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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e9042" ID-PMC="PMC5039340" ID-Pensoft-Pub="1314-2828-4-9042" ID-PubMed="27733812" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828-4-e9042" ModsDocOrigin="Biodiversity Data Journal 4" ModsDocTitle="Checklist of British and Irish Hymenoptera - Ichneumonidae" checkinTime="1467812366136" checkinUser="pensoft" docAuthor="Broad, Gavin R." docDate="2016" docId="453B140A62B86C9A4E1D1F129D60C02A" docLanguage="en" docName="BiodivDatJour 4: e9042" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e9042" docTitle="PIMPLINAE Wesmael 1845" docType="treatment" docVersion="5" lastPageNumber="9042" masterDocId="FFD7A327EB31FFE35536456BB66AFFFB" masterDocTitle="Checklist of British and Irish Hymenoptera - Ichneumonidae" masterLastPageNumber="9042" masterPageNumber="9042" pageNumber="9042" updateTime="1668123934241" updateUser="ExternalLinkService">
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<taxonomicName authority="Wesmael, 1845" authorityName="Wesmael" authorityYear="1845" class="Insecta" family="Ichneumonidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="subFamily" subFamily="PIMPLINAE">PIMPLINAE Wesmael, 1845</taxonomicName>
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<paragraph pageId="0" pageNumber="9042">
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Ephialtinae" kingdom="Animalia" lsidName="EPHIALTINAE" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="genus">EPHIALTINAE</taxonomicName>
<normalizedToken originalValue="Hellén">Hellen</normalizedToken>
, 1915
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<paragraph pageId="0" pageNumber="9042">Notes</paragraph>
<paragraph pageId="0" pageNumber="9042">
Distribution data from
<bibRefCitation author="Fitton, M. G." journalOrPublisher="Handbooks for the Identification of British Insects" pageId="0" pageNumber="9042" pagination="1 - 110" title="Pimpline Ichneumon-flies. Hymenoptera, Ichneumonidae (Pimplinae)" volume="7" year="1988">Fitton et al. (1988)</bibRefCitation>
,
<bibRefCitation author="O'Connor, J. P." journalOrPublisher="Entomologist's Monthly Magazine" pageId="0" pageNumber="9042" pagination=": 164" title="Notes on some interesting Irish Pimplinae (Hym., Ichneumonidae)" volume="128" year="1992">O'Connor and Butler (1992)</bibRefCitation>
and
<bibRefCitation author="Shaw, M. R." journalOrPublisher="British Journal of Entomology and Natural History" pageId="0" pageNumber="9042" pagination="217 - 238" title="Notes on British Pimplinae and Poemeniinae (Hymenoptera, Ichneumonidae), with additions to the British list" volume="19" year="2006">Shaw (2006a)</bibRefCitation>
, with further distribution data for the
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Polysphincta" kingdom="Animalia" lsidName="Polysphincta" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="genus">Polysphincta</taxonomicName>
genus-group taken from
<bibRefCitation author="Hudson, I. R." journalOrPublisher="British Journal of Entomology and Natural History" pageId="0" pageNumber="9042" pagination="77 - 78" title="Some further notes on species of Ichneumonidae reared as ectoparasites of spiders" volume="1" year="1988">Hudson (1988)</bibRefCitation>
. Additional distribution references are given. Tribal classification follows
<bibRefCitation author="Gauld, I. D." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="9042" pagination="421 - 485" title="The suprageneric groups of the Pimplinae (Hymenoptera: Ichneumonidae): a cladistic re-evaluation and evolutionary biological study" volume="136" year="2002">Gauld et al. (2002)</bibRefCitation>
.
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<mods:namePart>Calvo, Joel</mods:namePart>
<mods:affiliation>Instituto de Geografia, Facultad de Ciencias del Mar y Geografia, Pontificia Universidad Catolica de Valparaiso, Avenida Brasil 2241, 2362807 Valparaiso, Chile</mods:affiliation>
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<mods:namePart>Moreira-Munoz, Andres</mods:namePart>
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1.
