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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.888.38642" ID-GBIF-Dataset="6501eef7-37d1-4746-a91d-9f5dbe146919" ID-PMC="PMC6861340" ID-Pensoft-Pub="1313-2970-888-75" ID-Pensoft-UUID="E4E0C850139853D8ADD841592FD3A4F4" ID-PubMed="31754320" ID-ZooBank="6244FB2F0BFE4235BADED25AEF147F39" ModsDocID="1313-2970-888-75" checkinTime="1573555945410" checkinUser="pensoft" docAuthor="Narakusumo, Raden Pramesa, Balke, Michael &amp; Riedel, Alexander" docDate="2019" docId="45051FAD824059F8A4730F927DE4CF0E" docLanguage="en" docName="ZooKeys 888: 75-93" docOrigin="ZooKeys 888" docSource="http://dx.doi.org/10.3897/zookeys.888.38642" docTitle="Trigonopterus Fauvel 1862" docType="treatment" docVersion="5" id="E4E0C850139853D8ADD841592FD3A4F4" lastPageNumber="75" masterDocId="E4E0C850139853D8ADD841592FD3A4F4" masterDocTitle="Seven new species of Trigonopterus Fauvel (Coleoptera, Curculionidae) from the Tanimbar Archipelago" masterLastPageNumber="93" masterPageNumber="75" pageNumber="75" updateTime="1668168052672" updateUser="ExternalLinkService">
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<mods:title>Seven new species of Trigonopterus Fauvel (Coleoptera, Curculionidae) from the Tanimbar Archipelago</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Narakusumo, Raden Pramesa</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Balke, Michael</mods:namePart>
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<mods:role>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.888.38642</mods:identifier>
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<treatment ID-GBIF-Taxon="160342554" LSID="urn:lsid:plazi:treatment:45051FAD824059F8A4730F927DE4CF0E" httpUri="http://treatment.plazi.org/id/45051FAD824059F8A4730F927DE4CF0E" lastPageNumber="75" pageId="0" pageNumber="75">
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<taxonomicName LSID="45051FAD-8240-59F8-A473-0F927DE4CF0E" authority="Fauvel, 1862" authorityName="Fauvel" authorityYear="1862" class="Insecta" family="Curculionidae" genus="Trigonopterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trigonopterus" order="Coleoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="genus">Trigonopterus Fauvel, 1862</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="type species">
<paragraph pageId="0" pageNumber="75">Type species, by monotypy.</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName class="Insecta" family="Curculionidae" genus="Trigonopterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trigonopterus insignis" order="Coleoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="insignis">
<emphasis italics="true" pageId="0" pageNumber="75">Trigonopterus insignis</emphasis>
</taxonomicName>
Fauvel, 1862.
</paragraph>
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<subSubSection pageId="0" pageNumber="75" type="diagnosis">
<paragraph pageId="0" pageNumber="75">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="75">
Fully apterous genus of
<taxonomicName lsidName="" pageId="0" pageNumber="75" rank="subfamily" subfamily="Cryptorhynchinae">Cryptorhynchinae</taxonomicName>
s. s. Length 1.5-6.0 mm. Rostrum in repose not reaching middle of mesocoxa. Scutellar shield completely absent externally. Mesothoracic receptacle deep, posteriorly closed. Metanepisternum completely absent externally. Elytra with nine striae (sometimes superficially effaced). Tarsal claws minute. Usually body largely unclothed, without dense vestiture. For additional information, see http://species-id.net/wiki/Trigonopterus.
</paragraph>
<subSection pageId="0" pageNumber="75" type="descriptions of species">
<paragraph pageId="0" pageNumber="75">Descriptions of species</paragraph>
</subSection>
</subSubSection>
</treatment>
</document>

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<treatment id="450587FEFFE5F440FC774BA4FBF7FA62" ID-DOI="http://doi.org/10.5281/zenodo.5743730" ID-GBIF-Taxon="198839686" ID-Zenodo-Dep="5743730" LSID="urn:lsid:plazi:treatment:450587FEFFE5F440FC774BA4FBF7FA62" httpUri="http://treatment.plazi.org/id/450587FEFFE5F440FC774BA4FBF7FA62" lastPageNumber="91" pageId="9" pageNumber="91">
<subSubSection id="85B66563FFE5F440FC774BA4FA98FB68" box="[934,1333,1198,1222]" pageId="9" pageNumber="91" type="nomenclature">
<paragraph id="CD1336E8FFE5F440FC774BA4FA98FB68" blockId="9.[934,1333,1198,1222]" box="[934,1333,1198,1222]" pageId="9" pageNumber="91">
<heading id="965B8184FFE5F440FC774BA4FA98FB68" bold="true" box="[934,1333,1198,1222]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="91" reason="2">
<taxonomicName id="0AAC4D6BFFE5F440FC774BA4FA98FB68" ID-CoL="7BB7Q" authority="(Attems, 1911)" baseAuthorityName="Attems" baseAuthorityYear="1911" box="[934,1333,1198,1222]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="polyzonus">
<emphasis id="FFD8EAFAFFE5F440FC774BA4FA98FB68" bold="true" box="[934,1333,1198,1222]" pageId="9" pageNumber="91">
<emphasis id="FFD8EAFAFFE5F440FC774BA4FB26FB68" bold="true" box="[934,1163,1198,1222]" italics="true" pageId="9" pageNumber="91">Syrioiulus polyzonus</emphasis>
(
<bibRefCitation id="A93D4B19FFE5F440FB484BA5FA83FB68" author="Attems C." box="[1177,1326,1199,1222]" pageId="9" pageNumber="91" pagination="61 - 67" refId="ref6296" refString="Attems C. 1911. Description de Myriapodes nouveaux recueillis par M. Henri Gadeau de Kerville pendant son voyage zoologique en Syrie. Bulletin de la Societe des Amis des Sciences naturelles de Rouen 46 (5): 61 - 67." type="journal article" year="1911">Attems, 1911</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="85B66563FFE5F440FCEE4BD6FB79FAD0" pageId="9" pageNumber="91" type="reference_group">
<paragraph id="CD1336E8FFE5F440FCEE4BD6FB5FFB5F" blockId="9.[831,1437,1244,1406]" box="[831,1266,1244,1265]" pageId="9" pageNumber="91">
<treatmentCitationGroup id="EDBC11C6FFE5F440FCEE4BD6FB5FFB5F" box="[831,1266,1244,1265]" pageId="9" pageNumber="91">
<treatmentCitation id="4C0D10F9FFE5F440FCEE4BD6FB43FB5F" author="Attems C." box="[831,1262,1244,1265]" page="67" pageId="9" pageNumber="91" year="1911">
<taxonomicName id="0AAC4D6BFFE5F440FCEE4BD6FB43FB5F" ID-CoL="6DDBY" authority="Attems, 1911: 66 - 67" authorityName="Attems" authorityPageNumber="66 - 67" authorityYear="1911" box="[831,1262,1244,1265]" class="Diplopoda" family="Julidae" genus="Dolichoiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="polyzonus">
<emphasis id="FFD8EAFAFFE5F440FCEE4BD6FB8FFB5F" box="[831,1058,1244,1265]" italics="true" pageId="9" pageNumber="91">Dolichoiulus polyzonus</emphasis>
<bibRefCitation id="A93D4B19FFE5F440FBF74BD6FB43FB5F" author="Attems C." box="[1062,1262,1244,1265]" pageId="9" pageNumber="91" pagination="61 - 67" refId="ref6296" refString="Attems C. 1911. Description de Myriapodes nouveaux recueillis par M. Henri Gadeau de Kerville pendant son voyage zoologique en Syrie. Bulletin de la Societe des Amis des Sciences naturelles de Rouen 46 (5): 61 - 67." type="journal article" year="1911">Attems, 1911: 66-67</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="CD1336E8FFE5F440FCEE4BF2FADCFAA3" blockId="9.[831,1437,1244,1406]" box="[831,1393,1272,1293]" pageId="9" pageNumber="91">
<treatmentCitationGroup id="EDBC11C6FFE5F440FCEE4BF2FADCFAA3" box="[831,1393,1272,1293]" pageId="9" pageNumber="91">
<taxonomicName id="0AAC4D6BFFE5F440FCEE4BF2FB34FAA3" ID-CoL="6DDBY" authorityName="Attems" authorityYear="1911" box="[831,1177,1272,1293]" class="Diplopoda" family="Julidae" genus="Dolichoiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="polyzonus" subGenus="Syrioiulus">
<emphasis id="FFD8EAFAFFE5F440FCEE4BF2FB34FAA3" box="[831,1177,1272,1293]" italics="true" pageId="9" pageNumber="91">Dolichoiulus (Syrioiulus) polyzonus</emphasis>
</taxonomicName>
. –
<treatmentCitation id="4C0D10F9FFE5F440FB674BF2FAC3FAA3" author="Verhoeff K. W." box="[1206,1390,1272,1293]" page="64" pageId="9" pageNumber="91" year="1914">
<bibRefCitation id="A93D4B19FFE5F440FB674BF2FAC3FAA3" author="Verhoeff K. W." box="[1206,1390,1272,1293]" pageId="9" pageNumber="91" pagination="61 - 75" refId="ref7617" refString="Verhoeff K. W. 1914. Einige Chilognathen aus Palastina (Uber Diplopoden, 68. Aufsatz). Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien 64: 61 - 75." type="journal article" year="1914">Verhoeff, 1914: 64</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="CD1336E8FFE5F440FCEE4A1EFC4FFAE8" blockId="9.[831,1437,1244,1406]" pageId="9" pageNumber="91">
<treatmentCitationGroup id="EDBC11C6FFE5F440FCEE4A1EFC4FFAE8" pageId="9" pageNumber="91">
<taxonomicName id="0AAC4D6BFFE5F440FCEE4A1EFB62FA87" ID-CoL="4a9df6a5-0ff8-4b8c-b38f-8008ffa5588f" baseAuthorityName="Attems" baseAuthorityYear="1909" box="[831,1231,1300,1321]" class="Diplopoda" family="Julidae" genus="Micropachyiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="polyzonus" subGenus="Syrioiulus">
<emphasis id="FFD8EAFAFFE5F440FCEE4A1EFB62FA87" box="[831,1231,1300,1321]" italics="true" pageId="9" pageNumber="91">Micropachyiulus (Syrioiulus) polyzonus</emphasis>
</taxonomicName>
. –
<treatmentCitation id="4C0D10F9FFE5F440FB2F4A1EFC73FAE8" author="Verhoeff K. W." page="122" pageId="9" pageNumber="91" year="1923">
<bibRefCitation id="A93D4B19FFE5F440FB2F4A1EFC73FAE8" author="Verhoeff K. W." pageId="9" pageNumber="91" pagination="112 - 157" refId="ref7650" refString="Verhoeff K. W. 1923. Zur Kenntnis der Palastina-Chilognathen und uber einige andere mediterrane Formen. 93. Diplopoden-Aufsatz. Archiv fur Naturgeschichte 89 A (4): 112 - 157." type="journal article" year="1923">Verhoeff, 1923: 122-123</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="CD1336E8FFE5F440FCEE4A47FAD2FACC" blockId="9.[831,1437,1244,1406]" box="[831,1407,1357,1378]" pageId="9" pageNumber="91">
<treatmentCitationGroup id="EDBC11C6FFE5F440FCEE4A47FAD2FACC" box="[831,1407,1357,1378]" pageId="9" pageNumber="91">
<taxonomicName id="0AAC4D6BFFE5F440FCEE4A47FBBCFACC" ID-CoL="CLVD" authorityName="Attems" authorityYear="1926" box="[831,1041,1357,1378]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="polyzonus">
<emphasis id="FFD8EAFAFFE5F440FCEE4A47FBBCFACC" box="[831,1041,1357,1378]" italics="true" pageId="9" pageNumber="91">Amblyiulus polyzonus</emphasis>
</taxonomicName>
. –
<treatmentCitation id="4C0D10F9FFE5F440FBFF4A47FAA3FACC" author="Attems C." box="[1070,1294,1357,1378]" page="263" pageId="9" pageNumber="91" year="1926">
<bibRefCitation id="A93D4B19FFE5F440FBFF4A47FAA3FACC" author="Attems C." box="[1070,1294,1357,1378]" pageId="9" pageNumber="91" pagination="221 - 266" refId="ref6340" refString="Attems C. 1926. Etude sur les Myriopodes recueillis par M. Henri Gadeau de Kerville pendant son voyage zoologique en Syrie (Avril-Juin 1908) [pp. 221 - 266]. In: Bailliere J. - B. et fils (eds). Voyage zoologique d'Henri Gadeau de Kerville en Syrie (Avril-Juin 1908). Imprimerie Lecerf fils, Rouen, 365 pp." type="book chapter" year="1926">Attems, 1926: 263-264</bibRefCitation>
</treatmentCitation>
, figs 15-18.
</treatmentCitationGroup>
</paragraph>
<paragraph id="CD1336E8FFE5F440FCEE4A63FB79FAD0" blockId="9.[831,1437,1244,1406]" box="[831,1236,1385,1406]" pageId="9" pageNumber="91">
<treatmentCitationGroup id="EDBC11C6FFE5F440FCEE4A63FB79FAD0" box="[831,1236,1385,1406]" pageId="9" pageNumber="91">
<taxonomicName id="0AAC4D6BFFE5F440FCEE4A63FBAAFAD0" ID-CoL="7BB7Q" baseAuthorityName="Attems" baseAuthorityYear="1911" box="[831,1031,1385,1406]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="polyzonus">
<emphasis id="FFD8EAFAFFE5F440FCEE4A63FBAAFAD0" box="[831,1031,1385,1406]" italics="true" pageId="9" pageNumber="91">Syrioiulus polyzonus</emphasis>
</taxonomicName>
. –
<treatmentCitation id="4C0D10F9FFE5F440FBF44A63FB7DFAD0" author="Jeekel C. A. W." box="[1061,1232,1385,1406]" page="171" pageId="9" pageNumber="91" year="1970">
<bibRefCitation id="A93D4B19FFE5F440FBF44A63FB7DFAD0" author="Jeekel C. A. W." box="[1061,1232,1385,1406]" pageId="9" pageNumber="91" pagination="1 - 412" refId="ref6950" refString="Jeekel C. A. W. 1970. Nomenclator generum et familiarum Diplopodorum: A list of the genus and family-group names in the class Diplopoda from the 10 th edition of Linnaeus, 1758, to the end of 1957. Monografieen van de Nederlandse Entomologische Vereniging 5: 1 - 412." type="journal article" year="1970">Jeekel, 1970: 171</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFE5F440FCEE4A9CFBF7FA62" pageId="9" pageNumber="91" type="distribution">
<paragraph id="CD1336E8FFE5F440FCEE4A9CFBF7FA62" blockId="9.[831,1436,1430,1484]" pageId="9" pageNumber="91">
<emphasis id="FFD8EAFAFFE5F440FCEE4A9CFC53FA03" bold="true" box="[831,1022,1430,1453]" pageId="9" pageNumber="91">Known records:</emphasis>
<materialsCitation id="7DC43CB5FFE5F440FBDB4A9CFBF7FA62" ID-GBIF-Occurrence="3881331314" country="Syria" location="Berze" pageId="9" pageNumber="91" specimenCount="1">
<collectingCountry id="B5BB7678FFE5F440FBDB4A9CFBE8FA03" box="[1034,1093,1430,1453]" name="Syria" pageId="9" pageNumber="91">Syria</collectingCountry>
,
<location id="C8736033FFE5F440FB874A9CFB38FA03" LSID="urn:lsid:plazi:treatment:450587FEFFE5F440FC774BA4FBF7FA62:C8736033FFE5F440FB874A9CFB38FA03" box="[1110,1173,1430,1453]" country="Syria" name="Berze" pageId="9" pageNumber="91">Berzé</location>
[Barzeh, today part of Damascus] (type locality).
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72
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(type locality)
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<emphasis id="FFD8EAFAFFE5F440FF0E4BA4FE60FB68" bold="true" box="[223,461,1198,1222]" italics="true" pageId="9" pageNumber="91">Syrioiulus incarnatus</emphasis>
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<treatmentCitation id="4C0D10F9FFE5F440FF464BD6FDDCFB5F" author="Lohmander H." box="[151,625,1244,1265]" page="35" pageId="9" pageNumber="91" year="1932">
<taxonomicName id="0AAC4D6BFFE5F440FF464BD6FDDCFB5F" ID-CoL="7R295" authority="Lohmander, 1932 a: 33 - 35" authorityName="Lohmander" authorityPageNumber="33 - 35" authorityYear="1932" box="[151,625,1244,1265]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="incarnatus">
<emphasis id="FFD8EAFAFFE5F440FF464BD6FEDDFB5F" box="[151,368,1244,1265]" italics="true" pageId="9" pageNumber="91">Amblyiulus incarnatus</emphasis>
<bibRefCitation id="A93D4B19FFE5F440FEA74BD6FDDCFB5F" author="Lohmander H." box="[374,625,1244,1265]" pageId="9" pageNumber="91" pagination="1 - 44" refId="ref7038" refString="Lohmander H. 1932 a. Neue Diplopoden aus Persien. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles handlingar 5, Ser. B 3 (2): 1 - 44." type="journal article" year="1932">Lohmander, 1932a: 33-35</bibRefCitation>
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, fig. 26.
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<treatmentCitationGroup id="EDBC11C6FFE5F440FF464BF2FDFBFAA3" box="[151,598,1272,1293]" pageId="9" pageNumber="91">
?
<taxonomicName id="0AAC4D6BFFE5F440FF794BF2FEDBFAA3" ID-CoL="542P8" baseAuthorityName="Lohmander" baseAuthorityYear="1932" box="[168,374,1272,1293]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="incarnatus">
<emphasis id="FFD8EAFAFFE5F440FF794BF2FEDBFAA3" box="[168,374,1272,1293]" italics="true" pageId="9" pageNumber="91">Syrioiulus incarnatus</emphasis>
</taxonomicName>
.
<emphasis id="FFD8EAFAFFE5F440FE534BF2FE23FAA3" box="[386,398,1272,1293]" italics="true" pageId="9" pageNumber="91">–</emphasis>
<treatmentCitation id="4C0D10F9FFE5F440FE454BF2FDFFFAA3" author="Mauries J. - P." box="[404,594,1272,1293]" page="441" pageId="9" pageNumber="91" year="1982">
<bibRefCitation id="A93D4B19FFE5F440FE454BF2FDFFFAA3" author="Mauries J. - P." box="[404,594,1272,1293]" pageId="9" pageNumber="91" pagination="433 - 444" refId="ref7158" refString="Mauries J. - P. 1982. Dolichoiulus tongiorgii (Strasser), Diplopode halophile nouveau pour la faune de France. Remarques sur la classification des Pachyiulini (Myriapoda, Diplopoda, Iulida). Bulletin du Museum national d'Histoire naturelle 4 (3 - 4): 433 - 444." type="journal article" year="1982">Mauriès, 1982: 441</bibRefCitation>
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.
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<treatmentCitationGroup id="EDBC11C6FFE5F440FF464A1EFE05FAE8" pageId="9" pageNumber="91">
<emphasis id="FFD8EAFAFFE5F440FF464A1EFED9FA87" box="[151,372,1300,1321]" italics="true" pageId="9" pageNumber="91">
<taxonomicName id="0AAC4D6BFFE5F440FF464A1EFEDDFA87" ID-CoL="542P8" baseAuthorityName="Lohmander" baseAuthorityYear="1932" box="[151,368,1300,1321]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="incarnatus">Syrioiulus incarnatus</taxonomicName>
.
</emphasis>
–
<treatmentCitation id="4C0D10F9FFE5F440FE4B4A1EFDF6FA87" author="Mauries J. - P." box="[410,603,1300,1321]" page="48" pageId="9" pageNumber="91" year="1984">
<bibRefCitation id="A93D4B19FFE5F440FE4B4A1EFDF6FA87" author="Mauries J. - P." box="[410,603,1300,1321]" pageId="9" pageNumber="91" pagination="37 - 49" refId="ref7212" refString="Mauries J. - P. 1984. Deux especes nouvelles de Diplopodes cavernicoles des Cyclades: Hyleoglomeris beroni (Glomerida) et Syrioiulus andreevi (Iulida). Biologia Gallohellenica 11: 37 - 49." type="journal article" year="1984">Mauriès, 1984: 48</bibRefCitation>
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. – Enghoff &amp; Moravvej, 2005: 64.
