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<document ID-DOI="http://dx.doi.org/10.3897/zse.97.68000" ID-GBIF-Dataset="617dbf89-ec8b-48a2-a13e-ea581e42aecc" ID-GBIF-Taxon="184132871" ID-Pensoft-Pub="1860-0743-2-407" ID-Pensoft-UUID="9446F0CE40A752B4897F7B7B71BBFD29" ID-ZooBank="DC2EBA6293A141938ADC2A79F7D658B9" ModsDocID="1860-0743-97-2-407" checkinTime="1628790185545" checkinUser="pensoft" docAuthor="Tillack, Frank, Ruiter, Ronald de &amp; Roedel, Mark-Oliver" docDate="2021" docId="A03227687E8C5128B31BC0DCFD126E2A" docLanguage="en" docName="ZoosystEvol 97(2): 407-450" docOrigin="Zoosystematics and Evolution 97 (2)" docPubDate="2021-08-10" docSource="http://dx.doi.org/10.3897/zse.97.68000" docTitle="Hyperolius pulchromarmoratus Ahl 1931" docType="treatment" docVersion="3" id="9446F0CE40A752B4897F7B7B71BBFD29" lastPageNumber="407" masterDocId="9446F0CE40A752B4897F7B7B71BBFD29" masterDocTitle="A type catalogue of the reed frogs (Amphibia, Anura, Hyperoliidae) in the collection of the Museum fuer Naturkunde Berlin (ZMB) with comments on historical collectors and expeditions" masterLastPageNumber="450" masterPageNumber="407" pageId="0" pageNumber="407" updateTime="1643430994971" updateUser="ExternalLinkService">
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<mods:title>A type catalogue of the reed frogs (Amphibia, Anura, Hyperoliidae) in the collection of the Museum fuer Naturkunde Berlin (ZMB) with comments on historical collectors and expeditions</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tillack, Frank</mods:namePart>
<mods:affiliation>Museum fuer Naturkunde Berlin, Leibniz-Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, 10115 Berlin, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ruiter, Ronald de</mods:namePart>
<mods:affiliation>Nederlands Openluchtmuseum, Hoeferlaan 4, 6816 SG Arnhem, The Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Roedel, Mark-Oliver</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1666-195X</mods:nameIdentifier>
<mods:affiliation>Museum fuer Naturkunde Berlin, Leibniz-Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, 10115 Berlin, Germany</mods:affiliation>
<mods:nameIdentifier type="email">mo.roedel@mfn-berlin.de</mods:nameIdentifier>
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<mods:number>2021-08-10</mods:number>
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<mods:start>407</mods:start>
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<treatment ID-GBIF-Taxon="184132871" LSID="urn:lsid:plazi:treatment:A03227687E8C5128B31BC0DCFD126E2A" httpUri="http://treatment.plazi.org/id/A03227687E8C5128B31BC0DCFD126E2A" lastPageNumber="407" pageId="0" pageNumber="407">
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<taxonomicName LSID="A0322768-7E8C-5128-B31B-C0DCFD126E2A" authority="Ahl, 1931 a: 92." authorityName="Ahl" authorityYear="1931" class="Reptilia" family="Hyperoliidae" genus="Hyperolius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyperolius pulchromarmoratus" order="Anura" pageId="0" pageNumber="407" phylum="Chordata" rank="species" species="pulchromarmoratus">Hyperolius pulchromarmoratus Ahl, 1931a: 92.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="407" type="holotype">
<paragraph pageId="0" pageNumber="407">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="407">
ZMB 77751, &quot;Britisch Ostafrika&quot; [Kenya], coll. Richard Fritz Paul
<normalizedToken originalValue="Hübner">Huebner</normalizedToken>
[later Huebner].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="407" type="present name">
<paragraph pageId="0" pageNumber="407">Present name.</paragraph>
<paragraph pageId="0" pageNumber="407">
<taxonomicName authorityName="Peters" authorityYear="1878" class="Reptilia" family="Hyperoliidae" genus="Hyperolius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyperolius glandicolor" order="Anura" pageId="0" pageNumber="407" phylum="Chordata" rank="species" species="glandicolor">
<emphasis italics="true" pageId="0" pageNumber="407">Hyperolius glandicolor</emphasis>
</taxonomicName>
Peters, 1878.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="407" type="remarks">
<paragraph pageId="0" pageNumber="407">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="407">
Drawing in
<bibRefCitation DOI="https://doi.org/10.1515/9783111625430-fm" author="Ahl, E" journalOrPublisher="Mitteilungen aus dem Zoologischen Museum in Berlin" pageId="0" pageNumber="407" refId="B9" refString="Ahl, E, 1931b. Amphibia, Anura III, Polypedatidae. In: Hesse R (Ed.) Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. 55. Lieferung. Walter de Gruyter &amp; Co. Berlin und Leipzig, 477 pp. [+ (1-2)]. https://doi.org/10.1515/9783111625430-fm" title="Amphibia, Anura III, Polypedatidae. In: Hesse R (Ed.) Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. 55. Lieferung. Walter de Gruyter &amp; Co. Berlin und Leipzig, 477 pp. [+ (1 - 2)]." url="https://doi.org/10.1515/9783111625430-fm" year="1931 b">Ahl (1931b</bibRefCitation>
: 367, fig. 242). Huebner worked from 1894 to 1913 as a merchant, banker, farmer and administrator and from 1901 to 1903 as Municipal Commissioner of Nairobi in (British) East Africa. He was active in Zanzibar (1894-1896), Mombasa (1896-1899), Nairobi (1899-1905), Kibwezi (1905-1908, together with G. R. O. Scheffler), and Voi (1908-1913), and undertook a journey from Mombasa to Kampala from June to November 1899. In 1913 he travelled to Germany for a convalescent stay because of health problems. However, his already planned return to Kenya was thwarted by the beginning of the First World War. In his spare time he was engaged in nature observations and collected interesting zoological objects, which he sent to ZMB (
<bibRefCitation author="Sieberg, H" journalOrPublisher="Hildesheimer Universitaetsschriften, Band 3" pageId="0" pageNumber="407" refId="B200" refString="Sieberg, H, 1998. Zeitenumbruch in Ostafrika. Sansibar, Kenia und Uganda (1894-1913). Erinnerungen des Kaufmanns R. F. Paul Huebner. Hildesheimer Universitaetsschriften, Band 3" title="Zeitenumbruch in Ostafrika. Sansibar, Kenia und Uganda (1894 - 1913). Erinnerungen des Kaufmanns R. F. Paul Huebner." year="1998">Sieberg 1998</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="8875CD177F66EC171B693322F268014F">Chakra, a new genus of Scelioninae (Hymenoptera: Platygastridae) from India, along with description of a new species</mods:title>
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<treatment id="A0322C16FFD0FFE6FF23FD95FC14C19B" ID-DOI="http://doi.org/10.5281/zenodo.6143531" ID-GBIF-Taxon="119508034" ID-Zenodo-Dep="6143531" LSID="urn:lsid:plazi:treatment:A0322C16FFD0FFE6FF23FD95FC14C19B" httpUri="http://treatment.plazi.org/id/A0322C16FFD0FFE6FF23FD95FC14C19B" lastPageNumber="286" pageId="1" pageNumber="286">
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<paragraph id="28249D00FFD0FFE6FF23FD95FD44C47B" blockId="1.[151,750,544,605]" box="[151,750,544,571]" pageId="1" pageNumber="286">
<heading id="736C2A6CFFD0FFE6FF23FD95FD44C47B" bold="true" box="[151,750,544,571]" fontSize="11" level="1" pageId="1" pageNumber="286" reason="1">
<emphasis id="1AEF4112FFD0FFE6FF23FD95FD44C47B" bold="true" box="[151,750,544,571]" pageId="1" pageNumber="286">
<taxonomicName id="EF9BE683FFD0FFE6FF23FD95FDDEC47B" authority="Rajmohana and Veenakumari" authorityName="Rajmohana and Veenakumari" box="[151,628,544,571]" class="Hexapoda" family="Platygastridae" genus="Chakra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="1AEF4112FFD0FFE6FF23FD95FF58C47A" bold="true" box="[151,242,544,570]" italics="true" pageId="1" pageNumber="286">Chakra</emphasis>
Rajmohana and Veenakumari
</taxonomicName>
, gen. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="28249D00FFD0FFE6FF23FDF1FEB2C41C" blockId="1.[151,750,544,605]" box="[151,280,580,605]" pageId="1" pageNumber="286">
(
<figureCitation id="B0A08185FFD0FFE6FF2BFDF1FEA5C41C" box="[159,271,580,605]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 7 – 11" captionStartId-0="2.[151,264,1850,1873]" captionStartId-1="3.[151,264,1624,1647]" captionTargetBox-0="[159,1418,213,1777]" captionTargetBox-1="[190,1404,205,1592]" captionTargetId-0="figure@2.[143,1431,199,1792]" captionTargetId-1="figure@3.[175,1411,193,1604]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 – 6. Bright field images of Chakra sarvatra sp. nov. 1 — Habitus; 2 — Head, dorsal view; 3 — Head and mesosoma, dorsolateral view; 4 — Mesosoma, dorsal view 5 — Head, frontal view 6 — Mouthparts." captionText-1="FIGURES 7 – 11. Bright field images of Chakra sarvatra sp. nov. 7 — Male antennae and head; 8 — Female antennae showing rows of white sensilla; 9 — Fore wing; 10 — Ovipositor; 11 — Setigerous tubercles on head and mesosoma." httpUri-0="https://zenodo.org/record/225566/files/figure.png" httpUri-1="https://zenodo.org/record/225567/files/figure.png" pageId="1" pageNumber="286">Figs 1–11</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="28249D00FFD0FFE6FF23FD39FCE7C4E5" blockId="1.[151,1437,652,2011]" box="[151,845,652,677]" pageId="1" pageNumber="286">
<emphasis id="1AEF4112FFD0FFE6FF23FD39FE9EC4E5" bold="true" box="[151,308,652,677]" pageId="1" pageNumber="286">
<typeStatus id="F72023A2FFD0FFE6FF23FD39FF78C4E5" box="[151,210,652,677]" pageId="1" pageNumber="286">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="EF9BE683FFD0FFE6FE8FFD39FCE7C4E5" authority="Rajmohana and Veenakumari." authorityName="Rajmohana and Veenakumari." box="[315,845,652,677]" class="Hexapoda" family="Platygastridae" genus="Chakra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="286" phylum="Arthropoda" rank="species" species="sarvatra">
<emphasis id="1AEF4112FFD0FFE6FE8FFD39FE5FC4E4" box="[315,501,652,676]" italics="true" pageId="1" pageNumber="286">Chakra sarvatra</emphasis>
Rajmohana and Veenakumari.
</taxonomicName>
</paragraph>
<paragraph id="28249D00FFD0FFE6FF73FD1AFB29C4AD" blockId="1.[151,1437,652,2011]" pageId="1" pageNumber="286">
<emphasis id="1AEF4112FFD0FFE6FF73FD1AFDAFC488" bold="true" box="[199,517,687,712]" pageId="1" pageNumber="286">Derivation of genus name.</emphasis>
‘
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<emphasis id="1AEF4112FFD0FFE6FDA2FD05FDC0C488" box="[534,618,688,712]" italics="true" pageId="1" pageNumber="286">Chakra</emphasis>
</taxonomicName>
’ in Sanskrit means wheel, referring to the unique wheel-like setigerous flat-topped tubercles distributed all over the head and mesosoma. The gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="6081CE8BFFD0FFE6FF73FD42FDACC5A8" pageId="1" pageNumber="286" type="diagnosis">
<paragraph id="28249D00FFD0FFE6FF73FD42FDACC5A8" blockId="1.[151,1437,652,2011]" pageId="1" pageNumber="286">
<emphasis id="1AEF4112FFD0FFE6FF73FD42FEEBC550" bold="true" box="[199,321,759,784]" pageId="1" pageNumber="286">Diagnosis.</emphasis>
Although the specimens of the new genus key out to
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<emphasis id="1AEF4112FFD0FFE6FC0CFD4DFBF7C550" box="[952,1117,760,784]" italics="true" pageId="1" pageNumber="286">Opisthacantha</emphasis>
Ashmead
</taxonomicName>
at couplets 59 of
<bibRefCitation id="4C0AE0F1FFD0FFE6FF23FCA8FE97C575" author="Masner" box="[151,317,796,821]" pageId="1" pageNumber="286" refString="Masner, L. (1976) Revisionary notes and keys to world genera of Scelionidae (Hymenoptera: Proctotrupidae). Memoirs of the Entomological Society of Canada, 97, 1 - 87. http: // dx. doi. org / 10.4039 / entm 10897 fv" type="journal article" year="1976">Masner (1976)</bibRefCitation>
and 50 of
<bibRefCitation id="4C0AE0F1FFD0FFE6FE02FCA9FD53C574" author="Austin" box="[438,761,796,821]" pageId="1" pageNumber="286" refString="Galloway, I. D. &amp; Austin, A. D. (1984) Revision of the Scelioninae (Hymenoptera: Scelionidae) in Australia. Australian Journal of Zoology, Supplementary Series, 99, 1 - 138. http: // dx. doi. org / 10.1071 / ajzs 099" type="journal article" year="1984">Austin and Galloway (1984)</bibRefCitation>
, they do not fit into the concept of the genus
<taxonomicName id="EF9BE683FFD0FFE6FB43FCA9FE54C518" authority="Ashmead sensu Masner (1976)" authorityName="Ashmead sensu Masner" authorityYear="1976" class="Insecta" family="Scelionidae" genus="Opisthacantha" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="1AEF4112FFD0FFE6FB43FCA9FA36C574" box="[1271,1436,796,820]" italics="true" pageId="1" pageNumber="286">Opisthacantha</emphasis>
Ashmead
<emphasis id="1AEF4112FFD0FFE6FEBBFCF4FEE7C518" box="[271,333,833,856]" italics="true" pageId="1" pageNumber="286">sensu</emphasis>
<bibRefCitation id="4C0AE0F1FFD0FFE6FEE1FCF5FE54C518" author="Masner" box="[341,510,831,856]" pageId="1" pageNumber="286" refString="Masner, L. (1976) Revisionary notes and keys to world genera of Scelionidae (Hymenoptera: Proctotrupidae). Memoirs of the Entomological Society of Canada, 97, 1 - 87. http: // dx. doi. org / 10.4039 / entm 10897 fv" type="journal article" year="1976">Masner (1976)</bibRefCitation>
</taxonomicName>
. The new genus differs from
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<emphasis id="1AEF4112FFD0FFE6FCFDFCF5FC44C518" box="[841,1006,832,856]" italics="true" pageId="1" pageNumber="286">Opisthacantha</emphasis>
</taxonomicName>
in having frons, vertex, dorsal, lateral and ventral mesosomal surfaces with large, round, setigerous tubercles, whose diameter is almost equal to the diameter of the lateral ocelli (
<figureCitation id="B0A08185FFD0FFE6FE41FC32FDEBC5E0" box="[501,577,903,928]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1850,1873]" captionTargetBox="[159,1418,213,1777]" captionTargetId="figure@2.[143,1431,199,1792]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Bright field images of Chakra sarvatra sp. nov. 1 — Habitus; 2 — Head, dorsal view; 3 — Head and mesosoma, dorsolateral view; 4 — Mesosoma, dorsal view 5 — Head, frontal view 6 — Mouthparts." httpUri="https://zenodo.org/record/225566/files/figure.png" pageId="1" pageNumber="286">Figs 2</figureCitation>
,
<figureCitation id="B0A08185FFD0FFE6FDE5FC3DFDC6C5E0" box="[593,620,904,928]" captionStart="FIGURES 7 – 11" captionStartId="3.[151,264,1624,1647]" captionTargetBox="[190,1404,205,1592]" captionTargetId="figure@3.[175,1411,193,1604]" captionTargetPageId="3" captionText="FIGURES 7 – 11. Bright field images of Chakra sarvatra sp. nov. 7 — Male antennae and head; 8 — Female antennae showing rows of white sensilla; 9 — Fore wing; 10 — Ovipositor; 11 — Setigerous tubercles on head and mesosoma." httpUri="https://zenodo.org/record/225567/files/figure.png" pageId="1" pageNumber="286">11</figureCitation>
) and also in having remarkably small eyes that are hardly as long as temples, small ocellar triangle, with lateral ocelli much closer to median ocellus than to orbits (OOL&gt; POL) and raised interantennal prominence.
</paragraph>
</subSubSection>
<subSubSection id="6081CE8BFFD0FFE6FF73FC41FC14C19B" pageId="1" pageNumber="286" type="description">
<paragraph id="28249D00FFD0FFE6FF73FC41FC14C19B" blockId="1.[151,1437,652,2011]" pageId="1" pageNumber="286">
<emphasis id="1AEF4112FFD0FFE6FF73FC41FEF3C24D" bold="true" box="[199,345,1012,1037]" pageId="1" pageNumber="286">Description.</emphasis>
Body minute, 1.0––
<quantity id="EF6330E5FFD0FFE6FD8AFC41FD39C24C" box="[574,659,1012,1036]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="286" unit="mm" value="1.2">1.2 mm</quantity>
long. Head and body brown to blackish brown; Frons, vertex, dorsal, lateral and ventral mesosoma with large round setigerous tubercles, the diameter of the tubercles almost equal to diameter of lateral ocelli.
<emphasis id="1AEF4112FFD0FFE6FE7AFB89FDA4C215" bold="true" box="[462,526,1084,1109]" pageId="1" pageNumber="286">Head</emphasis>
distinctly transverse dorsally, subellipsoidal, densely setose; Occipital carina present, crenulate, extending throughout and continuing on lower margin of postgena; Eyes small, shorter than temples; Interommatidial setae short, 1.7 times as long as the diameter of a single ommatidium, comparable both in size and density to setae on frons and vertex; Temple visible in dorsal view; Ocelli clustered in a small triangle on vertex; OOL&gt;POL&gt;LOL (
<figureCitation id="B0A08185FFD0FFE6FE76FB79FDADC2A4" box="[450,519,1228,1253]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1850,1873]" captionTargetBox="[159,1418,213,1777]" captionTargetId="figure@2.[143,1431,199,1792]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Bright field images of Chakra sarvatra sp. nov. 1 — Habitus; 2 — Head, dorsal view; 3 — Head and mesosoma, dorsolateral view; 4 — Mesosoma, dorsal view 5 — Head, frontal view 6 — Mouthparts." httpUri="https://zenodo.org/record/225566/files/figure.png" pageId="1" pageNumber="286">Fig. 2</figureCitation>
); Frons convex without frontal depression; Interantennal process well developed with a pronounced curve (
<figureCitation id="B0A08185FFD0FFE6FE0BFB5AFE54C348" box="[447,510,1263,1288]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1850,1873]" captionTargetBox="[159,1418,213,1777]" captionTargetId="figure@2.[143,1431,199,1792]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Bright field images of Chakra sarvatra sp. nov. 1 — Habitus; 2 — Head, dorsal view; 3 — Head and mesosoma, dorsolateral view; 4 — Mesosoma, dorsal view 5 — Head, frontal view 6 — Mouthparts." httpUri="https://zenodo.org/record/225566/files/figure.png" pageId="1" pageNumber="286">Fig.3</figureCitation>
); Clypeus narrow, transverse, more than 3
<emphasis id="1AEF4112FFD0FFE6FC55FB5AFC5BC348" bold="true" box="[993,1009,1263,1288]" pageId="1" pageNumber="286">×</emphasis>
as long as wide; Mandibles short and wide (L: W = 39:48), subtridentate, median tooth highly reduced relative to lateral teeth (
<figureCitation id="B0A08185FFD0FFE6FBC3FAA1FB1CC36D" box="[1143,1206,1300,1325]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1850,1873]" captionTargetBox="[159,1418,213,1777]" captionTargetId="figure@2.[143,1431,199,1792]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Bright field images of Chakra sarvatra sp. nov. 1 — Habitus; 2 — Head, dorsal view; 3 — Head and mesosoma, dorsolateral view; 4 — Mesosoma, dorsal view 5 — Head, frontal view 6 — Mouthparts." httpUri="https://zenodo.org/record/225566/files/figure.png" pageId="1" pageNumber="286">Fig.6</figureCitation>
); Teeth acute; Facial striae short, 0.9 × length of malar sulcus; Malar sulcus wide near inner orbit of eye; Subocular suture present (
<figureCitation id="B0A08185FFD0FFE6FADBFA82FF0CC334" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1850,1873]" captionTargetBox="[159,1418,213,1777]" captionTargetId="figure@2.[143,1431,199,1792]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Bright field images of Chakra sarvatra sp. nov. 1 — Habitus; 2 — Head, dorsal view; 3 — Head and mesosoma, dorsolateral view; 4 — Mesosoma, dorsal view 5 — Head, frontal view 6 — Mouthparts." httpUri="https://zenodo.org/record/225566/files/figure.png" pageId="1" pageNumber="286">Fig. 3</figureCitation>
); Antenna with 12 antennomeres in both sexes, with 6 clavomeres in females (
<figureCitation id="B0A08185FFD0FFE6FBFFFAE9FB3BC334" box="[1099,1169,1372,1397]" captionStart="FIGURES 7 – 11" captionStartId="3.[151,264,1624,1647]" captionTargetBox="[190,1404,205,1592]" captionTargetId="figure@3.[175,1411,193,1604]" captionTargetPageId="3" captionText="FIGURES 7 – 11. Bright field images of Chakra sarvatra sp. nov. 7 — Male antennae and head; 8 — Female antennae showing rows of white sensilla; 9 — Fore wing; 10 — Ovipositor; 11 — Setigerous tubercles on head and mesosoma." httpUri="https://zenodo.org/record/225567/files/figure.png" pageId="1" pageNumber="286">Fig. 8</figureCitation>
) and filiform in males (
<figureCitation id="B0A08185FFD0FFE6FF2BFACAFF74C3D8" box="[159,222,1407,1432]" captionStart="FIGURES 7 – 11" captionStartId="3.[151,264,1624,1647]" captionTargetBox="[190,1404,205,1592]" captionTargetId="figure@3.[175,1411,193,1604]" captionTargetPageId="3" captionText="FIGURES 7 – 11. Bright field images of Chakra sarvatra sp. nov. 7 — Male antennae and head; 8 — Female antennae showing rows of white sensilla; 9 — Fore wing; 10 — Ovipositor; 11 — Setigerous tubercles on head and mesosoma." httpUri="https://zenodo.org/record/225567/files/figure.png" pageId="1" pageNumber="286">Fig.7</figureCitation>
), male A5 carinate.
