Add updates up until 2025-02-18 09:02:20
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<document id="0A170AD981303D65AD832B0DF61C56AF" ID-DOI="10.5252/zoosystema2025v47a5" ID-ISSN="1638-9387" ID-Zenodo-Dep="14886499" ID-ZooBank="102DD048-3E93-4147-B3E3-7559706C53A1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739868909815" checkinUser="plazi" docAuthor="Josiane, Natdanai Likhitrakarn, Sergei, Josiane Lips, Golovatch, Sergei I., Somsak, Ekgachai Jeratthitikul, Chirasak, Somsak Panha & Sutcharit, Chirasak" docDate="2025" docId="D87287BA7221FFC080B3139A047C2F9D" docLanguage="en" docName="zoosystema.2025.47.5.pdf" docOrigin="Zoosystema 47 (5)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2025v47a5.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.10:Zoosystema.2020-.journal_article.type1" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article.type1" docStyleVersion="10" docTitle="Tylopus namnonensis Likhitrakarn 2025, n. sp." docType="treatment" docVersion="2" lastPageNumber="83" masterDocId="244BFFC27226FFCA8055156B077A2C73" masterDocTitle="Three new species of the Oriental millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae) from China and Laos" masterLastPageNumber="88" masterPageNumber="75" pageNumber="80" updateTime="1739869334499" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="FF9DB730DBFC3C9BD5CF35F57A1B0EF7">
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<mods:title id="353A7DEC16DA478FB8D5C9F8C4FD9F0B">Three new species of the Oriental millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae) from China and Laos</mods:title>
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<mods:namePart id="73BF5E70EE48436A956321746EA1973E">Josiane, Natdanai Likhitrakarn</mods:namePart>
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<mods:namePart id="4EAD70A9D6B56BC9739A344F9A17DB7D">Sergei, Josiane Lips</mods:namePart>
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<mods:namePart id="706CAC3F635D6775D54900F744F5C384">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="13F10FDE6C22028CA7528D36D564919B">Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow, 119071 (Russia)</mods:affiliation>
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<mods:namePart id="36EE163292F8C836A2019E133A219F5F">Somsak, Ekgachai Jeratthitikul</mods:namePart>
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<mods:namePart id="6D346D95C32C23E036F56528DFE0EDAD">Chirasak, Somsak Panha</mods:namePart>
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<mods:namePart id="AC150EA8DD4BB7740F8806BEC02CD1BD">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="068FA2E5ED25B01FECB58F21BCDC0244">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330 (Thailand) chirasak. s @ chula. ac. th, jirasak 4 @ yahoo. com (corresponding author)</mods:affiliation>
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<mods:title id="955564305C055FE63481850A43954468">Zoosystema</mods:title>
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<mods:date id="C501D3D6F1D32CC495EE0A66E70ECC6E">2025</mods:date>
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<mods:number id="F6EB06C122BD88EBAC33054138556419">2025-02-17</mods:number>
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<mods:identifier id="B35963C64535F810728C6CD345191B43" type="ZooBank">102DD048-3E93-4147-B3E3-7559706C53A1</mods:identifier>
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<taxonomicName id="97DB4D2F7221FFCD80B3139A051A2B7F" authority="Likhitrakarn" authorityName="Likhitrakarn" authorityYear="2025" box="[230,608,1777,1804]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="80" phylum="Arthropoda" rank="species" species="namnonensis" status="sp. nov.">
|
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<emphasis id="62AFEABE7221FFCD80B3139A06B42B7F" bold="true" box="[230,462,1777,1804]" italics="true" pageId="7" pageNumber="80">Tylopus namnonensis</emphasis>
|
||||
Likhitrakarn
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="799C57C57221FFCD823F139905DF2B7F" box="[618,677,1778,1804]" pageId="7" pageNumber="80" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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<subSubSection id="18C165277221FFCD81C4127905962B13" pageId="7" pageNumber="80" type="description">
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<paragraph id="506436AC7221FFCD81C4127906832B5F" blockId="7.[230,677,1777,1836]" box="[401,505,1810,1836]" pageId="7" pageNumber="80">
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(
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<figureCitation id="C8E02A297221FFCD81CD1279068B2B5F" box="[408,497,1810,1836]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="8.[132,143,845,863]" captionStartId-1="8.[132,143,1964,1982]" captionStartId-2="9.[132,143,1009,1027]" captionTargetBox-0="[132,1455,216,802]" captionTargetBox-1="[160,1426,936,1920]" captionTargetBox-2="[298,1259,215,957]" captionTargetId-0="figure-18@8.[132,1455,216,802]" captionTargetId-1="figure-26@8.[160,1427,936,1920]" captionTargetId-2="figure-593@9.[284,1272,214,960]" captionTargetPageId-0="8" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIG. 4. — Tylopus namnonensis Likhitrakarn, n. sp.: A, Nam Non cave (the type locality), inside the cave entrance; B, ♀ paratype, dorsal view in live color pattern, pictures taken not to scale." captionText-1="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." captionText-2="FIG. 6. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype, left gonopod: A-D, mesal, lateral, caudal and suboral views, respectively; E, leg of ring 11. Scale bars: 0.2 mm. Abbreviations: fe, femur; l, lobe l; h, process h; pfe, prefemoral part; slm, solenomere; sph, solenophore; x, process x." figureDoi-0="http://doi.org/10.5281/zenodo.14886507" figureDoi-1="http://doi.org/10.5281/zenodo.14886509" figureDoi-2="http://doi.org/10.5281/zenodo.14886511" httpUri-0="https://zenodo.org/record/14886507/files/figure.png" httpUri-1="https://zenodo.org/record/14886509/files/figure.png" httpUri-2="https://zenodo.org/record/14886511/files/figure.png" pageId="7" pageNumber="80">Figs 4-6</figureCitation>
|
||||
)
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</paragraph>
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urn:lsid:zoobank.org:act:
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<subSubSection id="18C165277221FFCD80D112E805812B99" pageId="7" pageNumber="80" type="materials_examined">
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<paragraph id="506436AC7221FFCD80D112E805732BBC" blockId="7.[132,775,1922,2029]" pageId="7" pageNumber="80">
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<materialsCitation id="E0B33CF17221FFCD80D112E805732BBC" collectingDate="2023-02-28" collectionCode="TYPE, MATERIAL" collectorName="J. Lips" country="Laos" elevation="175" latitude="18.028889" location="Tham Nam Non" longLatPrecision="21" longitude="104.69056" municipality="Konglor" pageId="7" pageNumber="80" specimenCount="1" specimenCount-male="1" stateProvince="Khammouan" typeStatus="holotype">
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. —
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<typeStatus id="8F60880E7221FFCD811D12E806D32BE8" box="[328,425,1923,1947]" pageId="7" pageNumber="80" type="holotype">Holotype</typeStatus>
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.
