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<mods:title>Twenty-eight new species of the spider genus Merizocera Fage, 1912 (Araneae, Psilodercidae) from South and Southeast Asia</mods:title>
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<mods:title id="98A0233F96EF320D2CE34A1B5749E1AD">Twenty-eight new species of the spider genus Merizocera Fage, 1912 (Araneae, Psilodercidae) from South and Southeast Asia</mods:title>
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<mods:namePart>Chang, Wan-Jin</mods:namePart>
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<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China & Southeast Asia Biological Diversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar & University of Chinese Academy of Sciences, Beijing 100049, China</mods:affiliation>
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<mods:namePart>Yao, Zhiyuan</mods:namePart>
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<mods:namePart>Li, Shuqiang</mods:namePart>
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<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China & University of Chinese Academy of Sciences, Beijing 100049, China</mods:affiliation>
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<treatment ID-GBIF-Taxon="167284454" LSID="urn:lsid:zoobank.org:act:C706A307-0868-44DA-AFAB-57E83D0924FC" httpUri="http://treatment.plazi.org/id/C0C443C9F7325641ACBEC92557284972" lastPageNumber="41" pageId="0" pageNumber="41">
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<taxonomicName LSID="C0C443C9-F732-5641-ACBE-C92557284972" authority="Li" authorityName="Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Merizocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merizocera salawa" order="Araneae" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="salawa" status="sp. nov.">Merizocera salawa Li</taxonomicName>
|
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<taxonomicNameLabel pageId="0" pageNumber="41">sp. nov.</taxonomicNameLabel>
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||||
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Merizocera salawa sp. nov., holotype male. A Bulb, dorsal view B bulb, dorsal view, embolus and conductor distorted C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, CP = cymbial protrusion, EM = embolus." figureDoi="10.3897/zookeys.961.53058.figure35" httpUri="https://binary.pensoft.net/fig/443261" pageId="0" pageNumber="41">Figures 35</figureCitation>
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<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Merizocera salawa sp. nov., holotype male. A Habitus, dorsal view B habitus, ventral view." figureDoi="10.3897/zookeys.961.53058.figure36" httpUri="https://binary.pensoft.net/fig/443262" pageId="0" pageNumber="41">, 36</figureCitation>
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||||
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Distribution records of the new species of Merizocera from Sri Lanka. 1 M. colombo sp. nov. 2 M. galle sp. nov. 3 M. kandy sp. nov. 4 M. kurunegala sp. nov. 5 M. peraderiya sp. nov. 6 M. ratnapura sp. nov. 7 M. salawa sp. nov. 8 M. thenna sp. nov. 9 M. uva sp. nov." figureDoi="10.3897/zookeys.961.53058.figure52" httpUri="https://binary.pensoft.net/fig/443279" pageId="0" pageNumber="41">, 52</figureCitation>
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||||
<treatment id="C0C443C9F7325641ACBEC92557284972" ID-GBIF-Taxon="167284454" LSID="urn:lsid:zoobank.org:act:C706A307-0868-44DA-AFAB-57E83D0924FC" httpUri="http://treatment.plazi.org/id/C0C443C9F7325641ACBEC92557284972" lastPageNumber="41" pageId="0" pageNumber="41" scope_family="Psilodercidae" scope_order="Araneae">
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<subSubSection id="076B48A6F606E9DA3A06A496D588DBCA" pageId="0" pageNumber="41" type="nomenclature">
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||||
<paragraph id="E3F49939D982B636A08BD1827289A83F" pageId="0" pageNumber="41">
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||||
<taxonomicName id="B0C64081C832FC4F635F3DBA8A46B312" ID-CoL="98ZN8" LSID="C0C443C9-F732-5641-ACBE-C92557284972" authority="Li" authorityName="Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Merizocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merizocera salawa" order="Araneae" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="salawa" status="sp. nov.">Merizocera salawa Li</taxonomicName>
