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<emphasis id="9961E936AF68D0D5887A17F1E6A64577" italics="true">Leptotyphlops</emphasis>
cf.
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(Peters, 1854)
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<heading id="C20DF58265355135AEE3031A04C956D7" reason="title">Record.</heading>
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<paragraph id="AD2E95ED4F0FF23CE5A1714B41A4CB2F">
vic. Mamué [
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,
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,
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264756).
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<paragraph id="8B1BBE3AA9CBFF5F5667F96642BF1BCD">
<heading id="ACA13482798670457D81F7F8CFE82BC4" reason="title">Comments.</heading>
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<emphasis id="78CCE792F83761B9573C9CADEF1ABA12" italics="true">Leptotyphlops scutifrons</emphasis>
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is widely distributed in southern Africa. It was recorded from several localities in the Angolan Plateau (
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;
<bibRefCitation id="8D1FAC84047856B6A04B1F39AD020588" author="Conradie W &amp; Bills R &amp; Branch WR" refId="B50" refString="Conradie W, Bills R, Branch WR (2016 b) The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian and Reptile Conservation 10 (2) [Special Section]: 6 36.">Conradie et al. 2016 b</bibRefCitation>
;
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). Although some historical records may refer to
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<emphasis id="4D1625BD181FCC9E03338AB77D9C25B6" italics="true">Namibiana latifrons</emphasis>
(Sternfeld, 1908) (
<bibRefCitation id="A6EDB5B93B70B03FA9B4504418CD5F06" author="Marques" etAl="et al." firstAuthor="Marques" refId="B86" refString="Marques MP, Ceríaco LMP, Blackburn DC, Bauer AM (2018) Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola Atlas of Historical and Bibliographic Records (1840 2017). Proceedings of the California Academy of Sciences 4 (65, Supplement II): 1 501." year="2018">Marques et al. 2018</bibRefCitation>
)
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, our specimen agrees almost completely with the diagnosis of
<taxonomicName id="B1612105BDD347CF2879793E5B7DCB5C" baseAuthorityName="Peters" baseAuthorityYear="1854" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" phylum="Chordata" rank="species" species="scutifrons">
<emphasis id="0A8F06AD1E05E5BF76D4F1ACC6CFA831" italics="true">L. scutifrons</emphasis>
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that was provided by
<bibRefCitation id="C7995B3A7EE102973D0E6687CBEB703E" author="Broadley DG &amp; Broadley S" firstAuthor="Broadley" journalOrPublisher="Syntarsus" pagination="1-36" refId="B26" refString="Broadley DG, Broadley S (1999) A review of the African Worm Snakes from south of latitude 12 ºS (Serpentes: Leptotyphlopidae). Syntarsus 5: 1 36." title="A review of the African Worm Snakes from south of latitude 12 ºS (Serpentes: Leptotyphlopidae)." volume="5" year="1999">Broadley and Broadley (1999)</bibRefCitation>
. Nevertheless, this taxon should be treated as a species complex with multiple divergent lineages (
<bibRefCitation id="68E29D645CEA4220977DF75490A30B11" DOI="10.11646/zootaxa.2244.1.1" author="Adalsteinsson" etAl="et al." firstAuthor="Adalsteinsson" issue="1" journalOrPublisher="Zootaxa" pagination="1-50" refId="B1" refString="Adalsteinsson SA, Branch WR, Trape S, Vitt LJ, Hedges SB (2009) Molecular phylogeny, classification, and biogeography of snakes of the Family Leptotyphlopidae (Reptilia, Squamata). Zootaxa 2244 (1): 1 50. https://doi.org/10.11646/zootaxa.2244.1.1" title="Molecular phylogeny, classification, and biogeography of snakes of the Family Leptotyphlopidae (Reptilia, Squamata)." volume="2244" year="2009">Adalsteinsson et al. 2009</bibRefCitation>
). In
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, it is mostly restricted to the highlands of the Escarpment and Central Plateau (
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).
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<paragraph id="36E8B87DE224C7A69F9809C81E248C1A" pageId="19" pageNumber="20">
<pageBreakToken id="3B8DB81C782B2CBE011EAABF52360503" pageId="19" pageNumber="20" start="start">Taxon</pageBreakToken>
classification Animalia Stylommatophora Amphibulimidae
</paragraph>
</subSubSection>
<subSubSection id="DC98967538B0D7888DE9A3E1E9290F26" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="D8372CF274686D7AFE738CDCB84418AD" pageId="19" pageNumber="20">
<taxonomicName id="C1DC511ACCC1422A62761A5E17A640B0" ID-CoL="4K6XR" authority="Breure &amp; Araujo, 2015" authorityName="Breure &amp; Araujo" authorityYear="2015" class="Gastropoda" family="Amphibulimidae" genus="Plekocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plekocheilus (Plekocheilus) cecepeus" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="cecepeus" subGenus="Plekocheilus">Plekocheilus (Plekocheilus) cecepeus Breure &amp; Araujo, 2015</taxonomicName>
Figs 8
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</paragraph>
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<paragraph id="EB9B5F1768D7B3F100559734D78D47C1" pageId="19" pageNumber="20">
<taxonomicName id="F051160C7798ABF3A8B73E928A2CF35A" class="Gastropoda" family="Amphibulimidae" genus="Plekocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plekocheilus (Plekocheilus) cecepeus" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="cecepeus" subGenus="Plekocheilus">Plekocheilus (Plekocheilus) cecepeus</taxonomicName>
<bibRefCitation id="CD6AA5DC401ABA01BF824B0AACDAD1CD" author="Breure, ASH" journalOrPublisher="ZooKeys" pageId="84" pageNumber="85" pagination="85 - 93" title="A snail in the long tail: a new Plekocheilus species collected by the ' Comision Cientifica del Pacifico' (Mollusca, Gastropoda, Amphibulimidae)." url="10.3897/zookeys.516.10228" volume="516" year="2015">Breure and Araujo 2015</bibRefCitation>
: 89, fig. 2.
</paragraph>
</subSubSection>
<subSubSection id="5B17FB4CBB4937EB5203F69DE03223CB" pageId="19" pageNumber="20" type="type locality">
<paragraph id="A0EE256669EDD63B297E37B17BF51DD9" pageId="19" pageNumber="20">Type locality.</paragraph>
<paragraph id="770ACB686C3CBCC1A06E18A6AE1BDCA9" pageId="19" pageNumber="20">&quot;Ecuador, Quito&quot;.</paragraph>
</subSubSection>
<subSubSection id="1DFCC1555EF280C696B8079D38BCAF1A" pageId="19" pageNumber="20" type="type material">
<paragraph id="1C51DCED273ADD13098CF194FC7C0592" pageId="19" pageNumber="20">Type material.</paragraph>
<paragraph id="BB102B3AC9DCB0D2A6C7AEAF32BE20B1" pageId="19" pageNumber="20">MNCN 15.05/60013H, holotype.</paragraph>
</subSubSection>
<subSubSection id="EA44F7D5A6ABC65F44E250AAD0902074" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="968A56B9B13C22F3685A2461CB6654B3" pageId="19" pageNumber="20">Additional material.</paragraph>
<paragraph id="B89921CE7FDCE0393FA7778B3894F006" pageId="19" pageNumber="20">MNCN 15.05/60013P (5), MNCN 15.05/7477P (3), paratypes.</paragraph>
</subSubSection>
<subSubSection id="A7E09B0A49CFD0C6BEAB738C80897E0E" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="7914243D2DD07070C2B39E771ED7BE25" pageId="19" pageNumber="20">Diagnosis.</paragraph>
<paragraph id="71D0BD7E59D4E19267EFD328A7F7C36A" pageId="19" pageNumber="20">See above.</paragraph>
</subSubSection>
<subSubSection id="DA373B3EED15F82BF975C18F0DD17B3B" pageId="19" pageNumber="20" type="description">
<paragraph id="B59364FFAD5B93E924C948115C0FFB0B" pageId="19" pageNumber="20">Dimensions.</paragraph>
<paragraph id="2AFC6DAAE892CA0F59E33718652E36A0" pageId="19" pageNumber="20">Shell height 44.8, diameter 25.3 mm.</paragraph>
</subSubSection>
<subSubSection id="A70A76921B98A04980BF4FC41952FFB4" pageId="19" pageNumber="20" type="distribution">
<paragraph id="FF7D0E14375C595E6D6D8EBB7FE66FF9" pageId="19" pageNumber="20">Distribution.</paragraph>
<paragraph id="21F7DADB9D3594C3BBA614258CF31E86" pageId="19" pageNumber="20">Ecuador, without precise locality.</paragraph>
</subSubSection>
<subSubSection id="16AFE4CE2B87C21DFAEB39C84B4AC19F" pageId="19" pageNumber="20" type="remarks">
<paragraph id="602C2C9AF3A4A998541AA84214671DE7" pageId="19" pageNumber="20">Remarks.</paragraph>
<paragraph id="010B82EA71F80A3111957C77CF1B63FB" pageId="19" pageNumber="20">
This species was described on the basis of material collected by the
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<normalizedToken id="0FFDEF09DB5AEC6FD8903C8B85998BB5" originalValue="Científica">Cientifica</normalizedToken>
del
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with an imprecise locality. While more precise records are awaited, it is suggested that the eastern Cordillera could be a possible location where this species might occur.
</paragraph>
</subSubSection>
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<taxonomicName id="59FFFD74EFED539E91BC6C54CDCE58EB" ID-CoL="QDMB" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:228178" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" lsidName="Camponotus (Myrmosericus) rufoglaucus st. cinctellus" lsidName-HNS="Camponotus (Myrmosericus) rufoglaucus st. cinctellus" order="Hymenoptera" pageNumber="133" phylum="Arthropoda" rank="subSpecies" species="rufoglaucus" subGenus="Myrmosericus" subSpecies="cinctellus">C. (Myrmosericus) rufoglaucus, st. cinctellus Gerstaecker</taxonomicName>
.
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<paragraph id="60BCC604340F0DD076CB9E6D4D9BAE14" pageNumber="133">
Jordan, Madras. Journ. of Litt. Sc, vol. 17, p. 124 (1851), [[worker]] (
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); - Gerstaecker, Monatsb. Ak. Wiss. Berlin,
</paragraph>
<paragraph id="43E0567EA6A3E1F4A1A5341E0E291334" pageNumber="133">
p. 262 (1858), [[male]], [[queen]], [[worker]] (
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).
</paragraph>
</subSubSection>
<subSubSection id="7F9DF6B57FDA5711A83BA637E14D47A6" type="materials_examined">
<paragraph id="B181BA8EB742F40B6E86B291ADAD543B" pageNumber="133">Afrique orientale anglaise: region cotiere: Gazi (st. n° 6, nov. 1911), 1 [[worker]]; - Likoni (st. n° 4, nov. 1911), 5 [[worker]]; - Tiwi (st. n° 5, nov. 1911), 1 [[worker]]; - Mombasa (1909), 1 [[worker]]; - Bura, dans les Wa-Taita (1904), 2 [[worker]]; - Fort-Hall, dans lc pays Kikuyu (alt. 1.330 m., st. n° 30, janv. 1912), 2 [[worker]].</paragraph>
<paragraph id="7724727059A66882DE371C99C65788ED" pageNumber="133">Afrique orientale allemande:.Tanga (1909), [[worker]].</paragraph>
<paragraph id="EF7C26D2E22BED73FC726EEBB64A8E74" pageNumber="133">Ile de Zanzibar: riviere Mwera (1903), 2 [[worker]].</paragraph>
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<taxonomicName LSID="http://zoobank.org/DD5650C8-3A93-4088-88A2-300A07CE2557" class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes shakirae" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="shakirae">Aleiodes shakirae</taxonomicName>
<taxonomicNameLabel pageId="59" pageNumber="60">sp. n.</taxonomicNameLabel>
Figures 96-100, 120
</paragraph>
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<subSubSection lastPageId="61" lastPageNumber="62" pageId="59" pageNumber="60" type="description of holotype">
<paragraph pageId="59" pageNumber="60">Description of holotype.</paragraph>
<paragraph pageId="59" pageNumber="60">Female (holotype). Body length 6.0 mm; antenna length 7.4 mm; fore wing length 5.7 mm.</paragraph>
<paragraph pageId="59" pageNumber="60">
Color. Head yellow, ocellar triangle dark brown; antenna dark
<normalizedToken originalValue="brownblack">brown-black</normalizedToken>
, scapus lighter dorsally; mesosoma yellow, anterior corner of mesopleuron, metanotum and part of lunules, propodeum and metapleuron dark
<normalizedToken originalValue="brownblack">brown-black</normalizedToken>
, mesopleuron light
<normalizedToken originalValue="yellowwhitish">yellow-whitish</normalizedToken>
at broader with metapeluron but blackish in immediate border; fore and mid legs whitish at coxa, trochanter and trochantellus, darkening toward apex from light yellow to honey brown; hind leg: 1/2 basal coxa white, 1/2 apical black, trochanter white, trochantellus black with ventral white stripe, femur and tibia whitish with infuscate stains mid laterally and at apex, tibial spurs and telotarsi honey brown, remaining tarsi brown. Metasomal terga black, but T4-T7 white laterally; metasoma ventrally white; ovipositor sheaths dark brown on apical half, basal half whitish. Wings hyaline; stigma brown with central area lighter; most veins dark brown; vein C+SC+R black (extreme base whitish) connecting to a black parastigma with whitish central spot.
</paragraph>
<paragraph pageId="59" pageNumber="60">
Head. Antenna with 47 antennomeres, flagellomeres roughly 2.0
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, apical flagellomere with long
<normalizedToken originalValue="“bottle-nipple”">&quot;bottle-nipple&quot;</normalizedToken>
-shaped apex; malar space as long as basal width of mandible, and approximately 1/4 eye height; in dorsal view eyes 3.4
<normalizedToken originalValue="×">x</normalizedToken>
temples; occipital carina incomplete dorsally, curving toward lateral ocelli, laterally complete and meeting hypostomal carina; oral space small and circular, maximum width slightly equal to basal width of mandible; clypeus not swollen; ocellus moderate,
<normalizedToken originalValue="ocellocular">ocell-ocular</normalizedToken>
distance short, about 0.45
<normalizedToken originalValue="×">x</normalizedToken>
diameter of lateral ocellus; maxillary palp not swollen; head surface sculpturing finely granulate, occiput smooth and shining; higher face with some wrinkles just bellow toruli.
</paragraph>
<paragraph pageId="59" pageNumber="60">Mesosoma. Sculpturing finely granulate; propodeum with mid-longitudinal carina complete; notauli weak, shallow and smooth; posterior margin of mesoscutum with complete carina; scutellar sulcus with median carina plus two pairs of lateral carina.</paragraph>
<paragraph pageId="59" pageNumber="60">
Wings. Fore wing: stigma 4.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than high; vein r 1.27
<normalizedToken originalValue="×">x</normalizedToken>
vein 2RS, 1.2
<normalizedToken originalValue="×">x</normalizedToken>
vein RS+Mb, and as long as vein m-cu; vein 3RSa about 0.44
<normalizedToken originalValue="×">x</normalizedToken>
vein 3RSb, and 0.8
<normalizedToken originalValue="×">x</normalizedToken>
vein 2M; vein 1CUa 1.6
<normalizedToken originalValue="×">x</normalizedToken>
vein 1cu-a; vein 1CUb 2.2
<normalizedToken originalValue="×">x</normalizedToken>
vein 1CUa; vein 1M strongly curved at basal portion. Hind wing: m-cu virtually absent; vein M+CU 1.4
<normalizedToken originalValue="×">x</normalizedToken>
vein 1M; vein 1M 1.2
<normalizedToken originalValue="×">x</normalizedToken>
vein r-m; RS smoothly curved at middle; vein M dark brown, well pigmented; vein 2-1A absent.
</paragraph>
<paragraph pageId="60" pageNumber="61">
<pageBreakToken pageId="60" pageNumber="61" start="start">Legs</pageBreakToken>
. Hind tibia without comb of modified setae; tarsal claws pectinate basally, with several very short and tight bristles, longer and sparser apically, wide gap between apical bristles and claw; hind basitarsus 3
<normalizedToken originalValue="×">x</normalizedToken>
longer than inner apical spur on hind tibia.
</paragraph>
<paragraph pageId="60" pageNumber="61">
Metasoma. T1-T3
<normalizedToken originalValue="granularrugose">granular-rugose</normalizedToken>
; remainder terga granular; mid longitudinal carina complete from T1 throughout T3; ovipositor sheaths about as long as hind tarsomere II; T1 long and narrow, T1 length 1.7
<normalizedToken originalValue="×">x</normalizedToken>
its apical width.
</paragraph>
<paragraph pageId="60" pageNumber="61">Paratypes variation. Body length 4.5-6.2 mm; antenna with 47-50 segments; legs color somewhat darker in few paratypes, and/or mid trochantellus laterally infuscate; scutellar sulcus with 3 to 5 carina; other color and proportions with only minimal variation.</paragraph>
<paragraph pageId="60" pageNumber="61">
Male. Antenna with 44 or 46 segments; ocelli larger,
<normalizedToken originalValue="ocell">ocell-</normalizedToken>
cular distance 1/3 diameter of lateral ocellus; tergite 1 narrower, about 2
<normalizedToken originalValue="×">x</normalizedToken>
longer than apical width.
</paragraph>
<paragraph lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
Mummy. Length 14.0-18.7 mm, dark reddish brown medially, light brown anteriorly and posteriorly, thorax narrow and wrinkled, mummy withered posteriorly be
<pageBreakToken pageId="61" pageNumber="62" start="start">hind</pageBreakToken>
exit hole, general long and narrow aspect, curved down- or sideward
<normalizedToken originalValue="“V-shaped”">&quot;V-shaped&quot;</normalizedToken>
to almost straight, exit hole irregular, located postero-dorsally, anterior to prolegs, but one specimen with ventral exit hole.
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="62" type="type material">
<paragraph pageId="61" pageNumber="62">Type material.</paragraph>
<paragraph pageId="61" pageNumber="62">
Type-locality: ECUADOR, Napo Province, Yanayacu Biological Station, canopy Malaise trap on bamboo,
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-0.59833336">S00°35.9'</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.89">W77°53.4'</geoCoordinate>
, 2163 m, cloud forest, July 10-16, 2010, S.R: Shaw col.
</paragraph>
<paragraph pageId="61" pageNumber="62">
Type-specimen: Holotype female, point mounted. Top label: &quot;ECUADOR: Napo Province / Yanayacu Biological Station /
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-0.59833336">S00°35.9'</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.89">W77°53.4'</geoCoordinate>
2163m / 10-16 June 2010, Scott R. Shaw / canopy Malaise trap, bamboo / NSF-BSI-07-17458 expedition&quot;. (UWIM)
</paragraph>
<paragraph pageId="61" pageNumber="62">
Paratypes 5 females and 6 males (UWIM), same data as holotype, except: 1♀, June 1-8, 2007, Malaise trap, SRS-00047; 3♀ and 6 males, reared from
<taxonomicName family="Geometridae" lsidName="" pageId="61" pageNumber="62" rank="family">Geometridae</taxonomicName>
larvae on
<taxonomicName class="Liliopsida" family="Poaceae" genus="Chusquea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chusquea scandens" order="Poales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="scandens">Chusquea scandens</taxonomicName>
(
<taxonomicName genus="Poaceae" lsidName="Poaceae" pageId="61" pageNumber="62" rank="genus">Poaceae</taxonomicName>
): 1♂ June 2, 2006, YY-14220; 1♂ March 25, 2008, YY-29593; 1♂ August 26, 2009, YY-40478; 1♀ January 30, 2010, YY-44297; 1♂ July 29, 2010, YY-48878; 1♂ September 9, 2010, YY-50943; 1♂ November 16, 2010, YY-52497; 1♀, January 7, 2011, YY-52852; 1♀ March 10, 2011, YY-54224; 1♀ September 3, 2013, YY-78769.
