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<document ID-DOI="http://dx.doi.org/10.3897/ijm.5.2090" ID-GBIF-Dataset="67fa3f59-684b-4cc3-9a57-3436ea681205" ID-GBIF-Taxon="152016720" ID-Pensoft-Pub="1875-2543-5-1" LSID="urn:lsid:zoobank.org:act:8E73D97B-44EB-4CF7-B394-F471F5A3AD28" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1875-2543-5-1" ModsDocOrigin="International Journal of Myriapodology 5" ModsDocTitle="Review of the millipede genus Eustrongylosoma Silvestri, 1896 in the island of New Guinea, with descriptions of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451252547380" checkinUser="pensoft" docAuthor="Golovatch, Sergei I. &amp; Stoev, Pavel" docDate="2011" docId="AB3A2551968E944B1A9398FCFC386FD3" docLanguage="en" docName="InternJourMyriapod 5: 1-26" docOrigin="International Journal of Myriapodology 5" docSource="http://dx.doi.org/10.3897/ijm.5.2090" docTitle="Eustrongylosoma tumbuna Golovatch &amp; Stoev, 2011, sp. n." docType="treatment" docUuid="8E73D97B-44EB-4CF7-B394-F471F5A3AD28" docUuidSource="ZooBank" docVersion="3" lastPageId="5" lastPageNumber="6" masterDocId="724E36204F7968779912FFD2FFC3892C" masterDocTitle="Review of the millipede genus Eustrongylosoma Silvestri, 1896 in the island of New Guinea, with descriptions of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="26" masterPageNumber="1" pageId="4" pageNumber="5" updateTime="1643719993040" updateUser="ExternalLinkService">
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<mods:title>Review of the millipede genus Eustrongylosoma Silvestri, 1896 in the island of New Guinea, with descriptions of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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<mods:namePart>Stoev, Pavel</mods:namePart>
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<taxonomicName LSID="urn:lsid:zoobank.org:act:8E73D97B-44EB-4CF7-B394-F471F5A3AD28" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma tumbuna" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tumbuna">Eustrongylosoma tumbuna</taxonomicName>
<taxonomicNameLabel pageId="4" pageNumber="5">sp. n.</taxonomicNameLabel>
Figs 7-14
</paragraph>
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<subSubSection pageId="4" pageNumber="5" type="type material">
<paragraph pageId="4" pageNumber="5">Type material.</paragraph>
<paragraph pageId="4" pageNumber="5">Holotype♂(NMNHS),Papua New Guinea, Western Province, Mt Fugilil, near camp, 2980 m, 9 September 1975, B.S.E. (leg. Ph. Chapman &amp; P. Beron). - Paratype: 1 ♂(NMNHS), West Sepik Province, Telefomin area, 1700 m, September 1975, B.S.E. (leg. P. Beron).</paragraph>
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<paragraph pageId="4" pageNumber="5">Name.</paragraph>
<paragraph pageId="4" pageNumber="5">
<normalizedToken originalValue="“tumbuna”">&quot;tumbuna&quot;</normalizedToken>
means
<normalizedToken originalValue="“ancestor”">&quot;ancestor&quot;</normalizedToken>
or
<normalizedToken originalValue="“ancestors”">&quot;ancestors&quot;</normalizedToken>
in Pidgin English. A noun in apposition.
