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<document id="97E76365193FB4D372FD29CA56EDF5A5" ID-DOI="10.15298/rusentj.33.4.07" ID-ISSN="0132-8069" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737172247377" checkinUser="felipe" docAuthor="Petrov, A. V., Mandelshtam, M. Yu., Li, You, Potanina, S. O. &amp; Smith, S. M." docDate="2024" docId="038E3B521A28FFCEA7B3F9F9FC1F992B" docLanguage="en" docName="RussEntomolJ.33.4.463-465.pdf" docOrigin="Russian Entomological Journal 33 (4)" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.4:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="4" docTitle="Ernoporus eggersi Kurentsov 1941" docType="treatment" docVersion="1" lastPageNumber="465" masterDocId="FFB7432A1A2AFFCCA752FFE4FF909A07" masterDocTitle="Nomenclature changes in Palaearctic Scolytinae (Coleoptera: Curculionidae)" masterLastPageNumber="465" masterPageNumber="463" pageNumber="465" updateTime="1737593750220" updateUser="felipe" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7C514618F59EEB5066188988F211B2BC">Nomenclature changes in Palaearctic Scolytinae (Coleoptera: Curculionidae)</mods:title>
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<mods:namePart id="CDFA591A6E0C11AAA485D62CEEB4D240">Petrov, A. V.</mods:namePart>
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<mods:namePart id="ED2FD7CE3FA1F7E7BA0689D6D0A03EA5">Mandelshtam, M. Yu.</mods:namePart>
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<mods:namePart id="491FF4C2959C49281DB6AA3BF0EDAC01">Li, You</mods:namePart>
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<mods:namePart id="4F0CFCB133A0BB8B312A0F6920B03D4C">Potanina, S. O.</mods:namePart>
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<mods:namePart id="6BE52DADCE97F4608922439A6E06322F">Smith, S. M.</mods:namePart>
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<mods:title id="3ECF6B4C7E92116B77400FF7921189E7">Russian Entomological Journal</mods:title>
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<mods:date id="8BD256FAD20DB8CD50204046E6BB5F20">2024</mods:date>
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<treatment id="038E3B521A28FFCEA7B3F9F9FC1F992B" LSID="urn:lsid:plazi:treatment:038E3B521A28FFCEA7B3F9F9FC1F992B" httpUri="http://treatment.plazi.org/id/038E3B521A28FFCEA7B3F9F9FC1F992B" lastPageNumber="465" pageId="2" pageNumber="465">
<subSubSection id="C33DD9CF1A28FFCEA7B3F9F9FD7F9C33" box="[225,751,1565,1588]" pageId="2" pageNumber="465" type="nomenclature">
<paragraph id="8B988A441A28FFCEA7B3F9F9FD7F9C33" blockId="2.[225,751,1535,1618]" box="[225,751,1565,1588]" pageId="2" pageNumber="465">
<heading id="D0D03D281A28FFCEA7B3F9F9FD7F9C33" box="[225,751,1565,1588]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="465" reason="8">
<taxonomicName id="4C27F1C71A28FFCEA7B3F9F9FDCE9C33" ID-CoL="8J47S" authority="Kurentsov, 1941" authorityName="Kurentsov" authorityYear="1941" box="[225,606,1565,1588]" class="Insecta" family="Curculionidae" genus="Ernoporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="eggersi">
<emphasis id="B95356561A28FFCEA7B3F9F9FE389C33" box="[225,424,1565,1588]" italics="true" pageId="2" pageNumber="465">Ernoporus eggersi</emphasis>
<bibRefCitation id="EFB6F7B51A28FFCEA6FDF9F9FDCE9C33" author="Kurentsov A. I." box="[431,606,1565,1588]" pageId="2" pageNumber="465" refId="ref2327" refString="Kurentsov A. I. 1941. Koroedy Dal'nego Vostoka SSSR [Bark beetles of the USSR Far East] Moscow-Leningrad: Izdatel'stvo Akademii Nauk SSSR. 234 pp. [In Russian, with English descriptions of new species]" type="book" year="1941">Kurentsov, 1941</bibRefCitation>
</taxonomicName>
<emphasis id="B95356561A28FFCEA537F9F9FD789C33" bold="true" box="[613,744,1565,1588]" pageId="2" pageNumber="465">stat.ressurr</emphasis>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33DD9CF1A28FFCEA6E4F9DFFC1F992B" pageId="2" pageNumber="465" type="description">
<paragraph id="8B988A441A28FFCEA6E4F9DFFD8A9C55" blockId="2.[225,751,1535,1618]" box="[438,538,1595,1618]" pageId="2" pageNumber="465">
<figureCitation id="131C96C11A28FFCEA6E4F9DFFD869C55" box="[438,534,1595,1618]" captionStart="Figs 1–9" captionStartId="1.[188,226,1944,1963]" captionTargetBox="[221,1386,705,1887]" captionTargetId="figure-275@1.[221,1386,704,1907]" captionTargetPageId="1" captionText="Figs 1–9. Habitus and details of the species Scolytus and Ernoporus: 1, 3 — Scolytus emarginatus; 2, 4 — Scolytus schevyrewi; 5, 7 — Ernoporus eggersi; 6, 8 — Ernoporus tiliae; 1–2 — head, male; 3–4 — aedeagus without tegmen and spicule; 5–6 — habitus, males; 7–8 — aedeagus. Рис. 1–9. Габитус и детали строениЯ видов Scolytus и Ernoporus: 1, 3 — Scolytus emarginatus; 2, 4 — Scolytus schevyrewi; 5, 7 — Ernoporus eggersi; 6, 8 — Ernoporus tiliae; 1–2 — голова самца; 3–4 — гениталии самцов беЗ тегмена и спикулы; 5–6 — внешний вид самцов; 7–8 — гениталии самцов Ernoporus." pageId="2" pageNumber="465">Figs 5–8</figureCitation>
.
</paragraph>
<paragraph id="8B988A441A28FFCEA7B1F996FB42980B" blockId="2.[189,787,1650,2076]" lastBlockId="2.[824,1422,303,723]" pageId="2" pageNumber="465">
The name for the species was proposed by
<collectingCountry id="F330CAD41A28FFCEA53BF996FD059C80" box="[617,661,1650,1671]" name="Vietnam" pageId="2" pageNumber="465">V.N.</collectingCountry>
Stark [1936], but the morphological description of the species was provided for the first time in the monograph of A.I.
<bibRefCitation id="EFB6F7B51A28FFCEA531F94CFC9F9CBA" author="Kurentsov A. I." box="[611,783,1704,1725]" pageId="2" pageNumber="465" refId="ref2327" refString="Kurentsov A. I. 1941. Koroedy Dal'nego Vostoka SSSR [Bark beetles of the USSR Far East] Moscow-Leningrad: Izdatel'stvo Akademii Nauk SSSR. 234 pp. [In Russian, with English descriptions of new species]" type="book" year="1941">Kurentsov [1941]</bibRefCitation>
.
<collectingCountry id="F330CAD41A28FFCEA7EFF927FF799CDF" box="[189,233,1731,1752]" name="Vietnam" pageId="2" pageNumber="465">V.N.</collectingCountry>
<bibRefCitation id="EFB6F7B51A28FFCEA7A2F927FEF99CDF" author="Stark V. N." box="[240,361,1731,1752]" pageId="2" pageNumber="465" refId="ref2599" refString="Stark V. N. 1952. Koroedy // Fauna SSSR. Zhestkokrylye. Vol. 31. Moscow - Leningrad: Izdatel'stvo Akademii Nauk SSSR. 462 pp. [In Russian]" type="book" year="1952">Stark [1952]</bibRefCitation>
gave the following characteristics to distinguish
<taxonomicName id="4C27F1C71A28FFCEA7A5F93AFEC89CF4" authorityName="Kurentsov" authorityYear="1941" box="[247,344,1758,1779]" class="Insecta" family="Curculionidae" genus="Ernoporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="eggersi">
<emphasis id="B95356561A28FFCEA7A5F93AFEC89CF4" box="[247,344,1758,1779]" italics="true" pageId="2" pageNumber="465">E. eggersi</emphasis>
</taxonomicName>
from
<taxonomicName id="4C27F1C71A28FFCEA6C6F93AFDE59CF4" authority="(Panzer, 1793)" baseAuthorityName="Panzer" baseAuthorityYear="1793" box="[404,629,1758,1779]" class="Insecta" family="Curculionidae" genus="Ernoporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="tiliae">
<emphasis id="B95356561A28FFCEA6C6F93AFE709CF4" box="[404,480,1758,1779]" italics="true" pageId="2" pageNumber="465">E. tiliae</emphasis>
(Panzer, 1793)
</taxonomicName>
: more elongated body form and strong metallic shining luster of the body and broader elytral striae. In contemporary literature
<taxonomicName id="4C27F1C71A28FFCEA5CAF8F0FD6A9D2E" authorityName="Kurentsov" authorityYear="1941" box="[664,762,1812,1833]" class="Insecta" family="Curculionidae" genus="Ernoporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="eggersi">
<emphasis id="B95356561A28FFCEA5CAF8F0FD6A9D2E" box="[664,762,1812,1833]" italics="true" pageId="2" pageNumber="465">E. eggersi</emphasis>
</taxonomicName>
is considered a junior synonym of
<taxonomicName id="4C27F1C71A28FFCEA6BCF8CBFDA99D43" baseAuthorityName="Panzer" baseAuthorityYear="1793" box="[494,569,1839,1860]" class="Insecta" family="Curculionidae" genus="Ernoporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="tiliae">
<emphasis id="B95356561A28FFCEA6BCF8CBFDA99D43" box="[494,569,1839,1860]" italics="true" pageId="2" pageNumber="465">E. tiliae</emphasis>
</taxonomicName>
[
<bibRefCitation id="EFB6F7B51A28FFCEA516F8CBFC999D43" author="Johnson A. J. &amp; Hulcr J. &amp; Knizek M. &amp; Atkinson T. H. &amp; Mandelshtam M. Yu. &amp; Smith S. M. &amp; Cognato A. I. &amp; Park S. &amp; Li Y. &amp; Jordal B. H." box="[580,777,1839,1860]" pageId="2" pageNumber="465" pagination="1 - 81" refId="ref2122" refString="Johnson A. J., Hulcr J., Knizek M., Atkinson T. H., Mandelshtam M. Yu., Smith S. M., Cognato A. I., Park S., Li Y., Jordal B. H. 2020. Revision of the bark beetle genera within the former Cryphalini (Curculionidae: Scolytinae) // Insect Systematics and Diversity. No. 4 (3). P. 1 - 81. https: // doi. org / 10.1093 / isd / ixaa 002" type="book chapter" year="2020">
Johnson
<emphasis id="B95356561A28FFCEA5C8F8CBFD539D43" box="[666,707,1839,1860]" italics="true" pageId="2" pageNumber="465">et al</emphasis>
., 2020
</bibRefCitation>
]. In
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vicinity of Vladivostok (Bogataya River) A.V. Petrov collected a series of
<taxonomicName id="4C27F1C71A28FFCEA6D2F881FE709D7D" authorityName="Kurentsov" authorityYear="1941" box="[384,480,1893,1914]" class="Insecta" family="Curculionidae" genus="Ernoporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="eggersi">
<emphasis id="B95356561A28FFCEA6D2F881FE709D7D" box="[384,480,1893,1914]" italics="true" pageId="2" pageNumber="465">E. eggersi</emphasis>
</taxonomicName>
consisting of
<specimenCount id="9D2141CD1A28FFCEA53AF881FD789D7D" box="[616,744,1893,1914]" count="32" pageId="2" pageNumber="465" type="generic">32 specimens</specimenCount>
that allowed detailed morphological study of the species. In our opinion, features of outer morphology [
<bibRefCitation id="EFB6F7B51A28FFCEA564F87FFD389DB7" author="Stark V. N." box="[566,680,1947,1968]" pageId="2" pageNumber="465" refId="ref2599" refString="Stark V. N. 1952. Koroedy // Fauna SSSR. Zhestkokrylye. Vol. 31. Moscow - Leningrad: Izdatel'stvo Akademii Nauk SSSR. 462 pp. [In Russian]" type="book" year="1952">Stark, 1952</bibRefCitation>
] do not allow to separate the two closely related species (
<figureCitation id="131C96C11A28FFCEA5D7F852FD4C9DCC" box="[645,732,1974,1995]" captionStart="Figs 1–9" captionStartId="1.[188,226,1944,1963]" captionTargetBox="[221,1386,705,1887]" captionTargetId="figure-275@1.[221,1386,704,1907]" captionTargetPageId="1" captionText="Figs 1–9. Habitus and details of the species Scolytus and Ernoporus: 1, 3 — Scolytus emarginatus; 2, 4 — Scolytus schevyrewi; 5, 7 — Ernoporus eggersi; 6, 8 — Ernoporus tiliae; 1–2 — head, male; 3–4 — aedeagus without tegmen and spicule; 5–6 — habitus, males; 7–8 — aedeagus. Рис. 1–9. Габитус и детали строениЯ видов Scolytus и Ernoporus: 1, 3 — Scolytus emarginatus; 2, 4 — Scolytus schevyrewi; 5, 7 — Ernoporus eggersi; 6, 8 — Ernoporus tiliae; 1–2 — голова самца; 3–4 — гениталии самцов беЗ тегмена и спикулы; 5–6 — внешний вид самцов; 7–8 — гениталии самцов Ernoporus." pageId="2" pageNumber="465">Figs 5–6</figureCitation>
). The only reliable feature to distinguish
<taxonomicName id="4C27F1C71A28FFCEA555F835FDF79DE1" authorityName="Kurentsov" authorityYear="1941" box="[519,615,2001,2022]" class="Insecta" family="Curculionidae" genus="Ernoporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="eggersi">
<emphasis id="B95356561A28FFCEA555F835FDF79DE1" box="[519,615,2001,2022]" italics="true" pageId="2" pageNumber="465">E. eggersi</emphasis>
</taxonomicName>
is the structure of the male genitalia. Males of
<taxonomicName id="4C27F1C71A28FFCEA69EF808FDBC9206" authorityName="Kurentsov" authorityYear="1941" box="[460,556,2028,2049]" class="Insecta" family="Curculionidae" genus="Ernoporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="eggersi">
<emphasis id="B95356561A28FFCEA69EF808FDBC9206" box="[460,556,2028,2049]" italics="true" pageId="2" pageNumber="465">E. eggersi</emphasis>
</taxonomicName>
possess a broad tegmen of semicircular form; apophyses very short, penis body is 2.3 times as long as apophyses; penis tube elongated, 2.9 times as long as wide; supporting penis structure is located not on penis apex, but closer to penis middle (
<figureCitation id="131C96C11A28FFCEA339FE8CFB349B7A" box="[1131,1188,360,381]" captionStart="Figs 1–9" captionStartId="1.[188,226,1944,1963]" captionTargetBox="[221,1386,705,1887]" captionTargetId="figure-275@1.[221,1386,704,1907]" captionTargetPageId="1" captionText="Figs 1–9. Habitus and details of the species Scolytus and Ernoporus: 1, 3 — Scolytus emarginatus; 2, 4 — Scolytus schevyrewi; 5, 7 — Ernoporus eggersi; 6, 8 — Ernoporus tiliae; 1–2 — head, male; 3–4 — aedeagus without tegmen and spicule; 5–6 — habitus, males; 7–8 — aedeagus. Рис. 1–9. Габитус и детали строениЯ видов Scolytus и Ernoporus: 1, 3 — Scolytus emarginatus; 2, 4 — Scolytus schevyrewi; 5, 7 — Ernoporus eggersi; 6, 8 — Ernoporus tiliae; 1–2 — голова самца; 3–4 — гениталии самцов беЗ тегмена и спикулы; 5–6 — внешний вид самцов; 7–8 — гениталии самцов Ernoporus." pageId="2" pageNumber="465">Fig. 7</figureCitation>
). In males of
<taxonomicName id="4C27F1C71A28FFCEA270FE8CFAFB9B7A" baseAuthorityName="Panzer" baseAuthorityYear="1793" box="[1314,1387,360,381]" class="Insecta" family="Curculionidae" genus="Ernoporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="tiliae">
<emphasis id="B95356561A28FFCEA270FE8CFAFB9B7A" box="[1314,1387,360,381]" italics="true" pageId="2" pageNumber="465">E. tiliae</emphasis>
</taxonomicName>
the tegmen is narrow, apophyses are short, penis body is 2.0 times as long as apophyses; penis tube more broad, 1.7 times as long as wide; supporting penis structure is located on the penis apex (
<figureCitation id="131C96C11A28FFCEA46DFE3EFCEA9BE8" box="[831,890,474,495]" captionStart="Figs 1–9" captionStartId="1.[188,226,1944,1963]" captionTargetBox="[221,1386,705,1887]" captionTargetId="figure-275@1.[221,1386,704,1907]" captionTargetPageId="1" captionText="Figs 1–9. Habitus and details of the species Scolytus and Ernoporus: 1, 3 — Scolytus emarginatus; 2, 4 — Scolytus schevyrewi; 5, 7 — Ernoporus eggersi; 6, 8 — Ernoporus tiliae; 1–2 — head, male; 3–4 — aedeagus without tegmen and spicule; 5–6 — habitus, males; 7–8 — aedeagus. Рис. 1–9. Габитус и детали строениЯ видов Scolytus и Ernoporus: 1, 3 — Scolytus emarginatus; 2, 4 — Scolytus schevyrewi; 5, 7 — Ernoporus eggersi; 6, 8 — Ernoporus tiliae; 1–2 — голова самца; 3–4 — гениталии самцов беЗ тегмена и спикулы; 5–6 — внешний вид самцов; 7–8 — гениталии самцов Ernoporus." pageId="2" pageNumber="465">Fig. 8</figureCitation>
).
<taxonomicName id="4C27F1C71A28FFCEA4DDFE3EFBD09BE8" authorityName="Kurentsov" authorityYear="1941" box="[911,1088,474,495]" class="Insecta" family="Curculionidae" genus="Ernoporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="eggersi">
<emphasis id="B95356561A28FFCEA4DDFE3EFBD09BE8" box="[911,1088,474,495]" italics="true" pageId="2" pageNumber="465">Ernoporus eggersi</emphasis>
</taxonomicName>
thus is restored as a valid species and removed from synonymy with
<taxonomicName id="4C27F1C71A28FFCEA3D0FE13FB5C980B" baseAuthorityName="Panzer" baseAuthorityYear="1793" box="[1154,1228,503,524]" class="Insecta" family="Curculionidae" genus="Ernoporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="tiliae">
<emphasis id="B95356561A28FFCEA3D0FE13FB5C980B" box="[1154,1228,503,524]" italics="true" pageId="2" pageNumber="465">E. tiliae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B988A441A28FFCEA40CFDF7FBBF98D4" blockId="2.[824,1422,303,723]" pageId="2" pageNumber="465">
From the long syntypic series of
<taxonomicName id="4C27F1C71A28FFCEA3C9FDF7FA1D982F" authority="(ZIN)" baseAuthorityName="ZIN" box="[1179,1421,531,552]" class="Insecta" family="Curculionidae" genus="Ernoporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="eggersi">
<emphasis id="B95356561A28FFCEA3C9FDF7FADF982F" box="[1179,1359,531,552]" italics="true" pageId="2" pageNumber="465">Ernoporus eggersi</emphasis>
(ZIN)
</taxonomicName>
we selected the
<typeStatus id="549C34E61A28FFCEA487FDD4FBBD9842" box="[981,1069,560,581]" pageId="2" pageNumber="465" type="lectotype">lectotype</typeStatus>
(sex unknown) with the geographic label: “239. Майхинский ЛПХ [ЛеспромхоЗ], Соловейцев ключ,
<taxonomicName id="4C27F1C71A28FFCEA428FD8DFB9F9879" box="[890,1039,617,638]" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" pageId="2" pageNumber="465" phylum="Tracheophyta" rank="species" species="amurensis">
<emphasis id="B95356561A28FFCEA428FD8DFB9F9879" box="[890,1039,617,638]" italics="true" pageId="2" pageNumber="465">Tilia amurensis</emphasis>
</taxonomicName>
, 20.IX. [19]31” (239. Maikhe Forestry Enterprise, Soloveytsev stream, 20.IX. [19]31”) bearing also the additional label “collected by Kurentsov on Far East” probably added by
<collectingCountry id="F330CAD41A28FFCEA496FD5AFC6198D4" box="[964,1009,702,723]" name="Vietnam" pageId="2" pageNumber="465">V.N.</collectingCountry>
Stark.
</paragraph>
<paragraph id="8B988A441A28FFCEA40CFD13FC1F992B" blockId="2.[824,1421,759,812]" pageId="2" pageNumber="465">
<emphasis id="B95356561A28FFCEA40CFD13FBA4990B" bold="true" box="[862,1076,759,780]" pageId="2" pageNumber="465">Competing interests.</emphasis>
The authors declare no competing interests.
</paragraph>
</subSubSection>
</treatment>
</document>

278
data/03/FA/C7/03FAC7406140FFD6B4BFFBC0C2B3F95F.xml Normal file
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@ -0,0 +1,278 @@
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<mods:title id="77375EE85222A7D8ACC7CAD8458504F7">Seven new species of dance flies of the subgenus Polyblepharis Bezzi, 1909 (Diptera: Empididae) from Mongolia and East Siberia</mods:title>
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<treatment id="03FAC7406140FFD6B4BFFBC0C2B3F95F" LSID="urn:lsid:plazi:treatment:03FAC7406140FFD6B4BFFBC0C2B3F95F" httpUri="http://treatment.plazi.org/id/03FAC7406140FFD6B4BFFBC0C2B3F95F" lastPageId="5" lastPageNumber="501" pageId="3" pageNumber="499">
<subSubSection id="C34925DD6140FFD0B4BFFBC0C1BEFBA5" box="[217,758,1080,1103]" pageId="3" pageNumber="499" type="nomenclature">
<paragraph id="8BEC76566140FFD0B4BFFBC0C1BEFBA5" blockId="3.[217,758,1080,1132]" box="[217,758,1080,1103]" pageId="3" pageNumber="499">
<heading id="D0A4C13A6140FFD0B4BFFBC0C1BEFBA5" box="[217,758,1080,1103]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="499" reason="8">
<taxonomicName id="4C530DD56140FFD0B4BFFBC0C1FFFBA5" authority="Shamshev" authorityName="Shamshev" box="[217,695,1080,1103]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="499" phylum="Arthropoda" rank="species" species="mongolica" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA446140FFD0B4BFFBC0C254FBA5" box="[217,284,1080,1103]" italics="true" pageId="3" pageNumber="499">Empis</emphasis>
(
<emphasis id="B927AA446140FFD0B54DFBC0C289FBA5" box="[299,449,1080,1103]" italics="true" pageId="3" pageNumber="499">Polyblepharis</emphasis>
)
<emphasis id="B927AA446140FFD0B5B6FBC0C109FBA5" box="[464,577,1080,1103]" italics="true" pageId="3" pageNumber="499">mongolica</emphasis>
Shamshev
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,
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<taxonomicNameLabel id="A214173F6140FFD0B6A7FBC0C1BEFBA5" box="[705,758,1080,1103]" pageId="3" pageNumber="499" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6140FFD0B5D0FBADC152FB86" box="[438,538,1109,1132]" pageId="3" pageNumber="499" type="description">
<paragraph id="8BEC76566140FFD0B5D0FBADC152FB86" blockId="3.[217,758,1080,1132]" box="[438,538,1109,1132]" pageId="3" pageNumber="499">
<figureCitation id="13686AD36140FFD0B5D0FBADC15EFB86" box="[438,534,1109,1132]" captionStart="Figs 6–9" captionStartId="4.[190,228,1932,1951]" captionTargetBox="[216,1380,547,1855]" captionTargetId="figure-103@4.[200,1410,527,1877]" captionTargetPageId="4" captionText="Figs 6–9. Empis (Polyblepharis) mongolica Shamshev, sp.n., male: 6 — habitus, holotype, lateral view; 7 — head and thorax, holotype, anterodorsal view; 8 — hypopygium, lateral view; 9 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 6–9. Empis (Polyblepharis) mongolica Shamshev, sp.n., самец: 6 — габитус, голотип, вид сбоку; 7 — голова и грудь, голотип, вид спереди-сверху; 8 — гипопигий, вид сбоку; 9 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="3" pageNumber="499">Figs 6–9</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6140FFD0B485FB77C18BFA93" pageId="3" pageNumber="499" type="materials_examined">
<paragraph id="8BEC76566140FFD0B485FB77C2CDFAE5" blockId="3.[189,788,1167,2068]" pageId="3" pageNumber="499">
<materialsCitation id="3B3B7C0B6140FFD0B485FB77C2CDFAE5" collectionCode="TYPE, MATERIAL, ZISP, INS" country="Mongolia" location="Zaitzev" municipality="Khasagt-Khayrkhan Ridge" pageId="3" pageNumber="499" specimenCount="1" specimenCount-male="1" stateProvince="Govi-Altay" typeStatus="holotype">
<collectionCode id="ED42EE936140FFD0B485FB77C2E0FB4E" box="[227,424,1167,1188]" pageId="3" pageNumber="499">TYPE MATERIAL</collectionCode>
.
<typeStatus id="54E8C8F46140FFD0B5D0FB77C15AFB4E" box="[438,530,1167,1188]" pageId="3" pageNumber="499" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D55BDDF6140FFD0B647FB77C17BFB4E" box="[545,563,1167,1188]" count="1" pageId="3" pageNumber="499" type="male">♂</specimenCount>
, labelled [in Cyrillic, Russian]:
<collectingCountry id="F34436C66140FFD0B544FB52C2CAFB55" box="[290,386,1194,1215]" name="Mongolia" pageId="3" pageNumber="499">Mongolia</collectingCountry>
, G.-Alt. aym. [=Gobi-Altayskiy aymak, now
<collectingRegion id="4997B8B46140FFD0B493FB3CC215FB33" box="[245,349,1220,1241]" country="Mongolia" name="Govi-Altay" pageId="3" pageNumber="499">Govi-Altai</collectingRegion>
aymag (province)], /
<collectingMunicipality id="6B88EC2C6140FFD0B637FB3CC3BEFB1E" pageId="3" pageNumber="499">Khasagt-Khayrkhan Ridge</collectingMunicipality>
/
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<quantity id="4CABDBB36140FFD0B576FB27C218FB1E" box="[272,336,1247,1268]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="499" unit="km" value="15.0">15 km</quantity>
S Dzhargalan
</locationDeviation>
/ 14.viii.[1]970
<location id="8E8C208D6140FFD0B6E2FB27C184FB1E" LSID="urn:lsid:plazi:treatment:03FAC7406140FFD6B4BFFBC0C2B3F95F:8E8C208D6140FFD0B6E2FB27C184FB1E" box="[644,716,1247,1268]" country="Mongolia" municipality="Khasagt-Khayrkhan Ridge" name="Zaitzev" pageId="3" pageNumber="499" stateProvince="Govi-Altay">Zaitzev</location>
(
<collectionCode id="ED42EE936140FFD0B6BAFB27C047FB1E" box="[732,783,1247,1268]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="3" pageNumber="499">ZISP</collectionCode>
,
<collectionCode id="ED42EE936140FFD0B4DBFB02C3ABFAE5" box="[189,227,1274,1295]" pageId="3" pageNumber="499">INS</collectionCode>
_DIP_0000628).
</materialsCitation>
</paragraph>
<paragraph id="8BEC76566140FFD0B485FAECC18BFA93" blockId="3.[189,788,1167,2068]" pageId="3" pageNumber="499">
<materialsCitation id="3B3B7C0B6140FFD0B485FAECC119FAAE" collectingDate="1970-08-14" collectionCode="TYPE, MATERIAL, ZISP, INS" country="Mongolia" location="Nartshuk" municipality="Khasagt-Khayrkhan Ridge" pageId="3" pageNumber="499" specimenCount="1" specimenCount-male="1" stateProvince="Govi-Altay" typeStatus="paratype">
<typeStatus id="54E8C8F46140FFD0B485FAECC20BFAC3" box="[227,323,1300,1321]" pageId="3" pageNumber="499" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F34436C66140FFD0B52AFAECC29AFAC3" box="[332,466,1300,1321]" name="Mongolia" pageId="3" pageNumber="499">MONGOLIA</collectingCountry>
.
<collectingRegion id="4997B8B46140FFD0B5BCFAECC10AFAC3" box="[474,578,1300,1321]" country="Mongolia" name="Govi-Altay" pageId="3" pageNumber="499">Govi-Altai</collectingRegion>
aymag: same locality as holotype,
<date id="FFED50966140FFD0B55AFAD7C2F8FAAE" box="[316,432,1327,1348]" pageId="3" pageNumber="499" value="1970-08-14">
<collectingDate id="EFA9A97E6140FFD0B55AFAD7C2F8FAAE" box="[316,432,1327,1348]" pageId="3" pageNumber="499" value="1970-08-14">14.viii.1970</collectingDate>
</date>
<location id="8E8C208D6140FFD0B5DEFAD7C158FAAE" LSID="urn:lsid:plazi:treatment:03FAC7406140FFD6B4BFFBC0C2B3F95F:8E8C208D6140FFD0B5DEFAD7C158FAAE" box="[440,528,1327,1348]" country="Mongolia" name="Nartshuk" pageId="3" pageNumber="499" stateProvince="Govi-Altay">Nartshuk</location>
(
<specimenCount id="9D55BDDF6140FFD0B644FAD7C10FFAAE" box="[546,583,1327,1348]" count="1" pageId="3" pageNumber="499" type="male">1 ♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="3B3B7C0B6140FFD0B638FAD7C1F7FA93" collectingDate="1969-08-07" collectingDateMax="1969-08-08" collectingDateMin="1969-08-07" collectionCode="ZISP" collectorName="N Bayan-Gobi" country="Mongolia" elevation="2450" location="Bayankhongor" pageId="3" pageNumber="499" specimenCount="1" specimenCount-male="1" stateProvince="Bayanhongor" typeStatus="paratype">
<collectingRegion id="4997B8B46140FFD0B638FAD7C1A3FAAE" box="[606,747,1327,1348]" country="Mongolia" name="Bayanhongor" pageId="3" pageNumber="499">Bayankhongor</collectingRegion>
aymag: southern slope of Ikh-Bogdo-Ul,
<collectorName id="26A613806140FFD0B65CFAB2C183FAB5" box="[570,715,1354,1375]" pageId="3" pageNumber="499">N Bayan-Gobi</collectorName>
,
<quantity id="4CABDBB36140FFD0B6B0FAB2C240FA93" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="3" pageNumber="499" unit="m" value="2450.0" valueMax="2700.0" valueMin="2200.0">
<elevation id="007E91656140FFD0B6B0FAB2C240FA93" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="3" pageNumber="499" unit="m" value="2450.0" valueMax="2700.0" valueMin="2200.0">2200– 2700 m</elevation>
</quantity>
,
<date id="FFED50966140FFD0B574FA9CC2DAFA93" box="[274,402,1380,1401]" pageId="3" pageNumber="499" value="1969-08-07" valueMax="1969-08-08" valueMin="1969-08-07">
<collectingDate id="EFA9A97E6140FFD0B574FA9CC2DAFA93" box="[274,402,1380,1401]" pageId="3" pageNumber="499" value="1969-08-07" valueMax="1969-08-08" valueMin="1969-08-07">7–8.viii.1969</collectingDate>
</date>
Zaitzev (
<specimenCount id="9D55BDDF6140FFD0B589FA9CC15AFA93" box="[495,530,1380,1401]" count="1" pageId="3" pageNumber="499" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED42EE936140FFD0B678FA9CC11AFA93" box="[542,594,1380,1401]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="3" pageNumber="499">ZISP</collectionCode>
; dissected)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6140FFD0B485FA87C1D0F9DE" pageId="3" pageNumber="499" type="diagnosis">
<paragraph id="8BEC76566140FFD0B485FA87C1D0F9DE" blockId="3.[189,788,1167,2068]" pageId="3" pageNumber="499">
DIAGNOSIS. Mid-sized species (body length about
<quantity id="4CABDBB36140FFD0B4DBFA62C3B2FA45" box="[189,250,1434,1455]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="499" unit="mm" value="6.0">6 mm</quantity>
); male eyes holoptic; palpus yellow; mesoscutum with 3 indistinct, brownish vittae; acrostichal and dorsocentral setae multiserial, pale in presutural part, laterotergite with pale setae; fore and mid legs mostly brownish, hind leg mostly brownish yellow; wing hyaline; abdominal tergites
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lateral view uniformly brownish pruinose; hypopygium large.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6140FFD6B485F9C2C1C9F98F" lastPageId="5" lastPageNumber="501" pageId="3" pageNumber="499" type="description">
<paragraph id="8BEC76566140FFD0B485F9C2C75CFEB5" blockId="3.[189,788,1167,2068]" lastBlockId="3.[824,1422,303,2057]" pageId="3" pageNumber="499">
DESCRIPTION. Body length 6.0–
<quantity id="4CABDBB36140FFD0B678F9C2C12CF9A5" box="[542,612,1594,1615]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="499" unit="mm" value="6.3">6.3 mm</quantity>
, wing
<quantity id="4CABDBB36140FFD0B6C6F9C2C058F9A5" box="[672,784,1594,1615]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="5.8" metricValueMin="5.7" pageId="3" pageNumber="499" unit="mm" value="5.75" valueMax="5.8" valueMin="5.7">5.7–5.8 mm</quantity>
.
