Add updates up until 2025-03-17 17:54:28

This commit is contained in:
ggserver 2025-03-17 17:59:32 +00:00
parent 955a7f1e72
commit acf44a3306
9 changed files with 3619 additions and 213 deletions

View file

@ -0,0 +1,783 @@
<document id="1A03D3199091FAC65D422C76D7450B25" ID-DOI="10.3897/zookeys.1232.141693" ID-ZooBank="0AA099A2-EA57-4E27-937A-0976B9F8A4B5" ID-publisher-id="141693" URI-arpha="A9904ED3-9BDA-52DF-B936-1C42EC2713C5" URI-zoobank="http://zoobank.org/0AA099A2-EA57-4E27-937A-0976B9F8A4B5" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1742225059022" checkinUser="pensoft" docAuthor="Arredondo, Brandon S. &amp; Guerrero, Roberto J." docDate="2025" docId="3DBF20F901035E869B89A4EE102E7899" docLanguage="en" docName="ZooKeys 1232: 131-172" docOrigin="ZooKeys 1232" docSource="https://zookeys.pensoft.net/article/141693/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nesomyrmex asper" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="172" masterDocId="A9904ED39BDA52DFB9361C42EC2713C5" masterDocTitle="The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data" masterLastPageNumber="172" masterPageNumber="131" pageNumber="131" updateTime="1742234045672" updateUser="admin">
<mods:mods id="78C9C7A2518D895CFF6D8B3306DE50C3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F661CC9CE522213A365E8636C34296BE">
<mods:title id="6A86DD02E364BB0E979F40FEC7D34D22">The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data</mods:title>
</mods:titleInfo>
<mods:name id="9305AB3CA1EE2EE1BBF186B2FC15E594" type="personal">
<mods:role id="0E4C748A47BDF59B053A01F5594C6BB8">
<mods:roleTerm id="AB83BF4AC56E031CDE08942DD31163BC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D5579BF95A606F052BC050721D52883F">Arredondo, Brandon S.</mods:namePart>
<mods:nameIdentifier id="97E4EC0B9DDFA81CC129BE79EAE071B3" type="ORCID">0000-0003-2777-5909</mods:nameIdentifier>
<mods:affiliation id="0BBD481B207668798BA613744EC72645">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:name id="46CE221A9307F5E84A189CFE9A148192" type="personal">
<mods:role id="45E408DFD05F0C5AA79D689FB90260A6">
<mods:roleTerm id="B7B2D15BD39FA084F1DDFFCDEE365A0E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F9280E63579D06A45FBDFB0729F77343">Guerrero, Roberto J.</mods:namePart>
<mods:nameIdentifier id="6660F712A6FA1FB56DCCF8C38257B8EF" type="ORCID">0000-0003-3244-2754</mods:nameIdentifier>
<mods:affiliation id="F604CFFA431102B60699A5FC3FBBB977">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="93AE6EC37A527A25A113E7B1393CCA0C">text</mods:typeOfResource>
<mods:relatedItem id="15BFCBFADE0A41C33B5853711907E9A7" type="host">
<mods:titleInfo id="BC8E04DA508F3F4E0611E681AD3A7D0E">
<mods:title id="9CB5ED045AB9A7CF786EE18147626D5E">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="6FEC70548F62A54A361D3B30BCA2D6B8">
<mods:date id="3BB99EC89367429AD5BB97B4B5B696FE">2025</mods:date>
<mods:detail id="D1EF1DD88595FD6075D78878C6C72D75" type="pubDate">
<mods:number id="7319556F396F6292A7249CF0D25CECF3">2025-03-17</mods:number>
</mods:detail>
<mods:detail id="13BCC71FB5EDBD8C2AF08FE493F3E378" type="volume">
<mods:number id="8E4041685F4EC6B206F2063955B19EB3">1232</mods:number>
</mods:detail>
<mods:extent id="7D2469CAD2251623D088C0BC2FE7BEC6" unit="page">
<mods:start id="A87587C52398C66CD22ACE9536A88169">131</mods:start>
<mods:end id="9177E84C1524EAD87CF15E91B313538C">172</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5041C76A561ABC7C76F19F3216C74B65">journal article</mods:classification>
<mods:identifier id="24ADB047EF412B51166BB5B92C6B6BD4" type="DOI">10.3897/zookeys.1232.141693</mods:identifier>
<mods:identifier id="25EC59CD3728B9D08FF36A3E0D4AEE55" type="ZooBank">0AA099A2-EA57-4E27-937A-0976B9F8A4B5</mods:identifier>
</mods:mods>
<treatment id="3DBF20F901035E869B89A4EE102E7899" ID-arpha="3DBF20F9-0103-5E86-9B89-A4EE102E7899" LSID="urn:lsid:plazi:treatment:3DBF20F901035E869B89A4EE102E7899" httpUri="http://treatment.plazi.org/id/3DBF20F901035E869B89A4EE102E7899" scope_family="Formicidae" scope_subFamily="Myrmicinae">
<subSubSection id="FB7DBD6901C6BE62B07AED9B8F00899C" type="nomenclature">
<paragraph id="663910B115BF8D90FF29D0145B53A82D">
<taxonomicName id="9D30A9E0E53BB2109BAAF22075F391E4" ID-CoL="CM7WW" authority="(Mayr, 1887)" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="A36CC7576B4CB3E49AB0F31A8AA7A368" italics="true">Nesomyrmex asper</emphasis>
(
<bibRefCitation id="87ED0F6CB05DE02FCCC95811A162B062" author="Mayr G" journalOrPublisher="Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien" pagination="511-632" refId="B41" refString="Mayr G (1887) Sudamerikanische Formiciden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 37: 511-632." title="Südamerikanische Formiciden." volume="37" year="1887">Mayr, 1887</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="52AE3BECB622A527532401B6C5A5CB78" type="description">
<paragraph id="9203CD22772FE626C592F6171662C97B">
<figureCitation id="AF440227E9DFA1A16A7CAA2C59398E3D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Variation of lateral petiole tubercles in Nesomyrmex asper. White arrows indicate the tubercles." figureDoi="10.3897/zookeys.1232.141693.figure18" httpUri="https://binary.pensoft.net/fig/1282988">Figs 18</figureCitation>
,
<figureCitation id="D8C12ADE9F2F85419116785E19A31329" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Lateral petiole and postpetiole tubercles (Setigerous tubercles). Abbreviations: lpptt (lateral postpetiolar tubercles), lptt (lateral petiolar tubercles), pt (petiole), ppt (postpetiole), set (seta)." figureDoi="10.3897/zookeys.1232.141693.figure19" httpUri="https://binary.pensoft.net/fig/1282989">19</figureCitation>
,
<figureCitation id="96938132EA9790ADF78A2D4B8178C053" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Nesomyrmex asper worker (ICN 106553) A full-face view B lateral view C dorsal view. Scale bars: 0.5 mm (A); 1.0 mm (B)." figureDoi="10.3897/zookeys.1232.141693.figure20" httpUri="https://binary.pensoft.net/fig/1282990">20</figureCitation>
,
<figureCitation id="8C8744A4D34F2D8DB067593AA2D76828" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distributional map of Nesomyrmex species from the tropical dry forest in Colombia." figureDoi="10.3897/zookeys.1232.141693.figure29" httpUri="https://binary.pensoft.net/fig/1282999">29</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="772C65A692E35690B92720231FD91320" type="reference_group">
<paragraph id="C22976DA5723D456AAE2F7DFFE282116">
<treatmentCitationGroup id="E09ABC888634C09559D9775BC967BA6A">
<treatmentCitation id="BE0921460E9D2A5D01F1B3E0C433F3C5" author="Mayr G" page="618" year="1887">
<taxonomicName id="817B98540F1C8B54553913FA3A1570EB" ID-CoL="3TXX9" authority="Mayr, 1887: 618" authorityName="Mayr" authorityPageNumber="618" authorityYear="1887" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="DD767261C691731E26426CA0D85F48D0" italics="true">Leptothorax asper</emphasis>
<comment id="B1A9B9037DFC53DBAE0FADA8626F618D">
<bibRefCitation id="12335E9D03F801B911CCAA935AE86F52" author="Mayr G" journalOrPublisher="Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien" pagination="511-632" refId="B41" refString="Mayr G (1887) Sudamerikanische Formiciden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 37: 511-632." title="Südamerikanische Formiciden." volume="37" year="1887">Mayr, 1887: 618</bibRefCitation>
.
<typeStatus id="EFF80CD675C4EDD14426450BE0A9100E" type="syntype">Syntype</typeStatus>
series (worker, queen, male):
<collectingCountry id="A799998F4F3B639A6754375553154285" name="Brazil">Brazil</collectingCountry>
,
<collectingRegion id="E25A671C343486E562C77794F9C97CE0" country="Brazil" name="Santa Catarina">Santa Catarina</collectingRegion>
. [
<named-content id="2AAE05EB9642112A3F0EEC3C57BFA432" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum, Vienna">
<collectionCode id="DAD4AD6C86787630F956CAD95EE06352" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
</named-content>
] (AntWeb image examined,
<ext-link id="04BC008116C8141D3969EFC4B0E59E5B" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0901801" xlink_type="simple">
<specimenCode id="F353D03F455FF770700C42A201129FFC">CASENT 0901801</specimenCode>
</ext-link>
).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="939D83B2E9C9F4D3F7751AB87B87C85C">
Combination in
<taxonomicName id="4E5DE05BDD8CF16E5759D8D2B1F1BEE2" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Goniothorax">
<emphasis id="6275780EA4AEEBC96546D48E315679C8" italics="true">Leptothorax</emphasis>
(
<emphasis id="95E32DBAF2EA430D8F6C9D4DB6FF11B9" italics="true">Goniothorax</emphasis>
)
</taxonomicName>
:
<bibRefCitation id="14507507281123479C870E96A5117106" author="Emery C" firstAuthor="Emery" journalOrPublisher="Bullettino della Società Entomologica Italiana" page="59" pagination="33-107" refId="B23" refString="Emery C (1896) Studi sulle formiche della fauna neotropica. XVII-XXV. Bullettino della Società Entomologica Italiana 28: 33107." title="Studi sulle formiche della fauna neotropica. XVII-XXV." volume="28" year="1896">Emery 1896: 59</bibRefCitation>
.
</paragraph>
<paragraph id="977FB7CAB86A0883A897889E5C553C76">
<treatmentCitationGroup id="79AB1AE10A445981A90A0236B70671A0">
<treatmentCitation id="8091038E80948935D5CDD565FCCF64E8" ID-DOI="http://doi.org/10.5281/zenodo.6292143" ID-Zenodo-Dep="6292143" author="Emery C" httpUri="http://treatment.plazi.org/id/178BB12BD21ADF9DD2AFA7AFACBF5AC0" page="61" year="1896">
<taxonomicName id="5B853070AFC36F9AF297871D0D197276" authority="Emery, 1896: 61" authorityName="Emery" authorityPageNumber="61" authorityYear="1896" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="variety" species="asper" subGenus="Goniothorax" variety="rufa">
<emphasis id="C3046E698945A6C1DA2E85390AE024C0" italics="true">Leptothorax</emphasis>
(
<emphasis id="86048BFFD1855EE496B7054B23732389" italics="true">Goniothorax</emphasis>
)
<emphasis id="031BAD103B98E72DFA81D23CBF32CDD3" italics="true">asper var. rufa</emphasis>
<comment id="15CF9F2572448D2B88F43EDDA8392757">
<bibRefCitation id="6C0AC6EEAABEE0405D1FE8526DFC18DF" author="Emery C" journalOrPublisher="Bullettino della Società Entomologica Italiana" pagination="33-107" refId="B23" refString="Emery C (1896) Studi sulle formiche della fauna neotropica. XVII-XXV. Bullettino della Societa Entomologica Italiana 28: 33-107." title="Studi sulle formiche della fauna neotropica. XVII-XXV." volume="28" year="1896">Emery, 1896: 61</bibRefCitation>
.
<typeStatus id="0DC5D7201637F13A3EB7DF22EA877AAF" type="syntype">Syntype</typeStatus>
series (queen, worker):
<collectingCountry id="AE1D5FE4999B599272D414F44E9D0F4F" name="Brazil">Brazil</collectingCountry>
, Pará. [
<named-content id="9A1F4748C9E1F427056829C18024CE4B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-civico-di-storia-naturale-di-genova-giacomo-doria" xlink_title="Museo di Storia Naturale Giacomo Doria'">
<collectionCode id="3C72927C249CA40CA9CE9D8F24999477" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" type="">MSNG</collectionCode>
</named-content>
]. (AntWeb image examined,
<ext-link id="4B1B0DBF0399C18419888B047D352B2E" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0904722" xlink_type="simple">
<specimenCode id="DFF2C1DB65E11168FD7BC534A0462A14">CASENT 0904722</specimenCode>
</ext-link>
). Synonymy by
<treatmentCitation id="ED5754A486917E4379A0358E0B25A30D" author="Kempf WW" page="414" year="1959">
<bibRefCitation id="A57FCBDC091F02EE9C0D9E23FB82FB46" author="Kempf WW" firstAuthor="Kempf" journalOrPublisher="Studia Entomologica" page="414" pagination="391-432" refId="B37" refString="Kempf WW (1959) A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae). Studia Entomologica 2: 391432." title="A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae)." volume="2" year="1959">Kempf 1959: 414</bibRefCitation>
</treatmentCitation>
.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="4C5148C8F4741F88165D46D7019676C6">
<treatmentCitationGroup id="5D3B67DD6088A61C169D71595E1490F3">
<treatmentCitation id="7C1B344978088DF048AB289141F0A2D6" author="Emery C" page="61" year="1896">
<taxonomicName id="906E0E0F9CF24FCCC0190AF09B41DA17" ID-CoL="3TY7L" authority="Emery, 1896: 61" authorityName="Emery" authorityPageNumber="61" authorityYear="1896" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tristani" subGenus="Goniothorax">
<emphasis id="EDBE9B98DDF7A8F0B762FACA9D64553D" italics="true">Leptothorax</emphasis>
(
<emphasis id="63981F84213BE4B97BC1577C9DB72F62" italics="true">Goniothorax</emphasis>
)
<emphasis id="053065969AE71EFDE478F1FB39663CDB" italics="true">tristani</emphasis>
<comment id="16980140A908E7512E3A04A295EBCF2B">
<bibRefCitation id="47FF1F2D9C637B03543E4403927071B2" author="Emery C" journalOrPublisher="Bullettino della Società Entomologica Italiana" pagination="33-107" refId="B23" refString="Emery C (1896) Studi sulle formiche della fauna neotropica. XVII-XXV. Bullettino della Societa Entomologica Italiana 28: 33-107." title="Studi sulle formiche della fauna neotropica. XVII-XXV." volume="28" year="1896">Emery, 1896: 61</bibRefCitation>
.
<typeStatus id="028B61F9437B3ED1C516A5E8DB6EBD8C" type="syntype">Syntype</typeStatus>
series (queen, worker):
<collectingCountry id="818444A17FEF48FF5FBAC869AC18AD2B" name="Costa Rica">Costa Rica</collectingCountry>
. [
<named-content id="C2082F00D6DFC793351744349D521635" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-civico-di-storia-naturale-di-genova-giacomo-doria" xlink_title="Museo di Storia Naturale Giacomo Doria'">
<collectionCode id="4478BE48E31A4A6E2ECFBE2167198C09" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" type="">MSNG</collectionCode>
</named-content>
]. (AntWeb image examined,
<ext-link id="458AF95AA9331F6F5A4FDE2A3ED754DA" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0904724" xlink_type="simple">
<specimenCode id="B86B477B7B2362565B3D9B9D47E28575">CASENT 0904724</specimenCode>
</ext-link>
). Synonymy by
<treatmentCitation id="413895FB0C4310EFD6F48BBE6E139541" author="Longino JT" page="136" year="2006">
<bibRefCitation id="2C60DF87BB4CDF4CA57F922F3D401B7D" author="Longino JT" firstAuthor="Longino" journalOrPublisher="Myrmecological News" page="136" pagination="131-143" refId="B40" refString="Longino JT (2006) New species and nomenclatural changes for the Costa Rican ant fauna (Hymenoptera: Formicidae). Myrmecological News 8: 131143." title="New species and nomenclatural changes for the Costa Rican ant fauna (Hymenoptera: Formicidae)." volume="8" year="2006">Longino 2006: 136</bibRefCitation>
</treatmentCitation>
.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="97AA22CC17DA41E1B3A1346D3983B731">
<treatmentCitationGroup id="3E441E0CBC2AFE3D003D6DB9E11DEE7F">
<treatmentCitation id="230C8050087381DDE6C99215C0615948" author="Forel A" page="18" year="1912">
<taxonomicName id="0F770D69DF912EEC35CF773108D9AC12" authority="Forel, 1912: 18" authorityName="Forel" authorityPageNumber="18" authorityYear="1912" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="variety" species="asper" variety="antoniensis">
<emphasis id="DB9ACFE4559FA5B805148F420882F790" italics="true">Leptothorax asper var. antoniensis</emphasis>
<comment id="494040B446D6CDC206C26EDEDB1BBA07">
<bibRefCitation id="8D409D20EAC02D340D1A1D506238411C" DOI="10.5962/bhl.part.19251" author="Forel A" journalOrPublisher="Mémoires de la Société Entomologique de Belgique" pagination="1-32" refId="B30" refString="Forel A (1912) Formicides neotropiques. Part IV. 3 eme sous-famille Myrmicinae Lep. (suite). Memoires de la Societe Entomologique de Belgique 20: 1-32. https://doi.org/10.5962/bhl.part.19251" title="Formicides néotropiques. Part IV. 3 eme sous-famille Myrmicinae Lep. (suite)." volume="20" year="1912">Forel, 1912: 18</bibRefCitation>
.
<typeStatus id="7FD0BDF1CBBA31FCCD51B3DAD4D572A2" type="syntype">Syntype</typeStatus>
series. (worker):
<collectingCountry id="AE4DE7CB4EA2C76149FF7830D3D050DE" name="Colombia">Colombia</collectingCountry>
, Sierra Nevada de Santa Marta, San Antonio. [
<abbrev id="ABBRID0EPPAG" xlink_title="Muséum dHistoire Naturelle, Geneva">
<collectionCode id="B100CFFAB046BDCE92147B52E239A11F" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" type="Museum">MHNG</collectionCode>
</abbrev>
]. (AntWeb image examined,
<ext-link id="A674AEA962DBB02A67013C9F45B2885F" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0909002" xlink_type="simple">
<specimenCode id="E057D6944DEF42B43111F26A7D09A1D3">CASENT 0909002</specimenCode>
</ext-link>
). syn. nov.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="F106419748F47027A3D5CCA638734C51">
<treatmentCitationGroup id="529C5CB7F811A2D7DDD9F7F3126173F9">
<treatmentCitation id="203CCB507198BD891FC8BDAEE97B12D3" author="Forel A" page="18" year="1912">
<taxonomicName id="FE34E7B09F95F880748A71499DB70103" authority="Forel, 1912: 18" authorityName="Forel" authorityPageNumber="18" authorityYear="1912" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="variety" species="asper" variety="sulfurea">
<emphasis id="BE7E68D86AFE8EE9F9F73B5A7B7E53E4" italics="true">Leptothorax asper var. sulfurea</emphasis>
<comment id="566E8AABB57BD7F24BA70A965355D2E4">
<bibRefCitation id="0676E01B59D0F632E37F459CBAB3432E" DOI="10.5962/bhl.part.19251" author="Forel A" journalOrPublisher="Mémoires de la Société Entomologique de Belgique" pagination="1-32" refId="B30" refString="Forel A (1912) Formicides neotropiques. Part IV. 3 eme sous-famille Myrmicinae Lep. (suite). Memoires de la Societe Entomologique de Belgique 20: 1-32. https://doi.org/10.5962/bhl.part.19251" title="Formicides néotropiques. Part IV. 3 eme sous-famille Myrmicinae Lep. (suite)." volume="20" year="1912">Forel, 1912: 18</bibRefCitation>
.
<typeStatus id="EAE6CB8736B09FD55A1E180ACC387C7E" type="syntype">Syntype</typeStatus>
series (worker):
<collectingCountry id="EA5A8DBC6C9C5662404653BBD60F1088" name="Brazil">Brazil</collectingCountry>
. [
<abbrev id="ABBRID0EMQAG" xlink_title="Muséum dHistoire Naturelle, Geneva">
<collectionCode id="FE533312CCA81B92E8C61EBE99905686" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" type="Museum">MHNG</collectionCode>
</abbrev>
]. (AntWeb image examined,
<ext-link id="0395ADE4213A129C49A501FDC770121B" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0909003" xlink_type="simple">
<specimenCode id="B2BF40D062E3B5EB8826242574251932">CASENT 0909003</specimenCode>
</ext-link>
). Synonymy by
<treatmentCitation id="C6B1C48CF2424A5B29C9E826224F4DE4" author="Kempf WW" page="414" year="1959">
<bibRefCitation id="1EDA6583AA3BD7D77CB22FF5A70A6852" author="Kempf WW" firstAuthor="Kempf" journalOrPublisher="Studia Entomologica" page="414" pagination="391-432" refId="B37" refString="Kempf WW (1959) A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae). Studia Entomologica 2: 391432." title="A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae)." volume="2" year="1959">Kempf (1959: 414)</bibRefCitation>
</treatmentCitation>
.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="2146A6CE0A13707E841A078025E70A35">
Combination in
<taxonomicName id="5596F85BA2F2ED501A8489470586CA53" authorityName="Wheeler" authorityYear="1910" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Nesomyrmex">
<emphasis id="1AE47B5038589EC0DAD9B917B658344E" italics="true">Leptothorax</emphasis>
(
<emphasis id="3AF95A75781EADCD1FD922D9CDDFDA22" italics="true">Nesomyrmex</emphasis>
)
</taxonomicName>
:
<bibRefCitation id="C302AA550B24923A57686C2B0FB37AB2" author="Kempf WW" firstAuthor="Kempf" journalOrPublisher="Studia Entomologica" page="414" pagination="391-432" refId="B37" refString="Kempf WW (1959) A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae). Studia Entomologica 2: 391432." title="A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae)." volume="2" year="1959">Kempf 1959: 414</bibRefCitation>
.
</paragraph>
<paragraph id="25E33D52892E6B20278011AC45D4A99B">
Combination in
<taxonomicName id="81AB5E4B9EF04C742D6DD91DCEDB0622" ID-CoL="CM5NR" authorityName="Wheeler" authorityYear="1910" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="109994097B54A8D2AE4EFEEEFEEBF61A" italics="true">Nesomyrmex</emphasis>
</taxonomicName>
:
<bibRefCitation id="41BFDF6528212EB27CEA112162E440A1" author="Bolton B" firstAuthor="Bolton" journalOrPublisher="Memoirs of the American Entomological Society" page="272" pagination="1-370" refId="B5" refString="Bolton B (2003) Synopsis and classification of Formicidae. Memoirs of the American Entomological Society 71: 1370." title="Synopsis and classification of Formicidae." volume="71" year="2003">Bolton 2003: 272</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGSAG" type="description">
<paragraph id="349004CECEA813DE22F59B5F7C9E144C">
<heading id="E015AE770C6488C829EC4042A0557056" reason="title">Worker measurements.</heading>
</paragraph>
<paragraph id="5F516A0A97AC54A4A91F2DB8FD7F10A9">
(
<emphasis id="B13C45BD75931851A1D5E4629E0BAF71" italics="true">n</emphasis>
= 8)
<abbrev id="ABBRID0EOSAG" xlink_title="Head length">HL</abbrev>
0.861.08,
<abbrev id="ABBRID0ESSAG" xlink_title="Head width">HW</abbrev>
0.780.99,
<abbrev id="ABBRID0EWSAG" xlink_title="Scape length">SL</abbrev>
0.680.84, ML 0.410.53,
<abbrev id="ABBRID0E1SAG" xlink_title="Eye length">EL</abbrev>
0.190.24,
<abbrev id="ABBRID0E5SAG" xlink_title="Pronotal width">PW</abbrev>
0.510.66,
<abbrev id="ABBRID0ECTAG" xlink_title="Petiole width">PTW</abbrev>
0.210.31,
<abbrev id="ABBRID0EGTAG" xlink_title="Postpetiole width">PPW</abbrev>
0.310.44,
<abbrev id="ABBRID0EKTAG" xlink_title="Webers length">WL</abbrev>
1.051.38,
<abbrev id="ABBRID0EOTAG" xlink_title="Pronotal height">PH</abbrev>
0.290.34,
<abbrev id="ABBRID0ESTAG" xlink_title="Petiole length">PTL</abbrev>
0.370.46,
<abbrev id="ABBRID0EWTAG" xlink_title="Petiole height">PTH</abbrev>
0.230.29,
<abbrev id="ABBRID0E1TAG" xlink_title="Postpetiole length">PPL</abbrev>
0.240.34,
<abbrev id="ABBRID0E5TAG" xlink_title="Postpetiole height">PPH</abbrev>
0.260.32,
<abbrev id="ABBRID0ECUAG" xlink_title="Gaster length">GL</abbrev>
0.881.01.
<abbrev id="ABBRID0EGUAG" xlink_title="Cephalic index">CI</abbrev>
9092,
<abbrev id="ABBRID0EKUAG" xlink_title="Scape index">SI</abbrev>
8487.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOUAG" type="Geographic range">
<paragraph id="59D0C3B7C4000D1A79931ACA88AFCD5E">
<heading id="137EE09D05FD382DF666B465EEA3512B" reason="title">Geographic range.</heading>
</paragraph>
<paragraph id="C722BE2038674EA42894BE9535FEE93A">
<collectingCountry id="54FEEA3A7AEB1E009FBC2CADCCE75BF4" name="Argentina">Argentina</collectingCountry>
,
<collectingCountry id="DB3CD849356441110D9A999C1F13F160" name="Bolivia">Bolivia</collectingCountry>
,
<collectingCountry id="F70BDD0E2CB0EF39060F2958975B9939" name="Colombia">Colombia</collectingCountry>
,
<collectingCountry id="89F0AB8D35F30F82AC8922A7034799E3" name="Ecuador">Ecuador</collectingCountry>
, French Guaina,
<collectingCountry id="7E25D41C6240ED0B670221EDD7E26BF4" name="Nicaragua">Nicaragua</collectingCountry>
,
<collectingCountry id="624B4C2C5342B326DA07890E2CD66E41" name="Panama">Panama</collectingCountry>
,
<collectingCountry id="5231E98EFEEC743758CEA02B975FBF7E" name="Paraguay">Paraguay</collectingCountry>
,
<collectingCountry id="80C25C6FCE6651799A21DEEC56167EB6" name="Trinidad and Tobago">Trinidad and Tobago</collectingCountry>
,
<collectingCountry id="EE5883FD8E4497378C371334FD150115" name="Venezuela">Venezuela</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETUAG" type="material">
<paragraph id="B5C8C8518D0B53F6C2B59173563E45AB">
<heading id="88D7EF487E785077456B435D174F5DAA" reason="title">Examined material.</heading>
</paragraph>
<paragraph id="D5379779BC7FE36B70ACD6A30D397C16">
<materialsCitation id="8C4C28AF11E57C25A8346EEAFB4C1576" collectingDate="2023-03-28" collectingDateMax="2023-03-30" collectingDateMin="2023-03-28" collectingMethod="Winkler" collectionCode="CBUMAG" collectorName="J. Camargo &amp; H. Sierra &amp; S. De la Hoz" country="Colombia" elevation="155" latitude="10.75198" location="Vda. Luriza" longLatPrecision="0" longitude="-75.0" municipality="Usicuri" specimenCode="CBUMAG:ENT:54657" specimenCount="1" specimenCount-worker="1" stateProvince="Atlantico">
<collectingCountry id="50E1816343ECF7EB4F26F35394A7DA6C" name="Colombia">Colombia</collectingCountry>
<specimenCount id="4A054CAD00B1DC3426AC517C0B17C6DE" count="1" type="worker">1 worker</specimenCount>
;
<collectingRegion id="FBA9B32F22D7BFC7F48BFB4F888CD3D9" country="Colombia" name="Atlantico">Atlántico</collectingRegion>
,
<collectingMunicipality id="6AF3E780DDF1EC73F3625F50C6DF21E3">Usicurí</collectingMunicipality>
,
<location id="EF689C7D2C91887549AADB5B4486BA0D" LSID="urn:lsid:plazi:treatment:3DBF20F901035E869B89A4EE102E7899:EF689C7D2C91887549AADB5B4486BA0D" country="Colombia" latitude="10.75198" longLatPrecision="0" longitude="-75.0" municipality="Usicuri" name="Vda. Luriza" stateProvince="Atlantico">Vda. Luriza</location>
, CIALU;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3UAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-75.030750,10.751980]}">
<geoCoordinate id="15818835EBEADC8BC1577B847ABFAD3F" degrees="10.75198" direction="north" orientation="latitude" precision="1" value="10.75198">10.75198°N</geoCoordinate>
,
<geoCoordinate id="AB32392CE754DCEB54DF8BDBF8C6165F" degrees="75,03075" direction="west" orientation="longitude" precision="1" value="75,03075">75,030 75°W</geoCoordinate>
</named-content>
;
<elevation id="C4D9C6012233FF56E9969EFFF6993FDC" metricMagnitude="2" metricUnit="m" metricValue="1.55" unit="m" value="155.0">
<quantity id="5655CC1EE9D9F94F94EE55748E92B585" metricMagnitude="2" metricUnit="m" metricValue="1.55" unit="m" value="155.0">155 m</quantity>
a. s. l.
</elevation>
;
<date id="58F38F2CA2A054A3277271ED11521C21" value="2023-03-28" valueMax="2023-03-30" valueMin="2023-03-28">
<collectingDate id="CA23DF1C5B6D68170A2A30DE4A43C8B7" value="2023-03-28" valueMax="2023-03-30" valueMin="2023-03-28">2830 Mar. 2023</collectingDate>
</date>
;
<collectorName id="FB3FA56CD2C69566E2AFA6D767FFCF6E">J. Camargo</collectorName>
,
<collectorName id="1B0120D0A59FE59EBB07315954B6B751">H. Sierra</collectorName>
,
<collectorName id="0553D53A826E17DDFC781AE81B8857A0">S. De la Hoz</collectorName>
legs.;
<collectingMethod id="A0B78558A2CD55AEB865981B40BDA37C">Winkler</collectingMethod>
;
<specimenCode id="D2777746C4718B1C93826DFF86BEF34E">
<named-content id="AB913038DFA1A77253F3356E910EDE52" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:54657
</specimenCode>
</materialsCitation>
<materialsCitation id="5D9643BDCA665424C457DC9CCC3B679E" collectedFrom="secondary forest" collectingDate="2015-07-05" collectionCode="ICN" collectorName="A. Sagoval &amp; C. Cantor" country="Colombia" county="Bolivar" elevation="102" latitude="10.371944" location="Finca el Huerto" longLatPrecision="1" longitude="-75.34967" municipality="Turbaco" specimenCode="ICN 106553" specimenCount="1" specimenCount-worker="1" stateProvince="Atlantico">
<specimenCount id="8CD362F013662DF1354047742AA2EBDD" count="1" type="worker">1 worker</specimenCount>
;
<collectingCounty id="3EBC53789456E8A8907DBD244A902752">Bolívar</collectingCounty>
,
<collectingMunicipality id="BE8E67A7317C52357BEBAFAFBDF33C2C">Turbaco</collectingMunicipality>
,
<location id="999A850CB81CFD242A8D711C1062A137" LSID="urn:lsid:plazi:treatment:3DBF20F901035E869B89A4EE102E7899:999A850CB81CFD242A8D711C1062A137" country="Colombia" county="Bolivar" latitude="10.371944" longLatPrecision="1" longitude="-75.34967" municipality="Turbaco" name="Finca el Huerto" stateProvince="Atlantico">Finca el Huerto</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIVAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-75.349667,10.371944]}">
<geoCoordinate id="11BA0320D44B3089FAC7D8364A0F827F" degrees="10.371944" direction="north" orientation="latitude" precision="1" value="10.371944">10.371944°N</geoCoordinate>
,
<geoCoordinate id="FB8E671261CB592E214351A3990922F8" degrees="75.349667" direction="west" orientation="longitude" precision="1" value="-75.34967">75.349667°W</geoCoordinate>
</named-content>
;
<elevation id="C90A1A45B7AC592C6FEF317DFE4ABD1C" metricMagnitude="2" metricUnit="m" metricValue="1.02" unit="m" value="102.0">
<quantity id="ACC7A104A6A85DB95F91C0A3A2CDEE85" metricMagnitude="2" metricUnit="m" metricValue="1.02" unit="m" value="102.0">102 m</quantity>
a. s. l.
