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<document id="66AB9296AFFB1668871AC540D828A5B4" ID-DOI="10.5281/zenodo.11087990" ID-ISSN="1560-8425" ID-Zenodo-Dep="11087990" ID-ZooBank="2BA83ED2-C35A-49BC-8731-09A5F4BBB6C5" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1714419669962" checkinUser="felipe" docAuthor="Sirenko, Boris I." docDate="2018" docId="03A087CEBB70316AFE32FBC9FC26288C" docLanguage="en" docName="BullRusFarEastMalacolSoc.22.1-2.55-62.pdf" docOrigin="The Bulletin of the Russian Far East Malacological Society 22 (1 / 2)" docStyle="DocumentStyle:456D1B725C169D1BF26BD62A764ABD09.2:BullRusFarEastMalacolSoc.2013-2019.journal_article.english" docStyleId="456D1B725C169D1BF26BD62A764ABD09" docStyleName="BullRusFarEastMalacolSoc.2013-2019.journal_article.english" docStyleVersion="2" docTitle="Leptochitonidae Dall 1889" docType="treatment" docVersion="3" lastPageNumber="56" masterDocId="FF99FFB6BB71316BFFE2FFFFFFF82C7E" masterDocTitle="A new small chiton (Mollusca: Polyplacophora) from Guadelupe" masterLastPageNumber="62" masterPageNumber="55" pageNumber="56" updateTime="1727806048532" updateUser="valdenar" zenodo-license-document="CC-BY-4.0">
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<mods:title id="577829A775AA43D0C762519411B93972">A new small chiton (Mollusca: Polyplacophora) from Guadelupe</mods:title>
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<mods:namePart id="EC56E2C28F47D79C182E0A502E97BE41">Sirenko, Boris I.</mods:namePart>
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<heading id="D0FE81B4BB70316AFE32FBC9FC82282E" box="[464,890,1077,1104]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="56" reason="2">
Family
<taxonomicName id="4C094D5BBB70316AFDC8FBCAFC82282E" ID-CoL="7NJBT" authority="Dall, 1889" authorityName="Dall" authorityYear="1889" box="[554,890,1077,1104]" class="Polyplacophora" family="Leptochitonidae" kingdom="Animalia" order="Lepidopleurida" pageId="1" pageNumber="56" phylum="Mollusca" rank="family">
<emphasis id="B97DEACABB70316AFDC8FBCAFD0E2831" bold="true" box="[554,758,1077,1103]" pageId="1" pageNumber="56">Leptochitonidae</emphasis>
<bibRefCitation id="EF984B29BB70316AFD1FFBC9FC82282E" author="Dall W." box="[765,890,1078,1104]" pageId="1" pageNumber="56" pagination="1 - 492" refId="ref2379" refString="Dall W. 1889. Report on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. coast survey steamer &quot; Blake &quot;, Lieutenant-Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., commanding, 29, Report on the Mollusca. 2, Gastropoda and Scaphopoda // Bulletin of the Museum of Comparative Zoology, Harvard University. V. 18. P. 1 - 492." type="journal article" year="1889">Dall, 1889</bibRefCitation>
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</paragraph>
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Genus
<taxonomicName id="4C094D5BBB70316AFDA1FBA6FCA1280D" authority="Gray, 1847" authorityName="Gray" authorityYear="1847" box="[579,857,1113,1139]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
<emphasis id="B97DEACABB70316AFDA1FBA6FD37280D" box="[579,719,1113,1139]" italics="true" pageId="1" pageNumber="56">Leptochiton</emphasis>
Gray, 1847
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</paragraph>
<paragraph id="8BB636D8BB70316AFF01FB6DFB7D28B1" blockId="1.[166,1186,1169,1266]" pageId="1" pageNumber="56">
Ty p e s p e c i e s.
<emphasis id="B97DEACABB70316AFE2AFB6EFD7928D5" box="[456,641,1169,1195]" italics="true" pageId="1" pageNumber="56">Chiton cinereus</emphasis>
Montagu, 1803 (non Linne, 1767) =
<taxonomicName id="4C094D5BBB70316AFBD0FB6EFE1928B1" authority="(Gmelin, 1791)" baseAuthorityName="Gmelin" baseAuthorityYear="1791" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="1" pageNumber="56" phylum="Mollusca" rank="species" species="asellus">
<emphasis id="B97DEACABB70316AFBD0FB6EFEDC28B1" italics="true" pageId="1" pageNumber="56">Leptochiton asellus</emphasis>
(Gmelin, 1791)
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fide Lovén, 1846, subsequent designation by Gray, 1847.
