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<mods:title id="010BB5DECCEF9594521655CF40459F31">Revision of the genus Cenophengus LeConte, 1881 (Coleoptera, Phengodidae), with the description of four new species, new geographic records and a new synonymy</mods:title>
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<mods:namePart>Morrone, Juan J.</mods:namePart>
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<mods:affiliation>Museo de Zoologia " Alfonso L. Herrera ", Departamento de Biologia Evolutiva, Facultad de Ciencias, Universidad Nacional Autonoma de Mexico (UNAM), 04510 Mexico City, Mexico</mods:affiliation>
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<treatment ID-GBIF-Taxon="189997970" LSID="urn:lsid:plazi:treatment:16CFB7AD019054B79F70206AB5941961" httpUri="http://treatment.plazi.org/id/16CFB7AD019054B79F70206AB5941961" lastPageNumber="73" pageId="0" pageNumber="73">
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<taxonomicName LSID="16CFB7AD-0190-54B7-9F70-206AB5941961" authority="Wittmer, 1986" authorityName="Wittmer" authorityYear="1986" class="Insecta" family="Phengodidae" genus="Cenophengus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cenophengus niger" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="niger">Cenophengus niger Wittmer, 1986</taxonomicName>
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<treatment id="16CFB7AD019054B79F70206AB5941961" ID-GBIF-Taxon="189997970" LSID="urn:lsid:plazi:treatment:16CFB7AD019054B79F70206AB5941961" httpUri="http://treatment.plazi.org/id/16CFB7AD019054B79F70206AB5941961" lastPageNumber="73" pageId="0" pageNumber="73" scope_family="Phengodidae" scope_order="Coleoptera">
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<paragraph id="841F8F66F3D7828A54F85D92174BA372" pageId="0" pageNumber="73">
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<taxonomicName id="640C2A719183E251A11F818D06D3F274" ID-CoL="S5WS" LSID="16CFB7AD-0190-54B7-9F70-206AB5941961" authority="Wittmer, 1986" authorityName="Wittmer" authorityYear="1986" class="Insecta" family="Phengodidae" genus="Cenophengus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cenophengus niger" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="niger">Cenophengus niger Wittmer, 1986</taxonomicName>
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</paragraph>
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<subSubSection pageId="0" pageNumber="73" type="description">
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<paragraph pageId="0" pageNumber="73">
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Cenophengus niger Wittmer, 1986, male. Habitus: A dorsal B ventral C head dorsal D pronotum dorsal E hind wing. Wing venation: CR = Radial Cell; r 4 = radial 4 vein; RP = Posterior Radial vein; MP 1 + 2 = Posterior Median vein; CuA = Cubital vein; AA and AP = Anterior and Posterior Anal veins. Aedeagus: F dorsal view G lateral view H ventral view." figureDoi="10.3897/zookeys.1068.70295.figure22" httpUri="https://binary.pensoft.net/fig/607303" pageId="0" pageNumber="73">Fig. 22A-H</figureCitation>
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<paragraph id="C902D52B5AA716ED4FDE86F27FF50C26" pageId="0" pageNumber="73">
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<figureCitation id="40CC3676E9B1F17C46D60D0B4D8D7A26" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Cenophengus niger Wittmer, 1986, male. Habitus: A dorsal B ventral C head dorsal D pronotum dorsal E hind wing. Wing venation: CR = Radial Cell; r 4 = radial 4 vein; RP = Posterior Radial vein; MP 1 + 2 = Posterior Median vein; CuA = Cubital vein; AA and AP = Anterior and Posterior Anal veins. Aedeagus: F dorsal view G lateral view H ventral view." figureDoi="10.3897/zookeys.1068.70295.figure22" httpUri="https://binary.pensoft.net/fig/607303" pageId="0" pageNumber="73">Fig. 22A-H</figureCitation>
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</paragraph>
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<subSubSection pageId="0" pageNumber="73" type="reference_group">
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<paragraph pageId="0" pageNumber="73">
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<taxonomicName authorityName="Wittmer" authorityYear="1986" class="Insecta" family="Phengodidae" genus="Cenophengus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cenophengus niger" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="niger">Cenophengus niger</taxonomicName>
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<subSubSection id="64A36BDCBF4F3E3EAA0565FF6974031E" pageId="0" pageNumber="73" type="reference_group">
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<paragraph id="2182ED9FC1F1BB28DE09DEB229EE9CFD" pageId="0" pageNumber="73">
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<taxonomicName id="B7D7F2532182C3170B5144EF86B52590" authorityName="Wittmer" authorityYear="1986" class="Insecta" family="Phengodidae" genus="Cenophengus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cenophengus niger" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="niger">Cenophengus niger</taxonomicName>