|
||||
<taxonomicNameLabel id="36AB0629DD2BE8BEC8B98E84E388D630" pageId="0" pageNumber="41">sp. nov.</taxonomicNameLabel>
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||||
<figureCitation id="03AA7482C735A8502F614BCA47DF3AB3" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Merizocera salawa sp. nov., holotype male. A Bulb, dorsal view B bulb, dorsal view, embolus and conductor distorted C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, CP = cymbial protrusion, EM = embolus." figureDoi="10.3897/zookeys.961.53058.figure35" httpUri="https://binary.pensoft.net/fig/443261" pageId="0" pageNumber="41">Figures 35</figureCitation>
|
||||
<figureCitation id="FF10844DF23FA16788649D11BA47D442" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Merizocera salawa sp. nov., holotype male. A Habitus, dorsal view B habitus, ventral view." figureDoi="10.3897/zookeys.961.53058.figure36" httpUri="https://binary.pensoft.net/fig/443262" pageId="0" pageNumber="41">, 36</figureCitation>
|
||||
<figureCitation id="C878A6A30B484762A1EE6A08ACACCDFF" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Distribution records of the new species of Merizocera from Sri Lanka. 1 M. colombo sp. nov. 2 M. galle sp. nov. 3 M. kandy sp. nov. 4 M. kurunegala sp. nov. 5 M. peraderiya sp. nov. 6 M. ratnapura sp. nov. 7 M. salawa sp. nov. 8 M. thenna sp. nov. 9 M. uva sp. nov." figureDoi="10.3897/zookeys.961.53058.figure52" httpUri="https://binary.pensoft.net/fig/443279" pageId="0" pageNumber="41">, 52</figureCitation>
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||||
</paragraph>
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||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="41" type="type material">
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||||
<paragraph pageId="0" pageNumber="41">Type material.</paragraph>
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||||
<paragraph pageId="0" pageNumber="41">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Holotype</emphasis>
|
||||
<subSubSection id="ECB006950371A50CD24B0EFA5650E5EA" pageId="0" pageNumber="41" type="type material">
|
||||
<paragraph id="F778CD65C3C919FA2EEFE4DCCB4D5D4B" pageId="0" pageNumber="41">Type material.</paragraph>
|
||||
<paragraph id="D8BC9C6B64E4CC58FEDCAC7429E6ADE2" pageId="0" pageNumber="41">
|
||||
<emphasis id="71A0E8C58D5569BCFECEF0425FE25F3E" bold="true" italics="true" pageId="0" pageNumber="41">Holotype</emphasis>
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: male (IZCAS), Lenawara Lena (= cave) (
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<geoCoordinate degrees="6" direction="north" minutes="56.77" orientation="latitude" precision="9" value="6.9461665">6°56.77'N</geoCoordinate>
|
||||
<geoCoordinate id="A7E72060C017C55E6D865E1AB8D2BD4E" degrees="6" direction="north" minutes="56.77" orientation="latitude" precision="9" value="6.9461665">6°56.77'N</geoCoordinate>
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||||
,
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<geoCoordinate degrees="80" direction="east" minutes="6.76" orientation="longitude" precision="9" value="80.11266">80°6.76'E</geoCoordinate>
|
||||
<geoCoordinate id="BF38CCC5999F1FE5DEC698195175CA5F" degrees="80" direction="east" minutes="6.76" orientation="longitude" precision="9" value="80.11266">80°6.76'E</geoCoordinate>
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, elevation 66 m), Salawa Village, Avissawella Town, Maniyangama, Colombo District,
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<emphasis bold="true" pageId="0" pageNumber="41">Western Province</emphasis>
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||||
<emphasis id="25E0F3F5214A9C5E4F8640DF3F80F988" bold="true" pageId="0" pageNumber="41">Western Province</emphasis>
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||||
,
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<emphasis bold="true" pageId="0" pageNumber="41">Sri Lanka</emphasis>
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<emphasis id="B8B4D639B567F815638077C86D4DDF82" bold="true" pageId="0" pageNumber="41">Sri Lanka</emphasis>
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, 25 September 2014, S. Kosala leg.
