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<document id="3D4F59677F5ACD22F311B8B9CE980FBC" ID-DOI="10.5852/ejt.2025.989.2887" ID-ISSN="2118-9773" ID-Zenodo-Dep="15351100" ID-ZooBank="C2687BEC-66D1-4516-B847-56977033AA49" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1746545402734" checkinUser="plazi" docAuthor="Johnston, Nikolas P., Szpila, Krzysztof, Pape, Thomas, Meiklejohn, Kelly A., Foley, Liam B. &amp; Wallman, James F." docDate="2025" docId="03C3E220FFE5FFF1D0BABC64FAFEE801" docLanguage="en" docName="EJT.2025.989.94-118.pdf" docOrigin="European Journal of Taxonomy 989" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2887/13093" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Calliphora clarki Malloch 1927" docType="key" docVersion="1" lastPageNumber="100" masterDocId="FFFA9A58FFE3FFF7D007BE13FF89EF4C" masterDocTitle="A species-group framework to unravel blowfly diversity: integrative revision of the Calliphora clarki-group (Diptera: Calliphoridae)" masterLastPageNumber="118" masterPageNumber="94" pageNumber="100" updateTime="1746708283737" updateUser="guilherme" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F691405ACED54DF6F3952E71BA75B68C">A species-group framework to unravel blowfly diversity: integrative revision of the Calliphora clarki-group (Diptera: Calliphoridae)</mods:title>
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<mods:namePart id="7E131A694ABA9690C92FE1A4422E00D6">Johnston, Nikolas P.</mods:namePart>
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<mods:affiliation id="04B61BFD641E007597F2EEF59A6A3870">Molecular Horizons, School of Science, University of Wollongong Northfields Ave, Wollongong, NSW, 2500, Australia. &amp; School of Life Sciences, University of Technology Sydney, 15 Broadway, Ultimo, NSW, 2007, Australia.</mods:affiliation>
<mods:nameIdentifier id="FA081B47E3876F16127CCAB52ECC72E6" type="email">nikolaspjohnston@gmail.com</mods:nameIdentifier>
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<mods:namePart id="09442FFBB8807A7F2575B6B21D5630FC">Szpila, Krzysztof</mods:namePart>
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<mods:affiliation id="20178B0814AAD89FA274565E44FDD989">Department of Ecology and Biogeography, Nicolaus Copernicus University in Toruń, Lwowska 1, 87 - 100 Toruń, Poland.</mods:affiliation>
<mods:nameIdentifier id="68DF7936DA7E93137024ECB2C11289CF" type="email">szpila@umk.pl</mods:nameIdentifier>
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<mods:namePart id="89C2D7806128FDD808121B25F8B99EFC">Pape, Thomas</mods:namePart>
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<mods:affiliation id="EA472D1A1534206FC94BAD288B7A44C1">Natural History Museum of Denmark, Universitetsparken 15, 2100 Copenhagen, Denmark.</mods:affiliation>
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<mods:namePart id="882E3734A187502C15CD9892E70316B5">Meiklejohn, Kelly A.</mods:namePart>
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<mods:affiliation id="D12F610958822AFE36ECBB980803DED8">Department of Population Health and Pathobiology, North Carolina State University, 1060 William Moore Drive, Raleigh, NC, 27606 USA.</mods:affiliation>
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<mods:namePart id="DF0442EEF3579220591D8A7F58A49F49">Foley, Liam B.</mods:namePart>
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<mods:affiliation id="04239628CB52564796F775957AE6C0EB">Molecular Horizons, School of Science, University of Wollongong Northfields Ave, Wollongong, NSW, 2500, Australia. &amp; School of Life Sciences, University of Technology Sydney, 15 Broadway, Ultimo, NSW, 2007, Australia.</mods:affiliation>
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<mods:namePart id="4AA267770D95C12EC82946FE4656AF28">Wallman, James F.</mods:namePart>
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<mods:date id="BAA49FD31B03C19FA0EBF04C90DFF124">2025</mods:date>
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<treatment id="03C3E220FFE5FFF1D0BABC64FAFEE801" LSID="urn:lsid:plazi:treatment:03C3E220FFE5FFF1D0BABC64FAFEE801" httpUri="http://treatment.plazi.org/id/03C3E220FFE5FFF1D0BABC64FAFEE801" lastPageNumber="100" pageId="6" pageNumber="100">
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<emphasis id="B91E8F24FFE5FFF1D0BABC64FDAFEDDD" bold="true" box="[189,550,630,657]" pageId="6" pageNumber="100">
Diagnosis of the
<taxonomicName id="4C6A28B5FFE5FFF1D18CBC65FE58EDDD" ID-CoL="8B9ZX" authorityName="Malloch" authorityYear="1927" box="[395,465,630,657]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="clarki">
<emphasis id="B91E8F24FFE5FFF1D18CBC65FE58EDDD" bold="true" box="[395,465,630,657]" italics="true" pageId="6" pageNumber="100">clarki</emphasis>
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-group
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The
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-species-group can be separated from all other Australian
<taxonomicName id="4C6A28B5FFE5FFF1D3CFBCB7FBE1EDF2" box="[968,1128,676,702]" class="Insecta" family="Calliphoridae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="subFamily" subFamily="Calliphorinae">Calliphorinae</taxonomicName>
by the presence of (1) a reduced anterior spiracle, smaller at widest part than width of proepisternum (
<figureCitation id="13514FB3FFE5FFF1D45EBCD6FB55EDAC" box="[1113,1244,709,736]" captionStart="Fig" captionStartId="13.[189,232,1928,1954]" captionTargetBox="[200,1398,639,1887]" captionTargetId="figure-99@13.[189,1398,639,1887]" captionTargetPageId="13" captionText="Fig. 3. Calliphora ignicera sp. nov., paratype, ♀, Flinders Is., TAS, 20 May 1991, C. Tann leg. (ANIC). A. Habitus, dorsal view. B. Head, anterior view. C. Habitus, lateral view. D. Calypter, lateral view. Black scale bars = 1 mm; red scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351108" httpUri="https://zenodo.org/record/15351108/files/figure.png" pageId="6" pageNumber="100">Fig. 3C, D</figureCitation>
); (2) hyaline or lightly coloured upper calypter with contrasting dark brown or orange fringe in combination with an infuscated lower calypter (
<figureCitation id="13514FB3FFE5FFF1D1F2BD1AFDDAEC6F" box="[501,595,777,803]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="6" pageNumber="100">Figs 1E</figureCitation>
,
<figureCitation id="13514FB3FFE5FFF1D267BD1AFD0DEC6F" box="[608,644,777,803]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetBox="[255,1343,265,1730]" captionTargetId="figure-17@15.[245,1343,265,1744]" captionTargetPageId="15" captionText="Fig. 4. A–C. Calliphora niveata sp. nov., paratype, ♂, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Paratype, ♂, Guthega, NSW, 4 Apr. 1984, K.R. Norris leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = plural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351112" httpUri="https://zenodo.org/record/15351112/files/figure.png" pageId="6" pageNumber="100">4E</figureCitation>
); (3) postpedicel at least partially orange (
<figureCitation id="13514FB3FFE5FFF1D474BD1AFB5BEC6F" box="[1139,1234,777,803]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="6" pageNumber="100">Figs 1B</figureCitation>