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||||
Wittmer, 1986: 160.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="type locality">
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||||
<paragraph pageId="0" pageNumber="73">Type locality.</paragraph>
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||||
<paragraph pageId="0" pageNumber="73">Monteverde, Costa Rica.</paragraph>
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<subSubSection id="9B7A7C9D6BAEF3694194BA97C4926C68" pageId="0" pageNumber="73" type="type locality">
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<paragraph id="ECDE343A15C3E6F0D6AD69199464D5D4" pageId="0" pageNumber="73">Type locality.</paragraph>
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<paragraph id="D6E8B7DF1C43076FAF2C8D87E19EE571" pageId="0" pageNumber="73">Monteverde, Costa Rica.</paragraph>
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||||
</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="type material">
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<paragraph pageId="0" pageNumber="73">Type material examined.</paragraph>
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||||
<paragraph pageId="0" pageNumber="73">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Holotype</emphasis>
|
||||
<subSubSection id="33B3BBE1B300BB1FA1C140C9A9BECA8F" pageId="0" pageNumber="73" type="type material">
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||||
<paragraph id="720FACE47E8A237E79884461D16192AF" pageId="0" pageNumber="73">Type material examined.</paragraph>
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||||
<paragraph id="2C5686724C2DD8ECA46B651BCCE1437E" pageId="0" pageNumber="73">
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<emphasis id="866486F69A25FF48BB1F1BC2E3AE39D7" bold="true" italics="true" pageId="0" pageNumber="73">Holotype</emphasis>
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♂: "COSTA RICA: Punt. / Monteverde. 1400 m/ 23 May 1979/ H & A Howden" "
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<emphasis italics="true" pageId="0" pageNumber="73">
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<taxonomicName authorityName="LeConte" authorityYear="1881" class="Insecta" family="Phengodidae" genus="Cenophengus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cenophengus" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Cenophengus</taxonomicName>
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||||
<emphasis id="325E14EA014CE1786D62679782A7EFC3" italics="true" pageId="0" pageNumber="73">
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||||
<taxonomicName id="547FFC47583FF5F01B914EC4364998E9" authorityName="LeConte" authorityYear="1881" class="Insecta" family="Phengodidae" genus="Cenophengus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cenophengus" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Cenophengus</taxonomicName>
|
||||
/
|
||||
<taxonomicName authorityName="Wittmer" authorityYear="1986" class="Insecta" family="Phengodidae" genus="Cenophengus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cenophengus niger" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="niger">Cenophengus niger</taxonomicName>
|
||||
<taxonomicName id="142D5184BAEA6B30453D6FE59C067155" authorityName="Wittmer" authorityYear="1986" class="Insecta" family="Phengodidae" genus="Cenophengus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cenophengus niger" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="niger">Cenophengus niger</taxonomicName>
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||||
</emphasis>
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Wittmer" "
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||||
<taxonomicName authorityName="LeConte" authorityYear="1861" class="Insecta" family="Phengodidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="family">PHENGODIDAE</taxonomicName>
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<taxonomicName id="7AF540E15C0E9014E92D1DBB0019C421" authorityName="LeConte" authorityYear="1861" class="Insecta" family="Phengodidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="family">PHENGODIDAE</taxonomicName>
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/ PHENG00000347" | NHMB.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="remarks">
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<paragraph pageId="0" pageNumber="73">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="73">
|
||||
<taxonomicName authorityName="Wittmer" authorityYear="1986" class="Insecta" family="Phengodidae" genus="Cenophengus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cenophengus niger" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis italics="true" pageId="0" pageNumber="73">Cenophengus niger</emphasis>
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||||
<subSubSection id="01CBE3A696AF610E36C6C9434969285B" pageId="0" pageNumber="73" type="remarks">
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||||
<paragraph id="B74FD28A02917A404F80CA7B66508F47" pageId="0" pageNumber="73">Remarks.</paragraph>
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||||