</elevation>
;
<date id="94F1633187707053B04CDF909B1A6555" value="2015-07-05">
<collectingDate id="1028C47BFFBE1C7009B0F25BFAEB4538" value="2015-07-05">05 Jul. 2015</collectingDate>
</date>
;
<collectorName id="976E48AE63710EF5976C40E5FCF626E1">A. Sagoval</collectorName>
,
<collectorName id="A0C25B80BFA781126F2F24C6B4F8458D">C. Cantor</collectorName>
legs.;
<collectedFrom id="B196508F3AAB8655D097248087557888">secondary forest</collectedFrom>
;
<specimenCode id="ED77691C865240D8E36982CFD756C1CD" collectionCode="ICN" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" type="">
<abbrev id="ABBRID0ENVAG" xlink_title="Colección de Entomología del Instituto de Ciencias Naturales, Universidad Nacional de Colombia">ICN</abbrev>
106553
</specimenCode>
</materialsCitation>
. •
<materialsCitation id="F843D9E590E9BB2BF5A4D10A7D273A78" collectingMethod="pitfall" collectionCode="ICN" collectorName="F. Fernandez" country="Colombia" elevation="150" latitude="9.561111" location="La Jagua de Ibirico" longLatPrecision="1" longitude="-73.33639" specimenCode="ICN 019802" specimenCount="1" specimenCount-worker="1" stateProvince="Cesar">
<specimenCount id="94D183C0EE78A8CA8B3058BE719BD855" count="1" type="worker">1 worker</specimenCount>
;
<collectingRegion id="C79A0C821A5AE404DAD73CC85CC119A9" country="Colombia" name="Cesar">Cesar</collectingRegion>
,
<location id="F63BE233629EC5D18B8AEEA2F18095FF" LSID="urn:lsid:plazi:treatment:3DBF20F901035E869B89A4EE102E7899:F63BE233629EC5D18B8AEEA2F18095FF" country="Colombia" latitude="9.561111" longLatPrecision="1" longitude="-73.33639" name="La Jagua de Ibirico" stateProvince="Cesar">La Jagua de Ibirico</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EUVAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-73.336389,9.561111]}">
<geoCoordinate id="EE4B292F22BDBA6D3D292E0B2ED22DE1" degrees="9.561111" direction="north" orientation="latitude" precision="1" value="9.561111">9.561111°N</geoCoordinate>
,
<geoCoordinate id="D55EC49CADACF71EE22EF72804669169" degrees="73.336389" direction="west" orientation="longitude" precision="1" value="-73.33639">73.336389°W</geoCoordinate>
</named-content>
;
<elevation id="C7C9E101DBD6B541D6217EECAA618008" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">
<quantity id="D4D05FDB7C0399018405F3792D17DD38" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150 m</quantity>
a. s. l.
</elevation>
; 2007;
<collectorName id="2095E56BD4461E8A2F876264CD820320">F. Fernández</collectorName>
leg.;
<collectingMethod id="4E7F933EC758A342F974DC52C3C24BA5">pitfall</collectingMethod>
;
<specimenCode id="5087649F83C775DD87A3B727E34035E6" collectionCode="ICN" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" type="">
<abbrev id="ABBRID0EZVAG" xlink_title="Colección de Entomología del Instituto de Ciencias Naturales, Universidad Nacional de Colombia">ICN</abbrev>
019802
</specimenCode>
</materialsCitation>
. •
<materialsCitation id="6917920027A620AA4F1956C973FC343F" collectingDate="2017-11-25" collectingMethod="Winkler" collectionCode="IAvH" collectorName="L. Arcila" country="Morocco" county="Aipe" elevation="841" latitude="3.345356" location="Vda. San Isidro" longLatPrecision="1" longitude="-75.30578" municipality="Cuenca Rio Aipe" specimenCount="1" specimenCount-worker="1" stateProvince="Huila">
<specimenCount id="0ACB1A32C14EBA8CC49EF6965D6AC06A" count="1" type="worker">1 worker</specimenCount>
;
<collectingRegion id="B4A72FE16474BFB57CEFADE3FD1D28B6" country="Colombia" name="Huila">Huila</collectingRegion>
,
<collectingCounty id="9B8967472764026854D7355CA9ACEF99">Aipe</collectingCounty>
,
<collectingMunicipality id="83205009B95C862E86BBBF0A44C87E3D">Cuenca Río Aipe</collectingMunicipality>
,
<location id="34A8FD0A16AF7D19A68AFEDEEFFAAA24" LSID="urn:lsid:plazi:treatment:3DBF20F901035E869B89A4EE102E7899:34A8FD0A16AF7D19A68AFEDEEFFAAA24" country="Morocco" county="Aipe" latitude="3.345356" longLatPrecision="1" longitude="-75.30578" municipality="Cuenca Rio Aipe" name="Vda. San Isidro" stateProvince="Huila">Vda. San Isidro</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EAWAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-75.305778,3.345356]}">
<geoCoordinate id="57A376D96CABAE6F76769B5B46188C17" degrees="3.3453561" direction="north" orientation="latitude" precision="1" value="3.345356">3.3453561°N</geoCoordinate>
,
<geoCoordinate id="4198E890E9FAF67B21E3C170079D2051" degrees="75.3057781" direction="west" orientation="longitude" precision="1" value="-75.30578">75.3057781°W</geoCoordinate>
</named-content>
;
<elevation id="D74407A5E5372A9348047A4951A47860" metricMagnitude="2" metricUnit="m" metricValue="8.41" unit="m" value="841.0">
<quantity id="E93678E99437FD60C38701746DB32094" metricMagnitude="2" metricUnit="m" metricValue="8.41" unit="m" value="841.0">841 m</quantity>
a. s. l.
</elevation>
;
<date id="F152DD23F84A35BCA51681C53F9A3271" value="2017-11-25">
<collectingDate id="4281943B04870A2EF99C1081D8587BC3" value="2017-11-25">25 Nov. 2017</collectingDate>
</date>
;
<collectorName id="04BCCF4ECBACB78EF323336F76C494B6">L. Arcila</collectorName>
leg.;
<collectingMethod id="C9FA490086DC8454EAA9192FE506C3D1">Winkler</collectingMethod>
;
<abbrev id="ABBRID0EFWAG" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">
<collectionCode id="D28DA9CD17A62CAE0AC7D4A12F435120">IAvH</collectionCode>
</abbrev>
</materialsCitation>
. •
<materialsCitation id="ACD58FD00E8A02C84E659979A798A762" collectingDate="2021-08-17" collectingMethod="Pitfall" collectionCode="IAvH" collectorName="L. Arcila" country="Morocco" county="Valle del Cauca" elevation="1071" latitude="3.623869" location="Cuenca Rio Dagua" longLatPrecision="1" longitude="-75.6945" municipality="Dagua" specimenCount="1" specimenCount-worker="1">
<specimenCount id="24171CB3B2B0D7C64497DA93789A0BD7" count="1" type="worker">1 worker</specimenCount>
;
<collectingCounty id="7C083192DC77A78F98E4BBF40D42EDC6">Valle del Cauca</collectingCounty>
,
<collectingMunicipality id="367CFDDB39288011E2ECDFB0101E8924">Dagua</collectingMunicipality>
,
<location id="962386EF0E807941BC134B5589E967C8" LSID="urn:lsid:plazi:treatment:3DBF20F901035E869B89A4EE102E7899:962386EF0E807941BC134B5589E967C8" country="Morocco" county="Valle del Cauca" latitude="3.623869" longLatPrecision="1" longitude="-75.6945" municipality="Dagua" name="Cuenca Rio Dagua">Cuenca Río Dagua</location>
,
<location id="8261455850D7A0000EA5617E60B74F24" LSID="urn:lsid:plazi:treatment:3DBF20F901035E869B89A4EE102E7899:8261455850D7A0000EA5617E60B74F24" country="Morocco" county="Valle del Cauca" latitude="3.623869" longLatPrecision="1" longitude="-75.6945" municipality="Dagua" name="Vda. Limonar">Vda. Limonar</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMWAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-75.694507,3.623869]}">
<geoCoordinate id="C62A19AC9F5ED9EFAF9001F0DE903CB4" degrees="3.6238689" direction="north" orientation="latitude" precision="1" value="3.623869">3.6238689°N</geoCoordinate>
,
<geoCoordinate id="D07721419DF6C6993CCA3684FAE560A1" degrees="75.6945069" direction="west" orientation="longitude" precision="1" value="-75.6945">75.6945069°W</geoCoordinate>
</named-content>
;
<elevation id="C3D46F64C2AD1B298EA41C8A592EBAE8" metricMagnitude="3" metricUnit="m" metricValue="1.071" unit="m" value="1071.0">
<quantity id="DC86CD2E44058ADD77E9139D961D3476" metricMagnitude="3" metricUnit="m" metricValue="1.071" unit="m" value="1071.0">1071 m</quantity>
a. s. l.
</elevation>
;
<date id="4F60A5F35B87003D0B9B91B4594D12F3" value="2021-08-17">
<collectingDate id="6BFB40A677F32563F570CB22AE4F5887" value="2021-08-17">17 Aug. 2021</collectingDate>
</date>
;
<collectorName id="8618ED3F030561F7FB183A56404AD5FC">L. Arcila</collectorName>
leg.;
<collectingMethod id="73E717A90968517692359F5942323B80">Pitfall</collectingMethod>
;
<abbrev id="ABBRID0ERWAG" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">
<collectionCode id="59503AFF063E69B26734AF1DE26967D1">IAvH</collectionCode>
</abbrev>
</materialsCitation>
. •
<materialsCitation id="2FDFCC6D613545EFF4AE238E78DFC4CD" collectedFrom="meadow" collectingDate="2021-08-17" collectionCode="CBUMAG" collectorName="M. A. Bautista-Giraldo &amp; I. Armbrecht" country="Morocco" county="Cauca" elevation="909" latitude="3.1096" location="Las Chatas" longLatPrecision="7" longitude="-76.5265" municipality="Santander de Quilichao" specimenCode="CBUMAG:ENT:40121, CBUMAG:ENT:40122" specimenCount="2" specimenCount-worker="2">
<specimenCount id="9DAB24E6810E10D6E62DCFC79D9D1CD7" count="2" type="worker">2 workers</specimenCount>
;
<collectingCounty id="42283D107D3F90A00F07A9AAF3B659BF">Cauca</collectingCounty>
,
<collectingMunicipality id="3A8FFF1B3E54051236FB3DCED7C80850">Santander de Quilichao</collectingMunicipality>
,
<location id="2BB0AAE3852E6B0E2656988E5469C730" LSID="urn:lsid:plazi:treatment:3DBF20F901035E869B89A4EE102E7899:2BB0AAE3852E6B0E2656988E5469C730" country="Morocco" county="Cauca" latitude="3.1096" longLatPrecision="7" longitude="-76.5265" municipality="Santander de Quilichao" name="Las Chatas">Las Chatas</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYWAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-76.526500,3.109600]}">
<geoCoordinate id="DADE8CB4353C96D5E1840BFC69AD09A0" degrees="3.1096" direction="north" orientation="latitude" precision="5" value="3.1096">3.1096°N</geoCoordinate>
,
<geoCoordinate id="4BABD55498E60A6673B1687F19329425" degrees="76.5265" direction="west" orientation="longitude" precision="5" value="-76.5265">76.5265°W</geoCoordinate>
</named-content>
;
<elevation id="9684BCD07562DD89E9CD0B6E5120FB0C" metricMagnitude="2" metricUnit="m" metricValue="9.09" unit="m" value="909.0">
<quantity id="D73B25B3835128C9D0020E048C2ECE82" metricMagnitude="2" metricUnit="m" metricValue="9.09" unit="m" value="909.0">909 m</quantity>
a. s. l.
</elevation>
;
<date id="8F3AA3167B34D61D6E87FD1202EFEC2A" value="2021-08-17">
<collectingDate id="83DEFC06B23D88E9ED6FFE667A88AB2A" value="2021-08-17">17 Aug. 2021</collectingDate>
</date>
;
<collectorName id="AC7C4A1282DA92BB3BCED8BF5DB3E983">M. A. Bautista-Giraldo</collectorName>
;
<collectorName id="624DBCCE7AB56762F971BB83093258A4">I. Armbrecht</collectorName>
legs.;
<collectedFrom id="ADE92A59BC6839AA8E161964496C161A">meadow</collectedFrom>
;
<specimenCode id="720218387A688FC6D47AB6C5626BB47B">
<named-content id="FD7034409440B61E4FEF22D844C7F543" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:40121
</specimenCode>
,
<specimenCode id="3D15D11479937564A645DA0961BE142C">
<named-content id="511782AB9CF0483A3C8F30BEFB0BA2B8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:40122
</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFXAG" type="natural history">
<paragraph id="0784CBB4D697964D09B78DA2727C8A7B">
<heading id="4D4CF70DD71217551ADD29986982CFD5" reason="title">Natural history.</heading>
</paragraph>
<paragraph id="AFA6A095797C6B372784DFB311F925CC">
Some workers were collected foraging during the day on vegetation in
<abbrev id="ABBRID0ELXAG" xlink_title="tropical dry forest">TDF</abbrev>
fragments in the
<collectingRegion id="24E812E0902D0BCF683367A78F39DD97" country="Colombia" name="Valle del Cauca">Valle del Cauca</collectingRegion>
. Additionally,
<specimenCount id="970C43352D0DCF47341F0780AAD148A5" count="1" type="worker">one worker</specimenCount>
was collected foraging on the grassland soil using pitfall traps.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPXAG" type="comments">
<paragraph id="8D63A5A60E46F8B1D251C9B6D5C18CD5">
<heading id="1B8119E462B42442C26D7C393ED8D1EA" reason="title">Comments.</heading>
</paragraph>
<paragraph id="08525D2215AD8EF0AF53B0482F393675">
This species is recognized by the set of the following characters: 11 antennomeres, clypeus sculpture ranges from smooth and shiny to longitudinally costate or reticulate, inner area of dorsal surface of mandibles finely reticulate and opaque, propodeal lobe angulate, apex blunt.
<taxonomicName id="B16699B4E414DAB0D1CED1BBD408C4C6" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="78CE0B0D1E0E2EB4401F570D3550771E" italics="true">Nesomyrmex asper</emphasis>
</taxonomicName>
presents intraspecific morphological variation in some traits throughout its distribution in the Colombian
<abbrev id="ABBRID0EAYAG" xlink_title="tropical dry forest">TDF</abbrev>
. The lateral tubercles of the petiole and postpetiole are variable characteristics, both in quantity and size, between populations of the species (Fig.
<figureCitation id="3E6F5734626A72055CF7C4BB5640A822" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Variation of lateral petiole tubercles in Nesomyrmex asper. White arrows indicate the tubercles." figureDoi="10.3897/zookeys.1232.141693.figure18" httpUri="https://binary.pensoft.net/fig/1282988">18</figureCitation>
). For example, workers from populations in the
<collectingRegion id="C399993EFD56A9D15BA2390362D08E8D" country="Colombia" name="Valle del Cauca">Valle del Cauca</collectingRegion>
(western
<collectingCountry id="3D27379BBE31B1D3B6527BD52638A43B" name="Colombia">Colombia</collectingCountry>
) have no lateral tubercles on the petiole and postpetiole (Fig.
<figureCitation id="A8A35FE30009B2BF2E0E1597172C2CA9" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Variation of lateral petiole tubercles in Nesomyrmex asper. White arrows indicate the tubercles." figureDoi="10.3897/zookeys.1232.141693.figure18" httpUri="https://binary.pensoft.net/fig/1282988">18 A</figureCitation>
) while in more northern populations such as those from
<collectingRegion id="D116CCBA478A4783EAD9AB45C576F5DE" country="Colombia" name="La Guajira">La Guajira</collectingRegion>
and
<collectingRegion id="86BEBF2449DBDF25957778F3C84AA300" country="Colombia" name="Magdalena">Magdalena</collectingRegion>
there are between one and three tubercles distributed asymmetrically on the sides of the petiole (Fig.
<figureCitation id="1960085CD112BFBBD39D51EA9E74BAAB" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Variation of lateral petiole tubercles in Nesomyrmex asper. White arrows indicate the tubercles." figureDoi="10.3897/zookeys.1232.141693.figure18" httpUri="https://binary.pensoft.net/fig/1282988">18 D, E</figureCitation>
). Analysis of
<taxonomicName id="594E5963EB501E7B5F27CA1260BAE642" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="072D4413E2CB1A28AC10CA83F4132DD8" italics="true">N. asper</emphasis>
</taxonomicName>
specimens across the Colombian
<abbrev id="ABBRID0E2YAG" xlink_title="tropical dry forest">TDF</abbrev>
suggests that populations of this species are morphologically cohesive in terms of the diagnostic characteristics that define it, but traits such as lateral tubercles on the petiole / postpetiole show extensive phenotypic plasticity.
</paragraph>
<caption id="9BB22AD92E46E210BC05C80290450D76" ID-DOI="10.3897/zookeys.1232.141693.figure18" ID-arpha="F49B02C6-D3FF-58E1-8C9C-07AC028AF7FF" httpUri="https://binary.pensoft.net/fig/1282988" startId="F18">
<paragraph id="AB1C0F17A153B6FD9735C42AD1C003EF">
<label id="97A4E4740FBC74A6822FC09B03A44104">Figure 18.</label>
</paragraph>
<paragraph id="B2F6F5895DD75B3AC84C6D9D304A5D4D">
Variation of lateral petiole tubercles in
<taxonomicName id="A7BD69CF77E2E99DA8353BF9758F92AF" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="7491E3D1D96F4B0246694549D6511729" italics="true">Nesomyrmex asper</emphasis>
</taxonomicName>
. White arrows indicate the tubercles.
</paragraph>
</caption>
<paragraph id="CCF87F8D9D05F629772A9C64165EBA2C">
Our novel observations of variability in number and shape of lateral petiole and postpetiole tubercles in
<taxonomicName id="668FC2537051BF1528EEF0962F2E27C0" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="B9CC06F992B22EF12C5B34ADA00F85D1" italics="true">N. asper</emphasis>
</taxonomicName>
contrasts with the use of presence and absence of tubercles as a diagnostic trait for some nomenclatural changes in
<taxonomicName id="6A3AAF1F8D5638958137F81FB109C2D5" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="72CE6F0F941AAF143847EF103BDAAEAE" italics="true">N. asper</emphasis>
</taxonomicName>
(
<bibRefCitation id="B8BDD252A41F8453475C6321FAC94694" author="Kempf" firstAuthor="Kempf" journalOrPublisher="Anais da Academia Brasileira de Ciencias" pagination="91-102" refId="B36" refString="Kempf WW (1958) Sôbre algumas formigas neotrópicas do gênero Leptothorax Mayr (Hymenoptera: Formicidae). Anais da Academia Brasileira de Ciencias 30: 91102." title="Sôbre algumas formigas neotrópicas do gênero Leptothorax Mayr (Hymenoptera: Formicidae)." volume="30" year="1958">Kempf 1958</bibRefCitation>
,
<bibRefCitation id="2B33311087AB076685CAF9EE314B5B12" author="Kempf" firstAuthor="Kempf" journalOrPublisher="Studia Entomologica" pagination="391-432" refId="B37" refString="Kempf WW (1959) A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae). Studia Entomologica 2: 391432." title="A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae)." volume="2" year="1959">1959</bibRefCitation>
,
<bibRefCitation id="AB00CADE78F6F470B7B3905AD0B328E9" author="Kempf" firstAuthor="Kempf" journalOrPublisher="Studia Entomologica" pagination="341-380" refId="B38" refString="Kempf WW (1975) Miscellaneous studies on neotropical ants. VI. (Hymenoptera, Formicidae). Studia Entomologica 18: 341380." title="Miscellaneous studies on neotropical ants. VI. (Hymenoptera, Formicidae)." volume="18" year="1975">1975</bibRefCitation>
;
<bibRefCitation id="71365F3F9763C8191882950C386B4270" author="Longino" firstAuthor="Longino" journalOrPublisher="Myrmecological News" pagination="131-143" refId="B40" refString="Longino JT (2006) New species and nomenclatural changes for the Costa Rican ant fauna (Hymenoptera: Formicidae). Myrmecological News 8: 131143." title="New species and nomenclatural changes for the Costa Rican ant fauna (Hymenoptera: Formicidae)." volume="8" year="2006">Longino 2006</bibRefCitation>
).
<bibRefCitation id="46823039A95D0BBE1FFFC5DEDAE26D3A" author="Longino JT" firstAuthor="Longino" journalOrPublisher="Myrmecological News" pagination="131-143" refId="B40" refString="Longino JT (2006) New species and nomenclatural changes for the Costa Rican ant fauna (Hymenoptera: Formicidae). Myrmecological News 8: 131143." title="New species and nomenclatural changes for the Costa Rican ant fauna (Hymenoptera: Formicidae)." volume="8" year="2006">Longino (2006)</bibRefCitation>
proposed
<taxonomicName id="8DCD6856918F79AEA1E379FEC76B3485" authority="(Emery, 1896)" baseAuthorityName="Emery" baseAuthorityYear="1896" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tristani">
<emphasis id="47ADEB0C94A5BD748A8766FEC00BCCA4" italics="true">Nesomyrmex tristani</emphasis>
(
<bibRefCitation id="BBAE89DE3722C58D92080FB263D2FA22" author="Emery C" journalOrPublisher="Bullettino della Società Entomologica Italiana" pagination="33-107" refId="B23" refString="Emery C (1896) Studi sulle formiche della fauna neotropica. XVII-XXV. Bullettino della Societa Entomologica Italiana 28: 33-107." title="Studi sulle formiche della fauna neotropica. XVII-XXV." volume="28" year="1896">Emery, 1896</bibRefCitation>
)
</taxonomicName>
as a junior synonym of
<taxonomicName id="8AC315C429B1E09F6347AEAA63E0F343" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="1B6F81B7BBADDB17E16B5A8F3BEF7F4A" italics="true">Nesomyrmex asper</emphasis>
</taxonomicName>
considering the similarity of the lateral processes (i. e., lateral tubercles) of the petiole between the workers of both taxa. The same author used the differentiation of the lateral processes of the postpetiole (described there as “ acuminate teeth ”: p. 136) observed between the workers of
<taxonomicName id="D9001925DD4F080C5C5D34C4E315EB96" authority="(Forel, 1912)" baseAuthorityName="Forel" baseAuthorityYear="1912" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="variety" species="asper" variety="antoniensis">
<emphasis id="B8E7DF3E70EC43F9305E215AB3B61887" italics="true">Nesomyrmex asper var. antoniensis</emphasis>
(
<bibRefCitation id="78EFEC749F30E7A2818DD1EC09DEEFA0" DOI="10.5962/bhl.part.19251" author="Forel A" journalOrPublisher="Mémoires de la Société Entomologique de Belgique" pagination="1-32" refId="B30" refString="Forel A (1912) Formicides neotropiques. Part IV. 3 eme sous-famille Myrmicinae Lep. (suite). Memoires de la Societe Entomologique de Belgique 20: 1-32. https://doi.org/10.5962/bhl.part.19251" title="Formicides néotropiques. Part IV. 3 eme sous-famille Myrmicinae Lep. (suite)." volume="20" year="1912">Forel, 1912</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="EE9E2E6D0B88B037BBB4A3670CB09577" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="E3C8660A8E41A19F8AC03E11CBFABCC5" italics="true">N. asper</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DE512F0E21584431DA3839AF1E93E6EB" sensu="stricto">s. str</taxonomicNameLabel>
, to recognize
<taxonomicName id="726EA55BDC36E86B7CC366A683FF8BA0" authority="(Forel, 1912)" baseAuthorityName="Forel" baseAuthorityYear="1912" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="antoniensis">
<emphasis id="22B6316ECA04B0710E54AD72EBFD19A5" italics="true">N. antoniensis</emphasis>
(
<bibRefCitation id="028EF39363DB3CE4B896953BA9A3ABA5" DOI="10.5962/bhl.part.19251" author="Forel A" journalOrPublisher="Mémoires de la Société Entomologique de Belgique" pagination="1-32" refId="B30" refString="Forel A (1912) Formicides neotropiques. Part IV. 3 eme sous-famille Myrmicinae Lep. (suite). Memoires de la Societe Entomologique de Belgique 20: 1-32. https://doi.org/10.5962/bhl.part.19251" title="Formicides néotropiques. Part IV. 3 eme sous-famille Myrmicinae Lep. (suite)." volume="20" year="1912">Forel, 1912</bibRefCitation>
)
</taxonomicName>
as a valid species. Considering the intraspecific morphological variation in the lateral tubercles of
<taxonomicName id="E06F7A2EBB6C00E1A703226C27E9EA02" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="43D63AE13E8D29965789D3C29929D5BC" italics="true">N. asper</emphasis>
</taxonomicName>
workers, we compared this trait with the type worker of
<taxonomicName id="F86F74876721643236ADF9B472B09F32" baseAuthorityName="Forel" baseAuthorityYear="1912" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="antoniensis">
<emphasis id="5584A130C8B553413325F4C57F42250E" italics="true">N. antoniensis</emphasis>
</taxonomicName>
(
<ext-link id="04998A17DF85642916C03DBE9CB86734" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0909002" xlink_type="simple">
<specimenCode id="B392676653069864DC1EBEAE821FD64E">
<collectionCode id="96580DDDFD21FC00A2E328AED4489765">CASENT</collectionCode>
0909002
</specimenCode>
</ext-link>
), finding that the latter falls within the high intraspecific variability of
<taxonomicName id="93E3979547DB46820E215E4D9D1FA9DC" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="602036B3EFC8D36E9A7A9713DB08D834" italics="true">N. asper</emphasis>
</taxonomicName>
. Likewise, the type specimens of both
<taxonomicName id="ED07E71C49CFE1204F49658F39B12D32" baseAuthorityName="Forel" baseAuthorityYear="1912" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="antoniensis">
<emphasis id="09A15F6F26BE8A4905235A3069B3F009" italics="true">N. antoniensis</emphasis>
</taxonomicName>
(
<ext-link id="A74E6CA2767EDA0F2C42B6EDBE86D0BB" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0909002" xlink_type="simple">
<specimenCode id="5D8F241CD22794920C9B85291F561442">
<collectionCode id="D180F1E4636732497A478B59F7DFEDB6">CASENT</collectionCode>
0909002
</specimenCode>
</ext-link>
) and
<taxonomicName id="BCE42077B89AB36FF8C26AE1F3660EE3" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="4AB973681749E2326272BB191DA8DCE0" italics="true">N. asper</emphasis>
</taxonomicName>
(
<ext-link id="2D5E5684126A60043511A4E1D41F6914" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0901801" xlink_type="simple">
<specimenCode id="5542110E351EC55E7418D9C9D1862F46">
<collectionCode id="697F486841B7E37F3709A48BD3BF28B6">CASENT</collectionCode>
0901801
</specimenCode>
</ext-link>
), and the workers of
<taxonomicName id="5208AD09B822D61D74FEF66FB016C0E7" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="0E1D669B35D65D6B5152334DC2340D26" italics="true">N. asper</emphasis>
</taxonomicName>
present in
<collectingCountry id="8C1326EECF9BF7C8EB9A6DCA60F660C1" name="Colombia">Colombia</collectingCountry>
and the Sierra Nevada de Santa Marta (type locality of
<taxonomicName id="21AF56803CDCCDF15DFF794D945F2B46" baseAuthorityName="Forel" baseAuthorityYear="1912" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="antoniensis">
<emphasis id="09BF378E0713F8C08C66DE46BD1915D1" italics="true">N. antoniensis</emphasis>
</taxonomicName>
) were measured, finding that the type of
<taxonomicName id="805EEC5A5C31D0EFFAF841E9B86B06C8" baseAuthorityName="Forel" baseAuthorityYear="1912" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="antoniensis">
<emphasis id="213BE90F140CF8BDB68297D5D633FB1D" italics="true">N. antoniensis</emphasis>
</taxonomicName>
(
<abbrev id="ABBRID0EHABG" xlink_title="Head length">HL</abbrev>
0.89,
<abbrev id="ABBRID0ELABG" xlink_title="Head width">HW</abbrev>
0.83,
<abbrev id="ABBRID0EPABG" xlink_title="Scape length">SL</abbrev>
0.72, ML 0.43) matches the morphometric range of
<taxonomicName id="5282B41A20CCD5E3DFC1636FD1CD3667" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="15E67414B50166E74C6336157D2CA5DA" italics="true">N. asper</emphasis>
</taxonomicName>
(
<abbrev id="ABBRID0E5ABG" xlink_title="Head length">HL</abbrev>
0.861.08,
<abbrev id="ABBRID0ECBBG" xlink_title="Head width">HW</abbrev>
0.780.99,
<abbrev id="ABBRID0EGBBG" xlink_title="Scape length">SL</abbrev>
0.680.84, ML 0.410.53). Based on the evidence, we found that the morphological variation between
<taxonomicName id="A0C3150A3F247E30D383A2047FCADEAE" baseAuthorityName="Forel" baseAuthorityYear="1912" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="antoniensis">
<emphasis id="57C58730F30D4168288768B67505907D" italics="true">N. antoniensis</emphasis>
</taxonomicName>
and
<taxonomicName id="F77369F8894E9A6E1219B814E53E16CE" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="EF0BAFBCAC4D99128D49F500A0E7991C" italics="true">N. asper</emphasis>
</taxonomicName>
is not distinct, with the form referred to as
<taxonomicName id="7A51EE8CF8409638BD071A7668E1AAB2" baseAuthorityName="Forel" baseAuthorityYear="1912" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="antoniensis">
<emphasis id="EC4BCC956264032000039FA0FDCCCA82" italics="true">N. antoniensis</emphasis>
</taxonomicName>
falling within the variability observed in
<taxonomicName id="046EB1ED7DCD214BD35C5F0D9BB392E0" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="F8FF99B7D88A6C7D08896FD6D0A52EC0" italics="true">N. asper</emphasis>
</taxonomicName>
. Therefore, we propose
<taxonomicName id="E78408C92130AF79A5B92A2956257FEA" baseAuthorityName="Forel" baseAuthorityYear="1912" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="antoniensis">
<emphasis id="D70F57C8FDC7C84BF1D6AC2E75DCCD62" italics="true">N. antoniensis</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="3993A43176F7A9F4029150E879D5205A" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="050A6C8AD686C1E9DC0FF450DB2C3F3F" italics="true">Nesomyrmex asper</emphasis>
</taxonomicName>
. The highly variable number and shape of tubercles on lateral petiole and postpetiole in
<taxonomicName id="B4A49DAECEE1B52E2799A9FFB3A738BE" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="2B2DBB62E762FF2895B89F232529D0E9" italics="true">N. asper</emphasis>
</taxonomicName>
should not be used to separate taxonomic units in these species groups. Interestingly, the lateral processes of the petiole and postpetiolar node described as “ acuminate teeth ” by
<bibRefCitation id="C22905057FC6EF940A0B2C12ED81966D" author="Longino JT" firstAuthor="Longino" journalOrPublisher="Myrmecological News" page="136" pagination="131-143" refId="B40" refString="Longino JT (2006) New species and nomenclatural changes for the Costa Rican ant fauna (Hymenoptera: Formicidae). Myrmecological News 8: 131143." title="New species and nomenclatural changes for the Costa Rican ant fauna (Hymenoptera: Formicidae)." volume="8" year="2006">Longino (2006: 136)</bibRefCitation>
are setigerous tubercles (Fig.
<figureCitation id="E0F1B26F15D57B7C2335F8A907367ED3" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Lateral petiole and postpetiole tubercles (Setigerous tubercles). Abbreviations: lpptt (lateral postpetiolar tubercles), lptt (lateral petiolar tubercles), pt (petiole), ppt (postpetiole), set (seta)." figureDoi="10.3897/zookeys.1232.141693.figure19" httpUri="https://binary.pensoft.net/fig/1282989">19</figureCitation>
; also see
<bibRefCitation id="C87B6EE63FC1FAC4ECE51DCCE4F92002" author="Kempf" firstAuthor="Kempf" journalOrPublisher="Studia Entomologica" pagination="391-432" refId="B37" refString="Kempf WW (1959) A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae). Studia Entomologica 2: 391432." title="A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae)." volume="2" year="1959">Kempf 1959</bibRefCitation>
) and their variation in shape is due to projection and orientation relative to the body.
