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<document id="C2273B830373621355D3C37A58C3191D" ID-DOI="10.5281/zenodo.5356893" ID-ISSN="2345-7600" ID-Zenodo-Dep="5356893" ID-ZooBank="F7024292-9770-4958-845B-EA3BA4B468AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630218444165" checkinUser="valdenar" docAuthor="Kottelat, Maurice" docDate="2017" docId="E77387A63D74FFF3FEBFFB1D66CFF79D" docLanguage="en" docName="65rbz341-356.pdf" docOrigin="Raffles Bulletin of Zoology 65" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Schistura colossa Kottelat 2017, new species" docType="treatment" docUuid="AE9DD9F6-BA4F-424F-98B7-7D53BF86E3E7" docUuidSource="ZooBank" docVersion="4" lastPageNumber="343" masterDocId="1B4AFFDE3D75FFF1FFB9FF94630BFF9B" masterDocTitle="Schistura colossa and S. klydonion, two new species of loaches from Bolaven Plateau, southern Laos (Teleostei: Nemacheilidae)" masterLastPageNumber="356" masterPageNumber="341" pageNumber="342" updateTime="1699190615416" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7632888735A6A49A7A0F9A35CD5521C3">Schistura colossa and S. klydonion, two new species of loaches from Bolaven Plateau, southern Laos (Teleostei: Nemacheilidae)</mods:title>
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<mods:date id="99FA69FC7B49F2B6A7370FC37DA3A2E9">2017</mods:date>
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<taxonomicName id="A8DA4D333D74FFF0FEBFFB1D62CCFB3A" authority="Kottelat, 2017" authorityName="Kottelat" authorityYear="2017" box="[262,455,1161,1185]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="342" phylum="Chordata" rank="species" species="colossa" status="sp. nov.">
<emphasis id="5DAEEAA23D74FFF0FEBFFB1D62CCFB3A" bold="true" box="[262,455,1161,1185]" italics="true" pageId="1" pageNumber="342">Schistura colossa</emphasis>
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,
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<paragraph id="6F6536B03D74FFF0FECCFB3E62E6FB5A" blockId="1.[262,604,1161,1218]" box="[373,493,1193,1218]" pageId="1" pageNumber="342">
(
<figureCitation id="F7E12A353D74FFF0FEC4FB3E62EEFB5A" box="[381,485,1193,1218]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="2.[135,170,865,887]" captionStartId-1="3.[114,149,2064,2086]" captionStartId-2="4.[136,171,1927,1949]" captionStartId-3="5.[113,148,616,638]" captionStartId-4="5.[113,148,1100,1122]" captionTargetBox-0="[135,1473,189,850]" captionTargetBox-1="[248,1315,189,2047]" captionTargetBox-2="[136,1474,1537,1912]" captionTargetBox-3="[113,1451,189,601]" captionTargetBox-4="[113,1451,693,1085]" captionTargetId-0="figure-305@2.[135,1473,189,850]" captionTargetId-1="figure-10@3.[248,1317,189,2047]" captionTargetId-2="figure-249@4.[136,1474,1537,1912]" captionTargetId-3="figure-10@5.[113,1451,189,601]" captionTargetId-4="figure-29@5.[113,1451,693,1085]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="5" captionText-0="Fig. 2. Schistura colossa, MHNG 2767.084, holotype, 98.0 mm SL; Laos: Xe Pian River on Bolaven Plateau." captionText-1="Fig. 3. Schistura colossa, paratypes; Laos: Xe Pian on Bolaven Plateau; a, CMK 23378, 33.0 mm SL; b, CMK 23313, 43.1 mm SL; c, CMK 23313, 51.5 mm SL; and d, CMK 23313, 71.9 mm SL." captionText-2="Fig. 4. Schistura colossa, CMK 23390, paratype, 51.4 mm SL; Laos: Houay Champi on Bolaven Plateau." captionText-3="Fig. 5. Schistura colossa, CMK 23397, paratype, 77.7 mm SL; Laos: Xe Set on Bolaven Plateau." captionText-4="Fig. 6. Schistura colossa, CMK 23443, 36.0 mm SL; Laos: Houay Makchan-Gnai on Bolaven Plateau." figureDoi-0="http://doi.org/10.5281/zenodo.5356897" figureDoi-1="http://doi.org/10.5281/zenodo.5356899" figureDoi-2="http://doi.org/10.5281/zenodo.5356901" figureDoi-3="http://doi.org/10.5281/zenodo.5356903" figureDoi-4="http://doi.org/10.5281/zenodo.5356905" httpUri-0="https://zenodo.org/record/5356897/files/figure.png" httpUri-1="https://zenodo.org/record/5356899/files/figure.png" httpUri-2="https://zenodo.org/record/5356901/files/figure.png" httpUri-3="https://zenodo.org/record/5356903/files/figure.png" httpUri-4="https://zenodo.org/record/5356905/files/figure.png" pageId="1" pageNumber="342">Figs. 26</figureCitation>
)
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<paragraph id="6F6536B03D74FFF0FFC8FB7E6167FAF9" blockId="1.[112,752,1257,1378]" pageId="1" pageNumber="342">
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<typeStatus id="B06188123D74FFF0FFC8FB7E63D1FA99" box="[113,218,1258,1282]" pageId="1" pageNumber="342" type="holotype">Holotype</typeStatus>
.