<paragraph id="80D485797D8605FBEAC9041DA9B40E96" pageId="0" pageNumber="73">
|
||||
<taxonomicName id="08D9C54F2227ECDA26C6A16C1D3A5ACB" authorityName="Wittmer" authorityYear="1986" class="Insecta" family="Phengodidae" genus="Cenophengus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cenophengus niger" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="3CA66829D07EFB4CCEFD8DCD8A71E7C1" italics="true" pageId="0" pageNumber="73">Cenophengus niger</emphasis>
|
||||
</taxonomicName>
|
||||
is morphologically similar to
|
||||
<taxonomicName lsidName="C. howdeni" pageId="0" pageNumber="73" rank="species" species="howdeni">
|
||||
<emphasis italics="true" pageId="0" pageNumber="73">C. howdeni</emphasis>
|
||||
<taxonomicName id="5034474437090831A58ABF526340EF63" lsidName="C. howdeni" pageId="0" pageNumber="73" rank="species" species="howdeni">
|
||||
<emphasis id="A0C43ECF4C41C9732577ACF07D893DB8" italics="true" pageId="0" pageNumber="73">C. howdeni</emphasis>
|
||||
</taxonomicName>
|
||||
, but can be distinguished by the length of antennomere 1 and the diameters of the punctures. In
|
||||
<taxonomicName lsidName="C. niger" pageId="0" pageNumber="73" rank="species" species="niger">
|
||||
<emphasis italics="true" pageId="0" pageNumber="73">C. niger</emphasis>
|
||||
<taxonomicName id="76C9A5ECBEE12EFDBFE83BCC9AC0D5AB" lsidName="C. niger" pageId="0" pageNumber="73" rank="species" species="niger">
|
||||
<emphasis id="BE97CC9F8012CE9199368CA5D4FFCE4D" italics="true" pageId="0" pageNumber="73">C. niger</emphasis>
|
||||
</taxonomicName>
|
||||
, antennomere 1 is equal to the length of antennomeres 2 and 3 combined, whereas in
|
||||
<taxonomicName lsidName="C. howdeni" pageId="0" pageNumber="73" rank="species" species="howdeni">
|
||||
<emphasis italics="true" pageId="0" pageNumber="73">C. howdeni</emphasis>
|
||||
<taxonomicName id="9F71798768388352C2358194CCB308DB" lsidName="C. howdeni" pageId="0" pageNumber="73" rank="species" species="howdeni">
|
||||
<emphasis id="95C8D2C80A2BF7C8C9C6F92AFD424700" italics="true" pageId="0" pageNumber="73">C. howdeni</emphasis>
|
||||
</taxonomicName>
|
||||
, it is shorter than antennomeres 2 and 3 combined. In
|
||||
<taxonomicName lsidName="C. niger" pageId="0" pageNumber="73" rank="species" species="niger">
|
||||
<emphasis italics="true" pageId="0" pageNumber="73">C. niger</emphasis>
|
||||
<taxonomicName id="BDC84B02B0020B9138C01D4081D44E1E" lsidName="C. niger" pageId="0" pageNumber="73" rank="species" species="niger">
|
||||
<emphasis id="69981F5B9E1EAAA2F516DC80BAC4A5D9" italics="true" pageId="0" pageNumber="73">C. niger</emphasis>
|
||||
</taxonomicName>
|
||||
, the punctures are twice as long as eye facets and separated by approximately 0.2 punctured diameters, whereas in
|
||||
<taxonomicName lsidName="C. howdeni" pageId="0" pageNumber="73" rank="species" species="howdeni">
|
||||
<emphasis italics="true" pageId="0" pageNumber="73">C. howdeni</emphasis>
|
||||
<taxonomicName id="5809F9C4E805A98BB54B6E2B85A83BD3" lsidName="C. howdeni" pageId="0" pageNumber="73" rank="species" species="howdeni">
|
||||
<emphasis id="C97892D78FFBC1FCAFB2CB1CFDC9E26F" italics="true" pageId="0" pageNumber="73">C. howdeni</emphasis>
|
||||
</taxonomicName>
|
||||
, they are as long as eye facets and separated by approximately 1 punctured diameter.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="73" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="73">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="73">Integument chagreened, head less wide than pronotum, antennae less than twice the length of the pronotum, antennal rami twice as long as the respective antennomere and each elytron 3.5 times as long as wide; aedeagus with three teeth at the inner apex of paramere.</paragraph>
|
||||
<subSubSection id="EEA256BD3C7DA3E141FED19C632D3FE9" pageId="0" pageNumber="73" type="diagnosis">
|
||||
<paragraph id="3DE77285BA424AA0D987B86DF68609D2" pageId="0" pageNumber="73">Diagnosis.</paragraph>
|
||||
<paragraph id="C249F91351A3C9914F8CF035408175BD" pageId="0" pageNumber="73">Integument chagreened, head less wide than pronotum, antennae less than twice the length of the pronotum, antennal rami twice as long as the respective antennomere and each elytron 3.5 times as long as wide; aedeagus with three teeth at the inner apex of paramere.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="73" type="redescription">
|
||||
<paragraph pageId="0" pageNumber="73">Redescription.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="73">
|
||||
<emphasis bold="true" pageId="0" pageNumber="73">Male.</emphasis>
|
||||
<subSubSection id="A3C49C164452C5B0EA1EF3BDC7DCFFEF" pageId="0" pageNumber="73" type="redescription">
|
||||
<paragraph id="EEB7C524CE35F06E2A6F0373E22FC7F9" pageId="0" pageNumber="73">Redescription.</paragraph>
|
||||
<paragraph id="B467371EA964D0E2268D17CB22A4AAAA" pageId="0" pageNumber="73">
|
||||
<emphasis id="3A6F9A738869ADFAE478B18B1C423263" bold="true" pageId="0" pageNumber="73">Male.</emphasis>
|
||||
Body length 6.0-6.3 mm, maximum body width 0.80-0.85 mm (pronotum). Body dark brown, only mouthparts, three first antennomeres, two last abdominal segments, all legs with trochanter and coxae yellowish (Fig.
