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<document id="707CEEC942AF709D82B1E1B056DBFEF9" ID-DOI="10.26515/rzsi/v122/i3/2022/169417" ID-ISSN="2581-8686" ID-Zenodo-Dep="13185566" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722611788751" checkinUser="felipe" docAuthor="Saikia, Bhaskar, Laskar, Mostaque A., Dinesh, K. P., Shabnam, A. &amp; Sinha, Bikramjit" docDate="2022" docId="190C8795FFE2FF8F30FFFCB6FA8CC09C" docLanguage="en" docName="RecZoolSurvIndia.122.3.247-266.pdf" docOrigin="Records of the Zoological Survey of India 122 (3)" docStyle="DocumentStyle:30C78A88DFAFB68AE5A0604D5A6B2FFB.3:RecZoolSurvIndia.2017-.journal_article" docStyleId="30C78A88DFAFB68AE5A0604D5A6B2FFB" docStyleName="RecZoolSurvIndia.2017-.journal_article" docStyleVersion="3" docTitle="Amolops tawang Saikia, Laskar, Dinesh, Shabnam &amp; Sinha, 2022, sp. nov." docType="treatment" docUuid="0E8EF24C-3FCD-4D1C-9245-3716E44044C6" docUuidSource="ZooBank" docVersion="2" lastPageNumber="256" masterDocId="E535FFEDFFE5FF86305FFFEBFFBDC062" masterDocTitle="Description of two new species of Amolops (Anura: Ranidae) from Arunachal Pradesh, Northeast India under the morphological Viridimaculatus species group" masterLastPageNumber="266" masterPageNumber="247" pageNumber="254" updateTime="1722662603149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3E2CC2E882D6A67BF6A6FD5E7AD0EBF3">Description of two new species of Amolops (Anura: Ranidae) from Arunachal Pradesh, Northeast India under the morphological Viridimaculatus species group</mods:title>
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<mods:namePart id="F8C9FBAD4C42D27E65DFC1A1CDC9F55F">Saikia, Bhaskar</mods:namePart>
<mods:affiliation id="4A39CF346D9B5818E333981A3D16377E">Zoological Survey of India, North Eastern Regional Centre, Shillong - 793003, Meghalaya, India; Email: bj. sinha @ gov. in</mods:affiliation>
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<mods:namePart id="CBD99A15B8A113EC0AF3C2981FC52189">Laskar, Mostaque A.</mods:namePart>
<mods:affiliation id="08BEFFEF5F1B94648253BA1D04C3B037">Department of Biotechnology, St. Anthonys College, Shillong - 793001, Meghalaya, India</mods:affiliation>
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<mods:namePart id="4F41C0601B11D6750D150ED10BFB4582">Dinesh, K. P.</mods:namePart>
<mods:affiliation id="122B09A67ABC75DA3A0CCB400005D405">Zoological Survey of India, Western Regional Centre, Vidyanagar, Pune - 411044, Maharashtra, India</mods:affiliation>
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<mods:namePart id="C55322D125E31AC054EF10E79623E4CC">Shabnam, A.</mods:namePart>
<mods:affiliation id="4E21CD2503D3BACD5FA29E479BC57DCE">Zoological Survey of India, Western Regional Centre, Vidyanagar, Pune - 411044, Maharashtra, India</mods:affiliation>
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<mods:namePart id="9728AF63B87F95F277B0B4B658634623">Sinha, Bikramjit</mods:namePart>
<mods:affiliation id="DE93096A56553B132AAEA1F6A42AB299">Zoological Survey of India, North Eastern Regional Centre, Shillong - 793003, Meghalaya, India; Email: bj. sinha @ gov. in</mods:affiliation>
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<mods:date id="CD992C636C4B66445048F2F53A3A0768">2022</mods:date>
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<taxonomicName id="56A54D00FFE2FF8130FFFCB6FEE3C31A" authorityName="Saikia &amp; Laskar &amp; Dinesh &amp; Shabnam &amp; Sinha" authorityYear="2022" box="[160,350,861,888]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="7" pageNumber="254" phylum="Chordata" rank="species" species="tawang" status="sp. nov.">
<emphasis id="A3D1EA91FFE2FF8130FFFCB6FEE3C31A" bold="true" box="[160,350,861,888]" italics="true" pageId="7" pageNumber="254">Amolops tawang</emphasis>
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<paragraph id="911A3683FFE2FF8130FFFC69FE1BC3FE" blockId="7.[160,797,861,1176]" box="[160,422,898,924]" pageId="7" pageNumber="254">(Tawang Cascade Frog)</paragraph>
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urn:lsid:zoobank.org:act:
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<materialsCitation id="21CD3CDEFFE2FF813097FC05FCA0C432" collectionCode="V, NERC" collectorName="Bikramjit Sinha" latitude="25.925459" location="Gomkyaleng village" pageId="7" pageNumber="254" specimenCode="ZSI/1772" specimenCount="1" typeStatus="holotype">
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:
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/A/
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, adult male (
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), collected by
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and party on 08 March, 2018 from the
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(
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; E
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91.780033; alt.:
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),
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,
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,
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,
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.
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<paragraph id="911A3683FFE2FF8133DAFF3CFB5DC36E" blockId="7.[861,1499,214,1176]" pageId="7" pageNumber="254">
<emphasis id="A3D1EA91FFE2FF8133DAFF3CFB07C092" box="[901,1210,214,240]" italics="true" pageId="7" pageNumber="254">
Diagnosis:
<taxonomicName id="56A54D00FFE2FF81345FFF3DFB07C092" authorityName="Saikia &amp; Laskar &amp; Dinesh &amp; Shabnam &amp; Sinha" authorityYear="2022" box="[1024,1210,214,240]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="7" pageNumber="254" phylum="Chordata" rank="species" species="tawang" status="sp. nov.">Amolops tawang</taxonomicName>
</emphasis>
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<emphasis id="A3D1EA91FFE2FF81349BFF3DFAA4C092" bold="true" box="[1220,1305,214,240]" pageId="7" pageNumber="254">sp. nov</emphasis>
.
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is a member of the morphological
<taxonomicName id="56A54D00FFE2FF81346EFF11FB5FC176" baseAuthorityName="Jiang" baseAuthorityYear="1983" box="[1073,1250,250,276]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="7" pageNumber="254" phylum="Chordata" rank="species" species="viridimaculatus">
<emphasis id="A3D1EA91FFE2FF81346EFF11FB5FC176" box="[1073,1250,250,276]" italics="true" pageId="7" pageNumber="254">Viridimaculatus</emphasis>
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species group isolated in geographical space from its phylogenetic sister species having a distinct morphological set of characters. The new species can be diagnosed based on the following set of morphological characters: adult male size large (SVL
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); head wider than long (HW&gt;HL); snout rounded, snout longer than eye length (SL&gt;EL); inter-orbital distance equals the upper eyelid width but smaller than the inter-narial distance (IUE=UEW&lt;IN); tympanum small and extremely depressed, about 20% of eye length; finger I disk slightly dilated, all disks with circum-marginal grooves, but absent in finger I; tibia longer than femur and foot (TiL&gt;ShL), about 2/3
<superScript id="66D09BCBFFE2FF8134ABFD42FABFC2D5" attach="left" box="[1268,1282,681,695]" fontSize="6" pageId="7" pageNumber="254">rd</superScript>
of SVL; tibio-tarsal articulation reaches up to snout; supra-tympanic fold strong; dorso-lateral skin granular.
</paragraph>
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<emphasis id="A3D1EA91FFE2FF8133DAFCFCFB79C352" box="[901,1220,790,816]" italics="true" pageId="7" pageNumber="254">
Description of the
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: Adult male (SVL
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), body robust, head wider than long (HW&gt;HL), flat above; snout rounded, protruding in lateral view; snout longer than the eye diameter (SL&gt;EL); canthus rostralis distinct, loreal region depressed; inter-orbital space flat, equals the width of the eyelid, but narrower than the inter-narial distance (IUE=UEW&lt;IN); the nostrils laterally placed, equidistant between snout and eye (NE=NS); pupil horizontal; tympanum small and extremely depressed, separated from the eye by a distance of more than twice its diameter; pineal ocellus visible; a pair of vomerine teeth situated between the choanae; tongue bilobed. The vocal sac faintly visible.
</paragraph>
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<emphasis id="A3D1EA91FFE2FF8130FFF8B7FEB5C714" bold="true" box="[160,264,1884,1910]" pageId="7" pageNumber="254">Figure 6.</emphasis>
Holotype image of
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<emphasis id="A3D1EA91FFE2FF8131B5F8B7FD22C714" box="[490,671,1884,1910]" italics="true" pageId="7" pageNumber="254">Amolops tawang</emphasis>
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<emphasis id="A3D1EA91FFE2FF8132FAF8B7FD41C714" bold="true" box="[677,764,1884,1910]" pageId="7" pageNumber="254">
<taxonomicNameLabel id="B8E257EAFFE2FF8132FAF8B7FD41C714" box="[677,764,1884,1910]" pageId="7" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
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in preserved condition (
<emphasis id="A3D1EA91FFE2FF81344FF8B7FB97C714" bold="true" box="[1040,1066,1884,1910]" pageId="7" pageNumber="254">A:</emphasis>
dorsal view;
<emphasis id="A3D1EA91FFE2FF8134E1F8B7FB6BC714" bold="true" box="[1214,1238,1884,1910]" pageId="7" pageNumber="254">B:</emphasis>
ventral view).