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="62" type="biology">
<paragraph pageId="61" pageNumber="62">Biology.</paragraph>
<paragraph pageId="61" pageNumber="62">
Reared from
<taxonomicName family="Geometridae" lsidName="" pageId="61" pageNumber="62" rank="family">Geometridae</taxonomicName>
caterpillar feeding on
<taxonomicName class="Liliopsida" family="Poaceae" genus="Chusquea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chusquea scandens" order="Poales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="scandens">Chusquea scandens</taxonomicName>
(
<taxonomicName genus="Poaceae" lsidName="Poaceae" pageId="61" pageNumber="62" rank="genus">Poaceae</taxonomicName>
). Most host caterpillars were commonly named &quot;palito
<normalizedToken originalValue="café">cafe</normalizedToken>
chusquea.&quot; Considering associated caterpillar pictures, rearing information, and similar mummy morphology, it is likely that all the hosts were conspecific. Host caterpillars were collected in 2nd and 3rd instars. Time span, from pupation until adult emergence, varied from 2.5 up to 6 weeks. As the mummy dries, the middle section bends and dries in different ways, resulting in different but distinctive bent mummies.
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="62" type="discussion">
<paragraph pageId="61" pageNumber="62">Discussion.</paragraph>
<paragraph pageId="61" pageNumber="62">
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes shakirae" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="shakirae">Aleiodes shakirae</taxonomicName>
sp. n. belongs to circumscriptus/gastritor species group. The
<normalizedToken originalValue="“V-shaped”">&quot;V-shaped&quot;</normalizedToken>
mummies from
<taxonomicName family="Geometridae" lsidName="" pageId="61" pageNumber="62" rank="family">Geometridae</taxonomicName>
and some color features resemble
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes townsendi" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="townsendi">Aleiodes townsendi</taxonomicName>
sp. n.; however,
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes shakirae" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="shakirae">Aleiodes shakirae</taxonomicName>
sp. n. differs from
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes townsendi" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="townsendi">Aleiodes townsendi</taxonomicName>
sp. n. in the
<normalizedToken originalValue="ocellocular">ocell-ocular</normalizedToken>
distance, which is about half the diameter of the lateral ocellus, but is nearly the same length as the lateral ocellus in
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes townsendi" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="townsendi">Aleiodes townsendi</taxonomicName>
sp. n. In
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes shakirae" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="shakirae">Aleiodes shakirae</taxonomicName>
sp. n. the metapeluron and metasoma are dorsally black (in
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes townsendi" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="townsendi">Aleiodes townsendi</taxonomicName>
sp. n. the metapleuron and T1 are mostly white with black markings), the hind coxa basally white and apically black (colors inverted in
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes townsendi" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="townsendi">Aleiodes townsendi</taxonomicName>
sp. n.), and hind tibia and tarsi are lighter in
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes shakirae" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="shakirae">Aleiodes shakirae</taxonomicName>
sp. n., the hind tibial spurs are yellowish to honey yellow (as compared with black in
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes townsendi" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="townsendi">Aleiodes townsendi</taxonomicName>
). The sculpturing of metasomal terga 1-3 in
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes shakirae" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="shakirae">Aleiodes shakirae</taxonomicName>
sp. n. is
<normalizedToken originalValue="granularrugose">granular-rugose</normalizedToken>
(
<normalizedToken originalValue="rugosestriate">rugose-striate</normalizedToken>
in
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes townsendi" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="townsendi">Aleiodes townsendi</taxonomicName>
sp. n.), the metasoma is much slender in
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes shakirae" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="shakirae">Aleiodes shakirae</taxonomicName>
sp. n., petiole is 1.7-2.0 times longer than apical width (as compared with 1.2 times in
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes townsendi" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="townsendi">Aleiodes townsendi</taxonomicName>
sp. n.), and the hind wing m-cu is absent in
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes shakirae" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="shakirae">Aleiodes shakirae</taxonomicName>
sp. n. (present as a short pigmented stub in
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes townsendi" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="townsendi">Aleiodes townsendi</taxonomicName>
sp. n.). The host species &quot;palito
<normalizedToken originalValue="café">cafe</normalizedToken>
chusquea&quot; (
<taxonomicName family="Geometridae" lsidName="" pageId="61" pageNumber="62" rank="family">Geometridae</taxonomicName>
) is the same species attacked by
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes nubicola" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="nubicola">Aleiodes nubicola</taxonomicName>
sp. n. and
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes mirandae" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="mirandae">Aleiodes mirandae</taxonomicName>
sp. n.
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="62" type="etymology">
<paragraph pageId="61" pageNumber="62">Etymology.</paragraph>
<paragraph pageId="61" pageNumber="62">This species is named after the famous Colombian singer Shakira. Since parasitism by this species causes the host caterpillar to bend and twist its abdomen in various ways, and Shakira is also famous for her belly-dancing, the name seems particulary appropriate for this species.</paragraph>
<caption pageId="61" pageNumber="62">
<paragraph pageId="61" pageNumber="62">
Figures 96-100.
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes shakirae" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="shakirae">Aleiodes shakirae</taxonomicName>
sp. n. 96 host larva (
<taxonomicName family="Geometridae" lsidName="" pageId="61" pageNumber="62" rank="family">Geometridae</taxonomicName>
) 97 host mummy after parasitoid emergence 98 habitus 99 head, dorsal 100 metasoma, dorsal. Figures 101-103.
<taxonomicName class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes speciosus" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="speciosus">Aleiodes speciosus</taxonomicName>
Townsend. 101 head frontal 102 head, occiput and vertex 103 metasoma, dorsal.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<mods:namePart>Noyes, John S.</mods:namePart>
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<mods:role>
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</mods:role>
<mods:namePart>Livermore, Laurence</mods:namePart>
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<mods:role>
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</mods:role>
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<mods:date>2016</mods:date>
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<mods:number>4</mods:number>
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<taxonomicName authority="Erdoes, 1951" authorityName="Erdoes" authorityYear="1951" class="Insecta" family="Eulophidae" genus="Diaulinopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaulinopsis arenaria" order="Hymenoptera" pageId="0" pageNumber="8013" phylum="Arthropoda" rank="species" species="arenaria">
Diaulinopsis arenaria (
<normalizedToken originalValue="Erdös">Erdoes</normalizedToken>
, 1951)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8013" type="reference_group">
<paragraph pageId="0" pageNumber="8013">
<taxonomicName class="Insecta" family="Eulophidae" genus="Cycloscapus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cycloscapus arenarius" order="Hymenoptera" pageId="0" pageNumber="8013" phylum="Arthropoda" rank="species" species="arenarius">Cycloscapus arenarius</taxonomicName>
<normalizedToken originalValue="Erdös">Erdoes</normalizedToken>
, 1951
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8013" type="distribution">
<paragraph pageId="0" pageNumber="8013">Distribution</paragraph>
<paragraph pageId="0" pageNumber="8013">England</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8013" type="notes">
<paragraph pageId="0" pageNumber="8013">Notes</paragraph>
<paragraph pageId="0" pageNumber="8013">BMNH, det. Hansson, added here</paragraph>
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.91.709" ID-GBIF-Dataset="78741a70-2be5-4536-a819-3a51f9d89cde" ID-PMC="PMC3084493" ID-Pensoft-Pub="1313-2970-91-1" ID-PubMed="21594108" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-91-1" ModsDocOrigin="ZooKeys 91" ModsDocTitle="Taxonomic revision and phylogenetic analysis of the flightless Mancallinae (Aves, Pan-Alcidae)" checkinTime="1451250487905" checkinUser="pensoft" docAuthor="Smith, Neil Adam" docDate="2011" docId="A008D2293B5663E014871A788AADF201" docLanguage="en" docName="ZooKeys 91: 1-116" docOrigin="ZooKeys 91" docSource="http://dx.doi.org/10.3897/zookeys.91.709" docTitle="Mancalla lucasi Smith, 2011, sp. n." docType="treatment" docVersion="5" lastPageNumber="11" masterDocId="FF8FFFC9A904277FD17A5E6B442A302C" masterDocTitle="Taxonomic revision and phylogenetic analysis of the flightless Mancallinae (Aves, Pan-Alcidae)" masterLastPageNumber="116" masterPageNumber="1" pageNumber="8" updateTime="1668168425552" updateUser="ExternalLinkService">
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<mods:title>Taxonomic revision and phylogenetic analysis of the flightless Mancallinae (Aves, Pan-Alcidae)</mods:title>
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<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>91</mods:number>
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<mods:identifier type="Pensoft-Pub">1313-2970-91-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="159364928" LSID="urn:lsid:zoobank.org:act:31389B4B-0A03-48E5-8A0B-C71CDBCE7164" httpUri="http://treatment.plazi.org/id/A008D2293B5663E014871A788AADF201" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8">
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<paragraph pageId="7" pageNumber="8">
<taxonomicName LSID="urn:lsid:zoobank.org:act:31389B4B-0A03-48E5-8A0B-C71CDBCE7164" class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla lucasi" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="lucasi">Mancalla lucasi</taxonomicName>
<taxonomicNameLabel pageId="7" pageNumber="8">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="holotype">
<paragraph pageId="7" pageNumber="8">Holotype.</paragraph>
<paragraph pageId="7" pageNumber="8">SDSNH 25237: a partial postcranial skeleton comprising the following elements: right and left scapulae, partial sternum, right and left humeri, left femur (Fig. 3; Tables 1, 2 and 3). The holotype specimen was collected by H. M. Wagner in April, 1980.</paragraph>
<caption pageId="7" pageNumber="8">
<paragraph pageId="7" pageNumber="8">
Figure 3. Holotype specimen of
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla lucasi" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="lucasi">Mancalla lucasi</taxonomicName>
(SDSNH 25237). A Fragment of anterior sternum in anterior view B Carinal apex of sternum in right lateral view C Right humerus in anterior view D Left humerus in proximal view E Left humerus in distal view F Left humerus in posterior view G Left scapula in lateral view H Right scapula in medial view I Left femur in anterior view. Anatomical abbreviations: a acromion process bc bicipital crest c caput ca carinal apex ce caudal extremity of scapula cg capital groove cs coracoidal sulcus ct coracoidal tubercle d deltopectoral crest dc dorsal condyle dsp dorsal supracondylar dst dorsal supracondylar tubercle fh femoral head fp flexor process gp glenoid process hs humerotricipital sulcus le lateral epicondyle pf1 primary pneumotricipital fossa ps pectoralis scar sc supracoracoidal crest si sulcus intercondylaris sr sternal rostrum ss scapulotricipital sulcus st scapulotricipital tubercle tc trochanteric crest tls transverse ligament sulcus vc ventral condyle vst ventral supracondylar tubercle vt ventral tubercle.
</paragraph>
</caption>
<caption pageId="7" pageNumber="8">
<paragraph pageId="7" pageNumber="8">
Table 2. Measurements of
<taxonomicName lsidName="" pageId="7" pageNumber="8" rank="subFamily" subFamily="Mancallinae">Mancallinae</taxonomicName>
holotype humeri (mm). Abbreviations: (Glh) greatest length of humerus; (Bph) breadth of proximal humerus; (Diph) diagonal of proximal humerus; (Whs) width of humeral shaft; (Bdh) breadth of distal humerus; (Ddh) depth of distal humerus. Measurements according to
<bibRefCitation author="Von den Driesch, A" journalOrPublisher="Peabody Museum Bulletin" pageId="42" pageNumber="43" pagination="103 - 129" title="A guide to the measurement of animal bones from archaeological sites." volume="1" year="1976">Von den Driesch (1976)</bibRefCitation>
.
<normalizedToken originalValue="~">'~'</normalizedToken>
signifies approximate measurement due to damage.
<normalizedToken originalValue="-">'-'</normalizedToken>
signifies missing data due to damage.
</paragraph>
</caption>
<paragraph pageId="7" pageNumber="8">
<table pageId="7" pageNumber="8">
<tr pageId="7" pageNumber="8">
<th colspan="1" pageId="7" pageNumber="8" rowspan="1">Species</th>
<th colspan="1" pageId="7" pageNumber="8" rowspan="1">Specimen #</th>
<th colspan="1" pageId="7" pageNumber="8" rowspan="1">Glh</th>
<th colspan="1" pageId="7" pageNumber="8" rowspan="1">Bph</th>
<th colspan="1" pageId="7" pageNumber="8" rowspan="1">Diph</th>
<th colspan="1" pageId="7" pageNumber="8" rowspan="1">Whs</th>
<th colspan="1" pageId="7" pageNumber="8" rowspan="1">Bdh</th>
<th colspan="1" pageId="7" pageNumber="8" rowspan="1">Ddh</th>
</tr>
<tr pageId="7" pageNumber="8">
<td colspan="1" pageId="7" pageNumber="8" rowspan="1">
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla californiensis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="californiensis">Mancalla californiensis</taxonomicName>
</td>
</tr>
<tr pageId="7" pageNumber="8">
<td colspan="1" pageId="7" pageNumber="8" rowspan="1">
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla cedrosensis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cedrosensis">Mancalla cedrosensis</taxonomicName>
</td>
</tr>
<tr pageId="7" pageNumber="8">
<td colspan="1" pageId="7" pageNumber="8" rowspan="1">
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla lucasi" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="lucasi">Mancalla lucasi</taxonomicName>
</td>
</tr>
<tr pageId="7" pageNumber="8">
<td colspan="1" pageId="7" pageNumber="8" rowspan="1">
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
</td>
</tr>
<tr pageId="7" pageNumber="8">
<td colspan="1" pageId="7" pageNumber="8" rowspan="1">
<taxonomicName class="Aves" family="Alcidae" genus="Miomancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Miomancalla wetmorei" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="wetmorei">Miomancalla wetmorei</taxonomicName>
</td>
</tr>
<tr pageId="7" pageNumber="8">
<td colspan="1" pageId="7" pageNumber="8" rowspan="1">
<taxonomicName class="Aves" family="Alcidae" genus="Miomancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Miomancalla howardi" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="howardi">Miomancalla howardi</taxonomicName>
</td>
</tr>
<tr pageId="7" pageNumber="8">
<td colspan="1" pageId="7" pageNumber="8" rowspan="1">
<taxonomicName class="Aves" family="Alcidae" genus="Miomancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Miomancalla howardi" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="howardi">Miomancalla howardi</taxonomicName>
</td>
</tr>
</table>
</paragraph>
<caption ID-Table-UUID="DB9751960DA2F222F3FCDBE0C2204D19" httpUri="http://table.plazi.org/id/DB9751960DA2F222F3FCDBE0C2204D19" pageId="7" pageNumber="8">
<paragraph pageId="7" pageNumber="8">
Table 3. Measurements of new associated
<taxonomicName lsidName="" pageId="7" pageNumber="8" rank="subFamily" subFamily="Mancallinae">Mancallinae</taxonomicName>
holotype specimens (in mm).
<normalizedToken originalValue="-">'-'</normalizedToken>
= missing data due to damage or lack of comparable element.
</paragraph>
</caption>
<paragraph pageId="7" pageNumber="8">
<table pageId="7" pageNumber="8">
<tr pageId="7" pageNumber="8">
<td colspan="1" pageId="7" pageNumber="8" rowspan="1">
<taxonomicName class="Aves" family="Alcidae" genus="Miomancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Miomancalla howardi" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="howardi">Miomancalla howardi</taxonomicName>
</td>
<td colspan="1" pageId="7" pageNumber="8" rowspan="1">
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla lucasi" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="lucasi">Mancalla lucasi</taxonomicName>
</td>
<td colspan="1" pageId="7" pageNumber="8" rowspan="1">
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="etymology">
<paragraph pageId="7" pageNumber="8">Etymology.</paragraph>
<paragraph pageId="7" pageNumber="8">
This new species is named in honor of Frederic A. Lucas who described the first known remains of
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Mancalla</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="locality and horizon">
<paragraph pageId="7" pageNumber="8">Locality and horizon.</paragraph>
<paragraph pageId="7" pageNumber="8">Late Pliocene or Early Pleistocene (Zanclean or Calabrian) Niguel Formation of Orange County, California. Latitude, longitude, and elevation data are on file at SDSNH (locality 3202). Details of the geologic setting are provided in Appendix 6.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="referred specimen">
<paragraph pageId="7" pageNumber="8">Referred specimen.</paragraph>
<paragraph pageId="7" pageNumber="8">SDSNH 59049: a complete left humerus from the Middle Pliocene to Early Pleistocene San Diego Formation (SDSNH locality 3506; Fig. 4E).</paragraph>
<caption pageId="7" pageNumber="8">
<paragraph pageId="7" pageNumber="8">
Figure 4.
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Mancalla</taxonomicName>
referred humeri in anterior view. A
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
SDSNH 28152 B
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
SDSNH 42534 C
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
SDSNH 75051 D
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
SDSNH 42532 E
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla lucasi" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="lucasi">Mancalla lucasi</taxonomicName>
SDSNH 59049.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="differential diagnosis">
<paragraph pageId="7" pageNumber="8">Differential diagnosis.</paragraph>
<paragraph pageId="7" pageNumber="8">
Scar extending into primary pneumotricipital fossa is raised in relief to the floor of the primary pneumotricipital fossa and the humeral shaft as in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla cedrosensis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cedrosensis">Mancalla cedrosensis</taxonomicName>
,rather than an excavated pit as in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
sp. n. and
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla californiensis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="californiensis">Mancalla californiensis</taxonomicName>
<bibRefCitation author="Lucas, FA" journalOrPublisher="Proceedings of the United States National Museum" pageId="38" pageNumber="39" pagination="113 - 134" title="A Flightless auk, Mancalla californiensis, from the Miocene of California." volume="24" year="1901">Lucas 1901</bibRefCitation>
(121:1; Fig. 5); dorsal and ventral edges of scar extending into primary pneumotricipital fossa taper to a point as in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
, rather than remaining parallel as in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla californiensis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="californiensis">Mancalla californiensis</taxonomicName>
and
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla cedrosensis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cedrosensis">Mancalla cedrosensis</taxonomicName>
(123:1); humerus longer than
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla cedrosensis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cedrosensis">Mancalla cedrosensis</taxonomicName>
,
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla californiensis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="californiensis">Mancalla californiensis</taxonomicName>
,and
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
(Tables 2, 3).
</paragraph>
<caption pageId="7" pageNumber="8">
<paragraph pageId="7" pageNumber="8">
Figure 5. Line drawings for comparison of
<taxonomicName lsidName="" pageId="7" pageNumber="8" rank="subFamily" subFamily="Mancallinae">Mancallinae</taxonomicName>
proximal humeri in posterior view (not to scale). A
<taxonomicName class="Aves" family="Alcidae" genus="Miomancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Miomancalla wetmorei" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="wetmorei">Miomancalla wetmorei</taxonomicName>
B
<taxonomicName class="Aves" family="Alcidae" genus="Miomancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Miomancalla howardi" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="howardi">Miomancalla howardi</taxonomicName>
C
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla californiensis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="californiensis">Mancalla californiensis</taxonomicName>
D
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla cedrosensis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cedrosensis">Mancalla cedrosensis</taxonomicName>
E
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
F
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla lucasi" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="lucasi">Mancalla lucasi</taxonomicName>
. Anatomical abbreviations: ms mancalline scar pf1 primary pneumotricipital fossa sc supracoracoidal crest vt ventral tubercle.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8" type="anatomical description">
<paragraph pageId="7" pageNumber="8">Anatomical description.</paragraph>
<paragraph lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Both scapulae are preserved (Fig. 3G, H). As in all
<taxonomicName family="Alcidae" lsidName="" pageId="7" pageNumber="8" rank="family">Alcidae</taxonomicName>
, the scapular shaft is mediolaterally compressed throughout its entire length. The proximal end of the scapular shaft is more rounded in other Charadriiformes. As in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
, the acromion projects farther anteriorly than that of
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla cedrosensis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cedrosensis">Mancalla cedrosensis</taxonomicName>
and other alcids (e.g.,
<taxonomicName class="Aves" family="Alcidae" genus="Uria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uria" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Uria</taxonomicName>
,
<taxonomicName class="Aves" family="Alcidae" genus="Aethia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aethia" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Aethia</taxonomicName>
). As in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla cedrosensis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cedrosensis">Mancalla cedrosensis</taxonomicName>
, the coracoidal tubercle is less pronounced than in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
. As in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
and
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla cedrosensis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cedrosensis">Mancalla cedrosensis</taxonomicName>
, a scapulotricipital tubercle is present just distal to the glenoid process on the ventral margin of the scapular shaft. This feature is also present in other flightless wing-propelled divers such as
<taxonomicName family="Spheniscidae" lsidName="" pageId="7" pageNumber="8" rank="family">Spheniscidae</taxonomicName>
and
<taxonomicName class="Aves" family="Alcidae" genus="Pinguinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinguinus" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Pinguinus</taxonomicName>
, but is not known in any volant alcid. As in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
, the scapular shaft, including the caudal extremity, is slightly more robust than in other alcids (e.g.,
<taxonomicName class="Aves" family="Alcidae" genus="Alca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alca" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Alca</taxonomicName>
,
<taxonomicName class="Aves" family="Alcidae" genus="Aethia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aethia" order="Charadriiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Aethia</taxonomicName>
). The caudal
<pageBreakToken pageId="8" pageNumber="9" start="start">extremity</pageBreakToken>
is less dorsoventrally expanded than in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
. The caudal extremity is not known for
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla cedrosensis" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cedrosensis">Mancalla cedrosensis</taxonomicName>
.