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<collectingCountry id="28CC763C7221FFCD81E612E8069B2BE8" box="[435,481,1923,1947]" name="Laos" pageId="7" pageNumber="80">Laos</collectingCountry>
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</emphasis>
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•
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<specimenCount id="46DDFD257221FFCD81AC12EC05772BED" box="[505,525,1927,1950]" count="1" pageId="7" pageNumber="80" type="male">♂</specimenCount>
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;
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<collectingRegion id="921FF84E7221FFCD824F12E904792BE9" box="[538,771,1922,1946]" country="Laos" name="Khammouan" pageId="7" pageNumber="80">Khammouane Province</collectingRegion>
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,
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||||
<collectingMunicipality id="B000ACD67221FFCD80D012F607AC2BC6" box="[133,214,1949,1973]" pageId="7" pageNumber="80">Konglor</collectingMunicipality>
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,
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;
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a.s.l.
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;
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,
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;
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<collectingDate id="3421E9847221FFCD80D112D3078A2BA3" box="[132,240,1976,2000]" pageId="7" pageNumber="80" value="2023-02-28">28.II.2023</collectingDate>
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;
|
||||
<collectorName id="FD2E537A7221FFCD80AE12D306402BBC" box="[251,314,1975,1999]" pageId="7" pageNumber="80">J. Lips</collectorName>
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leg.; CUMZ-27205.
|
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</materialsCitation>
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<paragraph id="506436AC7221FFCD80D112B905812B99" blockId="7.[132,775,1922,2029]" box="[132,763,2002,2029]" pageId="7" pageNumber="80">
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<materialsCitation id="E0B33CF17221FFCD80D112B9058D2B99" box="[132,759,2002,2029]" collectingDate="2023-02-28" collectionCode="TYPE, MATERIAL" collectorName="J. Lips" country="Laos" elevation="175" latitude="18.028889" location="Tham Nam Non" longLatPrecision="21" longitude="104.69056" municipality="Konglor" pageId="7" pageNumber="80" specimenCount="1" specimenCount-female="1" stateProvince="Khammouan" typeStatus="paratype">
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<emphasis id="62AFEABE7221FFCD80D112B9066C2B99" bold="true" box="[132,278,2002,2026]" pageId="7" pageNumber="80">
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<typeStatus id="8F60880E7221FFCD80D112B907A72B99" box="[132,221,2002,2026]" pageId="7" pageNumber="80" type="paratype">Paratype</typeStatus>
|
||||
.
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||||
<collectingCountry id="28CC763C7221FFCD80BD12B9066C2B99" box="[232,278,2002,2026]" name="Laos" pageId="7" pageNumber="80">Laos</collectingCountry>
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</emphasis>
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•
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<specimenCount id="46DDFD257221FFCD817912B906372B9E" box="[300,333,2002,2029]" count="1" pageId="7" pageNumber="80" type="female">1 ♀</specimenCount>
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; same data as the holotype; CUMZ-27205
|
||||
</materialsCitation>
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||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
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<subSubSection id="18C165277221FFCD83781053022F291A" pageId="7" pageNumber="80" type="etymology">
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. — To emphasize Nam Non cave, the
|
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<typeStatus id="8F60880E7221FFCD8545105302432923" box="[1296,1337,1336,1360]" pageId="7" pageNumber="80">type</typeStatus>
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||||
locality, one of the biggest caves in
|
||||
<collectingRegion id="921FF84E7221FFCD8453103A0394291A" box="[1030,1262,1361,1385]" country="Laos" name="Khammouan" pageId="7" pageNumber="80">Khammouane Province</collectingRegion>
|
||||
; adjective.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="18C165277221FFCD837810E803432A4A" pageId="7" pageNumber="80" type="diagnosis">
|
||||
<paragraph id="506436AC7221FFCD837810E803432A4A" blockId="7.[811,1456,1411,1593]" pageId="7" pageNumber="80">
|
||||
<smallCapsWord id="5682A0707221FFCD837810E804ED29EA" baselines="1428,1429" box="[813,919,1411,1434]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="7" pageNumber="80">DIAGNOSIS</smallCapsWord>
|
||||
. — This new species closely resembles
|
||||
<taxonomicName id="97DB4D2F7221FFCD854510E8030029C6" authority="Likhitrakarn, Golovatch & Panha, 2016" authorityName="Likhitrakarn, Golovatch & Panha" authorityYear="2016" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="80" phylum="Arthropoda" rank="species" species="dorsalis">
|
||||
<emphasis id="62AFEABE7221FFCD854510E8021429E8" box="[1296,1390,1411,1435]" italics="true" pageId="7" pageNumber="80">T. dorsalis</emphasis>
|
||||
<bibRefCitation id="344A4B5D7221FFCD852610E8030029C6" author="LIKHITRAKARN N. & GOLOVATCH S. I. & PANHA S." pageId="7" pageNumber="80" pagination="1 - 47" refId="ref9482" refString="LIKHITRAKARN N., GOLOVATCH S. I. & PANHA S. 2016. - The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina. European Journal of Taxonomy 195: 1-47. https://doi.org/10.5852/ejt.2016.195" type="journal article" year="2016">Likhitrakarn, Golovatch & Panha, 2016</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, particularly in gonopod conformation. However, this new species is distinguished by the metaterga being roughly microgranulate (vs smooth), the distal position of process x on the solenophore (vs median), the pleurosternal carinae in
|
||||
<specimenCount id="46DDFD257221FFCD833A136004F92A51" box="[879,899,1547,1570]" count="1" pageId="7" pageNumber="80" type="male">♂</specimenCount>