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</paragraph>
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||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="41" type="etymology">
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||||
<paragraph pageId="0" pageNumber="41">Etymology.</paragraph>
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||||
<paragraph pageId="0" pageNumber="41">The specific name refers to the type locality; noun in apposition.</paragraph>
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||||
<subSubSection id="0D1A7799482067305658C411BD405D2F" pageId="0" pageNumber="41" type="etymology">
|
||||
<paragraph id="2093CBD867B863024C1DBFCF7EE353C4" pageId="0" pageNumber="41">Etymology.</paragraph>
|
||||
<paragraph id="BD287CCDBD0FF1E98D8D11F18C05812E" pageId="0" pageNumber="41">The specific name refers to the type locality; noun in apposition.</paragraph>
|
||||
</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="41" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="41">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="41">
|
||||
<subSubSection id="9C062DE03DC2B682DE1552A7AB36A29A" pageId="0" pageNumber="41" type="diagnosis">
|
||||
<paragraph id="7F672E7C5C0AD8179A42DC5EAD516B00" pageId="0" pageNumber="41">Diagnosis.</paragraph>
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||||
<paragraph id="3EA4AC00A1F397854387C989BABAE965" pageId="0" pageNumber="41">
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Diagnosis features of males are discussed in
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||||
<taxonomicName lsidName="M. mandai" pageId="0" pageNumber="41" rank="species" species="mandai">
|
||||
<emphasis italics="true" pageId="0" pageNumber="41">M. mandai</emphasis>
|
||||
<taxonomicName id="E4FAC1B46D89BE5F5B03058C9FFE73C0" lsidName="M. mandai" pageId="0" pageNumber="41" rank="species" species="mandai">
|
||||
<emphasis id="955A2D1F85C1DF0118806CB16946E9E0" italics="true" pageId="0" pageNumber="41">M. mandai</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. Bulb with a widely separated bifurcate conductor, and a curved embolus (Fig.
|
||||
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Merizocera salawa sp. nov., holotype male. A Bulb, dorsal view B bulb, dorsal view, embolus and conductor distorted C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, CP = cymbial protrusion, EM = embolus." figureDoi="10.3897/zookeys.961.53058.figure35" httpUri="https://binary.pensoft.net/fig/443261" pageId="0" pageNumber="41">35B</figureCitation>
|
||||
<figureCitation id="009A867E62EF14E809B8D248AB2C1209" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Merizocera salawa sp. nov., holotype male. A Bulb, dorsal view B bulb, dorsal view, embolus and conductor distorted C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, CP = cymbial protrusion, EM = embolus." figureDoi="10.3897/zookeys.961.53058.figure35" httpUri="https://binary.pensoft.net/fig/443261" pageId="0" pageNumber="41">35B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.961.53058.figure35" httpUri="https://binary.pensoft.net/fig/443261" pageId="0" pageNumber="41" start="Figure 35" startId="F35">
|
||||
<paragraph pageId="0" pageNumber="41">
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">Figure 35.</emphasis>
|
||||
<taxonomicName authorityName="Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Merizocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merizocera salawa" order="Araneae" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="salawa">
|
||||
<emphasis italics="true" pageId="0" pageNumber="41">Merizocera salawa</emphasis>
|
||||
<caption id="C962E5FADCBE8B50CCD4556B8A9C2E75" doi="10.3897/zookeys.961.53058.figure35" httpUri="https://binary.pensoft.net/fig/443261" pageId="0" pageNumber="41" start="Figure 35" startId="F35">
|
||||
<paragraph id="C98E07216B0D37369CC943BC6790B327" pageId="0" pageNumber="41">
|
||||
<emphasis id="A381F3C1099DC690CBF8FBC6A77B9681" bold="true" pageId="0" pageNumber="41">Figure 35.</emphasis>
|
||||
<taxonomicName id="76D2DC641F1DCCABAC91700E39D40081" authorityName="Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Merizocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merizocera salawa" order="Araneae" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="salawa">
|
||||
<emphasis id="B0AAB1FB293B6EC8FAB47BBCD1E65CB8" italics="true" pageId="0" pageNumber="41">Merizocera salawa</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., holotype male.
