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<document id="0BAFAA30D48547F0066AD7E61289455D" ID-ISSN="1732-2421" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1723142841851" checkinUser="felipe" docAuthor="Mergl, Michal &amp; Massa, Dominique" docDate="2005" docId="AE188799B819FFF0FC9AFB29FEB8098F" docLanguage="en" docName="ActaPalaeontolPol.50.2.397-402.pdf" docOrigin="Acta Palaeontologica Polonica 50 (2)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Gigadiscina lessardi Mergl &amp; Massa, 2005, sp. nov." docType="treatment" docVersion="2" lastPageNumber="400" masterDocId="5221FFE1B818FFF3FFB6FF9EFFBE0C44" masterDocTitle="A new giant discinoid brachiopod from the Lower Devonian of Algeria" masterLastPageNumber="402" masterPageNumber="397" pageNumber="398" updateTime="1725041265664" updateUser="karina" zenodo-license-document="CC-BY-4.0">
<mods:mods id="7AFEC4489B95942D0CD6FD5FAB4AB46E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5C8690A1C8B04DC9E092BB53DDAC96CC">
<mods:title id="1BCCFAE5AF7131987FDF10B06FC08BC6">A new giant discinoid brachiopod from the Lower Devonian of Algeria</mods:title>
</mods:titleInfo>
<mods:name id="937974AA3C1C308860B688B86AEA63BA" type="personal">
<mods:role id="7D94492E353DE4946E18125279134BF1">
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</mods:role>
<mods:namePart id="F4B8728C6C02492702DA6CB0BDC37D43">Mergl, Michal</mods:namePart>
</mods:name>
<mods:name id="DC2C253A60E382C345A12BCF423C9BC3" type="personal">
<mods:role id="8F45E0ECCF4C20F4B1C978470E513A3B">
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<mods:namePart id="F8287556B30BBFFCE661DD515508EFE6">Massa, Dominique</mods:namePart>
</mods:name>
<mods:typeOfResource id="03872EFB5609ED70EF9E8B06421AA509">text</mods:typeOfResource>
<mods:relatedItem id="E4F5EAB4D79478BFE240F6E8AC3C3ED7" type="host">
<mods:titleInfo id="4D06D83D10E01B57DDE380AF62BAE2CD">
<mods:title id="B72CBFD446CD84A703307C91ABD949B2">Acta Palaeontologica Polonica</mods:title>
</mods:titleInfo>
<mods:part id="35A8912F70EB9144E46B94C564090A39">
<mods:date id="3A00DD970B7825CBE9FEF74B8F69FE4E">2005</mods:date>
<mods:detail id="AF07D41EBFA604A71D270D97DB0331C6" type="volume">
<mods:number id="A5A66E5993A0FF093B10C708156E2667">50</mods:number>
</mods:detail>
<mods:detail id="09C63B08420FD2B12AE5F26138A0FE91" type="issue">
<mods:number id="51289B8DCF31DF6BF1BDBD9C392C4CE7">2</mods:number>
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<mods:classification id="D68D11874E0FE2892F82BFC511FA5E5A">journal article</mods:classification>
<mods:identifier id="BCD1A01E01CB60AA3649E89FDD88A242" type="ISSN">1732-2421</mods:identifier>
</mods:mods>
<treatment id="AE188799B819FFF0FC9AFB29FEB8098F" LSID="urn:lsid:plazi:treatment:AE188799B819FFF0FC9AFB29FEB8098F" httpUri="http://treatment.plazi.org/id/AE188799B819FFF0FC9AFB29FEB8098F" lastPageId="3" lastPageNumber="400" pageId="1" pageNumber="398">
<subSubSection id="6EAB6504B819FFF2FC9AFB29FC2908B1" pageId="1" pageNumber="398" type="nomenclature">
<paragraph id="260E368FB819FFF2FC9AFB29FB1C0897" blockId="1.[812,1186,1207,1269]" box="[812,1186,1207,1235]" pageId="1" pageNumber="398">
<heading id="7D4681E3B819FFF2FC9AFB29FB1C0897" box="[812,1186,1207,1235]" fontSize="12" level="2" pageId="1" pageNumber="398" reason="9">
<taxonomicName id="E1B14D0CB819FFF2FC9AFB29FB880897" authorityName="Mergl &amp; Massa" authorityYear="2005" box="[812,1078,1207,1235]" class="Lingulata" family="Discinidae" genus="Gigadiscina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lingulida" pageId="1" pageNumber="398" phylum="Brachiopoda" rank="species" species="lessardi" status="sp. nov.">
<emphasis id="14C5EA9DB819FFF2FC9AFB29FB880897" box="[812,1078,1207,1235]" italics="true" pageId="1" pageNumber="398">Gigadiscina lessardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0FF657E6B819FFF2FB88FB29FB1C0897" box="[1086,1186,1207,1235]" pageId="1" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="260E368FB819FFF2FC9AFB7EFC2908B1" blockId="1.[812,1186,1207,1269]" box="[812,919,1248,1269]" pageId="1" pageNumber="398">Fig. 1A–E.</paragraph>
</subSubSection>
<subSubSection id="6EAB6504B819FFF0FC9AFA97FEB8098F" lastPageId="3" lastPageNumber="400" pageId="1" pageNumber="398" type="materials_examined">
<paragraph id="260E368FB819FFF2FC9AFA97FBEC097D" blockId="1.[812,1485,1288,1527]" pageId="1" pageNumber="398">
<materialsCitation id="96D93CD2B819FFF2FC9AFA97FBEC097D" collectionCode="ZCM" collectorName="External &amp; Massa-Mergl" pageId="1" pageNumber="398" specimenCode="S 03734, MAME 001" specimenCount="1" typeStatus="holotype">
<emphasis id="14C5EA9DB819FFF2FC9AFA97FC3A095A" box="[812,900,1289,1310]" italics="true" pageId="1" pageNumber="398">
<typeStatus id="F90A882DB819FFF2FC9AFA97FC3A095A" box="[812,900,1289,1310]" pageId="1" pageNumber="398" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectorName id="8B445359B819FFF2FC39FA96FC5E0959" box="[911,992,1288,1309]" pageId="1" pageNumber="398">External</collectorName>
mould of ventral valve (Fig. 1A), coll.