</paragraph>
</caption>
<paragraph id="911A3683FFEDFF8E30F2FEF5FD5CC1AA" blockId="8.[133,771,214,1897]" pageId="8" pageNumber="255">Arms meaty, forearm length shorter than the hand length; fingers free, tips ending in disk, finger I disk slightly dilated, all disks with circum-marginal grooves except finger I, relative length of fingers I&lt;II&lt;IV&lt;III; sub-articular tubercles prominent, circular; prepollex oval.</paragraph>
<paragraph id="911A3683FFEDFF8E30F2FE39FF69C2C2" blockId="8.[133,771,214,1897]" pageId="8" pageNumber="255">Hindlimbs long and strong, tibia longer than both femur and foot (TiL&gt;FOL&gt;ShL); toes long, tips of all toes dilated into disks with circum-marginal grooves, toes completely webbed; sub-articular tubercles domed shaped; inner metatarsal tubercle oval shaped, outer absent.</paragraph>
<paragraph id="911A3683FFEDFF8E30F2FD41FE42C36E" blockId="8.[133,771,214,1897]" pageId="8" pageNumber="255">The dorsal skin is smooth, slightly granular towards dorso-lateral sides; the supra-tympanic fold is strong; lateral and ventral skin is smooth.</paragraph>
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Colouration of the
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:
</emphasis>
(
<emphasis id="A3D1EA91FFEDFF8E31B3FCFCFD9AC352" box="[492,551,790,816]" italics="true" pageId="8" pageNumber="255">in life</emphasis>
) Dorsal color olive-green, spotted with large, irregular shaped dark-brown spots, brown spots enclosing a number of small olive-green dots; laterally same as dorsally; ventral color dark, speckled with lighter dots and dashes. Dorso-laterally granular along with white-tipped minute tubercles. Tibia banded; thighs mottled with dark brown and olive-green (
<figureCitation id="099E2A06FFEDFF8E30D2FBF9FF56C44F" box="[141,235,1042,1069]" captionStart="Figure 5" captionStartId="6.[134,210,1884,1910]" captionTargetBox="[144,1461,1000,1857]" captionTargetId="figure-370@6.[144,1461,1000,1857]" captionTargetPageId="6" captionText="Figure 5. Live colour image of holotype of Amolops tawang sp. nov. (Photo: Bikramjit Sinha)." figureDoi="http://doi.org/10.5281/zenodo.13185577" httpUri="https://zenodo.org/record/13185577/files/figure.png" pageId="8" pageNumber="255">Figure 5</figureCitation>
).
<emphasis id="A3D1EA91FFEDFF8E30A0FBF8FE20C44E" box="[255,413,1043,1068]" italics="true" pageId="8" pageNumber="255">In preservative</emphasis>
same as in live condition, except that all the olive-green colours are faded (
<figureCitation id="099E2A06FFEDFF8E320CFBDDFD09C432" box="[595,692,1078,1104]" captionStart="Figure 6" captionStartId="7.[160,236,1884,1910]" captionTargetBox="[160,1499,1224,1829]" captionTargetId="figure-426@7.[157,1502,1219,1857]" captionTargetPageId="7" captionText="Figure 6. Holotype image of Amolops tawang sp. nov. in preserved condition (A: dorsal view; B: ventral view)." figureDoi="http://doi.org/10.5281/zenodo.13185579" httpUri="https://zenodo.org/record/13185579/files/figure.png" pageId="8" pageNumber="255">Figure 6</figureCitation>
).
</paragraph>
<paragraph id="911A3683FFEDFF8E30F2FBB1FD03C4DE" blockId="8.[133,771,214,1897]" pageId="8" pageNumber="255">
<emphasis id="A3D1EA91FFEDFF8E30F2FBB1FE25C416" box="[173,408,1114,1140]" italics="true" pageId="8" pageNumber="255">
Condition of the
<typeStatus id="4E1E8821FFEDFF8E313BFBB0FE2FC416" box="[356,402,1115,1140]" pageId="8" pageNumber="255">type</typeStatus>
:
</emphasis>
The
<typeStatus id="4E1E8821FFEDFF8E3191FBB1FE43C416" box="[462,510,1114,1140]" pageId="8" pageNumber="255">type</typeStatus>
is in good condition. A ventral incision was made in the
<typeStatus id="4E1E8821FFEDFF8E31ADFB95FD9FC4FA" box="[498,546,1150,1176]" pageId="8" pageNumber="255">type</typeStatus>
specimen to extract liver tissue, which was used in the molecular study.
</paragraph>
<paragraph id="911A3683FFEDFF8E30F2FB2DFEA4C5DA" blockId="8.[133,771,214,1897]" pageId="8" pageNumber="255">
<emphasis id="A3D1EA91FFEDFF8E30F2FB2DFED9C482" box="[173,356,1222,1248]" italics="true" pageId="8" pageNumber="255">Natural history:</emphasis>
The lone specimen of
<taxonomicName id="56A54D00FFEDFF8E32D4FB2DFCBEC482" authorityName="Saikia &amp; Laskar &amp; Dinesh &amp; Shabnam &amp; Sinha" authorityYear="2022" box="[651,771,1222,1248]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="tawang" status="sp. nov.">
<emphasis id="A3D1EA91FFEDFF8E32D4FB2DFCBEC482" box="[651,771,1222,1248]" italics="true" pageId="8" pageNumber="255">A. tawang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8E257EAFFEDFF8E30DAFB01FF55C567" box="[133,232,1258,1285]" pageId="8" pageNumber="255" rank="species">
<emphasis id="A3D1EA91FFEDFF8E30DAFB01FF5FC566" bold="true" box="[133,226,1258,1284]" pageId="8" pageNumber="255">sp. nov</emphasis>
.
</taxonomicNameLabel>
was collected during the mid-day from a torrential hill stream with low water volume, which was characterised by large boulders. The specimen was resting under a boulder in a shallow trickle of water. A single specimen of
<taxonomicName id="56A54D00FFEDFF8E314DFA90FE6DC5F6" baseAuthorityName="Gunther" baseAuthorityYear="1860" box="[274,464,1402,1428]" class="Amphibia" family="Dicroglossidae" genus="Nanorana" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="liebigii">
<emphasis id="A3D1EA91FFEDFF8E314DFA90FE6DC5F6" box="[274,464,1402,1428]" italics="true" pageId="8" pageNumber="255">Nanorana liebigii</emphasis>
</taxonomicName>
was also collected from the same habitat.
</paragraph>
<paragraph id="911A3683FFEDFF8E30F2FA29FE93C646" blockId="8.[133,771,214,1897]" pageId="8" pageNumber="255">
<emphasis id="A3D1EA91FFEDFF8E30F2FA29FE87C5BE" box="[173,314,1474,1500]" italics="true" pageId="8" pageNumber="255">Distribution:</emphasis>
Currently, this species is known only from its
<typeStatus id="4E1E8821FFEDFF8E30AAFA0CFE98C663" box="[245,293,1511,1537]" pageId="8" pageNumber="255">type</typeStatus>
locality in Tawang District,
<collectingRegion id="5361F861FFEDFF8E32D3FA0DFF5CC646" country="India" name="Arunachal Pradesh" pageId="8" pageNumber="255">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="E9B27613FFEDFF8E30B3F9E1FE97C646" box="[236,298,1546,1572]" name="India" pageId="8" pageNumber="255">India</collectingCountry>
.
</paragraph>
<paragraph id="911A3683FFEDFF8E30F2F9C5FD32C60F" blockId="8.[133,771,214,1897]" pageId="8" pageNumber="255">
<emphasis id="A3D1EA91FFEDFF8E30F2F9C5FEF9C62A" box="[173,324,1582,1608]" italics="true" pageId="8" pageNumber="255">Comparisons:</emphasis>
The new species is compared with the other species of
<taxonomicName id="56A54D00FFEDFF8E3166F9B9FE57C60E" baseAuthorityName="Jiang" baseAuthorityYear="1983" box="[313,490,1618,1644]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="viridimaculatus">
<emphasis id="A3D1EA91FFEDFF8E3166F9B9FE57C60E" box="[313,490,1618,1644]" italics="true" pageId="8" pageNumber="255">Viridimaculatus</emphasis>
</taxonomicName>
species group.