<emphasis id="B927AA446140FFD0B4DBF9ACC3B9F983" bold="true" box="[189,241,1620,1641]" pageId="3" pageNumber="499">Male</emphasis>
(
<figureCitation id="13686AD36140FFD0B567F9ACC277F983" box="[257,319,1620,1641]" captionStart="Figs 6–9" captionStartId="4.[190,228,1932,1951]" captionTargetBox="[216,1380,547,1855]" captionTargetId="figure-103@4.[200,1410,527,1877]" captionTargetPageId="4" captionText="Figs 6–9. Empis (Polyblepharis) mongolica Shamshev, sp.n., male: 6 — habitus, holotype, lateral view; 7 — head and thorax, holotype, anterodorsal view; 8 — hypopygium, lateral view; 9 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 6–9. Empis (Polyblepharis) mongolica Shamshev, sp.n., самец: 6 — габитус, голотип, вид сбоку; 7 — голова и грудь, голотип, вид спереди-сверху; 8 — гипопигий, вид сбоку; 9 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="3" pageNumber="499">Fig. 6</figureCitation>
). Head capsule regions mostly densely greyish pruinose; face on lower margin and entire clypeus shiny. Holoptic, upper ommatidia enlarged. Frons represented by very small subtriangular space just below ocellar triangle and larger space above antennae; bare. Face broad, bare (in
<specimenCount id="9D55BDDF6140FFD0B6F1F947C05BF93E" box="[663,787,1727,1748]" count="1" pageId="3" pageNumber="499" type="generic" typeStatus="paratype">one paratype</specimenCount>
1 setula present just below antennae). Ocellar triangle with several moderately long, mostly pale (1–2 brownish to black setae usually present), fine setae; ocellars undifferentiated. Occiput mostly with numerous, pale, hair-like setae, bearing moderately long, strong, black setae dorsally; postoculars moderately long, fine, black and pale on upper part, very short laterally; postgena with pale hair-like setae. Antenna with scape and pedicel brown, postpedicel and stylus black; scape slightly longer than subglobular pedicel, both with very short setulae; postpedicel nearly 3X as long as basal width, with straight margins; stylus slightly longer than postpedicel basal width. Proboscis with labrum brownish yellow, nearly 1.5X longer than eye height; palpus short, pale yellow, with rather numerous pale setulae.
</paragraph>
<paragraph id="8BEC76566140FFD0B738FE9CC6C6FBFE" blockId="3.[824,1422,303,2057]" pageId="3" pageNumber="499">
Thorax black in ground-colour, mostly densely greyish pruinose; mesoscutum (
<figureCitation id="13686AD36140FFD0B04EFE87C720FE7E" box="[1064,1128,383,404]" captionStart="Figs 6–9" captionStartId="4.[190,228,1932,1951]" captionTargetBox="[216,1380,547,1855]" captionTargetId="figure-103@4.[200,1410,527,1877]" captionTargetPageId="4" captionText="Figs 6–9. Empis (Polyblepharis) mongolica Shamshev, sp.n., male: 6 — habitus, holotype, lateral view; 7 — head and thorax, holotype, anterodorsal view; 8 — hypopygium, lateral view; 9 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 6–9. Empis (Polyblepharis) mongolica Shamshev, sp.n., самец: 6 — габитус, голотип, вид сбоку; 7 — голова и грудь, голотип, вид спереди-сверху; 8 — гипопигий, вид сбоку; 9 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="3" pageNumber="499">Fig. 7</figureCitation>
) with 3 indistinct, brownish vittae along acrostichal and dorsocentral setae. Antepronotum with 4–5 strong, black setae dorsally and long, hair-like setae laterally on each side (dorsal setae sometimes brownish yellow to pale). Postpronotal lobe with long, dense, fine, pale setae, 1 similar black seta. Proepisternum with tuft of numerous, pale, hair-like setae on lower part. Prosternum bare. Mesonotal setae: acrostichals pale and black on about anterior 1/3 of mesoscutum (sometimes mostly pale or mostly dark) and black posteriorly, arranged in 4–6 irregular rows, occupying anterior half of prescutellar depression, moderately long, dense; presutural dorsocentral setae black and pale, multiserial, similar to acrostichals; postsutural dorsocentrals black, becoming stronger, longer and less numerous towards scutellum, not separated by bare space from acrostichals on about anterior half of prescutellar depression, 3–4 prescutellars longest; presutural supra-alar space entirely covered with dense, long, pale, hair-like setae not separated by bare space from dorsocentrals (sometimes 1–2 black hair-like setae present); 4 black notopleurals (anterior seta usually slightly shorter); 1 similar postsutural supra-alar (with some additional short, fine, black and hair-like, pale setae); 1 long and 1 minute postalars, 4 scutellars of subequal lengths (lateral setae only slightly shorter than apicals). Laterotergite with numerous pale to pale yellow setae. Anterior and posterior spiracles pale yellow.
</paragraph>
<paragraph id="8BEC76566140FFD0B738FBE2C79EF8F3" blockId="3.[824,1422,303,2057]" pageId="3" pageNumber="499">Legs with coxae densely greyish pruinose, remaining podomeres subshiny; mostly black setose; Leg colour: coxae and trochanters brown; fore and mid femora mostly brownish (yellowish at extreme apex and base; fore femur sometimes entirely brownish), hind femur brownish yellow; fore and mid tibiae mostly brownish (brownish yellow close to base), hind tibia brownish yellow (slightly darker at apex); fore and mid tarsi almost entirely brownish (only basitarsi paler at extreme base), hind basitarsus mostly brownish yellow (brownish at apex), remaining hind tarsomeres brownish. Fore coxa covered with pale, long, dense, hair-like setae anteriorly. All femora without whitish pilosity ventrally. Fore femur with row of mostly minute, fine anteroventral setae; covered with hair-like, moderately long, pale setae posteroventrally and posteriorly. Fore tibia with 3–5 short, anterodorsal setae (position and robustness variable). Mid femur with complete rows of equally short, strong anteroventral and posteroventral setae. Mid tibia slender; with 3–4 anterodorsal and 2 similar posterodorsal, moderately long setae. Hind femur slender; with row of 3–4 moderately long anterodorsal setae in about apical third; complete row of rather short and scattered anteroventral setae; fine short setae posteroventrally (longer close to base). Hind tibia simple, slender; with 7–9 moderately long, strong anterodorsal and 3–4 similar posterodorsal setae; simple setulae ventrally; 1 short seta in posteroapical comb. Tarsomeres of fore and mid legs slender, hind basitarsus slightly thickened at base; basitarsi with circlet of short, subapical setae, bearing scattered, spine-like setae ventrally; in addition, hind basitarsus with 2 short anterodorsal setae on about middle.</paragraph>
<paragraph id="8BEC76566140FFD7B738F8E7C19CFEB5" blockId="3.[824,1422,303,2057]" lastBlockId="4.[189,787,303,457]" lastPageId="4" lastPageNumber="500" pageId="3" pageNumber="499">
Wing membrane hyaline; entirely covered with microtrichia; veins mostly yellowish (costa between anterior margin of pterostigma and R 5 as well as apical portion of radial veins brownish to brownish yellow), well-sclerotized. Veins R 5 and M
<subScript id="17D774136140FFD0B71EF860C037F84E" attach="left" box="[888,895,1944,1956]" fontSize="5" pageId="3" pageNumber="499">1</subScript>
divergent towards wing margin; radial fork acute; R
<subScript id="17D774136140FFD0B1E0F860C6C5F84E" attach="left" box="[1414,1421,1944,1956]" fontSize="5" pageId="3" pageNumber="499">5</subScript>
meeting costa before wing apex; CuA+CuP complete (slightly weakened beyond middle). Cell dm moderately large, with strongly elongate apex; apical portion of M
<subScript id="17D774136140FFD0B085F810C7A2F81E" attach="left" box="[1251,1258,2024,2036]" fontSize="5" pageId="3" pageNumber="499">4</subScript>
nearly 3X long- er than its middle portion. Pterostigma inconspicuous. Basal costal seta absent. Anal lobe well-developed; axillary incision very acute. Squama yellow, pale fringed. Halter yellow.
</paragraph>
<paragraph id="8BEC76566147FFD6B485FE9CC127FD81" blockId="4.[189,787,303,457]" lastBlockId="5.[189,787,303,1717]" lastPageId="5" lastPageNumber="501" pageId="4" pageNumber="500">Abdomen entirely black in ground colour. Pruinescence: tergites 1–5 densely brownish grey pruinose, tergites 6–7 with yellowish brown iridescent spot dorsally, greyish laterally (dorsal view); sternites 1–7 densely greyish pruinose; sternite 8 faintly pruinose, contrastingly black (upper antero-lateral projection shiny). Chaetotaxy: tergites 1–5 covered with black, minute, numerous setulae dorsally, bearing pale, dense, hair-like and yellowish posteromarginal setae laterally; tergites 6–7 only with pale lateral setae; sternite 1 with 2 long, fine, pale setae, sternites 2–8 with numerous similar setae (slightly shorter on sternite 8), in addition, sternite 8 with moderately long, black posteromarginal setae. Pregenital segments: segment 6 unmodified; tergite 7 slightly concave posteriorly, sternite 7 with slightly produced postero-lateral corner, pleuron of segment 7 slightly sclerotised posteriorly; segment 8 with separated tergite and sternite; tergite 8 somewhat flattened, represented by two subtriangular sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 large, scoop-shaped, somewhat constricted anteriorly, with 2 small, antero-lateral projections separated by depression (upper projection rounded, lower projection subconical) and large, broad projection on posterior margin of depression.</paragraph>
<caption id="DF2C26DE6147FFD7B4D8F874C00AF7FD" pageId="4" pageNumber="500" startId="4.[190,228,1932,1951]" targetBox="[216,1380,547,1855]" targetPageId="4" targetType="figure">
<paragraph id="8BEC76566147FFD7B4D8F874C029F825" blockId="4.[189,1422,1932,2071]" pageId="4" pageNumber="500">
<emphasis id="B927AA446147FFD7B4D8F874C258F875" bold="true" box="[190,272,1932,1951]" pageId="4" pageNumber="500">Figs 6–9.</emphasis>
<taxonomicName id="4C530DD56147FFD7B571F874C1D2F875" authority="Shamshev" authorityName="Shamshev" box="[279,666,1932,1951]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="500" phylum="Arthropoda" rank="species" species="mongolica" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA446147FFD7B571F874C204F875" box="[279,332,1932,1951]" italics="true" pageId="4" pageNumber="500">Empis</emphasis>
(
<emphasis id="B927AA446147FFD7B53CF874C29AF875" box="[346,466,1932,1951]" italics="true" pageId="4" pageNumber="500">Polyblepharis</emphasis>
)
<emphasis id="B927AA446147FFD7B586F874C172F875" box="[480,570,1932,1951]" italics="true" pageId="4" pageNumber="500">mongolica</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA446147FFD7B6C2F874C187F875" bold="true" box="[676,719,1932,1951]" pageId="4" pageNumber="500">
<taxonomicNameLabel id="A214173F6147FFD7B6C2F874C187F875" box="[676,719,1932,1951]" pageId="4" pageNumber="500" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, male: 6 — habitus, holotype, lateral view; 7 — head and thorax, holotype, anterodorsal view; 8 — hypopygium, lateral view; 9 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm.
</paragraph>
<paragraph id="8BEC76566147FFD7B4DBF82CC00AF7FD" blockId="4.[189,1422,1932,2071]" pageId="4" pageNumber="500">
<emphasis id="B927AA446147FFD7B4DBF82CC25BF80D" bold="true" box="[189,275,2004,2023]" pageId="4" pageNumber="500">Рис. 6–9.</emphasis>
<taxonomicName id="4C530DD56147FFD7B57AF82CC1EDF80D" authority="Shamshev" authorityName="Shamshev" box="[284,677,2004,2023]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="500" phylum="Arthropoda" rank="species" species="mongolica" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA446147FFD7B57AF82CC219F80D" box="[284,337,2004,2023]" italics="true" pageId="4" pageNumber="500">Empis</emphasis>
(
<emphasis id="B927AA446147FFD7B504F82CC292F80D" box="[354,474,2004,2023]" italics="true" pageId="4" pageNumber="500">Polyblepharis</emphasis>
)
<emphasis id="B927AA446147FFD7B58FF82CC10BF80D" box="[489,579,2004,2023]" italics="true" pageId="4" pageNumber="500">mongolica</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA446147FFD7B6D7F82CC194F80D" bold="true" box="[689,732,2004,2023]" pageId="4" pageNumber="500">
<taxonomicNameLabel id="A214173F6147FFD7B6D7F82CC194F80D" box="[689,732,2004,2023]" pageId="4" pageNumber="500" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, самец: 6 — габитус, голотип, вид сбоку; 7 — голова и грудь, голотип, вид спереди- сверху; 8 — гипопигий, вид сбоку; 9 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм.
</paragraph>
</caption>
<paragraph id="8BEC76566146FFD6B485FD89C23CFB39" blockId="5.[189,787,303,1717]" pageId="5" pageNumber="501">
Hypopygium (
<figureCitation id="13686AD36146FFD6B514FD89C283FD6C" box="[370,459,625,646]" captionStart="Figs 6–9" captionStartId="4.[190,228,1932,1951]" captionTargetBox="[216,1380,547,1855]" captionTargetId="figure-103@4.[200,1410,527,1877]" captionTargetPageId="4" captionText="Figs 6–9. Empis (Polyblepharis) mongolica Shamshev, sp.n., male: 6 — habitus, holotype, lateral view; 7 — head and thorax, holotype, anterodorsal view; 8 — hypopygium, lateral view; 9 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 6–9. Empis (Polyblepharis) mongolica Shamshev, sp.n., самец: 6 — габитус, голотип, вид сбоку; 7 — голова и грудь, голотип, вид спереди-сверху; 8 — гипопигий, вид сбоку; 9 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="5" pageNumber="501">Figs 8, 9</figureCitation>
) large; epandrium and basal half of phallus brown, apical half of phallus and cerci brownish yellow; epandrial lamella shiny on middle part, with short pale and black setae along lower margin and near apex, bearing spinule-like setae at tip; cercus covered with dense setulae dorsally and near apex laterally. Epandrium entirely broadly divided (epandrial bridge absent); epandrial lamella (
<figureCitation id="13686AD36146FFD6B6A9FCE9C042FCCC" box="[719,778,785,806]" captionStart="Figs 6–9" captionStartId="4.[190,228,1932,1951]" captionTargetBox="[216,1380,547,1855]" captionTargetId="figure-103@4.[200,1410,527,1877]" captionTargetPageId="4" captionText="Figs 6–9. Empis (Polyblepharis) mongolica Shamshev, sp.n., male: 6 — habitus, holotype, lateral view; 7 — head and thorax, holotype, anterodorsal view; 8 — hypopygium, lateral view; 9 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 6–9. Empis (Polyblepharis) mongolica Shamshev, sp.n., самец: 6 — габитус, голотип, вид сбоку; 7 — голова и грудь, голотип, вид спереди-сверху; 8 — гипопигий, вид сбоку; 9 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="5" pageNumber="501">Fig. 8</figureCitation>
) subtriangular, with slightly produced and bent inward apex (lateral view). Hypandrium separated from epandrium; mostly membranous, undivided, narrowly sclerotized along margin, subtriangular in ventral view; bare; gonocoxal apodeme small. Cerci separated with each other and from epandrium; cercus elongate ovate (lateral view), with apex extending to level of epandrial lamella apex, without dorsolateral, lobe-like projection in basal part (dorsal view), with serrate inner margin. Phallus (
<figureCitation id="13686AD36146FFD6B5F4FBFBC298FBF2" box="[402,464,1027,1048]" captionStart="Figs 6–9" captionStartId="4.[190,228,1932,1951]" captionTargetBox="[216,1380,547,1855]" captionTargetId="figure-103@4.[200,1410,527,1877]" captionTargetPageId="4" captionText="Figs 6–9. Empis (Polyblepharis) mongolica Shamshev, sp.n., male: 6 — habitus, holotype, lateral view; 7 — head and thorax, holotype, anterodorsal view; 8 — hypopygium, lateral view; 9 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 6–9. Empis (Polyblepharis) mongolica Shamshev, sp.n., самец: 6 — габитус, голотип, вид сбоку; 7 — голова и грудь, голотип, вид спереди-сверху; 8 — гипопигий, вид сбоку; 9 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="5" pageNumber="501">Fig. 9</figureCitation>
) almost entirely hidden; zigzag bent, strongly constricted beyond middle; its basal portion gently curved, thick at base and becoming slightly narrower towards constriction; apical bend portion almost circular (its tip pointing to rear), long, almost uniformly slender (except slightly thickened extreme base). Ejaculatory apodeme moderately large, extended far beyond basal curvature of phallus, with lateral wings.
</paragraph>
<paragraph id="8BEC76566146FFD6B485FB21C2D7FB04" blockId="5.[189,787,303,1717]" box="[227,415,1241,1262]" pageId="5" pageNumber="501">
<emphasis id="B927AA446146FFD6B485FB21C27BFB04" bold="true" box="[227,307,1241,1262]" pageId="5" pageNumber="501">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BEC76566146FFD6B485FB0CC1C9F98F" blockId="5.[189,787,303,1717]" pageId="5" pageNumber="501">
DIFFERENTIAL DIAGNOSIS. The new species is similar to
<taxonomicName id="4C530DD56146FFD6B493FAF6C2A6FAC9" authority="Chvala, 1999" authorityName="Chvala" authorityYear="1999" box="[245,494,1294,1315]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="501" phylum="Arthropoda" rank="species" species="indigirca">
<emphasis id="B927AA446146FFD6B493FAF6C24CFAC9" box="[245,260,1294,1315]" italics="true" pageId="5" pageNumber="501">E</emphasis>
.
<emphasis id="B927AA446146FFD6B576FAF6C22FFAC9" box="[272,359,1294,1315]" italics="true" pageId="5" pageNumber="501">indigirca</emphasis>
<bibRefCitation id="EFC20BA76146FFD6B50BFAF6C2A6FAC9" author="Chvala M." box="[365,494,1294,1315]" pageId="5" pageNumber="501" pagination="113 - 225" refId="ref15584" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999</bibRefCitation>
</taxonomicName>
(known from Eastern Siberia) and
<taxonomicName id="4C530DD56146FFD6B48EFAD1C212FAD4" box="[232,346,1321,1342]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="501" phylum="Arthropoda" rank="species" species="subcurta" status="sp.n.">
<emphasis id="B927AA446146FFD6B48EFAD1C3BFFAD4" box="[232,247,1321,1342]" italics="true" pageId="5" pageNumber="501">E</emphasis>
.
<emphasis id="B927AA446146FFD6B560FAD1C212FAD4" box="[262,346,1321,1342]" italics="true" pageId="5" pageNumber="501">subcurta</emphasis>
</taxonomicName>
<emphasis id="B927AA446146FFD6B505FAD1C2DBFAD4" bold="true" box="[355,403,1321,1342]" pageId="5" pageNumber="501">
<taxonomicNameLabel id="A214173F6146FFD6B505FAD1C2DBFAD4" box="[355,403,1321,1342]" pageId="5" pageNumber="501" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
described herein.
<taxonomicName id="4C530DD56146FFD6B62DFAD1C1AFFAD4" authorityName="Chvala" authorityYear="1999" box="[587,743,1321,1342]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="501" phylum="Arthropoda" rank="species" species="indigirca">
<emphasis id="B927AA446146FFD6B62DFAD1C1AFFAD4" box="[587,743,1321,1342]" italics="true" pageId="5" pageNumber="501">Empis indigirca</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C530DD56146FFD6B57CFABCC2D1FAB3" box="[282,409,1348,1369]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="501" phylum="Arthropoda" rank="species" species="mongolica" status="sp.n.">
<emphasis id="B927AA446146FFD6B57CFABCC261FAB3" box="[282,297,1348,1369]" italics="true" pageId="5" pageNumber="501">E</emphasis>
.
<emphasis id="B927AA446146FFD6B552FABCC2D1FAB3" box="[308,409,1348,1369]" italics="true" pageId="5" pageNumber="501">mongolica</emphasis>
</taxonomicName>
<emphasis id="B927AA446146FFD6B5F9FABCC287FAB3" bold="true" box="[415,463,1348,1369]" pageId="5" pageNumber="501">
<taxonomicNameLabel id="A214173F6146FFD6B5F9FABCC287FAB3" box="[415,463,1348,1369]" pageId="5" pageNumber="501" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
primarily by entirely black acrostichal and dorsocentral setae, faintly brownish infuscate wings, the presence of black posteromarginal setae on abdominal tergites 1–5 and by smaller male hypopygium [
<bibRefCitation id="EFC20BA76146FFD6B6E4FA6CC041FA43" author="Chvala M." box="[642,777,1428,1449]" pageId="5" pageNumber="501" pagination="113 - 225" refId="ref15584" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999</bibRefCitation>
].
<taxonomicName id="4C530DD56146FFD6B4DBFA57C21EFA2E" box="[189,342,1455,1476]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="501" phylum="Arthropoda" rank="species" species="subcurta" status="sp. nov.">
<emphasis id="B927AA446146FFD6B4DBFA57C21EFA2E" box="[189,342,1455,1476]" italics="true" pageId="5" pageNumber="501">Empis subcurta</emphasis>
</taxonomicName>
<emphasis id="B927AA446146FFD6B538FA57C2C6FA2E" bold="true" box="[350,398,1455,1476]" pageId="5" pageNumber="501">
<taxonomicNameLabel id="A214173F6146FFD6B538FA57C2C6FA2E" box="[350,398,1455,1476]" pageId="5" pageNumber="501" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from the new species primarily by entirely dark brown legs, entirely pale acrostichal and dorsocentral setae and smaller male hypopygium. In addition, in
<taxonomicName id="4C530DD56146FFD6B4DBF9F8C267F9FF" box="[189,303,1536,1557]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="501" phylum="Arthropoda" rank="species" species="subcurta" status="sp.n.">
<emphasis id="B927AA446146FFD6B4DBF9F8C384F9FF" box="[189,204,1536,1557]" italics="true" pageId="5" pageNumber="501">E</emphasis>
.
<emphasis id="B927AA446146FFD6B4BDF9F8C267F9FF" box="[219,303,1536,1557]" italics="true" pageId="5" pageNumber="501">subcurta</emphasis>
</taxonomicName>
<emphasis id="B927AA446146FFD6B55EF9F8C220F9FF" bold="true" box="[312,360,1536,1557]" pageId="5" pageNumber="501">
<taxonomicNameLabel id="A214173F6146FFD6B55EF9F8C220F9FF" box="[312,360,1536,1557]" pageId="5" pageNumber="501" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
strong mesonotal setae usually are yellow (except notopleurals) and abdominal tergites 1–5 are densely brownish pruinose dorsally and grey pruinose laterally (in dorsal view abdomen with broad brownish stripe).
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6146FFD6B485F993C239F971" pageId="5" pageNumber="501" type="etymology">
<paragraph id="8BEC76566146FFD6B485F993C239F971" blockId="5.[189,787,303,1717]" pageId="5" pageNumber="501">
ETYMOLOGY. The species is named after country of its origin (
<collectingCountry id="F34436C66146FFD6B565F97EC22EF971" box="[259,358,1670,1691]" name="Mongolia" pageId="5" pageNumber="501">Mongolia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6146FFD6B485F958C2B3F95F" box="[227,507,1696,1717]" pageId="5" pageNumber="501" type="distribution">
<paragraph id="8BEC76566146FFD6B485F958C2B3F95F" blockId="5.[189,787,303,1717]" box="[227,507,1696,1717]" pageId="5" pageNumber="501">
DISTRIBUTION.
<collectingCountry id="F34436C66146FFD6B5F1F958C2BFF95F" box="[407,503,1696,1717]" name="Mongolia" pageId="5" pageNumber="501">Mongolia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

279
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<mods:title id="37896CD6F9348D6FE12DAE123940D555">Seven new species of dance flies of the subgenus Polyblepharis Bezzi, 1909 (Diptera: Empididae) from Mongolia and East Siberia</mods:title>
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<taxonomicName id="4C530DD56142FFD2B4BAFC93C1FCFC68" authority="Shamshev" authorityName="Shamshev" box="[220,692,875,898]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="497" phylum="Arthropoda" rank="species" species="bylyrensis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA446142FFD2B4BAFC93C257FC68" box="[220,287,875,898]" italics="true" pageId="1" pageNumber="497">Empis</emphasis>
(
<emphasis id="B927AA446142FFD2B548FC93C28CFC68" box="[302,452,875,898]" italics="true" pageId="1" pageNumber="497">Polyblepharis</emphasis>
)
<emphasis id="B927AA446142FFD2B5B5FC93C176FC68" box="[467,574,875,898]" italics="true" pageId="1" pageNumber="497">bylyrensis</emphasis>
Shamshev
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<paragraph id="8BEC76566142FFD2B5D0FC70C151FC75" blockId="1.[220,755,875,927]" box="[438,537,904,927]" pageId="1" pageNumber="497">
<figureCitation id="13686AD36142FFD2B5D0FC70C15EFC75" box="[438,534,904,927]" captionStart="Figs 1–5" captionStartId="2.[189,227,1879,1898]" captionTargetBox="[216,1393,500,1807]" captionTargetId="figure-72@2.[200,1410,481,1827]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) bylyrensis Shamshev, sp.n.: 1–4 — male, holotype: 1 — hypopygium, lateral view; 2 — hypandrium, ventral view; 3 — cerci, dorsal view; 4 — phallus, lateral view; 5 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; goncx apod — gonocoxal apodeme; hypd — hypandrium; ph — phallus; ph pl — phallic plate; spm sac — sperm sac. Scale bar is 0.1 mm. Рис. 1–5. Empis (Polyblepharis) bylyrensis Shamshev, sp.n.: 1–4 — самец, голотип: 1 — гипопигий, вид сбоку; 2 — гипандрий, вид сниЗу; 3 — церки, вид сниЗу; 4 — фаллус, вид сбоку; 5 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; goncx apod — гонококсальнаЯ аподема; hypd — гипандрий; ph — фаллус; ph pl — пластина фаллуса; spm sac — сперматоЗоидный мешок. Масштаб 0,1 мм." pageId="1" pageNumber="497">Figs 1–5</figureCitation>
.
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, male, labelled [in Cyrillic, Russian]: Bylyra [
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,
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],
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[=Chitinskaya oblast, now Zabaykalskiy Territory] / step. skl. [=stepnoy sklon, steppe slope] i kolki [and small forest] /
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20 vi 975 (
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,
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_DIP_0000631; dissected).
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:
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, right shore of
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,
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,
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</materialsCitation>
.
<materialsCitation id="3B3B7C0B6142FFD2B662FB9AC1F6FB78" collectingDate="1975-06-17" collectionCode="ZISP, V" country="Russia" location="Kyra" municipality="Richter" pageId="1" pageNumber="497" specimenCount="2" specimenCount-female="2" stateProvince="Buryatia" typeStatus="paratype">
<collectingMunicipality id="6B88EC2C6142FFD2B662FB9AC103FB9D" box="[516,587,1122,1143]" pageId="1" pageNumber="497">Richter</collectingMunicipality>
(
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,
<collectionCode id="ED42EE936142FFD2B6C1FB9AC195FB9D" box="[679,733,1122,1143]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="1" pageNumber="497">ZISP</collectionCode>
). Zabaykalskiy Terr.:
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, forest-steppe,
<date id="FFED50966142FFD2B651FB85C1E8FB78" box="[567,672,1149,1170]" pageId="1" pageNumber="497" value="1975-06-17">
<collectingDate id="EFA9A97E6142FFD2B651FB85C1E8FB78" box="[567,672,1149,1170]" pageId="1" pageNumber="497" value="1975-06-17">17.vi.1975</collectingDate>
</date>
,
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</materialsCitation>
.
<materialsCitation id="3B3B7C0B6142FFD2B6AAFB85C27AFB46" collectionCode="ZISP" country="Russia" location="Richter" pageId="1" pageNumber="497" specimenCount="1" specimenCount-female="1" stateProvince="Buryatia" typeStatus="paratype">
<location id="8E8C208D6142FFD2B6AAFB85C05BFB78" LSID="urn:lsid:plazi:treatment:03FAC7406142FFD0B4BAFC93C3BBFBFE:8E8C208D6142FFD2B6AAFB85C05BFB78" box="[716,787,1149,1170]" country="Russia" name="Richter" pageId="1" pageNumber="497" stateProvince="Buryatia">Richter</location>
(
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,
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6142FFD2B485FB4AC27AFAA6" pageId="1" pageNumber="497" type="diagnosis">
<paragraph id="8BEC76566142FFD2B485FB4AC27AFAA6" blockId="1.[189,787,962,2076]" pageId="1" pageNumber="497">
DIAGNOSIS. Mid-sized species (body about
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); all femora bearing rows of anteroventral and posteroventral spines, all tibiae with row of anteroventral spinules; mesoscutum with 4 vittae. Male: eyes holoptic, almost touching on middle part of frons. Female: eyes dichoptic, separated by broad frons.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6142FFD0B485FAAAC10FFC43" lastPageId="3" lastPageNumber="499" pageId="1" pageNumber="497" type="description">
<paragraph id="8BEC76566142FFD2B485FAAAC190F8F8" blockId="1.[189,787,962,2076]" pageId="1" pageNumber="497">
DESCRIPTION. Body nearly
<quantity id="4CABDBB36142FFD2B677FAAAC129FA8D" box="[529,609,1362,1383]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="497" unit="mm" value="5.5">5.5 mm</quantity>
, wing
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.
<emphasis id="B927AA446142FFD2B4DBFA95C3B9FA68" bold="true" box="[189,241,1389,1410]" pageId="1" pageNumber="497">Male</emphasis>
. Head capsule regions mostly densely greyish pruinose; face on lower margin and entire clypeus shiny; head setation mostly black. Eyes almost touching on middle part of frons, upper ommatidia enlarged. Frons represented by small subtriangular spaces below ocellar triangle and above antennae; bare. Face broad, bare. Two short, fine ocellar setae. Occiput with numerous moderately long setae laterally; postocular setae very short (upper setae somewhat stronger); postgena with pale hair-like setae. Antenna with scape and pedicel brown, postpedicel and stylus black; scape slightly longer than subglobular pedicel, both with very short setulae; postpedicel nearly 2X as long as basal width, with gently concave subapical part of lower margin; stylus moderately long, slightly longer than postpedicel basal width. Proboscis with labrum brownish yellow, nearly 1.5X longer than eye height; palpus short, pale yellow; with scattered, dark and pale setulae.