|
||||
extending to ring 13 (vs 16), and the tarsal brushes in
|
||||
<specimenCount id="46DDFD257221FFCD85CE136002D52A51" box="[1435,1455,1547,1570]" count="1" pageId="7" pageNumber="80" type="male">♂</specimenCount>
|
||||
present until ring 12 (vs 6).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="18C165277221FFC08378131806F12E83" lastPageId="10" lastPageNumber="83" pageId="7" pageNumber="80" type="description">
|
||||
<paragraph id="506436AC7221FFCD8378131804C72AF8" blockId="7.[811,1457,1651,2028]" box="[813,957,1651,1677]" pageId="7" pageNumber="80">
|
||||
<smallCapsWord id="5682A0707221FFCD8378131804C72AF8" baselines="1670,1670" box="[813,957,1651,1677]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="7" pageNumber="80">DESCRIPTION</smallCapsWord>
|
||||
</paragraph>
|
||||
<paragraph id="506436AC7221FFCD837813F904D42A9F" blockId="7.[811,1457,1651,2028]" pageId="7" pageNumber="80">
|
||||
Length
|
||||
<quantity id="97239B497221FFCD832A13F904982ADE" box="[895,994,1682,1709]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="7" pageNumber="80" unit="mm" value="22.5">22.5 mm</quantity>
|
||||
(
|
||||
<specimenCount id="46DDFD257221FFCD83A413FD037D2ADF" box="[1009,1031,1686,1708]" count="1" pageId="7" pageNumber="80" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="97239B497221FFCD844B13F903FB2ADE" box="[1054,1153,1682,1709]" metricMagnitude="-2" metricUnit="m" metricValue="2.93" pageId="7" pageNumber="80" unit="mm" value="29.3">29.3 mm</quantity>
|
||||
(
|
||||
<specimenCount id="46DDFD257221FFCD84C513FD03D92ADF" box="[1168,1187,1686,1708]" count="1" pageId="7" pageNumber="80" type="female">♀</specimenCount>
|
||||
), width of midbody pro- and metazona 2.2 and
|
||||
<quantity id="97239B497221FFCD847613D903032ABE" box="[1059,1145,1714,1741]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="80" unit="mm" value="2.8">2.8 mm</quantity>
|
||||
(
|
||||
<specimenCount id="46DDFD257221FFCD84DC13DE03E52AB8" box="[1161,1183,1717,1739]" count="1" pageId="7" pageNumber="80" type="male">♂</specimenCount>
|
||||
) or 2.7 and
|
||||
<quantity id="97239B497221FFCD857313D902062ABE" box="[1318,1404,1714,1741]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="80" unit="mm" value="3.5">3.5 mm</quantity>
|
||||
(
|
||||
<specimenCount id="46DDFD257221FFCD85D913DE02E52AB8" box="[1420,1439,1717,1739]" count="1" pageId="7" pageNumber="80" type="female">♀</specimenCount>
|
||||
), respectively.
|
||||
</paragraph>
|
||||
<paragraph id="506436AC7221FFC383161399047928DD" blockId="7.[811,1457,1651,2028]" lastBlockId="9.[130,778,1108,2028]" lastPageId="9" lastPageNumber="82" pageId="7" pageNumber="80">
|
||||
Coloration of live animals light brown (
|
||||
<figureCitation id="C8E02A297221FFCD84B01399024D2B7F" box="[1253,1335,1778,1804]" captionStart="FIG" captionStartId="8.[132,143,845,863]" captionTargetBox="[132,1455,216,802]" captionTargetId="figure-18@8.[132,1455,216,802]" captionTargetPageId="8" captionText="FIG. 4. — Tylopus namnonensis Likhitrakarn, n. sp.: A, Nam Non cave (the type locality), inside the cave entrance; B, ♀ paratype, dorsal view in live color pattern, pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.14886507" httpUri="https://zenodo.org/record/14886507/files/figure.png" pageId="7" pageNumber="80">Fig. 4B</figureCitation>
|
||||
), with contrasting dark brown to blackish collar covering both pro- and metazona, head, collum, rings 2 and 3 dark brown to blackish, antennae and epiproct light brown, antennomere 7 dark brown, paraterga, venter and legs yellowish brown; coloration of alcohol material after one year of preservation dark brown; paraterga, legs, antennae and epiproct light yellow-brown, antennomere 7 dark brown, head and collum brownish, following terga with a light brown triangle and a blackish collar covering both pro- and metazona (
|
||||
<figureCitation id="C8E02A29722FFFC381B3111F056528FC" box="[486,543,1140,1167]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5</figureCitation>
|
||||
A-F), venter and a few basal podomeres light yellowish to pallid (
|
||||
<figureCitation id="C8E02A29722FFFC3821211FF05C228DD" box="[583,696,1172,1199]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5A, B</figureCitation>
|
||||
, E-G).
|
||||
</paragraph>
|
||||
<caption id="04A46624722EFFC280D1162606122F06" ID-DOI="http://doi.org/10.5281/zenodo.14886507" ID-Zenodo-Dep="14886507" httpUri="https://zenodo.org/record/14886507/files/figure.png" pageId="8" pageNumber="81" startId="8.[132,143,845,863]" targetBox="[132,1455,216,802]" targetPageId="8" targetType="figure">
|
||||
<paragraph id="506436AC722EFFC280D1162606122F06" blockId="8.[132,1457,845,885]" pageId="8" pageNumber="81">
|
||||
<smallCapsWord id="5682A070722EFFC280D1162607E72F2E" baselines="858,858" box="[132,157,845,863]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="8" pageNumber="81">FIG</smallCapsWord>
|
||||
. 4. —
|
||||
<taxonomicName id="97DB4D2F722EFFC28086162606952F2C" authority="Likhitrakarn" authorityName="Likhitrakarn" authorityYear="2025" box="[211,495,845,863]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="81" phylum="Arthropoda" rank="species" species="namnonensis" status="sp. nov.">
|
||||
<emphasis id="62AFEABE722EFFC28086162606FF2F2C" box="[211,389,845,863]" italics="true" pageId="8" pageNumber="81">Tylopus namnonensis</emphasis>
|
||||
Likhitrakarn
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="799C57C5722EFFC281A01626055D2F2C" box="[501,551,845,863]" pageId="8" pageNumber="81" rank="species">n. sp.</taxonomicNameLabel>
|
||||
:
|
||||
<emphasis id="62AFEABE722EFFC28278162605402F2C" bold="true" box="[557,570,845,863]" pageId="8" pageNumber="81">A</emphasis>
|
||||
, Nam Non cave (the type locality), inside the cave entrance;
|
||||
<emphasis id="62AFEABE722EFFC28478162603402F2C" bold="true" box="[1069,1082,845,863]" pageId="8" pageNumber="81">B</emphasis>
|
||||
, ♀ paratype, dorsal view in live color pattern, pictures taken not to scale.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="04A46624722EFFC280D112C705BF2B98" ID-DOI="http://doi.org/10.5281/zenodo.14886509" ID-Zenodo-Dep="14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="8" pageNumber="81" startId="8.[132,143,1964,1982]" targetBox="[160,1426,936,1920]" targetPageId="8" targetType="figure">