<collectorName id="8B445359B819FFF2FAFBFA96FA730959" box="[1357,1485,1288,1309]" pageId="1" pageNumber="398">Massa−Mergl</collectorName>
(
<collectionCode id="40A0AE4AB819FFF2FC85FABAFCD7097D" box="[819,873,1316,1337]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14031" name="Shetland Museum" pageId="1" pageNumber="398" type="Herbarium">ZCM</collectionCode>
<specimenCode id="76179EF4B819FFF2FCD8FABAFC00097D" box="[878,958,1316,1337]" collectionCode="ZCM" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14031" name="Shetland Museum" pageId="1" pageNumber="398" type="Herbarium">S 03734</specimenCode>
/
<specimenCode id="76179EF4B819FFF2FC72FABAFB82097D" box="[964,1084,1316,1337]" collectionCode="MAME" pageId="1" pageNumber="398">MAME 001</specimenCode>
a).
</materialsCitation>
</paragraph>
<paragraph id="260E368FB819FFF2FC9AFAD6FC5A093D" blockId="1.[812,1485,1288,1527]" pageId="1" pageNumber="398">
<emphasis id="14C5EA9DB819FFF2FC9AFAD6FC140919" box="[812,938,1352,1373]" italics="true" pageId="1" pageNumber="398">Type horizon</emphasis>
: Devonian, Pragian (Siegenian), equivalent of the Upper Tadrart Formation.
</paragraph>
<paragraph id="260E368FB819FFF2FC9AFA19FCC909FC" blockId="1.[812,1485,1288,1527]" pageId="1" pageNumber="398">
<emphasis id="14C5EA9DB819FFF2FC9AFA19FC1409D8" box="[812,938,1415,1436]" italics="true" pageId="1" pageNumber="398">Type locality</emphasis>
: Southern
<collectingCountry id="5EA6761FB819FFF2FBA1FA19FBDF09D8" box="[1047,1121,1415,1436]" name="Algeria" pageId="1" pageNumber="398">Algeria</collectingCountry>
, Tamesna Basin, In Rahir−Tim Amzi section.
</paragraph>
<paragraph id="260E368FB819FFF2FC9AFA58FB1309B3" blockId="1.[812,1485,1288,1527]" pageId="1" pageNumber="398">
<emphasis id="14C5EA9DB819FFF2FC9AFA58FC2D099F" box="[812,915,1478,1499]" italics="true" pageId="1" pageNumber="398">Etymology</emphasis>
: After French geologist Dr. L. Lessard, who made the pioneering field study in the Tamesna area.
</paragraph>
<paragraph id="260E368FB819FFF2FC9AF995FC570A27" blockId="1.[812,1485,1546,1635]" pageId="1" pageNumber="398">
<emphasis id="14C5EA9DB819FFF2FC9AF995FC300A67" box="[812,910,1547,1571]" italics="true" pageId="1" pageNumber="398">Material</emphasis>
.—Fourteen complete shells mostly preserved as corroded internal moulds, partially with remains of original phosphatic shell.
</paragraph>
<paragraph id="260E368FB819FFF2FC9AF9E8FADC0B4A" blockId="1.[812,1485,1653,1806]" pageId="1" pageNumber="398">
<emphasis id="14C5EA9DB819FFF2FC9AF9E8FC200AC9" box="[812,926,1654,1677]" italics="true" pageId="1" pageNumber="398">Diagnosis</emphasis>
.—Species of
<taxonomicName id="E1B14D0CB819FFF2FB89F9E8FB760ACA" authorityName="Mergl &amp; Massa" authorityYear="2005" box="[1087,1224,1654,1678]" class="Lingulata" family="Discinidae" genus="Gigadiscina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lingulida" pageId="1" pageNumber="398" phylum="Brachiopoda" rank="genus">
<emphasis id="14C5EA9DB819FFF2FB89F9E8FB760ACA" box="[1087,1224,1654,1678]" italics="true" pageId="1" pageNumber="398">Gigadiscina</emphasis>
</taxonomicName>
characterised by posteriorly weakly tapering shell, large pedicle foramen with small and short pedicle tube and rudimentary listrium; differs from
<taxonomicName id="E1B14D0CB819FFF2FCD8F948FC490AAA" authorityName="Kayser" authorityYear="1892" box="[878,1015,1750,1774]" class="Lingulata" family="Discinidae" genus="Discina" kingdom="Animalia" order="Lingulida" pageId="1" pageNumber="398" phylum="Brachiopoda" rank="species" species="anomala">
<emphasis id="14C5EA9DB819FFF2FCD8F948FC490AAA" box="[878,1015,1750,1774]" italics="true" pageId="1" pageNumber="398">G. anomala</emphasis>
</taxonomicName>
by finer and denser concentric rugellae (8–12 per mm
<emphasis id="14C5EA9DB819FFF2FC64F969FBA70B4A" box="[978,1049,1783,1806]" italics="true" pageId="1" pageNumber="398">versus</emphasis>
3–5 per mm in
<taxonomicName id="E1B14D0CB819FFF2FB7BF968FAEF0B4A" authorityName="Kayser" authorityYear="1892" box="[1229,1361,1782,1806]" class="Lingulata" family="Discinidae" genus="Discina" kingdom="Animalia" order="Lingulida" pageId="1" pageNumber="398" phylum="Brachiopoda" rank="species" species="anomala">
<emphasis id="14C5EA9DB819FFF2FB7BF968FAEF0B4A" box="[1229,1361,1782,1806]" italics="true" pageId="1" pageNumber="398">G. anomala</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="260E368FB819FFF2FFD0F8D5FDB10BAD" blockId="1.[102,1528,1866,2029]" pageId="1" pageNumber="398">
Fig. 1.