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<emphasis id="FFD8EAFAFFE5F440FF464A57FEF9FADA" bold="true" box="[151,340,1373,1396]" pageId="9" pageNumber="91">Known records:</emphasis>
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<collectingCountry id="B5BB7678FFE5F440FE8F4A57FE21FADA" box="[350,396,1373,1396]" name="Iran" pageId="9" pageNumber="91">Iran</collectingCountry>
,
<location id="C8736033FFE5F440FE4B4A57FD59FADA" LSID="urn:lsid:plazi:treatment:450587FEFFE5F440FF0E4BA4FE65F9EE:C8736033FFE5F440FE4B4A57FD59FADA" box="[410,756,1373,1396]" country="Iran" name="eastern slopes of Bogrov Dagh" pageId="9" pageNumber="91">eastern slopes of Bogrov Dāgh</location>
[Talysh Mts], “Altschaly” (type locality)
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.
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<emphasis id="FFD8EAFAFFE5F440FF464AA1FF78FA6C" bold="true" box="[151,213,1451,1474]" pageId="9" pageNumber="91">Note:</emphasis>
Despite the unknown male characters, the vulval structure in this species is so similar with that observed in the other three blind representatives of
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known from
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that there can be no doubt that
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is their congener.
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<emphasis id="FFD8EAFAFFE5F440FF244839FE6BF8E5" bold="true" box="[245,454,1843,1867]" italics="true" pageId="9" pageNumber="91">Syrioiulus persicus</emphasis>
(
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<bibRefCitation id="A93D4B19FFE5F440FE8E486BFDF3F8D8" author="Golovatch S. I." box="[351,606,1889,1910]" pageId="9" pageNumber="91" pagination="157 - 169" refId="ref6753" refString="Golovatch S. I. 1983. A contribution to the millipede fauna of Iran (Diplopoda). Annalen des Naturhistorischen Museums in Wien (Serie B) 85: 157 - 169." type="journal article" year="1983">Golovatch, 1983: 161-162</bibRefCitation>
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, figs 3-7.
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<emphasis id="FFD8EAFAFFE5F440FF464874FEE2F83D" box="[151,335,1918,1939]" italics="true" pageId="9" pageNumber="91">Syrioiulus persicus</emphasis>
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. –
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,
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[Karaj] (
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159
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?
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(
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, 123-125, figs 4-6.
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<emphasis id="FFD8EAFAFFE5F440FCEE4973FB84F920" box="[831,1065,1657,1678]" italics="true" pageId="9" pageNumber="91">Amblyiulus posthirsutus</emphasis>
</taxonomicName>
.
<emphasis id="FFD8EAFAFFE5F440FBE84973FBE8F920" box="[1081,1093,1657,1678]" italics="true" pageId="9" pageNumber="91">–</emphasis>
<treatmentCitation id="4C0D10F9FFE5F440FB9F4973FA99F920" author="Attems C." box="[1102,1332,1657,1678]" page="258" pageId="9" pageNumber="91" year="1926">
<bibRefCitation id="A93D4B19FFE5F440FB9F4973FA99F920" author="Attems C." box="[1102,1332,1657,1678]" pageId="9" pageNumber="91" pagination="221 - 266" refId="ref6340" refString="Attems C. 1926. Etude sur les Myriopodes recueillis par M. Henri Gadeau de Kerville pendant son voyage zoologique en Syrie (Avril-Juin 1908) [pp. 221 - 266]. In: Bailliere J. - B. et fils (eds). Voyage zoologique d'Henri Gadeau de Kerville en Syrie (Avril-Juin 1908). Imprimerie Lecerf fils, Rouen, 365 pp." type="book chapter" year="1926">Attems, 1926: 258-259</bibRefCitation>
</treatmentCitation>
. –
<treatmentCitation id="4C0D10F9FFE5F440FA874973FB91F905" author="Bodenheimer F. S." page="233" pageId="9" pageNumber="91" year="1937">
<bibRefCitation id="A93D4B19FFE5F440FA874973FB91F905" author="Bodenheimer F. S." pageId="9" pageNumber="91" pagination="1 - 286" refId="ref6524" refString="Bodenheimer F. S. 1937. Prodromus faunae Palestinae. Essai sur les elements zoogeographiques et historiques du sud-ouest du sous-regne palearctique. Memoires presentes a l'Institut d'Egypte 33 (1 - 2): 1 - 286." type="journal article" year="1937">Bodenheimer, 1937: 233</bibRefCitation>
</treatmentCitation>
. –
<treatmentCitation id="4C0D10F9FFE5F440FB86499CFABFF905" author="Tabacaru I." box="[1111,1298,1686,1707]" page="24" pageId="9" pageNumber="91" year="1995">
<bibRefCitation id="A93D4B19FFE5F440FB86499CFABFF905" author="Tabacaru I." box="[1111,1298,1686,1707]" pageId="9" pageNumber="91" pagination="19 - 28" refId="ref7508" refString="Tabacaru I. 1995. Diplopodes d'Israel. 1. Libanaphe adonis galilaensis n. ssp. avec une liste revisee des Diplopodes signales en Israel [pp. 19 - 28]. In: Por F. D., Decu V., Negrea S., Dimentman C. (eds.). Soil Fauna of Israel. Volume 1. Editura Academiei Romane, Bucharest, 156 pp." type="book chapter" year="1995">Tabacaru, 1995: 24</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="CD1336E8FFE5F440FCEE49B8FC6FF94A" blockId="9.[831,1437,1601,1851]" pageId="9" pageNumber="91">
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<taxonomicName id="0AAC4D6BFFE5F440FCEE49B8FB45F969" ID-CoL="8N7XG" baseAuthorityName="Verhoeff" baseAuthorityYear="1923" box="[831,1256,1714,1735]" class="Diplopoda" family="Julidae" genus="Trichopachyiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="posthirsutus" subGenus="Iudaeoiulus">
<emphasis id="FFD8EAFAFFE5F440FCEE49B8FB45F969" box="[831,1256,1714,1735]" italics="true" pageId="9" pageNumber="91">Trichopachyiulus (Iudaeoiulus) posthirsutus</emphasis>
</taxonomicName>
.
<emphasis id="FFD8EAFAFFE5F440FB2549B8FAADF969" box="[1268,1280,1714,1735]" italics="true" pageId="9" pageNumber="91">–</emphasis>
<treatmentCitation id="4C0D10F9FFE5F440FAD749B8FC13F94A" author="Verhoeff K. W." page="1670" pageId="9" pageNumber="91" year="1930">
<bibRefCitation id="A93D4B19FFE5F440FAD749B8FC13F94A" author="Verhoeff K. W." pageId="9" pageNumber="91" pagination="1523 - 1674" refId="ref7685" refString="Verhoeff K. W. 1930. 2. Buch: Diplopoda. 10. Lieferung [pp. 1523 - 1674]. In: Bronn H. G. (ed.). Klassen und Ordnungen des Tierreichs, 5 (2). Akademische Verlagsgesellschaft, Leipzig, 2083 pp." type="book chapter" year="1930">Verhoeff, 1930: 1670</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
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<emphasis id="FFD8EAFAFFE5F440FCEE49E6FC0DF8AF" box="[831,928,1772,1793]" italics="true" pageId="9" pageNumber="91">Syrioiulus</emphasis>
</taxonomicName>
sp.
<emphasis id="FFD8EAFAFFE5F440FC1949E6FC79F8AF" box="[968,980,1772,1793]" italics="true" pageId="9" pageNumber="91">–</emphasis>
<treatmentCitation id="4C0D10F9FFE5F440FC0B49E6FABAF8AF" author="Golovatch S. I. &amp; Wytwer J." box="[986,1303,1772,1793]" page="170" pageId="9" pageNumber="91" year="2009">
<bibRefCitation id="A93D4B19FFE5F440FC0B49E6FABAF8AF" author="Golovatch S. I. &amp; Wytwer J." box="[986,1303,1772,1793]" pageId="9" pageNumber="91" pagination="167 - 170" refId="ref6858" refString="Golovatch S. I., Wytwer J. 2009. A remarkable new species of the millipede family Julidae from Israel (Diplopoda, Julida) [pp. 167 - 170]. In: Roble S. M., Mitchell J. C. (eds.). A lifetime of contributions to myriapodology and the natural history of Virginia: a Festschrift in honor of Richard L. Hoffman's 80 th birthday. Virginia Museum of Natural History, Martinsville, VA, 458 pp." type="book chapter" year="2009">Golovatch &amp; Wytwer, 2009: 170</bibRefCitation>
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.
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<emphasis id="FFD8EAFAFFE5F440FCEE4803FBB0F8B0" box="[831,1053,1801,1822]" italics="true" pageId="9" pageNumber="91">Syrioiulus posthirsutus</emphasis>
</taxonomicName>
. –
<treatmentCitation id="4C0D10F9FFE5F440FBEE4803FAECF8B0" author="Golovatch S. I." box="[1087,1345,1801,1822]" page="792" pageId="9" pageNumber="91" year="2018">
<bibRefCitation id="A93D4B19FFE5F440FBEE4803FAECF8B0" author="Golovatch S. I." box="[1087,1345,1801,1822]" pageId="9" pageNumber="91" pagination="791 - 805" refId="ref6818" refString="Golovatch S. I. 2018. The millipede subfamily Pachyiulinae in Israel, with the description of a new species (Diplopoda, Julida, Julidae). Zoologicheskii zhurnal 97 (7): 791 - 805." type="journal article" year="2018">Golovatch, 2018: 792-794</bibRefCitation>
</treatmentCitation>
, figs 1A- C, 2A-E.
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<paragraph id="CD1336E8FFE5F440FCEE4859FBF8F7A4" blockId="9.[831,1436,1875,2058]" pageId="9" pageNumber="91">
<emphasis id="FFD8EAFAFFE5F440FCEE4859FBA9F8C4" bold="true" box="[831,1028,1875,1898]" pageId="9" pageNumber="91">Known records:</emphasis>
All from
<collectingCountry id="B5BB7678FFE5F440FB5D4859FB61F8C4" box="[1164,1228,1875,1898]" name="Israel" pageId="9" pageNumber="91">Israel</collectingCountry>
: Rehovot, Hulda, Nahal Rubin National Park, by the Sea of Galilee (original localities); Judean Mts, Devir (
<bibRefCitation id="A93D4B19FFE5F440FADB4899FC48F864" author="Golovatch S. I. &amp; Wytwer J." pageId="9" pageNumber="91" pagination="167 - 170" refId="ref6858" refString="Golovatch S. I., Wytwer J. 2009. A remarkable new species of the millipede family Julidae from Israel (Diplopoda, Julida) [pp. 167 - 170]. In: Roble S. M., Mitchell J. C. (eds.). A lifetime of contributions to myriapodology and the natural history of Virginia: a Festschrift in honor of Richard L. Hoffman's 80 th birthday. Virginia Museum of Natural History, Martinsville, VA, 458 pp." type="book chapter" year="2009">Golovatch &amp; Wytwer, 2009</bibRefCitation>
;
<bibRefCitation id="A93D4B19FFE5F440FC2A48B9FB12F864" author="Golovatch S. I." box="[1019,1215,1971,1994]" pageId="9" pageNumber="91" pagination="791 - 805" refId="ref6818" refString="Golovatch S. I. 2018. The millipede subfamily Pachyiulinae in Israel, with the description of a new species (Diplopoda, Julida, Julidae). Zoologicheskii zhurnal 97 (7): 791 - 805." type="journal article" year="2018">Golovatch, 2018</bibRefCitation>
); Timrat, Allonim, Nahal Mearot Nature Preserve, Giveat Yeshaiahu, Regba (
<bibRefCitation id="A93D4B19FFE5F440FC4348F9FBE7F7A4" author="Golovatch S. I." box="[914,1098,2035,2058]" pageId="9" pageNumber="91" pagination="791 - 805" refId="ref6818" refString="Golovatch S. I. 2018. The millipede subfamily Pachyiulinae in Israel, with the description of a new species (Diplopoda, Julida, Julidae). Zoologicheskii zhurnal 97 (7): 791 - 805." type="journal article" year="2018">Golovatch, 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
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<paragraph id="CD1336E8FFE6F443FF464E08FD22FE4F" blockId="10.[151,756,258,481]" pageId="10" pageNumber="92">
<emphasis id="FFD8EAFAFFE6F443FF464E08FF7BFEB7" bold="true" box="[151,214,258,281]" pageId="10" pageNumber="92">Note:</emphasis>
Athough the solenomere in this species seems to correspond with the current diagnosis of
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<emphasis id="FFD8EAFAFFE6F443FD504E29FD43FE94" box="[641,750,291,314]" italics="true" pageId="10" pageNumber="92">Syrioiulus</emphasis>
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, the intermediate lobe connecting the proximal sections of mesomeral process and solenomere, and especially the complete absence of apical denticles on the promere are characters that set
<taxonomicName id="0AAC4D6BFFE6F443FE704EA3FDEAFE6E" baseAuthorityName="Verhoeff" baseAuthorityYear="1923" box="[417,583,425,448]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="posthirsutus">
<emphasis id="FFD8EAFAFFE6F443FE704EA3FDEAFE6E" box="[417,583,425,448]" italics="true" pageId="10" pageNumber="92">S. posthirsutus</emphasis>
</taxonomicName>
apart from all remaining congeners, hence the question mark.
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?
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(Lohmander, 1932)
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<emphasis id="FFD8EAFAFFE6F443FF464C76FECDFC3F" box="[151,352,892,913]" italics="true" pageId="10" pageNumber="92">Amblyiulus zarudnyi</emphasis>
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, figs 21-25. –
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<bibRefCitation id="A93D4B19FFE6F443FF324C90FE1BFC01" author="Golovatch S. I." box="[227,438,922,943]" pageId="10" pageNumber="92" pagination="157 - 169" refId="ref6753" refString="Golovatch S. I. 1983. A contribution to the millipede fauna of Iran (Diplopoda). Annalen des Naturhistorischen Museums in Wien (Serie B) 85: 157 - 169." type="journal article" year="1983">Golovatch, 1983: 161</bibRefCitation>
.
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<emphasis id="FFD8EAFAFFE6F443FF464CB2FEE5FC63" box="[151,328,952,973]" italics="true" pageId="10" pageNumber="92">Syrioiulus zarudni</emphasis>
</taxonomicName>
(sic!). –
<treatmentCitation id="4C0D10F9FFE6F443FE4E4CB2FDCCFC63" author="Mauries J. - P." box="[415,609,952,973]" page="441" pageId="10" pageNumber="92" year="1982">
<bibRefCitation id="A93D4B19FFE6F443FE4E4CB2FDF0FC63" author="Mauries J. - P." box="[415,605,952,973]" pageId="10" pageNumber="92" pagination="433 - 444" refId="ref7158" refString="Mauries J. - P. 1982. Dolichoiulus tongiorgii (Strasser), Diplopode halophile nouveau pour la faune de France. Remarques sur la classification des Pachyiulini (Myriapoda, Diplopoda, Iulida). Bulletin du Museum national d'Histoire naturelle 4 (3 - 4): 433 - 444." type="journal article" year="1982">Mauriès, 1982: 441</bibRefCitation>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="CD1336E8FFE6F443FF464CDDFE0AFBA4" blockId="10.[151,756,892,1034]" pageId="10" pageNumber="92">
<treatmentCitationGroup id="EDBC11C6FFE6F443FF464CDDFE0AFBA4" pageId="10" pageNumber="92">
<taxonomicName id="0AAC4D6BFFE6F443FF464CDDFEF1FC42" box="[151,348,983,1004]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="zarydnyi">
<emphasis id="FFD8EAFAFFE6F443FF464CDDFEF1FC42" box="[151,348,983,1004]" italics="true" pageId="10" pageNumber="92">Syrioiulus zarydnyi</emphasis>
</taxonomicName>
. –
<treatmentCitation id="4C0D10F9FFE6F443FE5C4CDDFDF9FC42" author="Mauries J. - P." box="[397,596,983,1004]" page="48" pageId="10" pageNumber="92" year="1984">
<bibRefCitation id="A93D4B19FFE6F443FE5C4CDDFDF9FC42" author="Mauries J. - P." box="[397,596,983,1004]" pageId="10" pageNumber="92" pagination="37 - 49" refId="ref7212" refString="Mauries J. - P. 1984. Deux especes nouvelles de Diplopodes cavernicoles des Cyclades: Hyleoglomeris beroni (Glomerida) et Syrioiulus andreevi (Iulida). Biologia Gallohellenica 11: 37 - 49." type="journal article" year="1984">Mauriès, 1984: 48</bibRefCitation>
</treatmentCitation>
.
