1979)</bibRefCitation>
+redescription and drawings of the male genitalia of the specimens alleged to be 
+<taxonomicName id="12A90CD7EB2B0F2A5AC4A805B6184888" authorityName="Zimmermann" authorityYear="1921" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="octoguttatus">
+<emphasis id="5D51C55924C5419062167F10B65BB8A9" italics="true">L. octoguttatus</emphasis>
+</taxonomicName>
+(Figs 
+<figureCitation id="FCBB4D8B7A0A6B799AD71A99CF74EAC7" captionStart="Figures 9–12" captionStartId="F2" captionText="Figures 9–12. Male genitalia of Liocanthydrus nanops Baca et al., 2014, modified from drawings in Grosso (1979) 9 segment IX, right lateral aspect 10 right lateral lobe, right lateral aspect 11 left lateral lobe, right lateral aspect 12 median lobe, right lateral aspect. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1231.144746.figures9-12" httpUri="https://binary.pensoft.net/fig/1279878">9–12</figureCitation>
+, modified from original drawings), we can observe that the left lateral lobe has a well-projected distal angle (Fig. 
+<figureCitation id="5EEB4322D5505F7C61CB9A4031A04C51" captionStart="Figures 9–12" captionStartId="F2" captionText="Figures 9–12. Male genitalia of Liocanthydrus nanops Baca et al., 2014, modified from drawings in Grosso (1979) 9 segment IX, right lateral aspect 10 right lateral lobe, right lateral aspect 11 left lateral lobe, right lateral aspect 12 median lobe, right lateral aspect. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1231.144746.figures9-12" httpUri="https://binary.pensoft.net/fig/1279878">11</figureCitation>
+). The only known species of 
+<taxonomicName id="E9B6433C0B36974DBF086D7564FDD7C8" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
+<emphasis id="7575795C62503B07C0B8DC930F01DF11" italics="true">Liocanthydrus</emphasis>
+</taxonomicName>
+in which the left lateral lobe has this shape is 
+<taxonomicName id="08C280804B27631091229647BC058F94" authority="(Baca et al. 2014)" baseAuthorityName="Baca" baseAuthorityYear="2014" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="8F4E2E1B0BBB3CBE979D341DB5EB7DCC" italics="true">L nanops</emphasis>
+(
+<bibRefCitation id="BBE1FB9AD2E13676581A4CDBA378CAFF" DOI="10.11646/zootaxa.3793.2.3" author="Baca" etAl="et al." firstAuthor="Baca" journalOrPublisher="Zootaxa" pagination="231-246" refId="B1" refString="Baca SM, Gustafson GT, Toledo M, Miller KB (2014) Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species. Zootaxa 3793: 231–246. https://doi.org/10.11646/zootaxa.3793.2.3" title="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species." volume="3793" year="2014">Baca et al. 2014</bibRefCitation>
+)
+</taxonomicName>
+(Fig. 
+<figureCitation id="093681C2EDF3103E8287BCA754E1AA96" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1–8. Habitus (1, 2) and genitalia (3–8) of Liocanthydrus nanops Baca et al., 2014, male from Argentina, Misiones Province (LEBA) 1 dorsal aspect 2 ventral aspect 3 segment IX, right lateral aspect 4 right lateral lobe, right lateral aspect 5 left lateral lobe, right lateral aspect 6 median lobe, left lateral aspect 7 median lobe, dorsal aspect 8 median lobe, right lateral aspect. Scale bars: 1.00 mm (1, 2); 0.25 mm (3–8)." figureDoi="10.3897/zookeys.1231.144746.figures1-8" httpUri="https://binary.pensoft.net/fig/1279877">5</figureCitation>
+), which is also the only species distributed near the area where the specimens studied by Grosso were collected (Fig. 
+<figureCitation id="42B9C5539A29DBE5AC262FBFA682CBAA" captionStart="Figure 17" captionStartId="F4" captionText="Figure 17. Known distributional data for the southernmost Liocanthydrus species with references of records." figureDoi="10.3897/zookeys.1231.144746.figure17" httpUri="https://binary.pensoft.net/fig/1279880">17</figureCitation>
+). This evidence, summed to the shape of the apex of the median lobe in lateral view (Fig. 
+<figureCitation id="ED84590FCB5150B09D7A92D311CC86F0" captionStart="Figures 9–12" captionStartId="F2" captionText="Figures 9–12. Male genitalia of Liocanthydrus nanops Baca et al., 2014, modified from drawings in Grosso (1979) 9 segment IX, right lateral aspect 10 right lateral lobe, right lateral aspect 11 left lateral lobe, right lateral aspect 12 median lobe, right lateral aspect. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1231.144746.figures9-12" httpUri="https://binary.pensoft.net/fig/1279878">12</figureCitation>
+) led us to conclude that the specimens studied by Grosso belong to 
+<taxonomicName id="918BE751364C85A521FF149DCB64BB8A" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="72EA429F6921225F21A4C220C1908DEB" italics="true">L. nanops</emphasis>
+</taxonomicName>
+. It should be noted that, when Grosso identified his material, 
+<taxonomicName id="D4F4684908E8331CFDC4EF539EF738B6" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="09C4EA7B826D63B94515A9E31AD8A0A5" italics="true">L. nanops</emphasis>
+</taxonomicName>
+had not yet been described. Grosso likely relied on 
+<bibRefCitation id="D18D8AF36443AFC14B84373C668FFEE5" author="Zimmermann A" firstAuthor="Zimmermann's" journalOrPublisher="Archiv für Naturgeschichte" pagination="181-206" refId="B16" refString="Zimmermann A (1921) Beiträge zur Kenntnis der südamerikanischen Schwimmkäferfauna nebst 41 Neubeschreibungen. Archiv für Naturgeschichte 87: 181–206." title="Beiträge zur Kenntnis der südamerikanischen Schwimmkäferfauna nebst 41 Neubeschreibungen." volume="87" year="1921">Zimmermann’s (1921)</bibRefCitation>
+treatment of 
+<taxonomicName id="40E9BA00E71C672096C7C50D62875B84" authorityName="Zimmermann" authorityYear="1921" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="octoguttatus">
+<emphasis id="D359F5C3EB2B1435698C1F140172AA2C" italics="true">L. octoguttatus</emphasis>
+</taxonomicName>
+, which lacks a description of the male genitalia (a crucial feature to recognize that it belongs to a distinct species given the very similar external appearance of both species).
+</paragraph>
+<caption id="07CF707268E4CC581585C0087BC06FDF" ID-DOI="10.3897/zookeys.1231.144746.figure17" ID-arpha="D5609982-AD52-548F-A1E0-685C965CE054" httpUri="https://binary.pensoft.net/fig/1279880" startId="F4">
+<paragraph id="E9537DF05554E0CD7E43B99D66781BDA">
+<label id="BC648098EA3E0AF42AC7556DB28ED516">Figure 17.</label>
+</paragraph>
+<paragraph id="0DD56588FA825628C04ADB9E133563AE">
+Known distributional data for the southernmost 
+<taxonomicName id="C7A43B6DDCC19786EB7F660D366D0617">
+<emphasis id="55B99903CB0D8283052B6F897F269997" italics="true">Liocanthydrus</emphasis>
+species
+</taxonomicName>
+with references of records.
+</paragraph>
+</caption>
+<caption id="4F1EAC8C4AFBBB9CBB9AB761DFFDE92B" ID-DOI="10.3897/zookeys.1231.144746.figures18-20" ID-arpha="1626F16B-0C2E-502A-A0C3-9A136DF43148" httpUri="https://binary.pensoft.net/fig/1279881" startId="F5">
+<paragraph id="02DE08C1E838857CD2B0B987EA508E7F">
+<label id="A3B2BE488F3A27DE31FE3B881E550215">Figures 18–20.</label>
+</paragraph>
+<paragraph id="6C39973117582997C7863C3B3F86EA0C">
+<taxonomicName id="A1C68D80542C30433C962242C4B68EDB" authority="Baca et al., 2014" authorityName="Baca" authorityYear="2014" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="311B038F848B99CDC65085913459F18F" italics="true">Liocanthydrus nanops</emphasis>
+<bibRefCitation id="5575522B3D321E758294BA52A5193D49" DOI="10.11646/zootaxa.3793.2.3" author="Baca" etAl="et al." firstAuthor="Baca" journalOrPublisher="Zootaxa" pagination="231-246" refId="B1" refString="Baca SM, Gustafson GT, Toledo M, Miller KB (2014) Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species. Zootaxa 3793: 231–246. https://doi.org/10.11646/zootaxa.3793.2.3" title="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species." volume="3793" year="2014">Baca et al., 2014</bibRefCitation>
+</taxonomicName>
+, instar I 
+<emphasis id="27459805918E51776A517EB8DE25D8F1" bold="true">18</emphasis>
+habitus, dorsal aspect 
+<emphasis id="EAAEC48F0C60B5BE8C5A4B836B697E57" bold="true">19</emphasis>
+cephalic capsule, dorsal aspect 
+<emphasis id="AED362E4D4270962ABEA97B0F0FFA5FC" bold="true">20</emphasis>
+cephalic capsule, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. Solid square indicates additional seta. EB: egg burster, FR: frontoclypeus, PA: parietal, TP: tentorial pit. Scale bars: 0.50 mm (
+<emphasis id="109C9B59256E0C6732751674C33A67D8" bold="true">18</emphasis>
+); 0.10 mm (
+<emphasis id="D185437FE26F3A059469A7A77E319626" bold="true">19, 20</emphasis>
+).
+</paragraph>
+</caption>
+<caption id="31C89086F54E71C98AB2EA3C08C2B7A2" ID-DOI="10.3897/zookeys.1231.144746.figures21-27" ID-arpha="795BBADD-F555-5900-8481-02DC60388C68" httpUri="https://binary.pensoft.net/fig/1279882" startId="F6">
+<paragraph id="BE43B94CBE9B945AB7C48716F625D775">
+<label id="2ADB849A626DFBBABBE84D4C5F49B280">Figures 21–27.</label>
+</paragraph>
+<paragraph id="7C7CB5AC58CBFA1E8914639BE52046EA">
+<taxonomicName id="3BD75508F45198DA027CE7E41CA41707" authority="Baca et al., 2014" authorityName="Baca" authorityYear="2014" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="5665F9914077E7686B13CB01243758F5" italics="true">Liocanthydrus nanops</emphasis>
+<bibRefCitation id="B7B15AA98ABFE04B20B7C8454F00C9FC" DOI="10.11646/zootaxa.3793.2.3" author="Baca" etAl="et al." firstAuthor="Baca" journalOrPublisher="Zootaxa" pagination="231-246" refId="B1" refString="Baca SM, Gustafson GT, Toledo M, Miller KB (2014) Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species. Zootaxa 3793: 231–246. https://doi.org/10.11646/zootaxa.3793.2.3" title="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species." volume="3793" year="2014">Baca et al., 2014</bibRefCitation>
+</taxonomicName>
+, instar I 
+<emphasis id="6936B048991AFC2D0C8967CCE6BB5B03" bold="true">21</emphasis>
+left antenna, dorsal aspect 
+<emphasis id="8D886FCEAAA29F7D75EE22F9AA7DA80D" bold="true">22</emphasis>
+right antenna, ventral aspect 
+<emphasis id="D33A998EFFADBA01C93295B6B18D099C" bold="true">23</emphasis>
+left mandible, dorsal aspect 
+<emphasis id="769C578BF25E16E420C1A9403A6864C9" bold="true">24</emphasis>
+right maxilla, dorsal aspect 
+<emphasis id="F54ACF523D8B5EF533137C047EEFC607" bold="true">25</emphasis>
+left maxilla, ventral aspect 
+<emphasis id="482A3F2E2A61FB3EBBE9F77A14500FE2" bold="true">26</emphasis>
+labium, dorsal aspect 
+<emphasis id="598DB67D165C87F71CCD7EFBAA384445" bold="true">27</emphasis>
+labium, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AN: antenna, LA: labium, MN: mandible, MX: maxilla. Scale bars: 0.04 mm.