|
||||
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Cenophengus niger Wittmer, 1986, male. Habitus: A dorsal B ventral C head dorsal D pronotum dorsal E hind wing. Wing venation: CR = Radial Cell; r 4 = radial 4 vein; RP = Posterior Radial vein; MP 1 + 2 = Posterior Median vein; CuA = Cubital vein; AA and AP = Anterior and Posterior Anal veins. Aedeagus: F dorsal view G lateral view H ventral view." figureDoi="10.3897/zookeys.1068.70295.figure22" httpUri="https://binary.pensoft.net/fig/607303" pageId="0" pageNumber="73">22A, B</figureCitation>
|
||||
<figureCitation id="BF4B69598B5951EF4459FC27DD28FA42" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Cenophengus niger Wittmer, 1986, male. Habitus: A dorsal B ventral C head dorsal D pronotum dorsal E hind wing. Wing venation: CR = Radial Cell; r 4 = radial 4 vein; RP = Posterior Radial vein; MP 1 + 2 = Posterior Median vein; CuA = Cubital vein; AA and AP = Anterior and Posterior Anal veins. Aedeagus: F dorsal view G lateral view H ventral view." figureDoi="10.3897/zookeys.1068.70295.figure22" httpUri="https://binary.pensoft.net/fig/607303" pageId="0" pageNumber="73">22A, B</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="73">
|
||||
<emphasis italics="true" pageId="0" pageNumber="73">Head</emphasis>
|
||||
<emphasis id="4353814ADFAF49B444C64AA3273D6EE8" bold="true" pageId="0" pageNumber="73">
|
||||
<emphasis id="96BE375A0E23353B1156D3ABB222ADE5" italics="true" pageId="0" pageNumber="73">Head</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Wider (0.80-0.83 mm) (0.81
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="3D15B7DCFB38B05850A726ED25CA428D" originalValue="±">+/-</normalizedToken>
|
||||
0.0.17 mm, n = 3) than long (0.55-0.60 mm) (0.56
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="E29F217443171D4294AE7FB2B32E0A5F" originalValue="±">+/-</normalizedToken>
|
||||
0.028 mm, n = 3) (Fig.
|
||||
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Cenophengus niger Wittmer, 1986, male. Habitus: A dorsal B ventral C head dorsal D pronotum dorsal E hind wing. Wing venation: CR = Radial Cell; r 4 = radial 4 vein; RP = Posterior Radial vein; MP 1 + 2 = Posterior Median vein; CuA = Cubital vein; AA and AP = Anterior and Posterior Anal veins. Aedeagus: F dorsal view G lateral view H ventral view." figureDoi="10.3897/zookeys.1068.70295.figure22" httpUri="https://binary.pensoft.net/fig/607303" pageId="0" pageNumber="73">22C</figureCitation>
|
||||
<figureCitation id="A6872D32F4D4B7AF5905B5DCB163A2FA" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Cenophengus niger Wittmer, 1986, male. Habitus: A dorsal B ventral C head dorsal D pronotum dorsal E hind wing. Wing venation: CR = Radial Cell; r 4 = radial 4 vein; RP = Posterior Radial vein; MP 1 + 2 = Posterior Median vein; CuA = Cubital vein; AA and AP = Anterior and Posterior Anal veins. Aedeagus: F dorsal view G lateral view H ventral view." figureDoi="10.3897/zookeys.1068.70295.figure22" httpUri="https://binary.pensoft.net/fig/607303" pageId="0" pageNumber="73">22C</figureCitation>
|
||||
), at eye level, less wide than the pronotum, integument chagreened, punctures twice as long as eye facets and separated by approximately 0.2 punctured diameters, each puncture bearing a yellow-brown seta; interantennal distance (0.08-0.10 mm) (0.089
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="FCC8618D4D77DF4101AEBE1E07776BE7" originalValue="±">+/-</normalizedToken>
|
||||
0.11 mm, n = 3) less than the length of antennomere 1 (0.14-0.15 mm) (0.146
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="68ED54B4815831E929E1FC6BB89324C5" originalValue="±">+/-</normalizedToken>
|
||||
0.005 mm, n = 3); eyes 1/2 as long as head in lateral view, longer (0.26-0.28 mm) (0.266
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="B3F9D224868BD8867964CB5328B05A70" originalValue="±">+/-</normalizedToken>
|
||||
0.011 mm, n = 3) than wide (0.12-0.14 mm) (0.126
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="6BCAB6660D49CBD2340EEC97AA6FEF48" originalValue="±">+/-</normalizedToken>
|
||||
0.011 mm, n = 3); interocular distance (0.40-0.42 mm) (0.403
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="6181AC12977A875085A0C1934B3D67AC" originalValue="±">+/-</normalizedToken>
|
||||
0.011 mm, n = 3) 3 times eye width; antennae short (2.2-2.3 mm) (2.26
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="B0DA5DA84642D4A95FF5781886F881A5" originalValue="±">+/-</normalizedToken>
|
||||
0.057 mm, n = 3) less than twice the length of the pronotum; antennomere 1 (0.14-0.15 mm) (0.146
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="09DD4BE799CB9DAAE04553FC8782AFD6" originalValue="±">+/-</normalizedToken>
|
||||
0.005 mm, n = 3) equal to the length of the next two combined, antennomere 3 cup-shaped, 4 (0.13-0.14 mm) (0.136
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="4B93DC23246CFFC95996101199FA3EB8" originalValue="±">+/-</normalizedToken>
|
||||
0.005 mm, n = 3) shorter than following antennomeres, 5 to 11 about equal in length (0.15-0.16) (0.156
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="F780527A6DA1964FEFF5357B32BA0972" originalValue="±">+/-</normalizedToken>
|
||||
0.005, n = 3), 12 (terminal) (0.20-0.21 mm) (0.206
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="671953F0A471B294E5117489304849B7" originalValue="±">+/-</normalizedToken>
|
||||
0.005 mm, n = 3), antennal rami lanceolate in lateral view, twice as long as the respective antennomere; terminal maxillary palpomere robust, securiform (0.25-0.27 mm) (0.263
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="F5773CCB4C4103DA54537E930CE73C80" originalValue="±">+/-</normalizedToken>
|
||||
0.011 mm, n = 3), as long as the preceding three combined; terminal labial palpomere spindle-shaped (0.05-0.06 mm) (0.056
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="80F10FBFC22002BE218F9C88556A0B9A" originalValue="±">+/-</normalizedToken>
|
||||
0.005 mm, n = 3), 3 times as long as preceding one (0.02 mm) (0.02
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="C16C02B354164E594ADC42190B79AA4E" originalValue="±">+/-</normalizedToken>
|
||||
0 mm, n = 3).