</paragraph>
<paragraph id="911A3683FFEDFF8E30F2F99CFBD4C62A" blockId="8.[133,771,214,1897]" lastBlockId="8.[835,1473,214,1897]" pageId="8" pageNumber="255">
From the Indian
<taxonomicName id="56A54D00FFEDFF8E31CCF99DFDF9C6F2" baseAuthorityName="Jiang" baseAuthorityYear="1983" box="[403,580,1654,1680]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="viridimaculatus">
<emphasis id="A3D1EA91FFEDFF8E31CCF99DFDF9C6F2" box="[403,580,1654,1680]" italics="true" pageId="8" pageNumber="255">Viridimaculatus</emphasis>
</taxonomicName>
species group,
<taxonomicName id="56A54D00FFEDFF8E30DAF971FEFCC6D6" authorityName="Saikia &amp; Laskar &amp; Dinesh &amp; Shabnam &amp; Sinha" authorityYear="2022" box="[133,321,1690,1716]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="tawang" status="sp. nov.">
<emphasis id="A3D1EA91FFEDFF8E30DAF971FEFCC6D6" box="[133,321,1690,1716]" italics="true" pageId="8" pageNumber="255">Amolops tawang</emphasis>
</taxonomicName>
<emphasis id="A3D1EA91FFEDFF8E3112F971FE17C6D6" bold="true" box="[333,426,1690,1716]" pageId="8" pageNumber="255">
<taxonomicNameLabel id="B8E257EAFFEDFF8E3112F971FE17C6D6" box="[333,426,1690,1716]" pageId="8" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="56A54D00FFEDFF8E3212F971FD68C6D6" baseAuthorityName="Gunther" baseAuthorityYear="1876" box="[589,725,1690,1716]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="formosus">
<emphasis id="A3D1EA91FFEDFF8E3212F971FD68C6D6" box="[589,725,1690,1716]" italics="true" pageId="8" pageNumber="255">A. formosus</emphasis>
</taxonomicName>
(in parentheses) in having adult male larger (vs. smaller), tympanum small, distinct and depressed (vs. about the size of finger III disk), inner-metatarsal tubercles distinct (vs. indistinct), circum-marginal groove on finger I absent (vs. present), outer metatarsal tubercle absent (vs. present), dorsal color olive-green, spotted with large, irregular shaped dark-brown spots (vs. bright green with large dark spots), limbs with incomplete bands (vs. uniformly banded); from
<taxonomicName id="56A54D00FFEDFF8E3402FEA9FAB5C13E" baseAuthorityName="Boulenger" baseAuthorityYear="1888" box="[1117,1288,322,348]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="himalayanus">
<emphasis id="A3D1EA91FFEDFF8E3402FEA9FAB5C13E" box="[1117,1288,322,348]" italics="true" pageId="8" pageNumber="255">A. himalayanus</emphasis>
</taxonomicName>
(in parentheses) in having larger male SVL&gt;
<quantity id="565D9B66FFEDFF8E34D6FE8DFB61C1E3" box="[1161,1244,358,385]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="255" unit="mm" value="82.0">82 mm</quantity>
(vs. smaller &lt;
<quantity id="565D9B66FFEDFF8E35DFFE8DFCC9C1C6" metricMagnitude="-2" metricUnit="m" metricValue="7.45" metricValueMax="7.5" metricValueMin="7.4" pageId="8" pageNumber="255" unit="mm" value="74.5" valueMax="75.0" valueMin="74.0">74-75 mm</quantity>
), nostril equidistant (vs. nearer to eye), tympanum small, distinct and depressed (vs. slightly visible), visible sac faintly visible (vs. externally visible vocal sac present), tibia 2/3
<superScript id="66D09BCBFFEDFF8E3457FE1EFBABC261" attach="left" box="[1032,1046,501,515]" fontSize="6" pageId="8" pageNumber="255">rd</superScript>
of SVL (vs. almost equal), tibio-tarsal articulation reached up to the snout (vs. beyond the tip of the snout), dorsal color olive-green, spotted with large, irregular shaped dark-brown spots (vs. olive with darker spots); from
<taxonomicName id="56A54D00FFEDFF8E3381FD6DFBC0C2C2" authorityName="Biju, Mahony &amp; Kamei" authorityYear="2010" box="[990,1149,646,672]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="nidorbellus">
<emphasis id="A3D1EA91FFEDFF8E3381FD6DFBC0C2C2" box="[990,1149,646,672]" italics="true" pageId="8" pageNumber="255">A. nidorbellus</emphasis>
</taxonomicName>
(in parentheses) in having larger male SVL&gt;
<quantity id="565D9B66FFEDFF8E3457FD41FBEAC2A6" box="[1032,1111,682,708]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="255" unit="mm" value="82.0">82 mm</quantity>
(vs. smaller &lt;
<quantity id="565D9B66FFEDFF8E34B0FD41FAFDC2A6" box="[1263,1344,682,708]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="8" pageNumber="255" unit="mm" value="77.0">77 mm</quantity>
), IUEUEW (vs. IUE&gt;EUW), pineal ocellus distinct (vs. barely visible), vocal sac faintly visible (vs. externally visible vocal sacs absent), dorsal color olive-green, spotted with large, irregular shaped dark-brown spots (vs. brown with small, irregular cobalt green spots), ventral color dark, speckled with lighter dots and dashes (vs. pale brown with scattered small, white spots); from
<taxonomicName id="56A54D00FFEDFF8E34B3FC4DFA7DC3A2" baseAuthorityName="Jiang" baseAuthorityYear="1983" box="[1260,1472,934,960]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="viridimaculatus">
<emphasis id="A3D1EA91FFEDFF8E34B3FC4DFA7DC3A2" box="[1260,1472,934,960]" italics="true" pageId="8" pageNumber="255">A. viridimaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8E257EAFFEDFF8E331CFC21FC6CC386" box="[835,977,970,996]" pageId="8" pageNumber="255" sensu="stricto">sensu stricto</taxonomicNameLabel>
(in parentheses) in having larger male SVL&gt;
<quantity id="565D9B66FFEDFF8E330DFC05FC19C46A" box="[850,932,1006,1032]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="255" unit="mm" value="82.0">82 mm</quantity>
(vs. smaller &lt;
<quantity id="565D9B66FFEDFF8E3419FC05FB26C46A" box="[1094,1179,1006,1032]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="8" pageNumber="255" unit="mm" value="77.0">77 mm</quantity>
), head wider (vs. longer), tympanum distinct (vs. indistinct), outer metatarsal tubercle absent (vs. present), limbs not distinctly banded (vs. distinctly banded), dorsal color olive-green, spotted with large, irregular shaped dark-brown spots (vs. maroon red with large green blotches); from
<taxonomicName id="56A54D00FFEDFF8E3567FB49FA7DC4DE" authorityName="Saikia &amp; Laskar &amp; Dinesh &amp; Shabnam &amp; Sinha" authorityYear="2022" box="[1336,1472,1186,1212]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="chanakya" status="sp. nov.">
<emphasis id="A3D1EA91FFEDFF8E3567FB49FA7DC4DE" box="[1336,1472,1186,1212]" italics="true" pageId="8" pageNumber="255">A. chanakya</emphasis>
</taxonomicName>
<emphasis id="A3D1EA91FFEDFF8E331CFB2DFC21C482" bold="true" box="[835,924,1222,1248]" pageId="8" pageNumber="255">
<taxonomicNameLabel id="B8E257EAFFEDFF8E331CFB2DFC21C482" box="[835,924,1222,1248]" pageId="8" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(in parentheses) in having larger male SVL&gt;
<quantity id="565D9B66FFEDFF8E35FAFB2DFCCCC566" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="255" unit="mm" value="82.0">82 mm</quantity>
(vs. smaller &lt;
<quantity id="565D9B66FFEDFF8E3451FB01FBDFC566" box="[1038,1122,1258,1284]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="8" pageNumber="255" unit="mm" value="77.0">77 mm</quantity>
), nostril equidistant (vs. nearer to the eye), IUE=UEW (vs. IUE&lt;UEW), tympanum extremely depressed and small (vs. larger), TYD = ½TE (vs. TYD ≈ TE), external vocal sac faintly visible (vs. distinctly visible), tibio-tarsal articulation reaches up to snout (vs. nostril), dorsal color olive-green, spotted with large, irregular shaped dark-brown spots (vs. dorsum dull brick-red, spotted with irregular cocoa-brown spots), ventral color dark, speckled with lighter dots and dashes (vs. ventral cocoa-brown).
</paragraph>
<paragraph id="911A3683FFEDFF8F3334F9B8FD64C5BE" blockId="8.[835,1473,214,1897]" lastBlockId="9.[160,798,214,1788]" lastPageId="9" lastPageNumber="256" pageId="8" pageNumber="255">
From the
<taxonomicName id="56A54D00FFEDFF8E33A7F9B9FB14C60E" baseAuthorityName="Jiang" baseAuthorityYear="1983" box="[1016,1193,1618,1644]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="viridimaculatus">
<emphasis id="A3D1EA91FFEDFF8E33A7F9B9FB14C60E" box="[1016,1193,1618,1644]" italics="true" pageId="8" pageNumber="255">Viridimaculatus</emphasis>
</taxonomicName>
species group found outside
<collectingCountry id="E9B27613FFEDFF8E33FBF99DFC5CC6F2" box="[932,993,1654,1680]" name="India" pageId="8" pageNumber="255">India</collectingCountry>
,
<taxonomicName id="56A54D00FFEDFF8E33AAF99DFB09C6F2" authorityName="Saikia &amp; Laskar &amp; Dinesh &amp; Shabnam &amp; Sinha" authorityYear="2022" box="[1013,1204,1654,1680]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="tawang" status="sp. nov.">
<emphasis id="A3D1EA91FFEDFF8E33AAF99DFB09C6F2" box="[1013,1204,1654,1680]" italics="true" pageId="8" pageNumber="255">Amolops tawang</emphasis>
</taxonomicName>
<emphasis id="A3D1EA91FFEDFF8E349BF99DFA99C6F2" bold="true" box="[1220,1316,1654,1680]" pageId="8" pageNumber="255">
<taxonomicNameLabel id="B8E257EAFFEDFF8E349BF99DFA99C6F2" box="[1220,1316,1654,1680]" pageId="8" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="56A54D00FFEDFF8E331CF971FC52C6D6" box="[835,1007,1690,1716]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="beibengensis">
<emphasis id="A3D1EA91FFEDFF8E331CF971FC52C6D6" box="[835,1007,1690,1716]" italics="true" pageId="8" pageNumber="255">A. beibengensis</emphasis>
</taxonomicName>
(in parentheses) in having larger male SVL&gt;
<quantity id="565D9B66FFEDFF8E33D3F955FC62C6BB" box="[908,991,1726,1753]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="255" unit="mm" value="82.0">82 mm</quantity>
(vs. &lt;
<quantity id="565D9B66FFEDFF8E3476F955FB3DC6BA" box="[1065,1152,1726,1752]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="8" pageNumber="255" unit="mm" value="76.0">76 mm</quantity>
), head wider than long (vs. HL=HW), tibio-tarsal articulation reached up to the snout (vs. between nose and eye), vocal sac faintly visible (vs. absent), dorsal color olive-green, spotted with large, irregular shaped dark-brown spots (vs. dorsum brown with green spots); from
<taxonomicName id="56A54D00FFECFF8F31E9FF3DFDE3C092" box="[438,606,214,240]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="9" pageNumber="256" phylum="Chordata" rank="species" species="gyirongensis">
<emphasis id="A3D1EA91FFECFF8F31E9FF3DFDE3C092" box="[438,606,214,240]" italics="true" pageId="9" pageNumber="256">A. gyirongensis</emphasis>
</taxonomicName>
(in parentheses) in having larger male SVL&gt;
<quantity id="565D9B66FFECFF8F3180FF11FD8CC176" box="[479,561,250,276]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="9" pageNumber="256" unit="mm" value="82.0">82 mm</quantity>
(vs. &lt;
<quantity id="565D9B66FFECFF8F3229FF11FD76C176" box="[630,715,250,276]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="9" pageNumber="256" unit="mm" value="64.0">64 mm</quantity>
), tibio-tarsal articulation reached up to the snout (vs. nostril), vocal sac faintly visible (vs. absent); dorsal color olive-green, spotted with large, irregular shaped dark-brown spots (vs. bright green with dark brown irregular large spots); from
<taxonomicName id="56A54D00FFECFF8F3173FE45FE0EC1AA" baseAuthorityName="Smith" baseAuthorityYear="1940" box="[300,435,430,456]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="9" pageNumber="256" phylum="Chordata" rank="species" species="kaulbacki">
<emphasis id="A3D1EA91FFECFF8F3173FE45FE0EC1AA" box="[300,435,430,456]" italics="true" pageId="9" pageNumber="256">A. kaulbacki</emphasis>
</taxonomicName>
(in parentheses) in having larger male SVL&gt;
<quantity id="565D9B66FFECFF8F3174FE39FEC3C18E" box="[299,382,466,492]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="9" pageNumber="256" unit="mm" value="82.0">82 mm</quantity>
(vs. up to
<quantity id="565D9B66FFECFF8F325BFE39FDE7C18E" box="[516,602,466,492]" metricMagnitude="-2" metricUnit="m" metricValue="7.2" pageId="9" pageNumber="256" unit="mm" value="72.0">72 mm</quantity>
), IN&gt;IUE=UEW (vs. IN&gt;IUE&gt;UEW), tibio-tarsal articulation reached up to the snout (vs. beyond snout), vocal sac faintly visible (vs. absent), dorsal color olive-green, spotted with large, irregular shaped dark-brown spots (vs. green with black spots and marbling); from
<taxonomicName id="56A54D00FFECFF8F3198FD6DFDD9C2C2" baseAuthorityName="Andersson" baseAuthorityYear="1939" box="[455,612,646,672]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="9" pageNumber="256" phylum="Chordata" rank="species" species="longimanus">
<emphasis id="A3D1EA91FFECFF8F3198FD6DFDD9C2C2" box="[455,612,646,672]" italics="true" pageId="9" pageNumber="256">A. longimanus</emphasis>
</taxonomicName>
(in parentheses) in not having distinctive dorsolateral fold (vs. present), absence of parotid-like swelling above tympanum (vs. present); from
<taxonomicName id="56A54D00FFECFF8F3114FD19FE54C36E" authorityName="Li &amp; Rao in Zhao, Rao, Lu &amp; Dong" authorityYear="2005" box="[331,489,754,780]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="9" pageNumber="256" phylum="Chordata" rank="species" species="medogensis">
<emphasis id="A3D1EA91FFECFF8F3114FD19FE54C36E" box="[331,489,754,780]" italics="true" pageId="9" pageNumber="256">A. medogensis</emphasis>
</taxonomicName>
(in parentheses) in having smaller male SVL of
<quantity id="565D9B66FFECFF8F31D9FCFDFE68C352" box="[390,469,790,816]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="9" pageNumber="256" unit="mm" value="82.0">82 mm</quantity>
(vs.