+</paragraph>
+</caption>
+<caption id="013867C358B3188922760F6169D33417" ID-DOI="10.3897/zookeys.1231.144746.figures28-29" ID-arpha="D5CDE0BC-0DEB-594D-B473-05633538B349" httpUri="https://binary.pensoft.net/fig/1279883" startId="F7">
+<paragraph id="33517E76F2F6652AD2E0162FB6C9F95A">
+<label id="60FF319CDA7E7739EB6956C02E5D965A">Figures 28–29.</label>
+</paragraph>
+<paragraph id="3365937C11056844B973E7096885E687">
+<taxonomicName id="46E95BE06AE81B31696394E9AE5C483A" authority="Baca et al., 2014" authorityName="Baca" authorityYear="2014" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="F3500B3F614CA76C9B69D5CA79BCDF3C" italics="true">Liocanthydrus nanops</emphasis>
+<bibRefCitation id="63BB347454D292E41546F79916534E04" DOI="10.11646/zootaxa.3793.2.3" author="Baca" etAl="et al." firstAuthor="Baca" journalOrPublisher="Zootaxa" pagination="231-246" refId="B1" refString="Baca SM, Gustafson GT, Toledo M, Miller KB (2014) Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species. Zootaxa 3793: 231–246. https://doi.org/10.11646/zootaxa.3793.2.3" title="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species." volume="3793" year="2014">Baca et al., 2014</bibRefCitation>
+</taxonomicName>
+, instar I 
+<emphasis id="845B31663D4B0DD63B3679AA2F9F1437" bold="true">28</emphasis>
+left metathoracic leg, anterior aspect 
+<emphasis id="46233FED6A4197B87640FB5D8796BD7D" bold="true">29</emphasis>
+right metathoracic leg, posterior aspect. Numbers and lowercase letters indicate primary setae and pores respectively. Solid square indicates additional seta. CO: coxa, FE: femur, PT: pretarsus, TA: tarsus, TI: tibia, TR: trochanter. Scale bar: 0.10 mm.
+</paragraph>
+</caption>
+</subSubSection>
+<subSubSection id="SECID0EROAE" type="habitat">
+<paragraph id="BCE21AFAC20F9F744E3C3C9BD9FDC594">
+<heading id="31637EE5D818A9EC5E8C1CBDAF830B17" reason="title">Habitat and co-occurring taxa.</heading>
+</paragraph>
+<paragraph id="FC759C9ED6FF7587D11C00D7E79D1680">
+Adults and larvae of 
+<taxonomicName id="7ED9062D08CB74215B461CADA9DAED63" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="D221780FA0AC16E804F0F96148F926F6" italics="true">L. nanops</emphasis>
+</taxonomicName>
+were collected in a stream floodplain with other noterid species: 
+<taxonomicName id="C84028017FA651FE1CF6EC90A0AE024E" authority="Sahlberg, 1844" authorityName="Sahlberg" authorityYear="1844" class="Insecta" family="Noteridae" genus="Hydrocanthus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="socius">
+<emphasis id="9D7039507B3A4D16A3E79C002EF3D2E9" italics="true">Hydrocanthus socius</emphasis>
+Sahlberg, 1844
+</taxonomicName>
+, 
+<taxonomicName id="1E3DFD47ACCE1118ED9E7BFB0979D1D6" authority="(Regimbart, 1889)" baseAuthorityName="Regimbart" baseAuthorityYear="1889" class="Insecta" family="Noteridae" genus="Suphisellus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="balzani">
+<emphasis id="474934A2FB2B4A543319B9708AC79875" italics="true">Suphisellus balzani</emphasis>
+(Régimbart, 1889)
+</taxonomicName>
+, and 
+<taxonomicName id="0195B6FAB2DA6835CB21675BAA48D4E9" authority="(Sharp, 1882)" baseAuthorityName="Sharp" baseAuthorityYear="1882" class="Insecta" family="Noteridae" genus="Suphisellus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rufipes">
+<emphasis id="D3661A16713B567871FFB64528472C92" italics="true">S. rufipes</emphasis>
+(Sharp, 1882)
+</taxonomicName>
+. The sampling site was mostly exposed to sunlight, had a muddy bottom, shallow depth, slow current, cool water, and abundant emergent vegetation (Figs 
+<figureCitation id="E757B4298A25E46B529E3F5D43D40511" captionStart="Figures 13–16" captionStartId="F3" captionText="Figures 13–16. Habitat of Liocanthydrus nanops Baca et al., 2014, Iguazú NP, Misiones Province, Argentina 13 general view of the stream’s flooding site 14 collecting of beetles 15 details of the stream 16 instar III larva of L. nanops. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1231.144746.figures13-16" httpUri="https://binary.pensoft.net/fig/1279879">13–15</figureCitation>
+).
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0EHQAE" type="distribution">
+<paragraph id="DB78BD8ABCC969B94113173BD9511AB0">
+<heading id="1ABA8F89BBC2547B9923FCF39AD6E282" reason="title">Distribution.</heading>
+</paragraph>
+<paragraph id="743F916F087ED2B7367739DA370AEA42">
+<collectingCountry id="43537DCCA3F9EB11F2406BB4D83800C0" name="Argentina">Argentina</collectingCountry>
+(
+<collectingCountry id="6050DC8B00F2032952F0EEBB2E6E285C" name="Taiwan">Formosa</collectingCountry>
+, Misiones) (new record), 
+<collectingCountry id="C8928386C57020CF103E5DB76E10C772" name="Brazil">Brazil</collectingCountry>
+, and 
+<collectingCountry id="E5EA45FC0BF0316242523DDA149D9936" name="Paraguay">Paraguay</collectingCountry>
+(Fig. 
+<figureCitation id="A00279F779B2CF4F034A82AA6E2E9BDB" captionStart="Figure 17" captionStartId="F4" captionText="Figure 17. Known distributional data for the southernmost Liocanthydrus species with references of records." figureDoi="10.3897/zookeys.1231.144746.figure17" httpUri="https://binary.pensoft.net/fig/1279880">17</figureCitation>
+).
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0ERQAE" type="description">
+<paragraph id="73D304619FC23764B29F5EFECC2C729E">
+<heading id="AB7C815A32B1A8856CCB2782862A2524" reason="title">Description of larva.</heading>
+</paragraph>
+<paragraph id="083ABCFD3FADB6EEE2218555774E4791">
+<emphasis id="C2DBD5866BCB195E2DE07217F7F03196" bold="true">Instar I</emphasis>
+(Figs 
+<figureCitation id="2BAD3A1603BDD4478494A8C310C35CFD" captionStart="Figures 18–20" captionStartId="F5" captionText="Figures 18–20. Liocanthydrus nanops Baca et al., 2014, instar I 18 habitus, dorsal aspect 19 cephalic capsule, dorsal aspect 20 cephalic capsule, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. Solid square indicates additional seta. EB: egg burster, FR: frontoclypeus, PA: parietal, TP: tentorial pit. Scale bars: 0.50 mm (18); 0.10 mm (19, 20)." figureDoi="10.3897/zookeys.1231.144746.figures18-20" httpUri="https://binary.pensoft.net/fig/1279881">18</figureCitation>
+– 
+<figureCitation id="3D40022C6627CBA59FDF3B5FE593C890" captionStart="Figures 30–33" captionStartId="F8" captionText="Figures 30–33. Liocanthydrus nanops Baca et al., 2014, instar I 30 abdominal segment VIII, dorsal aspect 31 abdominal segment VIII and urogomphi, ventral aspect 32 urogomphi, dorsal aspect 33 urogomphi, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AB: abdominal segment VIII, UR: urogomphus. Scale bars: 0.10 mm (30, 31); 0.02 mm (32, 33)." figureDoi="10.3897/zookeys.1231.144746.figures30-33" httpUri="https://binary.pensoft.net/fig/1279884">33</figureCitation>
+).
+</paragraph>
+<caption id="3B2A4C805F1976AE46F1D809E581B4E9" ID-DOI="10.3897/zookeys.1231.144746.figures30-33" ID-arpha="D26D065B-814A-59B0-9F8F-62714FFC6BDA" httpUri="https://binary.pensoft.net/fig/1279884" startId="F8">
+<paragraph id="A939C4EA97B2714B533911DB019EE969">
+<label id="498E3AC254DFEB54B2548B35C9EF41DE">Figures 30–33.</label>
+</paragraph>
+<paragraph id="A672CE07A88D1418484436B881FFFC96">
+<taxonomicName id="ADCA5845A681988C27AFA914E6D52805" authority="Baca et al., 2014" authorityName="Baca" authorityYear="2014" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="0D0F36F7414B7FEFDED8AD37944C335C" italics="true">Liocanthydrus nanops</emphasis>
+<bibRefCitation id="877CFCFAFC0E16D62742F4BCDB453217" DOI="10.11646/zootaxa.3793.2.3" author="Baca" etAl="et al." firstAuthor="Baca" journalOrPublisher="Zootaxa" pagination="231-246" refId="B1" refString="Baca SM, Gustafson GT, Toledo M, Miller KB (2014) Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species. Zootaxa 3793: 231–246. https://doi.org/10.11646/zootaxa.3793.2.3" title="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species." volume="3793" year="2014">Baca et al., 2014</bibRefCitation>
+</taxonomicName>
+, instar I 
+<emphasis id="1AA0CBC77A0AEFA68FA26C632E3FC1EE" bold="true">30</emphasis>
+abdominal segment VIII, dorsal aspect 
+<emphasis id="5C060EB05EE4595E7B4F140DB812FDD9" bold="true">31</emphasis>
+abdominal segment VIII and urogomphi, ventral aspect 
+<emphasis id="93537110F36D871BB58C670D7695DC09" bold="true">32</emphasis>
+urogomphi, dorsal aspect 
+<emphasis id="A27DFA5F382F1833449EE21032638633" bold="true">33</emphasis>
+urogomphi, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AB: abdominal segment VIII, UR: urogomphus. Scale bars: 0.10 mm (
+<emphasis id="C69987DCD1936F7611B18B816F68AD83" bold="true">30, 31</emphasis>
+); 0.02 mm (
+<emphasis id="E8932FADEEC1D92D8A9AB8A2504DEC58" bold="true">32, 33</emphasis>
+).
+</paragraph>
+</caption>
+<paragraph id="4DF4DBC3FB7ABA647530DC154FE104EE">
+<emphasis id="DA7412AB696C1661ECC2AE32D218C0B4" bold="true">
+<emphasis id="5D7BEEF63D500DA458C9955F1A0F0AA5" bold="true" italics="true">Color</emphasis>
+.
+</emphasis>
+Entirely testaceous.
+</paragraph>
+<paragraph id="DAD2B1A0953B74A8F786FBDB922A7798">
+<emphasis id="2DBA3B584F7336081349D22DEB075732" bold="true">
+<emphasis id="525FCA764BB3080A2B0409EB29C9C762" bold="true" italics="true">Body</emphasis>
+.