<emphasis id="14C5EA9DB819FFF2FF15F8D4FF700B1A" bold="true" box="[163,206,1866,1887]" pageId="1" pageNumber="398">A–E</emphasis>
.
<taxonomicName id="E1B14D0CB819FFF2FF6EF8D5FE2D0B1B" authorityName="Mergl &amp; Massa" authorityYear="2005" box="[216,403,1867,1887]" class="Lingulata" family="Discinidae" genus="Gigadiscina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lingulida" pageId="1" pageNumber="398" phylum="Brachiopoda" rank="species" species="lessardi" status="sp. nov.">
<emphasis id="14C5EA9DB819FFF2FF6EF8D5FE2D0B1B" box="[216,403,1867,1887]" italics="true" pageId="1" pageNumber="398">Gigadiscina lessardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0FF657E6B819FFF2FE21F8D2FE630B1B" box="[407,477,1868,1887]" pageId="1" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
from Pragian (Lower Devonian) of Tamesna Basin, In Rahir−Tim Amzi section.
<emphasis id="14C5EA9DB819FFF2FB1DF8D4FB050B1A" bold="true" box="[1195,1211,1866,1886]" pageId="1" pageNumber="398">A</emphasis>
.
<typeStatus id="F90A882DB819FFF2FB73F8D5FAA20B1B" box="[1221,1308,1867,1887]" pageId="1" pageNumber="398" type="holotype">Holotype</typeStatus>
, ventral valve, latex
<emphasis id="14C5EA9DB819FFF2FA6AF8D0FA460B18" box="[1500,1528,1870,1884]" italics="true" pageId="1" pageNumber="398">Ą</emphasis>
cast of external mould, ZCM S 03734a/
<collectionCode id="40A0AE4AB819FFF2FE73F8F8FDB20B3E" box="[453,524,1894,1914]" pageId="1" pageNumber="398">MAME</collectionCode>
001a.
<emphasis id="14C5EA9DB819FFF2FDF0F8F9FDEB0B3E" bold="true" box="[582,597,1895,1914]" pageId="1" pageNumber="398">B</emphasis>
. Incomplete ventral valve, latex cast of external mould, ZCM S 03734b/
<collectionCode id="40A0AE4AB819FFF2FB6EF8F8FAA10B3E" box="[1240,1311,1894,1914]" pageId="1" pageNumber="398">MAME</collectionCode>
001b.
<emphasis id="14C5EA9DB819FFF2FAECF8F8FAD40B3E" bold="true" box="[1370,1386,1894,1914]" pageId="1" pageNumber="398">C</emphasis>
. Complete shell, ventral valve (C
<subScript id="BA3534CAB819FFF2FE9FF812FE8F0BDE" attach="left" box="[297,305,1932,1946]" fontSize="6" pageId="1" pageNumber="398">1</subScript>
), side view (C
<subScript id="BA3534CAB819FFF2FE04F812FE040BDE" attach="left" box="[434,442,1932,1946]" fontSize="6" pageId="1" pageNumber="398">2</subScript>
), dorsal valve (C
<subScript id="BA3534CAB819FFF2FDE5F812FDE50BDE" attach="left" box="[595,603,1932,1946]" fontSize="6" pageId="1" pageNumber="398">3</subScript>
), and detail of ventral valve exterior (C
<subScript id="BA3534CAB819FFF2FC0EF812FC7E0BDE" attach="left" box="[952,960,1932,1946]" fontSize="6" pageId="1" pageNumber="398">4</subScript>
), ZCM S 03735/
<collectionCode id="40A0AE4AB819FFF2FBE0F81CFB230BD2" box="[1110,1181,1922,1942]" pageId="1" pageNumber="398">MAME</collectionCode>
002.
<emphasis id="14C5EA9DB819FFF2FB7BF81DFB630BD2" bold="true" box="[1229,1245,1923,1942]" pageId="1" pageNumber="398">D</emphasis>
. Complete shell, side view (D
<subScript id="BA3534CAB819FFF2FFC8F839FF380BF1" attach="right" box="[126,134,1959,1973]" fontSize="6" pageId="1" pageNumber="398">1</subScript>
), dorsal valve (D
<subScript id="BA3534CAB819FFF2FE92F839FE920BF1" attach="right" box="[292,300,1959,1973]" fontSize="6" pageId="1" pageNumber="398">2</subScript>
), ventral valve (D
<subScript id="BA3534CAB819FFF2FE67F839FE670BF1" attach="left" box="[465,473,1959,1973]" fontSize="6" pageId="1" pageNumber="398">3</subScript>
), and detail of intervascular ridges in ventral mould (D
<subScript id="BA3534CAB819FFF2FC7AF839FC6A0BF1" attach="left" box="[972,980,1959,1973]" fontSize="6" pageId="1" pageNumber="398">4</subScript>
), ZCM S 03736/
<collectionCode id="40A0AE4AB819FFF2FBD9F800FB0B0BF6" box="[1135,1205,1950,1970]" pageId="1" pageNumber="398">MAME</collectionCode>
003.
<emphasis id="14C5EA9DB819FFF2FB5EF800FB490BF5" bold="true" box="[1256,1271,1950,1969]" pageId="1" pageNumber="398">E</emphasis>
. Ventral valve, internal mould, ZCM S 03737/
<collectionCode id="40A0AE4AB819FFF2FE85F827FEC40B89" box="[307,378,1977,1997]" pageId="1" pageNumber="398">MAME</collectionCode>
0004.
