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<mods:title>Twenty-eight new species of the spider genus Merizocera Fage, 1912 (Araneae, Psilodercidae) from South and Southeast Asia</mods:title>
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<mods:title id="FAB46696F846DF818624290B9938535C">Twenty-eight new species of the spider genus Merizocera Fage, 1912 (Araneae, Psilodercidae) from South and Southeast Asia</mods:title>
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<mods:namePart>Chang, Wan-Jin</mods:namePart>
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<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China & Southeast Asia Biological Diversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar & University of Chinese Academy of Sciences, Beijing 100049, China</mods:affiliation>
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<mods:namePart>Yao, Zhiyuan</mods:namePart>
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<mods:namePart>Li, Shuqiang</mods:namePart>
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<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China & University of Chinese Academy of Sciences, Beijing 100049, China</mods:affiliation>
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<treatment ID-GBIF-Taxon="167284470" LSID="urn:lsid:zoobank.org:act:B1A4B9F0-9C4F-4E0E-B5D8-9EAC98A73FA6" httpUri="http://treatment.plazi.org/id/5E7A813C917E5007A863D25AED5A843D" lastPageNumber="41" pageId="0" pageNumber="41">
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<taxonomicName LSID="5E7A813C-917E-5007-A863-D25AED5A843D" authority="Chang & Yao & Li, 2020" authorityName="Chang & Yao & Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Merizocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merizocera nyingchi" order="Araneae" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="nyingchi" status="sp. nov.">Merizocera nyingchi</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="41">sp. nov.</taxonomicNameLabel>
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||||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merizocera nyingchi sp. nov., holotype female. A Endogyne, dorsal view B female epigastric area, ventral view C female habitus, dorsal view D female habitus, ventral view E female habitus, lateral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.961.53058.figure24" httpUri="https://binary.pensoft.net/fig/443250" pageId="0" pageNumber="41">Figures 24</figureCitation>
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<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Distribution records of the new species of Merizocera from China and Myanmar. 1 M. baoshan sp. nov. 2 M. hponkanrazi sp. nov. 3 M. kachin sp. nov. 4 M. lincang sp. nov. 5 M. mainling sp. nov. 6 M. nyingchi sp. nov. 7 M. putao sp. nov. 8 M. tengchong sp. nov. 9 M. wenshan sp. nov. 10 M. wui sp. nov. 11 M. yuxi sp. nov." figureDoi="10.3897/zookeys.961.53058.figure53" httpUri="https://binary.pensoft.net/fig/443280" pageId="0" pageNumber="41">, 53</figureCitation>
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<treatment id="5E7A813C917E5007A863D25AED5A843D" ID-GBIF-Taxon="167284470" LSID="urn:lsid:zoobank.org:act:B1A4B9F0-9C4F-4E0E-B5D8-9EAC98A73FA6" httpUri="http://treatment.plazi.org/id/5E7A813C917E5007A863D25AED5A843D" lastPageNumber="41" pageId="0" pageNumber="41" scope_family="Psilodercidae" scope_order="Araneae">
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<subSubSection id="982F295A2E6AECA18C0F98466D5857DE" pageId="0" pageNumber="41" type="nomenclature">
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<paragraph id="8895A7F04B273B64791398020C168387" pageId="0" pageNumber="41">
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||||
<taxonomicName id="790063B937F8A73EBADCD13442E65965" ID-CoL="98ZNM" LSID="5E7A813C-917E-5007-A863-D25AED5A843D" authority="Chang & Yao & Li, 2020" authorityName="Chang & Yao & Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Merizocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merizocera nyingchi" order="Araneae" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="nyingchi" status="sp. nov.">Merizocera nyingchi</taxonomicName>
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<taxonomicNameLabel id="80E742A0EA076DFFE88039F3FB46BF7B" pageId="0" pageNumber="41">sp. nov.</taxonomicNameLabel>
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<figureCitation id="C9FD20A821A5AD2BB6CCEF63F8322569" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merizocera nyingchi sp. nov., holotype female. A Endogyne, dorsal view B female epigastric area, ventral view C female habitus, dorsal view D female habitus, ventral view E female habitus, lateral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.961.53058.figure24" httpUri="https://binary.pensoft.net/fig/443250" pageId="0" pageNumber="41">Figures 24</figureCitation>
|
||||
<figureCitation id="FCDF03B5DA23F43C7D86E2956EB28706" captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Distribution records of the new species of Merizocera from China and Myanmar. 1 M. baoshan sp. nov. 2 M. hponkanrazi sp. nov. 3 M. kachin sp. nov. 4 M. lincang sp. nov. 5 M. mainling sp. nov. 6 M. nyingchi sp. nov. 7 M. putao sp. nov. 8 M. tengchong sp. nov. 9 M. wenshan sp. nov. 10 M. wui sp. nov. 11 M. yuxi sp. nov." figureDoi="10.3897/zookeys.961.53058.figure53" httpUri="https://binary.pensoft.net/fig/443280" pageId="0" pageNumber="41">, 53</figureCitation>
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||||
</paragraph>
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||||
</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="41" type="type material">
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||||