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<specimenCode id="3F7C9ECB3D74FFF0FF5DFB7E62ABFA9A" box="[228,416,1257,1282]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="342" type="Museum">MHNG 2767.084</specimenCode>
, 98.0 mm SL;
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:
<collectingRegion id="AD1EF8523D74FFF0FDCCFB7E63D8FAB9" country="Laos" name="Champasak" pageId="1" pageNumber="342">Champasak Province</collectingRegion>
:
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:
<collectingMunicipality id="8F01ACCA3D74FFF0FE1AFA9E62F1FAB9" box="[419,506,1290,1314]" pageId="1" pageNumber="342">Xe Pian</collectingMunicipality>
at dam site, near
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;
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;
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;
<collectorName id="C22F53663D74FFF0FD68FABE63C3FAF9" pageId="1" pageNumber="342">M. Kottelat</collectorName>
&amp;
<collectorName id="C22F53663D74FFF0FF57FADE62AFFAF9" box="[238,420,1354,1378]" pageId="1" pageNumber="342">T. Phommavong</collectorName>
,
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.
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<typeStatus id="B06188123D74FFF0FFC8FA1E63EEFA39" box="[113,229,1418,1442]" pageId="1" pageNumber="342" type="paratype">Paratypes</typeStatus>
.
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<collectionCode id="09CBAE753D74FFF0FF48FA1E6225FA39" box="[241,302,1418,1442]" pageId="1" pageNumber="342">CMK</collectionCode>
23313,
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,
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SL
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;
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<collectionCode id="09CBAE753D74FFF0FDD5FA1E61ABFA39" box="[620,672,1418,1442]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="342">ZRC</collectionCode>
26222,
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SL; same data as holotype
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. —
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, 33.0
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SL;
<collectingCountry id="17CD76203D74FFF0FEE6FA5E629CFA7A" box="[351,407,1482,1505]" name="Laos" pageId="1" pageNumber="342">Laos</collectingCountry>
:
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:
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:
<collectingMunicipality id="8F01ACCA3D74FFF0FF61FA7E623DF999" box="[216,310,1514,1538]" pageId="1" pageNumber="342">Xe Pian</collectingMunicipality>
north of
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;
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;
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;
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&amp;
<collectorName id="C22F53663D74FFF0FD8BF99E61E6F9B9" box="[562,749,1546,1570]" pageId="1" pageNumber="342">T. Phommavong</collectorName>
,
<date id="1B6410703D74FFF0FFCBF9BE623AF9D9" box="[114,305,1578,1602]" pageId="1" pageNumber="342" value="2013-01-17">
<collectingDate id="0B20E9983D74FFF0FFCBF9BE623AF9D9" box="[114,305,1578,1602]" pageId="1" pageNumber="342" value="2013-01-17">17 January 2013</collectingDate>
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.
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<collectionCode id="09CBAE753D74FFF0FEEFF9BE6298F9D9" box="[342,403,1578,1602]" pageId="1" pageNumber="342">CMK</collectionCode>
23397,
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,
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SL;
<collectingCountry id="17CD76203D74FFF0FD0DF9BE61E6F9DA" box="[692,749,1578,1601]" name="Laos" pageId="1" pageNumber="342">Laos</collectingCountry>
:
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:
<collectingCounty id="86044E3C3D74FFF0FEDFF9DE6110F9F9" box="[358,539,1610,1634]" pageId="1" pageNumber="342">Bolaven Plateau</collectingCounty>
:
<collectingMunicipality id="8F01ACCA3D74FFF0FD9EF9DE6398F919" pageId="1" pageNumber="342">Tad Set on Houay Set</collectingMunicipality>
(Xe Set, tributary of Xe Don), near
<location id="6A05606B3D74FFF0FD9DF9FE61E7F919" LSID="urn:lsid:plazi:treatment:E77387A63D74FFF3FEBFFB1D66CFF79D:6A05606B3D74FFF0FD9DF9FE61E7F919" box="[548,748,1642,1666]" country="Laos" county="Bolaven Plateau" latitude="15.300834" longLatPrecision="21" longitude="106.30639" municipality="Tad Set on Houay Set" name="Ban Nonghinkhao" pageId="1" pageNumber="342" stateProvince="Champasak">Ban Nonghinkhao</location>
;
<geoCoordinate id="0AEE50773D74FFF0FFCAF91C63FCF93A" box="[115,247,1672,1697]" degrees="15" direction="north" minutes="18" orientation="latitude" pageId="1" pageNumber="342" precision="15" seconds="03" value="15.300834">15°1803″N</geoCoordinate>
<geoCoordinate id="0AEE50773D74FFF0FEBBF91C629FF93A" box="[258,404,1672,1697]" degrees="106" direction="east" minutes="18" orientation="longitude" pageId="1" pageNumber="342" precision="15" seconds="23" value="106.30639">106°1823″E</geoCoordinate>
;
<quantity id="A8229B553D74FFF0FE1AF91D6116F939" box="[419,541,1673,1698]" metricMagnitude="3" metricUnit="m" metricValue="1.136" pageId="1" pageNumber="342" unit="m" value="1136.0">
<elevation id="E4F7D1833D74FFF0FE1AF91D6116F939" box="[419,541,1673,1698]" metricMagnitude="3" metricUnit="m" metricValue="1.136" pageId="1" pageNumber="342" unit="m" value="1136.0">1136 masl</elevation>
</quantity>
;
<collectorName id="C22F53663D74FFF0FD95F91E61BBF939" box="[556,688,1674,1698]" pageId="1" pageNumber="342">M. Kottelat</collectorName>
&amp;
<collectorName id="C22F53663D74FFF0FD62F91E6201F959" pageId="1" pageNumber="342">T. Phommavong</collectorName>
,
<date id="1B6410703D74FFF0FEA1F93E62DAF959" box="[280,465,1706,1730]" pageId="1" pageNumber="342" value="2013-01-19">
<collectingDate id="0B20E9983D74FFF0FEA1F93E62DAF959" box="[280,465,1706,1730]" pageId="1" pageNumber="342" value="2013-01-19">19 January 2013</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="DFB23CED3D74FFF0FDBEF93E610DF8F9" accessionNumber="CMK 23390" collectingDate="2013-01-18" collectionCode="CMK" collectorName="M. Kottelat &amp; T. Phommavong" country="Laos" county="Bolaven Plateau" elevation="938" latitude="15.202499" location="Houay Champi" longLatPrecision="21" longitude="106.13084" municipality="Tad Champi" pageId="1" pageNumber="342" specimenCode="1" specimenCount="1" stateProvince="Champasak" typeStatus="paratype">
<collectionCode id="09CBAE753D74FFF0FDBEF93E614FF959" box="[519,580,1706,1730]" pageId="1" pageNumber="342">CMK</collectionCode>
23390,
<specimenCode id="3F7C9ECB3D74FFF0FD1BF93E61A4F959" box="[674,687,1706,1730]" collectionCode="CMK" pageId="1" pageNumber="342">1</specimenCode>
,
<quantity id="A8229B553D74FFF0FD78F93D6391F97A" metricMagnitude="-2" metricUnit="m" metricValue="5.14" pageId="1" pageNumber="342" unit="mm" value="51.4">51.4 mm</quantity>
SL;
<collectingCountry id="17CD76203D74FFF0FF6BF95E6201F97A" box="[210,266,1738,1761]" name="Laos" pageId="1" pageNumber="342">Laos</collectingCountry>
:
<collectingRegion id="AD1EF8523D74FFF0FEACF95E62F7F979" box="[277,508,1738,1762]" country="Laos" name="Champasak" pageId="1" pageNumber="342">Champasak Province</collectingRegion>
:
<collectingCounty id="86044E3C3D74FFF0FDB1F95E61B7F979" box="[520,700,1738,1762]" pageId="1" pageNumber="342">Bolaven Plateau</collectingCounty>
:
<collectingMunicipality id="8F01ACCA3D74FFF0FD71F95E63CEF899" pageId="1" pageNumber="342">Tad Champi</collectingMunicipality>
(waterfall) on