</paragraph>
<paragraph pageId="8" pageNumber="9">
Fragments of the sternum preserve the sternal rostrum, coracoidal sulci, and the carinal apex (Fig. 3A, B). These features are not preserved in
<taxonomicName class="Aves" family="Alcidae" genus="Miomancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Miomancalla howardi" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="howardi">Miomancalla howardi</taxonomicName>
and comparisons are therefore limited to extant alcids and specimens of
<taxonomicName lsidName="" pageId="8" pageNumber="9" rank="subFamily" subFamily="Mancallinae">Mancallinae</taxonomicName>
that are not presently referable to species. The morphology of the sternal rostrum is consistent with that of all other
<taxonomicName family="Alcidae" lsidName="" pageId="8" pageNumber="9" rank="family">Alcidae</taxonomicName>
. Although no coracoid is preserved in the holotype specimen of
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla lucasi" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="lucasi">Mancalla lucasi</taxonomicName>
, the shape of the coracoidal sulci of the sternum is consistent with the ~150° angle of the sternal articulation of the coracoid in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla cedrosensis" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cedrosensis">Mancalla cedrosensis</taxonomicName>
and
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
. The sternal articulation of the coracoid, and the coracoidal sulci of the sternum in other alcids curves more acutely (e.g., ~90° in
<taxonomicName class="Aves" family="Alcidae" genus="Alca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alca torda" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="torda">Alca torda</taxonomicName>
; Fig. 6).
</paragraph>
<caption pageId="8" pageNumber="9">
<paragraph pageId="8" pageNumber="9">
Figure 6. Comparison of sternal facet curvature in charadriiform left coracoids (sternal view; not to scale). A
<taxonomicName class="Aves" family="Stercorariidae" genus="Stercorarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stercorarius" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">Stercorarius</taxonomicName>
B
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">Mancalla</taxonomicName>
C
<taxonomicName class="Aves" family="Alcidae" genus="Alca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alca" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">Alca</taxonomicName>
D
<taxonomicName class="Aves" family="Alcidae" genus="Aethia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aethia" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">Aethia</taxonomicName>
.
</paragraph>
</caption>
<paragraph lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
Complete right and left humeri are preserved (Fig. 3C, D, E and F). Based upon humeral proportions,
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla lucasi" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="lucasi">Mancalla lucasi</taxonomicName>
represents the largest known species of
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">Mancalla</taxonomicName>
(Table 2). As in other
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">Mancalla</taxonomicName>
species, the ventral margin of the ventral tubercle is convex, and the capital groove is relatively narrower than other
<taxonomicName family="Alcidae" lsidName="" pageId="8" pageNumber="9" rank="family">Alcidae</taxonomicName>
. The ventral tubercle does not project as far ventrally as in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla californiensis" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="californiensis">Mancalla californiensis</taxonomicName>
(Fig. 5). The distal end of the deltopectoral crest transitions to the shaft more abruptly than in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
.
<pageBreakToken pageId="9" pageNumber="10" start="start">As</pageBreakToken>
in other
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla" order="Charadriiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">Mancalla</taxonomicName>
, the humeral head is rotated anteriorly, and the supracoracoideus muscle scar does not broaden proximally.
<taxonomicName lsidName="" pageId="9" pageNumber="10" rank="subFamily" subFamily="Mancallinae">Mancallinae</taxonomicName>
is characterized by a scar of unknown function that is positioned adjacent to the primary pneumotricipital fossa (hereafter referred to as the 'mancalline
<normalizedToken originalValue="scar">scar'</normalizedToken>
; Fig. 5). The position of the 'mancalline
<normalizedToken originalValue="scar">scar'</normalizedToken>
suggests an accessory insertion of m. humerotriceps (
<bibRefCitation author="Howard, H" journalOrPublisher="Carnegie Institute of Washington Publications, Contributions to Paleontology" pageId="35" pageNumber="36" pagination="177 - 199" title="New avian records for the Pliocene of California." volume="584" year="1949">Howard 1949</bibRefCitation>
), which can be divided into as many as four separate heads in some birds (
<bibRefCitation author="Baumel, J" editor="Baumel, JJ" journalOrPublisher="Nuttall Ornithology Club" pageId="32" pageNumber="33" pagination="45 - 132" title="Osteologica" volumeTitle="Handbook of avian anatomy: Nomina Anatomica Avium, 2 nd edition" year="1993">Baumel and Witmer 1993</bibRefCitation>
). Other potentially homologous muscle scars include m. coracobrachialis cranialis, which is well developed in penguins (
<bibRefCitation author="Ksepka, DT" journalOrPublisher="Journal of Anatomy" pageId="37" pageNumber="38" pagination="131 - 147" title="Osteology of Icadyptes salasi, a giant penguin from the Eocene of Peru." volume="213" year="2008">Ksepka et al. 2008</bibRefCitation>
), or m. scapulocranialis caudalis (see
<bibRefCitation author="Matsuoka, H" journalOrPublisher="Bulletin of the Gunma Museum of Natural History" pageId="38" pageNumber="39" pagination="7 - 14" title="Myology and osteology of the Whooper Swan Cygnus Cygnus (Aves: Anatidae) Part I. Muscles attached to the sternum, coracoid, clavicle, scapula and humerus." volume="11" year="2007">Matsuoka and Hasegawa 2007</bibRefCitation>
). However, the exact function of this feature is unknown because it is not present in any other charadriiform. The shape, position, and development of this scar is variable in
<taxonomicName lsidName="" pageId="9" pageNumber="10" rank="subFamily" subFamily="Mancallinae">Mancallinae</taxonomicName>
(Fig. 5). The 'mancalline
<normalizedToken originalValue="scar">scar'</normalizedToken>
of
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla lucasi" order="Charadriiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="lucasi">Mancalla lucasi</taxonomicName>
is raised in relief like that of
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla cedrosensis" order="Charadriiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="cedrosensis">Mancalla cedrosensis</taxonomicName>
, rather than excavated as in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla californiensis" order="Charadriiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="californiensis">Mancalla californiensis</taxonomicName>
and
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
(Fig. 5). As in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla vegrandis" order="Charadriiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="vegrandis">Mancalla vegrandis</taxonomicName>
, the scar extends from a point just proximal to the junction of the bicipital crest with the humeral shaft, tapers to a point, and extends into the primary pneumotricipital fossa (Fig. 5). The dorsal and ventral margins of the 'mancalline
<normalizedToken originalValue="scar">scar'</normalizedToken>
remain approximately parallel in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla californiensis" order="Charadriiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="californiensis">Mancalla californiensis</taxonomicName>
and
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla cedrosensis" order="Charadriiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="cedrosensis">Mancalla cedrosensis</taxonomicName>
(Fig. 5). As in all
<taxonomicName lsidName="" pageId="9" pageNumber="10" rank="subFamily" subFamily="Mancallinae">Mancallinae</taxonomicName>
, the humeral shaft is arced rather than sigmoidal or straight. As in other
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla" order="Charadriiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">Mancalla</taxonomicName>
, the dorsal supracondylar process is separated from the dorsal epicondyle by a small notch. A tubercle or papilla is present on the posterior side of the distal end of the humerus adjacent to the dorsal condyle (
<bibRefCitation author="Howard, H" journalOrPublisher="Smithsonian Contributions to Paleobiology" pageId="36" pageNumber="37" pagination="141 - 146" title="A new species of flightless auk from the Miocene of California (Alcidae: Mancallinae). In: Olson SL (Ed) Collected papers in avian paleontology honoring the 90 th birthday of Alexander Wetmore." volume="27" year="1976">Howard 1976</bibRefCitation>
). As with all
<taxonomicName lsidName="" pageId="9" pageNumber="10" rank="subFamily" subFamily="Mancallinae">Mancallinae</taxonomicName>
, the anterior surface of the ventral condyle is rounded, rather than flattened as in all other alcids. Rounded fossae are present at the proximal ends of
<pageBreakToken pageId="10" pageNumber="11" start="start">the</pageBreakToken>
humerotricipital and scapulotricipital grooves. The flexor process extends distal to the ventral condyle as in all
<taxonomicName lsidName="" pageId="10" pageNumber="11" rank="subFamily" subFamily="Mancallinae">Mancallinae</taxonomicName>
and
<taxonomicName class="Aves" family="Alcidae" genus="Pinguinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinguinus" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Pinguinus</taxonomicName>
.
</paragraph>
<paragraph pageId="10" pageNumber="11">
The left femur is preserved (Fig. 3I) and is smaller (~15%; Table 2)than in
<taxonomicName class="Aves" family="Alcidae" genus="Miomancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Miomancalla howardi" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="howardi">Miomancalla howardi</taxonomicName>
sp. n. (Table 3), and larger (~19%) than in
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla cedrosensis" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="cedrosensis">Mancalla cedrosensis</taxonomicName>
(Howard, 1971). Extant alcids do not display statistically significant degrees of sexual dimorphism in their size, plumage, or osteological morphology (
<bibRefCitation author="Storer, RW" journalOrPublisher="University of California Publications in Zoology" pageId="41" pageNumber="42" pagination="121 - 222" title="A comparison of variation, behavior and evolution in the sea bird genera Uria and Cepphus." volume="52" year="1952">Storer 1952</bibRefCitation>
;
<bibRefCitation author="Nettleship, DN" journalOrPublisher="Academic Press, London and Orlando" pageId="39" pageNumber="40" title="The Atlantic Alcidae: the evolution, distribution, and biology of the auks inhabiting the Atlantic Ocean and adjacent water areas" year="1985">Nettleship and Birkhead 1985</bibRefCitation>
;
<bibRefCitation author="Szekely, T" journalOrPublisher="Evolution" pageId="41" pageNumber="42" pagination="1404 - 1413" title="Sexual size dimorphism in shorebirds, gulls, and alcids: the influence of sexual and natural selection." volume="54" year="2000">Szekely et al. 2000</bibRefCitation>
). Thus, it can be reasonably assumed that extinct alcids were also not sexually dimorphic as the proposed sister taxon of all alcids, the
<taxonomicName family="Stercorariidae" lsidName="" pageId="10" pageNumber="11" rank="family">Stercorariidae</taxonomicName>
(
<bibRefCitation author="Ericson, PGP" journalOrPublisher="BMC Evolutionary Biology" pageId="34" pageNumber="35" title="Inter-familial relationships of the shorebirds (Aves: Charadriiformes) based on nuclear DNA sequence data" url="doi:10.1186/1471-2148-3-16" volume="3" year="2003">Ericson et al. 2003</bibRefCitation>
;
<bibRefCitation author="Thomas, G" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="41" pageNumber="42" pagination="516 - 526" title="Phylogeny of shorebirds, gulls and alcids (Aves: Charadrii) from the cytochrome-b gene: parsimony, Baysian inference, minimum evolution and quartet puzzling" url="doi:10.1016/S1055-7903(03)00222-7" volume="30" year="2004">Thomas et al. 2004</bibRefCitation>
;
<bibRefCitation author="Baker, AJ" journalOrPublisher="Biology Letters" pageId="31" pageNumber="32" pagination="205 - 209" title="Phylogenetic relationships and divergence times of Charadriiformes genera: multigene evidence for the Cretaceous origin of 14 clades of shorebirds" url="doi:10.1098/rsbl.2006.0606" volume="3" year="2007">Baker et al. 2007</bibRefCitation>
;
<bibRefCitation author="Pereira, SL" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="40" pageNumber="41" pagination="430 - 455" title="DNA evidence for a Paleocene origin of the Alcidae (Aves: Charadriiformes) in the Pacific and multiple dispersals across northern oceans" url="doi:10.1016/j.ympev.2007.11.020" volume="46" year="2008">Pereira and Baker 2008</bibRefCitation>
), as well as the closely related
<taxonomicName family="Laridae" lsidName="" pageId="10" pageNumber="11" rank="family">Laridae</taxonomicName>
are also not sexually dimorphic. This range of size between
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Mancalla</taxonomicName>
species is greater than the range of intraspecific variation documented for other alcids (~1-5%), including the flightless Great Auk (
<bibRefCitation author="Moen, SM" journalOrPublisher="Auk" pageId="39" pageNumber="40" pagination="755 - 763" title="Morphologic and genetic variation among breeding colonies of the Atlantic Puffin (Fratercula arctica)." volume="108" year="1991">Moen 1991</bibRefCitation>
;
<bibRefCitation author="Burness, GP" journalOrPublisher="Polar Biology" pageId="33" pageNumber="34" pagination="545 - 551" title="Oceanographic-related variation in the bone sizes of Great Auks" url="doi:10.1007/BF00237947" volume="11" year="1992">Burness and Montevecchi 1992</bibRefCitation>
). As in
<taxonomicName class="Aves" family="Alcidae" genus="Alle" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alle" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Alle</taxonomicName>
,
<taxonomicName class="Aves" family="Alcidae" genus="Cepphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cepphus" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Cepphus</taxonomicName>
<normalizedToken originalValue="Mörhing">Moerhing</normalizedToken>
, 1758,
<taxonomicName class="Aves" family="Alcidae" genus="Synthliboramphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Synthliboramphus" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Synthliboramphus</taxonomicName>
Brandt, 1837, and
<taxonomicName class="Aves" family="Alcidae" genus="Brachyramphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachyramphus" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Brachyramphus</taxonomicName>
Brandt, 1837, the femoral trochanter projects anteriorly in lateral view. The femoral trochanter in
<taxonomicName class="Aves" family="Alcidae" genus="Uria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uria" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Uria</taxonomicName>
,
<taxonomicName class="Aves" family="Alcidae" genus="Aethia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aethia" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Aethia</taxonomicName>
Merrem, 1788,
<taxonomicName class="Aves" family="Alcidae" genus="Alca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alca" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Alca</taxonomicName>
, and
<taxonomicName class="Aves" family="Alcidae" genus="Pinguinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinguinus" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Pinguinus</taxonomicName>
is not projected anteriorly (i.e., straight), and the trochanter is concave in lateral view in
<taxonomicName class="Aves" family="Alcidae" genus="Fratercula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fratercula" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Fratercula</taxonomicName>
Brisson, 1760 and
<taxonomicName class="Aves" family="Alcidae" genus="Cerorhinca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerorhinca" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Cerorhinca</taxonomicName>
Bonaparte, 1828. Femora of
<taxonomicName class="Aves" family="Alcidae" genus="Miocepphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Miocepphus" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Miocepphus</taxonomicName>
are not known. No diagnostic characteristics of the femur of
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla lucasi" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="lucasi">Mancalla lucasi</taxonomicName>
were identified.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="remarks">
<paragraph pageId="10" pageNumber="11">Remarks.</paragraph>
<paragraph pageId="10" pageNumber="11">
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla lucasi" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="lucasi">Mancalla lucasi</taxonomicName>
corresponds in size and some humeral characteristics with material previously referred to
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla diegensis" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="diegensis">Mancalla diegensis</taxonomicName>
. However,
<taxonomicName class="Aves" family="Alcidae" genus="Mancalla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mancalla diegensis" order="Charadriiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="diegensis">Mancalla diegensis</taxonomicName>
is considered
<taxonomicName family="Alcidae" lsidName="" pageId="10" pageNumber="11" rank="family">Alcidae</taxonomicName>
incertae sedis (see Appendix 1 for details of the taxonomic revision).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,149 @@
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<emphasis id="1AD58A60FFE0531CFF20FF60AB1933AE" bold="true" box="[151,522,221,247]" pageId="1" pageNumber="41">
<taxonomicName id="EFA12DF1FFE0531CFF20FF60A88A33AE" ID-CoL="62KV8" authority="Navas, 1921" authorityName="Navas" authorityYear="1921" box="[151,409,221,247]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="1AD58A60FFE0531CFF20FF60A9EF33AE" bold="true" box="[151,252,221,247]" italics="true" pageId="1" pageNumber="41">Capicua</emphasis>
<bibRefCitation id="4C302B83FFE0531CFEB4FF60A88A33AE" author="Navas" box="[259,409,221,247]" pageId="1" pageNumber="41" refString="Navas, L. (1921) Comunicaciones entomologicas. 4. Insectos exoticos nuevos, criticos o poco conocidos. Revista de la Real Academia de Ciencias Exactas Fisico-Quimicas y Naturales de Zaragoza, 6, 61 - 81." type="journal article" year="1921">Navás, 1921</bibRefCitation>
</taxonomicName>
stat. rev.
</emphasis>
</heading>
</paragraph>
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<typeStatus id="F71AE8D0FFE0531CFF20FE98A9DB3263" box="[151,200,293,314]" pageId="1" pageNumber="41">Type</typeStatus>
species
<taxonomicName id="EFA12DF1FFE0531CFEA8FE99AB783260" authority="Navas, 1921" authorityName="Navas" authorityYear="1921" box="[287,619,291,314]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="fulvicauda">
<emphasis id="1AD58A60FFE0531CFEA8FE99A8F53263" box="[287,486,292,314]" italics="true" pageId="1" pageNumber="41">Capicua fulvicauda</emphasis>
<bibRefCitation id="4C302B83FFE0531CFE5BFE9EAB783260" author="Navas" box="[492,619,291,313]" pageId="1" pageNumber="41" refString="Navas, L. (1921) Comunicaciones entomologicas. 4. Insectos exoticos nuevos, criticos o poco conocidos. Revista de la Real Academia de Ciencias Exactas Fisico-Quimicas y Naturales de Zaragoza, 6, 61 - 81." type="journal article" year="1921">Navás, 1921</bibRefCitation>
</taxonomicName>
by monotypy.
</paragraph>
<paragraph id="281E5672FFE0531CFF20FEDBAC713227" blockId="1.[151,1437,358,1102]" box="[151,1378,358,383]" pageId="1" pageNumber="41">
The genus
<taxonomicName id="EFA12DF1FFE0531CFEA5FEDAA8623227" box="[274,369,359,382]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="1AD58A60FFE0531CFEA5FEDAA8623227" box="[274,369,359,382]" italics="true" pageId="1" pageNumber="41">Capicua</emphasis>
</taxonomicName>
, synonymised with the genus
<taxonomicName id="EFA12DF1FFE0531CFD73FEDAAA013227" box="[708,786,359,382]" class="Insecta" family="Myrmeleontidae" genus="Geyria" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="1AD58A60FFE0531CFD73FEDAAA013227" box="[708,786,359,382]" italics="true" pageId="1" pageNumber="41">Geyria</emphasis>
</taxonomicName>
by
<bibRefCitation id="4C302B83FFE0531CFC8CFEDBAAC83227" author="Stange" box="[827,987,358,382]" pageId="1" pageNumber="41" refString="Stange, L. A. (2004) A systematic catalog, bibliography and classification of the world antlions (Insecta: Neuroptera: Myrmeleontidae). Memoirs of the American Entomological Institute, 74, 1 - 565." type="journal article" year="2004">Stange (2004)</bibRefCitation>
, is reinstated here as a valid taxon.