<emphasis id="1AEF4112FFD0FFE6FE72FACAFDEEC3D8" bold="true" box="[454,580,1407,1432]" pageId="1" pageNumber="286">Mesosoma</emphasis>
almost as long as wide in dorsal view, both mesosoma and mesoscutellum with similar sculpture as head; Skaphion and notaulus absent (
<figureCitation id="B0A08185FFD0FFE6FCE2FA11FC3FC3FD" box="[854,917,1444,1469]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1850,1873]" captionTargetBox="[159,1418,213,1777]" captionTargetId="figure@2.[143,1431,199,1792]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Bright field images of Chakra sarvatra sp. nov. 1 — Habitus; 2 — Head, dorsal view; 3 — Head and mesosoma, dorsolateral view; 4 — Mesosoma, dorsal view 5 — Head, frontal view 6 — Mouthparts." httpUri="https://zenodo.org/record/225566/files/figure.png" pageId="1" pageNumber="286">Fig.4</figureCitation>
); Mesoscutellum 2 × as wide as long, anterior margin crenulate; Lateral axillar area wide; Metascutellum foveolate with a broadly based triangular, medially pointed spine; Scutoscutellar sulcus wide; Epomial carina present; Netrion almost smooth, closed near fore coxa; Acetabular area setose, sculpture partially effaced in mesopleural depression; Mesepimeral sulcus foveolate; Metapleuron smooth dorsally, ventrally sculptured similar to mesonotum; Lateral propodeal area setose, with similar sculpture to that of head and mesoscutum; Propodeum with posterior margin concave medially, Posterior propodeal projection pointed downwards, with sparse setae (
<figureCitation id="B0A08185FFD0FFE6FC88F9C9FCD0C0D5" box="[828,890,1660,1685]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1850,1873]" captionTargetBox="[159,1418,213,1777]" captionTargetId="figure@2.[143,1431,199,1792]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Bright field images of Chakra sarvatra sp. nov. 1 — Habitus; 2 — Head, dorsal view; 3 — Head and mesosoma, dorsolateral view; 4 — Mesosoma, dorsal view 5 — Head, frontal view 6 — Mouthparts." httpUri="https://zenodo.org/record/225566/files/figure.png" pageId="1" pageNumber="286">Fig.4</figureCitation>
); Tibial spur formula 1:2:2. Fore wing extending beyond tip of metasoma in flexed position; Fore wing submarginal vein not reaching anterior margin and not curved posteriorly or broken before fore wing marginal vein; Fore wing marginal vein shorter than fore wing stigmal vein and fore wing postmarginal vein; Fore wing stigmal vein placed at a broad angle of 45 0 from fore wing margin (
<figureCitation id="B0A08185FFD0FFE6FF42F8BBFE9FC167" box="[246,309,1806,1831]" captionStart="FIGURES 7 – 11" captionStartId="3.[151,264,1624,1647]" captionTargetBox="[190,1404,205,1592]" captionTargetId="figure@3.[175,1411,193,1604]" captionTargetPageId="3" captionText="FIGURES 7 – 11. Bright field images of Chakra sarvatra sp. nov. 7 — Male antennae and head; 8 — Female antennae showing rows of white sensilla; 9 — Fore wing; 10 — Ovipositor; 11 — Setigerous tubercles on head and mesosoma." httpUri="https://zenodo.org/record/225567/files/figure.png" pageId="1" pageNumber="286">Fig.9</figureCitation>
); other fore wing veins absent; Marginal ciliae present;
<collectingCountry id="508CDD90FFD0FFE6FC18F8BBFC4EC167" box="[940,996,1806,1831]" name="India" pageId="1" pageNumber="286">Hind</collectingCountry>
wing with submarginal vein complete.
<emphasis id="1AEF4112FFD0FFE6FF23F887FEBEC10B" bold="true" box="[151,276,1842,1867]" pageId="1" pageNumber="286">Metasoma</emphasis>
pedunculate or spoon shaped (
<figureCitation id="B0A08185FFD0FFE6FD37F887FD62C10B" box="[643,712,1842,1867]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1850,1873]" captionTargetBox="[159,1418,213,1777]" captionTargetId="figure@2.[143,1431,199,1792]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Bright field images of Chakra sarvatra sp. nov. 1 — Habitus; 2 — Head, dorsal view; 3 — Head and mesosoma, dorsolateral view; 4 — Mesosoma, dorsal view 5 — Head, frontal view 6 — Mouthparts." httpUri="https://zenodo.org/record/225566/files/figure.png" pageId="1" pageNumber="286">Fig. 1</figureCitation>
), widest at its apical two-third, nearly as wide as mesosoma, shorter than head and mesosoma combined; Metasomal tergites transverse, T1 without horn; T1 and T2 costate, costae reaching posterior margin of tergite; T3 longest and widest; 7 tergites and 8 mediosternites in both sexes; Laterotergites narrow; Submarginal ridge well impressed; T6 visible as a thin strip with lower margin emarginated; T7 represented by a short lobe, external and not exerted with ovipositor.
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="6081CE8BFFD2FFE4FF23F96BFEB2C159" pageId="3" pageNumber="288" type="nomenclature">
<paragraph id="28249D00FFD2FFE4FF23F96BFCE3C0B8" blockId="3.[151,841,1758,1818]" box="[151,841,1758,1784]" pageId="3" pageNumber="288">
<heading id="736C2A6CFFD2FFE4FF23F96BFCE3C0B8" bold="true" box="[151,841,1758,1784]" fontSize="11" level="1" pageId="3" pageNumber="288" reason="1">
<emphasis id="1AEF4112FFD2FFE4FF23F96BFCE3C0B8" bold="true" box="[151,841,1758,1784]" pageId="3" pageNumber="288">
<taxonomicName id="EF9BE683FFD2FFE4FF23F96BFD74C0B8" authority="Rajmohana and Veenakumari" authorityName="Rajmohana and Veenakumari" box="[151,734,1758,1784]" class="Hexapoda" family="Platygastridae" genus="Chakra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="288" phylum="Arthropoda" rank="species" species="sarvatra" status="sp. nov.">
<emphasis id="1AEF4112FFD2FFE4FF23F96BFEF7C0B8" bold="true" box="[151,349,1758,1784]" italics="true" pageId="3" pageNumber="288">Chakra sarvatra</emphasis>
Rajmohana and Veenakumari
</taxonomicName>
,
<taxonomicNameLabel id="01DCFC69FFD2FFE4FD5FF96BFCE3C0B8" box="[747,841,1758,1784]" pageId="3" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="28249D00FFD2FFE4FF23F8B4FEB2C159" blockId="3.[151,841,1758,1818]" box="[151,280,1793,1818]" pageId="3" pageNumber="288">
(
<figureCitation id="B0A08185FFD2FFE4FF2BF8B4FEA5C159" box="[159,271,1793,1818]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 7 – 11" captionStartId-0="2.[151,264,1850,1873]" captionStartId-1="3.[151,264,1624,1647]" captionTargetBox-0="[159,1418,213,1777]" captionTargetBox-1="[190,1404,205,1592]" captionTargetId-0="figure@2.[143,1431,199,1792]" captionTargetId-1="figure@3.[175,1411,193,1604]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 – 6. Bright field images of Chakra sarvatra sp. nov. 1 — Habitus; 2 — Head, dorsal view; 3 — Head and mesosoma, dorsolateral view; 4 — Mesosoma, dorsal view 5 — Head, frontal view 6 — Mouthparts." captionText-1="FIGURES 7 – 11. Bright field images of Chakra sarvatra sp. nov. 7 — Male antennae and head; 8 — Female antennae showing rows of white sensilla; 9 — Fore wing; 10 — Ovipositor; 11 — Setigerous tubercles on head and mesosoma." httpUri-0="https://zenodo.org/record/225566/files/figure.png" httpUri-1="https://zenodo.org/record/225567/files/figure.png" pageId="3" pageNumber="288">Figs 1–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6081CE8BFFD2FFE3FF23F8FCFF46C420" lastPageId="4" lastPageNumber="289" pageId="3" pageNumber="288" type="materials_examined">
<paragraph id="28249D00FFD2FFE4FF23F8FCFEC2C122" blockId="3.[151,1437,1865,2034]" box="[151,360,1865,1890]" pageId="3" pageNumber="288">
<typeStatus id="F72023A2FFD2FFE4FF23F8FCFEAEC122" box="[151,260,1865,1890]" pageId="3" pageNumber="288" type="holotype">Holotype</typeStatus>
: Female.
</paragraph>
<paragraph id="28249D00FFD2FFE3FF73F8D8FF46C420" blockId="3.[151,1437,1865,2034]" lastBlockId="4.[151,1437,151,1282]" lastPageId="4" lastPageNumber="289" pageId="3" pageNumber="288">
<emphasis id="1AEF4112FFD2FFE4FF73F8D8FEF0C1C6" bold="true" box="[199,346,1901,1926]" pageId="3" pageNumber="288">Body length</emphasis>
=
<quantity id="EF6330E5FFD2FFE4FEC9F8D8FD8CC1C5" box="[381,550,1901,1926]" metricMagnitude="-3" metricUnit="m" metricValue="1.155" metricValueMax="1.25" metricValueMin="1.06" pageId="3" pageNumber="288" unit="mm" value="1.155" valueMax="1.25" valueMin="1.06">1.06–1.25 mm</quantity>
.
<emphasis id="1AEF4112FFD2FFE4FD80F8D8FDD7C1C6" bold="true" box="[564,637,1901,1926]" pageId="3" pageNumber="288">Color.</emphasis>
Head, mesonotum, T3–T6 blackish brown; T3 pale medially; Lower frons, lower gena, metascutellum, propodeum, lateral mesosoma, T1, anterior margin of T2 and legs including coxae yellowish brown; Antenna yellowish brown except blackish brown clava; Antennal setae white; Interocellar area, posterior margin of mesoscutellum and lateral T2 darker than rest of the body; Mandible pale with edges of teeth dark brown; Wings hyaline, veins brown.
<emphasis id="1AEF4112FFD5FFE3FD71FF22FCA6C6F0" bold="true" box="[709,780,151,176]" pageId="4" pageNumber="289">Head.</emphasis>
FCI=1.27; LCI=1.41; EL:EW=13:8.5, eyes placed wide apart on frons, inter orbital space 1.9 × eye height; Frons with two vertical rows of 3 setigerous tubercles medially above torular triangle from which small strips of striae radiate; OOL:POL:LOL =74:67:36; Radicle 0.18 × length of A1; Clava with six antennomeres; A3–A4, A7–A12 with four rows of sensilla; clava 1.2 × as long as A1; Ratio of length and width of antennomeres from A1 to A6 = 19.6:4.3, 4.7:3.5, 4.7:3.8, 3.8:3.7, 2.2:3, 2.2:3 and clava = 22.3:7.
<emphasis id="1AEF4112FFD5FFE3FF58FEF9FEDBC725" bold="true" box="[236,369,332,357]" pageId="4" pageNumber="289">Mesosoma.</emphasis>
(ML: MW = 34.7:34.4); FWL: FWW = 73.7:26.4; HWL: HWW = 64.3:11.4); 12–13 long and stiff brown bristles present along entire length of subcosta; Ratio of length of marginal vein: stigmal vein: postmarginal vein of fore wing = 3.9: 8.4:13.3.
<collectingCountry id="508CDD90FFD5FFE3FD75FE21FD53C7ED" box="[705,761,404,429]" name="India" pageId="4" pageNumber="289">Hind</collectingCountry>
wing cilia 0.57× width of hind wing.
<emphasis id="1AEF4112FFD5FFE3FB0CFE21FA96C7ED" bold="true" box="[1208,1340,404,429]" pageId="4" pageNumber="289">Metasoma.</emphasis>
(L:W = 55.3:33); T1 and T2 costate, costae reaching posterior margin of tergites; T2 foveolate on anterior margin; T3 smooth with faint traces of striae confined anterolaterally and with a light brown convex band on posterior end; T3 with an irregular row of setae on posterior margin; T4–T5 with dense setae relative to other tergites; T6 and T7 visible externally; Ratio of length and width of T1–T7 = 7.5:12.7, 12:26.4, 22.5:30.7, 3.4:28.9, 5.3:23.5, 2.0:17.1, 2.6:3.7.
</paragraph>
</subSubSection>
<subSubSection id="6081CE8BFFD5FFE3FF73FDD9FB1AC48D" pageId="4" pageNumber="289" type="description">
<paragraph id="28249D00FFD5FFE3FF73FDD9FB1AC48D" blockId="4.[151,1437,151,1282]" pageId="4" pageNumber="289">
<emphasis id="1AEF4112FFD5FFE3FF73FDD9FEA7C4C5" bold="true" box="[199,269,620,645]" pageId="4" pageNumber="289">Male:</emphasis>
Length =
<quantity id="EF6330E5FFD5FFE3FE3BFDD9FE45C4C4" box="[399,495,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="4" pageNumber="289" unit="mm" value="1.17">1.17mm</quantity>
. Similar to female except shape of antennomeres. Ratio of length and width of antennomeres A1–A12 = 16.5:3.9, 4.2:3.7, 5.3:4.0, 4.3:3.5, 5.0:3.3, 5.0:3.2, 5.5:3.2, 5.7:3.2, 5.7:3.2, 5.9:3.2, 5.9:3.2, 8.9:3.2; Antenna with short brownish-white setae and sparse long white setae (
<figureCitation id="B0A08185FFD5FFE3FBD1FD01FB0EC48D" box="[1125,1188,692,717]" captionStart="FIGURES 7 – 11" captionStartId="3.[151,264,1624,1647]" captionTargetBox="[190,1404,205,1592]" captionTargetId="figure@3.[175,1411,193,1604]" captionTargetPageId="3" captionText="FIGURES 7 – 11. Bright field images of Chakra sarvatra sp. nov. 7 — Male antennae and head; 8 — Female antennae showing rows of white sensilla; 9 — Fore wing; 10 — Ovipositor; 11 — Setigerous tubercles on head and mesosoma." httpUri="https://zenodo.org/record/225567/files/figure.png" pageId="4" pageNumber="289">Fig.7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6081CE8BFFD5FFE3FF73FD62FC4FC555" pageId="4" pageNumber="289" type="etymology">
<paragraph id="28249D00FFD5FFE3FF73FD62FC4FC555" blockId="4.[151,1437,151,1282]" pageId="4" pageNumber="289">
<emphasis id="1AEF4112FFD5FFE3FF73FD62FEE4C4B0" bold="true" box="[199,334,727,752]" pageId="4" pageNumber="289">Etymology.</emphasis>
The species is named ‘
<taxonomicName id="EF9BE683FFD5FFE3FDC7FD6CFD78C4B0" box="[627,722,729,752]" class="Hexapoda" family="Platygastridae" genus="Chakra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="sarvatra">
<emphasis id="1AEF4112FFD5FFE3FDC7FD6CFD78C4B0" box="[627,722,729,752]" italics="true" pageId="4" pageNumber="289">sarvatra</emphasis>
</taxonomicName>
’ meaning ‘everywhere’ in Sanskrit, due to the presence of extensive tuberculate sculpturing on the head and mesosoma of the species.
</paragraph>
</subSubSection>
<subSubSection id="6081CE8BFFD5FFE3FF73FCAAFC6FC341" pageId="4" pageNumber="289" type="materials_examined">
<paragraph id="28249D00FFD5FFE3FF73FCAAFE4EC2F4" blockId="4.[151,1437,151,1282]" pageId="4" pageNumber="289">
<emphasis id="1AEF4112FFD5FFE3FF73FCAAFD25C578" bold="true" box="[199,655,799,824]" pageId="4" pageNumber="289">
Material examined.
<typeStatus id="F72023A2FFD5FFE3FE03FCAAFD8CC578" box="[439,550,799,824]" pageId="4" pageNumber="289" type="holotype">Holotype</typeStatus>
, Female,
</emphasis>
(Reg.No.ZSI/
<collectionCode id="4E8A05C5FFD5FFE3FC9FFC95FCD5C578" box="[811,895,800,824]" httpUri="http://grbio.org/cool/z6tk-8dmw" name="Western Ghat Regional Station of the Zoological Survey of India at Calicut" pageId="4" pageNumber="289">WGRS</collectionCode>
/IR.INV. 2656),
<collectingCountry id="508CDD90FFD5FFE3FB81FC95FB2EC578" box="[1077,1156,800,824]" name="India" pageId="4" pageNumber="289">INDIA</collectingCountry>
: South Andaman Island, Garacharma, Central Agricultural Research Institute, yellow pan trap,
<date id="5C25BBC0FFD5FFE3FC12FCF3FBB0C51F" box="[934,1050,838,863]" pageId="4" pageNumber="289" value="2012-02-22">22.ii.2012</date>
, 11 0 61’37.48’’ N, 92 0 71’63.79’’ E.
<typeStatus id="F72023A2FFD5FFE3FF23FCDEFEA3C5C3" box="[151,265,875,899]" pageId="4" pageNumber="289" type="paratype">Paratypes</typeStatus>
: (Reg.No.ZSI/
<collectionCode id="4E8A05C5FFD5FFE3FE1AFCDEFDA8C5C3" box="[430,514,875,899]" httpUri="http://grbio.org/cool/z6tk-8dmw" name="Western Ghat Regional Station of the Zoological Survey of India at Calicut" pageId="4" pageNumber="289">WGRS</collectionCode>
/IR.INV. 2657),
<collectingCountry id="508CDD90FFD5FFE3FD7DFCDEFCB2C5C3" box="[713,792,875,899]" name="India" pageId="4" pageNumber="289">INDIA</collectingCountry>
: South Andaman Island: Sippighat, yellow pan trap,
<date id="5C25BBC0FFD5FFE3FF23FC24FEA1C5EA" box="[151,267,913,938]" pageId="4" pageNumber="289" value="2012-02-23">23.ii.2012</date>
, 11 0 60’71.20” N, 92 0 66’90.57’’ E, male; (Reg. No.
<collectionCode id="4E8A05C5FFD5FFE3FC30FC27FC61C5EA" box="[900,971,914,938]" pageId="4" pageNumber="289">ICAR</collectionCode>
/
<collectionCode id="4E8A05C5FFD5FFE3FC67FC27FB8EC5EA" box="[979,1060,914,938]" pageId="4" pageNumber="289">NBAII</collectionCode>
/P149), South Andaman Island: Sippighat, 11 0 60’71.20” N, 92 0 66’90.57’’ E, yellow pan trap,
<date id="5C25BBC0FFD5FFE3FCDCFC0DFC76C591" box="[872,988,952,977]" pageId="4" pageNumber="289" value="2012-02-23">23.ii.2012</date>
, female; (Reg.No.
<collectionCode id="4E8A05C5FFD5FFE3FB18FC0DFB59C591" box="[1196,1267,952,977]" pageId="4" pageNumber="289">ICAR</collectionCode>
/
<collectionCode id="4E8A05C5FFD5FFE3FB48FC0DFAE4C591" box="[1276,1358,952,977]" pageId="4" pageNumber="289">NBAII</collectionCode>
/P150), South Andaman Island: Sippighat, 11 0 60’71.20” N, 92 0 66’90.57’’ E, yellow pan trap,
<date id="5C25BBC0FFD5FFE3FB0BFC6AFA99C5B8" box="[1215,1331,991,1016]" pageId="4" pageNumber="289" value="2012-02-23">23.ii.2012</date>
, female; (Reg.No.
<collectionCode id="4E8A05C5FFD5FFE3FF4CFBB6FEEAC25C" box="[248,320,1027,1052]" pageId="4" pageNumber="289">ICAR</collectionCode>
/
<collectionCode id="4E8A05C5FFD5FFE3FEFDFBB6FE31C25C" box="[329,411,1027,1052]" pageId="4" pageNumber="289">NBAII</collectionCode>
/P151), South Andaman Island: Garacharma, Central Agricultural Research Institute, 11 0 61’37.48’’ N, 92 0 71’63.79’’ E, yellow pan trap,
<date id="5C25BBC0FFD5FFE3FD66FB9FFC94C203" box="[722,830,1066,1091]" pageId="4" pageNumber="289" value="2013-01-26">26.i.2013</date>
, male; (Reg.No.
<collectionCode id="4E8A05C5FFD5FFE3FC44FB9FFB9DC203" box="[1008,1079,1066,1091]" pageId="4" pageNumber="289">ICAR</collectionCode>
/
<collectionCode id="4E8A05C5FFD5FFE3FBF4FB9FFB38C203" box="[1088,1170,1066,1091]" pageId="4" pageNumber="289">NBAII</collectionCode>
/P152), South Andaman Island: Ferrargunj, 11 0 72’19.17’’ N, 92 0 65’63.51’’ E, sweep net,
<date id="5C25BBC0FFD5FFE3FC35FBE5FC5FC229" box="[897,1013,1104,1129]" pageId="4" pageNumber="289" value="2012-02-22">22.ii.2012</date>
, male; (Reg.No.
<collectionCode id="4E8A05C5FFD5FFE3FB19FBE5FB5FC229" box="[1197,1269,1104,1129]" pageId="4" pageNumber="289">ICAR</collectionCode>
/
<collectionCode id="4E8A05C5FFD5FFE3FB49FBE5FAE5C229" box="[1277,1359,1104,1129]" pageId="4" pageNumber="289">NBAII</collectionCode>
/P153), South Andaman Island: Sippighat, 11 0 60’71.20” N, 92 0 66’90.57’’ E, Malaise trap,
<date id="5C25BBC0FFD5FFE3FB89FBC2FB03C2D0" box="[1085,1193,1143,1168]" pageId="4" pageNumber="289" value="2013-01-31">31.i.2013</date>
, male; All specimens collected by K. Veenakumari.
</paragraph>
<paragraph id="28249D00FFD5FFE3FF73FB76FC6FC341" blockId="4.[151,1437,151,1282]" pageId="4" pageNumber="289">
(Reg.No.ZSI/
<collectionCode id="4E8A05C5FFD5FFE3FEEFFB76FE05C29B" box="[347,431,1219,1243]" httpUri="http://grbio.org/cool/z6tk-8dmw" name="Western Ghat Regional Station of the Zoological Survey of India at Calicut" pageId="4" pageNumber="289">WGRS</collectionCode>
/IR.INV. 2658)
<collectingCountry id="508CDD90FFD5FFE3FDDAFB77FD06C29B" box="[622,684,1218,1243]" name="India" pageId="4" pageNumber="289">India</collectingCountry>
: Kerala:
<collectingCountry id="508CDD90FFD5FFE3FCAAFB77FCD9C29B" box="[798,883,1218,1243]" name="India" pageId="4" pageNumber="289">Calicut</collectingCountry>
: Peruvayal (Paddy ecosystem), 11 0 15’178” N, 75 0 66’54.237’’ E, Malaise trap,
<date id="5C25BBC0FFD5FFE3FE49FB5CFDF1C342" box="[509,603,1257,1282]" pageId="4" pageNumber="289" value="2009-01-02">2.i.2009</date>
, female, coll. by Rajmohana, K.