</paragraph>
<paragraph id="8BEC76566142FFD2B485F8EFC037FE0C" blockId="1.[189,787,962,2076]" lastBlockId="1.[824,1422,303,2076]" pageId="1" pageNumber="497">Thorax black in ground-colour, densely greyish pruinose; mostly black setose; mesoscutum with 4 brown distinct vittae. Antepronotum with 4–5 black, strong setae dorsally and pale, hair-like setae laterally. Postpronotal lobe with 1 long and several short setae. Proepisternum with 5–6 black, strong and more numerous pale, hair-like setae on lower part. Prosternum bare. Mesonotal chaetotaxy reduced: acrostichal setae present in presutural part only, arranged in 2 close irregular rows, scattered, very short; dorsocentrals uniserial, very short (including prescutellars); 1 short presutural intra-alar, 1 similar presutural supra-alar, 3 long, strong notopleurals (sometimes 1 or 2 on one side), no postsutural supra-alar seta(e), 1 long and 1 minute postalars, 4 short scutellars (apical setae longer and stronger); notopleuron with scattered black and pale setulae anteriorly. Laterotergite with 3–4 strong, black and more numerous, hair-like, pale setae. Anterior and posterior spiracles yellow.</paragraph>
<paragraph id="8BEC76566142FFD2B738FE14C081FBB8" blockId="1.[824,1422,303,2076]" pageId="1" pageNumber="497">Legs with coxae densely greyish pruinose, remaining podomeres subshiny; mostly black setose. Leg colour: mostly yellow to dusky yellow; coxae, trochanters at apex, most part of fore femur (except extreme base and about apical third) and tarsomeres brownish (tarsomere 1 and basal part of tarsomere 2 sometimes paler). Fore coxa with scattered, strong, black setae and pale setulae anteriorly. All femora without whitish pilosity ventrally. All femora bearing rows of anteroventral and posteroventral spines: fore femur in about apical 1/2; mid femur in about apical 3/4; hind femur in about apical 2/3. In addition, all femora almost bare in lower half anteriorly; hind femur bearing 1 short anterior seta close to apex. Hind femur rather slender, only slightly broader than fore and mid femora. All tibiae lacking outstanding setae, only some anterodorsal and posterodorsal setulae slightly longer; fore tibia bearing row of anteroventral spinule-lake setae in about apical 1/2; mid and hind tibiae with almost complete row of similar anteroventral spinules (except close to base). Hind tibia rather slender; no seta in posteroapical comb. Hind basitarsus slightly thickened basally, becoming gently narrow towards apex (not broader than hind tibia at apex); remaining tarsomeres of all legs slender; fore and hind basitarsi with short, spinulae-like setae ventrally.</paragraph>
<paragraph id="8BEC76566142FFD2B738FBA0C73AFAB5" blockId="1.[824,1422,303,2076]" pageId="1" pageNumber="497">
Wing membrane hyaline; entirely covered with microtrichia; veins mostly brownish (brownish yellow near extreme base), well-sclerotized. Veins R
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and M
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slightly divergent towards wing margin; radial fork acute; R
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meeting costa before wing apex; CuA+CuP complete. Cell dm moderately large, with moderately elongate apex; apical portion of M
<subScript id="17D774136142FFD2B144FB15C661FB13" attach="left" box="[1314,1321,1261,1273]" fontSize="5" pageId="1" pageNumber="497">4</subScript>
nearly 2X longer than its middle portion. Pterostigma brownish yellow. Basal costal seta absent. Anal lobe well-developed; axillary incision very acute. Squama yellow, pale fringed. Halter with yellow stalk and brownish knob.
</paragraph>
<paragraph id="8BEC76566142FFD2B738FA9DC75BF949" blockId="1.[824,1422,303,2076]" pageId="1" pageNumber="497">Abdomen entirely brown in ground colour; tergites extensively shiny dorsally, narrowly densely greyish pruinose laterally; sternites densely greyish pruinose. Abdominal tergites covered with scattered minute setulae dorsally, tergites 1–3 bearing long, fine, pale setae laterally, with additional fine, brownish to black setae (including posteromarginals; also present on remaining tergites); sternites with scattered pale setulae. Pregenital segments: segments 6 and 7 unmodified; segment 8 with separated tergite and sternite; tergite 8 represented by two sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 somewhat enlarged, without prominent projections.</paragraph>
<paragraph id="8BEC76566142FFD1B738F951C72EFEB5" blockId="1.[824,1422,303,2076]" lastBlockId="2.[824,1421,303,404]" lastPageId="2" lastPageNumber="498" pageId="1" pageNumber="497">
Hypopygium (
<figureCitation id="13686AD36142FFD2B78CF951C708F954" box="[1002,1088,1705,1726]" captionStart="Figs 1–5" captionStartId="2.[189,227,1879,1898]" captionTargetBox="[216,1393,500,1807]" captionTargetId="figure-72@2.[200,1410,481,1827]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) bylyrensis Shamshev, sp.n.: 1–4 — male, holotype: 1 — hypopygium, lateral view; 2 — hypandrium, ventral view; 3 — cerci, dorsal view; 4 — phallus, lateral view; 5 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; goncx apod — gonocoxal apodeme; hypd — hypandrium; ph — phallus; ph pl — phallic plate; spm sac — sperm sac. Scale bar is 0.1 mm. Рис. 1–5. Empis (Polyblepharis) bylyrensis Shamshev, sp.n.: 1–4 — самец, голотип: 1 — гипопигий, вид сбоку; 2 — гипандрий, вид сниЗу; 3 — церки, вид сниЗу; 4 — фаллус, вид сбоку; 5 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; goncx apod — гонококсальнаЯ аподема; hypd — гипандрий; ph — фаллус; ph pl — пластина фаллуса; spm sac — сперматоЗоидный мешок. Масштаб 0,1 мм." pageId="1" pageNumber="497">Figs 1–4</figureCitation>
) moderately large; epandrium and cerci black; epandrial lamella with short, black, rather sparse setae in apical part; cercus covered with dark setulae. Epandrium entirely broadly divided (epandrial bridge absent); epandrial lamella (
<figureCitation id="13686AD36142FFD2B78AF8EDC76CF8C0" box="[1004,1060,1813,1834]" captionStart="Figs 1–5" captionStartId="2.[189,227,1879,1898]" captionTargetBox="[216,1393,500,1807]" captionTargetId="figure-72@2.[200,1410,481,1827]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) bylyrensis Shamshev, sp.n.: 1–4 — male, holotype: 1 — hypopygium, lateral view; 2 — hypandrium, ventral view; 3 — cerci, dorsal view; 4 — phallus, lateral view; 5 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; goncx apod — gonocoxal apodeme; hypd — hypandrium; ph — phallus; ph pl — phallic plate; spm sac — sperm sac. Scale bar is 0.1 mm. Рис. 1–5. Empis (Polyblepharis) bylyrensis Shamshev, sp.n.: 1–4 — самец, голотип: 1 — гипопигий, вид сбоку; 2 — гипандрий, вид сниЗу; 3 — церки, вид сниЗу; 4 — фаллус, вид сбоку; 5 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; goncx apod — гонококсальнаЯ аподема; hypd — гипандрий; ph — фаллус; ph pl — пластина фаллуса; spm sac — сперматоЗоидный мешок. Масштаб 0,1 мм." pageId="1" pageNumber="497">Fig. 1</figureCitation>
) rather subrectangular (lateral view), slightly narrower at apex. Hypandrium (
<figureCitation id="13686AD36142FFD2B0DEF8C8C7BAF8AF" box="[1208,1266,1840,1861]" captionStart="Figs 1–5" captionStartId="2.[189,227,1879,1898]" captionTargetBox="[216,1393,500,1807]" captionTargetId="figure-72@2.[200,1410,481,1827]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) bylyrensis Shamshev, sp.n.: 1–4 — male, holotype: 1 — hypopygium, lateral view; 2 — hypandrium, ventral view; 3 — cerci, dorsal view; 4 — phallus, lateral view; 5 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; goncx apod — gonocoxal apodeme; hypd — hypandrium; ph — phallus; ph pl — phallic plate; spm sac — sperm sac. Scale bar is 0.1 mm. Рис. 1–5. Empis (Polyblepharis) bylyrensis Shamshev, sp.n.: 1–4 — самец, голотип: 1 — гипопигий, вид сбоку; 2 — гипандрий, вид сниЗу; 3 — церки, вид сниЗу; 4 — фаллус, вид сбоку; 5 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; goncx apod — гонококсальнаЯ аподема; hypd — гипандрий; ph — фаллус; ph pl — пластина фаллуса; spm sac — сперматоЗоидный мешок. Масштаб 0,1 мм." pageId="1" pageNumber="497">Fig. 2</figureCitation>
) separated from epandrium; subtriangular in ventral view, with pointed apex, bare; gonocoxal apodeme small. Cerci (
<figureCitation id="13686AD36142FFD2B0A0F89EC669F891" box="[1222,1313,1894,1915]" captionStart="Figs 1–5" captionStartId="2.[189,227,1879,1898]" captionTargetBox="[216,1393,500,1807]" captionTargetId="figure-72@2.[200,1410,481,1827]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) bylyrensis Shamshev, sp.n.: 1–4 — male, holotype: 1 — hypopygium, lateral view; 2 — hypandrium, ventral view; 3 — cerci, dorsal view; 4 — phallus, lateral view; 5 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; goncx apod — gonocoxal apodeme; hypd — hypandrium; ph — phallus; ph pl — phallic plate; spm sac — sperm sac. Scale bar is 0.1 mm. Рис. 1–5. Empis (Polyblepharis) bylyrensis Shamshev, sp.n.: 1–4 — самец, голотип: 1 — гипопигий, вид сбоку; 2 — гипандрий, вид сниЗу; 3 — церки, вид сниЗу; 4 — фаллус, вид сбоку; 5 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; goncx apod — гонококсальнаЯ аподема; hypd — гипандрий; ph — фаллус; ph pl — пластина фаллуса; spm sac — сперматоЗоидный мешок. Масштаб 0,1 мм." pageId="1" pageNumber="497">Figs 1, 3</figureCitation>
) separated with each other and from epandrium; cercus elongate ovate (lateral view), with apex extending to level of epandrial lamella apex, without lobe-like, dorsolateral projection at basal part (dorsal view), with smoothed inner margin. Phallus (
<figureCitation id="13686AD36142FFD2B12FF829C6CEF80C" box="[1353,1414,2001,2022]" captionStart="Figs 1–5" captionStartId="2.[189,227,1879,1898]" captionTargetBox="[216,1393,500,1807]" captionTargetId="figure-72@2.[200,1410,481,1827]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) bylyrensis Shamshev, sp.n.: 1–4 — male, holotype: 1 — hypopygium, lateral view; 2 — hypandrium, ventral view; 3 — cerci, dorsal view; 4 — phallus, lateral view; 5 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; goncx apod — gonocoxal apodeme; hypd — hypandrium; ph — phallus; ph pl — phallic plate; spm sac — sperm sac. Scale bar is 0.1 mm. Рис. 1–5. Empis (Polyblepharis) bylyrensis Shamshev, sp.n.: 1–4 — самец, голотип: 1 — гипопигий, вид сбоку; 2 — гипандрий, вид сниЗу; 3 — церки, вид сниЗу; 4 — фаллус, вид сбоку; 5 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; goncx apod — гонококсальнаЯ аподема; hypd — гипандрий; ph — фаллус; ph pl — пластина фаллуса; spm sac — сперматоЗоидный мешок. Масштаб 0,1 мм." pageId="1" pageNumber="497">Fig. 4</figureCitation>
) almost entirely hidden; zigzag bent, not constricted beyond middle; almost uniformly thickened (ventral and dorsal tubercles present beyond middle) and gently arcuate between basal curvature and apical bend portion; apical bend portion semicircular (its tip pointing to rear), moderately long, mostly slender (except slightly thickened extreme base). Ejaculatory apodeme small, situated far beyond basal curvature of phallus, with lateral wings; sperm sac enlarged.
</paragraph>
<paragraph id="8BEC76566141FFD0B738FE9CC14BFD83" blockId="2.[824,1421,303,404]" lastBlockId="3.[189,787,303,1044]" lastPageId="3" lastPageNumber="499" pageId="2" pageNumber="498">
<emphasis id="B927AA446141FFD1B738FE9CC0E1FE93" bold="true" box="[862,937,356,377]" pageId="2" pageNumber="498">Female</emphasis>
(
<figureCitation id="13686AD36141FFD1B7D0FE9CC0BAFE93" box="[950,1010,356,377]" captionStart="Figs 1–5" captionStartId="2.[189,227,1879,1898]" captionTargetBox="[216,1393,500,1807]" captionTargetId="figure-72@2.[200,1410,481,1827]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) bylyrensis Shamshev, sp.n.: 1–4 — male, holotype: 1 — hypopygium, lateral view; 2 — hypandrium, ventral view; 3 — cerci, dorsal view; 4 — phallus, lateral view; 5 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; goncx apod — gonocoxal apodeme; hypd — hypandrium; ph — phallus; ph pl — phallic plate; spm sac — sperm sac. Scale bar is 0.1 mm. Рис. 1–5. Empis (Polyblepharis) bylyrensis Shamshev, sp.n.: 1–4 — самец, голотип: 1 — гипопигий, вид сбоку; 2 — гипандрий, вид сниЗу; 3 — церки, вид сниЗу; 4 — фаллус, вид сбоку; 5 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; goncx apod — гонококсальнаЯ аподема; hypd — гипандрий; ph — фаллус; ph pl — пластина фаллуса; spm sac — сперматоЗоидный мешок. Масштаб 0,1 мм." pageId="2" pageNumber="498">Fig. 5</figureCitation>
). Similar to male except as follows. Frons broad, with marginal setulae. Occiput and proepisternum with slightly shorter setae. Hind femur with row of short (slightly shorter than posteroventral spines), pennate setae just behind row of posteroventral spines in about apical half; hind tibia with fringe of short, slightly flattened posteroventral setae in about apical half. Abdomen covered with scattered, pale setulae (tergites 1–2 with longer setae laterally); mostly densely greyish pruinose; tergites 2–5 extensively shiny dorsally and pruinose laterally; in addition, tergite 2 pruinose narrowly anteriorly and tergite 5 broader posteriorly; tergite 6 mostly pruinose, narrowly shiny anteriorly (shiny space hidden by tergite 5); tergite 7 and segment 8 mostly shiny; cercus long, slender, covered with dark minute setulae.
</paragraph>
<caption id="DF2C26DE6141FFD1B4DBF8AFC10CF7F8" pageId="2" pageNumber="498" startId="2.[189,227,1879,1898]" targetBox="[216,1393,500,1807]" targetPageId="2" targetType="figure">
<paragraph id="8BEC76566141FFD1B4DBF8AFC2DAF858" blockId="2.[189,1421,1879,2066]" pageId="2" pageNumber="498">
<emphasis id="B927AA446141FFD1B4DBF8AFC246F880" bold="true" box="[189,270,1879,1898]" pageId="2" pageNumber="498">Figs 1–5.</emphasis>
<taxonomicName id="4C530DD56141FFD1B572F8AFC1D9F880" authority="Shamshev" authorityName="Shamshev" box="[276,657,1879,1898]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="498" phylum="Arthropoda" rank="species" species="bylyrensis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA446141FFD1B572F8AFC201F880" box="[276,329,1879,1898]" italics="true" pageId="2" pageNumber="498">Empis</emphasis>
(
<emphasis id="B927AA446141FFD1B531F8AFC287F880" box="[343,463,1879,1898]" italics="true" pageId="2" pageNumber="498">Polyblepharis</emphasis>
)
<emphasis id="B927AA446141FFD1B5BAF8AFC17AF880" box="[476,562,1879,1898]" italics="true" pageId="2" pageNumber="498">bylyrensis</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA446141FFD1B6FCF8AFC18CF880" bold="true" box="[666,708,1879,1898]" pageId="2" pageNumber="498">
<taxonomicNameLabel id="A214173F6141FFD1B6FCF8AFC18CF880" box="[666,708,1879,1898]" pageId="2" pageNumber="498" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 1–4 — male, holotype: 1 — hypopygium, lateral view; 2 — hypandrium, ventral view; 3 — cerci, dorsal view; 4 — phallus, lateral view; 5 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; goncx apod — gonocoxal apodeme; hypd — hypandrium; ph — phallus; ph pl — phallic plate; spm sac — sperm sac. Scale bar is 0.1 mm.
</paragraph>
<paragraph id="8BEC76566141FFD1B4DBF84FC10CF7F8" blockId="2.[189,1421,1879,2066]" pageId="2" pageNumber="498">
<emphasis id="B927AA446141FFD1B4DBF84FC247F820" bold="true" box="[189,271,1975,1994]" pageId="2" pageNumber="498">Рис. 1–5.</emphasis>
<taxonomicName id="4C530DD56141FFD1B573F84FC1C6F820" authority="Shamshev" authorityName="Shamshev" box="[277,654,1975,1994]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="498" phylum="Arthropoda" rank="species" species="bylyrensis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA446141FFD1B573F84FC202F820" box="[277,330,1975,1994]" italics="true" pageId="2" pageNumber="498">Empis</emphasis>
(
<emphasis id="B927AA446141FFD1B530F84FC286F820" box="[342,462,1975,1994]" italics="true" pageId="2" pageNumber="498">Polyblepharis</emphasis>
)
<emphasis id="B927AA446141FFD1B5BCF84FC178F820" box="[474,560,1975,1994]" italics="true" pageId="2" pageNumber="498">bylyrensis</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA446141FFD1B6F0F84FC189F820" bold="true" box="[662,705,1975,1994]" pageId="2" pageNumber="498">
<taxonomicNameLabel id="A214173F6141FFD1B6F0F84FC189F820" box="[662,705,1975,1994]" pageId="2" pageNumber="498" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 1–4 — самец, голотип: 1 — гипопигий, вид сбоку; 2 — гипандрий, вид сниЗу; 3 — церки, вид сниЗу; 4 — фаллус, вид сбоку; 5 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; goncx apod — гонококсальнаЯ аподема; hypd — гипандрий; ph — фаллус; ph pl — пластина фаллуса; spm sac — сперматоЗоидный мешок. Масштаб 0,1 мм.
</paragraph>
</caption>
<paragraph id="8BEC76566140FFD0B485FD97C10FFC43" blockId="3.[189,787,303,1044]" pageId="3" pageNumber="499">
DIFFERENTIAL DIAGNOSIS. The new species differs from all species assigned to
<taxonomicName id="4C530DD56140FFD0B580FD72C125FD75" authorityName="Bezzi" authorityYear="1909" box="[486,621,650,671]" pageId="3" pageNumber="499" rank="subGenus" subGenus="Polyblepharis">
<emphasis id="B927AA446140FFD0B580FD72C125FD75" box="[486,621,650,671]" italics="true" pageId="3" pageNumber="499">Polyblepharis</emphasis>
</taxonomicName>
by an armature of tibiae and femora of both sexes. In
<taxonomicName id="4C530DD56140FFD0B65AFD5CC1F1FD53" box="[572,697,676,697]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="499" phylum="Arthropoda" rank="species" species="bylyrensis" status="sp.n.">
<emphasis id="B927AA446140FFD0B65AFD5CC103FD53" box="[572,587,676,697]" italics="true" pageId="3" pageNumber="499">E</emphasis>
.
<emphasis id="B927AA446140FFD0B63FFD5CC1F1FD53" box="[601,697,676,697]" italics="true" pageId="3" pageNumber="499">bylyrensis</emphasis>
</taxonomicName>
<emphasis id="B927AA446140FFD0B6A4FD5CC1BAFD53" bold="true" box="[706,754,676,697]" pageId="3" pageNumber="499">
<taxonomicNameLabel id="A214173F6140FFD0B6A4FD5CC1BAFD53" box="[706,754,676,697]" pageId="3" pageNumber="499" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
all femora bear rows of anteroventral and posteroventral spines as well as all tibiae possess a row of anteroventral spinules. In addition, the abdominal sternite 8 of the male of the new species lacks prominent projections (usually present in species of
<taxonomicName id="4C530DD56140FFD0B4B1FCD2C216FCD5" authorityName="Bezzi" authorityYear="1909" box="[215,350,810,831]" pageId="3" pageNumber="499" rank="subGenus" subGenus="Polyblepharis">
<emphasis id="B927AA446140FFD0B4B1FCD2C216FCD5" box="[215,350,810,831]" italics="true" pageId="3" pageNumber="499">Polyblepharis</emphasis>
</taxonomicName>
). The shape of the phallus may suggest close relationships with
<taxonomicName id="4C530DD56140FFD0B50BFCBCC17CFCB3" box="[365,564,836,857]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="499" phylum="Arthropoda" rank="species" species="subtransbaicalica" status="sp.n.">
<emphasis id="B927AA446140FFD0B50BFCBCC234FCB3" box="[365,380,836,857]" italics="true" pageId="3" pageNumber="499">E</emphasis>
.
<emphasis id="B927AA446140FFD0B5EEFCBCC17CFCB3" box="[392,564,836,857]" italics="true" pageId="3" pageNumber="499">subtransbaicalica</emphasis>
</taxonomicName>
<emphasis id="B927AA446140FFD0B65FFCBCC122FCB3" bold="true" box="[569,618,836,857]" pageId="3" pageNumber="499">
<taxonomicNameLabel id="A214173F6140FFD0B65FFCBCC122FCB3" box="[569,618,836,857]" pageId="3" pageNumber="499" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
described herein. In addition to the characters noted above,
<taxonomicName id="4C530DD56140FFD0B63BFCA7C192FC9E" box="[605,730,863,884]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="499" phylum="Arthropoda" rank="species" species="bylyrensis" status="sp.n.">
<emphasis id="B927AA446140FFD0B63BFCA7C124FC9E" box="[605,620,863,884]" italics="true" pageId="3" pageNumber="499">E</emphasis>
.
<emphasis id="B927AA446140FFD0B61CFCA7C192FC9E" box="[634,730,863,884]" italics="true" pageId="3" pageNumber="499">bylyrensis</emphasis>
</taxonomicName>
<emphasis id="B927AA446140FFD0B684FCA7C05BFC9E" bold="true" box="[738,787,863,884]" pageId="3" pageNumber="499">
<taxonomicNameLabel id="A214173F6140FFD0B684FCA7C05BFC9E" box="[738,787,863,884]" pageId="3" pageNumber="499" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C530DD56140FFD0B55EFC82C148FC65" box="[312,512,890,911]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="499" phylum="Arthropoda" rank="species" species="subtransbaicalica" status="sp.n.">
<emphasis id="B927AA446140FFD0B55EFC82C20FFC65" box="[312,327,890,911]" italics="true" pageId="3" pageNumber="499">E</emphasis>
.
<emphasis id="B927AA446140FFD0B532FC82C148FC65" box="[340,512,890,911]" italics="true" pageId="3" pageNumber="499">subtransbaicalica</emphasis>
</taxonomicName>
<emphasis id="B927AA446140FFD0B661FC82C170FC65" bold="true" box="[519,568,890,911]" pageId="3" pageNumber="499">
<taxonomicNameLabel id="A214173F6140FFD0B661FC82C170FC65" box="[519,568,890,911]" pageId="3" pageNumber="499" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
by brown knob of the halter and by very short mesonotal setae.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6140FFD0B485FC57C2E0FC35" pageId="3" pageNumber="499" type="etymology">
<paragraph id="8BEC76566140FFD0B485FC57C2E0FC35" blockId="3.[189,787,303,1044]" pageId="3" pageNumber="499">
ETYMOLOGY. The epithet refers to the
<typeStatus id="54E8C8F46140FFD0B61AFC57C1EDFC2E" box="[636,677,943,964]" pageId="3" pageNumber="499">type</typeStatus>
locality of the new species, Bylyra.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6140FFD0B485FC1CC3BBFBFE" pageId="3" pageNumber="499" type="distribution">
<paragraph id="8BEC76566140FFD0B485FC1CC3BBFBFE" blockId="3.[189,787,303,1044]" pageId="3" pageNumber="499">
DISTRIBUTION.
<collectingCountry id="F34436C66140FFD0B5FDFC1CC293FC13" box="[411,475,996,1017]" name="Russia" pageId="3" pageNumber="499">Russia</collectingCountry>
(
<collectingRegion id="4997B8B46140FFD0B58AFC1CC10AFC13" box="[492,578,996,1017]" country="Russia" name="Buryatiya" pageId="3" pageNumber="499">Buryatia</collectingRegion>
, Zabaykalskiy Territory).
</paragraph>
</subSubSection>
</treatment>
</document>

473
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<document id="E7FED370980B0725B5016DD345A2AEFD" ID-DOI="10.15298/rusentj.33.4.13" ID-ISSN="0132-8069" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737172459620" checkinUser="felipe" docAuthor="Shamshev, I. V." docDate="2024" docId="03FAC7406144FFDAB4BDF8F7C7FCFC46" docLanguage="en" docName="RussEntomolJ.33.4.496-512.pdf" docOrigin="Russian Entomological Journal 33 (3)" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.4:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="4" docTitle="Empis (Polyblepharis) subdepilis Shamshev, sp.n." docType="treatment" docVersion="1" lastPageNumber="505" masterDocId="FFC3BF386143FFD3B466FFF8C348FFEA" masterDocTitle="Seven new species of dance flies of the subgenus Polyblepharis Bezzi, 1909 (Diptera: Empididae) from Mongolia and East Siberia" masterLastPageNumber="512" masterPageNumber="496" pageNumber="503" updateTime="1737594035090" updateUser="felipe" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9C5640794F76D9B7121EBB6D0D5AF363">Seven new species of dance flies of the subgenus Polyblepharis Bezzi, 1909 (Diptera: Empididae) from Mongolia and East Siberia</mods:title>
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<mods:namePart id="9DE13B25FFC2892B9758F849A9D38005">Shamshev, I. V.</mods:namePart>
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<mods:title id="5FCBA41D2C0D0B8A85F283EAC9525469">Russian Entomological Journal</mods:title>
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<mods:date id="316B7DE324E1BF4D0DFC2FC5705A6F6C">2024</mods:date>
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<mods:identifier id="72716237DA5B40EE64536EFA70C25909" type="ISSN">0132-8069</mods:identifier>
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<treatment id="03FAC7406144FFDAB4BDF8F7C7FCFC46" LSID="urn:lsid:plazi:treatment:03FAC7406144FFDAB4BDF8F7C7FCFC46" httpUri="http://treatment.plazi.org/id/03FAC7406144FFDAB4BDF8F7C7FCFC46" lastPageId="9" lastPageNumber="505" pageId="7" pageNumber="503">
<subSubSection id="C34925DD6144FFD4B4BDF8F7C1BCF8CC" box="[219,756,1807,1830]" pageId="7" pageNumber="503" type="nomenclature">
<paragraph id="8BEC76566144FFD4B4BDF8F7C1BCF8CC" blockId="7.[219,756,1807,1860]" box="[219,756,1807,1830]" pageId="7" pageNumber="503">
<heading id="D0A4C13A6144FFD4B4BDF8F7C1BCF8CC" box="[219,756,1807,1830]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="503" reason="8">
<taxonomicName id="4C530DD56144FFD4B4BDF8F7C1FCF8CC" authority="Shamshev" authorityName="Shamshev" box="[219,692,1807,1830]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="503" phylum="Arthropoda" rank="species" species="subdepilis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA446144FFD4B4BDF8F7C256F8CC" box="[219,286,1807,1830]" italics="true" pageId="7" pageNumber="503">Empis</emphasis>
(
<emphasis id="B927AA446144FFD4B54BF8F7C28BF8CC" box="[301,451,1807,1830]" italics="true" pageId="7" pageNumber="503">Polyblepharis</emphasis>
)
<emphasis id="B927AA446144FFD4B5B4F8F7C176F8CC" box="[466,574,1807,1830]" italics="true" pageId="7" pageNumber="503">subdepilis</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA446144FFD4B6D8F8F7C1BCF8CC" bold="true" box="[702,756,1807,1830]" pageId="7" pageNumber="503">
<taxonomicNameLabel id="A214173F6144FFD4B6D8F8F7C1BCF8CC" box="[702,756,1807,1830]" pageId="7" pageNumber="503" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6144FFD4B5CEF8D5C16FF8AE" box="[424,551,1837,1860]" pageId="7" pageNumber="503" type="description">
<paragraph id="8BEC76566144FFD4B5CEF8D5C16FF8AE" blockId="7.[219,756,1807,1860]" box="[424,551,1837,1860]" pageId="7" pageNumber="503">
<figureCitation id="13686AD36144FFD4B5CEF8D5C16AF8AE" box="[424,546,1837,1860]" captionStart="Figs 13–16" captionStartId="8.[189,227,1937,1956]" captionTargetBox="[215,1392,649,1860]" captionTargetId="figure-184@8.[200,1410,619,1887]" captionTargetPageId="8" captionText="Figs 13–16. Empis (Polyblepharis) subdepilis Shamshev, sp.n.: 13–15 — male: 13 — habitus, holotype, lateral view; 14 — hypopygium, lateral view; 15 — phallus, lateral view; 16 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 13–16. Empis (Polyblepharis) subdepilis Shamshev, sp.n.: 13–15 — самец: 13 — габитус, голотип, вид сбоку; 14 — гипопигий, вид сбоку; 15 — фаллус, вид сбоку; 16 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="7" pageNumber="503">Figs 13–16</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6144FFD4B485F89FC79AFD19" pageId="7" pageNumber="503" type="materials_examined">
<paragraph id="8BEC76566144FFD4B485F89FC1F6F80D" blockId="7.[189,786,1895,2076]" pageId="7" pageNumber="503">
<materialsCitation id="3B3B7C0B6144FFD4B485F89FC1F6F80D" collectionCode="TYPE, MATERIAL, ZISP, INS" country="Russia" latitude="51.466667" location="Zinovjev" longLatPrecision="1090" longitude="127.583336" municipality="Klimoutsy" pageId="7" pageNumber="503" specimenCount="1" specimenCount-male="1" stateProvince="Amur" typeStatus="holotype">
<collectionCode id="ED42EE936144FFD4B485F89FC2E0F896" box="[227,424,1895,1916]" pageId="7" pageNumber="503">TYPE MATERIAL</collectionCode>
.
<typeStatus id="54E8C8F46144FFD4B5D1F89FC15BF896" box="[439,531,1895,1916]" pageId="7" pageNumber="503" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D55BDDF6144FFD4B645F89FC17DF896" box="[547,565,1895,1916]" count="1" pageId="7" pageNumber="503" type="male">♂</specimenCount>
, labelled: [printed in Cyrillic, Russian]: [
<collectingCountry id="F34436C66144FFD4B518F879C290F87C" box="[382,472,1921,1942]" name="Russia" pageId="7" pageNumber="503">RUSSIA</collectingCountry>
, Amurskaya Province]
<collectingMunicipality id="6B88EC2C6144FFD4B6A7F879C391F85B" pageId="7" pageNumber="503">Klimoutsy</collectingMunicipality>
[
<geoCoordinate id="EE6710916144FFD4B481F864C270F85B" box="[231,312,1948,1969]" degrees="51" direction="north" minutes="28" orientation="latitude" pageId="7" pageNumber="503" precision="925" value="51.466667">51°28′N</geoCoordinate>
<geoCoordinate id="EE6710916144FFD4B526F864C2D4F85B" box="[320,412,1948,1969]" degrees="127" direction="east" minutes="35" orientation="longitude" pageId="7" pageNumber="503" precision="925" value="127.583336">127°35′E</geoCoordinate>
]
<collectingRegion id="4997B8B46144FFD4B5CEF864C146F85B" box="[424,526,1948,1969]" country="Russia" name="Amur" pageId="7" pageNumber="503">Amur obl.</collectingRegion>
[= Amurskaya Province] /
<locationDeviation id="9A5E6F746144FFD4B4DBF84FC2D0F826" box="[189,408,1975,1996]" location="Svobodny" pageId="7" pageNumber="503">
<quantity id="4CABDBB36144FFD4B4DBF84FC3B2F826" box="[189,250,1975,1996]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="503" unit="km" value="40.0">40 km</quantity>
W of Svobodny
</locationDeviation>
/
<location id="8E8C208D6144FFD4B5CAF84FC14BF826" LSID="urn:lsid:plazi:treatment:03FAC7406144FFDAB4BDF8F7C7FCFC46:8E8C208D6144FFD4B5CAF84FC14BF826" box="[428,515,1975,1996]" country="Russia" latitude="51.466667" longLatPrecision="1090" longitude="127.583336" municipality="Klimoutsy" name="Zinovjev" pageId="7" pageNumber="503" stateProvince="Amur">Zinovjev</location>
6.vi.[1]958 // oak-larch forest / on flowers of
<taxonomicName id="4C530DD56144FFD4B509F82AC2E5F80D" box="[367,429,2002,2023]" class="Magnoliopsida" family="Rosaceae" genus="Spirea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="503" phylum="Tracheophyta" rank="genus">
<emphasis id="B927AA446144FFD4B509F82AC2E5F80D" box="[367,429,2002,2023]" italics="true" pageId="7" pageNumber="503">Spirea</emphasis>
</taxonomicName>
(
<collectionCode id="ED42EE936144FFD4B5DFF82AC2A4F80D" box="[441,492,2002,2023]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="7" pageNumber="503">ZISP</collectionCode>
,
<collectionCode id="ED42EE936144FFD4B593F82AC153F80D" box="[501,539,2002,2023]" pageId="7" pageNumber="503">INS</collectionCode>
_DIP_0000630).