</paragraph>
<caption id="AE41BB8F77A6BFB971B5EC70745DEF11" ID-DOI="10.3897/zookeys.1232.141693.figure19" ID-arpha="89A725E2-02F9-5253-8549-668F531ED24A" httpUri="https://binary.pensoft.net/fig/1282989" startId="F19">
<paragraph id="99BBCBE93EA94B88D10706D6F585BC88">
<label id="36F9F0C60F4205A26B3FFBC68465F592">Figure 19.</label>
</paragraph>
<paragraph id="4BA4C27F07AA460F3C2014BC81AA5418">Lateral petiole and postpetiole tubercles (Setigerous tubercles). Abbreviations: lpptt (lateral postpetiolar tubercles), lptt (lateral petiolar tubercles), pt (petiole), ppt (postpetiole), set (seta).</paragraph>
</caption>
<caption id="A02961B5D6EB0A89DEE38D1A607FD457" ID-DOI="10.3897/zookeys.1232.141693.figure20" ID-arpha="AB8A29E1-3088-5F70-B7C8-5417397A7F2B" httpUri="https://binary.pensoft.net/fig/1282990" startId="F20">
<paragraph id="9F6DCCB190DC265BC6F934A2ECC21F3B">
<label id="2C3D518FD36D403D9C36B19C43C2DFE4">Figure 20.</label>
</paragraph>
<paragraph id="0473D32C6DF5A61126C03A0DFA897B1E">
<taxonomicName id="1B057822C4594EC1423110F5EB1C70E8" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="0A77C1C9783F0BB5AA34ACC93C57CB14" italics="true">Nesomyrmex asper</emphasis>
</taxonomicName>
worker (
<abbrev id="ABBRID0ECFBG" xlink_title="Colección de Entomología del Instituto de Ciencias Naturales, Universidad Nacional de Colombia">ICN</abbrev>
106553)
<emphasis id="BC6F94694211D8B88D6A59E94F2055B1" bold="true">A</emphasis>
full-face view
<emphasis id="7D1C308298FA1010BACFFB4FEE234659" bold="true">B</emphasis>
lateral view
<emphasis id="C23F583ED101FBD36D0B8BB3BB92E511" bold="true">C</emphasis>
dorsal view. Scale bars: 0.5 mm (
<emphasis id="7D2C64B40DD20C14B7694C210B789FFB" bold="true">A</emphasis>
); 1.0 mm (
<emphasis id="9D2FBE5CE99D95CAE150C40E613CF9BE" bold="true">B</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

View file

@ -1,48 +1,48 @@
<document id="13C9045663E3E03870B8A073764598BE" ID-DOI="10.3897/zookeys.1232.141693" ID-ZooBank="0AA099A2-EA57-4E27-937A-0976B9F8A4B5" ID-publisher-id="141693" URI-arpha="A9904ED3-9BDA-52DF-B936-1C42EC2713C5" URI-zoobank="http://zoobank.org/0AA099A2-EA57-4E27-937A-0976B9F8A4B5" article-type="research-article" checkinTime="1742225059022" checkinUser="pensoft" docAuthor="Arredondo, Brandon S. &amp; Guerrero, Roberto J." docDate="2025" docId="50DBCF393F2D54C583DC84C22407947F" docLanguage="en" docName="ZooKeys 1232: 131-172" docOrigin="ZooKeys 1232" docSource="https://zookeys.pensoft.net/article/141693/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nesomyrmex vargasi Longino 2006" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="172" masterDocId="A9904ED39BDA52DFB9361C42EC2713C5" masterDocTitle="The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data" masterLastPageNumber="172" masterPageNumber="131" pageNumber="131" updateTime="1742225485493" updateUser="ExternalLinkService">
<mods:mods id="555958E9F94080683384FBCAC679C8B2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D9EB319A602238FA8CBB5B29BF879960">
<mods:title id="D47029D577D896B8EC5B55EC506AB682">The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data</mods:title>
<document id="BAA9EADB5C2E6BA10DA447F9F0D4C7FD" ID-DOI="10.3897/zookeys.1232.141693" ID-ZooBank="0AA099A2-EA57-4E27-937A-0976B9F8A4B5" ID-publisher-id="141693" URI-arpha="A9904ED3-9BDA-52DF-B936-1C42EC2713C5" URI-zoobank="http://zoobank.org/0AA099A2-EA57-4E27-937A-0976B9F8A4B5" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1742225059022" checkinUser="pensoft" docAuthor="Arredondo, Brandon S. &amp; Guerrero, Roberto J." docDate="2025" docId="50DBCF393F2D54C583DC84C22407947F" docLanguage="en" docName="ZooKeys 1232: 131-172" docOrigin="ZooKeys 1232" docSource="https://zookeys.pensoft.net/article/141693/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nesomyrmex vargasi Longino 2006" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="172" masterDocId="A9904ED39BDA52DFB9361C42EC2713C5" masterDocTitle="The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data" masterLastPageNumber="172" masterPageNumber="131" pageNumber="131" updateTime="1742234045672" updateUser="admin">
<mods:mods id="7A5D1D9A0912425A1B770DCE3066BB82" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9C163D068E231277D1AA94180C782911">
<mods:title id="F854F59EA94656F37948632576ED7C19">The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data</mods:title>
</mods:titleInfo>
<mods:name id="88E2B10A1E1A4FCF8661BC067F6AE2B5" type="personal">
<mods:role id="E3CF40CBD367BBB4DB323936C594FF48">
<mods:roleTerm id="8CD16C56A289F4A05456AC403691606B">Author</mods:roleTerm>
<mods:name id="936FB149D7511EEF962EFEF9273F2A30" type="personal">
<mods:role id="D3B1B2498D4C04456ED66E556A9DB154">
<mods:roleTerm id="9009D87516DF5292EF3F4936007BBD11">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BF62EF5EAADEB33F2F8FB21CF3F8B56F">Arredondo, Brandon S.</mods:namePart>
<mods:nameIdentifier id="5CF717FE4638B55B4DC93F2B58A35107" type="ORCID">0000-0003-2777-5909</mods:nameIdentifier>
<mods:affiliation id="7A55A632447708A0FE8982E49DC169BE">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
<mods:namePart id="871AC5A7F7C1B087694911F1A507CF64">Arredondo, Brandon S.</mods:namePart>
<mods:nameIdentifier id="3DB3F4B8CF424598D8391E43E043B6A5" type="ORCID">0000-0003-2777-5909</mods:nameIdentifier>
<mods:affiliation id="5AF8BC6FC652C6539850DFFB0DBD4DDB">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:name id="216D813D202BE2C08BC1BC937898FF12" type="personal">
<mods:role id="205371D860ECC8911D351A0D55386BE6">
<mods:roleTerm id="851566159923A5F285E2337A5F985342">Author</mods:roleTerm>
<mods:name id="57CF2644819709FEF696EF295615812B" type="personal">
<mods:role id="BE4BF6DA6A6BCC3E9850D1E580DD5464">
<mods:roleTerm id="452C4492E2E105E57CBC201B1A7FB5C3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D570A24A8E842091B103399E81CF808C">Guerrero, Roberto J.</mods:namePart>
<mods:nameIdentifier id="28C6A87D2572DFD2AFAA7637DB7188AD" type="ORCID">0000-0003-3244-2754</mods:nameIdentifier>
<mods:affiliation id="09C78E0F98F7BC82DB15505CFB3C75DB">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
<mods:namePart id="CABD7E98A230D836470B4BFB67293B78">Guerrero, Roberto J.</mods:namePart>
<mods:nameIdentifier id="918DD54D080B5E2F91B909ED00F1868A" type="ORCID">0000-0003-3244-2754</mods:nameIdentifier>
<mods:affiliation id="28E38AEEAED7D7A947B20D30CCBBEA1A">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6CA2EDB6622BCB141B20A0FC519B8BF5">text</mods:typeOfResource>
<mods:relatedItem id="15CFA1FDF31EFB81238CC5DEE7AE3F9C" type="host">
<mods:titleInfo id="84A5FAC6129E9A2165F88BC163D9CE8A">
<mods:title id="D4662EA9377F550889C0BA88CA1183CD">ZooKeys</mods:title>
<mods:typeOfResource id="19E2D1DFE5C78A6762F1907A268B2233">text</mods:typeOfResource>
<mods:relatedItem id="C9B483F705466564FB74EC50C3455996" type="host">
<mods:titleInfo id="003D2CA8C13EEAE844C84EE0CB137A32">
<mods:title id="2A309603B6182646F37C83931CECC846">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="0D695F10BB2B89BA7628C88B336296DD">
<mods:date id="0F8AA4711CA987525E76FC6EED68C0BE">2025</mods:date>
<mods:detail id="96193E15D667215DCAF60F260FAE16A2" type="pubDate">
<mods:number id="D4CE2F307A6468074D0032129A3C75B7">2025-03-17</mods:number>
<mods:part id="BC7446CC6E977A4A873A9AAC80F31455">
<mods:date id="1643262809E2E5A921642B9ACCB0645A">2025</mods:date>
<mods:detail id="DE77C7B27A4FF311E366DEE3A150D0FA" type="pubDate">
<mods:number id="1FD4B8E40EDDEA964B1EE6C6797277A0">2025-03-17</mods:number>
</mods:detail>
<mods:detail id="D4A0495AE19B9A6E677637339DC3D018" type="volume">
<mods:number id="0677E2923CB97022385D7C0DC31CF717">1232</mods:number>
<mods:detail id="576287249A6302A4A7110FF23134CEB2" type="volume">
<mods:number id="3F0E1BF514D93EBB874D1DFF107F5D56">1232</mods:number>
</mods:detail>
<mods:extent id="79053B6DBD8FA72A9B7E21A4C5291348" unit="page">
<mods:start id="BE5BE81061C55C27D13F0E9E1535D704">131</mods:start>
<mods:end id="D4FA651520B01257300C3297EF689C59">172</mods:end>
<mods:extent id="5E387B0C04546285031E397FA4C9D479" unit="page">
<mods:start id="50D7E449548AE027C7154FD564C2CB00">131</mods:start>
<mods:end id="455BD9DC84FCF0CB21F167242DB1EB6A">172</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="576A5F8C5F6C79970A13841199F90B94">journal article</mods:classification>
<mods:identifier id="CCA91D9E1784F0B8BC233C9A671A9BDA" type="DOI">10.3897/zookeys.1232.141693</mods:identifier>
<mods:identifier id="32F0E3978DD57AA6CDF1E91956DF3B81" type="ZooBank">0AA099A2-EA57-4E27-937A-0976B9F8A4B5</mods:identifier>
<mods:classification id="688191153F2416C8B262130C2776353C">journal article</mods:classification>
<mods:identifier id="D5B1E29B773520E7EAD26906816510F9" type="DOI">10.3897/zookeys.1232.141693</mods:identifier>
<mods:identifier id="29AAE517335266AD5B980D88C77D599C" type="ZooBank">0AA099A2-EA57-4E27-937A-0976B9F8A4B5</mods:identifier>
</mods:mods>
<treatment id="50DBCF393F2D54C583DC84C22407947F" ID-DOI="http://doi.org/10.5281/zenodo.15040789" ID-Zenodo-Dep="15040789" ID-arpha="50DBCF39-3F2D-54C5-83DC-84C22407947F" LSID="urn:lsid:plazi:treatment:50DBCF393F2D54C583DC84C22407947F" httpUri="http://treatment.plazi.org/id/50DBCF393F2D54C583DC84C22407947F">
<treatment id="50DBCF393F2D54C583DC84C22407947F" ID-DOI="http://doi.org/10.5281/zenodo.15040789" ID-Zenodo-Dep="15040789" ID-arpha="50DBCF39-3F2D-54C5-83DC-84C22407947F" LSID="urn:lsid:plazi:treatment:50DBCF393F2D54C583DC84C22407947F" httpUri="http://treatment.plazi.org/id/50DBCF393F2D54C583DC84C22407947F" scope_family="Formicidae" scope_subFamily="Myrmicinae">
<subSubSection id="26AE868FB0E9DF5D3446F4E94AAA7CD8" type="nomenclature">
<paragraph id="FACD52B8398CE5A632DC882F268949E5">
<taxonomicName id="63804FCDBF2BAA2981FE156734865A5D" ID-CoL="CM82F" authority="Longino, 2006" authorityName="Longino" authorityYear="2006" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="vargasi">
@ -64,7 +64,6 @@
<treatmentCitation id="2487A4B408D62EC729C8605FAF55CE94" author="Longino JT" page="136" year="2006">
<taxonomicName id="430AEBB65BCB41485E0A73F5A96082FA" ID-CoL="CM82F" authority="Longino, 2006: 136" authorityName="Longino" authorityPageNumber="136" authorityYear="2006" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="vargasi">
<emphasis id="4FAC4A4E26B8D986071313B4AB966188" italics="true">Nesomyrmex vargasi</emphasis>
<materialsCitation id="18913B53FE86E6E9255BB5F83760E997" collectionCode="INBC" country="Costa Rica" location="Heredia Prov." specimenCount="1" stateProvince="Heredia" typeStatus="holotype">
<comment id="08BC00C739F3E79DFD36D8A8BF0263B8">
<bibRefCitation id="0EA43550ACDBE5706A3699A4EF36BAC0" author="Longino JT" journalOrPublisher="Myrmecological News" pagination="131-143" refId="B40" refString="Longino JT (2006) New species and nomenclatural changes for the Costa Rican ant fauna (Hymenoptera: Formicidae). Myrmecological News 8: 131-143." title="New species and nomenclatural changes for the Costa Rican ant fauna (Hymenoptera: Formicidae)." volume="8" year="2006">Longino, 2006: 136</bibRefCitation>
.
@ -77,7 +76,6 @@ worker:
<collectionCode id="E91A22144DE817A4237CB65B1F4A23DD" country="Costa Rica" name="Instituto Nacional de Biodiversidad (INBio)">INBC</collectionCode>
]. (AntWeb image examined, JTLC 000008517, JTLC 000008518, LACMENT 144699).
</comment>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
@ -107,9 +105,7 @@ worker:
0.250.27,
<abbrev id="ABBRID0ED2BI" xlink_title="Webers length">WL</abbrev>
0.880.96,
<abbrev id="ABBRID0EH2BI" xlink_title="Pronotal height">
<collectingCountry id="F8C16F901E267D7A4DE7DD6395BD9A7F" name="Philippines">PH</collectingCountry>
</abbrev>
<abbrev id="ABBRID0EH2BI" xlink_title="Pronotal height">PH</abbrev>
0.240.30,
<abbrev id="ABBRID0EL2BI" xlink_title="Petiole length">PTL</abbrev>
0.330.38,
@ -135,8 +131,11 @@ worker:
<emphasis id="70210D1B8445DC6C48A64EAA2BF4D510" italics="true">Nesomyrmex vargasi</emphasis>
</taxonomicName>
worker (
<specimenCode id="EFF31BDCEE9F2E1E096732A61E907259">
<named-content id="C05164F917353C754C88BD37EB872578" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
: ENT: 41665)
:ENT:41665
</specimenCode>
)
<emphasis id="91848298F7199F3349C65BE1F6660316" bold="true">A</emphasis>
full-face view
<emphasis id="ACBD45C6FFB384D3A7C2171243F74F83" bold="true">B</emphasis>
@ -162,40 +161,42 @@ dorsal view. Scale bars: 0.5 mm.
<heading id="F9E33144918CD13EBE72F4A668C37207" reason="title">Examined material.</heading>
</paragraph>
<paragraph id="A4A9647A550A4500471D092290FC8DF7">
<materialsCitation id="20B541A0DC16E6BD401D24F618A5113E" collectionCode="IAvH" country="Colombia" latitude="9.744721" location="Zambrano" longLatPrecision="1" longitude="-74.82421" municipality="Bolivar" specimenCount="1" specimenCount-worker="1">
<materialsCitation id="20B541A0DC16E6BD401D24F618A5113E" collectingMethod="Pitfall" collectionCode="IAvH" country="Colombia" latitude="9.744721" location="Zambrano" longLatPrecision="1" longitude="-74.82421" specimenCode="IAvH-E-251258" specimenCount="1" specimenCount-worker="1" stateProvince="Bolivar">
<collectingCountry id="375ED02D78228AFDC60208418B438DD4" name="Colombia">Colombia</collectingCountry>
<specimenCount id="6822A6D193AEC03E059D30023BF8EEE7" count="1" type="worker">1 worker</specimenCount>
;
<collectingMunicipality id="A51867A2252922C2B8EDF0CAA3F5D8A5">Bolivar</collectingMunicipality>
<collectingRegion id="3915D2BA1FF4C5C19D9C5AF173017A1A">Bolivar</collectingRegion>
,
<location id="430005DDEAB1045B43BD76A41001665C" LSID="urn:lsid:plazi:treatment:50DBCF393F2D54C583DC84C22407947F:430005DDEAB1045B43BD76A41001665C" country="Colombia" latitude="9.744721" longLatPrecision="1" longitude="-74.82421" municipality="Bolivar" name="Zambrano">Zambrano</location>
<location id="430005DDEAB1045B43BD76A41001665C" LSID="urn:lsid:plazi:treatment:50DBCF393F2D54C583DC84C22407947F:430005DDEAB1045B43BD76A41001665C" country="Colombia" latitude="9.744721" longLatPrecision="1" longitude="-74.82421" municipality="Bolivar" name="Zambrano" stateProvince="Bolivar">Zambrano</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EX4BI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-74.824210,9.744721]}">
<geoCoordinate id="C5D8F2FD512E00DFB54AB8BCF8EA7C9F" degrees="9.744721" direction="north" orientation="latitude" precision="1" value="9.744721">9.744721 ° N</geoCoordinate>
<geoCoordinate id="C5D8F2FD512E00DFB54AB8BCF8EA7C9F" degrees="9.744721" direction="north" orientation="latitude" precision="1" value="9.744721">9.744721°N</geoCoordinate>
,
<geoCoordinate id="6825716AE75371EC9EDE22CC06E2DBC6" degrees="74.82421" direction="west" orientation="longitude" precision="1" value="-74.82421">74.82421 ° W</geoCoordinate>
<geoCoordinate id="6825716AE75371EC9EDE22CC06E2DBC6" degrees="74.82421" direction="west" orientation="longitude" precision="1" value="-74.82421">74.82421°W</geoCoordinate>
</named-content>
; 19931994;
<collectingMethod id="03CF671860596E0686BC8C58442A65CE">Pitfall</collectingMethod>
;
<abbrev id="ABBRID0E34BI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">
<collectionCode id="ED5041837E47E9D8E27759262BB12C75">IAvH</collectionCode>
</abbrev>
- E- 251258
<specimenCode id="03948344064D953F3407D8B32F2B7C5D">
<abbrev id="ABBRID0E34BI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">IAvH</abbrev>
-E-251258
</specimenCode>
</materialsCitation>
<materialsCitation id="A53EBBDE01553CA9FCB9E9544B22221D" collectingDate="2016-03-31" collectionCode="CBUMAG" collectorName="R. Achury" country="Colombia" elevation="881" latitude="10.241965" location="Valledupar" longLatPrecision="1" longitude="-73.761215" specimenCount="1" specimenCount-worker="1" stateProvince="Cesar">
<materialsCitation id="A53EBBDE01553CA9FCB9E9544B22221D" collectingDate="2016-03-31" collectingMethod="Pitfall" collectionCode="CBUMAG" collectorName="R. Achury" country="Colombia" elevation="881" latitude="10.241965" location="Vda. Tierras nuevas" longLatPrecision="1" longitude="-73.761215" municipality="Valledupar" specimenCode="CBUMAG:ENT:41665" specimenCount="1" specimenCount-worker="1" stateProvince="Cesar">
<specimenCount id="7F6A049F84F9F1293866FEF5411B063E" count="1" type="worker">1 worker</specimenCount>
;
<collectingRegion id="3CA3A9B47A9D77B18C55A2265D6A926C" country="Colombia" name="Cesar">Cesar</collectingRegion>
,
<location id="76CD37A91005A8C60BF6D834EF634A49" LSID="urn:lsid:plazi:treatment:50DBCF393F2D54C583DC84C22407947F:76CD37A91005A8C60BF6D834EF634A49" country="Colombia" latitude="10.241965" longLatPrecision="1" longitude="-73.761215" name="Valledupar" stateProvince="Cesar">Valledupar</location>
, Vda. Tierras nuevas;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ED5BI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-73.761216,10.241965]}">
<geoCoordinate id="A19BBE573E75F1449D8339BFD07B3A95" degrees="10.241965" direction="north" orientation="latitude" precision="1" value="10.241965">10.241965 ° N</geoCoordinate>
<collectingMunicipality id="2DDC0C130EDD92981B963AD51899936D">Valledupar</collectingMunicipality>
,
<geoCoordinate id="1519CCD71BE7452F39B5ACA4F96CD24D" degrees="73.761216" direction="west" orientation="longitude" precision="1" value="-73.761215">73.761216 ° W</geoCoordinate>
<location id="AC74D85139FAC66BEF23BE10CCAEC810" LSID="urn:lsid:plazi:treatment:50DBCF393F2D54C583DC84C22407947F:AC74D85139FAC66BEF23BE10CCAEC810" country="Colombia" latitude="10.241965" longLatPrecision="1" longitude="-73.761215" municipality="Valledupar" name="Vda. Tierras nuevas" stateProvince="Cesar">Vda. Tierras nuevas</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ED5BI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-73.761216,10.241965]}">
<geoCoordinate id="A19BBE573E75F1449D8339BFD07B3A95" degrees="10.241965" direction="north" orientation="latitude" precision="1" value="10.241965">10.241965°N</geoCoordinate>
,
<geoCoordinate id="1519CCD71BE7452F39B5ACA4F96CD24D" degrees="73.761216" direction="west" orientation="longitude" precision="1" value="-73.761215">73.761216°W</geoCoordinate>
</named-content>
;
<elevation id="DF6A1A10A0B2B4BED845547EAD539DAA" metricMagnitude="2" metricUnit="m" metricValue="8.809999999999999" unit="m" value="881.0">
@ -211,25 +212,27 @@ a. s. l.
leg.;
<collectingMethod id="CCAACC241D468B12091E23F77D68635F">Pitfall</collectingMethod>
;
<named-content id="8238A32F86A0C54E30C2B90CE2696FF1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="D96D4DF1AFB6A60500BBC3FB33B28D1D">CBUMAG</collectionCode>
</named-content>
: ENT: 41665
<specimenCode id="432D7A6097B0265D505C28C57429F3CE">
<named-content id="8238A32F86A0C54E30C2B90CE2696FF1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:41665
</specimenCode>
</materialsCitation>
. •
<materialsCitation id="FDE5BB209FE4817CB08683F6FC55BC42" collectingDate="2004-02-08" collectingDateMax="2004-02-10" collectingDateMin="2004-02-08" collectionCode="IAvH" collectorName="Parque Nacional Natural el Tuparro &amp; I. Quintero &amp; E. Gonzalez &amp; Winkler" country="Colombia" elevation="135" latitude="5.331667" location="Cumaribo" longLatPrecision="1" longitude="-67.89083" specimenCount="1" specimenCount-worker="1" stateProvince="Vichada">
<materialsCitation id="FDE5BB209FE4817CB08683F6FC55BC42" collectingDate="2004-02-08" collectingDateMax="2004-02-10" collectingDateMin="2004-02-08" collectingMethod="Winkler" collectionCode="IAvH" collectorName="I. Quintero &amp; E. Gonzalez" country="Colombia" county="Cumaribo" elevation="135" latitude="5.331667" location="Parque Nacional Natural el Tuparro" longLatPrecision="1" longitude="-67.89083" municipality="Cgto. Santa Rita" specimenCode="IAvH-E-79867" specimenCount="1" specimenCount-worker="1" stateProvince="Vichada">
<specimenCount id="CAFBF5667CC690A8E30A7FB0C860A794" count="1" type="worker">1 worker</specimenCount>
;
<collectingRegion id="D91291BBDBC2D23CB7E50EFE1C66AAEA" country="Colombia" name="Vichada">Vichada</collectingRegion>
,
<location id="E7A96A17EAAFC7304B83F4418F449EF9" LSID="urn:lsid:plazi:treatment:50DBCF393F2D54C583DC84C22407947F:E7A96A17EAAFC7304B83F4418F449EF9" country="Colombia" latitude="5.331667" longLatPrecision="1" longitude="-67.89083" name="Cumaribo" stateProvince="Vichada">Cumaribo</location>
, Cgto. Santa Rita,
<collectorName id="33DB2C3251022F091EEE755E2EF9BD08">Parque Nacional Natural el Tuparro</collectorName>
<collectingCounty id="D09B58C1F092D2DE9D6F816E1066220E">Cumaribo</collectingCounty>
,
<collectingMunicipality id="4A78C8430BCD8773D5720A5A59E1D490">Cgto. Santa Rita</collectingMunicipality>
,
<location id="CAB5382728DBB6478630164408D2BF86" LSID="urn:lsid:plazi:treatment:50DBCF393F2D54C583DC84C22407947F:CAB5382728DBB6478630164408D2BF86" country="Colombia" county="Cumaribo" latitude="5.331667" longLatPrecision="1" longitude="-67.89083" municipality="Cgto. Santa Rita" name="Parque Nacional Natural el Tuparro" stateProvince="Vichada">Parque Nacional Natural el Tuparro</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EP5BI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-67.890833,5.331667]}">
<geoCoordinate id="94E1EC9B4DE11CDF7BD5FA9856015CF2" degrees="5.331667" direction="north" orientation="latitude" precision="1" value="5.331667">5.331667 ° N</geoCoordinate>
<geoCoordinate id="94E1EC9B4DE11CDF7BD5FA9856015CF2" degrees="5.331667" direction="north" orientation="latitude" precision="1" value="5.331667">5.331667°N</geoCoordinate>
,
<geoCoordinate id="275D56DE02F19615DEB1976CE443907E" degrees="67.890833" direction="west" orientation="longitude" precision="1" value="-67.89083">67.890833 ° W</geoCoordinate>
<geoCoordinate id="275D56DE02F19615DEB1976CE443907E" degrees="67.890833" direction="west" orientation="longitude" precision="1" value="-67.89083">67.890833°W</geoCoordinate>
</named-content>
;
<elevation id="725157A2AA1177286EA24A7EA17FEB77" metricMagnitude="2" metricUnit="m" metricValue="1.35" unit="m" value="135.0">
@ -245,12 +248,12 @@ a. s. l.
;
<collectorName id="B09DB89B9BEA079A38A94C4DB06A7454">E. González</collectorName>
legs.;
<collectorName id="D45D5A3B787977A64E1E6814C755BEC1">Winkler</collectorName>
<collectingMethod id="EAE24B4CA15D0F406E53F0E90E8C38A0">Winkler</collectingMethod>
;
<abbrev id="ABBRID0EU5BI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">
<collectionCode id="C4EE41EB494BAF2817E474B70192BAA2">IAvH</collectionCode>
</abbrev>
- E- 79867
<specimenCode id="549CCF7DA26F389D2536B6D0D4FDE8E5">
<abbrev id="ABBRID0EU5BI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">IAvH</abbrev>
-E-79867
</specimenCode>
</materialsCitation>
.