|
||||
<emphasis bold="true" pageId="0" pageNumber="73">
|
||||
<emphasis italics="true" pageId="0" pageNumber="73">Thorax</emphasis>
|
||||
<emphasis id="A3534CC70F809907DF0DB962714A68C6" bold="true" pageId="0" pageNumber="73">
|
||||
<emphasis id="26AA41247AA2B49A1647F8671BBB6093" italics="true" pageId="0" pageNumber="73">Thorax</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Pronotum longer (0.95-0.98 mm) (0.966
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="F184B647A7080B6EA34629BE907819A1" originalValue="±">+/-</normalizedToken>
|
||||
0.015 mm, n = 3) than wide (0.80-0.85 mm) (0.833
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="31EAF54E5323404D395219FF55A3D6F4" originalValue="±">+/-</normalizedToken>
|
||||
0.0.11 mm, n = 3) (Fig.
|
||||
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Cenophengus niger Wittmer, 1986, male. Habitus: A dorsal B ventral C head dorsal D pronotum dorsal E hind wing. Wing venation: CR = Radial Cell; r 4 = radial 4 vein; RP = Posterior Radial vein; MP 1 + 2 = Posterior Median vein; CuA = Cubital vein; AA and AP = Anterior and Posterior Anal veins. Aedeagus: F dorsal view G lateral view H ventral view." figureDoi="10.3897/zookeys.1068.70295.figure22" httpUri="https://binary.pensoft.net/fig/607303" pageId="0" pageNumber="73">22D</figureCitation>
|
||||
<figureCitation id="EC237ACA11B46287DC3A94009E0D6B4B" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Cenophengus niger Wittmer, 1986, male. Habitus: A dorsal B ventral C head dorsal D pronotum dorsal E hind wing. Wing venation: CR = Radial Cell; r 4 = radial 4 vein; RP = Posterior Radial vein; MP 1 + 2 = Posterior Median vein; CuA = Cubital vein; AA and AP = Anterior and Posterior Anal veins. Aedeagus: F dorsal view G lateral view H ventral view." figureDoi="10.3897/zookeys.1068.70295.figure22" httpUri="https://binary.pensoft.net/fig/607303" pageId="0" pageNumber="73">22D</figureCitation>
|
||||
); integument chagreened, punctures twice as long as eye facets and separated by approximately 0.2 punctured diameters, with a yellow-brown seta in each puncture; disc convex, weakly elevated dorsally forming a small depression in the basal part of each side, posterior margin curved, sides almost straight, anterior angles rounded and posterior angles acute; mesosternal suture complete; scutellum with posterior margin rounded; each elytron 3.5 times as long (1.62-1.74 mm) (1.68
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="DF7A71F2F2031A4D519150B1D52F0E01" originalValue="±">+/-</normalizedToken>
|
||||
0.061 mm, n = 3) as wide (0.46-0.48 mm) (0.473
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="CE09562F54E803FEEABDB522476A8575" originalValue="±">+/-</normalizedToken>
|
||||
0.011 mm, n = 3), convex, without longitudinal costae, elytral apex rounded; hind wings with posterior radial vein (RP) length 6.5 times less than the length of MP1+2, radial cell closed, r3 vein absent, r4 vein developed (reaching the radial cell), those of the anterior anal and posterior anal sectors, evident (Fig.
|
||||
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Cenophengus niger Wittmer, 1986, male. Habitus: A dorsal B ventral C head dorsal D pronotum dorsal E hind wing. Wing venation: CR = Radial Cell; r 4 = radial 4 vein; RP = Posterior Radial vein; MP 1 + 2 = Posterior Median vein; CuA = Cubital vein; AA and AP = Anterior and Posterior Anal veins. Aedeagus: F dorsal view G lateral view H ventral view." figureDoi="10.3897/zookeys.1068.70295.figure22" httpUri="https://binary.pensoft.net/fig/607303" pageId="0" pageNumber="73">22E</figureCitation>
|
||||
<figureCitation id="0D5D4E8823ED24241663DF9337C12BD8" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Cenophengus niger Wittmer, 1986, male. Habitus: A dorsal B ventral C head dorsal D pronotum dorsal E hind wing. Wing venation: CR = Radial Cell; r 4 = radial 4 vein; RP = Posterior Radial vein; MP 1 + 2 = Posterior Median vein; CuA = Cubital vein; AA and AP = Anterior and Posterior Anal veins. Aedeagus: F dorsal view G lateral view H ventral view." figureDoi="10.3897/zookeys.1068.70295.figure22" httpUri="https://binary.pensoft.net/fig/607303" pageId="0" pageNumber="73">22E</figureCitation>
|
||||
). Legs: tarsomere 1 of pro-, meso- and metathoracic legs is longer than 2.