</paragraph>
</subSubSection>
<subSubSection id="60BB05F9FFE0531CFF70FE37AD093717" pageId="1" pageNumber="41" type="description">
<paragraph id="281E5672FFE0531CFF70FE37AA00310F" blockId="1.[151,1437,358,1102]" pageId="1" pageNumber="41">
<emphasis id="1AD58A60FFE0531CFF70FE37A86732FA" bold="true" box="[199,372,394,419]" pageId="1" pageNumber="41">Redescription.</emphasis>
<taxonomicName id="EFA12DF1FFE0531CFECCFE36A8C932FB" box="[379,474,395,418]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="1AD58A60FFE0531CFECCFE36A8C932FB" box="[379,474,395,418]" italics="true" pageId="1" pageNumber="41">Capicua</emphasis>
</taxonomicName>
exhibits synapomorphies that characterize the tribe
<taxonomicName id="EFA12DF1FFE0531CFBA8FE37ADAE32FB" box="[1055,1213,394,418]" class="Insecta" family="Myrmeleontidae" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="tribe" tribe="Nemoleontini">Nemoleontini</taxonomicName>
. In the forewing the posterior cubital vein CuP originates at the basal crossvein and the anal vein A2 runs close to A1 for short distance, then bends at sharp angle toward A3. In the hind wing the anterior cubital vein CuA is connected to the posterior fork of posterior median MP2 by crossveins and there is only one presectoral crossvein. Moreover, the tarsal claws are not toothed or angled and the ventral surface of the distal tarsomeres has stout black setae placing
<taxonomicName id="EFA12DF1FFE0531CFAA8FDA6AC6D316B" box="[1311,1406,539,562]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="1AD58A60FFE0531CFAA8FDA6AC6D316B" box="[1311,1406,539,562]" italics="true" pageId="1" pageNumber="41">Capicua</emphasis>
</taxonomicName>
in the sub-tribe Neuroleontina as defined by
<bibRefCitation id="4C302B83FFE0531CFDD9FD83AA1C310F" author="Stange" box="[622,783,574,598]" pageId="1" pageNumber="41" refString="Stange, L. A. (2004) A systematic catalog, bibliography and classification of the world antlions (Insecta: Neuroptera: Myrmeleontidae). Memoirs of the American Entomological Institute, 74, 1 - 565." type="journal article" year="2004">Stange (2004)</bibRefCitation>
.
</paragraph>
<paragraph id="281E5672FFE0531CFF70FDDFABBF302F" blockId="1.[151,1437,358,1102]" pageId="1" pageNumber="41">
Diagnosis of the genus
<taxonomicName id="EFA12DF1FFE0531CFE66FDDEAB233123" box="[465,560,611,634]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="1AD58A60FFE0531CFE66FDDEAB233123" box="[465,560,611,634]" italics="true" pageId="1" pageNumber="41">Capicua</emphasis>
</taxonomicName>
, including the two new species described below can be expressed as follows. Small species. Body coloration brown or black. Wings narrow, lacking pillula axillaris. Fore coxa and fore femora without a tuft of long white setae.
<collectingCountry id="50B616E2FFE0531CFD94FD17AB48319B" box="[547,603,682,706]" name="India" pageId="1" pageNumber="41">Hind</collectingCountry>
femora of male with a tuft of long erect setae on dorsal surface (
<figureCitation id="B09A4AF7FFE0531CFAFFFD17AC9D319B" box="[1352,1422,682,706]" captionStart="FIGURES 4 14" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[224,1358,193,1850]" captionTargetId="figure@3.[224,1358,193,1861]" captionTargetPageId="3" captionText="FIGURES 4 14. Capicua nigra. Fig. 4. Holotype with labels; Fig. 5. Face; Fig. 6. Thorax, ventral view; Fig. 7. Right labial palpus (scanning electron picture); Fig. 8. Right hind leg of male; Fig. 9. Tibial spur of hind leg (scanning electron picture); Fig. 10. Ectoprocts of male, dorsal view; Fig. 11. Left ectoproct of male, lateral view; Fig. 12. Gonarcus-paramere complexe, right lateral view; Fig. 13. Gonarcus-paramere complexe, ventral view; Fig. 14. Hypandrium internum, lateral view." httpUri="https://zenodo.org/record/278720/files/figure.png" pageId="1" pageNumber="41">Fig. 8</figureCitation>
). Tibial spurs absent or at most as long as first tarsomere in fore legs and middle legs, shorter than first tarsomere in hind legs. Postventral lobe of male ectoprocts clearly more than 4 times longer than wide, with a few erect stout setae (this character must be verified for
<taxonomicName id="EFA12DF1FFE0531CFDC0FCAAAA023077" box="[631,785,790,814]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="fulvicauda">
<emphasis id="1AD58A60FFE0531CFDC0FCAAAA023077" box="[631,785,790,814]" italics="true" pageId="1" pageNumber="41">C. fulvicauda</emphasis>
</taxonomicName>
). In his description Navás mentions “his [ectoprocts] in parte horizontali cylindricis, tenuibus, longiter pilosis (
<figureCitation id="B09A4AF7FFE0531CFCACFC87AA71300B" box="[795,866,826,850]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1214,1237]" captionTargetBox="[151,1436,270,1150]" captionTargetId="figure@2.[151,1436,270,1190]" captionTargetPageId="2" captionText="FIGURES 1 3. Capicua fulvicauda. Fig. 1. Lectotype with labels; Fig. 2. Left middle leg of lectotype; Fig. 3. Extremity of male abdomen (a) and tarsus (b) (after Navás, 1921)." httpUri="https://zenodo.org/record/278719/files/figure.png" pageId="1" pageNumber="41">Fig. 3</figureCitation>
a). Parameres hook shaped. Gonarcus U shaped. Female anterior gonapophyses absent (
<figureCitation id="B09A4AF7FFE0531CFDFCFCE3AB8C302F" box="[587,671,862,886]" captionStart="FIGURES 15 25" captionStartId="5.[151,264,1892,1915]" captionTargetBox="[219,1369,193,1871]" captionTargetId="figure@5.[219,1371,193,1885]" captionTargetPageId="5" captionText="FIGURES 15 25. Capicua acalcarata. Fig. 15. Holotype with labels; Fig. 16. Face; Fig. 17. Thorax, ventral view; Fig. 18. Right labial palpus (scanning electron picture); Fig. 19. Extremity of tibia and first tarsomere of right hind leg (scanning electron picture); Fig. 20. Ectoprocts of male, dorsal view; Fig. 21. Left ectoproct of male, lateral view; Fig. 22. Gonarcus-paramere complexe, right lateral view; Fig. 23. Gonarcus-paramere complexe, ventral view; Fig. 24. Hypandrium internum, lateral view; Fig. 25. Female terminalia, left lateral view." httpUri="https://zenodo.org/record/278721/files/figure.png" pageId="1" pageNumber="41">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="281E5672FFE0531CFF70FC3EAD093717" blockId="1.[151,1437,358,1102]" pageId="1" pageNumber="41">
<taxonomicName id="EFA12DF1FFE0531CFF70FC3EA83530C3" box="[199,294,899,922]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="1AD58A60FFE0531CFF70FC3EA83530C3" box="[199,294,899,922]" italics="true" pageId="1" pageNumber="41">Capicua</emphasis>
</taxonomicName>
can be differentiated from the genus
<taxonomicName id="EFA12DF1FFE0531CFD70FC3EAA0630C3" box="[711,789,899,922]" class="Insecta" family="Myrmeleontidae" genus="Geyria" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="1AD58A60FFE0531CFD70FC3EAA0630C3" box="[711,789,899,922]" italics="true" pageId="1" pageNumber="41">Geyria</emphasis>
</taxonomicName>
by its larger size, the absence of a tuft of long white setae on the fore coxae and fore femora, the clearly shorter tibial spurs at most as long as first tarsomere, the narrow wings and dark coloration of the body. The absence of tibial spurs or at most as long as first tarsomere distinguishes
<taxonomicName id="EFA12DF1FFE0531CFF20FC52A9E5375F" box="[151,246,1007,1030]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="1AD58A60FFE0531CFF20FC52A9E5375F" box="[151,246,1007,1030]" italics="true" pageId="1" pageNumber="41">Capicua</emphasis>
</taxonomicName>
from
<taxonomicName id="EFA12DF1FFE0531CFE89FC4DAB6A375F" authority="Costa, 1855" authorityName="Costa" authorityYear="1855" box="[318,633,1006,1031]" class="Insecta" family="Myrmeleontidae" genus="Macronemurus" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="1AD58A60FFE0531CFE89FC4DA8FB375E" box="[318,488,1008,1031]" italics="true" pageId="1" pageNumber="41">Macronemurus</emphasis>
Costa, 1855
</taxonomicName>
and
<taxonomicName id="EFA12DF1FFE0531CFD04FC4DAAF8375F" authority="Navas, 1920" authorityName="Navas" authorityYear="1920" box="[691,1003,1006,1031]" class="Insecta" family="Myrmeleontidae" genus="Mesonemurus" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="1AD58A60FFE0531CFD04FC4DAA43375E" box="[691,848,1008,1031]" italics="true" pageId="1" pageNumber="41">Mesonemurus</emphasis>
Navás, 1920
</taxonomicName>
. The length of the postventral lobe of the male ectoprocts which is clearly more than 4 times longer than wide and the basitarsus of the hind legs being shorter than distal tarsomere differentiate
<taxonomicName id="EFA12DF1FFE0531CFDDCFB8AABD93717" box="[619,714,1079,1102]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="1AD58A60FFE0531CFDDCFB8AABD93717" box="[619,714,1079,1102]" italics="true" pageId="1" pageNumber="41">Capicua</emphasis>
</taxonomicName>
from
<taxonomicName id="EFA12DF1FFE0531CFCB9FB8BAD053717" authority="Navas, 1912" authorityName="Navas" authorityYear="1912" box="[782,1046,1078,1102]" class="Insecta" family="Myrmeleontidae" genus="Delfimeus" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="1AD58A60FFE0531CFCB9FB8BAA933717" box="[782,896,1078,1102]" italics="true" pageId="1" pageNumber="41">Delfimeus</emphasis>
Navás, 1912
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,198 @@
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<paragraph id="281E5672FFE0531CFF20FB1FAB1C37E2" blockId="1.[151,527,1186,1246]" box="[151,527,1186,1211]" pageId="1" pageNumber="41">
<heading id="7356E11EFFE0531CFF20FB1FAB1C37E2" bold="true" box="[151,527,1186,1211]" fontSize="10" level="2" pageId="1" pageNumber="41" reason="2">
<taxonomicName id="EFA12DF1FFE0531CFF20FB1FAB1C37E2" ID-CoL="5X9JP" authority="Navas, 1921" authorityName="Navas" authorityYear="1921" box="[151,527,1186,1211]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="fulvicauda">
<emphasis id="1AD58A60FFE0531CFF20FB1FAB1C37E2" bold="true" box="[151,527,1186,1211]" pageId="1" pageNumber="41">
<emphasis id="1AD58A60FFE0531CFF20FB1FA86A37E2" bold="true" box="[151,377,1186,1211]" italics="true" pageId="1" pageNumber="41">Capicua fulvicauda</emphasis>
<bibRefCitation id="4C302B83FFE0531CFE37FB1FAB1C37E2" author="Navas" box="[384,527,1186,1211]" pageId="1" pageNumber="41" refString="Navas, L. (1921) Comunicaciones entomologicas. 4. Insectos exoticos nuevos, criticos o poco conocidos. Revista de la Real Academia de Ciencias Exactas Fisico-Quimicas y Naturales de Zaragoza, 6, 61 - 81." type="journal article" year="1921">Navás, 1921</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="281E5672FFE0531CFF20FB7BA8013787" blockId="1.[151,527,1186,1246]" box="[151,274,1222,1246]" pageId="1" pageNumber="41">
(
<figureCitation id="B09A4AF7FFE0531CFF28FB7BA81A3787" box="[159,265,1222,1246]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1214,1237]" captionTargetBox="[151,1436,270,1150]" captionTargetId="figure@2.[151,1436,270,1190]" captionTargetPageId="2" captionText="FIGURES 1 3. Capicua fulvicauda. Fig. 1. Lectotype with labels; Fig. 2. Left middle leg of lectotype; Fig. 3. Extremity of male abdomen (a) and tarsus (b) (after Navás, 1921)." httpUri="https://zenodo.org/record/278719/files/figure.png" pageId="1" pageNumber="41">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="60BB05F9FFE0531CFF20FAB2A9E1354E" pageId="1" pageNumber="41" type="reference_group">
<paragraph id="281E5672FFE0531CFF20FAB2AB10367D" blockId="1.[151,681,1294,1348]" box="[151,515,1294,1317]" pageId="1" pageNumber="41">
<treatmentCitationGroup id="08B1715CFFE0531CFF20FAB2AB10367D" box="[151,515,1294,1317]" pageId="1" pageNumber="41">
<taxonomicName id="EFA12DF1FFE0531CFF20FAB2AB10367D" ID-CoL="5X9JP" authority="Navas, 1921:66" authorityName="Navas" authorityPageNumber="66" authorityYear="1921" box="[151,515,1294,1317]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="fulvicauda">
<emphasis id="1AD58A60FFE0531CFF20FAB2A84D367C" box="[151,350,1295,1317]" italics="true" pageId="1" pageNumber="41">Capicua fulvicauda</emphasis>
<treatmentCitation id="A9007063FFE0531CFED3FAB3AB10367D" author="Navas" box="[356,515,1294,1316]" page="66" pageId="1" pageNumber="41" year="1921">
<bibRefCitation id="4C302B83FFE0531CFED3FAB3A8F7367D" author="Navas" box="[356,484,1294,1316]" pageId="1" pageNumber="41" refString="Navas, L. (1921) Comunicaciones entomologicas. 4. Insectos exoticos nuevos, criticos o poco conocidos. Revista de la Real Academia de Ciencias Exactas Fisico-Quimicas y Naturales de Zaragoza, 6, 61 - 81." type="journal article" year="1921">Navás, 1921</bibRefCitation>
:66
</treatmentCitation>
</taxonomicName>
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</paragraph>
<paragraph id="281E5672FFE0531CFF20FA93ABBA361A" blockId="1.[151,681,1294,1348]" box="[151,681,1325,1348]" pageId="1" pageNumber="41">
<treatmentCitationGroup id="08B1715CFFE0531CFF20FA93ABBA361A" box="[151,681,1325,1348]" pageId="1" pageNumber="41">
<taxonomicName id="EFA12DF1FFE0531CFF20FA93A8F4361D" ID-CoL="3FYH3" authority="Navas, 1921" authorityName="Navas" authorityYear="1921" box="[151,487,1325,1348]" class="Insecta" family="Myrmeleontidae" genus="Geyria" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="fulvicauda">
<emphasis id="1AD58A60FFE0531CFF20FA93A85C361A" box="[151,335,1325,1347]" italics="true" pageId="1" pageNumber="41">Geyria fulvicauda</emphasis>
(
<bibRefCitation id="4C302B83FFE0531CFEECFA90A8CC361D" author="Navas" box="[347,479,1325,1348]" pageId="1" pageNumber="41" refString="Navas, L. (1921) Comunicaciones entomologicas. 4. Insectos exoticos nuevos, criticos o poco conocidos. Revista de la Real Academia de Ciencias Exactas Fisico-Quimicas y Naturales de Zaragoza, 6, 61 - 81." type="journal article" year="1921">Navás, 1921</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="A9007063FFE0531CFE46FA93ABB6361A" author="Stange" box="[497,677,1325,1347]" page="176" pageId="1" pageNumber="41" year="2004">
<bibRefCitation id="4C302B83FFE0531CFE46FA93AB69361A" author="Stange" box="[497,634,1325,1347]" pageId="1" pageNumber="41" refString="Stange, L. A. (2004) A systematic catalog, bibliography and classification of the world antlions (Insecta: Neuroptera: Myrmeleontidae). Memoirs of the American Entomological Institute, 74, 1 - 565." type="journal article" year="2004">Stange, 2004</bibRefCitation>
:176
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="281E5672FFE0531CFF20FAD2A9E1354E" blockId="1.[151,1436,1391,2028]" pageId="1" pageNumber="41">
Navás neither indicated how many specimens he examined in describing the species nor did he designate the specimen in Muséum National dHistoire Naturelle, Paris as
<typeStatus id="F71AE8D0FFE0531CFCADFA29AA9336F5" box="[794,896,1428,1452]" pageId="1" pageNumber="41" type="holotype">holotype</typeStatus>
. Consequently this specimen is designated here as
<typeStatus id="F71AE8D0FFE0531CFF00FA0AA80D3696" box="[183,286,1463,1487]" pageId="1" pageNumber="41" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="B09A4AF7FFE0531CFE98FA0AA8613696" box="[303,370,1463,1487]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1214,1237]" captionTargetBox="[151,1436,270,1150]" captionTargetId="figure@2.[151,1436,270,1190]" captionTargetPageId="2" captionText="FIGURES 1 3. Capicua fulvicauda. Fig. 1. Lectotype with labels; Fig. 2. Left middle leg of lectotype; Fig. 3. Extremity of male abdomen (a) and tarsus (b) (after Navás, 1921)." httpUri="https://zenodo.org/record/278719/files/figure.png" pageId="1" pageNumber="41">Fig. 1</figureCitation>
). This male specimen is lacking both antennal flagella, tarsomeres IVV of the right foreleg and IIIV of the left foreleg, the entire right middle leg and abdomen (
<figureCitation id="B09A4AF7FFE0531CFC1EFA61AAF936AD" box="[937,1002,1500,1524]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1214,1237]" captionTargetBox="[151,1436,270,1150]" captionTargetId="figure@2.[151,1436,270,1190]" captionTargetPageId="2" captionText="FIGURES 1 3. Capicua fulvicauda. Fig. 1. Lectotype with labels; Fig. 2. Left middle leg of lectotype; Fig. 3. Extremity of male abdomen (a) and tarsus (b) (after Navás, 1921)." httpUri="https://zenodo.org/record/278719/files/figure.png" pageId="1" pageNumber="41">Fig. 1</figureCitation>
). The left middle leg is glued to a card (
<figureCitation id="B09A4AF7FFE0531CFF28FA42A9F6354E" box="[159,229,1535,1559]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1214,1237]" captionTargetBox="[151,1436,270,1150]" captionTargetId="figure@2.[151,1436,270,1190]" captionTargetPageId="2" captionText="FIGURES 1 3. Capicua fulvicauda. Fig. 1. Lectotype with labels; Fig. 2. Left middle leg of lectotype; Fig. 3. Extremity of male abdomen (a) and tarsus (b) (after Navás, 1921)." httpUri="https://zenodo.org/record/278719/files/figure.png" pageId="1" pageNumber="41">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="60BB05F9FFE0531CFF70F99EAA3034B5" pageId="1" pageNumber="41" type="description">
<paragraph id="281E5672FFE0531CFF70F99EAA3034B5" blockId="1.[151,1436,1391,2028]" pageId="1" pageNumber="41">
<emphasis id="1AD58A60FFE0531CFF70F99EA8BF3565" bold="true" box="[199,428,1571,1596]" pageId="1" pageNumber="41">Redescription. Size</emphasis>
: small (forewing:
<quantity id="EF59FB97FFE0531CFDCBF999ABF13565" box="[636,738,1572,1596]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="1" pageNumber="41" unit="mm" value="21.5">21.5 mm</quantity>
, hind wing:
<quantity id="EF59FB97FFE0531CFCD9F999AAC53565" box="[878,982,1572,1596]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="1" pageNumber="41" unit="mm" value="21.5">21.5 mm</quantity>
).
<emphasis id="1AD58A60FFE0531CFC5DF999AD233565" bold="true" box="[1002,1072,1572,1596]" pageId="1" pageNumber="41">Head.</emphasis>
Vertex slightly raised. Face yellow with black chevron-shaped mark under scapes and brown above. No yellow transverse line between bases of antennae. Vertex brown limited anteriorly by a black line. Palpi yellow. Apical segment of labial palp very slightly enlarged.
<emphasis id="1AD58A60FFE0531CFEB2F92DA87635F1" bold="true" box="[261,357,1680,1704]" pageId="1" pageNumber="41">Thorax.</emphasis>
Brown with darker markings.
<emphasis id="1AD58A60FFE0531CFD76F932AA0035F1" bold="true" box="[705,787,1679,1704]" pageId="1" pageNumber="41">Wings.</emphasis>
Narrow without markings. Veins light brown. Longitudinal veins slightly darker at junction with crossveins. Forewing: 15 crossveins between origin of RS and pterostigma.