,
<figureCitation id="13514FB3FFE5FFF1D4E7BD1AFA8CEC6F" box="[1248,1285,777,803]" captionStart="Fig" captionStartId="11.[189,232,1857,1883]" captionTargetBox="[229,1369,265,1800]" captionTargetId="figure-17@11.[218,1369,265,1815]" captionTargetPageId="11" captionText="Fig. 2. A–C, F–H. Calliphora ampyx sp. nov., paratype, ♂, Canberra, ACT, 6 Mar. 1984, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Mt Stromlo, ACT, 26 Oct. 1993, D.K. Yeates leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351104" httpUri="https://zenodo.org/record/15351104/files/figure.png" pageId="6" pageNumber="100">2B</figureCitation>
,
<figureCitation id="13514FB3FFE5FFF1D516BD1AFABFEC6F" box="[1297,1334,777,803]" captionStart="Fig" captionStartId="13.[189,232,1928,1954]" captionTargetBox="[200,1398,639,1887]" captionTargetId="figure-99@13.[189,1398,639,1887]" captionTargetPageId="13" captionText="Fig. 3. Calliphora ignicera sp. nov., paratype, ♀, Flinders Is., TAS, 20 May 1991, C. Tann leg. (ANIC). A. Habitus, dorsal view. B. Head, anterior view. C. Habitus, lateral view. D. Calypter, lateral view. Black scale bars = 1 mm; red scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351108" httpUri="https://zenodo.org/record/15351108/files/figure.png" pageId="6" pageNumber="100">3B</figureCitation>
); and (4) male terminalia with weak or no sclerotisation in distal part of mesohypophallus (
<figureCitation id="13514FB3FFE5FFF1D4AFBD39FA83EC08" box="[1192,1290,810,836]" captionStart="Fig" captionStartId="11.[189,232,1857,1883]" captionTargetBox="[229,1369,265,1800]" captionTargetId="figure-17@11.[218,1369,265,1815]" captionTargetPageId="11" captionText="Fig. 2. A–C, F–H. Calliphora ampyx sp. nov., paratype, ♂, Canberra, ACT, 6 Mar. 1984, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Mt Stromlo, ACT, 26 Oct. 1993, D.K. Yeates leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351104" httpUri="https://zenodo.org/record/15351104/files/figure.png" pageId="6" pageNumber="100">Figs 2H</figureCitation>
,
<figureCitation id="13514FB3FFE5FFF1D511BD39FABFEC08" box="[1302,1334,810,836]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetBox="[255,1343,265,1730]" captionTargetId="figure-17@15.[245,1343,265,1744]" captionTargetPageId="15" captionText="Fig. 4. A–C. Calliphora niveata sp. nov., paratype, ♂, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Paratype, ♂, Guthega, NSW, 4 Apr. 1984, K.R. Norris leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = plural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351112" httpUri="https://zenodo.org/record/15351112/files/figure.png" pageId="6" pageNumber="100">4F</figureCitation>
,
<figureCitation id="13514FB3FFE5FFF1D545BD39FAE0EC09" box="[1346,1385,810,837]" captionStart="Fig" captionStartId="17.[189,232,1822,1848]" captionTargetBox="[244,1354,268,1763]" captionTargetId="figure-17@17.[234,1354,268,1777]" captionTargetPageId="17" captionText="Fig. 5. A–C. Calliphora xanthocera Malloch, 1927, ♂, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view.D–E. Female, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male, Kosciuszko, NSW, 25 Mar. 1965, D.E. Havenstein leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = pleural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351114" httpUri="https://zenodo.org/record/15351114/files/figure.png" pageId="6" pageNumber="100">5H</figureCitation>
). The species of this group were previously placed under the genera
<taxonomicName id="4C6A28B5FFE5FFF1D3CABD5FFBA9EC2A" authorityName="Robineau-Desvoidy" authorityYear="1830" box="[973,1056,844,870]" class="Insecta" family="Calliphoridae" genus="Onesia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="B91E8F24FFE5FFF1D3CABD5FFBA9EC2A" box="[973,1056,844,870]" italics="true" pageId="6" pageNumber="100">Onesia</emphasis>
</taxonomicName>
and
<emphasis id="B91E8F24FFE5FFF1D45EBD5FFB6BEC2A" box="[1113,1250,844,870]" italics="true" pageId="6" pageNumber="100">
<taxonomicName id="4C6A28B5FFE5FFF1D45EBD5FFB57EC2A" authorityName="Robineau-Desvoidy" authorityYear="1830" box="[1113,1246,844,870]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="genus">Calliphora</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37000BDFFE5FFF1D0BABD9FFD84ECB8" pageId="6" pageNumber="100" type="multiple">
<paragraph id="8BD55336FFE5FFF1D0BABD9FFD32ECEA" blockId="6.[189,699,907,934]" box="[189,699,907,934]" pageId="6" pageNumber="100">
<heading id="D09DE45AFFE5FFF1D0BABD9FFD32ECEA" bold="true" box="[189,699,907,934]" fontSize="11" level="3" pageId="6" pageNumber="100" reason="3">
<emphasis id="B91E8F24FFE5FFF1D0BABD9FFD32ECEA" bold="true" box="[189,699,907,934]" pageId="6" pageNumber="100">
Key to species of the
<taxonomicName id="4C6A28B5FFE5FFF1D1C5BD98FD81ECEA" authorityName="Malloch" authorityYear="1927" box="[450,520,907,934]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="clarki">
<emphasis id="B91E8F24FFE5FFF1D1C5BD98FD81ECEA" bold="true" box="[450,520,907,934]" italics="true" pageId="6" pageNumber="100">clarki</emphasis>
</taxonomicName>
species-group
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD55336FFE5FFF1D0BABDAAFD84ECB8" blockId="6.[189,1399,953,1012]" pageId="6" pageNumber="100">
Abbreviations for states and territories are provided within couplets for species that are restricted to specific regions of
<collectingCountry id="F37D13A6FFE5FFF1D19DBDC9FD80ECB8" box="[410,521,986,1012]" name="Australia" pageId="6" pageNumber="100">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37000BDFFE5FFF1D0BABA09FAFEE801" pageId="6" pageNumber="100" type="key">
<key id="FFFB0407FFE5FFF1D0BABA09FAFEE801" pageId="6" pageNumber="100">
<keyStep id="309E4B93FFE5FFF1D0BABA09FAFEEB1A" pageId="6" pageNumber="100">
<paragraph id="8BD55336FFE5FFF1D0BABA09FAFEEB78" blockId="6.[189,1399,1050,1110]" box="[189,1399,1050,1077]" pageId="6" pageNumber="100">
<keyLead id="309BF003FFE5FFF1D0BABA09FAFEEB78" box="[189,1399,1050,1077]" pageId="6" pageNumber="100">
1. Basicosta ground colour black (
<figureCitation id="13514FB3FFE5FFF1D259BA09FD34EB78" box="[606,701,1050,1076]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetBox="[255,1343,265,1730]" captionTargetId="figure-17@15.[245,1343,265,1744]" captionTargetPageId="15" captionText="Fig. 4. A–C. Calliphora niveata sp. nov., paratype, ♂, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Paratype, ♂, Guthega, NSW, 4 Apr. 1984, K.R. Norris leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = plural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351112" httpUri="https://zenodo.org/record/15351112/files/figure.png" pageId="6" pageNumber="100">Figs 4E</figureCitation>
,
<figureCitation id="13514FB3FFE5FFF1D2CEBA09FD63EB79" box="[713,746,1050,1077]" captionStart="Fig" captionStartId="17.[189,232,1822,1848]" captionTargetBox="[244,1354,268,1763]" captionTargetId="figure-17@17.[234,1354,268,1777]" captionTargetPageId="17" captionText="Fig. 5. A–C. Calliphora xanthocera Malloch, 1927, ♂, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view.D–E. Female, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male, Kosciuszko, NSW, 25 Mar. 1965, D.E. Havenstein leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = pleural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351114" httpUri="https://zenodo.org/record/15351114/files/figure.png" pageId="6" pageNumber="100">5E</figureCitation>
