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<document ENCODING="UTF8" ID-DOI="10.5281/zenodo.11581" ID-GBIF-Dataset="2eed386a-5aba-4ebb-869b-f28fc7adcd33" checkinTime="1424029685315" checkinUser="donat" docAuthor="Torsten Dikow &amp; Jason G. H. Londt" docDate="2000" docId="AB0C1FCBEEEBCF9FF62D61A12148ACB1" docLanguage="en" docName="58B9661576FE0FF007961939B25C81C2" docOrigin="Ann. Natal Mus. 41" docTitle="Anasillomos" docType="treatment" docVersion="17" lastPageNumber="111" masterDocId="58B9661576FE0FF007961939B25C81C2" masterDocTitle="A review of the genera Anasillomos Londt, 1983, Oratostylum Ricardo, 1925, and Remotomyia Londt, 1983, with description of a new genus and two new species (Diptera: Asilidae: Stenopogoninae)" masterLastPageNumber="121" masterPageNumber="107" pageNumber="111" updateTime="1682087710716" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A review of the genera Anasillomos Londt, 1983, Oratostylum Ricardo, 1925, and Remotomyia Londt, 1983, with description of a new genus and two new species (Diptera: Asilidae: Stenopogoninae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Torsten Dikow</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jason G. H. Londt</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Ann. Natal Mus.</mods:title>
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<mods:date>2000</mods:date>
<mods:detail type="volume">
<mods:number>41</mods:number>
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<mods:start>107</mods:start>
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<mods:identifier type="DOI">10.5281/zenodo.11581</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6280019" ID-GBIF-Taxon="125136127" ID-Zenodo-Dep="6280019" LSID="urn:lsid:plazi:treatment:AB0C1FCBEEEBCF9FF62D61A12148ACB1" httpUri="http://treatment.plazi.org/id/AB0C1FCBEEEBCF9FF62D61A12148ACB1" lastPageNumber="111" pageNumber="111">
<subSubSection type="nomenclature">
<paragraph pageId="4" pageNumber="111" type="mainText">
<taxonomicName ID-CoL="V97" ID-ENA="1717515" class="Insecta" family="Asilidae" genus="Anasillomos" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="111" phylum="Arthropoda" rank="genus">
<pageBreakToken pageId="4" pageNumber="111" start="start">Anasillomos</pageBreakToken>
</taxonomicName>
Londt, 1983
</paragraph>
</subSubSection>
<subSubSection type="reference_group">
<paragraph pageId="4" pageNumber="111" type="mainText">
<taxonomicName class="Insecta" family="Asilidae" genus="Anasillomos" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="111" phylum="Arthropoda" rank="genus">Anasillomos</taxonomicName>
Londt, 1983: 284;
<typeStatus>Type</typeStatus>
species:
<taxonomicName class="Insecta" family="Asilidae" genus="Anasillomos" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="111" phylum="Arthropoda" rank="species" species="chrysopos">Anasillomos chrysopos</taxonomicName>
Londt, 1983, by original designation.
</paragraph>
</subSubSection>
<subSubSection type="diagnosis">
<paragraph pageId="4" pageNumber="111" type="mainText">Generic diagnosis (based on the original diagnosis of Londt 1983):</paragraph>
</subSubSection>
<subSubSection type="description">
<paragraph pageId="4" pageNumber="111" type="mainText">
Antennal style with three elements (2 segments and a terminal seta-like element); postpedicel cylindrical, longer than scape and pedicel combined; palpi shorter than half length of proboscis; facial swelling pronounced on lower margin, moderately developed in upper part; proepisternum with strong macrosetae and weaker setae; pronotum with strong macrosetae; basal postpronotal setae well-developed and strong; 1-2 pairs of dorsocentral setae anterior of transverse suture; cell r
<sub pageId="4" pageNumber="111">5</sub>
closed and stalked; cell m
<sub pageId="4" pageNumber="111">3</sub>
narrowly open or just closed at wing margin; terga with median stripe apparently free of setae, laterally-situated setae directed posterolaterally; T6-7 of
<specimenCount type="female"></specimenCount>
abdomen with setae strongly developed and anteriorly directed (directed posteriorly on
<collectionCode>T</collectionCode>
1-5); epandrial lobes long and curved.
</paragraph>
</subSubSection>
</treatment>
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.439.7789" ID-GBIF-Dataset="8b3eb75d-2b12-43ec-a236-8a4f043e2a06" ID-PMC="PMC4196254" ID-Pensoft-Pub="1313-2970-439-19" ID-PubMed="25317055" ID-ZBK="30AC763E9C3146E28CAA8D6CD8461F60" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-439-19" ModsDocOrigin="ZooKeys 439" ModsDocTitle="New Chinese record of the genus Spinonychiurus (Collembola, Onychiuridae), with the description of a new species" checkinTime="1451245384479" checkinUser="pensoft" docAuthor="Sun, Xin &amp; Li, Yu" docDate="2014" docId="AB0C23D47EAE15E953CB2CDCE42FD224" docLanguage="en" docName="ZooKeys 439: 19-26" docOrigin="ZooKeys 439" docSource="http://dx.doi.org/10.3897/zookeys.439.7789" docTitle="Spinonychiurus sinensis Sun &amp; Li, 2014, sp. n." docType="treatment" docUuid="BC4B8586-C7AE-4DB2-A1FB-E02B17174A8A" docUuidSource="ZooBank" docVersion="5" lastPageNumber="23" masterDocId="EC11FFCFFFF7FF80FF87FFAFFFB2FFDC" masterDocTitle="New Chinese record of the genus Spinonychiurus (Collembola, Onychiuridae), with the description of a new species" masterLastPageNumber="26" masterPageNumber="19" pageNumber="21" updateTime="1668159270872" updateUser="ExternalLinkService">
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<mods:title>New Chinese record of the genus Spinonychiurus (Collembola, Onychiuridae), with the description of a new species</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sun, Xin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Yu</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>439</mods:number>
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<mods:extent unit="page">
<mods:start>19</mods:start>
<mods:end>26</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.439.7789</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.439.7789</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-439-19</mods:identifier>
<mods:identifier type="ZBK">30AC763E9C3146E28CAA8D6CD8461F60</mods:identifier>
<mods:identifier type="ZooBank">30AC763E9C3146E28CAA8D6CD8461F60</mods:identifier>
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<treatment ID-GBIF-Taxon="152055474" LSID="urn:lsid:zoobank.org:act:BC4B8586-C7AE-4DB2-A1FB-E02B17174A8A" httpUri="http://treatment.plazi.org/id/AB0C23D47EAE15E953CB2CDCE42FD224" lastPageId="4" lastPageNumber="23" pageId="2" pageNumber="21">
<subSubSection pageId="2" pageNumber="21" type="multiple">
<paragraph pageId="2" pageNumber="21">Taxon classification Animalia Poduromorpha Onychiuridae</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="21" type="nomenclature">
<paragraph pageId="2" pageNumber="21">
<taxonomicName LSID="http://zoobank.org/BC4B8586-C7AE-4DB2-A1FB-E02B17174A8A" class="Collembola" family="Onychiuridae" genus="Spinonychiurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinonychiurus sinensis" order="Poduromorpha" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="sinensis">Spinonychiurus sinensis</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="21">sp. n.</taxonomicNameLabel>
Figs 1-2
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="21" type="type material">
<paragraph pageId="2" pageNumber="21">Type material.</paragraph>
<paragraph pageId="2" pageNumber="21">
Holotype: female; paratypes: 3 females and 1 male on slides - China, Jilin, Changbai Mountain Range (alt. 689 m,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.0376">43.0376°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="128.9965">128.9965°E</geoCoordinate>
), 3.Oct.2011, litter and soil, Berlese extraction, leg. Tang Xuguang.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="21" type="diagnosis">
<paragraph pageId="2" pageNumber="21">Diagnosis.</paragraph>
<paragraph pageId="2" pageNumber="21">Pso formula as 32/133/33343 dorsally, absent ventrally, subcoxa 1 of legs I-III with 1, 1 and 1 pso respectively; psx formula as 11/000/122211m ventrally, absent dorsally, subcoxa 1 of legs I-III with 1, 1 and 1 psx respectively; S-chaetae formula as 1/011/111021 dorsally, 11/000/000110 ventrally; sterna of Th. I, II, and III with 0+0, 2+2, 2+2 chaetae respectively; Abd. IV tergum with axial chaeta p0, Abd. V tergum with a0 and m0, Abd. VI tergum with a0; the distal row of tibiotarsi with 7 chaetae; male ventral organ absent; anal spines present, 0.8 times as long as inner edge of hind unguis.</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="23" pageId="2" pageNumber="21" type="description">
<paragraph pageId="2" pageNumber="21">Description.</paragraph>
<paragraph pageId="2" pageNumber="21">
Body white in alcohol. Size 970-1200
<normalizedToken originalValue="µm">µm</normalizedToken>
in females, 900
<normalizedToken originalValue="µm">µm</normalizedToken>
in male; holotype: 1200
<normalizedToken originalValue="µm">µm</normalizedToken>
. Body subcylindrical, body sides parallel.