<emphasis id="1AD58A60FFE0531CFE9EF965A87B35A9" bold="true" box="[297,360,1752,1776]" pageId="1" pageNumber="41">Legs.</emphasis>
Yellow. Three long white slightly spatulated setae on right fore coxa and four on left fore coxa. No tuft of long white setae on fore femora. Male hind femora with a tuft of long erect setae on dorsal surface. Tibial spurs as long as first tarsomere in fore legs and middle legs (
<figureCitation id="B09A4AF7FFE0531CFCF1F8A2AA9B346E" box="[838,904,1823,1847]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1214,1237]" captionTargetBox="[151,1436,270,1150]" captionTargetId="figure@2.[151,1436,270,1190]" captionTargetPageId="2" captionText="FIGURES 1 3. Capicua fulvicauda. Fig. 1. Lectotype with labels; Fig. 2. Left middle leg of lectotype; Fig. 3. Extremity of male abdomen (a) and tarsus (b) (after Navás, 1921)." httpUri="https://zenodo.org/record/278719/files/figure.png" pageId="1" pageNumber="41">Fig. 3</figureCitation>
b) and shorter than first tarsomere in hind legs.
<collectingCountry id="50B616E2FFE0531CFF20F8F9A9DC3405" box="[151,207,1860,1884]" name="India" pageId="1" pageNumber="41">Hind</collectingCountry>
legs more slender than fore legs and middle legs.
<emphasis id="1AD58A60FFE0531CFCBBF8F9AA753405" bold="true" box="[780,870,1860,1884]" pageId="1" pageNumber="41">Foreleg</emphasis>
: femur
<quantity id="EF59FB97FFE0531CFC76F8F9AD3B3405" box="[961,1064,1860,1884]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="1" pageNumber="41" unit="mm" value="0.98">0.98 mm</quantity>
, tibia
<quantity id="EF59FB97FFE0531CFBD8F8F9ADC53405" box="[1135,1238,1860,1884]" metricMagnitude="-3" metricUnit="m" metricValue="1.27" pageId="1" pageNumber="41" unit="mm" value="1.27">1.27 mm</quantity>
, tarsomere I
<quantity id="EF59FB97FFE0531CFADCF8F9A9D534D9" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="41" unit="mm" value="0.25">0.25 mm</quantity>
, II
<quantity id="EF59FB97FFE0531CFF5AF8D5A84634D9" box="[237,341,1896,1920]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="41" unit="mm" value="0.16">0.16 mm</quantity>
, III
<quantity id="EF59FB97FFE0531CFE32F8D5A8FE34D9" box="[389,493,1896,1920]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="41" unit="mm" value="0.16">0.16 mm</quantity>
, tibial spur
<quantity id="EF59FB97FFE0531CFDC4F8D5ABC834D9" box="[627,731,1896,1920]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="41" unit="mm" value="0.25">0.25 mm</quantity>
.
<emphasis id="1AD58A60FFE0531CFD50F8DAAA7534D9" bold="true" box="[743,870,1895,1920]" pageId="1" pageNumber="41">Middle leg</emphasis>
: femur
<quantity id="EF59FB97FFE0531CFC75F8D5AD3B34D9" box="[962,1064,1896,1920]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="1" pageNumber="41" unit="mm" value="1.11">1.11 mm</quantity>
, tibia
<quantity id="EF59FB97FFE0531CFBD8F8D5ADC534D9" box="[1135,1238,1896,1920]" metricMagnitude="-3" metricUnit="m" metricValue="1.27" pageId="1" pageNumber="41" unit="mm" value="1.27">1.27 mm</quantity>
, tarsomere I
<quantity id="EF59FB97FFE0531CFADCF8D5A9D634FD" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="41" unit="mm" value="0.25">0.25 mm</quantity>
, II
<quantity id="EF59FB97FFE0531CFF5CF831A84234FD" box="[235,337,1932,1956]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="41" unit="mm" value="0.16">0.16 mm</quantity>
, III
<quantity id="EF59FB97FFE0531CFE37F831A8F534FD" box="[384,486,1932,1956]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="41" unit="mm" value="0.16">0.16 mm</quantity>
, IV
<quantity id="EF59FB97FFE0531CFDA0F831AB6E34FD" box="[535,637,1932,1956]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="41" unit="mm" value="0.16">0.16 mm</quantity>
, V
<quantity id="EF59FB97FFE0531CFD13F831AA1834FD" box="[676,779,1932,1956]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="1" pageNumber="41" unit="mm" value="0.54">0.54 mm</quantity>
, tibial spur
<quantity id="EF59FB97FFE0531CFC39F831AAE534FD" box="[910,1014,1932,1956]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="41" unit="mm" value="0.25">0.25 mm</quantity>
.
<emphasis id="1AD58A60FFE0531CFBB7F836AD7534FD" bold="true" box="[1024,1126,1931,1956]" pageId="1" pageNumber="41">
<collectingCountry id="50B616E2FFE0531CFBB7F836AD2F34FD" box="[1024,1084,1931,1956]" name="India" pageId="1" pageNumber="41">Hind</collectingCountry>
leg
</emphasis>
: femur
<quantity id="EF59FB97FFE0531CFB77F831AC3534FD" box="[1216,1318,1932,1956]" metricMagnitude="-3" metricUnit="m" metricValue="1.46" pageId="1" pageNumber="41" unit="mm" value="1.46">1.46 mm</quantity>
, tibia
<quantity id="EF59FB97FFE0531CFADCF831A9D63491" metricMagnitude="-3" metricUnit="m" metricValue="1.49" pageId="1" pageNumber="41" unit="mm" value="1.49">1.49 mm</quantity>
, tarsomere I
<quantity id="EF59FB97FFE0531CFEE0F80DA8AD3491" box="[343,446,1968,1992]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="41" unit="mm" value="0.32">0.32 mm</quantity>
, II
<quantity id="EF59FB97FFE0531CFE53F80DAB593491" box="[484,586,1968,1992]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="41" unit="mm" value="0.19">0.19 mm</quantity>
, III
<quantity id="EF59FB97FFE0531CFDCDF80DABF33491" box="[634,736,1968,1992]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="41" unit="mm" value="0.19">0.19 mm</quantity>
, IV
<quantity id="EF59FB97FFE0531CFCA6F80DAA6A3491" box="[785,889,1968,1992]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="41" unit="mm" value="0.19">0.19 mm</quantity>
, V
<quantity id="EF59FB97FFE0531CFC28F80DAD153491" box="[927,1030,1968,1992]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="1" pageNumber="41" unit="mm" value="0.54">0.54 mm</quantity>
, tibial spur:
<quantity id="EF59FB97FFE0531CFB24F80DADE93491" box="[1171,1274,1968,1992]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="41" unit="mm" value="0.22">0.22 mm</quantity>
. Ectoprocts of male clearly more than 4 times longer than wide (
<figureCitation id="B09A4AF7FFE0531CFD73F869AA1534B5" box="[708,774,2004,2028]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1214,1237]" captionTargetBox="[151,1436,270,1150]" captionTargetId="figure@2.[151,1436,270,1190]" captionTargetPageId="2" captionText="FIGURES 1 3. Capicua fulvicauda. Fig. 1. Lectotype with labels; Fig. 2. Left middle leg of lectotype; Fig. 3. Extremity of male abdomen (a) and tarsus (b) (after Navás, 1921)." httpUri="https://zenodo.org/record/278719/files/figure.png" pageId="1" pageNumber="41">Fig. 3</figureCitation>
a).
</paragraph>
</subSubSection>
<subSubSection id="60BB05F9FFE3531FFF70FF26ABEF338E" pageId="2" pageNumber="42" type="materials_examined">
<paragraph id="281E5672FFE3531FFF70FF26ABEF338E" blockId="2.[151,1435,151,215]" pageId="2" pageNumber="42">
<emphasis id="1AD58A60FFE3531FFF70FF26A8E433EA" bold="true" box="[199,503,154,179]" pageId="2" pageNumber="42">
<typeStatus id="F71AE8D0FFE3531FFF70FF26A81133EA" box="[199,258,155,179]" pageId="2" pageNumber="42">Type</typeStatus>
material.
<typeStatus id="F71AE8D0FFE3531FFE36FF26A8E433EA" box="[385,503,155,179]" pageId="2" pageNumber="42" type="lectotype">Lectotype</typeStatus>
</emphasis>
3—
<collectingCountry id="50B616E2FFE3531FFD85FF27AB9033EB" box="[562,643,154,178]" name="Chad" pageId="2" pageNumber="42">CHAD</collectingCountry>
, Fort Archambault (presently Sarh) (
<geoCoordinate id="4D9530B5FFE3531FFB9AFF27AD9C33EB" box="[1069,1167,154,178]" direction="north" orientation="latitude" pageId="2" pageNumber="42" precision="925" value="9.15">09°09N</geoCoordinate>
<geoCoordinate id="4D9530B5FFE3531FFB2EFF27ADEA33EB" box="[1177,1273,154,178]" direction="east" orientation="longitude" pageId="2" pageNumber="42" precision="925" value="18.383333">18°23E</geoCoordinate>
),
<date id="5C1F70B2FFE3531FFAA6FF27A9C2338E" pageId="2" pageNumber="42" value="1904-06-01" valueMax="1904-06-15" valueMin="1904-06-01">1st15th June 1904</date>
, Chari-Chad mission, Dr. J. Decorse rec. (
<figureCitation id="B09A4AF7FFE3531FFD1DFF02ABF8338E" box="[682,747,191,215]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1214,1237]" captionTargetBox="[151,1436,270,1150]" captionTargetId="figure@2.[151,1436,270,1190]" captionTargetPageId="2" captionText="FIGURES 1 3. Capicua fulvicauda. Fig. 1. Lectotype with labels; Fig. 2. Left middle leg of lectotype; Fig. 3. Extremity of male abdomen (a) and tarsus (b) (after Navás, 1921)." httpUri="https://zenodo.org/record/278719/files/figure.png" pageId="2" pageNumber="42">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,277 @@
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<heading id="7356E11EFFE3531FFF20FA82A88E360E" bold="true" box="[151,413,1343,1368]" fontSize="10" level="2" pageId="2" pageNumber="42" reason="2">
<emphasis id="1AD58A60FFE3531FFF20FA82A88E360E" bold="true" box="[151,413,1343,1368]" pageId="2" pageNumber="42">
<taxonomicName id="EFA12DF1FFE3531FFF20FA82A82F3601" ID-CoL="68XJP" box="[151,316,1343,1368]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
<emphasis id="1AD58A60FFE3531FFF20FA82A82F3601" bold="true" box="[151,316,1343,1368]" italics="true" pageId="2" pageNumber="42">Capicua nigra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="01E6371BFFE3531FFEF3FA82A88E360E" box="[324,413,1343,1367]" pageId="2" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="281E5672FFE3531FFF20FADEA8333622" blockId="2.[151,413,1343,1403]" box="[151,288,1379,1403]" pageId="2" pageNumber="42">
(
<figureCitation id="B09A4AF7FFE3531FFF28FADEA80B3622" box="[159,280,1379,1403]" captionStart="FIGURES 4 14" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[224,1358,193,1850]" captionTargetId="figure@3.[224,1358,193,1861]" captionTargetPageId="3" captionText="FIGURES 4 14. Capicua nigra. Fig. 4. Holotype with labels; Fig. 5. Face; Fig. 6. Thorax, ventral view; Fig. 7. Right labial palpus (scanning electron picture); Fig. 8. Right hind leg of male; Fig. 9. Tibial spur of hind leg (scanning electron picture); Fig. 10. Ectoprocts of male, dorsal view; Fig. 11. Left ectoproct of male, lateral view; Fig. 12. Gonarcus-paramere complexe, right lateral view; Fig. 13. Gonarcus-paramere complexe, ventral view; Fig. 14. Hypandrium internum, lateral view." httpUri="https://zenodo.org/record/278720/files/figure.png" pageId="2" pageNumber="42">Figs. 414</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="60BB05F9FFE3531FFF20FA17AA933576" pageId="2" pageNumber="42" type="diagnosis">
<paragraph id="281E5672FFE3531FFF20FA17AA933576" blockId="2.[151,1437,1450,2015]" pageId="2" pageNumber="42">
<emphasis id="1AD58A60FFE3531FFF20FA17A802369A" bold="true" box="[151,273,1450,1475]" pageId="2" pageNumber="42">Diagnosis.</emphasis>
General body coloration black (
<figureCitation id="B09A4AF7FFE3531FFDCCFA16ABD2369A" box="[635,705,1451,1475]" captionStart="FIGURES 4 14" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[224,1358,193,1850]" captionTargetId="figure@3.[224,1358,193,1861]" captionTargetPageId="3" captionText="FIGURES 4 14. Capicua nigra. Fig. 4. Holotype with labels; Fig. 5. Face; Fig. 6. Thorax, ventral view; Fig. 7. Right labial palpus (scanning electron picture); Fig. 8. Right hind leg of male; Fig. 9. Tibial spur of hind leg (scanning electron picture); Fig. 10. Ectoprocts of male, dorsal view; Fig. 11. Left ectoproct of male, lateral view; Fig. 12. Gonarcus-paramere complexe, right lateral view; Fig. 13. Gonarcus-paramere complexe, ventral view; Fig. 14. Hypandrium internum, lateral view." httpUri="https://zenodo.org/record/278720/files/figure.png" pageId="2" pageNumber="42">Fig. 4</figureCitation>
). Base of legs yellow contrasting with the black coloration of the thoracic sternites (
<figureCitation id="B09A4AF7FFE3531FFED0FA72A8BE36BE" box="[359,429,1487,1511]" captionStart="FIGURES 4 14" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[224,1358,193,1850]" captionTargetId="figure@3.[224,1358,193,1861]" captionTargetPageId="3" captionText="FIGURES 4 14. Capicua nigra. Fig. 4. Holotype with labels; Fig. 5. Face; Fig. 6. Thorax, ventral view; Fig. 7. Right labial palpus (scanning electron picture); Fig. 8. Right hind leg of male; Fig. 9. Tibial spur of hind leg (scanning electron picture); Fig. 10. Ectoprocts of male, dorsal view; Fig. 11. Left ectoproct of male, lateral view; Fig. 12. Gonarcus-paramere complexe, right lateral view; Fig. 13. Gonarcus-paramere complexe, ventral view; Fig. 14. Hypandrium internum, lateral view." httpUri="https://zenodo.org/record/278720/files/figure.png" pageId="2" pageNumber="42">Fig. 6</figureCitation>
). Tibial spurs clearly shorter than first tarsomere (
<figureCitation id="B09A4AF7FFE3531FFC5EFA72AD3C36BE" box="[1001,1071,1487,1511]" captionStart="FIGURES 4 14" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[224,1358,193,1850]" captionTargetId="figure@3.[224,1358,193,1861]" captionTargetPageId="3" captionText="FIGURES 4 14. Capicua nigra. Fig. 4. Holotype with labels; Fig. 5. Face; Fig. 6. Thorax, ventral view; Fig. 7. Right labial palpus (scanning electron picture); Fig. 8. Right hind leg of male; Fig. 9. Tibial spur of hind leg (scanning electron picture); Fig. 10. Ectoprocts of male, dorsal view; Fig. 11. Left ectoproct of male, lateral view; Fig. 12. Gonarcus-paramere complexe, right lateral view; Fig. 13. Gonarcus-paramere complexe, ventral view; Fig. 14. Hypandrium internum, lateral view." httpUri="https://zenodo.org/record/278720/files/figure.png" pageId="2" pageNumber="42">Fig. 9</figureCitation>
). No tuft of long white setae on fore femora. Wings narrow. Male ectoprocts with postventral lobe well developed, black proximally then yellow with four or five stout black setae inserted in the apical two-thirds.
</paragraph>
</subSubSection>
<subSubSection id="60BB05F9FFE35319FF70F987AD4B3195" lastPageId="4" lastPageNumber="44" pageId="2" pageNumber="42" type="description">
<paragraph id="281E5672FFE3531FFF70F987A83335BA" blockId="2.[151,1437,1450,2015]" pageId="2" pageNumber="42">
<emphasis id="1AD58A60FFE3531FFF70F987A8B6350A" bold="true" box="[199,421,1594,1619]" pageId="2" pageNumber="42">Description. Head.</emphasis>
Vertex raised. Face yellow with a black chevron-shaped mark under the scapes. A thin yellow line between bases of antennae and grey above. Vertex grey delimited anteriorly by a black line (
<figureCitation id="B09A4AF7FFE3531FFAF1F9E2AC9D352E" box="[1350,1422,1631,1655]" captionStart="FIGURES 4 14" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[224,1358,193,1850]" captionTargetId="figure@3.[224,1358,193,1861]" captionTargetPageId="3" captionText="FIGURES 4 14. Capicua nigra. Fig. 4. Holotype with labels; Fig. 5. Face; Fig. 6. Thorax, ventral view; Fig. 7. Right labial palpus (scanning electron picture); Fig. 8. Right hind leg of male; Fig. 9. Tibial spur of hind leg (scanning electron picture); Fig. 10. Ectoprocts of male, dorsal view; Fig. 11. Left ectoproct of male, lateral view; Fig. 12. Gonarcus-paramere complexe, right lateral view; Fig. 13. Gonarcus-paramere complexe, ventral view; Fig. 14. Hypandrium internum, lateral view." httpUri="https://zenodo.org/record/278720/files/figure.png" pageId="2" pageNumber="42">Fig. 5</figureCitation>
). Occiput with black and yellow markings. Palpi yellow. Last segment of labial palp slightly enlarged (
<figureCitation id="B09A4AF7FFE3531FFAF2F93EAC9D35C2" box="[1349,1422,1667,1691]" captionStart="FIGURES 4 14" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[224,1358,193,1850]" captionTargetId="figure@3.[224,1358,193,1861]" captionTargetPageId="3" captionText="FIGURES 4 14. Capicua nigra. Fig. 4. Holotype with labels; Fig. 5. Face; Fig. 6. Thorax, ventral view; Fig. 7. Right labial palpus (scanning electron picture); Fig. 8. Right hind leg of male; Fig. 9. Tibial spur of hind leg (scanning electron picture); Fig. 10. Ectoprocts of male, dorsal view; Fig. 11. Left ectoproct of male, lateral view; Fig. 12. Gonarcus-paramere complexe, right lateral view; Fig. 13. Gonarcus-paramere complexe, ventral view; Fig. 14. Hypandrium internum, lateral view." httpUri="https://zenodo.org/record/278720/files/figure.png" pageId="2" pageNumber="42">Fig. 7</figureCitation>
). Antennae: scape brown; pedicel brown, finely bordered with yellow apically; flagellomeres brown; ventral face of club yellow.
</paragraph>
<paragraph id="281E5672FFE3531FFF72F952AB613472" blockId="2.[151,1437,1450,2015]" pageId="2" pageNumber="42">
<emphasis id="1AD58A60FFE3531FFF72F952A837345E" bold="true" box="[197,292,1775,1799]" pageId="2" pageNumber="42">Thorax.</emphasis>
Pronotum black, wider than long, with 3 to 5 long white setae forwardly curved. Mesothorax and metathorax black, brown at base of wings.
</paragraph>
<paragraph id="281E5672FFE35319FF72F88AA9E13266" blockId="2.[151,1437,1450,2015]" lastBlockId="4.[151,1437,151,1004]" lastPageId="4" lastPageNumber="44" pageId="2" pageNumber="42">
<emphasis id="1AD58A60FFE3531FFF72F88AA87F3416" bold="true" box="[197,364,1847,1871]" pageId="2" pageNumber="42">Legs. Foreleg.</emphasis>
Coxa, trochanter, femur and tibia brown, yellow ventrally (
<figureCitation id="B09A4AF7FFE3531FFBBFF88AAD5E3416" box="[1032,1101,1847,1871]" captionStart="FIGURES 4 14" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[224,1358,193,1850]" captionTargetId="figure@3.[224,1358,193,1861]" captionTargetPageId="3" captionText="FIGURES 4 14. Capicua nigra. Fig. 4. Holotype with labels; Fig. 5. Face; Fig. 6. Thorax, ventral view; Fig. 7. Right labial palpus (scanning electron picture); Fig. 8. Right hind leg of male; Fig. 9. Tibial spur of hind leg (scanning electron picture); Fig. 10. Ectoprocts of male, dorsal view; Fig. 11. Left ectoproct of male, lateral view; Fig. 12. Gonarcus-paramere complexe, right lateral view; Fig. 13. Gonarcus-paramere complexe, ventral view; Fig. 14. Hypandrium internum, lateral view." httpUri="https://zenodo.org/record/278720/files/figure.png" pageId="2" pageNumber="42">Fig. 6</figureCitation>
). Coxa with 2 or 3 long white setae. Tarsomeres IIV yellow with the apical half brown. Tarsomere V yellow, brown apically. Tibial spurs brown, shorter than first tarsomere.