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<paragraph pageId="4" pageNumber="5">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="5">Differs from congeners by the rather wide paratergal calluses strongly delimited not only dorsally, but also ventrally, by the distal part of the solenophore being dentate/lobulate, not split (see also Key below).</paragraph>
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<paragraph pageId="4" pageNumber="5">Description.</paragraph>
<paragraph pageId="4" pageNumber="5">Length of both holo- and paratype ca 20 mm, width of pro- and metazona 1.4 and 2.1 (holotype) or 1.2 and 1.8 mm (paratype), respectively. Coloration rather uniformly light brown to brown, only front 1/3 of body slightly more intensely brown; dorsal sides of prozona, calluses on paraterga, epiproct and venter light yellow-brown to nearly pallid. Sometimes (paratype) region near stricture between pro- and metazona slightly infuscate, brown, thus creating a faint cingulate pattern.</paragraph>
<paragraph pageId="5" pageNumber="6">
<pageBreakToken pageId="5" pageNumber="6" start="start">In</pageBreakToken>
width, head &gt;&gt; collum = segment 3 = 4 &lt;2 &lt;5-16; thereafter body gently tapering towards telson. Entire head densely setose. Antennae rather long, slightly clavate, reaching beyond midlength of segment 3 or 4 dorsally; antennomere 6 longer than 2nd. Collum transversely oval, lateral flaps small, regularly and broadly rounded. Tegument generally smooth and shining, only below paraterga faintly microgranulate; rear halves of metaterga sometimes faintly rugulose. Paraterga of postcollum segments moderately well developed (Figs 7, 8), set low (at about 1/3 midbody height), subhorizontal, evidently thicker in pore-bearing segments than in poreless ones; paratergum 2 with a small anterolateral denticle, its front margin straight; following paraterga without denticulations at lateral margin, sometimes with only a very faint undulation near midway. Caudal corner of all postcollum paraterga pointed to narrowly rounded, starting from 5th extending beyond rear tergal margin. Lateral calluses of paraterga rather wide, especially so on pore-bearing segments, each delimited by a sulcus both dorsally and ventrally. Ozopores lateral to dorsolateral, placed inside an ovoid groove. Tergal setae mostly abraded, retained ones about 1/5 the length of metaterga, pattern traceable at least as 1+1 paramedian insertion points in front row (Figs 7, 8). Axial line nearly wanting, vaguely traceable only here and there on metaterga. Transverse sulcus on metaterga rather superficial, nearly reaching bases of paraterga in metaterga 5-18. Stricture between pro- and metazona rather faintly striolate. Pleurosternal carinae small, complete or nearly so in segments 2-8, a sharp or rounded caudal denticle retained also in segments 9 and 10, thereafter entirely missing. Epiproct (Fig. 9) short, flattened dorsoventrally, digitiform; apical papillae small, tip faintly concave; subapical papillae evident, removed from tip. Hypoproct subtrapeziform, setigerous paramedian knobs at caudal margin well-separated and small.
</paragraph>
<paragraph pageId="5" pageNumber="6">Sterna rather densely setose; a rather high, linguiform, caudally rounded, ventral, setose lamina only between ♂ coxae 4 (Fig. 10). Legs relatively long, midbody ones ca 1.6-1.7 times longer than body height, apparently slightly incrassate and longer compared to ♀; ♂evident tarsal and distotibial brushes increasingly well reduced towards legs of segment 10.</paragraph>
<paragraph pageId="5" pageNumber="6">Gonopods as in Figs 11-14. Tip of solenophore simple, faintly trifid, with three apical denticles or lobules. Femorite with a large rounded lobe (l) somewhat hollow on mesal side.</paragraph>
<caption pageId="5" pageNumber="6">
<paragraph pageId="5" pageNumber="6">
Figures 7-14.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma tumbuna" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tumbuna">Eustrongylosoma tumbuna</taxonomicName>
sp. n., holotype. 7 segment 10, lateral view 8 left half of segment 10, dorsal view 9 tip of epiproct, dorsal view 10 sternal lobe between coxae 4, caudal view 11-14 left gonopod, mesal, ventral, sublateral and subdorsal views, respectively. Scale bars: 1.0 (7-10) and 0.3 mm (11-14).
</paragraph>
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<taxonomicName id="BC18A74A6709D2B978634BD52EBC3B81" ID-CoL="KJ46" authority="Bey-Bienko, 1958" authorityName="Bey-Bienko" authorityYear="1958" class="Insecta" family="Ectobiidae" genus="Balta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balta hwangorum" order="Blattodea" pageId="11" pageNumber="24" phylum="Arthropoda" rank="species" species="hwangorum">Balta hwangorum Bey-Bienko, 1958</taxonomicName>
Figs 9, 10, 64-75
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<taxonomicName id="9F933B32F73BDFE5C46F2381F17E6F13" class="Insecta" family="Ectobiidae" genus="Balta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balta hwangorum" order="Blattodea" pageId="11" pageNumber="24" phylum="Arthropoda" rank="species" species="hwangorum">Balta hwangorum</taxonomicName>
Bey-Bienko, 1958: 676, 688 (Type locality: Yunnan, China);
<bibRefCitation id="BE2D51C54BF433A65715AC2FA9CF563C" pageId="11" pageNumber="24">Princis 1969</bibRefCitation>
: 978.