</paragraph>
<paragraph pageId="2" pageNumber="21">
Pseudocellar formula: 32/133/33343 dorsally, absent ventrally (Figs 1A, G, 2A), subcoxa 1 of legs
<normalizedToken originalValue="IIII">I-III</normalizedToken>
with 1, 1 and 1 pso respectively. Parapseudocellar formula: 11/000/122211m ventrally, absent dorsally (Figs 1A, 1G, 2A), subcoxa 1 of legs
<normalizedToken originalValue="IIII">I-III</normalizedToken>
with 1, 1 and 1 psx respectively. Pseudopore formula: 0/011/11110 dorsally, -/111/- ventrally (Figs 1A, G, 2A).
</paragraph>
<caption pageId="2" pageNumber="21">
<paragraph pageId="2" pageNumber="21">
Figure 1.
<taxonomicName class="Collembola" family="Onychiuridae" genus="Spinonychiurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinonychiurus sinensis" order="Poduromorpha" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="sinensis">Spinonychiurus sinensis</taxonomicName>
sp. n. A dorsal chaetotaxy of body B maxillary palp C labium D postantennal organ E antenna F distal part of leg III G chaetotaxy of Abd.
<normalizedToken originalValue="IIVI">II-VI</normalizedToken>
sterna. Scale bars: 0.1 mm (A, E, G), 0.01 mm (
<normalizedToken originalValue="BD">B-D</normalizedToken>
, F).
</paragraph>
</caption>
<caption pageId="2" pageNumber="21">
<paragraph pageId="2" pageNumber="21">
Figure 2.
<taxonomicName class="Collembola" family="Onychiuridae" genus="Spinonychiurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinonychiurus sinensis" order="Poduromorpha" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="sinensis">Spinonychiurus sinensis</taxonomicName>
sp. n. A ventral chaetotaxy of head
<normalizedToken originalValue="BC">B-C</normalizedToken>
central part of abdominal sternum IV D dorsal chaetotaxy of Abd.
<normalizedToken originalValue="IVVI">IV-VI</normalizedToken>
. Scale bars: 0.1 mm (A, D), 0.01 mm (
<normalizedToken originalValue="BC">B-C</normalizedToken>
).
</paragraph>
</caption>
<paragraph lastPageId="3" lastPageNumber="22" pageId="2" pageNumber="21">
Head. Antennae short and distinctly segmented, as long as head. Length ratio of Ant. I: II: III: IV as about 1: 2: 2: 2. Ant. IV with two distinct thickened S-chaetae, subapical organite with globular apex; basolateral ms at approx. half length from base, above the second proximal row of chaetae (Fig. 1E). AIIIO composed of 5 papillae, 5 guard chaetae, 2 sensory rods and 2 smooth sensory clubs, the inner bigger than the outer, and a lateral ms (Fig. 1E). Ant. II with 15 chaetae. Ant. I with 9 chaetae. Antennal base not marked (Fig. 1D). PAO with 12-13 compound vesicles (Fig. 1D). Dorsal cephalic chaeta d0 absent (Fig. 1A). 3+3 p-chaetae present between two inner posterior pso, p1 in line with others (Fig. 1E). Mandible with strong molar fig and 4 apical teeth. Maxilla bearing 3 teeth and 6 lamellae. Maxillary palp simple with 1 basal chaeta, without sublobal hair (Fig. 1B). Labral formula 4/1,4,2; labium with 6 proxi
<pageBreakToken pageId="3" pageNumber="22" start="start">mal</pageBreakToken>
, 4 basomedian (E, F, G, f) and 6 basolateral (a, b, c, d, e,
<normalizedToken originalValue="e">e'</normalizedToken>
) chaetae (Fig. 2A); labial type A, papillae
<normalizedToken originalValue="AE">A-E</normalizedToken>
respectively with 1, 4, 0, 3 and 3 guard chaetae (Fig. 1C). Head ventrally with 4+4 postlabial chaetae along ventral groove (Fig. 2A).
</paragraph>
<paragraph pageId="3" pageNumber="22">
Body chaetotaxy. S-chaetae subcylindrical, apically rounded, 1/011/111021 dorsally, 11/000/000110 ventrally (Fig. 1A, G); subcoxae 2 of legs I, II and III with 0, 0, 1 S-chaeta respectively. Tiny and blunt ms, present on Th.
<normalizedToken originalValue="IIIII">II-III</normalizedToken>
(Fig. 1A). Ordinary chaetae differentiated into meso- and macrochaetae, ratio Sp: m1: p1 on Abd. V tergum = 1: 1: 2.5. Th. I tergum with 7+7 dorsal chaetae. Th.
<normalizedToken originalValue="IIAbd">II-Abd</normalizedToken>
. III terga with 3+3 chaetae along axis respectively (Fig. 1A). Abd. IV tergum with one axial chaeta (p0), Abd. V tergum with two axial chaetae (a0 and m0), Abd. VI tergum with one axial chaeta (a0) (Figs 1A, 2D). Sterna of Th. I, II, and III with 0+0, 2+2, 2+2 chaetae respectively.
</paragraph>
<paragraph pageId="4" pageNumber="23">
<pageBreakToken pageId="4" pageNumber="23" start="start">Appendages</pageBreakToken>
. Subcoxa 1 of legs
<normalizedToken originalValue="IIII">I-III</normalizedToken>
with 4, 5 and 5 chaetae, subcoxa 2 with 1, 4 and 4 chaetae respectively. Tibiotarsi of legs I, II and III with 16 (1, 8, 7), 16 (1, 8, 7) and 15 (1, 7, 7) chaetae each (Fig. 1F). Unguis without teeth. Unguiculus approx. 0.6 times as long as inner edge of unguis, with inner basal lamella (Fig. 1F). Ventral tube with 4+4 basal and 7+7 distal chaetae. Furca reduced to a field of fine granulation with 4 small dental chaetae arranged in 2 rows posteriorly; only one manubrial row of chaetae posterior to dental chaetae (Figs 1G, 2B, C).
</paragraph>
<paragraph pageId="4" pageNumber="23">Genital fig with 9-11 chaetae in females, 32 chaetae in male. Male ventral organ absent. Anal valves with numerous acuminate chaetae; each lateral valve with a0, 2a1, 2a2; upper valves with chaetae a0, 2b1, 2b2, c0, 2c1, 2c2. Anal spines set on distinct papillae, 0.8 times as long as inner edge of hind unguis.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="23" type="derivatio nominis">
<paragraph pageId="4" pageNumber="23">Derivatio nominis.</paragraph>
<paragraph pageId="4" pageNumber="23">
Named for the first recordof the genus
<taxonomicName class="Collembola" family="Onychiuridae" genus="Spinonychiurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinonychiurus" order="Poduromorpha" pageId="4" pageNumber="23" phylum="Arthropoda" rank="genus">Spinonychiurus</taxonomicName>
in China.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="23" type="ecology">
<paragraph pageId="4" pageNumber="23">Ecology.</paragraph>
<paragraph pageId="4" pageNumber="23">Found in the coniferous forest.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="23" type="discussion">
<paragraph pageId="4" pageNumber="23">Discussion.</paragraph>
<paragraph pageId="4" pageNumber="23">
The new species is closest to the genus
<taxonomicName class="Collembola" family="Onychiuridae" genus="Spinonychiurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinonychiurus" order="Poduromorpha" pageId="4" pageNumber="23" phylum="Arthropoda" rank="genus">Spinonychiurus</taxonomicName>
in two important characters: two subsegments on Abd. III sternum and the absence of d0 on the head. However, it does not fit the current definition of the genus as proposed by
<bibRefCitation author="Kaprus', IJ" journalOrPublisher="Vestnik Zoologii" pageId="5" pageNumber="24" pagination="173 - 178" title="New Collembola species from the floodplain forests of the Transcarpathian Lowland (Ukraine)." url="10.2478/v10058-009-0008-8" volume="43" year="2009">
<normalizedToken originalValue="Kaprus">Kaprus'</normalizedToken>
and Tsalan (2009)
</bibRefCitation>
in having 7 chaetae in the distal row of tibiotarsi instead of 11. The distal tibiotarsal chaetae have been verified to be an unstable character in the generic level (
<bibRefCitation author="Sun, X" journalOrPublisher="Zootaxa" pageId="6" pageNumber="25" pagination="20 - 38" title="Six new species of Thalassaphorura (Collembola, Onychiuridae) from southern China, with a key to world species of the genus." volume="2627" year="2010">Sun et al. 2010</bibRefCitation>
;
<bibRefCitation author="Sun, X" journalOrPublisher="Zookeys" pageId="6" pageNumber="25" pagination="27 - 41" title="Redefinition of the genus Allonychiurus Yoshii, 1995 (Collembola, Onychiuridae) with description of a new species from China." url="10.3897/zookeys.78.977" volume="78" year="2011">Sun et al. 2011</bibRefCitation>
;
<bibRefCitation author="Sun, X" journalOrPublisher="Journal of Natural History" pageId="6" pageNumber="25" pagination="1895 - 1904" title="Two new species of Onychiurus (Collembola: Onychiuridae) from Eastern China." url="10.1080/00222933.2012.707236" volume="46" year="2012">Sun and Zhang 2012</bibRefCitation>
;
<bibRefCitation author="Sun, X" journalOrPublisher="ZooKeys" pageId="6" pageNumber="25" pagination="1 - 9" title="Broadening the definition of the genus Thalassaphorura Bagnall, 1949 (Collembola, Onychiuridae) with a new aberrant species from China." url="10.3897/zookeys.364.6332" volume="364" year="2013">Sun et al. 2013</bibRefCitation>
), so we propose the placement of the new species in the genus
<taxonomicName class="Collembola" family="Onychiuridae" genus="Spinonychiurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinonychiurus" order="Poduromorpha" pageId="4" pageNumber="23" phylum="Arthropoda" rank="genus">Spinonychiurus</taxonomicName>
and broaden its diagnosis accordingly. The main diagnostic characters of all known species of the genus are given in Table 1 and a key to these species is provided below.
