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Subgenus
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species.
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<taxonomicName id="464C1F65FB14961BE43807D3CE82A074" authority="Risch, 1983" authorityName="Risch" authorityYear="1983" class="Amphibia" family="Salamandridae" genus="Cynops" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="orphicus">
<emphasis id="F7E527A403223DA2858A08B8C84B2EFA" italics="true">Cynops orphicus</emphasis>
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, by present designation.
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<heading id="331AA4D0DFD90CC317C04037E1F20A53" reason="title">Etymology.</heading>
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<paragraph id="F41A7F59F966FE433053CE26B6A5AAA0">
The nomen of
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is derived from Hakka, referring to its distribution in northern and eastern
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where is the settlement of Hakka people and generic nomen
<emphasis id="E32AB90C54362A4FF8C584797FBBA7BF" italics="true">Triton</emphasis>
Laurenti, 1768.
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<heading id="682A3477115A2A546369BF89B839C549" reason="title">Diagnosis.</heading>
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<paragraph id="E351736C833147733FD6ECB07AA14BB4">(1) small size; (2) gular fold absent; (3) mid-dorsal vertebral ridge continuous; (4) metacarpal and metatarsal tubercles on external side of hands and feet absent; (5) postocular orange spot absent; (6) blue lateral stripe on tail in breeding males absent; (7) irregular greyish-blue patches on dorsum and lateral tail in breeding males and females present.</paragraph>
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<paragraph id="B98A885C486A15513359EA1EA5790B19">
<heading id="9D976E63B5D67F4B7143E691FEDA8B97" reason="title">Content.</heading>
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Two species distributed in isolated regions in northern and eastern
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in southern
<collectingCountry id="63BF0E01A446F660DA71D2801D66C70B" name="China">China</collectingCountry>
, respectively (Fig.
<figureCitation id="CCD1FEAD29D15CCE81DBF9143F5B7756" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map showing the localities for congeners of genus Hypselotriton from Southeast Chinese Hilly Area. Inset map on top left zooms in eastern Guangdong Province, showing the localities for H. (Cynotriton) oolong sp. nov. and H. (Hakkatriton) orphicus: 1. Mt Fenghuang, Chaoan District, Chaozhou City (type locality of H. oolong sp. nov.); 2. Dayang Township, Jiexi County, Jieyang City (purported type locality of H. orphicus; see Remarks in the species account for locality delimitation); 3. Liangtian Township, Jiexi County (type locality of H. orphicus according to the original coordinate); 4. Mt Dabei, Jiexi County; 5. Mt Liwangzhang, Jiexi County; 6. Mt Hongtuzhang, Fengshun County, Meizhou City; 7. Mt Tongguzhang, Fengshun County; 8. Mt Shijiadong, Fengshun County; 9. Meiguang Village, Mt Lianhua, Wuhua County, Meizhou City (type locality of H. glaucus syn. nov.). The map is derived from Tianditu (www. tianditu. gov. cn)." figureDoi="10.3897/zse.100.127268.figure1" httpUri="https://binary.pensoft.net/fig/1113320">1</figureCitation>
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<taxonomicName id="BAFF7E380C82A1894056A97288394FB5" ID-CoL="6BVX4" authority="Risch, 1983" authorityName="Risch" authorityYear="1983" class="Amphibia" family="Salamandridae" genus="Cynops" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="orphicus">
<emphasis id="E290067C3F69645BE01278E59D093DF3" italics="true">Cynops orphicus</emphasis>
<bibRefCitation id="9CA5346685261E31EA9C7ED8094BCD8E" author="Risch JP" journalOrPublisher="Alytes" pagination="45-52" refId="B25" refString="Risch JP (1983) Cynops orphicus, a new salamander from Guangdong Province, South China (Amphibia, Caudata, Salamandridae). Alytes 2: 45 - 52." title="Cynops orphicus, a new salamander from Guangdong Province, South China (Amphibia, Caudata, Salamandridae)." volume="2" year="1983">Risch, 1983</bibRefCitation>
</taxonomicName>
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,
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<treatmentCitation id="2535183C89AE6EA01C4A6AE99F8F4E34">
<bibRefCitation id="4C9C8A9C4AF9F1D21AE258015E21BB1E" DOI="10.3724/SP.J.1245.2013.00116" author="Yuan" etAl="et al." firstAuthor="Yuan" issue="2" journalOrPublisher="Asian Herpetological Research" pagination="116-123" refId="B35" refString="Yuan ZY, Jiang K, Ding L, Zhang L, Che J (2013) A new newt of the genus Cynops (Caudata: Salamandridae) from Guangdong, China. Asian Herpetological Research 4 (2): 116123. https://doi.org/10.3724/SP.J.1245.2013.00116" title="A new newt of the genus Cynops (Caudata: Salamandridae) from Guangdong, China." volume="4" year="2013">Yuan et al. (2013</bibRefCitation>
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,
<treatmentCitation id="63B2A37A9BD190B23D61D80309CF3E13">
<bibRefCitation id="BEC50BFBDB6F0F0214ABFD5C09F2E404" DOI="10.24272/j.issn.2095-8137.2022.080" author="Yuan" firstAuthor="Yuan" journalOrPublisher="Zoological Research" pagination="706-718" refId="B36" refString="Yuan ZY, Wu YK, Yan F, Murphy RW, Papenfuss TJ, Wake DB, Zhang YP, Che J (2022) Comparative multi-locus assessment of modern Asian newts (Cynops, Paramesotriton, and Pachytriton: Salamandridae) in southern China suggests a shared biogeographic history. Zoological Research 43: 706718. https://doi.org/10.24272/j.issn.2095-8137.2022.080" title="Comparative multi-locus assessment of modern Asian newts (Cynops, Paramesotriton, and Pachytriton: Salamandridae) in southern China suggests a shared biogeographic history." volume="43" year="2022">2022</bibRefCitation>
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);
<treatmentCitation id="3B4A13BAD71AA41FC99D06C177F2F3A2">
<bibRefCitation id="ADF0B1FFE47CD6841EE832B0756CEEDC" DOI="10.1163/9789004285620" author="Sparreboom M" firstAuthor="Sparreboom" refId="B28" refString="Sparreboom M (2014) Salamanders of the Old World. The Salamanders of Europe, Asia and Northern Africa. Zeist, Netherlands: KNNV Publishing. https://doi.org/10.1163/9789004285620" year="2014">Sparreboom (2014)</bibRefCitation>
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;
<treatmentCitation id="5489C160AA968BCFDF59B4A175FE6C2C">
<bibRefCitation id="974CAA7050453DFD7C7ABE888ECBAA64" author="Fei L &amp; Ye CY" firstAuthor="Fei" refId="B9" refString="Fei L, Ye CY (2016) Amphibians of China (Vol. 1). Beijing, China: Science Press." year="2016">Fei and Ye (2016)</bibRefCitation>
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;
<treatmentCitation id="BCFC84EAC059391F17179EA97FC6D4BC">
<bibRefCitation id="E152DD7571629F5A7235B1DEFED79B67" DOI="10.16373/j.cnki.ahr.220059" author="Lyu ZT &amp; Qi S &amp; Zhang SY &amp; Dai KY &amp; Wang YY" journalOrPublisher="Asian Herpetological Research" pagination="41-53" refId="B20" refString="Lyu ZT, Qi S, Zhang SY, Dai KY, Wang YY (2023 a) Two new species of fire-bellied newts (Caudata, Salamandridae, Cynops) from southeastern China. Asian Herpetological Research 14: 4153. https://doi.org/10.16373/j.cnki.ahr.220059" title="Two new species of fire-bellied newts (Caudata, Salamandridae, Cynops) from southeastern China." volume="14">Lyu et al. (2023 a</bibRefCitation>
</treatmentCitation>
)
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</paragraph>
<paragraph id="0AFA19792C6E1F54ECCC2E595A401CD3">
<treatmentCitationGroup id="F10875AF1429DB22947729340E70437A">
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<taxonomicName id="A21C72B4C1537BA9439353E6DAB8322E" ID-CoL="3P3CC" baseAuthorityName="Risch" baseAuthorityYear="1983" class="Amphibia" family="Salamandridae" genus="Hypselotriton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="orphicus" subGenus="Cynotriton">
<emphasis id="E3DE48BA7FF19D2315F3918A0DB35C4E" italics="true">Hypselotriton</emphasis>
(
<emphasis id="8E8E926E0EC73E47663EDF7C3DB52961" italics="true">Cynotriton</emphasis>
)
<emphasis id="BDBFE12721DE2F1277515CAD9B82DB35" italics="true">orphicus</emphasis>
</taxonomicName>
<bibRefCitation id="D97AA1B2F5F28FC5487CF9BF74DF251E" author="Raffaëlli J" firstAuthor="Raffaelli" refId="B23" refString="Raffaëlli J (2022) Salamanders &amp; Newts of the World. Plumelec, France: Penclen Édition." year="2022">Raffaëlli (2022)</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQ3AK" type="material">
<paragraph id="19546FD00F55E3CB1A0D672D883DA2BC">
<heading id="FFD90314E71F0E80C414B2CB7A3B25FD" reason="title">Type.</heading>
</paragraph>
<paragraph id="3E9437721B23ABE02CB9561DD8D0CFB8">
<materialsCitation id="12F901B30A59B4CE075F4F9FD9E1E3CD" collectingDate="2019-09-21" collectionCode="CIB" collectorName="Wei" country="China" county="Chaozhou City" elevation="1320" latitude="23.96" location="Mt Fenghuang" longLatPrecision="752" longitude="116.64" municipality="Chao'an District" specimenCode="CIB 121430" specimenCount="1" specimenCount-male="1" stateProvince="Guangdong" typeStatus="holotype">
<emphasis id="1D4ED2B908B462DE30F502DB23D8F470" bold="true" italics="true">
<typeStatus id="EFE973F64567DEC54FBC37F7CF43DE92" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="753786FCF4146D6FAC699EFDDB93A4FD" name="China">China</collectingCountry>
<specimenCount id="CB8F673F08B585516A54DCE9C7CF8BE5" count="1" type="male"></specimenCount>
;
<collectingRegion id="8AD0B5B3F35319955A543493AF27C961" country="China" name="Guangdong">Guangdong Province</collectingRegion>
,
<collectingCounty id="A008830417BA3D75B060B590271B0F96">Chaozhou City</collectingCounty>
,
<collectingMunicipality id="B7137BEF1ECD9D23C7E03D641DABF2EF">Chaoan District</collectingMunicipality>
,
<location id="C62B2960905097D85F70C5BEA02A47C1" LSID="urn:lsid:plazi:treatment:4281FFE97F6B5473A7170C8385C3AA59:C62B2960905097D85F70C5BEA02A47C1" country="China" county="Chaozhou City" latitude="23.96" longLatPrecision="752" longitude="116.64" municipality="Chao'an District" name="Mt Fenghuang" stateProvince="Guangdong">Mt Fenghuang</location>
,
<location id="953B3A5A79D1CDB5432A8D1D51FC3928" LSID="urn:lsid:plazi:treatment:4281FFE97F6B5473A7170C8385C3AA59:953B3A5A79D1CDB5432A8D1D51FC3928" country="China" county="Chaozhou City" latitude="23.96" longLatPrecision="752" longitude="116.64" municipality="Chao'an District" name="Peak Wudongding" stateProvince="Guangdong">Peak Wudongding</location>
,
<location id="F5695571FB3B65858CE2FE02AA6A9D1D" LSID="urn:lsid:plazi:treatment:4281FFE97F6B5473A7170C8385C3AA59:F5695571FB3B65858CE2FE02AA6A9D1D" country="China" county="Chaozhou City" latitude="23.96" longLatPrecision="752" longitude="116.64" municipality="Chao'an District" name="Tianchi Lake" stateProvince="Guangdong">Tianchi Lake</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E53AK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[116.640000,23.960000]}">
<geoCoordinate id="C88275DD342EE118B1141A857F278B8F" degrees="23.96" direction="north" orientation="latitude" precision="555" value="23.96">23.96 ° N</geoCoordinate>
,
<geoCoordinate id="CDC959AAA587AF7C6BFD6E912D197E3A" degrees="116.64" direction="east" orientation="longitude" precision="555" value="116.64">116.64 ° E</geoCoordinate>
</named-content>
, ca.