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<paragraph id="8BB636D8BB70316AFF01FB27FC26288C" blockId="1.[166,1186,1169,1266]" box="[227,990,1240,1266]" pageId="1" pageNumber="56">G e n u s d i s t r i b u t i o n. Worldwide, Carboniferous Recent.</paragraph>
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</treatment>
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<taxonomicName id="4C094D5BBB70316AFE5AFAE4FCC8294B" ID-CoL="3TLXC" authority="Sirenko" authorityName="Sirenko" authorityYear="2018" box="[440,816,1307,1333]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="1" pageNumber="56" phylum="Mollusca" rank="species" species="guadelupei" status="sp. nov.">
<emphasis id="B97DEACABB70316AFE5AFAE4FD35294B" box="[440,717,1307,1333]" italics="true" pageId="1" pageNumber="56">Leptochiton guadelupei</emphasis>
Sirenko
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<taxonomicNameLabel id="A24E57B1BB70316AFCD5FAE3FC6A2948" box="[823,914,1308,1334]" pageId="1" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="13322A5DBB70316AFD8CFAC1FD232926" box="[622,731,1342,1368]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="2.[136,171,1604,1626]" captionStartId-1="3.[166,201,1632,1654]" captionStartId-2="4.[136,171,686,708]" captionStartId-3="5.[282,317,1370,1392]" captionTargetBox-0="[138,1156,387,1579]" captionTargetBox-1="[166,1186,720,1599]" captionTargetBox-2="[136,1156,227,658]" captionTargetBox-3="[222,1123,185,1336]" captionTargetId-0="figure-13@2.[120,1183,352,1617]" captionTargetId-1="figure-164@3.[149,1203,677,1632]" captionTargetId-2="figure-358@4.[118,1178,195,692]" captionTargetId-3="figure-67@5.[222,1123,180,1368]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="Fig. 1. Leptochition guadelupei, holotype (MNHN IM-2013-67001), BL 2.0 mm; A head valve, dorsal view; B valve II, dorsal view; C valve V, dorsal view; D tail valve, dorsal view; E valve V, ventral view; F surface of tegmentum in central area; G valve V, rostral view; H tail valve, lateral view." captionText-1="Fig. 2. Leptochition guadelupei, holotype (MNHN IM-2013-67001), BL 2.0 mm; A, C dorsal scales, marginal spicules and ventral scales; B dorsal scales; D part of radula." captionText-2="Fig. 3. Leptochition guadelupei, holotype (MNHN IM-2013-67001), BL 2.0 mm; A part of radula; B central and first lateral teeth of radula." captionText-3="Fig. 4. Leptochition guadelupei, holotype (MNHN IM-2013-67001), BL 2.0 mm; A sutural needle; B dorsal scale; C marginal needle; D ventral scale near margin; E ventral scale in middle part; F central and first lateral teeth of radula; G heads of major lateral teeth of radula; H aesthete group on central area. Scale bar 100 µm." figureDoi-0="http://doi.org/10.5281/zenodo.11087992" figureDoi-1="http://doi.org/10.5281/zenodo.11087994" figureDoi-2="http://doi.org/10.5281/zenodo.11087996" figureDoi-3="http://doi.org/10.5281/zenodo.11087998" httpUri-0="https://zenodo.org/record/11087992/files/figure.png" httpUri-1="https://zenodo.org/record/11087994/files/figure.png" httpUri-2="https://zenodo.org/record/11087996/files/figure.png" httpUri-3="https://zenodo.org/record/11087998/files/figure.png" pageId="1" pageNumber="56">Figs. 14</figureCitation>
</paragraph>
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<emphasis id="B97DEACABB70316AFF01FA8FFBF429F6" bold="true" box="[227,1036,1392,1416]" pageId="1" pageNumber="56">
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urn:lsid:zoobank.org:act:
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<materialsCitation id="3B613C85BB70316AFF01FA58FD072A79" collectionCode="MNHN, II, V, VIII" pageId="1" pageNumber="56" specimenCode="IM-2013-67001" specimenCount="1" typeStatus="holotype">
Ty p e m a t e r i a l.