|
||||
<emphasis bold="true" pageId="0" pageNumber="73">
|
||||
<emphasis italics="true" pageId="0" pageNumber="73">Abdomen</emphasis>
|
||||
<emphasis id="F0DC05D37E307DD0095DF8D6717E416A" bold="true" pageId="0" pageNumber="73">
|
||||
<emphasis id="4524C7F2F41612C1BEF896EB1AD0F66A" italics="true" pageId="0" pageNumber="73">Abdomen</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Integument shiny, punctured, with long dense setae, sternite 7 with margin sinuate, sternite 8 with margin notched; aedeagus with three teeth at the inner apex of paramere (Fig.
|
||||
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Cenophengus niger Wittmer, 1986, male. Habitus: A dorsal B ventral C head dorsal D pronotum dorsal E hind wing. Wing venation: CR = Radial Cell; r 4 = radial 4 vein; RP = Posterior Radial vein; MP 1 + 2 = Posterior Median vein; CuA = Cubital vein; AA and AP = Anterior and Posterior Anal veins. Aedeagus: F dorsal view G lateral view H ventral view." figureDoi="10.3897/zookeys.1068.70295.figure22" httpUri="https://binary.pensoft.net/fig/607303" pageId="0" pageNumber="73">22F-H</figureCitation>
|
||||
<figureCitation id="72B11B95253049BEB563F7467473F722" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Cenophengus niger Wittmer, 1986, male. Habitus: A dorsal B ventral C head dorsal D pronotum dorsal E hind wing. Wing venation: CR = Radial Cell; r 4 = radial 4 vein; RP = Posterior Radial vein; MP 1 + 2 = Posterior Median vein; CuA = Cubital vein; AA and AP = Anterior and Posterior Anal veins. Aedeagus: F dorsal view G lateral view H ventral view." figureDoi="10.3897/zookeys.1068.70295.figure22" httpUri="https://binary.pensoft.net/fig/607303" pageId="0" pageNumber="73">22F-H</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1068.70295.figure22" httpUri="https://binary.pensoft.net/fig/607303" pageId="0" pageNumber="73" start="Figure 22" startId="F22">
|
||||
<paragraph pageId="0" pageNumber="73">
|
||||
<emphasis bold="true" pageId="0" pageNumber="73">Figure 22.</emphasis>
|
||||
<taxonomicName authorityName="Wittmer" authorityYear="1986" class="Insecta" family="Phengodidae" genus="Cenophengus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cenophengus niger" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis italics="true" pageId="0" pageNumber="73">Cenophengus niger</emphasis>
|
||||
<caption id="89110A4E1F4DF0A9AF5DF5C926E2A41C" doi="10.3897/zookeys.1068.70295.figure22" httpUri="https://binary.pensoft.net/fig/607303" pageId="0" pageNumber="73" start="Figure 22" startId="F22">
|
||||
<paragraph id="E89A60A3F704FBDB5EFA3348DC3E414C" pageId="0" pageNumber="73">
|
||||
<emphasis id="43F591D1258E35620FD77BFB532CD67A" bold="true" pageId="0" pageNumber="73">Figure 22.</emphasis>
|
||||
<taxonomicName id="85E23EB37B5CA1C0672D7AE9290C4B1D" authorityName="Wittmer" authorityYear="1986" class="Insecta" family="Phengodidae" genus="Cenophengus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cenophengus niger" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="9F3B124D7600AF119A4E096969665F71" italics="true" pageId="0" pageNumber="73">Cenophengus niger</emphasis>
|
||||
</taxonomicName>
|
||||
Wittmer, 1986, male. Habitus:
|
||||
<emphasis bold="true" pageId="0" pageNumber="73">A</emphasis>
|
||||
<emphasis id="922C52BBDF44301080401BC0B507B242" bold="true" pageId="0" pageNumber="73">A</emphasis>
|
||||
dorsal
|
||||
<emphasis bold="true" pageId="0" pageNumber="73">B</emphasis>
|
||||
<emphasis id="FCEB6201DED2BB8C530C69F25AFA710E" bold="true" pageId="0" pageNumber="73">B</emphasis>
|
||||
ventral
|
||||
<emphasis bold="true" pageId="0" pageNumber="73">C</emphasis>
|
||||
<emphasis id="922C47A62B371889F859683A9B54CCF1" bold="true" pageId="0" pageNumber="73">C</emphasis>
|
||||
head dorsal
|
||||
<emphasis bold="true" pageId="0" pageNumber="73">D</emphasis>
|
||||
<emphasis id="4DFF57EE8F085B49EB552242D1F76571" bold="true" pageId="0" pageNumber="73">D</emphasis>
|
||||
pronotum dorsal
|
||||
<emphasis bold="true" pageId="0" pageNumber="73">E</emphasis>
|
||||
<emphasis id="815AB9BAD83E02C641C27DF8BE934CE3" bold="true" pageId="0" pageNumber="73">E</emphasis>
|
||||
hind wing. Wing venation: CR = Radial Cell; r4 = radial 4 vein; RP = Posterior Radial vein; MP1+2 = Posterior Median vein; CuA = Cubital vein; AA and AP = Anterior and Posterior Anal veins. Aedeagus:
|
||||
<emphasis bold="true" pageId="0" pageNumber="73">F</emphasis>
|
||||
<emphasis id="3A98F8010188F3259912C29B23B9D05F" bold="true" pageId="0" pageNumber="73">F</emphasis>
|
||||
dorsal view
|
||||
<emphasis bold="true" pageId="0" pageNumber="73">G</emphasis>
|
||||
<emphasis id="8C79D7E9D566F5B20307F1929CDAF1C1" bold="true" pageId="0" pageNumber="73">G</emphasis>
|
||||
lateral view
|
||||
<emphasis bold="true" pageId="0" pageNumber="73">H</emphasis>
|
||||
<emphasis id="E1059ED628F9B6F85AD83F777D9C347B" bold="true" pageId="0" pageNumber="73">H</emphasis>
|
||||
ventral view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="73" type="female and immatures">
|
||||
<paragraph pageId="0" pageNumber="73">Female and immatures.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="73">Unknown.</paragraph>
|
||||
<subSubSection id="16E54C00DFC0EDE2D11C980D9468F42C" pageId="0" pageNumber="73" type="female and immatures">
|
||||
<paragraph id="A11269602491110649E0FC0D592A9A51" pageId="0" pageNumber="73">Female and immatures.</paragraph>
|
||||
<paragraph id="41F0000AA797C1745DBBB401689AAE81" pageId="0" pageNumber="73">Unknown.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="73" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="73">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="73">
|
||||
<subSubSection id="442FEC473C22DCB56BC9C7982B8AEB03" pageId="0" pageNumber="73" type="distribution">
|
||||
<paragraph id="451045F7E48FF9D32FCDE535311535BA" pageId="0" pageNumber="73">Distribution.</paragraph>
|
||||
<paragraph id="BCE7D21A3598EF9BBCF3E99311A69BA4" pageId="0" pageNumber="73">
|
||||
Costa Rica: Monteverde, Heredia and Puntarenas (Fig.
|
||||
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Map of southern North America showing specimen localities for Cenophengus spp. (continued)." figureDoi="10.3897/zookeys.1068.70295.figure19" httpUri="https://binary.pensoft.net/fig/607300" pageId="0" pageNumber="73">19</figureCitation>
|
||||
<figureCitation id="73E87A8AFDC5E45E92EE83C8A82A337B" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Map of southern North America showing specimen localities for Cenophengus spp. (continued)." figureDoi="10.3897/zookeys.1068.70295.figure19" httpUri="https://binary.pensoft.net/fig/607300" pageId="0" pageNumber="73">19</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="73" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="73">Additional material examined.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="73">
|
||||
<subSubSection id="FAB3993A48EB51FBC06906050E011A2D" pageId="0" pageNumber="73" type="materials_examined">
|
||||
<paragraph id="0F1B7C8A2AD45AF0AB054FD70D609237" pageId="0" pageNumber="73">Additional material examined.</paragraph>
|
||||
<paragraph id="8F1BEA8B3789C7A42262E2CAAD46AD1E" pageId="0" pageNumber="73">
|
||||
"
|
||||
<materialsCitation collectingDate="1991-07" collectingDateMax="1992-09" collectingDateMin="1991-07" collectorName="P. Hansen, P. Hanson. Malaise" country="COSTA RICA" elevation="75" latitude="10.433333" location="La Selva" longLatPrecision="1298" longitude="-84.01667" specimenCount="1" stateProvince="Heredia">
|
||||
<collectingCountry name="Costa Rica">COSTA RICA</collectingCountry>
|
||||
<materialsCitation id="A2E053BE748628B2A064D3CABE02CA16" collectingDate="1991-07" collectingDateMax="1992-09" collectingDateMin="1991-07" collectorName="P. Hansen, P. Hanson. Malaise" country="COSTA RICA" elevation="75" latitude="10.433333" location="La Selva" longLatPrecision="1298" longitude="-84.01667" specimenCount="1" stateProvince="Heredia">
|
||||
<collectingCountry id="4D63A7C73E006613B3A5263C223717B3" name="Costa Rica">COSTA RICA</collectingCountry>
|
||||
:
|
||||
<collectingRegion country="Costa Rica" name="Heredia">Heredia</collectingRegion>
|
||||
<collectingRegion id="E48B01A76075217900D5672CCEBAB262" country="Costa Rica" name="Heredia">Heredia</collectingRegion>
|
||||
/
|
||||
<location LSID="urn:lsid:plazi:treatment:16CFB7AD019054B79F70206AB5941961:4247142B9DB60D7F3224264E12F57A47" country="COSTA RICA" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" name="La Selva" stateProvince="Heredia">La Selva</location>
|