</materialsCitation>
</paragraph>
<paragraph id="8BEC76566144FFD4B485F815C05AF7F6" blockId="7.[189,786,1895,2076]" pageId="7" pageNumber="503">
<materialsCitation id="3B3B7C0B6144FFD4B485F815C046F7F6" country="Mongolia" location="Khubsugulskiy" pageId="7" pageNumber="503" specimenCount="1" stateProvince="Hovsgol" typeStatus="paratype">
<typeStatus id="54E8C8F46144FFD4B485F815C20BF7E8" box="[227,323,2029,2050]" pageId="7" pageNumber="503" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F34436C66144FFD4B531F815C295F7E8" box="[343,477,2029,2050]" name="Mongolia" pageId="7" pageNumber="503">MONGOLIA</collectingCountry>
.
<collectingRegion id="4997B8B46144FFD4B597F815C106F7E8" box="[497,590,2029,2050]" country="Mongolia" name="Hovsgol" pageId="7" pageNumber="503">Khövsgöl</collectingRegion>
(
<collectingRegion id="4997B8B46144FFD4B600F815C1F2F7E8" box="[614,698,2029,2050]" country="Mongolia" name="Hovsgol" pageId="7" pageNumber="503">Hovsgol</collectingRegion>
) aymag [=
<location id="8E8C208D6144FFD4B4A9F7FFC22CF7F6" LSID="urn:lsid:plazi:treatment:03FAC7406144FFDAB4BDF8F7C7FCFC46:8E8C208D6144FFD4B4A9F7FFC22CF7F6" box="[207,356,2055,2076]" country="Mongolia" name="Khubsugulskiy" pageId="7" pageNumber="503" stateProvince="Hovsgol">Khubsugulskiy</location>
aymak (province)]:
<locationDeviation id="9A5E6F746144FFD4B649F7FFC046F7F6" box="[559,782,2055,2076]" location="Muren" pageId="7" pageNumber="503">
<quantity id="4CABDBB36144FFD4B649F7FFC127F7F6" box="[559,623,2055,2076]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="503" unit="km" value="25.0">25 km</quantity>
SSW of Muren
</locationDeviation>
</materialsCitation>
,
</paragraph>
<paragraph id="8BEC76566144FFD4B75EFED7C79AFD19" blockId="7.[824,1422,303,2076]" pageId="7" pageNumber="503">
<materialsCitation id="3B3B7C0B6144FFD4B75EFED7C0CFFE90" collectingDate="1975-07-23" collectionCode="ZISP" collectorName="Nartshuk" elevation="1566" latitude="50.66022" location="Hohoo Gol" longLatPrecision="1" longitude="100.73886" municipality="Chandmani-Ondor Soum" pageId="7" pageNumber="503" specimenCount="1" specimenCount-male="1">
<date id="FFED50966144FFD4B75EFED7C0E0FEAE" box="[824,936,303,324]" pageId="7" pageNumber="503" value="1975-07-23">
<collectingDate id="EFA9A97E6144FFD4B75EFED7C0E0FEAE" box="[824,936,303,324]" pageId="7" pageNumber="503" value="1975-07-23">23.vii.1975</collectingDate>
</date>
,
<collectorName id="26A613806144FFD4B7D5FED7C743FEAE" box="[947,1035,303,324]" pageId="7" pageNumber="503">Nartshuk</collectorName>
(
<specimenCount id="9D55BDDF6144FFD4B07BFED7C70AFEAE" box="[1053,1090,303,324]" count="1" pageId="7" pageNumber="503" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED42EE936144FFD4B036FED7C7CDFEAE" box="[1104,1157,303,324]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="7" pageNumber="503">ZISP</collectionCode>
);
<collectingMunicipality id="6B88EC2C6144FFD4B0FFFED7C6C5FEAE" box="[1177,1421,303,324]" pageId="7" pageNumber="503">Chandmani-Ondor Soum</collectingMunicipality>
// unnamed trib. of
<location id="8E8C208D6144FFD4B797FEB2C716FEB5" LSID="urn:lsid:plazi:treatment:03FAC7406144FFDAB4BDF8F7C7FCFC46:8E8C208D6144FFD4B797FEB2C716FEB5" box="[1009,1118,330,351]" latitude="50.66022" longLatPrecision="1" longitude="100.73886" municipality="Chandmani-Ondor Soum" name="Hohoo Gol" pageId="7" pageNumber="503">Hohoo Gol</location>
,
<geoCoordinate id="EE6710916144FFD4B00EFEB2C79EFEB5" box="[1128,1238,330,351]" degrees="50.66022" direction="north" orientation="latitude" pageId="7" pageNumber="503" precision="1" value="50.66022">N50.66022</geoCoordinate>
,
<geoCoordinate id="EE6710916144FFD4B086FEB2C61EFEB5" box="[1248,1366,330,351]" degrees="100.73886" direction="east" orientation="longitude" pageId="7" pageNumber="503" precision="1" value="100.73886">E100.73886</geoCoordinate>
, elev.
<quantity id="4CABDBB36144FFD4B75EFE9DC0CFFE90" box="[824,903,357,378]" metricMagnitude="3" metricUnit="m" metricValue="1.566" pageId="7" pageNumber="503" unit="m" value="1566.0">
<elevation id="007E91656144FFD4B75EFE9DC0CFFE90" box="[824,903,357,378]" metricMagnitude="3" metricUnit="m" metricValue="1.566" pageId="7" pageNumber="503" unit="m" value="1566.0">1566 m</elevation>
</quantity>
</materialsCitation>
,
<materialsCitation id="3B3B7C0B6144FFD4B7F0FE9DC082FE21" collectingDate="2005-07-18" collectionCode="ANSP, CNC" collectorName="J. Gelhaus &amp; Selenge Proj." county="Tov" location="Bogdo-ula Mt" municipality="Zaysan" pageId="7" pageNumber="503" specimenCount="5" specimenCount-female="2" specimenCount-male="3">
<date id="FFED50966144FFD4B7F0FE9DC74DFE90" box="[918,1029,357,378]" pageId="7" pageNumber="503" value="2005-07-18">
<collectingDate id="EFA9A97E6144FFD4B7F0FE9DC74DFE90" box="[918,1029,357,378]" pageId="7" pageNumber="503" value="2005-07-18">18.vii.2005</collectingDate>
</date>
, coll.
<collectorName id="26A613806144FFD4B02EFE9DC7F9FE90" box="[1096,1201,357,378]" pageId="7" pageNumber="503">J. Gelhaus</collectorName>
#1022;
<collectorName id="26A613806144FFD4B16EFE9DC6C6FE90" box="[1288,1422,357,378]" pageId="7" pageNumber="503">Selenge Proj.</collectorName>
SRP#05071802 (
<specimenCount id="9D55BDDF6144FFD4B7B8FE78C75EFE7F" box="[990,1046,384,405]" count="3" pageId="7" pageNumber="503" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D55BDDF6144FFD4B042FE78C714FE7F" box="[1060,1116,384,405]" count="2" pageId="7" pageNumber="503" type="female">2 ♀♀</specimenCount>
,
<collectionCode id="ED42EE936144FFD4B00FFE78C7EDFE7F" box="[1129,1189,384,405]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="7" pageNumber="503">ANSP</collectionCode>
and
<collectionCode id="ED42EE936144FFD4B0B1FE78C645FE7F" box="[1239,1293,384,405]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="7" pageNumber="503">CNC</collectionCode>
).
<collectingCounty id="628D0EDA6144FFD4B178FE78C60DFE7F" box="[1310,1349,384,405]" pageId="7" pageNumber="503">Töv</collectingCounty>
aymag [=Tsentralny aymak (province)]:
<collectingMunicipality id="6B88EC2C6144FFD4B0E1FE63C787FE5A" box="[1159,1231,411,432]" pageId="7" pageNumber="503">Zaysan</collectingMunicipality>
, southern slope of
<location id="8E8C208D6144FFD4B75EFE4EC082FE21" LSID="urn:lsid:plazi:treatment:03FAC7406144FFDAB4BDF8F7C7FCFC46:8E8C208D6144FFD4B75EFE4EC082FE21" box="[824,970,438,459]" county="Tov" municipality="Zaysan" name="Bogdo-ula Mt" pageId="7" pageNumber="503">Bogdo-ula Mt</location>
</materialsCitation>
.,
<materialsCitation id="3B3B7C0B6144FFD4B7BAFE4EC6C2FE0C" collectingDate="1967-07-04" collectionCode="ZISP" collectorName="Zaitzev" country="Russia" county="Tov" location="Amurskaya Prov." municipality="Zaysan" pageId="7" pageNumber="503" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<date id="FFED50966144FFD4B7BAFE4EC708FE21" box="[988,1088,438,459]" pageId="7" pageNumber="503" value="1967-07-04">
<collectingDate id="EFA9A97E6144FFD4B7BAFE4EC708FE21" box="[988,1088,438,459]" pageId="7" pageNumber="503" value="1967-07-04">4.vii.1967</collectingDate>
</date>
,
<collectorName id="26A613806144FFD4B029FE4EC7DFFE21" box="[1103,1175,438,459]" pageId="7" pageNumber="503">Zaitzev</collectorName>
(
<specimenCount id="9D55BDDF6144FFD4B0CBFE4EC79DFE21" box="[1197,1237,438,459]" count="1" pageId="7" pageNumber="503" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED42EE936144FFD4B080FE4EC654FE21" box="[1254,1308,438,459]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="7" pageNumber="503">ZISP</collectionCode>
).
<collectingCountry id="F34436C66144FFD4B157FE4EC6C2FE21" box="[1329,1418,438,459]" name="Russia" pageId="7" pageNumber="503">RUSSIA</collectingCountry>
.
<location id="8E8C208D6144FFD4B75EFE29C0A0FE0C" LSID="urn:lsid:plazi:treatment:03FAC7406144FFDAB4BDF8F7C7FCFC46:8E8C208D6144FFD4B75EFE29C0A0FE0C" box="[824,1000,465,486]" country="Russia" name="Amurskaya Prov." pageId="7" pageNumber="503">Amurskaya Prov.</location>
: same locality and collector as
<typeStatus id="54E8C8F46144FFD4B154FE29C6C2FE0C" box="[1330,1418,465,486]" pageId="7" pageNumber="503" type="holotype">holotype</typeStatus>
</materialsCitation>
:
<date id="FFED50966144FFD4B75EFE14C0DFFDEB" box="[824,919,492,513]" pageId="7" pageNumber="503" value="1957-05-25">25.v.1957</date>
(
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);
<date id="FFED50966144FFD4B784FE14C708FDEB" box="[994,1088,492,513]" pageId="7" pageNumber="503" value="1957-05-30">30.v.1957</date>
(
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);
<date id="FFED50966144FFD4B0EAFE14C7AEFDEB" box="[1164,1254,492,513]" pageId="7" pageNumber="503" value="1957-06-01">1.vi.1957</date>
(
<specimenCount id="9D55BDDF6144FFD4B09EFE14C654FDEB" box="[1272,1308,492,513]" count="1" pageId="7" pageNumber="503" type="male">1 ♂</specimenCount>
);
<date id="FFED50966144FFD4B154FE14C6C4FDEB" box="[1330,1420,492,513]" pageId="7" pageNumber="503" value="1957-06-04">4.vi.1957</date>
(
<specimenCount id="9D55BDDF6144FFD4B727FDFFC02CFDF6" box="[833,868,519,540]" count="1" pageId="7" pageNumber="503" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D55BDDF6144FFD4B716FDFFC0EEFDF6" box="[880,934,519,540]" count="2" pageId="7" pageNumber="503" type="female">2 ♀♀</specimenCount>
);
<date id="FFED50966144FFD4B7DDFDFFC751FDF6" box="[955,1049,519,540]" pageId="7" pageNumber="503" value="1958-05-22">22.v.1958</date>
(
<specimenCount id="9D55BDDF6144FFD4B04FFDFFC703FDF6" box="[1065,1099,519,540]" count="1" pageId="7" pageNumber="503" type="female">1 ♀</specimenCount>
);
<date id="FFED50966144FFD4B006FDFFC7F3FDF6" box="[1120,1211,519,540]" pageId="7" pageNumber="503" value="1958-06-01">1.vi.1958</date>
(
<specimenCount id="9D55BDDF6144FFD4B0ADFDFFC7A5FDF6" box="[1227,1261,519,540]" count="1" pageId="7" pageNumber="503" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D55BDDF6144FFD4B09FFDFFC655FDF6" box="[1273,1309,519,540]" count="1" pageId="7" pageNumber="503" type="female">1 ♀</specimenCount>
);
<date id="FFED50966144FFD4B154FDFFC6C5FDF6" box="[1330,1421,519,540]" pageId="7" pageNumber="503" value="1958-06-06">6.vi.1958</date>
(
<specimenCount id="9D55BDDF6144FFD4B759FDDAC031FDDD" box="[831,889,546,567]" count="2" pageId="7" pageNumber="503" type="male">2 ♂♂</specimenCount>
(one dissected),
<specimenCount id="9D55BDDF6144FFD4B043FDDAC716FDDD" box="[1061,1118,546,567]" count="4" pageId="7" pageNumber="503" type="female">4 ♀♀</specimenCount>
);
<date id="FFED50966144FFD4B010FDDAC799FDDD" box="[1142,1233,546,567]" pageId="7" pageNumber="503" value="1958-06-08">8.vi.1958</date>
(
<specimenCount id="9D55BDDF6144FFD4B087FDDAC652FDDD" box="[1249,1306,546,567]" count="3" pageId="7" pageNumber="503" type="female">3 ♀♀</specimenCount>
); 14.vi.195 (
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);
<date id="FFED50966144FFD4B71CFDC5C0A8FDB8" box="[890,992,573,594]" pageId="7" pageNumber="503" value="1958-06-31">31.vi.1958</date>
(
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);
<date id="FFED50966144FFD4B04FFDC5C7CCFDB8" box="[1065,1156,573,594]" pageId="7" pageNumber="503" value="1959-06-02">2.vi.1959</date>
(
<specimenCount id="9D55BDDF6144FFD4B0F4FDC5C781FDB8" box="[1170,1225,573,594]" count="3" pageId="7" pageNumber="503" type="female">3 ♀♀</specimenCount>
);
<date id="FFED50966144FFD4B0B8FDC5C671FDB8" box="[1246,1337,573,594]" pageId="7" pageNumber="503" value="1959-06-04">4.vi.1959</date>
(
<specimenCount id="9D55BDDF6144FFD4B121FDC5C636FDB8" box="[1351,1406,573,594]" count="2" pageId="7" pageNumber="503" type="male">2 ♂♂</specimenCount>
);
<date id="FFED50966144FFD4B75EFDA0C0DFFD87" box="[824,919,600,621]" pageId="7" pageNumber="503" value="1959-05-31">31.v.1959</date>
(
<specimenCount id="9D55BDDF6144FFD4B7C5FDA0C092FD87" box="[931,986,600,621]" count="2" pageId="7" pageNumber="503" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D55BDDF6144FFD4B780FDA0C753FD87" box="[998,1051,600,621]" count="3" pageId="7" pageNumber="503" type="female">3 ♀♀</specimenCount>
);
<date id="FFED50966144FFD4B056FDA0C7C3FD87" box="[1072,1163,600,621]" pageId="7" pageNumber="503" value="1959-06-04">4.vi.1959</date>
(
<specimenCount id="9D55BDDF6144FFD4B0FCFDA0C7F4FD87" box="[1178,1212,600,621]" count="1" pageId="7" pageNumber="503" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D55BDDF6144FFD4B0AEFDA0C7A4FD87" box="[1224,1260,600,621]" count="1" pageId="7" pageNumber="503" type="female">1 ♀</specimenCount>
);
<materialsCitation id="3B3B7C0B6144FFD4B166FDA0C730FD62" country="Russia" location="Bolshoy Ergel" municipality="Malaya Pera" pageId="7" pageNumber="503" specimenCount="1" typeStatus="holotype">
between rivers
<collectingMunicipality id="6B88EC2C6144FFD4B75EFD8BC0FCFD62" box="[824,948,627,648]" pageId="7" pageNumber="503">Malaya Pera</collectingMunicipality>
and
<location id="8E8C208D6144FFD4B78FFD8BC730FD62" LSID="urn:lsid:plazi:treatment:03FAC7406144FFDAB4BDF8F7C7FCFC46:8E8C208D6144FFD4B78FFD8BC730FD62" box="[1001,1144,627,648]" country="Russia" municipality="Malaya Pera" name="Bolshoy Ergel" pageId="7" pageNumber="503">Bolshoy Ergel</location>
</materialsCitation>
,
<date id="FFED50966144FFD4B0E3FD8BC7AAFD62" box="[1157,1250,627,648]" pageId="7" pageNumber="503" value="1958-06-06">6.vi.1958</date>
, Zinovjev (
<specimenCount id="9D55BDDF6144FFD4B13EFD8BC636FD62" box="[1368,1406,627,648]" count="1" pageId="7" pageNumber="503" type="female">1 ♀</specimenCount>
);
<materialsCitation id="3B3B7C0B6144FFD4B75EFD76C0F1FD49" box="[824,953,654,675]" country="Russia" location="Bolshoy Ergel" municipality="Malaya Pera" pageId="7" pageNumber="503" specimenCount="1" typeStatus="holotype">same locality</materialsCitation>
,
<date id="FFED50966144FFD4B7A5FD76C768FD49" box="[963,1056,654,675]" pageId="7" pageNumber="503" value="1958-06-09">9.vi.1958</date>
, Zinovjev (
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).
<materialsCitation id="3B3B7C0B6144FFD4B0ACFD76C72DFD54" country="Russia" location="Khamsura village" pageId="7" pageNumber="503" specimenCount="1" stateProvince="Buryatiya" typeStatus="holotype">
<collectingRegion id="4997B8B46144FFD4B0ACFD76C668FD49" box="[1226,1312,654,675]" country="Russia" name="Buryatiya" pageId="7" pageNumber="503">Buryatia</collectingRegion>
:
<location id="8E8C208D6144FFD4B14CFD76C034FD54" LSID="urn:lsid:plazi:treatment:03FAC7406144FFDAB4BDF8F7C7FCFC46:8E8C208D6144FFD4B14CFD76C034FD54" country="Russia" name="Khamsura village" pageId="7" pageNumber="503" stateProvince="Buryatiya">Khamsura village</location>
,
<locationDeviation id="9A5E6F746144FFD4B7E1FD51C72DFD54" box="[903,1125,681,702]" location="Zakamensk" pageId="7" pageNumber="503">
<quantity id="4CABDBB36144FFD4B7E1FD51C0F2FD54" box="[903,954,681,702]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="503" unit="km" value="7.0">7 km</quantity>
S of Zakamensk
</locationDeviation>
</materialsCitation>
,
<materialsCitation id="3B3B7C0B6144FFD4B017FD51C6CFFD54" box="[1137,1415,681,702]" collectingDate="1971-06-30" collectionCode="V" collectorName="Richter" country="Russia" pageId="7" pageNumber="503" specimenCount="1" specimenCount-male="1" stateProvince="Buryatiya" typeStatus="holotype">
<date id="FFED50966144FFD4B017FD51C791FD54" box="[1137,1241,681,702]" pageId="7" pageNumber="503" value="1971-06-30">
<collectingDate id="EFA9A97E6144FFD4B017FD51C791FD54" box="[1137,1241,681,702]" pageId="7" pageNumber="503" value="1971-06-30">30.vi.1971</collectingDate>
</date>
,
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.
<collectorName id="26A613806144FFD4B167FD51C600FD54" box="[1281,1352,681,702]" pageId="7" pageNumber="503">Richter</collectorName>
(
<specimenCount id="9D55BDDF6144FFD4B13CFD51C636FD54" box="[1370,1406,681,702]" count="1" pageId="7" pageNumber="503" type="male">1 ♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B6144FFD4B75EFD3CC0A1FD33" box="[824,1001,708,729]" country="Russia" location="40 km N of Terey" pageId="7" pageNumber="503" specimenCount="1" stateProvince="Buryatiya" typeStatus="holotype">
<location id="8E8C208D6144FFD4B75EFD3CC0A1FD33" LSID="urn:lsid:plazi:treatment:03FAC7406144FFDAB4BDF8F7C7FCFC46:8E8C208D6144FFD4B75EFD3CC0A1FD33" box="[824,1001,708,729]" country="Russia" name="40 km N of Terey" pageId="7" pageNumber="503" stateProvince="Buryatiya">
<quantity id="4CABDBB36144FFD4B75EFD3CC03FFD33" box="[824,887,708,729]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="503" unit="km" value="40.0">40 km</quantity>
N of Terey
</location>
</materialsCitation>
,
<date id="FFED50966144FFD4B793FD3CC715FD33" box="[1013,1117,708,729]" pageId="7" pageNumber="503" value="1961-06-19">19.vi.1961</date>
, L. Zimin (
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).
<materialsCitation id="3B3B7C0B6144FFD4B16DFD3CC0A7FD19" collectorName="Zabaykalskiy Terr." country="Russia" location="Kyra" pageId="7" pageNumber="503" specimenCount="1" specimenCount-male="1" stateProvince="Buryatiya" typeStatus="holotype">
<collectorName id="26A613806144FFD4B16DFD3CC020FD19" pageId="7" pageNumber="503">Zabaykalskiy Terr.</collectorName>
,
<specimenCount id="9D55BDDF6144FFD4B716FD26C0FBFD19" box="[880,947,734,755]" count="1" pageId="7" pageNumber="503" type="male">1 male</specimenCount>
,
<location id="8E8C208D6144FFD4B7DBFD26C0A7FD19" LSID="urn:lsid:plazi:treatment:03FAC7406144FFDAB4BDF8F7C7FCFC46:8E8C208D6144FFD4B7DBFD26C0A7FD19" box="[957,1007,734,755]" country="Russia" name="Kyra" pageId="7" pageNumber="503" stateProvince="Buryatiya">Kyra</location>
</materialsCitation>
,
<materialsCitation id="3B3B7C0B6144FFD4B79FFD26C786FD19" box="[1017,1230,734,755]" collectingDate="1975-06-17" collectionCode="V" country="Russia" location="Richter" pageId="7" pageNumber="503" specimenCount="1" stateProvince="Buryatiya" typeStatus="holotype">
<date id="FFED50966144FFD4B79FFD26C729FD19" box="[1017,1121,734,755]" pageId="7" pageNumber="503" value="1975-06-17">
<collectingDate id="EFA9A97E6144FFD4B79FFD26C729FD19" box="[1017,1121,734,755]" pageId="7" pageNumber="503" value="1975-06-17">17.vi.1975</collectingDate>
</date>
,
<collectionCode id="ED42EE936144FFD4B00DFD26C734FD19" box="[1131,1148,734,755]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="503" type="Museum">V</collectionCode>
.
<location id="8E8C208D6144FFD4B0E3FD26C786FD19" LSID="urn:lsid:plazi:treatment:03FAC7406144FFDAB4BDF8F7C7FCFC46:8E8C208D6144FFD4B0E3FD26C786FD19" box="[1157,1230,734,755]" country="Russia" name="Richter" pageId="7" pageNumber="503" stateProvince="Buryatiya">Richter</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6144FFD4B738FD01C60EFC21" pageId="7" pageNumber="503" type="diagnosis">
<paragraph id="8BEC76566144FFD4B738FD01C60EFC21" blockId="7.[824,1422,303,2076]" pageId="7" pageNumber="503">
DIAGNOSIS. Mid-sized species (body about
<quantity id="4CABDBB36144FFD4B17DFD01C61DFCE4" box="[1307,1365,761,782]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="503" unit="mm" value="6.0">6 mm</quantity>
) with palpus largely yellow; mesoscutum with 4 vittae, presutural acrostichals and dorsocentrals 1–2-serial, hair-like, minute, partly pale; legs largely brownish yellow; laterotergite with black and pale setae. Male: eyes holoptic; abdominal sternite 8 broadly yellow posteriorly. Female: abdominal tergites 1–6 light grey, narrowly brownish anteriorly and posteriorly (dorsal view); all femora and hind tibia with pennate setae.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6144FFDAB738FC29C7D1FD4A" lastPageId="9" lastPageNumber="505" pageId="7" pageNumber="503" type="description">
<paragraph id="8BEC76566144FFD4B738FC29C7B5FA5A" blockId="7.[824,1422,303,2076]" pageId="7" pageNumber="503">
DESCRIPTION.
<emphasis id="B927AA446144FFD4B06AFC29C708FC0C" bold="true" box="[1036,1088,977,998]" pageId="7" pageNumber="503">Male</emphasis>
(
<figureCitation id="13686AD36144FFD4B032FC29C7EAFC0C" box="[1108,1186,977,998]" captionStart="Figs 13–16" captionStartId="8.[189,227,1937,1956]" captionTargetBox="[215,1392,649,1860]" captionTargetId="figure-184@8.[200,1410,619,1887]" captionTargetPageId="8" captionText="Figs 13–16. Empis (Polyblepharis) subdepilis Shamshev, sp.n.: 13–15 — male: 13 — habitus, holotype, lateral view; 14 — hypopygium, lateral view; 15 — phallus, lateral view; 16 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 13–16. Empis (Polyblepharis) subdepilis Shamshev, sp.n.: 13–15 — самец: 13 — габитус, голотип, вид сбоку; 14 — гипопигий, вид сбоку; 15 — фаллус, вид сбоку; 16 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="7" pageNumber="503">Fig. 13</figureCitation>
). Body length 6.0–6.4 (
<typeStatus id="54E8C8F46144FFD4B758FC14C0DCFBEB" box="[830,916,1004,1025]" pageId="7" pageNumber="503" type="holotype">holotype</typeStatus>
6.1) mm; wing length 5.7–5.9 (
<typeStatus id="54E8C8F46144FFD4B0B2FC14C662FBEB" box="[1236,1322,1004,1025]" pageId="7" pageNumber="503" type="holotype">holotype</typeStatus>
5.9) mm. Head black. Head capsule regions mostly densely greyish pruinose; face shiny along lower margin. Eyes holoptic, upper ommatidia enlarged. Frons represented by small subtriangular space below ocellar triangle and larger space above antennae; bare. Face wide, almost parallel-sided, bare. Ocellar triangle with 2 moderately long, fine and several short setae. Occiput with numerous, long, black setae laterally; postoculars moderately long in upper part, very short laterally; postgena bearing numerous pale, long, hair-like setae. Antenna with scape and pedicel brownish, postpedicel and stylus black; scape about 1.5X longer than pedicel, both with short setulae; postpedicel moderately long, nearly 2.5X longer than wide, with straight margins; stylus short, about 1/3 as long as postpedicel. Proboscis moderately long, labrum brownish, nearly 1.5X as long as head height; palpus broadly yellow (brownish to brownish yellow near base), with scattered black setulae.
</paragraph>
<paragraph id="8BEC76566144FFD4B738FA4EC61BF80C" blockId="7.[824,1422,303,2076]" pageId="7" pageNumber="503">Thorax black, mostly densely greyish pruinose and black setose; mesoscutum with 4 distinct, broad, brownish vittae (dorsal view). Antepronotum with several strong setae dorsally and some pale, hair-like setae laterally. Postpronotal lobe with 1 long and numerous shorter strong setae, bearing additional pale, hair-like setulae. Proepisternum with numerous long, pale, hair-like setae on lower part (sometimes dark setae present), bare on upper part. Prosternum bare. Mesonotal setae: acrostichals hair-like, minute, sparse, arranged in 2 close irregular rows, partly pale, lacking on prescutellar depression; presutural dorsocentrals arranged in 2 close irregular rows, similar to acrostichals, postsuturals sparse, slightly longer and stronger, 1 pair of prescutellars moderately long; 1 short, presutural intra-alar; 1 moderately long presutural supra-alar, 3 long notopleurals (2 posterior setae longer), 1 moderately long postsutural supra-alar, 1 long and 1 very short postalars, 4 scutellars (apical pair long, lateral pair very short); numerous black and pale setulae present on notopleuron anteriorly (usually mostly pale). Laterotergite with numerous black setae anteriorly (number highly variable) and numerous pale setae posteriorly. Anterior and posterior spiracles pale yellow.</paragraph>
<paragraph id="8BEC76566144FFDAB738F814C1DBFE90" blockId="7.[824,1422,303,2076]" lastBlockId="9.[189,787,303,2076]" lastPageId="9" lastPageNumber="505" pageId="7" pageNumber="503">Legs rather long, slender; coxae densely greyish pruinose, remaining podomeres subshiny; mostly black setose. Leg colour: coxae black; trochanters brownish; otherwise legs extensively brownish yellow, fore femur (except extreme apex and base), tibiae in subapical part and tarsi brownish; in paler specimens tibiae almost uniformly brownish yellow and basitarsi brownish yellow in basal part. Fore coxa with numerous, pale, hair-like and several black, strong setae. Femora without whitish pilosity ventrally. Fore femur with short, strong setae in about apical half anteriorly and posteriorly; bearing row of mostly short anteroventral setae and dense, moderately long, rather fine setae posteroventrally. Fore tibia with 3–5 short anterodorsal and 3–5 similar posterodorsal setae. Mid femur with complete rows of short anteroventral and slightly long- er posteroventral spine-like setae; bearing 2–3 short, strong setae near apex anteriorly and posteriorly. Mid tibia with 3–4 short anterodorsal and 3–5 posterodorsal setae; bearing 2–3 less distinct posteroventral setae in apical part. Hind femur almost uniformly slender, only slightly broader than mid femur; with similar setation to mid femur. Hind tibia somewhat gently thickened towards apex; with 5–6 short anterodorsal and 7–8 posterodorsal setae; covered with simple setulae ventrally; 1 short seta in posteroapical comb. Fore and mid tarsomeres 1 slender, hind tarsomere 1 rather slightly thickened; all basitarsi with short, spine-like setae ventrally.</paragraph>
<caption id="DF2C26DE614BFFDBB4DBF869C1A7F7F6" pageId="8" pageNumber="504" startId="8.[189,227,1937,1956]" targetBox="[215,1392,649,1860]" targetPageId="8" targetType="figure">
<paragraph id="8BEC7656614BFFDBB4DBF869C046F83E" blockId="8.[189,1421,1937,2076]" pageId="8" pageNumber="504">
<emphasis id="B927AA44614BFFDBB4DBF869C26BF84E" bold="true" box="[189,291,1937,1956]" pageId="8" pageNumber="504">Figs 13–16.</emphasis>
<taxonomicName id="4C530DD5614BFFDBB54EF869C1EAF84E" authority="Shamshev" authorityName="Shamshev" box="[296,674,1937,1956]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="504" phylum="Arthropoda" rank="species" species="subdepilis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA44614BFFDBB54EF869C215F84E" box="[296,349,1937,1956]" italics="true" pageId="8" pageNumber="504">Empis</emphasis>
(
<emphasis id="B927AA44614BFFDBB50CF869C2AAF84E" box="[362,482,1937,1956]" italics="true" pageId="8" pageNumber="504">Polyblepharis</emphasis>
)
<emphasis id="B927AA44614BFFDBB588F869C10DF84E" box="[494,581,1937,1956]" italics="true" pageId="8" pageNumber="504">subdepilis</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA44614BFFDBB6CCF869C19DF84E" bold="true" box="[682,725,1937,1956]" pageId="8" pageNumber="504">
<taxonomicNameLabel id="A214173F614BFFDBB6CCF869C19DF84E" box="[682,725,1937,1956]" pageId="8" pageNumber="504" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 13–15 — male: 13 — habitus, holotype, lateral view; 14 — hypopygium, lateral view; 15 — phallus, lateral view; 16 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm.
</paragraph>
<paragraph id="8BEC7656614BFFDBB4DBF821C1A7F7F6" blockId="8.[189,1421,1937,2076]" pageId="8" pageNumber="504">
<emphasis id="B927AA44614BFFDBB4DBF821C26EF806" bold="true" box="[189,294,2009,2028]" pageId="8" pageNumber="504">Рис. 13–16.</emphasis>
<taxonomicName id="4C530DD5614BFFDBB54BF821C1E3F806" authority="Shamshev" authorityName="Shamshev" box="[301,683,2009,2028]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="504" phylum="Arthropoda" rank="species" species="subdepilis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA44614BFFDBB54BF821C22AF806" box="[301,354,2009,2028]" italics="true" pageId="8" pageNumber="504">Empis</emphasis>
(
<emphasis id="B927AA44614BFFDBB516F821C2A0F806" box="[368,488,2009,2028]" italics="true" pageId="8" pageNumber="504">Polyblepharis</emphasis>
)
<emphasis id="B927AA44614BFFDBB593F821C104F806" box="[501,588,2009,2028]" italics="true" pageId="8" pageNumber="504">subdepilis</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA44614BFFDBB6D2F821C196F806" bold="true" box="[692,734,2009,2028]" pageId="8" pageNumber="504">
<taxonomicNameLabel id="A214173F614BFFDBB6D2F821C196F806" box="[692,734,2009,2028]" pageId="8" pageNumber="504" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 13–15 — самец: 13 — габитус, голотип, вид сбоку; 14 — гипопигий, вид сбоку; 15 — фаллус, вид сбоку; 16 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм.
