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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.131.1921" ID-GBIF-Dataset="72a81528-63ef-4531-9f9c-ade551a651d8" ID-PMC="PMC3208436" ID-Pensoft-Pub="1313-2970-131-1" ID-PubMed="22140329" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-131-1" ModsDocOrigin="ZooKeys 131" ModsDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249839150" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2011" docId="4DE66A2B23AF0522AAFD7FA046EBB29A" docLanguage="en" docName="ZooKeys 131: 1-161" docOrigin="ZooKeys 131" docSource="http://dx.doi.org/10.3897/zookeys.131.1921" docTitle="Orthomorpha tuberculifera Likhitrakarn, Golovatch &amp; Panha, 2011, sp. n., variety from Khao Wong Phrachan Temple" docType="treatment" docVersion="6" lastPageNumber="24" masterDocId="FFFF0F54FF88BD1C371C965D22453928" masterDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="23" updateTime="1668152297771" updateUser="ExternalLinkService">
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<mods:title>Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2011</mods:date>
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<mods:number>131</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.131.1921</mods:url>
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<treatment ID-GBIF-Taxon="152031134" LSID="urn:lsid:zoobank.org:act:323ACC16-597D-4A54-AADD-903141FD4E54" httpUri="http://treatment.plazi.org/id/4DE66A2B23AF0522AAFD7FA046EBB29A" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<subSubSection pageId="22" pageNumber="23" type="nomenclature">
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculifera" order="Polydesmida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tuberculifera">Orthomorpha tuberculifera</taxonomicName>
<taxonomicNameLabel pageId="22" pageNumber="23">sp. n., variety from Khao Wong Phrachan Temple</taxonomicNameLabel>
Figs 1618
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<paragraph pageId="22" pageNumber="23">Description.</paragraph>
<paragraph pageId="22" pageNumber="23">
Length 22-30 mm (♂), 24-31 mm (♀), width of midbody pro- and metazona 2.6-2.9 and 3.9-4.2 mm (♂), 2.8-3.3 and 4.0-4.3 mm (♀), respectively. Colour pattern same as in the type series of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculifera" order="Polydesmida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tuberculifera">Orthomorpha tuberculifera</taxonomicName>
sp. n., but coloration of alcohol material darker, blackish.
</paragraph>
<paragraph pageId="22" pageNumber="23">
All other characters as in the typical
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculifera" order="Polydesmida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tuberculifera">Orthomorpha tuberculifera</taxonomicName>
sp. n. (Figs 16-18), except as follows.
</paragraph>
<paragraph pageId="23" pageNumber="24">
<pageBreakToken pageId="23" pageNumber="24" start="start">Venter</pageBreakToken>
and legs sometimes as pallid as paraterga and epiproct. Paraterga slightly less prominent, narrower (Fig. 16A, C &amp; F). Pattern of setigerous tubercles in caudal row on metatergum 17 sometimes as 4+3, on metatergum 18 as 4+4. Paraterga 2 sometimes with four small indentations on one side, with three ones on the other. Surface below paraterga microgranulate and faintly rugulose.
</paragraph>
<caption pageId="23" pageNumber="24">
<paragraph pageId="23" pageNumber="24">
Figure 16.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculifera" order="Polydesmida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="tuberculifera">Orthomorpha tuberculifera</taxonomicName>
sp. n., ♂, variety from Khao Wong Phrachan Temple. A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, dorsal and ventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
</paragraph>
</caption>
<caption pageId="23" pageNumber="24">
<paragraph pageId="23" pageNumber="24">
Figure 17.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculifera" order="Polydesmida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="tuberculifera">Orthomorpha tuberculifera</taxonomicName>
sp. n., ♂, variety from Khao Wong Phrachan Temple. A, B right gonopod, mesal and lateral views, respectively C-F distal part of right gonopod, mesal, lateral, subcaudal and suboral views, respectively. Scale bar: 0.2 mm.
</paragraph>
</caption>
<caption pageId="23" pageNumber="24">
<paragraph pageId="23" pageNumber="24">
Figure 18.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculifera" order="Polydesmida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="tuberculifera">Orthomorpha tuberculifera</taxonomicName>
sp. n., ♂, variety from Khao Wong Phrachan Temple. A, B right gonopod, lateral and mesal views, respectively. Scale bar: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<subSubSection id="003E4B49BE5C667B6489617AAC8D594F" type="nomenclature">
<paragraph id="67CFB1D5E48CC24D4C6D170A661C4813" pageNumber="257">
<taxonomicName id="629D27A01A1C92D4D9BF751A600EDF7F" ID-CoL="76D8" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" kingdom="Animalia" order="Sarcoptiformes" pageNumber="257" phylum="Arthropoda" rank="genus">Quadroppia Jacot</taxonomicName>
, 1939
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<paragraph id="0EFA532B94D33D892F99EF7043682B52" pageNumber="257">
Typ:
<taxonomicName id="C4A4733623ED0A612482C226A83F11EF" class="Insecta" family="Crambidae" genus="Notaspis" kingdom="Animalia" order="Lepidoptera" pageNumber="257" phylum="Arthropoda" rank="species" species="quadricarinata">Notaspis quadricarinata Michael</taxonomicName>
, 1885. - Syn.