</paragraph>

View file

@ -1,48 +1,48 @@
<document id="D37728EF4B2E4A0E3DFD5D00703353D3" ID-DOI="10.3897/zookeys.1232.141693" ID-ZooBank="0AA099A2-EA57-4E27-937A-0976B9F8A4B5" ID-publisher-id="141693" URI-arpha="A9904ED3-9BDA-52DF-B936-1C42EC2713C5" URI-zoobank="http://zoobank.org/0AA099A2-EA57-4E27-937A-0976B9F8A4B5" article-type="research-article" checkinTime="1742225059022" checkinUser="pensoft" docAuthor="Arredondo, Brandon S. &amp; Guerrero, Roberto J." docDate="2025" docId="6573CDD71D425FF0AF74C1A06F70EF7C" docLanguage="en" docName="ZooKeys 1232: 131-172" docOrigin="ZooKeys 1232" docSource="https://zookeys.pensoft.net/article/141693/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nesomyrmex iku Arredondo &amp; Guerrero, 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="172" masterDocId="A9904ED39BDA52DFB9361C42EC2713C5" masterDocTitle="The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data" masterLastPageNumber="172" masterPageNumber="131" pageNumber="131" updateTime="1742225485493" updateUser="ExternalLinkService">
<mods:mods id="C97439E74A79445572F9655E6BA8543D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D4B07A22D26735C7587F2481D665AAA1">
<mods:title id="C00FE27570C05E6A71B033A6AA213E21">The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data</mods:title>
<document id="BCAD0550E706F8002F741223053ADA84" ID-DOI="10.3897/zookeys.1232.141693" ID-ZooBank="0AA099A2-EA57-4E27-937A-0976B9F8A4B5" ID-publisher-id="141693" URI-arpha="A9904ED3-9BDA-52DF-B936-1C42EC2713C5" URI-zoobank="http://zoobank.org/0AA099A2-EA57-4E27-937A-0976B9F8A4B5" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1742225059022" checkinUser="pensoft" docAuthor="Arredondo, Brandon S. &amp; Guerrero, Roberto J." docDate="2025" docId="6573CDD71D425FF0AF74C1A06F70EF7C" docLanguage="en" docName="ZooKeys 1232: 131-172" docOrigin="ZooKeys 1232" docSource="https://zookeys.pensoft.net/article/141693/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nesomyrmex iku Arredondo &amp; Guerrero, 2025, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="172" masterDocId="A9904ED39BDA52DFB9361C42EC2713C5" masterDocTitle="The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data" masterLastPageNumber="172" masterPageNumber="131" pageNumber="131" updateTime="1742234045672" updateUser="admin">
<mods:mods id="6195FFF8143EDECDD7099895B0512F12" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="255A4B6F0168AC4495ACD27E9EE0B47B">
<mods:title id="B5B95906ED8D1B4DD3B40E77D1EA7A47">The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data</mods:title>
</mods:titleInfo>
<mods:name id="918A0F0A070170C509B94874E6FBEF7F" type="personal">
<mods:role id="4A7CC7106E2CA482F7FEC0A8855ED31C">
<mods:roleTerm id="9D6CBA9A27D4420437859B5FC9AC5EA3">Author</mods:roleTerm>
<mods:name id="DBAE268E6156B6172CA13747A3D88E5A" type="personal">
<mods:role id="D69313D97DE3BB0F96043B66DBAEAF6D">
<mods:roleTerm id="89BDA1172780CEA882F78C0EB6DB303B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="53B8441968C505552692C39171A73BB3">Arredondo, Brandon S.</mods:namePart>
<mods:nameIdentifier id="5C93693B3AD76EE336A3A1B978CD6D53" type="ORCID">0000-0003-2777-5909</mods:nameIdentifier>
<mods:affiliation id="662C4D35CCD39902F02804ACDE20BA15">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
<mods:namePart id="538305FD091BFECE3D8161F79A32CB51">Arredondo, Brandon S.</mods:namePart>
<mods:nameIdentifier id="B395D0136E8AA4FAB2A1582AEFBC73AF" type="ORCID">0000-0003-2777-5909</mods:nameIdentifier>
<mods:affiliation id="DE0A7C94DD05845D18D9D781F45DC2B1">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:name id="7108C4B34878711C70AA9421EB26D91B" type="personal">
<mods:role id="04D7C1EDB000440BF550296FF8FDB474">
<mods:roleTerm id="8D49DDB355C3650796DF9D7995D82F82">Author</mods:roleTerm>
<mods:name id="F79B50A8226419942C6FBFC4DD847FEC" type="personal">
<mods:role id="FEDE89074D0871D337C8B575CC6DE400">
<mods:roleTerm id="559258329F79077AC18E7DC17998A376">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ECB6586179C7571B17A6D4E23A53445B">Guerrero, Roberto J.</mods:namePart>
<mods:nameIdentifier id="81A18C037A61491E870715E47355F74F" type="ORCID">0000-0003-3244-2754</mods:nameIdentifier>
<mods:affiliation id="B356EF2ECE3CE48A8977FD7882819414">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
<mods:namePart id="742F10D41C1F12878578DC216089833B">Guerrero, Roberto J.</mods:namePart>
<mods:nameIdentifier id="91614807B6AF735F4FFF779D78F53452" type="ORCID">0000-0003-3244-2754</mods:nameIdentifier>
<mods:affiliation id="EE20B180921EDEC7704767D237E778BB">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="993E7651AF6567C6472FBBAAFC6EE442">text</mods:typeOfResource>
<mods:relatedItem id="77A25770F829C4AA5EEA993DD9209308" type="host">
<mods:titleInfo id="9BF3F2AB33F5045CBD652AFFC076BE61">
<mods:title id="99DCE7EA4F7981D1E58901721995CA27">ZooKeys</mods:title>
<mods:typeOfResource id="0682F5E0F33D3A27B6FF2C18A8871FBD">text</mods:typeOfResource>
<mods:relatedItem id="1E7AC9B3B95227596414F964E98D3C09" type="host">
<mods:titleInfo id="812E19DD3F5097A6F8677DDC418B1262">
<mods:title id="508A38718177DDB99BF173284999A3BF">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="09AD3098A4C0A483086115AD8978E5BF">
<mods:date id="AFD76A5258FEA5314751AE8C1A2F5954">2025</mods:date>
<mods:detail id="64A45DD7234C8EBC9E2DFBA5397F9BC8" type="pubDate">
<mods:number id="2A5533C29C871D6C3E282D901ADA66D5">2025-03-17</mods:number>
<mods:part id="F238933A448D1EA2ABF89AF46309621D">
<mods:date id="FE37A37BC4F93228104D0B38CA2D9366">2025</mods:date>
<mods:detail id="129D749C255EFB1F29A702321D632236" type="pubDate">
<mods:number id="175155D46D95006E869AD42C1CE12056">2025-03-17</mods:number>
</mods:detail>
<mods:detail id="9F3BB6BE2A16573C95B46571CDF47B12" type="volume">
<mods:number id="60A6794A6E49B7EEDCCD6FDE886689C4">1232</mods:number>
<mods:detail id="4BC809096C1997EC0FCEE776BCF80B2E" type="volume">
<mods:number id="F00B07683544446B07581D3361D1F6A1">1232</mods:number>
</mods:detail>
<mods:extent id="0292B341EA88940AC04FE18B3614C6B7" unit="page">
<mods:start id="F77931AC28A0E60CB240D6566165F88B">131</mods:start>
<mods:end id="22CCC1A151C2115E78424598E96F7A2C">172</mods:end>
<mods:extent id="798D42814CF7006F0A6577D940D51F00" unit="page">
<mods:start id="0BAD1ED8C07F0F6F3FA34A8F713E0701">131</mods:start>
<mods:end id="5AD1433E97E197BD0015BEBD2A8632F8">172</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D7D61534932146E8CC9198E4C99C845D">journal article</mods:classification>
<mods:identifier id="B6143FB211341F4A492DF4876B87429B" type="DOI">10.3897/zookeys.1232.141693</mods:identifier>
<mods:identifier id="2B5C8CB97402AF1660326B789901AAB3" type="ZooBank">0AA099A2-EA57-4E27-937A-0976B9F8A4B5</mods:identifier>
<mods:classification id="38803E690ED5B748932B077A0CBEC6C0">journal article</mods:classification>
<mods:identifier id="C5EEF294F715FDF20A5B2CBD13EDC7FC" type="DOI">10.3897/zookeys.1232.141693</mods:identifier>
<mods:identifier id="B1DF2D1D7ED83823E3A2F26405098AB2" type="ZooBank">0AA099A2-EA57-4E27-937A-0976B9F8A4B5</mods:identifier>
</mods:mods>
<treatment id="6573CDD71D425FF0AF74C1A06F70EF7C" ID-DOI="http://doi.org/10.5281/zenodo.15040777" ID-Zenodo-Dep="15040777" ID-arpha="6573CDD7-1D42-5FF0-AF74-C1A06F70EF7C" ID-zoobank="https://zoobank.org/44B9782F-3DBC-467D-A3F8-58CFB3F6A0E4" LSID="urn:lsid:plazi:treatment:6573CDD71D425FF0AF74C1A06F70EF7C" LSID-ZBK="urn:lsid:zoobank.org:act:44B9782F-3DBC-467D-A3F8-58CFB3F6A0E4" httpUri="http://treatment.plazi.org/id/6573CDD71D425FF0AF74C1A06F70EF7C">
<treatment id="6573CDD71D425FF0AF74C1A06F70EF7C" ID-DOI="http://doi.org/10.5281/zenodo.15040777" ID-Zenodo-Dep="15040777" ID-arpha="6573CDD7-1D42-5FF0-AF74-C1A06F70EF7C" ID-zoobank="https://zoobank.org/44B9782F-3DBC-467D-A3F8-58CFB3F6A0E4" LSID="urn:lsid:plazi:treatment:6573CDD71D425FF0AF74C1A06F70EF7C" LSID-ZBK="urn:lsid:zoobank.org:act:44B9782F-3DBC-467D-A3F8-58CFB3F6A0E4" httpUri="http://treatment.plazi.org/id/6573CDD71D425FF0AF74C1A06F70EF7C" scope_family="Formicidae" scope_subFamily="Myrmicinae">
<subSubSection id="2ADF73E4088F5C75677CB391437AC2C8" type="nomenclature">
<paragraph id="E3EF52AC472A2D3BDC865AD21CCE6E95">
<taxonomicName id="CA25F1C8D9C9AC3271CBD61B97658B04" authorityName="Arredondo &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="iku" status="sp. nov.">
@ -63,7 +63,7 @@
<heading id="966BD015235105168EB0B78DCC10BF6A" reason="title">Type material.</heading>
</paragraph>
<paragraph id="0A84FF54960C2E42E7656CEAF6C052F8">
<materialsCitation id="B6F0703EBAF32FE9D8E0D15E4292B7B8" collectionCode="IAvH" country="Colombia" latitude="10.816667" location="Sierra Nevada de Santa Marta" longLatPrecision="1" longitude="-73.65" municipality="Santa Marta" specimenCount="1" specimenCount-worker="1" stateProvince="Magdalena" typeStatus="holotype">
<materialsCitation id="B6F0703EBAF32FE9D8E0D15E4292B7B8" collectionCode="IAvH" country="Colombia" latitude="10.816667" location="Sierra Nevada de Santa Marta" longLatPrecision="1" longitude="-73.65" municipality="Santa Marta" specimenCode="IAvH-E-75014" specimenCount="1" specimenCount-worker="1" stateProvince="Magdalena" typeStatus="holotype">
<emphasis id="157548D0DD8DE308CE2637498BFFD672" bold="true" italics="true">
<typeStatus id="8F7F0BED7F7980D16E72132FCDBA9240" type="holotype">Holotype</typeStatus>
</emphasis>
@ -79,21 +79,23 @@
<location id="403C257A7B4D64CF0D89558F5E61EA27" LSID="urn:lsid:plazi:treatment:6573CDD71D425FF0AF74C1A06F70EF7C:403C257A7B4D64CF0D89558F5E61EA27" country="Colombia" latitude="10.816667" longLatPrecision="1" longitude="-73.65" municipality="Santa Marta" name="Sierra Nevada de Santa Marta" stateProvince="Magdalena">Sierra Nevada de Santa Marta</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EX6BG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-73.650000,10.816667]}">
<geoCoordinate id="9A452472A9E8DB0B40ECB16515B35009" degrees="10.816667" direction="north" orientation="latitude" precision="1" value="10.816667">10.816667 ° N</geoCoordinate>
<geoCoordinate id="9A452472A9E8DB0B40ECB16515B35009" degrees="10.816667" direction="north" orientation="latitude" precision="1" value="10.816667">10.816667°N</geoCoordinate>
,
<geoCoordinate id="58259A7796CAEF7FADAC9FB64845263D" degrees="73.650000" direction="west" orientation="longitude" precision="1" value="-73.65">73.650000 ° W</geoCoordinate>
<geoCoordinate id="58259A7796CAEF7FADAC9FB64845263D" degrees="73.650000" direction="west" orientation="longitude" precision="1" value="-73.65">73.650000°W</geoCoordinate>
</named-content>
;
<abbrev id="ABBRID0E36BG" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">
<collectionCode id="6865ACACA3C08CE3A8BF2D471DF1FEC4">IAvH</collectionCode>
</abbrev>
- E- 75014 [
<specimenCode id="E737072C2EF13FDDA36A9A7AD2F54932">
<abbrev id="ABBRID0E36BG" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">IAvH</abbrev>
-E-75014
</specimenCode>
[
<abbrev id="ABBRID0EBAAI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">
<collectionCode id="D48DB006530919F2E00B3029E088A81A">IAvH</collectionCode>
</abbrev>
]
</materialsCitation>
.
<materialsCitation id="C256D56140FBE3890F83FB63A8E7AA63" collectionCode="CBUMAG" country="Colombia" latitude="10.816667" location="Sierra Nevada de Santa Marta" longLatPrecision="1" longitude="-73.65" municipality="Santa Marta" specimenCode="IAvH-E-75007" specimenCount="1" specimenCount-worker="1" stateProvince="Magdalena" typeStatus="paratype">
<emphasis id="332FE974F26F1C86412ACE5348CAD0F2" bold="true" italics="true">
<typeStatus id="80B72097919CB7D5EE36D31E72A99A3E" type="paratype">Paratype</typeStatus>
</emphasis>
@ -101,17 +103,18 @@
<emphasis id="02D99451F7084FFDDA9A7384FEE3B186" italics="true">n</emphasis>
= 1).
<specimenCount id="476E0BA70770CCBC19086D4EA2D40138" count="1" type="worker">1 worker</specimenCount>
; same data as
<typeStatus id="65E7B0F694763B2048BA38812013CD58" type="holotype">holotype</typeStatus>
;
<abbrev id="ABBRID0EKAAI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">
<collectionCode id="45973134E20B3B35037BAFA8545E0E0B">IAvH</collectionCode>
</abbrev>
- E- 75007 (
; same data as holotype;
<specimenCode id="15ABD55A5850EFC1C598BCE9946BBB21">
<abbrev id="ABBRID0EKAAI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">IAvH</abbrev>
-E-75007
</specimenCode>
(
<named-content id="30D1DFA4E19F99DB852C3EEF23A894F9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="C11B4FA222B3A486967B9F57292CAA7C">CBUMAG</collectionCode>
</named-content>
).
)
</materialsCitation>
.
</paragraph>
<caption id="453A961C5C6853A4DAA5F08F5C0780B9" ID-DOI="10.3897/zookeys.1232.141693.figure22" ID-arpha="2C07BC09-B76E-5886-9BF7-4F67DBB144AC" httpUri="https://binary.pensoft.net/fig/1282992" startId="F22">
<paragraph id="AE83582B5BAF2762D0E1900CAFA35949">
@ -123,8 +126,11 @@
</taxonomicName>
<taxonomicNameLabel id="FC8406B15CB7328B27A7A0B10ADCBCE3" rank="species">sp. nov.</taxonomicNameLabel>
Holotype worker (
<specimenCode id="63D59776244043968A7FED47CFB795CA">
<abbrev id="ABBRID0EFBAI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">IAvH</abbrev>
- E- 75014)
-E-75014
</specimenCode>
)
<emphasis id="0FBB71867B9E0F28CC04B7846A8420AB" bold="true">A</emphasis>
full-face view
<emphasis id="CC163AA380D2D2D87F7E2390E073FB8D" bold="true">B</emphasis>
@ -167,9 +173,7 @@ worker measurements.
0.28,
<abbrev id="ABBRID0E2CAI" xlink_title="Webers length">WL</abbrev>
1.13,
<abbrev id="ABBRID0E6CAI" xlink_title="Pronotal height">
<collectingCountry id="42318770EE370FE0A2A76A6392C0B86F" name="Philippines">PH</collectingCountry>
</abbrev>
<abbrev id="ABBRID0E6CAI" xlink_title="Pronotal height">PH</abbrev>
0.34,
<abbrev id="ABBRID0EDDAI" xlink_title="Petiole length">PTL</abbrev>
0.47,
@ -214,9 +218,7 @@ worker measurements.
0.30,
<abbrev id="ABBRID0EDFAI" xlink_title="Webers length">WL</abbrev>
1.18,
<abbrev id="ABBRID0EHFAI" xlink_title="Pronotal height">
<collectingCountry id="E078DA7E529B9BE25648B3229C4F8895" name="Philippines">PH</collectingCountry>
</abbrev>
<abbrev id="ABBRID0EHFAI" xlink_title="Pronotal height">PH</abbrev>
0.35,
<abbrev id="ABBRID0ELFAI" xlink_title="Petiole length">PTL</abbrev>
0.48,
@ -298,9 +300,7 @@ Dorsal surface of body with flattened erect hairs, mostly separated by a distanc
<heading id="1DC8154EC3AB3F8C8560BB08F1E8BC6F" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="6333427DA814ADB0454FFC5FC400B1FD">
The name of this species refers to the Arhuaco people with ancestral lands close to the
<typeStatus id="F85073B50DC461734F96A22D9C94D065">type</typeStatus>
locality of the species. The Arhuaco people self-identify as Iku, which translates as “ people ” in the
The name of this species refers to the Arhuaco people with ancestral lands close to the type locality of the species. The Arhuaco people self-identify as Iku, which translates as “ people ” in the
<collectingRegion id="24096B48CA36AE6EF45F4A6731B60025" country="Peru" name="Ica">Ika</collectingRegion>
language (
<bibRefCitation id="9295FDBDFD312D19F45010C9F16A35CC" author="Tracy" firstAuthor="Tracy" refId="B55" refString="Tracy H (1997) Kaʼgɨmmɨri nivizaku ni (La tierra es nuestra madre: La importancia de la tierra para la sobrevivencia del grupo étnico Ika: Otro punto de vista sobre la cuestión de las tierras en Bogotá: Instituto Lingüístico de Verano. http://www.sil.org/americas/colombia/pubs/38351.pdf" url="http://www.sil.org/americas/colombia/pubs/38351.pdf" year="1997">Tracy 1997</bibRefCitation>
@ -312,15 +312,13 @@ language (
<heading id="1A2E78445AE1D51636BF704CEEEF16E8" reason="title">Comments.</heading>
</paragraph>
<paragraph id="3C60AEC6AB26C8C31191A3522FBADF5F">
This species is only known from the
<typeStatus id="CC04CA5271863C70BBD8C7E19F62C255">type</typeStatus>
material.
This species is only known from the type material.
<taxonomicName id="B92E94F3436F346E407A9772A3C70C68" authorityName="Arredondo &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="iku" status="sp. nov.">
<emphasis id="EC982C9ED933467941A53F5B458149AA" italics="true">Nesomyrmex iku</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A12B41699598A264A9ABEAFBEAA6D4F5" rank="species">sp. nov.</taxonomicNameLabel>
is easily recognized by the shape of the straight mesosomal profile and by the lateral dorsopropodeal processes short, not covering spiracle in dorsal view. Morphologically, the closest species is
<taxonomicName id="5643D48F99449953563D8E6A7BF32A40" authorityName="Arredondo &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="xerophilus" status="sp. nov.">
<taxonomicName id="5643D48F99449953563D8E6A7BF32A40" authorityName="Arredondo-H &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="xerophilus" status="sp. nov.">
<emphasis id="A84391AC812CB5E58A447AD09AE22F29" italics="true">Nesomyrmex xerophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09EB4E17242CD13F8946B021B08AC1C" rank="species">sp. nov.</taxonomicNameLabel>
@ -329,7 +327,7 @@ is easily recognized by the shape of the straight mesosomal profile and by the l
<emphasis id="DEE42D0C17460C6E6EEBE1F6CB09BAF4" italics="true">N. iku</emphasis>
</taxonomicName>
does not have such a projection on the frontal lobes. On the other hand, in
<taxonomicName id="4F0A85C5B66040DF86ADBFBC9F8D24A1" authorityName="Arredondo &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="xerophilus">
<taxonomicName id="4F0A85C5B66040DF86ADBFBC9F8D24A1" authorityName="Arredondo-H &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="xerophilus">
<emphasis id="FBEC8C396CD4E8F340D60EAED5C80223" italics="true">N. xerophilus</emphasis>
</taxonomicName>
the propodeal spiracles are not visible in dorsal view, since the lateral portion of the dorsopropodeal projects above them, whereas in
@ -343,23 +341,25 @@ Another species with similar morphology is
<taxonomicName id="11DD6EDD4BACC572D50537E58B3F7650" baseAuthorityName="Kempf" baseAuthorityYear="1958" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="CF360930289ACDB1F3F32D90CF8DC73D" italics="true">N. brasiliensis</emphasis>
</taxonomicName>
. When comparing the
<typeStatus id="79F0A91B9BAA1490C06D580CFA3DA5AB">type</typeStatus>
workers of
. When comparing the type workers of
<taxonomicName id="F332CDEE568F730B79D8C45CE59B38CB" authorityName="Arredondo &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="iku">
<emphasis id="1A7F0DD44A06890E815A4A9BF43F91C8" italics="true">N. iku</emphasis>
</taxonomicName>
(
<specimenCode id="2808092B68C39515802E8B0E60BCA52B">
<abbrev id="ABBRID0EBLAI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">
<collectionCode id="9E481180EB1E2660A2FB545208CE8460">IAvH</collectionCode>
</abbrev>
- E- 75014,
-E-75014
</specimenCode>
,
<specimenCode id="67030EE9AE085A2FF64E978630C7A33A">
<abbrev id="ABBRID0EFLAI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">
<collectionCode id="FC9F46BD12EE0428A2FE16AD0B0B5BF8">IAvH</collectionCode>
</abbrev>
- E- 75007) with the
<typeStatus id="FB0C3A5ED3A806C87DC658CF56B02F9F">type</typeStatus>
ones of
-E-75007
</specimenCode>
) with the type ones of
<taxonomicName id="6B8CFA315BE08B641108D9B31DDB1ECB" baseAuthorityName="Kempf" baseAuthorityYear="1958" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="47A8AA9C3A582ED9202E3E01A8330170" italics="true">N. brasiliensis</emphasis>
</taxonomicName>
@ -392,7 +392,7 @@ is foveated in contrast to the frontal area striated in
<emphasis id="E7386F911BD5755E79FC84E1BBC397B4" italics="true">N. iku</emphasis>
</taxonomicName>
, and
<taxonomicName id="6F1BACC0045D1B95FA9FA84F527BFFF2" authorityName="Arredondo &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="xerophilus">
<taxonomicName id="6F1BACC0045D1B95FA9FA84F527BFFF2" authorityName="Arredondo-H &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="xerophilus">
<emphasis id="473111FA88863FB7C3DFD7CE7723CCFB" italics="true">N. xerophilus</emphasis>
</taxonomicName>
will be possible when more material can be analyzed.

View file

@ -0,0 +1,284 @@
<document id="921E95A21A614FDD38389783DB3787EE" ID-DOI="10.3897/zookeys.1232.141693" ID-ZooBank="0AA099A2-EA57-4E27-937A-0976B9F8A4B5" ID-publisher-id="141693" URI-arpha="A9904ED3-9BDA-52DF-B936-1C42EC2713C5" URI-zoobank="http://zoobank.org/0AA099A2-EA57-4E27-937A-0976B9F8A4B5" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1742225059022" checkinUser="pensoft" docAuthor="Arredondo, Brandon S. &amp; Guerrero, Roberto J." docDate="2025" docId="67EE9217248554D499111ECCC135C492" docLanguage="en" docName="ZooKeys 1232: 131-172" docOrigin="ZooKeys 1232" docSource="https://zookeys.pensoft.net/article/141693/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nesomyrmex pleuriticus" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="172" masterDocId="A9904ED39BDA52DFB9361C42EC2713C5" masterDocTitle="The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data" masterLastPageNumber="172" masterPageNumber="131" pageNumber="131" updateTime="1742234045672" updateUser="admin">
<mods:mods id="6D69BAF54576CEBBDC401A74A99F0EA2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="02625E13009B49EA274FA46CC79F77E1">
<mods:title id="25D4FDE2ADBF2C6F8742796C9153EC9B">The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data</mods:title>
</mods:titleInfo>
<mods:name id="8E29462BB9B07172E0C36CDCA4F797E2" type="personal">
<mods:role id="9091B5D8A2F577D5802F2E1D9AB9FB44">
<mods:roleTerm id="5174861221AE377F7787F977AE13EB87">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1A7137C574C18EC2B644A8FE089CBF2C">Arredondo, Brandon S.</mods:namePart>
<mods:nameIdentifier id="C7839B434A05164E1B52335AA3963478" type="ORCID">0000-0003-2777-5909</mods:nameIdentifier>
<mods:affiliation id="23FD1C17102B6AA6DC84C08D62A8D8ED">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:name id="9F899933C298220BE2409F161F834864" type="personal">
<mods:role id="FA5818176878B646213C8E1FDB6D6E45">
<mods:roleTerm id="67142F9A91A981D28F79038A30E60F7C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="286C1D65156090430CF94F086CEC4832">Guerrero, Roberto J.</mods:namePart>
<mods:nameIdentifier id="DF291762379469F77BBABD9F6C8B546E" type="ORCID">0000-0003-3244-2754</mods:nameIdentifier>
<mods:affiliation id="95399096239533604DF82B2F78BCEF12">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="1113D7385B61708804DDC1F2DFCCCEEC">text</mods:typeOfResource>
<mods:relatedItem id="2463A7F854997D8974FE0201F8C04DC5" type="host">
<mods:titleInfo id="77EA10C3FD26552129568D53927411AA">
<mods:title id="CDF0437E4AF28EA6E8B09ECA8D73A4D5">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="D3C9C77F4B4494F6349A853784B444EF">
<mods:date id="6239CEC75FAAF82A945AF575F6CBD6A2">2025</mods:date>
<mods:detail id="7BBD52387771642BA1858901562ED69E" type="pubDate">
<mods:number id="6F0FFE511E27366C172948A9C4A75592">2025-03-17</mods:number>
</mods:detail>
<mods:detail id="4361E82F289AE56D97AEB53B031088C7" type="volume">
<mods:number id="FB1B78CEA3566CC278DB5A524BDF3D44">1232</mods:number>
</mods:detail>
<mods:extent id="4937FADE6697B8CCA8AF99A8644987C1" unit="page">
<mods:start id="0C66BD985D289D5BA8E407862CD85FC6">131</mods:start>
<mods:end id="42ADCD651CB23ADEB26463415FCD70BE">172</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2F1FD951E766D5E84D00F0D6F8B64FD5">journal article</mods:classification>
<mods:identifier id="1ADCFA93900B0D230B1DBB17F5F071EF" type="DOI">10.3897/zookeys.1232.141693</mods:identifier>
<mods:identifier id="718EB9A102DB09FEA4450FFDC222B116" type="ZooBank">0AA099A2-EA57-4E27-937A-0976B9F8A4B5</mods:identifier>
</mods:mods>
<treatment id="67EE9217248554D499111ECCC135C492" ID-DOI="http://doi.org/10.5281/zenodo.15040786" ID-Zenodo-Dep="15040786" ID-arpha="67EE9217-2485-54D4-9911-1ECCC135C492" LSID="urn:lsid:plazi:treatment:67EE9217248554D499111ECCC135C492" httpUri="http://treatment.plazi.org/id/67EE9217248554D499111ECCC135C492" scope_family="Formicidae" scope_subFamily="Myrmicinae">
<subSubSection id="A0E942B1AD1A936D7FBD43D7EBEBDBC5" type="nomenclature">
<paragraph id="C5D70E0A643EE7F5E06CF786758BE848">
<taxonomicName id="1199C2B9797196C04BABE9B9E896D8D8" ID-CoL="CM7ZG" authority="(Kempf, 1959)" baseAuthorityName="Kempf" baseAuthorityYear="1959" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="pleuriticus">
<emphasis id="938DC87DC3BBF0BC667B7ED89751D9D6" italics="true">Nesomyrmex pleuriticus</emphasis>
(
<bibRefCitation id="F2404F8D961FACD37DC935CEC2F9B972" author="Kempf WW" journalOrPublisher="Studia Entomologica" pagination="391-432" refId="B37" refString="Kempf WW (1959) A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae). Studia Entomologica 2: 391-432." title="A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae)." volume="2" year="1959">Kempf, 1959</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="51BFFDE7CB4A3568D50119C44EB75F77" type="description">
<paragraph id="A0264A5CFC98A4DD9318A5E1C4FE5E2B">
<figureCitation id="3EEEA2F81B1951FED00E7B1C2AFED568" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Nesomyrmex pleuriticus worker (IAvH - E- 79866) A full-face view B lateral view C dorsal view. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1232.141693.figure25" httpUri="https://binary.pensoft.net/fig/1282995">Figs 25</figureCitation>
,
<figureCitation id="7BE66DC0F07E1AF9750D773BACB831C2" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distributional map of Nesomyrmex species from the tropical dry forest in Colombia." figureDoi="10.3897/zookeys.1232.141693.figure29" httpUri="https://binary.pensoft.net/fig/1282999">29</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F06E6AD8AE71C0A03482037042D2BF29" type="reference_group">
<paragraph id="F6D07B9C7C69D8B00ED5A36EC28CD854">
<treatmentCitationGroup id="C46BFD5F41F664DE0BA1EDAD7EA06247">
<treatmentCitation id="B4B14DDA7A85437B4FE9813A02DD6EA3" author="Kempf WW" page="414" year="1959">
<taxonomicName id="82C2D70F2DA00DB157CEED2E46DB97E6" ID-CoL="3TY4Z" authority="Kempf, 1959: 414" authorityName="Kempf" authorityPageNumber="414" authorityYear="1959" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="pleuriticus" subGenus="Nesomyrmex">
<emphasis id="63FAA0C5AC88122949DD51A66D98F254" italics="true">Leptothorax</emphasis>
(
<emphasis id="0F079B6A190AC07B67E374B985FC9C62" italics="true">Nesomyrmex</emphasis>
)
<emphasis id="4B0188E41273A2B634E2318AC6241165" italics="true">pleuriticus</emphasis>
<comment id="7173209A39F338553ADA2E2974EDCB20">
<bibRefCitation id="5DFA3BAC4548309AC30936A29C3209CE" author="Kempf WW" journalOrPublisher="Studia Entomologica" pagination="391-432" refId="B37" refString="Kempf WW (1959) A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae). Studia Entomologica 2: 391-432." title="A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae)." volume="2" year="1959">Kempf, 1959: 414</bibRefCitation>
.
<typeStatus id="27E729EB1DA5C297F749FBA82415E206" type="syntype">Syntype</typeStatus>
series (male, queen, worker):
<collectingCountry id="E6EE7A673455484761DAE7923B96485D" name="Guyana">Guyana</collectingCountry>
, Kartabo. [
<collectionCode id="EDABE60BF75C71A2A7E90E2B3974E72A">MCZC</collectionCode>
,
<named-content id="66D9AA2EC3B2F43B6DA613374BAF9BBF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/sao-paulo-museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museo de Zoologia da Universidade de São Paulo">
<collectionCode id="B196A70C149A157FB0A80C8EAA30F67F" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" type="Museum">MZSP</collectionCode>
</named-content>
]. (image examined,
<collectionCode id="BFBF4B40FFD4A95AD4F5937679F39376" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" type="Museum">MZSP</collectionCode>
87353,
<collectionCode id="B33FBD4E802F82FBD0983CCF8B8D87EB" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" type="Museum">MZSP</collectionCode>
87354).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="B8FF7CD0580B2F6D156C4538B884DD5B">
Combination in
<taxonomicName id="BF03F99C909D4F272D06255A3DC8E3F3" ID-CoL="CM5NR" authorityName="Wheeler" authorityYear="1910" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="9C35DC31091DF0A6E4625F7023C3126A" italics="true">Nesomyrmex</emphasis>
</taxonomicName>
:
<bibRefCitation id="CD0D91E719DB211B0E4521B0207F233C" author="Bolton B" firstAuthor="Bolton" journalOrPublisher="Memoirs of the American Entomological Society" page="272" pagination="1-370" refId="B5" refString="Bolton B (2003) Synopsis and classification of Formicidae. Memoirs of the American Entomological Society 71: 1370." title="Synopsis and classification of Formicidae." volume="71" year="2003">Bolton 2003: 272</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIGBI" type="description">
<paragraph id="A2CD427FBCC7486362D9542602C4EE98">
<heading id="B29E9446F210EE2F2617F9C4DECCDAEA" reason="title">Worker measurements.</heading>
</paragraph>
<paragraph id="DE1FFF0D932F9591684F59A31BADF091">
(
<emphasis id="EE73F8A4D95EA15E7AE03B1FC9925EFA" italics="true">n</emphasis>
= 1)
<abbrev id="ABBRID0EQGBI" xlink_title="Head length">HL</abbrev>
0.61,
<abbrev id="ABBRID0EUGBI" xlink_title="Head width">HW</abbrev>
0.58,
<abbrev id="ABBRID0EYGBI" xlink_title="Scape length">SL</abbrev>
0.50, ML 0.25,
<abbrev id="ABBRID0E3GBI" xlink_title="Eye length">EL</abbrev>
0.15,
<abbrev id="ABBRID0EAHBI" xlink_title="Pronotal width">PW</abbrev>
0.36,
<abbrev id="ABBRID0EEHBI" xlink_title="Petiole width">PTW</abbrev>
0.16,
<abbrev id="ABBRID0EIHBI" xlink_title="Postpetiole width">PPW</abbrev>
0.22,
<abbrev id="ABBRID0EMHBI" xlink_title="Webers length">WL</abbrev>
0.77,
<abbrev id="ABBRID0EQHBI" xlink_title="Pronotal height">PH</abbrev>
0.21,
<abbrev id="ABBRID0EUHBI" xlink_title="Petiole length">PTL</abbrev>
0.242,
<abbrev id="ABBRID0EYHBI" xlink_title="Petiole height">PTH</abbrev>
0.16,
<abbrev id="ABBRID0E3HBI" xlink_title="Postpetiole length">PPL</abbrev>
0.18,
<abbrev id="ABBRID0EAIBI" xlink_title="Postpetiole height">PPH</abbrev>
0.19,
<abbrev id="ABBRID0EEIBI" xlink_title="Gaster length">GL</abbrev>
0.64.
</paragraph>
<caption id="8935D932088A0846E30B147A170ADE20" ID-DOI="10.3897/zookeys.1232.141693.figure25" ID-arpha="4BA193D4-6461-5170-8A93-78A82BD545A7" httpUri="https://binary.pensoft.net/fig/1282995" startId="F25">
<paragraph id="4278283AE1B1406BB092B70AAB9DE14A">
<label id="47B5E1B934227EF72A54B0CCAF9143C9">Figure 25.</label>
</paragraph>
<paragraph id="FD3E7CCDF733C4364F4D763D16D5B131">
<taxonomicName id="F4E7F425C906940B0C211BC7343F7C2C" baseAuthorityName="Kempf" baseAuthorityYear="1959" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="pleuriticus">
<emphasis id="B94650E713CD5AB06AB8365B75409D5B" italics="true">Nesomyrmex pleuriticus</emphasis>
</taxonomicName>
worker (
<specimenCode id="A6B683F50757888A9A5453B7A9E49691">
<abbrev id="ABBRID0E2IBI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">IAvH</abbrev>
-E-79866
</specimenCode>
)
<emphasis id="AE30E224F4B432B59729DA636679D666" bold="true">A</emphasis>
full-face view
<emphasis id="67CAA5E40E32D03C497E72960AB71B61" bold="true">B</emphasis>
lateral view
<emphasis id="D54EA99F5C2B3930BFD2763D91715FFD" bold="true">C</emphasis>
dorsal view. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EKJBI" type="Geographic range">
<paragraph id="A5CFD69A75D8AE2764E7D971B830C448">
<heading id="1191AAB5A11F2F5FF6739AB9B81ACB00" reason="title">Geographic range.</heading>
</paragraph>
<paragraph id="D87782EC04B919FFA3ED2622407047C1">
<collectingCountry id="A79D15FC2AEE6B4B6C940C4D382F3CE0" name="Bolivia">Bolivia</collectingCountry>
,
<collectingCountry id="2FD102FE710A443404E0C4C3B63A8F0D" name="Brazil">Brazil</collectingCountry>
,
<collectingCountry id="D11AE79110BF10EF626FC167C82FE2A3" name="Colombia">Colombia</collectingCountry>
,
<collectingCountry id="716D1E0410BF67C8C0721D7FF6BC9456" name="Costa Rica">Costa Rica</collectingCountry>
,
<collectingCountry id="F23ED8029525CF9E314C4EEB9A2B05C6" name="Ecuador">Ecuador</collectingCountry>
, French
<collectingCountry id="E187ABA5D91D123E727DF58EDACA4DD5" name="Guyana">Guyana</collectingCountry>
,
<collectingCountry id="68661438C0BE642C82E188C0380DE2A0" name="Guatemala">Guatemala</collectingCountry>
,
<collectingCountry id="777A8E1CC70A6D4A4B749E831B45AB72" name="Guyana">Guyana</collectingCountry>
,
<collectingCountry id="35E61C5C96DE96DD119D1F53E7F3E24F" name="Honduras">Honduras</collectingCountry>
,
<collectingCountry id="D8CAAF751797EE18A0A5A78D449166F8" name="Mexico">Mexico</collectingCountry>
,
<collectingCountry id="53B2C0BC690AB637EC8AC5DF21D3697F" name="Panama">Panama</collectingCountry>
,
<collectingCountry id="7E0A7BB09171EBD655C239CA51888F8A" name="Suriname">Surinam</collectingCountry>
,
<collectingCountry id="9701ACF699186E61ED17DE8A244646FB" name="Venezuela">Venezuela</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPJBI" type="material">
<paragraph id="FD96916C56B7613238F60ACED802BE97">
<heading id="592D04C501FA32F02CA663CA24F5AD68" reason="title">Examined material.</heading>
</paragraph>
<paragraph id="D478036823E7AFE4C7911582756E1677">
<materialsCitation id="3FF7871AD87396EE9C35C602376D25D2" collectingDate="2004-02-08" collectingDateMax="2004-02-10" collectingDateMin="2004-02-08" collectingMethod="Winkler" collectionCode="IAvH" collectorName="I. Quintero &amp; E. Gonzalez" country="Colombia" county="Cumaribo" elevation="135" latitude="5.331667" location="Parque Nacional Natural el Tuparro" longLatPrecision="1" longitude="-67.89083" municipality="Cgto. Santa Rita" specimenCode="IAvH-E-79866" specimenCount="1" specimenCount-worker="1" stateProvince="Vichada">
<collectingCountry id="115CCD15F7A708B1C88BE69201E72953" name="Colombia">Colombia</collectingCountry>
<specimenCount id="D06130DB6122BC1B7F0B51F859FA198F" count="1" type="worker">1 worker</specimenCount>
;
<collectingRegion id="AEFB3B26D83CFB6CA0B36D6509B476E9" country="Colombia" name="Vichada">Vichada</collectingRegion>
,
<collectingCounty id="33EA0ABCF9047FCF23D199C12B79DAEF">Cumaribo</collectingCounty>
,
<collectingMunicipality id="F803CB97B68D245B5EBFDB7A8940741B">Cgto. Santa Rita</collectingMunicipality>
,
<location id="C63BB21C22748024D905E61B2C9E00BF" LSID="urn:lsid:plazi:treatment:67EE9217248554D499111ECCC135C492:C63BB21C22748024D905E61B2C9E00BF" country="Colombia" county="Cumaribo" latitude="5.331667" longLatPrecision="1" longitude="-67.89083" municipality="Cgto. Santa Rita" name="Parque Nacional Natural el Tuparro" stateProvince="Vichada">Parque Nacional Natural el Tuparro</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYJBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-67.890833,5.331667]}">
<geoCoordinate id="FAB9779C78DCCFBD529DB20F7DBAD6C2" degrees="5.331667" direction="north" orientation="latitude" precision="1" value="5.331667">5.331667°N</geoCoordinate>
,
<geoCoordinate id="1068D8F0BDA65D7843A2754383E1036C" degrees="67.890833" direction="west" orientation="longitude" precision="1" value="-67.89083">67.890833°W</geoCoordinate>
</named-content>
;
<elevation id="64ACCA367D9840D04A77EE44B6D664DC" metricMagnitude="2" metricUnit="m" metricValue="1.35" unit="m" value="135.0">
<quantity id="905A2D5210ACB8E574FE7AF78431DB8E" metricMagnitude="2" metricUnit="m" metricValue="1.35" unit="m" value="135.0">135 m</quantity>
a. s. l.
