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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.8.e52190" ID-PMC="PMC7205841" ID-Pensoft-Pub="1314-2828-8-e52190" ID-Pensoft-UUID="4EE598BC99D8506FB10BD389A32B5A60" ID-PubMed="32398922" ModsDocID="1314-2828-8-e52190" checkinTime="1588265229323" checkinUser="pensoft" docAuthor="Plotkin, David &amp; Kawahara, Akito Y." docDate="2020" docId="3C5E68CB85C5519AB31EC21D13C33821" docLanguage="en" docName="BiodivDatJour 8: e52190" docOrigin="Biodiversity Data Journal 8" docSource="http://dx.doi.org/10.3897/BDJ.8.e52190" docTitle="Haruchlora Viidalepp &amp; Lindt 2014" docType="treatment" docVersion="2" id="4EE598BC99D8506FB10BD389A32B5A60" lastPageNumber="52190" masterDocId="4EE598BC99D8506FB10BD389A32B5A60" masterDocTitle="Review of recent taxonomic changes to the emerald moths (Lepidoptera: Geometridae: Geometrinae)" masterLastPageNumber="52190" masterPageNumber="52190" pageNumber="52190" updateTime="1668125462316" updateUser="ExternalLinkService">
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<mods:title>Review of recent taxonomic changes to the emerald moths (Lepidoptera: Geometridae: Geometrinae)</mods:title>
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<mods:namePart>Plotkin, David</mods:namePart>
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<mods:namePart>Kawahara, Akito Y.</mods:namePart>
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<taxonomicName LSID="3C5E68CB-85C5-519A-B31E-C21D13C33821" authority="Viidalepp &amp; Lindt, 2014 (&quot; gen. nov. &quot;)" authorityName="Viidalepp &amp; Lindt" authorityYear="2014" class="Insecta" family="Geometridae" genus="Haruchlora" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haruchlora" order="Lepidoptera" pageId="0" pageNumber="52190" phylum="Arthropoda" rank="genus">Haruchlora Viidalepp &amp; Lindt, 2014 (&quot;gen. nov.&quot;)</taxonomicName>
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<taxonomicName class="Insecta" family="Geometridae" genus="Haruchlora" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haruchlora maesi" order="Lepidoptera" pageId="0" pageNumber="52190" phylum="Arthropoda" rank="species" species="maesi">Haruchlora maesi</taxonomicName>
Viidalepp &amp; Lindt, 2014 (&quot;sp. nov.&quot;)
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<paragraph pageId="0" pageNumber="52190">Notes</paragraph>
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<taxonomicName authorityName="Viidalepp &amp; Lindt" authorityYear="2014" class="Insecta" family="Geometridae" genus="Haruchlora" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haruchlora" order="Lepidoptera" pageId="0" pageNumber="52190" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="52190">Haruchlora</emphasis>
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is currently a monotypic genus containing only the type species,
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<emphasis italics="true" pageId="0" pageNumber="52190">H. maesi</emphasis>
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; both the genus and the species were described by
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.
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</treatment>
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subsp.
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Wroughton 1899
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<emphasis id="3E2FA36FBBBFC1A3412C6FC4E9544908" italics="true" pageNumber="479">Pipistrellus (Pipistrellus) tenuis</emphasis>
subsp.
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Wroughton 1912
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;
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<emphasis id="876C1D681676DE9765D0B1D4D3F058B6" italics="true" pageNumber="479">Pipistrellus (Pipistrellus) tenuis</emphasis>
subsp.
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.
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<subSubSection id="81A4412084271AD6BC01A65CC4F677C9" pageId="0" pageNumber="125" type="description">
<paragraph id="F6C9FC74CB5945DDD474F84BC715F469" pageId="0" pageNumber="125">
<figureCitation id="BDC1418408F41736D700E4E95C96D696" captionStart="Figures 9–14" captionStartId="F2" captionText="Figures 9 - 14. Neotropical Cybistrinae species, habitus 9 Cybister festae 10 Cybister puncticollis 11 Nilssondytes diversus 12 Megadytes parvus 13 Megadytes latus 14 species &quot; IR 57 &quot; (Ribera et al. 2008). Scale bar: 10.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures9-14" httpUri="https://binary.pensoft.net/fig/963584" pageId="0" pageNumber="125">Figs 10</figureCitation>
<figureCitation id="DFB53E47800265841164CFEF9F9A316C" captionStart="Figures 15–29" captionStartId="F3" captionText="Figures 15 - 29. Neotropical Cybistrinae species, metatarsal claws and tarsomere VI of males and females 15, 16 Cybister festae 17, 18 Cybister puncticollis 19, 20 Nilssondytes diversus 21, 22 Megadytes latus 23, 24 Megadytes parvus 25 species &quot; IR 57 &quot; (Ribera et al. 2008) 26, 27 Paramegadytes glaucus 28, 29 Metaxydytes fraternus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures15-29" httpUri="https://binary.pensoft.net/fig/963585" pageId="0" pageNumber="125">, 17</figureCitation>
<figureCitation id="759A3F77D45C25A87B28B4A751A49872" captionStart="Figures 15–29" captionStartId="F3" captionText="Figures 15 - 29. Neotropical Cybistrinae species, metatarsal claws and tarsomere VI of males and females 15, 16 Cybister festae 17, 18 Cybister puncticollis 19, 20 Nilssondytes diversus 21, 22 Megadytes latus 23, 24 Megadytes parvus 25 species &quot; IR 57 &quot; (Ribera et al. 2008) 26, 27 Paramegadytes glaucus 28, 29 Metaxydytes fraternus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures15-29" httpUri="https://binary.pensoft.net/fig/963585" pageId="0" pageNumber="125">, 18</figureCitation>
<figureCitation id="45E714AF519DB0C81A6F3A40201C6A41" captionStart="Figures 30–50" captionStartId="F4" captionText="Figures 30 - 50. Neotropical Cybistrinae species, male genitalia 30 - 32 Cybister festae: 30 median lobe, right lateral aspect 31 median lobe, ventral aspect 32 right lateral lobe, right lateral aspect 33 - 35 Cybister puncticollis: 33 median lobe, right lateral aspect 34 median lobe, ventral aspect 35 right lateral lobe, right lateral aspect 36 - 38 Nilssondytes diversus: 36 median lobe, right lateral aspect 37 median lobe, ventral aspect 38 right lateral lobe, right lateral aspect 39 - 43 Megadytes parvus: 39 median lobe, right lateral aspect 40 median lobe, ventral aspect 41 right lateral lobe, right lateral aspect 42 apex of median lobe, right lateral aspect 43 apex of median lobe, ventral aspect 44 - 47 Megadytes latus: 44 median lobe, right lateral aspect 45 median lobe, ventral aspect 46 right lateral lobe, right lateral aspect 47 apex of median lobe, ventral aspect 48 - 50 species &quot; IR 57 &quot; (Ribera et al. 2008): 48 median lobe, right lateral aspect 49 median lobe, ventral aspect 50 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1188.110081.figures30-50" httpUri="https://binary.pensoft.net/fig/963586" pageId="0" pageNumber="125">, 33-35</figureCitation>