</paragraph>
</subSubSection>
</treatment>
</document>

141
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<taxonomicName id="EF9BB7C3FFD2FFC0FEB0FD4ED8D6E08E" ID-CoL="6SWLV" authority="Tian &amp; Deuve" authorityName="Tian &amp; Deuve" authorityYear="2003" box="[257,656,646,670]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="thaicus">
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<emphasis id="1AEF1052FFD2FFC0FEB0FD4EDBA3E08E" bold="true" box="[257,485,646,670]" italics="true" pageId="4" pageNumber="265">Orthogonius thaicus</emphasis>
Tian &amp; Deuve
</emphasis>
</taxonomicName>
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<emphasis id="1AEF1052FFD2FFC0FF38FD0DDB09E0CA" box="[137,335,709,730]" italics="true" pageId="4" pageNumber="265">Orthogonius thaicus</emphasis>
Tian &amp; Deuve, 2003a: 46
</taxonomicName>
.
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. –
</emphasis>
<materialsCitation id="98F3C61DFFD2FFC0FED7FD36DB7EE15D" collectingDate="2005-04-19" collectionCode="R" collectorName="P. Grootaert" country="Singapore" location="Nee Soon" municipality="Station" pageId="4" pageNumber="265" specimenCount="2" specimenCount-male="2">
<specimenCount id="3E9D07C9FFD2FFC0FED7FD36DBF5E103" box="[358,435,766,787]" pageId="4" pageNumber="265" type="male">2 males</specimenCount>
, “Coll. I.
<collectionCode id="4E8A5485FFD2FFC0FDA4FD36D86EE103" box="[533,552,766,787]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="265">R</collectionCode>
. Sc. N. B. /
<collectingCountry id="508C8CD0FFD2FFC0FD10FD36D943E103" box="[673,773,766,787]" name="Singapore" pageId="4" pageNumber="265">Singapore</collectingCountry>
,
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, Mal. trap 2, swamp forest,
<collectingMunicipality id="C840563AFFD2FFC0FE45FCD3D87EE120" box="[500,568,795,816]" pageId="4" pageNumber="265">Station</collectingMunicipality>
25068.
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<collectingDate id="4C611368FFD2FFC0FD37FCD3D943E120" box="[646,773,795,816]" pageId="4" pageNumber="265" value="2005-04-19">19 Apr. 2005</collectingDate>
</date>
.
<collectorName id="856EA996FFD2FFC0FF38FCF0DABAE15D" box="[137,252,824,845]" pageId="4" pageNumber="265">P. Grootaert</collectorName>
coll.”
</materialsCitation>
;
<materialsCitation id="98F3C61DFFD2FFC0FEF3FCF0DB07E1D0" collectingDate="2005-04-19" collectionCode="R, IRSNB, MNHN" collectorName="P. Grootaert &amp; All &amp; Tian &amp; Deuve" country="Singapore" county="Station" location="Nee Soon" municipality="Swamp forest" pageId="4" pageNumber="265" specimenCount="2" specimenCount-male="2">
<specimenCount id="3E9D07C9FFD2FFC0FEF3FCF0DBC8E15D" box="[322,398,824,845]" pageId="4" pageNumber="265" type="male">2 males</specimenCount>
, “Coll. I.
<collectionCode id="4E8A5485FFD2FFC0FE5CFCF0D846E15D" box="[493,512,824,845]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="265">R</collectionCode>
. Sc. N. B. /
<collectingCountry id="508C8CD0FFD2FFC0FDC5FCF0D89EE15D" box="[628,728,824,845]" name="Singapore" pageId="4" pageNumber="265">Singapore</collectingCountry>
,
<location id="2D449A9BFFD2FFC0FD50FCF0DAFBE179" LSID="urn:lsid:plazi:treatment:A0327D56FFD2FFC0FEB0FD4EDBFEE650:2D449A9BFFD2FFC0FD50FCF0DAFBE179" country="Singapore" county="Station" municipality="Swamp forest" name="Nee Soon" pageId="4" pageNumber="265">Nee Soon</location>
, Mal. trap 2,
<collectingMunicipality id="C840563AFFD2FFC0FEF3FC9CDB8DE179" box="[322,459,852,873]" pageId="4" pageNumber="265">Swamp forest</collectingMunicipality>
,
<collectingCounty id="C145B4CCFFD2FFC0FE67FC9CD85CE179" box="[470,538,852,873]" pageId="4" pageNumber="265">Station</collectingCounty>
25028.
<date id="5C25EA80FFD2FFC0FDDBFC9CD8ABE179" box="[618,749,852,873]" pageId="4" pageNumber="265" value="2005-04-19">
<collectingDate id="4C611368FFD2FFC0FDDBFC9CD8ABE179" box="[618,749,852,873]" pageId="4" pageNumber="265" value="2005-04-19">19 Apr. 2005</collectingDate>
</date>
.
<collectorName id="856EA996FFD2FFC0FD46FC9CDAA0E196" pageId="4" pageNumber="265">P. Grootaert</collectorName>
coll.” deposited in
<collectionCode id="4E8A5485FFD2FFC0FE01FCB9DBB1E196" box="[432,503,881,902]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="4" pageNumber="265">IRSNB</collectionCode>
except for one in
<collectionCode id="4E8A5485FFD2FFC0FD09FCB9D943E196" box="[696,773,881,902]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="265">MNHN</collectionCode>
.
<collectorName id="856EA996FFD2FFC0FF38FC46DAEEE1B3" box="[137,168,910,931]" pageId="4" pageNumber="265">All</collectorName>
the
<typeStatus id="F72072E2FFD2FFC0FF68FC46DB45E1B3" box="[217,259,910,931]" pageId="4" pageNumber="265">type</typeStatus>
materials were also studied (for detail see
<collectorName id="856EA996FFD2FFC0FD71FC46D8AAE1B3" box="[704,748,910,931]" pageId="4" pageNumber="265">Tian</collectorName>
&amp;
<collectorName id="856EA996FFD2FFC0FF38FC63DA8CE1D0" box="[137,202,939,960]" pageId="4" pageNumber="265">Deuve</collectorName>
, 2003a: 47)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="60819FCBFFD2FFC0FF38FC21DBFEE650" pageId="4" pageNumber="265" type="distribution">
<paragraph id="2824CC40FFD2FFC0FF38FC21DBFEE650" blockId="4.[137,776,1001,1088]" pageId="4" pageNumber="265">
<emphasis id="1AEF1052FFD2FFC0FF38FC21DB6BE611" bold="true" box="[137,301,1001,1025]" pageId="4" pageNumber="265">
<emphasis id="1AEF1052FFD2FFC0FF38FC21DB49E611" bold="true" box="[137,271,1001,1025]" italics="true" pageId="4" pageNumber="265">Distribution</emphasis>
. –
</emphasis>
This species was described from southern
<collectingCountry id="508C8CD0FFD2FFC0FF38FBC1DAAAE630" box="[137,236,1033,1056]" name="Thailand" pageId="4" pageNumber="265">Thailand</collectingCountry>
(Tian &amp; Deuve, 2003a), and is recorded from
<collectingCountry id="508C8CD0FFD2FFC0FF38FBE1DAB3E650" box="[137,245,1065,1088]" name="Singapore" pageId="4" pageNumber="265">Singapore</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>

205
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@ -0,0 +1,205 @@
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<mods:title id="61ECA93189F8775516781B0C2A2BA786">A Taxonomic Treatment Of The Species Of Orthogonius Macleay (Coleoptera: Carabidae: Orthogoniini) Of Singapore</mods:title>
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<paragraph id="2824CC40FFD2FFC0FF61F9FED887E45E" blockId="4.[208,705,1590,1614]" box="[208,705,1590,1614]" pageId="4" pageNumber="265">
<heading id="736C7B2CFFD2FFC0FF61F9FED887E45E" bold="true" box="[208,705,1590,1614]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="265" reason="2">
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Key to species of
<taxonomicName id="EF9BB7C3FFD2FFC0FE2AF9FED860E45E" ID-CoL="8KY5F" authorityName="Macleay" authorityYear="1825" box="[411,550,1590,1614]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="1AEF1052FFD2FFC0FE2AF9FED860E45E" bold="true" box="[411,550,1590,1614]" italics="true" pageId="4" pageNumber="265">Orthogonius</emphasis>
</taxonomicName>
of
<collectingCountry id="508C8CD0FFD2FFC0FDFFF9FED887E45E" box="[590,705,1590,1614]" name="Singapore" pageId="4" pageNumber="265">Singapore</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="60819FCBFFD2FFC0FF38F9BCDF84E199" pageId="4" pageNumber="265" type="key">
<keyStep id="936FD4E5FFD2FFC0FF38F9BCD941E4B6" pageId="4" pageNumber="265">
<paragraph id="2824CC40FFD2FFC0FF38F9BCD941E499" blockId="4.[137,775,1652,1702]" box="[137,775,1652,1673]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FF38F9BCD941E499" box="[137,775,1652,1673]" pageId="4" pageNumber="265">1. Clypeus quadrisetose ........................................................................ 2</keyLead>
</paragraph>
<paragraph id="2824CC40FFD2FFC0FF38F959D941E4B6" blockId="4.[137,775,1652,1702]" box="[137,775,1681,1702]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FF38F959D941E4B6" box="[137,775,1681,1702]" pageId="4" pageNumber="265">– Clypeus bisetose ............................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="936FD4E5FFD2FFC0FF38F903D941E509" pageId="4" pageNumber="265">
<paragraph id="2824CC40FFD2FFC0FF38F903D941E4ED" blockId="4.[137,777,1739,1817]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FF38F903D941E4ED" pageId="4" pageNumber="265">
2. Body rather flat and broad, reddish-brown, large (Length 20.0–21.0 mm) .............................................................................
<taxonomicName id="EF9BB7C3FFD2FFC0FD32F920D941E4ED" ID-CoL="6T99V" authority="Gray" authorityName="Gray, Indes Orient, C. G. Gray" authorityYear="1832" box="[643,775,1768,1789]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="hopei">
<emphasis id="1AEF1052FFD2FFC0FD32F920D894E4ED" box="[643,722,1768,1789]" italics="true" pageId="4" pageNumber="265">O. hopei</emphasis>
Gray
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2824CC40FFD2FFC0FF38F8CCD941E509" blockId="4.[137,777,1739,1817]" box="[137,775,1796,1817]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FF38F8CCD941E509" box="[137,775,1796,1817]" pageId="4" pageNumber="265">– Body more convex and slender, dark brown, medium sized ...... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="936FD4E5FFD2FFC0FF38F8F6D940E5B9" pageId="4" pageNumber="265">
<paragraph id="2824CC40FFD2FFC0FF38F8F6D941E560" blockId="4.[137,775,1854,1962]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FF38F8F6D941E560" pageId="4" pageNumber="265">
3. Elytral striae shallow, intervals flat. Length 14.0 mm .................... .......................................................................
<taxonomicName id="EF9BB7C3FFD2FFC0FD83F893D941E560" ID-CoL="6T8ZZ" authority="Andrewes" authorityName="Andrewes" authorityYear="1926" box="[562,775,1883,1904]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="saundersi">
<emphasis id="1AEF1052FFD2FFC0FD83F893D8E0E560" box="[562,678,1883,1904]" italics="true" pageId="4" pageNumber="265">O. saundersi</emphasis>
Andrewes
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2824CC40FFD2FFC0FF38F8B0D940E5B9" blockId="4.[137,775,1854,1962]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FF38F8B0D940E5B9" pageId="4" pageNumber="265">
– Elytral striae deep, intervals convex. Length
<quantity id="EF6361A5FFD2FFC0FDF4F8B0D8DEE59D" box="[581,664,1912,1933]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="4" pageNumber="265" unit="mm" value="14.5">14.5 mm</quantity>
................... ...................................................................
<taxonomicName id="EF9BB7C3FFD2FFC0FDAEF85DD940E5B9" ID-CoL="6SWLV" authority="Tian &amp; Deuve" authorityName="Tian &amp; Deuve" authorityYear="2003" box="[543,774,1940,1962]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="thaicus">
<emphasis id="1AEF1052FFD2FFC0FDAEF85DD83BE5BA" box="[543,637,1941,1962]" italics="true" pageId="4" pageNumber="265">O. thaicus</emphasis>
Tian &amp; Deuve
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="936FD4E5FFD2FFC0FF38F806D941EA29" pageId="4" pageNumber="265">
<paragraph id="2824CC40FFD2FFC0FF38F806D941EA10" blockId="4.[137,777,1998,2105]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FF38F806D941EA10" pageId="4" pageNumber="265">4. Ligula with glossal sclerite quadrisetose at apex, mentum with a median tooth, hind angle of pronotum angular ............................ 5</keyLead>
</paragraph>
<paragraph id="2824CC40FFD2FFC0FF38F7C0D941EA29" blockId="4.[137,777,1998,2105]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FF38F7C0D941EA29" pageId="4" pageNumber="265">– Glossal sclerite bisetose at apex, mentum without median tooth, hind angle of pronotum rounded .................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="936FD4E5FFD2FFC0FCF5FF09DF87E376" pageId="4" pageNumber="265">
<paragraph id="2824CC40FFD2FFC0FCF5FF09DF87E300" blockId="4.[836,1477,193,358]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FCF5FF09DF87E300" pageId="4" pageNumber="265">
5. Head impunctate, labrum straight at front, interval 6 of elytra impunctate near base, smaller. Length
<quantity id="EF6361A5FFD2FFC0FB7BFF16DF79E2E3" box="[1226,1343,222,243]" metricMagnitude="-2" metricUnit="m" metricValue="1.375" metricValueMax="1.4" metricValueMin="1.35" pageId="4" pageNumber="265" unit="mm" value="13.75" valueMax="14.0" valueMin="13.5">13.5–14 mm</quantity>
....................... ........................................................
<taxonomicName id="EF9BB7C3FFD2FFC0FB2EFF33DF87E300" ID-CoL="758M3" authority="Tian &amp; Deuve" authorityName="Tian &amp; Deuve" authorityYear="2006" box="[1183,1473,251,272]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="singaporensis">
<emphasis id="1AEF1052FFD2FFC0FB2EFF33DF7EE300" box="[1183,1336,251,272]" italics="true" pageId="4" pageNumber="265">O. singaporensis</emphasis>
Tian &amp; Deuve
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2824CC40FFD2FFC0FCF5FED0DF87E376" blockId="4.[836,1477,193,358]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FCF5FED0DF87E376" pageId="4" pageNumber="265">
– Head sparsely punctate on vertex, labrum slightly emarginate at front, interval 6 of elytra with a few punctures near base, larger. Length 16.0–
<quantity id="EF6361A5FFD2FFC0FC54FE99DE70E376" box="[997,1078,337,358]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="4" pageNumber="265" unit="mm" value="17.5">17.5 mm</quantity>
.......................
<taxonomicName id="EF9BB7C3FFD2FFC0FB70FE99DF87E376" ID-CoL="6SWXT" authority="Tian &amp; Deuve" authorityName="Tian &amp; Deuve" authorityYear="2006" box="[1217,1473,337,358]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="fairmairei">
<emphasis id="1AEF1052FFD2FFC0FB70FE99DF7EE376" box="[1217,1336,337,358]" italics="true" pageId="4" pageNumber="265">O. fairmairei</emphasis>
Tian &amp; Deuve
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="936FD4E5FFD2FFC0FCF5FE43DF84E3AD" pageId="4" pageNumber="265">
<paragraph id="2824CC40FFD2FFC0FCF5FE43DF84E3B0" blockId="4.[836,1474,395,445]" box="[836,1474,395,416]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FCF5FE43DF84E3B0" box="[836,1474,395,416]" pageId="4" pageNumber="265">6. Head punctate, interval 7 of elytra more or less carinate ............ 7</keyLead>
</paragraph>
<paragraph id="2824CC40FFD2FFC0FCF5FE60DF84E3AD" blockId="4.[836,1474,395,445]" box="[836,1474,424,445]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FCF5FE60DF84E3AD" box="[836,1474,424,445]" pageId="4" pageNumber="265">– Head impunctate, interval 7 of elytra not carinate ....................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="936FD4E5FFD2FFC0FCF5FE29DF87E020" pageId="4" pageNumber="265">
<paragraph id="2824CC40FFD2FFC0FCF5FE29DF84E3E6" blockId="4.[836,1474,481,560]" box="[836,1474,481,502]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FCF5FE29DF84E3E6" box="[836,1474,481,502]" pageId="4" pageNumber="265">7. Head densely punctate, pronotum impunctate .............................. 8</keyLead>
</paragraph>
<paragraph id="2824CC40FFD2FFC0FCF5FE36DF87E020" blockId="4.[836,1474,481,560]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FCF5FE36DF87E020" pageId="4" pageNumber="265">
– Head and pronotum sparsely punctate. Length 12.0 mm .............. ...........................................................
<taxonomicName id="EF9BB7C3FFD2FFC0FB03FDD3DF87E020" ID-CoL="6T8LS" authority="Tian &amp; Deuve" authorityName="Tian &amp; Deuve" authorityYear="2003" box="[1202,1473,539,560]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="solidicornis">
<emphasis id="1AEF1052FFD2FFC0FB03FDD3DF71E020" box="[1202,1335,539,560]" italics="true" pageId="4" pageNumber="265">O. solidicornis</emphasis>
Tian &amp; Deuve
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="936FD4E5FFD2FFC0FCF5FD9CDF87E0CD" pageId="4" pageNumber="265">
<paragraph id="2824CC40FFD2FFC0FCF5FD9CDF84E096" blockId="4.[836,1476,596,733]" pageId="4" pageNumber="265">
<keyLead id="936A6F75FFD2FFC0FCF5FD9CDF84E096" pageId="4" pageNumber="265">
8. Labrum straight at front,the elytral base obscurely bordered,aedeagus stout. Length 15.0 mm .....................
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– Labrum slightly convex at front, the elytral base unbordered against intervals 1–3, and aedeagus slender in), and aedeagus stouter.Length 13.0 mm ......................................
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<emphasis id="1AEF1052FFD2FFC0FB24FD00DF7EE0CD" box="[1173,1336,712,733]" italics="true" pageId="4" pageNumber="265">O. andrewesianus</emphasis>
Tian &amp; Deuve
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9. Elytral striae shallow, intervals slightly convex, impunctate. Length 13.5–14.0 mm ..................................
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<emphasis id="1AEF1052FFD2FFC0FB07FCD6DF7EE123" box="[1206,1336,798,819]" italics="true" pageId="4" pageNumber="265">O. dongnanya</emphasis>
Tian &amp; Deuve
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– Elytral striae deep, intervals moderately convex; intervals 3, 5, 6 and 8 sparsely punctate, punctures small. Length
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.......... ..............................................................
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<emphasis id="1AEF1052FFD2FFC0FB73FCBDDF0EE19A" box="[1218,1352,885,906]" italics="true" pageId="4" pageNumber="265">O. cyclothorax</emphasis>
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126
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Tian &amp; Deuve
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<emphasis id="1AEF1052FFD2FFC0FF38FB00DBDFE6CD" box="[137,409,1224,1245]" italics="true" pageId="4" pageNumber="265">Orthogonius andrewesianus</emphasis>
Tian &amp; Deuve, 2006b: 109
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.
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. –
</emphasis>
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, labeled “Coll. I.
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. Sc. N. B. /
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coll.”, deposited in
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.
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the
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materials were also studied (for detail see
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&amp;
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, 2006b: 109)
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.
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<emphasis id="1AEF1052FFD2FFC0FF38FA5EDB48E7BE" bold="true" box="[137,270,1430,1454]" italics="true" pageId="4" pageNumber="265">Distribution</emphasis>
. –
</emphasis>
This species was described from
<collectingCountry id="508C8CD0FFD2FFC0FD17FA5ED94EE7BD" box="[678,776,1430,1453]" name="Malaysia" pageId="4" pageNumber="265">Malaysia</collectingCountry>
(
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) and
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(
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), and is recorded from
<collectingCountry id="508C8CD0FFD2FFC0FF38FA1EDAB3E7FD" box="[137,245,1494,1517]" name="Singapore" pageId="4" pageNumber="265">Singapore</collectingCountry>
for the first time.
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183
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<mods:identifier id="B7FAD855274362D23E266B8095310E46" type="DOI">10.5281/zenodo.5333241</mods:identifier>
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<heading id="736C7B2CFFD4FFC6FC1EFD89DF1FE049" bold="true" box="[943,1369,577,601]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="263" reason="2">
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<taxonomicName id="EF9BB7C3FFD4FFC6FC1EFD89DE82E049" ID-CoL="6SWY3" authority="Tian &amp; Deuve, 2007" authorityName="Tian &amp; Deuve" authorityYear="2007" box="[943,1220,577,601]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="cyclothorax" status="sp. nov.">
<emphasis id="1AEF1052FFD4FFC6FC1EFD89DE82E049" bold="true" box="[943,1220,577,601]" italics="true" pageId="2" pageNumber="263">Orthogonius cyclothorax</emphasis>
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,
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</paragraph>
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<subSubSection id="60819FCBFFD4FFC6FBF0FDAADE80E069" box="[1089,1222,610,633]" pageId="2" pageNumber="263" type="description">
<paragraph id="2824CC40FFD4FFC6FBF0FDAADE80E069" blockId="2.[943,1369,577,633]" box="[1089,1222,610,633]" pageId="2" pageNumber="263">
(
<figureCitation id="B0A0D0C5FFD4FFC6FBF8FDAADEF8E069" box="[1097,1214,610,633]" captionStart="Figs" captionStartId="3.[114,158,2036,2057]" captionTargetBox="[118,1453,1116,1998]" captionTargetId="figure-453@3.[85,1483,1085,2014]" captionTargetPageId="3" captionText="Figs. 7–11. Line drawings of structural features of Orthogonius cyclothorax, new species: 7, left fore tibia, dorsal view; 8, apex of right elytron; 9, abdominal ventrite VII of male; 10, aedeagal median lobe, right lateral view; 11, apical portion of aedeagal median lobe, dorsal view. Scale bars: 1 mm. A for Figs. 8–9; B for Figs. 7, 10–11." figureDoi="http://doi.org/10.5281/zenodo.5333245" httpUri="https://zenodo.org/record/5333245/files/figure.png" pageId="2" pageNumber="263">Figs. 7–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="60819FCBFFD4FFC6FCF5FD68DE8AE11C" pageId="2" pageNumber="263" type="materials_examined">
<paragraph id="2824CC40FFD4FFC6FCF5FD68DE8AE11C" blockId="2.[836,1476,672,780]" pageId="2" pageNumber="263">
<emphasis id="1AEF1052FFD4FFC6FCF5FD68D9AEE0A5" bold="true" box="[836,1000,672,693]" pageId="2" pageNumber="263">
<emphasis id="1AEF1052FFD4FFC6FCF5FD68D988E0A5" bold="true" box="[836,974,672,693]" italics="true" pageId="2" pageNumber="263">Type material</emphasis>
. –
</emphasis>
<materialsCitation id="98F3C61DFFD4FFC6FC41FD68DE8AE11C" collectingDate="2005-04-19" collectionCode="R, IRSNB" collectorName="P. Grootaert" country="Singapore" location="Swamp forest" pageId="2" pageNumber="263" specimenCount="1" typeStatus="holotype">
<typeStatus id="F72072E2FFD4FFC6FC41FD68DE0BE0A5" box="[1008,1101,672,693]" pageId="2" pageNumber="263" type="holotype">Holotype</typeStatus>
: male, labeled “Coll. I.