</paragraph>
<caption pageId="4" pageNumber="23">
<paragraph pageId="4" pageNumber="23">
Table 1. Main diagnostic characters of world species of
<taxonomicName class="Collembola" family="Onychiuridae" genus="Spinonychiurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinonychiurus" order="Poduromorpha" pageId="4" pageNumber="23" phylum="Arthropoda" rank="genus">Spinonychiurus</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="4" pageNumber="23">
<table pageId="4" pageNumber="23">
<tr pageId="4" pageNumber="23">
<th colspan="1" pageId="4" pageNumber="23" rowspan="1" style="text-align:center">
<taxonomicName class="Collembola" family="Onychiuridae" genus="Spinonychiurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinonychiurus sinensis" order="Poduromorpha" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="sinensis">Spinonychiurus sinensis</taxonomicName>
sp. n.
</th>
<th colspan="1" pageId="4" pageNumber="23" rowspan="1" style="text-align:center">
<taxonomicName class="Collembola" family="Onychiuridae" genus="Spinonychiurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinonychiurus edinensis" order="Poduromorpha" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="edinensis">Spinonychiurus edinensis</taxonomicName>
*
</th>
<th colspan="1" pageId="4" pageNumber="23" rowspan="1" style="text-align:center">
<taxonomicName class="Collembola" family="Onychiuridae" genus="Spinonychiurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinonychiurus epaphius" order="Poduromorpha" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="epaphius">Spinonychiurus epaphius</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="23" rowspan="1" style="text-align:center">
<taxonomicName class="Collembola" family="Onychiuridae" genus="Spinonychiurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinonychiurus subedinensis" order="Poduromorpha" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="subedinensis">Spinonychiurus subedinensis</taxonomicName>
**
</th>
<th colspan="1" pageId="4" pageNumber="23" rowspan="1" style="text-align:center">
<taxonomicName class="Collembola" family="Onychiuridae" genus="Spinonychiurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinonychiurus vandeli" order="Poduromorpha" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="vandeli">Spinonychiurus vandeli</taxonomicName>
</th>
</tr>
<tr pageId="4" pageNumber="23">
<td colspan="1" pageId="4" pageNumber="23" rowspan="1">***</td>
<td colspan="1" pageId="4" pageNumber="23" rowspan="1">***</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="89D9717D04A6C17BD172EBBFA7D147EB">Micralestes (Characiformes, Alestidae) of the lower Congo River, with a description of a new species endemic to the lower Congo River rapids in the Democratic Republic of Congo.</mods:title>
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<mods:namePart id="821202247623299A727B6D676BF08459">Melanie L. J. Stiassny</mods:namePart>
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<paragraph id="268F068FE7282E28FE4AC381C2FC7204" pageNumber="19">AMNH 240421, 3, 1 C&amp;S;</paragraph>
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<mods:title id="33413B0AD10D6F4DFA4783431290A302">A new pearlfish, Onuxodon albometeori sp. nov. (Ophidiiformes: Carapidae), from Taiwan</mods:title>
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<taxonomicName id="E4A54D18D425FFC5FF06F6E6FE3E3FD8" authority="Koeda, 2019" authorityName="Koeda" authorityYear="2019" box="[151,418,225,252]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="1" phylum="Chordata" rank="species" species="albometeori" status="sp. nov.">
<emphasis id="11D1EA89D425FFC5FF06F6E6FE3E3FD8" bold="true" box="[151,418,225,252]" italics="true" pageId="1">Onuxodon albometeori</emphasis>
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<paragraph id="231A369BD425FFC5FF06F702FE803E3B" blockId="1.[151,515,225,287]" box="[151,284,261,287]" pageId="1">
<figureCitation id="BB9E2A1ED425FFC5FF06F702FE803E3B" box="[151,284,261,287]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1983,2008]" captionStartId-1="2.[151,250,674,699]" captionStartId-2="2.[151,250,1135,1160]" captionTargetBox-0="[151,1436,1677,1957]" captionTargetBox-2="[340,1247,756,1109]" captionTargetId-0="figure@1.[151,1436,1677,1957]" captionTargetId-1="figure@2.[340,1247,181,648]" captionTargetId-2="figure@2.[340,1247,756,1109]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionText-0="FIGURE 1. Holotype of Onuxodon albometeori sp. nov. from southwestern Taiwan, fresh condition, KAUMI. 115161, 107.6 mm TL, 105.5 mm SL, 12.6 mm HL." captionText-1="FIGURE 2. Head of holotype of Onuxodon albometeori sp. nov, fresh condition." captionText-2="FIGURE 3. Radiograph of holotype of Onuxodon albometeori sp. nov. Red arrow: sagittal otolith; blue arrow: rocker bone; yellow arrow: predorsal bone (outline surrounded by yellow dotted line)." figureDoi-0="http://doi.org/10.5281/zenodo.3562815" figureDoi-1="http://doi.org/10.5281/zenodo.3615366" figureDoi-2="http://doi.org/10.5281/zenodo.3562817" httpUri-0="https://zenodo.org/record/3562815/files/figure.png" httpUri-1="https://zenodo.org/record/3615366/files/figure.png" httpUri-2="https://zenodo.org/record/3562817/files/figure.png" pageId="1">Figures 13</figureCitation>
</paragraph>
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<paragraph id="231A369BD425FFC5FF06F749FF443E8B" blockId="1.[151,1437,333,1656]" pageId="1">
<materialsCitation id="93CD3CC6D425FFC5FF06F749FF443E8B" ID-GBIF-Occurrence="2462935302" collectingDate="2018-05-10" collectionCode="KAUM-I" country="Taiwan" latitude="22.716667" location="Kaohsiung" longLatPrecision="1258" longitude="120.23333" pageId="1" specimenCount="1" stateProvince="Kaohsiung" typeStatus="holotype">
<emphasis id="11D1EA89D425FFC5FF06F749FE973E4C" bold="true" box="[151,267,334,360]" pageId="1">
<typeStatus id="FC1E8839D425FFC5FF06F749FE9A3E4C" box="[151,262,334,360]" pageId="1" type="holotype">Holotype</typeStatus>
.
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<collectionCode id="45B4AE5ED425FFC5FE83F749FE1F3E4C" box="[274,387,334,360]" pageId="1">KAUMI</collectionCode>
.
<quantity id="E45D9B7ED425FFC5FE1FF74AFDFD3E4C" box="[398,609,333,360]" metricMagnitude="5" metricUnit="m" metricValue="1.151611076" pageId="1" unit="mm" value="1.151611076E8">115161, 107.6 mm</quantity>
TL,
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SL,
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HL, off
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(ca.
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,
<geoCoordinate id="4691505CD425FFC5FAB2F74AFA0C3E4C" box="[1315,1424,333,360]" degrees="120" direction="east" minutes="14" orientation="longitude" pageId="1" precision="925" value="120.23333">120°14ʹE</geoCoordinate>
), obtained at Ke-tzu-liao fish landing port, southwestern
<collectingCountry id="5BB2760BD425FFC5FC89F776FCF13EAF" box="[792,877,369,395]" name="Taiwan" pageId="1">Taiwan</collectingCountry>
,
<date id="571B105BD425FFC5FCEDF776FB873EAF" box="[892,1051,369,396]" pageId="1" value="2018-05-10">
<collectingDate id="475FE9B3D425FFC5FCEDF776FB873EAF" box="[892,1051,369,396]" pageId="1" value="2018-05-10">10 May 2018</collectingDate>
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, collected by commercial bottom trawl.