+</emphasis>
+Elongate, nearly parallel sided (Fig. 
+<figureCitation id="094F047B16E4525CB1BACD8D5CB17ADF" captionStart="Figures 18–20" captionStartId="F5" captionText="Figures 18–20. Liocanthydrus nanops Baca et al., 2014, instar I 18 habitus, dorsal aspect 19 cephalic capsule, dorsal aspect 20 cephalic capsule, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. Solid square indicates additional seta. EB: egg burster, FR: frontoclypeus, PA: parietal, TP: tentorial pit. Scale bars: 0.50 mm (18); 0.10 mm (19, 20)." figureDoi="10.3897/zookeys.1231.144746.figures18-20" httpUri="https://binary.pensoft.net/fig/1279881">18</figureCitation>
+). Measurements and ratios that characterize the body shape are given in Table 
+<tableCitation id="BC06037A4546E764EDA99DCF14E7292F" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements and ratios for the larval instars of Liocanthydrus nanops Baca et al. 2014 (n = 1)." httpUri="http://table.plazi.org/id/FA6E299B8663F3E9739748C1C56820F3" tableUuid="FA6E299B8663F3E9739748C1C56820F3">1</tableCitation>
+.
+</paragraph>
+<caption id="FA6E299B8663F3E9739748C1C56820F3" ID-Table-UUID="FA6E299B8663F3E9739748C1C56820F3" httpUri="http://table.plazi.org/id/FA6E299B8663F3E9739748C1C56820F3" startId="T1" targetIsTable="true">
+<paragraph id="2526C23EB3FD4BB315DEF8EE4B0AFDD7">
+<label id="E0966C5748C77D2C5A4A08289026333E">Table 1.</label>
+</paragraph>
+<paragraph id="20DEC41F6246704BBE32E760B3BD0210">
+Measurements and ratios for the larval instars of 
+<taxonomicName id="C4AA0E2BB7A9D500E3B1A6485AAD7D26" authority="Baca et al. 2014" authorityName="Baca" authorityYear="2014" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="1882F985242D206AB3F0C48F671F9BA9" italics="true">Liocanthydrus nanops</emphasis>
+<bibRefCitation id="118180BE6CCBE0165D0D310ADCED52D0" DOI="10.11646/zootaxa.3793.2.3" author="Baca" etAl="et al." firstAuthor="Baca" journalOrPublisher="Zootaxa" pagination="231-246" refId="B1" refString="Baca SM, Gustafson GT, Toledo M, Miller KB (2014) Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species. Zootaxa 3793: 231–246. https://doi.org/10.11646/zootaxa.3793.2.3" title="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species." volume="3793" year="2014">Baca et al. 2014</bibRefCitation>
+</taxonomicName>
+(
+<emphasis id="8B915723F6E8D9C2DF2AA668B7667647" italics="true">n</emphasis>
+= 1).
+</paragraph>
+</caption>
+<paragraph id="4EFE5D07E35721810B7593CA08EE09F1">
+<table id="TID0EC1AE" rules="all">
+<tbody id="F49DE14CE87BF0A4FF9677867CC42EBF">
+<tr id="484A53B05759D013660914D901CDF902">
+<th id="3A03829B1FA5B4D4461EEFBE8DF73D05" colspan="1" rowspan="1">Measure</th>
+<th id="AE1832ACA70EFD29681DD6FDF2703AEF" colspan="1" rowspan="1">Instar I</th>
+<th id="B7AF9C15CE723D5FD7EA88BE356ED3B2" colspan="1" rowspan="1">Instar III</th>
+<th id="7904EDC44EA60674D39E6D1736FDEB49" colspan="1" rowspan="1">Measure</th>
+<th id="D249645D76A6B046B410CC843266A81B" colspan="1" rowspan="1">Instar I</th>
+<th id="2656AAE23CBBDEDC7C2A0A137ABDBF88" colspan="1" rowspan="1">Instar III</th>
+</tr>
+<tr id="16815C581D217EBF2002E46CB00AC38B">
+<td id="8CC3935759C9D3E3F7227BBF4A108F4D" colspan="1" rowspan="1">
+<abbrev id="ABBRID0EVUAE" xlink_title="total length">TL</abbrev>
+(mm)
+</td>
+<td id="007E6DC0A2FFC1F6406E477C8253FEC8" colspan="1" rowspan="1">1.97</td>
+<td id="B52D7022B4B357E82D6190940F5534D3" colspan="1" rowspan="1">3.51</td>
+<td id="C03E38C01172F21EC89E01B3F07170D4" colspan="1" rowspan="1">MP 1 / MP 3</td>
+<td id="6EBF099CB42D0AABDC4768D281900242" colspan="1" rowspan="1">0.50</td>
+<td id="3F082C6A474EB28D7620C8F799ED1EE2" colspan="1" rowspan="1">0.69</td>
+</tr>
+<tr id="7557EB83CB57BD8862761F9739AFE3B7">
+<td id="6313BF0F4F5A468EA125DF968619C549" colspan="1" rowspan="1">MW (mm)</td>
+<td id="E21446CA4E2D3DEF4E45A8E5B94D4175" colspan="1" rowspan="1">0.38</td>
+<td id="6C87CE730CB71080397E187281AD26CF" colspan="1" rowspan="1">0.63</td>
+<td id="EAB89F4572FCDEC9422267EA62D4AB93" colspan="1" rowspan="1">MP 2 / MP 3</td>
+<td id="4A04137621CBFF5EA7CB1C64831CCB87" colspan="1" rowspan="1">0.60</td>
+<td id="FB48528098219F4F52B4EBD78751A578" colspan="1" rowspan="1">0.62</td>
+</tr>
+<tr id="860341308E10B33F39765E0157E4B5BC">
+<td id="63A013C7D843F43E3284745A077E03E6" colspan="1" rowspan="1">HL (mm)</td>
+<td id="CC34F21938AECD0824BBE07907ACC00A" colspan="1" rowspan="1">0.34</td>
+<td id="55B042DD0123952827CCC469E5C25D8E" colspan="1" rowspan="1">0.56</td>
+<td id="FE36129B6DD6C57CE03120047E6BCB24" colspan="1" rowspan="1">MP / LP</td>
+<td id="00F52BB6B666B48804C07C44AD251BD9" colspan="1" rowspan="1">2.10</td>
+<td id="26D5353FA3CBFA3E736D8ACA5CD4E048" colspan="1" rowspan="1">1.20</td>
+</tr>
+<tr id="8FEC700C299D9178D1A7353D4658CBC1">
+<td id="81CD7633AC51C833BAEF63C2A2DD032A" colspan="1" rowspan="1">
+<abbrev id="ABBRID0ETWAE" xlink_title="greatest width of head">HW</abbrev>
+(mm)
+</td>
+<td id="40B0C4696B759C73AB554C8E2B8869E7" colspan="1" rowspan="1">0.31</td>
+<td id="F8FE32ECFF41A7210B70AF618397CB83" colspan="1" rowspan="1">0.56</td>
+<td id="FA57A06343B89E790BB94FB9BB7DB97C" colspan="1" rowspan="1">LP 1 / LP 2</td>
+<td id="B083C0EFD3EDB3CD2E70C65B1CE32F0B" colspan="1" rowspan="1">0.67</td>
+<td id="96B251A2A5329A37CCA9E0E1B05779B3" colspan="1" rowspan="1">0.67</td>
+</tr>
+<tr id="3A755283ECED676C1BBB57EF6CDFAA1A">
+<td id="BC1902A7396CF18D4267D6B194C8D412" colspan="1" rowspan="1">FRL (mm)</td>
+<td id="7EBD9323996E8EC9E60E0EFDF2932F6F" colspan="1" rowspan="1">0.34</td>
+<td id="114A12EA7A252238DB5B0B9DDC85865F" colspan="1" rowspan="1">0.54</td>
+<td id="D2666900AFF41576A59A3D4DAF35D6DF" colspan="1" rowspan="1">L 3 (mm)</td>
+<td id="974E8FC5B72E9508761A344C3E5B8A7D" colspan="1" rowspan="1">0.74</td>
+<td id="48B46F245577EAA553C20F185D9049BC" colspan="1" rowspan="1">1.27</td>
+</tr>
+<tr id="6B953FECEB61F8567AAC781A6CE1F561">
+<td id="9C88A381E16E47C890CDFFDD3A30EA5B" colspan="1" rowspan="1">OCW (mm)</td>
+<td id="8B0A63819FF7FD0A1C5355AE7A419B7D" colspan="1" rowspan="1">0.25</td>
+<td id="ED6F5E4CF16B582EEA10E3B309FE8C27" colspan="1" rowspan="1">0.51</td>
+<td id="1FC642AC5025274E34BE22A4AB9A7161" colspan="1" rowspan="1">L 3 / L 1</td>
+<td id="AA302E2166C86ACAEA0F5BB2244D4F14" colspan="1" rowspan="1">1.25</td>
+<td id="4D36D04F964FF051CB6DA7EC641514FF" colspan="1" rowspan="1">1.27</td>
+</tr>
+<tr id="DF0AB96AE1E0AAAFB7A64AE3F3220549">
+<td id="DF73470166C17325B253B7C309E3B0AB" colspan="1" rowspan="1">
+HL / 
+<abbrev id="ABBRID0ERYAE" xlink_title="greatest width of head">HW</abbrev>
+</td>
+<td id="DED6CA99B0E405B319832CE7CA1BA74F" colspan="1" rowspan="1">1.10</td>
+<td id="126F40C294064271CD9421163171A8DF" colspan="1" rowspan="1">1.00</td>
+<td id="50804587D5282CDEBE286F6650ED8D9A" colspan="1" rowspan="1">L 3 / L 2</td>
+<td id="867BA26E5047F2D6586A015C41F5C45E" colspan="1" rowspan="1">1.09</td>
+<td id="64170CFA65C58C275D41C08621ED4A69" colspan="1" rowspan="1">1.10</td>
+</tr>
+<tr id="37269834708200A8C9C4FE2E198F5A75">
+<td id="58B7AFC75FAF88F6FC30C177E55F9C13" colspan="1" rowspan="1">
+<abbrev id="ABBRID0EIZAE" xlink_title="greatest width of head">HW</abbrev>
+/ OCW
+</td>
+<td id="470AD1960509F53EB7BFECE492479231" colspan="1" rowspan="1">1.24</td>
+<td id="E056A5F574C916FB8025F8E1DCC25C6D" colspan="1" rowspan="1">1.08</td>
+<td id="C5BBE75B60444A683ED5D0E6B9260485" colspan="1" rowspan="1">
+L 3 / 
+<abbrev id="ABBRID0EWZAE" xlink_title="greatest width of head">HW</abbrev>
+</td>
+<td id="15A815DA64F72D8C3E04A033F1B7137F" colspan="1" rowspan="1">2.37</td>
+<td id="D01BDBB3516DE0638D142C4BA937612A" colspan="1" rowspan="1">2.31</td>
+</tr>
+<tr id="A27FFB9E1AEB7F160842D81E4D94AAD4">
+<td id="8B662746E81BE30D92585252219D9DA4" colspan="1" rowspan="1">COL / HL</td>
+<td id="761980B8DC34CA7D76089713DCBBBDD0" colspan="1" rowspan="1">0.02</td>
+<td id="D028F9EBF5C7CBFC2851F481ACA27911" colspan="1" rowspan="1">0.02</td>
+<td id="FDF7DE378FA3E04EA9F7AFE35E01B8E0" colspan="1" rowspan="1">CO / FE (L 3)</td>
+<td id="0708DBBA55D20C122CD67436EC606D66" colspan="1" rowspan="1">1.85</td>
+<td id="342EBB135C3E8A287399A847A0182616" colspan="1" rowspan="1">1.70</td>
+</tr>
+<tr id="1ECFB8DA1AF650D77B3379F0E1DBEDF0">
+<td id="9781D44008842E9DC0B90E5081C31886" colspan="1" rowspan="1">FRL / HL</td>