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<taxonomicName id="35B9F487126BA982C4D9A9F09C6C9CAD" class="Insecta" family="Ectobiidae" genus="Balta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balta picea" order="Blattodea" pageId="11" pageNumber="24" phylum="Arthropoda" rank="species" species="picea">Balta picea</taxonomicName>
Bey-Bienko, 1958: 677 (Holotype, female) (Type locality: Yunnan, China). Syn. n.
</paragraph>
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<paragraph id="2112D1B9F34902B80961139768E92CED" pageId="11" pageNumber="24">Material examined.</paragraph>
<paragraph id="49CB92EC84A105C0FDF710F06BFF627B" pageId="11" pageNumber="24">
Deposited in IESWU. China, Yunnan Prov.: 3 males, Xishuangbanna, Damenglong, 650 m, 13 April 1958, coll. Zhizi Chen; 5 males, Xishuangbanna, Menghun, 650-750 m, 9 June 1958, coll. Xuwu Meng; 1 male and 1 female, Xishuangbanna, Jinghong, 650 m, 6-24 July 1958, coll. Junhua He; 1 male, Xishuangbanna, Menghun, 650-750m, 1 June 1958, coll. Leyi Zheng; 1 male, Xishuangbanna, Menghun, 650-1080 m, 7 June 1958, coll. Chunpei Hong and Shuyong Wang; 1 male, Xishuangbanna, Damenglong, 650 m, 14 April 1958, coll. Chunpei Hong; 1 male, Xishuangbanna, Menghun, 650-750 m, 3 February 1958, coll. Shuyong Wang; 2 males, Xishuangbanna, Menga, 1050-1080 m, 20 May 1958, coll. Fuji Pu; 1 male, Xishuangbanna, Menghun, 650-750 m, 13 June 1958, coll. Yiran Zhang; 1 male, Xishuangbanna, Menghun, 650-750 m, 18 April 1958, coll. Leyi Zheng; 1 male, Xishuangbanna, Xiaomengyang, 850 m, 13 June 1958, coll. Lingchao Zang; 2 males and 1 female, Xishuangbanna, Menglun, G213 (National road) bamboo forest, 627 m,
<geoCoordinate id="202FBC9824567C57F1F3972E1B579A0F" direction="north" orientation="latitude" precision="1" value="21.906334">21°54.380N</geoCoordinate>
,
<geoCoordinate id="1E815C1F8526C8F690856B1B7CC65971" direction="east" orientation="longitude" precision="1" value="101.28025">101°16.815E</geoCoordinate>
, 21-26 November 2009, coll. Guo Tang and Zhiyuan Yao; 2 males and 2 females, Xishuangbanna, Menglun, G213 (National road) bamboo forest, 627 m,
<geoCoordinate id="487115F422BDB956748B5D523D1231E1" direction="north" orientation="latitude" precision="1" value="21.906334">21°54.380N</geoCoordinate>
,
<geoCoordinate id="E1B99775D9B33CE6DEC5D237AEB7CD31" direction="east" orientation="longitude" precision="1" value="101.28025">101°16.815E</geoCoordinate>
, 22 November 2009, coll. Guo Tang and Zhiyuan Yao; 2 males and 2 females, Xishuangbanna, Menglun, G213 (National road) secondary forest, 644 m,
<geoCoordinate id="6918E0E74502B92C9F23CDC5332F724E" direction="north" orientation="latitude" precision="1" value="21.907316">21°54.439N</geoCoordinate>
,
<geoCoordinate id="01EFD622CFD756E704912F76CB8C373C" direction="east" orientation="longitude" precision="1" value="101.27925">101°16.755E</geoCoordinate>
, 20 November 2009, coll. Guo Tang and Zhiyuan Yao; 1 male, Mojiang, 1 May 2013, coll. Zongqing Wang. China: Guangxi: 1 female, Pingxiang, 850 m, 11 May 1963, coll. Sikong Liu.