<emphasis id="1AD58A60FFE3531FFE65F8C3AB4334CE" bold="true" box="[466,592,1918,1943]" pageId="2" pageNumber="42">Middle leg</emphasis>
. Black with yellow marking along the internal side. Trochanter brown, yellow ventrally. Femur brown, yellow basally and apically. Tibia yellowish with brown marking on dorsal face. Tibial spurs shorter than first tarsomere.
<emphasis id="1AD58A60FFE3531FFDF2F87BABA03486" bold="true" box="[581,691,1990,2015]" pageId="2" pageNumber="42">
<collectingCountry id="50B616E2FFE3531FFDF2F87BAB913486" box="[581,642,1990,2015]" name="India" pageId="2" pageNumber="42">Hind</collectingCountry>
leg.
</emphasis>
Similar to middle leg but with a tuft of long erect setae on dorsal face (
<figureCitation id="B09A4AF7FFE55319FF61FF2AA80F33F6" box="[214,284,151,175]" captionStart="FIGURES 4 14" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[224,1358,193,1850]" captionTargetId="figure@3.[224,1358,193,1861]" captionTargetPageId="3" captionText="FIGURES 4 14. Capicua nigra. Fig. 4. Holotype with labels; Fig. 5. Face; Fig. 6. Thorax, ventral view; Fig. 7. Right labial palpus (scanning electron picture); Fig. 8. Right hind leg of male; Fig. 9. Tibial spur of hind leg (scanning electron picture); Fig. 10. Ectoprocts of male, dorsal view; Fig. 11. Left ectoproct of male, lateral view; Fig. 12. Gonarcus-paramere complexe, right lateral view; Fig. 13. Gonarcus-paramere complexe, ventral view; Fig. 14. Hypandrium internum, lateral view." httpUri="https://zenodo.org/record/278720/files/figure.png" pageId="4" pageNumber="44">Fig. 8</figureCitation>
).
<emphasis id="1AD58A60FFE55319FE85FF2AAA1C33E9" bold="true" box="[306,783,151,176]" pageId="4" pageNumber="44">
Dimensions of a male
<typeStatus id="F71AE8D0FFE55319FD8BFF25ABBA33E9" box="[572,681,152,176]" pageId="4" pageNumber="44" type="paratype">paratype</typeStatus>
. Foreleg
</emphasis>
: femur
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, tibia
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, tarsomere I
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, II
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, III
<quantity id="EF59FB97FFE55319FE9BFF01A881338D" box="[300,402,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="44" unit="mm" value="0.13">0.13 mm</quantity>
, IV
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, V
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, tibial spur
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.
<emphasis id="1AD58A60FFE55319FC1BFF06AD56338D" bold="true" box="[940,1093,187,212]" pageId="4" pageNumber="44">Middle leg: f</emphasis>
emur
<quantity id="EF59FB97FFE55319FB31FF01ADFF338D" box="[1158,1260,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="4" pageNumber="44" unit="mm" value="1.17">1.17 mm</quantity>
, tibia
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, tarsomere I
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, II
<quantity id="EF59FB97FFE55319FE12FF62AB1933AE" box="[421,522,223,247]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="44" unit="mm" value="0.13">0.13 mm</quantity>
, III
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, IV
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, V
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, tibial spur
<quantity id="EF59FB97FFE55319FB89FF62ADB733AE" box="[1086,1188,223,247]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="44" unit="mm" value="0.19">0.19 mm</quantity>
.
<emphasis id="1AD58A60FFE55319FB19FF62AC0033A1" bold="true" box="[1198,1299,223,248]" pageId="4" pageNumber="44">
<collectingCountry id="50B616E2FFE55319FB19FF62ADF833A1" box="[1198,1259,223,248]" name="India" pageId="4" pageNumber="44">Hind</collectingCountry>
leg
</emphasis>
: femur
<quantity id="EF59FB97FFE55319FADCFF62A9D63245" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="4" pageNumber="44" unit="mm" value="1.13">1.13 mm</quantity>
, tibia
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, tarsomere I
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, II
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, III
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, IV
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, V
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, tibial spur
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(
<figureCitation id="B09A4AF7FFE55319FF28FE9AA9F73266" box="[159,228,295,319]" captionStart="FIGURES 4 14" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[224,1358,193,1850]" captionTargetId="figure@3.[224,1358,193,1861]" captionTargetPageId="3" captionText="FIGURES 4 14. Capicua nigra. Fig. 4. Holotype with labels; Fig. 5. Face; Fig. 6. Thorax, ventral view; Fig. 7. Right labial palpus (scanning electron picture); Fig. 8. Right hind leg of male; Fig. 9. Tibial spur of hind leg (scanning electron picture); Fig. 10. Ectoprocts of male, dorsal view; Fig. 11. Left ectoproct of male, lateral view; Fig. 12. Gonarcus-paramere complexe, right lateral view; Fig. 13. Gonarcus-paramere complexe, ventral view; Fig. 14. Hypandrium internum, lateral view." httpUri="https://zenodo.org/record/278720/files/figure.png" pageId="4" pageNumber="44">Fig. 9</figureCitation>
).
</paragraph>
<caption id="7CDE06FAFFE2531EFF20F8C7ADEE34B5" httpUri="https://zenodo.org/record/278720/files/figure.png" pageId="3" pageNumber="43" targetBox="[224,1358,193,1850]" targetPageId="3">
<paragraph id="281E5672FFE2531EFF20F8C7ADEE34B5" blockId="3.[151,1437,1914,2028]" pageId="3" pageNumber="43">
<emphasis id="1AD58A60FFE2531EFF20F8C7A85B34C9" bold="true" box="[151,328,1914,1937]" pageId="3" pageNumber="43">FIGURES 414.</emphasis>
<taxonomicName id="EFA12DF1FFE2531EFEE7F8C6A8F734C9" box="[336,484,1915,1936]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="43" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="1AD58A60FFE2531EFEE7F8C6A8F734C9" box="[336,484,1915,1936]" italics="true" pageId="3" pageNumber="43">Capicua nigra</emphasis>
</taxonomicName>
. Fig. 4. Holotype with labels; Fig. 5. Face; Fig. 6. Thorax, ventral view; Fig. 7. Right labial palpus (scanning electron picture); Fig. 8. Right hind leg of male; Fig. 9. Tibial spur of hind leg (scanning electron picture); Fig. 10. Ectoprocts of male, dorsal view; Fig. 11. Left ectoproct of male, lateral view; Fig. 12. Gonarcus-paramere complexe, right lateral view; Fig. 13. Gonarcus-paramere complexe, ventral view; Fig. 14. Hypandrium internum, lateral view.
</paragraph>
</caption>
<paragraph id="281E5672FFE55319FF72FEF6A8953165" blockId="4.[151,1437,151,1004]" pageId="4" pageNumber="44">
<emphasis id="1AD58A60FFE55319FF72FEF6A804323D" bold="true" box="[197,279,331,356]" pageId="4" pageNumber="44">Wings.</emphasis>
Narrow.
<emphasis id="1AD58A60FFE55319FE35FEF6A8E0323D" bold="true" box="[386,499,331,356]" pageId="4" pageNumber="44">Forewing</emphasis>
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, hind wing
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. Vein density low (in fore wing 811 crossveins between origin of Rs and stigma). Coloration of veins uniform. C, SC, R and Rs light brown, other longitudinal veins darker. Forewing with A2 simple, A3 forked. Membrane hyaline with dark markings at base of the 4 or 5 last crossveins between A1 and hind margin of wing (sometimes absent), at the extremity of A1 and posterior branch of CuA, and a small dot before extremity of M. Variable light dark markings along some gradates.
<emphasis id="1AD58A60FFE55319FB7BFE66AC5A32AD" bold="true" box="[1228,1353,475,500]" pageId="4" pageNumber="44">
<collectingCountry id="50B616E2FFE55319FB7BFE66AC1A32AD" box="[1228,1289,475,500]" name="India" pageId="4" pageNumber="44">Hind</collectingCountry>
wing
</emphasis>
. Membrane hyaline with a smoky area in the apical fourth along the posterior margin. Membrane of both wings sometimes slightly smoky.
</paragraph>
<paragraph id="281E5672FFE55319FF72FDFAAD4B3195" blockId="4.[151,1437,151,1004]" pageId="4" pageNumber="44">
<emphasis id="1AD58A60FFE55319FF72FDFAA8523139" bold="true" box="[197,321,583,608]" pageId="4" pageNumber="44">Abdomen.</emphasis>
Black. Postventral lobe of ectoprocts of male yellowish with 4 or 5 erect stout black setae located in the apical two-thirds, increasing in length towards apex (
<figureCitation id="B09A4AF7FFE55319FCA3FDD1AA8631DD" box="[788,917,620,644]" captionStart="FIGURES 4 14" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[224,1358,193,1850]" captionTargetId="figure@3.[224,1358,193,1861]" captionTargetPageId="3" captionText="FIGURES 4 14. Capicua nigra. Fig. 4. Holotype with labels; Fig. 5. Face; Fig. 6. Thorax, ventral view; Fig. 7. Right labial palpus (scanning electron picture); Fig. 8. Right hind leg of male; Fig. 9. Tibial spur of hind leg (scanning electron picture); Fig. 10. Ectoprocts of male, dorsal view; Fig. 11. Left ectoproct of male, lateral view; Fig. 12. Gonarcus-paramere complexe, right lateral view; Fig. 13. Gonarcus-paramere complexe, ventral view; Fig. 14. Hypandrium internum, lateral view." httpUri="https://zenodo.org/record/278720/files/figure.png" pageId="4" pageNumber="44">Figs 10, 11</figureCitation>
). Subgenital plate rounded at apex. Parameres hook-shaped with one long seta at base (one individual with two on one side) (
<figureCitation id="B09A4AF7FFE55319FBBCFD32AD9E31FE" box="[1035,1165,655,679]" captionStart="FIGURES 4 14" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[224,1358,193,1850]" captionTargetId="figure@3.[224,1358,193,1861]" captionTargetPageId="3" captionText="FIGURES 4 14. Capicua nigra. Fig. 4. Holotype with labels; Fig. 5. Face; Fig. 6. Thorax, ventral view; Fig. 7. Right labial palpus (scanning electron picture); Fig. 8. Right hind leg of male; Fig. 9. Tibial spur of hind leg (scanning electron picture); Fig. 10. Ectoprocts of male, dorsal view; Fig. 11. Left ectoproct of male, lateral view; Fig. 12. Gonarcus-paramere complexe, right lateral view; Fig. 13. Gonarcus-paramere complexe, ventral view; Fig. 14. Hypandrium internum, lateral view." httpUri="https://zenodo.org/record/278720/files/figure.png" pageId="4" pageNumber="44">Figs 12, 13</figureCitation>
). Hypandrium internum rounded (
<figureCitation id="B09A4AF7FFE55319FEB5FD09A8463195" box="[258,341,692,716]" captionStart="FIGURES 4 14" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[224,1358,193,1850]" captionTargetId="figure@3.[224,1358,193,1861]" captionTargetPageId="3" captionText="FIGURES 4 14. Capicua nigra. Fig. 4. Holotype with labels; Fig. 5. Face; Fig. 6. Thorax, ventral view; Fig. 7. Right labial palpus (scanning electron picture); Fig. 8. Right hind leg of male; Fig. 9. Tibial spur of hind leg (scanning electron picture); Fig. 10. Ectoprocts of male, dorsal view; Fig. 11. Left ectoproct of male, lateral view; Fig. 12. Gonarcus-paramere complexe, right lateral view; Fig. 13. Gonarcus-paramere complexe, ventral view; Fig. 14. Hypandrium internum, lateral view." httpUri="https://zenodo.org/record/278720/files/figure.png" pageId="4" pageNumber="44">Fig. 14</figureCitation>
). Female ectoprocts and lateral gonapophyses with stout black setae.
</paragraph>
</subSubSection>
<subSubSection id="60BB05F9FFE55319FF70FD65AA34309E" pageId="4" pageNumber="44" type="materials_examined">
<paragraph id="281E5672FFE55319FF70FD65AA34309E" blockId="4.[151,1437,151,1004]" pageId="4" pageNumber="44">
<emphasis id="1AD58A60FFE55319FF70FD65A8EA31A9" bold="true" box="[199,505,727,752]" pageId="4" pageNumber="44">
<typeStatus id="F71AE8D0FFE55319FF70FD65A81131A9" box="[199,258,728,752]" pageId="4" pageNumber="44">Type</typeStatus>
material.
<typeStatus id="F71AE8D0FFE55319FE3BFD65A8EA31A9" box="[396,505,728,752]" pageId="4" pageNumber="44" type="holotype">Holotype</typeStatus>
</emphasis>
♀,
<collectingCountry id="50B616E2FFE55319FD83FD65AB9331A9" box="[564,640,728,752]" name="Serbia" pageId="4" pageNumber="44">MALI</collectingCountry>
: Natiékoura (
<geoCoordinate id="4D9530B5FFE55319FC90FD6AAA9B31B6" box="[807,904,727,751]" direction="north" orientation="latitude" pageId="4" pageNumber="44" precision="925" value="11.333333">11°20N</geoCoordinate>
<geoCoordinate id="4D9530B5FFE55319FC20FD6AAAEC31B6" box="[919,1023,727,751]" direction="west" orientation="longitude" pageId="4" pageNumber="44" precision="925" value="-5.85">05°51W</geoCoordinate>
)
<date id="5C1F70B2FFE55319FBA1FD6AAD8131B6" box="[1046,1170,727,751]" pageId="4" pageNumber="44" value="1998-03-04">04.iii.1998</date>
, around
<specimenCount id="3EA79DFBFFE55319FAB4FD6AAC2731B6" box="[1283,1332,727,751]" pageId="4" pageNumber="44" type="generic">5 p.</specimenCount>
m. (coll.
<collectionCode id="4EB0CEB7FFE55319FF20FD41A9E0304D" box="[151,243,764,788]" httpUri="http://grbio.org/cool/5ikt-qrvp" name="Centre de Cooperation Internationale en Recherche Agronomique pour le Developpement" pageId="4" pageNumber="44">CIRAD</collectionCode>
).
<emphasis id="1AD58A60FFE55319FEBFFD41A894304D" bold="true" box="[264,391,764,788]" pageId="4" pageNumber="44">
<typeStatus id="F71AE8D0FFE55319FEBFFD41A891304D" box="[264,386,764,788]" pageId="4" pageNumber="44" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="50B616E2FFE55319FE25FD41A8CD304D" box="[402,478,764,788]" name="Serbia" pageId="4" pageNumber="44">MALI</collectingCountry>
: Natiékoura 13
<date id="5C1F70B2FFE55319FD28FD41AA02304D" box="[671,785,764,788]" pageId="4" pageNumber="44" value="1998-02-02">02.ii.1998</date>
between 4
<specimenCount id="3EA79DFBFFE55319FC17FD41AABF304D" box="[928,940,764,788]" pageId="4" pageNumber="44" type="generic">5</specimenCount>
p.m.;13
<date id="5C1F70B2FFE55319FBA9FD41AD80304D" box="[1054,1171,764,788]" pageId="4" pageNumber="44" value="1997-02-22">22.ii.1997</date>
; 13
<date id="5C1F70B2FFE55319FB78FD41AC50304D" box="[1231,1347,764,788]" pageId="4" pageNumber="44" value="1997-02-27">27.ii.1997</date>
,
<specimenCount id="3EA79DFBFFE55319FAE5FD41AC6D304D" box="[1362,1406,764,788]" pageId="4" pageNumber="44" type="generic">5 p.</specimenCount>
m. Farako (
<geoCoordinate id="4D9530B5FFE55319FF4FFCA2A84B306E" box="[248,344,799,823]" direction="north" orientation="latitude" pageId="4" pageNumber="44" precision="925" value="11.2">11°12N</geoCoordinate>
<geoCoordinate id="4D9530B5FFE55319FED4FCA2A8D8306E" box="[355,459,799,823]" direction="west" orientation="longitude" pageId="4" pageNumber="44" precision="925" value="-5.483333">05°29W</geoCoordinate>
) 1♀
<date id="5C1F70B2FFE55319FDBBFCA2AB9B306E" box="[524,648,799,823]" pageId="4" pageNumber="44" value="1997-03-04">04.iii.1997</date>
,
<specimenCount id="3EA79DFBFFE55319FD20FCA2ABD7306E" box="[663,708,799,823]" pageId="4" pageNumber="44" type="generic">9 p.</specimenCount>
m. light-trap. Around
<quantity id="EF59FB97FFE55319FC70FCA2AD17306E" box="[967,1028,799,823]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="44" unit="km" value="8.0">8 km</quantity>
from Sikasso (
<geoCoordinate id="4D9530B5FFE55319FB0DFCA2AC0A306E" box="[1210,1305,799,823]" direction="north" orientation="latitude" pageId="4" pageNumber="44" precision="925" value="11.35">11°21N</geoCoordinate>
<geoCoordinate id="4D9530B5FFE55319FA93FCA2AC9D306E" box="[1316,1422,799,823]" direction="west" orientation="longitude" pageId="4" pageNumber="44" precision="925" value="-5.6833334">05°41W</geoCoordinate>
), savanna along the road to Farako 1♀
<date id="5C1F70B2FFE55319FD8EFCF9ABA73005" box="[569,692,836,860]" pageId="4" pageNumber="44" value="1998-03-14">14.iii.1998</date>
, 11 a.m. Zégoua (
<geoCoordinate id="4D9530B5FFE55319FCCBFCF9AACE3005" box="[892,989,836,860]" direction="north" orientation="latitude" pageId="4" pageNumber="44" precision="925" value="10.5">10°30N</geoCoordinate>
<geoCoordinate id="4D9530B5FFE55319FC53FCF9AD583005" box="[996,1099,836,860]" direction="west" orientation="longitude" pageId="4" pageNumber="44" precision="925" value="-5.65">05°39W</geoCoordinate>
) 1♀
<date id="5C1F70B2FFE55319FB32FCF9ADEA3005" box="[1157,1273,836,860]" pageId="4" pageNumber="44" value="1998-02-23">23.ii.1998</date>
, at the frontier post in the morning on a wall under a lamp. All collected by B. Michel.
<emphasis id="1AD58A60FFE55319FC7BFCDAADAC30D9" bold="true" box="[972,1215,871,896]" pageId="4" pageNumber="44">Additional material.</emphasis>
<collectingCountry id="50B616E2FFE55319FB70FCD5AC8A3026" box="[1223,1433,871,896]" name="Burkina Faso" pageId="4" pageNumber="44">BURKINA FASO</collectingCountry>
: Forest of the Mou (
<geoCoordinate id="4D9530B5FFE55319FEDAFC31A8DE30FD" box="[365,461,908,932]" direction="north" orientation="latitude" pageId="4" pageNumber="44" precision="925" value="11.1">11°06N</geoCoordinate>
<geoCoordinate id="4D9530B5FFE55319FE63FC31AB2D30FD" box="[468,574,908,932]" direction="west" orientation="longitude" pageId="4" pageNumber="44" precision="925" value="-3.65">03°39W</geoCoordinate>
), region of Koumbia, 13
<date id="5C1F70B2FFE55319FCEBFC31AAC430FD" box="[860,983,908,932]" pageId="4" pageNumber="44" value="2010-03-12">12.iii.2010</date>
, 0 9:
<specimenCount id="3EA79DFBFFE55319FBBEFC31AD5030FD" box="[1033,1091,908,932]" pageId="4" pageNumber="44" type="generic">16 p.</specimenCount>
m. light-trap, 2♀♀
<date id="5C1F70B2FFE55319FAACFC31AC8430FD" box="[1307,1431,908,932]" pageId="4" pageNumber="44" value="2010-03-12">12.iii.2010</date>
, between 8:00 and 8:
<specimenCount id="3EA79DFBFFE55319FEC2FC12A8A3309E" box="[373,432,943,967]" pageId="4" pageNumber="44" type="generic">30 p.</specimenCount>
m. All collected by M. Akoudjin.