) ................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BD55336FFE5FFF1D0C6BA2FFAFEEB1A" blockId="6.[189,1399,1050,1110]" box="[193,1399,1084,1110]" pageId="6" pageNumber="100">
<keyLead id="309BF003FFE5FFF1D0C6BA2FFAFEEB1A" box="[193,1399,1084,1110]" pageId="6" pageNumber="100">
– Basicosta ground colour brown, orange or yellow (
<figureCitation id="13514FB3FFE5FFF1D338BA2FFC15EB1A" box="[831,924,1084,1110]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="6" pageNumber="100">Figs 1E</figureCitation>
,
<figureCitation id="13514FB3FFE5FFF1D3AEBA2FFC42EB1A" box="[937,971,1084,1110]" captionStart="Fig" captionStartId="11.[189,232,1857,1883]" captionTargetBox="[229,1369,265,1800]" captionTargetId="figure-17@11.[218,1369,265,1815]" captionTargetPageId="11" captionText="Fig. 2. A–C, F–H. Calliphora ampyx sp. nov., paratype, ♂, Canberra, ACT, 6 Mar. 1984, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Mt Stromlo, ACT, 26 Oct. 1993, D.K. Yeates leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351104" httpUri="https://zenodo.org/record/15351104/files/figure.png" pageId="6" pageNumber="100">2E</figureCitation>
) ..................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309E4B93FFE5FFF1D0BABA6FFAFEEB95" pageId="6" pageNumber="100">
<paragraph id="8BD55336FFE5FFF1D0BABA6FFAFEEBDA" blockId="6.[189,1399,1147,1241]" box="[189,1399,1147,1174]" pageId="6" pageNumber="100">
<keyLead id="309BF003FFE5FFF1D0BABA6FFAFEEBDA" box="[189,1399,1147,1174]" pageId="6" pageNumber="100">
2. Arista plumose, 1.5 × postpedicel length (
<figureCitation id="13514FB3FFE5FFF1D2D6BA6FFCBBEBDA" box="[721,818,1148,1174]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetBox="[255,1343,265,1730]" captionTargetId="figure-17@15.[245,1343,265,1744]" captionTargetPageId="15" captionText="Fig. 4. A–C. Calliphora niveata sp. nov., paratype, ♂, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Paratype, ♂, Guthega, NSW, 4 Apr. 1984, K.R. Norris leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = plural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351112" httpUri="https://zenodo.org/record/15351112/files/figure.png" pageId="6" pageNumber="100">Figs 4C</figureCitation>
,
<figureCitation id="13514FB3FFE5FFF1D339BA68FCEAEBDA" box="[830,867,1147,1174]" captionStart="Fig" captionStartId="17.[189,232,1822,1848]" captionTargetBox="[244,1354,268,1763]" captionTargetId="figure-17@17.[234,1354,268,1777]" captionTargetPageId="17" captionText="Fig. 5. A–C. Calliphora xanthocera Malloch, 1927, ♂, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view.D–E. Female, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male, Kosciuszko, NSW, 25 Mar. 1965, D.E. Havenstein leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = pleural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351114" httpUri="https://zenodo.org/record/15351114/files/figure.png" pageId="6" pageNumber="100">5C</figureCitation>
); NSW ........................................................ 3
</keyLead>
</paragraph>
<paragraph id="8BD55336FFE5FFF1D0C6BA8DFAFEEB95" blockId="6.[189,1399,1147,1241]" pageId="6" pageNumber="100">
<keyLead id="309BF003FFE5FFF1D0C6BA8DFAFEEB95" pageId="6" pageNumber="100">
<emphasis id="B91E8F24FFE5FFF1D0C6BA8DFF59EBFB" box="[193,208,1182,1207]" italics="true" pageId="6" pageNumber="100">–</emphasis>
Arista almost bare, with only a few very short and sparse setulae near base, 2.0 postpedicel length (
<figureCitation id="13514FB3FFE5FFF1D0F3BAACFED8EB95" box="[244,337,1215,1241]" captionStart="Fig" captionStartId="13.[189,232,1928,1954]" captionTargetBox="[200,1398,639,1887]" captionTargetId="figure-99@13.[189,1398,639,1887]" captionTargetPageId="13" captionText="Fig. 3. Calliphora ignicera sp. nov., paratype, ♀, Flinders Is., TAS, 20 May 1991, C. Tann leg. (ANIC). A. Habitus, dorsal view. B. Head, anterior view. C. Habitus, lateral view. D. Calypter, lateral view. Black scale bars = 1 mm; red scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351108" httpUri="https://zenodo.org/record/15351108/files/figure.png" pageId="6" pageNumber="100">Fig. 3C</figureCitation>
); TAS .....................................................................................................
<taxonomicName id="4C6A28B5FFE5FFF1D488BAACFA9CEB95" authorityName="Johnston &amp; Szpila &amp; Pape &amp; Meiklejohn &amp; Foley &amp; Wallman" authorityYear="2025" box="[1167,1301,1215,1241]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="ignicera" status="sp. nov.">
<emphasis id="B91E8F24FFE5FFF1D488BAACFA9CEB95" bold="true" box="[1167,1301,1215,1241]" italics="true" pageId="6" pageNumber="100">C. ignicera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D325FFFE5FFF1D51BBAACFAFEEB95" box="[1308,1399,1215,1241]" pageId="6" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309E4B93FFE5FFF1D0BABAECFAFEE948" pageId="6" pageNumber="100">
<paragraph id="8BD55336FFE5FFF1D0BABAECFAFEEA32" blockId="6.[189,1399,1279,1541]" pageId="6" pageNumber="100">
<keyLead id="309BF003FFE5FFF1D0BABAECFAFEEA32" pageId="6" pageNumber="100">
3. Acrostichal setae 2+2 (rarely 2 +3); abdominal tergites ventrally with very weak or no microtomentum; pleural suture between anepisternum and anepimeron not pronounced, similar in colour to surrounding structures (
<figureCitation id="13514FB3FFE5FFF1D205BB51FDD4EA10" box="[514,605,1346,1372]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetBox="[255,1343,265,1730]" captionTargetId="figure-17@15.[245,1343,265,1744]" captionTargetPageId="15" captionText="Fig. 4. A–C. Calliphora niveata sp. nov., paratype, ♂, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Paratype, ♂, Guthega, NSW, 4 Apr. 1984, K.R. Norris leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = plural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351112" httpUri="https://zenodo.org/record/15351112/files/figure.png" pageId="6" pageNumber="100">Fig. 4E</figureCitation>
);
<emphasis id="B91E8F24FFE5FFF1D274BB51FD78EA10" box="[627,753,1346,1372]" italics="true" pageId="6" pageNumber="100">males only</emphasis>
, cercus appears hook-shaped in lateral view, with only very slight concavity in its basal half (
<figureCitation id="13514FB3FFE5FFF1D2ADBB70FC8DEA32" box="[682,772,1379,1406]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetBox="[255,1343,265,1730]" captionTargetId="figure-17@15.[245,1343,265,1744]" captionTargetPageId="15" captionText="Fig. 4. A–C. Calliphora niveata sp. nov., paratype, ♂, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Paratype, ♂, Guthega, NSW, 4 Apr. 1984, K.R. Norris leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = plural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351112" httpUri="https://zenodo.org/record/15351112/files/figure.png" pageId="6" pageNumber="100">Fig. 4F</figureCitation>
); NSW .........................................