<taxonomicName id="D6A129FF98ECFB7FAC8600A93059A276" authority="Ohkubo, 1995" authorityName="Ohkubo" authorityYear="1995" class="Arachnida" family="Quadroppiidae" genus="Coronoquadroppia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oribatida" pageNumber="257" phylum="Arthropoda" rank="genus">Coronoquadroppia</taxonomicName>
Ohkubo, 1995.
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<subSubSection id="B2BCBF44354E34F82371A20CD1476175" type="discussion">
<paragraph id="33CD3623424321B0EC4B87AF12ED612D" pageNumber="257">
Die rezente Artenaufgliederung und Synonymisierungen sind teils noch in der Diskussion. Es werden auch Arten aus
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in den
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aufgenommen, die bisher nicht im Untersuchungsgebiet nachgewiesen wurden, da mit neuen Nachweisen zu rechnen ist. Auch wurde erst durch Luxton (1987) die Morphologie der Typusart
<taxonomicName id="032E4860D80DB64E976C1B3A503F3E52" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" kingdom="Animalia" order="Sarcoptiformes" pageNumber="257" phylum="Arthropoda" rank="species" species="quadricarinata">Q. quadricarinata</taxonomicName>
(Michael, 1885) durch Typenstudium
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. Damit sind bisherige
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Angaben zu der Art
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. Die etwas erhabene Ring- oder Hufeisenstruktur, wie bei
<taxonomicName id="F2F40656327EB5FB1E8198F9899A24C1" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" kingdom="Animalia" order="Sarcoptiformes" pageNumber="257" phylum="Arthropoda" rank="species" species="monstruosa">Q. monstruosa</taxonomicName>
, hat Ohkubo (1995)
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, eine neue Gattung
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abzuspalten. Das Merkmal ist jedoch m. E. nicht hochwertig genug und ist zu variabel bei mehreren Arten, um eine Gattung sicher zu
<normalizedToken id="97B5231BC365BDC46A0CCDE5F478033A" originalValue="begründen" pageNumber="257">begruenden</normalizedToken>
.
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<paragraph id="67642D50B9886B0C8AA2B27D9853DD96" pageNumber="257">
1. Auf Rostrum-Mitte vor der Translamelle mit Ring- oder Hufeisenstruktur [wie 138g], seitlich daneben und neben den Lamellen
<normalizedToken id="EB01E490C7ACD7B00C8C3F8A850B8DD6" originalValue="zusätzliche" pageNumber="257">zusaetzliche</normalizedToken>
Kiele, die meist winkelig zur Seite ziehen ......................................................................................2
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<paragraph id="E0B944F3A2D0649F0B72E77855036C7B" pageNumber="257">
- Auf Rostrum-Mitte vor der Translamelle ohne Ringstruktur, seitlich neben den Lamellen
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Kiele, die meist
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nach vorn ziehen ......................................4
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2. (1) Auf Rostrum-Mitte mit
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Ringstruktur [138g]. (+) Tarsus II dorsal mit 2 Solenidien...........................................................3
</paragraph>
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<pageStartToken id="30E37E210F2265723D17632EBAC38323" pageNumber="258">-</pageStartToken>
Auf Rostrum-Mitte mit Hufeisen-Struktur, die vorn gebogen und hinten vor der Translamelle gerade ist. (+) Tarsus II dorsal mit 1 Solenidium; Lamellarborsten kurz ..................................................................... 7
</paragraph>
<paragraph id="4E835200DB859799C6C394E96FAA7CB5" pageNumber="258">
3. (2) Vor Genitaldeckeln zwischen den Epimeren 3 und 4 mit einem breiten Sternalfeld, dieses ist seitlich parallel und vorn gerundet [138i]; Sensillus nur schwach bedornt. (+) Lamellenspitzen vorn nach innen
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; Translamelle nur schwach angedeutet; Notogasterborsten kurz;
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170-200 µm. [138g-i] ............................... ....................