||||
<location id="4247142B9DB60D7F3224264E12F57A47" LSID="urn:lsid:plazi:treatment:16CFB7AD019054B79F70206AB5941961:4247142B9DB60D7F3224264E12F57A47" country="COSTA RICA" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" name="La Selva" stateProvince="Heredia">La Selva</location>
|
||||
,
|
||||
<quantity metricMagnitude="1" metricUnit="m" metricValue="7.5" unit="m" value="75.0">
|
||||
<elevation metricMagnitude="1" metricUnit="m" metricValue="7.5" unit="m" value="75.0">75 m</elevation>
|
||||
<quantity id="C3CD49BFC94AD6F2D76DEF686AB76CAF" metricMagnitude="1" metricUnit="m" metricValue="7.5" unit="m" value="75.0">
|
||||
<elevation id="D2E058F8863E2E941783D4C136F7E01B" metricMagnitude="1" metricUnit="m" metricValue="7.5" unit="m" value="75.0">75 m</elevation>
|
||||
</quantity>
|
||||
/
|
||||
<geoCoordinate degrees="10" direction="north" minutes="26" orientation="latitude" precision="925" value="10.433333">10°26'N</geoCoordinate>
|
||||
<geoCoordinate id="8BB6751CE8CE1521CE4F2ECF435FAC57" degrees="10" direction="north" minutes="26" orientation="latitude" precision="925" value="10.433333">10°26'N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="84" direction="west" minutes="01" orientation="longitude" precision="925" value="-84.01667">84°01'W</geoCoordinate>
|
||||
<geoCoordinate id="10CFB7B982E1CAC87C6461E38B016C81" degrees="84" direction="west" minutes="01" orientation="longitude" precision="925" value="-84.01667">84°01'W</geoCoordinate>
|
||||
/
|
||||
<collectingDate value="1992-09">Sept 1992</collectingDate>
|
||||
<collectingDate id="18ED8BA22F7741288B4C46325107E88C" value="1992-09">Sept 1992</collectingDate>
|
||||
/
|
||||
<collectorName>P. Hansen</collectorName>
|
||||
<collectorName id="9A6600D7F235E33D7421D581EC2F9A1D">P. Hansen</collectorName>
|
||||
, Malaise" "From the Michael/ Ivie Collection" (2) | MTEC; "
|
||||
<collectingCountry name="Costa Rica">COSTA RICA</collectingCountry>
|
||||
<collectingCountry id="F2409260F6D5E916FFF263444BE5671E" name="Costa Rica">COSTA RICA</collectingCountry>
|
||||
:
|
||||
<collectingRegion country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
|
||||
<collectingRegion id="C44B7A14388E05C5C8BB3E3D99364664" country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
|
||||
/
|
||||
<locationDeviation location="Rincon">
|
||||
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.0" unit="km" value="3.0">3 km</quantity>
|
||||
<locationDeviation id="36618C78A508595CC632AC1D065B9128" location="Rincon">
|
||||
<quantity id="49866433EDC99186C141EB20C4016E49" metricMagnitude="3" metricUnit="m" metricValue="3.0" unit="km" value="3.0">3 km</quantity>
|
||||
SW
|
||||
<normalizedToken originalValue="Rincón">Rincon</normalizedToken>
|
||||
<normalizedToken id="F0A595C370A874E30B3B6A41EA806626" originalValue="Rincón">Rincon</normalizedToken>
|
||||
</locationDeviation>
|
||||
/
|
||||
<geoCoordinate degrees="8.683" direction="north" orientation="latitude" precision="55" value="8.683">8.683°N</geoCoordinate>
|
||||
<geoCoordinate id="D9CCE097354F7BA26A4AE9A41C145158" degrees="8.683" direction="north" orientation="latitude" precision="55" value="8.683">8.683°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="83.483" direction="west" orientation="longitude" precision="55" value="-83.483">83.483°W</geoCoordinate>
|
||||
<collectingDate value="1991-07">
|
||||
<geoCoordinate id="EB5686EE61E91E63B26318F4E6F6A34D" degrees="83.483" direction="west" orientation="longitude" precision="55" value="-83.483">83.483°W</geoCoordinate>
|
||||
<collectingDate id="4D72249199A4A007838A5C73F96BC40A" value="1991-07">
|
||||
July
|
||||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.9910999999999999" unit="m" value="1991.1">1991. 10 m</quantity>
|
||||
<quantity id="5EE96B65334D2FCC6EF0186CBC3CF2C0" metricMagnitude="3" metricUnit="m" metricValue="1.9910999999999999" unit="m" value="1991.1">1991. 10 m</quantity>
|
||||
</collectingDate>
|
||||
/
|
||||
<collectorName>P. Hanson. Malaise</collectorName>
|
||||
<collectorName id="7AE4BFF189EB2AEE9AB7ADCFA00719CB">P. Hanson. Malaise</collectorName>
|
||||
" "From the Michael/ Ivie Collection" (1) | MTEC
|
||||
</materialsCitation>
|
||||
.
|
||||
|
|
|
|||
Loading…
Add table
Add a link
Reference in a new issue