|
||||
<paragraph id="506436AC722EFFC280D112C705BF2B98" blockId="8.[132,1456,1964,2027]" pageId="8" pageNumber="81">
|
||||
<smallCapsWord id="5682A070722EFFC280D112C707E72BCE" baselines="1977,1978" box="[132,157,1964,1982]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="8" pageNumber="81">FIG</smallCapsWord>
|
||||
. 5. —
|
||||
<taxonomicName id="97DB4D2F722EFFC2808012C706882BCD" authority="Likhitrakarn" authorityName="Likhitrakarn" authorityYear="2025" box="[213,498,1964,1982]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="81" phylum="Arthropoda" rank="species" species="namnonensis" status="sp. nov.">
|
||||
<emphasis id="62AFEABE722EFFC2808012C706F22BCD" box="[213,392,1964,1982]" italics="true" pageId="8" pageNumber="81">Tylopus namnonensis</emphasis>
|
||||
Likhitrakarn
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="799C57C5722EFFC281AC12C705562BCD" box="[505,556,1964,1982]" pageId="8" pageNumber="81" rank="species">n. sp.</taxonomicNameLabel>
|
||||
, ♂ holotype:
|
||||
<emphasis id="62AFEABE722EFFC282CF12C705DD2BCD" bold="true" box="[666,679,1964,1982]" pageId="8" pageNumber="81">A</emphasis>
|
||||
,
|
||||
<emphasis id="62AFEABE722EFFC282E712C705C52BCD" bold="true" box="[690,703,1964,1982]" pageId="8" pageNumber="81">B</emphasis>
|
||||
, anterior part of body, dorsal and lateral views, respectively;
|
||||
<emphasis id="62AFEABE722EFFC284E812C703B12BCD" bold="true" box="[1213,1227,1964,1982]" pageId="8" pageNumber="81">C</emphasis>
|
||||
,
|
||||
<emphasis id="62AFEABE722EFFC2848312C7039E2BCD" bold="true" box="[1238,1252,1964,1982]" pageId="8" pageNumber="81">D</emphasis>
|
||||
, rings 10 and 11, dorsal and lateral views, respectively;
|
||||
<emphasis id="62AFEABE722EFFC281DF12A806D12BA7" bold="true" box="[394,427,1986,2005]" pageId="8" pageNumber="81">E -G</emphasis>
|
||||
, posterior part of body, lateral, dorsal and ventral views, respectively;
|
||||
<emphasis id="62AFEABE722EFFC283AC12A8037D2BA6" bold="true" box="[1017,1031,1987,2005]" pageId="8" pageNumber="81">H</emphasis>
|
||||
,
|
||||
<emphasis id="62AFEABE722EFFC2844712A803622BA6" bold="true" box="[1042,1048,1987,2005]" pageId="8" pageNumber="81">I</emphasis>
|
||||
, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="04A46624722FFFC380D1169A03AD2869" ID-DOI="http://doi.org/10.5281/zenodo.14886511" ID-Zenodo-Dep="14886511" httpUri="https://zenodo.org/record/14886511/files/figure.png" pageId="9" pageNumber="82" startId="9.[132,143,1009,1027]" targetBox="[298,1259,215,957]" targetPageId="9" targetType="figure">
|
||||
<paragraph id="506436AC722FFFC380D1169A03AD2869" blockId="9.[132,1455,1009,1050]" pageId="9" pageNumber="82">
|
||||
<smallCapsWord id="5682A070722FFFC380D1169A07E72871" baselines="1022,1023" box="[132,157,1009,1027]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="9" pageNumber="82">FIG</smallCapsWord>
|
||||
. 6. —
|
||||
<taxonomicName id="97DB4D2F722FFFC38081169A06952870" authority="Likhitrakarn" authorityName="Likhitrakarn" authorityYear="2025" box="[212,495,1009,1027]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="namnonensis" status="sp. nov.">
|
||||
<emphasis id="62AFEABE722FFFC38081169A06FF2870" box="[212,389,1009,1027]" italics="true" pageId="9" pageNumber="82">Tylopus namnonensis</emphasis>
|
||||
Likhitrakarn
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="799C57C5722FFFC381A3169A055D2870" box="[502,551,1009,1027]" pageId="9" pageNumber="82" rank="species">n. sp.</taxonomicNameLabel>
|
||||
, ♂ holotype, left gonopod:
|
||||
<emphasis id="62AFEABE722FFFC38352169A04532870" bold="true" box="[775,809,1009,1027]" pageId="9" pageNumber="82">A -D</emphasis>
|
||||
, mesal, lateral, caudal and suboral views, respectively;
|
||||
<emphasis id="62AFEABE722FFFC384A0169A027B2870" bold="true" box="[1269,1281,1009,1027]" pageId="9" pageNumber="82">E</emphasis>
|
||||
, leg of ring 11. Scale bars: 0.2 mm. Abbreviations:
|
||||
<emphasis id="62AFEABE722FFFC3812C116306F02869" bold="true" box="[377,394,1032,1050]" pageId="9" pageNumber="82">fe</emphasis>
|
||||
, femur;
|
||||
<emphasis id="62AFEABE722FFFC3819A116306AE2869" bold="true" box="[463,468,1032,1050]" pageId="9" pageNumber="82">l</emphasis>
|
||||
, lobe l;
|
||||
<emphasis id="62AFEABE722FFFC382431163055B2869" bold="true" box="[534,545,1032,1050]" pageId="9" pageNumber="82">h</emphasis>
|
||||
, process h;
|
||||
<emphasis id="62AFEABE722FFFC382DD116305DE2869" bold="true" box="[648,676,1032,1050]" pageId="9" pageNumber="82">pfe</emphasis>
|
||||
, prefemoral part;
|
||||
<emphasis id="62AFEABE722FFFC3836C116304232869" bold="true" box="[825,857,1032,1050]" pageId="9" pageNumber="82">slm</emphasis>
|
||||
, solenomere;
|
||||
<emphasis id="62AFEABE722FFFC3839B116304942869" bold="true" box="[974,1006,1032,1050]" pageId="9" pageNumber="82">sph</emphasis>
|
||||
, solenophore;
|
||||
<emphasis id="62AFEABE722FFFC3843E1163030F2869" bold="true" box="[1131,1141,1032,1050]" pageId="9" pageNumber="82">x</emphasis>
|
||||
, process x.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="506436AC722FFFC380CE11DF057E299E" blockId="9.[130,778,1108,2028]" pageId="9" pageNumber="82">
|
||||
Clypeolabral region and vertex rather densely setose, epicranial suture distinct. Antennae relatively short (
|
||||
<figureCitation id="C8E02A29722FFFC382DC11BF0580289D" box="[649,762,1235,1262]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5A, B</figureCitation>
|
||||
), reaching to body ring 3 when stretched dorsally (
|
||||
<specimenCount id="46DDFD25722FFFC382CA119C05CF297E" box="[671,693,1271,1293]" count="1" pageId="9" pageNumber="82" type="male">♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="46DDFD25722FFFC38291119C05AD297E" box="[708,727,1271,1293]" count="1" pageId="9" pageNumber="82" type="female">♀</specimenCount>
|
||||
). In width, head <ring 3 <4 <collum <ring 2 <5 <6 <7-16 (
|
||||
<specimenCount id="46DDFD25722FFFC380D8105C07D9293E" box="[141,163,1335,1357]" count="1" pageId="9" pageNumber="82" type="male">♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="46DDFD25722FFFC380FA105C07B8293E" box="[175,194,1335,1357]" count="1" pageId="9" pageNumber="82" type="female">♀</specimenCount>
|
||||