<quantity id="654A9D72CB5C0AE4261BA35C1CBE0947" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" unit="m" value="1320.0">
<elevation id="DED80F107542E1C5AB1B8F04181CBFD7" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" unit="m" value="1320.0">1320 m</elevation>
</quantity>
elev.;
<date id="B8A323EC9F1EE1FA1E6F9689DAFDBD59" value="2019-09-21">
<collectingDate id="5CFE1321AB56B55F8F7E313B2E2331FF" value="2019-09-21">21 Sep. 2019</collectingDate>
</date>
; Tian-Li
<collectorName id="825D76EE82893C884863060B378EA168">Wei</collectorName>
leg.;
<specimenCode id="2233AD044689EC771C884F36ABFDDA09" collectionCode="CIB" country="China" httpUri="http://grbio.org/cool/vxkr-andy" name="Chengdu Institute of Biology" type="">
<named-content id="4E0F79BF92F4886860AD63087FF1D678" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chengdu-institute-biology-0" xlink_title="Herpetological Museum, Chengdu Institute of Biology, the Chinese Academy of Sciences, Chengdu, China">CIB</named-content>
121430
</specimenCode>
[field number
<abbrev id="ABBRID0EH4AK" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering, Foshan, China">GEP</abbrev>
a 192] (Fig.
<figureCitation id="F2B9937D1C62F848E755B08520536158" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. The holotype CIB 121430 of Hypselotriton (Cynotriton) oolong sp. nov. in life. A. Dorsal view; B. Ventral view; C. Lateral view of head; D. Dorsal view of head; E. Ventral view of hand; F. Ventral view of foot; G. Cloaca slit; H. Lateral view of tail." figureDoi="10.3897/zse.100.127268.figure4" httpUri="https://binary.pensoft.net/fig/1113323">4</figureCitation>
).
</materialsCitation>
</paragraph>
<caption id="2B0E76FE154B2AF68D5E9FED2402FEDE" ID-DOI="10.3897/zse.100.127268.figure4" ID-arpha="9CF8438F-8374-5D57-8DCA-64DCDCB10224" httpUri="https://binary.pensoft.net/fig/1113323" startId="F4">
<paragraph id="A1C8DA793FDCF6203DE23848C471A23B">
<label id="476DD697DE811E5B9768CE7F09C3CD26">Figure 4.</label>
</paragraph>
<paragraph id="E7687D0FBD203A093A476E593A1EA5C1">
The holotype
<named-content id="02024702D13B8935EBEE3095EEC8C491" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chengdu-institute-biology-0" xlink_title="Herpetological Museum, Chengdu Institute of Biology, the Chinese Academy of Sciences, Chengdu, China">CIB</named-content>
121430 of
<taxonomicName id="4276D3C05E5EF724A478F70DA293BEA6" authorityName="Wang &amp; Zeng &amp; Wei &amp; Lyu" authorityYear="2024" class="Amphibia" family="Salamandridae" genus="Hypselotriton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="oolong" status="sp. nov." subGenus="Cynotriton">
<emphasis id="6207DACC3272C58F7CEBDDC21FEE442A" italics="true">Hypselotriton</emphasis>
(
<emphasis id="16DE571101D65978C0C3BDB5CE10B276" italics="true">Cynotriton</emphasis>
)
<emphasis id="094420BB04A5C57078854FE0D789FCA1" italics="true">oolong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="146EFE148146B36A6E7C694F61DEA335" rank="species">sp. nov.</taxonomicNameLabel>
in life.
<emphasis id="CD9D6CBF70D7BCC2A2DA346A828DB232" bold="true">A.</emphasis>
Dorsal view;
<emphasis id="B035D72DF5516242C70756EB7CDC5A45" bold="true">B.</emphasis>
Ventral view;
<emphasis id="BF25042CE3BE555888A63D64E54B98CD" bold="true">C.</emphasis>
Lateral view of head;
<emphasis id="AA553C482823E69FEB4CB90A82A2B79E" bold="true">D.</emphasis>
Dorsal view of head;
<emphasis id="67D99DF610FB805A83E41215017A76D7" bold="true">E.</emphasis>
Ventral view of hand;
<emphasis id="1E2EFAB50ED3F015EBFED61F82273487" bold="true">F.</emphasis>
Ventral view of foot;
<emphasis id="1548265A65E856BE9B613BD74277F4A7" bold="true">G.</emphasis>
Cloaca slit;
<emphasis id="2241E9BFC52CDAD9531B9A97B0FCFDBA" bold="true">H.</emphasis>
Lateral view of tail.