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
|
||||
<emphasis id="042EB140DE6D195DA48342A93D1E69EE" bold="true" pageId="0" pageNumber="41">A</emphasis>
|
||||
Bulb, dorsal view
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
|
||||
<emphasis id="047302DE295B616843CD22E7121FF28D" bold="true" pageId="0" pageNumber="41">B</emphasis>
|
||||
bulb, dorsal view, embolus and conductor distorted
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
|
||||
<emphasis id="8F9F486F75CA9EF211A993B5694D3A65" bold="true" pageId="0" pageNumber="41">C</emphasis>
|
||||
palp, prolateral view
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">D</emphasis>
|
||||
<emphasis id="559F6F5DC0F22CFB125A76B382792728" bold="true" pageId="0" pageNumber="41">D</emphasis>
|
||||
palp, retrolateral view. Abbreviations: CO = conductor, CP = cymbial protrusion, EM = embolus.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="41" type="description">
|
||||
<paragraph pageId="0" pageNumber="41">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="41">
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">Male</emphasis>
|
||||
<subSubSection id="1CED24AB6E85D79ED81B6EFBE02ABBE4" pageId="0" pageNumber="41" type="description">
|
||||
<paragraph id="5138CE75658E7F02AEDF3532F7A85B9E" pageId="0" pageNumber="41">Description.</paragraph>
|
||||
<paragraph id="C40DDDDEC9F7AC60CDAFD228418CBD2A" pageId="0" pageNumber="41">
|
||||
<emphasis id="473C331DB5573D43AF853956645A8D09" bold="true" pageId="0" pageNumber="41">Male</emphasis>
|
||||
(holotype). Total length 1.52; carapace 0.61 long, 0.54 wide; abdomen 0.88 long, 0.41 wide. Carapace circular, brownish, with dark brown marks laterally and dark brown median stripe on anterior half (Fig.
|
||||
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Merizocera salawa sp. nov., holotype male. A Habitus, dorsal view B habitus, ventral view." figureDoi="10.3897/zookeys.961.53058.figure36" httpUri="https://binary.pensoft.net/fig/443262" pageId="0" pageNumber="41">36C</figureCitation>
|
||||
<figureCitation id="3F10266651CBE4654FC5E9AAF73CC95A" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Merizocera salawa sp. nov., holotype male. A Habitus, dorsal view B habitus, ventral view." figureDoi="10.3897/zookeys.961.53058.figure36" httpUri="https://binary.pensoft.net/fig/443262" pageId="0" pageNumber="41">36C</figureCitation>
|
||||
). Fovea shallow. Thoracic region distinctly elevated medially. Clypeus brownish, with dark brown marks medially. Labium dark brown. Sternum dark brown but brownish medially. Abdomen elongated, dark grey, with dark brown marks dorsally and ventrally. Legs missing. Palp (Fig.