<taxonomicName LSID="453B585F-2467-59CF-8D51-DE937BE09F7B" authority="(Cuatrec.) V. A. Funk, Novon 7 (3): 238. 1997." class="Magnoliopsida" family="Asteraceae" genus="Xenophyllum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Xenophyllum acerosum" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="acerosum">Xenophyllum acerosum (Cuatrec.) V.A.Funk, Novon 7(3): 238. 1997.</taxonomicName>
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<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Werneria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Werneria acerosa" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="acerosa">Werneria acerosa</taxonomicName>
Cuatrec., Brittonia 8: 45. 1954. Type. Ecuador. Azuay:
<normalizedToken originalValue="“Oriente”">&quot;Oriente&quot;</normalizedToken>
border, eastern Cordillera, between
<normalizedToken originalValue="Oña">Ona</normalizedToken>
and the
<normalizedToken originalValue="río">rio</normalizedToken>
Yacuambi, 3050-3415 m, 10/19 Sep 1945,
<emphasis italics="true" pageId="0" pageNumber="1">F. Prieto 280</emphasis>
(holotype: F-1402673!; isotypes: G-00305681 (digital image!), GH s.n.!, K-000527742 (digital image!), MO-1652045!, NY s.n.!, P-02088566 (digital image!), S-R-6520 (digital image!), UC-986455!, US-00037296!, VEN-34483 (digital image!)).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Suffruticose plant, forming distant, rather solitary, erect stems.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Rhizomes</emphasis>
6-10
<normalizedToken originalValue="×">x</normalizedToken>
0.2-0.3 cm, horizontal to oblique, glabrous.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Stems</emphasis>
12-20 cm tall, usually simple, sometimes branched at the upper part, covered with adpressed, arachnoid trichomes, with leaves restricted to the upper part.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Leaves</emphasis>
imbricate, extending into a sheath-like base that bears arachnoid trichomes; leaf laminas 4.6-5
<normalizedToken originalValue="×">x</normalizedToken>
ca. 0.9 mm, linear-acicular, aristate at the apex (arista up to 1 mm long, purplish), entire, terete in cross section, glabrous, unconspicuously nerved on both faces, somewhat fleshy, drying coriaceous, matte, papillose.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Capitula</emphasis>
radiate, erect, sessile, partially enclosed among the leaves.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Involucres</emphasis>
ca. 8.6
<normalizedToken originalValue="×">x</normalizedToken>
7.3 mm, cupuliform; involucral bracts ca. 13, 5.2-5.5
<normalizedToken originalValue="×">x</normalizedToken>
1.1-1.6 mm, rather acute at the apex, dark-purplish.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Ray florets</emphasis>
ca. 13; corollas ca. 12
<normalizedToken originalValue="×">x</normalizedToken>
2.3-2.5 mm, 4 to 5-veined, subentire to 3-toothed at the apex, conspicuously surpassing the involucre, white, purplish beneath.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Disc florets</emphasis>
ca. 50; corollas 3.8-4.2 mm long, pale yellow to creamy; style branches truncate with a crown of sweeping trichomes, yellowish.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Achenes</emphasis>
cylindrical, with scattered arachnoid trichomes (immature); pappus 3.1-3.3 mm long, barbellate, whitish. Chromosome number unknown. Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Xenophyllum acerosum A habit B stem apical part and capitulum C detail of leaf surface D adaxial leaf surface E ray corolla and style F disc corolla and stamens (vertically sectioned, style removed) G disc corolla and style branches H style I immature achene with pappus. All details drawn from Prieto 280 (US). Illustration by Alice Tangerini." figureDoi="10.3897/phytokeys.158.50848.figure5" httpUri="https://binary.pensoft.net/fig/448885" pageId="0" pageNumber="1">5</figureCitation>
.
</paragraph>
<caption doi="10.3897/phytokeys.158.50848.figure5" httpUri="https://binary.pensoft.net/fig/448885" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Xenophyllum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Xenophyllum acerosum" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="acerosum">
<emphasis italics="true" pageId="0" pageNumber="1">Xenophyllum acerosum</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
habit
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
stem apical part and capitulum
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
detail of leaf surface
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
adaxial leaf surface
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
ray corolla and style
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
disc corolla and stamens (vertically sectioned, style removed)
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
disc corolla and style branches
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
style
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
immature achene with pappus. All details drawn from
<emphasis italics="true" pageId="0" pageNumber="1">Prieto 280</emphasis>
(US). Illustration by Alice Tangerini.