</paragraph>
</caption>
<paragraph id="14A9EAF3092ECAE2D869C5C27D84C610">
<materialsCitation id="15ABED8E56AD34C27D4529AF03079A60" collectingDate="2019-09-21" collectionCode="CIB" collectorName="Wei" country="China" county="Chaozhou City" elevation="1320" latitude="23.96" location="Mt Fenghuang" longLatPrecision="752" longitude="116.64" municipality="Chao'an District" specimenCode="CIB 121429" specimenCount="2" specimenCount-male="2" stateProvince="Guangdong" typeStatus="paratype">
<emphasis id="292985A4A0E3B6F3FCA05FE92292D381" bold="true" italics="true">
<typeStatus id="DE0251405851866AF6B41E5299780891" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="B742969AB68E72A94BD4254339970FFD" name="China">China</collectingCountry>
<specimenCount id="AD2818CE71BE0B7957967D9F099FDE4D" count="2" type="male">2 ♂♂</specimenCount>
; same data as for holotype;
<specimenCode id="CBA5F185FBDF75088B60E54A82128997" collectionCode="CIB" country="China" httpUri="http://grbio.org/cool/vxkr-andy" name="Chengdu Institute of Biology" type="">
<named-content id="6435307672CD46224ED998284E33C1DF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chengdu-institute-biology-0" xlink_title="Herpetological Museum, Chengdu Institute of Biology, the Chinese Academy of Sciences, Chengdu, China">CIB</named-content>
121429
</specimenCode>
[field number
<abbrev id="ABBRID0ES6AK" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering, Foshan, China">GEP</abbrev>
a 191],
<named-content id="DE366BF6D7047198A12652B8F022ECC8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zhongshan-sun-yatsen-university" xlink_title="The Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYS</named-content>
a 009274 [field number
<abbrev id="ABBRID0E16AK" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering, Foshan, China">GEP</abbrev>
a 190]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E56AK" type="etymology">
<paragraph id="C7C642A5336CB74724BD7F565B863D89">
<heading id="C6FC8E43D42479967D39F69DD83773C5" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="FB89036A709DE52FCADC440DF1AAC125">
The specific name
<emphasis id="CF49B58A828416945FA00931817A333F" italics="true">oolong</emphasis>
is used as a noun in apposition, derived from oolong tea. The
<typeStatus id="B3EB65E882C26DA1B6793B2EA4C20805">type</typeStatus>
locality of this species, Mt Fenghuang, is famous for the cultivation and production of the Phoenix Oolong Tea. Yet, the developments of tea cultivation have affected and threatened the habitats of this species. We name this new species after the most famous local economic output in the hope that it would bring attention on the green and sustainable development as well as the harmonious co-existence between humanity and nature. This species name is also in memory of the Japanese manga artist Akira Toriyama (19552024). His most famous work,
<emphasis id="116B7BFA04D7B44B58F10292F8CEC2BB" italics="true">Dragon Ball</emphasis>
, was originally inspired by Chinese culture and one of the characters is named as Oolong who makes the first shown wish with the Dragon Balls.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELBBK" type="Common name">
<paragraph id="16F3B7632AB967291C7A9F41BBE3052E">
<heading id="19E7B87893CD1DB0DF3AECAD0223CF2A" reason="title">Common name.</heading>
</paragraph>
<paragraph id="B80D5EDE6DB83CC9C8C8AB630495A4F8">Oolong Fire-bellied Newt (in English) / wū lóng róng yuán (乌龙蝾螈 in Chinese).</paragraph>
</subSubSection>
<subSubSection id="SECID0EQBBK" type="diagnosis">
<paragraph id="AB303AA3BFB19A47FE2A9B3ADC118CF1">
<heading id="F2640CFF086AB5DFA4C9CA30C014452E" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="964B7194CFBF03844A6B856CF7FFBB8D">
(1) small body size,
<abbrev id="ABBRID0EWBBK" xlink_title="total length">TOL</abbrev>
<quantity id="5064027F2FD3E4E0F39FF19B2518AF41" metricMagnitude="-2" metricUnit="m" metricValue="7.03" metricValueMax="7.07" metricValueMin="6.99" unit="mm" value="70.3" valueMax="70.7" valueMin="69.9">69.970.7 mm</quantity>
in adult males; (2) parotoid gland inconspicuous; (3) postocular orange spot absent; (4) surface rough and granulated, gular fold present; (5) interrupted vertebral ridge conspicuous and bulged; (6) fingers and toes overlapping when fore-limb and hind-limb adpressed towards each other along body; (7) ground colour dark brown to olive brown with black spots; (8) lateral tail with black spots; (9) ground colour of venter dark brown with irregular bright orange patches, bright orange blotches on chin, base of ventral limbs and anterior half of cloaca; (10) ventral tail with a bright orange stripe.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1BBK" type="description">
<paragraph id="624A316B461EF486FF72DD426FA673AD">
<heading id="EBA28000C21E8AEEE10BBFB261FDA185" reason="title">
Description of the
<typeStatus id="1A64B7543FC130DF9A267C71F1B208A9" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="9A6F111EC34539F525A663B26202090D">
Body slender and small-sized,
<abbrev id="ABBRID0EACBK" xlink_title="total length">TOL</abbrev>
<quantity id="8674B45547F6EEAFB3FDD38CCAF5D36E" metricMagnitude="-2" metricUnit="m" metricValue="7.07" unit="mm" value="70.7">70.7 mm</quantity>
. Head oval in dorsal view; snout truncate, projecting slightly beyond mandible; nostril small, but conspicuous; labial fold well developed on posterior part of upper jaw; tongue elongate, enlarged anteriorly, with free lateral margin; vomerine tooth patch “ ^ ” - shaped; eye small, not extending beyond lateral margins of head; a conspicuous longitudinal ridge found posterior to each eye; parotoid gland inconspicuous, gill remnants absent; gular fold present.
</paragraph>
<paragraph id="4DFDC8BFFF7263F2B67AADFAFD5DC3D9">Surface rough, dense tapered granules and tiny spines on dorsum, flanks, limbs and tail; dense granules and inconspicuous wrinkles on venter; interrupted vertebral ridge conspicuous and bulged; cloacal opening oval, slightly protruding.</paragraph>
<paragraph id="15F629E472FB9F3EC749525BE4B3E634">
Limbs slender, fingers and toes overlapping when fore-limb and hind-limb adpressed towards each other along body; four fingers and five toes, slender and elongated, lack webbing; relative length of fingers I &lt;
<collectionCode id="1CD1862561A175BE94C960239C13983C">IV</collectionCode>
&lt;
<collectionCode id="706BBBD7AA4CEF29969152024538C221">II</collectionCode>
&lt;
<collectionCode id="26CE1C3B47E464ADC61EA74F881C2367">III</collectionCode>
; relative length of toes I &lt;
<collectionCode id="CE1450F9FD18008499E12F480B8AAE69" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
&lt;
<collectionCode id="7CCA2FABCCC12B1E2D6D37CE9A7E7E03">II</collectionCode>
&lt;
<collectionCode id="BB96C68E584A3CE1E5F1F4ACAD6F5061">IV</collectionCode>
&lt;
<collectionCode id="63DBF3A131AB27AEFEBE80E808013F01">III</collectionCode>
. Tail laterally compressed, tapers posteriorly; caudal fin distinct; tail tip bluntly pointed.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGCBK" type="Colouration of the holotype">
<paragraph id="121257B5048D18B8D15C8B927EDA268C">
<heading id="3DA021DBC5E5269688FE0AF20D66124E" reason="title">
Colouration of the
<typeStatus id="0D1796122E2D3B719A2C636792268E6B" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="0D1AC625F27964768498A297277BC042">In life, ground colour dark brown with black spots; vertebral ridge and upper margin of tail yellowish-brown; lateral tail with black spots; tips of digits light yellow; irregular bright orange stripe bordering dark patches on ventral trunk, bright orange blotches on chin, base of ventral limbs and anterior half of cloaca; ventral tail with a bright orange stripe.</paragraph>
<paragraph id="487A817AB068560839969DF2421B96BC">In preservation, ground colour faded, greyish, black spots absent; vertebral ridge, upper margin of tail and digits dark grey; black spots on lateral tail indistinct; bright orange stripe on ventral trunk and tail, bright orange blotches on chin, ventral limbs and anterior half of cloaca slightly faded, dark patches more distinct.</paragraph>
</subSubSection>
<subSubSection id="SECID0EMCBK" type="variation">
<paragraph id="7A4481DF5A526F00210D75098F4679F9">
<heading id="E7511FF1604A981897CDEE42054141A9" reason="title">Variations.</heading>
</paragraph>
<paragraph id="8888EFCF8526548BAB4FA959A78B5833">
Measurements of the
<typeStatus id="B5941FB55BBEE2CCFDDD7FB30AD9D4CF">type</typeStatus>
series are given in Table
<tableCitation id="BD4985FAAD65A292F33205DED30D789A" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Measurements (in mm) of Hypselotriton (Hakkatriton) orphicus and H. (Cynotriton) oolong sp. nov." httpUri="http://table.plazi.org/id/309576DAD536BC7FAE1130E5C8C4E033" tableUuid="309576DAD536BC7FAE1130E5C8C4E033">2</tableCitation>
. All male specimens are similar in body proportions; however, the number, shape and position of ventral orange blotches vary amongst individuals.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWCBK" type="distribution">
<paragraph id="0458460183AA06A7B4670AB7950D8305">
<heading id="2239FF52D0013DFF361F07E97DA7B859" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="AC7ECACD088344455E4A8F44296B564A">
This species is known only from Tianchi Lake and surrounding streams of Mt Fenghuang at elevations of ca.
<quantity id="E9AF8619F0CAAF4EDBA797A09D6196AF" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</quantity>
. This locality is situated on the east of the Hanjiang River in eastern
<collectingRegion id="539A273D86D049FD00A990CF547E662F" country="China" name="Guangdong">Guangdong</collectingRegion>
, while the populations of
<taxonomicName id="54666E69E9D120CA1C8505EDCDCB42EB" baseAuthorityName="Risch" baseAuthorityYear="1983" class="Amphibia" family="Salamandridae" genus="Hypselotriton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="orphicus" subGenus="Hakkatriton">
<emphasis id="455F74E964F0D89D041794DDBB494656" italics="true">H.</emphasis>
(
<emphasis id="AB0E79C690052840BF1021E6FDE3F178" italics="true">Ha.</emphasis>
)
<emphasis id="B491162CF3FE84D009D66E4A107EC0E6" italics="true">orphicus</emphasis>
</taxonomicName>
were all discovered from the west of the Hanjiang River (Fig.
<figureCitation id="EA319A8F3ECA468A060E737FC97D9B7D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map showing the localities for congeners of genus Hypselotriton from Southeast Chinese Hilly Area. Inset map on top left zooms in eastern Guangdong Province, showing the localities for H. (Cynotriton) oolong sp. nov. and H. (Hakkatriton) orphicus: 1. Mt Fenghuang, Chaoan District, Chaozhou City (type locality of H. oolong sp. nov.); 2. Dayang Township, Jiexi County, Jieyang City (purported type locality of H. orphicus; see Remarks in the species account for locality delimitation); 3. Liangtian Township, Jiexi County (type locality of H. orphicus according to the original coordinate); 4. Mt Dabei, Jiexi County; 5. Mt Liwangzhang, Jiexi County; 6. Mt Hongtuzhang, Fengshun County, Meizhou City; 7. Mt Tongguzhang, Fengshun County; 8. Mt Shijiadong, Fengshun County; 9. Meiguang Village, Mt Lianhua, Wuhua County, Meizhou City (type locality of H. glaucus syn. nov.). The map is derived from Tianditu (www. tianditu. gov. cn)." figureDoi="10.3897/zse.100.127268.figure1" httpUri="https://binary.pensoft.net/fig/1113320">1</figureCitation>
).
</paragraph>
<paragraph id="A550BD785D09297794989A0D9FF53928">
The adult individuals inhabit puddles and slow streams that are surrounded by bushes and weeds.