</paragraph>
</subSubSection>
<subSubSection id="F44E545B6D5BF2B4E3245E64A6C7B387" lastPageId="12" lastPageNumber="25" pageId="11" pageNumber="24" type="redescription">
<paragraph id="2193605855541E0629B66BD32D115396" pageId="11" pageNumber="24">Redescription.</paragraph>
<paragraph id="85EF43F771FE9E954C6CCE5039F61EEB" lastPageId="12" lastPageNumber="25" pageId="11" pageNumber="24">
Male. Body yellowish brown (Figs 9, 10). Vertex to anterior margin of antennal sockets reddish brown, face yellowish brown to blackish brown (Figs 9, 10). Pronotum with inconspicuous black brindle or two oblique and symmetrical stripes, with disk reddish brown to black brown, two lateral borders, anterior and posterior margin yellowish brown and hyaline (Fig. 66). Legs yellowish brown to blackish
<pageBreakToken id="21BE62E9F819A9B3D9B6C654629B4988" pageId="12" pageNumber="25" start="start">brown</pageBreakToken>
. Abdomen blackish brown. Apex of subgenital plate with reddish brown spots or without. Fourth and fifth maxillary palpomeres approximately same length, both distinctly shorter than the third (Fig. 65). Tegmen with M and CuA slightly oblique (Fig. 67). Hind wing with M without branches, CuA with five complete branches and without incomplete ones (Fig. 68). Front femur type C2 (Fig. 69), tarsal claws strongly asymmetrical and unspecialized (Fig. 70). Abdominal tergites unspecialized.
</paragraph>
<caption id="BAFD67B744F95003BF2BF4223A0B946A" pageId="12" pageNumber="25">
<paragraph id="E6536FB89A7B4EA9DA38CC7C257AF90E" pageId="12" pageNumber="25">
Figures 64-75.
<taxonomicName id="AAB365BF0AD8C66661F1DE8906D7856E" class="Insecta" family="Ectobiidae" genus="Balta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balta hwangorum" order="Blattodea" pageId="12" pageNumber="25" phylum="Arthropoda" rank="species" species="hwangorum">Balta hwangorum</taxonomicName>
Bey-Bienko, 1958. 64 head 65 maxillary palps 66 pronotum; 67 tegmen 68 hind wing 69 front femur 70 tarsal claws and arolium 71 supra-anal plate and paraprocts, ventral view 72 subgenital plate, dorsal view 73 right phallomere 74 median phallomere 75 left phallomere. Scale bars: 0.5 mm (64-66, 69-75), 2.0 mm (67, 68).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="EE56D23BCF6EC2A7EC14E3EAD4AD245C" pageId="12" pageNumber="25" type="male genitalia">
<paragraph id="BD893B23DFB2A9DF04DB3A0B5914C5EE" pageId="12" pageNumber="25">Male genitalia.</paragraph>
<paragraph id="8449B32D321324AE5641FA427FE2D181" pageId="12" pageNumber="25">Supra-anal plate in ventral view short, hind margin arched, slightly emarginated in the middle; right and left paraprocts similar (Fig. 71). Subgenital plate symmetrical throughout except at the base, hind margin with deep V-shape emargination, styli similar, conical, arising on lateral lobes (Fig. 72). Hook-like phallomere on right side, hook slender (Fig. 73). Median phallomere sticklike, slightly curved, apex acuminate, near the apex part with a sclerite, base of arc appendage with brush-like structure (Fig. 74). Left phallomere complex, apex with a bifurcate spinous structure and a slender hairs structure (Fig. 75).</paragraph>
</subSubSection>
<subSubSection id="99CC73944F0A32B5811967577E3E6AFA" pageId="12" pageNumber="25" type="female">
<paragraph id="ED217D70413203CF668D8D76A55E3F47" pageId="12" pageNumber="25">Female.</paragraph>
<paragraph id="51725E0A13B612D3AE8785B153BE97FF" pageId="12" pageNumber="25">Slight sexual dimorphism in that the female body is smaller than the male. Supra-anal plate symmetrical, rounded, hind margin divided in the middle. Subgenital plate broad and rounded.</paragraph>
</subSubSection>
<subSubSection id="99D8DED80591B347FC33E7BAE1AACC6F" pageId="12" pageNumber="25" type="description">
<paragraph id="91C0F6334EA2B70C7D8B5ED5061C551F" pageId="12" pageNumber="25">Measurement (mm).</paragraph>
<paragraph id="1EEA8DBE939710F9C30496F970EE06C6" pageId="12" pageNumber="25">
Overall length male 15.0-18.5, female 14.0-17.1; tegmen length male 13.5-16.0, female 11.2-14.0; pronotum length
<normalizedToken id="1EF18F3999C2D16B06C4379D8838EDC4" originalValue="×">x</normalizedToken>
width male 2.9-3.5
<normalizedToken id="A7DB6BB5253C6F5FAC8F54492142E172" originalValue="×">x</normalizedToken>
4.1-4.9, female 3.0-3.1
<normalizedToken id="6A6EBA30FFC5CDE3D7E7B50031D1DE22" originalValue="×">x</normalizedToken>
5.0-5.1.