), thereafter body gently and gradually tapering towards telson. Collum with three transverse rows of strong setae: 4+4 anterior, 1+ 1 intermediate, and 4+ 4 posterior; a small, lateral, setigerous incision near midway; caudal corner very broadly rounded, paraterga declined ventrad, not produced past rear tergal margin (
|
||||
<figureCitation id="C8E02A29722FFFC381D010B8068D299E" box="[389,503,1491,1518]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5A, B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="506436AC722FFFC380CE109803A82BD8" blockId="9.[130,778,1108,2028]" lastBlockId="9.[811,1457,1108,2028]" pageId="9" pageNumber="82">
|
||||
Tegument dull and shining, prozona finely shagreened, metaterga often roughly microgranulate and rugulose, leathery, surface below paraterga more delicately, but still sufficiently clearly microgranulate and rugulose (
|
||||
<figureCitation id="C8E02A29722FFFC38244133905342A1E" box="[529,590,1618,1645]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5</figureCitation>
|
||||
A-F). Postcollum metaterga with two transverse rows of rather long setae: 2 +
|
||||
<quantity id="97239B49722FFFC382AC131907E02ADF" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="9" pageNumber="82" unit="in" value="2.0">2 in</quantity>
|
||||
anterior and 3+
|
||||
<quantity id="97239B49722FFFC3811A13F906002ADF" box="[335,378,1682,1708]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="82" unit="in" value="3.0">3 in</quantity>
|
||||
posterior row, the latter often abrad- ed, but then readily traceable as insertion points. Tergal setae long, strong, slender, about 1/4 metatergal length. Axial line well visible on metazona, traceable also on prozona. Paraterga strongly developed (
|
||||
<figureCitation id="C8E02A29722FFFC3810F127906EC2B5F" box="[346,406,1809,1836]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5</figureCitation>
|
||||
A-G), especially so in
|
||||
<specimenCount id="46DDFD25722FFFC382D4127E05ED2B58" box="[641,663,1813,1835]" count="1" pageId="9" pageNumber="82" type="male">♂</specimenCount>
|
||||
, set rather high (at upper 1/3 body height), slightly upturned, but lying below dorsum; anterior edge broadly rounded and narrowly bordered, fused to callus; caudal corner rather narrowly round- ed, extending increasingly past rear tergal margin, especially well curved mesad on rings 16-19, posterior edge oblique (
|
||||
<figureCitation id="C8E02A29722FFFC380DB12BA07872B98" box="[142,253,2001,2028]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5F, G</figureCitation>
|
||||
); paraterga very thin blunt blades in lateral view, a little thicker only on pore-bearing rings (
|
||||
<figureCitation id="C8E02A29722FFFC384BB113F023F281C" box="[1262,1349,1108,1135]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5D</figureCitation>
|
||||
). Calluses on paraterga delimited by a sulcus both dorsally and ventrally. Paraterga 2 broad, horizontal, anterior edge angular, lateral edge with three evident incisions: two in anterior 1/3, one at midway, and the smallest one near a very broadly rounded caudal corner. Paraterga 3 and 4 each with two small incisions at lateral edge (
|
||||
<figureCitation id="C8E02A29722FFFC3839C10780361295D" box="[969,1051,1299,1326]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5A</figureCitation>
|
||||
), one in anterior 1/3, the other at posterior 1/3. Following paraterga each with one evident lateral incision in anterior 1/3 (
|
||||
<figureCitation id="C8E02A29722FFFC38461103803F0291D" box="[1076,1162,1363,1390]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5C</figureCitation>
|
||||
). Ozopores evident, lateral, lying in an ovoid groove at about 1/3 metatergal length in front of posterior edge of metaterga (
|
||||
<figureCitation id="C8E02A29722FFFC3842610F8039C29DE" box="[1139,1254,1427,1454]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5B, D</figureCitation>
|
||||
). Transverse sulcus usually distinct (
|
||||
<figureCitation id="C8E02A29722FFFC3838A10D8030F29BE" box="[991,1141,1459,1486]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5A, C, F</figureCitation>
|
||||
), slightly incomplete on ring 19, complete and clearly visible on metaterga 5-18, narrow, deep, reaching the bases of paraterga, ribbed at bottom. Stricture between pro- and metazona wide, deep, ribbed at bottom down to base of paraterga (
|
||||
<figureCitation id="C8E02A29722FFFC38468135903BF2A3E" box="[1085,1221,1586,1613]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5A, C, F</figureCitation>
|
||||
). Pleurosternal carinae complete crests on rings 2-4 (
|
||||
<specimenCount id="46DDFD25722FFFC38436133D03032A1F" box="[1123,1145,1622,1644]" count="1" pageId="9" pageNumber="82" type="male">♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="46DDFD25722FFFC384DC133D03E62A1F" box="[1161,1180,1622,1644]" count="1" pageId="9" pageNumber="82" type="female">♀</specimenCount>
|
||||
), clearly increasing in size on rings 5-7, thereafter broken into an anterior small bulge and a small and sharp caudal tooth, both growing increasingly reduced until ring 13, thereafter missing (
|
||||
<specimenCount id="46DDFD25722FFFC384B613DE03832AB8" box="[1251,1273,1717,1739]" count="1" pageId="9" pageNumber="82" type="male">♂</specimenCount>
|
||||
) (
|
||||
<figureCitation id="C8E02A29722FFFC3854513D902DA2ABF" box="[1296,1440,1714,1741]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5B, D, E</figureCitation>
|
||||
), or increasingly reduced and remaining only a small caudal tooth until ring 8, thereafter missing (
|
||||
<specimenCount id="46DDFD25722FFFC38491139E03AD2B78" box="[1220,1239,1781,1803]" count="1" pageId="9" pageNumber="82" type="female">♀</specimenCount>
|
||||
). Epiproct (
|
||||
<figureCitation id="C8E02A29722FFFC3850F139902EC2B7E" box="[1370,1430,1778,1805]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5</figureCitation>
|
||||
EG) conical, flattened dorsoventrally, subtruncate, with two evident apical papillae directed caudally, both pointed at tip; pre-apical papillae small, but evident, lying close to tip. Hypoproct subtrapeziform (
|
||||
<figureCitation id="C8E02A29722FFFC3846F121A03EA2BFF" box="[1082,1168,1905,1932]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5G</figureCitation>
|
||||
), small setiferous knobs at caudal edge well-separated and evident.