<bibRefCitation id="908351A8A776074AF6DABB99F5FE5A70" author="Fei L &amp; Hu SQ &amp; Ye CY &amp; Huang YZ" etAl="et al." firstAuthor="Fei" refId="B10" refString="Fei L, Hu SQ, Ye CY, Huang YZ (2006) Fauna Sinica. Amphibia. Volume 1. General Accounts of Gymnophiona and Urodela. Beijing, China: Science Press." year="2006">Fei et al. (2006)</bibRefCitation>
reported that
<specimenCount id="914223DBAC77E6F39302289B1C55E8CF" count="26" type="adult">26 adults</specimenCount>
were observed from Mt Fenghuang during a survey in
<date id="9292FA7EAE7B3BF5BB8AD2BB55CD18A3" value="2002-07">July 2002</date>
. However, only
<specimenCount id="286467B183F1F1902F053D8F36D9755C" count="9" type="adult">nine adults</specimenCount>
were observed in
<date id="30A42F3BF3611940ABBE08F99F9B48F6" value="2019-09">September 2019</date>
. The
<typeStatus id="48E236A3F4F34BB55DA195731D05C251">type</typeStatus>
locality is being threatened by the developments of tourism and tea planting, which might cause negative effects on this species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPEBK" type="remarks">
<paragraph id="39025B4ECEE4D47981861E437841603A">
<heading id="3A7E0A792B2CF096B1FFC00B046A56D0" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="1B301E284661B3E9328C2E9DAC113D62">
The
<taxonomicName id="33729FC4E86A1548B1BE1717A7933801" authorityName="Wolterstorff" authorityYear="1934" class="Amphibia" family="Salamandridae" genus="Hypselotriton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="genus">
<emphasis id="E0908CE8C736E097C0DACF2BD24C4FD4" italics="true">Hypselotriton</emphasis>
</taxonomicName>
populations from central
<collectingRegion id="CDBD9432C5831C888F77427C4AC64DDC" country="China" name="Fujian">Fujian</collectingRegion>
were originally reported as
<taxonomicName id="C64304A057C25192001E5ABDB8142963" baseAuthorityName="David" baseAuthorityYear="1873" class="Amphibia" family="Salamandridae" genus="Cynops" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="84E3073BF02C406C2B3A803F967BEE8E" italics="true">Cynops orientalis</emphasis>
</taxonomicName>
(
<bibRefCitation id="0947DED37EF5F008DE3CCE8CB109571A" author="Hu" etAl="et al." firstAuthor="Hu" journalOrPublisher="Materials of Herpetological Research" pagination="22-29" refId="B16" refString="Hu SQ, Fei L, Ye CY (1978) Investigation report of amphibians in Fujian. Materials of Herpetological Research 4: 2229." title="Investigation report of amphibians in Fujian." volume="4" year="1978">Hu et al. 1978</bibRefCitation>
;
<bibRefCitation id="190F88BE47A65E810AA3B61E0454D396" author="Risch" firstAuthor="Risch" journalOrPublisher="Alytes" pagination="45-52" refId="B25" refString="Risch JP (1983) Cynops orphicus, a new salamander from Guangdong Province, South China (Amphibia, Caudata, Salamandridae). Alytes 2: 4552." title="Cynops orphicus, a new salamander from Guangdong Province, South China (Amphibia, Caudata, Salamandridae)." volume="2" year="1983">Risch 1983</bibRefCitation>
).
<bibRefCitation id="D15A02080488B26D77D096AD665223BF" author="Fei L &amp; Hu SQ &amp; Ye CY &amp; Huang YZ" etAl="et al." firstAuthor="Fei" refId="B10" refString="Fei L, Hu SQ, Ye CY, Huang YZ (2006) Fauna Sinica. Amphibia. Volume 1. General Accounts of Gymnophiona and Urodela. Beijing, China: Science Press." year="2006">Fei et al. (2006)</bibRefCitation>
re-identified the specimens from Mt Daiyun as
<taxonomicName id="52CC510A479470C9D105B22651EB2108" baseAuthorityName="Risch" baseAuthorityYear="1983" class="Amphibia" family="Salamandridae" genus="Hypselotriton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="orphicus">
<emphasis id="DE981833A55EBFE38A15D62A2F1CBE81" italics="true">C. orphicus</emphasis>
</taxonomicName>
, based on the ventral colouration that is similar to the
<typeStatus id="CEFC8BAED358EA2E920FD2D76B4C29E6" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="D52C9660D59B34383E768A5ACA9AEC76" baseAuthorityName="Risch" baseAuthorityYear="1983" class="Amphibia" family="Salamandridae" genus="Hypselotriton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="orphicus">
<emphasis id="2A33DCD2F95521698B12AE17919522C4" italics="true">C. orphicus</emphasis>
</taxonomicName>
preserved in
<named-content id="993DE32F31EEB89628FFC88FC2BB3B12" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Museum national dHistoire naturelle, Paris, France">
<collectionCode id="B8B95A45550EF848DAEB91FB5FE6FD3F" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
. However, our morphological examinations found that such colouration could not be a distinguished character between the two species, which led to the misidentification. The phylogenetic analysis revealed close relationships between the central
<collectingRegion id="CDE91FBAF3E809664326D51F48BD23A7" country="China" name="Fujian">Fujian</collectingRegion>
populations and the new species, however, with distinct genetic divergences. Their distribution areas are also distantly isolated (Fig.
<figureCitation id="E475ADDD64C50C7B8BE9E12CF66468C1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map showing the localities for congeners of genus Hypselotriton from Southeast Chinese Hilly Area. Inset map on top left zooms in eastern Guangdong Province, showing the localities for H. (Cynotriton) oolong sp. nov. and H. (Hakkatriton) orphicus: 1. Mt Fenghuang, Chaoan District, Chaozhou City (type locality of H. oolong sp. nov.); 2. Dayang Township, Jiexi County, Jieyang City (purported type locality of H. orphicus; see Remarks in the species account for locality delimitation); 3. Liangtian Township, Jiexi County (type locality of H. orphicus according to the original coordinate); 4. Mt Dabei, Jiexi County; 5. Mt Liwangzhang, Jiexi County; 6. Mt Hongtuzhang, Fengshun County, Meizhou City; 7. Mt Tongguzhang, Fengshun County; 8. Mt Shijiadong, Fengshun County; 9. Meiguang Village, Mt Lianhua, Wuhua County, Meizhou City (type locality of H. glaucus syn. nov.). The map is derived from Tianditu (www. tianditu. gov. cn)." figureDoi="10.3897/zse.100.127268.figure1" httpUri="https://binary.pensoft.net/fig/1113320">1</figureCitation>
). As there is a lack of detailed morphological data of the
<collectingRegion id="C46B51DFA0817B9A5817FCA0543C717D" country="China" name="Fujian">Fujian</collectingRegion>
populations for accurate identification, we prudently label them as
<taxonomicName id="A937E8191BDFFE059D21DE4E7639562E" class="Amphibia" family="Salamandridae" genus="Hypselotriton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="71177986DCB60439B7B842FF7F0BF0B2" italics="true">Hypselotriton</emphasis>
sp.
</taxonomicName>
in this work provisionally.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,151 @@
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<mods:title id="1776C5F85006B6DB4517A6D578B746C2">Taxonomic determination of Hypselotriton populations distributed in eastern Guangdong, China (Caudata, Salamandridae), with description of a new species and a new subgenus</mods:title>
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<treatment id="8403D6596A3D558F9394B22E574BE00E" ID-DOI="http://doi.org/10.5281/zenodo.13322664" ID-Zenodo-Dep="13322664" ID-arpha="8403D659-6A3D-558F-9394-B22E574BE00E" LSID="urn:lsid:plazi:treatment:8403D6596A3D558F9394B22E574BE00E" httpUri="http://treatment.plazi.org/id/8403D6596A3D558F9394B22E574BE00E">
<subSubSection id="AB3FCC3FFB081AB3ECA73528089924E3" type="nomenclature">
<paragraph id="1B60ABB3F7467382F19BB0CB6A9973CB">
<emphasis id="5DAE36CE4D52BF48830FD4BA8CE5C11E" italics="true">
Subgenus
<taxonomicName id="313484D6BA62454AB655AF644C46749F" ID-CoL="52ZK" authority="Wolterstorff, 1934" authorityName="Wolterstorff" authorityYear="1934" class="Amphibia" family="Salamandridae" genus="Hypselotriton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="genus">Hypselotriton Wolterstorff, 1934</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQCCI" type="type species">
<paragraph id="E4A64B1D84E9FD7785B8B03D9867D9CC">
<heading id="E487AC607DB10D69F5ECC8A7330FD30E" reason="title">
<typeStatus id="5DA7BAD2CEE7E83699D63B6FE56CBB5E">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="DDFD3FB653AE17D6E25F0C34124FA9FC">
<taxonomicName id="649497F872E913FAD8A73BD15B15E72B" authority="Boulenger, 1905" authorityName="Boulenger" authorityYear="1905" class="Amphibia" family="Salamandridae" genus="Molge" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="wolterstorffi">
<emphasis id="11E3E7AB918276644F92C8681FA4B329" italics="true">Molge wolterstorffi</emphasis>
<bibRefCitation id="79B717FC180808B59563ACCB793A0089" author="Boulenger GA" refId="B3" refString="Boulenger GA (1905) Description of a new newt from Yunnan. Proceedings of the Zoological Society of London 1905: 277." year="1905">Boulenger, 1905</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBDCI" type="diagnosis">
<paragraph id="785D2F35EBB39AF5972A3D78EFCD36EB">
<heading id="D5A4606D82BC798574C2A96A8F590CD0" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="5640B061A8DB6E73E69CBD1BEAC3819D">(1) small to large size; (2) gular fold present; (3) mid-dorsal vertebral ridge continuous; (4) metacarpal and metatarsal tubercles on external side of hands and feet present; (5) postocular orange spot present; (6) blue lateral stripe on tail in breeding males present.</paragraph>
</subSubSection>
<subSubSection id="SECID0EGDCI" type="Content and remarks">
<paragraph id="B9A126B5D575D690F778114394AAFB79">
<heading id="7836EBA45EB7220ED7173787869B9654" reason="title">Content and remarks.</heading>
</paragraph>
<paragraph id="F0639F93564BA19BD5A56926C0F58E62">
This subgenus includes all known Fire-bellied Newts populations distributed in the Yunnan-Guizhou Plateau in southwestern
<collectingCountry id="9501620889F0AA6FF836E88B3B47B3DB" name="China">China</collectingCountry>
. Currently, four nomenclatures have been provisionally documented, i. e.