</paragraph>
</caption>
<paragraph id="8BEC7656614AFFDAB485FE78C159FDEB" blockId="9.[189,787,303,2076]" pageId="9" pageNumber="505">Wing membrane faintly brownish infuscate. Pterostigma brownish yellow, narrow. Basal costal seta absent. Veins R 5 and M 1+2 divergent subapically; radial fork acute. Anal angle very acute, subsequently anal lobe well developed. Calypter yellow, pale fringed. Halter yellow.</paragraph>
<paragraph id="8BEC7656614AFFDAB485FDFFC22DFBF6" blockId="9.[189,787,303,2076]" pageId="9" pageNumber="505">Abdomen almost entirely black, sternite 7 brownish yellow posteriorly and sternite 8 extensively yellowish. Pruinescence: in lateral view tergites 1–6 densely light grey pruinose on lower part (concolourous with sternites) and brownish on upper part, in dorsal view tergites 2–7 velvety brown, tergites 6–7 with slightly darker iridescent spots. Chaetotaxy: tergites 1–3 with long, pale setae laterally, tergites 1–6 with strong, black posteromarginal setae laterally, otherwise tergites covered with black setulae; sternites 1–5 with scattered pale setulae, sternites 6–8 with more numerous black setulae, sternite 8 with several moderately long posteromarginal setae. Pregenital segments: segment 6 unmodified; tergite 7 with straight posterior margin, bearing scattered setulae; sternite 7 somewhat broader posteriorly than sternite 6, with simple setae; tergite and sternite of segment 8 separated; tergite 8 divided medially, represented by two small, subtriangular sclerites bearing scattered setulae posteriorly; sternite 8 scoop-shaped, constricted near base, each side close to base with horn-like tubercle and elongate oval projection (ventral view) separated by deep subovate depression.</paragraph>
<paragraph id="8BEC7656614AFFDAB485FBDAC13DFB1E" blockId="9.[189,787,303,2076]" pageId="9" pageNumber="505">
Hypopygium (
<figureCitation id="13686AD3614AFFDAB51EFBDAC2B0FBDD" box="[376,504,1058,1079]" captionStart="Figs 13–16" captionStartId="8.[189,227,1937,1956]" captionTargetBox="[215,1392,649,1860]" captionTargetId="figure-184@8.[200,1410,619,1887]" captionTargetPageId="8" captionText="Figs 13–16. Empis (Polyblepharis) subdepilis Shamshev, sp.n.: 13–15 — male: 13 — habitus, holotype, lateral view; 14 — hypopygium, lateral view; 15 — phallus, lateral view; 16 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 13–16. Empis (Polyblepharis) subdepilis Shamshev, sp.n.: 13–15 — самец: 13 — габитус, голотип, вид сбоку; 14 — гипопигий, вид сбоку; 15 — фаллус, вид сбоку; 16 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="9" pageNumber="505">Figs 14, 15</figureCitation>
) moderately large, largely brownish, epandrial lamella reddish brown on upper part and phallus yellowish; cercus and epandrial lamella with black setulae. Epandrial lamella (
<figureCitation id="13686AD3614AFFDAB5A7FB8BC14EFB62" box="[449,518,1139,1160]" captionStart="Figs 13–16" captionStartId="8.[189,227,1937,1956]" captionTargetBox="[215,1392,649,1860]" captionTargetId="figure-184@8.[200,1410,619,1887]" captionTargetPageId="8" captionText="Figs 13–16. Empis (Polyblepharis) subdepilis Shamshev, sp.n.: 13–15 — male: 13 — habitus, holotype, lateral view; 14 — hypopygium, lateral view; 15 — phallus, lateral view; 16 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 13–16. Empis (Polyblepharis) subdepilis Shamshev, sp.n.: 13–15 — самец: 13 — габитус, голотип, вид сбоку; 14 — гипопигий, вид сбоку; 15 — фаллус, вид сбоку; 16 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="9" pageNumber="505">Fig. 14</figureCitation>
) rather trapezium-like, with scattered short setae on lower part closer to apex. Hypandrium rounded apically, bare. Cercus elongate oval (lateral view), with smoothed inner margin, without dorsolateral projection in basal part (dorsal view). Phallus as in
<figureCitation id="13686AD3614AFFDAB64CFB27C138FB1E" box="[554,624,1247,1268]" captionStart="Figs 13–16" captionStartId="8.[189,227,1937,1956]" captionTargetBox="[215,1392,649,1860]" captionTargetId="figure-184@8.[200,1410,619,1887]" captionTargetPageId="8" captionText="Figs 13–16. Empis (Polyblepharis) subdepilis Shamshev, sp.n.: 13–15 — male: 13 — habitus, holotype, lateral view; 14 — hypopygium, lateral view; 15 — phallus, lateral view; 16 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 13–16. Empis (Polyblepharis) subdepilis Shamshev, sp.n.: 13–15 — самец: 13 — габитус, голотип, вид сбоку; 14 — гипопигий, вид сбоку; 15 — фаллус, вид сбоку; 16 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="9" pageNumber="505">Fig. 15</figureCitation>
.
</paragraph>
<paragraph id="8BEC7656614AFFDAB485FB02C2C5F7F6" blockId="9.[189,787,303,2076]" pageId="9" pageNumber="505">
<emphasis id="B927AA44614AFFDAB485FB02C266FAE5" bold="true" box="[227,302,1274,1295]" pageId="9" pageNumber="505">Female</emphasis>
(
<figureCitation id="13686AD3614AFFDAB55BFB02C2CFFAE5" box="[317,391,1274,1295]" captionStart="Figs 13–16" captionStartId="8.[189,227,1937,1956]" captionTargetBox="[215,1392,649,1860]" captionTargetId="figure-184@8.[200,1410,619,1887]" captionTargetPageId="8" captionText="Figs 13–16. Empis (Polyblepharis) subdepilis Shamshev, sp.n.: 13–15 — male: 13 — habitus, holotype, lateral view; 14 — hypopygium, lateral view; 15 — phallus, lateral view; 16 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 13–16. Empis (Polyblepharis) subdepilis Shamshev, sp.n.: 13–15 — самец: 13 — габитус, голотип, вид сбоку; 14 — гипопигий, вид сбоку; 15 — фаллус, вид сбоку; 16 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="9" pageNumber="505">Fig. 16</figureCitation>
). Body length
<quantity id="4CABDBB3614AFFDAB67CFB02C1DAFAE5" box="[538,658,1274,1295]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="6.1" metricValueMin="5.6" pageId="9" pageNumber="505" unit="mm" value="5.85" valueMax="6.1" valueMin="5.6">5.6–6.1 mm</quantity>
; wing length 6.7–
<quantity id="4CABDBB3614AFFDAB48EFAEDC272FAC0" box="[232,314,1301,1322]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="9" pageNumber="505" unit="mm" value="6.3">6.3 mm</quantity>
. Similar to male except as follows. Eyes dichoptic, ommatidia of equal size. Frons broad, parallel-sided, usually bare (sometimes scattered, minute setulae present in lower part). Occiput with sparse setae. Notopleuron with minute, mostly black setulae anteriorly; acrostichals and dorsocentrals slightly stronger, uniformly black. Fore femur with fringe of short pennate setae anterodorsally and longer pennate setae posteroventrally (anterodorsal setae absent close to base). Mid femur with similar fringes of pennate setae as fore femur. Hind femur with fringe of short, pennate setae in about apical 1/3 dorsally; bearing long, pennate, posteroventral setae in about apical 2/3. Fore tibia and tarsus with simple setae. Mid tibia with 3–4 anteroventral and 2–3 posteroventral setae; sometimes bearing scattered, short, slightly flattened setae close to apex ventrally. Hind tibia and hind basitarsus slen- der; hind tibia with short, pennate, anterodorsal setae in apical 1/2–2/3 (in addition to rows of anterodorsal and posterodorsal simple, strong setae); row of short anteroventral setae; bearing moderately long, pennate, posteroventral setae in about apical 3/4. Wing membrane almost hyaline, slightly brownish infuscate at apex. Abdominal tergites 1–6 largely densely light grey pruinose, tergites 2–5 narrowly brownish pruinose anteriorly and posteriorly (dorsal view), sternites 1–6 entirely densely light grey pruinose, segments 7–8 brownish shiny, segment 10 and cercus contrastingly black, subshiny. Abdominal chaetotaxy very reduced, tergites mostly covered with scattered, minute setulae, sternites almost bare; only tergite 1 with numerous, short, black and pale, fine setae. Cercus short, clothed in dense dark setulae.
</paragraph>
<paragraph id="8BEC7656614AFFDAB738FED7C7D1FD4A" blockId="9.[824,1422,303,940]" pageId="9" pageNumber="505">
DIFFERENTIAL DIAGNOSIS. The new species is similar to
<taxonomicName id="4C530DD5614AFFDAB736FEB2C76AFEB5" authority="Loew, 1873" authorityName="Loew" authorityYear="1873" box="[848,1058,330,351]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="505" phylum="Arthropoda" rank="species" species="depilis">
<emphasis id="B927AA44614AFFDAB736FEB2C0E3FEB5" box="[848,939,330,351]" italics="true" pageId="9" pageNumber="505">E. depilis</emphasis>
Loew, 1873
</taxonomicName>
known only from the southern steppe areas of European
<collectingCountry id="F34436C6614AFFDAB78DFE9DC763FE90" box="[1003,1067,357,378]" name="Russia" pageId="9" pageNumber="505">Russia</collectingCountry>
[
<bibRefCitation id="EFC20BA7614AFFDAB050FE9DC611FE90" author="Berezhnova O. N. &amp; Shamshev I. V." box="[1078,1369,357,378]" pageId="9" pageNumber="505" pagination="225 - 229" refId="ref15532" refString="Berezhnova O. N., Shamshev I. V. 2006. New data on the genus Empis L. (Diptera: Empididae) from the forest-steppe and steppe areas of European Russia // International Journal of Dipterological Research. Vol. 17. P. 225 - 229." type="journal article" year="2006">Berezhnova, Shamshev, 2006</bibRefCitation>
]. The male of
<taxonomicName id="4C530DD5614AFFDAB7E1FE78C74BFE7F" box="[903,1027,384,405]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="505" phylum="Arthropoda" rank="species" species="subdepilis" status="sp.n.">
<emphasis id="B927AA44614AFFDAB7E1FE78C0DEFE7F" box="[903,918,384,405]" italics="true" pageId="9" pageNumber="505">E</emphasis>
.
<emphasis id="B927AA44614AFFDAB7C4FE78C74BFE7F" box="[930,1027,384,405]" italics="true" pageId="9" pageNumber="505">subdepilis</emphasis>
</taxonomicName>
<emphasis id="B927AA44614AFFDAB06CFE78C773FE7F" bold="true" box="[1034,1083,384,405]" pageId="9" pageNumber="505">
<taxonomicNameLabel id="A214173F614AFFDAB06CFE78C773FE7F" box="[1034,1083,384,405]" pageId="9" pageNumber="505" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C530DD5614AFFDAB127FE78C007FE45" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="505" phylum="Arthropoda" rank="species" species="depilis">
<emphasis id="B927AA44614AFFDAB127FE78C007FE45" italics="true" pageId="9" pageNumber="505">E. depilis</emphasis>
</taxonomicName>
by the following characters: acrostichal setae partly pale; presutural dorsocentral setae 2-serial; fore femur covered with dense, moderately long, rather fine setae posteroventrally. In addition, in the female of
<taxonomicName id="4C530DD5614AFFDAB05FFE13C7FFFDEA" box="[1081,1207,491,512]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="505" phylum="Arthropoda" rank="species" species="subdepilis" status="sp.n.">
<emphasis id="B927AA44614AFFDAB05FFE13C700FDEA" box="[1081,1096,491,512]" italics="true" pageId="9" pageNumber="505">E</emphasis>
.
<emphasis id="B927AA44614AFFDAB030FE13C7FFFDEA" box="[1110,1207,491,512]" italics="true" pageId="9" pageNumber="505">subdepilis</emphasis>
</taxonomicName>
<emphasis id="B927AA44614AFFDAB0D9FE13C7A7FDEA" bold="true" box="[1215,1263,491,512]" pageId="9" pageNumber="505">
<taxonomicNameLabel id="A214173F614AFFDAB0D9FE13C7A7FDEA" box="[1215,1263,491,512]" pageId="9" pageNumber="505" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
all femora and hind tibia bear pennate setae. The male of
<taxonomicName id="4C530DD5614AFFDAB0B7FDFDC664FDF0" box="[1233,1324,517,538]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="505" phylum="Arthropoda" rank="species" species="depilis">
<emphasis id="B927AA44614AFFDAB0B7FDFDC664FDF0" box="[1233,1324,517,538]" italics="true" pageId="9" pageNumber="505">E. depilis</emphasis>
</taxonomicName>
has black acrostichal setae; uniserial presutural dorsocentral setae and fore femur with row of short, spine-like, posteroventral setae (longer in about apical half). In the female of
<taxonomicName id="4C530DD5614AFFDAB098FDAEC613FD81" box="[1278,1371,598,619]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="505" phylum="Arthropoda" rank="species" species="depilis">
<emphasis id="B927AA44614AFFDAB098FDAEC613FD81" box="[1278,1371,598,619]" italics="true" pageId="9" pageNumber="505">E. depilis</emphasis>
</taxonomicName>
only mid and hind femora bear short posteroventral pennate setae, hind tibia covered with simple setae.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614AFFDAB738FD5EC718FD3C" pageId="9" pageNumber="505" type="etymology">
<paragraph id="8BEC7656614AFFDAB738FD5EC718FD3C" blockId="9.[824,1422,303,940]" pageId="9" pageNumber="505">
ETYMOLOGY. The name of the new species refers to its similarity to
<taxonomicName id="4C530DD5614AFFDAB7D5FD39C718FD3C" authority="Loew." authorityName="Loew." box="[947,1104,705,726]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="505" phylum="Arthropoda" rank="species" species="depilis">
<emphasis id="B927AA44614AFFDAB7D5FD39C745FD3C" box="[947,1037,705,726]" italics="true" pageId="9" pageNumber="505">E. depilis</emphasis>
Loew.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614AFFDAB738FD24C7FCFC46" pageId="9" pageNumber="505" type="distribution">
<paragraph id="8BEC7656614AFFDAB738FD24C7FEFCE6" blockId="9.[824,1422,303,940]" pageId="9" pageNumber="505">
DISTRIBUTION.
<collectingCountry id="F34436C6614AFFDAB07EFD24C730FD1B" box="[1048,1144,732,753]" name="Mongolia" pageId="9" pageNumber="505">Mongolia</collectingCountry>
,
<collectingCountry id="F34436C6614AFFDAB0EEFD24C780FD1B" box="[1160,1224,732,753]" name="Russia" pageId="9" pageNumber="505">Russia</collectingCountry>
(Amurskaya Province,
<collectingRegion id="4997B8B4614AFFDAB70AFD0FC089FCE6" box="[876,961,759,780]" country="Russia" name="Buryatiya" pageId="9" pageNumber="505">Buryatia</collectingRegion>
, Zabaykalskiy Territory).
</paragraph>
<paragraph id="8BEC7656614AFFDAB738FCE9C7FCFC46" blockId="9.[824,1422,303,940]" pageId="9" pageNumber="505">
HABITAT AND SEASONAL OCCURRENCE. According to labels data, the species inhabits open biotopes (meadows, steppe areas, glades), from the end of May until the end of June. Several specimens were taken from flowers of
<taxonomicName id="4C530DD5614AFFDAB124FC9AC035FC78" class="Magnoliopsida" family="Rosaceae" genus="Spiraea" kingdom="Plantae" order="Rosales" pageId="9" pageNumber="505" phylum="Tracheophyta" rank="species" species="sericea">
<emphasis id="B927AA44614AFFDAB124FC9AC035FC78" italics="true" pageId="9" pageNumber="505">Spiraea sericea</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C530DD5614AFFDAB7C3FC85C614FC78" authority="(Turcz.) Maxim." authorityName="Maxim." baseAuthorityName="Turcz." box="[933,1372,893,914]" class="Magnoliopsida" family="Rosaceae" genus="Spiraea" kingdom="Plantae" order="Rosales" pageId="9" pageNumber="505" phylum="Tracheophyta" rank="variety" species="media" variety="sericea">
<emphasis id="B927AA44614AFFDAB7C3FC85C77CFC78" box="[933,1076,893,914]" italics="true" pageId="9" pageNumber="505">Spiraea media</emphasis>
var.
<emphasis id="B927AA44614AFFDAB00FFC85C7E6FC78" box="[1129,1198,893,914]" italics="true" pageId="9" pageNumber="505">sericea</emphasis>
(Turcz.) Maxim.
</taxonomicName>
) and
<specimenCount id="9D55BDDF614AFFDAB75EFC6FC0F4FC46" box="[824,956,919,940]" count="1" pageId="9" pageNumber="505" type="generic">one specimen</specimenCount>
from flowers of an apple.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:number id="BAD9D559BFA8D8519864A7534C98338D">3</mods:number>
</mods:detail>
<mods:extent id="735566E7093AA40CF03DDA79E92A7ECF" unit="page">
<mods:start id="07217EBD4661593217BF8099A3FAE9E9">496</mods:start>
<mods:end id="EE194046112044DCE408BEF49417B693">512</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="80FDB81D9CAB71A639F60E94F93744FE">journal article</mods:classification>
<mods:identifier id="CA00276A8BCD01BDD2D540614C807D02" type="DOI">10.15298/rusentj.33.4.13</mods:identifier>
<mods:identifier id="5E0B70B36C25F1318603F05D76DC3D87" type="ISSN">0132-8069</mods:identifier>
</mods:mods>
<treatment id="03FAC7406146FFD4B485F922C2B3F901" LSID="urn:lsid:plazi:treatment:03FAC7406146FFD4B485F922C2B3F901" httpUri="http://treatment.plazi.org/id/03FAC7406146FFD4B485F922C2B3F901" lastPageId="7" lastPageNumber="503" pageId="5" pageNumber="501">
<subSubSection id="C34925DD6146FFD6B485F922C1A4F91B" box="[227,748,1754,1777]" pageId="5" pageNumber="501" type="nomenclature">
<paragraph id="8BEC76566146FFD6B485F922C1A4F91B" blockId="5.[227,748,1754,1806]" box="[227,748,1754,1777]" pageId="5" pageNumber="501">
<heading id="D0A4C13A6146FFD6B485F922C1A4F91B" box="[227,748,1754,1777]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="501" reason="8">
<taxonomicName id="4C530DD56146FFD6B485F922C1E5F91B" authority="Shamshev" authorityName="Shamshev" box="[227,685,1754,1777]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="501" phylum="Arthropoda" rank="species" species="subcurta" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA446146FFD6B485F922C26EF91B" box="[227,294,1754,1777]" italics="true" pageId="5" pageNumber="501">Empis</emphasis>
(
<emphasis id="B927AA446146FFD6B553F922C283F91B" box="[309,459,1754,1777]" italics="true" pageId="5" pageNumber="501">Polyblepharis</emphasis>
)
<emphasis id="B927AA446146FFD6B5BCF922C17FF91B" box="[474,567,1754,1777]" italics="true" pageId="5" pageNumber="501">subcurta</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA446146FFD6B6D1F922C1A4F91B" bold="true" box="[695,748,1754,1777]" pageId="5" pageNumber="501">
<taxonomicNameLabel id="A214173F6146FFD6B6D1F922C1A4F91B" box="[695,748,1754,1777]" pageId="5" pageNumber="501" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6146FFD6B5CFF90FC16FF8E4" box="[425,551,1783,1806]" pageId="5" pageNumber="501" type="description">
<paragraph id="8BEC76566146FFD6B5CFF90FC16FF8E4" blockId="5.[227,748,1754,1806]" box="[425,551,1783,1806]" pageId="5" pageNumber="501">
<figureCitation id="13686AD36146FFD6B5CFF90FC16BF8E4" box="[425,547,1783,1806]" captionStart="Figs 10–12" captionStartId="6.[189,227,1936,1955]" captionTargetBox="[211,1398,588,1859]" captionTargetId="figure-149@6.[200,1410,574,1881]" captionTargetPageId="6" captionText="Figs 10–12. Empis (Polyblepharis) subcurta Shamshev, sp.n., male: 10 — habitus, holotype, lateral view; 11 — hypopygium, lateral view; 12 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 10–12. Empis (Polyblepharis) subcurta Shamshev, sp.n., самец: 10 — габитус, голотип, вид сбоку; 11 — гипопигий, вид сбоку; 12 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="5" pageNumber="501">Figs 10–12</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6146FFD6B485F8C9C643FEB5" pageId="5" pageNumber="501" type="materials_examined">
<paragraph id="8BEC76566146FFD6B485F8C9C2CDF85B" blockId="5.[189,787,1841,2076]" pageId="5" pageNumber="501">
<materialsCitation id="3B3B7C0B6146FFD6B485F8C9C2CDF85B" collectionCode="TYPE, MATERIAL, ZISP, INS" country="Mongolia" location="Kerzhner" municipality="Khasagt-Khayrkhan Ridge" pageId="5" pageNumber="501" specimenCount="1" specimenCount-male="1" stateProvince="Govi-Altay" typeStatus="holotype">
<collectionCode id="ED42EE936146FFD6B485F8C9C2E0F8AC" box="[227,424,1841,1862]" pageId="5" pageNumber="501">TYPE MATERIAL</collectionCode>
.
<typeStatus id="54E8C8F46146FFD6B5D0F8C9C15AF8AC" box="[438,530,1841,1862]" pageId="5" pageNumber="501" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D55BDDF6146FFD6B647F8C9C17BF8AC" box="[545,563,1841,1862]" count="1" pageId="5" pageNumber="501" type="male">♂</specimenCount>
, labelled [in Cyrillic, Russian]:
<collectingCountry id="F34436C66146FFD6B544F8B4C2CAF88B" box="[290,386,1868,1889]" name="Mongolia" pageId="5" pageNumber="501">Mongolia</collectingCountry>
, G.-Alt. aym. [=Gobi-Altayskiy aymak, now
<collectingRegion id="4997B8B46146FFD6B493F89FC215F896" box="[245,349,1895,1916]" country="Mongolia" name="Govi-Altay" pageId="5" pageNumber="501">Govi-Altai</collectingRegion>
aymag (province)], /
<collectingMunicipality id="6B88EC2C6146FFD6B637F89FC3BEF87C" pageId="5" pageNumber="501">Khasagt-Khayrkhan Ridge</collectingMunicipality>
/
<locationDeviation id="9A5E6F746146FFD6B56AF879C29CF87C" box="[268,468,1921,1942]" location="Dzhargalan" pageId="5" pageNumber="501">
<quantity id="4CABDBB36146FFD6B56AF879C202F87C" box="[268,330,1921,1942]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="501" unit="km" value="15.0">15 km</quantity>
S Dzhargalan
</locationDeviation>
/ 14.viii.[1]970
<location id="8E8C208D6146FFD6B613F879C186F87C" LSID="urn:lsid:plazi:treatment:03FAC7406146FFD4B485F922C2B3F901:8E8C208D6146FFD6B613F879C186F87C" box="[629,718,1921,1942]" country="Mongolia" municipality="Khasagt-Khayrkhan Ridge" name="Kerzhner" pageId="5" pageNumber="501" stateProvince="Govi-Altay">Kerzhner</location>
(
<collectionCode id="ED42EE936146FFD6B6BAF879C047F87C" box="[732,783,1921,1942]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="5" pageNumber="501">ZISP</collectionCode>
,
<collectionCode id="ED42EE936146FFD6B4DBF864C3ABF85B" box="[189,227,1948,1969]" pageId="5" pageNumber="501">INS</collectionCode>
_DIP_0000629).
</materialsCitation>
</paragraph>
<paragraph id="8BEC76566146FFD6B485F84FC643FEB5" blockId="5.[189,787,1841,2076]" lastBlockId="5.[824,1423,303,2076]" pageId="5" pageNumber="501">
<materialsCitation id="3B3B7C0B6146FFD6B485F84FC107F80D" collectionCode="TYPE, MATERIAL, ZISP, INS" country="Mongolia" location="Nartshuk" municipality="Khasagt-Khayrkhan Ridge" pageId="5" pageNumber="501" specimenCount="1" specimenCount-male="1" stateProvince="Govi-Altay" typeStatus="paratype">
<typeStatus id="54E8C8F46146FFD6B485F84FC20BF826" box="[227,323,1975,1996]" pageId="5" pageNumber="501" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F34436C66146FFD6B52AF84FC299F826" box="[332,465,1975,1996]" name="Mongolia" pageId="5" pageNumber="501">MONGOLIA</collectingCountry>
,
<collectingRegion id="4997B8B46146FFD6B5BCF84FC10AF826" box="[474,578,1975,1996]" country="Mongolia" name="Govi-Altay" pageId="5" pageNumber="501">Govi-Altai</collectingRegion>
aymag: same locality as holotype, 14.viii.970,
<location id="8E8C208D6146FFD6B5D3F82AC145F80D" LSID="urn:lsid:plazi:treatment:03FAC7406146FFD4B485F922C2B3F901:8E8C208D6146FFD6B5D3F82AC145F80D" box="[437,525,2002,2023]" country="Mongolia" name="Nartshuk" pageId="5" pageNumber="501" stateProvince="Govi-Altay">Nartshuk</location>
(
<specimenCount id="9D55BDDF6146FFD6B646F82AC10DF80D" box="[544,581,2002,2023]" count="1" pageId="5" pageNumber="501" type="male">1 ♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="3B3B7C0B6146FFD6B63BF82AC15BF7F6" collectingDate="1971-07-10" country="Mongolia" location="Kerzhner" pageId="5" pageNumber="501" specimenCount="1" specimenCount-male="1" stateProvince="Suhbaatar" typeStatus="paratype">
<collectingRegion id="4997B8B46146FFD6B63BF82AC181F80D" box="[605,713,2002,2023]" country="Mongolia" name="Suhbaatar" pageId="5" pageNumber="501">Sükhbaatar</collectingRegion>
aymag [=Sukhe-Batorskiy aymak (province)]: Shiliyn-Bogdo-Ula Mt.,
<date id="FFED50966146FFD6B497F7FFC228F7F6" box="[241,352,2055,2076]" pageId="5" pageNumber="501" value="1971-07-10">
<collectingDate id="EFA9A97E6146FFD6B497F7FFC228F7F6" box="[241,352,2055,2076]" pageId="5" pageNumber="501" value="1971-07-10">10.vii.1971</collectingDate>
</date>
,
<location id="8E8C208D6146FFD6B517F7FFC282F7F6" LSID="urn:lsid:plazi:treatment:03FAC7406146FFD4B485F922C2B3F901:8E8C208D6146FFD6B517F7FFC282F7F6" box="[369,458,2055,2076]" country="Mongolia" name="Kerzhner" pageId="5" pageNumber="501" stateProvince="Suhbaatar">Kerzhner</location>
(
<specimenCount id="9D55BDDF6146FFD6B586F7FFC141F7F6" box="[480,521,2055,2076]" count="1" pageId="5" pageNumber="501" type="male">1 ♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="3B3B7C0B6146FFD6B645F7FFC64FFEB5" collectingDate="1970-08-10" collectionCode="ZISP" country="Mongolia" location="Nartshuk" pageId="5" pageNumber="501" specimenCount="1" specimenCount-male="1" stateProvince="Uvs" typeStatus="paratype">
<collectingRegion id="4997B8B46146FFD6B645F7FFC1DEF7F6" box="[547,662,2055,2076]" country="Mongolia" name="Uvs" pageId="5" pageNumber="501">Uvs aymag</collectingRegion>
(=Ubsunurskiy aymak (province)):
<locationDeviation id="9A5E6F746146FFD6B056FED7C64EFEAE" box="[1072,1286,303,324]" location="Khyargas" pageId="5" pageNumber="501">
<quantity id="4CABDBB36146FFD6B056FED7C72DFEAE" box="[1072,1125,303,324]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="501" unit="km" value="5.0">5 km</quantity>
N of Khyargas
</locationDeviation>
,
<date id="FFED50966146FFD6B175FED7C6C1FEAE" box="[1299,1417,303,324]" pageId="5" pageNumber="501" value="1970-08-10">
<collectingDate id="EFA9A97E6146FFD6B175FED7C6C1FEAE" box="[1299,1417,303,324]" pageId="5" pageNumber="501" value="1970-08-10">10.viii.1970</collectingDate>
</date>
,
<location id="8E8C208D6146FFD6B75EFEB2C0D8FEB5" LSID="urn:lsid:plazi:treatment:03FAC7406146FFD4B485F922C2B3F901:8E8C208D6146FFD6B75EFEB2C0D8FEB5" box="[824,912,330,351]" country="Mongolia" name="Nartshuk" pageId="5" pageNumber="501" stateProvince="Uvs">Nartshuk</location>
(
<specimenCount id="9D55BDDF6146FFD6B7F9FEB2C08AFEB5" box="[927,962,330,351]" count="1" pageId="5" pageNumber="501" type="male">1 ♂</specimenCount>
, dissected, all
<typeStatus id="54E8C8F46146FFD6B036FEB2C7E4FEB5" box="[1104,1196,330,351]" pageId="5" pageNumber="501" type="paratype">paratypes</typeStatus>
in
<collectionCode id="ED42EE936146FFD6B0ACFEB2C648FEB5" box="[1226,1280,330,351]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="5" pageNumber="501">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6146FFD6B738FE9DC7A2FDF6" pageId="5" pageNumber="501" type="diagnosis">
<paragraph id="8BEC76566146FFD6B738FE9DC7A2FDF6" blockId="5.[824,1423,303,2076]" pageId="5" pageNumber="501">
DIAGNOSIS. Mid-sized species (body length about
<quantity id="4CABDBB36146FFD6B75EFE78C0CCFE7F" box="[824,900,384,405]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="501" unit="mm" value="5.5">5.5 mm</quantity>
); male eyes holoptic; palpus yellow; mesoscutum with 3 indistinct, brownish vittae; acrostichal and dorsocentral setae multiserial, pale, hair-like, strong setae usually yellow (notopleurals sometimes black), laterotergite with pale setae; legs brown; wing hyaline; abdominal tergites 1–5 densely brownish pruinose dorsally and grey pruinose laterally.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6146FFD4B738FDDAC11BF971" lastPageId="7" lastPageNumber="503" pageId="5" pageNumber="501" type="description">
<paragraph id="8BEC76566146FFD6B738FDDAC635FBEB" blockId="5.[824,1423,303,2076]" pageId="5" pageNumber="501">
DESCRIPTION.Body length about
<quantity id="4CABDBB36146FFD6B0F9FDDAC7ADFDDD" box="[1183,1253,546,567]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="501" unit="mm" value="5.5">5.5 mm</quantity>
, wing
<quantity id="4CABDBB36146FFD6B17BFDDAC6C2FDDD" box="[1309,1418,546,567]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.6" metricValueMin="5.5" pageId="5" pageNumber="501" unit="mm" value="5.55" valueMax="5.6" valueMin="5.5">5.5–5.6 mm</quantity>
.