<typeStatus id="54B2887ABB70316AFE3FFA58FDB229BF" box="[477,586,1447,1473]" pageId="1" pageNumber="56" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED18AE1DBB70316AFDB9FA58FD4F29BF" box="[603,695,1447,1473]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="56">MNHN</collectionCode>
<specimenCode id="DBAF9EA3BB70316AFD5DFA58FC8529BC" box="[703,893,1447,1474]" collectionCode="IM-" pageId="1" pageNumber="56">IM-2013-67001</specimenCode>
) now disarticulated consisting of SEM stub of valves I,
<collectionCode id="ED18AE1DBB70316AFDC3FA34FDCF299B" box="[545,567,1483,1509]" pageId="1" pageNumber="56">II</collectionCode>
,
<collectionCode id="ED18AE1DBB70316AFDA1FA34FDA0299B" box="[579,600,1483,1509]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="56" type="Museum">V</collectionCode>
,
<collectionCode id="ED18AE1DBB70316AFD81FA34FD60299B" box="[611,664,1483,1509]" pageId="1" pageNumber="56">VIII</collectionCode>
, part of perinotum and radula, mount of part of perinotum and radula and vial with other valves.
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</paragraph>
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<materialsCitation id="3B613C85BB70316AFF01F9EEFB5A2A55" box="[227,1186,1552,1579]" location="Western Atlantic" pageId="1" pageNumber="56" specimenCount="1" stateProvince="Guadeloupe">
Ty p e l o c a l i t y.
<location id="8ED66003BB70316AFE30F9EEFD622A54" LSID="urn:lsid:plazi:treatment:03A087CEBB70316DFE5AFAE4FCA82E45:8ED66003BB70316AFE30F9EEFD622A54" box="[466,666,1552,1579]" name="Western Atlantic" pageId="1" pageNumber="56" stateProvince="Guadeloupe">Western Atlantic</location>
, off
<collectingRegion id="49CDF83ABB70316AFD32F9EFFCAC2A54" box="[720,852,1552,1578]" country="France" name="Guadeloupe" pageId="1" pageNumber="56">Guadelupe</collectingRegion>
: Est Petite Terre, Campagne
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KARUBENTHOS 2012, stn. GD61,
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,
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, depth
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, BC5071,
<date id="FFB71018BB70316AFF44F9A8FED62A0C" box="[166,302,1623,1650]" pageId="1" pageNumber="56" value="2012-05-26">26.05.2012</date>
.
</paragraph>
<paragraph id="8BB636D8BB70316AFF01F984FD3A2AEA" blockId="1.[166,1187,1447,1685]" box="[227,706,1658,1685]" pageId="1" pageNumber="56">
E t y m o l o g y. Named after
<collectingRegion id="49CDF83ABB70316AFDD9F985FD462AEA" box="[571,702,1658,1684]" country="France" name="Guadeloupe" pageId="1" pageNumber="56">Guadelupe</collectingRegion>
.
</paragraph>
<paragraph id="8BB636D8BB733169FF27FF1BFCC72C83" blockId="2.[136,1157,227,359]" box="[197,831,227,254]" pageId="2" pageNumber="57">
D i s t r i b u t i o n. Known only from the
<typeStatus id="54B2887ABB733169FD4AFF1BFD232C80" box="[680,731,228,254]" pageId="2" pageNumber="57">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BB636D8BB733168FF27FEF8FEC72DD0" blockId="2.[136,1157,227,359]" lastBlockId="3.[166,1187,227,679]" lastPageId="3" lastPageNumber="58" pageId="2" pageNumber="57">D i a g n o s i s. Animal very small. Valves moderately elevated, carinated, not beaked. Lateral areas raised. Tail valve with mucro anterior. Pleural parts of central areas with well raised oval granules arranged in concentric as well in longitudinal rows. Head valve, lateral areas of intermediate valves and postmucronal area of tail valve with similar granules arranged quincuncially. Each granule with one megalaesthete and four micraesthetes. Dorsal scales wide, obtusely pointed, with 1617 distinct, double ribs, interstices very narrow. Radula with numerous transverse rows of very small teeth. Major lateral teeth of radula with unidentate cusp, which has deep longitudinal groove on each side.</paragraph>
<caption id="DF766650BB733169FF6AF9BBFBA02AEA" ID-DOI="http://doi.org/10.5281/zenodo.11087992" ID-Zenodo-Dep="11087992" httpUri="https://zenodo.org/record/11087992/files/figure.png" pageId="2" pageNumber="57" startId="2.[136,171,1604,1626]" targetBox="[138,1156,387,1579]" targetPageId="2" targetType="figure">
<paragraph id="8BB636D8BB733169FF6AF9BBFBA02AEA" blockId="2.[136,1157,1604,1684]" pageId="2" pageNumber="57">
<emphasis id="B97DEACABB733169FF6AF9BBFF3A2A27" bold="true" box="[136,194,1604,1626]" pageId="2" pageNumber="57">Fig. 1</emphasis>
.