<taxonomicName id="B03C8D490189267515B1962D43D3EA9E" authority="(Kou &amp; Xing, 1983)" baseAuthorityName="Kou &amp; Xing" baseAuthorityYear="1983" class="Amphibia" family="Salamandridae" genus="Hypselotriton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="chenggongensis" subGenus="Hy.">
<emphasis id="A755A2D7EAC32F517389349C957E34E9" italics="true">H.</emphasis>
(
<emphasis id="88D2F838F6EC95D764EB6EEAA907BCDC" italics="true">Hy.</emphasis>
)
<emphasis id="9435E4D813A4708A487215BC31A399FB" italics="true">chenggongensis</emphasis>
(
<bibRefCitation id="FAA18178F80AB5805A271F69EB37444E" author="Kou ZT &amp; Xing YL" journalOrPublisher="New Series" pagination="51-54" refId="B17" refString="Kou ZT, Xing YL (1983) A new species of Cynops from Yunnan. Acta Herpetologica Sinica. New Series 2: 51 - 54." title="A new species of Cynops from Yunnan. Acta Herpetologica Sinica." volume="2" year="1983">Kou &amp; Xing, 1983</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="3FCDB97D6AAB42D35CE4CAD15DE481F0" authority="(Liu, Hu &amp; Yang, 1962)" baseAuthorityName="Liu, Hu &amp; Yang" baseAuthorityYear="1962" class="Amphibia" family="Salamandridae" genus="Hypselotriton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="cyanurus" subGenus="Hy.">
<emphasis id="5715D5AAA5C62C2CCC7A52B5FBDD8A3F" italics="true">H.</emphasis>
(
<emphasis id="D162678518B3135FA74C1277E8506C7C" italics="true">Hy.</emphasis>
)
<emphasis id="7FE186DE4ADBD21D6480C9DF1B1BCAEB" italics="true">cyanurus</emphasis>
(
<bibRefCitation id="F5FFE5AF18AD67CD47E13B7E9073DC4C" author="Liu CC &amp; Hu SQ &amp; Yang FH" journalOrPublisher="Dong Wu Xue Bao" pagination="381-392" refId="B18" refString="Liu CC, Hu SQ, Yang FH (1962) Preliminary report of Amphibia from western Kweichow. Dong Wu Xue Bao 14: 381 - 392." title="Preliminary report of Amphibia from western Kweichow." volume="14" year="1962">Liu, Hu &amp; Yang, 1962</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="DEAF0142D8F7DD95809E5F17CFEF58DF" authority="(Boulenger, 1905)" baseAuthorityName="Boulenger" baseAuthorityYear="1905" class="Amphibia" family="Salamandridae" genus="Molge" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="wolterstorffi" subGenus="Hy.">
<emphasis id="74790CBB345020185FB09F2B7E9B6AD2" italics="true">H.</emphasis>
(
<emphasis id="4AC17A11AE0750984A0D3E636E4A5C0B" italics="true">Hy.</emphasis>
)
<emphasis id="4B60A47E315C11FCE2D7A422A82C4D45" italics="true">wolterstorffi</emphasis>
(
<bibRefCitation id="A9A6039AE92EE1F2EA14CE2AB5640397" author="Boulenger GA" refId="B3" refString="Boulenger GA (1905) Description of a new newt from Yunnan. Proceedings of the Zoological Society of London 1905: 277." year="1905">Boulenger, 1905</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="574EC615A4C1F91CF6B9489FBA8ADC71" authority="(Yang, 1979) (Lyu et al. 2023 a)" authorityName="Lyu et al." authorityYear="2023" baseAuthorityName="Yang" baseAuthorityYear="1979" class="Amphibia" family="Salamandridae" genus="Hypselotriton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="yunnanensis" subGenus="Hy.">
<emphasis id="624C09959BEF2D8A34244C355B704AD9" italics="true">H.</emphasis>
(
<emphasis id="2730F50BCD341F8E8BBC65FB888875C2" italics="true">Hy.</emphasis>
)
<emphasis id="4830A0719B118E19EC35B0A550FDB52E" italics="true">yunnanensis</emphasis>
(Yang, 1979) (
<bibRefCitation id="C4B93D1CD329C871847FDE3260575443" DOI="10.16373/j.cnki.ahr.220059" author="Lyu ZT &amp; Qi S &amp; Zhang SY &amp; Dai KY &amp; Wang YY" journalOrPublisher="Asian Herpetological Research" pagination="41-53" refId="B20" refString="Lyu ZT, Qi S, Zhang SY, Dai KY, Wang YY (2023 a) Two new species of fire-bellied newts (Caudata, Salamandridae, Cynops) from southeastern China. Asian Herpetological Research 14: 4153. https://doi.org/10.16373/j.cnki.ahr.220059" title="Two new species of fire-bellied newts (Caudata, Salamandridae, Cynops) from southeastern China." volume="14">Lyu et al. 2023 a</bibRefCitation>
)
</taxonomicName>
. However, the exact taxonomic status for these populations / species is still unresolved (
<bibRefCitation id="8A1AD13A80A930F2B0A9A4F56CCC1AD9" author="Raffaelli" firstAuthor="Raffaelli" refId="B23" refString="Raffaëlli J (2022) Salamanders &amp; Newts of the World. Plumelec, France: Penclen Édition." year="2022">Raffaëlli 2022</bibRefCitation>
;
<bibRefCitation id="D74C1E468A4DC4AAE764C734B8CD1466" DOI="10.16373/j.cnki.ahr.220059" author="Lyu ZT &amp; Qi S &amp; Zhang SY &amp; Dai KY &amp; Wang YY" journalOrPublisher="Asian Herpetological Research" pagination="41-53" refId="B20" refString="Lyu ZT, Qi S, Zhang SY, Dai KY, Wang YY (2023 a) Two new species of fire-bellied newts (Caudata, Salamandridae, Cynops) from southeastern China. Asian Herpetological Research 14: 4153. https://doi.org/10.16373/j.cnki.ahr.220059" title="Two new species of fire-bellied newts (Caudata, Salamandridae, Cynops) from southeastern China." volume="14">Lyu et al. 2023 a</bibRefCitation>
). Thus, it is improper to present a key to these nomenclatures before a comprehensive study to precisely delimitate their taxonomic placements.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="58799B8F862FC596F4C63FC884FE8C86">
<emphasis id="D027E9C55E3188F97D84415C86C0D767" italics="true">
Subgenus
<taxonomicName id="A94D9571B66B6C88A85D18B019D984AD" authority="Dubois &amp; Raffaelli, 2011" authorityName="Dubois &amp; Raffaelli" authorityYear="2011" class="Amphibia" family="Salamandridae" higherTaxonomySource="CoL" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="subGenus" subGenus="Cynotriton">
Cynotriton
<bibRefCitation id="EE60B7EC499151FE2DB87054AA480D90" author="Dubois A &amp; Raffaëlli J" journalOrPublisher="Alytes" pagination="151-153" refId="B6" refString="Dubois A, Raffaelli J (2011) Subgeneric taxonomy and nomenclature of the genus Hypselotriton Wolterstorff, 1934 (Amphibia, Urodela). Alytes 27: 151 - 153." title="Subgeneric taxonomy and nomenclature of the genus Hypselotriton Wolterstorff, 1934 (Amphibia, Urodela)." volume="27" year="2011">Dubois &amp; Raffaëlli, 2011</bibRefCitation>
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<paragraph id="E93E08B3EAD1634D1AD61A48D609EFC0">
<heading id="3D06B24EFBED828FED1E9E7F9BAA382D" reason="title">
<typeStatus id="973D939D87BEA80CE53BDD693358B323">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="0A45577D38E1CD136AA19550FC2D2AC4">
<taxonomicName id="B67C78DC93878DA953240316A3692529" authority="David, 1875" authorityName="David" authorityYear="1875" class="Amphibia" family="Salamandridae" genus="Triton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="orientalis" subGenus="Cynops">
<emphasis id="51DFC4D090EC488E799E78A3AC73AD49" italics="true">Triton</emphasis>
(
<emphasis id="5620909288D0160D5357877BBEA8C003" italics="true">Cynops</emphasis>
)
<emphasis id="53247149BD98B56F875B5400B638380D" italics="true">orientalis</emphasis>
David, 1875
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, by original designation.