<quantity id="565D9B66FFECFF8F3257FCFDFDE7C352" box="[520,602,790,817]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="9" pageNumber="256" unit="mm" value="95.0">95 mm</quantity>
), vocal sac faintly visible (vs. absent), dorsal color olive-green, spotted with large, irregular shaped dark-brown spots (vs. olive green with brown markings/dark brown with small yellow-green spots); from
<taxonomicName id="56A54D00FFECFF8F31D5FC4DFD93C3A2" authorityName="Qi, Zhou, Lyu, Lu &amp; Li" authorityYear="2019" box="[394,558,934,960]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="9" pageNumber="256" phylum="Chordata" rank="species" species="pallasitatus">
<emphasis id="A3D1EA91FFECFF8F31D5FC4DFD93C3A2" box="[394,558,934,960]" italics="true" pageId="9" pageNumber="256">A. pallasitatus</emphasis>
</taxonomicName>
(in parentheses) in having tibio-tarsal articulation reached up to the snout (vs. posterior eye corner), dorsal color olive-green, spotted with large, irregular shaped dark-brown spots (vs. yellow green with irregular dark brown blotches); from
<taxonomicName id="56A54D00FFECFF8F30FFFBB1FEE4C416" authorityName="Orlov &amp; Ho" authorityYear="2007" box="[160,345,1114,1140]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="9" pageNumber="256" phylum="Chordata" rank="species" species="splendissimus">
<emphasis id="A3D1EA91FFECFF8F30FFFBB1FEE4C416" box="[160,345,1114,1140]" italics="true" pageId="9" pageNumber="256">A. splendissimus</emphasis>
</taxonomicName>
(in parentheses) in having larger male SVL&gt;
<quantity id="565D9B66FFECFF8F30B8FB95FE85C4FA" box="[231,312,1150,1176]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="9" pageNumber="256" unit="mm" value="82.0">82 mm</quantity>
(vs.
<quantity id="565D9B66FFECFF8F3130FB95FE7FC4FA" box="[367,450,1150,1176]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="9" pageNumber="256" unit="mm" value="76.0">76 mm</quantity>
), IUE=UWD (vs. IUE&lt;UEW), supra-tympanic fold strong (vs. poorly developed), vocal sac faintly visible (vs. absent), dorsal color olive-green, spotted with large, irregular shaped dark-brown spots (vs. black-brown spangled thickly with bright yellow spots); from
<taxonomicName id="56A54D00FFECFF8F30BBFAD9FE31C52E" box="[228,396,1330,1356]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="9" pageNumber="256" phylum="Chordata" rank="species" species="wangyufani">
<emphasis id="A3D1EA91FFECFF8F30BBFAD9FE31C52E" box="[228,396,1330,1356]" italics="true" pageId="9" pageNumber="256">A. wangyufani</emphasis>
</taxonomicName>
(in parentheses) in having larger male SVL&gt;
<quantity id="565D9B66FFECFF8F317DFABDFECFC513" box="[290,370,1366,1393]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="9" pageNumber="256" unit="mm" value="82.0">82 mm</quantity>
(vs. &lt;
<quantity id="565D9B66FFECFF8F31EBFABDFDBAC512" box="[436,519,1366,1392]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="256" unit="mm" value="70.0">70 mm</quantity>
), tibio-tarsal articulation reached up to the snout (vs. beyond snout), dorsal color olive-green, spotted with large, irregular shaped dark-brown spots (vs. brown with sparse green patches).
</paragraph>
<paragraph id="911A3683FFECFF8F3097FA0DFF58C69E" blockId="9.[160,798,214,1788]" pageId="9" pageNumber="256">
<emphasis id="A3D1EA91FFECFF8F3097FA0DFEFEC662" box="[200,323,1510,1536]" italics="true" pageId="9" pageNumber="256">Etymology:</emphasis>
The species epithet is a toponym named after the district Tawang, from where the
<typeStatus id="4E1E8821FFECFF8F32ECF9E0FD5EC647" box="[691,739,1547,1573]" pageId="9" pageNumber="256">type</typeStatus>
was collected. Located in the westernmost part of
<collectingRegion id="5361F861FFECFF8F32F8F9C5FF41C60E" country="India" name="Arunachal Pradesh" pageId="9" pageNumber="256">Arunachal Pradesh</collectingRegion>
, Tawang is renowned for Tawang Monastery, which is the largest in
<collectingCountry id="E9B27613FFECFF8F31CAF99DFE6DC6F2" box="[405,464,1654,1680]" name="India" pageId="9" pageNumber="256">India</collectingCountry>
as well as being the birthplace of the 6
<superScript id="66D09BCBFFECFF8F315FF972FEB3C6C5" attach="left" box="[256,270,1689,1703]" fontSize="6" pageId="9" pageNumber="256">th</superScript>
Dalai Lama, Tsangyang Gyatso. The species epithet is treated as a noun in apposition to the generic name.
</paragraph>
<paragraph id="911A3683FFECFF8F30FFF8C1FCA0C70A" blockId="9.[160,797,1834,1896]" pageId="9" pageNumber="256">
<emphasis id="A3D1EA91FFECFF8F30FFF8C1FE71C726" box="[160,460,1834,1860]" italics="true" pageId="9" pageNumber="256">Suggested Common Name:</emphasis>
We suggest
<emphasis id="A3D1EA91FFECFF8F323EF8C1FF6CC705" bold="true" pageId="9" pageNumber="256">Tawang cascade frog</emphasis>
as a common English name and तवांग झरना मढक
</paragraph>
<paragraph id="911A3683FFECFF8F3302FF0EFA8CC09C" blockId="9.[861,1329,228,255]" box="[861,1329,228,255]" pageId="9" pageNumber="256">as a common Hindi name for this species.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,367 @@
<document id="BEF40B7474C4A604A9D36A4C531231E5" ID-DOI="10.26515/rzsi/v122/i3/2022/169417" ID-ISSN="2581-8686" ID-Zenodo-Dep="13185566" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722611788751" checkinUser="felipe" docAuthor="Saikia, Bhaskar, Laskar, Mostaque A., Dinesh, K. P., Shabnam, A. &amp; Sinha, Bikramjit" docDate="2022" docId="190C8795FFE6FF813302F901FCBCC352" docLanguage="en" docName="RecZoolSurvIndia.122.3.247-266.pdf" docOrigin="Records of the Zoological Survey of India 122 (3)" docStyle="DocumentStyle:30C78A88DFAFB68AE5A0604D5A6B2FFB.3:RecZoolSurvIndia.2017-.journal_article" docStyleId="30C78A88DFAFB68AE5A0604D5A6B2FFB" docStyleName="RecZoolSurvIndia.2017-.journal_article" docStyleVersion="3" docTitle="Amolops chanakya Saikia, Laskar, Dinesh, Shabnam &amp; Sinha, 2022, sp. nov." docType="treatment" docUuid="1CF810F3-5B54-4D2C-9263-1EEC222A641F" docUuidSource="ZooBank" docVersion="2" lastPageNumber="254" masterDocId="E535FFEDFFE5FF86305FFFEBFFBDC062" masterDocTitle="Description of two new species of Amolops (Anura: Ranidae) from Arunachal Pradesh, Northeast India under the morphological Viridimaculatus species group" masterLastPageNumber="266" masterPageNumber="247" pageNumber="250" updateTime="1722662603149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A41F1E5EBB61550CD643BB399A89395B">Description of two new species of Amolops (Anura: Ranidae) from Arunachal Pradesh, Northeast India under the morphological Viridimaculatus species group</mods:title>
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<mods:namePart id="3A84DA11B074239D4D2019367A6D7856">Saikia, Bhaskar</mods:namePart>
<mods:affiliation id="B37C89BB915486DC43326947820825DB">Zoological Survey of India, North Eastern Regional Centre, Shillong - 793003, Meghalaya, India; Email: bj. sinha @ gov. in</mods:affiliation>
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<mods:namePart id="71E4EA0BE02CDE1B0D59259284628558">Laskar, Mostaque A.</mods:namePart>
<mods:affiliation id="1BA657C2828A3B03A0494D8AEDCB7669">Department of Biotechnology, St. Anthonys College, Shillong - 793001, Meghalaya, India</mods:affiliation>
</mods:name>
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<mods:namePart id="0CB5D8DBE274C5E735F71A0E9754B7E2">Dinesh, K. P.</mods:namePart>
<mods:affiliation id="05A18E8E428A80CF1AC78C04D971C1D7">Zoological Survey of India, Western Regional Centre, Vidyanagar, Pune - 411044, Maharashtra, India</mods:affiliation>
</mods:name>
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<mods:namePart id="0BA4788999640456A75C4550A44B2B46">Shabnam, A.</mods:namePart>
<mods:affiliation id="1D6477B3D6894B68A5A0F274AF265193">Zoological Survey of India, Western Regional Centre, Vidyanagar, Pune - 411044, Maharashtra, India</mods:affiliation>
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</mods:role>
<mods:namePart id="CE15E7E023DE1CF75DCC8720F30F4701">Sinha, Bikramjit</mods:namePart>
<mods:affiliation id="39B4880E9CA15F6E3C21A8007045896A">Zoological Survey of India, North Eastern Regional Centre, Shillong - 793003, Meghalaya, India; Email: bj. sinha @ gov. in</mods:affiliation>