<emphasis id="FFD8EAFAFFE6F443FDB64CDDFDDEFC42" box="[615,627,983,1004]" italics="true" pageId="10" pageNumber="92">–</emphasis>
<treatmentCitation id="4C0D10F9FFE6F443FD524CDDFE09FBA4" author="Enghoff H. &amp; Moravvej, S. A." page="65" pageId="10" pageNumber="92" year="2005">Enghoff &amp; Moravvej, 2005: 65</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="CD1336E8FFE6F443FCEE4E08FAF8FED7" blockId="10.[831,1436,258,377]" pageId="10" pageNumber="92">
<materialsCitation id="7DC43CB5FFE6F443FCEE4E08FAB1FE97" ID-GBIF-Occurrence="3881331301" collectorName="Canyon Sefidruda" country="Iran" isTailEnumeration="true" location="Khuzestan Province" pageId="10" pageNumber="92" specimenCount="1" stateProvince="Khuzestan Province" typeStatus="type">
<emphasis id="FFD8EAFAFFE6F443FCEE4E08FBADFEB7" bold="true" box="[831,1024,258,281]" pageId="10" pageNumber="92">Known records:</emphasis>
All from
<collectingCountry id="B5BB7678FFE6F443FBAF4E08FB01FEB7" box="[1150,1196,258,281]" name="Iran" pageId="10" pageNumber="92">Iran</collectingCountry>
,
<collectingRegion id="0F68F80AFFE6F443FB6F4E08FA31FEB7" box="[1214,1436,258,281]" country="Iran" name="Khuzestan" pageId="10" pageNumber="92">Khuzestan Province</collectingRegion>
(type locality) [exact locality unknown];
</materialsCitation>
<materialsCitation id="7DC43CB5FFE6F443FAFC4E28FB5AFEF7" ID-GBIF-Occurrence="3881331304" collectorName="Canyon Sefidruda" country="Iran" isTailEnumeration="true" location="NW Iran" pageId="10" pageNumber="92" specimenCount="1" stateProvince="Yezd">
<location id="C8736033FFE6F443FAFC4E28FA35FE97" LSID="urn:lsid:plazi:treatment:450587FEFFE6F442FF314C47FE22FEF7:C8736033FFE6F443FAFC4E28FA35FE97" box="[1325,1432,290,313]" country="Iran" name="NW Iran" pageId="10" pageNumber="92" stateProvince="Yezd">NW Iran</location>
,
<location id="C8736033FFE6F443FCEE4E48FBA9FEF7" LSID="urn:lsid:plazi:treatment:450587FEFFE6F442FF314C47FE22FEF7:C8736033FFE6F443FCEE4E48FBA9FEF7" box="[831,1028,322,345]" country="Iran" name="Canyon Sefidruda" pageId="10" pageNumber="92" stateProvince="Yezd">Canyon Sefidruda</location>
[on Sefid-rud River];
</materialsCitation>
<materialsCitation id="7DC43CB5FFE6F443FAD34E48FAF8FED7" ID-GBIF-Occurrence="3881331315" collectorName="Canyon Sefidruda" country="Iran" isTailEnumeration="true" location="end of Keredj" pageId="10" pageNumber="92" specimenCount="1" stateProvince="Yezd">
<locationDeviation id="DCA12FCAFFE6F443FAD34E48FCD5FED7" pageId="10" pageNumber="92">
<quantity id="0A549B0DFFE6F443FAD34E48FAE4FEF7" box="[1282,1353,322,345]" metricMagnitude="4" metricUnit="m" metricValue="6.8" pageId="10" pageNumber="92" unit="km" value="68.0">68 km</quantity>
SW of Yezd
</locationDeviation>
; N
<location id="C8736033FFE6F443FC4C4E68FB9DFED7" LSID="urn:lsid:plazi:treatment:450587FEFFE6F442FF314C47FE22FEF7:C8736033FFE6F443FC4C4E68FB9DFED7" box="[925,1072,354,377]" country="Iran" name="end of Keredj" pageId="10" pageNumber="92" stateProvince="Yezd">end of Keredj</location>
[Karaj] (
<bibRefCitation id="A93D4B19FFE6F443FB404E68FAE5FED7" author="Golovatch S. I." box="[1169,1352,354,377]" pageId="10" pageNumber="92" pagination="157 - 169" refId="ref6753" refString="Golovatch S. I. 1983. A contribution to the millipede fauna of Iran (Diplopoda). Annalen des Naturhistorischen Museums in Wien (Serie B) 85: 157 - 169." type="journal article" year="1983">Golovatch, 1983</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFE6F443FCEE4E9BFE8CFBD3" pageId="10" pageNumber="92" type="discussion">
<paragraph id="CD1336E8FFE6F443FCEE4E9BFBB7FD46" blockId="10.[831,1437,401,1032]" pageId="10" pageNumber="92">
<emphasis id="FFD8EAFAFFE6F443FCEE4E9BFCD3FE06" bold="true" box="[831,894,401,424]" pageId="10" pageNumber="92">Note:</emphasis>
This is another species of
<taxonomicName id="0AAC4D6BFFE6F443FB6E4E9BFA81FE06" authorityName="Verhoeff" authorityYear="1914" box="[1215,1324,401,424]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFE6F443FB6E4E9BFA81FE06" box="[1215,1324,401,424]" italics="true" pageId="10" pageNumber="92">Syrioiulus</emphasis>
</taxonomicName>
deviating from the majority of congeners by the conformations of promere and mesomeral processes which are alike those of
<taxonomicName id="0AAC4D6BFFE6F443FC634EFBFBCDFDA6" authorityName="Lohmander" authorityYear="1939" box="[946,1120,497,520]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="cappadocius">
<emphasis id="FFD8EAFAFFE6F443FC634EFBFBCDFDA6" box="[946,1120,497,520]" italics="true" pageId="10" pageNumber="92">S. cappadocius</emphasis>
</taxonomicName>
. Furthermore,
<taxonomicName id="0AAC4D6BFFE6F443FAC94EFBFA31FDA6" authorityName="Lohmander" authorityYear="1932" box="[1304,1436,497,520]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="zarudnyi">
<emphasis id="FFD8EAFAFFE6F443FAC94EFBFA31FDA6" box="[1304,1436,497,520]" italics="true" pageId="10" pageNumber="92">S. zarudnyi</emphasis>
</taxonomicName>
has a prominent anterior lamella of the solenomere (albeit apically not drawn into a process) which is also untypical of the genus. However, before a comprehensive revision and phylogeny of
<taxonomicName id="0AAC4D6BFFE6F443FAFE4D7BFA31FD26" authorityName="Verhoeff" authorityYear="1914" box="[1327,1436,625,648]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFE6F443FAFE4D7BFA31FD26" box="[1327,1436,625,648]" italics="true" pageId="10" pageNumber="92">Syrioiulus</emphasis>
</taxonomicName>
and related genera is made, it seems better that this and the other two aforementioned doubtful species are placed in
<taxonomicName id="0AAC4D6BFFE6F443FC764DDBFBBEFD46" authorityName="Verhoeff" authorityYear="1914" box="[935,1043,721,744]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFE6F443FC764DDBFBBEFD46" box="[935,1043,721,744]" italics="true" pageId="10" pageNumber="92">Syrioiulus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="CD1336E8FFE6F443FCEE4DFBFA31FBA6" blockId="10.[831,1437,401,1032]" pageId="10" pageNumber="92">
According to
<bibRefCitation id="A93D4B19FFE6F443FC0C4DFBFB1AFCA6" author="Lohmander H." box="[989,1207,753,776]" pageId="10" pageNumber="92" pagination="1 - 44" refId="ref7038" refString="Lohmander H. 1932 a. Neue Diplopoden aus Persien. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles handlingar 5, Ser. B 3 (2): 1 - 44." type="journal article" year="1932">Lohmander (1932a)</bibRefCitation>
the
<typeStatus id="1217884AFFE6F443FB214DFBFAB3FCA6" box="[1264,1310,753,776]" pageId="10" pageNumber="92">type</typeStatus>
locality of
<taxonomicName id="0AAC4D6BFFE6F443FCEE4C1BFC1AFC86" authorityName="Lohmander" authorityYear="1932" box="[831,951,785,808]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="zarudnyi">
<emphasis id="FFD8EAFAFFE6F443FCEE4C1BFC1AFC86" box="[831,951,785,808]" italics="true" pageId="10" pageNumber="92">S. zarudnyi</emphasis>
</taxonomicName>
is the “Tschemle-Rogan”
<location id="C8736033FFE6F443FB1F4C1BFA30FC86" LSID="urn:lsid:plazi:treatment:450587FEFFE6F442FF314C47FE22FEF7:C8736033FFE6F443FB1F4C1BFA30FC86" box="[1230,1437,785,808]" country="Iran" name="Spring in Arabistan" pageId="10" pageNumber="92" stateProvince="Khuzestan">Spring in Arabistan</location>
(today
<collectingRegion id="0F68F80AFFE6F443FC434C3BFBD8FCE6" box="[914,1141,817,840]" country="Iran" name="Khuzestan" pageId="10" pageNumber="92">Khuzestan Province</collectingRegion>
). The toponym obviously refers to “Tscheschme-Rogan” [“Cheshme” (English transliteration) meaning “spring” in Persian], “in the land of the Bakhtiary people”. This is one of the sites visited by the original collector of this species, the Russian zoologist Nikolai Zarudny, during his forth expedition to
<collectingCountry id="B5BB7678FFE6F443FAA04CDBFA31FC46" box="[1393,1436,977,1000]" name="Iran" pageId="10" pageNumber="92">Iran</collectingCountry>
(
<bibRefCitation id="A93D4B19FFE6F443FC964CFBFC7FFBA6" author="Birula A." box="[839,978,1009,1032]" pageId="10" pageNumber="92" pagination="119 - 148" refId="ref6495" refString="Birula A. 1905. Beitrage zur Kenntniss der Skorpionenfauna Persiens. Bulletin de l'Academie Imperiale des Sciences 23 (1 - 2): 119 - 148." type="journal article" year="1905">Birula, 1905</bibRefCitation>
). The exact location of that spring remains
</paragraph>
<paragraph id="CD1336E8FFE6F443FF464B4CFE8CFBD3" blockId="10.[151,1436,1093,1149]" pageId="10" pageNumber="92">
<emphasis id="FFD8EAFAFFE6F443FF464B4CFE8CFBD3" bold="true" pageId="10" pageNumber="92">
Key to the species of
<taxonomicName id="0AAC4D6BFFE6F443FE5D4B4FFE50FBF3" authorityName="Verhoeff" authorityYear="1914" box="[396,509,1093,1117]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFE6F443FE5D4B4FFE50FBF3" bold="true" box="[396,509,1093,1117]" italics="true" pageId="10" pageNumber="92">Syrioiulus</emphasis>
</taxonomicName>
known from
<collectingCountry id="B5BB7678FFE6F443FD484B4CFD66FBF3" box="[665,715,1094,1117]" name="Iran" pageId="10" pageNumber="92">Iran</collectingCountry>
based on characters of gonopods, vulvae and external somatic morphology
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFE6F442FF464BACFE22FEF7" lastPageId="11" lastPageNumber="93" pageId="10" pageNumber="92" type="key">
<key id="B93D61D9FFE6F443FF464BACFA34F7A3" pageId="10" pageNumber="92">
<keyStep id="76582E4DFFE6F443FF464BACFA37FB73" pageId="10" pageNumber="92">
<paragraph id="CD1336E8FFE6F443FF464BACFA37FB13" blockId="10.[151,1434,1190,1245]" box="[151,1434,1190,1213]" pageId="10" pageNumber="92">
<keyLead id="765D95DDFFE6F443FF464BACFA37FB13" box="[151,1434,1190,1213]" pageId="10" pageNumber="92">1A With ommatidia ....................................................................................................................................................... 2</keyLead>
</paragraph>
<paragraph id="CD1336E8FFE6F443FF464BCCFA37FB73" blockId="10.[151,1434,1190,1245]" box="[151,1434,1222,1245]" pageId="10" pageNumber="92">
<keyLead id="765D95DDFFE6F443FF464BCCFA37FB73" box="[151,1434,1222,1245]" pageId="10" pageNumber="92">1B Without ommatidia .................................................................................................................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="76582E4DFFE6F443FF464BFCFA37FAE3" pageId="10" pageNumber="92">
<paragraph id="CD1336E8FFE6F443FF464BFCFA34FA83" blockId="10.[151,1434,1270,1357]" pageId="10" pageNumber="92">
<keyLead id="765D95DDFFE6F443FF464BFCFA34FA83" pageId="10" pageNumber="92">
2A Opisthomere with a poorly developed mesocaudal lamella and a thick, apically finely dentate, anterior lamella .... ................................................................................................................................
<taxonomicName id="0AAC4D6BFFE6F443FBEF4A1CFA34FA83" authority="(Lohmander, 1932)" baseAuthorityName="Lohmander" baseAuthorityYear="1932" box="[1086,1433,1301,1326]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="zarudnyi">
<emphasis id="FFD8EAFAFFE6F443FBEF4A1CFA34FA83" bold="true" box="[1086,1433,1301,1326]" pageId="10" pageNumber="92">
<emphasis id="FFD8EAFAFFE6F443FBEF4A1CFB17FA83" bold="true" box="[1086,1210,1301,1326]" italics="true" pageId="10" pageNumber="92">S. zarudnyi</emphasis>
(Lohmander, 1932)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="CD1336E8FFE6F443FF464A3CFA37FAE3" blockId="10.[151,1434,1270,1357]" box="[151,1434,1334,1357]" pageId="10" pageNumber="92">
<keyLead id="765D95DDFFE6F443FF464A3CFA37FAE3" box="[151,1434,1334,1357]" pageId="10" pageNumber="92">2B Opisthomere with a well-developed mesocaudal lamella and a rather indistinct anterior lamella .......................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="76582E4DFFE6F443FF464A6CFA37F993" pageId="10" pageNumber="92">
<paragraph id="CD1336E8FFE6F443FF464A6CFA34FA73" blockId="10.[151,1436,1382,1598]" pageId="10" pageNumber="92">
<keyLead id="765D95DDFFE6F443FF464A6CFA34FA73" pageId="10" pageNumber="92">
3A Mesocaudal lamella apically forming a freely protruding pointed outgrowth, this being subequal to or slightly exceeding solenomere; ozopores in anterior part of body set at some distance behind pro-metazonal suture; metazonal hind margins without setae; walking legs in males without adhesive pads .............................................. .................................................................................................................................
<taxonomicName id="0AAC4D6BFFE6F443FB964ACCFA34FA73" ID-CoL="542P7" authority="(Lohmander, 1932)" baseAuthorityName="Lohmander" baseAuthorityYear="1932" box="[1095,1433,1477,1502]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="FFD8EAFAFFE6F443FB964ACCFA34FA73" bold="true" box="[1095,1433,1477,1502]" pageId="10" pageNumber="92">
<emphasis id="FFD8EAFAFFE6F443FB964ACCFB17FA73" bold="true" box="[1095,1210,1477,1502]" italics="true" pageId="10" pageNumber="92">S. discolor</emphasis>
(Lohmander, 1932)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="CD1336E8FFE6F443FF464AECFA37F993" blockId="10.[151,1436,1382,1598]" pageId="10" pageNumber="92">
<keyLead id="765D95DDFFE6F443FF464AECFA37F993" pageId="10" pageNumber="92">
3B Mesocaudal lamella without freely protruding apical part but ending bluntly much lower than tip of solenomere; ozopores in anterior part of body set right behind pro-metazonal suture; metazonal hind margins with a whorl of short setae; walking legs in males with adhesive pads on tibiae ............................
<taxonomicName id="0AAC4D6BFFE6F443FB92492CFA37F993" ID-CoL="84DND" authority="(Attems, 1903)" baseAuthorityName="Attems" baseAuthorityYear="1903" box="[1091,1434,1573,1598]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="continentalis">
<emphasis id="FFD8EAFAFFE6F443FB92492CFA3CF993" bold="true" box="[1091,1425,1573,1598]" pageId="10" pageNumber="92">
<emphasis id="FFD8EAFAFFE6F443FB92492CFB40F993" bold="true" box="[1091,1261,1573,1598]" italics="true" pageId="10" pageNumber="92">S. continentalis</emphasis>
(
<bibRefCitation id="A93D4B19FFE6F443FB2A492CFA3CF993" author="Attems C. G." box="[1275,1425,1574,1597]" pageId="10" pageNumber="92" pagination="63 - 154" refId="ref6264" refString="Attems C. G. 1903. Beitrage zur Myriopodenkunde. Zoologische Jahrbucher - Abteilung fur Systematik, Okologie und Geographie der Tiere 18 (1): 63 - 154." type="journal article" year="1903">Attems, 1903</bibRefCitation>
</emphasis>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="76582E4DFFE6F443FF46495CFA37F903" pageId="10" pageNumber="92">
<paragraph id="CD1336E8FFE6F443FF46495CFA37F923" blockId="10.[151,1437,1622,1709]" pageId="10" pageNumber="92">
<keyLead id="765D95DDFFE6F443FF46495CFA37F923" pageId="10" pageNumber="92">
4A Metazonal hind margins with a whorl of setae; vulval operculum only slightly exceeding bursa; promere with a rounded, lobe-like mesal denticle .............................................................................
<taxonomicName id="0AAC4D6BFFE6F443FB82497CFA37F923" ID-CoL="542P9" authority="(Golovatch, 1983)" baseAuthorityName="Golovatch" baseAuthorityYear="1983" box="[1107,1434,1654,1678]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="persicus">
<emphasis id="FFD8EAFAFFE6F443FB82497CFA37F923" bold="true" box="[1107,1434,1654,1678]" pageId="10" pageNumber="92">
<emphasis id="FFD8EAFAFFE6F443FB82497CFB64F920" bold="true" box="[1107,1225,1654,1678]" italics="true" pageId="10" pageNumber="92">S. persicus</emphasis>
(
<bibRefCitation id="A93D4B19FFE6F443FB09497CFA3EF923" author="Golovatch S. I." box="[1240,1427,1654,1677]" pageId="10" pageNumber="92" pagination="157 - 169" refId="ref6753" refString="Golovatch S. I. 1983. A contribution to the millipede fauna of Iran (Diplopoda). Annalen des Naturhistorischen Museums in Wien (Serie B) 85: 157 - 169." type="journal article" year="1983">Golovatch, 1983</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="CD1336E8FFE6F443FF46499CFA37F903" blockId="10.[151,1437,1622,1709]" box="[151,1434,1686,1709]" pageId="10" pageNumber="92">
<keyLead id="765D95DDFFE6F443FF46499CFA37F903" box="[151,1434,1686,1709]" pageId="10" pageNumber="92">4B Metazonal hind margins without setae; vulval operculum considerably exceeding bursa ...................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="76582E4DFFE6F443FF4649CCFA37F8F3" pageId="10" pageNumber="92">
<paragraph id="CD1336E8FFE6F443FF4649CCFA37F8B3" blockId="10.[151,1436,1734,1885]" pageId="10" pageNumber="92">
<keyLead id="765D95DDFFE6F443FF4649CCFA37F8B3" pageId="10" pageNumber="92">
5A Ozopores in anterior part of body set at 1/4-1/3 of metazonal length behind pro-metazonal suture; apical margin of vulval operculum with a distinct incision mid-laterally (
<figureCitation id="55972A6DFFE6F443FCEB49ECFC20F953" box="[826,909,1766,1789]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="10" pageNumber="92">Figs 10</figureCitation>
,
<figureCitation id="55972A6DFFE6F443FC4849ECFC1BF953" box="[921,950,1766,1789]" captionStart="Figs 19-21" captionStartId="9.[151,192,1067,1088]" captionTargetBox="[151,1436,256,1015]" captionTargetId="figure-299@9.[518,886,541,826]" captionTargetPageId="9" captionText="Figs 19-21. Semidiagrammatic drawings of vulvae in caudal views of Syrioiulus spp. (19) S. astrabadensis. (20) S. incarnatus. (21) S. lohmanderi sp. nov. (Figs 19-20 redrawn from Lohmander, 1932a). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743595" httpUri="https://zenodo.org/record/5743595/files/figure.png" pageId="10" pageNumber="92">21</figureCitation>
); mesomeral process subequal to solenomere (
<figureCitation id="55972A6DFFE6F443FF38480CFEC9F8B3" box="[233,356,1798,1821]" captionStart="Figs 11-14" captionStartId="5.[151,192,1052,1073]" captionTargetBox="[179,1412,258,1005]" captionTargetId="figure-18@5.[518,886,510,772]" captionTargetPageId="5" captionText="Figs 11-14. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (11) Left promere, caudal view. (12) Right pair of gonopods, mesal view. (13) Left opisthomere, caudal, slightly lateral view. (14) Distal parts of left opisthomere, lateral view.Abbreviations: cl = caudomesal lamella, ff = central filiform process, fp = fan-like process; m = mesomeral process; P = promere; s = solenomere; sp = shield-like process. Scale bar of Figs 11-13: 0.1 mm; Figs 14-15: 0.05 mm; Fig. 16: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.5743581" httpUri="https://zenodo.org/record/5743581/files/figure.png" pageId="10" pageNumber="92">Figs 12, 14</figureCitation>
,
<figureCitation id="55972A6DFFE6F443FEBE480CFE1FF8B3" box="[367,434,1798,1821]" captionStart="Figs 15-16" captionStartId="5.[151,192,2028,2049]" captionTargetBox="[151,1436,1266,1989]" captionTargetId="figure-137@5.[151,1436,1266,1989]" captionTargetPageId="5" captionText="Figs 15-16. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (15) Distal parts of left opisthomere, caudo-lateral view. (16) Solenomere, same aspect." figureDoi="http://doi.org/10.5281/zenodo.5743585" httpUri="https://zenodo.org/record/5743585/files/figure.png" pageId="10" pageNumber="92">15, 16</figureCitation>
) ..............................................................................................................
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<taxonomicName id="0AAC4D6BFFE6F443FB73480CFA93F8B3" ID-CoL="31ed5e90-30a8-413d-9108-6f6c61e4af18" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[1186,1342,1797,1822]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFE6F443FB73480CFA93F8B3" bold="true" box="[1186,1342,1797,1822]" italics="true" pageId="10" pageNumber="92">S. lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFE6F443FA94480CFA37F8B3" box="[1349,1434,1798,1821]" pageId="10" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
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<keyLead id="765D95DDFFE6F443FF46482CFA37F8F3" pageId="10" pageNumber="92">5B Ozopores in anterior part of body set on or right behind pro-metazonal sutures; apical margin of vulval operculum more or less smooth ................................................................................................................................................. 6</keyLead>
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</keyStep>
<keyStep id="76582E4DFFE6F443FF46487CFA34F7A3" pageId="10" pageNumber="92">
<paragraph id="CD1336E8FFE6F443FF46487CFA37F803" blockId="10.[151,1437,1910,2062]" pageId="10" pageNumber="92">
<keyLead id="765D95DDFFE6F443FF46487CFA37F803" pageId="10" pageNumber="92">
6A Vulva with its operculum exceeding bursa by 1/4-1/3 of total vulval height (
<figureCitation id="55972A6DFFE6F443FBC4487CFBCBF823" box="[1045,1126,1910,1933]" captionStart="Figs 19-21" captionStartId="9.[151,192,1067,1088]" captionTargetBox="[151,1436,256,1015]" captionTargetId="figure-299@9.[518,886,541,826]" captionTargetPageId="9" captionText="Figs 19-21. Semidiagrammatic drawings of vulvae in caudal views of Syrioiulus spp. (19) S. astrabadensis. (20) S. incarnatus. (21) S. lohmanderi sp. nov. (Figs 19-20 redrawn from Lohmander, 1932a). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743595" httpUri="https://zenodo.org/record/5743595/files/figure.png" pageId="10" pageNumber="92">Fig. 20</figureCitation>
); adult females
<quantity id="0A549B0DFFE6F443FAC0487CFA2DF823" box="[1297,1408,1910,1933]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" pageId="10" pageNumber="92" unit="mm" value="31.0" valueMax="32.0" valueMin="30.0">30-32 mm</quantity>
in length and
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in diameter .........................