<taxonomicName id="4C094D5BBB733169FF2DF9BBFE432A27" authorityName="Sirenko" authorityYear="2018" box="[207,443,1604,1625]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochition" kingdom="Animalia" order="Lepidopleurida" pageId="2" pageNumber="57" phylum="Mollusca" rank="species" species="guadelupei">
<emphasis id="B97DEACABB733169FF2DF9BBFE432A27" box="[207,443,1604,1625]" italics="true" pageId="2" pageNumber="57">Leptochition guadelupei</emphasis>
</taxonomicName>
,
<emphasis id="B97DEACABB733169FE25F9BBFDE72A27" bold="true" box="[455,543,1604,1625]" pageId="2" pageNumber="57">holotype</emphasis>
(MNHN IM-2013-67001), BL 2.0 mm;
<emphasis id="B97DEACABB733169FC4BF9BBFC422A27" bold="true" box="[937,954,1604,1625]" pageId="2" pageNumber="57">A</emphasis>
head valve, dorsal view;
<emphasis id="B97DEACABB733169FF26F99EFF2C2A08" bold="true" box="[196,212,1633,1654]" pageId="2" pageNumber="57">B</emphasis>
valve II, dorsal view;
<emphasis id="B97DEACABB733169FE5DF99EFE282A09" bold="true" box="[447,464,1633,1655]" pageId="2" pageNumber="57">C</emphasis>
valve V, dorsal view;
<emphasis id="B97DEACABB733169FD58F99EFD332A08" bold="true" box="[698,715,1633,1654]" pageId="2" pageNumber="57">D</emphasis>
tail valve, dorsal view;
<emphasis id="B97DEACABB733169FC27F99EFC2D2A08" bold="true" box="[965,981,1633,1654]" pageId="2" pageNumber="57">E</emphasis>
valve V, ventral view;
<emphasis id="B97DEACABB733169FF21F981FF2A2AED" bold="true" box="[195,210,1662,1683]" pageId="2" pageNumber="57">F</emphasis>
surface of tegmentum in central area;
<emphasis id="B97DEACABB733169FDB4F982FD912AED" bold="true" box="[598,617,1661,1683]" pageId="2" pageNumber="57">G</emphasis>
valve V, rostral view;
<emphasis id="B97DEACABB733169FCB1F981FC9E2AED" bold="true" box="[851,870,1662,1683]" pageId="2" pageNumber="57">H</emphasis>
tail valve, lateral view.
</paragraph>
</caption>
<paragraph id="8BB636D8BB723168FF01FE47FD392E67" blockId="3.[166,1187,227,679]" pageId="3" pageNumber="58">
<materialsCitation id="3B613C85BB723168FF01FE47FD392E67" collectionCode="V" pageId="3" pageNumber="58" specimenCount="1" typeStatus="holotype">
D e s c r i p t i o n.
<typeStatus id="54B2887ABB723168FE55FE47FDDC2DAC" box="[439,548,440,466]" pageId="3" pageNumber="58" type="holotype">Holotype</typeStatus>
very small, BL about 2.0 mm, elongate-oval, moderately elevated (elevation ratio in valve
<collectionCode id="ED18AE1DBB723168FD8EFE23FD792D88" box="[620,641,476,502]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="58" type="Museum">V</collectionCode>
0.45). Valves rather thick for the genus, carinated, not beaked. Color of tegmentum white.
</materialsCitation>
</paragraph>
<paragraph id="8BB636D8BB723168FF01FDDDFBE92ED9" blockId="3.[166,1187,227,679]" pageId="3" pageNumber="58">Head valve semicircular, wider than tail valve. Intermediate valves broadly rectangular, short and wide, anterior and posterior margins nearly straight, not beaked, lateral margins rounded. Lateral areas raised. Tail valve high, transversely elliptical with anterior mucro, antemucronal area convex, postmucronal area deeply concave.</paragraph>
<caption id="DF766650BB723168FF44F99FFC942AED" ID-DOI="http://doi.org/10.5281/zenodo.11087994" ID-Zenodo-Dep="11087994" httpUri="https://zenodo.org/record/11087994/files/figure.png" pageId="3" pageNumber="58" startId="3.[166,201,1632,1654]" targetBox="[166,1186,720,1599]" targetPageId="3" targetType="figure">
<paragraph id="8BB636D8BB723168FF44F99FFC942AED" blockId="3.[166,1187,1632,1683]" pageId="3" pageNumber="58">
<emphasis id="B97DEACABB723168FF44F99FFF102A0B" bold="true" box="[166,232,1632,1654]" pageId="3" pageNumber="58">Fig. 2.</emphasis>
<taxonomicName id="4C094D5BBB723168FF12F99EFE242A08" authorityName="Sirenko" authorityYear="2018" box="[240,476,1633,1654]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochition" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="58" phylum="Mollusca" rank="species" species="guadelupei">
<emphasis id="B97DEACABB723168FF12F99EFE242A08" box="[240,476,1633,1654]" italics="true" pageId="3" pageNumber="58">Leptochition guadelupei</emphasis>
</taxonomicName>
,
<emphasis id="B97DEACABB723168FE08F99EFDBA2A08" bold="true" box="[490,578,1633,1654]" pageId="3" pageNumber="58">holotype</emphasis>
(MNHN IM-2013-67001), BL 2.0 mm;
<emphasis id="B97DEACABB723168FC36F99FFC1D2A0B" bold="true" box="[980,997,1632,1653]" pageId="3" pageNumber="58">A</emphasis>
,
<emphasis id="B97DEACABB723168FC11F99FFBFC2A08" bold="true" box="[1011,1028,1632,1654]" pageId="3" pageNumber="58">C</emphasis>
dorsal scales, marginal spicules and ventral scales;
<emphasis id="B97DEACABB723168FDEEF982FDE42AEC" bold="true" box="[524,540,1661,1682]" pageId="3" pageNumber="58">B</emphasis>
dorsal scales;
<emphasis id="B97DEACABB723168FD58F982FD332AEC" bold="true" box="[698,715,1661,1682]" pageId="3" pageNumber="58">D</emphasis>
part of radula.