</paragraph>
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<subSubSection id="SECID0ESTAK" type="diagnosis">
<paragraph id="B842C2C5FC919030A77D573DB1B16A31">
<heading id="5CD45701D8871917CF7B45D1A508FA07" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="F9F2A292A6FD1E7D3A7BB850E9F577A2">(1) small size; (2) gular fold present or absent; (3) mid-dorsal vertebral ridge interrupted; (4) metacarpal and metatarsal tubercles on external side of hands and feet absent; (5) postocular orange spot absent; (6) blue lateral stripe on tail in breeding males absent; (7) irregular greyish-blue patches on dorsum and lateral tail in breeding males and females absent.</paragraph>
</subSubSection>
<subSubSection id="SECID0EXTAK" type="Content">
<paragraph id="7812DC6D4C75617434377FBA08835463">
<heading id="F181621253679CE337243A4B835A03C7" reason="title">Content.</heading>
</paragraph>
<paragraph id="E7DE00C2EF91706DF3B0A13625460B1D">
Four recognised species distributed in eastern
<collectingCountry id="29E9AAA9958152C40F7EC7111A148139" name="China">China</collectingCountry>
(Fig.
<figureCitation id="DCA1924EB44FD6CE696F565CE0F99A47" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map showing the localities for congeners of genus Hypselotriton from Southeast Chinese Hilly Area. Inset map on top left zooms in eastern Guangdong Province, showing the localities for H. (Cynotriton) oolong sp. nov. and H. (Hakkatriton) orphicus: 1. Mt Fenghuang, Chaoan District, Chaozhou City (type locality of H. oolong sp. nov.); 2. Dayang Township, Jiexi County, Jieyang City (purported type locality of H. orphicus; see Remarks in the species account for locality delimitation); 3. Liangtian Township, Jiexi County (type locality of H. orphicus according to the original coordinate); 4. Mt Dabei, Jiexi County; 5. Mt Liwangzhang, Jiexi County; 6. Mt Hongtuzhang, Fengshun County, Meizhou City; 7. Mt Tongguzhang, Fengshun County; 8. Mt Shijiadong, Fengshun County; 9. Meiguang Village, Mt Lianhua, Wuhua County, Meizhou City (type locality of H. glaucus syn. nov.). The map is derived from Tianditu (www. tianditu. gov. cn)." figureDoi="10.3897/zse.100.127268.figure1" httpUri="https://binary.pensoft.net/fig/1113320">1</figureCitation>
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</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="70AEA977FE48EAF234E408C93C09E6EF">journal article</mods:classification>
<mods:identifier id="F5767E1AC1ED8AF30954A3E856ED35C3" type="DOI">10.3897/zse.100.129455</mods:identifier>
<mods:identifier id="D073223DBB7C316EA9542454E9A56EC5" type="ZooBank">11F9B5B3-CEF8-4C81-943D-E23C9FF823FB</mods:identifier>
</mods:mods>
<treatment id="EFE0ABCF5E3E55CEA3CB8DFAA24FE1B2" ID-DOI="http://doi.org/10.5281/zenodo.13322655" ID-Zenodo-Dep="13322655" ID-arpha="EFE0ABCF-5E3E-55CE-A3CB-8DFAA24FE1B2" ID-zoobank="https://zoobank.org/953A9F87-2E64-4A11-AFD6-061303E94409" LSID="urn:lsid:zoobank.org:act:953A9F87-2E64-4A11-AFD6-061303E94409" LSID-ZBK="urn:lsid:zoobank.org:act:953A9F87-2E64-4A11-AFD6-061303E94409" httpUri="http://treatment.plazi.org/id/EFE0ABCF5E3E55CEA3CB8DFAA24FE1B2">
<subSubSection id="C77F444AB81F0960A992646322B3576D" type="nomenclature">
<paragraph id="CAC3A7E677CFACF8235E152E03C24979">
<taxonomicName id="7E50D687959096699B9FE7D481412A91" authority="Pholyotha &amp; Panha" authorityName="Pholyotha &amp; Sutcharit &amp; Panha &amp; Tongkerd" authorityYear="2024" class="Gastropoda" family="Ariophantidae" genus="Janbinmorpha" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="622DDA0219132A4C83E99E8FD8DB524D" italics="true">Janbinmorpha</emphasis>
Pholyotha &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="128259656FCCE63FC4E4D032F6ED916C" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EC3DK" type="type species">
<paragraph id="14B10431E966CDB04E5F3F4B98AB7324">
<heading id="40AF03F7C607173DA124D6A5A90C250F" reason="title">
<typeStatus id="2FAF98AC7CE4E60F7D7E7A3E4C7E9E69">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="8C0F389C82951375B8A1664BDCC6BCC6">
<taxonomicName id="D55B1D9153266CF6F93F48E0AD8D83AE" authority="Martens, 1883" authorityName="Martens" authorityYear="1883" class="Gastropoda" family="Trochomorphidae" genus="Trochomorpha" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="sculpticarina">
<emphasis id="D880065CA5FFA621E76F0F3D50EAEBE1" italics="true">Trochomorpha sculpticarina</emphasis>
<bibRefCitation id="9E90FFC2D6BE2AFCF53B6D66F12A9CD7" author="Martens von E" journalOrPublisher="Conchologische Mittheilungen" pagination="129-138" refId="B28" refString="Martens von E (1883) Conchylien von Salanga. Conchologische Mittheilungen 2: 129 - 138. https: // www. biodiversitylibrary. org / page / 15865880" title="Conchylien von Salanga." url="https://www.biodiversitylibrary.org/page/15865880" volume="2" year="1883">Martens, 1883</bibRefCitation>
</taxonomicName>
; here designated.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ET3DK" type="etymology">
<paragraph id="80267DFFF693C8EBB83C8CD819B0489C">
<heading id="81F0A381EA42F34AC974E4317FD984AF" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="1DA8F988FD9EE1214C139D1B488AF082">
The name combines “
<emphasis id="A0EF1590671E612117FE580B1CDD9E81" italics="true">Janbin</emphasis>
” in reference to the shape of an unidentified flying object (
<abbrev id="ABBRID0EA4DK" xlink_title="unidentified flying object">UFO</abbrev>
) in the Thai language, and similar to the shell shape of this new genus, and “
<emphasis id="77AE0F5DA27BED5AE115BA230C47EA6F" italics="true">morpha</emphasis>
” in reference to the similarity of shell morphology between this new genus and the genus
<taxonomicName id="B2ABDC94A3A4EDA07DA7850BB95A2F27" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Trochomorphidae" genus="Trochomorpha" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="D77ED5CA3945738F1E20408422B08E07" italics="true">Trochomorpha</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EN4DK" type="diagnosis">
<paragraph id="01ECA5E1FC4030EFEBA00EDFD8674106">
<heading id="39FD55AD07D8060CF6788FBBC6AEE755" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="4EDB54840D474E45A0B70B8B66503927">Shell dextral, umbilicated, depressed, lenticular, ribbed, with keeled last whorl. Animal with three dorsal lobes; foot tripartite; caudal foss present; caudal horn very reduced. Genitalia having epiphallic caecum, flagellum and dart apparatus; proximal epiphallus encircled with thickened muscular epiphallic sheath; gametolytic organ without duct and gametolytic sac consisting of two bulbs. Radula with tricuspid central tooth; unicuspid or bicuspid lateral teeth; unicuspid marginal teeth.</paragraph>
</subSubSection>
<subSubSection id="SECID0EZ4DK" type="description">
<paragraph id="A4CE5B715D6754D03EE53AC12DEBFF90">
<heading id="04E59B8A270760E6173C13C35257A0CE" reason="title">Description.</heading>
</paragraph>
<paragraph id="701A22E19C2DDAA8BF941E3974C833D9">
See below under the
<typeStatus id="C3DB585421EAAAE11145DA8A7DFAC0EE">type</typeStatus>
species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E54DK" type="Constituent species">
<paragraph id="52753E6E46D50DB07306A508436C7433">
<heading id="BF3D5FE503E198B5E7C8563AC4BBF686" reason="title">Constituent species.</heading>
</paragraph>
<paragraph id="31866DAC609B912CB94D3C74448F3AB8">
This new genus contains only the
<typeStatus id="CE66AF8EAFB8F79DFA9E2B216F3DF7F5">type</typeStatus>
species,
<taxonomicName id="C72708B9AFF42ECDB86D675AF4003421" authorityName="Pholyotha &amp; Sutcharit &amp; Panha &amp; Tongkerd" authorityYear="2024" baseAuthorityName="Pholyotha &amp; Sutcharit &amp; Panha &amp; Tongkerd" baseAuthorityYear="1883" class="Gastropoda" family="Trochomorphidae" genus="Trochomorpha" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="sculpticarina" status="comb. nov.">
<emphasis id="757BD88FCF19E5EC8F5B6ED591FDD2A0" italics="true">J. sculpticarina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C9BF41C923D64C5F05D0393217E2E630" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EP5DK" type="distribution">
<paragraph id="B0BC40B51BA812C96D5EA5C84504340D">
<heading id="4F541B9BBD7B70EB9343A818D6B16179" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="211D622DFD4E8DA9E21677CE7F855D36">Currently known only from the Malay Peninsula.</paragraph>
</subSubSection>
<subSubSection id="SECID0EU5DK" type="remarks">
<paragraph id="67C4F321C40DE112A74D29AC4265CAB0">
<heading id="32A4B66C684E58E7E4F04CEE7C08AE03" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="E7D949904A199A8089EF7F925CB0EDD1">
<taxonomicName id="0F0C0440AF0C0A37C702BCE4E60154B4" authorityName="Pholyotha &amp; Sutcharit &amp; Panha &amp; Tongkerd" authorityYear="2024" class="Gastropoda" family="Trochomorphidae" genus="Janbinmorpha" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="721314E4C22186BA192A7E1E6E7FE61A" italics="true">Janbinmorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CB56EC304D5B982D70FCD659789EF2CD" rank="genus">gen. nov.</taxonomicNameLabel>
is clearly discriminated from all other Southeast Asian limacoid genera (e. g.,
<taxonomicName id="A3C9362999FD08978423602F353D406A" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="4935722E93A3E62FCB39D0FA7D56A5AF" italics="true">Hemiplecta</emphasis>
</taxonomicName>
,
<taxonomicName id="259E698478E8DBC9CCCB3AB0E0ADB27F" authority="Godwin-Austen, 1908" authorityName="Godwin-Austen" authorityYear="1908" class="Gastropoda" family="Helicarionidae" genus="Holkeion" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="DCA3AF010B0D2AED6FCC92495273571A" italics="true">Holkeion</emphasis>
Godwin-Austen, 1908