<paragraph pageId="0" pageNumber="41">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="41">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Holotype</emphasis>
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||||
<subSubSection id="EE2ED167BCB340182E1432764F67AC50" pageId="0" pageNumber="41" type="type material">
|
||||
<paragraph id="74C207FF8A8494885B8FD697B856B902" pageId="0" pageNumber="41">Type material.</paragraph>
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||||
<paragraph id="5E9EECC3620461ADC23FC8ABA444A24F" pageId="0" pageNumber="41">
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||||
<emphasis id="8E6F6DE6F9BEC043ED9952962CD00889" bold="true" italics="true" pageId="0" pageNumber="41">Holotype</emphasis>
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: female (IZCAS), mountain behind a farmhouse resort (
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<geoCoordinate degrees="29" direction="north" minutes="19.09" orientation="latitude" precision="9" value="29.318167">29°19.09'N</geoCoordinate>
|
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<geoCoordinate id="DF740FDFC6DEFA4A990A35D454C4CAFE" degrees="29" direction="north" minutes="19.09" orientation="latitude" precision="9" value="29.318167">29°19.09'N</geoCoordinate>
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,
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<geoCoordinate degrees="95" direction="east" minutes="18.88" orientation="longitude" precision="9" value="95.31467">95°18.88'E</geoCoordinate>
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||||
<geoCoordinate id="FA7E08F61B401183AF7003797DB90EAD" degrees="95" direction="east" minutes="18.88" orientation="longitude" precision="9" value="95.31467">95°18.88'E</geoCoordinate>
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, elevation 1280 m), Medog County, Nyingchi,
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<emphasis bold="true" pageId="0" pageNumber="41">Tibet</emphasis>
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<emphasis id="CF41DE5279C0175A593DB53409893D21" bold="true" pageId="0" pageNumber="41">Tibet</emphasis>
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,
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<emphasis bold="true" pageId="0" pageNumber="41">China</emphasis>
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<emphasis id="D0D7706206C087480608458A516B532D" bold="true" pageId="0" pageNumber="41">China</emphasis>
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, 4 August 2013, L. Lin leg.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Paratype</emphasis>
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<emphasis id="826580982D56492A1AAF7F1271C47DC0" bold="true" italics="true" pageId="0" pageNumber="41">Paratype</emphasis>
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: 1 female (IZCAS), same data as holotype.
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</paragraph>
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</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="41" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="41">Etymology.</paragraph>
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||||
<paragraph pageId="0" pageNumber="41">The specific name refers to the type locality; noun in apposition.</paragraph>
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||||
<subSubSection id="6B4D2740D513409F9C051942C2D00AA3" pageId="0" pageNumber="41" type="etymology">
|
||||
<paragraph id="93304F2D26F30FD72EC2B500345781C3" pageId="0" pageNumber="41">Etymology.</paragraph>
|
||||
<paragraph id="8E4E005F21680E8B1E567F7B384E107C" pageId="0" pageNumber="41">The specific name refers to the type locality; noun in apposition.</paragraph>
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||||
</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="41" type="diagnosis">
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||||
<paragraph pageId="0" pageNumber="41">Diagnosis.</paragraph>
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||||
<paragraph pageId="0" pageNumber="41">
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||||
<subSubSection id="0E637555D295CA4D7B8546700FFE1176" pageId="0" pageNumber="41" type="diagnosis">
|
||||
<paragraph id="8374FF31D4D2786CBB6796F18B7B709A" pageId="0" pageNumber="41">Diagnosis.</paragraph>
|
||||
<paragraph id="46D820D42FB15AB2C84B13977B316A7C" pageId="0" pageNumber="41">
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Females can be distinguished from congeners by two pairs of twisted stalked spermathecae bearing globose distal ends, where lateral pairs are at least half as short as median pairs and have globose ends twice the size of the former (Fig.
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merizocera nyingchi sp. nov., holotype female. A Endogyne, dorsal view B female epigastric area, ventral view C female habitus, dorsal view D female habitus, ventral view E female habitus, lateral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.961.53058.figure24" httpUri="https://binary.pensoft.net/fig/443250" pageId="0" pageNumber="41">24A</figureCitation>
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||||
<figureCitation id="46D23C89554B2D42B0EC6D6D5B7C6626" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merizocera nyingchi sp. nov., holotype female. A Endogyne, dorsal view B female epigastric area, ventral view C female habitus, dorsal view D female habitus, ventral view E female habitus, lateral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.961.53058.figure24" httpUri="https://binary.pensoft.net/fig/443250" pageId="0" pageNumber="41">24A</figureCitation>
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||||
).