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</paragraph>
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<subSubSection id="6BBF6510D425FFC5FF56F7BEFCE43D3F" pageId="1" type="diagnosis">
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<emphasis id="11D1EA89D425FFC5FF56F7BEFEDE3EF0" bold="true" box="[199,322,441,468]" pageId="1">Diagnosis.</emphasis>
A species of
<taxonomicName id="E4A54D18D425FFC5FE49F7BEFDD13EF7" authorityName="Smith" authorityYear="1955" box="[472,589,441,467]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="1" phylum="Chordata" rank="genus">
<emphasis id="11D1EA89D425FFC5FE49F7BEFDD13EF7" box="[472,589,441,467]" italics="true" pageId="1">Onuxodon</emphasis>
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with the following combination of characters: precaudal vertebrae 23; body very elongated, its depth 56.9% HL and 6.7% TL at anal-fin origin; head width 40.0% HL; snout length 15.1% HL; body uniformly translucent whitish, without black pigmented tail.
</paragraph>
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<subSubSection id="6BBF6510D425FFC5FF56F422FEE1390B" pageId="1" type="description">
<paragraph id="231A369BD425FFC5FF56F422FC0E3D8F" blockId="1.[151,1437,333,1656]" pageId="1">
<emphasis id="11D1EA89D425FFC5FF56F422FEC53D64" bold="true" box="[199,345,549,576]" pageId="1">Description.</emphasis>
Dorsal-fin rays 136, 49 anterior to 30th vertebra and 0 anterior to vertical through anal-fin origin; anal-fin rays 135, 50 anterior to 30th vertebra and 2 anterior to vertical through dorsal-fin origin; pectoral-fin rays 13 (14 on right side); total vertebrae 75 (=23 precaudal + 52 caudal), 7 anterior to dorsal-fin origin, 6 anterior to anal-fin origin, 6 anterior to predorsal bone; developed gill rakers 3.
</paragraph>
<paragraph id="231A369BD425FFC5FF56F4B2FD853C44" blockId="1.[151,1437,333,1656]" pageId="1">Morphometrics: head length 11.7% TL; body depth 6.7% TL; pre-dorsal-fin length 16.2% TL; pre-anal-fin length 13.9% TL; pre-anus length 13.2% TL. Body depth 56.9% HL; pre-dorsal-fin length 127.8% HL; pre-anal-fin length 118.0% HL; pre-anus length 112.1% HL; snout length 15.1% HL; eye diameter 17.4% HL; bony interorbital width 16.3 HL; upper-jaw length 49.6% HL; lower-jaw length 47.7% HL; pectoral-fin length 47.9% HL; head depth 53.9% HL; head width 40.0% HL.</paragraph>
<paragraph id="231A369BD425FFC5FF56F56EFDD53CEF" blockId="1.[151,1437,333,1656]" pageId="1">Body very elongate, its depth greatest at gill opening, gradually becoming more shallow posteriorly. Body and head slightly compressed laterally, progressively more to tip of tail; caudal part much longer than body, snout to anus distance about 1/8 of total length.</paragraph>
<paragraph id="231A369BD425FFC5FF56F5D2FECA3B5B" blockId="1.[151,1437,333,1656]" pageId="1">Snout short, rounded. Eye large, circular, its diameter greater than snout length. Mouth slightly oblique, posterior end of upper lip well beyond posterior margin of eye. Villiform tooth band with small, numerous, conical teeth in 610 rows and 46 rows on upper and lower jaws, respectively, but not anteriorly on either jaw; a pair of single large, strong, curved fang-like teeth anteriorly on each jaw; tooth on vomer and palatines forming in 3 elongated, rounded patches.</paragraph>
<paragraph id="231A369BD425FFC5FF56F28EFBFC3BE3" blockId="1.[151,1437,333,1656]" pageId="1">Pectoral fins small, rounded, just behind gill opening; origin of dorsal fin slightly posterior to origin of anal fin; pelvic fin absent; dorsal and anal fins low, with very long bases; caudal fin very small.</paragraph>
<paragraph id="231A369BD425FFC5FF56F2D6FBCC3A73" blockId="1.[151,1437,333,1656]" pageId="1">
Sagittal otolith triangular, as large as eye (
<figureCitation id="BB9E2A1ED425FFC5FD36F2D6FD703BCF" box="[679,748,1233,1259]" captionStart="FIGURE 3" captionStartId="2.[151,250,1135,1160]" captionTargetBox="[340,1247,756,1109]" captionTargetId="figure@2.[340,1247,756,1109]" captionTargetPageId="2" captionText="FIGURE 3. Radiograph of holotype of Onuxodon albometeori sp. nov. Red arrow: sagittal otolith; blue arrow: rocker bone; yellow arrow: predorsal bone (outline surrounded by yellow dotted line)." figureDoi="http://doi.org/10.5281/zenodo.3562817" httpUri="https://zenodo.org/record/3562817/files/figure.png" pageId="1">Fig. 3</figureCitation>
); rocker bone positioned below 3rd vertebra; predorsal bone above 6th and 7th vertebrae; usually two proximal pterygiophores of dorsal and anal fins inserted between neural and haemal spines, respectively; anal-fin radials not elongate; numerous small distal radials supporting pectoral-fin rays; surface of posterior swim bladder smooth, lacking ventral patch of tunic ridges.
</paragraph>
<paragraph id="231A369BD425FFC5FF56F366FE8A3AC3" blockId="1.[151,1437,333,1656]" pageId="1">
<emphasis id="11D1EA89D425FFC5FF56F366FE1F3A5F" box="[199,387,1377,1403]" italics="true" pageId="1">Fresh coloration</emphasis>
(
<figureCitation id="BB9E2A1ED425FFC5FE00F366FE413A5F" box="[401,477,1377,1403]" captionStart="FIGURE 1" captionStartId="1.[151,250,1983,2008]" captionTargetBox="[151,1436,1677,1957]" captionTargetId="figure@1.[151,1436,1677,1957]" captionTargetPageId="1" captionText="FIGURE 1. Holotype of Onuxodon albometeori sp. nov. from southwestern Taiwan, fresh condition, KAUMI. 115161, 107.6 mm TL, 105.5 mm SL, 12.6 mm HL." figureDoi="http://doi.org/10.5281/zenodo.3562815" httpUri="https://zenodo.org/record/3562815/files/figure.png" pageId="1">Figs. 1</figureCitation>
,
<figureCitation id="BB9E2A1ED425FFC5FE7AF366FE603A5F" box="[491,508,1377,1403]" captionStart="FIGURE 2" captionStartId="2.[151,250,674,699]" captionTargetId="figure@2.[340,1247,181,648]" captionTargetPageId="2" captionText="FIGURE 2. Head of holotype of Onuxodon albometeori sp. nov, fresh condition." figureDoi="http://doi.org/10.5281/zenodo.3615366" httpUri="https://zenodo.org/record/3615366/files/figure.png" pageId="1">2</figureCitation>
). Body and head translucent whitish, without distinct pigmentation although faintly pinkish posteriorly; several areas of pink pigmentation on snout and anterior part of lower jaw; blackish pigments on brain and vertebrae visible through translucent skin; outer margins of dorsal and anal fins very faintly dusky posteriorly.
</paragraph>
<paragraph id="231A369BD425FFC5FF56F3F6FEE1390B" blockId="1.[151,1437,333,1656]" pageId="1">
<emphasis id="11D1EA89D425FFC5FF56F3F6FE2A392F" box="[199,438,1521,1547]" italics="true" pageId="1">Preserved coloration</emphasis>
. Head and body uniformly white; pigmentation on brain and posterior vertebral region visible through skin.
</paragraph>
</subSubSection>
<subSubSection id="6BBF6510D425FFC5FF56F03DFC4E3953" pageId="1" type="etymology">
<paragraph id="231A369BD425FFC5FF56F03DFC4E3953" blockId="1.[151,1437,333,1656]" pageId="1">
<emphasis id="11D1EA89D425FFC5FF56F03DFED23970" bold="true" box="[199,334,1594,1620]" pageId="1">Etymology.</emphasis>
The specific name,
<taxonomicName id="E4A54D18D425FFC5FDBDF03EFD2F3977" authorityName="Koeda" authorityYear="2019" box="[556,691,1593,1619]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="1" phylum="Chordata" rank="species" species="albometeori">
<emphasis id="11D1EA89D425FFC5FDBDF03EFD2F3977" box="[556,691,1593,1619]" italics="true" pageId="1">albometeori</emphasis>
</taxonomicName>
, combines the sense of “white” and “meteor” in Latin, in reference to the whitish coloration and very elongated, posteriorly shallowing body.