<emphasis id="B927AA446146FFD6B75EFDC5C024FDB8" bold="true" box="[824,876,573,594]" pageId="5" pageNumber="501">Male</emphasis>
(
<figureCitation id="13686AD36146FFD6B71BFDC5C080FDB8" box="[893,968,573,594]" captionStart="Figs 10–12" captionStartId="6.[189,227,1936,1955]" captionTargetBox="[211,1398,588,1859]" captionTargetId="figure-149@6.[200,1410,574,1881]" captionTargetPageId="6" captionText="Figs 10–12. Empis (Polyblepharis) subcurta Shamshev, sp.n., male: 10 — habitus, holotype, lateral view; 11 — hypopygium, lateral view; 12 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 10–12. Empis (Polyblepharis) subcurta Shamshev, sp.n., самец: 10 — габитус, голотип, вид сбоку; 11 — гипопигий, вид сбоку; 12 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="5" pageNumber="501">Fig. 10</figureCitation>
). Head capsule regions mostly densely greyish pruinose; face on lower margin and entire clypeus shiny; entirely pale setose (sometimes occiput with 2–3 moderately long, fine, brownish yellow to black setae dorsally). Eyes holoptic, upper ommatidia enlarged. Frons represented by very small subtriangular space just below ocellar triangle and larger space above antennae; bare. Face broad, bare. Ocellar triangle with several moderately long, hair-like setae; ocellars undifferentiated. Occiput and postgena covered with numerous, long, hair-like setae; postoculars moderately long on upper part and very short laterally. Antenna with scape and pedicel brown, postpedicel and stylus black; scape slightly longer than subglobular pedicel, both with very short setulae; postpedicel nearly 2X as long as basal width, with straight margins; stylus nearly as long as postpedicel basal width. Proboscis with labrum brownish yellow, nearly 1.5X longer than eye height; palpus short, pale yellow, with rather numerous pale setulae.
</paragraph>
<paragraph id="8BEC76566146FFD6B738FBFFC679FA0C" blockId="5.[824,1423,303,2076]" pageId="5" pageNumber="501">Thorax black in ground-colour, mostly densely greyish pruinose; mesoscutum with 3 indistinct, brownish vittae along acrostichal and dorsocentral setae; almost entirely pale setose, strong setae yellow (some notopleural setae often black).Antepronotum with stronger setae dorsally and long, hair-like setae laterally on each side. Postpronotal lobe with long, dense, hair-like setae. Proepisternum with tuft of numerous pale, hair-like setae on low- er part. Prosternum bare. Mesonotal setae: acrostichals arranged in 4–5 irregular rows, lacking on prescutellar depression, long, dense, hair-like; dorsocentral setae separated from acrostichals by narrow bare space, presutural dorsocentrals multiserial, similar to acrostichals, postsuturals 1–3-serial, 2 long prescutellars; presutural supra-alar space entirely covered with dense, long, hair-like setae not separated by bare space from dorsocentrals; 3–4 notopleurals (posterior seta sometimes finer); 1 postsutural supra-alar; 1 long and 1 minute postalars, 4 scutellars (lateral setae only slightly shorter than apicals). Laterotergite with numerous pale to pale yellow setae. Anterior and posterior spiracles pale.</paragraph>
<paragraph id="8BEC76566146FFD5B738FA14C13BFE45" blockId="5.[824,1423,303,2076]" lastBlockId="6.[189,786,303,516]" lastPageId="6" lastPageNumber="502" pageId="5" pageNumber="501">
Legs with coxae densely greyish pruinose, remaining podomeres subshiny; mostly black setose. Leg colour: entirely brown (in
<specimenCount id="9D55BDDF6146FFD6B7F9F9DAC755F9DD" box="[927,1053,1570,1591]" count="1" pageId="5" pageNumber="501" type="generic" typeStatus="paratype">one paratype</specimenCount>
hind tibia and femur somewhat paler close to base). Fore coxa covered with pale, long, dense, hair-like setae anteriorly. All femora without whitish pilosity ventrally. Fore femur with row of mostly minute, fine anteroventral setae; covered with hair-like, moderately long, pale setae posteroventrally and posteriorly. Fore tibia with 3–5 short, anterodorsal setae (position and robustness variable). Mid femur with complete rows of sparse, mostly very short, anteroventral and posteroventral setae (slightly longer close to base). Mid tibia slender; with row of 7–9 moderately long, rather fine, anterodorsal and 2–3 similar posterodorsal setae; 2–3 hardly prominent setae ventrally; setae of subapical circlet long. Hind femur slender; with row of 3–4 short anterodorsal setae in about apical third; complete row of rather short and scattered, anteroventral setae; fine, short setae posteroventrally (longer close to base); pale, long, hair-like setae near base posteriorly. Hind tibia slightly, gently thickened towards apex; with 6–7 moderately long, anterodorsal and 3–4 similar posterodorsal setae; simple setulae ventrally; 1 short seta in posteroapical comb. Tarsomeres of fore and mid legs slender, hind basitarsus slightly thickened at base and narrowed towards apex; basitarsi with circlet of moderately long, fine, subapical setae, bearing several spine-like setae ventrally; in addition, hind basitarsus with 2–3 moderately long, fine setae dorsally.
</paragraph>
<paragraph id="8BEC76566145FFD5B485FE4CC606FE15" blockId="6.[189,786,303,516]" lastBlockId="6.[824,1421,303,511]" pageId="6" pageNumber="502">
Wing membrane hyaline; entirely covered with microtrichia; veins mostly yellowish (costa between anterior margin of pterostigma and R
<subScript id="17D774136145FFD5B5D3FE00C2F4FDEE" attach="left" box="[437,444,504,516]" fontSize="5" pageId="6" pageNumber="502">5</subScript>
as well as apical portion of radial veins brownish to brownish yellow), well-sclerotized. Veins R 5 and M
<subScript id="17D774136145FFD5B71EFEA0C037FE8E" attach="left" box="[888,895,344,356]" fontSize="5" pageId="6" pageNumber="502">1</subScript>
divergent towards wing margin; radial fork acute; R
<subScript id="17D774136145FFD5B1E0FEA0C6C5FE8E" attach="left" box="[1414,1421,344,356]" fontSize="5" pageId="6" pageNumber="502">5</subScript>
meeting costa before wing apex; CuA+CuP complete (slightly weakened beyond middle). Cell dm moderately large, with strongly elongate apex; apical portion of M
<subScript id="17D774136145FFD5B084FE50C7A1FE5E" attach="left" box="[1250,1257,424,436]" fontSize="5" pageId="6" pageNumber="502">4</subScript>
nearly 3X long- er than its middle portion. Pterostigma inconspicuous. Basal costal seta absent. Anal lobe well-developed; axillary incision very acute. Squama yellow, pale fringed. Halter yellow.
</paragraph>
<caption id="DF2C26DE6145FFD5B4DBF868C21EF7F1" pageId="6" pageNumber="502" startId="6.[189,227,1936,1955]" targetBox="[211,1398,588,1859]" targetPageId="6" targetType="figure">
<paragraph id="8BEC76566145FFD5B4DBF868C224F839" blockId="6.[189,1422,1936,2075]" pageId="6" pageNumber="502">
<emphasis id="B927AA446145FFD5B4DBF868C26AF849" bold="true" box="[189,290,1936,1955]" pageId="6" pageNumber="502">Figs 10–12.</emphasis>
<taxonomicName id="4C530DD56145FFD5B541F868C1DDF849" authority="Shamshev" authorityName="Shamshev" box="[295,661,1936,1955]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="502" phylum="Arthropoda" rank="species" species="subcurta" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA446145FFD5B541F868C214F849" box="[295,348,1936,1955]" italics="true" pageId="6" pageNumber="502">Empis</emphasis>
(
<emphasis id="B927AA446145FFD5B50FF868C2A9F849" box="[361,481,1936,1955]" italics="true" pageId="6" pageNumber="502">Polyblepharis</emphasis>
)
<emphasis id="B927AA446145FFD5B58AF868C17FF849" box="[492,567,1936,1955]" italics="true" pageId="6" pageNumber="502">subcurta</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA446145FFD5B6FAF868C18FF849" bold="true" box="[668,711,1936,1955]" pageId="6" pageNumber="502">
<taxonomicNameLabel id="A214173F6145FFD5B6FAF868C18FF849" box="[668,711,1936,1955]" pageId="6" pageNumber="502" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, male: 10 — habitus, holotype, lateral view; 11 — hypopygium, lateral view; 12 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm.
</paragraph>
<paragraph id="8BEC76566145FFD5B4DBF820C21EF7F1" blockId="6.[189,1422,1936,2075]" pageId="6" pageNumber="502">
<emphasis id="B927AA446145FFD5B4DBF820C26DF801" bold="true" box="[189,293,2008,2027]" pageId="6" pageNumber="502">Рис. 10–12.</emphasis>
<taxonomicName id="4C530DD56145FFD5B54DF820C1D2F801" authority="Shamshev" authorityName="Shamshev" box="[299,666,2008,2027]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="502" phylum="Arthropoda" rank="species" species="subcurta" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA446145FFD5B54DF820C228F801" box="[299,352,2008,2027]" italics="true" pageId="6" pageNumber="502">Empis</emphasis>
(
<emphasis id="B927AA446145FFD5B50BF820C2ADF801" box="[365,485,2008,2027]" italics="true" pageId="6" pageNumber="502">Polyblepharis</emphasis>
)
<emphasis id="B927AA446145FFD5B597F820C174F801" box="[497,572,2008,2027]" italics="true" pageId="6" pageNumber="502">subcurta</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA446145FFD5B6C4F820C185F801" bold="true" box="[674,717,2008,2027]" pageId="6" pageNumber="502">
<taxonomicNameLabel id="A214173F6145FFD5B6C4F820C185F801" box="[674,717,2008,2027]" pageId="6" pageNumber="502" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, самец: 10 — габитус, голотип, вид сбоку; 11 — гипопигий, вид сбоку; 12 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм.
</paragraph>
</caption>
<paragraph id="8BEC76566144FFD4B485FED7C1A6FCAB" blockId="7.[189,787,303,1771]" pageId="7" pageNumber="503">Abdomen entirely black in ground colour. Pruinescence: tergites 1–5 densely brownish pruinose dorsally and grey pruinose laterally (in dorsal view abdomen with broad brownish stripe); tergites 6–7 with yellowish brown iridescent spot dorsally, grey laterally (lateral view); sternites 1–7 densely grey pruinose; sternite 8 faintly pruinose, with shiny upper antero-lateral projection. Chaetotaxy: abdomen covered with only pale setae; tergites almost bare dorsally (with scattered minute setulae), covered with numerous long fine setae laterally (including slightly stronger posteromarginals); sternite 1 bare, remaining sternites with numerous long setae. Pregenital segments: segment 6 unmodified; tergite 7 slightly concave posteriorly, sternite 7 with slightly produced postero-lateral corner, pleuron of segment 7 slightly sclerotised posteriorly; segment 8 with separated tergite and sternite; tergite 8 somewhat flattened, represented by two subtriangular sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 large, scoop-shaped, somewhat constricted anteriorly, with 2 small, antero-lateral projections separated by depression and larger, broad projection on posterior margin of depression.</paragraph>
<paragraph id="8BEC76566144FFD4B485FCBFC14FFA9E" blockId="7.[189,787,303,1771]" pageId="7" pageNumber="503">
Hypopygium (
<figureCitation id="13686AD36144FFD4B514FCBFC2AEFCB6" box="[370,486,839,860]" captionStart="Figs 10–12" captionStartId="6.[189,227,1936,1955]" captionTargetBox="[211,1398,588,1859]" captionTargetId="figure-149@6.[200,1410,574,1881]" captionTargetPageId="6" captionText="Figs 10–12. Empis (Polyblepharis) subcurta Shamshev, sp.n., male: 10 — habitus, holotype, lateral view; 11 — hypopygium, lateral view; 12 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 10–12. Empis (Polyblepharis) subcurta Shamshev, sp.n., самец: 10 — габитус, голотип, вид сбоку; 11 — гипопигий, вид сбоку; 12 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="7" pageNumber="503">Figs 11, 12</figureCitation>
) moderately large; epandrium and basal half of phallus brown, apical half of phallus and cerci brownish yellow; epandrial lamella shiny on middle part, with sparse, short, pale setae along lower margin and at apex; cercus with black setulae denser towards apex. Epandrium entirely broadly divided (epandrial bridge absent); epandrial lamella (
<figureCitation id="13686AD36144FFD4B4A2FC10C243FC17" box="[196,267,1000,1021]" captionStart="Figs 10–12" captionStartId="6.[189,227,1936,1955]" captionTargetBox="[211,1398,588,1859]" captionTargetId="figure-149@6.[200,1410,574,1881]" captionTargetPageId="6" captionText="Figs 10–12. Empis (Polyblepharis) subcurta Shamshev, sp.n., male: 10 — habitus, holotype, lateral view; 11 — hypopygium, lateral view; 12 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 10–12. Empis (Polyblepharis) subcurta Shamshev, sp.n., самец: 10 — габитус, голотип, вид сбоку; 11 — гипопигий, вид сбоку; 12 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="7" pageNumber="503">Fig. 11</figureCitation>
) subtriangular (lateral view). Hypandrium separated from epandrium; mostly membranous, undivided, narrowly sclerotized along margin, subrectangular in ventral view; bare; gonocoxal apodeme small. Cerci separated with each other and from epandrium; cercus elongate ovate (lateral view), with apex extending to level of epandrial lamella apex, without dorsolateral, lobe-like projection in basal part (dorsal view), with serrate inner margin. Phallus (
<figureCitation id="13686AD36144FFD4B645FB5BC125FB52" box="[547,621,1187,1208]" captionStart="Figs 10–12" captionStartId="6.[189,227,1936,1955]" captionTargetBox="[211,1398,588,1859]" captionTargetId="figure-149@6.[200,1410,574,1881]" captionTargetPageId="6" captionText="Figs 10–12. Empis (Polyblepharis) subcurta Shamshev, sp.n., male: 10 — habitus, holotype, lateral view; 11 — hypopygium, lateral view; 12 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 10–12. Empis (Polyblepharis) subcurta Shamshev, sp.n., самец: 10 — габитус, голотип, вид сбоку; 11 — гипопигий, вид сбоку; 12 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="7" pageNumber="503">Fig. 12</figureCitation>
) almost entirely hidden; zigzag bent, strongly constricted beyond middle; its basal portion gently curved, thick at base and becoming slightly narrower towards constriction; apical bend portion broadly semicircular (its tip pointing to rear), long, almost uniformly slender (except slightly thickened extreme base). Ejaculatory apodeme moderately large, extended far beyond basal curvature of phallus, with lateral wings.
</paragraph>
<paragraph id="8BEC76566144FFD4B485FA82C2D7FA65" blockId="7.[189,787,303,1771]" box="[227,415,1402,1423]" pageId="7" pageNumber="503">
<emphasis id="B927AA446144FFD4B485FA82C27BFA65" bold="true" box="[227,307,1402,1423]" pageId="7" pageNumber="503">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BEC76566144FFD4B485FA6CC11BF971" blockId="7.[189,787,303,1771]" pageId="7" pageNumber="503">
DIFFERENTIAL DIAGNOSIS. The new species is very similar to
<taxonomicName id="4C530DD56144FFD4B547FA57C2A2FA2E" authority="Loew, 1869" authorityName="Loew" authorityYear="1869" box="[289,490,1455,1476]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="503" phylum="Arthropoda" rank="species" species="curta">
<emphasis id="B927AA446144FFD4B547FA57C278FA2E" box="[289,304,1455,1476]" italics="true" pageId="7" pageNumber="503">E</emphasis>
.
<emphasis id="B927AA446144FFD4B55BFA57C238FA2E" box="[317,368,1455,1476]" italics="true" pageId="7" pageNumber="503">curta</emphasis>
Loew, 1869
</taxonomicName>
known only from the
<typeStatus id="54E8C8F46144FFD4B6A1FA57C1B8FA2E" box="[711,752,1455,1476]" pageId="7" pageNumber="503">type</typeStatus>
locality (“Sarepta”,
<collectingRegion id="4997B8B46144FFD4B514FA32C290FA35" box="[370,472,1482,1503]" country="Russia" name="Volgograd" pageId="7" pageNumber="503">Volgograd</collectingRegion>
,
<collectingCountry id="F34436C66144FFD4B583FA32C161FA35" box="[485,553,1482,1503]" name="Russia" pageId="7" pageNumber="503">Russia</collectingCountry>
).
<taxonomicName id="4C530DD56144FFD4B658FA32C190FA35" box="[574,728,1482,1503]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="503" phylum="Arthropoda" rank="species" species="subcurta" status="sp. nov.">
<emphasis id="B927AA446144FFD4B658FA32C190FA35" box="[574,728,1482,1503]" italics="true" pageId="7" pageNumber="503">Empis subcurta</emphasis>
</taxonomicName>
<emphasis id="B927AA446144FFD4B684FA32C05AFA35" bold="true" box="[738,786,1482,1503]" pageId="7" pageNumber="503">
<taxonomicNameLabel id="A214173F6144FFD4B684FA32C05AFA35" box="[738,786,1482,1503]" pageId="7" pageNumber="503" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C530DD56144FFD4B551FA1DC2CEFA10" authorityName="Loew" authorityYear="1869" box="[311,390,1509,1530]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="503" phylum="Arthropoda" rank="species" species="curta">
<emphasis id="B927AA446144FFD4B551FA1DC20EFA10" box="[311,326,1509,1530]" italics="true" pageId="7" pageNumber="503">E</emphasis>
.
<emphasis id="B927AA446144FFD4B535FA1DC2CEFA10" box="[339,390,1509,1530]" italics="true" pageId="7" pageNumber="503">curta</emphasis>
</taxonomicName>
primarily by mostly yellow to pale mesonotal setae and densely pruinose tergites 2–5 (versus shiny). In addition, the new species could be compared with
<taxonomicName id="4C530DD56144FFD4B6AFF9E2C231F9A0" authority="Chvala, 1999" authorityName="Chvala" authorityYear="1999" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="503" phylum="Arthropoda" rank="species" species="indigirca">
<emphasis id="B927AA446144FFD4B6AFF9E2C190F9C5" box="[713,728,1562,1583]" italics="true" pageId="7" pageNumber="503">E</emphasis>
.
<emphasis id="B927AA446144FFD4B683F9E2C3A7F9A0" italics="true" pageId="7" pageNumber="503">indigirca</emphasis>
<bibRefCitation id="EFC20BA76144FFD4B491F9CDC231F9A0" author="Chvala M." box="[247,377,1589,1610]" pageId="7" pageNumber="503" pagination="113 - 225" refId="ref15584" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999</bibRefCitation>
</taxonomicName>
(known from Eastern Siberia). The latter species has entirely black acrostichal and dorsocentral setae, partly brownish yellow legs and black posteromarginal setae on abdominal tergites 1–5 [
<bibRefCitation id="EFC20BA76144FFD4B5A4F97EC102F971" author="Chvala M." box="[450,586,1670,1691]" pageId="7" pageNumber="503" pagination="113 - 225" refId="ref15584" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6144FFD4B485F958C2C3F93A" pageId="7" pageNumber="503" type="etymology">
<paragraph id="8BEC76566144FFD4B485F958C2C3F93A" blockId="7.[189,787,303,1771]" pageId="7" pageNumber="503">
ETYMOLOGY. The epithet of the new species refers to its similarity to
<taxonomicName id="4C530DD56144FFD4B55EF943C2CDF93A" authorityName="Loew" authorityYear="1869" box="[312,389,1723,1744]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="503" phylum="Arthropoda" rank="species" species="curta">
<emphasis id="B927AA446144FFD4B55EF943C20FF93A" box="[312,327,1723,1744]" italics="true" pageId="7" pageNumber="503">E</emphasis>
.
<emphasis id="B927AA446144FFD4B534F943C2CDF93A" box="[338,389,1723,1744]" italics="true" pageId="7" pageNumber="503">curta</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6144FFD4B485F92EC2B3F901" box="[227,507,1750,1771]" pageId="7" pageNumber="503" type="distribution">
<paragraph id="8BEC76566144FFD4B485F92EC2B3F901" blockId="7.[189,787,303,1771]" box="[227,507,1750,1771]" pageId="7" pageNumber="503">
DISTRIBUTION.
<collectingCountry id="F34436C66144FFD4B5F1F92EC2BFF901" box="[407,503,1750,1771]" name="Mongolia" pageId="7" pageNumber="503">Mongolia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

292
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<subSubSection id="C34925DD614AFFDAB73FFC29C624FC02" box="[857,1388,977,1000]" pageId="9" pageNumber="505" type="nomenclature">
<paragraph id="8BEC7656614AFFDAB73FFC29C624FC02" blockId="9.[857,1388,977,1029]" box="[857,1388,977,1000]" pageId="9" pageNumber="505">
<heading id="D0A4C13A614AFFDAB73FFC29C624FC02" box="[857,1388,977,1000]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="505" reason="8">
<taxonomicName id="4C530DD5614AFFDAB73FFC29C664FC02" authority="Shamshev" authorityName="Shamshev" box="[857,1324,977,1000]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="505" phylum="Arthropoda" rank="species" species="subhaemi" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA44614AFFDAB73FFC29C0D4FC02" box="[857,924,977,1000]" italics="true" pageId="9" pageNumber="505">Empis</emphasis>
(
<emphasis id="B927AA44614AFFDAB7CDFC29C709FC02" box="[939,1089,977,1000]" italics="true" pageId="9" pageNumber="505">Polyblepharis</emphasis>
)
<emphasis id="B927AA44614AFFDAB036FC29C7FEFC02" box="[1104,1206,977,1000]" italics="true" pageId="9" pageNumber="505">subhaemi</emphasis>
Shamshev
</taxonomicName>
,
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<taxonomicNameLabel id="A214173F614AFFDAB151FC29C624FC02" box="[1335,1388,977,1000]" pageId="9" pageNumber="505" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614AFFDAB042FC16C7EAFBEF" box="[1060,1186,1006,1029]" pageId="9" pageNumber="505" type="description">
<paragraph id="8BEC7656614AFFDAB042FC16C7EAFBEF" blockId="9.[857,1388,977,1029]" box="[1060,1186,1006,1029]" pageId="9" pageNumber="505">
<figureCitation id="13686AD3614AFFDAB042FC16C7D6FBEF" box="[1060,1182,1006,1029]" captionStart="Figs 17–22" captionStartId="11.[190,228,1921,1940]" captionTargetBox="[217,1402,535,1842]" captionTargetId="figure-118@11.[202,1412,523,1873]" captionTargetPageId="11" captionText="Figs 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — male: 17 — habitus, holotype, lateral view; 18 — hypopygium, lateral view; 19 — hypandrium, ventral view; 20 — cerci, dorsal view; 21 — phallus, lateral view; 22 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — самец: 17 — габитус, голотип, вид сбоку; 18 — гипопигий, вид сбоку; 19 — гипандрий, вид сниЗу; 20 — церки, вид сверху; 21 — фаллус, вид сбоку; 22 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="9" pageNumber="505">Figs 17–22</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614AFFDAB738FBD0C09FFA5E" pageId="9" pageNumber="505" type="materials_examined">
<paragraph id="8BEC7656614AFFDAB738FBD0C78EFB67" blockId="9.[824,1422,1064,2076]" pageId="9" pageNumber="505">
<materialsCitation id="3B3B7C0B614AFFDAB738FBD0C78EFB67" collectionCode="TYPE, MATERIAL, ZISP, INS" country="Mongolia" county="Tsentralny" location="Kerzhner" municipality="Bogdoula" pageId="9" pageNumber="505" specimenCount="1" specimenCount-male="1" stateProvince="Tov" typeStatus="holotype">
<collectionCode id="ED42EE93614AFFDAB738FBD0C76EFBD7" box="[862,1062,1064,1085]" pageId="9" pageNumber="505">TYPE MATERIAL</collectionCode>
.
<typeStatus id="54E8C8F4614AFFDAB05EFBD0C7DCFBD7" box="[1080,1172,1064,1085]" pageId="9" pageNumber="505" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D55BDDF614AFFDAB0C0FBD0C7F0FBD7" box="[1190,1208,1064,1085]" count="1" pageId="9" pageNumber="505" type="male">♂</specimenCount>
labelled:
<collectingCountry id="F34436C6614AFFDAB14CFBD0C6C2FBD7" box="[1322,1418,1064,1085]" name="Mongolia" pageId="9" pageNumber="505">Mongolia</collectingCountry>
,
<collectingCounty id="628D0EDA614AFFDAB75EFBBBC0D5FBB2" box="[824,925,1091,1112]" pageId="9" pageNumber="505">Tsentralny</collectingCounty>
/ aymak [now
<collectingRegion id="4997B8B4614AFFDAB022FBBBC7FFFBB2" box="[1092,1207,1091,1112]" country="Mongolia" name="Tov" pageId="9" pageNumber="505">Töv aymag</collectingRegion>
(province)] sev. skl. [=northern slope] of
<collectingMunicipality id="6B88EC2C614AFFDAB068FBA6C723FB99" box="[1038,1131,1118,1139]" pageId="9" pageNumber="505">Bogdoula</collectingMunicipality>
near
<collectingRegion id="4997B8B4614AFFDAB0CDFBA6C65FFB99" box="[1195,1303,1118,1139]" country="Mongolia" name="Ulaanbaatar" pageId="9" pageNumber="505">Ulan-Bator</collectingRegion>
/
<location id="8E8C208D614AFFDAB152FBA6C6C5FB99" LSID="urn:lsid:plazi:treatment:03FAC740614AFFD8B73FFC29C6C5FE32:8E8C208D614AFFDAB152FBA6C6C5FB99" box="[1332,1421,1118,1139]" country="Mongolia" county="Tsentralny" municipality="Bogdoula" name="Kerzhner" pageId="9" pageNumber="505" stateProvince="Tov">Kerzhner</location>
14.vii.[1]967 (
<collectionCode id="ED42EE93614AFFDAB7A4FB80C0BDFB67" box="[962,1013,1144,1165]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="9" pageNumber="505">ZISP</collectionCode>
,
<collectionCode id="ED42EE93614AFFDAB798FB80C76CFB67" box="[1022,1060,1144,1165]" pageId="9" pageNumber="505">INS</collectionCode>
_DIP_0000627).
</materialsCitation>
</paragraph>
<paragraph id="8BEC7656614AFFDAB738FB6BC09FFA5E" blockId="9.[824,1422,1064,2076]" pageId="9" pageNumber="505">
<materialsCitation id="3B3B7C0B614AFFDAB738FB6BC74DFB29" collectionCode="TYPE, MATERIAL, ZISP, INS" country="Mongolia" county="Tsentralny" location="Kerzhner" municipality="Bogdoula" pageId="9" pageNumber="505" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Tov" typeStatus="paratype">
<typeStatus id="54E8C8F4614AFFDAB738FB6BC0F6FB42" box="[862,958,1171,1192]" pageId="9" pageNumber="505" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F34436C6614AFFDAB7AFFB6BC706FB42" box="[969,1102,1171,1192]" name="Mongolia" pageId="9" pageNumber="505">MONGOLIA</collectingCountry>
,
<collectingRegion id="4997B8B4614AFFDAB03FFB6BC783FB42" box="[1113,1227,1171,1192]" country="Mongolia" name="Tov" pageId="9" pageNumber="505">Töv aymag</collectingRegion>
: same data as holotype (
<specimenCount id="9D55BDDF614AFFDAB712FB56C0F8FB29" box="[884,944,1198,1219]" count="3" pageId="9" pageNumber="505" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D55BDDF614AFFDAB7A7FB56C0B4FB29" box="[961,1020,1198,1219]" count="4" pageId="9" pageNumber="505" type="female">4 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614AFFDAB070FB56C734FB34" collectingDate="1967-07-04" collectorName="Zaitzev" country="Mongolia" location="Bogdo-ula Mt." municipality="Zaysan" pageId="9" pageNumber="505" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<collectingMunicipality id="6B88EC2C614AFFDAB070FB56C716FB29" box="[1046,1118,1198,1219]" pageId="9" pageNumber="505">Zaysan</collectingMunicipality>
, southern slope of
<location id="8E8C208D614AFFDAB14EFB56C015FB34" LSID="urn:lsid:plazi:treatment:03FAC740614AFFD8B73FFC29C6C5FE32:8E8C208D614AFFDAB14EFB56C015FB34" country="Mongolia" municipality="Zaysan" name="Bogdo-ula Mt." pageId="9" pageNumber="505">Bogdo-ula Mt.</location>
,
<date id="FFED5096614AFFDAB70DFB31C086FB34" box="[875,974,1225,1246]" pageId="9" pageNumber="505" value="1967-07-04">
<collectingDate id="EFA9A97E614AFFDAB70DFB31C086FB34" box="[875,974,1225,1246]" pageId="9" pageNumber="505" value="1967-07-04">4.vii.1967</collectingDate>
</date>
,
<collectorName id="26A61380614AFFDAB7BBFB31C76DFB34" box="[989,1061,1225,1246]" pageId="9" pageNumber="505">Zaitzev</collectorName>
(
<specimenCount id="9D55BDDF614AFFDAB051FB31C73BFB34" box="[1079,1139,1225,1246]" count="2" pageId="9" pageNumber="505" type="female">2 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614AFFDAB0EAFB31C652FB13" collectingDate="1967-06-22" collectorName="Zaitzev" country="Mongolia" location="Bogdo-ula" pageId="9" pageNumber="505" specimenCount="1" specimenCount-female="1" stateProvince="Ulaanbaatar" typeStatus="paratype">
northern slope of
<location id="8E8C208D614AFFDAB123FB31C01DFB13" LSID="urn:lsid:plazi:treatment:03FAC740614AFFD8B73FFC29C6C5FE32:8E8C208D614AFFDAB123FB31C01DFB13" country="Mongolia" name="Bogdo-ula" pageId="9" pageNumber="505" stateProvince="Ulaanbaatar">Bogdo-ula</location>
near
<collectingRegion id="4997B8B4614AFFDAB7F0FB1CC74CFB13" box="[918,1028,1252,1273]" country="Mongolia" name="Ulaanbaatar" pageId="9" pageNumber="505">Ulan-Bator</collectingRegion>
,
<date id="FFED5096614AFFDAB074FB1CC733FB13" box="[1042,1147,1252,1273]" pageId="9" pageNumber="505" value="1967-06-22">
<collectingDate id="EFA9A97E614AFFDAB074FB1CC733FB13" box="[1042,1147,1252,1273]" pageId="9" pageNumber="505" value="1967-06-22">22.vi.1967</collectingDate>
</date>
,
<collectorName id="26A61380614AFFDAB0EDFB1CC79BFB13" box="[1163,1235,1252,1273]" pageId="9" pageNumber="505">Zaitzev</collectorName>
(
<specimenCount id="9D55BDDF614AFFDAB08EFB1CC658FB13" box="[1256,1296,1252,1273]" count="1" pageId="9" pageNumber="505" type="female">1 ♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="3B3B7C0B614AFFDAB14EFB1CC7A4FAC4" collectingDate="1970-07-31" collectorName="Megmarsuryan" country="Mongolia" location="Tevshrulekh" pageId="9" pageNumber="505" specimenCount="1" specimenCount-male="1" stateProvince="Arhangay" typeStatus="paratype">
<collectingRegion id="4997B8B4614AFFDAB14EFB1CC6C5FB13" box="[1320,1421,1252,1273]" country="Mongolia" name="Arhangay" pageId="9" pageNumber="505">Arkhangai</collectingRegion>
aymag [=Ara-Khangayskiy aymak (province)]:
<location id="8E8C208D614AFFDAB171FB06C6C5FAF9" LSID="urn:lsid:plazi:treatment:03FAC740614AFFD8B73FFC29C6C5FE32:8E8C208D614AFFDAB171FB06C6C5FAF9" box="[1303,1421,1278,1299]" country="Mongolia" name="Tevshrulekh" pageId="9" pageNumber="505" stateProvince="Arhangay">Tevshrulekh</location>
<quantity id="4CABDBB3614AFFDAB75EFAE1C03CFAC4" box="[824,884,1305,1326]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="505" unit="km" value="35.0">35 km</quantity>
SW,
<date id="FFED5096614AFFDAB7C1FAE1C75EFAC4" box="[935,1046,1305,1326]" pageId="9" pageNumber="505" value="1970-07-31">
<collectingDate id="EFA9A97E614AFFDAB7C1FAE1C75EFAC4" box="[935,1046,1305,1326]" pageId="9" pageNumber="505" value="1970-07-31">31.vii.1970</collectingDate>
</date>
,
<collectorName id="26A61380614AFFDAB079FAE1C7F8FAC4" box="[1055,1200,1305,1326]" pageId="9" pageNumber="505">Megmarsuryan</collectorName>
(
<specimenCount id="9D55BDDF614AFFDAB0A6FAE1C7AAFAC4" box="[1216,1250,1305,1326]" count="1" pageId="9" pageNumber="505" type="male">1 ♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="3B3B7C0B614AFFDAB093FAE1C60FFA8E" collectingDate="1967-07-28" collectingDateMax="1967-07-29" collectingDateMin="1967-07-28" collectorName="Zaitzev" country="Mongolia" location="Gurban-Saykhan" pageId="9" pageNumber="505" specimenCount="1" specimenCount-male="1" stateProvince="Omnogovi" typeStatus="paratype">
<collectingRegion id="4997B8B4614AFFDAB093FAE1C007FAA3" country="Mongolia" name="Omnogovi" pageId="9" pageNumber="505">Ömnögovi aimag</collectingRegion>
[=Yuzhno-Gobiyskiy aimak (province)]:
<location id="8E8C208D614AFFDAB083FACCC6C2FAA3" LSID="urn:lsid:plazi:treatment:03FAC740614AFFD8B73FFC29C6C5FE32:8E8C208D614AFFDAB083FACCC6C2FAA3" box="[1253,1418,1332,1353]" country="Mongolia" name="Gurban-Saykhan" pageId="9" pageNumber="505" stateProvince="Omnogovi">Gurban-Saykhan</location>
,
<locationDeviation id="9A5E6F74614AFFDAB75EFAB7C74EFA8E" box="[824,1030,1359,1380]" location="Bulgan" pageId="9" pageNumber="505">
<quantity id="4CABDBB3614AFFDAB75EFAB7C032FA8E" box="[824,890,1359,1380]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="505" unit="km" value="40.0">40 km</quantity>
S of
<collectingRegion id="4997B8B4614AFFDAB7D8FAB7C74EFA8E" box="[958,1030,1359,1380]" country="Mongolia" name="Bulgan" pageId="9" pageNumber="505">Bulgan</collectingRegion>
</locationDeviation>
,
<date id="FFED5096614AFFDAB073FAB7C7E0FA8E" box="[1045,1192,1359,1380]" pageId="9" pageNumber="505" value="1967-07-28" valueMax="1967-07-29" valueMin="1967-07-28">
<collectingDate id="EFA9A97E614AFFDAB073FAB7C7E0FA8E" box="[1045,1192,1359,1380]" pageId="9" pageNumber="505" value="1967-07-28" valueMax="1967-07-29" valueMin="1967-07-28">28–29.vii.1967</collectingDate>
</date>
,
<collectorName id="26A61380614AFFDAB0DEFAB7C648FA8E" box="[1208,1280,1359,1380]" pageId="9" pageNumber="505">Zaitzev</collectorName>
(
<specimenCount id="9D55BDDF614AFFDAB173FAB7C675FA8E" box="[1301,1341,1359,1380]" count="1" pageId="9" pageNumber="505" type="male">1 ♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="3B3B7C0B614AFFDAB130FAB7C09BFA5E" collectingDate="1977-08-10" collectionCode="V" collectorName="Kovalev" country="Russia" county="Zabaykalskiy Terr." location="Chernyshevsk" municipality="Kuenga River" pageId="9" pageNumber="505" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="F34436C6614AFFDAB130FAB7C029FA95" name="Russia" pageId="9" pageNumber="505">RUSSIA</collectingCountry>
,
<collectingCounty id="628D0EDA614AFFDAB70AFA92C76EFA95" box="[876,1062,1386,1407]" pageId="9" pageNumber="505">Zabaykalskiy Terr.</collectingCounty>
:
<collectingMunicipality id="6B88EC2C614AFFDAB049FA92C7FEFA95" box="[1071,1206,1386,1407]" pageId="9" pageNumber="505">Kuenga River</collectingMunicipality>
above
<location id="8E8C208D614AFFDAB099FA92C6C2FA95" LSID="urn:lsid:plazi:treatment:03FAC740614AFFD8B73FFC29C6C5FE32:8E8C208D614AFFDAB099FA92C6C2FA95" box="[1279,1418,1386,1407]" country="Russia" county="Zabaykalskiy Terr." municipality="Kuenga River" name="Chernyshevsk" pageId="9" pageNumber="505">Chernyshevsk</location>
, blooming herb meadow on a slope,
<date id="FFED5096614AFFDAB0FBFA7CC65CFA73" box="[1181,1300,1412,1433]" pageId="9" pageNumber="505" value="1977-08-10">
<collectingDate id="EFA9A97E614AFFDAB0FBFA7CC65CFA73" box="[1181,1300,1412,1433]" pageId="9" pageNumber="505" value="1977-08-10">10.viii.1977</collectingDate>
</date>
,
<collectionCode id="ED42EE93614AFFDAB179FA7CC678FA73" box="[1311,1328,1412,1433]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="9" pageNumber="505" type="Museum">V</collectionCode>
.