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<subSubSection id="D9BF6508FFE6FF853302F901FB34C74B" pageId="3" pageNumber="250" type="nomenclature">
<paragraph id="911A3683FFE6FF853302F901FB2DC767" blockId="3.[861,1499,1770,1906]" box="[861,1168,1770,1797]" pageId="3" pageNumber="250">
<heading id="CA5281EFFFE6FF853302F901FB2DC767" box="[861,1168,1770,1797]" fontSize="11" level="2" pageId="3" pageNumber="250" reason="4">
<taxonomicName id="56A54D00FFE6FF853302F901FB8BC767" authorityName="Saikia &amp; Laskar &amp; Dinesh &amp; Shabnam &amp; Sinha" authorityYear="2022" box="[861,1078,1770,1797]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="3" pageNumber="250" phylum="Chordata" rank="species" species="chanakya" status="sp. nov.">
<emphasis id="A3D1EA91FFE6FF853302F901FB8BC767" bold="true" box="[861,1078,1770,1797]" italics="true" pageId="3" pageNumber="250">Amolops chanakya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8E257EAFFE6FF853463F900FB2DC767" box="[1084,1168,1771,1797]" pageId="3" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="911A3683FFE6FF853302F8E4FB34C74B" blockId="3.[861,1499,1770,1906]" box="[861,1161,1807,1833]" pageId="3" pageNumber="250">(Chanakyas Cascade Frog)</paragraph>
</subSubSection>
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<paragraph id="911A3683FFE6FF853302F8D8FB20C710" blockId="3.[861,1499,1770,1906]" pageId="3" pageNumber="250">
Zoobank ID:
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urn:lsid:zoobank.org:act:
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<caption id="C5DA660BFFE1FF8230DAFBBEFB08C40D" ID-DOI="http://doi.org/10.5281/zenodo.13185573" ID-Zenodo-Dep="13185573" box="[133,1205,1109,1135]" httpUri="https://zenodo.org/record/13185573/files/figure.png" pageId="4" pageNumber="251" startId="4.[133,209,1109,1135]" targetBox="[165,1441,213,1082]" targetPageId="4" targetType="figure">
<paragraph id="911A3683FFE1FF8230DAFBBEFB08C40D" blockId="4.[133,1205,1109,1135]" box="[133,1205,1109,1135]" pageId="4" pageNumber="251">
<emphasis id="A3D1EA91FFE1FF8230DAFBBEFF50C40D" bold="true" box="[133,237,1109,1135]" pageId="4" pageNumber="251">Figure 3.</emphasis>
Live colour image of holotype of
<taxonomicName id="56A54D00FFE1FF823235FBBEFC84C40D" authorityName="Saikia &amp; Laskar &amp; Dinesh &amp; Shabnam &amp; Sinha" authorityYear="2022" box="[618,825,1109,1135]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="4" pageNumber="251" phylum="Chordata" rank="species" species="chanakya" status="sp. nov.">
<emphasis id="A3D1EA91FFE1FF823235FBBEFC84C40D" box="[618,825,1109,1135]" italics="true" pageId="4" pageNumber="251">Amolops chanakya</emphasis>
</taxonomicName>
<emphasis id="A3D1EA91FFE1FF823360FBBEFC2BC40D" bold="true" box="[831,918,1109,1135]" pageId="4" pageNumber="251">
<taxonomicNameLabel id="B8E257EAFFE1FF823360FBBEFC2BC40D" box="[831,918,1109,1135]" pageId="4" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Photo: Bikramjit Sinha).
</paragraph>
</caption>
<caption id="C5DA660BFFE1FF8230DAF8A7FAC9C704" ID-DOI="http://doi.org/10.5281/zenodo.13185575" ID-Zenodo-Dep="13185575" box="[133,1396,1868,1895]" httpUri="https://zenodo.org/record/13185575/files/figure.png" pageId="4" pageNumber="251" startId="4.[133,209,1868,1894]" targetBox="[133,1472,1207,1815]" targetPageId="4" targetType="figure">
<paragraph id="911A3683FFE1FF8230DAF8A7FAC9C704" blockId="4.[133,1396,1868,1895]" box="[133,1396,1868,1895]" pageId="4" pageNumber="251">
<emphasis id="A3D1EA91FFE1FF8230DAF8A7FF50C704" bold="true" box="[133,237,1868,1894]" pageId="4" pageNumber="251">Figure 4.</emphasis>
Holotype image of
<taxonomicName id="56A54D00FFE1FF823190F8A7FD23C704" authorityName="Saikia &amp; Laskar &amp; Dinesh &amp; Shabnam &amp; Sinha" authorityYear="2022" box="[463,670,1868,1894]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="4" pageNumber="251" phylum="Chordata" rank="species" species="chanakya" status="sp. nov.">
<emphasis id="A3D1EA91FFE1FF823190F8A7FD23C704" box="[463,670,1868,1894]" italics="true" pageId="4" pageNumber="251">Amolops chanakya</emphasis>
</taxonomicName>
<emphasis id="A3D1EA91FFE1FF8232FBF8A6FD46C705" bold="true" box="[676,763,1869,1895]" pageId="4" pageNumber="251">
<taxonomicNameLabel id="B8E257EAFFE1FF8232FBF8A6FD46C705" box="[676,763,1869,1895]" pageId="4" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in preserved condition (
<emphasis id="A3D1EA91FFE1FF823451F8A7FB94C704" bold="true" box="[1038,1065,1868,1894]" pageId="4" pageNumber="251">A:</emphasis>
dorsal view;
<emphasis id="A3D1EA91FFE1FF8234E3F8A6FB68C705" bold="true" box="[1212,1237,1869,1895]" pageId="4" pageNumber="251">B:</emphasis>
ventral view).
</paragraph>
</caption>
<paragraph id="911A3683FFE0FF833097FF3DFDC9C1E2" blockId="5.[160,798,214,1897]" pageId="5" pageNumber="252">
<materialsCitation id="21CD3CDEFFE0FF833097FF3DFDC9C1E2" collectingDate="2018-03-13" collectionCode="V, NERC" collectorName="Bikramjit Sinha" country="India" county="West Kameng district" elevation="1794" latitude="27.422089" location="Chakpa Village" longLatPrecision="1" longitude="92.21903" municipality="Dirang" pageId="5" pageNumber="252" specimenCode="ZSI/1771" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<emphasis id="A3D1EA91FFE0FF833097FF3DFE96C092" box="[200,299,214,240]" italics="true" pageId="5" pageNumber="252">
<typeStatus id="4E1E8821FFE0FF833097FF3DFE96C092" box="[200,299,214,240]" pageId="5" pageNumber="252" type="holotype">Holotype</typeStatus>
</emphasis>
:
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/A/
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/
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, adult male (
<figureCitation id="099E2A06FFE0FF83328CFF3DFF12C176" captionStart="Figure 3" captionStartId="4.[133,209,1109,1135]" captionTargetBox="[165,1441,213,1082]" captionTargetId="figure-28@4.[165,1441,213,1082]" captionTargetPageId="4" captionText="Figure 3. Live colour image of holotype of Amolops chanakya sp. nov. (Photo: Bikramjit Sinha)." figureDoi="http://doi.org/10.5281/zenodo.13185573" httpUri="https://zenodo.org/record/13185573/files/figure.png" pageId="5" pageNumber="252">Figure 3</figureCitation>
), collected by
<collectorName id="3C505355FFE0FF83310AFF11FDBBC176" box="[341,518,250,276]" pageId="5" pageNumber="252">Bikramjit Sinha</collectorName>
and party on
<date id="E51B1043FFE0FF8332F5FF11FF6BC15A" pageId="5" pageNumber="252" value="2018-03-13">
<collectingDate id="F55FE9ABFFE0FF8332F5FF11FF6BC15A" pageId="5" pageNumber="252" value="2018-03-13">March 13, 2018</collectingDate>
</date>
from
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(
<geoCoordinate id="F4915044FFE0FF833181FEF5FDCCC15A" box="[478,625,286,312]" degrees="27.422089" direction="north" orientation="latitude" pageId="5" pageNumber="252" precision="1" value="27.422089">N 27.422089</geoCoordinate>
;
<geoCoordinate id="F4915044FFE0FF833220FEF5FCA7C15A" box="[639,794,286,312]" degrees="92.2190308" direction="east" orientation="longitude" pageId="5" pageNumber="252" precision="1" value="92.21903">E 92.2190308</geoCoordinate>
; alt.:
<quantity id="565D9B66FFE0FF83308DFEA9FE96C13E" box="[210,299,322,348]" metricMagnitude="3" metricUnit="m" metricValue="1.794" pageId="5" pageNumber="252" unit="m" value="1794.0">
<elevation id="1A88D1B0FFE0FF83308DFEA9FE96C13E" box="[210,299,322,348]" metricMagnitude="3" metricUnit="m" metricValue="1.794" pageId="5" pageNumber="252" unit="m" value="1794.0">1794 m</elevation>
</quantity>
),
<collectingMunicipality id="717EACF9FFE0FF83311DFEA9FE2EC13E" box="[322,403,322,348]" pageId="5" pageNumber="252">Dirang</collectingMunicipality>
,
<collectingCounty id="787B4E0FFFE0FF8331FFFEA9FD27C13E" box="[416,666,322,348]" pageId="5" pageNumber="252">West Kameng district</collectingCounty>
,
<collectingRegion id="5361F861FFE0FF8332F8FEA9FF41C1E2" country="India" name="Arunachal Pradesh" pageId="5" pageNumber="252">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="E9B27613FFE0FF833158FE8DFEF9C1E2" box="[263,324,358,384]" name="India" pageId="5" pageNumber="252">India</collectingCountry>
; collected after dusk hours.