</paragraph>
</caption>
</subSubSection>
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<paragraph pageId="0" pageNumber="1">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">
Endemic to Ecuador (Azuay, Zamora-Chinchipe [expected]). It grows in marshy places of the paramo ecoregion, between elevations of 3050-3070 m (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution map of Xenophyllum acerosum (left hand) and X. humile (right hand)." figureDoi="10.3897/phytokeys.158.50848.figure6" httpUri="https://binary.pensoft.net/fig/448886" pageId="0" pageNumber="1">6</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Thus far,
<taxonomicName lsidName="X. acerosum" pageId="0" pageNumber="1" rank="species" species="acerosum">
<emphasis italics="true" pageId="0" pageNumber="1">X. acerosum</emphasis>
</taxonomicName>
is only known from the type locality in the paramo of Yacumbi, which is located above Saraguro on the border between the Ecuadorian provinces of Azuay and Zamora-Chinchipe.
</paragraph>
<caption doi="10.3897/phytokeys.158.50848.figure6" httpUri="https://binary.pensoft.net/fig/448886" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
Distribution map of
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Xenophyllum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Xenophyllum acerosum" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="acerosum">
<emphasis italics="true" pageId="0" pageNumber="1">Xenophyllum acerosum</emphasis>
</taxonomicName>
(left hand) and
<taxonomicName lsidName="X. humile" pageId="0" pageNumber="1" rank="species" species="humile">
<emphasis italics="true" pageId="0" pageNumber="1">X. humile</emphasis>
</taxonomicName>
(right hand).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="phenology">
<paragraph pageId="0" pageNumber="1">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="1">Collected in flower in September and November.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The adjective
<emphasis italics="true" pageId="0" pageNumber="1">acerosus -a -um</emphasis>
means needle-shaped, and it refers to the leaf shape of this species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="notes">
<paragraph pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Xenophyllum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Xenophyllum acerosum" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="acerosum">
<emphasis italics="true" pageId="0" pageNumber="1">Xenophyllum acerosum</emphasis>
</taxonomicName>
is unique in the genus in its distant, rather solitary, erect stems arising from long rhizomes and by its linear-acicular, aristate leaves. The stems only bear leaves in the upper part.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="additional specimen examined">
<paragraph pageId="0" pageNumber="1">Additional specimen examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Ecuador. Azuay</emphasis>
: Cordillera Cordoncillo, along road from Babes (ca. 2 km E of Urdaneta) towards 28 de Mayo,
<geoCoordinate degrees="3" direction="south" minutes="34" orientation="latitude" precision="925" value="-3.5666666">3°34'S</geoCoordinate>
,
<geoCoordinate degrees="79" direction="west" minutes="6" orientation="longitude" precision="925" value="-79.1">79°6'W</geoCoordinate>
, 10 Nov 2018,
<emphasis italics="true" pageId="0" pageNumber="1">
P.
<normalizedToken originalValue="Sklenář">Sklenar</normalizedToken>
15820
</emphasis>
(PRC n.v., QCA).