<taxonomicName id="4C6A28B5FFE5FFF1D49CBB70FA9CEA31" authorityName="Johnston &amp; Szpila &amp; Pape &amp; Meiklejohn &amp; Foley &amp; Wallman" authorityYear="2025" box="[1179,1301,1379,1405]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="niveata" status="sp. nov.">
<emphasis id="B91E8F24FFE5FFF1D49CBB70FA9CEA31" bold="true" box="[1179,1301,1379,1405]" italics="true" pageId="6" pageNumber="100">C. niveata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D325FFFE5FFF1D51BBB77FAFEEA32" box="[1308,1399,1380,1406]" pageId="6" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BD55336FFE5FFF1D0C6BB95FAFEE948" blockId="6.[189,1399,1279,1541]" pageId="6" pageNumber="100">
<keyLead id="309BF003FFE5FFF1D0C6BB95FAFEE948" pageId="6" pageNumber="100">
– Acrostichal setae 2+3 (
<figureCitation id="13514FB3FFE5FFF1D1FDBB96FDDFEAEC" box="[506,598,1413,1440]" captionStart="Fig" captionStartId="17.[189,232,1822,1848]" captionTargetBox="[244,1354,268,1763]" captionTargetId="figure-17@17.[234,1354,268,1777]" captionTargetPageId="17" captionText="Fig. 5. A–C. Calliphora xanthocera Malloch, 1927, ♂, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view.D–E. Female, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male, Kosciuszko, NSW, 25 Mar. 1965, D.E. Havenstein leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = pleural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351114" httpUri="https://zenodo.org/record/15351114/files/figure.png" pageId="6" pageNumber="100">Fig. 5A</figureCitation>
); abdominal tergites ventrally with grey microtomentum that changes colour with viewing angle; pleural suture between anepisternum and anepimeron pronounced, convex and dark red-brown, distinct from ground colour of pleura (
<figureCitation id="13514FB3FFE5FFF1D3F0BBDBFBD8EAAF" box="[1015,1105,1480,1507]" captionStart="Fig" captionStartId="17.[189,232,1822,1848]" captionTargetBox="[244,1354,268,1763]" captionTargetId="figure-17@17.[234,1354,268,1777]" captionTargetPageId="17" captionText="Fig. 5. A–C. Calliphora xanthocera Malloch, 1927, ♂, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view.D–E. Female, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male, Kosciuszko, NSW, 25 Mar. 1965, D.E. Havenstein leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = pleural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351114" httpUri="https://zenodo.org/record/15351114/files/figure.png" pageId="6" pageNumber="100">Fig. 5E</figureCitation>
);
<emphasis id="B91E8F24FFE5FFF1D462BBDBFB6AEAAE" box="[1125,1251,1480,1506]" italics="true" pageId="6" pageNumber="100">males only</emphasis>
, cercus with strong concavity in its basal half (lateral view) (
<figureCitation id="13514FB3FFE5FFF1D31DBBF9FCFDE949" box="[794,884,1514,1541]" captionStart="Fig" captionStartId="17.[189,232,1822,1848]" captionTargetBox="[244,1354,268,1763]" captionTargetId="figure-17@17.[234,1354,268,1777]" captionTargetPageId="17" captionText="Fig. 5. A–C. Calliphora xanthocera Malloch, 1927, ♂, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view.D–E. Female, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male, Kosciuszko, NSW, 25 Mar. 1965, D.E. Havenstein leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = pleural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351114" httpUri="https://zenodo.org/record/15351114/files/figure.png" pageId="6" pageNumber="100">Fig. 5F</figureCitation>
); NSW, VIC
<taxonomicName id="4C6A28B5FFE5FFF1D41EBBF9FAFEE948" authority="Malloch, 1927" authorityName="Malloch" authorityYear="1927" box="[1049,1399,1513,1540]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="xanthocera">
<emphasis id="B91E8F24FFE5FFF1D41EBBF9FB4DE948" bold="true" box="[1049,1220,1513,1540]" italics="true" pageId="6" pageNumber="100">C. xanthocera</emphasis>
<bibRefCitation id="EFFB2EC7FFE5FFF1D4CCBBF9FAFEE948" author="Malloch J. R." box="[1227,1399,1514,1540]" pageId="6" pageNumber="100" pagination="299 - 335" refId="ref11704" refString="Malloch J. R. 1927. Notes on Australian Diptera. No. XI. Proceedings of the Linnean Society of New South Wales 52: 299-335." type="journal article" year="1927">Malloch, 1927</bibRefCitation>
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</keyStep>
<keyStep id="309E4B93FFE5FFF1D0BAB839FAFEE9E5" pageId="6" pageNumber="100">
<paragraph id="8BD55336FFE5FFF1D0BAB839FAFEE92A" blockId="6.[189,1399,1578,1705]" pageId="6" pageNumber="100">
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4.
<emphasis id="B91E8F24FFE5FFF1D0EDB839FEFBE908" box="[234,370,1578,1604]" italics="true" pageId="6" pageNumber="100">Males only</emphasis>
, fronto-orbital and parafacial plate mottled gold at the height of antennal insertion (
<figureCitation id="13514FB3FFE5FFF1D0F4B858FEFDE929" box="[243,372,1611,1637]" captionStart="Fig" captionStartId="11.[189,232,1857,1883]" captionTargetBox="[229,1369,265,1800]" captionTargetId="figure-17@11.[218,1369,265,1815]" captionTargetPageId="11" captionText="Fig. 2. A–C, F–H. Calliphora ampyx sp. nov., paratype, ♂, Canberra, ACT, 6 Mar. 1984, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Mt Stromlo, ACT, 26 Oct. 1993, D.K. Yeates leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351104" httpUri="https://zenodo.org/record/15351104/files/figure.png" pageId="6" pageNumber="100">Fig. 2A–B</figureCitation>
);
<collectionCode id="ED7BCBF3FFE5FFF1D18FB858FE4DE929" box="[392,452,1611,1637]" pageId="6" pageNumber="100">ACT</collectionCode>
, NSW, VIC ................................................................................
<taxonomicName id="4C6A28B5FFE5FFF1D4A4B858FA9CE92A" authorityName="Johnston &amp; Szpila &amp; Pape &amp; Meiklejohn &amp; Foley &amp; Wallman" authorityYear="2025" box="[1187,1301,1611,1638]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="ampyx" status="sp. nov.">
<emphasis id="B91E8F24FFE5FFF1D4A4B858FA9CE92A" bold="true" box="[1187,1301,1611,1638]" italics="true" pageId="6" pageNumber="100">C. ampyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D325FFFE5FFF1D51BB85FFAFEE92A" box="[1308,1399,1612,1638]" pageId="6" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BD55336FFE5FFF1D0C6B87EFAFEE9E5" blockId="6.[189,1399,1578,1705]" pageId="6" pageNumber="100">
<keyLead id="309BF003FFE5FFF1D0C6B87EFAFEE9E5" pageId="6" pageNumber="100">
–
<emphasis id="B91E8F24FFE5FFF1D0EDB87EFEE4E9CB" box="[234,365,1645,1671]" italics="true" pageId="6" pageNumber="100">Males only</emphasis>
, fronto-orbital and parafacial plate grey, silver, white or black at the height of antennal insertion (
<figureCitation id="13514FB3FFE5FFF1D165B89DFE69E9E5" box="[354,480,1678,1705]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="6" pageNumber="100">Fig. 1A–B</figureCitation>
)......................................................................................................................... 5
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<keyStep id="309E4B93FFE5FFF1D0BAB8DDFAFEE801" pageId="6" pageNumber="100">
<paragraph id="8BD55336FFE5FFF1D0BAB8DDFAFEE846" blockId="6.[189,1399,1742,1870]" pageId="6" pageNumber="100">
<keyLead id="309BF003FFE5FFF1D0BAB8DDFAFEE846" pageId="6" pageNumber="100">
5. Pleural suture between anepisternum and anepimeron pronounced and dark red-brown, distinct from ground colour of pleura (
<figureCitation id="13514FB3FFE5FFF1D208B8E3FDE3E847" box="[527,618,1776,1803]" captionStart="Fig" captionStartId="17.[189,232,1822,1848]" captionTargetBox="[244,1354,268,1763]" captionTargetId="figure-17@17.[234,1354,268,1777]" captionTargetPageId="17" captionText="Fig. 5. A–C. Calliphora xanthocera Malloch, 1927, ♂, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view.D–E. Female, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male, Kosciuszko, NSW, 25 Mar. 1965, D.E. Havenstein leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = pleural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351114" httpUri="https://zenodo.org/record/15351114/files/figure.png" pageId="6" pageNumber="100">Fig. 5E</figureCitation>
); NSW, VIC .....................................