</materialsCitation>
</paragraph>
<paragraph id="911A3683FFE0FF833097FE60FDA3C3A2" blockId="5.[160,798,214,1897]" pageId="5" pageNumber="252">
<emphasis id="A3D1EA91FFE0FF833097FE60FE8EC1C6" box="[200,307,395,420]" italics="true" pageId="5" pageNumber="252">Diagnosis</emphasis>
:
<taxonomicName id="56A54D00FFE0FF83311FFE61FDB2C1C6" authorityName="Saikia &amp; Laskar &amp; Dinesh &amp; Shabnam &amp; Sinha" authorityYear="2022" box="[320,527,394,420]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="5" pageNumber="252" phylum="Chordata" rank="species" species="chanakya" status="sp. nov.">
<emphasis id="A3D1EA91FFE0FF83311FFE61FDB2C1C6" box="[320,527,394,420]" italics="true" pageId="5" pageNumber="252">Amolops chanakya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8E257EAFFE0FF833249FE61FDD3C1C6" box="[534,622,394,420]" pageId="5" pageNumber="252" rank="species">
<emphasis id="A3D1EA91FFE0FF833249FE61FDD5C1C6" bold="true" box="[534,616,394,420]" pageId="5" pageNumber="252">sp. nov</emphasis>
.
</taxonomicNameLabel>
is a member of the morphological
<taxonomicName id="56A54D00FFE0FF83312CFE45FD99C1AA" baseAuthorityName="Jiang" baseAuthorityYear="1983" box="[371,548,430,456]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="5" pageNumber="252" phylum="Chordata" rank="species" species="viridimaculatus">
<emphasis id="A3D1EA91FFE0FF83312CFE45FD99C1AA" box="[371,548,430,456]" italics="true" pageId="5" pageNumber="252">Viridimaculatus</emphasis>
</taxonomicName>
species group isolated in geographical space from its phylogenetic sister species and has a distinct set of morphological characters. The new species can be diagnosed based on the following set of morphological characters: adult male size large (SVL
<quantity id="565D9B66FFE0FF8332B0FDD5FF6CC21E" metricMagnitude="-2" metricUnit="m" metricValue="7.64" pageId="5" pageNumber="252" unit="mm" value="76.4">76.4 mm</quantity>
); head wider than long (HW&gt;HL); snout rounded, longer than eye length (SL&gt;EL); inter-orbital distance less than both upper eyelid width and inter-narial distance (IUE&lt;UEW&lt;IN); tympanum distinct, about 40% of eye length; finger I disk slightly dilated, all disks with circum-marginal grooves, only a hint of groove in finger I; tibia longer than femur and foot (TiL&gt;FOL&gt;ShL), about 2/3
<superScript id="66D09BCBFFE0FF833350FCD2FCA0C325" attach="none" box="[783,797,825,839]" fontSize="6" pageId="5" pageNumber="252">rd</superScript>
of SVL; tibio-tarsal articulation reaches up to the nostril; supra-tympanic fold strong; single row of discontinuous roundish tubercles dorso-laterally.
</paragraph>
<paragraph id="911A3683FFE0FF833097FC20FCA4C512" blockId="5.[160,798,214,1897]" pageId="5" pageNumber="252">
<emphasis id="A3D1EA91FFE0FF833097FC20FDBBC386" box="[200,518,970,996]" italics="true" pageId="5" pageNumber="252">
Description of the
<typeStatus id="4E1E8821FFE0FF8331FCFC21FDBBC386" box="[419,518,970,996]" pageId="5" pageNumber="252" type="holotype">Holotype</typeStatus>
</emphasis>
: Adult male (SVL
<quantity id="565D9B66FFE0FF8332B0FC21FF6CC46A" metricMagnitude="-2" metricUnit="m" metricValue="7.64" pageId="5" pageNumber="252" unit="mm" value="76.4">76.4 mm</quantity>
), body robust, head wider than long (HW&gt;HL), flat above; snout rounded, protruding in lateral view, snout longer than the eye diameter (SL&gt;EL); canthus rostralis distinct, loreal region depressed; inter-orbital distance less than both upper eyelid width and inter-narial distance (IUE&lt;UEW&lt;IN); the nostrils laterally placed, slightly nearer to the eyes (NS&gt;NE); pupil horizontal; tympanum distinct, circular and little depressed, separated from the eye by a distance slightly more than its diameter; pineal ocellus visible; a pair of vomerine teeth situated between the choanae; tongue bilobed. Vocal sac externally visible.
</paragraph>
<paragraph id="911A3683FFE0FF833097FA91FEF9C62A" blockId="5.[160,798,214,1897]" pageId="5" pageNumber="252">Arms strong, forearm length shorter than the hand length; fingers free, tips ending in disk, finger I disk slightly dilated; all disks with circum-marginal grooves, only a hint of groove in finger I, relative length of fingers I&lt;II&lt;IV&lt;III; sub-articular tubercles prominent, circular; prepollex oval.</paragraph>
<paragraph id="911A3683FFE0FF833097F9B9FD26C69E" blockId="5.[160,798,214,1897]" pageId="5" pageNumber="252">Hindlimbs long, tibia longer than femur and foot (TiL&gt;FOL&gt;ShL); toes long and thin, tips of all toes dilated into disks with circum-marginal grooves, toes completely webbed; sub-articular tubercles domed shaped; inner metatarsal tubercle oval shaped, outer absent.</paragraph>
<paragraph id="911A3683FFE0FF833097F8EDFC6EC256" blockId="5.[160,798,214,1897]" lastBlockId="5.[861,1499,214,1896]" pageId="5" pageNumber="252">
Dorsal skin smooth; a single row of discontinuous roundish tubercles dorso-laterally; laterally granular, supra-tympanic fold strong; ventral skin smooth.
<emphasis id="A3D1EA91FFE0FF8333DAFF3DFB55C092" box="[901,1256,214,240]" italics="true" pageId="5" pageNumber="252">
The Colouration of the
<typeStatus id="4E1E8821FFE0FF8334DAFF3DFB55C092" box="[1157,1256,214,240]" pageId="5" pageNumber="252" type="holotype">Holotype</typeStatus>
</emphasis>
: (
<emphasis id="A3D1EA91FFE0FF8334A0FF3CFA80C092" box="[1279,1341,214,240]" italics="true" pageId="5" pageNumber="252">in life</emphasis>
) Dorsal color dull brick-red, spotted with irregular cocoa-brown spots, these cocoa-brown spots enclosing a number of smaller dull brick-red spots; laterally spotted with neon yellow; ventral color cocoa-brown. Limbs distinctly banded; thighs with alternating dull brick-red and cocoa-brown (speckled with lighter dots) bands, hind limbs with neon yellow spots laterally (
<figureCitation id="099E2A06FFE0FF833404FE39FB03C18E" box="[1115,1214,466,492]" captionStart="Figure 3" captionStartId="4.[133,209,1109,1135]" captionTargetBox="[165,1441,213,1082]" captionTargetId="figure-28@4.[165,1441,213,1082]" captionTargetPageId="4" captionText="Figure 3. Live colour image of holotype of Amolops chanakya sp. nov. (Photo: Bikramjit Sinha)." figureDoi="http://doi.org/10.5281/zenodo.13185573" httpUri="https://zenodo.org/record/13185573/files/figure.png" pageId="5" pageNumber="252">Figure 3</figureCitation>
).
<emphasis id="A3D1EA91FFE0FF833489FE38FAC5C18E" box="[1238,1400,467,492]" italics="true" pageId="5" pageNumber="252">In preservative</emphasis>
same as in live condition, except that all the colours are faded (
<figureCitation id="099E2A06FFE0FF83333AFDF1FC7BC256" box="[869,966,538,564]" captionStart="Figure 4" captionStartId="4.[133,209,1868,1894]" captionTargetBox="[133,1472,1207,1815]" captionTargetId="figure-45@4.[130,1475,1202,1841]" captionTargetPageId="4" captionText="Figure 4. Holotype image of Amolops chanakya sp. nov. in preserved condition (A: dorsal view; B: ventral view)." figureDoi="http://doi.org/10.5281/zenodo.13185575" httpUri="https://zenodo.org/record/13185575/files/figure.png" pageId="5" pageNumber="252">Figure 4</figureCitation>
).
</paragraph>
<paragraph id="911A3683FFE0FF8333DAFDD5FB75C2C2" blockId="5.[861,1499,214,1896]" pageId="5" pageNumber="252">
<emphasis id="A3D1EA91FFE0FF8333DAFDD5FBCFC23A" box="[901,1138,574,600]" italics="true" pageId="5" pageNumber="252">
Condition of the
<typeStatus id="4E1E8821FFE0FF83341BFDD4FBCFC23A" box="[1092,1138,575,600]" pageId="5" pageNumber="252">type</typeStatus>
</emphasis>
: The
<typeStatus id="4E1E8821FFE0FF8334ECFDD4FB5EC23B" box="[1203,1251,575,601]" pageId="5" pageNumber="252">type</typeStatus>
is in good condition. Ventral incision was made to extract liver tissue which was used in the molecular study.
</paragraph>
<paragraph id="911A3683FFE0FF8333DAFD41FA64C28A" blockId="5.[861,1499,214,1896]" pageId="5" pageNumber="252">
<emphasis id="A3D1EA91FFE0FF8333DAFD41FB92C2A6" box="[901,1071,682,708]" italics="true" pageId="5" pageNumber="252">Natural history</emphasis>
: The
<typeStatus id="4E1E8821FFE0FF83342EFD41FB6EC2A6" box="[1137,1235,682,708]" pageId="5" pageNumber="252" type="holotype">holotype</typeStatus>
was collected after the dusk hours from a small hill stream from Chakpa village.