<collectionCode id="4E8A5485FFD4FFC6FA8AFD68DF08E0A5" box="[1339,1358,672,693]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="263">R</collectionCode>
. Sc. N. B. /
<collectingCountry id="508C8CD0FFD4FFC6FCF5FD75D9EEE0C2" box="[836,936,701,722]" name="Singapore" pageId="2" pageNumber="263">Singapore</collectingCountry>
, Nee Soon,
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2,
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, Station 25068.
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<collectingDate id="4C611368FFD4FFC6FCF5FD12D982E0FF" box="[836,964,730,751]" pageId="2" pageNumber="263" value="2005-04-19">19 Apr. 2005</collectingDate>
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.
<collectorName id="856EA996FFD4FFC6FC7CFD12DE79E0FF" box="[973,1087,730,751]" pageId="2" pageNumber="263">P. Grootaert</collectorName>
coll.”, collected at Nee Soon,
<collectingCountry id="508C8CD0FFD4FFC6FAD3FD12DF85E0FF" box="[1378,1475,730,751]" name="Singapore" pageId="2" pageNumber="263">Singapore</collectingCountry>
(type locality) and deposited in
<collectionCode id="4E8A5485FFD4FFC6FBCFFD3FDE8EE11C" box="[1150,1224,759,780]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="2" pageNumber="263">IRSNB</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="60819FCBFFD4FFC7FCF5FCFDD9FFE029" lastPageId="3" lastPageNumber="264" pageId="2" pageNumber="263" type="description">
<paragraph id="2824CC40FFD4FFC6FCF5FCFDDE01E17C" blockId="2.[836,1475,820,876]" pageId="2" pageNumber="263">
<emphasis id="1AEF1052FFD4FFC6FCF5FCFDD98DE15C" bold="true" box="[836,971,820,845]" pageId="2" pageNumber="263">
<emphasis id="1AEF1052FFD4FFC6FCF5FCFDD985E15D" bold="true" box="[836,963,821,845]" italics="true" pageId="2" pageNumber="263">Description</emphasis>
.
</emphasis>
Body: Length
<quantity id="EF6361A5FFD4FFC6FBC0FCFDDE93E15C" box="[1137,1237,821,844]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="263" unit="mm" value="10.5">10.5 mm</quantity>
; width
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. Form stout, moderately shiny.
</paragraph>
<paragraph id="2824CC40FFD4FFC6FCF5FC5DDF63E1FC" blockId="2.[836,1475,917,1004]" pageId="2" pageNumber="263">Color: Head dark brown, pronotum and elytra reddish brown; legs (except femora), ventral surface, palpi of mouthparts, antennae and labrum brown; femora yellow.</paragraph>
<paragraph id="2824CC40FFD4FFC6FCF5FBDDD9B0E67C" blockId="2.[836,1475,1045,1132]" pageId="2" pageNumber="263">Macrosculpture: Head and pronotum shallowly striate, impunctate, intervals 3, 5, 6 and 8 of elytra with sparse, small punctures.</paragraph>
<paragraph id="2824CC40FFD4FFC6FCF5FB5DDE38E6DC" blockId="2.[836,1475,1173,1228]" pageId="2" pageNumber="263">Microsculpture: Mesh pattern isodiametric on head, pronotum and elytra; microlines dense.</paragraph>
<paragraph id="2824CC40FFD4FFC6FCF5FB3DDE3AE7DC" blockId="2.[836,1476,1269,1484]" pageId="2" pageNumber="263">Head: Slightly narrower than long (HW/HL = 0.92), stout, evidently convex. Frontal impressions pit-like, small; clypeus bisetose, surface quite even. Eyes very large, prominent. Antennae rather long, extending to about one-fifths of elytra from base; pubescent from apical two-thirds of antennomere 4; antennomere 4 as long as antennomere 3, but less than two times of antennomere 2.</paragraph>
<paragraph id="2824CC40FFD4FFC6FCF5FA3DDFE1E4FC" blockId="2.[836,1476,1525,1772]" pageId="2" pageNumber="263">Mouthparts: Labrum six setose, anterior margin distinctly emarginate. Maxillae: palpi subcylindrical, palpomeres 3 and 4 subequal; palpomere 3 with several short setae at apex. Labium: submentum with one seta on either side; mentum without median tooth, asetose; glossal sclerite of ligula small, bisetose at apex; palpomeres with articles subcylindrical, palpomere 2 slightly longer than palpomere 3; palpomere 2 bisetose on inner margin with a few short setae at apex.</paragraph>
<paragraph id="2824CC40FFD4FFC6FCF5F8DDDE47EA1C" blockId="2.[836,1476,1813,2060]" pageId="2" pageNumber="263">Prothorax: Pronotum — form transverse (PW/PL = 1.82), widest at slightly posteriad middle; both anterior and posterior margins beaded; fore and hind angles round; sides evenly broadened; lateral expanded margins clearly defined, smooth and hardly reflexed; disc convex; anterior and posterior transverse impressions shallow, posteriolateral foveae small. Prosternum — intercoxal process distinctly bordered at apex.</paragraph>
<paragraph id="2824CC40FFD5FFC7FFC3FF0ADB9CE389" blockId="3.[114,754,194,409]" pageId="3" pageNumber="264">
Elytra: Form broadly ovate (EL/EW = 1.44), parallel at sides at middle portion, surface moderately convex; basal border complete; humeri square, apex broadly truncate, sutural angle nearly rectangular (
<figureCitation id="B0A0D0C5FFD5FFC7FE31FEEADB86E329" box="[384,448,290,313]" captionStart="Figs" captionStartId="3.[114,158,2036,2057]" captionTargetBox="[118,1453,1116,1998]" captionTargetId="figure-453@3.[85,1483,1085,2014]" captionTargetPageId="3" captionText="Figs. 7–11. Line drawings of structural features of Orthogonius cyclothorax, new species: 7, left fore tibia, dorsal view; 8, apex of right elytron; 9, abdominal ventrite VII of male; 10, aedeagal median lobe, right lateral view; 11, apical portion of aedeagal median lobe, dorsal view. Scale bars: 1 mm. A for Figs. 8–9; B for Figs. 7, 10–11." figureDoi="http://doi.org/10.5281/zenodo.5333245" httpUri="https://zenodo.org/record/5333245/files/figure.png" pageId="3" pageNumber="264">Fig. 8</figureCitation>
); striae rather deep, intervals quite flat, subequal in width at middle portion; interval 7 flat, not carinate; interval 3 with middle and hind setiferous punctures, basal puncture absent.
</paragraph>
<paragraph id="2824CC40FFD5FFC7FFC3FE0ADB37E0A9" blockId="3.[114,753,450,697]" pageId="3" pageNumber="264">
Legs: Middle coxae glabrous. Femora — hind femur moderate, with two setae on posterior margin of ventral surface. Tibiae — fore tibia stout, gradually enlarged at apex, apical margin sinuate, outer angle rectangular (
<figureCitation id="B0A0D0C5FFD5FFC7FD13FDEAD8A5E029" box="[674,739,546,569]" captionStart="Figs" captionStartId="3.[114,158,2036,2057]" captionTargetBox="[118,1453,1116,1998]" captionTargetId="figure-453@3.[85,1483,1085,2014]" captionTargetPageId="3" captionText="Figs. 7–11. Line drawings of structural features of Orthogonius cyclothorax, new species: 7, left fore tibia, dorsal view; 8, apex of right elytron; 9, abdominal ventrite VII of male; 10, aedeagal median lobe, right lateral view; 11, apical portion of aedeagal median lobe, dorsal view. Scale bars: 1 mm. A for Figs. 8–9; B for Figs. 7, 10–11." figureDoi="http://doi.org/10.5281/zenodo.5333245" httpUri="https://zenodo.org/record/5333245/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
); middle tibia slender; hind tibia hardly dilated at apex, apical spurs long and sharp. Tarsi — hind tarsomere 4 shorter than tarsomere 3, shallowly emarginate at apex; all tarsal claws pectinate, pectens long.
</paragraph>
<paragraph id="2824CC40FFD5FFC7FFC3FD2AD85EE109" blockId="3.[114,753,738,793]" pageId="3" pageNumber="264">
Abdomen: Abdominal ventrite VII of male complete, without emargination at apical margin (
<figureCitation id="B0A0D0C5FFD5FFC7FE76FCCAD84DE109" box="[455,523,770,793]" captionStart="Figs" captionStartId="3.[114,158,2036,2057]" captionTargetBox="[118,1453,1116,1998]" captionTargetId="figure-453@3.[85,1483,1085,2014]" captionTargetPageId="3" captionText="Figs. 7–11. Line drawings of structural features of Orthogonius cyclothorax, new species: 7, left fore tibia, dorsal view; 8, apex of right elytron; 9, abdominal ventrite VII of male; 10, aedeagal median lobe, right lateral view; 11, apical portion of aedeagal median lobe, dorsal view. Scale bars: 1 mm. A for Figs. 8–9; B for Figs. 7, 10–11." figureDoi="http://doi.org/10.5281/zenodo.5333245" httpUri="https://zenodo.org/record/5333245/files/figure.png" pageId="3" pageNumber="264">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="2824CC40FFD5FFC7FFC3FC8ADA83E1A9" blockId="3.[114,754,834,953]" pageId="3" pageNumber="264">
Male genitalia: Median lobe of aedeagus slightly expanded in middle portion, apex abruptly terminated and pointed; apical lamella small, longer than wide, obtuse at apex (
<figureCitation id="B0A0D0C5FFD5FFC7FD0DFC4ADAFFE1A9" captionStart="Figs" captionStartId="3.[114,158,2036,2057]" captionTargetBox="[118,1453,1116,1998]" captionTargetId="figure-453@3.[85,1483,1085,2014]" captionTargetPageId="3" captionText="Figs. 7–11. Line drawings of structural features of Orthogonius cyclothorax, new species: 7, left fore tibia, dorsal view; 8, apex of right elytron; 9, abdominal ventrite VII of male; 10, aedeagal median lobe, right lateral view; 11, apical portion of aedeagal median lobe, dorsal view. Scale bars: 1 mm. A for Figs. 8–9; B for Figs. 7, 10–11." figureDoi="http://doi.org/10.5281/zenodo.5333245" httpUri="https://zenodo.org/record/5333245/files/figure.png" pageId="3" pageNumber="264">Figs. 10–11</figureCitation>
).
</paragraph>
<paragraph id="2824CC40FFD5FFC7FFC3FC2ADEE6E329" blockId="3.[114,321,994,1017]" lastBlockId="3.[813,1454,193,313]" pageId="3" pageNumber="264">
Female: Unknown.
<emphasis id="1AEF1052FFD5FFC7FC9CFF09D986E2C9" bold="true" box="[813,960,193,217]" pageId="3" pageNumber="264">
<emphasis id="1AEF1052FFD5FFC7FC9CFF09D9E4E2C9" bold="true" box="[813,930,193,217]" italics="true" pageId="3" pageNumber="264">Etymology</emphasis>
. –
</emphasis>
The specific epithet is a combination of the Greek adjectival prefix “cyclo-”, meaning round, and feminine noun “thorax”, referring to the form of the pronotum which has rounded fore and hind angles.
</paragraph>
<paragraph id="2824CC40FFD5FFC7FC9CFEA9D9FFE029" blockId="3.[813,1454,353,569]" pageId="3" pageNumber="264">
<emphasis id="1AEF1052FFD5FFC7FC9CFEA9D991E369" bold="true" box="[813,983,353,377]" pageId="3" pageNumber="264">
<emphasis id="1AEF1052FFD5FFC7FC9CFEA9D9F8E369" bold="true" box="[813,958,353,377]" italics="true" pageId="3" pageNumber="264">Comparisons</emphasis>
. –
</emphasis>
This new species is similar to
<taxonomicName id="EF9BB7C3FFD5FFC7FAA2FEAADE4FE389" authority="Tian &amp; Deuve, 2006" authorityName="Tian &amp; Deuve" authorityYear="2006" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="medanensis">
<emphasis id="1AEF1052FFD5FFC7FAA2FEAADFE8E369" box="[1299,1454,354,377]" italics="true" pageId="3" pageNumber="264">O. medanensis</emphasis>
Tian &amp; Deuve, 2006
</taxonomicName>
of
<collectingRegion id="EA5F02A2FFD5FFC7FB9FFE4ADECCE389" box="[1070,1162,386,409]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="264">Sumatra</collectingRegion>
, but is markedly distinct in the following morphological features: its mentum is without seta (2 setae in
<taxonomicName id="EF9BB7C3FFD5FFC7FC5DFE0ADED3E3C9" authorityName="Tian &amp; Deuve" authorityYear="2006" box="[1004,1173,450,473]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="medanensis">
<emphasis id="1AEF1052FFD5FFC7FC5DFE0ADED3E3C9" box="[1004,1173,450,473]" italics="true" pageId="3" pageNumber="264">O. medanensis</emphasis>
</taxonomicName>
), interval 3 of elytra is without basal setiferous punctures (evident in
<taxonomicName id="EF9BB7C3FFD5FFC7FABAFE2ADFE3E3E9" authorityName="Tian &amp; Deuve" authorityYear="2006" box="[1291,1445,482,505]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="medanensis">
<emphasis id="1AEF1052FFD5FFC7FABAFE2ADFE3E3E9" box="[1291,1445,482,505]" italics="true" pageId="3" pageNumber="264">O. medanensis</emphasis>
</taxonomicName>
) and apical lamella of aedeagus is narrower (broader in
<taxonomicName id="EF9BB7C3FFD5FFC7FA25FDCAD9EFE029" authorityName="Tian &amp; Deuve" authorityYear="2006" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="medanensis">
<emphasis id="1AEF1052FFD5FFC7FA25FDCAD9EFE029" italics="true" pageId="3" pageNumber="264">O. medanensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="60819FCBFFD5FFC7FC9CFDA9D9B8E089" pageId="3" pageNumber="264" type="distribution">
<paragraph id="2824CC40FFD5FFC7FC9CFDA9D9B8E089" blockId="3.[813,1453,609,665]" pageId="3" pageNumber="264">
<emphasis id="1AEF1052FFD5FFC7FC9CFDA9D997E069" bold="true" box="[813,977,609,633]" pageId="3" pageNumber="264">
<emphasis id="1AEF1052FFD5FFC7FC9CFDA9D9F5E069" bold="true" box="[813,947,609,633]" italics="true" pageId="3" pageNumber="264">Distribution</emphasis>
. –
</emphasis>
This species is known only from the
<typeStatus id="F72072E2FFD5FFC7FACEFDAADFEBE069" box="[1407,1453,610,633]" pageId="3" pageNumber="264">type</typeStatus>
locality,
<collectingCountry id="508C8CD0FFD5FFC7FC3AFD4AD9BDE089" box="[907,1019,642,665]" name="Singapore" pageId="3" pageNumber="264">Singapore</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

265
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<mods:number id="3E435AAE2268DACF475809ECDF31BCF1">55</mods:number>
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<mods:number id="67D7F28037AF65A46E076468285691D5">2</mods:number>
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<mods:identifier id="463F6732C8A666CD5061D24412EF1146" type="ISSN">2345-7600</mods:identifier>
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<heading id="736C7B2CFFD5FFC7FC52FD29DEB1E0E9" bold="true" box="[995,1271,737,761]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="264" reason="2">
<taxonomicName id="EF9BB7C3FFD5FFC7FC52FD29DEB1E0E9" ID-CoL="6T99V" authority="Gray" authorityName="Gray, Indes Orient, C. G. Gray" authorityYear="1832" box="[995,1271,737,761]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="hopei">
<emphasis id="1AEF1052FFD5FFC7FC52FD29DEB1E0E9" bold="true" box="[995,1271,737,761]" pageId="3" pageNumber="264">
<emphasis id="1AEF1052FFD5FFC7FC52FD29DEF2E0E9" bold="true" box="[995,1204,737,761]" italics="true" pageId="3" pageNumber="264">Orthogonius hopei</emphasis>
Gray
</emphasis>
</taxonomicName>
</heading>
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<emphasis id="1AEF1052FFD5FFC7FC9CFCE8D9A2E125" box="[813,996,800,821]" italics="true" pageId="3" pageNumber="264">Orthogonius hopei</emphasis>
Gray
</taxonomicName>
, in
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</treatmentCitation>
;
<treatmentCitation id="A93AEA51FFD5FFC7FAA1FCE8D93CE142" author="Chaudoir, M" page="103" pageId="3" pageNumber="264" year="1871">
<bibRefCitation id="4C0AB1B1FFD5FFC7FAA1FCE8D93CE142" author="Chaudoir, M" pageId="3" pageNumber="264" pagination="95 - 130" refId="ref3709" refString="Chaudoir, M. de, 1871. Essai Monographique sur les Orthogoniens. Bulletin de la Societe entomologique de Belgique, 14: 95 - 130." type="journal article" year="1871">Chaudoir, 1871: 103</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="A93AEA51FFD5FFC7FC34FCF5DE71E142" author="Csiki, E" box="[901,1079,829,850]" page="1859" pageId="3" pageNumber="264" year="1932">
<bibRefCitation id="4C0AB1B1FFD5FFC7FC34FCF5DE71E142" author="Csiki, E" box="[901,1079,829,850]" pageId="3" pageNumber="264" pagination="1279 - 1598" refId="ref3737" refString="Csiki, E., 1932. Harpalinae VII. In: Junk, W. &amp; S. Schenkling (eds.), Coleopterorum Catalogus auspiciis et auxilio. Vol. 3, pars 124: 1279 - 1598. Berlin: W. Junk." type="journal article" year="1932">Csiki, 1932: 1859</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2824CC40FFD5FFC0FC9CFCBEDBD6E3AD" blockId="3.[813,1455,886,1052]" lastBlockId="4.[137,777,193,445]" lastPageId="4" lastPageNumber="265" pageId="3" pageNumber="264">
<emphasis id="1AEF1052FFD5FFC7FC9CFCBEDE54E19B" bold="true" box="[813,1042,886,907]" pageId="3" pageNumber="264">
<emphasis id="1AEF1052FFD5FFC7FC9CFCBED9BDE19B" bold="true" box="[813,1019,886,907]" italics="true" pageId="3" pageNumber="264">Specimens examined</emphasis>
. –
</emphasis>
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<specimenCount id="3E9D07C9FFD5FFC7FBA6FCBFDE2AE19C" box="[1047,1132,887,908]" pageId="3" pageNumber="264" type="female">1 female</specimenCount>
, “Coll. I.
<collectionCode id="4E8A5485FFD5FFC7FB77FCBFDE9FE19C" box="[1222,1241,887,908]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="264">R</collectionCode>
. Sc. N. B. /
<collectingCountry id="508C8CD0FFD5FFC7FAF7FCBFDFECE19C" box="[1350,1450,887,908]" name="Singapore" pageId="3" pageNumber="264">Singapore</collectingCountry>
,
<location id="2D449A9BFFD5FFC7FC9CFC5BD9C8E1B8" LSID="urn:lsid:plazi:treatment:A0327D56FFD5FFC0FC52FD29DB05E02D:2D449A9BFFD5FFC7FC9CFC5BD9C8E1B8" box="[813,910,915,936]" country="Singapore" county="Station" municipality="Mal." name="Nee Soon" pageId="3" pageNumber="264">Nee Soon</location>
,
<location id="2D449A9BFFD5FFC7FC29FC5BDE63E1B8" LSID="urn:lsid:plazi:treatment:A0327D56FFD5FFC0FC52FD29DB05E02D:2D449A9BFFD5FFC7FC29FC5BDE63E1B8" box="[920,1061,915,936]" country="Singapore" county="Station" municipality="Mal." name="Swamp Forest" pageId="3" pageNumber="264">Swamp Forest</location>
,
<collectingMunicipality id="C840563AFFD5FFC7FB9EFC5BDE1BE1B8" box="[1071,1117,915,936]" pageId="3" pageNumber="264">Mal.</collectingMunicipality>
trap 1,
<collectingCounty id="C145B4CCFFD5FFC7FB16FC5BDEADE1B8" box="[1191,1259,915,936]" pageId="3" pageNumber="264">Station</collectingCounty>
25028.
<collectorName id="856EA996FFD5FFC7FA8BFC5BDFE8E1B8" box="[1338,1454,915,936]" pageId="3" pageNumber="264">P. Grootaert</collectorName>
coll.”
</materialsCitation>
;
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<specimenCount id="3E9D07C9FFD5FFC7FCDCFC78D9E8E1D5" box="[877,942,944,965]" pageId="3" pageNumber="264" type="male">1 male</specimenCount>
and
<specimenCount id="3E9D07C9FFD5FFC7FC6CFC78DE72E1D5" box="[989,1076,944,965]" pageId="3" pageNumber="264" type="female">1 female</specimenCount>
, “Coll. I.
<collectionCode id="4E8A5485FFD5FFC7FB23FC78DEE3E1D5" box="[1170,1189,944,965]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="264">R</collectionCode>
. Sc. N. B. /
<collectingCountry id="508C8CD0FFD5FFC7FAA8FC78DF3BE1D5" box="[1305,1405,944,965]" name="Singapore" pageId="3" pageNumber="264">Singapore</collectingCountry>
,
<location id="2D449A9BFFD5FFC7FA36FC78D927E1F2" LSID="urn:lsid:plazi:treatment:A0327D56FFD5FFC0FC52FD29DB05E02D:2D449A9BFFD5FFC7FA36FC78D927E1F2" country="Singapore" county="Station" municipality="Swamp forest" name="Nee Soon" pageId="3" pageNumber="264">Nee Soon</location>
,
<collectingMunicipality id="C840563AFFD5FFC7FCDCFC05D9B3E1F2" box="[877,1013,973,994]" pageId="3" pageNumber="264">Swamp forest</collectingMunicipality>
,
<collectingMethod id="F1DAB457FFD5FFC7FBB0FC05DE13E1F2" box="[1025,1109,973,994]" pageId="3" pageNumber="264">Mal trap</collectingMethod>
2,
<collectingCounty id="C145B4CCFFD5FFC7FBC9FC05DEFAE1F2" box="[1144,1212,973,994]" pageId="3" pageNumber="264">Station</collectingCounty>
25068.
<date id="5C25EA80FFD5FFC7FABFFC05DFD4E1F2" box="[1294,1426,973,994]" pageId="3" pageNumber="264" value="2005-04-19">
<collectingDate id="4C611368FFD5FFC7FABFFC05DFD4E1F2" box="[1294,1426,973,994]" pageId="3" pageNumber="264" value="2005-04-19">19 Apr. 2005</collectingDate>
</date>
.