<location id="6A05606B3D74FFF0FEDBF97E6109F899" LSID="urn:lsid:plazi:treatment:E77387A63D74FFF3FEBFFB1D66CFF79D:6A05606B3D74FFF0FEDBF97E6109F899" box="[354,514,1770,1794]" country="Laos" county="Bolaven Plateau" latitude="15.202499" longLatPrecision="21" longitude="106.13084" municipality="Tad Champi" name="Houay Champi" pageId="1" pageNumber="342" stateProvince="Champasak">Houay Champi</location>
(tributary of Xe Don), off road from
<location id="6A05606B3D74FFF0FEB3F89E6243F8B9" LSID="urn:lsid:plazi:treatment:E77387A63D74FFF3FEBFFB1D66CFF79D:6A05606B3D74FFF0FEB3F89E6243F8B9" box="[266,328,1802,1826]" country="Laos" county="Bolaven Plateau" latitude="15.202499" longLatPrecision="21" longitude="106.13084" municipality="Tad Champi" name="Pakse" pageId="1" pageNumber="342" stateProvince="Champasak">Pakse</location>
to
<location id="6A05606B3D74FFF0FED5F89E62C2F8B9" LSID="urn:lsid:plazi:treatment:E77387A63D74FFF3FEBFFB1D66CFF79D:6A05606B3D74FFF0FED5F89E62C2F8B9" box="[364,457,1802,1826]" country="Laos" county="Bolaven Plateau" latitude="15.202499" longLatPrecision="21" longitude="106.13084" municipality="Tad Champi" name="Paksong" pageId="1" pageNumber="342" stateProvince="Champasak">Paksong</location>
, north at
<location id="6A05606B3D74FFF0FD96F89E6180F8B9" LSID="urn:lsid:plazi:treatment:E77387A63D74FFF3FEBFFB1D66CFF79D:6A05606B3D74FFF0FD96F89E6180F8B9" box="[559,651,1802,1826]" country="Laos" county="Bolaven Plateau" latitude="15.202499" longLatPrecision="21" longitude="106.13084" municipality="Tad Champi" name="Ban Lak" pageId="1" pageNumber="342" stateProvince="Champasak">Ban Lak</location>
38 [Km- 38 village];
<geoCoordinate id="0AEE50773D74FFF0FF49F8BC6264F8DA" box="[240,367,1832,1857]" degrees="15" direction="north" minutes="12" orientation="latitude" pageId="1" pageNumber="342" precision="15" seconds="09" value="15.202499">15°1209″N</geoCoordinate>
<geoCoordinate id="0AEE50773D74FFF0FECDF8BC610AF8DA" box="[372,513,1832,1857]" degrees="106" direction="east" minutes="07" orientation="longitude" pageId="1" pageNumber="342" precision="15" seconds="51" value="106.13084">106°0751″E</geoCoordinate>
;
<quantity id="A8229B553D74FFF0FDB3F8BE6166F8D9" box="[522,621,1834,1858]" metricMagnitude="2" metricUnit="m" metricValue="9.379999999999999" pageId="1" pageNumber="342" unit="m" value="938.0">
<elevation id="E4F7D1833D74FFF0FDB3F8BE6166F8D9" box="[522,621,1834,1858]" metricMagnitude="2" metricUnit="m" metricValue="9.379999999999999" pageId="1" pageNumber="342" unit="m" value="938.0">938 masl</elevation>
</quantity>
;
<collectorName id="C22F53663D74FFF0FDCFF8BE61FBF8D9" box="[630,752,1834,1858]" pageId="1" pageNumber="342">M. Kottelat</collectorName>
&amp;
<collectorName id="C22F53663D74FFF0FF36F8DE624EF8F9" box="[143,325,1866,1890]" pageId="1" pageNumber="342">T. Phommavong</collectorName>
,
<date id="1B6410703D74FFF0FEEBF8DE610DF8F9" box="[338,518,1866,1890]" pageId="1" pageNumber="342" value="2013-01-18">
<collectingDate id="0B20E9983D74FFF0FEEBF8DE610DF8F9" box="[338,518,1866,1890]" pageId="1" pageNumber="342" value="2013-01-18">18 January 2013</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="6F6536B03D74FFF0FFC8F81D61FBF79A" blockId="1.[113,752,1929,2050]" pageId="1" pageNumber="342">
<materialsCitation id="DFB23CED3D74FFF0FFC8F81D61C9F799" accessionNumber="CMK 23443" collectionCode="CMK" country="Laos" county="Bolaven Plateau" elevation="784" latitude="15.070833" location="Ban" longLatPrecision="21" longitude="106.54278" municipality="Houay Makchan-Gnai" pageId="1" pageNumber="342" specimenCode="1" specimenCount="2" stateProvince="Champasak">
<emphasis id="5DAEEAA23D74FFF0FFC8F81D62CBF83A" bold="true" box="[113,448,1929,1953]" pageId="1" pageNumber="342">
Other material (non
<typeStatus id="B06188123D74FFF0FECBF81D62B8F83A" box="[370,435,1929,1953]" pageId="1" pageNumber="342">types</typeStatus>
).