<taxonomicName id="0AAC4D6BFFE6F443FD2D489CFBC3F803" ID-CoL="542P8" authority="(Lohmander, 1932)" baseAuthorityName="Lohmander" baseAuthorityYear="1932" box="[764,1134,1942,1966]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="incarnatus">
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<emphasis id="FFD8EAFAFFE6F443FD2D489CFC3DF800" bold="true" box="[764,912,1942,1966]" italics="true" pageId="10" pageNumber="92">S. incarnatus</emphasis>
(Lohmander, 1932)
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(known only from females)
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<paragraph id="CD1336E8FFE6F443FF4648BCFA34F7A3" blockId="10.[151,1437,1910,2062]" pageId="10" pageNumber="92">
<keyLead id="765D95DDFFE6F443FF4648BCFA34F7A3" pageId="10" pageNumber="92">
6B Vulva with its operculum exceeding bursa by 1/5-1/4 of total vulval height (
<figureCitation id="55972A6DFFE6F443FBDD48BCFBF6F863" box="[1036,1115,1974,1997]" captionStart="Figs 19-21" captionStartId="9.[151,192,1067,1088]" captionTargetBox="[151,1436,256,1015]" captionTargetId="figure-299@9.[518,886,541,826]" captionTargetPageId="9" captionText="Figs 19-21. Semidiagrammatic drawings of vulvae in caudal views of Syrioiulus spp. (19) S. astrabadensis. (20) S. incarnatus. (21) S. lohmanderi sp. nov. (Figs 19-20 redrawn from Lohmander, 1932a). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743595" httpUri="https://zenodo.org/record/5743595/files/figure.png" pageId="10" pageNumber="92">Fig. 19</figureCitation>
); females
<quantity id="0A549B0DFFE6F443FB1848BCFA9AF863" box="[1225,1335,1974,1997]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.8" metricValueMin="2.4" pageId="10" pageNumber="92" unit="mm" value="26.0" valueMax="28.0" valueMin="24.0">24-28 mm</quantity>
in length and
<quantity id="0A549B0DFFE6F443FEDF48DCFE25F843" box="[270,392,2006,2029]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="10" pageNumber="92" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4-1.5 mm</quantity>
in diameter; males with expanded mandibular stipites; mesomeral process shorter than solenomere (
<figureCitation id="55972A6DFFE6F443FF3B48FCFE9AF7A3" box="[234,311,2038,2061]" captionStart="Fig" captionStartId="6.[151,186,1310,1331]" captionTargetBox="[215,694,269,1269]" captionTargetId="figure-544@6.[151,493,532,809]" captionTargetPageId="6" captionText="Fig. 17. Syrioiulus astrabadensis (Lohmander, 1932a), paratype male, in GNM, distal parts of right opisthomere in mesal view. Abbreviations: cl = caudomesal lamella; ff = central filiform process; m = mesomeral process; s = solenomere. Scale bar: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5743587" httpUri="https://zenodo.org/record/5743587/files/figure.png" pageId="10" pageNumber="92">Fig. 17</figureCitation>
) .........................................................................................................
<taxonomicName id="0AAC4D6BFFE6F443FBD848FCFA34F7A3" ID-CoL="542P6" authority="(Lohmander, 1932)" baseAuthorityName="Lohmander" baseAuthorityYear="1932" box="[1033,1433,2037,2062]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="astrabadensis">
<emphasis id="FFD8EAFAFFE6F443FBD848FCFA34F7A3" bold="true" box="[1033,1433,2037,2062]" pageId="10" pageNumber="92">
<emphasis id="FFD8EAFAFFE6F443FBD848FCFB16F7A3" bold="true" box="[1033,1211,2037,2062]" italics="true" pageId="10" pageNumber="92">S. astrabadensis</emphasis>
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unknown, but it is most likely situated in the eastern parts of
<collectingRegion id="0F68F80AFFE7F442FF694E28FE84FE97" box="[184,297,290,313]" country="Iran" name="Khuzestan" pageId="11" pageNumber="93">Khuzestan</collectingRegion>
which is inhabited predominantly by the Bakhtiari ethnic group.
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106
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, figs 336-338. –
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. –
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. –
<treatmentCitation id="4C0D10F9FFE6F443FEBC4D93FD86FD00" author="Mauries J. - P." box="[365,555,665,686]" page="441" pageId="10" pageNumber="92" year="1982">
<bibRefCitation id="A93D4B19FFE6F443FEBC4D93FD86FD00" author="Mauries J. - P." box="[365,555,665,686]" pageId="10" pageNumber="92" pagination="433 - 444" refId="ref7158" refString="Mauries J. - P. 1982. Dolichoiulus tongiorgii (Strasser), Diplopode halophile nouveau pour la faune de France. Remarques sur la classification des Pachyiulini (Myriapoda, Diplopoda, Iulida). Bulletin du Museum national d'Histoire naturelle 4 (3 - 4): 433 - 444." type="journal article" year="1982">Mauriès, 1982: 441</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFE6F443FF464DC2FE62FCAE" pageId="10" pageNumber="92" type="description">
<paragraph id="CD1336E8FFE6F443FF464DC2FE62FCAE" blockId="10.[151,756,712,768]" pageId="10" pageNumber="92">
<emphasis id="FFD8EAFAFFE6F443FF464DC2FEE2FD71" bold="true" box="[151,335,712,735]" pageId="10" pageNumber="92">Known records:</emphasis>
<materialsCitation id="7DC43CB5FFE6F443FE854DC2FE62FCAE" ID-GBIF-Occurrence="3881331303" country="Azerbaijan" location="area of Talysh Mts" pageId="10" pageNumber="92" specimenCount="1" stateProvince="Lenkoran">
<collectingCountry id="B5BB7678FFE6F443FE854DC2FE61FD71" box="[340,460,712,735]" name="Azerbaijan" pageId="10" pageNumber="92">Azerbaijan</collectingCountry>
,
<collectingRegion id="0F68F80AFFE6F443FE064DC2FD90FD71" box="[471,573,712,735]" country="Azerbaijan" name="Lankaran" pageId="10" pageNumber="92">Lenkoran</collectingRegion>
[Lankaran],
<location id="C8736033FFE6F443FD194DC2FE87FCAE" LSID="urn:lsid:plazi:treatment:450587FEFFE6F443FED74D27FE62FCAE:C8736033FFE6F443FD194DC2FE87FCAE" country="Azerbaijan" name="area of Talysh Mts" pageId="10" pageNumber="92" stateProvince="Lenkoran">area of Talysh Mts</location>
(type locality).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

230
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<mods:title id="190E0E687A9E0CB17CA0068EC3823395">A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae)</mods:title>
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<treatment id="450587FEFFEAF44EFCA94B82FEF4FD76" ID-DOI="http://doi.org/10.5281/zenodo.6975137" ID-GBIF-Taxon="198839710" ID-Zenodo-Dep="6975137" LSID="urn:lsid:plazi:treatment:450587FEFFEAF44EFCA94B82FEF4FD76" httpUri="http://treatment.plazi.org/id/450587FEFFEAF44EFCA94B82FEF4FD76" lastPageId="7" lastPageNumber="89" pageId="6" pageNumber="88">
<subSubSection id="85B66563FFEAF44FFCA94B82FAC9FB0E" box="[888,1380,1160,1184]" pageId="6" pageNumber="88" type="nomenclature">
<paragraph id="CD1336E8FFEAF44FFCA94B82FAC9FB0E" blockId="6.[888,1380,1160,1216]" box="[888,1380,1160,1184]" pageId="6" pageNumber="88">
<heading id="965B8184FFEAF44FFCA94B82FAC9FB0E" bold="true" box="[888,1380,1160,1184]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="88" reason="2">
<taxonomicName id="0AAC4D6BFFEAF44FFCA94B82FAC9FB0E" ID-CoL="542P6" authority="(Lohmander, 1932)" baseAuthorityName="Lohmander" baseAuthorityYear="1932" box="[888,1380,1160,1184]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="88" phylum="Arthropoda" rank="species" species="astrabadensis">
<emphasis id="FFD8EAFAFFEAF44FFCA94B82FAC9FB0E" bold="true" box="[888,1380,1160,1184]" pageId="6" pageNumber="88">
<emphasis id="FFD8EAFAFFEAF44FFCA94B82FB28FB0E" bold="true" box="[888,1157,1160,1184]" italics="true" pageId="6" pageNumber="88">Syrioiulus astrabadensis</emphasis>
(Lohmander, 1932)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEAF44FFB994BA3FB39FB6E" box="[1096,1172,1193,1216]" pageId="6" pageNumber="88" type="description">
<paragraph id="CD1336E8FFEAF44FFB994BA3FB39FB6E" blockId="6.[888,1380,1160,1216]" box="[1096,1172,1193,1216]" pageId="6" pageNumber="88">
<figureCitation id="55972A6DFFEAF44FFB994BA3FB39FB6E" box="[1096,1172,1193,1216]" captionStart="Fig" captionStartId="6.[151,186,1310,1331]" captionTargetBox="[215,694,269,1269]" captionTargetId="figure-544@6.[151,493,532,809]" captionTargetPageId="6" captionText="Fig. 17. Syrioiulus astrabadensis (Lohmander, 1932a), paratype male, in GNM, distal parts of right opisthomere in mesal view. Abbreviations: cl = caudomesal lamella; ff = central filiform process; m = mesomeral process; s = solenomere. Scale bar: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5743587" httpUri="https://zenodo.org/record/5743587/files/figure.png" pageId="6" pageNumber="88">Fig. 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEAF44FFCEE4BDDFC07FAA7" pageId="6" pageNumber="88" type="reference_group">
<paragraph id="CD1336E8FFEAF44FFCEE4BDDFC07FAA7" blockId="6.[831,1437,1239,1289]" pageId="6" pageNumber="88">
<treatmentCitationGroup id="EDBC11C6FFEAF44FFCEE4BDDFC07FAA7" pageId="6" pageNumber="88">
<treatmentCitation id="4C0D10F9FFEAF44FFCEE4BDDFAE9FB42" author="Lohmander H." box="[831,1348,1239,1260]" page="33" pageId="6" pageNumber="88" year="1932">
<taxonomicName id="0AAC4D6BFFEAF44FFCEE4BDDFAE9FB42" ID-CoL="7R28Z" authority="Lohmander, 1932 a: 30 - 33" authorityName="Lohmander" authorityPageNumber="30 - 33" authorityYear="1932" box="[831,1348,1239,1260]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="88" phylum="Arthropoda" rank="species" species="astrabadensis">
<emphasis id="FFD8EAFAFFEAF44FFCEE4BDDFB94FB42" box="[831,1081,1239,1260]" italics="true" pageId="6" pageNumber="88">Amblyiulus astrabadensis</emphasis>
<bibRefCitation id="A93D4B19FFEAF44FFB924BDDFAE9FB42" author="Lohmander H." box="[1091,1348,1239,1260]" pageId="6" pageNumber="88" pagination="1 - 44" refId="ref7038" refString="Lohmander H. 1932 a. Neue Diplopoden aus Persien. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles handlingar 5, Ser. B 3 (2): 1 - 44." type="journal article" year="1932">Lohmander, 1932a: 30-33</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 30- 32.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="99D36660FFEAF44FFF464A14FEFFFA6D" ID-DOI="http://doi.org/10.5281/zenodo.5743587" ID-Zenodo-Dep="5743587" httpUri="https://zenodo.org/record/5743587/files/figure.png" pageId="6" pageNumber="88" startId="6.[151,186,1310,1331]" targetBox="[215,694,269,1269]" targetPageId="6">
<paragraph id="CD1336E8FFEAF44FFF464A14FEFFFA6D" blockId="6.[151,757,1310,1475]" pageId="6" pageNumber="88">
Fig. 17.
<taxonomicName id="0AAC4D6BFFEAF44FFF274A14FD5DFA9D" authority="(Lohmander, 1932 a)" baseAuthorityName="Lohmander" baseAuthorityYear="1932" box="[246,752,1310,1331]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="88" phylum="Arthropoda" rank="species" species="astrabadensis">
<emphasis id="FFD8EAFAFFEAF44FFF274A14FE55FA9D" box="[246,504,1310,1331]" italics="true" pageId="6" pageNumber="88">Syrioiulus astrabadensis</emphasis>
(
<bibRefCitation id="A93D4B19FFEAF44FFDCB4A14FD44FA9D" author="Lohmander H." box="[538,745,1310,1331]" pageId="6" pageNumber="88" pagination="1 - 44" refId="ref7038" refString="Lohmander H. 1932 a. Neue Diplopoden aus Persien. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles handlingar 5, Ser. B 3 (2): 1 - 44." type="journal article" year="1932">Lohmander, 1932a</bibRefCitation>
)
</taxonomicName>
, paratype male, in GNM, distal parts of right opisthomere in mesal view. Abbreviations:
<emphasis id="FFD8EAFAFFEAF44FFD144A52FD7BFAC3" box="[709,726,1368,1389]" italics="true" pageId="6" pageNumber="88">cl</emphasis>
= caudomesal lamella;
<emphasis id="FFD8EAFAFFEAF44FFE1C4A7FFE77FA24" box="[461,474,1397,1418]" italics="true" pageId="6" pageNumber="88">ff</emphasis>
=
<emphasis id="FFD8EAFAFFEAF44FFDD14A7FFEAAFA09" italics="true" pageId="6" pageNumber="88">central filiform process; m</emphasis>
= mesomeral process;
<emphasis id="FFD8EAFAFFEAF44FFE3F4A98FE5AFA09" box="[494,503,1426,1447]" italics="true" pageId="6" pageNumber="88">s</emphasis>
= solenomere. Scale bar: 0.05 mm.
</paragraph>
</caption>
<subSubSection id="85B66563FFEAF44FFCEE4A28FB8BF905" pageId="6" pageNumber="88" type="materials_examined">
<paragraph id="CD1336E8FFEAF44FFCEE4A28FC4FF9F5" blockId="6.[831,1437,1314,1627]" pageId="6" pageNumber="88">
<emphasis id="FFD8EAFAFFEAF44FFCEE4A28FC00FA97" bold="true" box="[831,941,1314,1337]" pageId="6" pageNumber="88">Material:</emphasis>
<materialsCitation id="7DC43CB5FFEAF44FFC6F4A28FB42FAD7" ID-GBIF-Occurrence="3881331310" collectingDate="1905-05-02" collectionCode="GNM" country="Iran" location="Persien" pageId="6" pageNumber="88" specimenCount="2" typeStatus="syntype">
<collectionCode id="ABBDAE2DFFEAF44FFC6F4A28FBADFA97" box="[958,1024,1314,1337]" collectionName="Sweden, Goeteborg, Naturhistoriska Museet" country="Sweden" httpUri="http://grbio.org/cool/m262-zrs7" name="Gothenburg Museum of Natural History (Goteborgs Naturhistoriska Museum)" pageId="6" pageNumber="88">GNM</collectionCode>
;
<specimenCount id="DBAAFD61FFEAF44FFBC44A28FB8FFA97" box="[1045,1058,1314,1337]" pageId="6" pageNumber="88" type="generic">2</specimenCount>
(supposed) female
<typeStatus id="1217884AFFEAF44FFAC74A28FAD8FA97" box="[1302,1397,1314,1337]" pageId="6" pageNumber="88" type="syntype">syntypes</typeStatus>
of “
<taxonomicName id="0AAC4D6BFFEAF44FFC9A4A48FBC5FAF7" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[843,1128,1346,1369]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="88" phylum="Arthropoda" rank="species" species="astrabadensis" status="sp. nov.">
<emphasis id="FFD8EAFAFFEAF44FFC9A4A48FBC5FAF7" box="[843,1128,1346,1369]" italics="true" pageId="6" pageNumber="88">Amblyiulus astrabadensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFEAF44FFBA74A48FB14FAF7" box="[1142,1209,1346,1369]" pageId="6" pageNumber="88" rank="species">n. sp.</taxonomicNameLabel>
”;
<collectingCountry id="B5BB7678FFEAF44FFB094A48FA81FAF7" box="[1240,1324,1346,1369]" name="Iran" pageId="6" pageNumber="88">Persien</collectingCountry>
: Astrabad (nr. 28);
<date id="B9121028FFEAF44FFC4C4A68FB81FAD7" box="[925,1068,1378,1401]" pageId="6" pageNumber="88" value="1905-05-02">
<collectingDate id="A956E9C0FFEAF44FFC4C4A68FB81FAD7" box="[925,1068,1378,1401]" pageId="6" pageNumber="88" value="1905-05-02">02.May.1905</collectingDate>
</date>
, E. Filippowitsch.
</materialsCitation>
<materialsCitation id="7DC43CB5FFEAF44FFB244A68FB62FA74" ID-GBIF-Occurrence="3881331309" collectionCode="GNM" country="Iran" location="Astrabad" pageId="6" pageNumber="88" specimenCount="1" typeStatus="syntype">
–
<collectionCode id="ABBDAE2DFFEAF44FFAD84A68FAE6FAD7" box="[1289,1355,1378,1401]" collectionName="Sweden, Goeteborg, Naturhistoriska Museet" country="Sweden" httpUri="http://grbio.org/cool/m262-zrs7" name="Gothenburg Museum of Natural History (Goteborgs Naturhistoriska Museum)" pageId="6" pageNumber="88">GNM</collectionCode>
; 2 slide preparations of (supposed) female
<typeStatus id="1217884AFFEAF44FFB154A89FA8BFA34" box="[1220,1318,1411,1434]" pageId="6" pageNumber="88" type="syntype">syntypes</typeStatus>
:
<emphasis id="FFD8EAFAFFEAF44FFAE44A89FA3BFA34" box="[1333,1430,1411,1434]" italics="true" pageId="6" pageNumber="88">Amblyiul</emphasis>
. nr. 28, I: vulvae, legs, and mouthparts, and
<emphasis id="FFD8EAFAFFEAF44FFAE44AA9FA3BFA14" box="[1333,1430,1443,1466]" italics="true" pageId="6" pageNumber="88">Amblyiul</emphasis>
. nr. 28, II: mouthparts and vulvae.
</materialsCitation>
<materialsCitation id="7DC43CB5FFEAF44FFB0C4AC9FC4FF9F5" ID-GBIF-Occurrence="3881331321" collectionCode="GNM" country="Iran" location="Astrabad" pageId="6" pageNumber="88" specimenCount="1" typeStatus="syntype">
–
<collectionCode id="ABBDAE2DFFEAF44FFB284AC9FA96FA74" box="[1273,1339,1475,1498]" collectionName="Sweden, Goeteborg, Naturhistoriska Museet" country="Sweden" httpUri="http://grbio.org/cool/m262-zrs7" name="Gothenburg Museum of Natural History (Goteborgs Naturhistoriska Museum)" pageId="6" pageNumber="88">GNM</collectionCode>
; 2 slide preparations of (supposed) male
<typeStatus id="1217884AFFEAF44FFB114AE9FA8EFA54" box="[1216,1315,1507,1530]" pageId="6" pageNumber="88" type="syntype">syntypes</typeStatus>
:
<emphasis id="FFD8EAFAFFEAF44FFAE54AE9FA38FA54" box="[1332,1429,1507,1530]" italics="true" pageId="6" pageNumber="88">Amblyiul</emphasis>
. nr. 28: gonopods, mouthparts, legs, antennae, edges of pleurotergum 7, and
<emphasis id="FFD8EAFAFFEAF44FFB87492EFB1AF995" box="[1110,1207,1572,1595]" italics="true" pageId="6" pageNumber="88">Amblyiul</emphasis>
. nr. 28, II: labrum, mandible, legs.
</materialsCitation>
</paragraph>
<paragraph id="CD1336E8FFEAF44FFCEE4979FB8BF905" blockId="6.[831,1436,1651,1707]" pageId="6" pageNumber="88">
<emphasis id="FFD8EAFAFFEAF44FFCEE4979FC42F924" bold="true" box="[831,1007,1651,1674]" pageId="6" pageNumber="88">
<typeStatus id="1217884AFFEAF44FFCEE4979FCD5F924" box="[831,888,1651,1674]" pageId="6" pageNumber="88">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="B5BB7678FFEAF44FFBD64979FB9BF924" box="[1031,1078,1651,1674]" name="Iran" pageId="6" pageNumber="88">Iran</collectingCountry>
,
<collectingRegion id="0F68F80AFFEAF44FFB824979FA82F924" box="[1107,1327,1651,1674]" country="Iran" name="Golestan" pageId="6" pageNumber="88">Golestan province</collectingRegion>
, Gorgan (formerly Esterabad).