</taxonomicName>
,
<taxonomicName id="BA2647110DA5536EBD0AD073130C8EF2" authority="Pholyotha" authorityName="Pholyotha" authorityYear="2023" class="Gastropoda" family="Microcystidae" genus="Siamoconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="3C76215DF29303CC67AE8D47249B4355" italics="true">Siamoconus</emphasis>
Pholyotha
</taxonomicName>
in
<bibRefCitation id="625A683BB12DA71EEC39C3EEC51FCAD1" DOI="10.1071/IS23012" author="Pholyotha A &amp; Panha S &amp; Sutcharit C &amp; Jirapatrasilp P &amp; Seesamut T &amp; Liew T-S &amp; Tongkerd P" issue="8" journalOrPublisher="Invertebrate Systematics" pagination="571-605" refId="B44" refString="Pholyotha A, Panha S, Sutcharit C, Jirapatrasilp P, Seesamut T, Liew T-S, Tongkerd P (2023 a) Molecular phylogeny of the land snail family Euconulidae in Thailand and its position in the superfamily Trochomorphoidea (Stylommatophora: Limacoidei), with description of a new genus. Invertebrate Systematics 37 (8): 571605. https://doi.org/10.1071/IS23012" title="Molecular phylogeny of the land snail family Euconulidae in Thailand and its position in the superfamily Trochomorphoidea (Stylommatophora: Limacoidei), with description of a new genus." volume="37">Pholyotha et al. 2023 a</bibRefCitation>
,
<taxonomicName id="B609A2DD6A2B66F331669F25C60475EF" authorityName="Adams &amp; Adams" authorityYear="1855" class="Gastropoda" family="Trochomorphidae" genus="Videna" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="C59F182A7D86CB699FF6B65FF436E64D" italics="true">Videna</emphasis>
</taxonomicName>
and
<taxonomicName id="CD74C14D0EC7BB966D0484A6BBC6AC17" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Trochomorphidae" genus="Trochomorpha" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="D6279243D405F7333D85A21A892E47DF" italics="true">Trochomorpha</emphasis>
</taxonomicName>
) by the shape of the gametolytic sac and by the attachment of the penial retractor muscle. This new genus has no gametolytic duct and the gametolytic sac is divided into two lobes, while the gametolytic sac of all other limacoid genera is not divided into two lobes and most of them have a gametolytic duct. The penial retractor muscle of this new genus is attached to two regions (epiphallic caecum and epiphallic sheath), while all other limacoid genera have the retractor muscle attached to either the epiphallic caecum or epiphallus (
<bibRefCitation id="3894EDA356A718DBAFB31229713978E8" DOI="10.5962/bhl.title.13103" author="Blanford and Godwin-Austen" firstAuthor="Blanford" refId="B5" refString="Blanford WT, Godwin-Austen HH (1908) The Fauna of British India. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London, UK. https://doi.org/10.5962/bhl.title.13103" year="1908">Blanford and Godwin-Austen 1908</bibRefCitation>
;
<bibRefCitation id="753BED173E2968DF00A94CE1EE2B1705" author="Solem" firstAuthor="Solem" journalOrPublisher="Spolia Zoologica Musei Hauniensis" pagination="1-110" refId="B54" refString="Solem A (1966) Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Spolia Zoologica Musei Hauniensis 24: 1110." title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
;
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,
<bibRefCitation id="661C8148C07C2ED5D2B960EA86F10D04" author="Schileyko AA" refId="B52" refString="Schileyko AA (2002 b) Treatise on recent terrestrial pulmonate molluscs. Part 9. Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae. Ruthenica (Supplement 2): 11671307.">b</bibRefCitation>
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;
<bibRefCitation id="E2794D5DF6D4B106EFA73CD0D01042D8" DOI="10.1111/jzs.12420" author="Sutcharit C &amp; Jeratthitikul E &amp; Pholyotha A &amp; Lin A &amp; Panha S" issue="4" journalOrPublisher="Journal of Zoological Systematics and Evolutionary Research" pagination="957-981" refId="B58" refString="Sutcharit C, Jeratthitikul E, Pholyotha A, Lin A, Panha S (2020 a) Molecular phylogeny reveals high diversity and endemism in the limestone karst-restricted land snail genus Sophina Benson, 1859 from Myanmar (Eupulmonata: Helicarionidae), with description of four new species. Journal of Zoological Systematics and Evolutionary Research 58 (4): 957981. https://doi.org/10.1111/jzs.12420" title="Molecular phylogeny reveals high diversity and endemism in the limestone karst-restricted land snail genus Sophina Benson, 1859 from Myanmar (Eupulmonata: Helicarionidae), with description of four new species." volume="58">
Sutcharit et al. 2020
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</bibRefCitation>
;
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;
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,
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,
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,
<bibRefCitation id="47B223244C86ADEEE8B2CE5B100DFF9B" DOI="10.1127/arch.moll/152/269-280" author="Pholyotha A &amp; Sutcharit C &amp; Panha S" issue="2" journalOrPublisher="Archiv für Molluskenkunde" pagination="269-280" refId="B45" refString="Pholyotha A, Sutcharit C, Panha S (2023 b) Taxonomic revision of the land-snail genus Holkeion Godwin-Austen, 1908 (Eupulmonata: Ariophantidae), with a description of a new species, H. nordsiecki. Archiv für Molluskenkunde 152 (2): 269280. https://doi.org/10.1127/arch.moll/152/269-280" title="Taxonomic revision of the land-snail genus Holkeion Godwin-Austen, 1908 (Eupulmonata: Ariophantidae), with a description of a new species, H. nordsiecki." volume="152">2023 b</bibRefCitation>
,
<bibRefCitation id="C0AB1961B784905F33F375CDC7B85897" author="Pholyotha A &amp; Tongkerd P &amp; Sutcharit C" refId="B46" refString="Pholyotha A, Tongkerd P, Sutcharit C (2023 c) Siamochlamys panhai, a new limestone karst-restricted terrestrial snail from Thailand (Eupulmonata: Helicarionidae). Tropical Natural History (Supplement 7): 4150.">c</bibRefCitation>
).
</paragraph>
<paragraph id="1824A4FA52DB9CE4E7EDC556AEFEEAC1">
<taxonomicName id="F778523FCA8B44EE7732D67FBB3F57D9" authorityName="Pholyotha &amp; Sutcharit &amp; Panha &amp; Tongkerd" authorityYear="2024" class="Gastropoda" family="Trochomorphidae" genus="Janbinmorpha" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="F2E620C33B7E8691A40B4CBE33884540" italics="true">Janbinmorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="235590786CEBA64D032ADCAC72067A85" rank="genus">gen. nov.</taxonomicNameLabel>
also differs from
<taxonomicName id="9ED5B23AB030FF6DB1FAD9BC954628F3" authorityName="Adams &amp; Adams" authorityYear="1855" class="Gastropoda" family="Trochomorphidae" genus="Videna" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="2AEAAB1104B79A532C602277DC8490A0" italics="true">Videna</emphasis>
</taxonomicName>
in having a long dart apparatus and the position of gametolytic organ on female side, while the latter genus has no dart apparatus and gametolytic duct opening into the base of the penis (
<bibRefCitation id="32F86C172F1F76F0D5C9B93848A57C2F" author="Schileyko AA" refId="B51" refString="Schileyko AA (2002 a) Treatise on recent terrestrial pulmonate molluscs. Part 8. Punctidae, Helicodiscidae, Discidae, Cystopeltidae, Euconulidae, Trochomorphidae. Ruthenica (Supplement 2): 10351166.">Schileyko 2002 a</bibRefCitation>
).
</paragraph>
<paragraph id="D256A4A38FD3D80E40417C26E13DA272">
<taxonomicName id="18AE66E09EA51C49096DDA4EC369E52F" authorityName="Pholyotha &amp; Sutcharit &amp; Panha &amp; Tongkerd" authorityYear="2024" class="Gastropoda" family="Trochomorphidae" genus="Janbinmorpha" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="E956528470EB1D56B739218E4CCDC1DF" italics="true">Janbinmorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="14F75C9F824409B9960840E23486F4F7" rank="genus">gen. nov.</taxonomicNameLabel>
is conchologically very similar to the Indian helicarionoidean genus
<taxonomicName id="D28854338BFCC7D1FDD171E3B5DD2455" authority="Blanford, 1863" authorityName="Blanford" authorityYear="1863" class="Gastropoda" family="Helicarionidae" genus="Sivella" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="946D3C098DB84C8DC79E40AB3504D48B" italics="true">Sivella</emphasis>
Blanford, 1863
</taxonomicName>
. However, this new genus differs by having a long dart apparatus, a straight epiphallic caecum, and no gametolytic duct. In comparison,
<taxonomicName id="3B6DD106A0CF677A24137531124A1A42" authorityName="Blanford" authorityYear="1863" class="Gastropoda" family="Helicarionidae" genus="Sivella" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="384E5201E7FEBE734A9A792ABFEEFEA7" italics="true">Sivella</emphasis>
</taxonomicName>
has a relatively short gametolytic duct and a subglobular gametolytic sac, and does not have a dart apparatus or epiphallic caecum (
<bibRefCitation id="42F4F8AC253F8B4B2C348452EDCF5574" DOI="10.26515/rzsi/v8/i12/1918/163139" author="Godwin-Austen" firstAuthor="Godwin-Austen" issue="12" journalOrPublisher="Records of the Indian Museum" pagination="601-621" refId="B12" refString="Godwin-Austen HH (1918) Zoological results of the Abor Expedition, 19111912. XLIX. Mollusca, IX. Records of the Indian Museum 8 (12): 601621. https://doi.org/10.26515/rzsi/v8/i12/1918/163139" title="Zoological results of the Abor Expedition, 1911-1912. XLIX. Mollusca, IX." volume="8" year="1918">Godwin-Austen 1918</bibRefCitation>
;
<bibRefCitation id="AFD7463B7031E8E3EEE6865737AA805C" author="Schileyko" firstAuthor="Schileyko" refId="B53" refString="Schileyko AA (2003) Treatise on recent terrestrial pulmonate mollusks. Part 10. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae. Ruthenica (Supplement 2): 13091466." year="2003">Schileyko 2003</bibRefCitation>
).