</paragraph>
<paragraph id="911A3683FFE0FF8333DAFD19FB48C336" blockId="5.[861,1499,214,1896]" pageId="5" pageNumber="252">
<emphasis id="A3D1EA91FFE0FF8333DAFD19FBB6C36E" box="[901,1035,754,780]" italics="true" pageId="5" pageNumber="252">Distribution</emphasis>
: Known from its
<typeStatus id="4E1E8821FFE0FF83348AFD18FAB8C36F" box="[1237,1285,755,781]" pageId="5" pageNumber="252">type</typeStatus>
locality of Chakpa Village, Dirang Administrative Circle, West Kameng district of
<collectingRegion id="5361F861FFE0FF83338FFCD1FB14C336" box="[976,1193,826,852]" country="India" name="Arunachal Pradesh" pageId="5" pageNumber="252">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="E9B27613FFE0FF8334ECFCD1FB4CC336" box="[1203,1265,826,852]" name="India" pageId="5" pageNumber="252">India</collectingCountry>
.
</paragraph>
<paragraph id="911A3683FFE0FF8333DAFCB4FADAC3FF" blockId="5.[861,1499,214,1896]" pageId="5" pageNumber="252">
<emphasis id="A3D1EA91FFE0FF8333DAFCB4FBA8C31B" box="[901,1045,863,889]" italics="true" pageId="5" pageNumber="252">Comparisons</emphasis>
: The new species is compared with the other species of
<taxonomicName id="56A54D00FFE0FF83344EFC69FB7FC3FE" baseAuthorityName="Jiang" baseAuthorityYear="1983" box="[1041,1218,898,924]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="5" pageNumber="252" phylum="Chordata" rank="species" species="viridimaculatus">
<emphasis id="A3D1EA91FFE0FF83344EFC69FB7FC3FE" box="[1041,1218,898,924]" italics="true" pageId="5" pageNumber="252">Viridimaculatus</emphasis>
</taxonomicName>
species group.
</paragraph>
<paragraph id="911A3683FFE0FF8033DAFC4CFEFDC256" blockId="5.[861,1499,214,1896]" lastBlockId="6.[133,771,214,960]" lastPageId="6" lastPageNumber="253" pageId="5" pageNumber="252">
From the Indian
<taxonomicName id="56A54D00FFE0FF833434FC4DFAA1C3A2" baseAuthorityName="Jiang" baseAuthorityYear="1983" box="[1131,1308,934,960]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="5" pageNumber="252" phylum="Chordata" rank="species" species="viridimaculatus">
<emphasis id="A3D1EA91FFE0FF833434FC4DFAA1C3A2" box="[1131,1308,934,960]" italics="true" pageId="5" pageNumber="252">Viridimaculatus</emphasis>
</taxonomicName>
species group,
<taxonomicName id="56A54D00FFE0FF833302FC21FB92C386" authorityName="Saikia &amp; Laskar &amp; Dinesh &amp; Shabnam &amp; Sinha" authorityYear="2022" box="[861,1071,970,996]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="5" pageNumber="252" phylum="Chordata" rank="species" species="chanakya" status="sp. nov.">
<emphasis id="A3D1EA91FFE0FF833302FC21FB92C386" box="[861,1071,970,996]" italics="true" pageId="5" pageNumber="252">Amolops chanakya</emphasis>
</taxonomicName>
<emphasis id="A3D1EA91FFE0FF833467FC21FB2FC386" bold="true" box="[1080,1170,970,996]" pageId="5" pageNumber="252">
<taxonomicNameLabel id="B8E257EAFFE0FF833467FC21FB2FC386" box="[1080,1170,970,996]" pageId="5" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="56A54D00FFE0FF833574FC20FA0CC386" baseAuthorityName="Gunther" baseAuthorityYear="1876" box="[1323,1457,970,997]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="5" pageNumber="252" phylum="Chordata" rank="species" species="formosus">
<emphasis id="A3D1EA91FFE0FF833574FC20FA0CC386" box="[1323,1457,970,997]" italics="true" pageId="5" pageNumber="252">A. formosus</emphasis>
</taxonomicName>
(in parentheses) in having larger male SVL (vs. smaller), nostril nearer to eye (vs. equidistant), tympanum small, distinct (vs. about the size of finger III disk), TiL&gt;ShL (vs. TiL&lt;ShL), inner-metatarsal tubercles distinct (vs. indistinct), outer metatarsal tubercle absent (vs. present), dorsal color dull brick-red, spotted with irregular cocoa-brown spots (vs. bright green with large dark spots); from
<taxonomicName id="56A54D00FFE0FF8333FEFB00FBEEC566" baseAuthorityName="Boulenger" baseAuthorityYear="1888" box="[929,1107,1258,1285]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="5" pageNumber="252" phylum="Chordata" rank="species" species="himalayanus">
<emphasis id="A3D1EA91FFE0FF8333FEFB00FBEEC566" box="[929,1107,1258,1285]" italics="true" pageId="5" pageNumber="252">A. himalayanus</emphasis>
</taxonomicName>
(in parentheses) in having small tympanum but distinct and depressed (vs. slightly visible), tibia 2/3
<superScript id="66D09BCBFFE0FF833444FADAFB94C55D" attach="left" box="[1051,1065,1329,1343]" fontSize="6" pageId="5" pageNumber="252">rd</superScript>
of SVL (vs. almost equal), tibio-tarsal articulation reached up to nostril (vs. beyond the tip of the snout), dorsum dull brick-red, spotted with irregular cocoa-brown spots (vs. olive with darker spots); from
<taxonomicName id="56A54D00FFE0FF83359EFA75FC6BC5BE" authorityName="Biju, Mahony &amp; Kamei" authorityYear="2010" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="5" pageNumber="252" phylum="Chordata" rank="species" species="nidorbellus">
<emphasis id="A3D1EA91FFE0FF83359EFA75FC6BC5BE" italics="true" pageId="5" pageNumber="252">A. nidorbellus</emphasis>
</taxonomicName>
(in parentheses) having distinct pineal ocellus (vs. barely visible), IUE&lt;UEW (vs. IUE&gt;EUW), external vocal sac visible (vs. absent), dorsal color dull brick-red, spotted with irregular cocoa-brown spots (vs. brown with small, irregular cobalt green spots), ventral cocoa-brown (vs. pale brown with scattered small, white spots); from
<taxonomicName id="56A54D00FFE0FF8333C3F971FBD7C6D6" baseAuthorityName="Jiang" baseAuthorityYear="1983" box="[924,1130,1690,1716]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="5" pageNumber="252" phylum="Chordata" rank="species" species="viridimaculatus">
<emphasis id="A3D1EA91FFE0FF8333C3F971FBD7C6D6" box="[924,1130,1690,1716]" italics="true" pageId="5" pageNumber="252">A. viridimaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8E257EAFFE0FF83342DF971FABDC6D6" box="[1138,1280,1690,1716]" pageId="5" pageNumber="252" sensu="stricto">sensu stricto</taxonomicNameLabel>
(in parentheses) in having wider head (vs. longer), tympanum distinct (vs. indistinct), outer metatarsal tubercle absent (vs. present), dorsal color dull brick-red, spotted with irregular cocoa-brown spots (vs. maroon red with large green blotches); from
<taxonomicName id="56A54D00FFE0FF8333FBF8A5FBA0C70A" authorityName="Saikia &amp; Laskar &amp; Dinesh &amp; Shabnam &amp; Sinha" authorityYear="2022" box="[932,1053,1870,1896]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="5" pageNumber="252" phylum="Chordata" rank="species" species="tawang" status="sp. nov.">
<emphasis id="A3D1EA91FFE0FF8333FBF8A5FBA0C70A" box="[932,1053,1870,1896]" italics="true" pageId="5" pageNumber="252">A. tawang</emphasis>
</taxonomicName>
<emphasis id="A3D1EA91FFE0FF833473F8A5FB31C70A" bold="true" box="[1068,1164,1870,1896]" pageId="5" pageNumber="252">
<taxonomicNameLabel id="B8E257EAFFE0FF833473F8A5FB31C70A" box="[1068,1164,1870,1896]" pageId="5" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(in parentheses) in having smaller male SVL (vs. larger), nostril nearer to the eye (vs. equidistant), IUE&lt;UEW (vs. IUE=UEW), tympanum larger (vs. extremely depressed and small), TYD ≈ TE (vs. TYD= ½TE), external vocal sac distinctly visible (vs. faintly visible), tibio-tarsal articulation reaches up to nostril (vs. snout), dorsum dull brick-red, spotted with irregular cocoa-brown spots (vs. dorsal color olive-green, spotted with large, irregular shaped dark-brown spots), ventral color cocoa-brown (vs. dark, speckled with lighter dots and dashes)
</paragraph>
<paragraph id="911A3683FFE3FF8130F2FDD5FD64C1AA" blockId="6.[133,771,214,960]" lastBlockId="7.[160,798,214,816]" lastPageId="7" lastPageNumber="254" pageId="6" pageNumber="253">
From the
<taxonomicName id="56A54D00FFE3FF803164FDD5FE51C23A" baseAuthorityName="Jiang" baseAuthorityYear="1983" box="[315,492,574,600]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="6" pageNumber="253" phylum="Chordata" rank="species" species="viridimaculatus">
<emphasis id="A3D1EA91FFE3FF803164FDD5FE51C23A" box="[315,492,574,600]" italics="true" pageId="6" pageNumber="253">Viridimaculatus</emphasis>
</taxonomicName>
species group found outside
<collectingCountry id="E9B27613FFE3FF803081FD89FEA6C21E" box="[222,283,610,636]" name="India" pageId="6" pageNumber="253">India</collectingCountry>
,
<taxonomicName id="56A54D00FFE3FF803178FD89FE4AC21E" authorityName="Saikia &amp; Laskar &amp; Dinesh &amp; Shabnam &amp; Sinha" authorityYear="2022" box="[295,503,610,636]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="6" pageNumber="253" phylum="Chordata" rank="species" species="chanakya" status="sp. nov.">
<emphasis id="A3D1EA91FFE3FF803178FD89FE4AC21E" box="[295,503,610,636]" italics="true" pageId="6" pageNumber="253">Amolops chanakya</emphasis>
</taxonomicName>
<emphasis id="A3D1EA91FFE3FF8031A1FD89FDEBC21E" bold="true" box="[510,598,610,636]" pageId="6" pageNumber="253">
<taxonomicNameLabel id="B8E257EAFFE3FF8031A1FD89FDEBC21E" box="[510,598,610,636]" pageId="6" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="56A54D00FFE3FF8032B5FD89FEB1C2C2" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="6" pageNumber="253" phylum="Chordata" rank="species" species="beibengensis">