<collectorName id="856EA996FFD5FFC7FA2CFC05D9CFE1EF" pageId="3" pageNumber="264">P. Grootaert</collectorName>
coll.”
</materialsCitation>
;
<materialsCitation id="98F3C61DFFD5FFC0FC63FC22D876E2C6" collectingDate="2005-04-05" collectionCode="R, IRSNB" collectorName="P. Grootaert" country="Singapore" county="Station" lastPageId="4" lastPageNumber="265" location="Nee Soon" municipality="Swamp Forest" pageId="3" pageNumber="264" specimenCount="1" specimenCount-female="1">
<specimenCount id="3E9D07C9FFD5FFC7FC63FC22DE6FE1EF" box="[978,1065,1002,1023]" pageId="3" pageNumber="264" type="female">1 female</specimenCount>
, “Coll. I.
<collectionCode id="4E8A5485FFD5FFC7FB3AFC22DED8E1EF" box="[1163,1182,1002,1023]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="264">R</collectionCode>
. Sc. N. B. /
<collectingCountry id="508C8CD0FFD5FFC7FAA9FC22DF3AE1EF" box="[1304,1404,1002,1023]" name="Singapore" pageId="3" pageNumber="264">Singapore</collectingCountry>
,
<location id="2D449A9BFFD5FFC7FA36FC22D927E60C" LSID="urn:lsid:plazi:treatment:A0327D56FFD5FFC0FC52FD29DB05E02D:2D449A9BFFD5FFC7FA36FC22D927E60C" country="Singapore" county="Station" municipality="Swamp Forest" name="Nee Soon" pageId="3" pageNumber="264">Nee Soon</location>
,
<location id="2D449A9BFFD5FFC7FCDCFBCFD9DCE60C" LSID="urn:lsid:plazi:treatment:A0327D56FFD5FFC0FC52FD29DB05E02D:2D449A9BFFD5FFC7FCDCFBCFD9DCE60C" box="[877,922,1031,1052]" country="Singapore" county="Station" municipality="Swamp Forest" name="Mal." pageId="3" pageNumber="264">Mal.</location>
trap 1,
<collectingMunicipality id="C840563AFFD5FFC7FC58FBCFDE31E60C" box="[1001,1143,1031,1052]" pageId="3" pageNumber="264">Swamp Forest</collectingMunicipality>
,
<collectingCounty id="C145B4CCFFD5FFC7FB32FBCFDE81E60C" box="[1155,1223,1031,1052]" pageId="3" pageNumber="264">Station</collectingCounty>
25046.
<date id="5C25EA80FFD5FFC7FAABFBCFDFD4E60C" box="[1306,1426,1031,1052]" pageId="3" pageNumber="264" value="2005-04-05">
<collectingDate id="4C611368FFD5FFC7FAABFBCFDFD4E60C" box="[1306,1426,1031,1052]" pageId="3" pageNumber="264" value="2005-04-05">5 Apr. 2005</collectingDate>
</date>
.
<collectorName id="856EA996FFD5FFC0FA2CFBCFDAA3E2C6" lastPageId="4" lastPageNumber="265" pageId="3" pageNumber="264">P. Grootaert</collectorName>
coll.”, all are deposited in
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</materialsCitation>
except for
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<specimenCount id="3E9D07C9FFD2FFC0FD2FFF09D94EE2C6" box="[670,776,193,214]" pageId="4" pageNumber="265" type="female">one female</specimenCount>
in
<collectionCode id="4E8A5485FFD2FFC0FF17FF16DAB2E2E3" box="[166,244,222,243]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="265">MNHN</collectionCode>
</materialsCitation>
.
<materialsCitation id="98F3C61DFFD2FFC0FEB0FF16DAA0E300" collectionCode="MNHN" country="Singapore" pageId="4" pageNumber="265" specimenCount="1" specimenCount-female="1">
<specimenCount id="3E9D07C9FFD2FFC0FEB0FF16DB1BE2E3" box="[257,349,222,243]" pageId="4" pageNumber="265" type="female">1 female</specimenCount>
, “
<taxonomicName id="EF9BB7C3FFD2FFC0FEC5FF16D8B8E2E3" ID-CoL="6T99V" authority="Gray, Indes Orient, C. G. Gray" authorityName="Gray, Indes Orient, C. G. Gray" authorityYear="1832" box="[372,766,222,243]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="hopei">Hopei Gray, Indes Orient, C. G. Gray</taxonomicName>
” (
<collectionCode id="4E8A5485FFD2FFC0FF21FF33DA99E300" box="[144,223,251,272]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="265">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="98F3C61DFFD2FFC0FF43FF33D8E7E300" box="[242,673,251,272]" collectionCode="MNHN" country="Malaysia" location="Malacca" pageId="4" pageNumber="265" specimenCount="1" specimenCount-female="1">
<specimenCount id="3E9D07C9FFD2FFC0FF43FF33DB0CE300" box="[242,330,251,272]" pageId="4" pageNumber="265" type="female">1 female</specimenCount>
, “
<collectingCountry id="508C8CD0FFD2FFC0FEEEFF33DBF2E300" box="[351,436,251,272]" name="Malaysia" pageId="4" pageNumber="265">Malacca</collectingCountry>
, H. Deyralle” (
<collectionCode id="4E8A5485FFD2FFC0FDFAFF33D8DCE300" box="[587,666,251,272]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="265">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="98F3C61DFFD2FFC0FD1CFF33D866E33D" collectionCode="MNHN" collectorName="Coll. E. Steinheil" country="Malaysia" location="Malacca" pageId="4" pageNumber="265" specimenCount="1" specimenCount-female="1">
<specimenCount id="3E9D07C9FFD2FFC0FD1CFF33D943E300" box="[685,773,251,272]" pageId="4" pageNumber="265" type="female">1 female</specimenCount>
, “
<collectingCountry id="508C8CD0FFD2FFC0FF22FED0DAAFE33D" box="[147,233,280,301]" name="Malaysia" pageId="4" pageNumber="265">Malacca</collectingCountry>
”, “
<collectorName id="856EA996FFD2FFC0FEB9FED0DBF4E33D" box="[264,434,280,301]" pageId="4" pageNumber="265">Coll. E. Steinheil</collectorName>
” (
<collectionCode id="4E8A5485FFD2FFC0FE7BFED0D85FE33D" box="[458,537,280,301]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="265">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="98F3C61DFFD2FFC0FD9AFED0DBFFE359" collectionCode="MNHN" country="India" location="Inde" pageId="4" pageNumber="265" specimenCount="1" specimenCount-female="1">
<specimenCount id="3E9D07C9FFD2FFC0FD9AFED0D8C4E33D" box="[555,642,280,301]" pageId="4" pageNumber="265" type="female">1 female</specimenCount>
, “
<collectingCountry id="508C8CD0FFD2FFC0FD26FED0D880E33D" box="[663,710,280,301]" name="India" pageId="4" pageNumber="265">Inde</collectingCountry>
”, “Ex- Musaeo Mniszech” (
<collectionCode id="4E8A5485FFD2FFC0FED1FEFCDBF4E359" box="[352,434,308,329]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="265">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="98F3C61DFFD2FFC0FE76FEFCDB08E376" collectionCode="MNHN" country="India" pageId="4" pageNumber="265" specimenCount="1" specimenCount-male="1">
<specimenCount id="3E9D07C9FFD2FFC0FE76FEFCD856E359" box="[455,528,308,329]" pageId="4" pageNumber="265" type="male">1 male</specimenCount>
, “Malais”, “Ex-Musaeo Mniszech” (
<collectionCode id="4E8A5485FFD2FFC0FF48FE99DB00E376" box="[249,326,337,358]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="265">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="98F3C61DFFD2FFC0FEE4FE99D943E376" box="[341,773,337,358]" collectionCode="MNHN" country="India" pageId="4" pageNumber="265" specimenCount="1" specimenCount-female="1">
<specimenCount id="3E9D07C9FFD2FFC0FEE4FE99DBE1E376" box="[341,423,337,358]" pageId="4" pageNumber="265" type="female">1 female</specimenCount>
, “Ex-Musaeo L. Fairmaire” (
<collectionCode id="4E8A5485FFD2FFC0FD00FE99D8B9E376" box="[689,767,337,358]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="265">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="98F3C61DFFD2FFC0FF38FEA6D8EEE393" box="[137,680,366,387]" collectionCode="MNHN" country="Malaysia" location="Malacca" pageId="4" pageNumber="265" specimenCount="1" specimenCount-female="1">
<specimenCount id="3E9D07C9FFD2FFC0FF38FEA6DA98E393" box="[137,222,366,387]" pageId="4" pageNumber="265" type="female">1 female</specimenCount>
, “
<collectingCountry id="508C8CD0FFD2FFC0FF41FEA6DB00E393" box="[240,326,366,387]" name="Malaysia" pageId="4" pageNumber="265">Malacca</collectingCountry>
”, “Collection Cal. Pradier” (
<collectionCode id="4E8A5485FFD2FFC0FDE3FEA6D8E7E393" box="[594,673,366,387]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="265">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="98F3C61DFFD2FFC0FD01FEA6DB96E3B0" collectionCode="MNHN" country="Malaysia" pageId="4" pageNumber="265" specimenCount="1" specimenCount-female="1">
<specimenCount id="3E9D07C9FFD2FFC0FD01FEA6D943E393" box="[688,773,366,387]" pageId="4" pageNumber="265" type="female">1 female</specimenCount>
, “Ex-Musaeo Mniszech” (
<collectionCode id="4E8A5485FFD2FFC0FECBFE43DB8FE3B0" box="[378,457,395,416]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="265">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="98F3C61DFFD2FFC0FE69FE43DBCAE3AD" collectionCode="MNHN" country="Malaysia" location="Malacca" pageId="4" pageNumber="265" specimenCount="1" specimenCount-female="1">
<specimenCount id="3E9D07C9FFD2FFC0FE69FE43D86BE3B0" box="[472,557,395,416]" pageId="4" pageNumber="265" type="female">1 female</specimenCount>
, “
<collectingCountry id="508C8CD0FFD2FFC0FD8EFE43D8D3E3B0" box="[575,661,395,416]" name="Malaysia" pageId="4" pageNumber="265">Malacca</collectingCountry>
”, “
<collectingCountry id="508C8CD0FFD2FFC0FD01FE43D943E3B0" box="[688,773,395,416]" name="Malaysia" pageId="4" pageNumber="265">Malacca</collectingCountry>
, Oberthur, 1896” (
<collectionCode id="4E8A5485FFD2FFC0FE87FE60DBC3E3AD" box="[310,389,424,445]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="265">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="7CE49CC8FFD5FFC7FFC3F83CD89CEA53" ID-DOI="http://doi.org/10.5281/zenodo.5333245" ID-Zenodo-Dep="5333245" httpUri="https://zenodo.org/record/5333245/files/figure.png" pageId="3" pageNumber="264" startId="3.[114,158,2036,2057]" targetBox="[118,1453,1116,1998]" targetPageId="3">
<paragraph id="2824CC40FFD5FFC7FFC3F83CD89CEA53" blockId="3.[114,1454,2036,2115]" pageId="3" pageNumber="264">
Figs. 7–11. Line drawings of structural features of
<taxonomicName id="EF9BB7C3FFD5FFC7FDDBF83CD91AEA19" authority="Tian &amp; Deuve, 2007" authorityName="Tian &amp; Deuve" authorityYear="2007" box="[618,860,2036,2057]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="cyclothorax" status="sp. nov.">
<emphasis id="1AEF1052FFD5FFC7FDDBF83CD91AEA19" box="[618,860,2036,2057]" italics="true" pageId="3" pageNumber="264">Orthogonius cyclothorax</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="01DCAD29FFD5FFC7FCDBF83CD9A4EA19" box="[874,994,2036,2057]" pageId="3" pageNumber="264" rank="species">new species</taxonomicNameLabel>
: 7, left fore tibia, dorsal view; 8, apex of right elytron; 9, abdominal ventrite VII of male; 10, aedeagal median lobe, right lateral view; 11, apical portion of aedeagal median lobe, dorsal view. Scale bars: 1 mm. A for Figs. 8–9; B for Figs. 7, 10–11.
</paragraph>
</caption>
<subSubSection id="60819FCBFFD2FFC0FF38FE2EDB05E02D" pageId="4" pageNumber="265" type="distribution">
<paragraph id="2824CC40FFD2FFC0FF38FE2EDB05E02D" blockId="4.[137,776,486,573]" pageId="4" pageNumber="265">
<emphasis id="1AEF1052FFD2FFC0FF38FE2EDB6EE3EE" bold="true" box="[137,296,486,510]" pageId="4" pageNumber="265">
<emphasis id="1AEF1052FFD2FFC0FF38FE2EDB48E3EE" bold="true" box="[137,270,486,510]" italics="true" pageId="4" pageNumber="265">Distribution</emphasis>
. –
</emphasis>
This species has so far been cited from
<collectingCountry id="508C8CD0FFD2FFC0FD60FE2ED94EE3ED" box="[721,776,486,509]" name="India" pageId="4" pageNumber="265">India</collectingCountry>
and Malay Peninsula. Here, it is recorded from
<collectingCountry id="508C8CD0FFD2FFC0FD2DFDCED94EE00D" box="[668,776,518,541]" name="Singapore" pageId="4" pageNumber="265">Singapore</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>

189
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<mods:title id="EF983D6834E42FB31F5CDC35FC937B79">A Taxonomic Treatment Of The Species Of Orthogonius Macleay (Coleoptera: Carabidae: Orthogoniini) Of Singapore</mods:title>
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<mods:namePart id="30AD8D846722CF86255554B7B835B8ED">Tian, Ming-yi</mods:namePart>
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<mods:title id="FDF1CE7621FF7C6F4B354B41E0790FC0">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="DDF7D1032508617C30469FC080D6567B">
<mods:date id="7948D10AB1FFB9D4CBD5AB2F667B9E38">2007</mods:date>
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<mods:number id="FC8343B5B5F21CA865819F0A13D7F4C7">2007-08-31</mods:number>
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<subSubSection id="60819FCBFFD7FFC5FF68FE69D8CDE3A9" box="[217,651,417,441]" pageId="1" pageNumber="262" type="nomenclature">
<paragraph id="2824CC40FFD7FFC5FF68FE69D8CDE3A9" blockId="1.[217,651,417,472]" box="[217,651,417,441]" pageId="1" pageNumber="262">
<heading id="736C7B2CFFD7FFC5FF68FE69D8CDE3A9" bold="true" box="[217,651,417,441]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="262" reason="2">
<emphasis id="1AEF1052FFD7FFC5FF68FE69D8CDE3A9" bold="true" box="[217,651,417,441]" pageId="1" pageNumber="262">
<taxonomicName id="EF9BB7C3FFD7FFC5FF68FE69DBB1E3A9" ID-CoL="758Y4" authority="Tian &amp; Deuve, 2007" authorityName="Tian &amp; Deuve" authorityYear="2007" box="[217,503,417,441]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="emarginatus" status="sp. nov.">
<emphasis id="1AEF1052FFD7FFC5FF68FE69DBB1E3A9" bold="true" box="[217,503,417,441]" italics="true" pageId="1" pageNumber="262">Orthogonius emarginatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="01DCAD29FFD7FFC5FDB7FE69D8CDE3A9" box="[518,651,417,441]" pageId="1" pageNumber="262" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="60819FCBFFD7FFC5FEC7FE09DBA8E3C8" box="[374,494,449,472]" pageId="1" pageNumber="262" type="description">
<paragraph id="2824CC40FFD7FFC5FEC7FE09DBA8E3C8" blockId="1.[217,651,417,472]" box="[374,494,449,472]" pageId="1" pageNumber="262">
(
<figureCitation id="B0A0D0C5FFD7FFC5FECFFE09DBA2E3C8" box="[382,484,449,472]" captionStart="Figs" captionStartId="1.[116,160,2035,2056]" captionTargetBox="[114,1449,791,1979]" captionTargetId="figure-286@1.[70,1488,765,2003]" captionTargetPageId="1" captionText="Figs. 1–6. Line drawings of structural features of Orthogonius emarginatus, new species: 1, left fore tibia, dorsal view; 2, left mid femur, ventral view; 3, apex of right elytron; 4, abdominal ventrite VII of male, ventral view; 5, aedeagus, right lateral view; 6. apical portion of aedeagus, dorsal view. Scale bars: 1 mm. A for Figs. 3–4; B for Figs. 1–2 and 5–6." figureDoi="http://doi.org/10.5281/zenodo.5333243" httpUri="https://zenodo.org/record/5333243/files/figure.png" pageId="1" pageNumber="262">Figs. 1–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="60819FCBFFD7FFC5FFC3FDC8DBBCE07B" pageId="1" pageNumber="262" type="materials_examined">
<paragraph id="2824CC40FFD7FFC5FFC3FDC8DBBCE07B" blockId="1.[114,754,512,619]" pageId="1" pageNumber="262">
<emphasis id="1AEF1052FFD7FFC5FFC3FDC8DB50E005" box="[114,278,512,533]" italics="true" pageId="1" pageNumber="262">
<emphasis id="1AEF1052FFD7FFC5FFC3FDC8DB45E005" bold="true" box="[114,259,512,533]" italics="true" pageId="1" pageNumber="262">Type material.</emphasis>
–
</emphasis>
<materialsCitation id="98F3C61DFFD7FFC5FEAFFDC8DBBCE07B" collectingDate="2005-04-05" collectionCode="R, IRSNB" collectorName="P. Grootaert" country="Singapore" location="Swamp forest" pageId="1" pageNumber="262" specimenCount="1" typeStatus="holotype">
<typeStatus id="F72072E2FFD7FFC5FEAFFDC8DB3DE005" box="[286,379,512,533]" pageId="1" pageNumber="262" type="holotype">Holotype</typeStatus>
: male, labeled “Coll. I.
<collectionCode id="4E8A5485FFD7FFC5FDD8FDC8D83AE005" box="[617,636,512,533]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="262">R</collectionCode>
. Sc. N. B. /
<collectingCountry id="508C8CD0FFD7FFC5FFC3FDD5DA90E022" box="[114,214,541,562]" name="Singapore" pageId="1" pageNumber="262">Singapore</collectingCountry>
, Nee Soon,
<collectingMethod id="F1DAB457FFD7FFC5FEE2FDD5DBE1E022" box="[339,423,541,562]" pageId="1" pageNumber="262">Mal trap</collectingMethod>
1,
<location id="2D449A9BFFD7FFC5FE7DFDD5D813E022" LSID="urn:lsid:plazi:treatment:A0327D56FFD7FFC6FF68FE69D9FEE3E9:2D449A9BFFD7FFC5FE7DFDD5D813E022" box="[460,597,541,562]" country="Singapore" name="Swamp forest" pageId="1" pageNumber="262">Swamp forest</location>
, Station 25046.
<date id="5C25EA80FFD7FFC5FFC3FDF1DAAEE05E" box="[114,232,569,590]" pageId="1" pageNumber="262" value="2005-04-05">
<collectingDate id="4C611368FFD7FFC5FFC3FDF1DAAEE05E" box="[114,232,569,590]" pageId="1" pageNumber="262" value="2005-04-05">5 Apr. 2005</collectingDate>
</date>
.
<collectorName id="856EA996FFD7FFC5FF43FDF1DB20E05E" box="[242,358,569,590]" pageId="1" pageNumber="262">P. Grootaert</collectorName>
coll.”, collected at Nee Soon,
<collectingCountry id="508C8CD0FFD7FFC5FD21FDF1D8B7E05E" box="[656,753,569,590]" name="Singapore" pageId="1" pageNumber="262">Singapore</collectingCountry>
(type locality) and deposited in
<collectionCode id="4E8A5485FFD7FFC5FE1DFD9EDBB0E07B" box="[428,502,598,619]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="262">IRSNB</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="60819FCBFFD7FFC6FFC3FD5CDB1FE569" lastPageId="2" lastPageNumber="263" pageId="1" pageNumber="262" type="description">
<paragraph id="2824CC40FFD7FFC5FFC3FD5CDBFEE0DC" blockId="1.[114,753,660,716]" pageId="1" pageNumber="262">
<emphasis id="1AEF1052FFD7FFC5FFC3FD5CDB4AE0BC" bold="true" box="[114,268,660,684]" pageId="1" pageNumber="262">
<emphasis id="1AEF1052FFD7FFC5FFC3FD5CDABEE0BC" bold="true" box="[114,248,660,684]" italics="true" pageId="1" pageNumber="262">Description.</emphasis>
–
</emphasis>
Body: Length 15.0 mm; width 6.0 mm. Form stout; surface markedly shiny.
</paragraph>
<paragraph id="2824CC40FFD7FFC5FC9FFF09DE4EE2E8" blockId="1.[814,1454,193,248]" pageId="1" pageNumber="262">Color: Body black; antennae, palpi, legs and ventral surface reddish dark brown.</paragraph>
<paragraph id="2824CC40FFD7FFC5FC9FFEE9DFCDE3A8" blockId="1.[814,1454,289,440]" pageId="1" pageNumber="262">Macrosculpture. Head: dorsal surface without striation, but densely and moderately punctate; base of neck and median portion of vertex impunctate. Pronotum smooth, glabrous, shallowly striate. Elytra with intervals very shallowly and sparsely punctate. Ventral surface smooth and glabrous.</paragraph>
<paragraph id="2824CC40FFD7FFC5FC9FFE29D9E7E028" blockId="1.[814,1454,481,568]" pageId="1" pageNumber="262">Microsculpture: Mesh pattern isodiametric on dorsal surface of head, but densely and moderately transverse on pronotum and elytra.</paragraph>
<paragraph id="2824CC40FFD7FFC6FC9FFDA9D8AEE329" blockId="1.[814,1454,609,728]" lastBlockId="2.[138,776,194,313]" lastPageId="2" lastPageNumber="263" pageId="1" pageNumber="262">Head: Slightly wider than long (HW/HL = 1.15), convex. Frontal impressions large and deep, fovea-like, not extended to level of anterior pair of supraorbital setae; clypeus bisetose, surface uneven, slightly raised at base. Eyes very large and markedly prominent, longitudinal diameter of eye about twothirds distance between eyes. Antennae rather long, slightly extended over base of elytra; antennomere 4 longer than antennomere 3, almost twice as long as antennomere 2.</paragraph>
<caption id="7CE49CC8FFD7FFC5FFC5F83BD9E1EA52" ID-DOI="http://doi.org/10.5281/zenodo.5333243" ID-Zenodo-Dep="5333243" httpUri="https://zenodo.org/record/5333243/files/figure.png" pageId="1" pageNumber="262" startId="1.[116,160,2035,2056]" targetBox="[114,1449,791,1979]" targetPageId="1">
<paragraph id="2824CC40FFD7FFC5FFC5F83BD9E1EA52" blockId="1.[114,1454,2035,2114]" pageId="1" pageNumber="262">
Figs. 1–6. Line drawings of structural features of
<taxonomicName id="EF9BB7C3FFD7FFC5FDE9F83CD909EA19" authority="Tian &amp; Deuve, 2007" authorityName="Tian &amp; Deuve" authorityYear="2007" box="[600,847,2036,2057]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="emarginatus" status="sp. nov.">
<emphasis id="1AEF1052FFD7FFC5FDE9F83CD909EA19" box="[600,847,2036,2057]" italics="true" pageId="1" pageNumber="262">Orthogonius emarginatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="01DCAD29FFD7FFC5FCEDF83BD992EA18" box="[860,980,2035,2056]" pageId="1" pageNumber="262" rank="species">new species</taxonomicNameLabel>
: 1, left fore tibia, dorsal view; 2, left mid femur, ventral view; 3, apex of right elytron; 4, abdominal ventrite VII of male, ventral view; 5, aedeagus, right lateral view; 6. apical portion of aedeagus, dorsal view. Scale bars: 1 mm. A for Figs. 3–4; B for Figs. 1–2 and 5–6.