|
||||
</paragraph>
|
||||
<paragraph id="506436AC722FFFC0831612DA05AF2DC2" blockId="9.[811,1457,1108,2028]" lastBlockId="10.[130,777,215,752]" lastPageId="10" lastPageNumber="83" pageId="9" pageNumber="82">
|
||||
Sterna sparsely setose, without modifications (
|
||||
<figureCitation id="C8E02A29722FFFC3851212DA02DB2BBF" box="[1351,1441,1969,1996]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="9" pageNumber="82">Fig. 5G</figureCitation>
|
||||
); cross-impressions shallow; a large, central, slightly bifid, setose lobe between
|
||||
<specimenCount id="46DDFD25722CFFC0814C15B006552C82" box="[281,303,219,241]" count="1" pageId="10" pageNumber="83" type="male">♂</specimenCount>
|
||||
coxae 4 (
|
||||
<figureCitation id="C8E02A29722CFFC081C115BC05792C82" box="[404,515,215,242]" captionStart="FIG" captionStartId="8.[132,143,1964,1982]" captionTargetBox="[160,1426,936,1920]" captionTargetId="figure-26@8.[160,1427,936,1920]" captionTargetPageId="8" captionText="FIG. 5. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype: A, B, anterior part of body, dorsal and lateral views, respectively; C, D, rings 10 and 11, dorsal and lateral views, respectively; E-G, posterior part of body, lateral, dorsal and ventral views, respectively; H, I, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A-G, 0.1 mm; H, I, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14886509" httpUri="https://zenodo.org/record/14886509/files/figure.png" pageId="10" pageNumber="83">Fig. 5H, I</figureCitation>
|
||||
). Legs long and slender, midbody ones
|
||||
<emphasis id="62AFEABE722CFFC081711593064F2D62" box="[292,309,248,273]" italics="true" pageId="10" pageNumber="83">c.</emphasis>
|
||||
1.2-1.4 (
|
||||
<specimenCount id="46DDFD25722CFFC081CF159006CA2D62" box="[410,432,251,273]" count="1" pageId="10" pageNumber="83" type="male">♂</specimenCount>
|
||||
) or 0.9-1.1 times (
|
||||
<specimenCount id="46DDFD25722CFFC08228159005EA2D62" box="[637,656,251,273]" count="1" pageId="10" pageNumber="83" type="female">♀</specimenCount>
|
||||
) as long as body height; prefemora not swollen, legs on rings 6-8 each with a small adenostyle on postfemur and tibia; legs of rings 9-16 each with a small adenostyle on femur, postfemur, tibia and tarsus (
|
||||
<figureCitation id="C8E02A29722CFFC08157141C062C2DE2" box="[258,342,375,401]" captionStart="FIG" captionStartId="9.[132,143,1009,1027]" captionTargetBox="[298,1259,215,957]" captionTargetId="figure-593@9.[284,1272,214,960]" captionTargetPageId="9" captionText="FIG. 6. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype, left gonopod: A-D, mesal, lateral, caudal and suboral views, respectively; E, leg of ring 11. Scale bars: 0.2 mm. Abbreviations: fe, femur; l, lobe l; h, process h; pfe, prefemoral part; slm, solenomere; sph, solenophore; x, process x." figureDoi="http://doi.org/10.5281/zenodo.14886511" httpUri="https://zenodo.org/record/14886511/files/figure.png" pageId="10" pageNumber="83">Fig. 6E</figureCitation>
|
||||
); telopodites particularly densely setose ventrally (
|
||||
<figureCitation id="C8E02A29722CFFC080A714FD063E2DC3" box="[242,324,406,432]" captionStart="FIG" captionStartId="9.[132,143,1009,1027]" captionTargetBox="[298,1259,215,957]" captionTargetId="figure-593@9.[284,1272,214,960]" captionTargetPageId="9" captionText="FIG. 6. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype, left gonopod: A-D, mesal, lateral, caudal and suboral views, respectively; E, leg of ring 11. Scale bars: 0.2 mm. Abbreviations: fe, femur; l, lobe l; h, process h; pfe, prefemoral part; slm, solenomere; sph, solenophore; x, process x." figureDoi="http://doi.org/10.5281/zenodo.14886511" httpUri="https://zenodo.org/record/14886511/files/figure.png" pageId="10" pageNumber="83">Fig. 6E</figureCitation>
|
||||
), tarsal brushes present until ring 12.