<collectorName id="26A61380614AFFDAB15AFA7CC6C5FA73" box="[1340,1421,1412,1433]" pageId="9" pageNumber="505">Kovalev</collectorName>
(
<specimenCount id="9D55BDDF614AFFDAB727FA67C02CFA5E" box="[833,868,1439,1460]" count="1" pageId="9" pageNumber="505" type="male">1 ♂</specimenCount>
, dissected)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614AFFDAB738FA42C027F94F" pageId="9" pageNumber="505" type="diagnosis">
<paragraph id="8BEC7656614AFFDAB738FA42C027F94F" blockId="9.[824,1422,1064,2076]" pageId="9" pageNumber="505">
DIAGNOSIS. Mid-sized species (body about
<quantity id="4CABDBB3614AFFDAB156FA42C6CAFA25" box="[1328,1410,1466,1487]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="505" unit="mm" value="5.5">5.5 mm</quantity>
); palpus yellow; mesoscutum with 3 brownish vittae, acrostichal setae biserial and presutural dorsocentrals 3-serial; legs extensively brown, femora at apex and tibiae brownish yellow. Male: eyes holoptic; wings faintly brownish infuscate; tergites 1–6 densely brownish pruinose dorsally and subshiny laterally. Female: mid and hind femora with posteroventral pennate setae; hind tibia with anterodorsal and posteroventral pennate setae.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614AFFD8B738F953C644FE95" lastPageId="11" lastPageNumber="507" pageId="9" pageNumber="505" type="description">
<paragraph id="8BEC7656614AFFD9B738F953C1AAFE90" blockId="9.[824,1422,1064,2076]" lastBlockId="10.[189,788,303,2076]" lastPageId="10" lastPageNumber="506" pageId="9" pageNumber="505">
DESCRIPTION. Body length about
<quantity id="4CABDBB3614AFFDAB0B5F953C606F92A" box="[1235,1358,1707,1728]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="9" pageNumber="505" unit="mm" value="5.4" valueMax="5.5" valueMin="5.3">5.3–5.5 mm</quantity>
, wing
<quantity id="4CABDBB3614AFFDAB75EF93EC0FBF931" box="[824,947,1734,1755]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.6" metricValueMin="5.5" pageId="9" pageNumber="505" unit="mm" value="5.55" valueMax="5.6" valueMin="5.5">5.5–5.6 mm</quantity>
.
<emphasis id="B927AA44614AFFDAB7A7F93EC0BDF931" bold="true" box="[961,1013,1734,1755]" pageId="9" pageNumber="505">Male</emphasis>
(
<figureCitation id="13686AD3614AFFDAB061F93EC71BF931" box="[1031,1107,1734,1755]" captionStart="Figs 17–22" captionStartId="11.[190,228,1921,1940]" captionTargetBox="[217,1402,535,1842]" captionTargetId="figure-118@11.[202,1412,523,1873]" captionTargetPageId="11" captionText="Figs 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — male: 17 — habitus, holotype, lateral view; 18 — hypopygium, lateral view; 19 — hypandrium, ventral view; 20 — cerci, dorsal view; 21 — phallus, lateral view; 22 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — самец: 17 — габитус, голотип, вид сбоку; 18 — гипопигий, вид сбоку; 19 — гипандрий, вид сниЗу; 20 — церки, вид сверху; 21 — фаллус, вид сбоку; 22 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="9" pageNumber="505">Fig. 17</figureCitation>
). Head capsule regions mostly densely greyish pruinose; face on lower margin and entire clypeus shiny; mostly black setose. Eyes holoptic, upper ommatidia enlarged. Frons represented by very small subtriangular space just below ocellar triangle and larger space above antennae; bare. Face broad, bare. Ocellar triangle with 2 fine, moderately long and several slightly shorter setae. Occiput with numerous long setae; postoculars rather fine, moderately long on upper part and short laterally; postgena with numerous, pale, hair-like setae. Antenna with scape and pedicel brown, postpedicel and stylus black; scape slightly longer than subglobular pedicel, both with very short setulae; postpedicel nearly 2.5X as long as basal width, with straight margins; stylus nearly 1.5X as long as postpedicel basal width. Proboscis with labrum brownish yellow, nearly 1.7X longer than eye height; palpus short, yellow, with scattered black setulae.
</paragraph>
<paragraph id="8BEC76566149FFD9B485FE78C227FC5A" blockId="10.[189,788,303,2076]" pageId="10" pageNumber="506">Thorax black in ground-colour, mostly densely light grey pruinose and black setose; mesoscutum rather brownish grey pruinose, with 3 indistinct, brownish vittae along acrostichal and dorsocentral setae. Antepronotum with strong setae dorsally and long, hair-like setae laterally. Postpronotal lobe with 1 long seta surrounded by short, fine, black and pale setae. Proepisternum with tuft of numerous, pale, hair-like setae on lower part. Prosternum bare. Mesonotal setae: acrostichals arranged in 2 close irregular rows, lacking on prescutellar depression, rather long (nearly as long as antennal stylus), numerous; presutural dorsocentral setae mostly arranged in 3 irregular rows (sometimes more numerous opposite postpronotal lobe), similar to acrostichals, postsuturals 1–2-serial, longer (2–3 prescutellars longest); presutural intra-alar seta undistinguishable from dorsocentrals; 1 presutural supra-alar; 3 notopleurals; 1 postsutural supra-alar; 1 long and 1 minute postalars, 4 scutellars (lateral setae only slightly shorter than apicals); in addition, pale setulae present in anterior part of notopleuron and just behind postpronotal lobe. Laterotergite with numerous pale to pale yellow setae. Anterior and posterior spiracles pale.</paragraph>
<paragraph id="8BEC76566149FFD9B485FC4EC165F9B8" blockId="10.[189,788,303,2076]" pageId="10" pageNumber="506">Legs with coxae densely greyish pruinose, remaining podomeres subshiny; mostly black setose. Leg colour: extensively brown; femora brownish yellow narrowly at apex (also, sometimes mid and hind femora slightly paler near extreme base); fore and hind tibiae mostly brownish yellow (yellowish close to base), mid tibia rather uniformly yellowish. Fore coxa mostly covered with pale, hair-like setae anteriorly; with additional scattered, strong, black setae. Mid femur with whitish pilosity ventrally. Fore femur with row of minute, fine anteroventral setae; covered with hair-like, moderately long, pale setae posteroventrally and posteriorly. Fore tibia with 3–4 short, anterodorsal setae (position and robustness variable). Mid femur with complete rows of short, spine-like anteroventral and posteroventral setae. Mid tibia slender; with row of 4–5 moderately long, strong anterodorsals; 1 posterodorsal seta near middle (often absent); 2–3 short posteroventrals in apical half (position and robustness variable). Hind femur slender; 3–4 short anterodorsal setae in about apical 1/3; complete row of short, spine-like anteroventral setae; dense spinule-like setae posteroventrally. Hind tibia slightly gently thickened towards apex; with 6–7 short, anterodorsal and 3–4 similar posterodorsal setae; simple setulae ventrally; no seta in posteroapical comb. Tarsomeres of fore and mid legs slender, hind basitarsus slightly thickened at base and narrowed towards apex; basitarsi with several spine-like setae ventrally.</paragraph>
<paragraph id="8BEC76566149FFD9B485F9A0C1DFF88A" blockId="10.[189,788,303,2076]" pageId="10" pageNumber="506">
Wing membrane faintly infuscate; entirely covered with microtrichia; veins mostly brownish (brownish yellow close to base), well-sclerotized. Veins R 5 and M 1 divergent towards wing margin; radial fork acute; R
<subScript id="17D774136149FFD9B5AFF94FC298F929" attach="left" box="[457,464,1719,1731]" fontSize="5" pageId="10" pageNumber="506">5</subScript>
meeting costa before wing apex; CuA+CuP complete (slightly weakened beyond middle). Cell dm moderately large, with strongly elongate apex; apical portion of M
<subScript id="17D774136149FFD9B573F8F0C254F8FE" attach="left" box="[277,284,1800,1812]" fontSize="5" pageId="10" pageNumber="506">4</subScript>
nearly 3X longer than its middle portion. Pterostigma brownish yellow. Basal costal seta usually present, short (sometimes absent). Anal lobe well-developed; axillary incision very acute. Squama yellow, pale fringed. Halter yellow.
</paragraph>
<paragraph id="8BEC76566149FFD9B485F89EC76EFD33" blockId="10.[189,788,303,2076]" lastBlockId="10.[824,1423,303,2076]" pageId="10" pageNumber="506">Abdomen entirely black in ground colour. Pruinescence: tergites 1–6 densely brownish pruinose dorsally and subshiny laterally; tergites 6–7 with dark brown iridescent spot dorsally (better visible in lateral view); sternites 1–7 mostly densely greyish pruinose, tergites 2–7 faintly pruinose along posterior margin; sternite 8 faintly pruinose, subshiny posteriorly. Chaetotaxy: tergites 2–6 covered with black setulae dorsally (tergite 6 only anteriorly), tergite 7 with scattered setulae only laterally; tergites 1–3 with pale to yellowish, long, fine setae laterally, tergites 4–6 with intermixed black and pale setulae laterally; tergites 2–4 with stronger, yellowish to black posteromarginal setae laterally (long- er on tergite 2); sternite 1 bare, sternites 2–7 with very long, pale yellow, posteromarginal setae; sternite 8 only with black setae, including moderately long posteromarginals. Pregenital segments: segment 6 unmodified; tergite 7 slightly concave posteriorly, sternite 7 with slightly produced postero-lateral corner, pleuron of segment 7 slightly sclerotised posteriorly; segment 8 with separat- ed tergite and sternite; tergite 8 somewhat flattened, represented by two subtriangular sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 large, scoop-shaped, somewhat constricted anteriorly, with 2 small, antero-lateral, rounded projections separated by depression and similar projection on posterior margin of depression.</paragraph>
<paragraph id="8BEC76566149FFD9B738FD26C66CFB1E" blockId="10.[824,1423,303,2076]" pageId="10" pageNumber="506">
Hypopygium (
<figureCitation id="13686AD36149FFD9B78BFD26C717FD19" box="[1005,1119,734,755]" captionStart="Figs 17–22" captionStartId="11.[190,228,1921,1940]" captionTargetBox="[217,1402,535,1842]" captionTargetId="figure-118@11.[202,1412,523,1873]" captionTargetPageId="11" captionText="Figs 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — male: 17 — habitus, holotype, lateral view; 18 — hypopygium, lateral view; 19 — hypandrium, ventral view; 20 — cerci, dorsal view; 21 — phallus, lateral view; 22 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — самец: 17 — габитус, голотип, вид сбоку; 18 — гипопигий, вид сбоку; 19 — гипандрий, вид сниЗу; 20 — церки, вид сверху; 21 — фаллус, вид сбоку; 22 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="10" pageNumber="506">Figs 18–21</figureCitation>
) moderately large; epandrium brown, cerci yellowish brown; epandrial lamella with short, rather sparse setae along lower margin and near apex; cercus covered with black setulae. Epandrium entirely broadly divid- ed (epandrial bridge absent); epandrial lamella (
<figureCitation id="13686AD36149FFD9B168FCB2C61CFCB5" box="[1294,1364,842,863]" captionStart="Figs 17–22" captionStartId="11.[190,228,1921,1940]" captionTargetBox="[217,1402,535,1842]" captionTargetId="figure-118@11.[202,1412,523,1873]" captionTargetPageId="11" captionText="Figs 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — male: 17 — habitus, holotype, lateral view; 18 — hypopygium, lateral view; 19 — hypandrium, ventral view; 20 — cerci, dorsal view; 21 — phallus, lateral view; 22 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — самец: 17 — габитус, голотип, вид сбоку; 18 — гипопигий, вид сбоку; 19 — гипандрий, вид сниЗу; 20 — церки, вид сверху; 21 — фаллус, вид сбоку; 22 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="10" pageNumber="506">Fig. 18</figureCitation>
) subtriangular (lateral view). Hypandrium (
<figureCitation id="13686AD36149FFD9B0CAFC9DC7BAFC90" box="[1196,1266,869,890]" captionStart="Figs 17–22" captionStartId="11.[190,228,1921,1940]" captionTargetBox="[217,1402,535,1842]" captionTargetId="figure-118@11.[202,1412,523,1873]" captionTargetPageId="11" captionText="Figs 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — male: 17 — habitus, holotype, lateral view; 18 — hypopygium, lateral view; 19 — hypandrium, ventral view; 20 — cerci, dorsal view; 21 — phallus, lateral view; 22 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — самец: 17 — габитус, голотип, вид сбоку; 18 — гипопигий, вид сбоку; 19 — гипандрий, вид сниЗу; 20 — церки, вид сверху; 21 — фаллус, вид сбоку; 22 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="10" pageNumber="506">Fig. 19</figureCitation>
) separated from epandrium; mostly membranous, undivided, narrowly sclerotized along margin, rather subtriangular in ventral view (broad apically); bare; gonocoxal apodeme small. Cerci (
<figureCitation id="13686AD36149FFD9B175FC4EC611FC21" box="[1299,1369,950,971]" captionStart="Figs 17–22" captionStartId="11.[190,228,1921,1940]" captionTargetBox="[217,1402,535,1842]" captionTargetId="figure-118@11.[202,1412,523,1873]" captionTargetPageId="11" captionText="Figs 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — male: 17 — habitus, holotype, lateral view; 18 — hypopygium, lateral view; 19 — hypandrium, ventral view; 20 — cerci, dorsal view; 21 — phallus, lateral view; 22 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — самец: 17 — габитус, голотип, вид сбоку; 18 — гипопигий, вид сбоку; 19 — гипандрий, вид сниЗу; 20 — церки, вид сверху; 21 — фаллус, вид сбоку; 22 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="10" pageNumber="506">Fig. 20</figureCitation>
) separated with each other and from epandrium; cercus elongate ovate (lateral view), with apex extending to level of epandrial lamella apex, without lobe-like, dorsolateral projection at basal part (dorsal view), with serrate inner margin. Phallus (
<figureCitation id="13686AD36149FFD9B126FBDAC6CEFBDD" box="[1344,1414,1058,1079]" captionStart="Figs 17–22" captionStartId="11.[190,228,1921,1940]" captionTargetBox="[217,1402,535,1842]" captionTargetId="figure-118@11.[202,1412,523,1873]" captionTargetPageId="11" captionText="Figs 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — male: 17 — habitus, holotype, lateral view; 18 — hypopygium, lateral view; 19 — hypandrium, ventral view; 20 — cerci, dorsal view; 21 — phallus, lateral view; 22 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — самец: 17 — габитус, голотип, вид сбоку; 18 — гипопигий, вид сбоку; 19 — гипандрий, вид сниЗу; 20 — церки, вид сверху; 21 — фаллус, вид сбоку; 22 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="10" pageNumber="506">Fig. 21</figureCitation>
) almost entirely hidden; zigzag bent, strongly constricted beyond middle; its basal portion gently curved, thick at base and becoming slightly narrower towards constriction; apical bend portion broadly semicircular (its tip pointing to rear), long, almost uniformly slender (except slightly thickened extreme base). Ejaculatory apodeme moderately large, extended far beyond basal curvature of phallus, with lateral wings.
</paragraph>
<paragraph id="8BEC76566149FFD8B738FB02C1CCFE95" blockId="10.[824,1423,303,2076]" lastBlockId="11.[189,787,303,472]" lastPageId="11" lastPageNumber="507" pageId="10" pageNumber="506">
<emphasis id="B927AA446149FFD9B738FB02C0EFFAE5" bold="true" box="[862,935,1274,1295]" pageId="10" pageNumber="506">Female</emphasis>
(
<figureCitation id="13686AD36149FFD9B7D3FB02C0B6FAE5" box="[949,1022,1274,1295]" captionStart="Figs 17–22" captionStartId="11.[190,228,1921,1940]" captionTargetBox="[217,1402,535,1842]" captionTargetId="figure-118@11.[202,1412,523,1873]" captionTargetPageId="11" captionText="Figs 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — male: 17 — habitus, holotype, lateral view; 18 — hypopygium, lateral view; 19 — hypandrium, ventral view; 20 — cerci, dorsal view; 21 — phallus, lateral view; 22 — female, habitus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 17–22. Empis (Polyblepharis) subhaemi Shamshev, sp.n.: 17–21 — самец: 17 — габитус, голотип, вид сбоку; 18 — гипопигий, вид сбоку; 19 — гипандрий, вид сниЗу; 20 — церки, вид сверху; 21 — фаллус, вид сбоку; 22 — самка, габитус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="10" pageNumber="506">Fig. 22</figureCitation>
). Similar to male except as follows. Eyes dichoptic, ommatidia of equal size. Frons broad, parallel-sided, with marginal setulae. Occiput with sparser and somewhat stronger setae; also, ocellar setae slightly stronger. Proboscis with labrum nearly 2X longer than eye height. Thorax with shorter setae; acrostichals and dorsocentrals very short; laterotergite mostly with yellow setae, usually with some number of additional black setae (variable, sometimes setae entirely yellow). Fore femur covered with simple, short setulae. Fore tibia with 4–5 short, anterodorsal setae (position and robustness variable) and 1–2 short posteroventral setae near middle. Mid femur without whitish pilosity ventrally; with fine, very short anteroventral and posteroventral setae; bearing fringe of pennate, posteroventral setae (moderately long in about apical 1/3, becoming gradually shorter towards base and absent in about basal 1/4 of femur). Mid tibia slender; dorsal setae as in male, in addition with 3–4 anteroventral and 3–4 posteroventral setae (position and robustness variable). Hind femur slender; with scattered short setae in apical 2/3; bearing pennate posteroventral setae in about apical 2/3 (longer towards apex). Hind tibia bearing fringe of short, pennate, almost uniform, anterodorsal setae (absent near extreme base and close to apex of tibia) and similar, pennate, posteroventral setae in about apical 1/3; in addition, with row of 5–6 short, strong, anterodorsal setae situated just before pennate setae, 1–2 posterodorsal seta(e) in middle part (sometimes absent) and 3–4 short anteroventral setae (slightly shorter than anterodorsals). Wing almost hyaline. Abdomen with tergites 1–6 entirely densely greyish pruinose, tergites 7–8 shiny; covered with mostly minute setulae, only tergites 1–2 with somewhat longer setae laterally, no posteromarginal setae. Sternites 2–6 entirely densely greyish pruinose, with scattered, very short setae; sternite 7 faintly pruinose. Cercus brown, subglobular, covered with dense, pale, erect setulae.
</paragraph>
<paragraph id="8BEC76566148FFD8B485FE70C644FE95" blockId="11.[189,787,303,472]" lastBlockId="11.[824,1421,303,472]" pageId="11" pageNumber="507">
DIFFERENTIAL DIAGNOSIS. The new species is similar to
<taxonomicName id="4C530DD56148FFD8B491FE5DC28EFE50" authority="Loew, 1862" authorityName="Loew" authorityYear="1862" box="[247,454,421,442]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="507" phylum="Arthropoda" rank="species" species="haemi">
<emphasis id="B927AA446148FFD8B491FE5DC24EFE50" box="[247,262,421,442]" italics="true" pageId="11" pageNumber="507">E</emphasis>
.
<emphasis id="B927AA446148FFD8B574FE5DC205FE50" box="[274,333,421,442]" italics="true" pageId="11" pageNumber="507">haemi</emphasis>
Loew, 1862
</taxonomicName>
known from
<collectingCountry id="F34436C66148FFD8B62DFE5DC1D6FE50" box="[587,670,421,442]" name="Bulgaria" pageId="11" pageNumber="507">Bulgaria</collectingCountry>
and
<collectingCountry id="F34436C66148FFD8B6A8FE5DC05AFE50" box="[718,786,421,442]" name="Turkey" pageId="11" pageNumber="507">Turkey</collectingCountry>
[
<bibRefCitation id="EFC20BA76148FFD8B4A5FE3BC20FFE32" author="Chvala M." box="[195,327,451,472]" pageId="11" pageNumber="507" pagination="113 - 225" refId="ref15584" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999</bibRefCitation>
;Çiftçi, Hasbelni, 2007]. However,
<taxonomicName id="4C530DD56148FFD8B6C4FE3BC1B2FE32" box="[674,762,451,472]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="507" phylum="Arthropoda" rank="species" species="haemi">
<emphasis id="B927AA446148FFD8B6C4FE3BC1F9FE32" box="[674,689,451,472]" italics="true" pageId="11" pageNumber="507">E</emphasis>
.
<emphasis id="B927AA446148FFD8B6D9FE3BC1B2FE32" box="[703,762,451,472]" italics="true" pageId="11" pageNumber="507">haemi</emphasis>
</taxonomicName>
is larger (body
<quantity id="4CABDBB36148FFD8B7D2FED7C74EFEAE" box="[948,1030,303,324]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="11" pageNumber="507" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8–9 mm</quantity>
) and has longer proboscis (labrum nearly 3X longer than head height). In addition, in the female of
<taxonomicName id="4C530DD56148FFD8B75EFE92C0C5FE95" box="[824,909,362,383]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="507" phylum="Arthropoda" rank="species" species="haemi">
<emphasis id="B927AA446148FFD8B75EFE92C00FFE95" box="[824,839,362,383]" italics="true" pageId="11" pageNumber="507">E</emphasis>
.
<emphasis id="B927AA446148FFD8B734FE92C0C5FE95" box="[850,909,362,383]" italics="true" pageId="11" pageNumber="507">haemi</emphasis>
</taxonomicName>
the hind basitarsus bears pennate setae.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6148FFD8B738FE70C7D7FE50" pageId="11" pageNumber="507" type="etymology">
<paragraph id="8BEC76566148FFD8B738FE70C7D7FE50" blockId="11.[824,1421,303,472]" pageId="11" pageNumber="507">
ETYMOLOGY. The specific name refers to the similarity of the new species to
<taxonomicName id="4C530DD56148FFD8B061FE5DC7D7FE50" authority="Loew." authorityName="Loew." box="[1031,1183,421,442]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="507" phylum="Arthropoda" rank="species" species="haemi">
<emphasis id="B927AA446148FFD8B061FE5DC75EFE50" box="[1031,1046,421,442]" italics="true" pageId="11" pageNumber="507">E</emphasis>
.
<emphasis id="B927AA446148FFD8B044FE5DC715FE50" box="[1058,1117,421,442]" italics="true" pageId="11" pageNumber="507">haemi</emphasis>
Loew.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6148FFD8B738FE3BC6C5FE32" box="[862,1421,451,472]" pageId="11" pageNumber="507" type="distribution">
<paragraph id="8BEC76566148FFD8B738FE3BC6C5FE32" blockId="11.[824,1421,303,472]" box="[862,1421,451,472]" pageId="11" pageNumber="507">
DISTRIBUTION.
<collectingCountry id="F34436C66148FFD8B061FE3BC729FE32" box="[1031,1121,451,472]" name="Mongolia" pageId="11" pageNumber="507">Mongolia</collectingCountry>
,
<collectingCountry id="F34436C66148FFD8B00CFE3BC7EEFE32" box="[1130,1190,451,472]" name="Russia" pageId="11" pageNumber="507">Russia</collectingCountry>
(Zabaykalskiy Territory).
</paragraph>
</subSubSection>
</treatment>
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<taxonomicName id="4C530DD5614DFFDDB480FBD9C1E1FBD2" authority="Shamshev" authorityName="Shamshev" box="[230,681,1057,1080]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="510" phylum="Arthropoda" rank="species" species="tuvinica" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA44614DFFDDB480FBD9C261FBD2" box="[230,297,1057,1080]" italics="true" pageId="14" pageNumber="510">Empis</emphasis>
(
<emphasis id="B927AA44614DFFDDB55FFBD9C287FBD2" box="[313,463,1057,1080]" italics="true" pageId="14" pageNumber="510">Polyblepharis</emphasis>
)
<emphasis id="B927AA44614DFFDDB5B8FBD9C17CFBD2" box="[478,564,1057,1080]" italics="true" pageId="14" pageNumber="510">tuvinica</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA44614DFFDDB6D5FBD9C1A1FBD2" bold="true" box="[691,745,1057,1080]" pageId="14" pageNumber="510">
<taxonomicNameLabel id="A214173F614DFFDDB6D5FBD9C1A1FBD2" box="[691,745,1057,1080]" pageId="14" pageNumber="510" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614DFFDDB5CFFBC6C16FFBBF" box="[425,551,1086,1109]" pageId="14" pageNumber="510" type="description">
<paragraph id="8BEC7656614DFFDDB5CFFBC6C16FFBBF" blockId="14.[230,745,1057,1109]" box="[425,551,1086,1109]" pageId="14" pageNumber="510">
<figureCitation id="13686AD3614DFFDDB5CFFBC6C16BFBBF" box="[425,547,1086,1109]" captionStart="Figs 29–31" captionStartId="15.[190,228,1929,1948]" captionTargetBox="[211,1394,485,1883]" captionTargetId="figure-79@15.[200,1410,466,1892]" captionTargetPageId="15" captionText="Figs 29–31. Empis (Polyblepharis) tuvinica Shamshev, sp.n., male: 29 — habitus, holotype, lateral view; 30 — hypopygium, lateral view; 31 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 29–31. Empis (Polyblepharis) tuvinica Shamshev, sp.n., самец: 29 — габитус, голотип, вид сбоку; 30 — гипопигий, вид сбоку; 31 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="14" pageNumber="510">Figs 29–31</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614DFFDDB485FB80C23FFAA3" pageId="14" pageNumber="510" type="materials_examined">
<paragraph id="8BEC7656614DFFDDB485FB80C116FB29" blockId="14.[189,788,1144,2076]" pageId="14" pageNumber="510">
<materialsCitation id="3B3B7C0B614DFFDDB485FB80C116FB29" collectionCode="TYPE, MATERIAL, V, VI, ZISP, INS" country="Russia" latitude="52.133335" location="Richter" longLatPrecision="1085" longitude="93.916664" municipality="Turan" pageId="14" pageNumber="510" specimenCount="1" specimenCount-male="1" stateProvince="Tyva" typeStatus="holotype">
<collectionCode id="ED42EE93614DFFDDB485FB80C2E3FB67" box="[227,427,1144,1165]" pageId="14" pageNumber="510">TYPE MATERIAL</collectionCode>
.
<typeStatus id="54E8C8F4614DFFDDB5DBFB80C151FB67" box="[445,537,1144,1165]" pageId="14" pageNumber="510" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D55BDDF614DFFDDB64DFB80C175FB67" box="[555,573,1144,1165]" count="1" pageId="14" pageNumber="510" type="male">♂</specimenCount>
labelled: [
<collectingCountry id="F34436C6614DFFDDB6D2FB80C046FB67" box="[692,782,1144,1165]" name="Russia" pageId="14" pageNumber="510">RUSSIA</collectingCountry>
,
<collectingRegion id="4997B8B4614DFFDDB4DBFB6BC3B8FB42" box="[189,240,1171,1192]" country="Russia" name="Tyva" pageId="14" pageNumber="510">Tuva</collectingRegion>
]
<collectingMunicipality id="6B88EC2C614DFFDDB49CFB6BC27AFB42" box="[250,306,1171,1192]" pageId="14" pageNumber="510">Turan</collectingMunicipality>
[
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<geoCoordinate id="EE671091614DFFDDB5F0FB6BC2ABFB42" box="[406,483,1171,1192]" degrees="93" direction="east" minutes="55" orientation="longitude" pageId="14" pageNumber="510" precision="925" value="93.916664">93°55E</geoCoordinate>
],
<collectingRegion id="4997B8B4614DFFDDB597FB6BC169FB42" box="[497,545,1171,1192]" country="Russia" name="Tyva" pageId="14" pageNumber="510">Tuva</collectingRegion>
/ steppe slopes /
<collectionCode id="ED42EE93614DFFDDB6A5FB6BC19CFB42" box="[707,724,1171,1192]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="510" type="Museum">V</collectionCode>
.