</elevation>
;
<date id="3887DF7CBBBC8747C40D17A316AFF018" value="2004-02-08">
<collectingDate id="978EB3E350750881FF483C0413BFC0DD" value="2004-02-08">08 Feb. 2004</collectingDate>
</date>
<date id="8E48AC40F097B147FAD809186D29FDAC" value="2004-02-10">
<collectingDate id="CABFBE78007FE0788B36FC430D469B17" value="2004-02-10">10 Feb. 2004</collectingDate>
</date>
;
<collectorName id="2CF22131254CDE161813C87ECCAA2F32">I. Quintero</collectorName>
;
<collectorName id="68E2F5152ACEC7A33AC796D9B3D98516">E. González</collectorName>
legs.;
<collectingMethod id="8471D4D2201CBAE80D1E65FC5FC27726">Winkler</collectingMethod>
;
<specimenCode id="07ABA6435583A9C03C26A58C7705F812">
<abbrev id="ABBRID0E4JBI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">IAvH</abbrev>
-E-79866
</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBKBI" type="natural history">
<paragraph id="17BFC73E8210A4BDA605C9D491EBA414">
<heading id="CD995667C2FF204464D66FEBB5BE9BAA" reason="title">Natural history.</heading>
</paragraph>
<paragraph id="29462CFC91EC603116930011852BC468">Foragers were collected on vegetation.</paragraph>
</subSubSection>
<subSubSection id="SECID0EGKBI" type="comments">
<paragraph id="4A7FCDFA5CBCD8D8B31D6626E15FAA5C">
<heading id="C9DCC061925B01D6B4042BF08B262B9D" reason="title">Comments.</heading>
</paragraph>
<paragraph id="8996D43619B58E132E09B02CCBFE626D">This species can be distinguished from other ones by having a straight anterior margin of the pronotum, an opaque mesosoma with striate sculpture and imbricate microsculpture, a dorsum of the postpetiole with longitudinal striations, scapes and legs covered with long, erect hairs, and a smooth, shiny clypeus.</paragraph>
<paragraph id="A3CE256AA6D5FD66B0BDD145E3F66070">
The previous records of
<taxonomicName id="C5B2C9CA68EA03420451F792F61F750F" baseAuthorityName="Kempf" baseAuthorityYear="1959" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="pleuriticus">
<emphasis id="CDCC0A37AD0DE0D3E66613D4D6DE501F" italics="true">N. pleuriticus</emphasis>
</taxonomicName>
for the tropical dry forest in
<collectingCountry id="0845302DE70FD3D3D48A76802AED481B" name="Colombia">Colombia</collectingCountry>
are based on information recorded for the departments of
<collectingRegion id="B59724D3D3FFDEFFC3616943E10C51E5" country="Colombia" name="Cauca">Cauca</collectingRegion>
(
<bibRefCitation id="FFBA1BF52F930F8ED527E2DA7788B0C1" DOI="10.15517/rbt.v62i0.16341" author="Chacon de Ulloa" etAl="et al." firstAuthor="Chacon de Ulloa" issue="1" journalOrPublisher="Revista de Biología Tropical" pagination="277-287" refId="B11" refString="Chacón de Ulloa P, Valdes-Rodriguez S, Hurtado-Giraldo A, Cleopatra Pimienta M (2014) Arboreal ants of Gorgona National Park (Pacific of Colombia). Revista de Biología Tropical 62 (1): 277287. https://doi.org/10.15517/rbt.v62i0.16341" title="Arboreal ants of Gorgona National Park (Pacific of Colombia)." volume="62" year="2014">Chacón de Ulloa et al. 2014</bibRefCitation>
) and
<collectingRegion id="7971AB41EDD9D0DFEA498AD3906BCCD4" country="Colombia" name="Valle del Cauca">Valle del Cauca</collectingRegion>
(
<bibRefCitation id="058C0AE09B8F6BB2CE6B781CEF78FBD4" author="Armbrecht" etAl="et al." firstAuthor="Armbrecht" issue="3" journalOrPublisher="Pan-Pacific Entomologist" pagination="196-209" refId="B2" refString="Armbrecht I, Tischer I, Chacón P (2001) Nested subsets and partition patterns in ant assemblages (Hymenoptera, Formicidae) of Colombian dry forest fragments. Pan-Pacific Entomologist 77 (3): 196209." title="Nested subsets and partition patterns in ant assemblages (Hymenoptera, Formicidae) of Colombian dry forest fragments." volume="77" year="2001">Armbrecht et al. 2001</bibRefCitation>
,
<bibRefCitation id="C15D1FD7700413C2B1C360DFE1CDAAC7" author="Chacon de Ulloa" etAl="et al." firstAuthor="Chacon de Ulloa" issue="2" journalOrPublisher="Biota Colombiana" pagination="165-181" refId="B10" refString="Chacón de Ulloa PA, Osorio-García M, Achury R, Bermúdez-Rivas C (2012) Hormigas (Hymenoptera: Formicidae) del Bosque seco tropical (Bs-T) de la cuenca alta del Rio Cauca, Colombia. Biota Colombiana 13 (2): 165181." title="Hormigas (Hymenoptera: Formicidae) del Bosque seco tropical (Bs-T) de la cuenca alta del Rio Cauca, Colombia." volume="13" year="2012">Chacón de Ulloa et al. 2012</bibRefCitation>
) but we were unable to analyze these specimens to confirm the taxonomic identity of these ants (see comments on
<taxonomicName id="A3AFAB4544B4852E5A22488287012BE0" authorityName="Longino" authorityYear="2006" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="vargasi">
<emphasis id="79A2FD7E63CEEC5A78A5B5454B3DCA79" italics="true">N. vargasi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,371 @@
<document id="15958EE39AEE89DA8D8561DFC80491CA" ID-DOI="10.3897/zookeys.1232.141693" ID-ZooBank="0AA099A2-EA57-4E27-937A-0976B9F8A4B5" ID-publisher-id="141693" URI-arpha="A9904ED3-9BDA-52DF-B936-1C42EC2713C5" URI-zoobank="http://zoobank.org/0AA099A2-EA57-4E27-937A-0976B9F8A4B5" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1742225059022" checkinUser="pensoft" docAuthor="Arredondo, Brandon S. &amp; Guerrero, Roberto J." docDate="2025" docId="9409219A903D5F39BBB82133FBBCAC1C" docLanguage="en" docName="ZooKeys 1232: 131-172" docOrigin="ZooKeys 1232" docSource="https://zookeys.pensoft.net/article/141693/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nesomyrmex spininodis" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="172" masterDocId="A9904ED39BDA52DFB9361C42EC2713C5" masterDocTitle="The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data" masterLastPageNumber="172" masterPageNumber="131" pageNumber="131" updateTime="1742234045672" updateUser="admin">
<mods:mods id="1140956B328AF05A75154F5BD599F418" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DB6659A3865F19F789332541040F4161">
<mods:title id="648F697D2F4364BB9168F2448EF7B463">The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data</mods:title>
</mods:titleInfo>
<mods:name id="1FC8FA541DE6C00E4763480666CB2F24" type="personal">
<mods:role id="2E908BDDDA958C7859CC391D7DD648A5">
<mods:roleTerm id="33A8209F141840F96F55873AAD5C82ED">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9851B18F5D61669DE436012A3255712C">Arredondo, Brandon S.</mods:namePart>
<mods:nameIdentifier id="36BEB8EE92C63B05577D6AF7AE1C19DC" type="ORCID">0000-0003-2777-5909</mods:nameIdentifier>
<mods:affiliation id="2BCF6E99E0FFD309A0DBAC6DECE8AD14">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:name id="094A9656D129E6C172D76CBF70E6BB86" type="personal">
<mods:role id="C1368EDE0E77887EC099B414E65376D1">
<mods:roleTerm id="6A09A6A2D368E892F9ABA63931F4281E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6AC999980D91EFF35160EBC3C0B2F63B">Guerrero, Roberto J.</mods:namePart>
<mods:nameIdentifier id="B5598183506EEC0692CDE9521C5748A3" type="ORCID">0000-0003-3244-2754</mods:nameIdentifier>
<mods:affiliation id="74857081A0B2BE30757D76CD8E989123">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="FA9563E5FB1401F2024762C2C8FA0A16">text</mods:typeOfResource>
<mods:relatedItem id="CC723B923A9264AB5AEFB0461D3C0364" type="host">
<mods:titleInfo id="5CA2E866ABA0FA50228327F6CF43ACB1">
<mods:title id="68AE12702AC6B2A6D79A87026EC7E77F">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="F2B20DF3269832DFACDC515BF43C9020">
<mods:date id="802A147C3E1B3F94CF7F0E7091A6D8BA">2025</mods:date>
<mods:detail id="9E3B5D877DEB938C96AC8211C55EC5C5" type="pubDate">
<mods:number id="AB16E0816FC977C3C9DF456141FC97BF">2025-03-17</mods:number>
</mods:detail>
<mods:detail id="8B1F40114966B19D07CCFDFEABE3F1F8" type="volume">
<mods:number id="400A9D0E887764485A5353AD0224DAC1">1232</mods:number>
</mods:detail>
<mods:extent id="4F0ED57A3389529E2F86AF4AC5B74551" unit="page">
<mods:start id="82DD026DC811872D42DCBFBE9744B22B">131</mods:start>
<mods:end id="DD13DDD197A7C8147CFFBC3A10F396F0">172</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7A2F1743F77964EC73E25D0016FAAAC5">journal article</mods:classification>
<mods:identifier id="C25F859C2E5CDCF3000B8E72B6F3E8D1" type="DOI">10.3897/zookeys.1232.141693</mods:identifier>
<mods:identifier id="80AC90D27F07941F4F2CF433F319166E" type="ZooBank">0AA099A2-EA57-4E27-937A-0976B9F8A4B5</mods:identifier>
</mods:mods>
<treatment id="9409219A903D5F39BBB82133FBBCAC1C" ID-arpha="9409219A-903D-5F39-BBB8-2133FBBCAC1C" LSID="urn:lsid:plazi:treatment:9409219A903D5F39BBB82133FBBCAC1C" httpUri="http://treatment.plazi.org/id/9409219A903D5F39BBB82133FBBCAC1C" scope_family="Formicidae" scope_subFamily="Myrmicinae">
<subSubSection id="3FBCC6FA695190C86E1C0219370D423B" type="nomenclature">
<paragraph id="F73EDD139B376A845D006B69D2A5DF5F">
<taxonomicName id="AD7E30F90601D1263C66AEFF94421CAF" ID-CoL="CM823" authority="(Mayr, 1887)" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="spininodis">
<emphasis id="B5792E194092C9F050BAC252AB448ECB" italics="true">Nesomyrmex spininodis</emphasis>
(
<bibRefCitation id="0FC8B229AE468E736E25F241E2A24DEB" author="Mayr G" journalOrPublisher="Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien" pagination="511-632" refId="B41" refString="Mayr G (1887) Sudamerikanische Formiciden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 37: 511-632." title="Südamerikanische Formiciden." volume="37" year="1887">Mayr, 1887</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F46D06FA7FEC54EBE17B7163C3F8F310" type="description">
<paragraph id="C26CEC8A6D46A3D600C1C3FBB4D1926E">
<figureCitation id="E06CC6036BD1B8E1B0DC3DD3288DEBBD" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Nesomyrmex spininodis worker (IAvH - E- 251254) A full-face view B lateral view C dorsal view. Scale bars: 0.2 mm (A, B); 0.5 mm (C)." figureDoi="10.3897/zookeys.1232.141693.figure26" httpUri="https://binary.pensoft.net/fig/1282996">Figs 26</figureCitation>
,
<figureCitation id="9F9471B45E460B12D0C6EB161442516B" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distributional map of Nesomyrmex species from the tropical dry forest in Colombia." figureDoi="10.3897/zookeys.1232.141693.figure29" httpUri="https://binary.pensoft.net/fig/1282999">29</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5544BEFB1BD7667DC1F0DDDE02C7E3B5" type="reference_group">
<paragraph id="A21F64F314C9715E9834BD9C0738F647">
<treatmentCitationGroup id="9B07AD3111DA8270DBD3E8E04FE98D23">
<treatmentCitation id="7CC591DCB9F4AB061EDA923CC6B0BEC7" author="Mayr G" page="617" year="1887">
<taxonomicName id="645C5DA6E12BA12AFCF2A8B3673F1FF0" ID-CoL="3TY6K" authority="Mayr, 1887: 617" authorityName="Mayr" authorityPageNumber="617" authorityYear="1887" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="spininodis">
<emphasis id="956C8BE604BD1690427FE77DFF094918" italics="true">Leptothorax spininodis</emphasis>
<comment id="29C990D0F7512A6C7B7CDD417EB5CAA4">
<bibRefCitation id="59C1556C0E6AD2BF453FDBF910953C46" author="Mayr G" journalOrPublisher="Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien" pagination="511-632" refId="B41" refString="Mayr G (1887) Sudamerikanische Formiciden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 37: 511-632." title="Südamerikanische Formiciden." volume="37" year="1887">Mayr, 1887: 617</bibRefCitation>
.
<typeStatus id="5C0927A72F10415C4B49DF441586C483" type="lectotype">Lectotype</typeStatus>
worker:
<collectingCountry id="1B475891048DC4F323F7ECD6F8D1D84E" name="Brazil">Brazil</collectingCountry>
, Rio de Janeiro or
<collectingCountry id="BE6A91ED88368F0B981CB44CED108B1D" name="Chile">Chile</collectingCountry>
,
<collectingRegion id="0C72D0AA02899E9316C78E4728BBD516" country="Chile" name="Valparaiso">Valparaiso</collectingRegion>
. [
<named-content id="F7D5455F98A07A22C7F83FC512BC9886" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum, Vienna">
<collectionCode id="314AF77CA1323807B1E7DD87803D29DE" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
</named-content>
]. (AntWeb image examined,
<ext-link id="025A2021F43A1ACBC81447D39D3F8B86" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0919734" xlink_type="simple">
<specimenCode id="7FAFF6A5D12E7200FEEA47C5188121FE">CASENT 0919734</specimenCode>
</ext-link>
).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="101E44C15AD23835765D731F07CEDFE1">
Combination in
<taxonomicName id="92A447167F4B913BA459E9189999E0C3" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Goniothorax">
<emphasis id="D71FD7FB55806C274A79D21913C76CBB" italics="true">Leptothorax</emphasis>
(
<emphasis id="7FD00DD9A50D33BA010B5648CD1F2253" italics="true">Goniothorax</emphasis>
)
</taxonomicName>
:
<bibRefCitation id="DBF2E6A947A13343C4A205CD4C0E4C3B" author="Emery C" firstAuthor="Emery" journalOrPublisher="Bullettino della Società Entomologica Italiana" page="59" pagination="33-107" refId="B23" refString="Emery C (1896) Studi sulle formiche della fauna neotropica. XVII-XXV. Bullettino della Società Entomologica Italiana 28: 33107." title="Studi sulle formiche della fauna neotropica. XVII-XXV." volume="28" year="1896">Emery 1896: 59</bibRefCitation>
.
</paragraph>
<paragraph id="CB624B8D69E18142876E323EA1C9D08C">
<treatmentCitationGroup id="C42F25F06315706E9C1014D7AE73A1D5">
<treatmentCitation id="7E604B32B75F5BF7D6E830E392C6B8D5" author="Wheeler WM" page="160" year="1921">
<taxonomicName id="DE4F4F1FD45012348CD9CC12B820883B" ID-CoL="CM62W" authority="Wheeler, 1921: 160" authorityName="Wheeler" authorityPageNumber="160" authorityYear="1921" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="umbratilis" subGenus="Goniothorax">
<emphasis id="AEA71031F9CEC892C023CFCE332FBE8F" italics="true">Leptothorax</emphasis>
(
<emphasis id="31F60DFC4369C1082D1C8C90C0CFE288" italics="true">Goniothorax</emphasis>
)
<emphasis id="4669F20F7D92B5B7C652111EE1486621" italics="true">umbratilis</emphasis>
<comment id="A45730711231862FBE1A4286875EDD95">
<bibRefCitation id="5110DBF581AA01076693C8AF5B7E424C" DOI="10.5962/p.184627" author="Wheeler WM" journalOrPublisher="Zoologica (New York)" pagination="137-168" refId="B58" refString="Wheeler WM (1921) The Tachigalia ants. Zoologica (New York) 3: 137-168. https://doi.org/10.5962/p.184627" title="The Tachigalia ants." volume="3" year="1921">Wheeler, 1921: 160</bibRefCitation>
.
<typeStatus id="FAB3C4E970EB8BCFB44815D51EDFB777" type="syntype">Syntype</typeStatus>
(queen, worker):
<collectingCountry id="E20A5664794C7201765B09B13E67565B" name="Guyana">Guyana</collectingCountry>
, Bartica, Penal Settlement. [
<collectionCode id="FF88C7CB2C551CB5FEF620F8DCDB7F2B">MCZC</collectionCode>
,
<named-content id="C52C4C9A0CDE574089EFD9ADC24D7ADE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/sao-paulo-museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museo de Zoologia da Universidade de São Paulo">
<collectionCode id="49183271DE4B4BECD84ECDDB2E8E62F6" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" type="Museum">MZSP</collectionCode>
</named-content>
]. (Image examined,
<collectionCode id="9350586DBD90A185877D70DA0A6CD1CC" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" type="Museum">MZSP</collectionCode>
78366). Synonymy by
<treatmentCitation id="0BE8299599C7E2352A5E171F2A27FC35" author="Kempf WW" page="427" year="1959">
<bibRefCitation id="BEB81F69CCE91C425CC2D76CFA4AA14D" author="Kempf WW" firstAuthor="Kempf" journalOrPublisher="Studia Entomologica" page="427" pagination="391-432" refId="B37" refString="Kempf WW (1959) A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae). Studia Entomologica 2: 391432." title="A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae)." volume="2" year="1959">Kempf 1959: 427</bibRefCitation>
</treatmentCitation>
.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="CE16E4D204C0ECEA4B9020107A08D71D">
<treatmentCitationGroup id="488411D96FF6D63CF3D9CFDB6ED9E0E7">
<treatmentCitation id="8DB52549FC4744BB9B0AFA76145FFE18" author="Santschi F" page="252" year="1922">
<taxonomicName id="F1A70D9D2D270DBB5260A1039286372E" ID-CoL="CM5S4" authority="Santschi, 1922: 252" authorityName="Santschi" authorityPageNumber="252" authorityYear="1922" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="genualia" subGenus="Goniothorax">
<emphasis id="B18D9E85CB8F049446F4FB0E59BA9C54" italics="true">Leptothorax</emphasis>
(
<emphasis id="5C0B7AAAFB31E47639110A5E0D0004A4" italics="true">Goniothorax</emphasis>
)
<emphasis id="A50D0028D4E8BE18EC5795D109AA4E9E" italics="true">genualia</emphasis>
<comment id="AD77E1259C762DE703F1D3BA338D299F">
<bibRefCitation id="857C21544C634841187983826D18D576" author="Santschi F" journalOrPublisher="Anales de la Sociedad Cientifica Argentina" pagination="241-262" refId="B49" refString="Santschi F (1922) Description de nouvelles fourmis de l'Argentine et pays limitrophes. Anales de la Sociedad Cientifica Argentina 94: 241-262." title="Description de nouvelles fourmis de lArgentine et pays limitrophes." volume="94" year="1922">Santschi, 1922: 252</bibRefCitation>
.
<typeStatus id="64D30DE9E58613C4F416C16BD8D64133" type="holotype">Holotype</typeStatus>
(worker):
<collectingCountry id="FFE732084BEE8E24B18BC39E38BB60CD" name="Paraguay">Paraguay</collectingCountry>
,
<collectingRegion id="A3D573D36C4BAB2E04CF37AA3272EFC0" country="Paraguay" name="Asuncion">Asuncion</collectingRegion>
. [
<collectionCode id="7E943E7DD5F7D83CD85095C3B79AB1DB" country="0" name="Natural History Museum Bucharest">NHMB</collectionCode>
]. (AntWeb image examined,
<ext-link id="0CD85D31984F9B159514644C7F157374" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0913001" xlink_type="simple">
<specimenCode id="D7152012D00B93543479FD8FBD557DD4">CASENT 0913001</specimenCode>
</ext-link>
). Synonymy by
<treatmentCitation id="3E881765000AA5D174F4B202AB496F6E" author="Kempf WW" page="427" year="1959">
<bibRefCitation id="D68F220546A90942E80DE33D7F9429CB" author="Kempf WW" firstAuthor="Kempf" journalOrPublisher="Studia Entomologica" page="427" pagination="391-432" refId="B37" refString="Kempf WW (1959) A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae). Studia Entomologica 2: 391432." title="A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae)." volume="2" year="1959">Kempf 1959: 427</bibRefCitation>
</treatmentCitation>
.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="D8E46D3CFEE7A7F04C1DFD5C80966FCB">
Combination in
<taxonomicName id="F6532E006BC8E1181C6DC045BE6F8EEE" authorityName="Wheeler" authorityYear="1910" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Nesomyrmex">
<emphasis id="D6F29DC28ECE5F45AC528A45D440C194" italics="true">Leptothorax</emphasis>
(
<emphasis id="C04492470FA70374C0A5D82D5CDFDF45" italics="true">Nesomyrmex</emphasis>
)
</taxonomicName>
:
<bibRefCitation id="B6735DDA080178735788E635CD9C2698" author="Kempf WW" firstAuthor="Kempf" journalOrPublisher="Studia Entomologica" page="427" pagination="391-432" refId="B37" refString="Kempf WW (1959) A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae). Studia Entomologica 2: 391432." title="A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae)." volume="2" year="1959">Kempf 1959: 427</bibRefCitation>
</paragraph>
<paragraph id="C50215423D7C3004E8F40A2805939311">
Combination in
<taxonomicName id="2D8441FFEC823A5DB19446825D657A27" ID-CoL="CM5NR" authorityName="Wheeler" authorityYear="1910" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="0E27B8F5AEBD085A06098D2AFDB2F914" italics="true">Nesomyrmex</emphasis>
</taxonomicName>
:
<bibRefCitation id="75CE3AF6E1359B7F963A3DFABD5778D4" author="Bolton B" firstAuthor="Bolton" journalOrPublisher="Memoirs of the American Entomological Society" page="272" pagination="1-370" refId="B5" refString="Bolton B (2003) Synopsis and classification of Formicidae. Memoirs of the American Entomological Society 71: 1370." title="Synopsis and classification of Formicidae." volume="71" year="2003">Bolton 2003: 272</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMRBI" type="description">
<paragraph id="75CAE6860AF269B258AD377E81F55711">
<heading id="97C1B45D410BF2242A0D09BC0996E712" reason="title">Worker measurements.</heading>
</paragraph>
<paragraph id="F38399FC9C34FA2AB0C557B2E7A15411">
(
<emphasis id="A5C35689BFDE61A15BB97D8802379CAC" italics="true">n</emphasis>
= 1)
<abbrev id="ABBRID0EURBI" xlink_title="Head length">HL</abbrev>
0.68,
<abbrev id="ABBRID0EYRBI" xlink_title="Head width">HW</abbrev>
0.62,
<abbrev id="ABBRID0E3RBI" xlink_title="Scape length">SL</abbrev>
0.48, ML 0.29,
<abbrev id="ABBRID0EASBI" xlink_title="Eye length">EL</abbrev>
0.15,
<abbrev id="ABBRID0EESBI" xlink_title="Pronotal width">PW</abbrev>
0.44,
<abbrev id="ABBRID0EISBI" xlink_title="Petiole width">PTW</abbrev>
0.21,
<abbrev id="ABBRID0EMSBI" xlink_title="Postpetiole width">PPW</abbrev>
0.28,
<abbrev id="ABBRID0EQSBI" xlink_title="Webers length">WL</abbrev>
0.82,
<abbrev id="ABBRID0EUSBI" xlink_title="Pronotal height">PH</abbrev>
0.25,
<abbrev id="ABBRID0EYSBI" xlink_title="Petiole length">PTL</abbrev>
0.32,
<abbrev id="ABBRID0E3SBI" xlink_title="Petiole height">PTH</abbrev>
0.22,
<abbrev id="ABBRID0EATBI" xlink_title="Postpetiole length">PPL</abbrev>
0.22,
<abbrev id="ABBRID0EETBI" xlink_title="Postpetiole height">PPH</abbrev>
0.22,
<abbrev id="ABBRID0EITBI" xlink_title="Gaster length">GL</abbrev>
0.79,
<abbrev id="ABBRID0EMTBI" xlink_title="Cephalic index">CI</abbrev>
91,
<abbrev id="ABBRID0EQTBI" xlink_title="Scape index">SI</abbrev>
77.
</paragraph>
<caption id="74FAA92A0BFF95B611E7A9F26A5E811A" ID-DOI="10.3897/zookeys.1232.141693.figure26" ID-arpha="0C88224B-1C52-5788-ABAE-7E86CFDDE82A" httpUri="https://binary.pensoft.net/fig/1282996" startId="F26">
<paragraph id="4A6557AB3585CC955165C2578278F12F">
<label id="45C444749A9E19F7463C397B76B68372">Figure 26.</label>
</paragraph>
<paragraph id="2D4341E1F9D04F059D529A2EB540ED47">
<taxonomicName id="274BA01622BE9DB43CB814C964427A0A" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="spininodis">
<emphasis id="0514D63DF3AB4B4F4F8AF7C004638A6B" italics="true">Nesomyrmex spininodis</emphasis>
</taxonomicName>
worker (
<specimenCode id="84204C26D97B2D40B7407464C0711B3B">
<abbrev id="ABBRID0EHUBI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">IAvH</abbrev>
-E-251254
</specimenCode>
)
<emphasis id="92C282A60EDB454C4D3C8431E176E5D6" bold="true">A</emphasis>
full-face view
<emphasis id="AF5033FCFB5F158CC9743431777C5923" bold="true">B</emphasis>
lateral view
<emphasis id="C213A81B3613357BBFE2BA4D1784AB22" bold="true">C</emphasis>
dorsal view. Scale bars: 0.2 mm (
<emphasis id="CA1F87F7407B4DFCD720DE6F1B32B386" bold="true">A, B</emphasis>
); 0.5 mm (
<emphasis id="CBE4B76A565627990E93B907F6BF4A73" bold="true">C</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E1UBI" type="Geographic range">
<paragraph id="59D2C73016E7BC6F3910DECDCD38B8A7">
<heading id="AD44149AE45BFCE8DB5D437F8497004A" reason="title">Geographic range.</heading>
</paragraph>
<paragraph id="1C35C3B6979E685F71F2FD19B71D2CDD">
<collectingCountry id="988ADA6AEEB0449A5AE4C40554E3B838" name="Argentina">Argentina</collectingCountry>
,
<collectingCountry id="B588FC3331B8DC05B6CE728B6AEB9527" name="Bolivia">Bolivia</collectingCountry>
,
<collectingCountry id="D1E395BD5B1237285E1B86461E56C2FB" name="Brazil">Brazil</collectingCountry>
,
<collectingCountry id="C0EBDE402CF2B52A71B0850D7C5ABE7A" name="Colombia">Colombia</collectingCountry>
,
<collectingCountry id="DE9AC1E6BBB853BECBA14E455AF97B73" name="Costa Rica">Costa Rica</collectingCountry>
,
<collectingCountry id="690528A94B2D60E6A271CF4AF03273A9" name="Ecuador">Ecuador</collectingCountry>
,
<collectingCountry id="16F9FDE23F2AC18A461CF04F739C1847" name="French Guiana">French Guiana</collectingCountry>
,
<collectingCountry id="BF8ACF728B45D4F971670CD80A5B028D" name="Paraguay">Paraguay</collectingCountry>
,
<collectingCountry id="A1CCEB084AEFEB1632E59CE0FBFA209A" name="Suriname">Suriname</collectingCountry>
,
<collectingCountry id="B7E6F2D7E09AA53B7D2E8B93630CD9B1" name="Venezuela">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6UBI" type="material">
<paragraph id="98CBD9842E6D108F6120D85A2D463D99">
<heading id="9348AADC141CEB81C16BF0BCE3862BA2" reason="title">Examined material.</heading>
</paragraph>
<paragraph id="82F76D23CAFCF90169E8385B6740251D">
<materialsCitation id="35AE580D9371246BA3914B6D39164568" collectingDate="1995-04-12" collectingMethod="Manual coll." collectionCode="IAvH" collectorName="J. Munoz" country="Colombia" elevation="100" latitude="5.350278" location="Parque Nacional Natural Tuparro" longLatPrecision="1" longitude="-67.87666" municipality="Cumaribo" specimenCode="IAvH-E-251254" specimenCount="1" specimenCount-worker="1" stateProvince="Vichada">
<collectingCountry id="01712A035FAA8A7CD9B27CED9FB492AF" name="Colombia">Colombia</collectingCountry>
<specimenCount id="BE8C1203EA9CB27033A4E02D8E577949" count="1" type="worker">1 worker</specimenCount>
;
<collectingRegion id="9E41E9884FEC3479879693345FD224EF" country="Colombia" name="Vichada">Vichada</collectingRegion>
,
<collectingMunicipality id="4EA0CE6A61E7BDCB506EB812AC59ADEE">Cumaribo</collectingMunicipality>
,
<location id="02EE313C37544BC9F99039807E649792" LSID="urn:lsid:plazi:treatment:9409219A903D5F39BBB82133FBBCAC1C:02EE313C37544BC9F99039807E649792" country="Colombia" latitude="5.350278" longLatPrecision="1" longitude="-67.87666" municipality="Cumaribo" name="Parque Nacional Natural Tuparro" stateProvince="Vichada">Parque Nacional Natural Tuparro</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIVBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-67.876667,5.350278]}">
<geoCoordinate id="CAC87CB356A55AA75C5146604E543010" degrees="5.350278" direction="north" orientation="latitude" precision="1" value="5.350278">5.350278°N</geoCoordinate>
,
<geoCoordinate id="B57D7CC9C8199C016FABE13204DF404A" degrees="67.876667" direction="west" orientation="longitude" precision="1" value="-67.87666">67.876667°W</geoCoordinate>
</named-content>
;
<elevation id="91EDF64788702CFF58112EF199D02BA9" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<quantity id="0BF3748E564B141A2646932A4E827ABF" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</quantity>
a. s. l.