<taxonomicName id="E2C8EF32A03D14711042C683C7FB44A0" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" kingdom="Animalia" order="Sarcoptiformes" pageNumber="258" phylum="Arthropoda" rank="species" species="monstruosa">Quadroppia monstruosa Hammer</taxonomicName>
, 1979 (sensu Minguez et al. 1985)
</paragraph>
<paragraph id="EAA1266130BF9D5A08A7DD003F9D4410" pageNumber="258">
- Sternalfeld zwischen den Epimeren 3 und 4 nach vorn verengt, spitz zulaufend, gelegentlich mit kleinem, abgetrennten Teilfeld; Sensillus dicht bedornt. (+) Lamellenspitzen vorn nach innen
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; Translamelle nur schwach angedeutet; Notogasterborsten
<normalizedToken id="9365D1DD079554EBB3B7C5EC161B2DAC" originalValue="mäßig" pageNumber="258">maessig</normalizedToken>
lang;
<normalizedToken id="71379A318EAEDD29AAF605A39D0B1CDC" originalValue="Körperlänge" pageNumber="258">Koerperlaenge</normalizedToken>
175-205 µm...................................................................
<taxonomicName id="1F61D461F6D55921B8CEFC7D7E69C70F" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" kingdom="Animalia" order="Sarcoptiformes" pageNumber="258" phylum="Arthropoda" rank="species" species="michaeli">Quadroppia michaeli Mahunka</taxonomicName>
, 1977
</paragraph>
<paragraph id="B6A7D2410882CAE84661A95C82983805" pageNumber="258">
4. (1) Notogasterborsten lang (
<normalizedToken id="7EE5C182CB185881B81DD662C7F17825" originalValue="über" pageNumber="258">ueber</normalizedToken>
30 µm). (+) Sternalfeld zwischen den Epimeren 3 und 4 breit seitlich parallel und vorn + gerundet; Sensillus lang gestielt, Kopf dicht bedornt. Lamellenspitzen vorn breit; Translamelle schmal;
<normalizedToken id="E7C50C0DAB12E33703B33A1ADFC9E743" originalValue="Körperlänge" pageNumber="258">Koerperlaenge</normalizedToken>
185-230 µm ..........................................................
<taxonomicName id="6935C936CF2D6907F0416FED84F60FF8" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" kingdom="Animalia" order="Sarcoptiformes" pageNumber="258" phylum="Arthropoda" rank="species" species="longisetosa">Quadroppia longisetosa Minguez</taxonomicName>
et al., 1985
</paragraph>
<paragraph id="A0891CBF417EDD29F5B344F630BE4088" pageNumber="258">- Notogasterborsten kurz (unter 20 µm) ..................................................................5</paragraph>
<paragraph id="3F52A0B2D18364A0F05116261340766B" pageNumber="258">
5. (4) zentrale Notogasterborsten sehr kurz; breite Lamellenspitzen gegen Translamelle deutlich abgegrenzt, Translamelle undeutlich (als Chitinverdickung unter der
<normalizedToken id="A0EE370CD42023E51D883CB4344F6958" originalValue="Oberfläche" pageNumber="258">Oberflaeche</normalizedToken>
?). (+) Sternalfeld zwischen den Epimeren 3 und 4 breit seitlich parallel und vorn gerundet[138f: Pfeil]; Sensilluskopf dicht bedornt, Lamellarborsten kurz;
<normalizedToken id="1BC5E04026970AA516E4C5EFC1C2C2F3" originalValue="Körperlänge" pageNumber="258">Koerperlaenge</normalizedToken>
155-180 µm. [138d-f] .....................................................................
<taxonomicName id="633E738A196F4A95C03FF357D036D007" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" kingdom="Animalia" order="Sarcoptiformes" pageNumber="258" phylum="Arthropoda" rank="species" species="hammerae">Quadroppia hammerae Minguez</taxonomicName>
et al., 1985
</paragraph>
<caption id="9F578E0FA3EC056A4955E8C0523A89F2" pageNumber="258">
<paragraph id="8F68561B285666555E20401613473853" pageNumber="258">
Abb. 138: a)
<taxonomicName id="8B31CE47B28EFA4FF4BD32749E1CA551" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" kingdom="Animalia" order="Sarcoptiformes" pageNumber="258" phylum="Arthropoda" rank="species" species="quadricarinata">Quadroppia quadricarinata</taxonomicName>
: dorsal; b) Sensillus; c) ventral, Epimeren-Genital-Region. - d)
<taxonomicName id="3AF858074D0D27913A44EE458E5656BB" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" genus.outerRound="1" kingdom="Animalia" order="Sarcoptiformes" pageNumber="258" phylum="Arthropoda" rank="species" species="hammerae">Q. hammerae</taxonomicName>
: dorsal; e) Sensillus; f) ventral, Epimeren-Genital-Region. -g)