+<td id="73F641EEDEE27CE54FC1C9A0E7A45524" colspan="1" rowspan="1">0.99</td>
+<td id="8A8F0477F60485074AAA8F2A8144A3EC" colspan="1" rowspan="1">0.98</td>
+<td id="7C30A630570704CF96583989618F0E9B" colspan="1" rowspan="1">TI / FE (L 3)</td>
+<td id="714F3855A1712B57D7CBCC0CBE312FC7" colspan="1" rowspan="1">0.72</td>
+<td id="A886DCCE6CCB1DE6DEE66245540B83B5" colspan="1" rowspan="1">0.63</td>
+</tr>
+<tr id="A0DBA920EF02D5BB1ED737DBF0DD0508">
+<td id="0105BA784D2DC984E7D81A6A88C186AD" colspan="1" rowspan="1">
+A / 
+<abbrev id="ABBRID0EK2AE" xlink_title="greatest width of head">HW</abbrev>
+</td>
+<td id="0AD9C90BFD09B7BCA9C1A377ED490445" colspan="1" rowspan="1">0.60</td>
+<td id="E0CB3D227DDC734A09AE30A81F731A21" colspan="1" rowspan="1">0.53</td>
+<td id="0599733C776E3F00E27FDA40407339A0" colspan="1" rowspan="1">TA / FE (L 3)</td>
+<td id="A72149594D0F2292A71E511201DA5519" colspan="1" rowspan="1">0.57</td>
+<td id="B24B758CEE992ACD895C3EB46E1BA6D2" colspan="1" rowspan="1">0.44</td>
+</tr>
+<tr id="9D60651BC1E29B2328B86AFF8719EB26">
+<td id="753729F39A11E128EDDF071A8FC3709E" colspan="1" rowspan="1">A 1 / A 3</td>
+<td id="AD7E6ED938E4B471651D19DEFF23CAD8" colspan="1" rowspan="1">0.63</td>
+<td id="51B4CA9F6123337417039A9974A10E1F" colspan="1" rowspan="1">0.59</td>
+<td id="A4AD0C3A76F003FA4B7D4DEF67EBE21A" colspan="1" rowspan="1">CL / TA (L 3)</td>
+<td id="87633C90C6DA548681BE801526DCEC36" colspan="1" rowspan="1">1.38</td>
+<td id="F857662896FB4A9CE6143316AF79DA81" colspan="1" rowspan="1">1.44</td>
+</tr>
+<tr id="5BBAC707F02C651FDCAD977F2C8F7498">
+<td id="BA718FEF005082C11A2FA2DF09CBCC25" colspan="1" rowspan="1">A 2 / A 3</td>
+<td id="DCFCA8835468D3A522C4CF1CEF1154B5" colspan="1" rowspan="1">1.05</td>
+<td id="139C8A164B3137A8CC4EC4B5F69B1283" colspan="1" rowspan="1">1.29</td>
+<td id="9E74DAB7D1FB2C63F30C27E7A14BB715" colspan="1" rowspan="1">LAS (mm)</td>
+<td id="581B5FA3F743D1F9D4C4A1C9D290AA4A" colspan="1" rowspan="1">0.27</td>
+<td id="2971B4972792B6E4A35A14E2475FD955" colspan="1" rowspan="1">0.70</td>
+</tr>
+<tr id="1C533169F41ABFD04330F3CB1802534A">
+<td id="CE69F90680DD1C3782E0ED3F16CA9D61" colspan="1" rowspan="1">A 4 / A 3</td>
+<td id="F6FC42B98F20EE3DDD2DAD6E741409E4" colspan="1" rowspan="1">0.73</td>
+<td id="008C2F109974F9BDA44EC0728C72C814" colspan="1" rowspan="1">0.53</td>
+<td id="1FFFCB2D622934872D1FDF514884B8D3" colspan="1" rowspan="1">
+LAS / 
+<abbrev id="ABBRID0EQ4AE" xlink_title="greatest width of head">HW</abbrev>
+</td>
+<td id="6904BD8625B9D44E5182F05BC8ECCBD9" colspan="1" rowspan="1">0.87</td>
+<td id="FC54690B5886138E48362CF761E1320E" colspan="1" rowspan="1">1.26</td>
+</tr>
+<tr id="9AB02A7B46FF1F3D3F50531647088B9C">
+<td id="784E8FD0E4603480CFF84A8971BD5328" colspan="1" rowspan="1">MNL / MNW</td>
+<td id="0944BE5D7FD286547A43578102ECD420" colspan="1" rowspan="1">3.25</td>
+<td id="7D6BAC74975D8846D07EFDD734869E71" colspan="1" rowspan="1">2.53</td>
+<td id="11BA92EF73DE8CE0031A4D33482BFDE1" colspan="1" rowspan="1">U (mm)</td>
+<td id="C74DE7D76EC42EAEA08E2D21628985CE" colspan="1" rowspan="1">0.13</td>
+<td id="3E3A984BFA24BF37EB85B48CA264C71B" colspan="1" rowspan="1">0.17</td>
+</tr>
+<tr id="DB55B4703535E9A4281FD4CA567541EA">
+<td id="9B4A439DD185E9B02C012A70B8E50FA1" colspan="1" rowspan="1">MNL / HL</td>
+<td id="42601F9919F62C2C5E53D44173E8BA3B" colspan="1" rowspan="1">0.38</td>
+<td id="D234F9EC46E28460F7F2AA8D3CD3FFAB" colspan="1" rowspan="1">0.35</td>
+<td id="70A5B23F38E081C1E8F8285690E0C15C" colspan="1" rowspan="1">U / LAS</td>
+<td id="3D8C624BF4D541E15B1A97B22C0782A0" colspan="1" rowspan="1">0.48</td>
+<td id="69D8DC839F2C7937C30C76066E003FB6" colspan="1" rowspan="1">0.25</td>
+</tr>
+<tr id="FEDBA6C47E4650CEDC0B9A7B247E982C">
+<td id="01B89584D12E48DC34B17CC18321EED0" colspan="1" rowspan="1">A / MP</td>
+<td id="DC7B56CA23381108EC4404BF0E10960F" colspan="1" rowspan="1">1.79</td>
+<td id="85964032A990E992FE12C8B7D1A41533" colspan="1" rowspan="1">1.93</td>
+<td id="A99181F654BC66917576F4BB07A76ABB" colspan="1" rowspan="1">
+U / 
+<abbrev id="ABBRID0EN6AE" xlink_title="greatest width of head">HW</abbrev>
+</td>
+<td id="1269926990BAB7EE533304932F82D566" colspan="1" rowspan="1">0.42</td>
+<td id="E78F13DA86669452C71F30A1BE3D1CB0" colspan="1" rowspan="1">0.31</td>
+</tr>
+<tr id="F7D956D12EF1CD3726B7DBB6ABB6CDE6">
+<td id="1B1586CA26DBCE916DD5E9A889EDE39A" colspan="1" rowspan="1">GA / MP 1</td>
+<td id="68868CC47A8798759F9B0746F82D4D06" colspan="1" rowspan="1">1.20</td>
+<td id="7F383543E39BC7B03EB0818CD7D9B38E" colspan="1" rowspan="1">1.00</td>
+<td id="FAC25B52ED9490B1328E092D3601ACC5" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="F26A7BEF7B4FF7B896626EA438261615" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="F784CB536F99C3B76FD282EBDB4E68D6" colspan="1" isEmpty="true" rowspan="1">-</td>
+</tr>
+</tbody>
+</table>
+</paragraph>
+<paragraph id="DEDE5D1A6139F6ACA5217992FAE17107">
+<emphasis id="48909E3D1B9CB3D1D36C07BE3A81F8E4" bold="true">
+<emphasis id="AE54B315A867ED36F5AF12FBE3AA0331" bold="true" italics="true">Head</emphasis>
+.
+</emphasis>
+Prognathous; cephalic capsule (Figs 
+<figureCitation id="8E43B2753BF009D4F2561D62BD823DA3" captionStart="Figures 18–20" captionStartId="F5" captionText="Figures 18–20. Liocanthydrus nanops Baca et al., 2014, instar I 18 habitus, dorsal aspect 19 cephalic capsule, dorsal aspect 20 cephalic capsule, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. Solid square indicates additional seta. EB: egg burster, FR: frontoclypeus, PA: parietal, TP: tentorial pit. Scale bars: 0.50 mm (18); 0.10 mm (19, 20)." figureDoi="10.3897/zookeys.1231.144746.figures18-20" httpUri="https://binary.pensoft.net/fig/1279881">19</figureCitation>
+, 
+<figureCitation id="CB5BE981BDE4F56679B8E2B6A679C0D7" captionStart="Figures 18–20" captionStartId="F5" captionText="Figures 18–20. Liocanthydrus nanops Baca et al., 2014, instar I 18 habitus, dorsal aspect 19 cephalic capsule, dorsal aspect 20 cephalic capsule, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. Solid square indicates additional seta. EB: egg burster, FR: frontoclypeus, PA: parietal, TP: tentorial pit. Scale bars: 0.50 mm (18); 0.10 mm (19, 20)." figureDoi="10.3897/zookeys.1231.144746.figures18-20" httpUri="https://binary.pensoft.net/fig/1279881">20</figureCitation>
+) slightly longer than broad; maximum width posterior to stemmata; slightly narrowed posteriorly; occipital foramen large; coronal suture very short; ecdysial suture U-shaped; tentorial pits visible postero-ventrally, well separated from each other and from occipital foramen; six lateral stemmata arranged in two curved vertical rows at each side. Frontoclypeus elongate, roughly subovate, anterior margin rounded, with two spine-like egg bursters on basal third. 
+<emphasis id="516CA58AF724C417247E9DFB0F53FD9D" bold="true" italics="true">Antenna</emphasis>
+(Figs 
+<figureCitation id="4E4CCB47669BC425CC8C78AD007E5C52" captionStart="Figures 21–27" captionStartId="F6" captionText="Figures 21–27. Liocanthydrus nanops Baca et al., 2014, instar I 21 left antenna, dorsal aspect 22 right antenna, ventral aspect 23 left mandible, dorsal aspect 24 right maxilla, dorsal aspect 25 left maxilla, ventral aspect 26 labium, dorsal aspect 27 labium, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AN: antenna, LA: labium, MN: mandible, MX: maxilla. Scale bars: 0.04 mm." figureDoi="10.3897/zookeys.1231.144746.figures21-27" httpUri="https://binary.pensoft.net/fig/1279882">21</figureCitation>
+, 
+<figureCitation id="8F387DC56E1092E944DC07694C34EEFF" captionStart="Figures 21–27" captionStartId="F6" captionText="Figures 21–27. Liocanthydrus nanops Baca et al., 2014, instar I 21 left antenna, dorsal aspect 22 right antenna, ventral aspect 23 left mandible, dorsal aspect 24 right maxilla, dorsal aspect 25 left maxilla, ventral aspect 26 labium, dorsal aspect 27 labium, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AN: antenna, LA: labium, MN: mandible, MX: maxilla. Scale bars: 0.04 mm." figureDoi="10.3897/zookeys.1231.144746.figures21-27" httpUri="https://binary.pensoft.net/fig/1279882">22</figureCitation>
+). Short, robust, shorter than maximum head width, composed of four antennomeres; A 2 and A 3 longest, subequal; A 3 with a rugged area on distal portion; A 4 approximately 3 / 4 length of A 3; A 1 shortest. 