</paragraph>
</subSubSection>
<subSubSection id="60BB05F9FFE55319FF70FC69AA8130B5" box="[199,914,980,1004]" pageId="4" pageNumber="44" type="etymology">
<paragraph id="281E5672FFE55319FF70FC69AA8130B5" blockId="4.[151,1437,151,1004]" box="[199,914,980,1004]" pageId="4" pageNumber="44">
<emphasis id="1AD58A60FFE55319FF70FC69A85B30B5" bold="true" box="[199,328,980,1004]" pageId="4" pageNumber="44">Etymology</emphasis>
:
<taxonomicName id="EFA12DF1FFE55319FEE0FC68A88030B5" box="[343,403,981,1004]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="44" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="1AD58A60FFE55319FEE0FC68A88030B5" box="[343,403,981,1004]" italics="true" pageId="4" pageNumber="44">nigra</emphasis>
</taxonomicName>
, referring to the black coloration of the body.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,321 @@
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<mods:title id="DCD23B395427E472BE8237CE5010C77C">Reinstatement of the genus Capicua Navás with descriptions of two new species (Neuroptera, Myrmeleontidae)</mods:title>
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<mods:role id="46D4C27DF5F42D4E5BD55EE81D540E1D">
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<mods:namePart id="7979122112DE2337EFBDA6A2FB53C9C1">Michel, Bruno</mods:namePart>
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<treatment id="A008E764FFE5531BFF20FB82AB7F30B5" ID-DOI="http://doi.org/10.5281/zenodo.6184728" ID-GBIF-Taxon="119416761" ID-Zenodo-Dep="6184728" LSID="urn:lsid:plazi:treatment:A008E764FFE5531BFF20FB82AB7F30B5" httpUri="http://treatment.plazi.org/id/A008E764FFE5531BFF20FB82AB7F30B5" lastPageId="6" lastPageNumber="46" pageId="4" pageNumber="44">
<subSubSection id="60BB05F9FFE55319FF20FB82A83D3723" pageId="4" pageNumber="44" type="nomenclature">
<paragraph id="281E5672FFE55319FF20FB82A8F53700" blockId="4.[151,486,1087,1146]" box="[151,486,1087,1113]" pageId="4" pageNumber="44">
<heading id="7356E11EFFE55319FF20FB82A8F53700" bold="true" box="[151,486,1087,1113]" fontSize="11" level="1" pageId="4" pageNumber="44" reason="1">
<emphasis id="1AD58A60FFE55319FF20FB82A8F53700" bold="true" box="[151,486,1087,1113]" pageId="4" pageNumber="44">
<taxonomicName id="EFA12DF1FFE55319FF20FB82A8923700" ID-CoL="5WXJP" box="[151,385,1087,1113]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="44" phylum="Arthropoda" rank="species" species="acalcarata" status="sp. nov.">
<emphasis id="1AD58A60FFE55319FF20FB82A8923700" bold="true" box="[151,385,1087,1113]" italics="true" pageId="4" pageNumber="44">Capicua acalcarata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="01E6371BFFE55319FE3EFB82A8F53700" box="[393,486,1087,1113]" pageId="4" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="281E5672FFE55319FF20FBDFA83D3723" blockId="4.[151,486,1087,1146]" box="[151,302,1122,1146]" pageId="4" pageNumber="44">
(
<figureCitation id="B09A4AF7FFE55319FF28FBDFA8353723" box="[159,294,1122,1146]" captionStart="FIGURES 15 25" captionStartId="5.[151,264,1892,1915]" captionTargetBox="[219,1369,193,1871]" captionTargetId="figure@5.[219,1371,193,1885]" captionTargetPageId="5" captionText="FIGURES 15 25. Capicua acalcarata. Fig. 15. Holotype with labels; Fig. 16. Face; Fig. 17. Thorax, ventral view; Fig. 18. Right labial palpus (scanning electron picture); Fig. 19. Extremity of tibia and first tarsomere of right hind leg (scanning electron picture); Fig. 20. Ectoprocts of male, dorsal view; Fig. 21. Left ectoproct of male, lateral view; Fig. 22. Gonarcus-paramere complexe, right lateral view; Fig. 23. Gonarcus-paramere complexe, ventral view; Fig. 24. Hypandrium internum, lateral view; Fig. 25. Female terminalia, left lateral view." httpUri="https://zenodo.org/record/278721/files/figure.png" pageId="4" pageNumber="44">Figs. 1525</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="60BB05F9FFE55319FF20FB17A8613677" pageId="4" pageNumber="44" type="diagnosis">
<paragraph id="281E5672FFE55319FF20FB17A8613677" blockId="4.[151,1437,1194,2010]" pageId="4" pageNumber="44">
<emphasis id="1AD58A60FFE55319FF20FB17A802379A" bold="true" box="[151,273,1194,1219]" pageId="4" pageNumber="44">Diagnosis.</emphasis>
General body coloration dark brown (
<figureCitation id="B09A4AF7FFE55319FD77FB17AA00379B" box="[704,787,1194,1218]" captionStart="FIGURES 15 25" captionStartId="5.[151,264,1892,1915]" captionTargetBox="[219,1369,193,1871]" captionTargetId="figure@5.[219,1371,193,1885]" captionTargetPageId="5" captionText="FIGURES 15 25. Capicua acalcarata. Fig. 15. Holotype with labels; Fig. 16. Face; Fig. 17. Thorax, ventral view; Fig. 18. Right labial palpus (scanning electron picture); Fig. 19. Extremity of tibia and first tarsomere of right hind leg (scanning electron picture); Fig. 20. Ectoprocts of male, dorsal view; Fig. 21. Left ectoproct of male, lateral view; Fig. 22. Gonarcus-paramere complexe, right lateral view; Fig. 23. Gonarcus-paramere complexe, ventral view; Fig. 24. Hypandrium internum, lateral view; Fig. 25. Female terminalia, left lateral view." httpUri="https://zenodo.org/record/278721/files/figure.png" pageId="4" pageNumber="44">Fig. 15</figureCitation>
). Base of legs yellow contrasting with the dark coloration of the thoracic sternites (
<figureCitation id="B09A4AF7FFE55319FE18FB73AB1037BF" box="[431,515,1230,1254]" captionStart="FIGURES 15 25" captionStartId="5.[151,264,1892,1915]" captionTargetBox="[219,1369,193,1871]" captionTargetId="figure@5.[219,1371,193,1885]" captionTargetPageId="5" captionText="FIGURES 15 25. Capicua acalcarata. Fig. 15. Holotype with labels; Fig. 16. Face; Fig. 17. Thorax, ventral view; Fig. 18. Right labial palpus (scanning electron picture); Fig. 19. Extremity of tibia and first tarsomere of right hind leg (scanning electron picture); Fig. 20. Ectoprocts of male, dorsal view; Fig. 21. Left ectoproct of male, lateral view; Fig. 22. Gonarcus-paramere complexe, right lateral view; Fig. 23. Gonarcus-paramere complexe, ventral view; Fig. 24. Hypandrium internum, lateral view; Fig. 25. Female terminalia, left lateral view." httpUri="https://zenodo.org/record/278721/files/figure.png" pageId="4" pageNumber="44">Fig. 17</figureCitation>
). Tibial spurs absent (
<figureCitation id="B09A4AF7FFE55319FD4AFB73AA4237BF" box="[765,849,1230,1254]" captionStart="FIGURES 15 25" captionStartId="5.[151,264,1892,1915]" captionTargetBox="[219,1369,193,1871]" captionTargetId="figure@5.[219,1371,193,1885]" captionTargetPageId="5" captionText="FIGURES 15 25. Capicua acalcarata. Fig. 15. Holotype with labels; Fig. 16. Face; Fig. 17. Thorax, ventral view; Fig. 18. Right labial palpus (scanning electron picture); Fig. 19. Extremity of tibia and first tarsomere of right hind leg (scanning electron picture); Fig. 20. Ectoprocts of male, dorsal view; Fig. 21. Left ectoproct of male, lateral view; Fig. 22. Gonarcus-paramere complexe, right lateral view; Fig. 23. Gonarcus-paramere complexe, ventral view; Fig. 24. Hypandrium internum, lateral view; Fig. 25. Female terminalia, left lateral view." httpUri="https://zenodo.org/record/278721/files/figure.png" pageId="4" pageNumber="44">Fig. 19</figureCitation>
). No tuft of long white setae on fore femora. Wings narrow. Postventral lobe of male ectoprocts well developed, yellowish with four erect stout black setae inserted in the distal one-third.
</paragraph>
</subSubSection>
<subSubSection id="60BB05F9FFE5531BFF72FA87A8D332DE" lastPageId="6" lastPageNumber="46" pageId="4" pageNumber="44" type="description">
<paragraph id="281E5672FFE55319FF72FA87AC7F36E7" blockId="4.[151,1437,1194,2010]" pageId="4" pageNumber="44">
<emphasis id="1AD58A60FFE55319FF72FA87A8B0360A" bold="true" box="[197,419,1338,1363]" pageId="4" pageNumber="44">Description. Head.</emphasis>
Vertex raised. Face yellow with a black chevron-shaped mark under the scapes, grey above without yellow transverse line between bases of antennae (
<figureCitation id="B09A4AF7FFE55319FC83FAE3AA9A362F" box="[820,905,1374,1398]" captionStart="FIGURES 15 25" captionStartId="5.[151,264,1892,1915]" captionTargetBox="[219,1369,193,1871]" captionTargetId="figure@5.[219,1371,193,1885]" captionTargetPageId="5" captionText="FIGURES 15 25. Capicua acalcarata. Fig. 15. Holotype with labels; Fig. 16. Face; Fig. 17. Thorax, ventral view; Fig. 18. Right labial palpus (scanning electron picture); Fig. 19. Extremity of tibia and first tarsomere of right hind leg (scanning electron picture); Fig. 20. Ectoprocts of male, dorsal view; Fig. 21. Left ectoproct of male, lateral view; Fig. 22. Gonarcus-paramere complexe, right lateral view; Fig. 23. Gonarcus-paramere complexe, ventral view; Fig. 24. Hypandrium internum, lateral view; Fig. 25. Female terminalia, left lateral view." httpUri="https://zenodo.org/record/278721/files/figure.png" pageId="4" pageNumber="44">Fig. 16</figureCitation>
). Vertex grey to dark grey delimited anteriorly by a black line more or less interrupted. Occiput grey with black and yellowish marking variably contrasted. Antennae as in
<taxonomicName id="EFA12DF1FFE55319FEF3FA1AA8B136E7" box="[324,418,1447,1470]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="44" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="1AD58A60FFE55319FEF3FA1AA8B136E7" box="[324,418,1447,1470]" italics="true" pageId="4" pageNumber="44">C. nigra</emphasis>
</taxonomicName>
. Palpi yellow with the last segment of labial palp darker and clearly enlarged (
<figureCitation id="B09A4AF7FFE55319FAA4FA1BAC4C36E7" box="[1299,1375,1446,1470]" captionStart="FIGURES 15 25" captionStartId="5.[151,264,1892,1915]" captionTargetBox="[219,1369,193,1871]" captionTargetId="figure@5.[219,1371,193,1885]" captionTargetPageId="5" captionText="FIGURES 15 25. Capicua acalcarata. Fig. 15. Holotype with labels; Fig. 16. Face; Fig. 17. Thorax, ventral view; Fig. 18. Right labial palpus (scanning electron picture); Fig. 19. Extremity of tibia and first tarsomere of right hind leg (scanning electron picture); Fig. 20. Ectoprocts of male, dorsal view; Fig. 21. Left ectoproct of male, lateral view; Fig. 22. Gonarcus-paramere complexe, right lateral view; Fig. 23. Gonarcus-paramere complexe, ventral view; Fig. 24. Hypandrium internum, lateral view; Fig. 25. Female terminalia, left lateral view." httpUri="https://zenodo.org/record/278721/files/figure.png" pageId="4" pageNumber="44">Fig 18</figureCitation>
).
</paragraph>
<paragraph id="281E5672FFE55319FF72FA76A8143573" blockId="4.[151,1437,1194,2010]" pageId="4" pageNumber="44">
<emphasis id="1AD58A60FFE55319FF72FA76A83736BA" bold="true" box="[197,292,1483,1507]" pageId="4" pageNumber="44">Thorax.</emphasis>
Pronotum wider than long. Coloration grey and dark grey forming longitudinal lines. 45 long white lateral setae forwardly curved. Mesothorax and metathorax grey to dark grey with variably distinct longitudinal markings.
</paragraph>
<paragraph id="281E5672FFE55319FF72F98AA9D934CB" blockId="4.[151,1437,1194,2010]" pageId="4" pageNumber="44">
<emphasis id="1AD58A60FFE55319FF72F98AA87F3516" bold="true" box="[197,364,1591,1615]" pageId="4" pageNumber="44">Legs. Foreleg.</emphasis>
Coxa dark brown, yellowish ventrally with 23 long white setae. Trochanter yellowish. Femur dark brown, yellowish ventrally without long white setae. Tibia brown, yellowish dorsally with a dark transverse line at basal third. Tarsomeres yellowish, dark apically.
<emphasis id="1AD58A60FFE55319FC97F9C3AAB235CE" bold="true" box="[800,929,1662,1687]" pageId="4" pageNumber="44">Middle leg</emphasis>
. Coxa brown. Trochanter yellowish. Femur entirely brown. Tibia brown with two large yellow rings. Tarsomeres yellowish, dark apically.
<emphasis id="1AD58A60FFE55319FB51F91FAC5C35E2" bold="true" box="[1254,1359,1698,1723]" pageId="4" pageNumber="44">
<collectingCountry id="50B616E2FFE55319FB51F91FAC3035E2" box="[1254,1315,1698,1723]" name="India" pageId="4" pageNumber="44">Hind</collectingCountry>
leg
</emphasis>
. Coxa brown. Trochanter yellowish. Femur brown. Tibia pale, darker ventrally, with a transversal dark line at basal third. In male a tuft of long erected setae on dorsal face. Tarsomeres yellowish, dark apically.
<emphasis id="1AD58A60FFE55319FBCBF957A9E2347E" bold="true" pageId="4" pageNumber="44">
Dimensions of
<typeStatus id="F71AE8D0FFE55319FA99F956AC84345A" box="[1326,1431,1771,1795]" pageId="4" pageNumber="44" type="holotype">holotype</typeStatus>
. Foreleg
</emphasis>
: femur
<quantity id="EF59FB97FFE55319FEFEF8B3A8BC347F" box="[329,431,1806,1830]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="4" pageNumber="44" unit="mm" value="0.82">0.82 mm</quantity>
, tibia
<quantity id="EF59FB97FFE55319FE43F8B3AB49347F" box="[500,602,1806,1830]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="4" pageNumber="44" unit="mm" value="0.79">0.79 mm</quantity>
, tarsomere I
<quantity id="EF59FB97FFE55319FD5BF8B3AA41347F" box="[748,850,1806,1830]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="44" unit="mm" value="0.1">0.10 mm</quantity>
, II
<quantity id="EF59FB97FFE55319FCCFF8B3AACD347F" box="[888,990,1806,1830]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="44" unit="mm" value="0.1">0.10 mm</quantity>
, III
<quantity id="EF59FB97FFE55319FBBAF8B3AD60347F" box="[1037,1139,1806,1830]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="44" unit="mm" value="0.1">0.10 mm</quantity>
, IV
<quantity id="EF59FB97FFE55319FB13F8B3AC19347F" box="[1188,1290,1806,1830]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="44" unit="mm" value="0.1">0.10 mm</quantity>
, V.0
<quantity id="EF59FB97FFE55319FAF3F8B3AC8B347F" box="[1348,1432,1806,1830]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="44" unit="mm" value="38.0">38 mm</quantity>
.
<emphasis id="1AD58A60FFE55319FF20F88FA8053412" bold="true" box="[151,278,1842,1867]" pageId="4" pageNumber="44">Middle leg</emphasis>
: femur
<quantity id="EF59FB97FFE55319FEC5F88FA8CB3413" box="[370,472,1842,1866]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="4" pageNumber="44" unit="mm" value="0.82">0.82 mm</quantity>
, tibia
<quantity id="EF59FB97FFE55319FDA8F88FAB953413" box="[543,646,1842,1866]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="4" pageNumber="44" unit="mm" value="0.86">0.86 mm</quantity>
, tarsomere I
<quantity id="EF59FB97FFE55319FCACF88FAA913413" box="[795,898,1842,1866]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="44" unit="mm" value="0.16">0.16 mm</quantity>
, II
<quantity id="EF59FB97FFE55319FC1DF88FAD023413" box="[938,1041,1842,1866]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="44" unit="mm" value="0.08">0.08 mm</quantity>
, III
<quantity id="EF59FB97FFE55319FBF6F88FADBA3413" box="[1089,1193,1842,1866]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="44" unit="mm" value="0.08">0.08 mm</quantity>
, IV
<quantity id="EF59FB97FFE55319FB6CF88FAC513413" box="[1243,1346,1842,1866]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="44" unit="mm" value="0.08">0.08 mm</quantity>
, V
<quantity id="EF59FB97FFE55319FADCF88FA9D53437" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="44" unit="mm" value="0.38">0.38 mm</quantity>
.
<emphasis id="1AD58A60FFE55319FF67F8EBA8253436" bold="true" box="[208,310,1878,1903]" pageId="4" pageNumber="44">
<collectingCountry id="50B616E2FFE55319FF67F8EBA81E3436" box="[208,269,1878,1903]" name="India" pageId="4" pageNumber="44">Hind</collectingCountry>
leg
</emphasis>
: femur
<quantity id="EF59FB97FFE55319FE39F8EBA8E13437" box="[398,498,1878,1902]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="4" pageNumber="44" unit="mm" value="1.11">1.11 mm</quantity>
, tibia
<quantity id="EF59FB97FFE55319FD81F8EBAB883437" box="[566,667,1878,1902]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" pageId="4" pageNumber="44" unit="mm" value="1.24">1.24 mm</quantity>
, tarsomere I
<quantity id="EF59FB97FFE55319FC9BF8EBAA813437" box="[812,914,1878,1902]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="44" unit="mm" value="0.22">0.22 mm</quantity>
, II
<quantity id="EF59FB97FFE55319FC01F8EBAD0F3437" box="[950,1052,1878,1902]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="44" unit="mm" value="0.13">0.13 mm</quantity>
, III
<quantity id="EF59FB97FFE55319FBFDF8EBADBC3437" box="[1098,1199,1878,1902]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="44" unit="mm" value="0.1">0.10 mm</quantity>
, IV
<quantity id="EF59FB97FFE55319FB68F8EBAC573437" box="[1247,1348,1878,1902]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="44" unit="mm" value="0.1">0.10 mm</quantity>
, V
<quantity id="EF59FB97FFE55319FADDF8EBA9D534CB" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="44" unit="mm" value="0.38">0.38 mm</quantity>
.
</paragraph>
<paragraph id="281E5672FFE5531BFF72F823AC3C338D" blockId="4.[151,1437,1194,2010]" lastBlockId="6.[151,1437,151,1004]" lastPageId="6" lastPageNumber="46" pageId="4" pageNumber="44">
<emphasis id="1AD58A60FFE55319FF72F823A80434EE" bold="true" box="[197,279,1950,1975]" pageId="4" pageNumber="44">Wings.</emphasis>
Narrow.
<emphasis id="1AD58A60FFE55319FE3CF823A8EF34EE" bold="true" box="[395,508,1950,1975]" pageId="4" pageNumber="44">Forewing</emphasis>
<quantity id="EF59FB97FFE55319FDBFF823AB9A34EF" box="[520,649,1950,1974]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="4" pageNumber="44" unit="mm" value="17.0" valueMax="19.0" valueMin="15.0">1519 mm</quantity>
, hind wing
<quantity id="EF59FB97FFE55319FCABF823AA8D34EF" box="[796,926,1950,1974]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="4" pageNumber="44" unit="mm" value="16.0" valueMax="18.0" valueMin="14.0">1418 mm</quantity>
. Vein density low (generally 10 crossveins between origin of RS and pterostigma). Veins light brown. A2 simple, A3 forked. Membrane slightly smoky. Dark markings of variable extension at apex of crossveins between A1 and hind margin of wing and along all or only few gradates.
<emphasis id="1AD58A60FFE7531BFE84FF06A8A4338D" bold="true" box="[307,439,187,212]" pageId="6" pageNumber="46">
<collectingCountry id="50B616E2FFE7531BFE84FF06A863338D" box="[307,368,187,212]" name="India" pageId="6" pageNumber="46">Hind</collectingCountry>
wing.
</emphasis>
No marking. Membrane slightly smoky in the apical fourth along hind margin.