<figureCitation id="FF754457FD1E7D971352E89A20C83647" captionStart="Figures 51–57" captionStartId="F5" captionText="Figures 51 - 57. Neotropical Cybistrinae species, male sternite IX, ventral aspect 51 Cybister festae 52 Cybister puncticollis 53 Nilssondytes diversus 54 Megadytes latus 55 Megadytes parvus 56 Paramegadytes glaucus 57 Metaxydytes fraternus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures51-57" httpUri="https://binary.pensoft.net/fig/963587" pageId="0" pageNumber="125">, 52</figureCitation>
<figureCitation id="CBDBAC30A3352A8C2BA59C73DE2FE242" captionStart="Figures 58–61" captionStartId="F6" captionText="Figures 58 - 61. Neotropical Cybistrinae species, female genitalia including left gonocoxosternite, ventral aspect, except Fig. 60 without internal genitalia 58 Cybister festae 59 Cybister puncticollis 60 Megadytes latus 61 Nilssondytes diversus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures58-61" httpUri="https://binary.pensoft.net/fig/963588" pageId="0" pageNumber="125">, 59</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="17F7C8DC87146E3D38A05E59F7846BAC" pageId="0" pageNumber="125" type="reference_group">
<paragraph id="DC9FF32F2412C9311E83BEC09DFF2D0B" pageId="0" pageNumber="125">
<taxonomicName id="2FAF2360B77D17ED2C9BEF0AC5D5C687" class="Insecta" family="Dytiscidae" genus="Dyticus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dyticus puncticollis" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="puncticollis">Dyticus puncticollis</taxonomicName>
<normalizedToken id="6F75BED8ED034BE99A8E2E716C84F958" originalValue="Brullé">Brulle</normalizedToken>
, 1837: 46.
</paragraph>
<paragraph id="724D1C9EA11BF57481315C7225931CB2" pageId="0" pageNumber="125">
<taxonomicName id="56E9EB5026BB3EDE0E9F52AD85D4BBCF" baseAuthorityName="Brulle" baseAuthorityYear="1837" class="Insecta" family="Dytiscidae" genus="Cybister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybister puncticollis" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="puncticollis">Cybister puncticollis</taxonomicName>
:
<normalizedToken id="B40E7287393110E81AC184E17E824957" originalValue="Aubé">Aube</normalizedToken>
1838: 52.
</paragraph>
<paragraph id="70F3FDB63110D9A4159952A688E785D3" pageId="0" pageNumber="125">
<taxonomicName id="6CD5442115F44731040EBDA03DE638C0" class="Insecta" family="Dytiscidae" genus="Trogus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trogus puncticollis" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="puncticollis">Trogus puncticollis</taxonomicName>
: Gemminger and Harold 1868: 460.
</paragraph>
<paragraph id="66578601042381309165551889565A65" pageId="0" pageNumber="125">
<taxonomicName id="04E2B06C020601BD6924C4A906B0A2B1" class="Insecta" family="Dytiscidae" genus="Megadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megadytes puncticollis" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="puncticollis">Megadytes puncticollis</taxonomicName>
:
<bibRefCitation id="334492F9086D508C2E334EB4DCB85404" DOI="https://doi.org/10.5962/bhl.title.9530" author="Sharp, D" journalOrPublisher="Scientific Transactions of the Royal Dublin Society" pageId="0" pageNumber="125" pagination="179 - 1003" refId="B36" refString="Sharp, D, 1882. On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society 2: 179 - 1003, DOI: https://doi.org/10.5962/bhl.title.9530" title="On aquatic carnivorous Coleoptera or Dytiscidae." url="https://doi.org/10.5962/bhl.title.9530" volume="2" year="1882">Sharp 1882</bibRefCitation>
: 709; van den Branden 1885: 29; Regimbart 1889: 267; 1904: 225; Chatanay 1910: 434; Bruch 1915: 479; Zimmermann 1919: 235; 1920: 256; Wilke 1921: 24; Bruch 1927: 543;
<bibRefCitation id="63BC81C8FFE77850574E872FEDC70339" DOI="https://doi.org/10.5479/si.03629236.185.i" author="Blackwelder, RE" journalOrPublisher="Bulletin - United States National Museum" pageId="0" pageNumber="125" pagination="1 - 188" refId="B4" refString="Blackwelder, RE, 1944. Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America. Bulletin - United States National Museum 185: 1 - 188, DOI: https://doi.org/10.5479/si.03629236.185.i" title="Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America." url="https://doi.org/10.5479/si.03629236.185.i" volume="185" year="1944">Blackwelder 1944</bibRefCitation>
: 80;
<bibRefCitation id="C06AAAE30AD964173EC932B3CD83BE66" author="Brinck, P" journalOrPublisher="Kungliga Fysiografiska Saellskapets Handlingar" pageId="0" pageNumber="125" pagination="1 - 20" refId="B6" refString="Brinck, P, 1945. Nomenklatorische und systematische Studien ueber Dytisciden, III. Die Klassifikation der Cybisterinen. Kungliga Fysiografiska Saellskapets Handlingar 56: 1 - 20" title="Nomenklatorische und systematische Studien ueber Dytisciden, III. Die Klassifikation der Cybisterinen." volume="56" year="1945">Brinck 1945</bibRefCitation>
: 8;
<bibRefCitation id="A76E1E7BFB3CEC90DE714648F40DA89D" author="Mouchamps, R" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien" pageId="0" pageNumber="125" pagination="278 - 287" refId="B31" refString="Mouchamps, R, 1957. Contribution a la connaissance des Cybisterini (Col Dytiscidae) du Musee d'Histoire Naturelle de Vienne (9e note). Annalen des Naturhistorischen Museums in Wien 61: 278 - 287" title="Contribution a la connaissance des Cybisterini (Col Dytiscidae) du Musee d'Histoire Naturelle de Vienne (9 e note)." volume="61" year="1957">Mouchamps 1957</bibRefCitation>
: 283;
<bibRefCitation id="03CBF01E510513DBA3EE0945B3950339" author="Tremouilles, ER" journalOrPublisher="Revista de la Sociedad Entomologica Argentina" pageId="0" pageNumber="125" pagination="101 - 125" refId="B40" refString="Tremouilles, ER, Bachmann, AO, 1980. La tribu Cybisterini en la Argentina (Coleoptera, Dytiscidae). Revista de la Sociedad Entomologica Argentina 39: 101 - 125" title="La tribu Cybisterini en la Argentina (Coleoptera, Dytiscidae)." volume="39" year="1980">
<normalizedToken id="47EC453B6C0F9ACE75E43081B562F0D3" originalValue="Trémouilles">Tremouilles</normalizedToken>
and Bachmann 1980
</bibRefCitation>
: 105.