+<emphasis id="33E894EA982903B802C188CA630AF5FD" bold="true" italics="true">Mandible</emphasis>
+(Fig. 
+<figureCitation id="5C59ADFAA7E455DC2245E04F13BAC35D" captionStart="Figures 21–27" captionStartId="F6" captionText="Figures 21–27. Liocanthydrus nanops Baca et al., 2014, instar I 21 left antenna, dorsal aspect 22 right antenna, ventral aspect 23 left mandible, dorsal aspect 24 right maxilla, dorsal aspect 25 left maxilla, ventral aspect 26 labium, dorsal aspect 27 labium, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AN: antenna, LA: labium, MN: mandible, MX: maxilla. Scale bars: 0.04 mm." figureDoi="10.3897/zookeys.1231.144746.figures21-27" httpUri="https://binary.pensoft.net/fig/1279882">23</figureCitation>
+). Symmetrical, short, basal half broad, inner margin with strong subrectangular process, distal half slender, curved inwards, narrowing to pointed apex, inner margin smooth. 
+<emphasis id="DD1A44E4CDE9B60C85E119EA022F9F87" bold="true" italics="true">Maxilla</emphasis>
+(Figs 
+<figureCitation id="60CDD545CD3D6F0DE72C2E5E0A8CB5C9" captionStart="Figures 21–27" captionStartId="F6" captionText="Figures 21–27. Liocanthydrus nanops Baca et al., 2014, instar I 21 left antenna, dorsal aspect 22 right antenna, ventral aspect 23 left mandible, dorsal aspect 24 right maxilla, dorsal aspect 25 left maxilla, ventral aspect 26 labium, dorsal aspect 27 labium, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AN: antenna, LA: labium, MN: mandible, MX: maxilla. Scale bars: 0.04 mm." figureDoi="10.3897/zookeys.1231.144746.figures21-27" httpUri="https://binary.pensoft.net/fig/1279882">24</figureCitation>
+, 
+<figureCitation id="815DD9B302FCC036FC5BD65DABAA3A3D" captionStart="Figures 21–27" captionStartId="F6" captionText="Figures 21–27. Liocanthydrus nanops Baca et al., 2014, instar I 21 left antenna, dorsal aspect 22 right antenna, ventral aspect 23 left mandible, dorsal aspect 24 right maxilla, dorsal aspect 25 left maxilla, ventral aspect 26 labium, dorsal aspect 27 labium, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AN: antenna, LA: labium, MN: mandible, MX: maxilla. Scale bars: 0.04 mm." figureDoi="10.3897/zookeys.1231.144746.figures21-27" httpUri="https://binary.pensoft.net/fig/1279882">25</figureCitation>
+). Cardo small, suboval; stipes well developed, subtrapezoidal, bearing a galea on distal inner margin and a palpus on distal outer margin; galea well developed, composed of two galeomeres, GA 2 more slender and subequal in length to GA 1; palpifer not cleary differentiated from stipes, more evident in ventral view; palpus short, robust, composed of three palpomeres, MP 1 shortest, MP 3 longest. 
+<emphasis id="F274C816C7A179FD5F83EFE91D0F51D7" bold="true" italics="true">Labium</emphasis>
+(Figs 
+<figureCitation id="54069529A9161349E85000BB1CCC5BA4" captionStart="Figures 21–27" captionStartId="F6" captionText="Figures 21–27. Liocanthydrus nanops Baca et al., 2014, instar I 21 left antenna, dorsal aspect 22 right antenna, ventral aspect 23 left mandible, dorsal aspect 24 right maxilla, dorsal aspect 25 left maxilla, ventral aspect 26 labium, dorsal aspect 27 labium, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AN: antenna, LA: labium, MN: mandible, MX: maxilla. Scale bars: 0.04 mm." figureDoi="10.3897/zookeys.1231.144746.figures21-27" httpUri="https://binary.pensoft.net/fig/1279882">26</figureCitation>
+, 
+<figureCitation id="A441908840641F99A3C5D40471AE7B03" captionStart="Figures 21–27" captionStartId="F6" captionText="Figures 21–27. Liocanthydrus nanops Baca et al., 2014, instar I 21 left antenna, dorsal aspect 22 right antenna, ventral aspect 23 left mandible, dorsal aspect 24 right maxilla, dorsal aspect 25 left maxilla, ventral aspect 26 labium, dorsal aspect 27 labium, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AN: antenna, LA: labium, MN: mandible, MX: maxilla. Scale bars: 0.04 mm." figureDoi="10.3897/zookeys.1231.144746.figures21-27" httpUri="https://binary.pensoft.net/fig/1279882">27</figureCitation>
+). Prementum well developed, subrectangular, somewhat broader than long, anterior margin narrowly indented medially; palpus short, robust, composed of two palpomeres, LP 2 longer than LP 1.
+</paragraph>
+<caption id="5A71ACFDB817DF3EA14B3BD4BE5BC44F" ID-DOI="10.3897/zookeys.1231.144746.figures34-37" ID-arpha="540FC164-0CC3-505E-BF5E-78B0B5D05804" httpUri="https://binary.pensoft.net/fig/1279885" startId="F9">
+<paragraph id="049094184C3D1EC5EA3CF7DEAFCDCFCE">
+<label id="DEA1C6D488BD6908CE3F171BEA054DF6">Figures 34–37.</label>
+</paragraph>
+<paragraph id="043C5034AC92DB1C94DDABB27097DDEF">
+<taxonomicName id="BCA94CC6BCF97947CC3F52FEAA4387C4" authority="Baca et al., 2014" authorityName="Baca" authorityYear="2014" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="9841C477D8BA07689C4CC43EB138D117" italics="true">Liocanthydrus nanops</emphasis>
+<bibRefCitation id="366619EF3E77771F0EC5BD3ADF33AFCC" DOI="10.11646/zootaxa.3793.2.3" author="Baca" etAl="et al." firstAuthor="Baca" journalOrPublisher="Zootaxa" pagination="231-246" refId="B1" refString="Baca SM, Gustafson GT, Toledo M, Miller KB (2014) Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species. Zootaxa 3793: 231–246. https://doi.org/10.11646/zootaxa.3793.2.3" title="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species." volume="3793" year="2014">Baca et al., 2014</bibRefCitation>
+</taxonomicName>
+, instar III 
+<emphasis id="15CFA09E5B8548D1D3A767AA33438D37" bold="true">34</emphasis>
+head, dorsal aspect 
+<emphasis id="E679663E02B4CD85370FC9771030F6EC" bold="true">35</emphasis>
+left prothoracic leg, anterior aspect 
+<emphasis id="C83AD9633154D459EDCBCEF7EA0F5C35" bold="true">36</emphasis>
+right prothoracic leg, posterior aspect 
+<emphasis id="7441A3C2ECB080F2EF2A61FB582BFDB4" bold="true">37</emphasis>
+abdominal segment VIII, dorsal aspect. Numbers and lowercase letters indicate primary setae and pores, respectively (some setae on abdominal segment VIII could not be identified). Scale bars: 0.20 mm.
+</paragraph>
+</caption>
+<paragraph id="89DF35B5D60452934FEDD3F7172256BA">
+<emphasis id="3BA5D282199E535C12C78391ABBAB235" bold="true">
+<emphasis id="EBF82E29B7C6EE8599EC71FBBACF2299" bold="true" italics="true">Thorax</emphasis>
+.
+</emphasis>
+Terga fully sclerotised, convex (Fig. 
+<figureCitation id="0CB873E32B6101DCF2260CC0F1D293E0" captionStart="Figures 18–20" captionStartId="F5" captionText="Figures 18–20. Liocanthydrus nanops Baca et al., 2014, instar I 18 habitus, dorsal aspect 19 cephalic capsule, dorsal aspect 20 cephalic capsule, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. Solid square indicates additional seta. EB: egg burster, FR: frontoclypeus, PA: parietal, TP: tentorial pit. Scale bars: 0.50 mm (18); 0.10 mm (19, 20)." figureDoi="10.3897/zookeys.1231.144746.figures18-20" httpUri="https://binary.pensoft.net/fig/1279881">18</figureCitation>
+); pronotum about as long as meso- and metanotum combined, meso- and metanotum subequal in length, approximately as wide as pronotum; protergite subrectangular, lateral margins rounded, more developed than meso- and metatergite; meso- and metatergite with anterotransverse carina; ecdysial line absent. 
+<emphasis id="3FB2BA0D01A37C8456EE1C7F457E7396" bold="true" italics="true">Legs</emphasis>
+(Figs 
+<figureCitation id="AA1C8E05DCD66096ABF3A27626225305" captionStart="Figures 28–29" captionStartId="F7" captionText="Figures 28–29. Liocanthydrus nanops Baca et al., 2014, instar I 28 left metathoracic leg, anterior aspect 29 right metathoracic leg, posterior aspect. Numbers and lowercase letters indicate primary setae and pores respectively. Solid square indicates additional seta. CO: coxa, FE: femur, PT: pretarsus, TA: tarsus, TI: tibia, TR: trochanter. Scale bar: 0.10 mm." figureDoi="10.3897/zookeys.1231.144746.figures28-29" httpUri="https://binary.pensoft.net/fig/1279883">28</figureCitation>
+, 
+<figureCitation id="6E8C2EA68E4DE8DC7FE5ABF33E729A7F" captionStart="Figures 28–29" captionStartId="F7" captionText="Figures 28–29. Liocanthydrus nanops Baca et al., 2014, instar I 28 left metathoracic leg, anterior aspect 29 right metathoracic leg, posterior aspect. Numbers and lowercase letters indicate primary setae and pores respectively. Solid square indicates additional seta. CO: coxa, FE: femur, PT: pretarsus, TA: tarsus, TI: tibia, TR: trochanter. Scale bar: 0.10 mm." figureDoi="10.3897/zookeys.1231.144746.figures28-29" httpUri="https://binary.pensoft.net/fig/1279883">29</figureCitation>
+). Short, robust, composed of six articles, L 1 shortest, L 3 longest; coxa broad, elongate, trochanter lacking annulus, femur, tibia and tarsus short, subcylindrical, pretarsus with two long, slender, slightly curved claws, posterior claw slightly shorter than anterior claw.
+</paragraph>
+<paragraph id="C510A2ABF499180DDECA790DE0065B64">
+<emphasis id="D3DD9B27B58F5540C369F67950E38FE7" bold="true">
+<emphasis id="C7AA8CB77C276460A9DB1CC883A62CB6" bold="true" italics="true">Abdomen</emphasis>
+.
+</emphasis>
+Eight-segmented (Fig. 