<emphasis id="14C5EA9DB819FFF2FE0EF824FE780B89" bold="true" box="[440,454,1978,1997]" pageId="1" pageNumber="398">F</emphasis>
.
<taxonomicName id="E1B14D0CB819FFF2FE64F824FC9D0B89" authority="(Kayser, 1892)" baseAuthorityName="Kayser" baseAuthorityYear="1892" box="[466,803,1977,1998]" class="Lingulata" family="Discinidae" genus="Gigadiscina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lingulida" pageId="1" pageNumber="398" phylum="Brachiopoda" rank="species" species="anomala">
<emphasis id="14C5EA9DB819FFF2FE64F824FD2B0B8A" box="[466,661,1978,1998]" italics="true" pageId="1" pageNumber="398">Gigadiscina anomala</emphasis>
(
<bibRefCitation id="42204B7EB819FFF2FD17F827FCA20B89" author="Kayser, E." box="[673,796,1977,1997]" pageId="1" pageNumber="398" pagination="95 - 107" refId="ref4692" refString="Kayser, E. 1892. Beitrage zur Kenntnis der Fauna der Siegenischen Grauwacke. Jahrbuch der koninglichen preussischen geologischen Landesanstalt 2 (8): 95 - 107." type="journal article" year="1892">Kayser, 1892</bibRefCitation>
)
</taxonomicName>
from Pragian (Lower Devonian) of Siegen,
<collectingCountry id="5EA6761FB819FFF2FB03F827FAB20B89" box="[1205,1292,1977,1997]" name="Germany" pageId="1" pageNumber="398">Germany</collectingCountry>
. Complete shell, side view (F
<subScript id="BA3534CAB819FFF2FF1AF841FF0A0BA9" attach="left" box="[172,180,2015,2029]" fontSize="6" pageId="1" pageNumber="398">1</subScript>
), ventral valve (F
<subScript id="BA3534CAB819FFF2FEE0F841FEE00BA9" attach="left" box="[342,350,2015,2029]" fontSize="6" pageId="1" pageNumber="398">2</subScript>
),
<collectionCode id="40A0AE4AB819FFF2FEC7F848FE0B0BAD" box="[369,437,2006,2025]" pageId="1" pageNumber="398">BMNH</collectionCode>
B 42906.
</paragraph>
<paragraph id="260E368FB81BFFF0FFD0FF77FDCE0D25" blockId="3.[102,775,232,1186]" pageId="3" pageNumber="400">
<emphasis id="14C5EA9DB81BFFF0FFD0FF77FF540D44" box="[102,234,233,256]" italics="true" pageId="3" pageNumber="400">Description</emphasis>
.—The shell is
<quantity id="E1499B6AB81BFFF0FE21FF76FDAF0D44" box="[407,529,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="3" pageNumber="400" unit="mm" value="57.5" valueMax="60.0" valueMin="55.0">55–60 mm</quantity>
wide and thin−walled in a relation to the shell size. The outline is almost circular with less curved posterolateral margins. Commissure is rectimarginate. The shell width equals shell length.
</paragraph>
<paragraph id="260E368FB81BFFF0FF3AFEF6FEDD0E65" blockId="3.[102,775,232,1186]" pageId="3" pageNumber="400">The dorsal valve is asymmetrical and highly conical, with a rounded apex. The dorsal apex is subcentral (slightly anterior). The anterior slope is steep and gently convex in lateral profile, the posterior slope is similarly steep, straight near the apex but it tends to be concave near the posterior margin. Lateral slopes are straight.</paragraph>
<paragraph id="260E368FB81BFFF0FF3AFDB7FE200F26" blockId="3.[102,775,232,1186]" pageId="3" pageNumber="400">The ventral valve is flat, with dorsally curved posterior and posterolateral margins. The ventral apex is subcentral, situated between the posterior third and the midlength, embedded into flat surface of the valve. The pedicle foramen is large, located at the bottom of weakly depressed circular area. The pedicle foramen continues internally into a very short, narrow, and posteriorly extend pedicle tube. The interiors of both valves are devoid of well−defined visceral areas. Fine radially arranged vascular canals are impressed on the internal surface of the shell.</paragraph>
<paragraph id="260E368FB81BFFF0FF3AFCF7FE0308E6" blockId="3.[102,775,232,1186]" pageId="3" pageNumber="400">
The ornament consists of fine concentric rugellae with rounded crests. The size of rugellae is almost uniform over the whole surface, with more distant rugellae only in the apical region. There are 12–16 rugellae per
<quantity id="E1499B6AB81BFFF0FE51FC57FD980FA5" box="[487,550,969,994]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="400" unit="mm" value="5.0">5 mm</quantity>
in the anteromedian part of the shell. The beak region of the ventral valve is covered by inconspicuous rugellae; the earliest coarser rugellae appear
<quantity id="E1499B6AB81BFFF0FF03FBB7FEB10805" box="[181,271,1065,1090]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="3" pageNumber="400" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
from the apex. The rugellae are evenly spaced over the whole surface with the anteromedian sector of the shell exhibiting weak irregularities. New rugellae originate exclusively by an intercalation.
</paragraph>
<paragraph id="260E368FB81BFFF0FFD0FB2CFEB8098F" blockId="3.[102,775,1202,1483]" pageId="3" pageNumber="400">
<emphasis id="14C5EA9DB81BFFF0FFD0FB2CFF54088E" box="[102,234,1202,1226]" italics="true" pageId="3" pageNumber="400">Occurrence</emphasis>
.—All specimens of the new species were collected in In Rahir−Tim Amzi section (L. Lessard’s sample H 1035) in the Tenere Desert of the Tamesna Basin, south of the Ahaggar Massif in South
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. In this area, the Lower Palaeozoic comprises a sequence of Cambrian to Devonian strata in a narrow SW−NE trending belt. The basal transgressive Devonian unit bearing the new species is probably of Pragian (Siegenian) age and it rests unconformably on Silurian shale.