</paragraph>
</caption>
<caption id="DF766650BB75316FFF6AFD51FDE72E9F" ID-DOI="http://doi.org/10.5281/zenodo.11087996" ID-Zenodo-Dep="11087996" httpUri="https://zenodo.org/record/11087996/files/figure.png" pageId="4" pageNumber="59" startId="4.[136,171,686,708]" targetBox="[136,1156,227,658]" targetPageId="4" targetType="figure">
<paragraph id="8BB636D8BB75316FFF6AFD51FDE72E9F" blockId="4.[136,1157,686,737]" pageId="4" pageNumber="59">
<emphasis id="B97DEACABB75316FFF6AFD51FF332EBD" bold="true" box="[136,203,686,708]" pageId="4" pageNumber="59">Fig. 3.</emphasis>
<emphasis id="B97DEACABB75316FFF37FD51FE312EBD" box="[213,457,686,707]" italics="true" pageId="4" pageNumber="59">
<taxonomicName id="4C094D5BBB75316FFF37FD51FE3D2EBD" authorityName="Sirenko" authorityYear="2018" box="[213,453,686,707]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochition" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="59" phylum="Mollusca" rank="species" species="guadelupei">Leptochition guadelupei</taxonomicName>
,
</emphasis>
<emphasis id="B97DEACABB75316FFE30FD51FDD22EBD" bold="true" box="[466,554,686,707]" pageId="4" pageNumber="59">holotype</emphasis>
(MNHN IM-2013-67001), BL 2.0 mm;
<emphasis id="B97DEACABB75316FFC27FD51FC2E2EBD" bold="true" box="[965,982,686,707]" pageId="4" pageNumber="59">A</emphasis>
part of radula;
<emphasis id="B97DEACABB75316FFF6AFD34FF602E9E" bold="true" box="[136,152,715,736]" pageId="4" pageNumber="59">B</emphasis>
central and first lateral teeth of radula.
</paragraph>
</caption>
<paragraph id="8BB636D8BB75316FFF27FCF4FDBF2FAA" blockId="4.[136,1157,778,1685]" pageId="4" pageNumber="59">Tegmentum sculptured with well raised oval granules arranged in more or less quincuncial pattern on head valve, the lateral areas of intermediate valves and on the postmucronal area of tail valve, in longitudinal as well in concentric rows on pleural parts of central areas of intermediate valves and on antemucronal area of tail valve. Jugal areas with flattened granules arranged in a random manner. Each granule has one megalaesthete and four micraesthetes.</paragraph>
<paragraph id="8BB636D8BB75316FFF27FC22FBED2F89" blockId="4.[136,1157,778,1685]" box="[197,1045,989,1015]" pageId="4" pageNumber="59">Articulamentum well developed, apophyses small and widely separated.</paragraph>
<paragraph id="8BB636D8BB75316FFF27FBFFFEE028B4" blockId="4.[136,1157,778,1685]" pageId="4" pageNumber="59">Girdle very narrow, covered with wide and obtusely pointed dorsal scales (50x50 μm) with 1617 distinct double ribs. Sutures between valves armed with few long and ribbed spicules (210x22 μm). Marginal spicules elongate (110x18 μm) with 67 distinct ribs arranged around the spicule. Ventral scales in central part pointed, smooth (42x18 μm) those near margin have 45 short, weak ribs on the distal half in ventral side.</paragraph>
<paragraph id="8BB636D8BB75316FFF27FB2BFD332929" blockId="4.[136,1157,778,1685]" pageId="4" pageNumber="59">
Radula of the
<typeStatus id="54B2887ABB75316FFE95FB2BFE262890" box="[375,478,1236,1262]" pageId="4" pageNumber="59" type="holotype">holotype</typeStatus>
<quantity id="4CF19B3DBB75316FFE0BFB2BFDBF2890" box="[489,583,1236,1262]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="59" unit="mm" value="0.7">0.7 mm</quantity>
long with about 40 transverse rows of mature teeth. Central teeth bulbose near the point base, distal half narrow, first lateral teeth with a narrow cusp and an outward projecting wing, major lateral teeth with unidentate cusp, which has deep longitudinal groove on each side.