</paragraph>
<paragraph id="7DCF022133F56799C9CBD4345025F6B7" pageId="0" pageNumber="125">
<taxonomicName id="250301E1042C2DD9E6AAC3AB02F8A2A6" class="Insecta" family="Dytiscidae" genus="Cybister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybister (Meganectes) kemneri" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="kemneri" subGenus="Meganectes">Cybister (Meganectes) kemneri</taxonomicName>
Brinck, 1945: 18;
<bibRefCitation id="D8AC760095D1941EF570672EB4423D79" author="Tremouilles, ER" journalOrPublisher="Revista de la Sociedad Entomologica Argentina" pageId="0" pageNumber="125" pagination="101 - 125" refId="B40" refString="Tremouilles, ER, Bachmann, AO, 1980. La tribu Cybisterini en la Argentina (Coleoptera, Dytiscidae). Revista de la Sociedad Entomologica Argentina 39: 101 - 125" title="La tribu Cybisterini en la Argentina (Coleoptera, Dytiscidae)." volume="39" year="1980">
<normalizedToken id="E3F5C394DF0C5EFF1449BDFB07870595" originalValue="Trémouilles">Tremouilles</normalizedToken>
and Bachmann 1980
</bibRefCitation>
: 105.
</paragraph>
<paragraph id="0F9371233CBF390E47A43F34C4087635" pageId="0" pageNumber="125">
<taxonomicName id="28DB82AA35C7D049BEBF744EBF7CCDF0" baseAuthorityName="Brulle" baseAuthorityYear="1837" class="Insecta" family="Dytiscidae" genus="Cybister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybister (Meganectes) puncticollis" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="puncticollis" subGenus="Meganectes">Cybister (Meganectes) puncticollis</taxonomicName>
:
<bibRefCitation id="063A1C1F7EAA8213F6B77E6180C7BCCA" author="Tremouilles, ER" journalOrPublisher="Revista de la Sociedad Entomologica Argentina" pageId="0" pageNumber="125" pagination="101 - 125" refId="B40" refString="Tremouilles, ER, Bachmann, AO, 1980. La tribu Cybisterini en la Argentina (Coleoptera, Dytiscidae). Revista de la Sociedad Entomologica Argentina 39: 101 - 125" title="La tribu Cybisterini en la Argentina (Coleoptera, Dytiscidae)." volume="39" year="1980">
<normalizedToken id="5256BB13168132CA68CB26943A850F71" originalValue="Trémouilles">Tremouilles</normalizedToken>
and Bachmann 1980
</bibRefCitation>
: 105.
</paragraph>
<paragraph id="646C3DAE0DA19001842BEEDAAAE5F5BE" pageId="0" pageNumber="125">
<taxonomicName id="0606D0AECDD4C0D680037DB9AA3A81F4" baseAuthorityName="Brulle" baseAuthorityYear="1837" class="Insecta" family="Dytiscidae" genus="Cybister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybister (Neocybister) puncticollis" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="puncticollis" subGenus="Neocybister">Cybister (Neocybister) puncticollis</taxonomicName>
:
<bibRefCitation id="3689E8C34CA5C83B61B36C359A2CDFFD" DOI="https://doi.org/10.1111/j.1463-6409.2006.00254.x" author="Miller, KB" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="125" pagination="41 - 59" refId="B29" refString="Miller, KB, Bergsten, J, Whiting, MF, 2007. Phylogeny and classification of diving beetles in the tribe Cybistrini (Coleoptera, Dytiscidae, Dytiscinae). Zoologica Scripta 36 (1): 41 - 59, DOI: https://doi.org/10.1111/j.1463-6409.2006.00254.x" title="Phylogeny and classification of diving beetles in the tribe Cybistrini (Coleoptera, Dytiscidae, Dytiscinae)." url="https://doi.org/10.1111/j.1463-6409.2006.00254.x" volume="36" year="2007">Miller et al. 2007</bibRefCitation>
: 54;
<bibRefCitation id="C51B699431B66F9CC7D96536E77034AA" author="Nilsson, AN" journalOrPublisher="Canadian Journal of Zoology" pageId="0" pageNumber="125" refId="B33" refString="Nilsson, AN, Hajek, J, 2023. A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). 1.I.2023: 1-319." title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). 1. I. 2023: 1 - 319." year="2023">
Nilsson and
<normalizedToken id="F350D464ABE7F4AE94D33954E06B83B4" originalValue="Hájek">Hajek</normalizedToken>
2023
</bibRefCitation>
: 84.