</paragraph>
</subSubSection>
<caption id="77DA6613D425FFC5FF06F1B8FD8D38DF" ID-DOI="http://doi.org/10.5281/zenodo.3562815" ID-Zenodo-Dep="3562815" httpUri="https://zenodo.org/record/3562815/files/figure.png" pageId="1" startId="1.[151,250,1983,2008]" targetBox="[151,1436,1677,1957]" targetPageId="1">
<paragraph id="231A369BD425FFC5FF06F1B8FD8D38DF" blockId="1.[151,1436,1983,2044]" pageId="1">
<emphasis id="11D1EA89D425FFC5FF06F1B8FE9138FC" bold="true" box="[151,269,1983,2008]" pageId="1">FIGURE 1</emphasis>
. Holotype of
<taxonomicName id="E4A54D18D425FFC5FE09F1B8FD1E38F3" authority="Koeda, 2019" authorityName="Koeda" authorityYear="2019" box="[408,642,1983,2007]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="1" phylum="Chordata" rank="species" species="albometeori" status="sp. nov.">
<emphasis id="11D1EA89D425FFC5FE09F1B8FD1E38F3" box="[408,642,1983,2007]" italics="true" pageId="1">Onuxodon albometeori</emphasis>
</taxonomicName>
<emphasis id="11D1EA89D425FFC5FD19F1B8FD4538F3" bold="true" box="[648,729,1983,2007]" pageId="1">
<taxonomicNameLabel id="0AE257F2D425FFC5FD19F1B8FD4538F3" box="[648,729,1983,2007]" pageId="1" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from southwestern Taiwan, fresh condition, KAUMI. 115161, 107.6 mm TL, 105.5 mm SL, 12.6 mm HL.
</paragraph>
</caption>
<caption id="77DA6613D426FFC6FF06F4A5FC4C3D9F" ID-DOI="http://doi.org/10.5281/zenodo.3615366" ID-Zenodo-Dep="3615366" box="[151,976,674,699]" httpUri="https://zenodo.org/record/3615366/files/figure.png" pageId="2" targetBox="[340,1247,181,648]" targetPageId="2">
<paragraph id="231A369BD426FFC6FF06F4A5FC4C3D9F" blockId="2.[151,976,674,699]" box="[151,976,674,699]" pageId="2">
<emphasis id="11D1EA89D426FFC6FF06F4A5FE8F3D9F" bold="true" box="[151,275,674,699]" pageId="2">FIGURE 2.</emphasis>
Head of holotype of
<taxonomicName id="E4A54D18D426FFC6FE7AF4A5FD493D9E" authority="Koeda, 2019" authorityName="Koeda" authorityYear="2019" box="[491,725,674,698]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="albometeori" status="sp. nov.">
<emphasis id="11D1EA89D426FFC6FE7AF4A5FD493D9E" box="[491,725,674,698]" italics="true" pageId="2">Onuxodon albometeori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AE257F2D426FFC6FD4DF4A4FCBA3D9E" box="[732,806,675,698]" pageId="2" rank="species">sp. nov</taxonomicNameLabel>
, fresh condition.
</paragraph>
</caption>
<caption id="77DA6613D426FFC6FF06F268FCE73B88" ID-DOI="http://doi.org/10.5281/zenodo.3562817" ID-Zenodo-Dep="3562817" httpUri="https://zenodo.org/record/3562817/files/figure.png" pageId="2" startId="2.[151,250,1135,1160]" targetBox="[340,1247,756,1109]" targetPageId="2">
<paragraph id="231A369BD426FFC6FF06F268FCE73B88" blockId="2.[151,1436,1135,1196]" pageId="2">
<emphasis id="11D1EA89D426FFC6FF06F268FE883BAC" bold="true" box="[151,276,1135,1160]" pageId="2">FIGURE 3.</emphasis>
Radiograph of holotype of
<taxonomicName id="E4A54D18D426FFC6FDA5F277FCBC3BAC" authority="Koeda, 2019" authorityName="Koeda" authorityYear="2019" box="[564,800,1136,1160]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="albometeori" status="sp. nov.">
<emphasis id="11D1EA89D426FFC6FDA5F277FCBC3BAC" box="[564,800,1136,1160]" italics="true" pageId="2">Onuxodon albometeori</emphasis>
</taxonomicName>
<emphasis id="11D1EA89D426FFC6FCB9F277FCE63BAC" bold="true" box="[808,890,1136,1160]" pageId="2">
<taxonomicNameLabel id="0AE257F2D426FFC6FCB9F277FCE63BAC" box="[808,890,1136,1160]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Red arrow: sagittal otolith; blue arrow: rocker bone; yellow arrow: predorsal bone (outline surrounded by yellow dotted line).
</paragraph>
</caption>
<subSubSection id="6BBF6510D426FFC6FF56F2FAFF6D3A18" pageId="2" type="distribution">
<paragraph id="231A369BD426FFC6FF56F2FAFF6D3A18" blockId="2.[151,1437,1277,2025]" pageId="2">
<emphasis id="11D1EA89D426FFC6FF56F2FAFEFD3A3C" bold="true" box="[199,353,1277,1304]" pageId="2">Distribution.</emphasis>
<taxonomicName id="E4A54D18D426FFC6FEF8F2F9FDF23A3C" authority="Koeda, 2019" authorityName="Koeda" authorityYear="2019" box="[361,622,1278,1304]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="albometeori" status="sp. nov.">
<emphasis id="11D1EA89D426FFC6FEF8F2F9FDF23A3C" box="[361,622,1278,1304]" italics="true" pageId="2">Onuxodon albometeori</emphasis>
</taxonomicName>
<emphasis id="11D1EA89D426FFC6FDE6F2F9FD4F3A3C" bold="true" box="[631,723,1278,1304]" pageId="2">
<taxonomicNameLabel id="0AE257F2D426FFC6FDE6F2F9FD4F3A3C" box="[631,723,1278,1304]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="FC1E8839D426FFC6FC63F2F9FBC83A3C" box="[1010,1108,1278,1304]" pageId="2" type="holotype">holotype</typeStatus>
collcted in the southwestern
<collectingCountry id="5BB2760BD426FFC6FF06F325FF713A18" box="[151,237,1314,1340]" name="Taiwan" pageId="2">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6BBF6510D426FFC7FF56F341FCBD3DA3" lastPageId="3" pageId="2" type="discussion">
<paragraph id="231A369BD426FFC6FF56F341FC453858" blockId="2.[151,1437,1277,2025]" pageId="2">
<emphasis id="11D1EA89D426FFC6FF56F341FEA73A44" bold="true" box="[199,315,1350,1376]" pageId="2">Remarks.</emphasis>
<taxonomicName id="E4A54D18D426FFC6FED2F341FDDA3A44" authority="Koeda, 2019" authorityName="Koeda" authorityYear="2019" box="[323,582,1350,1376]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="albometeori" status="sp. nov.">
<emphasis id="11D1EA89D426FFC6FED2F341FDDA3A44" box="[323,582,1350,1376]" italics="true" pageId="2">Onuxodon albometeori</emphasis>
</taxonomicName>
<emphasis id="11D1EA89D426FFC6FDDFF341FD343A44" bold="true" box="[590,680,1350,1376]" pageId="2">
<taxonomicNameLabel id="0AE257F2D426FFC6FDDFF341FD343A44" box="[590,680,1350,1376]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shares the following characters with other congeners which distinguish the genus
<taxonomicName id="E4A54D18D426FFC6FEDAF36DFE5C3AA0" authorityName="Smith" authorityYear="1955" box="[331,448,1386,1412]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="genus">
<emphasis id="11D1EA89D426FFC6FEDAF36DFE5C3AA0" box="[331,448,1386,1412]" italics="true" pageId="2">Onuxodon</emphasis>
</taxonomicName>
from other carapids: lack of pelvic fins and ventral patch of posterior tunic ridges on the swim bladder and presence of a rocker bone, predorsal bone, and single large, symphysial fangs on each jaw. The new species resembles
<taxonomicName id="E4A54D18D426FFC6FE0AF3B5FD903AE8" baseAuthorityName="Smith" baseAuthorityYear="1955" box="[411,524,1458,1484]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="fowleri">
<emphasis id="11D1EA89D426FFC6FE0AF3B5FD903AE8" box="[411,524,1458,1484]" italics="true" pageId="2">O. fowleri</emphasis>
</taxonomicName>
, which is widely distributed in the Indo-Pacific; both species have a slender body with higher vertebral counts than those two other congeners (
<bibRefCitation id="47344B6AD426FFC6FCD3F3D1FBDC3AD4" author="Markle, D. F. &amp; Olney, J. E." box="[834,1088,1494,1520]" pageId="2" pagination="269 - 410" refId="ref2376" refString="Markle, D. F. &amp; Olney, J. E. (1990) Systematics of the pearlfishes (Pisces: Carapidae). Bulletin of Marine Science, 47 (2), 269 - 410." type="journal article" year="1990">Markle &amp; Olney 1990</bibRefCitation>
). However, a comparison of
<taxonomicName id="E4A54D18D426FFC6FA10F3D1FE823930" authorityName="Koeda" authorityYear="2019" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="albometeori">
<emphasis id="11D1EA89D426FFC6FA10F3D1FE823930" italics="true" pageId="2">O. albometeori</emphasis>
</taxonomicName>
with the data of
<typeStatus id="FC1E8839D426FFC6FE48F3FDFDDC3930" box="[473,576,1530,1556]" pageId="2" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="E4A54D18D426FFC6FDF5F3FDFD493930" baseAuthorityName="Smith" baseAuthorityYear="1955" box="[612,725,1530,1556]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="fowleri">
<emphasis id="11D1EA89D426FFC6FDF5F3FDFD493930" box="[612,725,1530,1556]" italics="true" pageId="2">O. fowleri</emphasis>
</taxonomicName>
(BPBM 5001) given by
<bibRefCitation id="47344B6AD426FFC6FC78F3FDFB6E3930" author="Markle, D. F. &amp; Olney, J. E." box="[1001,1266,1530,1556]" pageId="2" pagination="269 - 410" refId="ref2376" refString="Markle, D. F. &amp; Olney, J. E. (1990) Systematics of the pearlfishes (Pisces: Carapidae). Bulletin of Marine Science, 47 (2), 269 - 410." type="journal article" year="1990">Markle &amp; Olney (1990)</bibRefCitation>
indicated clear differences in the following characters: precaudal vertebrae
<quantity id="E45D9B7ED426FFC6FCA7F019FCF3391C" box="[822,879,1566,1592]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="2" unit="in" value="23.0">23 in</quantity>
<taxonomicName id="E4A54D18D426FFC6FCE7F019FB83391C" authorityName="Koeda" authorityYear="2019" box="[886,1055,1566,1592]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="albometeori">
<emphasis id="11D1EA89D426FFC6FCE7F019FB83391C" box="[886,1055,1566,1592]" italics="true" pageId="2">O. albometeori</emphasis>
</taxonomicName>
(vs.