<taxonomicName id="4C6A28B5FFE5FFF1D41EB8E3FAFEE846" authority="Malloch, 1927" authorityName="Malloch" authorityYear="1927" box="[1049,1399,1775,1802]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="xanthocera">
<emphasis id="B91E8F24FFE5FFF1D41EB8E3FB4DE846" bold="true" box="[1049,1220,1775,1802]" italics="true" pageId="6" pageNumber="100">C. xanthocera</emphasis>
<bibRefCitation id="EFFB2EC7FFE5FFF1D4CCB8E3FAFEE846" author="Malloch J. R." box="[1227,1399,1776,1802]" pageId="6" pageNumber="100" pagination="299 - 335" refId="ref11704" refString="Malloch J. R. 1927. Notes on Australian Diptera. No. XI. Proceedings of the Linnean Society of New South Wales 52: 299-335." type="journal article" year="1927">Malloch, 1927</bibRefCitation>
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<paragraph id="8BD55336FFE5FFF1D0C6B901FAFEE801" blockId="6.[189,1399,1742,1870]" pageId="6" pageNumber="100">
<keyLead id="309BF003FFE5FFF1D0C6B901FAFEE801" pageId="6" pageNumber="100">
– Pleural suture between anepisternum and anepimeron not pronounced and similar in colour to surrounding structures (
<figureCitation id="13514FB3FFE5FFF1D205B920FDD2E801" box="[514,603,1843,1869]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="6" pageNumber="100">Fig. 1E</figureCitation>
); WA ...........................................................
<taxonomicName id="4C6A28B5FFE5FFF1D45BB920FAFEE801" authority="Malloch, 1927" authorityName="Malloch" authorityYear="1927" box="[1116,1399,1843,1870]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="clarki">
<emphasis id="B91E8F24FFE5FFF1D45BB920FB4DE802" bold="true" box="[1116,1220,1843,1870]" italics="true" pageId="6" pageNumber="100">C. clarki</emphasis>
<bibRefCitation id="EFFB2EC7FFE5FFF1D4CCB920FAFEE801" author="Malloch J. R." box="[1227,1399,1843,1869]" pageId="6" pageNumber="100" pagination="299 - 335" refId="ref11704" refString="Malloch J. R. 1927. Notes on Australian Diptera. No. XI. Proceedings of the Linnean Society of New South Wales 52: 299-335." type="journal article" year="1927">Malloch, 1927</bibRefCitation>
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<paragraph id="8BD55336FFE6FFF2D214BC19FBA9ED68" blockId="5.[488,1099,416,548]" box="[531,1056,522,548]" pageId="5" pageNumber="99">
Genus
<taxonomicName id="4C6A28B5FFE6FFF2D261BC19FBA9ED68" ID-CoL="9CJFG" authority="Robineau-Desvoidy, 1830" authorityName="Robineau-Desvoidy" authorityYear="1830" box="[614,1056,522,548]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="B91E8F24FFE6FFF2D261BC19FD6EED68" box="[614,743,522,548]" italics="true" pageId="5" pageNumber="99">Calliphora</emphasis>
Robineau-Desvoidy, 1830
</taxonomicName>
</paragraph>
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<subSubSection id="C37000BDFFE6FFF2D0BABC58FDAFEDB8" pageId="5" pageNumber="99" type="diagnosis">
<paragraph id="8BD55336FFE6FFF2D0BABC58FDA7ED2A" blockId="5.[189,558,587,614]" box="[189,558,587,614]" pageId="5" pageNumber="99">
<heading id="D09DE45AFFE6FFF2D0BABC58FDA7ED2A" bold="true" box="[189,558,587,614]" fontSize="11" level="3" pageId="5" pageNumber="99" reason="3">
<emphasis id="B91E8F24FFE6FFF2D0BABC58FDA7ED2A" bold="true" box="[189,558,587,614]" pageId="5" pageNumber="99">
Diagnosis of genus
<taxonomicName id="4C6A28B5FFE6FFF2D1ABBC58FDA7ED2A" authorityName="Robineau-Desvoidy" authorityYear="1830" box="[428,558,587,614]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="B91E8F24FFE6FFF2D1ABBC58FDA7ED2A" bold="true" box="[428,558,587,614]" italics="true" pageId="5" pageNumber="99">Calliphora</emphasis>
</taxonomicName>
</emphasis>
</heading>
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<paragraph id="8BD55336FFE6FFF2D0BABC6BFDAFEDB8" blockId="5.[187,1398,632,757]" pageId="5" pageNumber="99">
The genus
<taxonomicName id="4C6A28B5FFE6FFF2D13EBC6BFE33EDDE" authorityName="Robineau-Desvoidy" authorityYear="1830" box="[313,442,632,658]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="B91E8F24FFE6FFF2D13EBC6BFE33EDDE" box="[313,442,632,658]" italics="true" pageId="5" pageNumber="99">Calliphora</emphasis>
</taxonomicName>
shares a combination of (a) abdomen ground colour metallic blue, purple or green, mottled gold or silver or entirely orange, and (b) stem vein bare dorsally with other genera of subfamily
<taxonomicName id="4C6A28B5FFE6FFF2D0BABCAAFEE9ED9F" box="[189,352,697,723]" class="Insecta" family="Calliphoridae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="99" phylum="Arthropoda" rank="subFamily" subFamily="Calliphorinae">Calliphorinae</taxonomicName>
, but differs by the combination of (c) lower calypter setose dorsally, and (d) subcostal sclerite not setose (pubescent).
</paragraph>
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<subSubSection id="C37000BDFFE6FFF2D0BABD0FFBE1EC7A" box="[189,1128,795,822]" pageId="5" pageNumber="99" type="multiple">
<paragraph id="8BD55336FFE6FFF2D0BABD0FFBE1EC7A" blockId="5.[189,1128,795,822]" box="[189,1128,795,822]" pageId="5" pageNumber="99">
<heading id="D09DE45AFFE6FFF2D0BABD0FFBE1EC7A" bold="true" box="[189,1128,795,822]" fontSize="11" level="3" pageId="5" pageNumber="99" reason="3">
<emphasis id="B91E8F24FFE6FFF2D0BABD0FFBE1EC7A" bold="true" box="[189,1128,795,822]" pageId="5" pageNumber="99">
Key to genera of Australian
<taxonomicName id="4C6A28B5FFE6FFF2D219BD08FD42EC79" box="[542,715,795,821]" class="Insecta" family="Calliphoridae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="99" phylum="Arthropoda" rank="subFamily" subFamily="Calliphorinae">Calliphorinae</taxonomicName>
and species-groups of
<taxonomicName id="4C6A28B5FFE6FFF2D3E1BD08FBE1EC7A" authorityName="Robineau-Desvoidy" authorityYear="1830" box="[998,1128,795,822]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="B91E8F24FFE6FFF2D3E1BD08FBE1EC7A" bold="true" box="[998,1128,795,822]" italics="true" pageId="5" pageNumber="99">Calliphora</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C37000BDFFE6FFF1D0BABD59FAFFED1E" lastPageId="6" lastPageNumber="100" pageId="5" pageNumber="99" type="key">
<key id="FFFB0407FFE6FFF1D0BABD59FAFFED1E" lastPageId="6" lastPageNumber="100" pageId="5" pageNumber="99">
<keyStep id="309E4B93FFE6FFF2D0BABD59FAFEECCB" pageId="5" pageNumber="99">
<paragraph id="8BD55336FFE6FFF2D0BABD59FAFEEC28" blockId="5.[189,1399,842,903]" box="[189,1399,842,868]" pageId="5" pageNumber="99">
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1. Subcostal sclerite setulose (Supp. file 1:
<figureCitation id="13514FB3FFE6FFF2D2C2BD59FCA2EC28" box="[709,811,842,868]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Fig. S1F</figureCitation>
) .......................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BD55336FFE6FFF2D0C6BD7EFAFEECCB" blockId="5.[189,1399,842,903]" box="[193,1399,876,903]" pageId="5" pageNumber="99">
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– Subcostal sclerite pubescent or bare (Supp. file 1:
<figureCitation id="13514FB3FFE6FFF2D331BD7FFC14ECCB" box="[822,925,876,903]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Fig. S1E</figureCitation>
) ........................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309E4B93FFE6FFF2D0BABDBEFAFEECA6" pageId="5" pageNumber="99">
<paragraph id="8BD55336FFE6FFF2D0BABDBEFAFEEC8B" blockId="5.[189,1399,941,1002]" box="[189,1399,941,968]" pageId="5" pageNumber="99">
<keyLead id="309BF003FFE6FFF2D0BABDBEFAFEEC8B" box="[189,1399,941,968]" pageId="5" pageNumber="99">
2. Eye densely and uniformly setulose (Supp. file 1:
<figureCitation id="13514FB3FFE6FFF2D333BDBEFC15EC8B" box="[820,924,941,967]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Fig. S1B</figureCitation>
) .........................