</elevation>
;
<date id="E982719125296835C2761A070DB36DE7" value="1995-04-12">
<collectingDate id="59A7F63551CEA54819FB816D161B6C09" value="1995-04-12">12 Apr. 1995</collectingDate>
</date>
;
<collectorName id="829CA5209DC86EF1813A2FD422E324BC">J. Muñoz</collectorName>
leg.;
<collectingMethod id="66B6E38DE64ABC6C71B98BC8CCBDC066">Manual coll.</collectingMethod>
;
<specimenCode id="5C527DA7B56EE7C1BC572832E55CAB4B">
<abbrev id="ABBRID0ENVBI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">IAvH</abbrev>
-E-251254
</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERVBI" type="natural history">
<paragraph id="45B7021C33C700F317C66462373C6F82">
<heading id="CD17DB12C386A887B6F6ADBB0BE3D4D6" reason="title">Natural history.</heading>
</paragraph>
<paragraph id="7B2C91F09BFBE5F7B5BA4CC71CB28880">Species collected by manual sampling on vegetation in Parque Nacional Natural Tuparro.</paragraph>
</subSubSection>
<subSubSection id="SECID0EWVBI" type="comments">
<paragraph id="383FA31AFAFF36BC534517FD529E5BB4">
<heading id="765A67C5C96671DC9EF6371263E14F9A" reason="title">Comments.</heading>
</paragraph>
<paragraph id="AE1DF6D4F2B010986A037FF37EB3DC95">This species can be recognized by 11 antennomeres, uniformly testaceous or yellowish-brown coloration, dorsal surface of head finely reticulate-punctate and longitudinally rugose, without presence of shiny areas, and basal third of first gastral tergite generally aciculate-striate in its sculpture.</paragraph>
<paragraph id="0060516C317AFAD96BF18129D6444D2A">
<bibRefCitation id="6D95AED6AF11E04CCEFC1C566C4AD0EE" author="Kempf WW" firstAuthor="Kempf" journalOrPublisher="Studia Entomologica" pagination="391-432" refId="B37" refString="Kempf WW (1959) A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae). Studia Entomologica 2: 391432." title="A synopsis of the New World species belonging to the Nesomyrmex - group of the ant genus Leptothorax Mayr (Hymenoptera: Formicidae)." volume="2" year="1959">Kempf (1959)</bibRefCitation>
synonymized all previously described species and subspecies within the
<emphasis id="85A9B36CD450A0862F6D54A0FF4CA51C" italics="true">echinatinodis</emphasis>
complex under the nominal species, except for
<taxonomicName id="3B83F076EDE5EDC0832D284F1FDAD65B" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="spininodis">
<emphasis id="0834460D6B0F5A462F4F4E2366D9F2DE" italics="true">N. spininodis</emphasis>
</taxonomicName>
. A potential revision of the
<taxonomicName id="9845960490CC6EC3C6A0658E6C4B3BF7" baseAuthorityName="Forel" baseAuthorityYear="1886" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="echinatinodis">
<emphasis id="F7B10F300021B47ACD45B9AE894EA6D3" italics="true">N. echinatinodis</emphasis>
</taxonomicName>
complex could either result in the synonymization of certain species or the division of the complex into multiple forms. It is common to find
<taxonomicName id="BE6C88D6BB8E8CC9D632A49635D09CCD" baseAuthorityName="Forel" baseAuthorityYear="1886" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="echinatinodis">
<emphasis id="4C01FD9C35F8E4400B7A0CB736784E0A" italics="true">N. echinatinodis</emphasis>
</taxonomicName>
exhibiting sculpture similar to that of
<taxonomicName id="D62CD4191931F810F2E5F482EE84B629" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="spininodis">
<emphasis id="98357A5D8BD789DF15AFDAC49194CA08" italics="true">N. spininodis</emphasis>
</taxonomicName>
, particularly in populations from
<collectingCountry id="B321DFC83A10A90F0C2FC122457AAC31" name="Colombia">Colombia</collectingCountry>
different from those living in the
<abbrev id="ABBRID0EXXBI" xlink_title="tropical dry forest">TDF</abbrev>
(Unpublished data). Additionally, coloration is highly variable within the
<emphasis id="B13E6F36CDA3F1FBEC351A1CA9F187DE" italics="true">echinatinodis</emphasis>
complex. For this reason, using coloration as the primary criterion for distinguishing between different forms is not recommended. Thus, the separation of both species should not rely solely on coloration; instead, the sculpture of the frontal area of the head and the first gastral tergum should also be compared.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,277 @@
<document id="9F31695F64DF7387C0F7DD0371EC72D5" ID-DOI="10.3897/zookeys.1232.141693" ID-ZooBank="0AA099A2-EA57-4E27-937A-0976B9F8A4B5" ID-publisher-id="141693" URI-arpha="A9904ED3-9BDA-52DF-B936-1C42EC2713C5" URI-zoobank="http://zoobank.org/0AA099A2-EA57-4E27-937A-0976B9F8A4B5" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1742225059022" checkinUser="pensoft" docAuthor="Arredondo, Brandon S. &amp; Guerrero, Roberto J." docDate="2025" docId="99D09D34BB6C56399A14680FE250C1B0" docLanguage="en" docName="ZooKeys 1232: 131-172" docOrigin="ZooKeys 1232" docSource="https://zookeys.pensoft.net/article/141693/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nesomyrmex pittieri" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="172" masterDocId="A9904ED39BDA52DFB9361C42EC2713C5" masterDocTitle="The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data" masterLastPageNumber="172" masterPageNumber="131" pageNumber="131" updateTime="1742234045672" updateUser="admin">
<mods:mods id="3BA3AF844CE35075384E81BF4F432D25" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D5434F60A7332E2EF8B78928CE7F3135">
<mods:title id="58221ACD1C5078EDC64D7EE7657463B8">The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data</mods:title>
</mods:titleInfo>
<mods:name id="8BFB86519F3992A830298C7D9D9A2CC2" type="personal">
<mods:role id="E480E5BDBFD3D77DBBCF79A7F35F8F2C">
<mods:roleTerm id="CA99A8B73EAA3FED702EEB8AAF3E1EBB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D8AC8988542491DD90D57719F24B853E">Arredondo, Brandon S.</mods:namePart>
<mods:nameIdentifier id="FB25E697868561C4055D8545ED345B0D" type="ORCID">0000-0003-2777-5909</mods:nameIdentifier>
<mods:affiliation id="85DF4D703B613D91BC1A79E75DF573A1">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:name id="1621943C6A4A3B12528D0887FDAD633C" type="personal">
<mods:role id="884BBD04F7DFCC01713824537D5AEEC5">
<mods:roleTerm id="5F72442E499329F493CA82F70D744B56">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BC796A0493C6E4D169B93C3031948F2B">Guerrero, Roberto J.</mods:namePart>
<mods:nameIdentifier id="69757D403D021E9596CBAA46CEB979E3" type="ORCID">0000-0003-3244-2754</mods:nameIdentifier>
<mods:affiliation id="8D5C2AE27289C77A5E2351B8F7CBC63F">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="53535DCBB37ABDD2ADD040DFF96B8BE1">text</mods:typeOfResource>
<mods:relatedItem id="301E46F0A9300D2E13F30A458B3BD3AB" type="host">
<mods:titleInfo id="7781BD0A1CB9A89961FC36F0DA2665BC">
<mods:title id="A298412D3A72E3FD2592CFF0544F0438">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="FF749B9A9E36A10FAA2368DCBCBABA4E">
<mods:date id="818432B8D3A59C3AE937E71811CFA32F">2025</mods:date>
<mods:detail id="1190F41C0F66867DAB011C103F4390E4" type="pubDate">
<mods:number id="0B61E5E5E786A12C1BC02AC4691E473E">2025-03-17</mods:number>
</mods:detail>
<mods:detail id="8E433CFFBA4160FF45F59B3FDAD09345" type="volume">
<mods:number id="E50C22F743D235FA3483F0F24A9663E3">1232</mods:number>
</mods:detail>
<mods:extent id="91572C6F9D2CF5F4FB2892FA2EA92CBC" unit="page">
<mods:start id="750CDF7D7EAF8FE3870AD7C456ED4859">131</mods:start>
<mods:end id="2C7A8F5E2B2D9E33E4230FB2DB1EC32B">172</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="EA904158D8EF26B697D07C94B2B4F508">journal article</mods:classification>
<mods:identifier id="0FF944BDC774570DCBE33E23F8F3931B" type="DOI">10.3897/zookeys.1232.141693</mods:identifier>
<mods:identifier id="4529A65C7226548E48E05DE8F4EA707B" type="ZooBank">0AA099A2-EA57-4E27-937A-0976B9F8A4B5</mods:identifier>
</mods:mods>
<treatment id="99D09D34BB6C56399A14680FE250C1B0" ID-arpha="99D09D34-BB6C-5639-9A14-680FE250C1B0" LSID="urn:lsid:plazi:treatment:99D09D34BB6C56399A14680FE250C1B0" httpUri="http://treatment.plazi.org/id/99D09D34BB6C56399A14680FE250C1B0" scope_family="Formicidae" scope_subFamily="Myrmicinae">
<subSubSection id="664828BE92957C107D5DB8C37A3BAD8B" type="nomenclature">
<paragraph id="A48E0B850BD9DD1C51C1DAD09BFE8082">
<taxonomicName id="9FB411C1CFE2D076FCE440801C98B5E4" ID-CoL="CM7ZF" authority="(Forel, 1899)" baseAuthorityName="Forel" baseAuthorityYear="1899" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="pittieri">
<emphasis id="69D8AA2F6D99BDEDCC554AD1AFE2E9D4" italics="true">Nesomyrmex pittieri</emphasis>
(
<bibRefCitation id="5E0128811AC786C357973436C5DF0B0E" author="Forel A" journalOrPublisher="Biologia Centrali-Americana Hymenoptera" pagination="25-56" refId="B29" refString="Forel A (1899) Formicidae. [part]. Biologia Centrali-Americana Hymenoptera 3: 25-56." title="Formicidae. [part]." volume="3" year="1899">Forel, 1899</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="0B23547FFAAD29CCB37E2F2AC4DFC47D" type="description">
<paragraph id="868BD7FB3021971ADFA44896F9F2D061">
<figureCitation id="F0615ED7EB8E02F3222A3238EF6F9A54" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Nesomyrmex pittieri worker (CBUMAG: ENT: 41664) A full-face view B lateral view C dorsal view. Scale bars: 0.2 mm (A); 0.5 mm (B, C)." figureDoi="10.3897/zookeys.1232.141693.figure24" httpUri="https://binary.pensoft.net/fig/1282994">Figs 24</figureCitation>
,
<figureCitation id="151520E607745169584F6F63B68F3E6B" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distributional map of Nesomyrmex species from the tropical dry forest in Colombia." figureDoi="10.3897/zookeys.1232.141693.figure29" httpUri="https://binary.pensoft.net/fig/1282999">29</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="74F9E4902AE489AA81C9921FC41B3B47" type="reference_group">
<paragraph id="D7DBC3FAEB92B9A4056F910DD86A8230">
<treatmentCitationGroup id="9109E2C76C842AF65AA0EC834C707F3C">
<treatmentCitation id="E66CE430F5D13965E2779EA064C69314" author="Forel A" page="56" year="1899">
<taxonomicName id="EC8F77AA7DA939FEF34C6F0E08280F3A" ID-CoL="3TY4W" authority="Forel, 1899: 56" authorityName="Forel" authorityPageNumber="56" authorityYear="1899" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="pittieri">
<emphasis id="BEB47F4EF2E0F5CAF43E6F66EEFA3EC2" italics="true">Leptothorax pittieri</emphasis>
<comment id="A91E139FC6819913D8230B4B9B713BD5">
<bibRefCitation id="786114D9C0DE77C0BD36DC35F93E6943" author="Forel A" journalOrPublisher="Biologia Centrali-Americana Hymenoptera" pagination="25-56" refId="B29" refString="Forel A (1899) Formicidae. [part]. Biologia Centrali-Americana Hymenoptera 3: 25-56." title="Formicidae. [part]." volume="3" year="1899">Forel, 1899: 56</bibRefCitation>
.
<typeStatus id="DD7FBC46D5F6801973AC0312787B1EDD" type="holotype">Holotype</typeStatus>
(worker):
<collectingCountry id="17F190A275E8E288D4FA43DFB4CB7E7C" name="Costa Rica">Costa Rica</collectingCountry>
. [
<abbrev id="ABBRID0ER3AI" xlink_title="Muséum dHistoire Naturelle, Geneva">
<collectionCode id="577C0D807831E198F27A8B9F1694A6C2" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" type="Museum">MHNG</collectionCode>
</abbrev>
] (AntWeb image examined,
<ext-link id="F812D629DD3B2438F8768F80C14B87F3" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0908997" xlink_type="simple">
<specimenCode id="E581EA39599FC2AB9A7EAE1952259D81">CASENT 0908997</specimenCode>
</ext-link>
).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="C943177699E9760498159504B773F994">
Combination in
<taxonomicName id="7DBE036113A21496D74EF30F2D4A6CF8" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Goniothorax">
<emphasis id="7400E2C4910B857EDC3B7DC067E198D7" italics="true">Leptothorax</emphasis>
(
<emphasis id="2DAA2A3CD005E5C0D06213A9BC738A88" italics="true">Goniothorax</emphasis>
)
</taxonomicName>
: Emery 1924: 250.
</paragraph>
<paragraph id="5B993C8997B3926E86BDD36553375633">
Combination in
<taxonomicName id="F880E1084B0AB136C0D65FCEDF915D65" authorityName="Wheeler" authorityYear="1910" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Nesomyrmex">
<emphasis id="303825E15E67AC1C51C30A2F8D277DBA" italics="true">Leptothorax</emphasis>
(
<emphasis id="66F866883F5EACDFC63A888F8BB40AFD" italics="true">Nesomyrmex</emphasis>
)
</taxonomicName>
:
<bibRefCitation id="D00A439DF52BFBDBDAA721EC4CE8792D" author="Kempf WW" firstAuthor="Kempf" journalOrPublisher="Anais da Academia Brasileira de Ciencias" page="93" pagination="91-102" refId="B36" refString="Kempf WW (1958) Sôbre algumas formigas neotrópicas do gênero Leptothorax Mayr (Hymenoptera: Formicidae). Anais da Academia Brasileira de Ciencias 30: 91102." title="Sôbre algumas formigas neotrópicas do gênero Leptothorax Mayr (Hymenoptera: Formicidae)." volume="30" year="1958">Kempf 1958: 93</bibRefCitation>
.
</paragraph>
<paragraph id="52131011DCF6694C8455B13049571115">
Combination in
<taxonomicName id="050CA5088BF1F60BFAD0CC18CA6661A3" ID-CoL="CM5NR" authorityName="Wheeler" authorityYear="1910" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="90662CFD4EE866681F80F871F041CD41" italics="true">Nesomyrmex</emphasis>
</taxonomicName>
:
<bibRefCitation id="ED60D2630001CE2390426EFA93957064" author="Bolton B" firstAuthor="Bolton" journalOrPublisher="Memoirs of the American Entomological Society" page="272" pagination="1-370" refId="B5" refString="Bolton B (2003) Synopsis and classification of Formicidae. Memoirs of the American Entomological Society 71: 1370." title="Synopsis and classification of Formicidae." volume="71" year="2003">Bolton 2003: 272</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E35AI" type="description">
<paragraph id="C53C41DD45CB76C7FF8FCCCB6C8BBE23">
<heading id="532174206D8FA811F49D1400C2850A38" reason="title">Worker measurements.</heading>
</paragraph>
<paragraph id="B6A7E1E2DE4F33DEC5CA82AF31E56968">
(
<emphasis id="038874235A26442F1E0CAA241FE327B3" italics="true">n</emphasis>
= 1)
<abbrev id="ABBRID0EE6AI" xlink_title="Head length">HL</abbrev>
0.81,
<abbrev id="ABBRID0EI6AI" xlink_title="Head width">HW</abbrev>
0.68,
<abbrev id="ABBRID0EM6AI" xlink_title="Scape length">SL</abbrev>
0.53, ML 0.33,
<abbrev id="ABBRID0EQ6AI" xlink_title="Eye length">EL</abbrev>
0.21,
<abbrev id="ABBRID0EU6AI" xlink_title="Pronotal width">PW</abbrev>
0.51,
<abbrev id="ABBRID0EY6AI" xlink_title="Petiole width">PTW</abbrev>
0.24,
<abbrev id="ABBRID0E36AI" xlink_title="Postpetiole width">PPW</abbrev>
0.36,
<abbrev id="ABBRID0EAABI" xlink_title="Webers length">WL</abbrev>
0.98,
<abbrev id="ABBRID0EEABI" xlink_title="Pronotal height">PH</abbrev>
0.3,
<abbrev id="ABBRID0EIABI" xlink_title="Petiole length">PTL</abbrev>
0.35,
<abbrev id="ABBRID0EMABI" xlink_title="Petiole height">PTH</abbrev>
0.26,
<abbrev id="ABBRID0EQABI" xlink_title="Postpetiole length">PPL</abbrev>
0.24,
<abbrev id="ABBRID0EUABI" xlink_title="Postpetiole height">PPH</abbrev>
0.24,
<abbrev id="ABBRID0EYABI" xlink_title="Gaster length">GL</abbrev>
0.78.
<abbrev id="ABBRID0E3ABI" xlink_title="Cephalic index">CI</abbrev>
83,
<abbrev id="ABBRID0EABBI" xlink_title="Scape index">SI</abbrev>
78.
</paragraph>
<caption id="56A9DC5498A61803D2A9F411A95DCA6D" ID-DOI="10.3897/zookeys.1232.141693.figure24" ID-arpha="C4A17195-2C15-599F-BCE7-977F8F67A029" httpUri="https://binary.pensoft.net/fig/1282994" startId="F24">
<paragraph id="6AE50551A22F2BF17B059D56957ADF88">
<label id="EDEA8788A428AE7C27A36E205CA78D0F">Figure 24.</label>
</paragraph>
<paragraph id="BE99133E98B8A895D7315E09862B34E4">
<taxonomicName id="1091C3122C24670AD3BE1BE56FDC87BF" baseAuthorityName="Forel" baseAuthorityYear="1899" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="pittieri">
<emphasis id="F260BA84CCB7E36746348FEE3928ED9F" italics="true">Nesomyrmex pittieri</emphasis>
</taxonomicName>
worker (
<specimenCode id="EF351D4541813B5CEB757F95D726703C">
<named-content id="16063912182B59A696E7547ED5D143C6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:41664
</specimenCode>
)
<emphasis id="5F9CD3AAE1699D682526024CBF97C67B" bold="true">A</emphasis>
full-face view
<emphasis id="58FF45B5B72D541573BB2B831CE99FD2" bold="true">B</emphasis>
lateral view
<emphasis id="F03B289F6BFF961D4B72085C3159DE03" bold="true">C</emphasis>
dorsal view. Scale bars: 0.2 mm (
<emphasis id="6A1428F09165EC40263C72E083E98D99" bold="true">A</emphasis>
); 0.5 mm (
<emphasis id="E9B3542A17096DF729CD36B549A9BE32" bold="true">B, C</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EKCBI" type="Geographic range">
<paragraph id="51BA830BE999F06CAD1D1D3D9439A440">
<heading id="D6310A731A7C87B5E9E64C557F7455F5" reason="title">Geographic range.</heading>
</paragraph>
<paragraph id="34834E8CC994069F82520F37D6D84B3D">
<collectingCountry id="F45678E25D69BA85AF3E9AC5409FD9C6" name="Colombia">Colombia</collectingCountry>
,
<collectingCountry id="A3C63BAC42873CA4B8B42BB85781E536" name="Costa Rica">Costa Rica</collectingCountry>
,
<collectingCountry id="40E66EA415A602BA6793849FB01D6430" name="Guatemala">Guatemala</collectingCountry>
,
<collectingCountry id="E6E5C741AFBAB3726023F22EC931F684" name="Honduras">Honduras</collectingCountry>
,
<collectingCountry id="8DBC75ECFF4E5022B2824E7A5BF0EC68" name="Mexico">Mexico</collectingCountry>
,
<collectingCountry id="4A56A59D75A71A5ABD89907EE0E47BBD" name="Nicaragua">Nicaragua</collectingCountry>
,
<collectingCountry id="B4C32276BA46D7D6BFED90114C013D92" name="Panama">Panama</collectingCountry>
,
<collectingCountry id="6F226B0F1F23757F952A76B1D50B2293" name="Trinidad and Tobago">Trinidad and Tobago</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPCBI" type="material">
<paragraph id="75F1B4ECCBF4B1454E81D6FF211AA618">
<heading id="04A971F120955D50C53D1A38F0F5534D" reason="title">Examined material.</heading>
</paragraph>
<paragraph id="0C29C53717A654B8EEA1402BDDBABC93">
<materialsCitation id="FC1AF084E46DACC3BB52998D4B6DDF25" collectingDate="2021-07-30" collectionCode="CBUMAG" collectorName="M. A. Bautista-Giraldo &amp; I. Armbrecht" country="Colombia" county="Jamundi" elevation="979" latitude="3.33931" location="Colindres" longLatPrecision="5" longitude="-76.5413" municipality="Vda. San Isidro" specimenCode="CBUMAG:ENT:41664" specimenCount="1" specimenCount-worker="1" stateProvince="Valle del Cauca">
<collectingCountry id="2EF01427CAA8DD45EFC02281FDB03510" name="Colombia">Colombia</collectingCountry>
<specimenCount id="1B46FAB14E70BF2AF97DB75B6C352638" count="1" type="worker">1 worker</specimenCount>
;
<collectingRegion id="89D7B5281231287CB93807C5968FECB6" country="Colombia" name="Valle del Cauca">Valle del Cauca</collectingRegion>
,
<collectingCounty id="8F558E4E9BDA02695FE41BD97D7EA0BC">Jamundí</collectingCounty>
,
<collectingMunicipality id="EF1F2B778FD22CFE727FDDB81C1BE135">Vda. San Isidro</collectingMunicipality>
,
<location id="61172E648DA55FAC121DD6C296611A26" LSID="urn:lsid:plazi:treatment:99D09D34BB6C56399A14680FE250C1B0:61172E648DA55FAC121DD6C296611A26" country="Colombia" county="Jamundi" latitude="3.33931" longLatPrecision="5" longitude="-76.5413" municipality="Vda. San Isidro" name="Colindres" stateProvince="Valle del Cauca">Colindres</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYCBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-76.541300,3.339310]}">
<geoCoordinate id="D770D52053202685F61245DD5545A06C" degrees="3.33931" direction="north" orientation="latitude" precision="1" value="3.33931">3.33931°N</geoCoordinate>
,
<geoCoordinate id="A666496E806581A41404516EE3AD02DC" degrees="76.5413" direction="west" orientation="longitude" precision="5" value="-76.5413">76.5413°W</geoCoordinate>
</named-content>
;
<elevation id="9C3CA6DE99E9F2961C4E1E7A32B2983C" metricMagnitude="2" metricUnit="m" metricValue="9.79" unit="m" value="979.0">
<quantity id="A2F425AA28CC8F4F54D7ADD1778BE96C" metricMagnitude="2" metricUnit="m" metricValue="9.79" unit="m" value="979.0">979 m</quantity>
a. s. l.
</elevation>
;
<date id="D48CFA8ADBDB23A628825AABD9B87ABB" value="2021-07-30">
<collectingDate id="DB77C58CBD2AC24CD0C69B92437FBE97" value="2021-07-30">30 Jul. 2021</collectingDate>
</date>
;
<collectorName id="D6BC3E205A1FB3290DB010CC54278015">M. A. Bautista-Giraldo</collectorName>
,
<collectorName id="BA164628E1E7CA2271962108C43987AC">I. Armbrecht</collectorName>
legs.;
<specimenCode id="822F1F1EE0B3A411B689798214F118EE">
<named-content id="7E9BF259CF47C21D48558757DD7158D5" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:41664
</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBDBI" type="natural history">
<paragraph id="72BBB1EABD0C893EA249FEB017F14082">
<heading id="4CD68D5EBF6152359DD95C71C4297C83" reason="title">Natural history.</heading>
</paragraph>
<paragraph id="849B39C3E8268F836787C37B5FF8EB5B">Species collected by manual sampling on vegetation in Parque Nacional Natural Tuparro.</paragraph>
</subSubSection>
<subSubSection id="SECID0EGDBI" type="comments">
<paragraph id="34EDB94A44B54FE52ADB90C39574B1FF">
<heading id="564F3823F0CDCCACC82442F630112CB9" reason="title">Comments.</heading>
</paragraph>
<paragraph id="24CF429EDC13EF75E3C2520EEF98505A">This species is recognized by the following characters: wide and deep foveae on the dorsal surface of the head, antennal scapes curved at their base, and propodeal spines as long as the distance between their tips.</paragraph>
<paragraph id="702E5C8D3B34AAD0D58623CB795B5D85">
The only specimen examined shows variation in coloration with respect to specimens from other forest types in
<collectingCountry id="0CCDD42F442104DCE811A2C134C037F5" name="Colombia">Colombia</collectingCountry>
(unpublished data). Populations found in premontane forest show bicoloration, with variations between reddish brown to dark. The sculpture of the frontal area is consistent in all populations (Fig.
<figureCitation id="D14650C2AF26A5B47B11CC5AB2470FE1" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Nesomyrmex pittieri worker (CBUMAG: ENT: 41664) A full-face view B lateral view C dorsal view. Scale bars: 0.2 mm (A); 0.5 mm (B, C)." figureDoi="10.3897/zookeys.1232.141693.figure24" httpUri="https://binary.pensoft.net/fig/1282994">24</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,48 +1,48 @@
<document id="AE6B7302C2BC33CEE1A1102C4B99A91F" ID-DOI="10.3897/zookeys.1232.141693" ID-ZooBank="0AA099A2-EA57-4E27-937A-0976B9F8A4B5" ID-publisher-id="141693" URI-arpha="A9904ED3-9BDA-52DF-B936-1C42EC2713C5" URI-zoobank="http://zoobank.org/0AA099A2-EA57-4E27-937A-0976B9F8A4B5" article-type="research-article" checkinTime="1742225059022" checkinUser="pensoft" docAuthor="Arredondo, Brandon S. &amp; Guerrero, Roberto J." docDate="2025" docId="B5A743066B13534EB7B3D9F3B0DC4E69" docLanguage="en" docName="ZooKeys 1232: 131-172" docOrigin="ZooKeys 1232" docSource="https://zookeys.pensoft.net/article/141693/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nesomyrmex konina Arredondo &amp; Guerrero, 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="172" masterDocId="A9904ED39BDA52DFB9361C42EC2713C5" masterDocTitle="The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data" masterLastPageNumber="172" masterPageNumber="131" pageNumber="131" updateTime="1742225485493" updateUser="ExternalLinkService">
<mods:mods id="B82BF54A5E3969F2F0EDA0B63D080E0E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A0B9D01785B6A701AA0175159A4D69C1">
<mods:title id="8FF9F8DAF09C0D708DFF13B2BBA63C95">The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data</mods:title>
<document id="4BFAF7EA5AD9E5AA57D24FDFBE0DB68C" ID-DOI="10.3897/zookeys.1232.141693" ID-ZooBank="0AA099A2-EA57-4E27-937A-0976B9F8A4B5" ID-publisher-id="141693" URI-arpha="A9904ED3-9BDA-52DF-B936-1C42EC2713C5" URI-zoobank="http://zoobank.org/0AA099A2-EA57-4E27-937A-0976B9F8A4B5" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1742225059022" checkinUser="pensoft" docAuthor="Arredondo, Brandon S. &amp; Guerrero, Roberto J." docDate="2025" docId="B5A743066B13534EB7B3D9F3B0DC4E69" docLanguage="en" docName="ZooKeys 1232: 131-172" docOrigin="ZooKeys 1232" docSource="https://zookeys.pensoft.net/article/141693/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nesomyrmex konina Arredondo &amp; Guerrero, 2025, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="172" masterDocId="A9904ED39BDA52DFB9361C42EC2713C5" masterDocTitle="The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data" masterLastPageNumber="172" masterPageNumber="131" pageNumber="131" updateTime="1742234045672" updateUser="admin">
<mods:mods id="8AE7B212D91DCC98B3A37E14BE4A0FE5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="33333C596DAF2354DE578386152B4F8C">
<mods:title id="78D1AD62B5CC7DD3C6047CACE72A657B">The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data</mods:title>
</mods:titleInfo>
<mods:name id="7BB48988B3E67B08935CDE5866ED6D71" type="personal">
<mods:role id="C96E5DA4BDDC97E4DD6B0F767A41CC02">
<mods:roleTerm id="6C27C2DA7AED7C910DF9AB473BAAA898">Author</mods:roleTerm>
<mods:name id="99CF420CE0DEAAFEFE758EAFE0D58E77" type="personal">
<mods:role id="4691A4B682DDBF764FF3CACE90B01AD0">
<mods:roleTerm id="DD23878332741C687A47CC2B83C10EA4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5F120A6A54ADB355D896843CB66F34D1">Arredondo, Brandon S.</mods:namePart>
<mods:nameIdentifier id="B5936EE4055D08F20A813C5D2C17D47C" type="ORCID">0000-0003-2777-5909</mods:nameIdentifier>
<mods:affiliation id="7802EFFCEB97197EED63FF6FA4524270">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
<mods:namePart id="0EB8DCDB4A05E475E9AA8D4A9382DCE6">Arredondo, Brandon S.</mods:namePart>
<mods:nameIdentifier id="9D9D5FAD2BE3C10C66797CE7312AB291" type="ORCID">0000-0003-2777-5909</mods:nameIdentifier>
<mods:affiliation id="A1260B3E5B5B12C13BEE6C143F8E43B3">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:name id="2C127C71A2EFC614C6358C0D14417146" type="personal">
<mods:role id="36CD005027B4C87D350D04146AD95C21">
<mods:roleTerm id="3D7CB40CB5136A2065AAE6FE0D5B9007">Author</mods:roleTerm>
<mods:name id="24639875956EA777ED3889AD3A16D05B" type="personal">
<mods:role id="03A414F4D54AD967539C4196FC772377">
<mods:roleTerm id="A41C2E7ED9E68A66EB9780547C36A237">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4A10573A5E4FFBC353F88B16F4181265">Guerrero, Roberto J.</mods:namePart>
<mods:nameIdentifier id="BEB4D7E61CD50BF7E2EC64A6A4FF6DC8" type="ORCID">0000-0003-3244-2754</mods:nameIdentifier>
<mods:affiliation id="13C245CCAE4E8245276A6C75BEAF19EB">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
<mods:namePart id="8025DB635359C6EA2A1E8DF02A241E92">Guerrero, Roberto J.</mods:namePart>
<mods:nameIdentifier id="89F69F0157BDFCABEE676D457F5503E8" type="ORCID">0000-0003-3244-2754</mods:nameIdentifier>
<mods:affiliation id="15CBC79020C998EF87E86F116E4FA0E9">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="F232D6EE270FB5601418014FA4012896">text</mods:typeOfResource>
<mods:relatedItem id="7C3631578E714AFAA64AB2591D4C1CFE" type="host">
<mods:titleInfo id="7CB923FE33FB300ACF717A7871D71795">
<mods:title id="C5CA7713AD6C409499C1170791370C7E">ZooKeys</mods:title>
<mods:typeOfResource id="53DBA8C85C6B92D2537E72740E59C643">text</mods:typeOfResource>
<mods:relatedItem id="619005118C07DE20A5D21EBEDC3BADBD" type="host">
<mods:titleInfo id="A0DC8D55072E038B947B3CAE58DC21B1">
<mods:title id="53AE9DA237EF235C3279377BA373291D">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="3D4A8238970B464871DE4E9F61928F1A">
<mods:date id="F656C56640CACD8212DD2EBE2CFDD959">2025</mods:date>
<mods:detail id="3D45D77F6C20F0914CE28172757419C7" type="pubDate">
<mods:number id="AB429BD31FAEA134C220FE772C28BF87">2025-03-17</mods:number>
<mods:part id="70CE232B901377DCFFF8B859EA5910B8">
<mods:date id="46B6E66387E50525A0894150329E85C4">2025</mods:date>
<mods:detail id="C81E4A39B05C840E6630A68B03ED9B83" type="pubDate">
<mods:number id="4D5DEFB183C3BE58E2FF477EFB6D008E">2025-03-17</mods:number>
</mods:detail>
<mods:detail id="2DBFD6185F0E28CBC10B0559660F6FA8" type="volume">
<mods:number id="6D5CE28F46618FA4960400852D0A8B8D">1232</mods:number>
<mods:detail id="2CD59367293782697E1BF38D25A65D76" type="volume">
<mods:number id="CBFDE8ACD0980D22FC5E780D4494D267">1232</mods:number>
</mods:detail>
<mods:extent id="27088E6EA2795C5309CCEB70F9FDB457" unit="page">
<mods:start id="D958CAB39FB9A4204579D820B9A58381">131</mods:start>
<mods:end id="9F7339FC9304B5CCFFD4BADFFB2CC525">172</mods:end>
<mods:extent id="B1D62C51A5C648AB07FEEFC3D46FCDBE" unit="page">
<mods:start id="B2C8A7599B3F873E05DA0E49B6DF01C0">131</mods:start>
<mods:end id="8547601775827149F26DCBF1E98C13EB">172</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="320B68183152C060AB4B34CD0C9CDC96">journal article</mods:classification>
<mods:identifier id="CE3ED056C3D0B7B256D6909856C3A045" type="DOI">10.3897/zookeys.1232.141693</mods:identifier>
<mods:identifier id="3155B5D5079660D6178CA6AC3F3CE626" type="ZooBank">0AA099A2-EA57-4E27-937A-0976B9F8A4B5</mods:identifier>
<mods:classification id="EDAFC67291C7B676937209169257BC8A">journal article</mods:classification>
<mods:identifier id="37393E60DBEBB16E06C3F9E9F8A5718B" type="DOI">10.3897/zookeys.1232.141693</mods:identifier>
<mods:identifier id="DF0309CE48AD96669A046E35B87D63B4" type="ZooBank">0AA099A2-EA57-4E27-937A-0976B9F8A4B5</mods:identifier>
</mods:mods>
<treatment id="B5A743066B13534EB7B3D9F3B0DC4E69" ID-DOI="http://doi.org/10.5281/zenodo.15040783" ID-Zenodo-Dep="15040783" ID-arpha="B5A74306-6B13-534E-B7B3-D9F3B0DC4E69" ID-zoobank="https://zoobank.org/BD83D862-28F6-41A7-B3D2-AB18EBB2371E" LSID="urn:lsid:plazi:treatment:B5A743066B13534EB7B3D9F3B0DC4E69" LSID-ZBK="urn:lsid:zoobank.org:act:BD83D862-28F6-41A7-B3D2-AB18EBB2371E" httpUri="http://treatment.plazi.org/id/B5A743066B13534EB7B3D9F3B0DC4E69">
<treatment id="B5A743066B13534EB7B3D9F3B0DC4E69" ID-DOI="http://doi.org/10.5281/zenodo.15040783" ID-Zenodo-Dep="15040783" ID-arpha="B5A74306-6B13-534E-B7B3-D9F3B0DC4E69" ID-zoobank="https://zoobank.org/BD83D862-28F6-41A7-B3D2-AB18EBB2371E" LSID="urn:lsid:plazi:treatment:B5A743066B13534EB7B3D9F3B0DC4E69" LSID-ZBK="urn:lsid:zoobank.org:act:BD83D862-28F6-41A7-B3D2-AB18EBB2371E" httpUri="http://treatment.plazi.org/id/B5A743066B13534EB7B3D9F3B0DC4E69" scope_family="Formicidae" scope_subFamily="Myrmicinae">
<subSubSection id="E22C0E2EF76B7470FD0C86A960BD4EFA" type="nomenclature">
<paragraph id="A943DC8131822929324762B641E9EC21">
<taxonomicName id="19350FF8672254ACDB66348294F91F05" authorityName="Arredondo &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="konina" status="sp. nov.">
@ -63,7 +63,7 @@
<heading id="C4C5D18F530CB2C6ADCE9900409F05ED" reason="title">Type material.</heading>
</paragraph>
<paragraph id="78553512C9A204267F62F403C41F0795">
<materialsCitation id="F1E0F028EBD09A8327CEBF29387125A5" collectingDate="2024-03-19" collectingDateMax="2024-03-21" collectingDateMin="2024-03-19" collectionCode="CBUMAG" collectorName="H. Sierra &amp; S. de La Hoz &amp; Winkler" country="Colombia" elevation="675" latitude="6.359289" location="Parcela del Humboldt" longLatPrecision="1" longitude="-71.89426" municipality="Tame" specimenCount="1" specimenCount-worker="1" stateProvince="Arauca" typeStatus="holotype">
<materialsCitation id="F1E0F028EBD09A8327CEBF29387125A5" collectingDate="2024-03-19" collectingDateMax="2024-03-21" collectingDateMin="2024-03-19" collectingMethod="Winkler" collectionCode="CBUMAG" collectorName="H. Sierra &amp; S. de La Hoz" country="Colombia" elevation="675" latitude="6.359289" location="Parcela del Humboldt" longLatPrecision="1" longitude="-71.89426" municipality="Tame" specimenCode="CBUMAG:ENT:35951" specimenCount="1" specimenCount-worker="1" stateProvince="Arauca" typeStatus="holotype">
<emphasis id="105052953D7D77DC4F621310DE599EA5" bold="true">
<emphasis id="2E218A722E67C783460180E4F14408B4" bold="true" italics="true">
<typeStatus id="66A5CF5FB2C9B8EACB45EC3297B15FBE" type="holotype">Holotype</typeStatus>
@ -81,9 +81,9 @@
<location id="A7C1D5698C857C3FD6CE5F217800EA73" LSID="urn:lsid:plazi:treatment:B5A743066B13534EB7B3D9F3B0DC4E69:A7C1D5698C857C3FD6CE5F217800EA73" country="Colombia" latitude="6.359289" longLatPrecision="1" longitude="-71.89426" municipality="Tame" name="Parcela del Humboldt" stateProvince="Arauca">Parcela del Humboldt</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6PAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-71.894258,6.359289]}">
<geoCoordinate id="B1FF165341B671DCFD89D35644F822C4" degrees="6.359289" direction="north" orientation="latitude" precision="1" value="6.359289">6.359289 ° N</geoCoordinate>
<geoCoordinate id="B1FF165341B671DCFD89D35644F822C4" degrees="6.359289" direction="north" orientation="latitude" precision="1" value="6.359289">6.359289°N</geoCoordinate>
,
<geoCoordinate id="195FAF6CACAED26A1447077F45FD4DFE" degrees="71.894258" direction="west" orientation="longitude" precision="1" value="-71.89426">71.894258 ° W</geoCoordinate>
<geoCoordinate id="195FAF6CACAED26A1447077F45FD4DFE" degrees="71.894258" direction="west" orientation="longitude" precision="1" value="-71.89426">71.894258°W</geoCoordinate>
</named-content>
;
<elevation id="B6BEE26A470C10DD12956AFD099F65F1" metricMagnitude="2" metricUnit="m" metricValue="6.75" unit="m" value="675.0">
@ -103,19 +103,20 @@ a. s. l.