<quantity id="E45D9B7ED426FFC6FBC7F019FB13391C" box="[1110,1167,1566,1592]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="2" unit="in" value="20.0">20 in</quantity>
<taxonomicName id="E4A54D18D426FFC6FB07F019FA9B391C" baseAuthorityName="Smith" baseAuthorityYear="1955" box="[1174,1287,1566,1592]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="fowleri">
<emphasis id="11D1EA89D426FFC6FB07F019FA9B391C" box="[1174,1287,1566,1592]" italics="true" pageId="2">O. fowleri</emphasis>
</taxonomicName>
); body depth 56.9% HL (vs. 74.0% HL) and 6.7% TL (9.2% TL) at anal-fin origin; head width 40.0% HL (vs. 31.1% HL); snout length 86.8% of eye diameter (vs. 118.0% of eye diameter); upper-jaw length 49.6% HL (vs. 57.8% HL); pre-dorsalfin length 137.8% HL (vs. 127.8% HL); pectoral-fin length 47.9% HL (vs. 37.8% HL). Some morphometric characters overlap with those of non-types (given to nearest 10%) given by
<bibRefCitation id="47344B6AD426FFC6FC28F0A9FB5B39EC" author="Markle, D. F. &amp; Olney, J. E." box="[953,1223,1710,1736]" pageId="2" pagination="269 - 410" refId="ref2376" refString="Markle, D. F. &amp; Olney, J. E. (1990) Systematics of the pearlfishes (Pisces: Carapidae). Bulletin of Marine Science, 47 (2), 269 - 410." type="journal article" year="1990">Markle &amp; Olney (1990)</bibRefCitation>
, e.g. pre-dorsal-fin length 118.0% HL (cf. 110140% HL in
<taxonomicName id="E4A54D18D426FFC6FDC9F0D5FD5439C8" baseAuthorityName="Smith" baseAuthorityYear="1955" box="[600,712,1746,1772]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="fowleri">
<emphasis id="11D1EA89D426FFC6FDC9F0D5FD5439C8" box="[600,712,1746,1772]" italics="true" pageId="2">O. fowleri</emphasis>
</taxonomicName>
) and head width 40.0% HL (cf. 3040% HL). These may suggest that some (but not all) characters vary with growth or simply reflect the condition of the specimens.
<bibRefCitation id="47344B6AD426FFC6FB77F0F1FF473810" author="Markle, D. F. &amp; Olney, J. E." pageId="2" pagination="269 - 410" refId="ref2376" refString="Markle, D. F. &amp; Olney, J. E. (1990) Systematics of the pearlfishes (Pisces: Carapidae). Bulletin of Marine Science, 47 (2), 269 - 410." type="journal article" year="1990">Markle &amp; Olney (1990</bibRefCitation>
, fig. 91) showed the relationship between total length and body depth in
<taxonomicName id="E4A54D18D426FFC6FC6EF11DFBF33810" baseAuthorityName="Smith" baseAuthorityYear="1955" box="[1023,1135,1818,1844]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="fowleri">
<emphasis id="11D1EA89D426FFC6FC6EF11DFBF33810" box="[1023,1135,1818,1844]" italics="true" pageId="2">O. fowleri</emphasis>
</taxonomicName>
, but that of
<taxonomicName id="E4A54D18D426FFC6FB65F11DFA003810" authorityName="Koeda" authorityYear="2019" box="[1268,1436,1818,1844]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="albometeori">
<emphasis id="11D1EA89D426FFC6FB65F11DFA003810" box="[1268,1436,1818,1844]" italics="true" pageId="2">O. albometeori</emphasis>
</taxonomicName>
is obviously outside the range. In addition, the coloration of the two species clearly differed,
<taxonomicName id="E4A54D18D426FFC6FB0AF139FADF387C" authorityName="Koeda" authorityYear="2019" box="[1179,1347,1854,1880]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="albometeori">
<emphasis id="11D1EA89D426FFC6FB0AF139FADF387C" box="[1179,1347,1854,1880]" italics="true" pageId="2">O. albometeori</emphasis>
</taxonomicName>
lacking black pigment on tail, whereas
<taxonomicName id="E4A54D18D426FFC6FE64F165FDFA3858" baseAuthorityName="Smith" baseAuthorityYear="1955" box="[501,614,1890,1916]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="fowleri">
<emphasis id="11D1EA89D426FFC6FE64F165FDFA3858" box="[501,614,1890,1916]" italics="true" pageId="2">O. fowleri</emphasis>
</taxonomicName>
has a completely pigmented tail.
</paragraph>
<paragraph id="231A369BD426FFC7FF56F181FE543E8B" blockId="2.[151,1437,1277,2025]" lastBlockId="3.[151,1437,153,647]" lastPageId="3" pageId="2">
<taxonomicName id="E4A54D18D426FFC6FF56F181FD9B3884" authority="Smith, 1955" authorityName="Smith" authorityYear="1955" box="[199,519,1925,1952]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="reedi">
<emphasis id="11D1EA89D426FFC6FF56F181FEE63884" box="[199,378,1926,1952]" italics="true" pageId="2">Onuxodon reedi</emphasis>
<bibRefCitation id="47344B6AD426FFC6FEEEF181FD9B3884" author="Smith, J. L. B." box="[383,519,1925,1952]" pageId="2" pagination="401 - 416" refId="ref2663" refString="Smith, J. L. B. (1955) The fishes of the family Carapidae in the western Indian Ocean. Annals and Magazine of Natural History (Series 12), 8, 401 - 416. https: // doi. org / 10.1080 / 00222935508656068" type="journal article" year="1955">Smith, 1955</bibRefCitation>
</taxonomicName>
was originally described on the basis of a subadult specimen (
<quantity id="E45D9B7ED426FFC6FB23F181FA9D3885" box="[1202,1281,1926,1953]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="2" unit="mm" value="70.0">70 mm</quantity>
TL) collected from
<collectingCountry id="5BB2760BD426FFC6FF42F1ADFEFF38E0" box="[211,355,1962,1988]" name="South Africa" pageId="2">South Africa</collectingCountry>
, but
<bibRefCitation id="47344B6AD426FFC6FE07F1ADFD0138E0" author="Markle, D. F. &amp; Olney, J. E." box="[406,669,1962,1988]" pageId="2" pagination="269 - 410" refId="ref2376" refString="Markle, D. F. &amp; Olney, J. E. (1990) Systematics of the pearlfishes (Pisces: Carapidae). Bulletin of Marine Science, 47 (2), 269 - 410." type="journal article" year="1990">Markle &amp; Olney (1990)</bibRefCitation>
doubted the validity of the species.