<taxonomicName id="4C6A28B5FFE6FFF2D46DBDBEFAFEEC8B" authority="Bezzi, 1927" authorityName="Bezzi" authorityYear="1927" box="[1130,1399,941,968]" class="Insecta" family="Calliphoridae" genus="Ptilonesia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="B91E8F24FFE6FFF2D46DBDBEFB6AEC84" bold="true" box="[1130,1251,941,968]" italics="true" pageId="5" pageNumber="99">Ptilonesia</emphasis>
Bezzi, 1927
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD55336FFE6FFF2D0C6BDC3FAFEECA6" blockId="5.[189,1399,941,1002]" box="[193,1399,975,1002]" pageId="5" pageNumber="99">
<keyLead id="309BF003FFE6FFF2D0C6BDC3FAFEECA6" box="[193,1399,975,1002]" pageId="5" pageNumber="99">
– Eye bare or with only sparse setulae (
<figureCitation id="13514FB3FFE6FFF2D2A6BDC3FCAFECA6" box="[673,806,976,1002]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Figs 1B, D</figureCitation>
,
<figureCitation id="13514FB3FFE6FFF2D335BDC3FCF0ECA6" box="[818,889,976,1002]" captionStart="Fig" captionStartId="11.[189,232,1857,1883]" captionTargetBox="[229,1369,265,1800]" captionTargetId="figure-17@11.[218,1369,265,1815]" captionTargetPageId="11" captionText="Fig. 2. A–C, F–H. Calliphora ampyx sp. nov., paratype, ♂, Canberra, ACT, 6 Mar. 1984, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Mt Stromlo, ACT, 26 Oct. 1993, D.K. Yeates leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351104" httpUri="https://zenodo.org/record/15351104/files/figure.png" pageId="5" pageNumber="99">2B, D</figureCitation>
) .................
<taxonomicName id="4C6A28B5FFE6FFF2D40EBDDCFAFEECA6" authority="Malloch, 1924" authorityName="Malloch" authorityYear="1924" box="[1033,1399,975,1002]" class="Insecta" family="Calliphoridae" genus="Xenocalliphora" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="B91E8F24FFE6FFF2D40EBDDCFB4DECA6" bold="true" box="[1033,1220,975,1002]" italics="true" pageId="5" pageNumber="99">Xenocalliphora</emphasis>
Malloch, 1924
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</paragraph>
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<keyStep id="309E4B93FFE6FFF2D0BABA02FAFEEBDF" pageId="5" pageNumber="99">
<paragraph id="8BD55336FFE6FFF2D0BABA02FAFEEB02" blockId="5.[189,1399,1041,1171]" pageId="5" pageNumber="99">
<keyLead id="309BF003FFE6FFF2D0BABA02FAFEEB02" pageId="5" pageNumber="99">
3. Male abdominal sternite 5 with postero-median margin extended into down-turned subapical spine; lower calypter bare dorsally (see
<bibRefCitation id="EFFB2EC7FFE6FFF2D277BA20FC8BEB02" author="Norris K. R." box="[624,770,1075,1102]" pageId="5" pageNumber="99" pagination="511 - 628" refId="ref12028" refString="Norris K. R. 1999. Establishment of a subfamily Aphyssurinae for the Australian genus Aphyssura Hardy (Diptera: Calliphoridae), with a review of known forms and descriptions of new species. Invertebrate Systematics 13 (4): 511-628. https://doi.org/10.1071/IT98007" type="journal article" year="1999">Norris 1999</bibRefCitation>
: fig. 1h–i) .............................
<taxonomicName id="4C6A28B5FFE6FFF2D459BA20FAFEEB02" authority="Hardy, 1940" authorityName="Hardy" authorityYear="1940" box="[1118,1399,1075,1102]" class="Insecta" family="Calliphoridae" genus="Aphyssura" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="B91E8F24FFE6FFF2D459BA20FB56EB02" bold="true" box="[1118,1247,1075,1102]" italics="true" pageId="5" pageNumber="99">Aphyssura</emphasis>
Hardy, 1940
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</paragraph>
<paragraph id="8BD55336FFE6FFF2D0C6BA44FAFEEBDF" blockId="5.[189,1399,1041,1171]" pageId="5" pageNumber="99">
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– Male abdominal sternite 5 with postero-median margin unmodified; lower calypter with setae dorsally ........................................................................................................................4 (
<taxonomicName id="4C6A28B5FFE6FFF2D4EDBA6BFAE5EBDF" authorityName="Robineau-Desvoidy" authorityYear="1830" box="[1258,1388,1144,1171]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="B91E8F24FFE6FFF2D4EDBA6BFAE5EBDF" bold="true" box="[1258,1388,1144,1171]" italics="true" pageId="5" pageNumber="99">Calliphora</emphasis>
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)
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</paragraph>
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<keyStep id="309E4B93FFE6FFF2D0BABAA9FAFEEBBB" pageId="5" pageNumber="99">
<paragraph id="8BD55336FFE6FFF2D0BABAA9FAFFEB98" blockId="5.[189,1399,1209,1271]" box="[189,1398,1209,1236]" pageId="5" pageNumber="99">
<keyLead id="309BF003FFE6FFF2D0BABAA9FAFFEB98" box="[189,1398,1209,1236]" pageId="5" pageNumber="99">
4. Eye densely setulose over entire surface (Supp. file 1:
<figureCitation id="13514FB3FFE6FFF2D360BAA9FC46EB98" box="[871,975,1210,1236]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Fig. S1B</figureCitation>
) .............................