</paragraph>
</subSubSection>
<subSubSection id="19F6F38365A7996E0C40F724976639B9" pageId="12" pageNumber="25" type="remarks">
<paragraph id="AA8A0AC46B2659D776E2DDE8F00AE2C6" pageId="12" pageNumber="25">Remarks.</paragraph>
<paragraph id="3409917E0DE60429BC6D55281AD4E28F" pageId="12" pageNumber="25">
Several specimens were separated as two species by Bey-Bienko, mainly based on the body color (1958). However, after examining a large number of specimens, the results show that body color of
<taxonomicName id="1FDF0FA3CC747B4B6C39720AB2B21047" lsidName="B. hwangorum" pageId="12" pageNumber="25" rank="species" species="hwangorum">B. hwangorum</taxonomicName>
varies. Moreover, in
<normalizedToken id="13858DEE0C34CE9133BB0BD27018A581" originalValue="Bey-Bienkos">Bey-Bienko's</normalizedToken>
records, in
<taxonomicName id="928EDE97A418E89A01848F895352DEFD" lsidName="B. hwangorum" pageId="12" pageNumber="25" rank="species" species="hwangorum">B. hwangorum</taxonomicName>
, the length of lobes of the subgenital plate is different among the species, but in fact we find no difference.
</paragraph>
</subSubSection>
<subSubSection id="BBACE95EB3D3EA11CF4FD1C5E7737879" pageId="12" pageNumber="25" type="distribution">
<paragraph id="99DB98E70032EA6922A53F20642189C1" pageId="12" pageNumber="25">Distribution.</paragraph>
<paragraph id="AB4FC221F6AAF6EDC2267D72A35A58BE" pageId="12" pageNumber="25">China (Yunnan, Guangxi).</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="6E0F546FA808A03BF30A865354D7801F" ID-CoL="4GY3G" authority="Habermehl, 1925" authorityName="Habermehl" authorityYear="1925" class="Insecta" family="Ichneumonidae" genus="Phthorima" kingdom="Animalia" lsidName="Phthorima picta" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="picta">Phthorima picta (Habermehl, 1925)</taxonomicName>
</paragraph>
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<paragraph id="FC09D8C727BE85350432A04EE4D40743" pageId="0" pageNumber="9042">
<taxonomicName id="080E7B0C8F9E3C184146301FCA37F05E" class="Insecta" family="Ichneumonidae" genus="Phthorimus" kingdom="Animalia" lsidName="Phthorimus picta" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="picta">Phthorimus picta</taxonomicName>
Habermehl, 1925
</paragraph>
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<taxonomicName id="0B75A28DFE9CAAC19AB3FCB22FC0DFF4" class="Insecta" family="Ichneumonidae" genus="Ethelurgina" kingdom="Animalia" lsidName="Ethelurgina gaullei" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="gaullei">gaullei</taxonomicName>
Seyrig, 1928
</paragraph>
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<paragraph id="EB46269B38E4FA2A2761DAE595BC990F" pageId="0" pageNumber="9042">England, Ireland</paragraph>
</subSubSection>
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<paragraph id="49E6B05A4E8FF9EC5C62E739FA57985F" pageId="0" pageNumber="9042">Notes</paragraph>
<paragraph id="0A31E9EAE43BAC42A151CAAFE9C12894" pageId="0" pageNumber="9042">added by Fitton and Boston (1988)</paragraph>
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<paragraph pageId="175" pageNumber="176">Taxon classification Animalia Coleoptera Carabidae</paragraph>
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<paragraph pageId="175" pageNumber="176">
(072)
<taxonomicName authority="Blackburn" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax angusticollis" order="Coleoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="angusticollis">Mecyclothorax angusticollis (Blackburn)</taxonomicName>
Figs 90C, 91
<normalizedToken originalValue="DE">D-E</normalizedToken>
, 92A, 93A, 94
</paragraph>
</subSubSection>
<subSubSection pageId="175" pageNumber="176" type="reference_group">
<paragraph pageId="175" pageNumber="176">
<taxonomicName class="Insecta" family="Carabidae" genus="Cyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclothorax angusticollis" order="Coleoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="angusticollis">Cyclothorax angusticollis</taxonomicName>