<bibRefCitation id="47344B6AD426FFC6FBB9F1ADFB5638E0" author="Markle, D. F." box="[1064,1226,1962,1988]" pageId="2" pagination="10 - 21" refId="ref2416" refString="Markle, D. F. (1999) Carapidae. In: Nielsen, J. G., Cohen, D. F., Makle, D. F. &amp; Robins, C. R. (Eds.), FAO species catalogue. Volume 18. Ophidiiform fishes of the world (Order Ophidiiformes). An annotated and illustrated catalogue of pearlfishes, cusk-eels, brotulas and other ophidiirorm fishes known to date. FAO (Food and Agriculture Organization of the United Nations) Fisheries Synopsis No. 125. FAO, Rome, pp. 10 - 21." type="book chapter" year="1999">Markle (1999)</bibRefCitation>
subsequently indicated that
<taxonomicName id="E4A54D18D426FFC6FE9BF1C9FEF938CC" authorityName="Smith" authorityYear="1955" box="[266,357,1998,2024]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="reedi">
<emphasis id="11D1EA89D426FFC6FE9BF1C9FEF938CC" box="[266,357,1998,2024]" italics="true" pageId="2">O. reedi</emphasis>
</taxonomicName>
was a junior synonym of either
<taxonomicName id="E4A54D18D426FFC6FD5FF1C9FC0538CC" baseAuthorityName="Smith" baseAuthorityYear="1955" box="[718,921,1998,2024]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="fowleri">
<emphasis id="11D1EA89D426FFC6FD5FF1C9FC0538CC" box="[718,921,1998,2024]" italics="true" pageId="2">Onuxodon fowleri</emphasis>
</taxonomicName>
or
<taxonomicName id="E4A54D18D426FFC6FC2FF1C9FB1A38CC" baseAuthorityName="Fowler" baseAuthorityYear="1927" box="[958,1158,1998,2024]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="parvibrachium">
<emphasis id="11D1EA89D426FFC6FC2FF1C9FB1A38CC" box="[958,1158,1998,2024]" italics="true" pageId="2">O. parvibrachium</emphasis>
</taxonomicName>
. The validity of
<taxonomicName id="E4A54D18D426FFC6FAD3F1C9FA0138CC" authorityName="Smith" authorityYear="1955" box="[1346,1437,1998,2024]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="2" phylum="Chordata" rank="species" species="reedi">
<emphasis id="11D1EA89D426FFC6FAD3F1C9FA0138CC" box="[1346,1437,1998,2024]" italics="true" pageId="2">O. reedi</emphasis>
</taxonomicName>
is still questionable, the species being regarded as a junior synonym of species of
<taxonomicName id="E4A54D18D427FFC7FBAEF69EFB283F97" authorityName="Smith" authorityYear="1955" box="[1087,1204,153,179]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="3" phylum="Chordata" rank="genus">
<emphasis id="11D1EA89D427FFC7FBAEF69EFB283F97" box="[1087,1204,153,179]" italics="true" pageId="3">Onuxodon</emphasis>
</taxonomicName>
. Some characters of
<taxonomicName id="E4A54D18D427FFC7FF06F6BAFF6E3FF3" authorityName="Smith" authorityYear="1955" box="[151,242,189,215]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="3" phylum="Chordata" rank="species" species="reedi">
<emphasis id="11D1EA89D427FFC7FF06F6BAFF6E3FF3" box="[151,242,189,215]" italics="true" pageId="3">O. reedi</emphasis>
</taxonomicName>
included in the original description were similar to those of
<taxonomicName id="E4A54D18D427FFC7FC09F6BAFBDD3FF3" authorityName="Koeda" authorityYear="2019" box="[920,1089,189,215]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="3" phylum="Chordata" rank="species" species="albometeori">
<emphasis id="11D1EA89D427FFC7FC09F6BAFBDD3FF3" box="[920,1089,189,215]" italics="true" pageId="3">O. albometeori</emphasis>
</taxonomicName>
, including a very slender body (depth 5% TL at anal-fin origin, cf. 6.6% TL in
<taxonomicName id="E4A54D18D427FFC7FD3CF6E6FCCA3FDF" authorityName="Koeda" authorityYear="2019" box="[685,854,225,251]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="3" phylum="Chordata" rank="species" species="albometeori">
<emphasis id="11D1EA89D427FFC7FD3CF6E6FCCA3FDF" box="[685,854,225,251]" italics="true" pageId="3">O. albometeori</emphasis>
</taxonomicName>
). However,
<bibRefCitation id="47344B6AD427FFC7FC70F6E6FBEA3FDF" author="Smith, J. L. B." box="[993,1142,224,251]" pageId="3" pagination="401 - 416" refId="ref2663" refString="Smith, J. L. B. (1955) The fishes of the family Carapidae in the western Indian Ocean. Annals and Magazine of Natural History (Series 12), 8, 401 - 416. https: // doi. org / 10.1080 / 00222935508656068" type="journal article" year="1955">Smith (1955)</bibRefCitation>
noted that the
<typeStatus id="FC1E8839D427FFC7FA8DF6E6FAE23FDF" box="[1308,1406,225,251]" pageId="3" type="holotype">holotype</typeStatus>
of
<taxonomicName id="E4A54D18D427FFC7FF06F702FF6E3E3B" authorityName="Smith" authorityYear="1955" box="[151,242,261,287]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="3" phylum="Chordata" rank="species" species="reedi">
<emphasis id="11D1EA89D427FFC7FF06F702FF6E3E3B" box="[151,242,261,287]" italics="true" pageId="3">O. reedi</emphasis>
</taxonomicName>
had a strongly compressed head, its depth 2.5 times the width, clearly different from that of
<taxonomicName id="E4A54D18D427FFC7FB6AF702FA013E3B" box="[1275,1437,261,287]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="3" phylum="Chordata" rank="species" species="albometeor">
<emphasis id="11D1EA89D427FFC7FB6AF702FA013E3B" box="[1275,1437,261,287]" italics="true" pageId="3">O. albometeor</emphasis>
</taxonomicName>
(1.3 times). In addition, the counts of pectoral-fin ray of the
<typeStatus id="FC1E8839D427FFC7FCD3F72EFC383E67" box="[834,932,297,323]" pageId="3" type="holotype">holotype</typeStatus>
of
<taxonomicName id="E4A54D18D427FFC7FC5AF72EFBBB3E67" authorityName="Smith" authorityYear="1955" box="[971,1063,297,323]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="3" phylum="Chordata" rank="species" species="reedi">
<emphasis id="11D1EA89D427FFC7FC5AF72EFBBB3E67" box="[971,1063,297,323]" italics="true" pageId="3">O. reedi</emphasis>
</taxonomicName>
(17) is distinctly differ from that of
<taxonomicName id="E4A54D18D427FFC7FF24F74AFEC23E43" authorityName="Koeda" authorityYear="2019" box="[181,350,333,359]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="3" phylum="Chordata" rank="species" species="albometeori">
<emphasis id="11D1EA89D427FFC7FF24F74AFEC23E43" box="[181,350,333,359]" italics="true" pageId="3">O. albometeori</emphasis>
</taxonomicName>
(13 or 14). While it is difficult to compare subadult characters with those of
<taxonomicName id="E4A54D18D427FFC7FB28F74AFAC03E43" box="[1209,1372,333,359]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="3" phylum="Chordata" rank="species" species="albometeor">
<emphasis id="11D1EA89D427FFC7FB28F74AFAC03E43" box="[1209,1372,333,359]" italics="true" pageId="3">O. albometeor</emphasis>
</taxonomicName>
, their morphological differences combined with geographic dislocation strongly support recognition of the Taiwanese specimen as a new species.
</paragraph>
<paragraph id="231A369BD427FFC7FF56F7BEFCBD3DA3" blockId="3.[151,1437,153,647]" pageId="3">
Several reports of
<taxonomicName id="E4A54D18D427FFC7FE06F7BEFD953EF7" baseAuthorityName="Smith" baseAuthorityYear="1955" box="[407,521,441,467]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="3" phylum="Chordata" rank="species" species="fowleri">
<emphasis id="11D1EA89D427FFC7FE06F7BEFD953EF7" box="[407,521,441,467]" italics="true" pageId="3">O. fowleri</emphasis>
</taxonomicName>
from the Indo-Pacific region have included photographs or drawings suitable for species identification (e.g.