</paragraph>
<paragraph id="8BB636D8BB75316FFF27FA9FFC282905" blockId="4.[136,1157,778,1685]" box="[197,976,1376,1403]" pageId="4" pageNumber="59">The gills were not counted as the specimen was dry and curled up.</paragraph>
<paragraph id="8BB636D8BB75316FFF27FA7BFC2F2A54" blockId="4.[136,1157,778,1685]" pageId="4" pageNumber="59">
<collectionCode id="ED18AE1DBB75316FFF27FA7BFF2129E0" box="[197,217,1412,1438]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="59">R</collectionCode>
e m a r k s. New species differs from other species of genus
<taxonomicName id="4C094D5BBB75316FFC42FA7BFBD429E0" authorityName="Gray" authorityYear="1847" box="[928,1068,1412,1438]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="59" phylum="Mollusca" rank="genus">
<emphasis id="B97DEACABB75316FFC42FA7BFBD429E0" box="[928,1068,1412,1438]" italics="true" pageId="4" pageNumber="59">Leptochiton</emphasis>
</taxonomicName>
known from the western Atlantic (
<taxonomicName id="4C094D5BBB75316FFE02FA57FD9129BF" baseAuthorityName="Loven" baseAuthorityYear="1846" box="[480,617,1447,1473]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="59" phylum="Mollusca" rank="species" species="alveolus">
<emphasis id="B97DEACABB75316FFE02FA57FD9129BF" box="[480,617,1447,1473]" italics="true" pageId="4" pageNumber="59">L. alveolus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C094D5BBB75316FFD9CFA57FCF029BF" authorityName="Plate" authorityYear="1899" box="[638,776,1447,1473]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="59" phylum="Mollusca" rank="species" species="medinae">
<emphasis id="B97DEACABB75316FFD9CFA57FCF029BF" box="[638,776,1447,1473]" italics="true" pageId="4" pageNumber="59">L. medinae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C094D5BBB75316FFCFEFA57FC2A29BF" authorityName="Dall" authorityYear="1889" box="[796,978,1448,1473]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="59" phylum="Mollusca" rank="species" species="pergranatus">
<emphasis id="B97DEACABB75316FFCFEFA57FC2A29BF" box="[796,978,1448,1473]" italics="true" pageId="4" pageNumber="59">L. pergranatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C094D5BBB75316FFC05FA57FB8529BF" baseAuthorityName="Boone" baseAuthorityYear="1928" box="[999,1149,1447,1473]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="59" phylum="Mollusca" rank="species" species="binghami">
<emphasis id="B97DEACABB75316FFC05FA57FB8529BF" box="[999,1149,1447,1473]" italics="true" pageId="4" pageNumber="59">L. binghami</emphasis>
</taxonomicName>
,
<taxonomicName id="4C094D5BBB75316FFF6AFA34FF0A299A" authorityName="Righi" authorityYear="1973" box="[136,242,1482,1508]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="59" phylum="Mollusca" rank="species" species="darioi">
<emphasis id="B97DEACABB75316FFF6AFA34FF0A299A" box="[136,242,1482,1508]" italics="true" pageId="4" pageNumber="59">L. darioi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C094D5BBB75316FFEE7FA34FE1E299A" authorityName="Kaas" authorityYear="1994" box="[261,486,1482,1508]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="59" phylum="Mollusca" rank="species" species="micropustulosus">
<emphasis id="B97DEACABB75316FFEE7FA34FE1E299A" box="[261,486,1482,1508]" italics="true" pageId="4" pageNumber="59">L. micropustulosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C094D5BBB75316FFDC7FA34FCFE299A" authorityName="Carmona Zalvide &amp; Urgorri" authorityYear="1999" box="[549,774,1482,1508]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="59" phylum="Mollusca" rank="species" species="compostellanum">
<emphasis id="B97DEACABB75316FFDC7FA34FCFE299A" box="[549,774,1482,1508]" italics="true" pageId="4" pageNumber="59">L. compostellanum</emphasis>
</taxonomicName>
) by having unique sculpture of tegmentum in the form of concentric rows of granules and unidentate cusp of major lateral tooth of radula, which has deep longitudinal groove on each side.