</paragraph>
</subSubSection>
</treatment>
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<mods:url id="90B8CD0F3A3769BEF1B22B19B1CE3452">http://antbase.org/ants/publications/4029/4029.pdf</mods:url>
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<mods:classification id="31972250F842FC4E0690AF3FB967D8B7">journal article</mods:classification>
<mods:identifier id="9217151E4B569542206342F83F990363" type="HNS-Pub">4029</mods:identifier>
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<treatment id="453BC3433E6ED3A200598D49D5C10F0C" ID-DOI="http://doi.org/10.5281/zenodo.6290386" ID-GBIF-Taxon="100129397" ID-Zenodo-Dep="6290386" LSID="urn:lsid:plazi:treatment:453BC3433E6ED3A200598D49D5C10F0C" httpUri="http://treatment.plazi.org/id/453BC3433E6ED3A200598D49D5C10F0C" lastPageNumber="301" pageNumber="301">
<subSubSection id="DB27DB77AA504AF01974AC33B3A43CD8" type="nomenclature">
<paragraph id="1DD4E3D0837607C185C4E358A0589D07">
<taxonomicName id="91A8654CE88DDD455D20BB084A70B707" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:245422" authority="Forel (Atopogyne)" authorityName="Forel (Atopogyne)" class="Insecta" family="Formicidae" genus="Cremastogaster" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-HNS="Cremastogaster montezumia var. functa" order="Hymenoptera" phylum="Arthropoda" rank="variety" species="montezumia" variety="functa">Cremastogaster Montezumia Sm. v. functa</taxonomicName>
<taxonomicNameLabel id="BF2F6DD5F50CFFE170DAE6AD2B4399AD" rank="variety">n. var.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="AA88B6C85CBD5F8D1F0251C234596CC9" type="description">
<paragraph id="BCE44906EB8F53C38B7F48CDA640B537" pageNumber="301">
<pageStartToken id="1CAF8C1C5D6AB5DDF9910B65EF6D6076" pageNumber="301">
<pageStartToken id="96C848C5726A4AE067D6AB9660ED213B" pageNumber="301">
<pageStartToken id="46B3525D7B0F1F2751BECE276B6A2751" pageNumber="301">[[</pageStartToken>
</pageStartToken>
</pageStartToken>
worker]]. Gleich
<normalizedToken id="E2F0F6898BCECF49840B261D681C1DDA" originalValue="groß">gross</normalizedToken>
wie die Stammart, aber hellbraun oder
<normalizedToken id="2CD10FF9383F408A4316B3CC241D879A" originalValue="rötlich">roetlich</normalizedToken>
braun. Der Kopf fast glatt und die Epinotumdornen sehr kurz, zahnartig, wie bei der
<taxonomicName id="C193CED1C09AB8459875E6BE91F54143" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239877" authority="Forel (Atopogyne)" authorityName="Forel (Atopogyne)" class="Insecta" family="Formicidae" genus="Cremastogaster" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-HNS="Cremastogaster sulcata var. ramulinida Forel" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="variety" species="salomonis" variety="ramulinida">var. ramulinida Forel</taxonomicName>
.
<normalizedToken id="09AD33C6514FB588888865332B6F6856" originalValue="Außerdem">Ausserdem</normalizedToken>
sind die Seiten des Epinotums glatt und
<normalizedToken id="4B6A55800A4673440D9DC409FDC843CC" originalValue="glänzend">glaenzend</normalizedToken>
. Der
<taxonomicName id="768F13A604110B7C3B0ADED7C3FD12EC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239877" authority="Forel (Atopogyne)" authorityName="Forel (Atopogyne)" class="Insecta" family="Formicidae" genus="Cremastogaster" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-HNS="Cremastogaster sulcata var. ramulinida Forel" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="variety" species="salomonis" variety="ramulinida">var. ramulinida For</taxonomicName>
. aus Columbien sehr nahe, aber viel heller
<normalizedToken id="5424EB464894FD0296F9A7D96F0A7D34" originalValue="gefärbt">gefaerbt</normalizedToken>
. Bei
<taxonomicName id="8F7FE9888420E0CBBFE90CB281261057" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239877" authority="Forel (Atopogyne)" authorityName="Forel (Atopogyne)" class="Insecta" family="Formicidae" genus="Cremastogaster" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-HNS="Cremastogaster sulcata var. ramulinida Forel" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="variety" species="salomonis" variety="ramulinida">v. ramulinida</taxonomicName>
sind die Seiten des Epinotums teilweise punktiert-genetzt.
</paragraph>
</subSubSection>
<subSubSection id="B9DB25EB432DF49EB4ED53B68EBE496C" type="materials_examined">
<paragraph id="46F90BAC57889AB747535FC20951658A" pageNumber="301">
Bahnhof Cubatao, Est. Sao Paulo (
<normalizedToken id="9B59CF6CE875E384B5C9449ABC103910" originalValue="Lüderwaldt">Luederwaldt</normalizedToken>
).
</paragraph>
</subSubSection>
</treatment>
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