</paragraph>
</caption>
<paragraph id="2824CC40FFD4FFC6FF3BFEAAD8A8E369" blockId="2.[138,750,354,377]" box="[138,750,354,377]" pageId="2" pageNumber="263">Mouthparts: Labrum six setose, straight at front margin.</paragraph>
<paragraph id="2824CC40FFD4FFC6FF3BFE6AD8E4E3A9" blockId="2.[138,674,418,441]" box="[138,674,418,441]" pageId="2" pageNumber="263">Maxillae: palpal articles subcylindrical, glabrous.</paragraph>
<paragraph id="2824CC40FFD4FFC6FF3BFE2ADA9BE069" blockId="2.[138,776,482,633]" pageId="2" pageNumber="263">Labium: submentum with a pair of setae; mentum without median tooth, with a pair of setae; glossal sclerite short, quite stout, bisetose at apex; palpal articles subcylindrical, glabrous; palpiger asetose; palpomere 2 bisetose on inner margin.</paragraph>
<paragraph id="2824CC40FFD4FFC6FF3BFD6ADB24E1A9" blockId="2.[138,777,674,953]" pageId="2" pageNumber="263">Prothorax: Pronotum — form transverse (PW/PL=1.66), widest at about middle; both fore and basal margins finely beaded; sides anteriorly and posteriorly evenly contracted; the lateral expanded margins comparatively narrow, quite even throughout, smooth and evidently reflexed; both fore and hind angles round; disc markedly convex; anterior and posterior transverse impressions distinct, deep and wide; posteriolateral foveae small. Prosternum — intercoxal process unbordered at apex.</paragraph>
<paragraph id="2824CC40FFD4FFC6FF3BFC2AD863E6A9" blockId="2.[138,777,994,1209]" pageId="2" pageNumber="263">
Elytra: Form elongate (EL/EW=1.5), parallel at sides in middle portion; surface rather flat; basal border indistinct basad intervals 1–5; humeri square; apex roundly truncate, sutural angle nearly rectangular (
<figureCitation id="B0A0D0C5FFD4FFC6FE55FB8AD865E649" box="[484,547,1090,1113]" captionStart="Figs" captionStartId="1.[116,160,2035,2056]" captionTargetBox="[114,1449,791,1979]" captionTargetId="figure-286@1.[70,1488,765,2003]" captionTargetPageId="1" captionText="Figs. 1–6. Line drawings of structural features of Orthogonius emarginatus, new species: 1, left fore tibia, dorsal view; 2, left mid femur, ventral view; 3, apex of right elytron; 4, abdominal ventrite VII of male, ventral view; 5, aedeagus, right lateral view; 6. apical portion of aedeagus, dorsal view. Scale bars: 1 mm. A for Figs. 3–4; B for Figs. 1–2 and 5–6." figureDoi="http://doi.org/10.5281/zenodo.5333243" httpUri="https://zenodo.org/record/5333243/files/figure.png" pageId="2" pageNumber="263">Fig. 3</figureCitation>
); striae fine and deep, intervals slightly convex, except interval 7 carinate in basal half; intervals subequal in width in middle portion; interval 3 with only basal setiferous puncture.
</paragraph>
<paragraph id="2824CC40FFD4FFC6FF3BFB2AD896E409" blockId="2.[138,777,1250,1561]" pageId="2" pageNumber="263">
Legs: Middle coxae glabrous. Femora — hind femur comparatively slender, with only one seta on posterior margin of ventral surface, at one-fourth from base. Tibiae — fore tibia slightly dilated at apex, latter rounded and sinuate near outer apical angle (
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); middle tibia robust, markedly dilated at apical portion, outer apical surface slightly but evidently emarginate in outer apical portion (
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); apical spurs of hind tibia long and sharp. Tarsi — hind tarsomere 3 much longer than tarsomere 4; tarsomere symmetrically bilobed at apex; all tarsal claws pectinate, pectens very short.
</paragraph>
<paragraph id="2824CC40FFD4FFC6FF3BF98ADBDDE469" blockId="2.[138,777,1602,1657]" pageId="2" pageNumber="263">
Abdomen:Abdominal ventrite VII of male deeply emarginate at apical margin (
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).
</paragraph>
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Male genitalia: Median lobe of aedeagus robust (
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), ventral margin markedly sinuate, evenly constricted toward apex, obtuse and somewhat tube-like; dorsal orifice long; apical lamella slightly narrowed toward apex, slightly longer than wide.
</paragraph>
<paragraph id="2824CC40FFD4FFC6FF3BF8AADB1FE569" blockId="2.[138,345,1890,1913]" box="[138,345,1890,1913]" pageId="2" pageNumber="263">Female: Unknown.</paragraph>
</subSubSection>
<subSubSection id="60819FCBFFD4FFC6FF3BF869DE6EE389" pageId="2" pageNumber="263" type="etymology">
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<emphasis id="1AEF1052FFD4FFC6FF3BF869DB5CE5A9" bold="true" box="[138,282,1953,1977]" pageId="2" pageNumber="263">
<emphasis id="1AEF1052FFD4FFC6FF3BF869DAB9E5A9" bold="true" box="[138,255,1953,1977]" italics="true" pageId="2" pageNumber="263">Etymology</emphasis>
. –
</emphasis>
The specific epithet, “emarginatus”, is a Latin adjective, nominative case, referring to the emargination of middle tibia in outer apical portion, which characterises this species.
</paragraph>
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<emphasis id="1AEF1052FFD4FFC6FCF5FF09D9B2E2C9" bold="true" box="[836,1012,193,217]" pageId="2" pageNumber="263">
<emphasis id="1AEF1052FFD4FFC6FCF5FF09D990E2C9" bold="true" box="[836,982,193,217]" italics="true" pageId="2" pageNumber="263">Comparisons</emphasis>
. –
</emphasis>
This new species is closely resembles to
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<emphasis id="1AEF1052FFD4FFC6FCF5FF2ADE4FE2E9" box="[836,1033,226,249]" italics="true" pageId="2" pageNumber="263">O. andrewesianus</emphasis>
Tian &amp; Deuve (2006d)
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, but it is easily distinguishable from the latter by its larger size, labrum that is straight at the front margin (slightly convex in
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<emphasis id="1AEF1052FFD4FFC6FCF5FE8ADE56E349" box="[836,1040,322,345]" italics="true" pageId="2" pageNumber="263">O. andrewesianus</emphasis>
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), base of elytra indistinctly bordered (unbordered against intervals
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) and the stouter aedeagus.
</paragraph>
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<subSubSection id="60819FCBFFD4FFC6FCF5FE09D9FEE3E9" pageId="2" pageNumber="263" type="distribution">
<paragraph id="2824CC40FFD4FFC6FCF5FE09D9FEE3E9" blockId="2.[836,1473,449,505]" pageId="2" pageNumber="263">
<emphasis id="1AEF1052FFD4FFC6FCF5FE09DE40E3C9" bold="true" box="[836,1030,449,473]" pageId="2" pageNumber="263">
<emphasis id="1AEF1052FFD4FFC6FCF5FE09D9A7E3C9" bold="true" box="[836,993,449,473]" italics="true" pageId="2" pageNumber="263">Distribution</emphasis>
. –
</emphasis>
This species is known only from
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.
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(Fig. 402)
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<paragraph id="753B411441D537EEFABED54A720CF4EB" pageNumber="679">HOLOTYPE WORKER. TL 2.4, HL 0.55, HW 0.37, CI 67, ML 0.26, MI 48, SL 0.34, SI 93, PW 0.25, AL 0.63. Characters of epulo-complex. Mandibles in full-face view with outer margins shallowly and evenly convex, the width approximately constant from the level of the proximal preapical tooth to near the base. In full-face view, upper scrobe margin narrowly concave (pinched in) immediately behind frontal lobes. Behind this, upper scrobe margin divergent and shallowly convex, not bordered by a rim or flange except for a thin lamella immediately behind frontal lobes, the eyes clearly visible. Apices of the upper scrobe margins confluent with the sides of the occipital lobes through an even curve, without an angle separating the two. Maximum diameter of eye slightly larger than maximum width of scape, with 4 - 5 ommatidia in longest row. Scape narrow and subcylindrical, shallowly and gently curved; hairs on leading edge narrow, flattened or spoon-shaped apically. Cephalic dorsum clothed with inconspicuous, curved, narrow, filiform ground-pilosity, the upper scrobe margin fringed with hairs which are the same shape and size as those on the dorsum. Upper scrobe margin with two laterally projecting flagellate hairs, one just posterior to eye, the other apicoscrobal. Cephalic dorsum with 2 pairs of standing fine, flagellate hairs close to the occipital margin and a more anteriorly situated flagellate pair. A shorter filiform hair is located at the apex of the occipital lobe. Pronotal humeral hair long, flagellate. Mesonotum with two pairs of standing hairs, a flagellate pair on the anterior margin and a filiform pair immediately behind them that curve gently toward the midline. Dorsal alitrunk without ground-pilosity. Dorsum of alitrunk in outline convex anteriorly and more or less flat posteriorly; posterior portion of mesonotum only slightly depressed below the anterior margin of propodeum. Anterior mesonotum with a narrow flange above the mesothoracic spiracle. Propodeal tooth broadly triangular and directed posteriorly, without a distinct lamella on declivity. Dorsal surfaces of alitrunk, petiole node and postpetiole disc more or less smooth and shiny, declivity of promesonotum with fine punctures which appear almost effaced. Sides of alitrunk smooth.</paragraph>
<paragraph id="61EE4B6FD3460EAF6FBF752B8AE3A797" pageNumber="680">In dorsal view petiole node longer than broad, dorsolateral margin of peduncle with a lamellate flange that extends to the petiolar disc. In profile the petiole with a well developed ventral spongiform curtain that is deeper than the peduncle of the petiole. In dorsal view disc of postpetiole completely surrounded by spongiform tissue. Basigastral costulae very short, more or less restricted to limbus. Dorsal surfaces of petiole, postpetiole and gaster with filiform and flagellate hairs. Colour brownish yellow.</paragraph>
<paragraph id="AA8A359F10FF37B9618853DDBE4CFFEA" pageNumber="680">PARATYPE WORKERS. TL 2.2 - 2.4, HL 0.51 - 0.56, HW 0.35 - 0.38, CI 67 - 71, ML 0.26 - 0.28, MI 46 - 53, SL 0.31 - 34, SI 85 - 90, PW 0.22 - 0.25, AL 0.57 - 0.64 (8 measured). As holotype.</paragraph>
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<paragraph id="52E60A86C020F4C0BBA94F7531D89082" pageNumber="680">Holotype worker, Madagascar: 6.3 km. S Ambanizana, Andranobe, 15 ° 41 ' S, 49 ° 57 ' E, 25 m., 14. xi. 1993, sifted litter (leaf mold, rotten wood), rainforest, # 886 (10) - 4 (B. L. Fisher) (MCZ).</paragraph>
<paragraph id="33BB72FE65DE5448DDB17D49C1E559E9" pageNumber="680">Paratypes. 8 workers and 1 queen (dealate) with same data as holotype but coded (12) - 12, (41) - 13 (BMNH, SAM).</paragraph>
<paragraph id="382B121372027F7661E1AAA5DD7C3656" pageNumber="680">NON-PARATYPIC MATERIAL EXAMINED. Madagascar: 5.3 km. SSE Ambanizana, Andranobe, 425 m. (B. L. Fisher); 19 km. SSE Maroantsetra, 350 m. (P. S. Vtxrd).</paragraph>
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<paragraph id="794E28F69EE628A6E53DA7D302CAEF34" pageNumber="680">Measurements of this material extend the range shown by the type-series: HL 0.52 - 0.59, HW 0.35 - 0.39, CI 65 - 70, ML 0.26 - 0.32, MI 50 - 54, SL 0.33 - 0.36, SI 90 - 94 (6 measured). The material for 19 km. SSE Maroantsetra differs from the type-material by having additional pairs of short filiform hairs on the posterior half of the cephalic dorsum.</paragraph>
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<paragraph id="3FEF8FDE58AF1AAA2BD69C19F38CED91" pageNumber="680">
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is easily distinguished from other species in the
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by the presence of flagellate hairs on the cephalic dorsum, pronotal humerus and anterior margin of mesonotum. The mostly smooth and shiny alitrunk dorsum, petiole and postpetiole also help distinguish this species.
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<mods:roleTerm id="CD08AC976070C1A5569B250739F76F93">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="940F284A530C727A5C860A79A6355132">D'Alessandro, Paola</mods:namePart>
<mods:nameIdentifier id="F920FBF3607E09CCFB4B957A9722CC24" type="ORCID">0000-0002-4481-9152</mods:nameIdentifier>
<mods:affiliation id="B561500CF18B06886ED8DBE057AD6328">Department of Life, Health and Environmental Sciences, University of L’Aquila, 67100 L’Aquila, Italy</mods:affiliation>
</mods:name>
<mods:typeOfResource id="45B22FBC0EF5C89B16515C3468CE2065">text</mods:typeOfResource>
<mods:relatedItem id="BD4B8F4D127FE03A8AF40AE41FB7E621" type="host">
<mods:titleInfo id="F9FD051E4D52B8C44854CC97895F4456">
<mods:title id="C72B0BFF2E0195E7DB9380B7473C4303">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="37F1E018BDA43E01BEC15D69FA51F94F">
<mods:date id="642649B200C958ECDC9484779ADCF9BB">2024</mods:date>
<mods:detail id="BF15F393E62C7EAE381866EDCC4A940F" type="pubDate">
<mods:number id="722B3CC1F27A2DAD5B08C372D646ADC6">2024-05-27</mods:number>
</mods:detail>
<mods:detail id="80ECCFD6C5946472FCF5C9936CFD7A05" type="volume">
<mods:number id="FEF0AC4FEB086718CC6E328FDBEDAA82">1202</mods:number>
</mods:detail>
<mods:extent id="4BC5B85D24C14CE462C5EE5877E5CADD" unit="page">
<mods:start id="1BDD643A84294EB3322859D976035841">303</mods:start>
<mods:end id="11036214332C7336AD32D1BD66D0CF5E">327</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="C7EBF7E7C3E22225FE1FDB309DD55B1B">journal article</mods:classification>
<mods:identifier id="8BDAFB003AD75BE663480A70BF400679" type="DOI">10.3897/zookeys.1202.122977</mods:identifier>
<mods:identifier id="547F16F3CD24CF4CAD9F451B8675548F" type="ZooBank">1C308A71-A36D-4B60-8CD0-45BD70283919</mods:identifier>
</mods:mods>
<treatment id="A032F1403BB75883B1B1F4D109645B23" ID-arpha="A032F140-3BB7-5883-B1B1-F4D109645B23" ID-zoobank="https://zoobank.org/D3E13A53-73C9-4BFC-9031-CA836A3E7775" LSID="urn:lsid:zoobank.org:act:D3E13A53-73C9-4BFC-9031-CA836A3E7775" LSID-ZBK="urn:lsid:zoobank.org:act:D3E13A53-73C9-4BFC-9031-CA836A3E7775" httpUri="http://treatment.plazi.org/id/A032F1403BB75883B1B1F4D109645B23">
<subSubSection id="9EF88920F77887B67E2D7CAB148633E8" type="nomenclature">
<paragraph id="D901D99C108717FA0AFA16D42473FDB1">
<taxonomicName id="4F0563793491F763AC9BC644942323A3" authority="Biondi &amp; Iannella &amp; D'Alessandro, 2024" authorityName="Biondi &amp; Iannella &amp; D'Alessandro" authorityYear="2024" class="Insecta" family="Chrysomelidae" genus="Argopistes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="keiseri" status="sp. nov.">
<emphasis id="6DB84A690EF377AACAB0D22C9F9F4998" italics="true">Argopistes keiseri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9382711A9C656506DE3B4ED02090F7FF" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="764A50C3D9F191FEA1AF25E19C585C42" type="description">
<paragraph id="EE49C17F0EECEC611DF94CB4CCE639B0">
<figureCitation id="C57907A0C6FE4671F904454B5D9FB6D0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">Figs 4</figureCitation>
,
<figureCitation id="47E968803B87233F3C10DDE301D7200F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution and habitat suitability of the Argopistes species in Madagascar. A occurrence locality for each species B probability of presence of the occurrence localities, and potential suitability areas for the genus in Madagascar from ENMs." figureDoi="10.3897/zookeys.1202.122977.figure8" httpUri="https://binary.pensoft.net/fig/1055978">8 A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2PAG" type="type material">
<paragraph id="F159AB1F83AE5BFEC7C04151BB0C0545">
<heading id="BC6DF0DBDEF3EB01116C77790B4DE344" reason="title">Type material.</heading>
</paragraph>
<paragraph id="9EA6CCB67C73F0613498E59C9E327084">
<materialsCitation id="27BB8D690A973A41565005A2A97BF642" collectingDate="1995-11-11" collectingDateMax="1995-11-12" collectingDateMin="1995-11-11" collectionCode="BAQ" collectorName="Ivo Jenis" country="Madagascar" latitude="-17.987667" location="Tamatave prov." longLatPrecision="13" longitude="46.90333" municipality="Manankazo" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="2DB6A01F0A9C285FD12E9E1F21D10EF9" bold="true" italics="true">
<typeStatus id="069FDFB8721E24379BC478604557E547" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="86482B7A1026FDB2C8FA0E84AA5ACA48" count="1" type="male">♂</specimenCount>
: “
<collectingCountry id="B701C14CE1AC9AF3904EBB28414A13D1" name="Madagascar">Madagascar</collectingCountry>
/
<location id="957962A86A6E1B52B3310A355606ECFB" LSID="urn:lsid:plazi:treatment:A032F1403BB75883B1B1F4D109645B23:957962A86A6E1B52B3310A355606ECFB" country="Madagascar" latitude="-17.987667" longLatPrecision="13" longitude="46.90333" municipality="Manankazo" name="Tamatave prov.">Tamatave prov.</location>
/
<collectingMunicipality id="4B2030B7ACFCE5FB6B6F861ABD0EF48F">Manankazo</collectingMunicipality>
env. /
<date id="51EE0DDBDBF56B8D6FFD35F047000260" value="1995-11-11" valueMax="1995-11-12" valueMin="1995-11-11">
<collectingDate id="20FF663FDAB5224A417BC97BF764068C" value="1995-11-11" valueMax="1995-11-12" valueMin="1995-11-11">11-12.11. 1995</collectingDate>
</date>
/
<collectorName id="C5BB5CB7FC23009C2989199205346A5A">Ivo Jeniš</collectorName>
leg. ” [printed on white card] [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJQAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[46.903333,-17.987667]}">
<geoCoordinate id="BA188FA40AD4D848571ED07D164CF929" degrees="17" direction="south" minutes="59.26" orientation="latitude" precision="9" value="-17.987667">17 ° 59.26 ' S</geoCoordinate>
;
<geoCoordinate id="3D9597BA5BE83758F1CF748AEFBB193F" degrees="46" direction="east" minutes="54.20" orientation="longitude" precision="9" value="46.90333">46 ° 54.20 ' E</geoCoordinate>
</named-content>
] (
<abbrev id="ABBRID0EOQAG" xlink_title="Italy, University of L’Aquila, Collection of M. Biondi">
<collectionCode id="989B52DE2D23597D5CC683B6B902E7CA">BAQ</collectionCode>
</abbrev>
)
</materialsCitation>
.