+<figureCitation id="E7896FDFF90686A3F5BEF8FEE33584BC" captionStart="Figures 18–20" captionStartId="F5" captionText="Figures 18–20. Liocanthydrus nanops Baca et al., 2014, instar I 18 habitus, dorsal aspect 19 cephalic capsule, dorsal aspect 20 cephalic capsule, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. Solid square indicates additional seta. EB: egg burster, FR: frontoclypeus, PA: parietal, TP: tentorial pit. Scale bars: 0.50 mm (18); 0.10 mm (19, 20)." figureDoi="10.3897/zookeys.1231.144746.figures18-20" httpUri="https://binary.pensoft.net/fig/1279881">18</figureCitation>
+); segments I – VIII completely sclerotised, ring-like, progressively narrowing to apex, with anterotransverse carina; segment VIII (Figs 
+<figureCitation id="0020BDDF02C8E09CF47A66E51D8CAF1D" captionStart="Figures 30–33" captionStartId="F8" captionText="Figures 30–33. Liocanthydrus nanops Baca et al., 2014, instar I 30 abdominal segment VIII, dorsal aspect 31 abdominal segment VIII and urogomphi, ventral aspect 32 urogomphi, dorsal aspect 33 urogomphi, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AB: abdominal segment VIII, UR: urogomphus. Scale bars: 0.10 mm (30, 31); 0.02 mm (32, 33)." figureDoi="10.3897/zookeys.1231.144746.figures30-33" httpUri="https://binary.pensoft.net/fig/1279884">30</figureCitation>
+, 
+<figureCitation id="4A19C87F060C52C60848540844BC652C" captionStart="Figures 30–33" captionStartId="F8" captionText="Figures 30–33. Liocanthydrus nanops Baca et al., 2014, instar I 30 abdominal segment VIII, dorsal aspect 31 abdominal segment VIII and urogomphi, ventral aspect 32 urogomphi, dorsal aspect 33 urogomphi, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AB: abdominal segment VIII, UR: urogomphus. Scale bars: 0.10 mm (30, 31); 0.02 mm (32, 33)." figureDoi="10.3897/zookeys.1231.144746.figures30-33" httpUri="https://binary.pensoft.net/fig/1279884">31</figureCitation>
+) the longest and narrowest, with two terminal dorsal projections, lacking a U-shaped wavy membranous area ventrally, contiguous to urogomphi; siphon absent. 
+<emphasis id="510717C883E9194843A580EB939816AA" bold="true" italics="true">Urogomphi</emphasis>
+(Figs 
+<figureCitation id="5E8A86D5CFDE2132069B99A7E572FF41" captionStart="Figures 30–33" captionStartId="F8" captionText="Figures 30–33. Liocanthydrus nanops Baca et al., 2014, instar I 30 abdominal segment VIII, dorsal aspect 31 abdominal segment VIII and urogomphi, ventral aspect 32 urogomphi, dorsal aspect 33 urogomphi, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AB: abdominal segment VIII, UR: urogomphus. Scale bars: 0.10 mm (30, 31); 0.02 mm (32, 33)." figureDoi="10.3897/zookeys.1231.144746.figures30-33" httpUri="https://binary.pensoft.net/fig/1279884">32</figureCitation>
+, 
+<figureCitation id="92FA3CAE0B3D4717F583E30A8F0A0EE6" captionStart="Figures 30–33" captionStartId="F8" captionText="Figures 30–33. Liocanthydrus nanops Baca et al., 2014, instar I 30 abdominal segment VIII, dorsal aspect 31 abdominal segment VIII and urogomphi, ventral aspect 32 urogomphi, dorsal aspect 33 urogomphi, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AB: abdominal segment VIII, UR: urogomphus. Scale bars: 0.10 mm (30, 31); 0.02 mm (32, 33)." figureDoi="10.3897/zookeys.1231.144746.figures30-33" httpUri="https://binary.pensoft.net/fig/1279884">33</figureCitation>
+). Short, cylindrical, visible in dorsal view (Fig. 
+<figureCitation id="54643B62824A2BD5423893CCB43A88DF" captionStart="Figures 18–20" captionStartId="F5" captionText="Figures 18–20. Liocanthydrus nanops Baca et al., 2014, instar I 18 habitus, dorsal aspect 19 cephalic capsule, dorsal aspect 20 cephalic capsule, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. Solid square indicates additional seta. EB: egg burster, FR: frontoclypeus, PA: parietal, TP: tentorial pit. Scale bars: 0.50 mm (18); 0.10 mm (19, 20)." figureDoi="10.3897/zookeys.1231.144746.figures18-20" httpUri="https://binary.pensoft.net/fig/1279881">18</figureCitation>
+), not fused to each other.
+</paragraph>
+<paragraph id="4BECE691A095A23F0175BF237DA6EF94">
+<emphasis id="AB2990FB1F6398A3B50110EFDEC43959" bold="true">
+<emphasis id="150042D6EDA5319C1ADD8552D9ED80D2" bold="true" italics="true">Chaetotaxy</emphasis>
+.
+</emphasis>
+Similar to that of 
+<taxonomicName id="26548A43E27EC903859384AF54C22951" authorityName="" baseAuthorityName="J. Balfour-Browne" baseAuthorityYear="1969" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="clayae">
+<emphasis id="8C9CBB5F3C7C3FF8BF1876F51F3D039D" italics="true">L. clayae</emphasis>
+</taxonomicName>
+(see 
+<bibRefCitation id="CE02E53D2546394611CC6B7B94086117" DOI="10.1080/00222933.2020.1842931" author="Urcola" etAl="et al." firstAuthor="Urcola" journalOrPublisher="Journal of Natural History" pagination="2285-2296" refId="B13" refString="Urcola JI, Benetti CJ, Alarie Y, Michat MC (2021) The unknown larva of the burrowing water beetle genus Liocanthydrus Guignot, 1957 (Coleoptera: Noteridae). Journal of Natural History 54: 2285–2296. https://doi.org/10.1080/00222933.2020.1842931" title="The unknown larva of the burrowing water beetle genus Liocanthydrus Guignot, 1957 (Coleoptera: Noteridae)." volume="54" year="2021">Urcola et al. 2021</bibRefCitation>
+) except for the following features: seta FR 1 very short (Fig. 
+<figureCitation id="2FD715B25EBEFB1BF70E707E0E1A804F" captionStart="Figures 18–20" captionStartId="F5" captionText="Figures 18–20. Liocanthydrus nanops Baca et al., 2014, instar I 18 habitus, dorsal aspect 19 cephalic capsule, dorsal aspect 20 cephalic capsule, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. Solid square indicates additional seta. EB: egg burster, FR: frontoclypeus, PA: parietal, TP: tentorial pit. Scale bars: 0.50 mm (18); 0.10 mm (19, 20)." figureDoi="10.3897/zookeys.1231.144746.figures18-20" httpUri="https://binary.pensoft.net/fig/1279881">19</figureCitation>
+); seta PA 9 absent (Fig. 
+<figureCitation id="2B46D4CEA866CF4826A22D8089D88098" captionStart="Figures 18–20" captionStartId="F5" captionText="Figures 18–20. Liocanthydrus nanops Baca et al., 2014, instar I 18 habitus, dorsal aspect 19 cephalic capsule, dorsal aspect 20 cephalic capsule, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. Solid square indicates additional seta. EB: egg burster, FR: frontoclypeus, PA: parietal, TP: tentorial pit. Scale bars: 0.50 mm (18); 0.10 mm (19, 20)." figureDoi="10.3897/zookeys.1231.144746.figures18-20" httpUri="https://binary.pensoft.net/fig/1279881">19</figureCitation>
+); seta AN 1 inserted more distally (Fig. 
+<figureCitation id="11D391DBEC79ED799ED04926FDC4E291" captionStart="Figures 21–27" captionStartId="F6" captionText="Figures 21–27. Liocanthydrus nanops Baca et al., 2014, instar I 21 left antenna, dorsal aspect 22 right antenna, ventral aspect 23 left mandible, dorsal aspect 24 right maxilla, dorsal aspect 25 left maxilla, ventral aspect 26 labium, dorsal aspect 27 labium, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AN: antenna, LA: labium, MN: mandible, MX: maxilla. Scale bars: 0.04 mm." figureDoi="10.3897/zookeys.1231.144746.figures21-27" httpUri="https://binary.pensoft.net/fig/1279882">21</figureCitation>
+); seta MN 1 short (Fig. 
+<figureCitation id="34C04F2769DA6F1D0C17DE21812281F3" captionStart="Figures 21–27" captionStartId="F6" captionText="Figures 21–27. Liocanthydrus nanops Baca et al., 2014, instar I 21 left antenna, dorsal aspect 22 right antenna, ventral aspect 23 left mandible, dorsal aspect 24 right maxilla, dorsal aspect 25 left maxilla, ventral aspect 26 labium, dorsal aspect 27 labium, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AN: antenna, LA: labium, MN: mandible, MX: maxilla. Scale bars: 0.04 mm." figureDoi="10.3897/zookeys.1231.144746.figures21-27" httpUri="https://binary.pensoft.net/fig/1279882">23</figureCitation>
+); MP 3 with several minute sensilla on surface (Figs 
+<figureCitation id="AA4DC485386FB4CF3A7BB809938B5A41" captionStart="Figures 21–27" captionStartId="F6" captionText="Figures 21–27. Liocanthydrus nanops Baca et al., 2014, instar I 21 left antenna, dorsal aspect 22 right antenna, ventral aspect 23 left mandible, dorsal aspect 24 right maxilla, dorsal aspect 25 left maxilla, ventral aspect 26 labium, dorsal aspect 27 labium, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AN: antenna, LA: labium, MN: mandible, MX: maxilla. Scale bars: 0.04 mm." figureDoi="10.3897/zookeys.1231.144746.figures21-27" httpUri="https://binary.pensoft.net/fig/1279882">24</figureCitation>
+, 
+<figureCitation id="C73D1CAA43D3F0705A753337D8967F0D" captionStart="Figures 21–27" captionStartId="F6" captionText="Figures 21–27. Liocanthydrus nanops Baca et al., 2014, instar I 21 left antenna, dorsal aspect 22 right antenna, ventral aspect 23 left mandible, dorsal aspect 24 right maxilla, dorsal aspect 25 left maxilla, ventral aspect 26 labium, dorsal aspect 27 labium, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AN: antenna, LA: labium, MN: mandible, MX: maxilla. Scale bars: 0.04 mm." figureDoi="10.3897/zookeys.1231.144746.figures21-27" httpUri="https://binary.pensoft.net/fig/1279882">25</figureCitation>
+); pore URa located on dorsal surface (Fig. 
+<figureCitation id="D86CBB83B5CC8692143F3C50D45C8EEE" captionStart="Figures 30–33" captionStartId="F8" captionText="Figures 30–33. Liocanthydrus nanops Baca et al., 2014, instar I 30 abdominal segment VIII, dorsal aspect 31 abdominal segment VIII and urogomphi, ventral aspect 32 urogomphi, dorsal aspect 33 urogomphi, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AB: abdominal segment VIII, UR: urogomphus. Scale bars: 0.10 mm (30, 31); 0.02 mm (32, 33)." figureDoi="10.3897/zookeys.1231.144746.figures30-33" httpUri="https://binary.pensoft.net/fig/1279884">32</figureCitation>
+). Setae on abdominal segment VIII were not named in instar II of 
+<taxonomicName id="D5D488EE47853C7117F84FEFC3796F14" authorityName="" baseAuthorityName="J. Balfour-Browne" baseAuthorityYear="1969" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="clayae">
+<emphasis id="BB714107E4CA6E9FEDBFB33583A4962E" italics="true">L. clayae</emphasis>
+</taxonomicName>
+due to the presence of secondary setae. Therefore, they are detailed here for 
+<taxonomicName id="851BB05014CF62E8DF6EEE5F49DC78C3" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="4C569E6E855B1342D75E221A18881520" italics="true">L. nanops</emphasis>
+</taxonomicName>
+: dorsal surface of segment VIII with one seta (AB 1) on basal region, one seta (AB 3) on distal third and one long (AB 4) and four minute (AB 5, AB 6, AB 7, AB 16) setae apically (Fig. 