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEAF44EFCEE49C9FEF4FD76" lastPageId="7" lastPageNumber="89" pageId="6" pageNumber="88" type="discussion">
<paragraph id="CD1336E8FFEAF44FFCEE49C9FC23F7B3" blockId="6.[831,1437,1731,2077]" pageId="6" pageNumber="88">
<emphasis id="FFD8EAFAFFEAF44FFCEE49C9FBA0F974" bold="true" box="[831,1037,1731,1754]" pageId="6" pageNumber="88">Taxonomic notes:</emphasis>
Promere virtually indistinguishable from that of
<taxonomicName id="0AAC4D6BFFEAF44FFC1849E9FBCDF954" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[969,1120,1763,1786]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="88" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFEAF44FFC1849E9FBCDF954" box="[969,1120,1763,1786]" italics="true" pageId="6" pageNumber="88">S. lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFEAF44FFBB649E9FB16F954" box="[1127,1211,1763,1786]" pageId="6" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
Caudomesal lamella (
<emphasis id="FFD8EAFAFFEAF44FFC98480EFCF1F8B5" box="[841,860,1796,1819]" italics="true" pageId="6" pageNumber="88">cl</emphasis>
) directed meso-anteriad rather than completely mesad. Mesomeral process (
<emphasis id="FFD8EAFAFFEAF44FFB5D482EFB32F895" box="[1164,1183,1828,1851]" italics="true" pageId="6" pageNumber="88">m</emphasis>
) considerably shorter than solenomere (
<emphasis id="FFD8EAFAFFEAF44FFBD5484EFBA3F8F5" box="[1028,1038,1860,1883]" italics="true" pageId="6" pageNumber="88">s</emphasis>
), without strongly protruding apical corners; solenomere apically forming more shallow concavity in comparison with
<taxonomicName id="0AAC4D6BFFEAF44FFB46488FFA9EF832" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[1175,1331,1925,1948]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="88" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFEAF44FFB46488FFA9EF832" box="[1175,1331,1925,1948]" italics="true" pageId="6" pageNumber="88">S. lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFEAF44FFAEF488FFA35F832" box="[1342,1432,1925,1948]" pageId="6" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
, and with an anterior process directed mostly distad rather than bent caudad. A very fine central filiform process (
<emphasis id="FFD8EAFAFFEAF44FFC7548EFFC1EF852" box="[932,947,2021,2044]" italics="true" pageId="6" pageNumber="88">ff</emphasis>
), similar to that of
<taxonomicName id="0AAC4D6BFFEAF44FFB4948EFFA99F852" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[1176,1332,2021,2044]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="88" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFEAF44FFB4948EFFA99F852" box="[1176,1332,2021,2044]" italics="true" pageId="6" pageNumber="88">S. lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFEAF44FFAEE48EFFA35F852" box="[1343,1432,2021,2044]" pageId="6" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
, visible.
</paragraph>
<paragraph id="CD1336E8FFEBF44EFF464E0BFEF4FD76" blockId="7.[151,756,257,728]" pageId="7" pageNumber="89">
<emphasis id="FFD8EAFAFFEBF44EFF464E0BFEA5FEB6" bold="true" box="[151,264,257,280]" pageId="7" pageNumber="89">Remarks:</emphasis>
Despite the conspicuous similarities between the gonopods of
<taxonomicName id="0AAC4D6BFFEBF44EFE9E4E2BFE52FE96" authorityName="Vagalinski" authorityYear="2020" box="[335,511,289,312]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="astrabadensis">
<emphasis id="FFD8EAFAFFEBF44EFE9E4E2BFE52FE96" box="[335,511,289,312]" italics="true" pageId="7" pageNumber="89">S. astrabadensis</emphasis>
</taxonomicName>
and
<taxonomicName id="0AAC4D6BFFEBF44EFDE74E2BFD60FE96" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[566,717,289,312]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFEBF44EFDE74E2BFD60FE96" box="[566,717,289,312]" italics="true" pageId="7" pageNumber="89">S. lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFEBF44EFD074E2BFF6AFEF6" pageId="7" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
, the two species can be rather easily distinguished by overall appearance:
<taxonomicName id="0AAC4D6BFFEBF44EFE164E6BFD27FED6" authorityName="Vagalinski" authorityYear="2020" box="[455,650,353,376]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="astrabadensis">
<emphasis id="FFD8EAFAFFEBF44EFE164E6BFD27FED6" box="[455,650,353,376]" italics="true" pageId="7" pageNumber="89">S. astrabadensis</emphasis>
</taxonomicName>
is less than
<quantity id="0A549B0DFFEBF44EFF044E8BFE84FE36" box="[213,297,385,408]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="89" unit="mm" value="25.0">25 mm</quantity>
in length and entirely pallid, whereas
<taxonomicName id="0AAC4D6BFFEBF44EFF464EABFE99FE16" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[151,308,417,440]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFEBF44EFF464EABFE99FE16" box="[151,308,417,440]" italics="true" pageId="7" pageNumber="89">S. lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFEBF44EFE904EABFE37FE16" box="[321,410,417,440]" pageId="7" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
is
<quantity id="0A549B0DFFEBF44EFE144EABFD96FE16" box="[453,571,417,440]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="2.9" metricValueMin="2.8" pageId="7" pageNumber="89" unit="mm" value="28.5" valueMax="29.0" valueMin="28.0">28-29 mm</quantity>
long and has a brownish pigmentation. Likewise, the very similar gonopod conformation in
<taxonomicName id="0AAC4D6BFFEBF44EFE014EEBFDE2FE56" baseAuthorityName="Golovatch" baseAuthorityYear="1983" box="[464,591,481,504]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="persicus">
<emphasis id="FFD8EAFAFFEBF44EFE014EEBFDE2FE56" box="[464,591,481,504]" italics="true" pageId="7" pageNumber="89">S. persicus</emphasis>
</taxonomicName>
is in a way countervailed by the setose metazonal hind margins in this species; both
<taxonomicName id="0AAC4D6BFFEBF44EFEB74D2BFDB7FD96" authorityName="Vagalinski" authorityYear="2020" box="[358,538,545,568]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="astrabadensis">
<emphasis id="FFD8EAFAFFEBF44EFEB74D2BFDB7FD96" box="[358,538,545,568]" italics="true" pageId="7" pageNumber="89">S. astrabadensis</emphasis>
</taxonomicName>
and
<taxonomicName id="0AAC4D6BFFEBF44EFD894D2BFD59FD96" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[600,756,545,568]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFEBF44EFD894D2BFD59FD96" box="[600,756,545,568]" italics="true" pageId="7" pageNumber="89">S. lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFEBF44EFF464D4BFF46FDF6" box="[151,235,577,600]" pageId="7" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
lack metazonal setae. Examples of species with (almost) completely identical gonopods but otherwise obvious differences in size and colouration are not uncommon in
<taxonomicName id="0AAC4D6BFFEBF44EFEE54DABFE00FD16" authorityName="Verhoeff" authorityYear="1899" box="[308,429,673,696]" class="Diplopoda" family="Julidae" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="tribe" tribe="Pachyiulini">Pachyiulini</taxonomicName>
(e.g.
<bibRefCitation id="A93D4B19FFEBF44EFE3D4DABFD5DFD16" author="Frederiksen S. B. &amp; Petersen G. &amp; Enghoff H." box="[492,752,673,696]" pageId="7" pageNumber="89" pagination="599 - 611" refId="ref6664" refString="Frederiksen S. B., Petersen G., Enghoff H. 2012. How many species are there of Pachyiulus? A contribution to the taxonomy of Europe's largest millipedes (Diplopoda: Julida: Julidae). Journal of Natural History 46 (9 - 10): 599 - 611." type="journal article" year="2012">
Frederiksen
<emphasis id="FFD8EAFAFFEBF44EFDA34DABFD0FFD16" box="[626,674,673,696]" italics="true" pageId="7" pageNumber="89">et al</emphasis>
., 2012
</bibRefCitation>
;
<bibRefCitation id="A93D4B19FFEBF44EFF464DCBFEE3FD76" author="Golovatch S. I." box="[151,334,705,728]" pageId="7" pageNumber="89" pagination="791 - 805" refId="ref6818" refString="Golovatch S. I. 2018. The millipede subfamily Pachyiulinae in Israel, with the description of a new species (Diplopoda, Julida, Julidae). Zoologicheskii zhurnal 97 (7): 791 - 805." type="journal article" year="2018">Golovatch, 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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126
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<taxonomicName id="0AAC4D6BFFEBF44EFC444A3CFAEAFAE0" ID-CoL="84DND" authority="(Attems, 1903)" baseAuthorityName="Attems" baseAuthorityYear="1903" box="[917,1351,1334,1358]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="continentalis">
<emphasis id="FFD8EAFAFFEBF44EFC444A3CFAEAFAE0" bold="true" box="[917,1351,1334,1358]" pageId="7" pageNumber="89">
<emphasis id="FFD8EAFAFFEBF44EFC444A3CFB37FAE0" bold="true" box="[917,1178,1334,1358]" italics="true" pageId="7" pageNumber="89">Syrioiulus continentalis</emphasis>
(
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)
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<treatmentCitation id="4C0D10F9FFEBF44EFCEE4A6EFC1CFA38" author="Attems C. G." page="148" pageId="7" pageNumber="89" year="1903">
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<emphasis id="FFD8EAFAFFEBF44EFCEE4A6EFB66FAD7" box="[831,1227,1380,1401]" italics="true" pageId="7" pageNumber="89">Pachyiulus (Dolichoiulus) continentalis</emphasis>
<bibRefCitation id="A93D4B19FFEBF44EFB074A6EFC1CFA38" author="Attems C. G." pageId="7" pageNumber="89" pagination="63 - 154" refId="ref6264" refString="Attems C. G. 1903. Beitrage zur Myriopodenkunde. Zoologische Jahrbucher - Abteilung fur Systematik, Okologie und Geographie der Tiere 18 (1): 63 - 154." type="journal article" year="1903">Attems, 1903: 147- 148</bibRefCitation>
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, figs 82-84.
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<treatmentCitationGroup id="EDBC11C6FFEBF44EFCEE4A94FB32FA7E" pageId="7" pageNumber="89">
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<emphasis id="FFD8EAFAFFEBF44EFCEE4A94FB80FA1D" box="[831,1069,1438,1459]" italics="true" pageId="7" pageNumber="89">Amblyiulus continentalis</emphasis>
</taxonomicName>
. –
<treatmentCitation id="4C0D10F9FFEBF44EFB9A4A94FA8BFA1D" author="Lohmander H." box="[1099,1318,1438,1459]" page="40" pageId="7" pageNumber="89" year="1932">
<bibRefCitation id="A93D4B19FFEBF44EFB9A4A94FA8BFA1D" author="Lohmander H." box="[1099,1318,1438,1459]" pageId="7" pageNumber="89" pagination="1 - 44" refId="ref7038" refString="Lohmander H. 1932 a. Neue Diplopoden aus Persien. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles handlingar 5, Ser. B 3 (2): 1 - 44." type="journal article" year="1932">Lohmander, 1932a: 40</bibRefCitation>
</treatmentCitation>
, 41, figs 33- 35. –
<treatmentCitation id="4C0D10F9FFEBF44EFC104AB1FB36FA7E" author="Bababekova L. A." box="[961,1179,1467,1488]" page="90" pageId="7" pageNumber="89" year="1996">
<bibRefCitation id="A93D4B19FFEBF44EFC104AB1FB36FA7E" author="Bababekova L. A." box="[961,1179,1467,1488]" pageId="7" pageNumber="89" pagination="89 - 97" refId="ref6434" refString="Bababekova L. A. 1996. The subphylum of tracheates - Tracheata [pp. 89 - 97]. In: Aliev S. V., Kasyimov A. G. (eds.). Animal life of Azerbaijan. Vol. 2: The phylum of arthropods. Elm Publishers, Baku, 412 pp." type="book chapter" year="1996">Bababekova, 1996: 90</bibRefCitation>
</treatmentCitation>
.
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<treatmentCitationGroup id="EDBC11C6FFEBF44EFCEE4AD2FC4AF9A7" pageId="7" pageNumber="89">
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<emphasis id="FFD8EAFAFFEBF44EFCEE4AD2FB84FA43" box="[831,1065,1496,1517]" italics="true" pageId="7" pageNumber="89">Syrioiulus continentalis</emphasis>
</taxonomicName>
. –
<treatmentCitation id="4C0D10F9FFEBF44EFB854AD2FA8DFA43" author="Mauries J. - P." box="[1108,1312,1496,1517]" page="441" pageId="7" pageNumber="89" year="1982">
<bibRefCitation id="A93D4B19FFEBF44EFB854AD2FA8DFA43" author="Mauries J. - P." box="[1108,1312,1496,1517]" pageId="7" pageNumber="89" pagination="433 - 444" refId="ref7158" refString="Mauries J. - P. 1982. Dolichoiulus tongiorgii (Strasser), Diplopode halophile nouveau pour la faune de France. Remarques sur la classification des Pachyiulini (Myriapoda, Diplopoda, Iulida). Bulletin du Museum national d'Histoire naturelle 4 (3 - 4): 433 - 444." type="journal article" year="1982">Mauriès, 1982: 441</bibRefCitation>
</treatmentCitation>
. –
<treatmentCitation id="4C0D10F9FFEBF44EFA994AD2FC4EF9A7" author="Mauries J. - P." page="48" pageId="7" pageNumber="89" year="1984">
<bibRefCitation id="A93D4B19FFEBF44EFA994AD2FC4EF9A7" author="Mauries J. - P." pageId="7" pageNumber="89" pagination="37 - 49" refId="ref7212" refString="Mauries J. - P. 1984. Deux especes nouvelles de Diplopodes cavernicoles des Cyclades: Hyleoglomeris beroni (Glomerida) et Syrioiulus andreevi (Iulida). Biologia Gallohellenica 11: 37 - 49." type="journal article" year="1984">Mauriès, 1984: 48</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<collectingCountry id="B5BB7678FFEBF44EFC344948FBBEF9F7" box="[997,1043,1602,1625]" name="Iran" pageId="7" pageNumber="89">Iran</collectingCountry>
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<emphasis id="FFD8EAFAFFEBF44EFC4649CAFBC8F976" bold="true" box="[919,1125,1728,1752]" italics="true" pageId="7" pageNumber="89">Syrioiulus discolor</emphasis>
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, figs 27-29.
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. –
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?
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<emphasis id="FFD8EAFAFFEBF44EFCEE4D6BFB80FDD8" box="[831,1069,609,630]" italics="true" pageId="7" pageNumber="89">Amblyiulus cappadocius</emphasis>
<bibRefCitation id="A93D4B19FFEBF44EFBE74D6BFAE9FDD8" author="Lohmander H." box="[1078,1348,609,630]" pageId="7" pageNumber="89" pagination="126 - 150" refId="ref7132" refString="Lohmander H. 1939. Uber einige neue Diplopoden aus dem ostlichen Anatolien. Verhandlungen der Naturforschenden Gesellschaft in Basel 50: 126 - 150." type="journal article" year="1939">Lohmander, 1939: 138-140</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 19- 22. –
<treatmentCitation id="4C0D10F9FFEBF44EFC104D77FBD2FD3C" author="Mauries J. - P." box="[961,1151,637,658]" page="441" pageId="7" pageNumber="89" year="1982">
<bibRefCitation id="A93D4B19FFEBF44EFC104D77FBD2FD3C" author="Mauries J. - P." box="[961,1151,637,658]" pageId="7" pageNumber="89" pagination="433 - 444" refId="ref7158" refString="Mauries J. - P. 1982. Dolichoiulus tongiorgii (Strasser), Diplopode halophile nouveau pour la faune de France. Remarques sur la classification des Pachyiulini (Myriapoda, Diplopoda, Iulida). Bulletin du Museum national d'Histoire naturelle 4 (3 - 4): 433 - 444." type="journal article" year="1982">Mauriès, 1982: 441</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="CD1336E8FFEBF44EFCEE4D90FA9AFD01" blockId="7.[831,1436,609,687]" box="[831,1335,666,687]" pageId="7" pageNumber="89">
<treatmentCitationGroup id="EDBC11C6FFEBF44EFCEE4D90FA9AFD01" box="[831,1335,666,687]" pageId="7" pageNumber="89">
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<emphasis id="FFD8EAFAFFEBF44EFCEE4D90FBB3FD01" box="[831,1054,666,687]" italics="true" pageId="7" pageNumber="89">Amblyiulus cappdoicus</emphasis>
</taxonomicName>
(sic!). –
<treatmentCitation id="4C0D10F9FFEBF44EFBA44D90FA9EFD01" author="Enghoff H." box="[1141,1331,666,687]" page="180" pageId="7" pageNumber="89" year="2006">
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</treatmentCitation>
.
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<paragraph id="CD1336E8FFEBF44EFCEE4DE2FC0DFCB1" blockId="7.[831,1436,744,799]" pageId="7" pageNumber="89">
<emphasis id="FFD8EAFAFFEBF44EFCEE4DE2FC50FD51" bold="true" box="[831,1021,744,767]" pageId="7" pageNumber="89">Known records:</emphasis>
<materialsCitation id="7DC43CB5FFEBF44EFBD84DE2FC0DFCB1" ID-GBIF-Occurrence="3881331318" country="Turkey" location="Kegubat near Kayseri" pageId="7" pageNumber="89" specimenCount="1">
<collectingCountry id="B5BB7678FFEBF44EFBD84DE2FBFAFD51" box="[1033,1111,744,767]" name="Turkey" pageId="7" pageNumber="89">Turkey</collectingCountry>
,
<location id="C8736033FFEBF44EFBB74DE2FAF4FD51" LSID="urn:lsid:plazi:treatment:450587FEFFEBF44EFC9E4D19FC3BFB40:C8736033FFEBF44EFBB74DE2FAF4FD51" box="[1126,1369,744,767]" country="Turkey" name="Kegubat near Kayseri" pageId="7" pageNumber="89">Keğubat near Kayseri</location>
(type locality).
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<subSubSection id="85B66563FFEBF44EFCEE4C3DFC3BFB40" pageId="7" pageNumber="89" type="discussion">
<paragraph id="CD1336E8FFEBF44EFCEE4C3DFC3BFB40" blockId="7.[831,1437,823,1262]" pageId="7" pageNumber="89">
<emphasis id="FFD8EAFAFFEBF44EFCEE4C3DFCD3FCE0" bold="true" box="[831,894,823,846]" pageId="7" pageNumber="89">Note:</emphasis>
The solenomere in this species ends up with a broad and blunt tip, while it lacks a rod-like mesal process, and the anterior lamella does not form an apical protrusion. This combination of characters does not correspond to the latest concept of
<taxonomicName id="0AAC4D6BFFEBF44EFAF24CBDFA30FC60" box="[1315,1437,951,974]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEBF44EFAF24CBDFA30FC60" box="[1315,1437,951,974]" italics="true" pageId="7" pageNumber="89">Amblyiulus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="A93D4B19FFEBF44EFCA74CDDFB83FC40" author="Golovatch S. I." box="[886,1070,983,1006]" pageId="7" pageNumber="89" pagination="791 - 805" refId="ref6818" refString="Golovatch S. I. 2018. The millipede subfamily Pachyiulinae in Israel, with the description of a new species (Diplopoda, Julida, Julidae). Zoologicheskii zhurnal 97 (7): 791 - 805." type="journal article" year="2018">Golovatch, 2018</bibRefCitation>
) but it fits the current diagnosis of
<taxonomicName id="0AAC4D6BFFEBF44EFCB34CFDFC62FBA0" authorityName="Verhoeff" authorityYear="1914" box="[866,975,1015,1038]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEBF44EFCB34CFDFC62FBA0" box="[866,975,1015,1038]" italics="true" pageId="7" pageNumber="89">Syrioiulus</emphasis>
</taxonomicName>
. Thus the species is here transferred to
<taxonomicName id="0AAC4D6BFFEBF44EFCEE4B1DFC01FB80" authorityName="Verhoeff" authorityYear="1914" box="[831,940,1047,1070]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEBF44EFCEE4B1DFC01FB80" box="[831,940,1047,1070]" italics="true" pageId="7" pageNumber="89">Syrioiulus</emphasis>
</taxonomicName>
. However, the pronouncedly spoon-shaped promere, with a lateral prominence and the mesomeral process tightly contiguous with the solenomere, partly concealing the latter in lateral view, are untypical of
<taxonomicName id="0AAC4D6BFFEBF44EFCEE4B9DFC01FB00" authorityName="Verhoeff" authorityYear="1914" box="[831,940,1175,1198]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEBF44EFCEE4B9DFC01FB00" box="[831,940,1175,1198]" italics="true" pageId="7" pageNumber="89">Syrioiulus</emphasis>
</taxonomicName>
, but are rather common within the genera
<taxonomicName id="0AAC4D6BFFEBF44EFCEE4BBDFC1AFB60" box="[831,951,1207,1230]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEBF44EFCEE4BBDFC1AFB60" box="[831,951,1207,1230]" italics="true" pageId="7" pageNumber="89">Amblyiulus</emphasis>
</taxonomicName>
and
<taxonomicName id="0AAC4D6BFFEBF44EFC204BBDFBD6FB60" authorityName="Verhoeff" authorityYear="1901" box="[1009,1147,1207,1230]" class="Diplopoda" family="Julidae" genus="Dolichoiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEBF44EFC204BBDFBD6FB60" box="[1009,1147,1207,1230]" italics="true" pageId="7" pageNumber="89">Dolichoiulus</emphasis>
</taxonomicName>
. Hence the question mark applied.
</paragraph>
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94
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<emphasis id="FFD8EAFAFFEBF44EFCA94E0AFB28FEB6" bold="true" box="[888,1157,256,280]" italics="true" pageId="7" pageNumber="89">Syrioiulus astrabadensis</emphasis>
(Lohmander, 1932)
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<bibRefCitation id="A93D4B19FFEBF44EFB924E25FAE9FEEA" author="Lohmander H." box="[1091,1348,303,324]" pageId="7" pageNumber="89" pagination="1 - 44" refId="ref7038" refString="Lohmander H. 1932 a. Neue Diplopoden aus Persien. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles handlingar 5, Ser. B 3 (2): 1 - 44." type="journal article" year="1932">Lohmander, 1932a: 30-33</bibRefCitation>
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, figs 30- 32.