</emphasis>
<collectionCode id="09CBAE753D74FFF0FE72F81E6102F839" box="[459,521,1930,1954]" pageId="1" pageNumber="342">CMK</collectionCode>
23443,
<specimenCode id="3F7C9ECB3D74FFF0FDD2F81E6172F839" box="[619,633,1930,1954]" collectionCode="CMK" pageId="1" pageNumber="342">1</specimenCode>
, 36.0 mm SL;
<collectingCountry id="17CD76203D74FFF0FF1AF83E63D7F85A" box="[163,220,1962,1985]" name="Laos" pageId="1" pageNumber="342">Laos</collectingCountry>
:
<collectingRegion id="AD1EF8523D74FFF0FF50F83E62DEF859" box="[233,469,1962,1986]" country="Laos" name="Champasak" pageId="1" pageNumber="342">Champasak Province</collectingRegion>
:
<collectingCounty id="86044E3C3D74FFF0FE5AF83E6190F859" box="[483,667,1962,1986]" pageId="1" pageNumber="342">Bolaven Plateau</collectingCounty>
:
<collectingMunicipality id="8F01ACCA3D74FFF0FD11F83E621AF879" pageId="1" pageNumber="342">Houay Makchan-Gnai</collectingMunicipality>
at bridge on road from
<location id="6A05606B3D74FFF0FD9BF85E6146F87A" LSID="urn:lsid:plazi:treatment:E77387A63D74FFF3FEBFFB1D66CFF79D:6A05606B3D74FFF0FD9BF85E6146F87A" box="[546,589,1994,2017]" country="Laos" county="Bolaven Plateau" latitude="15.070833" longLatPrecision="21" longitude="106.54278" municipality="Houay Makchan-Gnai" name="Ban" pageId="1" pageNumber="342" stateProvince="Champasak">Ban</location>
Ta-Od to
<location id="6A05606B3D74FFF0FD7CF85E6217F799" LSID="urn:lsid:plazi:treatment:E77387A63D74FFF3FEBFFB1D66CFF79D:6A05606B3D74FFF0FD7CF85E6217F799" country="Laos" county="Bolaven Plateau" latitude="15.070833" longLatPrecision="21" longitude="106.54278" municipality="Houay Makchan-Gnai" name="Ban Nongphanouan" pageId="1" pageNumber="342" stateProvince="Champasak">Ban Nongphanouan</location>
;
<geoCoordinate id="0AEE50773D74FFF0FE92F87C62A5F79A" box="[299,430,2024,2049]" degrees="15" direction="north" minutes="04" orientation="latitude" pageId="1" pageNumber="342" precision="15" seconds="15" value="15.070833">15°0415″N</geoCoordinate>
<geoCoordinate id="0AEE50773D74FFF0FE01F87C6143F79A" box="[440,584,2024,2049]" degrees="106" direction="east" minutes="32" orientation="longitude" pageId="1" pageNumber="342" precision="15" seconds="34" value="106.54278">106°3234″E</geoCoordinate>
;
<quantity id="A8229B553D74FFF0FDEEF87E61C9F799" box="[599,706,2026,2050]" metricMagnitude="2" metricUnit="m" metricValue="7.84" pageId="1" pageNumber="342" unit="m" value="784.0">
<elevation id="E4F7D1833D74FFF0FDEEF87E61C9F799" box="[599,706,2026,2050]" metricMagnitude="2" metricUnit="m" metricValue="7.84" pageId="1" pageNumber="342" unit="m" value="784.0">784 masl</elevation>
</quantity>
</materialsCitation>
; M.
</paragraph>
<paragraph id="6F6536B03D74FFF0FC94FB1E6069FA99" blockId="1.[813,1452,1162,1282]" pageId="1" pageNumber="342">
<materialsCitation id="DFB23CED3D74FFF0FC94FB1E6055FA99" accessionNumber="CMK 15931" collectingDate="2013-01-23" collectionCode="CMK" collectorName="T. Phommavong &amp; K. Vattahanatham" country="Laos" location="Xe Set" municipality="Xe Don" pageId="1" pageNumber="342" specimenCode="1" specimenCount="1" stateProvince="Salavan">
Kottelat &amp;
<collectorName id="C22F53663D74FFF0FC09FB1E6766FB39" box="[944,1133,1162,1186]" pageId="1" pageNumber="342">T. Phommavong</collectorName>
,
<date id="1B6410703D74FFF0FBC2FB1E663CFB39" box="[1147,1335,1162,1186]" pageId="1" pageNumber="342" value="2013-01-23">
<collectingDate id="0B20E9983D74FFF0FBC2FB1E663CFB39" box="[1147,1335,1162,1186]" pageId="1" pageNumber="342" value="2013-01-23">23 January 2013</collectingDate>
</date>
. —
<collectionCode id="09CBAE753D74FFF0FAD7FB1E66A0FB39" box="[1390,1451,1162,1186]" pageId="1" pageNumber="342">CMK</collectionCode>
15931,
<specimenCode id="3F7C9ECB3D74FFF0FC38FB3E6085FB59" box="[897,910,1194,1218]" collectionCode="CMK" pageId="1" pageNumber="342">1</specimenCode>
,
<quantity id="A8229B553D74FFF0FC26FB3E6709FB5A" box="[927,1026,1194,1218]" metricMagnitude="-2" metricUnit="m" metricValue="8.72" pageId="1" pageNumber="342" unit="mm" value="87.2">87.2 mm</quantity>
SL;
<collectingCountry id="17CD76203D74FFF0FB84FB3E677EFB5A" box="[1085,1141,1194,1217]" name="Laos" pageId="1" pageNumber="342">Laos</collectingCountry>
:
<collectingRegion id="AD1EF8523D74FFF0FB3AFB3E664DFB59" box="[1155,1350,1194,1218]" country="Laos" name="Salavan" pageId="1" pageNumber="342">Salavan Province</collectingRegion>
:
<collectingMunicipality id="8F01ACCA3D74FFF0FAEAFB3E66A0FB5A" box="[1363,1451,1194,1217]" pageId="1" pageNumber="342">Xe Don</collectingMunicipality>
basin,
<location id="6A05606B3D74FFF0FCCDFB5E60B5FB79" LSID="urn:lsid:plazi:treatment:E77387A63D74FFF3FEBFFB1D66CFF79D:6A05606B3D74FFF0FCCDFB5E60B5FB79" box="[884,958,1226,1250]" country="Laos" municipality="Xe Don" name="Xe Set" pageId="1" pageNumber="342" stateProvince="Salavan">Xe Set</location>
upstream of reservoir;
<collectorName id="C22F53663D74FFF0FB07FB5E668FFB79" box="[1214,1412,1226,1250]" pageId="1" pageNumber="342">K. Vattahanatham</collectorName>
, no date
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="27C0653B3D74FFF0FC94FABD6758F799" pageId="1" pageNumber="342" type="diagnosis">
<paragraph id="6F6536B03D74FFF0FC94FABD6718F979" blockId="1.[811,1452,1321,1762]" pageId="1" pageNumber="342">
<emphasis id="5DAEEAA23D74FFF0FC94FABD60ACFADA" bold="true" box="[813,935,1321,1345]" pageId="1" pageNumber="342">Diagnosis.</emphasis>
<taxonomicName id="A8DA4D333D74FFF0FC0AFABE6777FAD9" authorityName="Kottelat" authorityYear="2017" box="[947,1148,1322,1346]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="342" phylum="Chordata" rank="species" species="colossa">
<emphasis id="5DAEEAA23D74FFF0FC0AFABE6777FAD9" box="[947,1148,1322,1346]" italics="true" pageId="1" pageNumber="342">Schistura colossa</emphasis>
</taxonomicName>
is distinguished from the other species of the genus in Southeast Asia by its colour pattern: the body has 1621 bars, wider than interspaces, quite regularly shaped in juveniles; with increasing size, the bars become more irregular, some become interrupted or joined; in largest individuals (above
<quantity id="A8229B553D74FFF0FB43FA5D6641FA7A" box="[1274,1354,1481,1505]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="342" unit="mm" value="60.0">60 mm</quantity>
SL), on the caudal peduncle, the bars are broken up in blotches of irregular shape and size and irregularly set. The black pattern at the base of the caudal-fin is made of a vertically elongated blotch occupying the middle half of the fin base, sometimes with a constriction at the level of the lateral line; above and below, this blotch is continued by an arched band, along the base of the principal rays, not reaching the dorsal and ventral midlines.
</paragraph>
<paragraph id="6F6536B03D74FFF0FC94F89E6758F799" blockId="1.[813,1452,1802,2050]" pageId="1" pageNumber="342">
Additional diagnostic characters, not unique to the species are: relatively large size (up to at least
<quantity id="A8229B553D74FFF0FB72F8BE661DF8DA" box="[1227,1302,1834,1858]" metricMagnitude="-2" metricUnit="m" metricValue="9.8" pageId="1" pageNumber="342" unit="mm" value="98.0">98 mm</quantity>
SL); depth of caudal peduncle 1.31.6 times in its length; 78½ branched dorsal-fin rays; pelvic axillary lobe rudimentary and free; usually 9+8 branched caudal-fin rays; no suborbital flap; upper lip with a small median notch; processus dentiformis present, pointed; and origin of dorsal fin above or slightly behind origin of pelvic fin.
</paragraph>
</subSubSection>
<caption id="3BA566383D77FFF3FF3EFCF567CAFCEC" ID-DOI="http://doi.org/10.5281/zenodo.5356897" ID-Zenodo-Dep="5356897" box="[135,1217,865,887]" httpUri="https://zenodo.org/record/5356897/files/figure.png" pageId="2" pageNumber="343" startId="2.[135,170,865,887]" targetBox="[135,1473,189,850]" targetPageId="2">
<paragraph id="6F6536B03D77FFF3FF3EFCF567CAFCEC" blockId="2.[135,1217,865,887]" box="[135,1217,865,887]" pageId="2" pageNumber="343">
Fig. 2.
<taxonomicName id="A8DA4D333D77FFF3FF68FCF562DBFCED" authority=", MHNG" authorityName="MHNG" box="[209,464,865,887]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="343" phylum="Chordata" rank="species" species="colossa">
<emphasis id="5DAEEAA23D77FFF3FF68FCF56272FCED" box="[209,377,865,886]" italics="true" pageId="2" pageNumber="343">Schistura colossa</emphasis>
, MHNG
</taxonomicName>
2767.084, holotype, 98.0 mm SL; Laos: Xe Pian River on Bolaven Plateau.