<bibRefCitation author="Blackburn, T" journalOrPublisher="The Entomologist's Monthly Magazine" pageId="307" pageNumber="308" pagination="156 - 158" title="Characters of new genera and descriptions of new species of Geodephaga from the Hawaiian Islands." volume="15" year="1878 b">Blackburn 1878b</bibRefCitation>
: 156;
<bibRefCitation author="Blackburn, T" journalOrPublisher="Memoirs on the Coleoptera of the Hawaiian Islands (Ser 2)" pageId="307" pageNumber="308" pagination="119 - 290" volume="3" year="1885">Blackburn and Sharp 1885</bibRefCitation>
: 216.
</paragraph>
<paragraph pageId="175" pageNumber="176">
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax angusticollis" order="Coleoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="angusticollis">Mecyclothorax angusticollis</taxonomicName>
,
<bibRefCitation author="Sharp, D" editor="Sharp, D" journalOrPublisher="The University Press, Cambridge" pageId="310" pageNumber="311" pagination="175 - 292" title="Coleoptera II. Caraboidea." volumeTitle="Fauna Hawaiiensis" year="1903">Sharp 1903</bibRefCitation>
: 246;
<bibRefCitation author="Britton, EB" journalOrPublisher="Bishop Museum" pageId="307" pageNumber="308" pagination="107 - 166" title="A revision of the Hawaiian species of Mecyclothorax (Coleoptera: Carabidae). Occasional Papers of the Bernice P." volume="19" year="1948 b">Britton 1948b</bibRefCitation>
: 155.
</paragraph>
</subSubSection>
<subSubSection pageId="175" pageNumber="176" type="diagnosis">
<paragraph pageId="175" pageNumber="176">Diagnosis.</paragraph>
<paragraph pageId="175" pageNumber="176">
This species is distinguished from all others in the group by the transverse pronotum (Fig. 90C), MPW/PL = 1.21-1.28, that is broad relative to the elytra, MEW/MPW = 1.35-1.39. Most similar to
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax minor" order="Coleoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="minor">Mecyclothorax minor</taxonomicName>
,
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax angusticollis" order="Coleoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="angusticollis">Mecyclothorax angusticollis</taxonomicName>
can also be distinguished from that species by the distinct isodiametric sculpticells in transverse rows on the elytral disc and apex. Body coloration is darker as well, with the pronotal disc rufobrunneous to rufopiceous, and the elytra rufobrunneous, in some instances with an apical rufopiceous cloud. Setal formula 2 2 2 0. Standardized body length 3.2-3.7 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="175" pageNumber="176" type="identification">
<paragraph pageId="175" pageNumber="176">Identification</paragraph>
<paragraph pageId="175" pageNumber="176">
(n = 5). The eyes are relatively flat, ocular ratio = 1.32-1.39, and cover about
<normalizedToken originalValue="¾">3/4</normalizedToken>
of the ocular lobe, ocular lobe ratio = 0.75-0.79. The labrum is medially emarginate 1/5-1/6 of its length. The pronotal hind angles are obtuse rounded and little projected, the lateral margin only briefly subparallel before the angle. The pronotal median base is slightly depressed relative to the disc, but well set off by the dense elongate punctures and wrinkles, and the upraised isodiametric and transverse sculpticells over the surface. The elytral striae are well developed, with striae 1-6 and 8 complete and moderately deep on disc, stria 7 shallower. Striae 1-6 are distinctly punctate in basal
<normalizedToken originalValue="¾">3/4</normalizedToken>
of length. Sutural stria deep to apex, stria 2 shallower in apical half, except for deep portion parallel to deepened apex of stria 7. The 8th interval is broadly convex, slightly more so than inner intervals. The lateral elytral setae are arranged in an anterior series of 6 setae and a posterior series of 4 setae.