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</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="41" type="description">
|
||||
<paragraph pageId="0" pageNumber="41">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="41">
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">Female</emphasis>
|
||||
<subSubSection id="0EA4B1E1F27185DC8BAEDCDA87FFEA09" pageId="0" pageNumber="41" type="description">
|
||||
<paragraph id="FCF6A2A0C88827E03E2CA4F188B43C19" pageId="0" pageNumber="41">Description.</paragraph>
|
||||
<paragraph id="F8F0D6CA103093BAFB6040F496D2746F" pageId="0" pageNumber="41">
|
||||
<emphasis id="FC806132F3F783B1750B58F64C1AA8E3" bold="true" pageId="0" pageNumber="41">Female</emphasis>
|
||||
(holotype). Total length 1.42; carapace 0.65 long, 0.53 wide; abdomen 0.75 long, 0.53 wide. Carapace circular, brown, with dark brown radiating marks (Fig.
|
||||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merizocera nyingchi sp. nov., holotype female. A Endogyne, dorsal view B female epigastric area, ventral view C female habitus, dorsal view D female habitus, ventral view E female habitus, lateral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.961.53058.figure24" httpUri="https://binary.pensoft.net/fig/443250" pageId="0" pageNumber="41">24C</figureCitation>
|
||||
<figureCitation id="5881B7238ABB9B2DEF4D7FDD8B225F80" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merizocera nyingchi sp. nov., holotype female. A Endogyne, dorsal view B female epigastric area, ventral view C female habitus, dorsal view D female habitus, ventral view E female habitus, lateral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.961.53058.figure24" httpUri="https://binary.pensoft.net/fig/443250" pageId="0" pageNumber="41">24C</figureCitation>
|
||||
). Fovea shallow. Thoracic region distinctly elevated medially. Clypeus, labium, and sternum dark brown. Abdomen slightly elongated, dark brown (Fig.
|
||||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merizocera nyingchi sp. nov., holotype female. A Endogyne, dorsal view B female epigastric area, ventral view C female habitus, dorsal view D female habitus, ventral view E female habitus, lateral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.961.53058.figure24" httpUri="https://binary.pensoft.net/fig/443250" pageId="0" pageNumber="41">24B</figureCitation>
|
||||
<figureCitation id="ADDFE6F5598148EE9F770A3D1399A4C8" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merizocera nyingchi sp. nov., holotype female. A Endogyne, dorsal view B female epigastric area, ventral view C female habitus, dorsal view D female habitus, ventral view E female habitus, lateral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.961.53058.figure24" httpUri="https://binary.pensoft.net/fig/443250" pageId="0" pageNumber="41">24B</figureCitation>
|
||||
). Legs light brown; measurements: I, II, and III missing, IV 4.06 (1.05, 0.19, 1.19, 1.00, 0.63). Epigastric area (Fig.
|
||||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merizocera nyingchi sp. nov., holotype female. A Endogyne, dorsal view B female epigastric area, ventral view C female habitus, dorsal view D female habitus, ventral view E female habitus, lateral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.961.53058.figure24" httpUri="https://binary.pensoft.net/fig/443250" pageId="0" pageNumber="41">24B</figureCitation>
|
||||
<figureCitation id="34E47ABDCCD6BFDB019502C2E1BCFA23" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merizocera nyingchi sp. nov., holotype female. A Endogyne, dorsal view B female epigastric area, ventral view C female habitus, dorsal view D female habitus, ventral view E female habitus, lateral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.961.53058.figure24" httpUri="https://binary.pensoft.net/fig/443250" pageId="0" pageNumber="41">24B</figureCitation>
|
||||
): dark brown patch, medially with pale yellow horizontal strip. Endogyne (Fig.