</paragraph>
</caption>
<subSubSection id="27C0653B3D77FFF3FF31FC3A66CFF79D" pageId="2" pageNumber="343" type="description">
<paragraph id="6F6536B03D77FFF3FF31FC3A6102FAFD" blockId="2.[135,778,942,1383]" pageId="2" pageNumber="343">
<emphasis id="5DAEEAA23D77FFF3FF31FC3A6218FC5D" bold="true" box="[136,275,942,966]" pageId="2" pageNumber="343">Description.</emphasis>
See
<figureCitation id="F7E12A353D77FFF3FEFAFC3A62AEFC5D" box="[323,421,942,966]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="2.[135,170,865,887]" captionStartId-1="3.[114,149,2064,2086]" captionStartId-2="4.[136,171,1927,1949]" captionStartId-3="5.[113,148,616,638]" captionStartId-4="5.[113,148,1100,1122]" captionTargetBox-0="[135,1473,189,850]" captionTargetBox-1="[248,1315,189,2047]" captionTargetBox-2="[136,1474,1537,1912]" captionTargetBox-3="[113,1451,189,601]" captionTargetBox-4="[113,1451,693,1085]" captionTargetId-0="figure-305@2.[135,1473,189,850]" captionTargetId-1="figure-10@3.[248,1317,189,2047]" captionTargetId-2="figure-249@4.[136,1474,1537,1912]" captionTargetId-3="figure-10@5.[113,1451,189,601]" captionTargetId-4="figure-29@5.[113,1451,693,1085]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="5" captionText-0="Fig. 2. Schistura colossa, MHNG 2767.084, holotype, 98.0 mm SL; Laos: Xe Pian River on Bolaven Plateau." captionText-1="Fig. 3. Schistura colossa, paratypes; Laos: Xe Pian on Bolaven Plateau; a, CMK 23378, 33.0 mm SL; b, CMK 23313, 43.1 mm SL; c, CMK 23313, 51.5 mm SL; and d, CMK 23313, 71.9 mm SL." captionText-2="Fig. 4. Schistura colossa, CMK 23390, paratype, 51.4 mm SL; Laos: Houay Champi on Bolaven Plateau." captionText-3="Fig. 5. Schistura colossa, CMK 23397, paratype, 77.7 mm SL; Laos: Xe Set on Bolaven Plateau." captionText-4="Fig. 6. Schistura colossa, CMK 23443, 36.0 mm SL; Laos: Houay Makchan-Gnai on Bolaven Plateau." figureDoi-0="http://doi.org/10.5281/zenodo.5356897" figureDoi-1="http://doi.org/10.5281/zenodo.5356899" figureDoi-2="http://doi.org/10.5281/zenodo.5356901" figureDoi-3="http://doi.org/10.5281/zenodo.5356903" figureDoi-4="http://doi.org/10.5281/zenodo.5356905" httpUri-0="https://zenodo.org/record/5356897/files/figure.png" httpUri-1="https://zenodo.org/record/5356899/files/figure.png" httpUri-2="https://zenodo.org/record/5356901/files/figure.png" httpUri-3="https://zenodo.org/record/5356903/files/figure.png" httpUri-4="https://zenodo.org/record/5356905/files/figure.png" pageId="2" pageNumber="343">Figs. 26</figureCitation>
for general appearance and
<tableCitation id="2258030B3D77FFF3FD72FC3A639FFC7D" captionStart="Table 1" captionStartId="4.[135,190,189,211]" captionText="Table 1. Morphometric data of type material of Schistura colossa (n=10; MHNG 2767.084, holotype; CMK 23313, 6; CMK 23378, 3). Ranges and means include holotype data." pageId="2" pageNumber="343">Table 1</tableCitation>
for morphometric data of
<typeStatus id="B06188123D77FFF3FE06FC5A6117FC7D" box="[447,540,974,998]" pageId="2" pageNumber="343" type="holotype">holotype</typeStatus>
and
<specimenCount id="79DCFD393D77FFF3FDEEFC5A61D3FC7C" box="[599,728,974,999]" pageId="2" pageNumber="343" type="generic" typeStatus="paratypes">9 paratypes</specimenCount>
. An elongate nemacheilid with body depth slowly increasing up to dorsal-fin origin. Behind dorsal fin, body depth almost uniform until caudal-fin base. Dorsal profile continuous between head and body (no hump). Head slightly depressed; body slightly compressed anteriorly to very compressed posteriorly. Interorbital area flat. In lateral view, upper margin of eye flushed with dorsal profile of head. Cheeks not swollen. Snout pointed. Depth of caudal peduncle 1.31.6 times in its length. Low dorsal keel on posterior half of post-dorsal area. Low ventral keel on entire length of caudal peduncle. Dorsal keel continuous with upper margin of caudal fin. Largest recorded size 98.0 mm SL.