|
||||
</paragraph>
|
||||
<paragraph id="506436AC722CFFC080CE14DD06F12E83" blockId="10.[130,777,215,752]" pageId="10" pageNumber="83">
|
||||
Gonopods rather simple (
|
||||
<figureCitation id="C8E02A29722CFFC081E914DD06842DA3" box="[444,510,438,464]" captionStart="FIG" captionStartId="9.[132,143,1009,1027]" captionTargetBox="[298,1259,215,957]" captionTargetId="figure-593@9.[284,1272,214,960]" captionTargetPageId="9" captionText="FIG. 6. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype, left gonopod: A-D, mesal, lateral, caudal and suboral views, respectively; E, leg of ring 11. Scale bars: 0.2 mm. Abbreviations: fe, femur; l, lobe l; h, process h; pfe, prefemoral part; slm, solenomere; sph, solenophore; x, process x." figureDoi="http://doi.org/10.5281/zenodo.14886511" httpUri="https://zenodo.org/record/14886511/files/figure.png" pageId="10" pageNumber="83">Fig. 6</figureCitation>
|
||||
A-D). Prefemorite (pfe) densely setose, about 1/3 as long as femorite + “postfemoral” part. Femorite (fe) slightly curved and stout, with an evident mesal groove and a clear distolateral sulcus demarcating a postfemoral part; lobe l simple (
|
||||
<figureCitation id="C8E02A29722CFFC0818F175D05362E23" box="[474,588,566,592]" captionStart="FIG" captionStartId="9.[132,143,1009,1027]" captionTargetBox="[298,1259,215,957]" captionTargetId="figure-593@9.[284,1272,214,960]" captionTargetPageId="9" captionText="FIG. 6. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype, left gonopod: A-D, mesal, lateral, caudal and suboral views, respectively; E, leg of ring 11. Scale bars: 0.2 mm. Abbreviations: fe, femur; l, lobe l; h, process h; pfe, prefemoral part; slm, solenomere; sph, solenophore; x, process x." figureDoi="http://doi.org/10.5281/zenodo.14886511" httpUri="https://zenodo.org/record/14886511/files/figure.png" pageId="10" pageNumber="83">Fig. 6B, C</figureCitation>
|
||||
); process h rather short, mediodorsal, flattened, tip rounded and with a small notch (
|
||||
<figureCitation id="C8E02A29722CFFC08085171D063A2EE3" box="[208,320,630,656]" captionStart="FIG" captionStartId="9.[132,143,1009,1027]" captionTargetBox="[298,1259,215,957]" captionTargetId="figure-593@9.[284,1272,214,960]" captionTargetPageId="9" captionText="FIG. 6. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype, left gonopod: A-D, mesal, lateral, caudal and suboral views, respectively; E, leg of ring 11. Scale bars: 0.2 mm. Abbreviations: fe, femur; l, lobe l; h, process h; pfe, prefemoral part; slm, solenomere; sph, solenophore; x, process x." figureDoi="http://doi.org/10.5281/zenodo.14886511" httpUri="https://zenodo.org/record/14886511/files/figure.png" pageId="10" pageNumber="83">Fig. 6B, C</figureCitation>
|
||||
); solenophore (sph) long and slender, typically coiled, expanded distad, flattened and evidently bifid, with process x being elongated, apically rounded and directed forward (
|
||||
<figureCitation id="C8E02A29722CFFC080B317BE06042E83" box="[230,382,725,752]" captionStart="FIG" captionStartId="9.[132,143,1009,1027]" captionTargetBox="[298,1259,215,957]" captionTargetId="figure-593@9.[284,1272,214,960]" captionTargetPageId="9" captionText="FIG. 6. — Tylopus namnonensis Likhitrakarn, n. sp., ♂ holotype, left gonopod: A-D, mesal, lateral, caudal and suboral views, respectively; E, leg of ring 11. Scale bars: 0.2 mm. Abbreviations: fe, femur; l, lobe l; h, process h; pfe, prefemoral part; slm, solenomere; sph, solenophore; x, process x." figureDoi="http://doi.org/10.5281/zenodo.14886511" httpUri="https://zenodo.org/record/14886511/files/figure.png" pageId="10" pageNumber="83">Fig. 6A, C, D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="18C16527722CFFC080D1167D047C2F9D" pageId="10" pageNumber="83" type="discussion">
|
||||
<paragraph id="506436AC722CFFC080D1167D07922F5D" blockId="10.[130,777,790,1007]" box="[132,232,790,816]" pageId="10" pageNumber="83">
|
||||
<smallCapsWord id="5682A070722CFFC080D1167D07922F5D" baselines="809,809" box="[132,232,790,816]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="10" pageNumber="83">REMARKS</smallCapsWord>
|
||||
</paragraph>
|
||||
<paragraph id="506436AC722CFFC080D1165E047C2F9D" blockId="10.[130,777,790,1007]" pageId="10" pageNumber="83">
|
||||
The specimens were collected inside a cave (
|
||||
<figureCitation id="C8E02A29722CFFC08201165E05DD2F3C" box="[596,679,821,847]" captionStart="FIG" captionStartId="8.[132,143,845,863]" captionTargetBox="[132,1455,216,802]" captionTargetId="figure-18@8.[132,1455,216,802]" captionTargetPageId="8" captionText="FIG. 4. — Tylopus namnonensis Likhitrakarn, n. sp.: A, Nam Non cave (the type locality), inside the cave entrance; B, ♀ paratype, dorsal view in live color pattern, pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.14886507" httpUri="https://zenodo.org/record/14886507/files/figure.png" pageId="10" pageNumber="83">Fig. 4A</figureCitation>
|
||||
), around 500 meters away from the entrance. In the wet season, an underground river takes up all of the gallery. Big trees and a lot of wood remains are carried inside. The specimens were collected on wood debris.Despite formally coming from a cave, this new species seems to be nothing more than a trogloxene.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
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||||
212
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<mods:title id="B251A7A121EE6C4498DD70573FFF405C">Three new species of the Oriental millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae) from China and Laos</mods:title>
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.
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.
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.
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. —
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, by original designation.
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. — Body medium-sized to large (
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|
||||
<emphasis id="62AFEABE7225FFC9833012F6040F2BC7" box="[869,885,1949,1972]" italics="true" pageId="3" pageNumber="76">c.</emphasis>
|
||||
<quantity id="97239B497225FFC9832E12F704882BC7" box="[891,1010,1948,1972]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="5.1" metricValueMin="0.5" pageId="3" pageNumber="76" unit="mm" value="2.8" valueMax="5.1" valueMin="0.5">0.5-5.1 mm</quantity>
|
||||
wide), with 20 segments. Paraterga from virtually missing to strongly developed, usually with lateral incisions. Transverse metatergal sulcus distinct. Leg relatively long and slender, most species present adenostyles (= ventral outgrowths) at least on some of the male legs.
|
||||
<specimenCount id="46DDFD257222FFCE813D135B06062A34" box="[360,380,1584,1607]" count="1" pageId="4" pageNumber="77" type="male">♂</specimenCount>
|
||||
tarsal brushes usually present. A sternal lobe between
|
||||
<specimenCount id="46DDFD257222FFCE815E132106652A12" box="[267,287,1610,1633]" count="1" pageId="4" pageNumber="77" type="male">♂</specimenCount>
|
||||
coxae 4 present.