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105
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<emphasis id="14C5EA9DB81BFFF0FC9AFCC1FC760F3F" box="[812,968,863,891]" italics="true" pageId="3" pageNumber="400">Gigadiscina</emphasis>
cf.
<emphasis id="14C5EA9DB81BFFF0FC4FFCC1FB800F3F" box="[1017,1086,863,891]" italics="true" pageId="3" pageNumber="400">collis</emphasis>
(
<bibRefCitation id="42204B7EB81BFFF0FBF9FCC0FB490F3E" author="Clarke, J. M." box="[1103,1271,862,890]" pageId="3" pageNumber="400" pagination="1 - 353" refId="ref4263" refString="Clarke, J. M. 1913. Fosseis devonianos do Parana. Monographias do Servico Geologico e Mineralogico do Brasil 1: 1 - 353." type="journal article" year="1913">Clarke, 1913</bibRefCitation>
)
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<subSubSection id="6EAB6504B81BFFF0FC9AFC18FA330A1F" pageId="3" pageNumber="400" type="description">
<paragraph id="260E368FB81BFFF0FC9AFC18FCC20FDF" blockId="3.[812,1281,862,923]" box="[812,892,902,923]" pageId="3" pageNumber="400">
<figureCitation id="BE8A2A0AB81BFFF0FC9AFC18FCC70FDF" box="[812,889,902,923]" captionStart="Fig" captionStartId="4.[102,134,1294,1314]" captionTargetBox="[102,1484,225,1267]" captionTargetId="figure-320@4.[102,1485,225,1267]" captionTargetPageId="4" captionText="Fig. 2. A. Gigadiscina cf. collis (Clarke, 1913) from Upper Devonian of Falkland Islands, near Port Howard. External mould of ventral valve, BNMH B 60298. B. Gigadiscina sp. from Middle–Upper Devonian of Libya, Tripoli area (borehole BP, well C1−8). Ventral valve BMHN BD 3385b, external mould (B1), ventral valve BMNH BD 3385a, detail of ornament (B2), detail of microornament (B3), and detail of pitted radial bands alternating with smooth bands (B4)." pageId="3" pageNumber="400">Fig. 2A</figureCitation>
.
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<emphasis id="14C5EA9DB81BFFF0FC9AFC2EFC300F8C" box="[812,910,944,968]" italics="true" pageId="3" pageNumber="400">Material</emphasis>
.—Single ventral valve, preserved in ferruginous sandstone of Lower Devonian age, collected between Port Howard and Manybranch Harbour in the
<collectingCountry id="5EA6761FB81BFFF0FABDFC71FA77084C" box="[1291,1481,1007,1032]" name="Falkland Islands" pageId="3" pageNumber="400">Falkland Islands</collectingCountry>
, kept in the Natural History Museum, London (BMNH B 60298).
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<emphasis id="14C5EA9DB81BFFF0FC9AFBC4FC0E0835" box="[812,944,1114,1137]" italics="true" pageId="3" pageNumber="400">Description</emphasis>
.—The ventral valve is some
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long, flat with a shallow depression around the pedicle foramen. The shell is widest slightly anterior to the midlength. The posterior part of the valve is not preserved, but the courses of the rugellae indicate, that the posterior margin is slightly extended. The surface is ornamented by regular, rather fine and uniform concentric rugellae, numbering 14–15 per
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<emphasis id="14C5EA9DB81BFFF0FC9AFADCFC30091E" box="[812,910,1346,1370]" italics="true" pageId="3" pageNumber="400">Remarks</emphasis>
.—The illustrated herein ventral valve may belong to
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or another closely related species, although the
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of
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<emphasis id="14C5EA9DB81BFFF0FB28FA1CFAC809DE" box="[1182,1398,1410,1434]" italics="true" pageId="3" pageNumber="400">Orbiculoidea collis</emphasis>
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is known only from a single, dorsal valve (
<bibRefCitation id="42204B7EB81BFFF0FAFCFA3CFA7609FF" author="Reed, F. R. C." box="[1354,1480,1442,1467]" pageId="3" pageNumber="400" pagination="27 - 225" refId="ref4962" refString="Reed, F. R. C. 1925 Revision of the Fauna of the Bokkeveld Beds. Annals of the South African Museum 22: 27 - 225." type="journal article" year="1925">Reed 1925</bibRefCitation>
;
<bibRefCitation id="42204B7EB81BFFF0FC9AFA5DFBB5099F" author="Boucot, A. J. &amp; Rowell, A. J. &amp; Racheboeuf, P. &amp; Pereira, E. &amp; Goncalves de Melo, J. H. &amp; Peixoto de Siqueira, L." box="[812,1035,1474,1499]" pageId="3" pageNumber="400" pagination="109 - 120" refId="ref4185" refString="Boucot, A. J., Rowell, A. J., Racheboeuf, P., Pereira, E., Goncalves de Melo, J. H., and Peixoto de Siqueira, L. 2001. Position of the Malvinokaffric Realm's northern boundary (Early Devonian) based on newly discovered brachiopods from the Parecis Basin (Brazil). Journal of the Czech Geological Society 46: 109 - 120." type="journal article" year="2001">Boucot et al. 2001</bibRefCitation>
). We found out that our ventral valve shares the same style of sculpture hence can be conspecific. Because the size and morphology of the illustrated herein ventral valve is similar to that of
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<emphasis id="14C5EA9DB81BFFF0FB00F9BCFA1D0A7E" box="[1206,1443,1570,1594]" italics="true" pageId="3" pageNumber="400">Gigadiscina lessardi</emphasis>
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we propose to include
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<emphasis id="14C5EA9DB81BFFF0FBD5F9DCFB7C0A1E" box="[1123,1218,1602,1626]" italics="true" pageId="3" pageNumber="400">O. collis</emphasis>
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to the new genus.