<taxonomicName id="F6FBCF701C58310C2417F60308B7AA77" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" genus.outerRound="1" kingdom="Animalia" order="Sarcoptiformes" pageNumber="258" phylum="Arthropoda" rank="species" species="monstruosa">Q. monstruosa</taxonomicName>
: dorsal; h) Sensillus; i) ventral, Epimeren-Genital-Region. (der Pfeil weist auf das Sternalfeld)
</paragraph>
</caption>
<paragraph id="270A5FD05D23262DA7099F4D59E9C87F" pageNumber="259">
<pageStartToken id="7550CCBE2E7B9258033061FDA36664C6" pageNumber="259">-</pageStartToken>
zentrale Notogasterborsten nicht sehr kurz; breite Lamellenspitzen gegen schmale Translamelle meist nicht abgegrenzt.........................................................................6
</paragraph>
<paragraph id="A9EC4D13F04BA052BFB12EE8E3D81114" pageNumber="259">
6. (5) Sternalfeld zwischen den Epimeren 3 und 4 nach vorn wie ein Flaschenhals verengt, distal gerundet [138c: Pfeil]; Tarsus II dorsal mit 1 Solenidium. (+) Sensillus sehr schwach bedornt; Lamellarborsten relativ lang, wie etwa halber Abstand zueinander(sie stehen nicht auf den &quot;Cuspides&quot; der Lamellen, sondern etwas davor auf dem Prodorsum; Lamellenspitzen und Translamelle etwas variabel); Notogasterborsten
<normalizedToken id="D4354CB196DE3DCD76C133F01EF4FD75" originalValue="mäßig" pageNumber="259">maessig</normalizedToken>
lang;
<normalizedToken id="96BA7220184F6BFFE6E10A6B69C0F440" originalValue="Körperlänge" pageNumber="259">Koerperlaenge</normalizedToken>
190-230 µm. [138a-c] ................................................................
<taxonomicName id="AAF084FF58EE2A4EFE408CDEE0569EAB" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" kingdom="Animalia" order="Sarcoptiformes" pageNumber="259" phylum="Arthropoda" rank="species" species="quadricarinata">Quadroppia quadricarinata</taxonomicName>
(Michael, 1885)
</paragraph>
<paragraph id="F19812473B99A817D562D109CA195EF7" pageNumber="259">
- Sternalfeld zwischen den Epimeren 3 und 4 breit, seitlich parallel und vorn gerundet (
<normalizedToken id="A4D7EB66BB09AA7FCDEB035171564D33" originalValue="ähnlich" pageNumber="259">aehnlich</normalizedToken>
<taxonomicName id="80FFDD2E3115535E4AAA206D7365238E" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" kingdom="Animalia" order="Sarcoptiformes" pageNumber="259" phylum="Arthropoda" rank="species" species="hammerae">Q. hammerae</taxonomicName>
); Tarsus II dorsal mit 2 Solenidien. (+) Lamellarborsten kurz;
<normalizedToken id="F9163B8F30E6B63FB5D1CA7A8E37273E" originalValue="Körperlänge" pageNumber="259">Koerperlaenge</normalizedToken>
170-210 µm........................................................
<taxonomicName id="CC031F329921E86FCA64B65330585AFD" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" kingdom="Animalia" order="Sarcoptiformes" pageNumber="259" phylum="Arthropoda" rank="species" species="maritalis">Quadroppia maritalis Lions</taxonomicName>
, 1982
</paragraph>
<paragraph id="8F7B71E0C303792555DE6E4950678819" pageNumber="259">
7. (2) Sternalfeld zwischen den Epimeren 3 und 4 breit, seitlich parallel und vorn gerade; seitlicher Kiel des Schulterknotens relativ lang und gebogen; Sensillus dicht bedornt;
<normalizedToken id="059C26BDF3747EB9E70067255102EB15" originalValue="Körperlänge" pageNumber="259">Koerperlaenge</normalizedToken>
170-195 µm...............................................................
<taxonomicName id="523E3B5C730BA5F51EE3A487F62490E5" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" kingdom="Animalia" order="Sarcoptiformes" pageNumber="259" phylum="Arthropoda" rank="species" species="pseudocircumita">Quadroppia pseudocircumita Minguez</taxonomicName>
et al., 1985
</paragraph>
<paragraph id="64F75EA0F208F07699346991B0FAB1B8" pageNumber="259">
- Sternalfeld zwischen den Epimeren in der Mitte stark verengt, dann ganz vorn wieder verbreitert; seitlicher Kiel des Schulterknotens sehr kurz und gebogen; Sensillus nackt;
<normalizedToken id="2CD05A791D420F1FA0A3B2EFC7279656" originalValue="Körperlänge" pageNumber="259">Koerperlaenge</normalizedToken>
170-195 µm.........................................................
<taxonomicName id="6AB79D6D890785A8248E77271A85D516" class="Arachnida" family="Quadroppiidae" genus="Quadroppia" kingdom="Animalia" order="Sarcoptiformes" pageNumber="259" phylum="Arthropoda" rank="species" species="galaica">Quadroppia galaica Minguez</taxonomicName>
et al., 1985
</paragraph>
</subSubSection>
</treatment>
</document>