|
||||
</paragraph>
|
||||
<caption id="04A466247222FFCE80D110CA03342993" ID-DOI="http://doi.org/10.5281/zenodo.14886503" ID-Zenodo-Dep="14886503" httpUri="https://zenodo.org/record/14886503/files/figure.png" pageId="4" pageNumber="77" startId="4.[132,143,1441,1459]" targetBox="[213,1362,212,1398]" targetPageId="4" targetType="figure">
|
||||
<paragraph id="506436AC7222FFCE80D110CA03342993" blockId="4.[132,1456,1441,1504]" pageId="4" pageNumber="77">
|
||||
<smallCapsWord id="5682A0707222FFCE80D110CA07E729C2" baselines="1454,1454" box="[132,157,1441,1459]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="4" pageNumber="77">FIG</smallCapsWord>
|
||||
. 1. —
|
||||
<taxonomicName id="97DB4D2F7222FFCE808710CA06B329C0" authority="Likhitrakarn" authorityName="Likhitrakarn" authorityYear="2025" box="[210,457,1441,1459]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="tropicalis" status="sp. nov.">
|
||||
<emphasis id="62AFEABE7222FFCE808710CA061B29C0" box="[210,353,1441,1459]" italics="true" pageId="4" pageNumber="77">Tylopus tropicalis</emphasis>
|
||||
Likhitrakarn
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="799C57C57222FFCE819A10CA057B29C0" box="[463,513,1441,1459]" pageId="4" pageNumber="77" rank="species">n. sp.</taxonomicNameLabel>
|
||||
, ♂ holotype:
|
||||
<emphasis id="62AFEABE7222FFCE823E10CA050229C0" bold="true" box="[619,632,1441,1459]" pageId="4" pageNumber="77">A</emphasis>
|
||||
, habitus, live coloration;
|
||||
<emphasis id="62AFEABE7222FFCE831110CA042B29C0" bold="true" box="[836,849,1441,1459]" pageId="4" pageNumber="77">B</emphasis>
|
||||
,
|
||||
<emphasis id="62AFEABE7222FFCE830E10CA041329C0" bold="true" box="[859,873,1441,1459]" pageId="4" pageNumber="77">C</emphasis>
|
||||
, anterior part of body,dorsal and lateral views, respectively;
|
||||
<emphasis id="62AFEABE7222FFCE850C10CA021D29C0" bold="true" box="[1369,1383,1441,1459]" pageId="4" pageNumber="77">D</emphasis>
|
||||
,
|
||||
<emphasis id="62AFEABE7222FFCE852410CA020729C0" bold="true" box="[1393,1405,1441,1459]" pageId="4" pageNumber="77">E</emphasis>
|
||||
, rings 10 and 11, dorsal and lateral views, respectively;
|
||||
<emphasis id="62AFEABE7222FFCE824810DC054729BA" bold="true" box="[541,573,1463,1481]" pageId="4" pageNumber="77">F -H</emphasis>
|
||||
, posterior part of body, dorsal, ventral and lateral views, respectively;
|
||||
<emphasis id="62AFEABE7222FFCE84D710DC03F229BA" bold="true" box="[1154,1160,1463,1481]" pageId="4" pageNumber="77">I</emphasis>
|
||||
,
|
||||
<emphasis id="62AFEABE7222FFCE84C710DC03E629BA" bold="true" box="[1170,1180,1463,1481]" pageId="4" pageNumber="77">J</emphasis>
|
||||
, sternal cones between coxae 4, subcaudal and sublateral views, respectively. Scale bars: A, D, G, 0.5 mm; B, C, E, F, H, 1 mm; I, 0.2 mm; J, 0.3 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="506436AC7222FFCE80D1130B05F32B99" blockId="4.[130,775,1553,2027]" pageId="4" pageNumber="77">
|
||||
The gonopod telopodite typically features a prominent transverse ring, or cingulum, demarcating the postfemoral region which starts at the base of a free, flagelliform solenomere. This solenomere is largely sheathed by a slender, sigmoid solenophore that often bears basal outgrowths. The cingulum is rarely incomplete, usually due to a reduced sulcus at the base of a postfemoral lobe l. While lobe l itself is usually simple, sometimes it can be crowned with an additional outgrowth or process. Beyond lobe l, the postfemoral region usually possesses a medially positioned process h. Additional disto- and/or postfemoral outgrowth appearing even more sporadically is process z, more variable, albeit occasionally reduced or even absent. Only a few species show particularly complex gonopods, where not only is the postfemoral region supplied with a long, spiniform process z, but also with a small to elongated, knife- or spine-shaped outgrowth m on the distodorsal portion of the femorite (
|
||||
<bibRefCitation id="344A4B5D7222FFCE80D912B906EA2B98" author="GOLOVATCH S. I. & ENGHOFF H." box="[140,400,2002,2027]" pageId="4" pageNumber="77" pagination="85 - 125" refId="ref9104" refString="GOLOVATCH S. I. & ENGHOFF H. 1993. - Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85-125." type="journal article" year="1993">Golovatch & Enghoff 1993</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="344A4B5D7222FFCE81C912B905062B98" author="LIKHITRAKARN N. & GOLOVATCH S. I. & PRATEEPASEN R. & PANHA S." box="[412,636,2002,2027]" pageId="4" pageNumber="77" pagination="23 - 68" refId="ref9547" refString="LIKHITRAKARN N., GOLOVATCH S. I., PRATEEPASEN R. & PANHA S. 2010. - Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 72: 23-68. https://doi.org/10.3897/zookeys.72.744" type="journal article" year="2010">
|
||||
Likhitrakarn
|
||||
<emphasis id="62AFEABE7222FFCE824D12B8053F2B99" box="[536,581,2002,2026]" italics="true" pageId="4" pageNumber="77">et al.</emphasis>
|
||||
2010
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="18C165277222FFCE83781378025E2A1E" pageId="4" pageNumber="77" type="discussion">
|
||||
<paragraph id="506436AC7222FFCE8378137804FD2A58" blockId="4.[813,1455,1555,1645]" box="[813,903,1555,1581]" pageId="4" pageNumber="77">
|
||||
<smallCapsWord id="5682A0707222FFCE8378137804FD2A58" baselines="1574,1574" box="[813,903,1555,1581]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remark" pageId="4" pageNumber="77">REMARK</smallCapsWord>
|
||||
</paragraph>
|
||||
<paragraph id="506436AC7222FFCE83781359025E2A1E" blockId="4.[813,1455,1555,1645]" pageId="4" pageNumber="77">
|
||||
Full reviews of this genus were performed by
|
||||
<bibRefCitation id="344A4B5D7222FFCE8575135904AE2A1F" author="GOLOVATCH S. I. & ENGHOFF H." pageId="4" pageNumber="77" pagination="85 - 125" refId="ref9104" refString="GOLOVATCH S. I. & ENGHOFF H. 1993. - Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85-125." type="journal article" year="1993">Golovatch & Enghoff (1993)</bibRefCitation>
|
||||
and
|
||||
<bibRefCitation id="344A4B5D7222FFCE845E133902642A1F" author="LIKHITRAKARN N. & GOLOVATCH S. I. & PRATEEPASEN R. & PANHA S." box="[1035,1310,1618,1644]" pageId="4" pageNumber="77" pagination="23 - 68" refId="ref9547" refString="LIKHITRAKARN N., GOLOVATCH S. I., PRATEEPASEN R. & PANHA S. 2010. - Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 72: 23-68. https://doi.org/10.3897/zookeys.72.744" type="journal article" year="2010">
|
||||
Likhitrakarn
|
||||
<emphasis id="62AFEABE7222FFCE84CF133803B72A1F" box="[1178,1229,1618,1644]" italics="true" pageId="4" pageNumber="77">et al.</emphasis>
|
||||
(2010)
|
||||
</bibRefCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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Reference in a new issue