</paragraph>
<paragraph id="6F6536B03D77FFF3FF31FA1A6116F8DD" blockId="2.[136,776,1421,1862]" pageId="2" pageNumber="343">Dorsal fin with 4 unbranched and 7½ (1) or 8½ (9*) branched rays; distal margin convex; branched ray 2 or 3 longest. Pectoral fin with 1 unbranched and 8 (1) or 10 (9*) branched rays, rounded, reaching about halfway of distance to pelvic-fin base; origin over base of simple or first branched pelvic-fin rays. No axillary pectoral lobe. Pelvic fin with 1 unbranched and 7 branched rays; reaching about two thirds of distance to anus; rounded; posterior margin convex; axillary pelvic lobe rudimentary, entirely free. Anus situated about 2 eye diameters in front of anal fin. Anal fin with 3 unbranched and 5½ branched rays; distal margin rounded; branched ray 2 longest. Caudal fin with 9+8 (9*) or 8+8 (1) branched rays; dorsal and ventral procurrent rays cannot be counted; emarginate, lobes rounded, subequal.</paragraph>
<paragraph id="6F6536B03D77FFF3FF31F8FA67E6FB9D" blockId="2.[136,775,1902,2054]" lastBlockId="2.[835,1474,942,1030]" pageId="2" pageNumber="343">
Body entirely scaled, except belly in front of pelvic fins and predorsal area (anterior extremity in smaller specimens, entirely in specimens over about
<quantity id="A8229B553D77FFF3FDC6F83A61C5F85D" box="[639,718,1966,1990]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="343" unit="mm" value="60.0">60 mm</quantity>
SL). Scales embedded, deeply embedded in anterior predorsal area. Lateral line complete, with 92107 pores (difficult to count with accuracy). Cephalic lateral line system with 6 supraorbital, 4 + 1213 infraorbital, 911 preoperculomandibular and 3 supratemporal pores.
</paragraph>
<paragraph id="6F6536B03D77FFF3FCFAFBBA67E5FA7D" blockId="2.[834,1477,1070,1510]" pageId="2" pageNumber="343">
Anterior nostril pierced in front side of a pointed flap-like tube. Posterior nostril adjacent to anterior one. Mouth strongly arched, gape about 22½ times wider than long (
<figureCitation id="F7E12A353D77FFF3FAD4FBFA66B8FB1D" box="[1389,1459,1134,1158]" captionStart="Fig" captionStartId="6.[136,171,655,677]" captionTargetBox="[136,775,189,639]" captionTargetId="figure-28@6.[136,775,189,639]" captionTargetPageId="6" captionText="Fig. 7. Schistura colossa, CMK 23378, paratype, 81.1 mm SL; mouth." figureDoi="http://doi.org/10.5281/zenodo.5356907" httpUri="https://zenodo.org/record/5356907/files/figure.png" pageId="2" pageNumber="343">Fig. 7</figureCitation>
). Lips thick. Upper lip with small median notch, with a few shallow furrows in median area and near corner of mouth, edge not crenulated. Processus dentiformis present. Lower lip with narrow median interruption; median part with 13 shallow sulcus, lateral part smooth. Tip of lower jaw not exposed. A deep median concavity in lower jaw (in adults). Inner rostral barbel reaching corner of mouth; outer one almost reaching vertical of anterior margin of eye. Maxillary barbel reaching vertical of posterior margin of eye. Intestine with a bend behind stomach (
<figureCitation id="F7E12A353D77FFF3FB34FA3A67DAFA5D" box="[1165,1233,1454,1478]" captionStart="Fig" captionStartId="6.[835,870,959,981]" captionTargetBox="[979,1337,189,944]" captionTargetId="figure-30@6.[937,1371,172,958]" captionTargetPageId="6" captionText="Fig. 8. Schistura colossa, CMK 23378, paratype, 76.6 mm SL; digestive tract. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5356909" httpUri="https://zenodo.org/record/5356909/files/figure.png" pageId="2" pageNumber="343">Fig. 8</figureCitation>
). Air bladder without posterior chamber in abdominal cavity.
</paragraph>
<paragraph id="6F6536B03D77FFF3FCFAF99A609EF9DD" blockId="2.[835,1474,1550,1606]" pageId="2" pageNumber="343">
<emphasis id="5DAEEAA23D77FFF3FCFAF99A6721F9BD" bold="true" box="[835,1066,1550,1574]" pageId="2" pageNumber="343">Sexual dimorphism.</emphasis>
None observed. Ripe females deeper bodied.
</paragraph>
<paragraph id="6F6536B03D77FFF3FCFAF9FA66CFF79D" blockId="2.[834,1476,1646,2054]" pageId="2" pageNumber="343">
<emphasis id="5DAEEAA23D77FFF3FCFAF9FA60DFF91D" bold="true" box="[835,980,1646,1670]" pageId="2" pageNumber="343">Colouration.</emphasis>
About 3 weeks after fixation. Head and body background colour pale brown; throat, belly and lower part of caudal peduncle pale greyish. Except otherwise stated, all markings dark brown. Body with 1621 bars (67 predorsal, 34 subdorsal, 710 postdorsal), extending from dorsal midline to level of pectoral fin, wider than interspaces, some continuous across back with contralaterals, others dissociated into blotches in predorsal area. Bars of quite regular shape in juveniles. With increasing size, bars becoming more irregular in shape, some becoming interrupted or joined, more obvious posteriorly. In specimens above about
<quantity id="A8229B553D77FFF3FB64F83A662CF85D" box="[1245,1319,1966,1990]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="343" unit="mm" value="60.0">60 mm</quantity>
SL, on caudal peduncle, bars broken up in blotches of irregular shape and size and irregularly set. Axial inner stripe faint or indistinct.
</paragraph>
</subSubSection>
</treatment>
</document>