</paragraph>
</subSubSection>
<subSubSection id="FAAFA3A1E21A740E943FED89A7678B86" pageId="0" pageNumber="125" type="type locality">
<paragraph id="D7799B8C4065E7992994114476D4EA68" pageId="0" pageNumber="125">Type locality.</paragraph>
<paragraph id="0E44F33A6CD5CD5C04D1459792DC5DF2" pageId="0" pageNumber="125">
<taxonomicName id="023A5DA363291FF5AFEE77306D612E24" baseAuthorityName="Brulle" baseAuthorityYear="1837" class="Insecta" family="Dytiscidae" genus="Cybister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybister puncticollis" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="E67D53561C1837C51FEDA456A44A52D1" italics="true" pageId="0" pageNumber="125">Cybister puncticollis</emphasis>
</taxonomicName>
: Bolivia, San Miguel.
<taxonomicName id="D541439E3D8845B7A7098A6B50326A7A" class="Insecta" family="Dytiscidae" genus="Cybister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybister kemneri" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="kemneri">
<emphasis id="F393344B0801F2F540063A2BFD977523" italics="true" pageId="0" pageNumber="125">Cybister kemneri</emphasis>
</taxonomicName>
: Brazil, La Plata, Amazonas, Rio Autaz.
</paragraph>
</subSubSection>
<subSubSection id="E0C267A527A275AC4124151144A2D780" pageId="0" pageNumber="125" type="type specimen">
<paragraph id="1B556CF322B096D911D0D652ACE6C6EE" pageId="0" pageNumber="125">Type specimen.</paragraph>
<paragraph id="F78A9792742EF4B43FC505C8AA23DFAC" pageId="0" pageNumber="125">
<taxonomicName id="ECE3244A192A2A6F12D76F2A07441906" baseAuthorityName="Brulle" baseAuthorityYear="1837" class="Insecta" family="Dytiscidae" genus="Cybister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybister puncticollis" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="5A17A0FF3D550BDDDB3936E6C7E383C3" italics="true" pageId="0" pageNumber="125">Cybister puncticollis</emphasis>
</taxonomicName>
, in MNHN (not examined).
<taxonomicName id="353D4DAAAF9D3F370D6492DB53D6961C" class="Insecta" family="Dytiscidae" genus="Cybister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybister kemneri" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="kemneri">
<emphasis id="9479794E9D6DE87F2262E2A1AAF845DA" italics="true" pageId="0" pageNumber="125">Cybister kemneri</emphasis>
</taxonomicName>
, in MZLU (not examined).
</paragraph>
</subSubSection>
<subSubSection id="0D952CFD2CCCB6947792C0EB4A59C7E1" pageId="0" pageNumber="125" type="diagnosis">
<paragraph id="728FB99E8CA3CFD23CE86446DE208F7D" pageId="0" pageNumber="125">Diagnosis.</paragraph>
<paragraph id="3100C789D14A049B2948B5CB3AC040A8" pageId="0" pageNumber="125">
This species is larger (Fig.
<figureCitation id="2B653990BCF3535E3EC6E88ADFFFCB1A" captionStart="Figures 9–14" captionStartId="F2" captionText="Figures 9 - 14. Neotropical Cybistrinae species, habitus 9 Cybister festae 10 Cybister puncticollis 11 Nilssondytes diversus 12 Megadytes parvus 13 Megadytes latus 14 species &quot; IR 57 &quot; (Ribera et al. 2008). Scale bar: 10.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures9-14" httpUri="https://binary.pensoft.net/fig/963584" pageId="0" pageNumber="125">10</figureCitation>
) than the other Neotropical species in the genus,
<taxonomicName id="5F0D6B3E0B5C8D58E5782FC0854F9E3B" lsidName="C. festae" pageId="0" pageNumber="125" rank="species" species="festae">
<emphasis id="64A1BF59FF747348588E585BB52F32AE" italics="true" pageId="0" pageNumber="125">C. festae</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="8AB666DEFD2151A5F89760E3AF20EA04" captionStart="Figures 9–14" captionStartId="F2" captionText="Figures 9 - 14. Neotropical Cybistrinae species, habitus 9 Cybister festae 10 Cybister puncticollis 11 Nilssondytes diversus 12 Megadytes parvus 13 Megadytes latus 14 species &quot; IR 57 &quot; (Ribera et al. 2008). Scale bar: 10.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures9-14" httpUri="https://binary.pensoft.net/fig/963584" pageId="0" pageNumber="125">9</figureCitation>
) and the male genitalia are distinctly different (Figs
<figureCitation id="456342BEFC2F628DC0433D51BAE6F4D1" captionStart="Figures 30–50" captionStartId="F4" captionText="Figures 30 - 50. Neotropical Cybistrinae species, male genitalia 30 - 32 Cybister festae: 30 median lobe, right lateral aspect 31 median lobe, ventral aspect 32 right lateral lobe, right lateral aspect 33 - 35 Cybister puncticollis: 33 median lobe, right lateral aspect 34 median lobe, ventral aspect 35 right lateral lobe, right lateral aspect 36 - 38 Nilssondytes diversus: 36 median lobe, right lateral aspect 37 median lobe, ventral aspect 38 right lateral lobe, right lateral aspect 39 - 43 Megadytes parvus: 39 median lobe, right lateral aspect 40 median lobe, ventral aspect 41 right lateral lobe, right lateral aspect 42 apex of median lobe, right lateral aspect 43 apex of median lobe, ventral aspect 44 - 47 Megadytes latus: 44 median lobe, right lateral aspect 45 median lobe, ventral aspect 46 right lateral lobe, right lateral aspect 47 apex of median lobe, ventral aspect 48 - 50 species &quot; IR 57 &quot; (Ribera et al. 2008): 48 median lobe, right lateral aspect 49 median lobe, ventral aspect 50 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1188.110081.figures30-50" httpUri="https://binary.pensoft.net/fig/963586" pageId="0" pageNumber="125">33-35</figureCitation>
). See above under that species for details about this and additional diagnostic differences between these two species of
<taxonomicName id="4F977BF23A04E8057CF9B3810DBBAE97" authorityName="Curtis" authorityYear="1827" class="Insecta" family="Dytiscidae" genus="Cybister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybister" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="E06F5DA0CAF113CCC5ECBCFB19C72643" italics="true" pageId="0" pageNumber="125">Cybister</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="38B8A244B1A0954F71B4A9284AADED2C" pageId="0" pageNumber="125" type="description">
<paragraph id="DF230DCEBD31A497593F0AD97A8E71CE" pageId="0" pageNumber="125">Description.</paragraph>
<paragraph id="B20963A548AE267E7D1EB9E74FA8A907" pageId="0" pageNumber="125">
<emphasis id="B549E9BFB7954EC60E7A44183CA799AA" bold="true" pageId="0" pageNumber="125">
<emphasis id="75B087C7DE20E4FDB898011A03A863AE" italics="true" pageId="0" pageNumber="125">Measurements</emphasis>
.