+<figureCitation id="6779A90C8218EC5261F898812B57A627" captionStart="Figures 30–33" captionStartId="F8" captionText="Figures 30–33. Liocanthydrus nanops Baca et al., 2014, instar I 30 abdominal segment VIII, dorsal aspect 31 abdominal segment VIII and urogomphi, ventral aspect 32 urogomphi, dorsal aspect 33 urogomphi, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AB: abdominal segment VIII, UR: urogomphus. Scale bars: 0.10 mm (30, 31); 0.02 mm (32, 33)." figureDoi="10.3897/zookeys.1231.144746.figures30-33" httpUri="https://binary.pensoft.net/fig/1279884">30</figureCitation>
+); each terminal dorsal projection with one short medial seta (AB 14) and one long apical seta (AB 8); ventral surface of segment VIII with two setae (AB 12, AB 13) on basal region and four setae (AB 9, AB 10, AB 11, B 15) on distal region (Fig. 
+<figureCitation id="E5E791F75208FA466840E18C98698447" captionStart="Figures 30–33" captionStartId="F8" captionText="Figures 30–33. Liocanthydrus nanops Baca et al., 2014, instar I 30 abdominal segment VIII, dorsal aspect 31 abdominal segment VIII and urogomphi, ventral aspect 32 urogomphi, dorsal aspect 33 urogomphi, ventral aspect. Numbers and lowercase letters indicate primary setae and pores, respectively. AB: abdominal segment VIII, UR: urogomphus. Scale bars: 0.10 mm (30, 31); 0.02 mm (32, 33)." figureDoi="10.3897/zookeys.1231.144746.figures30-33" httpUri="https://binary.pensoft.net/fig/1279884">31</figureCitation>
+).
+</paragraph>
+<paragraph id="A94F505F8D93448190C71623F8AE20DB">
+<emphasis id="AAE0FB064A6DD43D728DA808433F5620" bold="true">Instar III</emphasis>
+(Figs 
+<figureCitation id="B6BA7AAD769923148D61062FE208F704" captionStart="Figures 13–16" captionStartId="F3" captionText="Figures 13–16. Habitat of Liocanthydrus nanops Baca et al., 2014, Iguazú NP, Misiones Province, Argentina 13 general view of the stream’s flooding site 14 collecting of beetles 15 details of the stream 16 instar III larva of L. nanops. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1231.144746.figures13-16" httpUri="https://binary.pensoft.net/fig/1279879">16</figureCitation>
+, 
+<figureCitation id="B67DE683E1619003541FE27998D1E58D" captionStart="Figures 34–37" captionStartId="F9" captionText="Figures 34–37. Liocanthydrus nanops Baca et al., 2014, instar III 34 head, dorsal aspect 35 left prothoracic leg, anterior aspect 36 right prothoracic leg, posterior aspect 37 abdominal segment VIII, dorsal aspect. Numbers and lowercase letters indicate primary setae and pores, respectively (some setae on abdominal segment VIII could not be identified). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1231.144746.figures34-37" httpUri="https://binary.pensoft.net/fig/1279885">34–37</figureCitation>
+)
+</paragraph>
+<paragraph id="227B12026A2BD806DA74DDB0CA57D596">As for instar I except for the following features:</paragraph>
+<paragraph id="79B3751A04A6FDF09AF6B1E8E7FF15C9">
+<emphasis id="E0D9F4CB44BBFB320FAACCD823A21B6F" bold="true">
+<emphasis id="355D68B9EC67C70314E5AF902EC8439C" bold="true" italics="true">Body</emphasis>
+.
+</emphasis>
+Measurements and ratios that characterise body shape are shown in Table 
+<tableCitation id="97667C9B8F5B8B44E7A25DE08C6B32F6" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements and ratios for the larval instars of Liocanthydrus nanops Baca et al. 2014 (n = 1)." httpUri="http://table.plazi.org/id/FA6E299B8663F3E9739748C1C56820F3" tableUuid="FA6E299B8663F3E9739748C1C56820F3">1</tableCitation>
+.
+</paragraph>
+<paragraph id="998D22D225BC0939FACEF9FA50BFC107">
+<emphasis id="D72438810441567BDB9A6BFB8B4DB262" bold="true">
+<emphasis id="088F656607516671652994CA9A186176" bold="true" italics="true">Head</emphasis>
+.
+</emphasis>
+Egg bursters absent; A 2 longer than A 3; A 4 shortest, approximately 1 / 2 length of A 3; mandible more robust, process less prominent (Fig. 
+<figureCitation id="A352CFF40DBC1B829831C44CE67CEA5A" captionStart="Figures 34–37" captionStartId="F9" captionText="Figures 34–37. Liocanthydrus nanops Baca et al., 2014, instar III 34 head, dorsal aspect 35 left prothoracic leg, anterior aspect 36 right prothoracic leg, posterior aspect 37 abdominal segment VIII, dorsal aspect. Numbers and lowercase letters indicate primary setae and pores, respectively (some setae on abdominal segment VIII could not be identified). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1231.144746.figures34-37" httpUri="https://binary.pensoft.net/fig/1279885">34</figureCitation>
+).
+</paragraph>
+<paragraph id="DC9A61C7545DDA2F288D99DBE268356A">
+<emphasis id="F4630467350B0B3D30DEF01A6D0B3E8A" bold="true">
+<emphasis id="23BBEAE0DA441A1E9733C9912287C086" bold="true" italics="true">Abdomen</emphasis>
+.
+</emphasis>
+Siphon relatively long, slender, apex truncated (Fig. 
+<figureCitation id="7A43BD1E9190C3F3254CB6E81BB749D0" captionStart="Figures 34–37" captionStartId="F9" captionText="Figures 34–37. Liocanthydrus nanops Baca et al., 2014, instar III 34 head, dorsal aspect 35 left prothoracic leg, anterior aspect 36 right prothoracic leg, posterior aspect 37 abdominal segment VIII, dorsal aspect. Numbers and lowercase letters indicate primary setae and pores, respectively (some setae on abdominal segment VIII could not be identified). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1231.144746.figures34-37" httpUri="https://binary.pensoft.net/fig/1279885">37</figureCitation>
+).
+</paragraph>
+<paragraph id="C78AA1DF960426CF4273055FA0AB81F7">
+<emphasis id="C6D45E19A38D789D8C353386339885C9" bold="true">
+<emphasis id="00E0F59795D1E52B6A23A7C82C3E1439" bold="true" italics="true">Chaetotaxy</emphasis>
+.
+</emphasis>
+Frontoclypeus with 14–21 minute secondary setae on anterior half and 3–4 minute secondary setae on posterior half; dorsal surface of parietal with seta PA 9 present (inserted close to seta PA 6), 0–4 minute secondary setae on anterior portion and 10–13 minute secondary setae on posterior portion (Fig. 
+<figureCitation id="43CF84AA4EC9A50453B1E2CFAFCEE479" captionStart="Figures 34–37" captionStartId="F9" captionText="Figures 34–37. Liocanthydrus nanops Baca et al., 2014, instar III 34 head, dorsal aspect 35 left prothoracic leg, anterior aspect 36 right prothoracic leg, posterior aspect 37 abdominal segment VIII, dorsal aspect. Numbers and lowercase letters indicate primary setae and pores, respectively (some setae on abdominal segment VIII could not be identified). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1231.144746.figures34-37" httpUri="https://binary.pensoft.net/fig/1279885">34</figureCitation>
+); ventral surface of parietal with 10–13 minute secondary setae on anterior half and 1–2 minute secondary setae on posterior half; secondary leg setation detailed in Table 
+<tableCitation id="27EA4AD3A253F2E16B4D95DFB34A31F5" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Number and position of secondary setae on the legs of larvae of Liocanthydrus nanops Baca et al. 2014. Numbers between slash marks refer to pro-, meso-, and metathoracic leg, respectively. A = anterior, PD = posterodorsal, PV = posteroventral; Total = total number of secondary setae on the article (i. e., excluding primary setae) (n = 1)." httpUri="http://table.plazi.org/id/40AE87F8CC844DB08E7B09B2CD312719" tableUuid="40AE87F8CC844DB08E7B09B2CD312719">2</tableCitation>
+and Figs 
+<figureCitation id="62D9B8ECA9BCD6AEE107C8614644A454" captionStart="Figures 34–37" captionStartId="F9" captionText="Figures 34–37. Liocanthydrus nanops Baca et al., 2014, instar III 34 head, dorsal aspect 35 left prothoracic leg, anterior aspect 36 right prothoracic leg, posterior aspect 37 abdominal segment VIII, dorsal aspect. Numbers and lowercase letters indicate primary setae and pores, respectively (some setae on abdominal segment VIII could not be identified). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1231.144746.figures34-37" httpUri="https://binary.pensoft.net/fig/1279885">35</figureCitation>
+, 
+<figureCitation id="8C595F790E1394CB38719F6D70BFCDB0" captionStart="Figures 34–37" captionStartId="F9" captionText="Figures 34–37. Liocanthydrus nanops Baca et al., 2014, instar III 34 head, dorsal aspect 35 left prothoracic leg, anterior aspect 36 right prothoracic leg, posterior aspect 37 abdominal segment VIII, dorsal aspect. Numbers and lowercase letters indicate primary setae and pores, respectively (some setae on abdominal segment VIII could not be identified). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1231.144746.figures34-37" httpUri="https://binary.pensoft.net/fig/1279885">36</figureCitation>
+; abdominal segments I – VII with several secondary setae; dorsal surface of abdominal segment VIII (Fig. 
+<figureCitation id="ABB22E6C2C217BEBA59613BDC056650A" captionStart="Figures 34–37" captionStartId="F9" captionText="Figures 34–37. Liocanthydrus nanops Baca et al., 2014, instar III 34 head, dorsal aspect 35 left prothoracic leg, anterior aspect 36 right prothoracic leg, posterior aspect 37 abdominal segment VIII, dorsal aspect. Numbers and lowercase letters indicate primary setae and pores, respectively (some setae on abdominal segment VIII could not be identified). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1231.144746.figures34-37" httpUri="https://binary.pensoft.net/fig/1279885">37</figureCitation>
+) with three elongate hair-like secondary setae and 2–3 minute secondary setae on anterior portion, 2–3 minute secondary setae on medial portion, and five hair like-like secondary seta and 4–5 minute secondary setae on posterior portion; ventral surface of abdominal segment VIII with two elongate hair-like secondary setae on anterior portion, one elongate hair-like secondary seta on medial portion, and one short and one long secondary setae on posterior portion.