<typeStatus id="A6163605AE28CB994B37F6D4BDD7B483" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="E1F6BA38C776523829F4FDA332C933BD" count="1" type="male">1 ♂</specimenCount>
and
<specimenCount id="610E511338358B2D7A8B01977561CFAC" count="1" type="female">1 ♀</specimenCount>
“
<collectingCountry id="7C74D40A71B6BD3A28662F54A46D4239" name="Madagascar">Madagascar</collectingCountry>
Tam. / Moramanga /
<date id="0540B810A42324231EBE85827D8EDC07" value="1957-12-20">20. xii. 1957</date>
F. Keiser ” [printed on pink card] // “ Non Cocc. Det. H. Fürsch ” [printed on white card] [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVQAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[48.224500,-18.948833]}">
<geoCoordinate id="BB8DF66C64A44114AB6B8F187F413924" degrees="18" direction="south" minutes="56.93" orientation="latitude" precision="9" value="-18.948833">18 ° 56.93 ' S</geoCoordinate>
;
<geoCoordinate id="F9DE0509BF0D5A097E8A67C82AD7AC65" degrees="48" direction="east" minutes="13.47" orientation="longitude" precision="9" value="48.2245">48 ° 13.47 ' E</geoCoordinate>
</named-content>
] (
<named-content id="A6A0B2845544E60E019A1114A603F57A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-basel-0" xlink_title="Switzerland, Basel, Naturhistorisches Museum">
<collectionCode id="933AEB6FA81F566CACA5C68010401561" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" type="">NHMB</collectionCode>
</named-content>
);
<specimenCount id="8C7B32986B63D47FFD554B6877974EBB" count="2" type="female">2 ♀♀</specimenCount>
, “
<collectingCountry id="51C89D1E13F5C3FF200C38B47BACDF6E" name="Madagascar">Madagascar</collectingCountry>
/ Tamatave prov. / Moramanga env. / 24.2-
<date id="3CD7D3EFD0F14417900EC8290405C7F9" value="1995-03-01">1.3. 1995</date>
/ Ivo Jeniš leg. ” [printed on white card] [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBRAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[48.224500,-18.948833]}">
<geoCoordinate id="71418EBB974B50FDF292451EB6974B93" degrees="18" direction="south" minutes="56.93" orientation="latitude" precision="9" value="-18.948833">18 ° 56.93 ' S</geoCoordinate>
;
<geoCoordinate id="03185565BC7A9E6BE0F06150B31B790C" degrees="48" direction="east" minutes="13.47" orientation="longitude" precision="9" value="48.2245">48 ° 13.47 ' E</geoCoordinate>
</named-content>
] (
<abbrev id="ABBRID0EGRAG" xlink_title="Italy, University of L’Aquila, Collection of M. Biondi">
<collectionCode id="01B268A6B935ACC783E5FF9A00EA8A1B">BAQ</collectionCode>
</abbrev>
);
<specimenCount id="12014DF78F55C364DE8BDA309B729FC8" count="2" type="female">2 ♀♀</specimenCount>
, “
<collectingCountry id="C9B5AF2081F340F774B5BDE32EB1B69E" name="Madagascar">Madagascar</collectingCountry>
/ Tamatave prov. / Moramanga env. /
<date id="3BA57D09B17F877DB6D1A757886A1E2D" value="1995-11-25" valueMax="1995-11-27" valueMin="1995-11-25">25-27.11. 1995</date>
/ Ivo Jeniš leg. ” [printed on white card] [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENRAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[48.224500,-18.948833]}">
<geoCoordinate id="74669747762928119DB8C65D6BB8FBCF" degrees="18" direction="south" minutes="56.93" orientation="latitude" precision="9" value="-18.948833">18 ° 56.93 ' S</geoCoordinate>
;
<geoCoordinate id="5EB77C6D5C13EC9B720BAD75A222B8ED" degrees="48" direction="east" minutes="13.47" orientation="longitude" precision="9" value="48.2245">48 ° 13.47 ' E</geoCoordinate>
</named-content>
] (
<abbrev id="ABBRID0ESRAG" xlink_title="Italy, University of L’Aquila, Collection of M. Biondi">
<collectionCode id="35661C5378ECC8D07CF939964F486861">BAQ</collectionCode>
</abbrev>
);
<specimenCount id="729BD445DB2B84DF8E21E9E57D19FE22" count="2" type="female">2 ♀♀</specimenCount>
, “
<collectingCountry id="3D09442A3A159E53A0A27162666039E7" name="Madagascar">Madagascar</collectingCountry>
/ Tamatave prov. / Moramanga env. /
<date id="C37D03CCD03944D41F418AA02DF10A70" value="1996-12-21" valueMax="1996-12-24" valueMin="1996-12-21">21-24.12. 1996</date>
/ Ivo Jeniš leg. ” [printed on white card] [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZRAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[48.224500,-18.948833]}">
<geoCoordinate id="8E692727F37D067119A3510686647A79" degrees="18" direction="south" minutes="56.93" orientation="latitude" precision="9" value="-18.948833">18 ° 56.93 ' S</geoCoordinate>
;
<geoCoordinate id="B5F6B5D85CF9C9593992208C046BF346" degrees="48" direction="east" minutes="13.47" orientation="longitude" precision="9" value="48.2245">48 ° 13.47 ' E</geoCoordinate>
</named-content>
] (
<abbrev id="ABBRID0E5RAG" xlink_title="Italy, University of L’Aquila, Collection of M. Biondi">
<collectionCode id="4AB5957B5BCC1EF95E38CB6FD51B57C1">BAQ</collectionCode>
</abbrev>
);
<specimenCount id="4EC1E84DADB0087F3BBC1E041EB2CDDE" count="1" type="male">1 ♂</specimenCount>
and
<specimenCount id="776E34EFC448F9DCF8100755EFCE26AB" count="1" type="female">1 ♀</specimenCount>
, “
<collectingCountry id="5E0C49E569F9D790DB8577CE0B334E06" name="Madagascar">Madagascar</collectingCountry>
/ Tamatave prov. / Maromizaha /
<date id="7710398E8EE4B95240E8303AA1C0093D" value="1995-02-21">21. II. 1995</date>
Ivo Jeniš ” [printed on white card] [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFSAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[48.465000,-18.976167]}">
<geoCoordinate id="176667EB27B65C6B5631225BAF9FC788" degrees="18" direction="south" minutes="58.57" orientation="latitude" precision="9" value="-18.976166">18 ° 58.57 ' S</geoCoordinate>
;
<geoCoordinate id="F4F272418775BAD03449B7501137A8E0" degrees="48" direction="east" minutes="27.90" orientation="longitude" precision="9" value="48.465">48 ° 27.90 ' E</geoCoordinate>
</named-content>
] (
<abbrev id="ABBRID0EKSAG" xlink_title="Italy, University of L’Aquila, Collection of M. Biondi">
<collectionCode id="F8DFE8189D0BF36B9D9DE2D5222AAED7">BAQ</collectionCode>
</abbrev>
);
<specimenCount id="D83596C3DFB6A047D510591C02931DF7" count="2" type="male">2 ♂</specimenCount>
and
<specimenCount id="3B9F554DDC72BFF1E48680A9BA6FBD81" count="3" type="female">3 ♀</specimenCount>
, “
<collectingCountry id="C64027EB9281FBADB3E1AAB71A687BDF" name="Madagascar">Madagascar</collectingCountry>
/
<collectingRegion id="98BAE710B1D3C1B8352301DA083C43FB" country="Madagascar" name="Fianarantsoa">Fianarantsoa prov.</collectingRegion>
/ Ranomafana env. /
<date id="7CAFD77D38E1A75730A9256E7BBAC34D" value="1995-01-28" valueMax="1995-02-06" valueMin="1995-01-28">28. I- 6. II. 1995</date>
Ivo Jeniš ” [printed on white card] [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERSAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[47.452000,-21.262667]}">
<geoCoordinate id="20A172E3A001B7C86D1F3BE031ACF909" degrees="21" direction="south" minutes="15.76" orientation="latitude" precision="9" value="-21.262667">21 ° 15.76 ' S</geoCoordinate>
;
<geoCoordinate id="B31616E888DF16820DA1A71B6B41A5D3" degrees="47" direction="east" minutes="27.12" orientation="longitude" precision="9" value="47.452">47 ° 27.12 ' E</geoCoordinate>
</named-content>
] (
<abbrev id="ABBRID0EWSAG" xlink_title="Italy, University of L’Aquila, Collection of M. Biondi">
<collectionCode id="6A08B048C3BB06800CAF032D9E4B8C49">BAQ</collectionCode>
</abbrev>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1SAG" type="diagnosis">
<paragraph id="A9F083FC71CFC1333629B913A6F2F3DB">
<heading id="1E5B189A46EA6409279BDD7F13310190" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="491112FACA6ABE6C81231A6F29B5D6F4">
<taxonomicName id="148B6AD29A889FF85AFB7D081E776617" authority="Biondi &amp; Iannella &amp; D'Alessandro, 2024" authorityName="Biondi &amp; Iannella &amp; D'Alessandro" authorityYear="2024" class="Insecta" family="Chrysomelidae" genus="Argopistes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="keiseri" status="sp. nov.">
<emphasis id="90760D69C07282D5C1742F6CA0484EF8" italics="true">Argopistes keiseri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83250AEE06C73B74640F1A9659D42962" rank="species">sp. nov.</taxonomicNameLabel>
shows major similarities with
<taxonomicName id="24E1D3083AC8E6E1F0D8127FE679F65F" authority="Biondi &amp; Iannella &amp; D'Alessandro, 2024" authorityName="Biondi &amp; Iannella &amp; D'Alessandro" authorityYear="2024" class="Insecta" family="Chrysomelidae" genus="Argopistes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="seyrigi" status="sp. nov.">
<emphasis id="9A0F00B004C319D1E1A48A18815E3B02" italics="true">A. seyrigi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AD4A4EB9B5FF2FEB550D7FE7E883D2D9" rank="species">sp. nov.</taxonomicNameLabel>
Both have the spur of hind tibiae distinctly elongate, extending significantly beyond the tibial apex (Figs
<figureCitation id="4AFFF6872E103152E183FD46F0ED1C64" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 B</figureCitation>
,
<figureCitation id="2E4DAEB663F9B9A891BE4B66D0F8CA89" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Argopistes seyrigi sp. nov. A holotype, habitus in dorsal view B ibid, ventral view C ibid, median lobe of the aedeagus. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 500 μm (C)." figureDoi="10.3897/zookeys.1202.122977.figure6" httpUri="https://binary.pensoft.net/fig/1055976">6 B</figureCitation>
), black dorsal integuments (Figs
<figureCitation id="4E1BCA91636C755BCF896D900AC5B1A4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 A</figureCitation>
,
<figureCitation id="034C079A9E336B070B47BBD4238E6CC1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Argopistes seyrigi sp. nov. A holotype, habitus in dorsal view B ibid, ventral view C ibid, median lobe of the aedeagus. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 500 μm (C)." figureDoi="10.3897/zookeys.1202.122977.figure6" httpUri="https://binary.pensoft.net/fig/1055976">6 A</figureCitation>
), and mostly confused elytral punctation.
<taxonomicName id="F0F2793EC191C67E69B95596A287DD18" authority="Biondi &amp; Iannella &amp; D'Alessandro, 2024" authorityName="Biondi &amp; Iannella &amp; D'Alessandro" authorityYear="2024" class="Insecta" family="Chrysomelidae" genus="Argopistes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="keiseri" status="sp. nov.">
<emphasis id="391A27DAD87451DA52FF5FA8439F42DB" italics="true">Argopistes keiseri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3757BDD3DF84AFA4E227EDD3E5496221" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished by the blackish abdomen and tibiae (mostly reddish brown in
<taxonomicName id="5DE5ECD138DBE21D75AFF2160301983B" authority="Biondi &amp; Iannella &amp; D'Alessandro, 2024" authorityName="Biondi &amp; Iannella &amp; D'Alessandro" authorityYear="2024" class="Insecta" family="Chrysomelidae" genus="Argopistes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="seyrigi" status="sp. nov.">
<emphasis id="78ED84A6020CD9F270FDDF3DFC66769E" italics="true">A. seyrigi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="93D0615399873A8BD8FA00CE11DAB7DC" rank="species">sp. nov.</taxonomicNameLabel>
) (Figs
<figureCitation id="D4A6CECB6BF1A5AABD191F62C3A80F41" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 B</figureCitation>
,
<figureCitation id="508725681F28D674E3C0266D70FF3BFE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Argopistes seyrigi sp. nov. A holotype, habitus in dorsal view B ibid, ventral view C ibid, median lobe of the aedeagus. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 500 μm (C)." figureDoi="10.3897/zookeys.1202.122977.figure6" httpUri="https://binary.pensoft.net/fig/1055976">6 B</figureCitation>
). Both the median lobe of aedeagus and spermatheca are diagnostic for
<taxonomicName id="67BA677D031E431E6B07617C6F331A8F" authority="Biondi &amp; Iannella &amp; D'Alessandro, 2024" authorityName="Biondi &amp; Iannella &amp; D'Alessandro" authorityYear="2024" class="Insecta" family="Chrysomelidae" genus="Argopistes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="keiseri" status="sp. nov.">
<emphasis id="169BBAB16BC5402080883308E219223D" italics="true">Argopistes keiseri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CF34FFEE25352E2C9C64C364FD059FE5" rank="species">sp. nov.</taxonomicNameLabel>
: median lobe (Fig.
<figureCitation id="1D6840E23A746BA059078723C8475778" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 D</figureCitation>
) is thickset, with irregular outline in ventral view, and clearly sinuate in lateral view; spermatheca (Fig.
<figureCitation id="FFC9BB020B6B71FDE2521C7A42BC06A7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 C</figureCitation>
) has sinuate basal part and elongate, U-shaped, and uncoiled ductus.
</paragraph>
<caption id="D6DF9503582112D8F2B74355896C806D" ID-DOI="10.3897/zookeys.1202.122977.figure4" ID-arpha="25847C34-D840-5B16-AB62-E1E416D6B8AE" httpUri="https://binary.pensoft.net/fig/1055974" startId="F4">
<paragraph id="657C8DEFAE5B97163AFAA997A0BCE034">
<label id="B76178DCD066EDB08F02CCCF11015EB4">Figure 4.</label>
</paragraph>
<paragraph id="3CBEF0A0197D29ED07AD3D2A5C287909">
<taxonomicName id="C6CFD59DEC869B587D7571BEBB2BD828" authority="Biondi &amp; Iannella &amp; D'Alessandro, 2024" authorityName="Biondi &amp; Iannella &amp; D'Alessandro" authorityYear="2024" class="Insecta" family="Chrysomelidae" genus="Argopistes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="keiseri" status="sp. nov.">
<emphasis id="D8765153D49587EAD50B2F130B750410" italics="true">Argopistes keiseri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38374E213BB3926BE1677FA8A9BD2E7F" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="670F6F01A36FB96FF9470B2377646A99" bold="true">A</emphasis>
habitus in dorsal view, male from Moramanga
<emphasis id="9062493EF65B38442061E88B8DF700BA" bold="true">B</emphasis>
ibid, ventral view
<emphasis id="A0DA54D4B25C6CD567061E91381FA70D" bold="true">C</emphasis>
spermatheca, from Ranomafana
<emphasis id="6288F760734C5C31A32655256335306B" bold="true">D</emphasis>
median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (
<emphasis id="75CA2E52DF30623D3B311CBDA0E85E1E" bold="true">A, B</emphasis>
); 300 μm (
<emphasis id="79C1764A4B0DCFBDE06A27ACAAD1BCEB" bold="true">C</emphasis>
); 500 μm (
<emphasis id="A75D6C37B22D312B4280D8EF8067BD29" bold="true">D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E4WAG" type="description">
<paragraph id="5F9A78E12E464E4358176F92437DB7F7">
<heading id="FC851BEC05CD407830D0D15E554B268D" reason="title">
Description of the
<typeStatus id="67F5F9554A9081A42F93F140804EC802" type="holotype">holotype</typeStatus>
</heading>
</paragraph>
<paragraph id="7813C7FFCB028E8915E08931540DEF32">
(
<specimenCount id="01576B985CA0B45EA5EEDECC586EF501" count="1" type="male">♂</specimenCount>
). Body roundish in dorsal view (Fig.
<figureCitation id="B2B17C0F3A56460CBFD3E2DEBD8A8369" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 A</figureCitation>
), strongly convex in lateral view; total length of body (
<abbrev id="ABBRID0EHXAG" xlink_title="total body length">
<collectionCode id="10A3C3DAD8540D2FEF7DCFB349FFFB14">LB</collectionCode>
</abbrev>
) =
<quantity id="956CC957549BC4B237BE5BAA12C6E2F0" metricMagnitude="-3" metricUnit="m" metricValue="3.48" unit="mm" value="3.48">3.48 mm</quantity>
; maximum pronotal width at the base (
<abbrev id="ABBRID0ELXAG" xlink_title="maximum width of pronotum">WP</abbrev>
=
<quantity id="653A9CB85582C9A1E1DE197393A4F82F" metricMagnitude="-3" metricUnit="m" metricValue="2.08" unit="mm" value="2.08">2.08 mm</quantity>
); maximum width of elytra in the middle (
<abbrev id="ABBRID0EPXAG" xlink_title="maximum width of elytra combined">WE</abbrev>
=
<quantity id="49A437CBA84EF131BCE81999024BBE3D" metricMagnitude="-3" metricUnit="m" metricValue="3.16" unit="mm" value="3.16">3.16 mm</quantity>
);
<abbrev id="ABBRID0ETXAG" xlink_title="maximum width of elytra combined">WE</abbrev>
/
<abbrev id="ABBRID0EXXAG" xlink_title="maximum width of pronotum">WP</abbrev>
= 1.52. Dorsal integuments (Fig.
<figureCitation id="A963304A67E492B49C1762F0F0BDD186" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 A</figureCitation>
) entirely black with weak metallic reflections; ventral parts (Fig.
<figureCitation id="6A190E20B54224A5D7538993C39F37C7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 B</figureCitation>
) entirely blackish; head black; frons and mouthparts black, with yellowish maxillary palpi; antennae yellowish (Fig.
<figureCitation id="8F162F4C6D76F56E120F6C95A5368980" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 B</figureCitation>
); legs, including articulations, black, with yellowish tarsi (Fig.
<figureCitation id="F8D72B8ABC9460E4BEECDD2363C5EC6D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 B</figureCitation>
). Head entirely hidden by the pronotum; vertex punctate, with a pair of large setiferous pores; area of frontal calli weakly raised; frons moderately elongate, roughly wrinkled; frontal ridge thin, weakly raised; frontogenal sutures thin, strongly raised; eyes large, elongate, slightly kidney-shaped; interantennal space clearly narrower than antennal sockets. Antennae (Fig.
<figureCitation id="909E1E5E0FEE3A93E837241C3B6C0170" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 B</figureCitation>
) filiform, distinctly shorter than 1 / 2 the body length (
<abbrev id="ABBRID0EPYAG" xlink_title="length of antennae">
<collectionCode id="0003333395E7C01AEC825BBC7D1C2A7C" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13468" name="Lancing College" type="Herbarium">LAN</collectionCode>
</abbrev>
=
<quantity id="895519145898FAA42A497BA7ECC70D80" metricMagnitude="-3" metricUnit="m" metricValue="1.44" unit="mm" value="1.44">1.44 mm</quantity>
;
<abbrev id="ABBRID0ETYAG" xlink_title="length of antennae">
<collectionCode id="9F0300049CAEE6422720A38B63378FBA" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13468" name="Lancing College" type="Herbarium">LAN</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EXYAG" xlink_title="total body length">
<collectionCode id="D0D8AE4B3B3F38EF9482EBE90D46F84D">LB</collectionCode>
</abbrev>
= 0.41); segments 1–2 thicker; segments 3–11 slightly and gradually flattened;
<abbrev id="ABBRID0E2YAG" xlink_title="numerical sequence from base to apex of each antennomere, proportional to the length of the first antennomere">LA</abbrev>
= 100: 51: 47: 53: 37: 48: 44: 64: 51: 50: 88. Pronotum (Fig.
<figureCitation id="137CC5861689E76E35BDFE02C846C176" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 A</figureCitation>
) distinctly transverse (
<abbrev id="ABBRID0EDZAG" xlink_title="medial length of pronotum">
<collectionCode id="AD0345059836ACEADD3E55E8602CBA61" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
= 1.00 mm;
<abbrev id="ABBRID0EHZAG" xlink_title="maximum width of pronotum">
<collectionCode id="6DBF2775B9B81988D3800169CB3B1D83">WP</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0ELZAG" xlink_title="medial length of pronotum">
<collectionCode id="2A7C4A7585AC965316DEA8DED87116B8" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
= 2.08); lateral margins strongly convergent anteriorly, straight, weakly expanded, not visible in dorsal view; basal margin arcuate and distinctly sinuate; surface sparsely micropunctate, with dense, small punctation; surface moderately raised parallel to the lateral margins; a large setiferous pore at the anterior angles. Scutellum small, subtriangular. Elytra (
<abbrev id="ABBRID0EPZAG" xlink_title="maximum length of elytra">
<collectionCode id="0E8A58EA96B826CDB7435CA96ADE93C8" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
=
<quantity id="BC4A8AF7D6A39FB1BDEAB5B884816557" metricMagnitude="-3" metricUnit="m" metricValue="3.2" unit="mm" value="3.2">3.20 mm</quantity>
;
<abbrev id="ABBRID0ETZAG" xlink_title="maximum length of elytra">
<collectionCode id="3C0A354754AA8B4E223C2BAF61BBF2B9" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EXZAG" xlink_title="medial length of pronotum">
<collectionCode id="46E7359E70C78B824E7FBC1EE6E31F0A" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
= 3.20) strongly curved laterally (Fig.
<figureCitation id="B4384A9CF973CFA04DC7A7D26AC81EDB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 A</figureCitation>
), approx. as long as wide (
<abbrev id="ABBRID0E6ZAG" xlink_title="maximum width of elytra combined">
<collectionCode id="1F60101841BC2FD3E42780A669A968BD">WE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0ED1AG" xlink_title="maximum length of elytra">
<collectionCode id="550DB5ABF07CC05AAD4524AD34B95651" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
= 1.02), jointly rounded apically; lateral margins finely bordered, visible in dorsal view; surface smooth; punctation small, dense, mostly confused, but arranged in some regular lines laterally, of which one is made of slightly larger punctures. Humeral calli moderately raised. Macropterous. Prosternum with posteriorly open procoxal cavities and large intercoxal prosternal process. Mesosternum very short. First abdominal sternite slightly longer than fifth (Fig.
<figureCitation id="3553A266E6C2DA7B314913B6DB182975" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 B</figureCitation>
); its central area bordered by ridges is quite wide, slightly narrowing posteriorly. Anterior and middle legs without modifications. Posterior femora greatly swollen (
<abbrev id="ABBRID0EL1AG" xlink_title="maximum width of hind femora">
<collectionCode id="664AAF37D604ECA0A6A8ABBEAB89C2B2">WF</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EP1AG" xlink_title="maximum length of hind femora">
<collectionCode id="C910E77A8A6CFD328F78BA2070B681C3">LF</collectionCode>
</abbrev>
= 0.68), elongate-subtriangular; posterior tibiae thick, distinctly shorter than femora, apically widened and prolonged into a spur-like process on inner side; outer side of hind tibia apically toothed; apical spur of hind tibiae simple, lanceolate, very elongate; first metatarsomere moderately enlarged. Median lobe of the aedeagus (
<abbrev id="ABBRID0ET1AG" xlink_title="length of median lobe of the aedeagus">
<collectionCode id="B064687B704A2FD1BC2D781D2AEF000D">LAED</collectionCode>
</abbrev>
=
<quantity id="507EEBBE7C52C00263444257A70A4567" metricMagnitude="-3" metricUnit="m" metricValue="1.26" unit="mm" value="1.26">1.26 mm</quantity>
;
<abbrev id="ABBRID0EX1AG" xlink_title="maximum length of elytra">
<collectionCode id="B0E99D56FD13F7208E135E8A9D35B784" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0E21AG" xlink_title="length of median lobe of the aedeagus">
<collectionCode id="BA06BC59658048F6F8255A7235681E6A">LAED</collectionCode>
</abbrev>
= 2.54) (Fig.