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<treatmentCitationGroup id="EDBC11C6FFEBF44EFCEE4E63FBFDFE35" pageId="7" pageNumber="89">
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<emphasis id="FFD8EAFAFFEBF44EFCEE4E63FB9DFED0" box="[831,1072,361,382]" italics="true" pageId="7" pageNumber="89">Syrioiulus astrabadensis</emphasis>
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. –
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<bibRefCitation id="A93D4B19FFEBF44EFB844E63FAA2FED0" author="Mauries J. - P." box="[1109,1295,361,382]" pageId="7" pageNumber="89" pagination="37 - 49" refId="ref7212" refString="Mauries J. - P. 1984. Deux especes nouvelles de Diplopodes cavernicoles des Cyclades: Hyleoglomeris beroni (Glomerida) et Syrioiulus andreevi (Iulida). Biologia Gallohellenica 11: 37 - 49." type="journal article" year="1984">Mauriès, 1984: 48</bibRefCitation>
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. – Enghoff &amp; Moravvej, 2005: 64.
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<emphasis id="FFD8EAFAFFEBF44EFCEE4EB9FC55FE64" bold="true" box="[831,1016,435,458]" pageId="7" pageNumber="89">Known records:</emphasis>
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,
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<taxonomicName id="0AAC4D6BFFEBF44EFED7482CFD29F890" ID-CoL="542P5" authority="Mauries, 1984" authorityName="Mauries" authorityYear="1984" box="[262,644,1830,1854]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="andreevi">
<emphasis id="FFD8EAFAFFEBF44EFED7482CFD29F890" bold="true" box="[262,644,1830,1854]" pageId="7" pageNumber="89">
<emphasis id="FFD8EAFAFFEBF44EFED7482CFE70F890" bold="true" box="[262,477,1830,1854]" italics="true" pageId="7" pageNumber="89">Syrioiulus andreevi</emphasis>
<bibRefCitation id="A93D4B19FFEBF44EFE32482DFD29F890" author="Mauries J. - P." box="[483,644,1831,1854]" pageId="7" pageNumber="89" pagination="37 - 49" refId="ref7212" refString="Mauries J. - P. 1984. Deux especes nouvelles de Diplopodes cavernicoles des Cyclades: Hyleoglomeris beroni (Glomerida) et Syrioiulus andreevi (Iulida). Biologia Gallohellenica 11: 37 - 49." type="journal article" year="1984">Mauriès, 1984</bibRefCitation>
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<taxonomicName id="0AAC4D6BFFEBF44EFF46485EFD82F8C7" ID-CoL="542P5" authority="Mauries, 1984: 44 - 48" authorityName="Mauries" authorityPageNumber="44 - 48" authorityYear="1984" box="[151,559,1876,1897]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="andreevi">
<emphasis id="FFD8EAFAFFEBF44EFF46485EFEFEF8C7" box="[151,339,1876,1897]" italics="true" pageId="7" pageNumber="89">Syrioiulus andreevi</emphasis>
<bibRefCitation id="A93D4B19FFEBF44EFE8A485EFD82F8C7" author="Mauries J. - P." box="[347,559,1876,1897]" pageId="7" pageNumber="89" pagination="37 - 49" refId="ref7212" refString="Mauries J. - P. 1984. Deux especes nouvelles de Diplopodes cavernicoles des Cyclades: Hyleoglomeris beroni (Glomerida) et Syrioiulus andreevi (Iulida). Biologia Gallohellenica 11: 37 - 49." type="journal article" year="1984">Mauriès, 1984: 44-48</bibRefCitation>
</taxonomicName>
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, figs 12-20. –
<treatmentCitation id="4C0D10F9FFEBF44EFD6C485EFE0BF828" author="Stoev P. &amp; Beron P." page="100" pageId="7" pageNumber="89" year="2001">
<bibRefCitation id="A93D4B19FFEBF44EFD6C485EFE0BF828" author="Stoev P. &amp; Beron P." pageId="7" pageNumber="89" pagination="91 - 102" refId="ref7398" refString="Stoev P., Beron P. 2001. On type specimens of Myriapoda (Chilopoda, Diplopoda) in the collection of the National Museum of Natural History in Sofia. Arthropoda Selecta 9 (2): 91 - 102." type="journal article" year="2001">Stoev &amp; Beron, 2001: 100</bibRefCitation>
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.
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<emphasis id="FFD8EAFAFFEBF44EFF464894FEFBF81B" bold="true" box="[151,342,1950,1973]" pageId="7" pageNumber="89">Known records:</emphasis>
<collectingCountry id="B5BB7678FFEBF44EFEB04894FE1DF81B" box="[353,432,1950,1973]" name="Greece" pageId="7" pageNumber="89">Greece</collectingCountry>
,
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(type locality).
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<taxonomicName id="0AAC4D6BFFEBF44EFF4F4C5AFDC1FCC6" authority="(Lohmander, 1936)" authorityName="Vagalinski" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[158,620,848,872]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="adsharicus" status="comb. nov.">
<emphasis id="FFD8EAFAFFEBF44EFF4F4C5AFE20FCC6" bold="true" box="[158,397,848,872]" italics="true" pageId="7" pageNumber="89">Syrioiulus adsharicus</emphasis>
(
<bibRefCitation id="A93D4B19FFEBF44EFE4A4C5BFDC8FCC6" author="Lohmander H." box="[411,613,849,872]" pageId="7" pageNumber="89" pagination="1 - 196" refId="ref7101" refString="Lohmander H. 1936. Uber die Diplopoden des Kaukasusgebietes. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles handlingar (Ser. B) 5 (1): 1 - 196." type="journal article" year="1936">Lohmander, 1936</bibRefCitation>
)
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<taxonomicNameLabel id="E4EB5781FFEBF44EFDA24C5BFD43FCC6" box="[627,750,849,872]" pageId="7" pageNumber="89" rank="species">comb. nov.</taxonomicNameLabel>
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<paragraph id="CD1336E8FFEBF44EFF464C65FE13FC0F" blockId="7.[151,757,848,958]" pageId="7" pageNumber="89">
<treatmentCitationGroup id="EDBC11C6FFEBF44EFF464C65FE13FC0F" pageId="7" pageNumber="89">
<treatmentCitation id="4C0D10F9FFEBF44EFF464C65FE99FC0F" author="Lohmander H." page="159" pageId="7" pageNumber="89" year="1936">
<taxonomicName id="0AAC4D6BFFEBF44EFF464C65FE99FC0F" ID-CoL="5TZYN" authority="Lohmander, 1936: 156 - 159" authorityName="Lohmander" authorityPageNumber="156 - 159" authorityYear="1936" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="adsharicus" subGenus="Heteropachyiulus">
<emphasis id="FFD8EAFAFFEBF44EFF464C65FD95FC2A" box="[151,568,879,900]" italics="true" pageId="7" pageNumber="89">Amblyiulus (Heteropachyiulus) adsharicus</emphasis>
<bibRefCitation id="A93D4B19FFEBF44EFD904C65FE99FC0F" author="Lohmander H." pageId="7" pageNumber="89" pagination="1 - 196" refId="ref7101" refString="Lohmander H. 1936. Uber die Diplopoden des Kaukasusgebietes. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles handlingar (Ser. B) 5 (1): 1 - 196." type="journal article" year="1936">Lohmander, 1936: 156-159</bibRefCitation>
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, figs 131-132.
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<treatmentCitationGroup id="EDBC11C6FFEBF44EFF464CA3FD6CFC10" box="[151,705,937,958]" pageId="7" pageNumber="89">
<taxonomicName id="0AAC4D6BFFEBF44EFF464CA3FEDFFC10" ID-CoL="5TZYN" authorityName="Lohmander" authorityYear="1936" box="[151,370,937,958]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="adsharicus">
<emphasis id="FFD8EAFAFFEBF44EFF464CA3FEDFFC10" box="[151,370,937,958]" italics="true" pageId="7" pageNumber="89">Amblyiulus adsharicus</emphasis>
</taxonomicName>
. –
<treatmentCitation id="4C0D10F9FFEBF44EFE404CA3FD10FC10" author="Kokhia M. S. &amp; Golovatch S. I." box="[401,701,937,958]" page="40" pageId="7" pageNumber="89" year="2018">
<bibRefCitation id="A93D4B19FFEBF44EFE404CA3FD10FC10" author="Kokhia M. S. &amp; Golovatch S. I." box="[401,701,937,958]" pageId="7" pageNumber="89" pagination="35 - 48" refId="ref7005" refString="Kokhia M. S., Golovatch S. I. 2018. A checklist of the millipedes of Georgia, Caucasus (Diplopoda). ZooKeys 741: 35 - 48." type="journal article" year="2018">Kokhia &amp; Golovatch, 2018: 40</bibRefCitation>
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.
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<paragraph id="CD1336E8FFEBF44EFF464CDCFD46FB83" blockId="7.[151,756,982,1069]" pageId="7" pageNumber="89">
<emphasis id="FFD8EAFAFFEBF44EFF464CDCFEF9FC43" bold="true" box="[151,340,982,1005]" pageId="7" pageNumber="89">Known records:</emphasis>
Ajara [geographic region], region of Batumi, “Bortschcha” [probably Borçka,
<collectingCountry id="B5BB7678FFEBF44EFD8F4CFCFD06FBA3" box="[606,683,1014,1037]" name="Turkey" pageId="7" pageNumber="89">Turkey</collectingCountry>
, about
<quantity id="0A549B0DFFEBF44EFF464B1CFF76FB83" box="[151,219,1046,1069]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="89" unit="km" value="15.0">15 km</quantity>
south of the border with
<collectingCountry id="B5BB7678FFEBF44EFE394B1CFDEDFB83" box="[488,576,1046,1069]" name="Georgia" pageId="7" pageNumber="89">Georgia</collectingCountry>
] (
<typeStatus id="1217884AFFEBF44EFD854B1CFD2EFB83" box="[596,643,1046,1069]" pageId="7" pageNumber="89">type</typeStatus>
locality).
</paragraph>
<paragraph id="CD1336E8FFEBF44EFF464B4FFDF4FB72" blockId="7.[151,756,1093,1244]" pageId="7" pageNumber="89">
<emphasis id="FFD8EAFAFFEBF44EFF464B4FFF7BFBF2" bold="true" box="[151,214,1093,1116]" pageId="7" pageNumber="89">Note:</emphasis>
The original gonopod drawings clearly show an apically concave solenomere which lacks both a rodlike mesal process and an anterior lamella, or the latter is very weakly pronounced. Thus, the species is here transferred from
<taxonomicName id="0AAC4D6BFFEBF44EFE9D4BCFFE69FB72" authorityName="Silvestri" authorityYear="1896" box="[332,452,1221,1244]" class="Diplopoda" family="Julidae" genus="Amblyiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEBF44EFE9D4BCFFE69FB72" box="[332,452,1221,1244]" italics="true" pageId="7" pageNumber="89">Amblyiulus</emphasis>
</taxonomicName>
to
<taxonomicName id="0AAC4D6BFFEBF44EFE374BCFFDFFFB72" authorityName="Verhoeff" authorityYear="1914" box="[486,594,1221,1244]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEBF44EFE374BCFFDFFFB72" box="[486,594,1221,1244]" italics="true" pageId="7" pageNumber="89">Syrioiulus</emphasis>
</taxonomicName>
.
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113
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Genus
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<bibRefCitation id="A93D4B19FFEEF44BFE074A8AFDD6FA39" author="Verhoeff K. W." box="[470,635,1408,1431]" pageId="2" pageNumber="84" pagination="61 - 75" refId="ref7617" refString="Verhoeff K. W. 1914. Einige Chilognathen aus Palastina (Uber Diplopoden, 68. Aufsatz). Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien 64: 61 - 75." type="journal article" year="1914">Verhoeff, 1914</bibRefCitation>
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<bibRefCitation id="A93D4B19FFEEF44BFED24AA7FE6DFA6C" author="Verhoeff K. W." box="[259,448,1453,1474]" pageId="2" pageNumber="84" pagination="61 - 75" refId="ref7617" refString="Verhoeff K. W. 1914. Einige Chilognathen aus Palastina (Uber Diplopoden, 68. Aufsatz). Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien 64: 61 - 75." type="journal article" year="1914">Verhoeff, 1914: 65</bibRefCitation>
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(established as a subgenus of
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).
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<typeStatus id="1217884AFFEEF44BFF464AF2FF62F9A1" box="[151,207,1528,1551]" pageId="2" pageNumber="84">Type</typeStatus>
species:
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<emphasis id="FFD8EAFAFFEEF44BFEE64AF2FD99F9A1" box="[311,564,1528,1551]" italics="true" pageId="2" pageNumber="84">Dolichoiulus polyzonus</emphasis>
<bibRefCitation id="A93D4B19FFEEF44BFDEB4AF2FD60F9A1" author="Attems C." box="[570,717,1528,1551]" pageId="2" pageNumber="84" pagination="61 - 67" refId="ref6296" refString="Attems C. 1911. Description de Myriapodes nouveaux recueillis par M. Henri Gadeau de Kerville pendant son voyage zoologique en Syrie. Bulletin de la Societe des Amis des Sciences naturelles de Rouen 46 (5): 61 - 67." type="journal article" year="1911">Attems, 1911</bibRefCitation>
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, by subsequent designation by
<bibRefCitation id="A93D4B19FFEEF44BFE664912FDE6F981" author="Jeekel C. A. W." box="[439,587,1560,1583]" pageId="2" pageNumber="84" pagination="1 - 412" refId="ref6950" refString="Jeekel C. A. W. 1970. Nomenclator generum et familiarum Diplopodorum: A list of the genus and family-group names in the class Diplopoda from the 10 th edition of Linnaeus, 1758, to the end of 1957. Monografieen van de Nederlandse Entomologische Vereniging 5: 1 - 412." type="journal article" year="1970">Jeekel (1970)</bibRefCitation>
.
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<emphasis id="FFD8EAFAFFEEF44BFF46494DFE2DF9F0" bold="true" box="[151,384,1607,1630]" pageId="2" pageNumber="84">Tentative diagnosis:</emphasis>
A genus of the tribe
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characterized by a (more or less distinctly) spoonshaped promere with two apical denticles, and with an opisthomere deeply divided into a mesomeral and a solenomeral branch, the latter lacking fovea or pseudofovea and pseudoflagellum. Different from the similar genera
<taxonomicName id="0AAC4D6BFFEEF44BFE6B480DFDF3F8B0" ID-CoL="7NN97" authorityName="Schubart" authorityYear="1960" box="[442,606,1799,1822]" class="Diplopoda" family="Julidae" genus="Afropachyiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEEF44BFE6B480DFDF3F8B0" box="[442,606,1799,1822]" italics="true" pageId="2" pageNumber="84">Afropachyiulus</emphasis>
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,
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<emphasis id="FFD8EAFAFFEEF44BFDA5480DFD40F8B0" box="[628,749,1799,1822]" italics="true" pageId="2" pageNumber="84">Amblyiulus</emphasis>
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,
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<emphasis id="FFD8EAFAFFEEF44BFF46482DFE8FF890" box="[151,290,1831,1854]" italics="true" pageId="2" pageNumber="84">Dolichoiulus</emphasis>
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, and
<taxonomicName id="0AAC4D6BFFEEF44BFEBA482DFE42F890" ID-CoL="f8c3126f-a5f1-4311-a36b-4bac663ec7e8" authorityName="Verhoeff" authorityYear="1937" box="[363,495,1831,1854]" class="Diplopoda" family="Julidae" genus="Japanoiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEEF44BFEBA482DFE42F890" box="[363,495,1831,1854]" italics="true" pageId="2" pageNumber="84">Japanoiulus</emphasis>
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mostly by a typically (but not always) weakly pronounced anterior lamella of the solenomere, apically never drawn into a distinct process, and by the solenomere apex being blunt or concave, sometimes bearing minute apical processes or/ and ciliae, but not tapering into a single, pointed tip.
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<subSubSection id="85B66563FFEEF44BFBEA4E2BFB0DFE96" box="[1083,1184,289,312]" pageId="2" pageNumber="84" type="description">
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<figureCitation id="55972A6DFFEEF44BFBEA4E2BFB0DFE96" box="[1083,1184,289,312]" captionStart-0="Figs 1-4" captionStart-1="Figs 5-10" captionStart-2="Figs 11-14" captionStart-3="Figs 15-16" captionStartId-0="3.[151,192,1996,2017]" captionStartId-1="4.[151,192,1539,1560]" captionStartId-2="5.[151,192,1052,1073]" captionStartId-3="5.[151,192,2028,2049]" captionTargetBox-0="[155,1436,615,1953]" captionTargetBox-1="[198,1386,263,1488]" captionTargetBox-2="[179,1412,258,1005]" captionTargetBox-3="[151,1436,1266,1989]" captionTargetId-0="figure-159@3.[153,1438,615,1953]" captionTargetId-1="figure-212@4.[515,880,803,1077]" captionTargetId-2="figure-18@5.[518,886,510,772]" captionTargetId-3="figure-137@5.[151,1436,1266,1989]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="5" captionText-0="Figs 1-4. Syrioiulus lohmanderi sp. nov., overall appearance of male holotype (1, 2, 4) and female paratype (3), in MHNG. (1) Head and body rings 1-5. (2) Mid-body rings. (3) Head and collum. (4) Telson. Photos not to scale." captionText-1="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." captionText-2="Figs 11-14. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (11) Left promere, caudal view. (12) Right pair of gonopods, mesal view. (13) Left opisthomere, caudal, slightly lateral view. (14) Distal parts of left opisthomere, lateral view.Abbreviations: cl = caudomesal lamella, ff = central filiform process, fp = fan-like process; m = mesomeral process; P = promere; s = solenomere; sp = shield-like process. Scale bar of Figs 11-13: 0.1 mm; Figs 14-15: 0.05 mm; Fig. 16: 0.01 mm." captionText-3="Figs 15-16. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (15) Distal parts of left opisthomere, caudo-lateral view. (16) Solenomere, same aspect." figureDoi-0="http://doi.org/10.5281/zenodo.5743573" figureDoi-1="http://doi.org/10.5281/zenodo.5743577" figureDoi-2="http://doi.org/10.5281/zenodo.5743581" figureDoi-3="http://doi.org/10.5281/zenodo.5743585" httpUri-0="https://zenodo.org/record/5743573/files/figure.png" httpUri-1="https://zenodo.org/record/5743577/files/figure.png" httpUri-2="https://zenodo.org/record/5743581/files/figure.png" httpUri-3="https://zenodo.org/record/5743585/files/figure.png" pageId="2" pageNumber="84">Figs 1-16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEEF44BFCEE4E6AFB0FFD79" pageId="2" pageNumber="84" type="materials_examined">
<paragraph id="CD1336E8FFEEF44BFCEE4E6AFB0FFD79" blockId="2.[831,1437,352,727]" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4E6AFC00FED9" bold="true" box="[831,941,352,375]" pageId="2" pageNumber="84">Material:</emphasis>
<materialsCitation id="7DC43CB5FFEEF44BFC6F4E6AFC58FDD9" ID-GBIF-Occurrence="3881331305" collectingDate="1973-07-06" collectionCode="MNHG" collectorName="A. Senglet." country="Iran" isTailEnumeration="true" location="Gilan" pageId="2" pageNumber="84" specimenCount="1" specimenCount-adult="1" specimenCount-juvenile="1" stateProvince="Gilan" typeStatus="holotype">
<collectionCode id="ABBDAE2DFFEEF44BFC6F4E6AFBB9FED9" box="[958,1044,352,375]" collectionName="MNHG" pageId="2" pageNumber="84">MNHG</collectionCode>
;
<specimenCount id="DBAAFD61FFEEF44BFBF94E6AFBF1FED9" box="[1064,1116,352,375]" pageId="2" pageNumber="84" type="male">male</specimenCount>
<typeStatus id="1217884AFFEEF44BFBBF4E6AFB60FED9" box="[1134,1229,352,375]" pageId="2" pageNumber="84" type="holotype">holotype</typeStatus>
(fragmented into head, rings 1-6, left part of ring 7 and rest of body; gonopods, left antenna, leg-pair 1, leg-pair 2, penis, right leg 3, right edge of ring 7, and left mid-body leg dissected; left opisthomere prepared for
<collectionCode id="ABBDAE2DFFEEF44BFAD14EEAFA96FE59" box="[1280,1339,480,503]" pageId="2" pageNumber="84">SEM</collectionCode>
), sample field number 7314; label “
<collectingCountry id="B5BB7678FFEEF44BFBB04D0AFB23FDB9" box="[1121,1166,512,535]" name="Iran" pageId="2" pageNumber="84">Iran</collectingCountry>
(
<collectingRegion id="0F68F80AFFEEF44BFB4E4D0AFB72FDB9" box="[1183,1247,512,535]" country="Iran" name="Gilan" pageId="2" pageNumber="84">Gilan</collectingRegion>
), Siahkal, tamis., mousse, creux gros arbres” [Iran, Gilan Province, Siahkal, sifting, moss, old, hollow trees];
<date id="B9121028FFEEF44BFADB4D4AFA35FDF9" box="[1290,1432,576,599]" pageId="2" pageNumber="84" value="1973-07-06">
<collectingDate id="A956E9C0FFEEF44BFADB4D4AFA35FDF9" box="[1290,1432,576,599]" pageId="2" pageNumber="84" value="1973-07-06">06.JUL.1973</collectingDate>
</date>
; leg.