</paragraph>
<paragraph pageId="175" pageNumber="176">
Male genitalia (n = 2). Aedeagal median lobe gracile, distance from parameral articulation to tip 3.9
<normalizedToken originalValue="×">x</normalizedToken>
depth at midlength (Fig. 91D); apex extended for twice breadth beyond ostial opening, dorsal surface broadly expanded before evenly rounded tip; median lobe curved rightward near apex in ventral view, right and left margins parallel before bluntly rounded apex that is offset to the right (Fig. 91E); internal sac with small, well-sclerotized ventral ostial microtrichial patch (on left of sac, Fig. 91E); flagellar plate of moderate size, length 0.40
<normalizedToken originalValue="×">x</normalizedToken>
parameral articulation-tip distance (estimated by viewing ovoid sclerotized plate in ventral view, Fig. 91E).
</paragraph>
<paragraph pageId="175" pageNumber="176">
Female reproductive tract (n = 1). Bursa copulatrix columnar with apical cap separated by constriction, overall length 0.67 mm, apical cap breadth 0.23 mm, medial shaft breadth 0.30 mm (Fig. 92A); bursal walls translucent with thin wrinkles; gonocoxite 1 with 3 small apical fringe setae and 6 very small setae on medial surface (Fig. 93A); gonocoxite 2 broadly subtriangular, apex evenly curved laterally, base evenly expanded laterally, 2 lateral ensiform setae, apical nematiform setae on medial surface at 0.67
<normalizedToken originalValue="×">x</normalizedToken>
gonocoxite length.
</paragraph>
<caption pageId="175" pageNumber="176">
<paragraph pageId="175" pageNumber="176">
Figure 92. Female bursa copulatrix and associated reproductive structures,
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax microps" order="Coleoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="microps">Mecyclothorax microps</taxonomicName>
group species, ventral view (for abbreviations see Table 2, p. 23). A
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax angusticollis" order="Coleoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="angusticollis">Mecyclothorax angusticollis</taxonomicName>
(nr. Ukulele Camp, 1525 m) B
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax contractus" order="Coleoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="contractus">Mecyclothorax contractus</taxonomicName>
(Kuhiwa, 1585 m). Scale bar = 0.50 mm.
</paragraph>
</caption>
<caption pageId="175" pageNumber="176">
<paragraph pageId="175" pageNumber="176">
Figure 93. Left female gonocoxa,
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax microps" order="Coleoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="microps">Mecyclothorax microps</taxonomicName>
group species, ventral view. A
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax angusticollis" order="Coleoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="angusticollis">Mecyclothorax angusticollis</taxonomicName>
(nr. Ukulele Camp, 1525 m) B
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax contractus" order="Coleoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="contractus">Mecyclothorax contractus</taxonomicName>
(Kuhiwa, 1585 m).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="175" pageNumber="176" type="lectotype">
<paragraph pageId="175" pageNumber="176">Lectotype.</paragraph>
<paragraph pageId="175" pageNumber="176">
Male (BMNH) hereby designated, labeled: mounting platen with Blackburn Maui code (
<bibRefCitation author="Zimmerman, EC" journalOrPublisher="University of Hawai'i Press, Honolulu" pageId="310" pageNumber="311" title="Ephemeroptera-Neuroptera-Trichoptera and Supplement to Volumes 1 to 5." volumeTitle="Insects of Hawai'i" year="1957">Zimmerman 1957</bibRefCitation>
: 210), angustic (on reverse) // Type // Hawaiian Is. Rev. T. Blackburn 1888-30. // Lectotype
<taxonomicName class="Insecta" family="Carabidae" genus="Cyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclothorax angusticollis" order="Coleoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="angusticollis">Cyclothorax angusticollis</taxonomicName>
Blackburn J.K.Liebherr 1998 (black-margined red label).