<emphasis id="B91E8F24FFE6FFF2D4BEBAAAFAA1EB98" bold="true" box="[1209,1320,1209,1236]" italics="true" pageId="5" pageNumber="99">ochracea</emphasis>
-group
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</paragraph>
<paragraph id="8BD55336FFE6FFF2D0C6BACEFAFEEBBB" blockId="5.[189,1399,1209,1271]" box="[193,1399,1244,1271]" pageId="5" pageNumber="99">
<keyLead id="309BF003FFE6FFF2D0C6BACEFAFEEBBB" box="[193,1399,1244,1271]" pageId="5" pageNumber="99">– Eye without setulae ........................................................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="309E4B93FFE6FFF2D0BABB0EFAFEEAA9" pageId="5" pageNumber="99">
<paragraph id="8BD55336FFE6FFF2D0BABB0EFAFFEAEC" blockId="5.[189,1399,1309,1509]" pageId="5" pageNumber="99">
<keyLead id="309BF003FFE6FFF2D0BABB0EFAFFEAEC" pageId="5" pageNumber="99">
5. Femora and tibiae at least partially orange or yellow (Supp. file 1:
<figureCitation id="13514FB3FFE6FFF2D427BB0EFBF6EA7B" box="[1056,1151,1309,1335]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Fig. 1A</figureCitation>
); abdominal tergites 1+2–5 with microtomentum mottled golden at least in the lateral 0.3 (Supp. file 1:
<figureCitation id="13514FB3FFE6FFF2D4B1BB53FAABEA16" box="[1206,1314,1344,1370]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Fig. S1D</figureCitation>
), some species with contrasting median metallic patches dorsally on
<collectionCode id="ED7BCBF3FFE6FFF2D3BEBB71FC45EA31" box="[953,972,1378,1405]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="99">T</collectionCode>
1+2–
<collectionCode id="ED7BCBF3FFE6FFF2D401BB71FB90EA31" box="[1030,1049,1378,1405]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="99">T</collectionCode>
5 (Supp. file 1:
<figureCitation id="13514FB3FFE6FFF2D4C9BB70FABEEA31" box="[1230,1335,1379,1405]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Fig. S1C</figureCitation>
) ..... .........................................................................................................................................
<taxonomicName id="4C6A28B5FFE6FFF2D4E5BB96FAA1EAD3" authorityName="Fabricius" authorityYear="1782" box="[1250,1320,1413,1439]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="99" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B91E8F24FFE6FFF2D4E5BB96FAA1EAD3" bold="true" box="[1250,1320,1413,1439]" italics="true" pageId="5" pageNumber="99">stygia</emphasis>
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<paragraph id="8BD55336FFE6FFF2D0C6BBBAFAFEEAA9" blockId="5.[189,1399,1309,1509]" pageId="5" pageNumber="99">
<keyLead id="309BF003FFE6FFF2D0C6BBBAFAFEEAA9" pageId="5" pageNumber="99">
– Femora and tibiae dark brown or black (
<figureCitation id="13514FB3FFE6FFF2D2B4BBBBFC99EA8E" box="[691,784,1448,1474]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Figs 1C</figureCitation>
,
<figureCitation id="13514FB3FFE6FFF2D31BBBBBFCCBEA8E" box="[796,834,1448,1474]" captionStart="Fig" captionStartId="11.[189,232,1857,1883]" captionTargetBox="[229,1369,265,1800]" captionTargetId="figure-17@11.[218,1369,265,1815]" captionTargetPageId="11" captionText="Fig. 2. A–C, F–H. Calliphora ampyx sp. nov., paratype, ♂, Canberra, ACT, 6 Mar. 1984, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Mt Stromlo, ACT, 26 Oct. 1993, D.K. Yeates leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351104" httpUri="https://zenodo.org/record/15351104/files/figure.png" pageId="5" pageNumber="99">2C</figureCitation>
); abdominal tergites 1+2–5 with microtomentum mottled golden, silver or absent (Supp. file 1:
<figureCitation id="13514FB3FFE6FFF2D303BBD8FCE7EAA9" box="[772,878,1483,1509]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Fig. S1D</figureCitation>
) ................................................................. 6
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<keyStep id="309E4B93FFE6FFF2D0BAB818FAFEE9C2" pageId="5" pageNumber="99">
<paragraph id="8BD55336FFE6FFF2D0BAB818FAFEE905" blockId="5.[189,1399,1547,1678]" pageId="5" pageNumber="99">
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6. Anterior spiracle reduced in size, smaller in width at widest part than proepisternum (Supp. file 1:
<figureCitation id="13514FB3FFE6FFF2D18DB83DFE7FE904" box="[394,502,1582,1608]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Fig. S1G</figureCitation>
;
<figureCitation id="13514FB3FFE6FFF2D204B83DFDEAE904" box="[515,611,1582,1608]" captionStart="Fig" captionStartId="13.[189,232,1928,1954]" captionTargetBox="[200,1398,639,1887]" captionTargetId="figure-99@13.[189,1398,639,1887]" captionTargetPageId="13" captionText="Fig. 3. Calliphora ignicera sp. nov., paratype, ♀, Flinders Is., TAS, 20 May 1991, C. Tann leg. (ANIC). A. Habitus, dorsal view. B. Head, anterior view. C. Habitus, lateral view. D. Calypter, lateral view. Black scale bars = 1 mm; red scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351108" httpUri="https://zenodo.org/record/15351108/files/figure.png" pageId="5" pageNumber="99">Figs 3D</figureCitation>
,
<figureCitation id="13514FB3FFE6FFF2D277B83DFD1BE905" box="[624,658,1582,1609]" captionStart="Fig" captionStartId="17.[189,232,1822,1848]" captionTargetBox="[244,1354,268,1763]" captionTargetId="figure-17@17.[234,1354,268,1777]" captionTargetPageId="17" captionText="Fig. 5. A–C. Calliphora xanthocera Malloch, 1927, ♂, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view.D–E. Female, Tinmine Hut Track, Snowy Mountains, NSW, 20 Mar. 1971, K.R. Norris leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male, Kosciuszko, NSW, 25 Mar. 1965, D.E. Havenstein leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = pleural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351114" httpUri="https://zenodo.org/record/15351114/files/figure.png" pageId="5" pageNumber="99">5E</figureCitation>
) ............................................................................................... 7
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</paragraph>
<paragraph id="8BD55336FFE6FFF2D0C6B842FAFEE9C2" blockId="5.[189,1399,1547,1678]" pageId="5" pageNumber="99">
<keyLead id="309BF003FFE6FFF2D0C6B842FAFEE9C2" pageId="5" pageNumber="99">
– Anterior spiracle regular in size, equal to or larger in width at widest part than proepisternum (Supp. file 1:
<figureCitation id="13514FB3FFE6FFF2D18DB860FE7DE9C2" box="[394,500,1651,1678]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Fig. S1H</figureCitation>
) ..................................................................................................................... 8
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<keyStep id="309E4B93FFE6FFF2D0BAB8A6FAFFE8D3" pageId="5" pageNumber="99">
<paragraph id="8BD55336FFE6FFF2D0BAB8A6FAFFE858" blockId="5.[189,1399,1716,1951]" pageId="5" pageNumber="99">
<keyLead id="309BF003FFE6FFF2D0BAB8A6FAFFE858" pageId="5" pageNumber="99">
7. Postpedicel at least partially orange (
<figureCitation id="13514FB3FFE6FFF2D29DB8A7FD73E983" box="[666,762,1716,1743]" captionStart="Fig" captionStartId="11.[189,232,1857,1883]" captionTargetBox="[229,1369,265,1800]" captionTargetId="figure-17@11.[218,1369,265,1815]" captionTargetPageId="11" captionText="Fig. 2. A–C, F–H. Calliphora ampyx sp. nov., paratype, ♂, Canberra, ACT, 6 Mar. 1984, K.R. Norris leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Mt Stromlo, ACT, 26 Oct. 1993, D.K. Yeates leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351104" httpUri="https://zenodo.org/record/15351104/files/figure.png" pageId="5" pageNumber="99">Figs 2B</figureCitation>
,
<figureCitation id="13514FB3FFE6FFF2D301B8A6FCA2E983" box="[774,811,1717,1743]" captionStart="Fig" captionStartId="13.[189,232,1928,1954]" captionTargetBox="[200,1398,639,1887]" captionTargetId="figure-99@13.[189,1398,639,1887]" captionTargetPageId="13" captionText="Fig. 3. Calliphora ignicera sp. nov., paratype, ♀, Flinders Is., TAS, 20 May 1991, C. Tann leg. (ANIC). A. Habitus, dorsal view. B. Head, anterior view. C. Habitus, lateral view. D. Calypter, lateral view. Black scale bars = 1 mm; red scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351108" httpUri="https://zenodo.org/record/15351108/files/figure.png" pageId="5" pageNumber="99">3B</figureCitation>
); upper calypter with dark orange or brown fringe (
<figureCitation id="13514FB3FFE6FFF2D0F5B8C4FEDBE9BD" box="[242,338,1751,1777]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Figs 1E</figureCitation>
,
<figureCitation id="13514FB3FFE6FFF2D165B8C4FE0FE9BD" box="[354,390,1751,1777]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetBox="[255,1343,265,1730]" captionTargetId="figure-17@15.[245,1343,265,1744]" captionTargetPageId="15" captionText="Fig. 4. A–C. Calliphora niveata sp. nov., paratype, ♂, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Paratype, ♂, Guthega, NSW, 4 Apr. 1984, K.R. Norris leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = plural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351112" httpUri="https://zenodo.org/record/15351112/files/figure.png" pageId="5" pageNumber="99">4E</figureCitation>
); male terminalia with cercus hooked with a weak concave region in basal half and weak to no sclerotisation in distal half of mesohypophallus (
<figureCitation id="13514FB3FFE6FFF2D3B7B8E9FBA0E858" box="[944,1065,1786,1812]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetBox="[255,1343,265,1730]" captionTargetId="figure-17@15.[245,1343,265,1744]" captionTargetPageId="15" captionText="Fig. 4. A–C. Calliphora niveata sp. nov., paratype, ♂, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Paratype, ♀, Rainbow Lake Walking track Kosciuszko, NSW, 30 Jan. 2023, N.P. Johnston, J.F. Wallman, A. Grzywacz and K. Szpila leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Paratype, ♂, Guthega, NSW, 4 Apr. 1984, K.R. Norris leg. (ANIC). F. Terminalia, lateral view. G. Terminalia, posterior view. H. Phallus, lateral view. Abbreviation: Pl = plural suture. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351112" httpUri="https://zenodo.org/record/15351112/files/figure.png" pageId="5" pageNumber="99">Fig. 4F, H</figureCitation>
) ......................