<collectorName id="6059533EFFEEF44BFCA54D6AFC58FDD9" box="[884,1013,608,631]" pageId="2" pageNumber="84">A. Senglet.</collectorName>
</materialsCitation>
<materialsCitation id="7DC43CB5FFEEF44BFBD24D6AFC59FD19" ID-GBIF-Occurrence="3881331302" collectionCode="MNHG" collectorName="A. Senglet." country="Iran" isTailEnumeration="true" location="Gilan" pageId="2" pageNumber="84" specimenCount-adult="1" specimenCount-juvenile="1" stateProvince="Gilan" typeStatus="paratype">
–
<collectionCode id="ABBDAE2DFFEEF44BFBCE4D6AFBD8FDD9" box="[1055,1141,608,631]" collectionName="MNHG" pageId="2" pageNumber="84">MNHG</collectionCode>
;
<specimenCount id="DBAAFD61FFEEF44BFB564D6AFB74FDD9" box="[1159,1241,608,631]" pageId="2" pageNumber="84" type="adult">1 adult</specimenCount>
<specimenCount id="DBAAFD61FFEEF44BFB394D6AFA9CFDD9" box="[1256,1329,608,631]" pageId="2" pageNumber="84" type="female">female</specimenCount>
<typeStatus id="1217884AFFEEF44BFAEE4D6AFA31FDD9" box="[1343,1436,608,631]" pageId="2" pageNumber="84" type="paratype">paratype</typeStatus>
(fragmented into head to ring 3 and rest of body; right vulva dissected);
</materialsCitation>
<materialsCitation id="7DC43CB5FFEEF44BFC2A4DAAFB33FD79" ID-GBIF-Occurrence="3881331311" collectingDate="1973-07-06" collectionCode="MNHG" collectorName="A. Senglet." country="Iran" isTailEnumeration="true" location="Gilan" pageId="2" pageNumber="84" specimenCount-juvenile="1" stateProvince="Gilan" typeStatus="paratype">
<specimenCount id="DBAAFD61FFEEF44BFC2A4DAAFBC8FD19" box="[1019,1125,672,695]" pageId="2" pageNumber="84" type="juvenile">1 juvenile</specimenCount>
female
<typeStatus id="1217884AFFEEF44BFB6D4DAAFAB5FD19" box="[1212,1304,672,695]" pageId="2" pageNumber="84" type="paratype">paratype</typeStatus>
(unbroken); same collecting data as
<typeStatus id="1217884AFFEEF44BFBEC4DCAFB33FD79" box="[1085,1182,704,727]" pageId="2" pageNumber="84" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEEF44BFCEE4DE5FAB3FCA8" box="[831,1310,751,774]" pageId="2" pageNumber="84" type="type_taxon">
<paragraph id="CD1336E8FFEEF44BFCEE4DE5FAB3FCA8" blockId="2.[831,1310,751,774]" box="[831,1310,751,774]" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4DE5FC77FCA8" bold="true" box="[831,986,751,774]" pageId="2" pageNumber="84">
<typeStatus id="1217884AFFEEF44BFCEE4DE5FCDAFCA8" box="[831,887,751,774]" pageId="2" pageNumber="84">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="B5BB7678FFEEF44BFC314DE5FBA3FCA8" box="[992,1038,751,774]" name="Iran" pageId="2" pageNumber="84">Iran</collectingCountry>
,
<collectingRegion id="0F68F80AFFEEF44BFBC84DE5FB10FCA8" box="[1049,1213,751,774]" country="Iran" name="Gilan" pageId="2" pageNumber="84">Gilan Province</collectingRegion>
, Siahkal.
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEEF44BFCEE4C14FAD3FCDB" pageId="2" pageNumber="84" type="etymology">
<paragraph id="CD1336E8FFEEF44BFCEE4C14FAD3FCDB" blockId="2.[831,1436,798,885]" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4C14FC68FC9B" bold="true" box="[831,965,798,821]" pageId="2" pageNumber="84">Etymology:</emphasis>
Named in memory of Hans Lohmander (1896-1961), a prominent millipede taxonomist who described six species currently assigned to
<taxonomicName id="0AAC4D6BFFEEF44BFADA4C54FADAFCDB" authorityName="Verhoeff" authorityYear="1914" box="[1291,1399,862,885]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEEF44BFADA4C54FADAFCDB" box="[1291,1399,862,885]" italics="true" pageId="2" pageNumber="84">Syrioiulus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEEF44BFCEE4C87FAA9FB6A" pageId="2" pageNumber="84" type="diagnosis">
<paragraph id="CD1336E8FFEEF44BFCEE4C87FAA9FB6A" blockId="2.[831,1437,909,1220]" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4C87FC1AFC0A" bold="true" box="[831,951,909,932]" pageId="2" pageNumber="84">Diagnosis:</emphasis>
A blind species of
<taxonomicName id="0AAC4D6BFFEEF44BFB714C87FAA0FC0A" authorityName="Verhoeff" authorityYear="1914" box="[1184,1293,909,932]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEEF44BFB714C87FAA0FC0A" box="[1184,1293,909,932]" italics="true" pageId="2" pageNumber="84">Syrioiulus</emphasis>
</taxonomicName>
, particularly similar to
<taxonomicName id="0AAC4D6BFFEEF44BFC6D4CA7FACAFC6A" authority="(Lohmander, 1932 a)" baseAuthorityName="Lohmander" baseAuthorityYear="1932" box="[956,1383,941,964]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="astrabadensis">
<emphasis id="FFD8EAFAFFEEF44BFC6D4CA7FBD8FC6A" box="[956,1141,941,964]" italics="true" pageId="2" pageNumber="84">S. astrabadensis</emphasis>
(
<bibRefCitation id="A93D4B19FFEEF44BFB5B4CA7FAF2FC6A" author="Lohmander H." box="[1162,1375,941,964]" pageId="2" pageNumber="84" pagination="1 - 44" refId="ref7038" refString="Lohmander H. 1932 a. Neue Diplopoden aus Persien. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles handlingar 5, Ser. B 3 (2): 1 - 44." type="journal article" year="1932">Lohmander, 1932a</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="0AAC4D6BFFEEF44BFCEE4CC7FB3EFC4A" authority="(Golovatch, 1983)" baseAuthorityName="Golovatch" baseAuthorityYear="1983" box="[831,1171,973,996]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="persicus">
<emphasis id="FFD8EAFAFFEEF44BFCEE4CC7FC16FC4A" box="[831,955,973,996]" italics="true" pageId="2" pageNumber="84">S. persicus</emphasis>
(
<bibRefCitation id="A93D4B19FFEEF44BFC1E4CC7FB26FC4A" author="Golovatch S. I." box="[975,1163,973,996]" pageId="2" pageNumber="84" pagination="157 - 169" refId="ref6753" refString="Golovatch S. I. 1983. A contribution to the millipede fauna of Iran (Diplopoda). Annalen des Naturhistorischen Museums in Wien (Serie B) 85: 157 - 169." type="journal article" year="1983">Golovatch, 1983</bibRefCitation>
)
</taxonomicName>
in gonopod and vulva structure. Different from
<taxonomicName id="0AAC4D6BFFEEF44BFB854CE7FAA8FBAA" baseAuthorityName="Lohmander" baseAuthorityYear="1932" box="[1108,1285,1005,1028]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="astrabadensis">
<emphasis id="FFD8EAFAFFEEF44BFB854CE7FAA8FBAA" box="[1108,1285,1005,1028]" italics="true" pageId="2" pageNumber="84">S. astrabadensis</emphasis>
</taxonomicName>
mostly by its larger size and darker colouration, and by mesomeral process being subequal to rather than significantly shorter than solenomere. Different from
<taxonomicName id="0AAC4D6BFFEEF44BFAD04B47FAD5FBCA" baseAuthorityName="Golovatch" baseAuthorityYear="1983" box="[1281,1400,1101,1124]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="persicus">
<emphasis id="FFD8EAFAFFEEF44BFAD04B47FAD5FBCA" box="[1281,1400,1101,1124]" italics="true" pageId="2" pageNumber="84">S. persicus</emphasis>
</taxonomicName>
by absence of metazonal setae, and by presence of typical, pointed mesal apical denticle of promere (the same being a rounded lobe in the latter species).
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEEF44FFCEE4BD6FA3DFD5B" lastPageId="6" lastPageNumber="88" pageId="2" pageNumber="84" type="description">
<paragraph id="CD1336E8FFEEF44BFCEE4BD6FC6EFB5D" blockId="2.[831,1437,1244,2067]" box="[831,963,1244,1267]" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4BD6FC6EFB5D" bold="true" box="[831,963,1244,1267]" pageId="2" pageNumber="84">Description</emphasis>
</paragraph>
<paragraph id="CD1336E8FFEEF44BFCEE4BF6FB50FA3D" blockId="2.[831,1437,1244,2067]" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4BF6FC77FABD" box="[831,986,1276,1299]" italics="true" pageId="2" pageNumber="84">Measurements</emphasis>
:
<typeStatus id="1217884AFFEEF44BFC244BF6FBF5FABD" box="[1013,1112,1276,1299]" pageId="2" pageNumber="84" type="holotype">Holotype</typeStatus>
with 60+1+T body rings, length
<quantity id="0A549B0DFFEEF44BFC5E4A16FC4EFA9D" box="[911,995,1308,1331]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="84" unit="mm" value="29.0">29 mm</quantity>
, mid-body vertical diameter
<quantity id="0A549B0DFFEEF44BFAE14A16FA34FA9D" box="[1328,1433,1308,1331]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="2" pageNumber="84" unit="mm" value="1.65">1.65 mm</quantity>
. Adult female
<typeStatus id="1217884AFFEEF44BFC094A36FB99FAFD" box="[984,1076,1340,1363]" pageId="2" pageNumber="84" type="paratype">paratype</typeStatus>
with 59+1+T body rings, length
<quantity id="0A549B0DFFEEF44BFCEE4A56FC3FFADD" box="[831,914,1372,1395]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="84" unit="mm" value="28.0">28 mm</quantity>
, mid-body vertical diameter
<quantity id="0A549B0DFFEEF44BFB0B4A56FA99FADD" box="[1242,1332,1372,1395]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="84" unit="mm" value="1.9">1.9 mm</quantity>
. Juvenile female
<typeStatus id="1217884AFFEEF44BFC5E4A76FC46FA3D" box="[911,1003,1404,1427]" pageId="2" pageNumber="84" type="paratype">paratype</typeStatus>
with 49+4+T body rings.
</paragraph>
<paragraph id="CD1336E8FFEEF44BFCEE4A96FB6EF9DD" blockId="2.[831,1437,1244,2067]" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4A96FC6CFA1D" box="[831,961,1436,1459]" italics="true" pageId="2" pageNumber="84">Colouration</emphasis>
(after more than 40 years of ethanol preservation) (
<figureCitation id="55972A6DFFEEF44BFC0D4AB6FB97FA7D" box="[988,1082,1468,1491]" captionStart="Figs 1-4" captionStartId="3.[151,192,1996,2017]" captionTargetBox="[155,1436,615,1953]" captionTargetId="figure-159@3.[153,1438,615,1953]" captionTargetPageId="3" captionText="Figs 1-4. Syrioiulus lohmanderi sp. nov., overall appearance of male holotype (1, 2, 4) and female paratype (3), in MHNG. (1) Head and body rings 1-5. (2) Mid-body rings. (3) Head and collum. (4) Telson. Photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743573" httpUri="https://zenodo.org/record/5743573/files/figure.png" pageId="2" pageNumber="84">Figs 1-4</figureCitation>
): Mostly ochre brown, prozonae greyish with purple tinges; head with a dark, blurred, oval spot mesally to each antenna, and with a dark, narrow, axial line on vertex; body without axial line; legs and antennae light yellow to completely pallid; pre-anal ring and paraprocts darker ventrally.
</paragraph>
<paragraph id="CD1336E8FFEEF44AFCEE4976FE17FE97" blockId="2.[831,1437,1244,2067]" lastBlockId="3.[151,756,258,537]" lastPageId="3" lastPageNumber="85" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4976FB8EF93D" box="[831,1059,1660,1683]" italics="true" pageId="2" pageNumber="84">General morphology</emphasis>
(
<figureCitation id="55972A6DFFEEF44BFBE44976FB38F93D" box="[1077,1173,1660,1683]" captionStart="Figs 1-4" captionStartId="3.[151,192,1996,2017]" captionTargetBox="[155,1436,615,1953]" captionTargetId="figure-159@3.[153,1438,615,1953]" captionTargetPageId="3" captionText="Figs 1-4. Syrioiulus lohmanderi sp. nov., overall appearance of male holotype (1, 2, 4) and female paratype (3), in MHNG. (1) Head and body rings 1-5. (2) Mid-body rings. (3) Head and collum. (4) Telson. Photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743573" httpUri="https://zenodo.org/record/5743573/files/figure.png" pageId="2" pageNumber="84">Figs 1-4</figureCitation>
,
<figureCitation id="55972A6DFFEEF44BFB744976FB18F93D" box="[1189,1205,1660,1683]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="2" pageNumber="84">5</figureCitation>
): Ommatidia absent; vertigial setae absent; 5 supralabral setae in the adult specimens, and
<quantity id="0A549B0DFFEEF44BFC2749B6FB8BF97D" box="[1014,1062,1724,1747]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="2" pageNumber="84" unit="in" value="4.0">4 in</quantity>
the juvenile; 20-26 labral setae. Antennae (
<figureCitation id="55972A6DFFEEF44BFC6849D6FC53F95D" box="[953,1022,1756,1779]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="2" pageNumber="84">Fig. 5</figureCitation>
) about 1.2 times as long as head in both sexes; antennomeres 3, 4, and 5 subequal in length, about 0.8 times as long as 2, and 1.5 times as long as 6, 5 considerably thicker than 6; a whorl of sensilla basiconica at distal margins of antennomeres 5 and 6, those on 5 nearly as long as the four apical cones, somewhat longer than those on 6. Gnathochilarium of normal julid appearance; promentum relatively short: considerably less than half the length of lingual lamellae; each lingual lamella bearing 4-5 setae. Collum smooth, some specimens with one to several very shallow grooves near posterolateral corners.
</paragraph>
<paragraph id="CD1336E8FFEFF44AFF464E48FA2CFE17" blockId="3.[151,756,258,537]" lastBlockId="3.[831,1437,258,537]" pageId="3" pageNumber="85">Prozonae with very short and shallow, scattered, longitudinal striae. Metazonae rather shallowly and somewhat irregularly striated, with 5-8 striae in a square with sides equal to metazonal length just below ozopore level; hind margins slightly turned outwards, bearing no setae. Ozopores set in metazonae, at 1/4-1/3 of metazonal length behind pro-metazonal suture; the latter indistinct, especially on dorsal side, recognizable only as a barely visible concentric groove running through middle of ring. Mid-body legs 1.55 and 0.95 times as long as midbody diameter in the adult male and in the adult female, respectively. Tarsus of mid-body legs 1.2-1.3 times as long as tibia, and about 3 times as long as apical claw.</paragraph>
<paragraph id="CD1336E8FFEFF44DFCEE4EC8FDB0F899" blockId="3.[831,1437,258,537]" lastBlockId="4.[151,757,1728,2071]" lastPageId="4" lastPageNumber="86" pageId="3" pageNumber="85">
Telson (
<figureCitation id="55972A6DFFEFF44AFC464EC8FC71FE77" box="[919,988,450,473]" captionStart="Figs 1-4" captionStartId="3.[151,192,1996,2017]" captionTargetBox="[155,1436,615,1953]" captionTargetId="figure-159@3.[153,1438,615,1953]" captionTargetPageId="3" captionText="Figs 1-4. Syrioiulus lohmanderi sp. nov., overall appearance of male holotype (1, 2, 4) and female paratype (3), in MHNG. (1) Head and body rings 1-5. (2) Mid-body rings. (3) Head and collum. (4) Telson. Photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743573" httpUri="https://zenodo.org/record/5743573/files/figure.png" pageId="3" pageNumber="85">Fig. 4</figureCitation>
): Pre-anal ring dorsally with a few short setae; epiproct absent; hypoproct short and broadly rounded, tightly fitting under paraprocts in both sexes, with a pair of disto-median setae and a transverse row of around 10 setae proximally to the former. Paraprocts densely covered with long setae, without rows of short and stiff setae along caudal margins.
</paragraph>
<caption id="99D36660FFEFF44AFF4648C6FBD8F850" ID-DOI="http://doi.org/10.5281/zenodo.5743573" ID-Zenodo-Dep="5743573" httpUri="https://zenodo.org/record/5743573/files/figure.png" pageId="3" pageNumber="85" startId="3.[151,192,1996,2017]" targetBox="[155,1436,615,1953]" targetPageId="3">
<paragraph id="CD1336E8FFEFF44AFF4648C6FBD8F850" blockId="3.[151,1437,1996,2046]" pageId="3" pageNumber="85">
Figs 1-4.
<taxonomicName id="0AAC4D6BFFEFF44AFF2748C6FE60F84F" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[246,461,1996,2017]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="85" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFEFF44AFF2748C6FE60F84F" box="[246,461,1996,2017]" italics="true" pageId="3" pageNumber="85">Syrioiulus lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFEFF44AFE0248C6FD8CF84F" box="[467,545,1996,2017]" pageId="3" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
, overall appearance of male holotype (1, 2, 4) and female paratype (3), in MHNG. (1) Head and body rings 1-5. (2) Mid-body rings. (3) Head and collum. (4) Telson. Photos not to scale.
</paragraph>
</caption>
<caption id="99D36660FFE8F44DFF464909FBBAF9C0" ID-DOI="http://doi.org/10.5281/zenodo.5743577" ID-Zenodo-Dep="5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="4" pageNumber="86" startId="4.[151,192,1539,1560]" targetBox="[198,1386,263,1488]" targetPageId="4">
<paragraph id="CD1336E8FFE8F44DFF464909FBBAF9C0" blockId="4.[151,1437,1539,1646]" pageId="4" pageNumber="86">
Figs 5-10.
<taxonomicName id="0AAC4D6BFFE8F44DFED84909FE4CF9B6" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[265,481,1539,1560]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="86" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFE8F44DFED84909FE4CF9B6" box="[265,481,1539,1560]" italics="true" pageId="4" pageNumber="86">Syrioiulus lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFE8F44DFE394909FD9BF9B6" box="[488,566,1539,1560]" pageId="4" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
, structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations:
<emphasis id="FFD8EAFAFFE8F44DFD364937FD55F9FC" box="[743,760,1597,1618]" italics="true" pageId="4" pageNumber="86">ct</emphasis>
= central tube;
<emphasis id="FFD8EAFAFFE8F44DFC5A4937FC31F9FC" box="[907,924,1597,1618]" italics="true" pageId="4" pageNumber="86">lv</emphasis>
= lateral valve;
<emphasis id="FFD8EAFAFFE8F44DFBE44937FBFFF9FC" box="[1077,1106,1597,1618]" italics="true" pageId="4" pageNumber="86">ma</emphasis>
= mesal ampulla;
<emphasis id="FFD8EAFAFFE8F44DFAD14937FAB5F9FC" box="[1280,1304,1597,1618]" italics="true" pageId="4" pageNumber="86">mt</emphasis>
= mesal tube;
<emphasis id="FFD8EAFAFFE8F44DFED84953FE88F9C0" box="[265,293,1625,1646]" italics="true" pageId="4" pageNumber="86">mv</emphasis>
= mesal valve;
<emphasis id="FFD8EAFAFFE8F44DFE6C4953FE78F9C0" box="[445,469,1625,1646]" italics="true" pageId="4" pageNumber="86">op</emphasis>
= operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm.