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Fig
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<taxonomicNameLabel id="0FF657E6B81BFFF0FEA6F9A0FEF80A17" box="[272,326,1598,1619]" pageId="3" pageNumber="400" rank="species">n. sp.</taxonomicNameLabel>
;
<treatmentCitation id="A710109EB81BFFF0FEE6F9A0FE4C0A17" author="Kayser, E." box="[336,498,1598,1619]" page="96" pageId="3" pageNumber="400" year="1892">
<bibRefCitation id="42204B7EB81BFFF0FEE6F9A0FE4C0A17" author="Kayser, E." box="[336,498,1598,1619]" pageId="3" pageNumber="400" pagination="95 - 107" refId="ref4692" refString="Kayser, E. 1892. Beitrage zur Kenntnis der Fauna der Siegenischen Grauwacke. Jahrbuch der koninglichen preussischen geologischen Landesanstalt 2 (8): 95 - 107." type="journal article" year="1892">Kayser 1892: 96</bibRefCitation>
</treatmentCitation>
: pl. 10: 1–3.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="6EAB6504B81BFFF0FFD0F9FDFAEA0F06" pageId="3" pageNumber="400" type="materials_examined">
<paragraph id="260E368FB81BFFF0FFD0F9FDFE480AEB" blockId="3.[102,775,1598,1807]" pageId="3" pageNumber="400">
<materialsCitation id="96D93CD2B81BFFF0FFD0F9FDFE480AEB" collectionCode="MGB" country="Germany" county="Marburg" location="Dorsal" municipality="Marburg University" pageId="3" pageNumber="400" specimenCode="MGB 2107" specimenCount="1" typeStatus="holotype">
<emphasis id="14C5EA9DB81BFFF0FFD0F9FDFF000A3C" box="[102,190,1635,1656]" italics="true" pageId="3" pageNumber="400">
<typeStatus id="F90A882DB81BFFF0FFD0F9FDFF000A3C" box="[102,190,1635,1656]" pageId="3" pageNumber="400" type="holotype">Holotype</typeStatus>
</emphasis>
:
<location id="236E6054B81BFFF0FF7EF9FCFEB60A33" LSID="urn:lsid:plazi:treatment:AE188799B81BFFF0FFD0FA6EFAEA0F06:236E6054B81BFFF0FF7EF9FCFEB60A33" box="[200,264,1634,1655]" country="Germany" county="Marburg" municipality="Marburg University" name="Dorsal" pageId="3" pageNumber="400">Dorsal</location>
valve, internal moulds with remains of original ventral valve, deposited in the palaeontological collection of the
<collectingMunicipality id="C66AACF5B81BFFF0FD34F9E0FF140AEB" pageId="3" pageNumber="400">Marburg University</collectingMunicipality>
,
<collectingCounty id="CF6F4E03B81BFFF0FF05F904FEB70AEB" box="[179,265,1690,1711]" pageId="3" pageNumber="400">Marburg</collectingCounty>
,
<collectingCountry id="5EA6761FB81BFFF0FEA5F904FED20AEB" box="[275,364,1690,1711]" name="Germany" pageId="3" pageNumber="400">Germany</collectingCountry>
(
<specimenCode id="76179EF4B81BFFF0FECFF904FE550AEB" box="[377,491,1690,1711]" collectionCode="MGB" country="Spain" httpUri="http://grbio.org/cool/d3z2-v7v8" name="Museo de Geologia (del Seminario Diocesano) de Barcelona" pageId="3" pageNumber="400">MGB 2107</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="260E368FB81BFFF0FFD0F922FCBB0A95" blockId="3.[102,775,1598,1807]" box="[102,773,1724,1745]" pageId="3" pageNumber="400">
<emphasis id="14C5EA9DB81BFFF0FFD0F922FF5D0A95" box="[102,227,1724,1745]" italics="true" pageId="3" pageNumber="400">Type horizon</emphasis>
: Devonian, Pragian (Siegenian), Siegenian Greywackes.
</paragraph>
<paragraph id="260E368FB81BFFF0FFD0F940FEB20B4B" blockId="3.[102,775,1598,1807]" pageId="3" pageNumber="400">
<emphasis id="14C5EA9DB81BFFF0FFD0F940FF5C0AB7" box="[102,226,1758,1779]" italics="true" pageId="3" pageNumber="400">Type locality</emphasis>
:
<collectingCountry id="5EA6761FB81BFFF0FF59F940FEF50AB7" box="[239,331,1758,1779]" name="Germany" pageId="3" pageNumber="400">Germany</collectingCountry>
, Siegen, quarry eastward of Siegen, near “Siegena“ ironworks.