</paragraph>
</subSubSection>
<subSubSection pageId="176" pageNumber="177" type="distribution and habitat">
<paragraph pageId="176" pageNumber="177">
<pageBreakToken pageId="176" pageNumber="177" start="start">Distribution</pageBreakToken>
and habitat.
</paragraph>
<paragraph pageId="176" pageNumber="177">
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax angusticollis" order="Coleoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="angusticollis">Mecyclothorax angusticollis</taxonomicName>
was collected repeatedly by Blackburn and Perkins in the late 19th Century (Perkins lots 112, 115, 120, 251, 350, 366, 367, 369, 371, 372, 597, 604, 605, 615, 680;
<bibRefCitation author="Anonymous," journalOrPublisher="Dept. of Entomology Library, The Natural History Museum, London" pageId="307" pageNumber="308" title="Key to numbers &amp; localities of Mr. Perkins' Hawaiian insects" year="N D">Anonymous N D</bibRefCitation>
). Yet the only 20th Century record was a single specimen collected by beating vegetation at 1830-1980 m elevation (E.C. Zimmerman, BPBM). This species appears to have been a species of the forest edge (Fig. 94), with the historical collecting sites now extensively disturbed, the old forest edge gone.
</paragraph>
<caption pageId="176" pageNumber="177">
<paragraph pageId="176" pageNumber="177">
Figure 94. Recorded geographic distributions of
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax microps" order="Coleoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="microps">Mecyclothorax microps</taxonomicName>
group species.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,104 @@
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<paragraph id="4C441D20CA818B0659E3420BEDAD144D" pageId="0" pageNumber="1066">Materials</paragraph>
<paragraph id="17FF3A79CFC3B155EE85D491BDD31C0B" pageId="0" pageNumber="1066">
<materialsCitation id="737BD6C8E5D0EA28B81EB963E2D5BA81" collectingDate="28 September 2013" collectorName="P. Decker &amp; M. Koehler" country="Germany" latitude="52.5102" location="Berlin" longitude="13.3366" pageId="0" pageNumber="1066" specimenCount="1" typeStatus="Other material">
Type status:
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. Occurrence: recordedBy:
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P. Decker &amp; M.
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; individualCount:
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; disposition: SMNG; Location: country:
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; locality:
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; verbatimLocality: Berlin Zoological Garden; decimalLatitude:
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; decimalLongitude:
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; geodeticDatum: WGS84; Event: eventDate:
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</paragraph>
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<materialsCitation id="24BEA270CAAA53BFCC2AA8E410776EC3" collectingDate="26 March 2013, 05 April 2013, 28 July 2013" collectorName="E. N. Lindner" country="Germany" latitude="51.3505" location="Leipzig" longitude="12.3716" pageId="0" pageNumber="1066" specimenCount="33" typeStatus="Other material">
Type status:
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. Occurrence: recordedBy:
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; individualCount:
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; disposition: SMNG; Location: country:
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; locality:
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; verbatimLocality: Zoological Garden Leipzig, Gondwanaland; decimalLatitude:
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; geodeticDatum: WGS84; Event: eventDate:
<collectingDate id="A7EB38082D9EF91B4C5E6224C419E017" pageId="0" pageNumber="1066" value="26 March 2013, 05 April 2013, 28 July 2013">26 March 2013, 05 April 2013, 28 July 2013</collectingDate>
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<paragraph id="D193A784FCAD86BDF8D0C7F569920028" pageId="0" pageNumber="1066">Distribution</paragraph>
<paragraph id="097D1D19B4B527348B9542104661DAFE" pageId="0" pageNumber="1066">Southeast Asia</paragraph>
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