</paragraph>
</caption>
<paragraph id="CD1336E8FFE8F44DFF46484AFA2BF839" blockId="4.[151,757,1728,2071]" lastBlockId="4.[831,1437,1728,2071]" pageId="4" pageNumber="86">
<emphasis id="FFD8EAFAFFE8F44DFF46484AFE3EF8F9" box="[151,403,1856,1879]" italics="true" pageId="4" pageNumber="86">Male sexual characters</emphasis>
: Mandibular stipites (see
<figureCitation id="55972A6DFFE8F44DFD7D484AFD47F8F9" box="[684,746,1856,1879]" captionStart="Figs 1-4" captionStartId="3.[151,192,1996,2017]" captionTargetBox="[155,1436,615,1953]" captionTargetId="figure-159@3.[153,1438,615,1953]" captionTargetPageId="3" captionText="Figs 1-4. Syrioiulus lohmanderi sp. nov., overall appearance of male holotype (1, 2, 4) and female paratype (3), in MHNG. (1) Head and body rings 1-5. (2) Mid-body rings. (3) Head and collum. (4) Telson. Photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743573" httpUri="https://zenodo.org/record/5743573/files/figure.png" pageId="4" pageNumber="86">Fig. 1</figureCitation>
) forming rather small, rounded, frontoventral lobes. Legpair 1 (
<figureCitation id="55972A6DFFE8F44DFF0E488AFEB0F839" box="[223,285,1920,1943]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="4" pageNumber="86">Fig. 6</figureCitation>
) developed as compact hooks pointing mostly mesad, with a tapering tibial outgrowth and a minute, knob-like, tarsal remnant. Leg-pair 2 (
<figureCitation id="55972A6DFFE8F44DFDEF48CAFD2CF879" box="[574,641,1984,2007]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="4" pageNumber="86">Fig. 7</figureCitation>
) with two crested adhesive pads, one each on postfemur and tibia. Leg-pair 3 (
<figureCitation id="55972A6DFFE8F44DFEC3470AFEFDF7B9" box="[274,336,2048,2071]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="4" pageNumber="86">Fig. 8</figureCitation>
) and following pairs additionally with a vestigial pad on femur; all pads gradually diminishing in size towards telson, completely disappearing in posterior third of body. Pleurotergum 7 (
<figureCitation id="55972A6DFFE8F44DFB5E480AFB7DF8B9" box="[1167,1232,1792,1815]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="4" pageNumber="86">Fig. 9</figureCitation>
) ventrally forming large, rounded lobes originating from the border between pro- and metazona, protruding mostly ventrad. Penis soft, hyaline, with deeply divided, conoidal lobes ending up in fingertip-like terminal lamellae turned slightly caudad.
</paragraph>
<paragraph id="CD1336E8FFE8F44FFCEE48AAFC7AFED6" blockId="4.[831,1437,1728,2071]" lastBlockId="6.[831,1437,258,757]" lastPageId="6" lastPageNumber="88" pageId="4" pageNumber="86">
<emphasis id="FFD8EAFAFFE8F44DFCEE48AAFC00F819" box="[831,941,1952,1975]" italics="true" pageId="4" pageNumber="86">Gonopods</emphasis>
(
<figureCitation id="55972A6DFFE8F44DFC6C48AAFB9BF819" box="[957,1078,1952,1975]" captionStart-0="Figs 11-14" captionStart-1="Figs 15-16" captionStartId-0="5.[151,192,1052,1073]" captionStartId-1="5.[151,192,2028,2049]" captionTargetBox-0="[179,1412,258,1005]" captionTargetBox-1="[151,1436,1266,1989]" captionTargetId-0="figure-18@5.[518,886,510,772]" captionTargetId-1="figure-137@5.[151,1436,1266,1989]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="Figs 11-14. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (11) Left promere, caudal view. (12) Right pair of gonopods, mesal view. (13) Left opisthomere, caudal, slightly lateral view. (14) Distal parts of left opisthomere, lateral view.Abbreviations: cl = caudomesal lamella, ff = central filiform process, fp = fan-like process; m = mesomeral process; P = promere; s = solenomere; sp = shield-like process. Scale bar of Figs 11-13: 0.1 mm; Figs 14-15: 0.05 mm; Fig. 16: 0.01 mm." captionText-1="Figs 15-16. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (15) Distal parts of left opisthomere, caudo-lateral view. (16) Solenomere, same aspect." figureDoi-0="http://doi.org/10.5281/zenodo.5743581" figureDoi-1="http://doi.org/10.5281/zenodo.5743585" httpUri-0="https://zenodo.org/record/5743581/files/figure.png" httpUri-1="https://zenodo.org/record/5743585/files/figure.png" pageId="4" pageNumber="86">Figs 11-16</figureCitation>
): Promere (
<figureCitation id="55972A6DFFE8F44DFB6448AAFAAEF819" box="[1205,1283,1952,1975]" captionStart="Figs 11-14" captionStartId="5.[151,192,1052,1073]" captionTargetBox="[179,1412,258,1005]" captionTargetId="figure-18@5.[518,886,510,772]" captionTargetPageId="5" captionText="Figs 11-14. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (11) Left promere, caudal view. (12) Right pair of gonopods, mesal view. (13) Left opisthomere, caudal, slightly lateral view. (14) Distal parts of left opisthomere, lateral view.Abbreviations: cl = caudomesal lamella, ff = central filiform process, fp = fan-like process; m = mesomeral process; P = promere; s = solenomere; sp = shield-like process. Scale bar of Figs 11-13: 0.1 mm; Figs 14-15: 0.05 mm; Fig. 16: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.5743581" httpUri="https://zenodo.org/record/5743581/files/figure.png" pageId="4" pageNumber="86">Fig. 11</figureCitation>
,
<emphasis id="FFD8EAFAFFE8F44DFAC048AAFA8CF819" box="[1297,1313,1952,1975]" italics="true" pageId="4" pageNumber="86">P</emphasis>
in
<figureCitation id="55972A6DFFE8F44DFA9748AAFA39F819" box="[1350,1428,1952,1975]" captionStart="Figs 11-14" captionStartId="5.[151,192,1052,1073]" captionTargetBox="[179,1412,258,1005]" captionTargetId="figure-18@5.[518,886,510,772]" captionTargetPageId="5" captionText="Figs 11-14. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (11) Left promere, caudal view. (12) Right pair of gonopods, mesal view. (13) Left opisthomere, caudal, slightly lateral view. (14) Distal parts of left opisthomere, lateral view.Abbreviations: cl = caudomesal lamella, ff = central filiform process, fp = fan-like process; m = mesomeral process; P = promere; s = solenomere; sp = shield-like process. Scale bar of Figs 11-13: 0.1 mm; Figs 14-15: 0.05 mm; Fig. 16: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.5743581" httpUri="https://zenodo.org/record/5743581/files/figure.png" pageId="4" pageNumber="86">Fig. 12</figureCitation>
) relatively slender and spoon-like, tapering, as high as opisthomere; width of shaft about 0.7 times that of bowl; ridge (
<emphasis id="FFD8EAFAFFE8F44DFC09470AFC4FF7B9" box="[984,994,2048,2071]" italics="true" pageId="4" pageNumber="86">r</emphasis>
) massive and rounded, without freely protruding apical part; apical denticles well developed, clearly separated, mesal one running vertical, lateral one somewhat oblique, both with blunt tips pointing caudobasad. Opisthomere (
<figureCitation id="55972A6DFFEAF44FFEA8496BFE43F9D6" box="[377,494,1633,1656]" captionStart-0="Figs 11-14" captionStart-1="Figs 15-16" captionStartId-0="5.[151,192,1052,1073]" captionStartId-1="5.[151,192,2028,2049]" captionTargetBox-0="[179,1412,258,1005]" captionTargetBox-1="[151,1436,1266,1989]" captionTargetId-0="figure-18@5.[518,886,510,772]" captionTargetId-1="figure-137@5.[151,1436,1266,1989]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="Figs 11-14. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (11) Left promere, caudal view. (12) Right pair of gonopods, mesal view. (13) Left opisthomere, caudal, slightly lateral view. (14) Distal parts of left opisthomere, lateral view.Abbreviations: cl = caudomesal lamella, ff = central filiform process, fp = fan-like process; m = mesomeral process; P = promere; s = solenomere; sp = shield-like process. Scale bar of Figs 11-13: 0.1 mm; Figs 14-15: 0.05 mm; Fig. 16: 0.01 mm." captionText-1="Figs 15-16. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (15) Distal parts of left opisthomere, caudo-lateral view. (16) Solenomere, same aspect." figureDoi-0="http://doi.org/10.5281/zenodo.5743581" figureDoi-1="http://doi.org/10.5281/zenodo.5743585" httpUri-0="https://zenodo.org/record/5743581/files/figure.png" httpUri-1="https://zenodo.org/record/5743585/files/figure.png" pageId="6" pageNumber="88">Figs 12-16</figureCitation>
) approximately halfway divided into mesomeral process (
<emphasis id="FFD8EAFAFFEAF44FFDC8498BFD81F936" box="[537,556,1665,1688]" italics="true" pageId="6" pageNumber="88">m</emphasis>
) and solenomere (
<emphasis id="FFD8EAFAFFEAF44FFF7149ABFF07F916" box="[160,170,1697,1720]" italics="true" pageId="6" pageNumber="88">s</emphasis>
), these being subequal in height, forming a broad sinus between them; mesomeral process running mostly straight, at considerable distance from solenomere, apically broadened and spade-like, with a strongly elongated mesal corner turned caudad, partly concealing solenomere, and with a nearly right-angled lateral corner bearing several minute spines/denticles; caudomesal lamella (
<emphasis id="FFD8EAFAFFEAF44FFF2F488EFEBCF835" box="[254,273,1924,1947]" italics="true" pageId="6" pageNumber="88">cl</emphasis>
) well-developed, forming a narrow ridge protruding completely mesad, thus being clearly visible in frontal and caudal views; solenomere slender, gradually bent anteriad, ending in two small processes [an anterior, fan-like (
<emphasis id="FFD8EAFAFFEAF44FFE55470CFE34F7B3" box="[388,409,2054,2077]" italics="true" pageId="6" pageNumber="88">fp</emphasis>
) and a posterior, shield-like one (
<emphasis id="FFD8EAFAFFEAF44FFC994E08FCCDFEB7" box="[840,864,258,281]" italics="true" pageId="6" pageNumber="88">sp</emphasis>
)] surrounding a deep central concavity; a minute central filiform process (
<emphasis id="FFD8EAFAFFEAF44FFB884E28FBC5FE97" box="[1113,1128,290,313]" italics="true" pageId="6" pageNumber="88">ff</emphasis>
) visible in certain aspects; anterior lamella vestigial, recognizable only by several apical fringes.
</paragraph>
<caption id="99D36660FFE9F44CFF464B16FED5FB0A" ID-DOI="http://doi.org/10.5281/zenodo.5743581" ID-Zenodo-Dep="5743581" httpUri="https://zenodo.org/record/5743581/files/figure.png" pageId="5" pageNumber="87" startId="5.[151,192,1052,1073]" targetBox="[179,1412,258,1005]" targetPageId="5">
<paragraph id="CD1336E8FFE9F44CFF464B16FED5FB0A" blockId="5.[151,1437,1052,1188]" pageId="5" pageNumber="87">
Figs 11-14.
<taxonomicName id="0AAC4D6BFFE9F44CFECD4B16FE5EFB9F" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[284,499,1052,1073]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="87" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFE9F44CFECD4B16FE5EFB9F" box="[284,499,1052,1073]" italics="true" pageId="5" pageNumber="87">Syrioiulus lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFE9F44CFE284B16FDEBFB9F" box="[505,582,1052,1073]" pageId="5" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
, gonopods of male holotype, in MHNG. (11) Left promere, caudal view. (12) Right pair of gonopods, mesal view. (13) Left opisthomere, caudal, slightly lateral view. (14) Distal parts of left opisthomere, lateral view. Abbreviations:
<emphasis id="FFD8EAFAFFE9F44CFE364B5FFE55FBC4" box="[487,504,1109,1130]" italics="true" pageId="5" pageNumber="87">cl</emphasis>
= caudomesal lamella,
<emphasis id="FFD8EAFAFFE9F44CFD084B5FFD4BFBC4" box="[729,742,1109,1130]" italics="true" pageId="5" pageNumber="87">ff</emphasis>
= central filiform process,
<emphasis id="FFD8EAFAFFE9F44CFC364B5FFC57FBC4" box="[999,1018,1109,1130]" italics="true" pageId="5" pageNumber="87">fp</emphasis>
= fan-like process;
<emphasis id="FFD8EAFAFFE9F44CFB644B5FFB6BFBC4" box="[1205,1222,1109,1130]" italics="true" pageId="5" pageNumber="87">m</emphasis>
= mesomeral process;
<emphasis id="FFD8EAFAFFE9F44CFECD4B78FE86FB29" box="[284,299,1138,1159]" italics="true" pageId="5" pageNumber="87">P</emphasis>
= promere;
<emphasis id="FFD8EAFAFFE9F44CFE734B78FE06FB29" box="[418,427,1138,1159]" italics="true" pageId="5" pageNumber="87">s</emphasis>
= solenomere;
<emphasis id="FFD8EAFAFFE9F44CFD934B78FDFAFB29" box="[578,599,1138,1159]" italics="true" pageId="5" pageNumber="87">sp</emphasis>
= shield-like process. Scale bar of Figs 11-13: 0.1 mm; Figs 14-15: 0.05 mm; Fig. 16: 0.01 mm.
</paragraph>
</caption>
<caption id="99D36660FFE9F44CFF4648E6FDD1F7B0" ID-DOI="http://doi.org/10.5281/zenodo.5743585" ID-Zenodo-Dep="5743585" httpUri="https://zenodo.org/record/5743585/files/figure.png" pageId="5" pageNumber="87" startId="5.[151,192,2028,2049]" targetBox="[151,1436,1266,1989]" targetPageId="5">
<paragraph id="CD1336E8FFE9F44CFF4648E6FDD1F7B0" blockId="5.[151,1436,2028,2078]" pageId="5" pageNumber="87">
Figs 15-16.
<taxonomicName id="0AAC4D6BFFE9F44CFECD48E6FE5CF7AF" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[284,497,2028,2049]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="87" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFE9F44CFECD48E6FE5CF7AF" box="[284,497,2028,2049]" italics="true" pageId="5" pageNumber="87">Syrioiulus lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFE9F44CFE2448E6FDECF7AF" box="[501,577,2028,2049]" pageId="5" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
, gonopods of male holotype, in MHNG. (15) Distal parts of left opisthomere, caudo-lateral view. (16) Solenomere, same aspect.
</paragraph>
</caption>
<paragraph id="CD1336E8FFEAF44FFCEE4E8BFA3DFD5B" blockId="6.[831,1437,258,757]" pageId="6" pageNumber="88">
<emphasis id="FFD8EAFAFFEAF44FFCEE4E8BFBF8FE36" box="[831,1109,385,408]" italics="true" pageId="6" pageNumber="88">Female sexual characters</emphasis>
: Leg-pair 1 considerably, legpair 2 slightly flattened antero-caudally. Vulva (
<figureCitation id="55972A6DFFEAF44FFA944EAAFA39FE19" box="[1349,1428,416,439]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="6" pageNumber="88">Fig. 10</figureCitation>
) strongly compressed in the sagittal plane; bursa mostly symmetric: lateral valve (
<emphasis id="FFD8EAFAFFEAF44FFBB14EEAFBDEFE59" box="[1120,1139,480,503]" italics="true" pageId="6" pageNumber="88">lv</emphasis>
) slightly higher and more narrow than mesal one (
<emphasis id="FFD8EAFAFFEAF44FFBB14D0AFBD2FDB9" box="[1120,1151,512,535]" italics="true" pageId="6" pageNumber="88">mv</emphasis>
); median cleft relatively broad; operculum (
<emphasis id="FFD8EAFAFFEAF44FFBC64D15FB9FFD98" box="[1047,1074,543,566]" italics="true" pageId="6" pageNumber="88">op</emphasis>
) by far exceeding bursa, apical margin with a distinct mid-lateral incision and with several bumps along mesal section; setation moderately dense, randomly spread across whole surface of vulva. Receptaculum seminis consisting of a narrow, nearly straight central tube (
<emphasis id="FFD8EAFAFFEAF44FFBFC4DB4FBEDFD7B" box="[1069,1088,702,725]" italics="true" pageId="6" pageNumber="88">ct</emphasis>
) supporting a very short mesal tube (
<emphasis id="FFD8EAFAFFEAF44FFCAC4DD4FC35FD5B" box="[893,920,734,757]" italics="true" pageId="6" pageNumber="88">mt</emphasis>
) ending in a small, ovoid, mesal ampulla (
<emphasis id="FFD8EAFAFFEAF44FFAB14DD4FA2CFD5B" box="[1376,1409,734,757]" italics="true" pageId="6" pageNumber="88">ma</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEAF44FFC914C06FBC7FBEF" pageId="6" pageNumber="88" type="discussion">
<paragraph id="CD1336E8FFEAF44FFC914C06FAC0FC6C" blockId="6.[832,1443,780,1089]" pageId="6" pageNumber="88">
<emphasis id="FFD8EAFAFFEAF44FFC914C06FC1DFC8D" bold="true" box="[832,944,780,803]" pageId="6" pageNumber="88">Remarks:</emphasis>
The
<typeStatus id="1217884AFFEAF44FFC3C4C06FBB6FC8D" box="[1005,1051,780,803]" pageId="6" pageNumber="88">type</typeStatus>
locality of the new species lies in the foothills of the
<collectingRegion id="0F68F80AFFEAF44FFBF64C26FB5BFCED" box="[1063,1270,812,835]" country="Iran" name="Alborz" pageId="6" pageNumber="88">Alborz Mountains</collectingRegion>
, the mountain range where most
<taxonomicName id="0AAC4D6BFFEAF44FFBC84C46FB2BFCCD" authorityName="Verhoeff" authorityYear="1914" box="[1049,1158,844,867]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEAF44FFBC84C46FB2BFCCD" box="[1049,1158,844,867]" italics="true" pageId="6" pageNumber="88">Syrioiulus</emphasis>
</taxonomicName>
species from
<collectingCountry id="B5BB7678FFEAF44FFAFE4C46FAF7FCCD" box="[1327,1370,844,867]" name="Iran" pageId="6" pageNumber="88">Iran</collectingCountry>
were exclusively recorded (
<figureCitation id="55972A6DFFEAF44FFBE94C66FB20FC2D" box="[1080,1165,876,899]" captionStart="Fig" captionStartId="8.[151,186,1806,1827]" captionTargetBox="[151,1435,257,1760]" captionTargetId="figure-4@8.[151,1436,257,1760]" captionTargetPageId="8" captionText="Fig. 18. Localities of Syrioiulus spp. in Iran. Ring: S. astrabadensis; cross: S. continentalis; square: S. discolor; pentagon: S. incarnatus; star: S. lohmanderi sp. nov.; filled circle: S. persicus; triangle: S. zarudnyi." figureDoi="http://doi.org/10.5281/zenodo.5743591" httpUri="https://zenodo.org/record/5743591/files/figure.png" pageId="6" pageNumber="88">Fig. 18</figureCitation>
). The only exception is
<taxonomicName id="0AAC4D6BFFEAF44FFC914C81FB05FC0C" authority="(Lohmander, 1932 a)" baseAuthorityName="Lohmander" baseAuthorityYear="1932" box="[832,1192,907,930]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="88" phylum="Arthropoda" rank="species" species="zarudnyi">
<emphasis id="FFD8EAFAFFEAF44FFC914C81FC10FC0C" box="[832,957,907,930]" italics="true" pageId="6" pageNumber="88">S. zarudnyi</emphasis>
(
<bibRefCitation id="A93D4B19FFEAF44FFC1E4C81FB0DFC0C" author="Lohmander H." box="[975,1184,907,930]" pageId="6" pageNumber="88" pagination="1 - 44" refId="ref7038" refString="Lohmander H. 1932 a. Neue Diplopoden aus Persien. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles handlingar 5, Ser. B 3 (2): 1 - 44." type="journal article" year="1932">Lohmander, 1932a</bibRefCitation>
)
</taxonomicName>
which is known also from two localities in the central part of the country.
</paragraph>
<paragraph id="CD1336E8FFEAF44FFC914CC1FBC7FBEF" blockId="6.[832,1443,780,1089]" pageId="6" pageNumber="88">
Considering the absence of ommatidia and the method (sifting dead organic matter) by which the studied specimens were collected,
<taxonomicName id="0AAC4D6BFFEAF44FFBA44B00FABEFB8F" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[1141,1299,1034,1057]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="88" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFEAF44FFBA44B00FABEFB8F" box="[1141,1299,1034,1057]" italics="true" pageId="6" pageNumber="88">S. lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFEAF44FFAF34B00FAD0FB8F" box="[1314,1405,1034,1057]" pageId="6" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
is likely an euedaphic species.
</paragraph>
</subSubSection>
</treatment>
</document>