|
||||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merizocera nyingchi sp. nov., holotype female. A Endogyne, dorsal view B female epigastric area, ventral view C female habitus, dorsal view D female habitus, ventral view E female habitus, lateral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.961.53058.figure24" httpUri="https://binary.pensoft.net/fig/443250" pageId="0" pageNumber="41">24A</figureCitation>
|
||||
<figureCitation id="12345D574A12F54A1A385E2F6C69CFF2" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merizocera nyingchi sp. nov., holotype female. A Endogyne, dorsal view B female epigastric area, ventral view C female habitus, dorsal view D female habitus, ventral view E female habitus, lateral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.961.53058.figure24" httpUri="https://binary.pensoft.net/fig/443250" pageId="0" pageNumber="41">24A</figureCitation>
|
||||
) with two pairs of twisted stalked spermathecae bearing globose distal ends, lateral pairs half as short as median pairs, globose ends of lateral pairs twice larger than median pairs.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.961.53058.figure24" httpUri="https://binary.pensoft.net/fig/443250" pageId="0" pageNumber="41" start="Figure 24" startId="F24">
|
||||
<paragraph pageId="0" pageNumber="41">
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">Figure 24.</emphasis>
|
||||
<taxonomicName authorityName="Chang & Yao & Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Merizocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merizocera nyingchi" order="Araneae" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="nyingchi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="41">Merizocera nyingchi</emphasis>
|
||||
<caption id="0250E3A3E6E0C60FC625BED78476BD31" doi="10.3897/zookeys.961.53058.figure24" httpUri="https://binary.pensoft.net/fig/443250" pageId="0" pageNumber="41" start="Figure 24" startId="F24">
|
||||
<paragraph id="2746C97A7102C6F7EC721614117C6620" pageId="0" pageNumber="41">
|
||||
<emphasis id="4FE620F38B8DA7F566FD82A8DF94B91C" bold="true" pageId="0" pageNumber="41">Figure 24.</emphasis>
|
||||
<taxonomicName id="D52E1CECD0DEFE3C01D9F2F0905899CD" authorityName="Chang & Yao & Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Merizocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merizocera nyingchi" order="Araneae" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="nyingchi">
|
||||
<emphasis id="76E006674A7F08B470C028710BC9B678" italics="true" pageId="0" pageNumber="41">Merizocera nyingchi</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., holotype female.
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
|
||||
<emphasis id="284610E3C3F2E2F416A39A9423715F4A" bold="true" pageId="0" pageNumber="41">A</emphasis>
|
||||
Endogyne, dorsal view
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
|
||||
<emphasis id="345D86FAC1C3E4AE8E0D8C38DB2C7115" bold="true" pageId="0" pageNumber="41">B</emphasis>
|
||||
female epigastric area, ventral view
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
|
||||
<emphasis id="1320379ABFEFB9DFD512F27A3F3A2A71" bold="true" pageId="0" pageNumber="41">C</emphasis>
|
||||
female habitus, dorsal view
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">D</emphasis>
|
||||
<emphasis id="0CDD9B560F41CFCC74DE6E2033D4C008" bold="true" pageId="0" pageNumber="41">D</emphasis>
|
||||
female habitus, ventral view
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">E</emphasis>
|
||||
<emphasis id="3EC15DF5C7841F9D3D6D1C29541DE5B6" bold="true" pageId="0" pageNumber="41">E</emphasis>
|
||||
female habitus, lateral view. Abbreviation: SP = spermatheca.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph pageId="0" pageNumber="41">
|
||||
<emphasis bold="true" pageId="0" pageNumber="41">Male.</emphasis>
|
||||
<paragraph id="90E7257F2DD7C01DAF8874816588BD3A" pageId="0" pageNumber="41">
|
||||
<emphasis id="63641FE0C1B1AAFCB626C32F6848D357" bold="true" pageId="0" pageNumber="41">Male.</emphasis>
|
||||
Unknown.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="41" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="41">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="41">
|
||||
<subSubSection id="AFB93830247D0D43BE97D4B9681B13C2" pageId="0" pageNumber="41" type="distribution">
|
||||
<paragraph id="02031E0CC03E05F309200BB0A4C59B4D" pageId="0" pageNumber="41">Distribution.</paragraph>
|
||||
<paragraph id="DBCF7A8A89BE68E2DDD91AA6660BF73B" pageId="0" pageNumber="41">
|
||||
Known only from the type locality (China; Fig.
|
||||
<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Distribution records of the new species of Merizocera from China and Myanmar. 1 M. baoshan sp. nov. 2 M. hponkanrazi sp. nov. 3 M. kachin sp. nov. 4 M. lincang sp. nov. 5 M. mainling sp. nov. 6 M. nyingchi sp. nov. 7 M. putao sp. nov. 8 M. tengchong sp. nov. 9 M. wenshan sp. nov. 10 M. wui sp. nov. 11 M. yuxi sp. nov." figureDoi="10.3897/zookeys.961.53058.figure53" httpUri="https://binary.pensoft.net/fig/443280" pageId="0" pageNumber="41">53</figureCitation>
|
||||
<figureCitation id="C87BC7AA7B21F7572C090B01D2FFAFA8" captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Distribution records of the new species of Merizocera from China and Myanmar. 1 M. baoshan sp. nov. 2 M. hponkanrazi sp. nov. 3 M. kachin sp. nov. 4 M. lincang sp. nov. 5 M. mainling sp. nov. 6 M. nyingchi sp. nov. 7 M. putao sp. nov. 8 M. tengchong sp. nov. 9 M. wenshan sp. nov. 10 M. wui sp. nov. 11 M. yuxi sp. nov." figureDoi="10.3897/zookeys.961.53058.figure53" httpUri="https://binary.pensoft.net/fig/443280" pageId="0" pageNumber="41">53</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
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