<bibRefCitation id="47344B6AD427FFC7FE5FF7DAFDE23ED3" author="Heemstra, P. C." box="[462,638,476,503]" pageId="3" pagination="350 - 354" refId="ref2313" refString="Heemstra, P. C. (1995) Family No. 97: Carapidae. In: Smith, M. M. &amp; Heemstra, P. C. (Eds.), S mith's Sea Fishes. Third impression. J. L. B. Smith Institute of Ichthyology, Grahamstown, pp. 350 - 354." type="book chapter" year="1995">Heemstra 1995</bibRefCitation>
;
<bibRefCitation id="47344B6AD427FFC7FD1CF7DAFCF33ED3" author="Randall, J. E. &amp; Allen, G. R. &amp; Steene, R." box="[653,879,477,503]" pageId="3" refId="ref2607" refString="Randall, J. E., Allen, G. R. &amp; Steene, R. C (1997) Fishes of the Great Barrier Reef and Coral Sea. 2 nd, revised edition. Crawford House Press, Bathurst NSW, xx + 557 pp., [7 plates.]" type="book" year="1997">
Randall
<emphasis id="11D1EA89D427FFC7FD7FF7D9FCBF3ED3" box="[750,803,477,503]" italics="true" pageId="3">et al</emphasis>
. 1997
</bibRefCitation>
;
<bibRefCitation id="47344B6AD427FFC7FCEFF7DAFB963ED3" author="Myers, R. F." box="[894,1034,477,503]" pageId="3" refId="ref2524" refString="Myers, R. F. (1999) Micronesian reef fishes. A comprehensive guide to the coral reef fishes of Micronesia. 3 rd revised edtion, Coral Graphics, Guam, vi + 330 pp. [192 plates.]" type="book" year="1999">Myers 1999</bibRefCitation>
;
<bibRefCitation id="47344B6AD427FFC7FB89F7DAFAAA3ED3" author="Amaoka, K. &amp; Yoseda, K." box="[1048,1334,476,503]" pageId="3" pagination="5 - 12" refId="ref2254" refString="Amaoka, K. &amp; Yoseda, K. (2005) Two species of pearl fishes, Onuxodon parvibrachium and O. fowleri (Ophidiiformes, Carapidae) from the honeycomb oyster (Hyotissa hyotis), from Ishigaki Island, Okinawa Prefecture. Bulletin of the Biogeographical Society of Japan, 60, 5 - 12." type="journal article" year="2005">Amaoka &amp; Yoseda 2005</bibRefCitation>
; Randall 2007;
<bibRefCitation id="47344B6AD427FFC7FF4CF406FE773D3F" author="Allen, G. R. &amp; Erdmann, M. V." box="[221,491,513,539]" pageId="3" refId="ref2217" refString="Allen, G. R. &amp; Erdmann, M. V. (2012) Reef fishes of the East Indies. Vol 1. Tropical Reef Research, Perth, xiii + 424 pp." type="book" year="2012">Allen &amp; Erdmann 2012</bibRefCitation>
), all of them show a deep body and distinctly black tail, and well agree with the diagnostic characters of
<taxonomicName id="E4A54D18D427FFC7FE04F422FD9A3D1B" baseAuthorityName="Smith" baseAuthorityYear="1955" box="[405,518,549,575]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="3" phylum="Chordata" rank="species" species="fowleri">
<emphasis id="11D1EA89D427FFC7FE04F422FD9A3D1B" box="[405,518,549,575]" italics="true" pageId="3">O. fowleri</emphasis>
</taxonomicName>
. Therefore, the proposed
<typeStatus id="FC1E8839D427FFC7FCBAF422FC113D1B" box="[811,909,549,575]" pageId="3" type="holotype">holotype</typeStatus>
of
<taxonomicName id="E4A54D18D427FFC7FC22F422FBC13D1B" authorityName="Koeda" authorityYear="2019" box="[947,1117,549,575]" class="Actinopterygii" family="Carapidae" genus="Onuxodon" kingdom="Animalia" order="Ophidiiformes" pageId="3" phylum="Chordata" rank="species" species="albometeori">
<emphasis id="11D1EA89D427FFC7FC22F422FBC13D1B" box="[947,1117,549,575]" italics="true" pageId="3">O. albometeori</emphasis>
</taxonomicName>
is considered to be the only known example of the new species. Because the specimen was retrieved from the floor of the fish landing port, the potential hosts of this symbiotic species remain unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,105 @@
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.90.20586" ID-PMC="PMC5784240" ID-Pensoft-Pub="1314-2003-90-1" ID-Pensoft-UUID="80067D29FFFB7D34FF80E95D553F4254" ID-PubMed="29391851" ID-Zenodo-Dep="1138341" ModsDocID="1314-2003-90-1" checkinTime="1510830429569" checkinUser="pensoft" docAuthor="Berry, Paul E., Kainulainen, Kent &amp; Ee, Benjamin W. van" docDate="2017" docId="AB0CBCDEA73E5B42AE25E46F9B75E640" docLanguage="en" docName="PhytoKeys 90: 1-87" docOrigin="PhytoKeys 90" docPubDate="2017-11-15" docSource="http://dx.doi.org/10.3897/phytokeys.90.20586" docTitle="Croton vatomandrensis Leandri, Ann. Mus. Colon. Marseille, sér. 5, 7 (1): 71. 1939" docType="treatment" docVersion="3" id="80067D29FFFB7D34FF80E95D553F4254" lastPageNumber="62" masterDocId="80067D29FFFB7D34FF80E95D553F4254" masterDocTitle="A Nomenclator of Croton (Euphorbiaceae) in Madagascar, the Comoros Archipelago, and the Mascarene Islands" masterLastPageNumber="87" masterPageNumber="1" pageNumber="62" updateTime="1668142129654" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A Nomenclator of Croton (Euphorbiaceae) in Madagascar, the Comoros Archipelago, and the Mascarene Islands</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Berry, Paul E.</mods:namePart>
<mods:affiliation>Herbarium, Department of Ecology and Evolutionary Biology, University of Michigan, 3600 Varsity Drive, Ann Arbor, Michigan 48108, U. S. A.</mods:affiliation>
<mods:nameIdentifier type="email">peberry@umich.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kainulainen, Kent</mods:namePart>
<mods:affiliation>Herbarium, Department of Ecology and Evolutionary Biology, University of Michigan, 3600 Varsity Drive, Ann Arbor, Michigan 48108, U. S. A.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ee, Benjamin W. van</mods:namePart>
<mods:affiliation>Department of Biology, Universidad de Puerto Rico, Recinto Universitario de Mayagueez, Mayagueez, PR 00680, Puerto Rico, U. S. A.</mods:affiliation>
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<mods:date>2017</mods:date>
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<mods:number>2017-11-15</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>90</mods:number>
</mods:detail>
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<mods:start>1</mods:start>
<mods:end>87</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.90.20586</mods:url>
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<treatment ID-GBIF-Taxon="182225587" LSID="urn:lsid:plazi:treatment:AB0CBCDEA73E5B42AE25E46F9B75E640" httpUri="http://treatment.plazi.org/id/AB0CBCDEA73E5B42AE25E46F9B75E640" lastPageNumber="62" pageId="61" pageNumber="62">
<subSubSection pageId="61" pageNumber="62" type="nomenclature">
<paragraph pageId="61" pageNumber="62">
124.
<taxonomicName LSID="AB0CBCDE-A73E-5B42-AE25-E46F9B75E640" authority="Leandri, Ann. Mus. Colon. Marseille, ser. 5, 7 (1): 71. 1939" authorityName="Leandri, Ann. Mus. Colon. Marseille, sér. 5, 7 (1): 71." authorityYear="1939" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton vatomandrensis" order="Malpighiales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="vatomandrensis">
Croton vatomandrensis Leandri, Ann. Mus. Colon. Marseille,
<normalizedToken originalValue="sér">ser</normalizedToken>
. 5, 7(1): 71. 1939
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="62" type="materials_examined">
<paragraph pageId="61" pageNumber="62">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="61" pageNumber="62">
<materialsCitation collectingDate="1921-11" collectorName="de Vatomandry, H. Perrier de la Bathie" country="Madagascar" location="Toamasina" specimenCount="1" stateProvince="Toamasina" typeStatus="lectotype">
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
. Prov.
<collectingRegion country="Madagascar" name="Toamasina">Toamasina</collectingRegion>
: environs
<collectorName>de Vatomandry</collectorName>
,
<normalizedToken originalValue="près">pres</normalizedToken>
de lagunes,
<collectingDate value="1921-11">Nov 1921</collectingDate>
,
<emphasis italics="true" pageId="61" pageNumber="62">
<collectorName>
H. Perrier de la
<normalizedToken originalValue="Bâthie">Bathie</normalizedToken>
</collectorName>
14084
</emphasis>
(
<typeStatus>lectotype</typeStatus>
, designated by
<bibRefCitation DOI="https://doi.org/10.15553/c2017v721a5" author="Kainulainen, K" journalOrPublisher="Candollea" pageId="72" pageNumber="73" pagination="35 - 44" refId="B30" refString="Kainulainen, K, Berry, PE, Van Ee, B, 2017a. Rediscovery of two species of Croton L. (Euphorbiaceae) from littoral habitats of eastern Madagascar. Candollea 72: 35 - 44, DOI: https://doi.org/10.15553/c2017v721a5" title="Rediscovery of two species of Croton L. (Euphorbiaceae) from littoral habitats of eastern Madagascar." url="https://doi.org/10.15553/c2017v721a5" volume="72" year="2017 a">Kainulainen et al. 2017a</bibRefCitation>
, pg. 39: P [P00248926]!; isolectotypes: P [P00154409]!, P [P00154410]!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="62" type="distribution">
<paragraph pageId="61" pageNumber="62">Habit and distribution.</paragraph>
<paragraph pageId="61" pageNumber="62">Shrubs or small trees; eastern littoral forests of Madagascar (Fianarantsoa, Toamasina).</paragraph>
</subSubSection>
</treatment>
</document>