</emphasis>
TL = 26.6-27.6 mm, GW = 15.0-15.9 mm, PW = 10.8-11.6 mm, HW = 6.5-6.8 mm, EW = 3.7-3.8 mm, TL/GW = 1.7-1.8, HW/EW = 1.7-1.8, WC/WV = 2.9-3.2. Body shape oval, widest slightly posterad of middle; lateral margins broadly curved, continuously curved between pronotum and elytron. Depressed and somewhat flattened in lateral aspect.
</paragraph>
<paragraph id="C300D706E3B95116AAB431CB21FF2B59" pageId="0" pageNumber="125">
<emphasis id="A10312500D90BF5336CEB884695030C1" bold="true" pageId="0" pageNumber="125">
<emphasis id="7228C3568CA6A9636E27837004531630" italics="true" pageId="0" pageNumber="125">Coloration</emphasis>
.
</emphasis>
Head dark green, clypeus and labrum pale yellow. Pronotum dark green, with broad lateral yellow marginal band (Fig.
<figureCitation id="6C6C9BE6C2DE25E4D6382719C2035F40" captionStart="Figures 9–14" captionStartId="F2" captionText="Figures 9 - 14. Neotropical Cybistrinae species, habitus 9 Cybister festae 10 Cybister puncticollis 11 Nilssondytes diversus 12 Megadytes parvus 13 Megadytes latus 14 species &quot; IR 57 &quot; (Ribera et al. 2008). Scale bar: 10.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures9-14" httpUri="https://binary.pensoft.net/fig/963584" pageId="0" pageNumber="125">10</figureCitation>
), medial margin of band distinctly delimited, band separated from posterolateral margin by narrow green strip (Fig.
<figureCitation id="FF95630992E89AB00B5AF4B3662EE1A6" captionStart="Figures 9–14" captionStartId="F2" captionText="Figures 9 - 14. Neotropical Cybistrinae species, habitus 9 Cybister festae 10 Cybister puncticollis 11 Nilssondytes diversus 12 Megadytes parvus 13 Megadytes latus 14 species &quot; IR 57 &quot; (Ribera et al. 2008). Scale bar: 10.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures9-14" httpUri="https://binary.pensoft.net/fig/963584" pageId="0" pageNumber="125">10</figureCitation>
). Elytron dark green with broad lateral yellow marginal band, medial margin of band distinctly delimited, lateral margin of band slightly remote from lateral elytral margin for medial portion of length, extending laterally to margin of elytron anteriorly and posteriorly, apex of yellow band diffusing into coloration of elytron, less distinct (Fig.
<figureCitation id="F1F93F083717D19FE8720CD6EA823802" captionStart="Figures 9–14" captionStartId="F2" captionText="Figures 9 - 14. Neotropical Cybistrinae species, habitus 9 Cybister festae 10 Cybister puncticollis 11 Nilssondytes diversus 12 Megadytes parvus 13 Megadytes latus 14 species &quot; IR 57 &quot; (Ribera et al. 2008). Scale bar: 10.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures9-14" httpUri="https://binary.pensoft.net/fig/963584" pageId="0" pageNumber="125">10</figureCitation>
). Ventral surfaces mostly black except antennae and palpi orange, pro- and mesothoracic legs mostly orange, tarsi dark orange, metathoracic legs testaceous, anterodorsal surface of tibia orange, propleuron and elytral epipleuron pale orange to testaceous, and with small orange maculae laterally on abdominal ventrites IV-VI.
</paragraph>
<paragraph id="9F57EEBECB72FAAF954D8CA90FA94643" pageId="0" pageNumber="125">
<emphasis id="F38EB5C32F9AD9A8AF84F8AF5A0B865A" bold="true" pageId="0" pageNumber="125">
<emphasis id="711AD85A254938C49C1375B19A30234D" italics="true" pageId="0" pageNumber="125">Sculpture and structure</emphasis>
.
</emphasis>
Head broad, anteriorly produced, eyes prominent; dorsal surface evenly covered with exceptionally fine micropunctures but smooth and shiny. Pronotum with lateral margins evenly and shallowly curved; surface of pronotum similar to surface of head in micropunctation; pronotum dorsally evenly curved. Elytron with margins very broadly curved, more strongly curved posteriorly; surface similar to surface of head in micropunctation and surface appearance. Prosternal process anteriorly distinctly emarginate medially, remaining surface nearly flat throughout and moderately broad, apex broadly elongate and sharply pointed. Metaventral wing narrow (WC/WV = 2.9-3.2); surface smooth, with extremely fine microsculpture, shiny. Lateral portion of metacoxa large, broad, surface smooth and shiny, with extremely fine microsculpture; metacoxal lines short and curved, extending anteriorly less than half distance across metacoxa. Abdominal ventrites smooth, relatively unsculptured.
</paragraph>
<paragraph id="D1BC8531988ABCEAF20535906D02DFFB" pageId="0" pageNumber="125">
<emphasis id="95ADF453CEF36375D8CABAD9B4466614" bold="true" pageId="0" pageNumber="125">
<emphasis id="243A53DAD37844AC272C3B76E419E109" italics="true" pageId="0" pageNumber="125">Male genitalia</emphasis>
.