</paragraph>
<paragraph id="8BB636D8BB75316DFF27F9CBFCA82E45" blockId="4.[136,1157,778,1685]" lastBlockId="6.[136,1157,227,571]" lastPageId="6" lastPageNumber="61" pageId="4" pageNumber="59">
<taxonomicName id="4C094D5BBB75316FFF27F9CBFE9E2A30" authority="Sirenko, 2018" authorityName="Sirenko" authorityYear="2018" box="[197,358,1588,1614]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochition" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="59" phylum="Mollusca" rank="species" species="guadelupei" status="sp. nov.">
<emphasis id="B97DEACABB75316FFF27F9CBFE9E2A30" box="[197,358,1588,1614]" italics="true" pageId="4" pageNumber="59">L. guadelupei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B1BB75316FFE8DF9CBFE322A30" box="[367,458,1588,1614]" pageId="4" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
is more similar to the eastern Atlantic species of the genus
<taxonomicName id="4C094D5BBB75316FFF6AF9A8FEEC2A0F" authorityName="Gray" authorityYear="1847" box="[136,276,1623,1649]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="59" phylum="Mollusca" rank="genus">
<emphasis id="B97DEACABB75316FFF6AF9A8FEEC2A0F" box="[136,276,1623,1649]" italics="true" pageId="4" pageNumber="59">Leptochiton</emphasis>
</taxonomicName>
(
<taxonomicName id="4C094D5BBB75316FFEC5F9A7FDD12A0F" authority="Kaas, 1979" authorityName="Kaas" authorityYear="1979" box="[295,553,1623,1649]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="59" phylum="Mollusca" rank="species" species="leloupi">
<emphasis id="B97DEACABB75316FFEC5F9A7FE612A0F" box="[295,409,1623,1649]" italics="true" pageId="4" pageNumber="59">L. leloupi</emphasis>
Kaas, 1979
</taxonomicName>
and
<taxonomicName id="4C094D5BBB75316FFD84F9A7FC982A0F" authority="Kaas, 1979" authorityName="Kaas" authorityYear="1979" box="[614,864,1623,1649]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="59" phylum="Mollusca" rank="species" species="tenuis">
<emphasis id="B97DEACABB75316FFD84F9A7FD352A0F" box="[614,717,1624,1649]" italics="true" pageId="4" pageNumber="59">L. tenuis</emphasis>
Kaas, 1979
</taxonomicName>
). Both are from the Bay of Biscay. The new species differs from the other ones by having 1617 double ribs on dorsal scales (vs. 1214 single ribs in
<taxonomicName id="4C094D5BBB74316EFD4EF9EEFCD82A54" authorityName="Kaas" authorityYear="1979" box="[684,800,1552,1578]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="60" phylum="Mollusca" rank="species" species="leloupi">
<emphasis id="B97DEACABB74316EFD4EF9EEFCD82A54" box="[684,800,1552,1578]" italics="true" pageId="5" pageNumber="60">L. leloupi</emphasis>
</taxonomicName>
), the anterior mucro (vs. about central in
<taxonomicName id="4C094D5BBB74316EFEC2F9CBFE6A2A33" authorityName="Kaas" authorityYear="1979" box="[288,402,1587,1613]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="60" phylum="Mollusca" rank="species" species="leloupi">
<emphasis id="B97DEACABB74316EFEC2F9CBFE6A2A33" box="[288,402,1587,1613]" italics="true" pageId="5" pageNumber="60">L. leloupi</emphasis>
</taxonomicName>
), pleural parts of the central areas with well raised oval granules arranged in concentric and longitudinal rows (vs central areas with rather widely separated longitudinal chains of irregularly shaped tubercles mostly composed of several granules cemented together in
<taxonomicName id="4C094D5BBB77316DFE1EFF1BFD962C83" authorityName="Kaas" authorityYear="1979" box="[508,622,227,253]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="leloupi">
<emphasis id="B97DEACABB77316DFE1EFF1BFD962C83" box="[508,622,227,253]" italics="true" pageId="6" pageNumber="61">L. leloupi</emphasis>
</taxonomicName>
). The new species differs from
<taxonomicName id="4C094D5BBB77316DFC17FF1BFBA52C83" authorityName="Kaas" authorityYear="1979" box="[1013,1117,228,253]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="tenuis">