</paragraph>
<paragraph id="260E368FB81BFFF0FFD0F8BEFE910BDD" blockId="3.[102,775,1824,1945]" pageId="3" pageNumber="400">
<emphasis id="14C5EA9DB81BFFF0FFD0F8BEFF780B7C" box="[102,198,1824,1848]" italics="true" pageId="3" pageNumber="400">Material</emphasis>
.—
<materialsCitation id="96D93CD2B81BFFF0FF50F8BEFE950BDD" collectionCode="BMNH" country="Germany" county="London" location="The" municipality="Natural History Museum" pageId="3" pageNumber="400" specimenCode="B 42906, B 42915" specimenCount="1" typeStatus="holotype">
<location id="236E6054B81BFFF0FF50F8BEFEAD0B7D" LSID="urn:lsid:plazi:treatment:AE188799B81BFFF0FFD0FA6EFAEA0F06:236E6054B81BFFF0FF50F8BEFEAD0B7D" box="[230,275,1824,1849]" country="Germany" county="London" municipality="Natural History Museum" name="The" pageId="3" pageNumber="400">The</location>
<typeStatus id="F90A882DB81BFFF0FEACF8BEFEC30B7D" box="[282,381,1824,1849]" pageId="3" pageNumber="400" type="holotype">holotype</typeStatus>
, and two complete shells, preserved in sandstone from the quarry east of
<location id="236E6054B81BFFF0FE59F8DEFD820B1D" LSID="urn:lsid:plazi:treatment:AE188799B81BFFF0FFD0FA6EFAEA0F06:236E6054B81BFFF0FE59F8DEFD820B1D" box="[495,572,1856,1881]" country="Germany" county="London" municipality="Natural History Museum" name="Siegen" pageId="3" pageNumber="400">Siegen</location>
,
<collectingCountry id="5EA6761FB81BFFF0FDF0F8DEFD110B1D" box="[582,687,1856,1881]" name="Germany" pageId="3" pageNumber="400">Germany</collectingCountry>
, housed in the
<collectingMunicipality id="C66AACF5B81BFFF0FF06F8FEFE720B3C" box="[176,460,1888,1913]" pageId="3" pageNumber="400">Natural History Museum</collectingMunicipality>
,
<collectingCounty id="CF6F4E03B81BFFF0FE6FF8FEFD910B3D" box="[473,559,1888,1913]" pageId="3" pageNumber="400">London</collectingCounty>
(
<collectionCode id="40A0AE4AB81BFFF0FDF6F8FEFD2B0B3D" box="[576,661,1888,1913]" pageId="3" pageNumber="400">BMNH</collectionCode>
<specimenCode id="76179EF4B81BFFF0FD28F8FFFCBD0B3D" box="[670,771,1888,1913]" collectionCode="BMNH" pageId="3" pageNumber="400">B 42906</specimenCode>
,
<collectionCode id="40A0AE4AB81BFFF0FFD0F81FFF070BDC" box="[102,185,1921,1944]" pageId="3" pageNumber="400">BMNH</collectionCode>
<specimenCode id="76179EF4B81BFFF0FF76F81FFE9D0BDD" box="[192,291,1920,1945]" collectionCode="BMNH" pageId="3" pageNumber="400">B 42915</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="260E368FB81BFFF0FFD0F834FE5B0B86" blockId="3.[102,775,1961,2026]" box="[102,485,1961,1986]" pageId="3" pageNumber="400">
<emphasis id="14C5EA9DB81BFFF0FFD0F834FF540B85" box="[102,234,1962,1985]" italics="true" pageId="3" pageNumber="400">Description</emphasis>
.—See
<bibRefCitation id="42204B7EB81BFFF0FE8AF837FE5E0B86" author="Kayser, E." box="[316,480,1961,1986]" pageId="3" pageNumber="400" pagination="95 - 107" refId="ref4692" refString="Kayser, E. 1892. Beitrage zur Kenntnis der Fauna der Siegenischen Grauwacke. Jahrbuch der koninglichen preussischen geologischen Landesanstalt 2 (8): 95 - 107." type="journal article" year="1892">Kayser (1892)</bibRefCitation>
.
</paragraph>
<paragraph id="260E368FB81BFFF0FFD0F84CFA3C0E45" blockId="3.[102,775,1961,2026]" lastBlockId="3.[812,1486,232,834]" pageId="3" pageNumber="400">
<emphasis id="14C5EA9DB81BFFF0FFD0F84CFF7B0BAE" box="[102,197,2002,2026]" italics="true" pageId="3" pageNumber="400">Remarks</emphasis>
.—The
<typeStatus id="F90A882DB81BFFF0FEAEF84FFEC90BAE" box="[280,375,2001,2026]" pageId="3" pageNumber="400" type="holotype">holotype</typeStatus>
is large, strongly asymmetrical, shell
<quantity id="E1499B6AB81BFFF0FC9AFF76FCC70D44" box="[812,889,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="400" unit="mm" value="43.0">43 mm</quantity>
long, by
<quantity id="E1499B6AB81BFFF0FC68FF76FB940D44" box="[990,1066,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="400" unit="mm" value="45.0">45 mm</quantity>
wide, and
<quantity id="E1499B6AB81BFFF0FB16FF76FB530D44" box="[1184,1261,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="400" unit="mm" value="20.0">20 mm</quantity>
high. Its ornamentation is preserved only in posterolateral slope, with ten dense rugellae per
<quantity id="E1499B6AB81BFFF0FC0EFEB6FC430D04" box="[952,1021,296,321]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="400" unit="mm" value="5.0">5 mm</quantity>
. Two other available complete specimens can be referred to
<taxonomicName id="E1B14D0CB81BFFF0FC59FED7FAEA0D25" authority="(Kayser, 1892)" baseAuthorityName="Kayser" baseAuthorityYear="1892" box="[1007,1364,328,353]" class="Lingulata" family="Discinidae" genus="Discina" kingdom="Animalia" order="Lingulida" pageId="3" pageNumber="400" phylum="Brachiopoda" rank="species" species="anomala">
<emphasis id="14C5EA9DB81BFFF0FC59FED7FB170D24" box="[1007,1193,328,352]" italics="true" pageId="3" pageNumber="400">Discina anomala</emphasis>
(
<bibRefCitation id="42204B7EB81BFFF0FB00FED6FAF20D25" author="Kayser, E." box="[1206,1356,328,353]" pageId="3" pageNumber="400" pagination="95 - 107" refId="ref4692" refString="Kayser, E. 1892. Beitrage zur Kenntnis der Fauna der Siegenischen Grauwacke. Jahrbuch der koninglichen preussischen geologischen Landesanstalt 2 (8): 95 - 107." type="journal article" year="1892">Kayser, 1892</bibRefCitation>
)
</taxonomicName>
. The larger specimen is
<quantity id="E1499B6AB81BFFF0FC0DFEF6FBB30DC4" box="[955,1037,360,385]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="400" unit="mm" value="60.0">60 mm</quantity>
long, with the slightly eccentric dorsal apex, a flat ventral valve with a depression around the pedicle foramen, and distinct concentric rugellae covering shell surface. The second specimen is smaller, with an irregular outline and a flat ventral valve covered by concentric rugellae.