</emphasis>
Male median lobe in lateral aspect moderately slender throughout, evenly curved, apically abruptly expanded with a narrowly rounded apex, ventral sclerite slender and evenly curved basally, apically straight and apically pointed (Fig.
<figureCitation id="F6B3D455D4A7B15509F777E02651B80B" captionStart="Figures 30–50" captionStartId="F4" captionText="Figures 30 - 50. Neotropical Cybistrinae species, male genitalia 30 - 32 Cybister festae: 30 median lobe, right lateral aspect 31 median lobe, ventral aspect 32 right lateral lobe, right lateral aspect 33 - 35 Cybister puncticollis: 33 median lobe, right lateral aspect 34 median lobe, ventral aspect 35 right lateral lobe, right lateral aspect 36 - 38 Nilssondytes diversus: 36 median lobe, right lateral aspect 37 median lobe, ventral aspect 38 right lateral lobe, right lateral aspect 39 - 43 Megadytes parvus: 39 median lobe, right lateral aspect 40 median lobe, ventral aspect 41 right lateral lobe, right lateral aspect 42 apex of median lobe, right lateral aspect 43 apex of median lobe, ventral aspect 44 - 47 Megadytes latus: 44 median lobe, right lateral aspect 45 median lobe, ventral aspect 46 right lateral lobe, right lateral aspect 47 apex of median lobe, ventral aspect 48 - 50 species &quot; IR 57 &quot; (Ribera et al. 2008): 48 median lobe, right lateral aspect 49 median lobe, ventral aspect 50 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1188.110081.figures30-50" httpUri="https://binary.pensoft.net/fig/963586" pageId="0" pageNumber="125">33</figureCitation>
); in dorsal aspect median lobe moderately broad basally, lateral margins broadly sinuate, apex broadly expanded, with apex broadly sub-truncate, medially broadly pointed (Fig.
<figureCitation id="A503C7DA535D650FFBBAD195C0FCD15C" captionStart="Figures 30–50" captionStartId="F4" captionText="Figures 30 - 50. Neotropical Cybistrinae species, male genitalia 30 - 32 Cybister festae: 30 median lobe, right lateral aspect 31 median lobe, ventral aspect 32 right lateral lobe, right lateral aspect 33 - 35 Cybister puncticollis: 33 median lobe, right lateral aspect 34 median lobe, ventral aspect 35 right lateral lobe, right lateral aspect 36 - 38 Nilssondytes diversus: 36 median lobe, right lateral aspect 37 median lobe, ventral aspect 38 right lateral lobe, right lateral aspect 39 - 43 Megadytes parvus: 39 median lobe, right lateral aspect 40 median lobe, ventral aspect 41 right lateral lobe, right lateral aspect 42 apex of median lobe, right lateral aspect 43 apex of median lobe, ventral aspect 44 - 47 Megadytes latus: 44 median lobe, right lateral aspect 45 median lobe, ventral aspect 46 right lateral lobe, right lateral aspect 47 apex of median lobe, ventral aspect 48 - 50 species &quot; IR 57 &quot; (Ribera et al. 2008): 48 median lobe, right lateral aspect 49 median lobe, ventral aspect 50 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1188.110081.figures30-50" httpUri="https://binary.pensoft.net/fig/963586" pageId="0" pageNumber="125">34</figureCitation>
). Lateral lobe moderately broad throughout length, evenly curved, apex narrowly rounded, with long series of setae along dorsal margin (Fig.
<figureCitation id="916B5039F049525BE14648CBE16E087C" captionStart="Figures 30–50" captionStartId="F4" captionText="Figures 30 - 50. Neotropical Cybistrinae species, male genitalia 30 - 32 Cybister festae: 30 median lobe, right lateral aspect 31 median lobe, ventral aspect 32 right lateral lobe, right lateral aspect 33 - 35 Cybister puncticollis: 33 median lobe, right lateral aspect 34 median lobe, ventral aspect 35 right lateral lobe, right lateral aspect 36 - 38 Nilssondytes diversus: 36 median lobe, right lateral aspect 37 median lobe, ventral aspect 38 right lateral lobe, right lateral aspect 39 - 43 Megadytes parvus: 39 median lobe, right lateral aspect 40 median lobe, ventral aspect 41 right lateral lobe, right lateral aspect 42 apex of median lobe, right lateral aspect 43 apex of median lobe, ventral aspect 44 - 47 Megadytes latus: 44 median lobe, right lateral aspect 45 median lobe, ventral aspect 46 right lateral lobe, right lateral aspect 47 apex of median lobe, ventral aspect 48 - 50 species &quot; IR 57 &quot; (Ribera et al. 2008): 48 median lobe, right lateral aspect 49 median lobe, ventral aspect 50 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1188.110081.figures30-50" httpUri="https://binary.pensoft.net/fig/963586" pageId="0" pageNumber="125">35</figureCitation>
).
</paragraph>
<paragraph id="5F1A89E25ED0808130CC4E98E98867FE" pageId="0" pageNumber="125">
<emphasis id="55AF7A4D841F8EB423B3060BA13F4708" bold="true" pageId="0" pageNumber="125">
<emphasis id="51EB4A121183B32A1599A3F03A79A1D8" italics="true" pageId="0" pageNumber="125">Female genitalia</emphasis>
.
</emphasis>
Vagina extremely elongate, slender; with enlarged area at base of common oviduct and spermatheca, with enlarged lateral sacs on this enlarged area; spermatheca extremely slender and elongate (Fig.
<figureCitation id="EA017BF7F95278065D0F1A39E9C7E3C5" captionStart="Figures 58–61" captionStartId="F6" captionText="Figures 58 - 61. Neotropical Cybistrinae species, female genitalia including left gonocoxosternite, ventral aspect, except Fig. 60 without internal genitalia 58 Cybister festae 59 Cybister puncticollis 60 Megadytes latus 61 Nilssondytes diversus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures58-61" httpUri="https://binary.pensoft.net/fig/963588" pageId="0" pageNumber="125">59</figureCitation>
); gonocoxae together slender and apically pointed (Fig.