<emphasis id="A3D1EA91FFE3FF8032B5FD89FEB1C2C2" italics="true" pageId="6" pageNumber="253">A. beibengensis</emphasis>
</taxonomicName>
(in parentheses) in having head wider than long (vs. HL=HW), tibio-tarsal articulation reached up to the nostril (vs. between nose and eye), vocal sac externally visible (vs. absent), dorsal color dull brick-red, spotted with irregular cocoa-brown spots (vs. dorsum brown with green spots); from
<taxonomicName id="56A54D00FFE3FF803133FCD1FDA5C336" box="[364,536,826,852]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="6" pageNumber="253" phylum="Chordata" rank="species" species="gyirongensis">
<emphasis id="A3D1EA91FFE3FF803133FCD1FDA5C336" box="[364,536,826,852]" italics="true" pageId="6" pageNumber="253">A. gyirongensis</emphasis>
</taxonomicName>
(in parentheses) in having larger male SVL&gt;
<quantity id="565D9B66FFE3FF8031FDFCB5FE4EC31A" box="[418,499,862,888]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="6" pageNumber="253" unit="mm" value="76.0">76 mm</quantity>
(vs. &lt;
<quantity id="565D9B66FFE3FF803268FCB5FD37C31A" box="[567,650,862,888]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="6" pageNumber="253" unit="mm" value="64.0">64 mm</quantity>
), vocal sac externally visible (vs. absent); dorsal color dull brick-red, spotted with irregular cocoa-brown spots (vs. bright green with dark brown irregular large spots); from
<taxonomicName id="56A54D00FFE3FF803568FF3DFA7DC092" baseAuthorityName="Smith" baseAuthorityYear="1940" box="[1335,1472,214,240]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="6" pageNumber="253" phylum="Chordata" rank="species" species="kaulbacki">
<emphasis id="A3D1EA91FFE3FF803568FF3DFA7DC092" box="[1335,1472,214,240]" italics="true" pageId="6" pageNumber="253">A. kaulbacki</emphasis>
</taxonomicName>
(in parentheses) in having larger male SVL&gt;
<quantity id="565D9B66FFE3FF80351DFF11FA2FC176" box="[1346,1426,250,276]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="6" pageNumber="253" unit="mm" value="76.0">76 mm</quantity>
(vs. up to
<quantity id="565D9B66FFE3FF8033D5FEF5FC5DC15A" box="[906,992,286,312]" metricMagnitude="-2" metricUnit="m" metricValue="7.2" pageId="6" pageNumber="253" unit="mm" value="72.0">72 mm</quantity>
), IUE&lt;UEW (vs. IUE&gt;UEW), tibio-tarsal articulation reached up to the nostril (vs. beyond snout), vocal sac externally visible (vs. absent), dorsal color dull brick-red, spotted with irregular cocoa-brown spots (vs. green with black spots and marbling); from
<taxonomicName id="56A54D00FFE3FF80357DFE45FA7DC1AA" baseAuthorityName="Andersson" baseAuthorityYear="1939" box="[1314,1472,430,456]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="6" pageNumber="253" phylum="Chordata" rank="species" species="longimanus">
<emphasis id="A3D1EA91FFE3FF80357DFE45FA7DC1AA" box="[1314,1472,430,456]" italics="true" pageId="6" pageNumber="253">A. longimanus</emphasis>
</taxonomicName>
(in parentheses) in not having a distinctive dorsolateral fold (vs. present), absence of parotid-like swelling above tympanum (vs. present); from
<taxonomicName id="56A54D00FFE3FF8034AAFDF1FA28C256" authorityName="Li &amp; Rao in Zhao, Rao, Lu &amp; Dong" authorityYear="2005" box="[1269,1429,538,564]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="6" pageNumber="253" phylum="Chordata" rank="species" species="medogensis">
<emphasis id="A3D1EA91FFE3FF8034AAFDF1FA28C256" box="[1269,1429,538,564]" italics="true" pageId="6" pageNumber="253">A. medogensis</emphasis>
</taxonomicName>
(in parentheses) in having smaller male SVL of
<quantity id="565D9B66FFE3FF803561FDD5FA2DC23B" box="[1342,1424,574,601]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="6" pageNumber="253" unit="mm" value="76.0">76 mm</quantity>
(vs.
<quantity id="565D9B66FFE3FF80331CFD89FC25C21E" box="[835,920,610,637]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="6" pageNumber="253" unit="mm" value="95.0">95 mm</quantity>
), vocal sac externally visible (vs. absent), dorsal color dull brick-red, spotted with irregular cocoa-brown spots (vs. olive green with brown markings/dark brown with small yellow-green spots); from
<taxonomicName id="56A54D00FFE3FF8034AAFD25FA28C28A" authorityName="Qi, Zhou, Lyu, Lu &amp; Li" authorityYear="2019" box="[1269,1429,718,744]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="6" pageNumber="253" phylum="Chordata" rank="species" species="pallasitatus">
<emphasis id="A3D1EA91FFE3FF8034AAFD25FA28C28A" box="[1269,1429,718,744]" italics="true" pageId="6" pageNumber="253">A. pallasitatus</emphasis>
</taxonomicName>
(in parentheses) in having tibio-tarsal articulation reached up to the nostril (vs. posterior eye corner), dorsal color dull brick-red, spotted with irregular cocoa-brown spots (vs. yellow green with irregular dark brown blotches); from
<taxonomicName id="56A54D00FFE3FF8033DEFC69FB88C3FE" authorityName="Orlov &amp; Ho" authorityYear="2007" box="[897,1077,898,924]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="6" pageNumber="253" phylum="Chordata" rank="species" species="splendissimus">
<emphasis id="A3D1EA91FFE3FF8033DEFC69FB88C3FE" box="[897,1077,898,924]" italics="true" pageId="6" pageNumber="253">A. splendissimus</emphasis>
</taxonomicName>
(in parentheses) in having a supra-tympanic fold strong (vs. poorly developed), dorsal color dull brick-red, spotted with irregular cocoa-brown spots (vs. black-brown spangled thickly with bright yellow spots); from
<taxonomicName id="56A54D00FFE2FF813166FEF5FE62C15A" box="[313,479,286,312]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="7" pageNumber="254" phylum="Chordata" rank="species" species="wangyufani">
<emphasis id="A3D1EA91FFE2FF813166FEF5FE62C15A" box="[313,479,286,312]" italics="true" pageId="7" pageNumber="254">A. wangyufani</emphasis>
</taxonomicName>
(in parentheses) in having larger male SVL&gt;
<quantity id="565D9B66FFE2FF8131DEFEA9FE64C13E" box="[385,473,322,348]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="254" unit="mm" value="76.0">76 mm</quantity>
(vs. &lt;
<quantity id="565D9B66FFE2FF813273FEA9FD3AC13E" box="[556,647,322,348]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="254" unit="mm" value="70.0">70 mm</quantity>
), tibio-tarsal articulation reached up to the nostril (vs. beyond snout), dorsal color dull brick-red, spotted with irregular cocoa-brown spots (vs. brown with sparse green patches).
</paragraph>
<caption id="C5DA660BFFE3FF8030D9F8B7FB21C714" ID-DOI="http://doi.org/10.5281/zenodo.13185577" ID-Zenodo-Dep="13185577" box="[134,1180,1884,1910]" httpUri="https://zenodo.org/record/13185577/files/figure.png" pageId="6" pageNumber="253" startId="6.[134,210,1884,1910]" targetBox="[144,1461,1000,1857]" targetPageId="6" targetType="figure">
<paragraph id="911A3683FFE3FF8030D9F8B7FB21C714" blockId="6.[134,1180,1884,1910]" box="[134,1180,1884,1910]" pageId="6" pageNumber="253">
<emphasis id="A3D1EA91FFE3FF8030D9F8B7FF50C714" bold="true" box="[134,237,1884,1910]" pageId="6" pageNumber="253">Figure 5.</emphasis>
Live colour image of holotype of
<taxonomicName id="56A54D00FFE3FF803235F8B7FC9DC714" authorityName="Saikia &amp; Laskar &amp; Dinesh &amp; Shabnam &amp; Sinha" authorityYear="2022" box="[618,800,1884,1910]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="6" pageNumber="253" phylum="Chordata" rank="species" species="tawang" status="sp. nov.">
<emphasis id="A3D1EA91FFE3FF803235F8B7FC9DC714" box="[618,800,1884,1910]" italics="true" pageId="6" pageNumber="253">Amolops tawang</emphasis>
</taxonomicName>
<emphasis id="A3D1EA91FFE3FF803379F8B7FCC0C714" bold="true" box="[806,893,1884,1910]" pageId="6" pageNumber="253">
<taxonomicNameLabel id="B8E257EAFFE3FF803379F8B7FCC0C714" box="[806,893,1884,1910]" pageId="6" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Photo: Bikramjit Sinha).
</paragraph>
</caption>
<paragraph id="911A3683FFE2FF813097FE39FF58C2A6" blockId="7.[160,798,214,816]" pageId="7" pageNumber="254">
<emphasis id="A3D1EA91FFE2FF813097FE39FE81C18E" box="[200,316,466,492]" italics="true" pageId="7" pageNumber="254">Etymology</emphasis>
: The species epithet is a patronym named after Chanakya (also known as Kautilya or Vishnugupta), a 4
<superScript id="66D09BCBFFE2FF813099FDF2FF69C245" attach="left" box="[198,212,537,551]" fontSize="6" pageId="7" pageNumber="254">th</superScript>
century BCE Indian polymath and administrator famous for Arthashastra, a Sanskrit treatise on economic policies, statecraft and military strategy. The species epithet is treated as a noun in apposition to the generic name.
</paragraph>
<paragraph id="911A3683FFE2FF813097FD25FCBCC352" blockId="7.[160,798,214,816]" pageId="7" pageNumber="254">
<emphasis id="A3D1EA91FFE2FF813097FD25FE48C28A" box="[200,501,718,744]" italics="true" pageId="7" pageNumber="254">Suggested Common Name</emphasis>
: We suggest
<emphasis id="A3D1EA91FFE2FF8132C4FD26FE89C369" bold="true" pageId="7" pageNumber="254">Chanakyas cascade frog</emphasis>
as a common English name and चाणक्य झरना मेंढक as a common Hindi name for this species.
</paragraph>
</subSubSection>
</treatment>
</document>