<figureCitation id="54E4E9D98804941020082B0CE352207A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 D</figureCitation>
) with smooth surface; in ventral view thickset, lanceolate but with slightly irregular outline; in lateral view median lobe moderately curved, thicker at the subapical part, with sinuate ventral outline; apex ventrally oriented; dorsal ligula formed by a central lobe medially incised in the apical part, and two thinner lateral lobes; its base at apical ~ 1 / 3.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ED2AG" type="variability">
<paragraph id="182138ECC9E0791455618B47C6456891">
<heading id="62313673C22F62BE19793F3B841F33C6" reason="title">Variability.</heading>
</paragraph>
<paragraph id="F107F1CD69A121355D1CDD0F59513A80">
Male (
<emphasis id="38357B6A4911B54C75C0EBAE608C1CA3" italics="true">n</emphasis>
= 5; mean and standard deviation; range):
<abbrev id="ABBRID0EL2AG" xlink_title="maximum length of elytra">
<collectionCode id="3A30505D938A27ADBF3914490D7BBE37" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
= 3.08 ±
<quantity id="D30DB02C6B6FF25698D8888FAA82ED59" metricMagnitude="-4" metricUnit="m" metricValue="1.3" unit="mm" value="0.13">0.13 mm</quantity>
(2.88 ≤
<abbrev id="ABBRID0EP2AG" xlink_title="maximum length of elytra">
<collectionCode id="98F234D692053B105A0CDB12A81BBEFF" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
≤
<quantity id="18A4E9236EEF2D65B54E169F3A51E9B8" metricMagnitude="-3" metricUnit="m" metricValue="3.2" unit="mm" value="3.2">3.20 mm</quantity>
);
<abbrev id="ABBRID0ET2AG" xlink_title="maximum width of elytra combined">WE</abbrev>
= 2.98 ±
<quantity id="D24F8473CB5435D5FB48812017B03DEF" metricMagnitude="-4" metricUnit="m" metricValue="1.3" unit="mm" value="0.13">0.13 mm</quantity>
(2.82 ≤
<abbrev id="ABBRID0EX2AG" xlink_title="maximum width of elytra combined">
<collectionCode id="634D50E7D9F18D62E5E3BE928718926A">WE</collectionCode>
</abbrev>
≤
<quantity id="9196BF52DCE83EFDC5CA738B7CF7D6F5" metricMagnitude="-3" metricUnit="m" metricValue="3.16" unit="mm" value="3.16">3.16 mm</quantity>
);
<abbrev id="ABBRID0E22AG" xlink_title="medial length of pronotum">
<collectionCode id="A437CBE5770471D02B46CEFB273FC665" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
= 0.92 ±
<quantity id="872F41329B38DF1115606DF16013E091" metricMagnitude="-5" metricUnit="m" metricValue="6.0" unit="mm" value="0.06">0.06 mm</quantity>
(0.86 ≤
<abbrev id="ABBRID0E62AG" xlink_title="medial length of pronotum">
<collectionCode id="450EFC5DC933D9BBAEC8E2542B062A69" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
≤ 1.00 mm);
<abbrev id="ABBRID0ED3AG" xlink_title="maximum width of pronotum">WP</abbrev>
= 1.98 ±
<quantity id="449B05307B4C09D3F6B3281813AA1241" metricMagnitude="-5" metricUnit="m" metricValue="9.0" unit="mm" value="0.09">0.09 mm</quantity>
(1.88 ≤
<abbrev id="ABBRID0EH3AG" xlink_title="maximum width of pronotum">
<collectionCode id="2ED0CC7A0191CF1794779190BE516410">WP</collectionCode>
</abbrev>
≤
<quantity id="3B35DD54CD9C92C9A23C06C5C778791E" metricMagnitude="-3" metricUnit="m" metricValue="2.08" unit="mm" value="2.08">2.08 mm</quantity>
);
<abbrev id="ABBRID0EL3AG" xlink_title="length of antennae">LAN</abbrev>
= 1.39 ±
<quantity id="73129C69189CAE39EE02FE0568FA92A1" metricMagnitude="-4" metricUnit="m" metricValue="1.5" unit="mm" value="0.15">0.15 mm</quantity>
(1.16 ≤
<abbrev id="ABBRID0EP3AG" xlink_title="length of antennae">
<collectionCode id="18000B354326B06E5933454DC0FC7BAB" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13468" name="Lancing College" type="Herbarium">LAN</collectionCode>
</abbrev>
≤
<quantity id="02DB6F4D7B7020067F45D7E7DA54503F" metricMagnitude="-3" metricUnit="m" metricValue="1.52" unit="mm" value="1.52">1.52 mm</quantity>
);
<abbrev id="ABBRID0ET3AG" xlink_title="length of median lobe of the aedeagus">
<collectionCode id="970E1528D2D087CDCCF6F890D7D7957B">LAED</collectionCode>
</abbrev>
= 1.38 ±
<quantity id="E1B1807BC1743F4BBA6E1A4D91F5E1D3" metricMagnitude="-5" metricUnit="m" metricValue="9.0" unit="mm" value="0.09">0.09 mm</quantity>
(1.26 ≤
<abbrev id="ABBRID0EX3AG" xlink_title="length of median lobe of the aedeagus">
<collectionCode id="0CD1DB93C8F4C7F617E29A5F29CD5EBD">LAED</collectionCode>
</abbrev>
≤
<quantity id="C9747653389A404E319B76531BFDD5A7" metricMagnitude="-3" metricUnit="m" metricValue="1.48" unit="mm" value="1.48">1.48 mm</quantity>
);
<abbrev id="ABBRID0E23AG" xlink_title="total body length">LB</abbrev>
= 3.37 ±
<quantity id="EE36E9354CAB2D122DD1E905EC4A04C7" metricMagnitude="-4" metricUnit="m" metricValue="1.2" unit="mm" value="0.12">0.12 mm</quantity>
(3.20 ≤
<abbrev id="ABBRID0E63AG" xlink_title="total body length">
<collectionCode id="C1B5B4A88B59370E6EB621EA3E448C7E">LB</collectionCode>
</abbrev>
≤
<quantity id="E2E40B0A8B81E56862C14048AE96F8A3" metricMagnitude="-3" metricUnit="m" metricValue="3.48" unit="mm" value="3.48">3.48 mm</quantity>
);
<abbrev id="ABBRID0ED4AG" xlink_title="maximum length of elytra">
<collectionCode id="6C9E5B6C70B717086065357F22677775" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EH4AG" xlink_title="medial length of pronotum">
<collectionCode id="32992645F8EA040B4D24E96BDEAEF8D1" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
= 3.35 ± 0.11 (3.20 ≤
<abbrev id="ABBRID0EL4AG" xlink_title="maximum length of elytra">
<collectionCode id="8A18DAA973B3A09A3EC75BF5F64A5D9B" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EP4AG" xlink_title="medial length of pronotum">
<collectionCode id="17BBB15F4936872A670F3F8C9405CFC9" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
≤ 3.50);
<abbrev id="ABBRID0ET4AG" xlink_title="maximum width of elytra combined">WE</abbrev>
/
<abbrev id="ABBRID0EX4AG" xlink_title="maximum width of pronotum">WP</abbrev>
= 1.50 ± 0.03 (1.44 ≤
<abbrev id="ABBRID0E24AG" xlink_title="maximum width of elytra combined">
<collectionCode id="24921FB859996A1DD84F7067CEDFFC50">WE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0E64AG" xlink_title="maximum width of pronotum">
<collectionCode id="4815B76F4BA38E4B15FF24C986AE8C87">WP</collectionCode>
</abbrev>
≤ 1.53);
<abbrev id="ABBRID0ED5AG" xlink_title="maximum width of pronotum">WP</abbrev>
/
<abbrev id="ABBRID0EH5AG" xlink_title="medial length of pronotum">
<collectionCode id="4BA40E65CBCCC4569DA4596BD00DF82D" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
= 2.16 ± 0.04 (2.08 ≤
<abbrev id="ABBRID0EL5AG" xlink_title="maximum width of pronotum">
<collectionCode id="84C1A2FB5F7B4A62D3FB60191EA9CF96">WP</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EP5AG" xlink_title="medial length of pronotum">
<collectionCode id="F70443642100F8BDF13E02224839B7FA" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
≤ 2.19);
<abbrev id="ABBRID0ET5AG" xlink_title="maximum width of elytra combined">WE</abbrev>
/
<abbrev id="ABBRID0EX5AG" xlink_title="maximum length of elytra">
<collectionCode id="6CB8D089E355D68A39E8517DFDA64DDD" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
= 0.97 ± 0.03 (0.94 ≤
<abbrev id="ABBRID0E25AG" xlink_title="maximum width of elytra combined">
<collectionCode id="D7DBAB6C3F3CFD9D17FEA5FF9AA4F71C">WE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0E65AG" xlink_title="maximum length of elytra">
<collectionCode id="0F1107B85295A2528FB5FD49A7C58821" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
≤ 0.99);
<abbrev id="ABBRID0ED6AG" xlink_title="length of antennae">LAN</abbrev>
/
<abbrev id="ABBRID0EH6AG" xlink_title="total body length">LB</abbrev>
= 0.41 ± 0.04 (0.35 ≤
<abbrev id="ABBRID0EL6AG" xlink_title="length of antennae">
<collectionCode id="42EC3158F0B878DB6E30E1C7ED00DAA5" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13468" name="Lancing College" type="Herbarium">LAN</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EP6AG" xlink_title="total body length">
<collectionCode id="22CBF7D83BD65B6BA85F26892B8BBDC6">LB</collectionCode>
</abbrev>
≤ 0.45);
<abbrev id="ABBRID0ET6AG" xlink_title="maximum length of elytra">
<collectionCode id="F3775357E01998D2E1C952C2AACED961" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EX6AG" xlink_title="length of median lobe of the aedeagus">
<collectionCode id="4F34028402668C497C3F87D1384DB934">LAED</collectionCode>
</abbrev>
= 2.24 ± 0.18 (2.04 ≤
<abbrev id="ABBRID0E26AG" xlink_title="maximum length of elytra">
<collectionCode id="EEE8979E280F5C19E035BA822E47B9A7" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0E66AG" xlink_title="length of median lobe of the aedeagus">
<collectionCode id="89C3B1CF8A43462B744BA820D563A719">LAED</collectionCode>
</abbrev>
≤ 2.54). Female (n = 11; mean and standard deviation; range):
<abbrev id="ABBRID0EDABG" xlink_title="maximum length of elytra">
<collectionCode id="8F5B8B0FC8ABFFB71ADB844890812050" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
= 3.35 ±
<quantity id="1E689A0ABEE7CF8A00AB3ACAEAAE0BB9" metricMagnitude="-5" metricUnit="m" metricValue="4.0" unit="mm" value="0.04">0.04 mm</quantity>
(3.32 ≤
<abbrev id="ABBRID0EHABG" xlink_title="maximum length of elytra">
<collectionCode id="82CB4490AF2546346C0A98BBC311F363" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
≤
<quantity id="18876591E5F9BDFAE8C2E1075065776A" metricMagnitude="-3" metricUnit="m" metricValue="3.4" unit="mm" value="3.4">3.40 mm</quantity>
);
<abbrev id="ABBRID0ELABG" xlink_title="maximum width of elytra combined">WE</abbrev>
= 3.20 ±
<quantity id="D88DA83462406FEAD754A31DBD3D1BDC" metricMagnitude="-5" metricUnit="m" metricValue="6.0" unit="mm" value="0.06">0.06 mm</quantity>
(3.12 ≤
<abbrev id="ABBRID0EPABG" xlink_title="maximum width of elytra combined">
<collectionCode id="A8CE697C4021BBE8345360E2BA43F1CB">WE</collectionCode>
</abbrev>
≤
<quantity id="4FC2F158CC0B0B9A7AE919AC305EB32D" metricMagnitude="-3" metricUnit="m" metricValue="3.24" unit="mm" value="3.24">3.24 mm</quantity>
);
<abbrev id="ABBRID0ETABG" xlink_title="medial length of pronotum">
<collectionCode id="C28B542EE3901724B709CEBC8A3BF6A7" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
= 1.02 ±
<quantity id="89B0132377EC654138DD0A80498DB246" metricMagnitude="-5" metricUnit="m" metricValue="3.0" unit="mm" value="0.03">0.03 mm</quantity>
(0.98 ≤
<abbrev id="ABBRID0EXABG" xlink_title="medial length of pronotum">
<collectionCode id="7C1AEA5E073211AF201CD029C3B76BCC" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
≤
<quantity id="080F25D4476A356198913C34EC2EE974" metricMagnitude="-3" metricUnit="m" metricValue="1.04" unit="mm" value="1.04">1.04 mm</quantity>
);
<abbrev id="ABBRID0E2ABG" xlink_title="maximum width of pronotum">WP</abbrev>
= 2.10 ±
<quantity id="EE672B42F98171EDFA26B3EE29141D23" metricMagnitude="-5" metricUnit="m" metricValue="5.0" unit="mm" value="0.05">0.05 mm</quantity>
(2.02 ≤
<abbrev id="ABBRID0E6ABG" xlink_title="maximum width of pronotum">
<collectionCode id="A57B9AD6488D5F4C0DF1BC717C4F3446">WP</collectionCode>
</abbrev>
≤
<quantity id="DCAA606A155CA561FA0C4E604CDC0773" metricMagnitude="-3" metricUnit="m" metricValue="2.12" unit="mm" value="2.12">2.12 mm</quantity>
);
<abbrev id="ABBRID0EDBBG" xlink_title="length of antennae">LAN</abbrev>
= 1.34 ±
<quantity id="1315A774E6B628D7BE62E1D90ACBC892" metricMagnitude="-5" metricUnit="m" metricValue="8.0" unit="mm" value="0.08">0.08 mm</quantity>
(1.28 ≤
<abbrev id="ABBRID0EHBBG" xlink_title="length of antennae">
<collectionCode id="6523F0095DCFC9D39423BF7FA23A3060" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13468" name="Lancing College" type="Herbarium">LAN</collectionCode>
</abbrev>
≤
<quantity id="EE5EEBF1A9B860AA1B5744DD56319C72" metricMagnitude="-3" metricUnit="m" metricValue="1.44" unit="mm" value="1.44">1.44 mm</quantity>
);
<abbrev id="ABBRID0ELBBG" xlink_title="maximum length of spermathecal capsule">
<collectionCode id="2204B1DBBB8CA1918E6B7CB8E2E6C971">LSPC</collectionCode>
</abbrev>
= 0.40 ±
<quantity id="45D133EA1E70218DFC3C9BA7773E61EA" metricMagnitude="-5" metricUnit="m" metricValue="1.0" unit="mm" value="0.01">0.01 mm</quantity>
(0.38 ≤
<abbrev id="ABBRID0EPBBG" xlink_title="maximum length of spermathecal capsule">
<collectionCode id="EFBD081DF7A23A6A15618612E2BE6840">LSPC</collectionCode>
</abbrev>
≤
<quantity id="510A0AD090FA4FA48A603C84B7001AF3" metricMagnitude="-4" metricUnit="m" metricValue="4.0" unit="mm" value="0.4">0.40 mm</quantity>
);
<abbrev id="ABBRID0ETBBG" xlink_title="total body length">LB</abbrev>
= 3.66 ±
<quantity id="0C047BF5D6239C6924AC4B842602C2C3" metricMagnitude="-5" metricUnit="m" metricValue="7" unit="mm" value="0.07">0.07 mm</quantity>
(3.58 ≤
<abbrev id="ABBRID0EXBBG" xlink_title="total body length">
<collectionCode id="8518D54E8DFE9039655F2EDA9E143D28">LB</collectionCode>
</abbrev>
≤
<quantity id="B932E1E5296DD68AFE6BFE1B5CBE1C10" metricMagnitude="-3" metricUnit="m" metricValue="3.72" unit="mm" value="3.72">3.72 mm</quantity>
);
<abbrev id="ABBRID0E2BBG" xlink_title="maximum length of elytra">
<collectionCode id="69C23A85B465ADA207A0ED8AB0E8DCD1" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0E6BBG" xlink_title="medial length of pronotum">
<collectionCode id="E0263363E6B07CCD7AFADC8B0C512B1E" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
= 3.30 ± 0.07 (3.23 ≤
<abbrev id="ABBRID0EDCBG" xlink_title="maximum length of elytra">
<collectionCode id="EFEEC294C3A8FF1C212054F9B35C2774" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EHCBG" xlink_title="medial length of pronotum">
<collectionCode id="46DEA9F310EBAEC80CFEE99297340845" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
≤ 3.39);
<abbrev id="ABBRID0ELCBG" xlink_title="maximum width of elytra combined">WE</abbrev>
/
<abbrev id="ABBRID0EPCBG" xlink_title="maximum width of pronotum">WP</abbrev>
= 1.53 ± 0.01 (1.51 ≤
<abbrev id="ABBRID0ETCBG" xlink_title="maximum width of elytra combined">
<collectionCode id="74E33807BA4CAC5B9B9922071F7B5E90">WE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EXCBG" xlink_title="maximum width of pronotum">
<collectionCode id="0B8FB8AFF6B2EA7174D927E2C562F428">WP</collectionCode>
</abbrev>
≤ 1.54);
<abbrev id="ABBRID0E2CBG" xlink_title="maximum width of pronotum">WP</abbrev>
/
<abbrev id="ABBRID0E6CBG" xlink_title="medial length of pronotum">
<collectionCode id="3912EC152AEAD1805A07CCD9B2E6B660" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
= 2.06 ± 0.04 (2.04 ≤
<abbrev id="ABBRID0EDDBG" xlink_title="maximum width of pronotum">
<collectionCode id="BDE8966EA9F6C09818274357036C32EB">WP</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EHDBG" xlink_title="medial length of pronotum">
<collectionCode id="7879FBB775B3B6508F59B484B5656BEC" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
</abbrev>
≤ 2.12);
<abbrev id="ABBRID0ELDBG" xlink_title="maximum width of elytra combined">WE</abbrev>
/
<abbrev id="ABBRID0EPDBG" xlink_title="maximum length of elytra">
<collectionCode id="FC0DCB32EA9E97DF2C201F841FD86A0C" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
= 0.96 ± 0.02 (0.94 ≤
<abbrev id="ABBRID0ETDBG" xlink_title="maximum width of elytra combined">
<collectionCode id="331CEE19C06D420F8111E406A0F4CB5A">WE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EXDBG" xlink_title="maximum length of elytra">
<collectionCode id="047B3D53E7206AD2D29B6E6FACC01035" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
≤ 0.98);
<abbrev id="ABBRID0E2DBG" xlink_title="length of antennae">LAN</abbrev>
/
<abbrev id="ABBRID0E6DBG" xlink_title="total body length">LB</abbrev>
= 0.37 ± 0.01 (0.35 ≤
<abbrev id="ABBRID0EDEBG" xlink_title="length of antennae">
<collectionCode id="BCD615C39614D0C696487D2D41298612" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13468" name="Lancing College" type="Herbarium">LAN</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EHEBG" xlink_title="total body length">
<collectionCode id="F313F37B5DA878A09C26D1165A970CA0">LB</collectionCode>
</abbrev>
≤ 0.39);
<abbrev id="ABBRID0ELEBG" xlink_title="maximum length of elytra">
<collectionCode id="AC67621C1B53DD92EB3D6247290459BE" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EPEBG" xlink_title="maximum length of spermathecal capsule">
<collectionCode id="3CD351B169B5FD3DE7766E7DBA007EEB">LSPC</collectionCode>
</abbrev>
= 8.49 ± 0.19 (8.30 ≤
<abbrev id="ABBRID0ETEBG" xlink_title="maximum length of elytra">
<collectionCode id="9E3979514F454564D0649DF6F9073084" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EXEBG" xlink_title="maximum length of spermathecal capsule">
<collectionCode id="E3CFF19EEEE49E797021FEFAE20C6675">LSPC</collectionCode>
</abbrev>
≤ 8.74).
</paragraph>
<paragraph id="C60BD62E900D6E22728AB06F8E1ED595">
Male and female
<typeStatus id="4DAA2566D05DA24D765F8FC5EA06DCD5" type="paratype">paratypes</typeStatus>
very similar in shape, size, and color to the
<typeStatus id="C27CD85D8E1A64D027E0479941F06C67" type="holotype">holotype</typeStatus>
. The arrangement of elytral punctation in 9 (+ 1 sutural) regular rows is better visible in some specimens. Spermatheca (Fig.
<figureCitation id="A5298BFD821BCC80626547AB911BE20E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Argopistes keiseri sp. nov. A habitus in dorsal view, male from Moramanga B ibid, ventral view C spermatheca, from Ranomafana D median lobe of the aedeagus, from left to right in dorsal, ventral, and lateral view, from Ranomafana, and two additional lateral view from Manankazo, and Maromizama. Abbreviations: as: apical spur of hind tibia. Scale bars: 3 mm (A, B); 300 μm (C); 500 μm (D)." figureDoi="10.3897/zookeys.1202.122977.figure4" httpUri="https://binary.pensoft.net/fig/1055974">4 C</figureCitation>
) with subcylindical, sinuate basal part; collum short; apical part moderately elongate, gradually narrowing, slightly wrinkled; ductus subventrally inserted, quite thickset, elongate, U-shaped, uncoiled.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBFBG" type="etymology">
<paragraph id="D68D7CBD831D1182FAD786AADC4E1C12">
<heading id="2A11517C857FFA0CC41556EFDF262E1B" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="4DC7130C80B0514D3F84319502850450">
The specific epithet refers to the first collector of the new species: Alfred “ Fred ” Kaiser (1895–1969) from
<collectingCountry id="473748CDFBFC5035278C939F8E6B73EA" name="Switzerland">Switzerland</collectingCountry>
, renowned expert on
<taxonomicName id="8F23540A531A4F7E56312A1911F9B9D4" class="Insecta" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
<taxonomicName id="CD766F930F2969D820E5F760A14E283A" authorityName="Rondani" authorityYear="1856" class="Insecta" family="Syrphidae" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="family">Syrphidae</taxonomicName>
from
<collectingCountry id="A5F98913BC702F78AE02038993F0CA29" name="Madagascar">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERFBG" type="distribution">
<paragraph id="43B6BB36542515D553EB931716CB6400">
<heading id="925447C52DC2A6E04252F9F13000595A" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="A28D1D170D504904A73DD3AAEBD210C3">
Central-Eastern
<collectingCountry id="EE8ED28D805608DE62A2CEF700D4867A" name="Madagascar">Madagascar</collectingCountry>
(
<collectingRegion id="1890763BC3D9B4906B3EF4F14429BBA7" country="Madagascar" name="Toamasina">Toamasina province</collectingRegion>
; Fig.
<figureCitation id="7E9BE15FCE82351BFC2D06FF344D1149" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution and habitat suitability of the Argopistes species in Madagascar. A occurrence locality for each species B probability of presence of the occurrence localities, and potential suitability areas for the genus in Madagascar from ENMs." figureDoi="10.3897/zookeys.1202.122977.figure8" httpUri="https://binary.pensoft.net/fig/1055978">8 A</figureCitation>
). Malagasy chorotype.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2FBG" type="Ecological notes">
<paragraph id="D202ABADC2098BA7AE0E3859A8BC73E2">
<heading id="7C9E4B74738D09414089A78319CCE53D" reason="title">Ecological notes.</heading>
</paragraph>
<paragraph id="271A1C5FA5A4C2D974133B5FB593684C">Host plant unknown. Collection localities fall within areas characterized by the vegetation division ‘ Malagasy Evergreen &amp; Semi-Evergreen Forest’.</paragraph>
</subSubSection>
</treatment>
</document>