,
<collectorName id="BA160801A5D779B698F8E7ABC1F94933">S. de La Hoz</collectorName>
legs.;
<collectorName id="CD589CCE0C447D860814B1D5CC389295">Winkler</collectorName>
<collectingMethod id="5BF478B4E27509FC0C0CB184227E51A6">Winkler</collectingMethod>
;
<named-content id="75DA532F656CE1B64393C5B6B4BB14EA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="18D2338C451E60A61131A0BEF3125048">CBUMAG</collectionCode>
</named-content>
: ENT: 35951 [
<specimenCode id="4E76896C5B6CAC8011CCA32CE13FB4D5">
<named-content id="75DA532F656CE1B64393C5B6B4BB14EA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:35951
</specimenCode>
[
<named-content id="ECC141552B63FA6F4ACB9544CF236F54" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="441E6104748699B7B4BF235CC2353E39">CBUMAG</collectionCode>
</named-content>
]
</materialsCitation>
.
<materialsCitation id="FC2A520A29B975E979AD26A73840B38E" collectingDate="2024-03-19" collectingDateMax="2024-03-21" collectingDateMin="2024-03-19" collectionCode="CBUMAG" collectorName="H. Sierra &amp; S. de La Hoz &amp; Winkler" country="Colombia" elevation="675" latitude="6.359289" location="Parcela del Humboldt" longLatPrecision="1" longitude="-71.89426" municipality="Tame" specimenCount="3" specimenCount-worker="3" stateProvince="Arauca" typeStatus="paratype">
<materialsCitation id="FC2A520A29B975E979AD26A73840B38E" collectingDate="2024-03-19" collectingDateMax="2024-03-21" collectingDateMin="2024-03-19" collectingMethod="Winkler" collectionCode="CBUMAG" collectorName="H. Sierra &amp; S. de La Hoz" country="Colombia" elevation="675" latitude="6.359289" location="Colombia" longLatPrecision="1" longitude="-71.89426" municipality="Tame" specimenCode="CBUMAG:ENT:40035, CBUMAG:ENT:40036, CBUMAG:ENT:41654" specimenCount="3" specimenCount-worker="3" stateProvince="Arauca" typeStatus="paratype">
<emphasis id="775969AEBF92017A15219E46DB638668" bold="true" italics="true">
<typeStatus id="AE5ECBB588B6784B8BB44E4F93FB8042" type="paratype">Paratypes</typeStatus>
</emphasis>
@ -126,35 +127,40 @@ legs.;
<specimenCount id="CEA4CDFC2199EFBFB0646F89139DF2AF" count="3" type="worker">3 workers</specimenCount>
; same data as holotype;
<named-content id="FDF3FA1173AFF4C29281F73265F8D53B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="3FE1FC56033C781204A3415584878C18">CBUMAG</collectionCode>
</named-content>
: ENT: 40035,
<named-content id="72FF00058CDD2777BAF63D9A8BD52576" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="84B009256517EE6990D8863EE2D85C76">CBUMAG</collectionCode>
</named-content>
: ENT: 40036,
<named-content id="400549CA0FC1F7661F607C1FAE445EFC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="4CB39801065CDEB83F22E78B8D98EF3B">CBUMAG</collectionCode>
</named-content>
: ENT: 41654 (
<specimenCode id="60ADC5FF3C9A0AB8CAE6FF136935DED9">
<named-content id="FDF3FA1173AFF4C29281F73265F8D53B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:40035
</specimenCode>
,
<specimenCode id="C736276CE316AC1634A2D5CB4321F0BC">
<named-content id="72FF00058CDD2777BAF63D9A8BD52576" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:40036
</specimenCode>
,
<specimenCode id="125910C3F757A9DA7AF4E6D9A0BFDC97">
<named-content id="400549CA0FC1F7661F607C1FAE445EFC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:41654
</specimenCode>
(
<named-content id="88FC423F6F3B2F82B984218A7C256815" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="F94C903F6EC8D9FF30D6567AB787F2F5">CBUMAG</collectionCode>
</named-content>
)
</materialsCitation>
. •
<materialsCitation id="B9FF214A1EF05594D62DF5270852ABC1" collectingDate="2019-02-07" collectingDateMax="2019-02-21" collectingDateMin="2019-02-07" collectionCode="MPUJ" country="Colombia" elevation="1043" latitude="4.512722" location="Medina" longLatPrecision="1" longitude="-73.426834" specimenCode="ENT 0064660" specimenCount="1" specimenCount-worker="1" stateProvince="Cundinamarca" typeStatus="paratype">
<materialsCitation id="B9FF214A1EF05594D62DF5270852ABC1" collectingDate="2019-02-07" collectingDateMax="2019-02-21" collectingDateMin="2019-02-07" collectionCode="MPUJ" country="Colombia" elevation="1043" latitude="4.512722" location="Vda. Periquito" longLatPrecision="1" longitude="-73.426834" municipality="Medina" specimenCode="MPUJ_ENT 0064660" specimenCount="1" specimenCount-worker="1" stateProvince="Cundinamarca" typeStatus="paratype">
<specimenCount id="8D13968F73051BCBD134FDE6768098E5" count="1" type="worker">1 worker</specimenCount>
;
<collectingRegion id="D18773E0F34609EEA8A99B48A58C989D" country="Colombia" name="Cundinamarca">Cundinamarca</collectingRegion>
,
<location id="614C9D963EEA0AA9C553847838443ED5" LSID="urn:lsid:plazi:treatment:B5A743066B13534EB7B3D9F3B0DC4E69:614C9D963EEA0AA9C553847838443ED5" country="Colombia" latitude="4.512722" longLatPrecision="1" longitude="-73.426834" name="Medina" stateProvince="Cundinamarca">Medina</location>
, Vda. Periquito;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EERAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-73.426833,4.512722]}">
<geoCoordinate id="14A2ED36D2685C8E75818CFA697E6D68" degrees="4.512722" direction="north" orientation="latitude" precision="1" value="4.512722">4.512722 ° N</geoCoordinate>
<collectingMunicipality id="3CBD051185D64CDBC376AA399FC2DE90">Medina</collectingMunicipality>
,
<geoCoordinate id="1A50EDCC46723C45D953CCB333610774" degrees="73.426833" direction="west" orientation="longitude" precision="1" value="-73.426834">73.426833 ° W</geoCoordinate>
<location id="BC900E926DD7DFD608635DF57534DFDD" LSID="urn:lsid:plazi:treatment:B5A743066B13534EB7B3D9F3B0DC4E69:BC900E926DD7DFD608635DF57534DFDD" country="Colombia" latitude="4.512722" longLatPrecision="1" longitude="-73.426834" municipality="Medina" name="Vda. Periquito" stateProvince="Cundinamarca">Vda. Periquito</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EERAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-73.426833,4.512722]}">
<geoCoordinate id="14A2ED36D2685C8E75818CFA697E6D68" degrees="4.512722" direction="north" orientation="latitude" precision="1" value="4.512722">4.512722°N</geoCoordinate>
,
<geoCoordinate id="1A50EDCC46723C45D953CCB333610774" degrees="73.426833" direction="west" orientation="longitude" precision="1" value="-73.426834">73.426833°W</geoCoordinate>
</named-content>
;
<elevation id="29FF17FAD84CDA2CBBD4DEDE1F78E4FF" metricMagnitude="3" metricUnit="m" metricValue="1.043" unit="m" value="1043.0">
@ -166,9 +172,7 @@ a. s. l.
<collectingDate id="8F69A68164760F7889CDC121AB12FE02" value="2019-02-07" valueMax="2019-02-21" valueMin="2019-02-07">0721 Feb. 2019</collectingDate>
</date>
;
<collectionCode id="D3A3B166FFA1AFA0585CD07F471174AC">MPUJ</collectionCode>
_
<specimenCode id="5A429B64AAF68CB36AA8F300B96A6361" collectionCode="MPUJ">ENT 0064660</specimenCode>
<specimenCode id="57D571B991DDB4E227689C08B2BCD6BF">MPUJ_ENT 0064660</specimenCode>
(
<named-content id="3C7E755229FA8417471AB9589DC02D73" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/pontificia-universidad-javeriana" xlink_title="Colección de Artrópodos del Museo de Historia Natural de la Pontificia Universidad Javeriana">
<collectionCode id="F0AEF2174F625448B7D95A70693C1E5D">MPUJ</collectionCode>
@ -187,8 +191,11 @@ _
</taxonomicName>
<taxonomicNameLabel id="B7702ED31724646A9F0859A9A36DD619" rank="species">sp. nov.</taxonomicNameLabel>
Holotype worker (
<specimenCode id="D08274C52A15A03571F47F5107BC40DB">
<named-content id="638548FDD12580874D4D967CF1ACEB36" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
: ENT: 35951)
:ENT:35951
</specimenCode>
)
<emphasis id="CEF4DEE3D80DC83188DFD9ACCB07D2F4" bold="true">A</emphasis>
full-face view
<emphasis id="8E12632495E3A0D2A8689B86DD4D4C92" bold="true">B</emphasis>
@ -226,9 +233,7 @@ worker measurements.
0.25,
<abbrev id="ABBRID0EVTAI" xlink_title="Webers length">WL</abbrev>
0.76,
<abbrev id="ABBRID0EZTAI" xlink_title="Pronotal height">
<collectingCountry id="9E23C9B9472C2C0189AE8BE328A159C5" name="Philippines">PH</collectingCountry>
</abbrev>
<abbrev id="ABBRID0EZTAI" xlink_title="Pronotal height">PH</abbrev>
0.24,
<abbrev id="ABBRID0E4TAI" xlink_title="Petiole length">PTL</abbrev>
0.23,
@ -273,9 +278,7 @@ worker measurements.
0.24 0.25,
<abbrev id="ABBRID0E4VAI" xlink_title="Webers length">WL</abbrev>
0.750.77,
<abbrev id="ABBRID0EBWAI" xlink_title="Pronotal height">
<collectingCountry id="23D0188D28D3342F364D527085CF2E4B" name="Philippines">PH</collectingCountry>
</abbrev>
<abbrev id="ABBRID0EBWAI" xlink_title="Pronotal height">PH</abbrev>
0.210.26,
<abbrev id="ABBRID0EFWAI" xlink_title="Petiole length">PTL</abbrev>
0.210.24,
@ -354,9 +357,7 @@ Dorsal surface of body, except head, with erect flattened hairs mostly separated
<heading id="77C91318ECAEE9B28307826D5BFDF660" reason="title">Natural history.</heading>
</paragraph>
<paragraph id="BC9050E950563E29041B99ED7E9031F2">
Several of the
<typeStatus id="EB0B8F516B363D3A0A45B531D63A8446">type</typeStatus>
workers were extracted from the leaf litter in a fragment of tropical dry forest that has been recovering for more than 20 years in eastern
Several of the type workers were extracted from the leaf litter in a fragment of tropical dry forest that has been recovering for more than 20 years in eastern
<collectingCountry id="FBEE2D43DC4C6F2A849EA09B4188B86D" name="Colombia">Colombia</collectingCountry>
.
</paragraph>

View file

@ -0,0 +1,645 @@
<document id="65A5CBB2EBDEA32F8B9CD1B916998A55" ID-DOI="10.3897/zookeys.1232.141693" ID-ZooBank="0AA099A2-EA57-4E27-937A-0976B9F8A4B5" ID-publisher-id="141693" URI-arpha="A9904ED3-9BDA-52DF-B936-1C42EC2713C5" URI-zoobank="http://zoobank.org/0AA099A2-EA57-4E27-937A-0976B9F8A4B5" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1742225059022" checkinUser="pensoft" docAuthor="Arredondo, Brandon S. &amp; Guerrero, Roberto J." docDate="2025" docId="BC051E13C8215581A9840E21F9F60A48" docLanguage="en" docName="ZooKeys 1232: 131-172" docOrigin="ZooKeys 1232" docSource="https://zookeys.pensoft.net/article/141693/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nesomyrmex xerophilus Arredondo-H &amp; Guerrero 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="172" masterDocId="A9904ED39BDA52DFB9361C42EC2713C5" masterDocTitle="The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data" masterLastPageNumber="172" masterPageNumber="131" pageNumber="131" updateTime="1742234045672" updateUser="admin">
<mods:mods id="0B757BB72C46E8A395CF0D73036F44D0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B1E90AA14BEA22C913EE79FC0E4E3FD9">
<mods:title id="2599C99859BF9A7F9CD129142BDE423A">The ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) from the threatened Colombian tropical dry forest: three new species, a new synonymy, and new distributional data</mods:title>
</mods:titleInfo>
<mods:name id="DB51452E3C0F4E91CC57474AB847163A" type="personal">
<mods:role id="F87930ECCEF9F81137D4213A8CF12B85">
<mods:roleTerm id="5E90AF87A85C0D9142E9541128DAEEBC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CD61458DE8BC3DC7571E5143F2A90647">Arredondo, Brandon S.</mods:namePart>
<mods:nameIdentifier id="56A5FA8B1A079A795B7AB78E89082876" type="ORCID">0000-0003-2777-5909</mods:nameIdentifier>
<mods:affiliation id="2D2FEAE49E4F34B6CCE765AC6AF70B0B">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:name id="50DAD2EF99D63BB383389560FF0F00B4" type="personal">
<mods:role id="B7D1EA0394F8C7EA0201BFB425F73516">
<mods:roleTerm id="5CF6BA8E4BE75C312FAF858B11841C66">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0A6D8262433ECEFD707F59F40F3497BA">Guerrero, Roberto J.</mods:namePart>
<mods:nameIdentifier id="CEA10CD2C1C5BE046E4935D3520B838E" type="ORCID">0000-0003-3244-2754</mods:nameIdentifier>
<mods:affiliation id="C940E643C113E6EC49462ECB8D06E224">Universidad del Magdalena, Santa Marta, Colombia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="65023F513FD1BFBB893765865C9D97B6">text</mods:typeOfResource>
<mods:relatedItem id="223AD3B621C57238E99A61BC130ABAB0" type="host">
<mods:titleInfo id="CD8CAF7D7302ED7A06D89CA37863CE61">
<mods:title id="5D3EBB4374C719410597ACA074A0DAC9">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="B6FE8E8553FA56E588903F656728388F">
<mods:date id="4F21253DCA4D63F1ACA7D1D02489143E">2025</mods:date>
<mods:detail id="E99DBC55346962BF4E14DF3B305D2038" type="pubDate">
<mods:number id="C45C46ACD8BF459E8CE4141CCE95A92A">2025-03-17</mods:number>
</mods:detail>
<mods:detail id="199BA2B533CB195B6224EA12F4AC44D0" type="volume">
<mods:number id="3870F6E91FFBB43C1A72F5BFF8649F59">1232</mods:number>
</mods:detail>
<mods:extent id="68B8619898751EC5B5C8191B172A7FCD" unit="page">
<mods:start id="6EFB76D6A772A3CB7FCDBF1D963AF82D">131</mods:start>
<mods:end id="CC12DD84C9F7D5B9AEEF3FDB4914DA37">172</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C38BBABDB40FFF0A7C0C8E7EC0EDF610">journal article</mods:classification>
<mods:identifier id="CB2D8122E9D76456EF8C5CB845225C34" type="DOI">10.3897/zookeys.1232.141693</mods:identifier>
<mods:identifier id="451427D012FE76749410F198F9DED866" type="ZooBank">0AA099A2-EA57-4E27-937A-0976B9F8A4B5</mods:identifier>
</mods:mods>
<treatment id="BC051E13C8215581A9840E21F9F60A48" ID-arpha="BC051E13-C821-5581-A984-0E21F9F60A48" ID-zoobank="https://zoobank.org/93376D41-ADB1-4AE0-AD39-E550F07516C7" LSID="urn:lsid:plazi:treatment:BC051E13C8215581A9840E21F9F60A48" LSID-ZBK="urn:lsid:zoobank.org:act:93376D41-ADB1-4AE0-AD39-E550F07516C7" httpUri="http://treatment.plazi.org/id/BC051E13C8215581A9840E21F9F60A48" scope_family="Formicidae" scope_subFamily="Myrmicinae">
<subSubSection id="7722E2013932C2991ECB374D59C5E5F2" type="nomenclature">
<paragraph id="C9059F799B4DB070CA848A857BAA1C85">
<taxonomicName id="AAA337E4D05283FE4FFBD9408FF9423C" authority="Arredondo-H &amp; Guerrero" authorityName="Arredondo-H &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="xerophilus" status="sp. nov.">
<emphasis id="8F63126A58C8DFB2850D54BCA75FFA10" italics="true">Nesomyrmex xerophilus</emphasis>
Arredondo-H &amp; Guerrero
</taxonomicName>
<taxonomicNameLabel id="ADBF5CE9D41BB3D9050F754D4A052A9A" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="5742EE3738EAABBE59CE4E9F82508CD5" type="description">
<paragraph id="7FFA252ED0CDA9FDBCEDA4E75ACF3FA7">
<figureCitation id="1E6E14453D9B33F8CC308D8A7A6C8356" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Nesomyrmex xerophilus sp. nov. Holotype worker (CBUMAG: ENT 12440) A full-face view B lateral view C dorsal view. Scale bars: 0.2 mm (A); 0.5 mm (B, C)." figureDoi="10.3897/zookeys.1232.141693.figure28" httpUri="https://binary.pensoft.net/fig/1282998">Figs 28</figureCitation>
,
<figureCitation id="342B7801AD3E2D19DF8FAFB35CE0063D" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distributional map of Nesomyrmex species from the tropical dry forest in Colombia." figureDoi="10.3897/zookeys.1232.141693.figure29" httpUri="https://binary.pensoft.net/fig/1282999">29</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERHCI" type="type material">
<paragraph id="6EB7F07CAE4ADF60BAEE1B0DD90B59CA">
<heading id="674FEEB17C8B2BFF872A8894E130C816" reason="title">Type material.</heading>
</paragraph>
<paragraph id="123466620F00BE66CFBD5B5E55F71868">
<materialsCitation id="93558FFB0CD4AE7A41F25A807BA0FCB3" collectingDate="2019-11-02" collectingMethod="Manual capture" collectionCode="CBUMAG" collectorName="M. Escarraga" country="Colombia" county="Santa Marta" latitude="11.176972" location="Reserva la Iguana Verde" longLatPrecision="1" longitude="-74.185165" municipality="Vda. Puerto Mosquito" specimenCode="CBUMAG:ENT:12440" specimenCount="1" specimenCount-worker="1" stateProvince="Magdalena" typeStatus="holotype">
<emphasis id="E8B6319B8FB1BA3BE21250CB25E52BBF" bold="true">
<emphasis id="B4705EC34DA4EA8BFBA337A34ED03B1D" bold="true" italics="true">
<typeStatus id="7C33BE94583B78D2B4851FD78F9CA3C1" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="B02D097C92E140B2AF2CE151F4DDBD6B" name="Colombia">Colombia</collectingCountry>
<specimenCount id="299843256B71B717CE8870D06BE7A4E2" count="1" type="worker">1 worker</specimenCount>
;
<collectingRegion id="0298C85540A04564AABFF7AAAFAA535D" country="Colombia" name="Magdalena">Magdalena</collectingRegion>
,
<collectingCounty id="AB78AE0A4C69347DBCA1C889BEF18196">Santa Marta</collectingCounty>
,
<collectingMunicipality id="BFFA8F8DF442371614D5A087DE867665">Vda. Puerto Mosquito</collectingMunicipality>
,
<location id="03A4EE777BA3E7A3EB1908995FAB1A30" LSID="urn:lsid:plazi:treatment:BC051E13C8215581A9840E21F9F60A48:03A4EE777BA3E7A3EB1908995FAB1A30" country="Colombia" county="Santa Marta" latitude="11.176972" longLatPrecision="1" longitude="-74.185165" municipality="Vda. Puerto Mosquito" name="Reserva la Iguana Verde" stateProvince="Magdalena">Reserva la Iguana Verde</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6HCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-74.185167,11.176972]}">
<geoCoordinate id="068A29D66CFF80F0CD9376A5C82351DD" degrees="11.176972" direction="north" orientation="latitude" precision="1" value="11.176972">11.176972°N</geoCoordinate>
,
<geoCoordinate id="B08A5DA24EC4CDC8FCA25D6CB73E1C7B" degrees="74.185167" direction="west" orientation="longitude" precision="1" value="-74.185165">74.185167°W</geoCoordinate>
</named-content>
;
<date id="A551770BA12A8913DC59485410045656" value="2019-11-02">
<collectingDate id="8FD15934DBE41A319D097A863F361AC6" value="2019-11-02">02 Nov. 2019</collectingDate>
</date>
;
<collectorName id="6AE7707B5B48FB3E14C8DF7CEC344842">M. Escárraga</collectorName>
leg.;
<collectingMethod id="CCB606319E4935946CDDFB81AE1950EA">Manual capture</collectingMethod>
;
<specimenCode id="2C115A2759FD3D37DBB935E0813CE770">
<named-content id="4F79464A1342F9949FAD1F564311EB98" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:12440
</specimenCode>
[
<named-content id="0722A3F47F0FE45B51411D19D05A75A1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="10C7B8B691099FF80763E9FECCAF8563">CBUMAG</collectionCode>
</named-content>
]
</materialsCitation>
.
<materialsCitation id="4D0FD1ABD7F9CD32BACCED03913CDD32" collectingDate="2019-11-02" collectingMethod="Manual capture" collectionCode="ICN, CBUMAG, MEFLG" collectorName="M. Escarraga &amp; Manual" country="Colombia" county="Santa Marta" latitude="11.176972" location="Colombia" longLatPrecision="1" longitude="-74.185165" municipality="Santa Marta" specimenCode="CBUMAG:ENT:12441, CBUMAG:ENT:41656, CBUMAG:ENT:42097, CBUMAG:ENT:42098, CBUMAG:ENT:42099, CBUMAG:ENT:42100, CBUMAG:ENT:42101, CBUMAG:ENT:42102" specimenCount="8" specimenCount-worker="8" stateProvince="Magdalena" typeStatus="paratype">
<emphasis id="8F6DFAE7008883AF447E3305B09D2B0F" bold="true" italics="true">
<typeStatus id="0D5473B8225EBF6FFBAB073378EFC393" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<emphasis id="E2FCD12F3A49EE01AF859179D4F5927A" italics="true">n</emphasis>
= 14).
<collectingCountry id="23EFFEDA28A96E4F2967E87DAFE1F605" name="Colombia">Colombia</collectingCountry>
<specimenCount id="C038D3BFFE1DA33F5952429717AAD0F6" count="8" type="worker">8 workers</specimenCount>
; same data as holotype;
<specimenCode id="684917BE7AF8F558C2AB51A6BBC326A9">
<named-content id="E346AC551FFDD0067C0013F897616109" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:12441
</specimenCode>
(
<abbrev id="ABBRID0EVICI" xlink_title="Colección de Entomología del Instituto de Ciencias Naturales, Universidad Nacional de Colombia">
<collectionCode id="E691DABD18E839FD272FA43F5E1649E1" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" type="">ICN</collectionCode>
</abbrev>
),
<specimenCode id="2B9AAEA059F5AC0BAFBB567F5704AFE0">
<named-content id="6314DA1128FDDDBD9717F3BC2F50CF17" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:41656
</specimenCode>
(
<named-content id="0313751F35A075FF354652CDBE40D276" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="9A6D4BBB647C2C621F159803C56FB1C1">CBUMAG</collectionCode>
</named-content>
),
<specimenCode id="13BAAA2F2AE3F4669D2D8607CF0158A3">
<named-content id="F6823084FC8D56EF596B99902EC065B3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:42097
</specimenCode>
(
<named-content id="7C4AA364552E675FEAA606860F5D4939" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="0B9F12B5E764810FC911585DD72CFC9D">CBUMAG</collectionCode>
</named-content>
),
<specimenCode id="BED142AB428C8B93066D5ADD9688F982">
<named-content id="AD1C1ED5F11B889ACE1330DBA4CD9309" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:42098
</specimenCode>
(
<named-content id="E8324A32E8E07C971E89B0C3BC612CEA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="FC875941AC087CCD75B83B4DE32B7DAF">CBUMAG</collectionCode>
</named-content>
),
<specimenCode id="AE97C37592CC1BC98FAA04E0EFFF5A82">
<named-content id="226FE6E7DE2F179673872AE735BA986A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:42099
</specimenCode>
(
<named-content id="1CBE264AADE9B64006CCBC16604DE080" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="CF099CF78470FAEDFCD4308FAE149E19">CBUMAG</collectionCode>
</named-content>
),
<specimenCode id="63A2E7794E592837C52B9B32F39B7FA0">
<named-content id="ACBA62BA093CE3763BB16C0BB82EEAA6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:42100
</specimenCode>
(
<named-content id="8E3CF0A74D16C69DEB1FC22885628D56" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="3A368568B076053E94FF37C18FBC4667">CBUMAG</collectionCode>
</named-content>
),
<specimenCode id="7F53DA2AFEDADBB126D25ADE4EC2C28C">
<named-content id="2872FB607F4408A5691BF4F7D1A22DD4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:42101
</specimenCode>
(
<named-content id="B807E0929F43185233C63E03CDA2CB3D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="5ED295D742F9E1B895FB128274EF4C57">CBUMAG</collectionCode>
</named-content>
),
<specimenCode id="96FDF308D70670B4DA075E7974FEEBC2">
<named-content id="6603D4E67E1F1918387C5C3812750E38" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:42102
</specimenCode>
(
<named-content id="D26B45EF6A96FAEB6E608897098497EC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-entomologico-francisco-luis-gallego" xlink_title="Museo Entomológico Francisco Luis Gallego, Universidad Nacional de Colombia">
<collectionCode id="3B8BE399224EC92433E2741B80449917" country="Colombia" httpUri="http://grbio.org/cool/ct7t-ipa9" name="Museo Entomologico Francisco Luis Gallego" type="">MEFLG</collectionCode>
</named-content>
)
</materialsCitation>
. •
<materialsCitation id="511871E63C76CCBF793644E30938D9F1" collectingDate="2023-09-16" collectingMethod="Winkler" collectionCode="CBUMAG" collectorName="H. Sierra &amp; L. Ramos &amp; R. Guerrero &amp; S. De La Hoz" country="Colombia" county="Valledupar" elevation="584" latitude="10.572165" location="PNR Los Besotes" longLatPrecision="1" longitude="-73.27122" municipality="Cgto. Los Corazones" specimenCode="CBUMAG:ENT:41655" specimenCount="1" specimenCount-worker="1" stateProvince="Cesar" typeStatus="paratype">
<specimenCount id="5493450E99B5799C5BB6C624E0D63D93" count="1" type="worker">1 worker</specimenCount>
;
<collectingRegion id="E45E2781CA7EAD4311530F77E46DB73D" country="Colombia" name="Cesar">Cesar</collectingRegion>
,
<collectingCounty id="61AE7B064000E85950C70D649913926C">Valledupar</collectingCounty>
,
<collectingMunicipality id="E9DA5B4169F6A4F90677DB86061964B4">Cgto. Los Corazones</collectingMunicipality>
,
<location id="F042A2EA2C46DD098C780CF50F4B6C52" LSID="urn:lsid:plazi:treatment:BC051E13C8215581A9840E21F9F60A48:F042A2EA2C46DD098C780CF50F4B6C52" country="Colombia" county="Valledupar" latitude="10.572165" longLatPrecision="1" longitude="-73.27122" municipality="Cgto. Los Corazones" name="PNR Los Besotes" stateProvince="Cesar">PNR Los Besotes</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EUKCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-73.271218,10.572165]}">
<geoCoordinate id="2BECAA03C70B8028E93EE519E3378439" degrees="10.572165" direction="north" orientation="latitude" precision="1" value="10.572165">10.572165°N</geoCoordinate>
,
<geoCoordinate id="BA25AF604817CF9C49AE30B7DB9A0F7F" degrees="73.271218" direction="west" orientation="longitude" precision="1" value="-73.27122">73.271218°W</geoCoordinate>
</named-content>
;
<elevation id="F72AB77DA3B310ADF79AD38EB5A5936F" metricMagnitude="2" metricUnit="m" metricValue="5.84" unit="m" value="584.0">
<quantity id="17D769577615E0F449A3946EDDC268E0" metricMagnitude="2" metricUnit="m" metricValue="5.84" unit="m" value="584.0">584 m</quantity>
a. s. l.