<figureCitation id="FA5CE39EF6020B481D052140F2A8E095" captionStart="Figures 58–61" captionStartId="F6" captionText="Figures 58 - 61. Neotropical Cybistrinae species, female genitalia including left gonocoxosternite, ventral aspect, except Fig. 60 without internal genitalia 58 Cybister festae 59 Cybister puncticollis 60 Megadytes latus 61 Nilssondytes diversus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures58-61" httpUri="https://binary.pensoft.net/fig/963588" pageId="0" pageNumber="125">59</figureCitation>
); gonocoxosternite broad, anterolateral lobe broad, medial margin distinctly, narrowly emarginate, basomedially with field of setae (Fig.
<figureCitation id="E5CE17A4FB30B227C7EFFE686A42EA23" captionStart="Figures 58–61" captionStartId="F6" captionText="Figures 58 - 61. Neotropical Cybistrinae species, female genitalia including left gonocoxosternite, ventral aspect, except Fig. 60 without internal genitalia 58 Cybister festae 59 Cybister puncticollis 60 Megadytes latus 61 Nilssondytes diversus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures58-61" httpUri="https://binary.pensoft.net/fig/963588" pageId="0" pageNumber="125">59</figureCitation>
).
</paragraph>
<paragraph id="7275FC4BD0BBD38150251CCF9FCAC6B6" pageId="0" pageNumber="125">
<emphasis id="52CDCD10483518494AFEAA070E08CB76" bold="true" pageId="0" pageNumber="125">
<emphasis id="18B8BCC0754554259EA2EE1CF5AD85E0" italics="true" pageId="0" pageNumber="125">Sexual dimorphism</emphasis>
.
</emphasis>
Males have a broad protarsal palette with ventral adhesive setae with apical structures in the form of elongate flattened spatulate surfaces; males also have mesotarsomeres with posteroventral fields of setae which are absent in females. Females lack natatory setae along the ventral margins of the metatarsomeres, but these are present in males. Males have a single metatarsal claw (the anterior claw, Fig.
<figureCitation id="8680DCE311D14398ACAE0358446C2E3C" captionStart="Figures 15–29" captionStartId="F3" captionText="Figures 15 - 29. Neotropical Cybistrinae species, metatarsal claws and tarsomere VI of males and females 15, 16 Cybister festae 17, 18 Cybister puncticollis 19, 20 Nilssondytes diversus 21, 22 Megadytes latus 23, 24 Megadytes parvus 25 species &quot; IR 57 &quot; (Ribera et al. 2008) 26, 27 Paramegadytes glaucus 28, 29 Metaxydytes fraternus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures15-29" httpUri="https://binary.pensoft.net/fig/963585" pageId="0" pageNumber="125">17</figureCitation>
), whereas females have a second, posterior claw that is ~ 1/2 the length of the anterior, is abruptly curved apically and sharply pointed (Fig.
<figureCitation id="F0107EAABEEBB06D4DFEE77D39029457" captionStart="Figures 15–29" captionStartId="F3" captionText="Figures 15 - 29. Neotropical Cybistrinae species, metatarsal claws and tarsomere VI of males and females 15, 16 Cybister festae 17, 18 Cybister puncticollis 19, 20 Nilssondytes diversus 21, 22 Megadytes latus 23, 24 Megadytes parvus 25 species &quot; IR 57 &quot; (Ribera et al. 2008) 26, 27 Paramegadytes glaucus 28, 29 Metaxydytes fraternus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures15-29" httpUri="https://binary.pensoft.net/fig/963585" pageId="0" pageNumber="125">18</figureCitation>
).
</paragraph>
<paragraph id="A4E1A89D23F9384E5E579D7C48A95718" pageId="0" pageNumber="125">
<emphasis id="29708DBA057FC4A308BF3BD326A3FEB2" bold="true" pageId="0" pageNumber="125">
<emphasis id="5D282BD94D7DBE397FAF511772C9007C" italics="true" pageId="0" pageNumber="125">Variation</emphasis>
.
</emphasis>
Specimens vary somewhat in size and coloration, with some specimens more greenish and others darker, less greenish.
</paragraph>
</subSubSection>
<subSubSection id="88115869A76B2A464E7FFD5A8062CAC1" pageId="0" pageNumber="125" type="distribution">
<paragraph id="B1C28BE5C21729BCFBAD7C7CDA98B608" pageId="0" pageNumber="125">Distribution.</paragraph>
<paragraph id="9153ADA0820EC71F45E1A280A85D20A7" pageId="0" pageNumber="125">This species is known from Argentina, Bolivia, Brazil, French Guyana, and Peru. There is also a previously unpublished record from Paraguay.</paragraph>
</subSubSection>
<subSubSection id="DC5EFAAF6F4C9A35DFD0F87ED59508A2" pageId="0" pageNumber="125" type="materials_examined">
<paragraph id="B940E42A71578C4A3164BDDBF3CE2F29" pageId="0" pageNumber="125">Material examined.</paragraph>
<paragraph id="9F2B69FD7D08F2E74EC40EC418EA303A" pageId="0" pageNumber="125">
Specimens were examined from
<collectingCountry id="CD8F53658CE9C5B26D537C4B78DD88A3" name="Argentina">Argentina</collectingCountry>
,
<collectingCountry id="F202DF6CE505C06F3BFEB3FF4862BC0E" name="Bolivia">Bolivia</collectingCountry>
,
<collectingCountry id="DB8AD9152134A843A0956F277D7BFD30" name="Peru">Peru</collectingCountry>
, and
<collectingCountry id="9F717BAD2C047819865C38A90B817CBF" name="Paraguay">Paraguay</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

113
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<paragraph id="2035DBBC532F3AC94EE53F2A2A64BD9A" pageNumber="638">