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<taxonomicName id="4C6A28B5FFE6FFF2D4E5B8EAFAA1E858" authorityName="Malloch" authorityYear="1927" box="[1250,1320,1785,1812]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="99" phylum="Arthropoda" rank="species" species="clarki">
<emphasis id="B91E8F24FFE6FFF2D4E5B8EAFAA1E858" bold="true" box="[1250,1320,1785,1812]" italics="true" pageId="5" pageNumber="99">clarki</emphasis>
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group
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<paragraph id="8BD55336FFE6FFF2D0C6B90EFAFFE8D3" blockId="5.[189,1399,1716,1951]" pageId="5" pageNumber="99">
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– Postpedicel orange, brown or black (Supp. file 1:
<figureCitation id="13514FB3FFE6FFF2D36AB90FFB8EE87B" box="[877,1031,1820,1847]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Fig. S1I, M</figureCitation>
); upper calypter with yellow or hyaline fringe (Supp. file 1:
<figureCitation id="13514FB3FFE6FFF2D272B92CFD6EE815" box="[629,743,1855,1881]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Fig. S1G</figureCitation>
); male terminalia with weakly curved cercus (lateral view) and mesohypophallus with distinct break or very weak region of sclerotisation medially (Supp. file 1:
<figureCitation id="13514FB3FFE6FFF2D18DB997FE63E8D3" box="[394,490,1924,1951]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="5" pageNumber="99">Fig. S1J</figureCitation>
) ..............................................................................................
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<emphasis id="B91E8F24FFE6FFF2D4B2B997FAA0E8D3" bold="true" box="[1205,1321,1924,1951]" italics="true" pageId="5" pageNumber="99">flexipenis</emphasis>
-
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group
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8. Male terminalia with mesohypophallus strongly curved (c-shaped) in distal half, without break in sclerotisation (Supp. file 1:
<figureCitation id="13514FB3FFE5FFF1D229BF3FFD11EE0A" box="[558,664,300,326]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="6" pageNumber="100">Fig. S1K</figureCitation>
) ........................................................................
<taxonomicName id="4C6A28B5FFE5FFF1D4C5BF3FFAA0EE0B" authorityName="Malloch" authorityYear="1932" box="[1218,1321,300,327]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="sternalis">
<emphasis id="B91E8F24FFE5FFF1D4C5BF3FFAA0EE0B" bold="true" box="[1218,1321,300,327]" italics="true" pageId="6" pageNumber="100">sternalis</emphasis>
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<keyLead id="309BF003FFE5FFF1D0C6BF43FAFEEE25" box="[193,1399,335,362]" pageId="6" pageNumber="100">– Male terminalia with mesohypophallus sclerotised and straight or gently curved in distal half ...... 9</keyLead>
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<paragraph id="8BD55336FFE5FFF1D0BABF9CFAFFEEA1" blockId="6.[189,1399,399,594]" pageId="6" pageNumber="100">
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9. Epandrium elongated, twice length of cercus from base to insertion point of cerci; acrophallus ~2 × as long as paraphallus; cercus broad in basal half, at least twice width at tip; surstylus distally with swollen lobe (Supp. file 1:
<figureCitation id="13514FB3FFE5FFF1D221BFC1FD18EEA1" box="[550,657,466,493]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="6" pageNumber="100">Fig. S1N</figureCitation>
) ............................................................................
<taxonomicName id="4C6A28B5FFE5FFF1D4D0BFC1FAA1EEA1" box="[1239,1320,466,493]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B91E8F24FFE5FFF1D4D0BFC1FAA1EEA1" bold="true" box="[1239,1320,466,493]" italics="true" pageId="6" pageNumber="100">tibialis</emphasis>
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<paragraph id="8BD55336FFE5FFF1D0C6BFE6FAFFED1E" blockId="6.[189,1399,399,594]" pageId="6" pageNumber="100">
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– Epandrium not elongated, similar in length to cercus; acrophallus equal to length of paraphallus at most extending slightly beyond tips; cercus slender for entire length, similar in width to surstylus; surstylus distally without a lobe (Supp. file 1:
<figureCitation id="13514FB3FFE5FFF1D300BC24FCE7ED1D" box="[775,878,567,593]" captionStart="Fig" captionStartId="8.[189,232,1752,1778]" captionTargetBox="[266,1330,265,1697]" captionTargetId="figure-12@8.[257,1330,265,1710]" captionTargetPageId="8" captionText="Fig. 1. A–C. Calliphora clarki Malloch, 1927, ♂, Yallingup, WA, 9 Nov. 1958, E.F. Riek leg. (ANIC). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, anterior view. D–E. Female, Augusta, WA, 3 Oct. 1970, D.H. Colless leg. (ANIC). D. Head, anterior view. E. Calypter, lateral view. F–H. Male terminalia and phallus, Circular Pool, Normalup WA, 9 Aug. 1970, D.H. Colless leg. (ANIC). F. Lateral view. G. Posterior view. H. Phallus, lateral view. Abbreviations: Acr = acrophallus; Ce = cercus; Epi = epiphallus; Hy = hypophallic lobe; M = mesohypophallus; Par = paraphallus; Pl = pleural suture; PoG = postgonite; PrG = pregonite; Su = surstylus. Black scale bars = 1 mm; red scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.15351102" httpUri="https://zenodo.org/record/15351102/files/figure.png" pageId="6" pageNumber="100">Fig. S1L</figureCitation>
) ...............................................
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<emphasis id="B91E8F24FFE5FFF1D4E6BC24FAA0ED1D" bold="true" box="[1249,1321,567,593]" italics="true" pageId="6" pageNumber="100">typica</emphasis>
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