</paragraph>
<caption id="7CDE06FAFFE45318FF20F8D9AB4734AC" httpUri="https://zenodo.org/record/278721/files/figure.png" pageId="5" pageNumber="45" targetBox="[219,1369,193,1871]" targetPageId="5">
<paragraph id="281E5672FFE45318FF20F8D9AB4734AC" blockId="5.[151,1436,1892,2037]" pageId="5" pageNumber="45">
<emphasis id="1AD58A60FFE45318FF20F8D9A8463420" bold="true" box="[151,341,1892,1915]" pageId="5" pageNumber="45">FIGURES 1525.</emphasis>
<taxonomicName id="EFA12DF1FFE45318FEEAF8D8AB3A3423" box="[349,553,1892,1914]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="45" phylum="Arthropoda" rank="species" species="acalcarata">
<emphasis id="1AD58A60FFE45318FEEAF8D8AB3A3423" box="[349,553,1892,1914]" italics="true" pageId="5" pageNumber="45">Capicua acalcarata</emphasis>
</taxonomicName>
. Fig. 15. Holotype with labels; Fig. 16. Face; Fig. 17. Thorax, ventral view; Fig. 18. Right labial palpus (scanning electron picture); Fig. 19. Extremity of tibia and first tarsomere of right hind leg (scanning electron picture); Fig. 20. Ectoprocts of male, dorsal view; Fig. 21. Left ectoproct of male, lateral view; Fig. 22. Gonarcus-paramere complexe, right lateral view; Fig. 23. Gonarcus-paramere complexe, ventral view; Fig. 24. Hypandrium internum, lateral view; Fig. 25. Female terminalia, left lateral view.
</paragraph>
</caption>
<paragraph id="281E5672FFE7531BFF72FF62A8D332DE" blockId="6.[151,1437,151,1004]" pageId="6" pageNumber="46">
<emphasis id="1AD58A60FFE7531BFF72FF62A85233A1" bold="true" box="[197,321,223,248]" pageId="6" pageNumber="46">Abdomen.</emphasis>
Dark brown. Male ectoprocts with a vertical yellow line on the basal part. Postventral lobe light brown to yellowish with 34 erect stout black setae generally inserted in the distal third (
<figureCitation id="B09A4AF7FFE7531BFB25FEB9AC043245" box="[1170,1303,260,284]" captionStart="FIGURES 15 25" captionStartId="5.[151,264,1892,1915]" captionTargetBox="[219,1369,193,1871]" captionTargetId="figure@5.[219,1371,193,1885]" captionTargetPageId="5" captionText="FIGURES 15 25. Capicua acalcarata. Fig. 15. Holotype with labels; Fig. 16. Face; Fig. 17. Thorax, ventral view; Fig. 18. Right labial palpus (scanning electron picture); Fig. 19. Extremity of tibia and first tarsomere of right hind leg (scanning electron picture); Fig. 20. Ectoprocts of male, dorsal view; Fig. 21. Left ectoproct of male, lateral view; Fig. 22. Gonarcus-paramere complexe, right lateral view; Fig. 23. Gonarcus-paramere complexe, ventral view; Fig. 24. Hypandrium internum, lateral view; Fig. 25. Female terminalia, left lateral view." httpUri="https://zenodo.org/record/278721/files/figure.png" pageId="6" pageNumber="46">Figs 20, 21</figureCitation>
). Length of these setae increasing towards apex. Subgenital plate rounded at apex. Parameres hook-shaped with one long seta at base (
<figureCitation id="B09A4AF7FFE7531BFF41FEF1A86F323D" box="[246,380,332,356]" captionStart="FIGURES 15 25" captionStartId="5.[151,264,1892,1915]" captionTargetBox="[219,1369,193,1871]" captionTargetId="figure@5.[219,1371,193,1885]" captionTargetPageId="5" captionText="FIGURES 15 25. Capicua acalcarata. Fig. 15. Holotype with labels; Fig. 16. Face; Fig. 17. Thorax, ventral view; Fig. 18. Right labial palpus (scanning electron picture); Fig. 19. Extremity of tibia and first tarsomere of right hind leg (scanning electron picture); Fig. 20. Ectoprocts of male, dorsal view; Fig. 21. Left ectoproct of male, lateral view; Fig. 22. Gonarcus-paramere complexe, right lateral view; Fig. 23. Gonarcus-paramere complexe, ventral view; Fig. 24. Hypandrium internum, lateral view; Fig. 25. Female terminalia, left lateral view." httpUri="https://zenodo.org/record/278721/files/figure.png" pageId="6" pageNumber="46">Figs 22, 23</figureCitation>
). Hypandrium internum rounded (
<figureCitation id="B09A4AF7FFE7531BFCB6FEF1AA44323D" box="[769,855,332,356]" captionStart="FIGURES 15 25" captionStartId="5.[151,264,1892,1915]" captionTargetBox="[219,1369,193,1871]" captionTargetId="figure@5.[219,1371,193,1885]" captionTargetPageId="5" captionText="FIGURES 15 25. Capicua acalcarata. Fig. 15. Holotype with labels; Fig. 16. Face; Fig. 17. Thorax, ventral view; Fig. 18. Right labial palpus (scanning electron picture); Fig. 19. Extremity of tibia and first tarsomere of right hind leg (scanning electron picture); Fig. 20. Ectoprocts of male, dorsal view; Fig. 21. Left ectoproct of male, lateral view; Fig. 22. Gonarcus-paramere complexe, right lateral view; Fig. 23. Gonarcus-paramere complexe, ventral view; Fig. 24. Hypandrium internum, lateral view; Fig. 25. Female terminalia, left lateral view." httpUri="https://zenodo.org/record/278721/files/figure.png" pageId="6" pageNumber="46">Fig. 24</figureCitation>
). Female ectoprocts and lateral gonapophyses with stout black setae (
<figureCitation id="B09A4AF7FFE7531BFEE8FED2A8A732DE" box="[351,436,367,391]" captionStart="FIGURES 15 25" captionStartId="5.[151,264,1892,1915]" captionTargetBox="[219,1369,193,1871]" captionTargetId="figure@5.[219,1371,193,1885]" captionTargetPageId="5" captionText="FIGURES 15 25. Capicua acalcarata. Fig. 15. Holotype with labels; Fig. 16. Face; Fig. 17. Thorax, ventral view; Fig. 18. Right labial palpus (scanning electron picture); Fig. 19. Extremity of tibia and first tarsomere of right hind leg (scanning electron picture); Fig. 20. Ectoprocts of male, dorsal view; Fig. 21. Left ectoproct of male, lateral view; Fig. 22. Gonarcus-paramere complexe, right lateral view; Fig. 23. Gonarcus-paramere complexe, ventral view; Fig. 24. Hypandrium internum, lateral view; Fig. 25. Female terminalia, left lateral view." httpUri="https://zenodo.org/record/278721/files/figure.png" pageId="6" pageNumber="46">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="60BB05F9FFE7531BFF72FE29A81A31FE" pageId="6" pageNumber="46" type="materials_examined">
<paragraph id="281E5672FFE7531BFF72FE29A81A31FE" blockId="6.[151,1437,151,1004]" pageId="6" pageNumber="46">
<emphasis id="1AD58A60FFE7531BFF72FE29A8F532F5" bold="true" box="[197,486,403,428]" pageId="6" pageNumber="46">
<typeStatus id="F71AE8D0FFE7531BFF72FE29A81332F5" box="[197,256,404,428]" pageId="6" pageNumber="46">Type</typeStatus>
material.
<typeStatus id="F71AE8D0FFE7531BFECEFE29A8F532F5" box="[377,486,404,428]" pageId="6" pageNumber="46" type="holotype">Holotype</typeStatus>
</emphasis>
3,
<collectingCountry id="50B616E2FFE7531BFDB8FE29AB4832F5" box="[527,603,404,428]" name="Serbia" pageId="6" pageNumber="46">MALI</collectingCountry>
: Natiékoura
<date id="5C1F70B2FFE7531BFD5EFE29AA4F32F5" box="[745,860,404,428]" pageId="6" pageNumber="46" value="1997-02-24">24.ii.1997</date>
, around
<specimenCount id="3EA79DFBFFE7531BFC0BFE29AAF632F5" box="[956,997,404,428]" pageId="6" pageNumber="46" type="generic">5 p.</specimenCount>
m. (coll.
<collectionCode id="4EB0CEB7FFE7531BFBFDFE29ADB532F5" box="[1098,1190,404,428]" httpUri="http://grbio.org/cool/5ikt-qrvp" name="Centre de Cooperation Internationale en Recherche Agronomique pour le Developpement" pageId="6" pageNumber="46">CIRAD</collectionCode>
).
<typeStatus id="F71AE8D0FFE7531BFB0EFE29AC2232F5" box="[1209,1329,404,428]" pageId="6" pageNumber="46" type="paratype">
<emphasis id="1AD58A60FFE7531BFB0EFE29AC2232F5" bold="true" box="[1209,1329,404,428]" pageId="6" pageNumber="46">Paratypes</emphasis>
</typeStatus>
<collectingCountry id="50B616E2FFE7531BFAF9FE29AC8B32F5" box="[1358,1432,404,428]" name="Serbia" pageId="6" pageNumber="46">MALI</collectingCountry>
: Natiékoura 13
<date id="5C1F70B2FFE7531BFEF2FE0AA8A13296" box="[325,434,439,463]" pageId="6" pageNumber="46" value="1997-01-08">08.i.1997</date>
;
<date id="5C1F70B2FFE7531BFE09FE0AAB303296" box="[446,547,439,464]" pageId="6" pageNumber="46" value="1916-03-02">2 3 16</date>
.i.1997; 13
<date id="5C1F70B2FFE7531BFD1EFE0AAA053296" box="[681,790,439,463]" pageId="6" pageNumber="46" value="1997-01-30">30.i.1997</date>
; 13 2♀♀
<date id="5C1F70B2FFE7531BFC3BFE0AAD133296" box="[908,1024,439,463]" pageId="6" pageNumber="46" value="1998-02-02">02.ii.1998</date>
, between 4
<specimenCount id="3EA79DFBFFE7531BFB38FE0AAD883296" box="[1167,1179,439,463]" pageId="6" pageNumber="46" type="generic">5</specimenCount>
p.m.; 2♀♀
<date id="5C1F70B2FFE7531BFA93FE0AAC843296" box="[1316,1431,439,463]" pageId="6" pageNumber="46" value="1997-02-11">11.ii.1997</date>
; 13
<date id="5C1F70B2FFE7531BFF70FE61A82F32AD" box="[199,316,476,500]" pageId="6" pageNumber="46" value="1996-02-16">16.ii.1996</date>
; 1♀
<date id="5C1F70B2FFE7531BFECBFE61A8E232AD" box="[380,497,476,500]" pageId="6" pageNumber="46" value="1997-02-22">22.ii.1997</date>
; 23 1♀
<date id="5C1F70B2FFE7531BFDC0FE61ABF832AD" box="[631,747,476,500]" pageId="6" pageNumber="46" value="1997-02-25">25.ii.1997</date>
, around
<specimenCount id="3EA79DFBFFE7531BFCE0FE61AA9432AD" box="[855,903,476,500]" pageId="6" pageNumber="46" type="generic">5 p.</specimenCount>
m.; 13 1♀
<date id="5C1F70B2FFE7531BFBAFFE61AD8632AD" box="[1048,1173,476,500]" pageId="6" pageNumber="46" value="1998-03-04">04.iii.1998</date>
; 2♀♀
<date id="5C1F70B2FFE7531BFB5CFE61AC7432AD" box="[1259,1383,476,500]" pageId="6" pageNumber="46" value="1996-03-18">18.iii.1996</date>
; 13
<date id="5C1F70B2FFE7531BFF20FE42A800314E" box="[151,275,511,535]" pageId="6" pageNumber="46" value="1998-03-23">23.iii.1998</date>
; 13
<date id="5C1F70B2FFE7531BFEFEFE42A8D0314E" box="[329,451,511,535]" pageId="6" pageNumber="46" value="1996-04-03">03.iv.1996</date>
; 1♀
<date id="5C1F70B2FFE7531BFE4FFE42AB61314E" box="[504,626,511,535]" pageId="6" pageNumber="46" value="1996-04-05">05.iv.1996</date>
; 13
<date id="5C1F70B2FFE7531BFD10FE42AA32314E" box="[679,801,511,535]" pageId="6" pageNumber="46" value="1998-04-07">07.iv.1998</date>
. Around
<quantity id="EF59FB97FFE7531BFC3FFE42AADC314E" box="[904,975,511,535]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="46" unit="km" value="11.0">11 km</quantity>
from Sikasso, savanna along the track to Longorola (
<geoCoordinate id="4D9530B5FFE7531BFE97FD99A8A53165" box="[288,438,548,572]" direction="north" orientation="latitude" pageId="6" pageNumber="46" precision="15" value="11.442223">11°2632N-</geoCoordinate>
<geoCoordinate id="4D9530B5FFE7531BFE01FD99AB5A3165" box="[438,585,548,572]" direction="west" orientation="longitude" pageId="6" pageNumber="46" precision="15" value="-5.6530557">05°3911W</geoCoordinate>
) 13
<date id="5C1F70B2FFE7531BFD3BFD99ABEB3165" box="[652,760,548,572]" pageId="6" pageNumber="46" value="1997-01-15">15.i.1997</date>
; 23 1♀
<date id="5C1F70B2FFE7531BFCCCFD99AAF43165" box="[891,999,548,572]" pageId="6" pageNumber="46" value="1996-01-24">24.i.1996</date>
; 13
<date id="5C1F70B2FFE7531BFB91FD99AD803165" box="[1062,1171,548,572]" pageId="6" pageNumber="46" value="1996-01-27">27.i.1996</date>
; 2♀♀
<date id="5C1F70B2FFE7531BFB50FD99AC403165" box="[1255,1363,548,572]" pageId="6" pageNumber="46" value="1996-01-29">29.i.1996</date>
; 23 2♀♀
<date id="5C1F70B2FFE7531BFF60FDFAA85F3106" box="[215,332,583,607]" pageId="6" pageNumber="46" value="1996-02-05">05.ii.1996</date>
; 1♀
<date id="5C1F70B2FFE7531BFE33FDFAA8EA3106" box="[388,505,583,607]" pageId="6" pageNumber="46" value="1996-02-23">23.ii.1996</date>
; 1♀
<date id="5C1F70B2FFE7531BFD86FDFAABB53106" box="[561,678,583,607]" pageId="6" pageNumber="46" value="1996-02-28">28.ii.1996</date>
; 13
<date id="5C1F70B2FFE7531BFD69FDFAAA483106" box="[734,859,583,607]" pageId="6" pageNumber="46" value="1998-03-13">13.iii.1998</date>
; 1♀
<date id="5C1F70B2FFE7531BFC24FDFAAD1E3106" box="[915,1037,583,607]" pageId="6" pageNumber="46" value="1998-04-04">04.iv.1998</date>
. All collected by B. Michel.
<emphasis id="1AD58A60FFE7531BFAE0FDFAA84031DD" bold="true" pageId="6" pageNumber="46">Additional material.</emphasis>
<collectingCountry id="50B616E2FFE7531BFEE9FDD1AB2131DD" box="[350,562,620,644]" name="Burkina Faso" pageId="6" pageNumber="46">BURKINA FASO</collectingCountry>
: Forest of the Mou 13
<date id="5C1F70B2FFE7531BFCE7FDD1AADF31DD" box="[848,972,620,644]" pageId="6" pageNumber="46" value="2010-03-10">10.iii.2010</date>
, 10:09 p.m. light-trap. Collected by M. Akoudjin.
</paragraph>
</subSubSection>
<subSubSection id="60BB05F9FFE7531BFF72FD09AA993195" box="[197,906,692,716]" pageId="6" pageNumber="46" type="etymology">
<paragraph id="281E5672FFE7531BFF72FD09AA993195" blockId="6.[151,1437,151,1004]" box="[197,906,692,716]" pageId="6" pageNumber="46">
<emphasis id="1AD58A60FFE7531BFF72FD09A8583195" bold="true" box="[197,331,692,716]" pageId="6" pageNumber="46">Etymology.</emphasis>
<taxonomicName id="EFA12DF1FFE7531BFEE5FD09A8D83195" box="[338,459,692,716]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="acalcarata">
<emphasis id="1AD58A60FFE7531BFEE5FD09A8D83195" box="[338,459,692,716]" italics="true" pageId="6" pageNumber="46">acalcarata</emphasis>
</taxonomicName>
, referring to the absence of tibial spurs.
</paragraph>
</subSubSection>
<subSubSection id="60BB05F9FFE7531BFF72FD65AD60304D" pageId="6" pageNumber="46" type="discussion">
<paragraph id="281E5672FFE7531BFF72FD65AD60304D" blockId="6.[151,1437,151,1004]" pageId="6" pageNumber="46">
<emphasis id="1AD58A60FFE7531BFF72FD65A82A31A9" bold="true" box="[197,313,728,752]" pageId="6" pageNumber="46">Remarks.</emphasis>
<taxonomicName id="EFA12DF1FFE7531BFE88FD65A8CA31A9" box="[319,473,728,752]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="acalcarata">
<emphasis id="1AD58A60FFE7531BFE88FD65A8CA31A9" box="[319,473,728,752]" italics="true" pageId="6" pageNumber="46">C. acalcarata</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="EFA12DF1FFE7531BFCBFFD65AA7531A9" box="[776,870,728,752]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="1AD58A60FFE7531BFCBFFD65AA7531A9" box="[776,870,728,752]" italics="true" pageId="6" pageNumber="46">C. nigra</emphasis>
</taxonomicName>
by the smaller size, the absence of tibial spurs and the more distal insertion of the stout setae of the postventral lobe of the male ectoprocts.
</paragraph>
</subSubSection>
<subSubSection id="60BB05F9FFE7531BFF70FCA2AB7F30B5" pageId="6" pageNumber="46" type="distribution">
<paragraph id="281E5672FFE7531BFF70FCA2AB7F30B5" blockId="6.[151,1437,151,1004]" pageId="6" pageNumber="46">
<emphasis id="1AD58A60FFE7531BFF70FCA2A8E63061" bold="true" box="[199,501,799,824]" pageId="6" pageNumber="46">Distribution and ecology.</emphasis>
The two new species described in this paper are syntopic and occur in savanna or tree savanna with lateritic soil covered by dry grass. Adults emerge during the dry season which goes from November to end of April.
<taxonomicName id="EFA12DF1FFE7531BFEC4FCD5A8C330D9" box="[371,464,872,896]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="1AD58A60FFE7531BFEC4FCD5A8C330D9" box="[371,464,872,896]" italics="true" pageId="6" pageNumber="46">C. nigra</emphasis>
</taxonomicName>
was collected in February and March and
<taxonomicName id="EFA12DF1FFE7531BFC1FFCD5AD5230D9" box="[936,1089,872,896]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="acalcarata">
<emphasis id="1AD58A60FFE7531BFC1FFCD5AD5230D9" box="[936,1089,872,896]" italics="true" pageId="6" pageNumber="46">C. acalcarata</emphasis>
</taxonomicName>
from January to end of March. After bushfires adults rest on short twigs of grass darkened by smoke where their coloration makes them difficult to see. Disturbed, they fly close to the ground. Recorded from southern
<collectingCountry id="50B616E2FFE7531BFC15FC12AAC4309E" box="[930,983,943,967]" name="Serbia" pageId="6" pageNumber="46">Mali</collectingCountry>
and
<collectingCountry id="50B616E2FFE7531BFBB9FC12ADB63091" box="[1038,1189,943,968]" name="Burkina Faso" pageId="6" pageNumber="46">Burkina Faso</collectingCountry>
only.
<taxonomicName id="EFA12DF1FFE7531BFB5CFC0DAC903091" box="[1259,1411,944,968]" class="Insecta" family="Myrmeleontidae" genus="Capicua" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="fulvicauda">
<emphasis id="1AD58A60FFE7531BFB5CFC0DAC903091" box="[1259,1411,944,968]" italics="true" pageId="6" pageNumber="46">C. fulvicauda</emphasis>
</taxonomicName>
is known from
<collectingCountry id="50B616E2FFE7531BFE90FC69A87130B5" box="[295,354,980,1004]" name="Chad" pageId="6" pageNumber="46">Chad</collectingCountry>
only. Larvae unknown.
</paragraph>
</subSubSection>
</treatment>
</document>