<emphasis id="B97DEACABB77316DFC17FF1BFBA52C83" box="[1013,1117,228,253]" italics="true" pageId="6" pageNumber="61">L. tenuis</emphasis>
</taxonomicName>
by having broader dorsal scales 50 μm long, 50 μm wide, with 1617 double ribs on the dorsal scales (vs. 6070 μm long, 3540 μm wide with 1012 single ribs in
<taxonomicName id="4C094D5BBB77316DFBF2FED5FB8B2D3D" authorityName="Kaas" authorityYear="1979" box="[1040,1139,298,323]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="tenuis">
<emphasis id="B97DEACABB77316DFBF2FED5FB8B2D3D" box="[1040,1139,298,323]" italics="true" pageId="6" pageNumber="61">L. tenuis</emphasis>
</taxonomicName>
), the anterior mucro (vs. the mucro behind the center in
<taxonomicName id="4C094D5BBB77316DFD16FEB1FCAF2D19" authorityName="Kaas" authorityYear="1979" box="[756,855,334,359]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="tenuis">
<emphasis id="B97DEACABB77316DFD16FEB1FCAF2D19" box="[756,855,334,359]" italics="true" pageId="6" pageNumber="61">L. tenuis</emphasis>
</taxonomicName>
), carinated valves (vs. not carinated in
<taxonomicName id="4C094D5BBB77316DFEFAFE8EFE852DF4" authorityName="Kaas" authorityYear="1979" box="[280,381,369,394]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="tenuis">
<emphasis id="B97DEACABB77316DFEFAFE8EFE852DF4" box="[280,381,369,394]" italics="true" pageId="6" pageNumber="61">L. tenuis</emphasis>
</taxonomicName>
).
<taxonomicName id="4C094D5BBB77316DFE74FE8EFDCE2DF4" authorityName="Sirenko" authorityYear="2018" box="[406,566,368,394]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochition" kingdom="Animalia" order="Lepidopleurida" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="guadelupei">
<emphasis id="B97DEACABB77316DFE74FE8EFDCE2DF4" box="[406,566,368,394]" italics="true" pageId="6" pageNumber="61">L. guadelupei</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="4C094D5BBB77316DFCE4FE8EFF3B2DD3" authority="Schwabe et Lozouet, 2006" authorityName="Schwabe et Lozouet" authorityYear="2006" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="hiriensis">
<emphasis id="B97DEACABB77316DFCE4FE8EFC712DF4" box="[774,905,368,394]" italics="true" pageId="6" pageNumber="61">L. hiriensis</emphasis>
Schwabe et Lozouet, 2006
</taxonomicName>
from Rapa Island, southern Pacific by the sculpture of tegmentum, but differs by having the unidentate cusp of the major lateral tooth of radula, which has a deep longitudinal groove on each side (vs. tridentate cusp in
<taxonomicName id="4C094D5BBB77316DFD58FE24FCC52D8A" authorityName="Schwabe et Lozouet" authorityYear="2006" box="[698,829,474,500]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="hiriensis">
<emphasis id="B97DEACABB77316DFD58FE24FCC52D8A" box="[698,829,474,500]" italics="true" pageId="6" pageNumber="61">L. hiriensis</emphasis>
</taxonomicName>
), 1617 distinct double ribs (vs. up to 15 simple ribs in
<taxonomicName id="4C094D5BBB77316DFE2EFE01FDA82E69" authorityName="Schwabe et Lozouet" authorityYear="2006" box="[460,592,509,536]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="hiriensis">
<emphasis id="B97DEACABB77316DFE2EFE01FE242E69" box="[460,476,510,535]" italics="true" pageId="6" pageNumber="61">L</emphasis>
.
<emphasis id="B97DEACABB77316DFE09FE02FDA82E69" box="[491,592,509,535]" italics="true" pageId="6" pageNumber="61">hiriensis</emphasis>
</taxonomicName>
) and the high tail valve with the anterior mucro (vs. the low tail valve with the central mucro in
<taxonomicName id="4C094D5BBB77316DFD59FDDEFCC72E44" authorityName="Schwabe et Lozouet" authorityYear="2006" box="[699,831,544,571]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="hiriensis">
<emphasis id="B97DEACABB77316DFD59FDDEFD332E44" box="[699,715,545,570]" italics="true" pageId="6" pageNumber="61">L</emphasis>
.
<emphasis id="B97DEACABB77316DFD38FDDFFCC72E44" box="[730,831,544,570]" italics="true" pageId="6" pageNumber="61">hiriensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>