+</paragraph>
+<caption id="40AE87F8CC844DB08E7B09B2CD312719" ID-Table-UUID="40AE87F8CC844DB08E7B09B2CD312719" httpUri="http://table.plazi.org/id/40AE87F8CC844DB08E7B09B2CD312719" startId="T2" targetIsTable="true">
+<paragraph id="E2DFDAD39B2618F7733408BCD7ACED89">
+<label id="13B1F781E1BBD64908FB6CE5BF4DEAA2">Table 2.</label>
+</paragraph>
+<paragraph id="612DF6BD2F4CF88F0F6BD0F3CF02D49F">
+Number and position of secondary setae on the legs of larvae of 
+<taxonomicName id="5C3665738EB0D16B6D694BA08B8A1B3F" authority="Baca et al. 2014" authorityName="Baca" authorityYear="2014" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="E8F9394543A451CF4658387292274A01" italics="true">Liocanthydrus nanops</emphasis>
+<bibRefCitation id="1D6874D84B8F708CF03AE38D7044759B" DOI="10.11646/zootaxa.3793.2.3" author="Baca" etAl="et al." firstAuthor="Baca" journalOrPublisher="Zootaxa" pagination="231-246" refId="B1" refString="Baca SM, Gustafson GT, Toledo M, Miller KB (2014) Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species. Zootaxa 3793: 231–246. https://doi.org/10.11646/zootaxa.3793.2.3" title="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species." volume="3793" year="2014">Baca et al. 2014</bibRefCitation>
+</taxonomicName>
+. Numbers between slash marks refer to pro-, meso-, and metathoracic leg, respectively. A = anterior, 
+<abbrev id="ABBRID0ETKAG" xlink_title="posterodorsal">PD</abbrev>
+= posterodorsal, 
+<abbrev id="ABBRID0EXKAG" xlink_title="posteroventral">PV</abbrev>
+= posteroventral; Total = total number of secondary setae on the article (i. e., excluding primary setae) (
+<emphasis id="3AC5F1748C5FD3DD54146ECF3B325EDF" italics="true">n</emphasis>
+= 1).
+</paragraph>
+</caption>
+<paragraph id="2D1B02E0876C8663B927395DDE3EAE35">
+<table id="TID0EGMAG" rules="all">
+<tbody id="D5471149941B2E41C4CF63917EEAC361">
+<tr id="235A6CF66495DDDC37BA7F38FF7B6A29">
+<th id="BBE667C5C631DF3E41E8938408D83D84" colspan="1" rowspan="1">Article</th>
+<th id="05C21B95B18483A045B12B07D21FD3B9" colspan="1" rowspan="1">Position</th>
+<th id="7196CF462AA4EE18DC07BFDDCA8EB644" colspan="1" rowspan="1">Instar III</th>
+</tr>
+<tr id="B492B02E29BA1E27040C4EB65C0ABCBC">
+<td id="3F7592CF388D10B8DF925C29B5E140BD" colspan="1" rowspan="4">Coxa</td>
+<td id="793AA3EBD40C0AB1D9EB39D0F9DF1E9A" colspan="1" rowspan="1">A</td>
+<td id="8992A443B0C959F2E7328568E81C5179" colspan="1" rowspan="1">0 / 2 / 2</td>
+</tr>
+<tr id="3BB1DC8773236FD9FAB95CBFDF0FC81E">
+<td id="C40015864BFFD2D30D1E95BA25D42145" colspan="1" rowspan="1">
+<abbrev id="ABBRID0E1LAG" xlink_title="posterodorsal">PD</abbrev>
+</td>
+<td id="4CE9492366B1EF7E26EE14E4C68BE891" colspan="1" rowspan="1">1 / 1 / 1</td>
+</tr>
+<tr id="17DA47FA7FB96207662E1497AFEE76FE">
+<td id="E3E28A89E1ECA1CA0D6ABF4C2F6F2099" colspan="1" rowspan="1">
+<abbrev id="ABBRID0EGMAG" xlink_title="posteroventral">PV</abbrev>
+</td>
+<td id="2D9EB18D2DBEF725002C467664B64600" colspan="1" rowspan="1">0 / 1 / 1</td>
+</tr>
+<tr id="64C85B6142AC4810ABFC77084CF2C497">
+<td id="EF78776A24D35E3B00FD78DC1561FFE5" colspan="1" rowspan="1">Total</td>
+<td id="AD61875777081C5AFD8A9161CA27AE36" colspan="1" rowspan="1">1 / 4 / 4</td>
+</tr>
+</tbody>
+</table>
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0EUMAG" type="remarks">
+<paragraph id="6D9F4D037853E3A81C42D58FDEE106C1">
+<heading id="4C93A94F86B1B8EEED12B0195A0816D7" reason="title">Remarks.</heading>
+</paragraph>
+<paragraph id="0C1F4BFC512C1CB044974809D3EC5ED6">
+When comparing the first and third instars of 
+<taxonomicName id="3D503DCEAB357A4C19A64696E12D6B0B" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="F494F588DBFE766EF2ABF6657D0FB730" italics="true">L. nanops</emphasis>
+</taxonomicName>
+with the supposedly third instar of 
+<taxonomicName id="8D049D3BD27F4575C0CB31FE3C9CCE05" authority="(Urcola et al. 2021)" baseAuthorityName="Urcola" baseAuthorityYear="2021" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="clayae">
+<emphasis id="CBE432078BB70BC5E490B0BDD03CF79B" italics="true">L. clayae</emphasis>
+(
+<bibRefCitation id="AEFCA6A908615A030059B1E1161F43D7" DOI="10.1080/00222933.2020.1842931" author="Urcola" etAl="et al." firstAuthor="Urcola" journalOrPublisher="Journal of Natural History" pagination="2285-2296" refId="B13" refString="Urcola JI, Benetti CJ, Alarie Y, Michat MC (2021) The unknown larva of the burrowing water beetle genus Liocanthydrus Guignot, 1957 (Coleoptera: Noteridae). Journal of Natural History 54: 2285–2296. https://doi.org/10.1080/00222933.2020.1842931" title="The unknown larva of the burrowing water beetle genus Liocanthydrus Guignot, 1957 (Coleoptera: Noteridae)." volume="54" year="2021">Urcola et al. 2021</bibRefCitation>
+)
+</taxonomicName>
+, we can conclude that the larva of this latter species is actually a second instar. This conclusion is based on the following evidence: the head of the larva of 
+<taxonomicName id="A370A13B8E46FAE74F8865DDFE50A398" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="clayae">
+<emphasis id="FBFD0CBBC8BECAD2E9E9CE3C8F0569D6" italics="true">L. clayae</emphasis>
+</taxonomicName>
+(as expressed in the head width) exhibits an intermediate size between the first and third instars of 
+<taxonomicName id="FBA6E58B1693C4A78CF55A67C67C2FB7" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="A9ABB37A60206BAA7C4D507B92E7B9EA" italics="true">L. nanops</emphasis>
+</taxonomicName>
+; the mandibles in 
+<taxonomicName id="18EB07765757121DC57C897A0B8B6BE4" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="clayae">
+<emphasis id="156008C090F9D97CCF8319658FFD9DCD" italics="true">L. clayae</emphasis>
+</taxonomicName>
+are not as robust as those of the third instar of 
+<taxonomicName id="E7E26C06651884732039ACEBB5732904" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="DFF5C9DE2DD035024D7C6AA64CCE2084" italics="true">L. nanops</emphasis>
+</taxonomicName>
+; and the siphon in 
+<taxonomicName id="7DBFBA8940D01131695ED214E24E5BAE" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="clayae">
+<emphasis id="653D3C5143B416BDA39F77F1DB47E06B" italics="true">L. clayae</emphasis>
+</taxonomicName>
+is more developed than that of the first instar of 
+<taxonomicName id="E61F971D1111C86CF052A0D722727A49" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="B205EBA15FF0DD43DC5B375D18B142F3" italics="true">L. nanops</emphasis>
+</taxonomicName>
+but not as strongly developed as that of the third instar of this species. Regarding chaetotaxy, seta PA 9 is absent on the parietal of the first instar of 
+<taxonomicName id="FCAF41C3717E7BE3A7C49DE49E54F5B1" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="4B203E9ABBAE682747D7BEE33854512C" italics="true">L. nanops</emphasis>
+</taxonomicName>
+. This conspicuous sensillum, however, is present in the third instar of this species (Fig. 
+<figureCitation id="80D952BFF7FF0849DE125CACAA813AED" captionStart="Figures 34–37" captionStartId="F9" captionText="Figures 34–37. Liocanthydrus nanops Baca et al., 2014, instar III 34 head, dorsal aspect 35 left prothoracic leg, anterior aspect 36 right prothoracic leg, posterior aspect 37 abdominal segment VIII, dorsal aspect. Numbers and lowercase letters indicate primary setae and pores, respectively (some setae on abdominal segment VIII could not be identified). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1231.144746.figures34-37" httpUri="https://binary.pensoft.net/fig/1279885">34</figureCitation>
+), as well as in all noterid larvae known in detail (e. g. 
+<bibRefCitation id="D28303ECFAFD9307002A59435F3B0A91" DOI="10.11646/zootaxa.4619.1.5" author="Urcola" etAl="et al." firstAuthor="Urcola" journalOrPublisher="Zootaxa" pagination="121-138" refId="B11" refString="Urcola JI, Alarie Y, Benetti CJ, Rodriguez G, Michat MC (2019) Larval morphology and analysis of primary chaetotaxy in the genus Suphis Aubé, 1836 (Coleoptera: Noteridae). Zootaxa 4619: 121–138. https://doi.org/10.11646/zootaxa.4619.1.5" title="Larval morphology and analysis of primary chaetotaxy in the genus Suphis Aubé, 1836 (Coleoptera: Noteridae)." volume="4619" year="2019">Urcola et al. 2019</bibRefCitation>
+, 
+<bibRefCitation id="ECE1FF53E8868D50A6853208369D775F" DOI="10.3161/00034541ANZ2020.70.4.012" author="Urcola" firstAuthor="Urcola" journalOrPublisher="Annales Zoologici" pagination="687-695" refId="B12" refString="Urcola JI, Alarie Y, Benetti CJ, Michat MC (2020) Description of the larval stages of Hydrocanthus socius Sahlberg, 1844 (Coleoptera: Noteridae) with chaetotaxy analysis. Annales Zoologici 70: 687–695. https://doi.org/10.3161/00034541ANZ2020.70.4.012" title="Description of the larval stages of Hydrocanthus socius Sahlberg, 1844 (Coleoptera: Noteridae) with chaetotaxy analysis." volume="70" year="2020">2020</bibRefCitation>
+, 
+<bibRefCitation id="FBCC9A316CA35F7572BAA62B9408F892" DOI="10.1080/00222933.2020.1842931" author="Urcola" firstAuthor="Urcola" journalOrPublisher="Journal of Natural History" pagination="2285-2296" refId="B13" refString="Urcola JI, Benetti CJ, Alarie Y, Michat MC (2021) The unknown larva of the burrowing water beetle genus Liocanthydrus Guignot, 1957 (Coleoptera: Noteridae). Journal of Natural History 54: 2285–2296. https://doi.org/10.1080/00222933.2020.1842931" title="The unknown larva of the burrowing water beetle genus Liocanthydrus Guignot, 1957 (Coleoptera: Noteridae)." volume="54" year="2021">2021</bibRefCitation>
+). Since we examined only a single specimen of each instar of 
+<taxonomicName id="26357829EEC905D94A9EBACD59B1400D" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nanops">
+<emphasis id="548C042B5F44A9C5537EB9B88B6FF147" italics="true">L. nanops</emphasis>
+</taxonomicName>
+, we prefer not to consider the absence of PA 
+<quantity id="81458A5E4E66D15291AE24587B99D810" metricMagnitude="-1" metricUnit="m" metricValue="2.286" unit="in" value="9.0">9 in</quantity>
+instar I as a diagnostic character for the species until more material can be studied.
+</paragraph>
+</subSubSection>
+</treatment>
+</document>
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