<taxonomicName id="745415F9BD4D51081E044AEDD4F6BBA2" ID-CoL="523H" authority="Thomas 1904" authorityName="Thomas" authorityYear="1904" class="Mammalia" family="Suidae" genus="Hylochoerus" kingdom="Animalia" order="Artiodactyla" pageNumber="638" phylum="Chordata" rank="genus">
<emphasis id="7F683C3E6828CD75CCE40646FB6D86F2" italics="true" pageNumber="638">Hylochoerus</emphasis>
Thomas 1904
</taxonomicName>
</paragraph>
</subSubSection>
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<paragraph id="5779D03E6E0E74DB88F090EDDBDA3F39" pageNumber="638">
<treatmentCitation id="FF43A504B64F162094B774C91EA1676D" author="Thomas" authority="Thomas 1904" authorityName="Thomas" authorityYear="1904" class="Mammalia" family="Suidae" genus="Hylochoerus" journal="Nature" kingdom="Animalia" order="Artiodactyla" page="577" pageNumber="638" phylum="Chordata" rank="genus" title="Hylochoerus" volume="70" year="1904">
<taxonomicName id="B56B5B466DF109C576E612715404214D" ID-CoL="523H" authority="Thomas 1904" authorityName="Thomas" authorityYear="1904" class="Mammalia" family="Suidae" genus="Hylochoerus" kingdom="Animalia" order="Artiodactyla" pageNumber="638" phylum="Chordata" rank="genus">
<emphasis id="4633ED3F120747EB0A7B90B9608FA450" italics="true" pageNumber="638">Hylochoerus</emphasis>
Thomas 1904
</taxonomicName>
,
<bibCitation id="6718219AFFB52AA4D8655777DEAAD87D" author="Thomas" journal="Nature" pageNumber="638" pagination="577" title="Hylochoerus" volume="70" year="1904">Nature, 70: 577</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C449304AD0CD547ED751DF598D89975C" pageNumber="638" type="type_taxon">
<paragraph id="B5D3421ABBCA253A98B60A69CD7BF28C" pageNumber="638">
<emphasis id="64FE353AA27D98C7DBE0CC8750208B1B" inLineHeading="true" pageNumber="638">Type Species:</emphasis>
<taxonomicName id="C7066341C183184EFA700A35EEC883A1" authority="Thomas 1904" authorityName="Thomas" authorityYear="1904" class="Mammalia" family="Suidae" genus="Hylochoerus" kingdom="Animalia" order="Artiodactyla" pageNumber="638" phylum="Chordata" rank="species" species="meinertzhageni">
<emphasis id="FC4F0819BA6E91A97D0E6D68AB48BDF7" italics="true" pageNumber="638">Hylochoerus meinertzhageni</emphasis>
Thomas 1904
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="1A8E18B69E15DE0A7BF532531BB3BF81" pageNumber="638" type="child_taxa">
<paragraph id="46F63D1445E7050DE1770AD8C72C78A3" pageNumber="638">
<emphasis id="99BE429B4C6C1BADD8F677D6F8EA96C2" inLineHeading="true" pageNumber="638">Species and subspecies:</emphasis>
1 species with 3 subspecies:
</paragraph>
<paragraph id="B2EB9090AE7BD30AB766640BE42AB9C8" pageNumber="638">
Species
<taxonomicName id="E4602239F800482CCED6730C142ABD6A" authority="Thomas 1904" authorityName="Thomas" authorityYear="1904" class="Mammalia" family="Suidae" genus="Hylochoerus" kingdom="Animalia" order="Artiodactyla" pageNumber="638" phylum="Chordata" rank="species" species="meinertzhageni">
<emphasis id="8929094569469C1FAC551E6344F5740F" italics="true" pageNumber="638">Hylochoerus meinertzhageni</emphasis>
Thomas 1904
</taxonomicName>
</paragraph>
<paragraph id="03CCB36588453D7C31DC8F223E2E6087" pageNumber="638">
Subspecies
<taxonomicName id="F18805197D1EF72F01182A25F3720E2B" authority="Thomas 1904" authorityName="Thomas" authorityYear="1904" class="Mammalia" family="Suidae" genus="Hylochoerus" kingdom="Animalia" order="Artiodactyla" pageNumber="638" phylum="Chordata" rank="subSpecies" species="meinertzhageni" subSpecies="meinertzhageni">
<emphasis id="8727D0089E5AAA3128CCFD9E8D650A82" italics="true" pageNumber="638">Hylochoerus meinertzhageni</emphasis>
subsp.
<emphasis id="8AF9BAC8E162C46ED4818E67EF1E7F08" italics="true" pageNumber="638">meinertzhageni</emphasis>
Thomas 1904
</taxonomicName>
</paragraph>
<paragraph id="FC6832A71B399996F62526B9B776A029" pageNumber="638">
Subspecies
<taxonomicName id="671E2801740647A54126F9B3DE72AE21" authority="Thomas 1906" authorityName="Thomas" authorityYear="1906" class="Mammalia" family="Suidae" genus="Hylochoerus" kingdom="Animalia" order="Artiodactyla" pageNumber="638" phylum="Chordata" rank="subSpecies" species="meinertzhageni" subSpecies="rimator">
<emphasis id="220CFBD5D4C757E7FC81AF00EEA20E0C" italics="true" pageNumber="638">Hylochoerus meinertzhageni</emphasis>
subsp.
<emphasis id="B36BBF078D77066F015E9B3C7C6FD65C" italics="true" pageNumber="638">rimator</emphasis>
Thomas 1906
</taxonomicName>
</paragraph>
<paragraph id="5902BB78C4E580B110EEF46270BF05BB" pageNumber="638">
Subspecies
<taxonomicName id="9190CF70F5B87C2D8536ADB8356930B5" authority="Bouet and Neuville 1930" authorityName="Bouet and Neuville" authorityYear="1930" class="Mammalia" family="Suidae" genus="Hylochoerus" kingdom="Animalia" order="Artiodactyla" pageNumber="638" phylum="Chordata" rank="subSpecies" species="meinertzhageni" subSpecies="ivoriensis">
<emphasis id="6047EF010574995D0BA0BE0CE47B66F6" italics="true" pageNumber="638">Hylochoerus meinertzhageni</emphasis>
subsp.
<emphasis id="C7793CC1F2CADC9A13254FD5A8958790" italics="true" pageNumber="638">ivoriensis</emphasis>
Bouet and Neuville 1930
</taxonomicName>
</paragraph>
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