</elevation>
; 14
<date id="56B256AC9A3607F3D46B87FF8FE56F28" value="2023-09-16">
<collectingDate id="DCCB6CB5D4A0EBC96134D42CD8AC532C" value="2023-09-16">16 Sep. 2023</collectingDate>
</date>
;
<collectorName id="045ABF1B2E870258993D0527B333A1CF">H. Sierra</collectorName>
,
<collectorName id="A21B0E98F0D6337800A5E0AC2AD2B9F1">L. Ramos</collectorName>
,
<collectorName id="C7B141DBBE77D8CA02E475DB89E4959E">R. Guerrero</collectorName>
y
<collectorName id="1E3DDA9E70FE6D12B13CD75F9E56C68F">S. De La Hoz</collectorName>
legs.;
<collectingMethod id="0F9EC42570254507AE8EDF8F33D79D39">Winkler</collectingMethod>
;
<specimenCode id="83FE34F5B92CCC0C712A517CEB738EDE">
<named-content id="17FBD92F63C3B7BC3DBEB356B8942849" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:41655
</specimenCode>
(
<named-content id="2491DEA4764D34EF00BB9BB9B52D15C3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="E2EC6695DDB85736D4DF27FBDE77B536">CBUMAG</collectionCode>
</named-content>
)
</materialsCitation>
. •
<materialsCitation id="779817C231692BA8D2E702BD20B3E77F" collectingDate="2018-11-27" collectionCode="CBUMAG" collectorName="M. Escarraga" country="Colombia" latitude="11.213139" location="Reserva Caoba" longLatPrecision="1" longitude="-74.10167" municipality="Santa Marta" specimenCode="CBUMAG:ENT:12164, CBUMAG:ENT:12165" specimenCount="2" specimenCount-worker="2" stateProvince="Magdalena" typeStatus="paratype">
<specimenCount id="5CDB3996C4A32E94F870F0ED186F1542" count="2" type="worker">2 workers</specimenCount>
;
<collectingRegion id="1A6B5571BC2431B1E214ECD763488974" country="Colombia" name="Magdalena">Magdalena</collectingRegion>
,
<collectingMunicipality id="1DFE769CD006636B6870122EA97E54C2">Santa Marta</collectingMunicipality>
,
<location id="912BA70A78734F3EF6E48DABEF5DD5AA" LSID="urn:lsid:plazi:treatment:BC051E13C8215581A9840E21F9F60A48:912BA70A78734F3EF6E48DABEF5DD5AA" country="Colombia" latitude="11.213139" longLatPrecision="1" longitude="-74.10167" municipality="Santa Marta" name="Reserva Caoba" stateProvince="Magdalena">Reserva Caoba</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EELCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-74.101667,11.213139]}">
<geoCoordinate id="7AAA2EC51743795DBD4F62EA43D54200" degrees="11.213139" direction="north" orientation="latitude" precision="1" value="11.213139">11.213139°N</geoCoordinate>
,
<geoCoordinate id="07F493EED6DAF2377F938289FD3B54F8" degrees="74.101667" direction="west" orientation="longitude" precision="1" value="-74.10167">74.101667°W</geoCoordinate>
</named-content>
;
<date id="1A8435121A787E5D073E09BABBBB9ECD" value="2018-11-27">
<collectingDate id="EC89034F91A8731C6004AEA934634561" value="2018-11-27">27 Nov. 2018</collectingDate>
</date>
;
<collectorName id="0774680E4DBAEC786A12CAA82B1BFC13">M. Escárraga</collectorName>
leg.;
<specimenCode id="D550F58F302278FB034FF35130262127">
<named-content id="DFEDAEA7CFAE53C2E384CEF6AA1E3911" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:12164
</specimenCode>
,
<specimenCode id="BF8BDEDB990220C31F4B407FF1453B96">
<named-content id="5F91D283CC892E3E31A7E89EFEE5F0D6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:12165
</specimenCode>
(
<named-content id="FE0DCB0D2D75983CDF95F20261BA5E68" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">
<collectionCode id="8862B6327964E4D257E6A44AF2E1ECAF">CBUMAG</collectionCode>
</named-content>
)
</materialsCitation>
. •
<materialsCitation id="642348D3255D959901252B4B97FDAB2C" collectingDate="2007-03-03" collectingMethod="Pitfall" collectionCode="IAvH" collectorName="D. Ramirez &amp; D. Olivero-G" country="Colombia" elevation="117" latitude="11.259111" location="Cerro Taganga" longLatPrecision="1" longitude="-74.18014" municipality="Santa Marta" specimenCode="CBUMAG:ENT:12163" specimenCount="1" specimenCount-worker="1" stateProvince="Magdalena" typeStatus="paratype">
<specimenCount id="ED0CE64B9FFD13F840B73A009D0002A3" count="1" type="worker">1 worker</specimenCount>
;
<collectingRegion id="283D118E50865516983D37AFF9724EBB" country="Colombia" name="Magdalena">Magdalena</collectingRegion>
,
<collectingMunicipality id="AC35A5CACD44EEE60A46347E9AF695E0">Santa Marta</collectingMunicipality>
,
<location id="12FDCE44750535F03E76C08C24E563F5" LSID="urn:lsid:plazi:treatment:BC051E13C8215581A9840E21F9F60A48:12FDCE44750535F03E76C08C24E563F5" country="Colombia" latitude="11.259111" longLatPrecision="1" longitude="-74.18014" municipality="Santa Marta" name="Cerro Taganga" stateProvince="Magdalena">Cerro Taganga</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYLCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-74.180139,11.259111]}">
<geoCoordinate id="9D9EEC3D628E38E4F8F1B7345FEDCD9A" degrees="11.259111" direction="north" orientation="latitude" precision="1" value="11.259111">11.259111°N</geoCoordinate>
,
<geoCoordinate id="A3820F96EB7B0F179B10586A9DF2C295" degrees="74.180139" direction="west" orientation="longitude" precision="1" value="-74.18014">74.180139°W</geoCoordinate>
</named-content>
;
<elevation id="8DF953E4E69E6F2CC8B0E5288EFF9DCE" metricMagnitude="2" metricUnit="m" metricValue="1.173" unit="m" value="117.3">
<quantity id="18E214061F4238BA8F8E7FBE771BF726" metricMagnitude="2" metricUnit="m" metricValue="1.173" unit="m" value="117.3">117.3 m</quantity>
a. s. l.
</elevation>
;
<date id="14D363667B6C8C9B100BD71A2C5A16E5" value="2007-03-03">
<collectingDate id="E4C249A7BF67BC70B4FDCFA8F6549B0A" value="2007-03-03">03 Mar. 2007</collectingDate>
</date>
;
<collectorName id="6BBD9A224588726FDA0C8624446ABD02">D. Ramírez</collectorName>
y
<collectorName id="7D6D725DCDD51CCCA211EEB4AE2B9BB6">D. Olivero-G</collectorName>
legs.;
<collectingMethod id="099E6906A2012F9F2B5925159E8BFA84">Pitfall</collectingMethod>
;
<specimenCode id="2C1DCAF0F3C64CD515D5FAB5C52EE827">
<named-content id="96182449A80BCE4AB4F8C7A5B99BB8AD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
:ENT:12163
</specimenCode>
(
<abbrev id="ABBRID0EBMCI" xlink_title="Colección de Entomología del Instituto de Investigaciones de recursos biológicos Alexander von Humboldt">
<collectionCode id="9E9CDF783A0DCFFAF92B725C8814CB49">IAvH</collectionCode>
</abbrev>
)
</materialsCitation>
. •
<materialsCitation id="0D6C0F68435AE600CFBD02AC35BE66FB" collectingDate="1985-08-16" collectionCode="ICN" collectorName="P. Ward" country="Colombia" county="Santa Marta" elevation="100" latitude="10.883306" location="Rio Frio" longLatPrecision="1" longitude="-74.13333" municipality="Cienaga" specimenCode="ICN 019704" specimenCount="2" specimenCount-worker="2" stateProvince="Magdalena" typeStatus="paratype">
<specimenCount id="876CBF28879BAAA1758019812031CB9A" count="2" type="worker">2 workers</specimenCount>
.
<collectingRegion id="51EDE8D15C3DB73CF22D63BD05256941" country="Colombia" name="Magdalena">Magdalena</collectingRegion>
,
<collectingCounty id="BEC3B311913BF6A0F8726C671789F53D">Santa Marta</collectingCounty>
,
<collectingMunicipality id="6E46F064482261656CA4FD90F79E783F">Ciénaga</collectingMunicipality>
,
<locationDeviation id="9A5F239F204354F0F733FC9288849F95">
<quantity id="AA75D09F6A72773238A89488F538E5BE" metricMagnitude="3" metricUnit="m" metricValue="5.0" unit="km" value="5.0">5 km</quantity>
SE
</locationDeviation>
,
<location id="E4E8DAD77C34DB9364DA6E0C22CB165F" LSID="urn:lsid:plazi:treatment:BC051E13C8215581A9840E21F9F60A48:E4E8DAD77C34DB9364DA6E0C22CB165F" country="Colombia" county="Santa Marta" latitude="10.883306" longLatPrecision="1" longitude="-74.13333" municipality="Cienaga" name="Rio Frio" stateProvince="Magdalena">Río Frío</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIMCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-74.133333,10.883306]}">
<geoCoordinate id="8226AA704D0CC364FDDD89843C8A8EFE" degrees="10.883306" direction="north" orientation="latitude" precision="1" value="10.883306">10.883306°N</geoCoordinate>
,
<geoCoordinate id="FB166ADFC8A9F503AC4CEDF414BAABA5" degrees="74.133333" direction="west" orientation="longitude" precision="1" value="-74.13333">74.133333W</geoCoordinate>
</named-content>
;
<elevation id="659B8B5D5B7645522B1BFD53C205C38C" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<quantity id="4C9C29AFEEF83E6D105B61BA6822CD77" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</quantity>
a. s. l.
</elevation>
;
<date id="2FF9A46FCFAACCC9E98EB68883D12C72" value="1985-08-16">
<collectingDate id="600C4C8E5FE49DD2572732D2D86A9F3E" value="1985-08-16">16 ago. 1985</collectingDate>
</date>
;
<collectorName id="2879A5F06F8CA133260EF94546A38891">P. Ward</collectorName>
leg.;
<specimenCode id="A6ADD1611818778E04D28CE77AEB8E48" collectionCode="ICN" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" type="">
<abbrev id="ABBRID0ENMCI" xlink_title="Colección de Entomología del Instituto de Ciencias Naturales, Universidad Nacional de Colombia">ICN</abbrev>
019704
</specimenCode>
(
<abbrev id="ABBRID0ERMCI" xlink_title="Colección de Entomología del Instituto de Ciencias Naturales, Universidad Nacional de Colombia">
<collectionCode id="1AD6A4C3A5BD4BC3BA027BBCBAB5F444" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" type="">ICN</collectionCode>
</abbrev>
)
</materialsCitation>
.
</paragraph>
<caption id="941C1F3BF9E10A81212BBD4B1643E1D4" ID-DOI="10.3897/zookeys.1232.141693.figure28" ID-arpha="C5E943EA-1B86-53D3-BFDD-CDCA393A43B8" httpUri="https://binary.pensoft.net/fig/1282998" startId="F28">
<paragraph id="FDC737CCA62F5C2FFD61D2716A525A95">
<label id="CDC62EDC3E3D19E2F612E2F187B8F5BC">Figure 28.</label>
</paragraph>
<paragraph id="77C200405FB2A50AD922203B1A2DAC7B">
<taxonomicName id="1BF0BE28C4B0CA4729A1AB7B23740B06" authorityName="Arredondo-H &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="xerophilus" status="sp. nov.">
<emphasis id="59EE1730134EB58F0F56FE77AADBD769" italics="true">Nesomyrmex xerophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43894C4FE70B71EA3EC83C53F2F24994" rank="species">sp. nov.</taxonomicNameLabel>
Holotype worker (
<named-content id="CF050FB460B71C2A3AAB5AF368C4A0B7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-del-magdalena" xlink_title="Centro de Colecciones Biológicas de la Universidad del Magdalena">CBUMAG</named-content>
: ENT 12440)
<emphasis id="978504E49D0277A9045D1B3C770AB70B" bold="true">A</emphasis>
full-face view
<emphasis id="49A120B69779D92F594C511E019ACF4A" bold="true">B</emphasis>
lateral view
<emphasis id="2A99FCAD7BD49615EA795C3D6D9CFF1C" bold="true">C</emphasis>
dorsal view. Scale bars: 0.2 mm (
<emphasis id="0416F1D093A21A4FB9EDE32F5A8C8277" bold="true">A</emphasis>
); 0.5 mm (
<emphasis id="56F37C514F91CE2EA31B09E55DA5A3AE" bold="true">B, C</emphasis>
).
</paragraph>
</caption>
<caption id="F0BAB5D978008B8B26B268EE6A72086F" ID-DOI="10.3897/zookeys.1232.141693.figure29" ID-arpha="655CAE3E-289A-5F41-A9FE-2D27B5753502" httpUri="https://binary.pensoft.net/fig/1282999" startId="F29">
<paragraph id="498C4736965B3FC154842479D155A2B1">
<label id="0B19C115A458C1F13C5B028A8EAA7EE7">Figure 29.</label>
</paragraph>
<paragraph id="3348E751BA4482600D8DD78051B5C8B8">
Distributional map of
<taxonomicName id="30435167579A4E37E0716BB3D8201BCD" class="Insecta" family="Formicidae" genus="Nesomyrmex" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="49694660237488AC31F184E92F135350" italics="true">Nesomyrmex</emphasis>
</taxonomicName>
species from the tropical dry forest in Colombia.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EPOCI" type="Geographic range">
<paragraph id="456E2A1657DB02043FCE38F465A2D223">
<heading id="CD051B74802E25CD50D5B0496554001D" reason="title">Geographic range.</heading>
</paragraph>
<paragraph id="DEABB0571775264D8FCABABC964CB179">
<collectingCountry id="E87910F487E4BEBCB4942927257C62A7" name="Colombia">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUOCI" type="description">
<paragraph id="3DF726197DBF1E0C6339ABEC95BBAA8D">
<heading id="DE63A2E237145492F4BAF82E20A0121A" reason="title">
<typeStatus id="0C86674A253E9AEA79522BC7D5E9C482" type="holotype">Holotype</typeStatus>
worker measurements.
</heading>
</paragraph>
<paragraph id="CB8C8B6AEEADF7B34088DD72EE093794">
<abbrev id="ABBRID0E1OCI" xlink_title="Head length">HL</abbrev>
0.69,
<abbrev id="ABBRID0E5OCI" xlink_title="Head width">HW</abbrev>
0.6,
<abbrev id="ABBRID0ECPCI" xlink_title="Scape length">SL</abbrev>
0.44, ML 0.29,
<abbrev id="ABBRID0EGPCI" xlink_title="Eye length">EL</abbrev>
0.15,
<abbrev id="ABBRID0EKPCI" xlink_title="Pronotal width">PW</abbrev>
0.43,
<abbrev id="ABBRID0EOPCI" xlink_title="Petiole width">PTW</abbrev>
0.28,
<abbrev id="ABBRID0ESPCI" xlink_title="Postpetiole width">PPW</abbrev>
0.37,
<abbrev id="ABBRID0EWPCI" xlink_title="Webers length">WL</abbrev>
0.88,
<abbrev id="ABBRID0E1PCI" xlink_title="Pronotal height">PH</abbrev>
0.23,
<abbrev id="ABBRID0E5PCI" xlink_title="Petiole length">PTL</abbrev>
0.23,
<abbrev id="ABBRID0ECQCI" xlink_title="Petiole height">PTH</abbrev>
0.25,
<abbrev id="ABBRID0EGQCI" xlink_title="Postpetiole length">PPL</abbrev>
0.22,
<abbrev id="ABBRID0EKQCI" xlink_title="Postpetiole height">PPH</abbrev>
0.23,
<abbrev id="ABBRID0EOQCI" xlink_title="Gaster length">GL</abbrev>
0.66
<abbrev id="ABBRID0ESQCI" xlink_title="Cephalic index">CI</abbrev>
86,
<abbrev id="ABBRID0EWQCI" xlink_title="Scape index">SI</abbrev>
0.73.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1QCI" type="description">
<paragraph id="A4CD70BCDCD278AB67DA1A0C0A03A55D">
<heading id="9CE5B2B17BBEA0119C2B16CA0E3A3DAF" reason="title">
<typeStatus id="0D488390DA52CED4F9CE1AC2D49D3435" type="paratype">Paratype</typeStatus>
worker measurements.
</heading>
</paragraph>
<paragraph id="C35B8AFCCF30A1C3136B7C80851DCD1B">
(
<emphasis id="E29D62AA69EFD3E4D70F2A2A2C8004C5" italics="true">n</emphasis>
= 4)
<abbrev id="ABBRID0ECRCI" xlink_title="Head length">HL</abbrev>
0.690.7,
<abbrev id="ABBRID0EGRCI" xlink_title="Head width">HW</abbrev>
0.590.6,
<abbrev id="ABBRID0EKRCI" xlink_title="Scape length">SL</abbrev>
0.430.44, ML 0.29,
<abbrev id="ABBRID0EORCI" xlink_title="Eye length">EL</abbrev>
0.150.16,
<abbrev id="ABBRID0ESRCI" xlink_title="Pronotal width">PW</abbrev>
0.430.44,
<abbrev id="ABBRID0EWRCI" xlink_title="Petiole width">PTW</abbrev>
0.280.29,
<abbrev id="ABBRID0E1RCI" xlink_title="Postpetiole width">PPW</abbrev>
0.360.37,
<abbrev id="ABBRID0E5RCI" xlink_title="Webers length">WL</abbrev>
0.90.91,
<abbrev id="ABBRID0ECSCI" xlink_title="Pronotal height">PH</abbrev>
0.220.23,
<abbrev id="ABBRID0EGSCI" xlink_title="Petiole length">PTL</abbrev>
0.230.27,
<abbrev id="ABBRID0EKSCI" xlink_title="Petiole height">PTH</abbrev>
0.240.25,
<abbrev id="ABBRID0EOSCI" xlink_title="Postpetiole length">PPL</abbrev>
0.220.23,
<abbrev id="ABBRID0ESSCI" xlink_title="Postpetiole height">PPH</abbrev>
0.230.24,
<abbrev id="ABBRID0EWSCI" xlink_title="Gaster length">GL</abbrev>
0.660.68,
<abbrev id="ABBRID0E1SCI" xlink_title="Cephalic index">CI</abbrev>
8586,
<abbrev id="ABBRID0E5SCI" xlink_title="Scape index">SI</abbrev>
7273.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECTCI" type="diagnosis">
<paragraph id="0C6DFF01A09E0431B5B26FCA7EFA6DE2">
<heading id="D6D4655E50E2C5C54D7800892E7ADD2C" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="01B559C350196C56556244DD358D62F0">Frontal lobe projected laterally, covering the antennal insertions. Frontal carina extending to posterior cephalic vertex, forming a weak scrobus. Dorsum of mesosoma straight and flattened. Lateral margin of pronotum straight, converging towards mesonotum. Lateral projection of the mesonotum dentate. Lateral dorsopropodeal processes long, covering propodeal spiracle in dorsal view.</paragraph>
</subSubSection>
<subSubSection id="SECID0EHTCI" type="description">
<paragraph id="A60E2F3F49C6125122A94DD23F8C8897">
<heading id="D9D42430EC3BC46088CA45B13A3B348D" reason="title">Description.</heading>
</paragraph>
<paragraph id="C7B607F9DBD500128D182359B5888DAD">
<emphasis id="6FF0669E88316DBCA5E95EE2E10B5AEE" bold="true">Worker.</emphasis>
In full-face view, head longer than wide (
<abbrev id="ABBRID0EPTCI" xlink_title="Cephalic index">CI</abbrev>
8586), slightly narrowing behind the eyes; occipital margin weakly convex; occipital angles rounded; lateral margins slightly curved posteriorly; anterior margin of clypeus weakly convex, projecting over mandibles; eyes lateral, with 10 ommatidia at greatest diameter; frontal lobes laterally expanded, antennal insertions completely hidden by frontal lobes; frontal carina extending to posterior cephalic vertex, as if forming a weak scrobus; antenna with 12 antennomeres; mandible triangular; inner margin with five teeth, separated from basal margin by fifth tooth; scapes short (
<abbrev id="ABBRID0ETTCI" xlink_title="Scape index">SI</abbrev>
7273) barely reaching posterior margin of eye, curved towards mid-length; pedicel longer than wide, and almost as long as next three antennomeres together; antennal club with three antennomeres.
</paragraph>
<paragraph id="25C7E9B238B8AFECD945AEF05B5F6E7E">
<emphasis id="21083337321EB5067CAA2D726AC63832" bold="true">
<emphasis id="48C69EA21EF0F5FF39A19E1E40CF5552" bold="true" italics="true">Mesosoma</emphasis>
.
</emphasis>
In lateral view, mesosomal profile straight; in dorsal view, pronotum wider than long, with its maximum width towards anterior margin, the latter slightly convex, humeral angle angulated, lateral margins of pronotum curved; in lateral view, promesonotal suture present, in dorsal view, absent; in dorsal view, mesosomal lateral projection angulate; dorsopropodeal lateral projection angulate, projecting over propodeal spiracles; propodeal spines short, less than half the distance between their apices in length, and divergent; in lateral view, propodeal spiracle small (diameter =
<quantity id="F710D8C23A543884B947F2E8D395F5B1" metricMagnitude="-5" metricUnit="m" metricValue="5.7" unit="mm" value="0.057">0.057 mm</quantity>
), approximately equal to the length of the 10
<superScript id="F4505DD85AFCEDDEEFCC41D7E21DFAF1">th</superScript>
antennomere, circular, projected posterolaterally; propodeal lobe short and uniformly rounded.
</paragraph>
<paragraph id="A7140ACBABF409FB046B9FEF090B8EF8">
<emphasis id="C73F0810F4E4EC448D03305B4F5BEE88" bold="true">
<emphasis id="CF7DDE2AEBD487148323BECE868993AC" bold="true" italics="true">Metasoma</emphasis>
.
</emphasis>
In dorsal view, petiole trapezoidal, anterior margin shorter than posterior margin; posterior margin with two mesial tubercles of equal length; lateral margins diverging from anterior to posterior margin; lateral margin with two posterolaterals tubercles; in anteroposterior direction, first tubercle poorly developed, length less than half the total length of the following tubercle; in lateral view, petiole subquadrate, peduncle and petiolar node without apparent differentiation, forming a continuous outline, which extends to the dorsal surface of the petiolar node; anterior surface of petiole with anteromedial petiolar spine on each side, superior to the petiolar spiracle; dorsal surface weakly rounded and short; anteroventral process of petiole acute; in dorsal view, postpetiole hexagonal, twice as wide as long; in lateral view postpetiole globose, taller than long.
</paragraph>
<paragraph id="94DCB30CE01F3EB4CDB42517C20B99B1">
<emphasis id="68D8955E2503F13961F289AD7C3EEFC5" bold="true">
<emphasis id="E2829006BD717ABF1E32F9EA339D0C9E" bold="true" italics="true">Sculpture</emphasis>
.
</emphasis>
Body generally opaque, with smooth, shiny areas restricted to gaster; dorsal surface of body rugose-reticulate, excluding gaster; lateral surface of mesosoma reticulate; clypeus smooth and shiny, with medial longitudinal carina; dorsal surface of mandibles shiny, with weakly marked striation; scape and legs shiny with superficial sculpture; gaster weakly imbricate.
</paragraph>
<paragraph id="A224870A85E15AAF4DFBE143919DDE23">
<emphasis id="81AD188962965961BFDCE09E02E9D3AD" bold="true">
<emphasis id="FE7303E93D72E4AA8A50D007AA19354D" bold="true" italics="true">Pilosity and color</emphasis>
.
</emphasis>
Dorsal surface of body with flattened erect hairs, mostly separated by a distance equal to or greater than its length; hairs as long as the base of the propodeal spines; in full-face view, head capsule with erect hairs restricted to the clypeus, area between the frontal carinae and vertex; scapes covered with short decumbent hairs, without erect hairs; propodeal spines with erect hairs; legs with simple, appressed, sparse hairs restricted to the apex of the femora; gaster with abundant erect hairs, separated by a distance equal to its length. Body pale yellow, legs slightly paler, base and apex of femur and tibia darker.
</paragraph>
<paragraph id="43982B910007A0258642873276CCD8D4">
<emphasis id="3E358B8AD0AAD7E885C0999C88C36AD9" bold="true">Queen and male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZUCI" type="natural history">
<paragraph id="BCBE0BD1F4B248E940B0FF7D5BA829AB">
<heading id="1DEC46F7FEB551A60F9740AE96639213" reason="title">Natural history.</heading>
</paragraph>
<paragraph id="456EB63D397AE1266E945B68759CE89B">
This species lives within dead branches of shrub vegetation. Populations are distributed in fragments ~
<quantity id="544EE9510C9AB4741D08949C9C7E8B4C" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</quantity>
altitude. In all cases, the ants were found within dry forest with dense vegetation.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5UCI" type="etymology">
<paragraph id="69307D81828444CB420244EA01B8CBC5">
<heading id="281B7E2CEB20ED7115D734BEE2DF6EE5" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="6C46E1F8BEDC565F6033F3071A898BF7">
The epithet
<taxonomicName id="084017DEB3C4606B27CB8FEF89451FD0" authorityName="Arredondo-H &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="xerophilus">
<emphasis id="EA444EE64B99B89886216D4892329404" italics="true">xerophilus</emphasis>
</taxonomicName>
is a word composed of the prefix
<emphasis id="6416A8344EC1D9D7B677CBC9B2B2A4B9" italics="true">xero</emphasis>
- (from ancient Greek
<emphasis id="535FB39BA3AD78CE5EFAC176C77B11FD" italics="true">ξηρο</emphasis>
- meaning dry) and the suffix -
<emphasis id="A360E97A3BD51AAE6B40A8436CE9ED16" italics="true">philo</emphasis>
or -
<emphasis id="0D682E2456E3BD0CDEAE34B5E2882F82" italics="true">philus</emphasis>
(from ancient Greek
<emphasis id="EDA55EEFA1F056FF338F6FA8D7D4F92D" italics="true">φίλος</emphasis>
meaning attraction towards). The species epithet refers to the dry habitat where these ants live, that is, the tropical dry forest, a life zone highly threatened by the high rate of deforestation in
<collectingCountry id="24C9A85DCCA167009516237873CF932E" name="Colombia">Colombia</collectingCountry>
. It is a noun in apposition and thus invariant.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYVCI" type="comments">
<paragraph id="CF5AB62A7A840D1B2A6A4D49668DAB39">
<heading id="3F87F3F18BC1464F76586122DC674C0E" reason="title">Comments.</heading>
</paragraph>
<paragraph id="89118E6B29203D043DD7E5E12DDA221E">
Species easily recognizable by the lateral expansion of the frontal lobes and the lateral projection of the mesosoma and dorsopropodeum in the form of an angled lobe. Morphologically, the closest species to
<taxonomicName id="C0ECF6E571737BA0577A4A8E47CC5F1D" authorityName="Arredondo-H &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="xerophilus">
<emphasis id="6CFE5DC3607A9F79F359EC7CB9B9EBC1" italics="true">N. xerophilus</emphasis>
</taxonomicName>
is
<taxonomicName id="255E36574DF458B7AE57E0A93BD05D25" authorityName="Arredondo &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="iku">
<emphasis id="55CFD769DE7D86523E1884B9FFC9C5AE" italics="true">N. iku</emphasis>
</taxonomicName>
(see comments on
<taxonomicName id="9A6D4D5F477EE0E24EB1C2A9801AA8C8" authorityName="Arredondo &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="iku" status="sp. nov.">
<emphasis id="FF2A7EEDE393D52C5836A3E7E838C06A" italics="true">Nesomyrmex iku</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3FBD15E7D02A70AB7AC828AB9A796113" rank="species">sp. nov.</taxonomicNameLabel>
).
<taxonomicName id="CDACC878239B57EA8A87B359CE0E929B" authorityName="Arredondo-H &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="xerophilus">
<emphasis id="01C0F1614AA44CA334ADC80B53F62D15" italics="true">N. xerophilus</emphasis>
</taxonomicName>
is morphologically similar to
<taxonomicName id="C51885775554D2A1CEA488228870E46D" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="wilda">
<emphasis id="015D996825D2E6270DF24BEE7F51489F" italics="true">N. wilda</emphasis>
</taxonomicName>
, with both species exhibiting a lateral projection of the mesonotum dentate, long lateral dorsopropodeal processes covering of the propodeal spiracle in dorsal view. Additionally, they share the shape of the propodeal lobes, coloration, and sculpture. Despite these similarities,
<taxonomicName id="1CC0AB9FCEE469880D02D4FE7EAEF17A" authorityName="Arredondo-H &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="xerophilus">
<emphasis id="9123B73E7783EEEA9CE1EEE7BAA8A08F" italics="true">N. xerophilus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="20D6DD1E11E806E6195BBBE2923AB037" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="wilda">
<emphasis id="9B3A412D7F67B4CFBB060345AB049B3F" italics="true">N. wilda</emphasis>
</taxonomicName>
in the number of antennomeres, as well as in the shape of the frontal lobes, petiole, and postpetiole.
</paragraph>
<paragraph id="97A52C0B449750E869D55CECC41E3620">
Unlike
<taxonomicName id="D82FA4ACEC2EA09D9A13E9C0FE62BA7D" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="5D4391A46E11249337DF944766C7C95B" italics="true">N. asper</emphasis>
</taxonomicName>
, the position and number of petiolar and postpetiolar tubercles in
<taxonomicName id="07E70C872D5F03CDF51EEDD0A1266B7E" authorityName="Arredondo-H &amp; Guerrero" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="xerophilus">
<emphasis id="B9088126C2A25C52B609A4A069308FF7" italics="true">N. xerophilus</emphasis>
</taxonomicName>
remain consistent across its populations, with no variation observed in this characteristic. This stability may be attributed to its limited distribution in the tropical dry forest of the Colombian Caribbean region. In contrast, the variation in the position and number of petiolar and postpetiolar tubercles in
<taxonomicName id="4597C0AEFA41D0251542D2F0F92151E2" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="FF8F7694AAF5E7286E72618A6E1F0160" italics="true">N. asper</emphasis>
</taxonomicName>
appears to be linked to its altitudinal distribution and geographic location (see comments on
<taxonomicName id="42315D27111D5F719674A2B4E330A1AF" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="65E463FAC1FA09B1BE588169ACA2402F" italics="true">N. asper</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

File diff suppressed because it is too large Load diff