</paragraph>
<paragraph id="260E368FB81BFFF0FCE4FD97FAEA0F06" blockId="3.[812,1486,232,834]" pageId="3" pageNumber="400">
Although closely related, the species
<taxonomicName id="E1B14D0CB81BFFF0FAB0FD97FBAE0E06" authority="(Kayser, 1892)" baseAuthorityName="Kayser" baseAuthorityYear="1892" class="Lingulata" family="Discinidae" genus="Gigadiscina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lingulida" pageId="3" pageNumber="400" phylum="Brachiopoda" rank="species" species="anomala">
<emphasis id="14C5EA9DB81BFFF0FAB0FD97FCDA0E05" italics="true" pageId="3" pageNumber="400">Gigadiscina anomala</emphasis>
(
<bibRefCitation id="42204B7EB81BFFF0FCC7FDB7FBB60E06" author="Kayser, E." box="[881,1032,553,578]" pageId="3" pageNumber="400" pagination="95 - 107" refId="ref4692" refString="Kayser, E. 1892. Beitrage zur Kenntnis der Fauna der Siegenischen Grauwacke. Jahrbuch der koninglichen preussischen geologischen Landesanstalt 2 (8): 95 - 107." type="journal article" year="1892">Kayser, 1892</bibRefCitation>
)
</taxonomicName>
differs from
<taxonomicName id="E1B14D0CB81BFFF0FB29FDB7FAA70E05" authorityName="Mergl &amp; Massa" authorityYear="2005" box="[1183,1305,553,577]" class="Lingulata" family="Discinidae" genus="Gigadiscina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lingulida" pageId="3" pageNumber="400" phylum="Brachiopoda" rank="species" species="lessardi" status="sp. nov.">
<emphasis id="14C5EA9DB81BFFF0FB29FDB7FAA70E05" box="[1183,1305,553,577]" italics="true" pageId="3" pageNumber="400">G. lessardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0FF657E6B81BFFF0FAA8FDB4FACB0E05" box="[1310,1397,554,577]" pageId="3" pageNumber="400" rank="species">sp. nov.</taxonomicNameLabel>
by having coarser and more distant concentric rugellae on the shell surface. The larger British specimen (
<collectionCode id="40A0AE4AB81BFFF0FB61FDF7FA930EC6" box="[1239,1325,617,642]" pageId="3" pageNumber="400">BMNH</collectionCode>
B 42906) has the dorsal apex situated posteriorly while the apex of
<taxonomicName id="E1B14D0CB81BFFF0FA04FD17FC3B0E85" authorityName="Mergl &amp; Massa" authorityYear="2005" class="Lingulata" family="Discinidae" genus="Gigadiscina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lingulida" pageId="3" pageNumber="400" phylum="Brachiopoda" rank="species" species="lessardi">
<emphasis id="14C5EA9DB81BFFF0FA04FD17FC3B0E85" italics="true" pageId="3" pageNumber="400">G. lessardi</emphasis>
</taxonomicName>
lies anteriorly to midlength. The limited material of
<taxonomicName id="E1B14D0CB81BFFF0FC9AFD57FC0C0EA5" baseAuthorityName="Kayser" baseAuthorityYear="1892" box="[812,946,713,737]" class="Lingulata" family="Discinidae" genus="Gigadiscina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lingulida" pageId="3" pageNumber="400" phylum="Brachiopoda" rank="species" species="anomala">
<emphasis id="14C5EA9DB81BFFF0FC9AFD57FC0C0EA5" box="[812,946,713,737]" italics="true" pageId="3" pageNumber="400">G. anomala</emphasis>
</taxonomicName>
does not allow us to satisfactorily compare the shell outline but, judging from the drawings of
<bibRefCitation id="42204B7EB81BFFF0FACBFD74FCC70F66" author="Kayser, E." pageId="3" pageNumber="400" pagination="95 - 107" refId="ref4692" refString="Kayser, E. 1892. Beitrage zur Kenntnis der Fauna der Siegenischen Grauwacke. Jahrbuch der koninglichen preussischen geologischen Landesanstalt 2 (8): 95 - 107." type="journal article" year="1892">Kayser (1892)</bibRefCitation>
, the
<typeStatus id="F90A882DB81BFFF0FC1BFC97FBAC0F66" box="[941,1042,777,802]" pageId="3" pageNumber="400" type="holotype">holotype</typeStatus>
, and two additional specimens, the species
<taxonomicName id="E1B14D0CB81BFFF0FCE9FCB7FC5D0F05" baseAuthorityName="Kayser" baseAuthorityYear="1892" box="[863,995,809,833]" class="Lingulata" family="Discinidae" genus="Gigadiscina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lingulida" pageId="3" pageNumber="400" phylum="Brachiopoda" rank="species" species="anomala">
<emphasis id="14C5EA9DB81BFFF0FCE9FCB7FC5D0F05" box="[863,995,809,833]" italics="true" pageId="3" pageNumber="400">G. anomala</emphasis>
</taxonomicName>
has slightly more elongate shell.
</paragraph>
</subSubSection>
</treatment>
</document>