</paragraph>
<paragraph id="72EDDCD9B1261CB6478039294051F6BC">
In addition, the molecular phylogeny (Fig.
<figureCitation id="4D18DE5CAF47B7F908F1C34CF5290355" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood phylogram based on analyses of concatenated COI, 16 S, and 28 S sequences. Numbers on branches indicate nodal support by ML bootstrap (BS) and Bayesian posterior probabilities (PP) analyses. Asterisk indicates the type species of its corresponding genus. The sample code in front of species name indicates voucher samples as shown in Table 1. Living specimen of Janbinmorpha sculpticarina (Martens, 1883), comb. nov. from Phang-nga Province, not to scale." figureDoi="10.3897/zse.100.129455.figure1" httpUri="https://binary.pensoft.net/fig/1113481">1</figureCitation>
) supports a distinct lineage of the monotypic
<taxonomicName id="63BFBE885563F49D223C31012FC1108C" authorityName="Pholyotha &amp; Sutcharit &amp; Panha &amp; Tongkerd" authorityYear="2024" class="Gastropoda" family="Trochomorphidae" genus="Janbinmorpha" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="0F7FD252695AA3E8F0059E2775F4ED69" italics="true">Janbinmorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8C208EC057B92618CF86D3800DF1200C" rank="genus">gen. nov.</taxonomicNameLabel>
from genera
<taxonomicName id="3F4A4F9A064A754855851058747826D4" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="1ADC7EBBD9EDEAD49C6AE9E92B9CFA34" italics="true">Hemiplecta</emphasis>
</taxonomicName>
,
<taxonomicName id="05A7CE6412CDE234C6B3501D516B0DE4" authorityName="Pholyotha" authorityYear="2023" class="Gastropoda" family="Microcystidae" genus="Siamoconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="0A3A8DAF6D39A52EC070D66FD675516F" italics="true">Siamoconus</emphasis>
</taxonomicName>
,
<taxonomicName id="825BFDDBD5DBF56BBA9250B3125CB7C0" authorityName="Adams &amp; Adams" authorityYear="1855" class="Gastropoda" family="Trochomorphidae" genus="Videna" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="76F8D1EFA4583CA556475CE65031F7E9" italics="true">Videna</emphasis>
</taxonomicName>
and
<taxonomicName id="04C4F760C2DE4893A3A45BF085DF1960" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Trochomorphidae" genus="Trochomorpha" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="323209CF8D2E5C01882D9F9EE1228EC2" italics="true">Trochomorpha</emphasis>
</taxonomicName>
. Currently, genetic sequences of
<taxonomicName id="57AA8AF2B333AE0F818F4CF83C6FD183" authorityName="Godwin-Austen" authorityYear="1908" class="Gastropoda" family="Helicarionidae" genus="Holkeion" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="0CEC3F45B7D52776CFE6847D2B764090" italics="true">Holkeion</emphasis>
</taxonomicName>
and
<taxonomicName id="F446F62DBC798AB02374B8553E6F5FC6" authorityName="Blanford" authorityYear="1863" class="Gastropoda" family="Helicarionidae" genus="Sivella" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="543A117AC638A4D6496DAB4CE086DD2E" italics="true">Sivella</emphasis>
</taxonomicName>
are not available in the online GenBank sequence database for phylogenetic analysis.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,128 @@
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<mods:title id="6949399C56F7B63F9C46E0EFD7FF5A20">Taxonomic determination of Hypselotriton populations distributed in eastern Guangdong, China (Caudata, Salamandridae), with description of a new species and a new subgenus</mods:title>
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<treatment id="F8B933C9BD955387AC78A5E3786F8099" ID-DOI="http://doi.org/10.5281/zenodo.13322662" ID-Zenodo-Dep="13322662" ID-arpha="F8B933C9-BD95-5387-AC78-A5E3786F8099" LSID="urn:lsid:plazi:treatment:F8B933C9BD955387AC78A5E3786F8099" httpUri="http://treatment.plazi.org/id/F8B933C9BD955387AC78A5E3786F8099">
<subSubSection id="C539CE475980B35B7E5D0FEADA824B46" type="nomenclature">
<paragraph id="AF069E02D5158C985DAB3DC6AA88E9E9">
<heading id="6982EB55C0487BF5A10EACA5451CCD77" reason="label">Genus</heading>
<taxonomicName id="B42C44FCB5664B6B44424CBFED922884" ID-CoL="52ZK" authority="Wolterstorff, 1934" authorityName="Wolterstorff" authorityYear="1934" class="Amphibia" family="Salamandridae" genus="Hypselotriton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="genus">
<emphasis id="1C651C7C8493A3010B83A88FCF181C5A" italics="true">Hypselotriton</emphasis>
Wolterstorff, 1934
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EO3BI" type="type species">
<paragraph id="A8EAB33B463E4C0552798B7E9D94E6BE">
<heading id="EC88EE59A97EA5A618AB7513B28AE8FD" reason="title">
<typeStatus id="BA853B9D87C17325F65C658A7DF5372C">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="703835BEA1E071AC49715B39724A6D7C">
<taxonomicName id="AD838D3CCCA9569F3EDD24AD1B2E4243" authority="Boulenger, 1905" authorityName="Boulenger" authorityYear="1905" class="Amphibia" family="Salamandridae" genus="Molge" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="species" species="wolterstorffi">
<emphasis id="8FB3DCCE34749F4FED1ED26577294245" italics="true">Molge wolterstorffi</emphasis>
<bibRefCitation id="2767F5D3D2A8EC8D0E98C5C146701DDA" author="Boulenger GA" refId="B3" refString="Boulenger GA (1905) Description of a new newt from Yunnan. Proceedings of the Zoological Society of London 1905: 277." year="1905">Boulenger, 1905</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E63BI" type="Common name">
<paragraph id="BEA2229F74E30A2BB5BF46AA4598C70F">
<heading id="CF12D5F41B7FD634974AEA3C74E40573" reason="title">Common name.</heading>
</paragraph>
<paragraph id="9DE7D610C6EA8EBDF9BE68DEAF388682">Chinese Fire-bellied Newts (in English) / diān yuán shǔ (滇螈属 in Chinese).</paragraph>
</subSubSection>
<subSubSection id="SECID0EE4BI" type="distribution">
<paragraph id="16E1C6C914DEA679E0B9399104756517">
<heading id="2FA9F7B4E525B85C802B60147B467C99" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="2912A37DB4734F8D98A02758236B32A3">
Endemic in mainland
<collectingCountry id="C6650A1DD90383A94A1251FA0EEB6717" name="China">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJ4BI" type="remarks">
<paragraph id="F8DAFD63B43FF6F4C1A82C393E3124E9">
<heading id="75903F655C408900830FE09CF99ABF7C" reason="title">Remark.</heading>
</paragraph>
<paragraph id="F4AF27D0C97BD79A59AAC2D2CAAA8206">
In spite of the paraphyletic relationships and distinct geographical isolation, species of the mainland genus
<taxonomicName id="5E4CCE88FFBDFE7F36B44FDF23012A4D" authorityName="Wolterstorff" authorityYear="1934" class="Amphibia" family="Salamandridae" genus="Hypselotriton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="genus">
<emphasis id="614CFB3771FD0BDAB2BFED0B309698DB" italics="true">Hypselotriton</emphasis>
</taxonomicName>
are very similar to those of insular genus
<taxonomicName id="C388D266471EC82F8490E8A91C4EE3C1" authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Salamandridae" genus="Cynops" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="genus">
<emphasis id="9636914C590572C755612D78D2117407" italics="true">Cynops</emphasis>
</taxonomicName>
in morphology, resulting in the prolonged categorising of these Fire-bellied Newts in a single genus.
<taxonomicName id="8F242A86703D601330F2915EEF65CBCF" authorityName="Wolterstorff" authorityYear="1934" class="Amphibia" family="Salamandridae" genus="Hypselotriton" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="genus">
<emphasis id="A997FC89E1808DE973CFE8F76C3DF37B" italics="true">Hypselotriton</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="21D7A7C7F00D1CBFC2EE8DCED617373F" authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Salamandridae" genus="Cynops" kingdom="Animalia" order="Caudata" phylum="Chordata" rank="genus">
<emphasis id="672DB55E0883BE7FD7C5F80333D2E21F" italics="true">Cynops</emphasis>
</taxonomicName>
by the inconspicuous parotoid gland (vs. well developed) and smooth or slightly granular skin (vs. distinctly granular).
</paragraph>
</subSubSection>
</treatment>
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