|
||||
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Merizocera salawa sp. nov., holotype male. A Bulb, dorsal view B bulb, dorsal view, embolus and conductor distorted C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, CP = cymbial protrusion, EM = embolus." figureDoi="10.3897/zookeys.961.53058.figure35" httpUri="https://binary.pensoft.net/fig/443261" pageId="0" pageNumber="41">35A-D</figureCitation>
|
||||
<figureCitation id="488789E681F102E6A8B9D05D66F89B24" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Merizocera salawa sp. nov., holotype male. A Bulb, dorsal view B bulb, dorsal view, embolus and conductor distorted C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, CP = cymbial protrusion, EM = embolus." figureDoi="10.3897/zookeys.961.53058.figure35" httpUri="https://binary.pensoft.net/fig/443261" pageId="0" pageNumber="41">35A-D</figureCitation>
|
||||
): femur slender, four times longer than patella; patella not swollen; tibia not swollen, 1/2 femur length; cymbium with distal protrusion, 1/2 femur length, length ratio of dorsal elongation and cymbium: 0.63; bulb pale yellow, pyriform with embolus and conductor emerging distally; embolus arising laterally, bent, with consistent width; conductor bifurcate, arising laterally, similar width as embolus, resembles a widely open crescent-shape.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.961.53058.figure36" httpUri="https://binary.pensoft.net/fig/443262" pageId="0" pageNumber="41" start="Figure 36" startId="F36">
|
||||
<paragraph pageId="0" pageNumber="41">
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">Figure 36.</emphasis>
|
||||
<taxonomicName authorityName="Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Merizocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merizocera salawa" order="Araneae" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="salawa">
|
||||
<emphasis italics="true" pageId="0" pageNumber="41">Merizocera salawa</emphasis>
|
||||
<caption id="B3BEA304C118963424A0699A99BE1A8E" doi="10.3897/zookeys.961.53058.figure36" httpUri="https://binary.pensoft.net/fig/443262" pageId="0" pageNumber="41" start="Figure 36" startId="F36">
|
||||
<paragraph id="9616A215D15F47AD7CAEA6CDDBA5CF85" pageId="0" pageNumber="41">
|
||||
<emphasis id="F96BBA83A3E4F11FD97B546F3D97FCDF" bold="true" pageId="0" pageNumber="41">Figure 36.</emphasis>
|
||||
<taxonomicName id="9AC86DB350499519452DF7398A04DEB6" authorityName="Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Merizocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merizocera salawa" order="Araneae" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="salawa">
|
||||
<emphasis id="10E2102926898844AF7D034D5C5A7C3C" italics="true" pageId="0" pageNumber="41">Merizocera salawa</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., holotype male.
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
|
||||
<emphasis id="A3701AC22D740EF852225C5F7F522132" bold="true" pageId="0" pageNumber="41">A</emphasis>
|
||||
Habitus, dorsal view
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
|
||||
<emphasis id="3F1D5AC53CD65BD79FD54AB939517F1D" bold="true" pageId="0" pageNumber="41">B</emphasis>
|
||||
habitus, ventral view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph pageId="0" pageNumber="41">
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">Female.</emphasis>
|
||||
<paragraph id="508609704B274873F4C751A12A038426" pageId="0" pageNumber="41">
|
||||
<emphasis id="57DDEF46D555E3DC2AB1EE5B8CB1BA0B" bold="true" pageId="0" pageNumber="41">Female.</emphasis>
|
||||
Unknown.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="41" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="41">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="41">
|
||||
<subSubSection id="4C79C51F6C3C5F71F714085EEF0C7C11" pageId="0" pageNumber="41" type="distribution">
|
||||
<paragraph id="16B69961A08D3D2F90590E919923ABFF" pageId="0" pageNumber="41">Distribution.</paragraph>
|
||||
<paragraph id="71BE8AE8791F90DDCA9451F82C983BB2" pageId="0" pageNumber="41">
|
||||
Known only from the type locality (Sri Lanka; Fig.
|
||||
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Distribution records of the new species of Merizocera from Sri Lanka. 1 M. colombo sp. nov. 2 M. galle sp. nov. 3 M. kandy sp. nov. 4 M. kurunegala sp. nov. 5 M. peraderiya sp. nov. 6 M. ratnapura sp. nov. 7 M. salawa sp. nov. 8 M. thenna sp. nov. 9 M. uva sp. nov." figureDoi="10.3897/zookeys.961.53058.figure52" httpUri="https://binary.pensoft.net/fig/443279" pageId="0" pageNumber="41">52</figureCitation>
|
||||
<figureCitation id="F72D407D1A63077C4C0B58587D06CB70" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Distribution records of the new species of Merizocera from Sri Lanka. 1 M. colombo sp. nov. 2 M. galle sp. nov. 3 M. kandy sp. nov. 4 M. kurunegala sp. nov. 5 M. peraderiya sp. nov. 6 M. ratnapura sp. nov. 7 M. salawa sp. nov. 8 M. thenna sp. nov. 9 M. uva sp. nov." figureDoi="10.3897/zookeys.961.53058.figure52" httpUri="https://binary.pensoft.net/fig/443279" pageId="0" pageNumber="41">52</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
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Add table
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