<location id="8E8C208D614DFFDDB6BBFB6BC39EFB29" LSID="urn:lsid:plazi:treatment:03FAC740614DFFC3B480FBD9C16CFA4E:8E8C208D614DFFDDB6BBFB6BC39EFB29" country="Russia" latitude="52.133335" longLatPrecision="1085" longitude="93.916664" municipality="Turan" name="Richter" pageId="14" pageNumber="510" stateProvince="Tyva">Richter</location>
1
<collectionCode id="ED42EE93614DFFDDB488FB56C24FFB29" box="[238,263,1198,1219]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="14" pageNumber="510">VI</collectionCode>
[1]975 (
<collectionCode id="ED42EE93614DFFDDB53CFB56C2C5FB29" box="[346,397,1198,1219]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="14" pageNumber="510">ZISP</collectionCode>
,
<collectionCode id="ED42EE93614DFFDDB5F0FB56C2F4FB29" box="[406,444,1198,1219]" pageId="14" pageNumber="510">INS</collectionCode>
_DIP_0000626).
</materialsCitation>
</paragraph>
<paragraph id="8BEC7656614DFFDDB485FB31C23FFAA3" blockId="14.[189,788,1144,2076]" pageId="14" pageNumber="510">
<materialsCitation id="3B3B7C0B614DFFDDB485FB31C275FB13" collectionCode="ZISP" country="Russia" latitude="52.133335" location="Richter" longLatPrecision="1085" longitude="93.916664" municipality="Turan" pageId="14" pageNumber="510" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Tyva" typeStatus="paratype">
<typeStatus id="54E8C8F4614DFFDDB485FB31C20BFB34" box="[227,323,1225,1246]" pageId="14" pageNumber="510" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F34436C6614DFFDDB536FB31C2E0FB34" box="[336,424,1225,1246]" name="Russia" pageId="14" pageNumber="510">RUSSIA</collectingCountry>
,
<collectingRegion id="4997B8B4614DFFDDB5D2FB31C2A0FB34" box="[436,488,1225,1246]" country="Russia" name="Tyva" pageId="14" pageNumber="510">Tuva</collectingRegion>
: same data as holotype (
<specimenCount id="9D55BDDF614DFFDDB680FB31C044FB34" box="[742,780,1225,1246]" count="1" pageId="14" pageNumber="510" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D55BDDF614DFFDDB4DBFB1CC3BCFB13" box="[189,244,1252,1273]" count="2" pageId="14" pageNumber="510" type="female">2 ♀♀</specimenCount>
,
<collectionCode id="ED42EE93614DFFDDB566FB1CC27DFB13" box="[256,309,1252,1273]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="14" pageNumber="510">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614DFFDDB521FB1CC18AFB13" box="[327,706,1252,1273]" collectingDate="1975-06-05" collectionCode="V" country="Russia" latitude="52.133335" location="Richter" longLatPrecision="1085" longitude="93.916664" municipality="Turan" pageId="14" pageNumber="510" specimenCount="1" stateProvince="Tuva" typeStatus="paratype">
same locality as holotype,
<date id="FFED5096614DFFDDB62CFB1CC1EEFB13" box="[586,678,1252,1273]" pageId="14" pageNumber="510" value="1975-06-05">
<collectingDate id="EFA9A97E614DFFDDB62CFB1CC1EEFB13" box="[586,678,1252,1273]" pageId="14" pageNumber="510" value="1975-06-05">5.vi.1975</collectingDate>
</date>
,
<collectionCode id="ED42EE93614DFFDDB6D7FB1CC18AFB13" box="[689,706,1252,1273]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="510" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B3B7C0B614DFFDDB6AAFB1CC20BFAF9" collectionCode="ZISP" country="Russia" location="Richter" pageId="14" pageNumber="510" specimenCount="2" specimenCount-female="2" stateProvince="Tuva" typeStatus="paratype">
<location id="8E8C208D614DFFDDB6AAFB1CC05BFB13" LSID="urn:lsid:plazi:treatment:03FAC740614DFFC3B480FBD9C16CFA4E:8E8C208D614DFFDDB6AAFB1CC05BFB13" box="[716,787,1252,1273]" country="Russia" name="Richter" pageId="14" pageNumber="510" stateProvince="Tuva">Richter</location>
(
<specimenCount id="9D55BDDF614DFFDDB4A2FB06C3B3FAF9" box="[196,251,1278,1299]" count="2" pageId="14" pageNumber="510" type="female">2 ♀♀</specimenCount>
,
<collectionCode id="ED42EE93614DFFDDB561FB06C274FAF9" box="[263,316,1278,1299]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="14" pageNumber="510">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614DFFDDB52BFB06C18AFAF9" box="[333,706,1278,1299]" collectingDate="1975-06-06" collectionCode="V" country="Russia" latitude="52.133335" location="Richter" longLatPrecision="1085" longitude="93.916664" municipality="Turan" pageId="14" pageNumber="510" specimenCount="1" stateProvince="Tuva" typeStatus="paratype">
same locality as holotype,
<date id="FFED5096614DFFDDB62DFB06C1E0FAF9" box="[587,680,1278,1299]" pageId="14" pageNumber="510" value="1975-06-06">
<collectingDate id="EFA9A97E614DFFDDB62DFB06C1E0FAF9" box="[587,680,1278,1299]" pageId="14" pageNumber="510" value="1975-06-06">6.vi.1975</collectingDate>
</date>
,
<collectionCode id="ED42EE93614DFFDDB6D7FB06C18AFAF9" box="[689,706,1278,1299]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="510" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B3B7C0B614DFFDDB6AAFB06C27DFAC4" collectionCode="ZISP" country="Russia" location="Richter" pageId="14" pageNumber="510" specimenCount="1" specimenCount-female="1" stateProvince="Tuva" typeStatus="paratype">
<location id="8E8C208D614DFFDDB6AAFB06C05BFAF9" LSID="urn:lsid:plazi:treatment:03FAC740614DFFC3B480FBD9C16CFA4E:8E8C208D614DFFDDB6AAFB06C05BFAF9" box="[716,787,1278,1299]" country="Russia" name="Richter" pageId="14" pageNumber="510" stateProvince="Tuva">Richter</location>
(
<specimenCount id="9D55BDDF614DFFDDB4A0FAE1C3A3FAC4" box="[198,235,1305,1326]" count="1" pageId="14" pageNumber="510" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED42EE93614DFFDDB49FFAE1C266FAC4" box="[249,302,1305,1326]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="14" pageNumber="510">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614DFFDDB527FAE1C188FAC4" box="[321,704,1305,1326]" collectingDate="1975-06-07" collectionCode="V" country="Russia" latitude="52.133335" location="Richter" longLatPrecision="1085" longitude="93.916664" municipality="Turan" pageId="14" pageNumber="510" specimenCount="1" stateProvince="Tuva" typeStatus="paratype">
same locality as holotype,
<date id="FFED5096614DFFDDB621FAE1C1ECFAC4" box="[583,676,1305,1326]" pageId="14" pageNumber="510" value="1975-06-07">
<collectingDate id="EFA9A97E614DFFDDB621FAE1C1ECFAC4" box="[583,676,1305,1326]" pageId="14" pageNumber="510" value="1975-06-07">7.vi.1975</collectingDate>
</date>
,
<collectionCode id="ED42EE93614DFFDDB6C9FAE1C188FAC4" box="[687,704,1305,1326]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="510" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B3B7C0B614DFFDDB6AAFAE1C23CFAA3" collectionCode="ZISP" country="Russia" location="Richter" pageId="14" pageNumber="510" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Tuva" typeStatus="paratype">
<location id="8E8C208D614DFFDDB6AAFAE1C05BFAC4" LSID="urn:lsid:plazi:treatment:03FAC740614DFFC3B480FBD9C16CFA4E:8E8C208D614DFFDDB6AAFAE1C05BFAC4" box="[716,787,1305,1326]" country="Russia" name="Richter" pageId="14" pageNumber="510" stateProvince="Tuva">Richter</location>
(
<specimenCount id="9D55BDDF614DFFDDB4A0FACCC3A1FAA3" box="[198,233,1332,1353]" count="1" pageId="14" pageNumber="510" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D55BDDF614DFFDDB493FACCC263FAA3" box="[245,299,1332,1353]" count="2" pageId="14" pageNumber="510" type="female">2 ♀♀</specimenCount>
,
<collectionCode id="ED42EE93614DFFDDB551FACCC225FAA3" box="[311,365,1332,1353]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="14" pageNumber="510">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614DFFDDB485FAB7C26AF99A" pageId="14" pageNumber="510" type="diagnosis">
<paragraph id="8BEC7656614DFFDDB485FAB7C26AF99A" blockId="14.[189,788,1144,2076]" pageId="14" pageNumber="510">
DIAGNOSIS. Mid-sized species (body length about
<quantity id="4CABDBB3614DFFDDB6A8FAB7C040FA8E" box="[718,776,1359,1380]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="510" unit="mm" value="5.0">5 mm</quantity>
); male eyes dichoptic but frons very narrow at middle, narrow- er than anterior ocellus; postpedicel nearly 2.5X longer than basal width; labrum nearly 2X longer than eye height, palpus yellow; mesoscutum with 4 brown vittae, acrostichals biserial, dorsocentrals uniserial, laterotergite with mostly black setae; legs extensively yellow to brownish yellow; abdomen with sternite 8 yellow posteriorly, tergites 2–5 mostly shiny, tergites 6 and 7 with iridescent spots dorsally, tergites 1–5 bearing black posteromarginal setae laterally; female legs without pennate setae.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614DFFC3B485F98DC2FBFAD3" lastPageId="16" lastPageNumber="512" pageId="14" pageNumber="510" type="description">
<paragraph id="8BEC7656614DFFDDB485F98DC784FEB5" blockId="14.[189,788,1144,2076]" lastBlockId="14.[824,1422,303,2076]" pageId="14" pageNumber="510">
DESCRIPTION. Body length about
<quantity id="4CABDBB3614DFFDDB64BF98DC12CF960" box="[557,612,1653,1674]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="510" unit="mm" value="5.0">5 mm</quantity>
, wing
<quantity id="4CABDBB3614DFFDDB6C6F98DC058F960" box="[672,784,1653,1674]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="4.9" metricValueMin="4.7" pageId="14" pageNumber="510" unit="mm" value="4.8" valueMax="4.9" valueMin="4.7">4.7–4.9 mm</quantity>
.
<emphasis id="B927AA44614DFFDDB4DBF968C3B9F94F" bold="true" box="[189,241,1680,1701]" pageId="14" pageNumber="510">Male</emphasis>
(
<figureCitation id="13686AD3614DFFDDB564F968C206F94F" box="[258,334,1680,1701]" captionStart="Figs 29–31" captionStartId="15.[190,228,1929,1948]" captionTargetBox="[211,1394,485,1883]" captionTargetId="figure-79@15.[200,1410,466,1892]" captionTargetPageId="15" captionText="Figs 29–31. Empis (Polyblepharis) tuvinica Shamshev, sp.n., male: 29 — habitus, holotype, lateral view; 30 — hypopygium, lateral view; 31 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 29–31. Empis (Polyblepharis) tuvinica Shamshev, sp.n., самец: 29 — габитус, голотип, вид сбоку; 30 — гипопигий, вид сбоку; 31 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="14" pageNumber="510">Fig. 29</figureCitation>
). Head capsule regions mostly densely greyish pruinose; face on lower margin and entire clypeus shiny; head setation mostly black. Eyes dichoptic, ommatidia equally small. Frons slightly broadened just below ocellar triangle, very narrow before middle (narrower than anterior ocellus), broadened towards antennae on remaining portion; bare. Face broad, bare. Ocellar triangle with 2 long and several short fine setae. Occiput with almost regular row of moderately long setae; postoculars moderately long on upper part and very short laterally; postgena with pale hair-like setae. Antenna with scape and pedicel brown, postpedicel and stylus black; scape slightly longer than subglobular pedicel, both with very short setulae; postpedicel nearly 2.5X as long as basal width, with straight margins; stylus slightly longer than postpedicel basal width. Proboscis with labrum brownish yellow, nearly 2X longer than eye height; palpus short, mostly yellow, brownish close to base; with scattered, dark setulae.
</paragraph>
<paragraph id="8BEC7656614DFFDDB738FE9DC72CFCB5" blockId="14.[824,1422,303,2076]" pageId="14" pageNumber="510">Thorax black in ground-colour, mostly densely greyish pruinose and black setose; mesoscutum with 4 brown vittae (lateral vittae less distinct). Antepronotum with 4–5 black setae dorsally on each side. Postpronotal lobe with 1 long and several shorter setae of different lengths (some of them only slightly shorter than postpronotal seta). Proepisternum with tuft of several, pale, fine setae on lower part. Prosternum bare. Mesonotal setae: acrostichals arranged in 2 close irregular rows, short, absent on prescutellar depression; dorsocentral setae uniserial, presuturals rather long (slightly longer than antennal stylus), 3 long prescutellars; 1 presutural intra-alar (nearly as long as presutural dorsocentrals), 1 long presutural supra-alar, 3–4 notopleurals, 1 postsutural supra-alar (with 1–2 short, fine seta(e) anteriorly), 1 long and 1 minute postalars, 4 scutellars (lateral setae only slightly shorter than apicals); Laterotergite usually with a few strong black setae (sometimes some setae brownish to brownish yellow and additional short, fine, pale setulae present). Anterior and posterior spiracles yellow.</paragraph>
<paragraph id="8BEC7656614DFFDDB738FC9DC7F7FA21" blockId="14.[824,1422,303,2076]" pageId="14" pageNumber="510">Legs with coxae densely greyish pruinose, remaining podomeres subshiny; mostly black setose; Leg colour: extensively yellow to brownish yellow; coxae (except apex), most part of fore femur (except apex and sometimes extreme base), tarsomere 1 at apex, most part of tarsomere 2 (except base) and entire tarsomeres 3–5 brownish. Fore coxa with scattered strong setae and pale setulae anteriorly. All femora whitish pilose ventrally. Fore femur with rows of minute anteroventral and posteroventral setae. Fore tibia with 2–4 short, anterodorsal setae (position and robustness variable). Mid femur with short setae in about apical half anterodorsally and anteriorly; bearing rows of minute, fine anteroventral and posteroventral setae becoming slightly longer and denser close to base; with similar setae on ventral face near base. Mid tibia rather slightly thickened; with 2–4 anterodorsal and 2–3 posterodorsal, short, strong setae (position and robustness variable). Hind femur slender; bearing complete row of short, numerous, anteroventral setae (longest setae shorter than half of femur maximal width); covered with short, finer setae posteroventrally. Hind tibia simple, slender; with 2–3 anterodorsal and 3 posterodorsal, moderately long setae; simple setulae ventrally; no seta in posteroapical comb. Tarsomeres of all legs slender; basitarsi with scattered, spine-like setae ventrally.</paragraph>
<paragraph id="8BEC7656614DFFDDB738FA29C726F933" blockId="14.[824,1422,303,2076]" pageId="14" pageNumber="510">Wing membrane faintly brownish infuscate; entirely covered with microtrichia; veins yellowish brown, well-sclerotized. Veins R 5 and M 1 almost parallel towards wing margin; radial fork acute; R 5 meeting costa before wing apex; CuA+CuP complete (slightly weakened beyond middle). Cell dm moderately large, with strongly elongate apex; apical portion of M 4 nearly 3X longer than its middle portion. Pterostigma inconspicuous. Basal costal seta absent. Anal lobe well-developed; axillary incision nearly 90°. Squama yellow, pale fringed. Halter pale yellow.</paragraph>
<paragraph id="8BEC7656614DFFC3B738F927C2E6FEB5" blockId="14.[824,1422,303,2076]" lastBlockId="16.[189,787,303,1444]" lastPageId="16" lastPageNumber="512" pageId="14" pageNumber="510">Abdomen almost entirely brown in ground colour (tergites of pregenital segments somewhat darker), only sternite 8 yellow posteriorly (size of yellow space variable). Pruinescence: tergite 1 entirely densely greyish pruinose; tergites 2–5 mostly shiny (narrowly faintly pruinose anteriorly); tergites 6–7 with yellowish grey iridescent spot dorsally, shiny laterally (dorsal view); sternites 1–7 densely greyish pruinose, sternite 8 faintly pruinose. Chaetotaxy: tergites 1–5 with black and pale setulae dorsally and longer, fine, pale setae laterally, bearing long, black posteromarginal setae laterally (sometimes some posteromarginal setae brownish yellow to yellow); tergites 6–7 with scattered setulae; sternite 1 bare, sternites 2–7 with short, pale setae, sternite 8 with black setae (posteromarginals moderatelylong).Pregenitalsegments:segment 6unmodified;tergite 7 broadly concave posteriorly, sternite 7 unmodified; segment 8 with separated tergite and sternite; tergite 8 somewhat flattened, represented by two subglobular sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 large, scoop-shaped, somewhat constricted anteriorly, with 2 small, antero-lateral projections separated by depression and larger, subconical projection on posterior margin of depression (slightly curved at apex).</paragraph>
<caption id="DF2C26DE614CFFDCB4D8F871C210F7FE" pageId="15" pageNumber="511" startId="15.[190,228,1929,1948]" targetBox="[211,1394,485,1883]" targetPageId="15" targetType="figure">
<paragraph id="8BEC7656614CFFDCB4D8F871C226F826" blockId="15.[190,1424,1929,2068]" pageId="15" pageNumber="511">
<emphasis id="B927AA44614CFFDCB4D8F871C26CF876" bold="true" box="[190,292,1929,1948]" pageId="15" pageNumber="511">Figs 29–31.</emphasis>
<taxonomicName id="4C530DD5614CFFDCB54FF871C1D9F876" authority="Shamshev" authorityName="Shamshev" box="[297,657,1929,1948]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="511" phylum="Arthropoda" rank="species" species="tuvinica" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA44614CFFDCB54FF871C216F876" box="[297,350,1929,1948]" italics="true" pageId="15" pageNumber="511">Empis</emphasis>
(
<emphasis id="B927AA44614CFFDCB50DF871C2ABF876" box="[363,483,1929,1948]" italics="true" pageId="15" pageNumber="511">Polyblepharis</emphasis>
)
<emphasis id="B927AA44614CFFDCB588F871C17BF876" box="[494,563,1929,1948]" italics="true" pageId="15" pageNumber="511">tuvinica</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA44614CFFDCB6FFF871C18CF876" bold="true" box="[665,708,1929,1948]" pageId="15" pageNumber="511">
<taxonomicNameLabel id="A214173F614CFFDCB6FFF871C18CF876" box="[665,708,1929,1948]" pageId="15" pageNumber="511" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, male: 29 — habitus, holotype, lateral view; 30 — hypopygium, lateral view; 31 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm.
</paragraph>
<paragraph id="8BEC7656614CFFDCB4D8F829C210F7FE" blockId="15.[190,1424,1929,2068]" pageId="15" pageNumber="511">
<emphasis id="B927AA44614CFFDCB4D8F829C26FF80E" bold="true" box="[190,295,2001,2020]" pageId="15" pageNumber="511">Рис. 29–31.</emphasis>
<taxonomicName id="4C530DD5614CFFDCB54AF829C1DFF80E" authority="Shamshev" authorityName="Shamshev" box="[300,663,2001,2020]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="511" phylum="Arthropoda" rank="species" species="tuvinica" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA44614CFFDCB54AF829C229F80E" box="[300,353,2001,2020]" italics="true" pageId="15" pageNumber="511">Empis</emphasis>
(
<emphasis id="B927AA44614CFFDCB509F829C2AFF80E" box="[367,487,2001,2020]" italics="true" pageId="15" pageNumber="511">Polyblepharis</emphasis>
)
<emphasis id="B927AA44614CFFDCB595F829C170F80E" box="[499,568,2001,2020]" italics="true" pageId="15" pageNumber="511">tuvinica</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA44614CFFDCB6F9F829C182F80E" bold="true" box="[671,714,2001,2020]" pageId="15" pageNumber="511">
<taxonomicNameLabel id="A214173F614CFFDCB6F9F829C182F80E" box="[671,714,2001,2020]" pageId="15" pageNumber="511" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, самец: 29 — габитус, голотип, вид сбоку; 30 — гипопигий, вид сбоку; 31 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм.
</paragraph>
</caption>
<paragraph id="8BEC76566153FFC3B485FE9CC14FFC9E" blockId="16.[189,787,303,1444]" pageId="16" pageNumber="512">
Hypopygium (
<figureCitation id="13686AD36153FFC3B50BFE9CC290FE93" box="[365,472,356,377]" captionStart="Figs 29–31" captionStartId="15.[190,228,1929,1948]" captionTargetBox="[211,1394,485,1883]" captionTargetId="figure-79@15.[200,1410,466,1892]" captionTargetPageId="15" captionText="Figs 29–31. Empis (Polyblepharis) tuvinica Shamshev, sp.n., male: 29 — habitus, holotype, lateral view; 30 — hypopygium, lateral view; 31 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 29–31. Empis (Polyblepharis) tuvinica Shamshev, sp.n., самец: 29 — габитус, голотип, вид сбоку; 30 — гипопигий, вид сбоку; 31 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="16" pageNumber="512">Figs 30, 31</figureCitation>
) moderately large; yellow (epandrium mostly brownish yellow in
<typeStatus id="54E8C8F46153FFC3B663FE87C111FE7E" box="[517,601,383,404]" pageId="16" pageNumber="512" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D55BDDF6153FFC3B6EEFE87C040FE7E" box="[648,776,383,404]" count="1" pageId="16" pageNumber="512" type="generic" typeStatus="paratype">one paratype</specimenCount>
); epandrial lamella with short, black setae more numerous along lower margin; cercus covered with black setulae. Epandrium entirely broadly divided (epandrial bridge absent); epandrial lamella (
<figureCitation id="13686AD36153FFC3B571FE12C22BFE15" box="[279,355,490,511]" captionStart="Figs 29–31" captionStartId="15.[190,228,1929,1948]" captionTargetBox="[211,1394,485,1883]" captionTargetId="figure-79@15.[200,1410,466,1892]" captionTargetPageId="15" captionText="Figs 29–31. Empis (Polyblepharis) tuvinica Shamshev, sp.n., male: 29 — habitus, holotype, lateral view; 30 — hypopygium, lateral view; 31 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 29–31. Empis (Polyblepharis) tuvinica Shamshev, sp.n., самец: 29 — габитус, голотип, вид сбоку; 30 — гипопигий, вид сбоку; 31 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="16" pageNumber="512">Fig. 30</figureCitation>
) subtriangular (lateral view). Hypandrium separated from epandrium; mostly membranous, undivided, narrowly sclerotized along margin, subtriangular in ventral view; bare; gonocoxal apodeme small. Cerci separated with each other and from epandrium; cercus elongate ovate (lateral view), shorter than epandrial lamella, somewhat lobe-like produced at about basal half dorsally (dorsal view), with 2 small, pointed projections beyond middle of inner margin. Phallus (
<figureCitation id="13686AD36153FFC3B4A2FD47C246FD3E" box="[196,270,703,724]" captionStart="Figs 29–31" captionStartId="15.[190,228,1929,1948]" captionTargetBox="[211,1394,485,1883]" captionTargetId="figure-79@15.[200,1410,466,1892]" captionTargetPageId="15" captionText="Figs 29–31. Empis (Polyblepharis) tuvinica Shamshev, sp.n., male: 29 — habitus, holotype, lateral view; 30 — hypopygium, lateral view; 31 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 29–31. Empis (Polyblepharis) tuvinica Shamshev, sp.n., самец: 29 — габитус, голотип, вид сбоку; 30 — гипопигий, вид сбоку; 31 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="16" pageNumber="512">Fig. 31</figureCitation>
) almost entirely hidden; zigzag bent, strongly constricted beyond middle; its basal portion gently curved, thick at base and becoming gradually narrower towards constriction; apical bend portion semicircular (its tip pointing to rear), rather mostly thicken, with short, slender subapical part. Ejaculatory apodeme moderately large, extended far beyond basal curvature of phallus, with lateral wings.
</paragraph>
<paragraph id="8BEC76566153FFC3B485FC82C139FBA3" blockId="16.[189,787,303,1444]" pageId="16" pageNumber="512">
<emphasis id="B927AA446153FFC3B485FC82C27BFC65" bold="true" box="[227,307,890,911]" pageId="16" pageNumber="512">Female.</emphasis>
Similar to male except as follows. Frons broad, slightly broadened towards antennae; with scattered marginal setulae. Mid tibia slender; hind femur slightly thickened. Mid and hind femora mostly covered with minute setulae ventrally; hind femur with 4–5 short anteroventral setae in about apical 1/3. Abdomen almost entirely greyish pruinose; tergite 8 black, sternite 8 brownish yellow, both sclerites shiny. Cercus brown, short, covered with dense, pale, erect setulae.
</paragraph>
<paragraph id="8BEC76566153FFC3B485FBB7C2FBFAD3" blockId="16.[189,787,303,1444]" pageId="16" pageNumber="512">
DIFFERENTIAL DIAGNOSIS. The new species is similar to
<taxonomicName id="4C530DD56153FFC3B49AFB92C15AFB95" authority="Shamshev" authorityName="Shamshev" authorityYear="2006" box="[252,530,1130,1151]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="512" phylum="Arthropoda" rank="species" species="transbaicalica">
<emphasis id="B927AA446153FFC3B49AFB92C243FB95" box="[252,267,1130,1151]" italics="true" pageId="16" pageNumber="512">E</emphasis>
.
<emphasis id="B927AA446153FFC3B57DFB92C2EEFB95" box="[283,422,1130,1151]" italics="true" pageId="16" pageNumber="512">transbaicalica</emphasis>
Shamshev
</taxonomicName>
and
<taxonomicName id="4C530DD56153FFC3B62EFB92C05AFB95" box="[584,786,1130,1151]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="512" phylum="Arthropoda" rank="species" species="subtransbaicalica" status="sp.n.">
<emphasis id="B927AA446153FFC3B62EFB92C11FFB95" box="[584,599,1130,1151]" italics="true" pageId="16" pageNumber="512">E</emphasis>
.
<emphasis id="B927AA446153FFC3B600FB92C05AFB95" box="[614,786,1130,1151]" italics="true" pageId="16" pageNumber="512">subtransbaicalica</emphasis>
</taxonomicName>
<emphasis id="B927AA446153FFC3B4DBFB7CC3A5FB73" bold="true" box="[189,237,1156,1177]" pageId="16" pageNumber="512">
<taxonomicNameLabel id="A214173F6153FFC3B4DBFB7CC3A5FB73" box="[189,237,1156,1177]" pageId="16" pageNumber="512" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
described herein.
<taxonomicName id="4C530DD56153FFC3B5D3FB7CC1C2FB73" authorityName="Shamshev" authorityYear="2006" box="[437,650,1156,1177]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="512" phylum="Arthropoda" rank="species" species="transbaicalica">
<emphasis id="B927AA446153FFC3B5D3FB7CC1C2FB73" box="[437,650,1156,1177]" italics="true" pageId="16" pageNumber="512">Empis transbaicalica</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C530DD56153FFC3B4DBFB67C26BFB5E" box="[189,291,1183,1204]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="512" phylum="Arthropoda" rank="species" species="tuvinica" status="sp.n.">
<emphasis id="B927AA446153FFC3B4DBFB67C384FB5E" box="[189,204,1183,1204]" italics="true" pageId="16" pageNumber="512">E</emphasis>
.
<emphasis id="B927AA446153FFC3B4B0FB67C26BFB5E" box="[214,291,1183,1204]" italics="true" pageId="16" pageNumber="512">tuvinica</emphasis>
</taxonomicName>
<emphasis id="B927AA446153FFC3B541FB67C21FFB5E" bold="true" box="[295,343,1183,1204]" pageId="16" pageNumber="512">
<taxonomicNameLabel id="A214173F6153FFC3B541FB67C21FFB5E" box="[295,343,1183,1204]" pageId="16" pageNumber="512" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
primarily by broader frons of the male (slightly broader than anterior ocellus) and by the absence of an iridescent spot on the male abdominal tergite 6 [
<bibRefCitation id="EFC20BA76153FFC3B605FB2CC041FB03" author="Shamshev I. V." box="[611,777,1236,1257]" pageId="16" pageNumber="512" pagination="231 - 264" refId="ref15746" refString="Shamshev I. V. 2006. Revision of the genus Empis Linnaeus (Diptera: Empididae) from Russia and neighbouring lands. 3. Descriptions of thirteen new species of the subgenus Polyblepharis Bezzi // International Journal of Dipterological Research. Vol. 17. P. 231 - 264." type="journal article" year="2006">Shamshev, 2006</bibRefCitation>
].
<taxonomicName id="4C530DD56153FFC3B4DBFB17C2E6FAEE" box="[189,430,1263,1284]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="512" phylum="Arthropoda" rank="species" species="subtransbaicalica" status="sp. nov.">
<emphasis id="B927AA446153FFC3B4DBFB17C2E6FAEE" box="[189,430,1263,1284]" italics="true" pageId="16" pageNumber="512">Empis subtransbaicalica</emphasis>
</taxonomicName>
<emphasis id="B927AA446153FFC3B5D1FB17C2AFFAEE" bold="true" box="[439,487,1263,1284]" pageId="16" pageNumber="512">
<taxonomicNameLabel id="A214173F6153FFC3B5D1FB17C2AFFAEE" box="[439,487,1263,1284]" pageId="16" pageNumber="512" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C530DD56153FFC3B608FB17C191FAEE" box="[622,729,1263,1284]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="512" phylum="Arthropoda" rank="species" species="tuvinica" status="sp.n.">
<emphasis id="B927AA446153FFC3B608FB17C135FAEE" box="[622,637,1263,1284]" italics="true" pageId="16" pageNumber="512">E</emphasis>
.
<emphasis id="B927AA446153FFC3B6EAFB17C191FAEE" box="[652,729,1263,1284]" italics="true" pageId="16" pageNumber="512">tuvinica</emphasis>
</taxonomicName>
<emphasis id="B927AA446153FFC3B684FB17C05AFAEE" bold="true" box="[738,786,1263,1284]" pageId="16" pageNumber="512">
<taxonomicNameLabel id="A214173F6153FFC3B684FB17C05AFAEE" box="[738,786,1263,1284]" pageId="16" pageNumber="512" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
primarily by pale laterotergal setae, 2 scutellar setae (sian 4) and pale setose abdomen.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6153FFC3B485FAC7C204FA63" pageId="16" pageNumber="512" type="etymology">
<paragraph id="8BEC76566153FFC3B485FAC7C204FA63" blockId="16.[189,787,303,1444]" pageId="16" pageNumber="512">
ETYMOLOGY. The specific epithet refers to the territory of the origin of the new species,
<collectingRegion id="4997B8B46153FFC3B595FAA2C16BFA85" box="[499,547,1370,1391]" country="Russia" name="Tyva" pageId="16" pageNumber="512">Tuva</collectingRegion>
(
<collectingRegion id="4997B8B46153FFC3B656FAA2C12AFA85" box="[560,610,1370,1391]" country="Russia" name="Tyva" pageId="16" pageNumber="512">Tyva</collectingRegion>
) Republic of
<collectingCountry id="F34436C66153FFC3B683FAA2C390FA63" name="Russia" pageId="16" pageNumber="512">Russia</collectingCountry>
Federation.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD6153FFC3B485FA77C16CFA4E" box="[227,548,1423,1444]" pageId="16" pageNumber="512" type="distribution">
<paragraph id="8BEC76566153FFC3B485FA77C16CFA4E" blockId="16.[189,787,303,1444]" box="[227,548,1423,1444]" pageId="16" pageNumber="512">
DISTRIBUTION.
<collectingCountry id="F34436C66153FFC3B5F1FA77C29FFA4E" box="[407,471,1423,1444]" name="Russia" pageId="16" pageNumber="512">Russia</collectingCountry>
(
<collectingRegion id="4997B8B46153FFC3B582FA77C152FA4E" box="[484,538,1423,1444]" country="Russia" name="Tyva" pageId="16" pageNumber="512">Tuva</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:detail id="E4D561116CD44A0A185065134A94862D" type="issue">
<mods:number id="AF5EE7B68D9578ADF07DF13EA1082526">3</mods:number>
</mods:detail>
<mods:extent id="82717F8FAA4F2910145E06B0A2C5450F" unit="page">
<mods:start id="4324A0AEE7AF29BAC8E622A29604FA5C">496</mods:start>
<mods:end id="F8ABA40A3405BC8DEC3FD4A5A1C86B34">512</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="CE3ECF6A5643BB6541BA6E073E138310">journal article</mods:classification>
<mods:identifier id="0C6AAACF76B8A1CD14278679D215E482" type="DOI">10.15298/rusentj.33.4.13</mods:identifier>
<mods:identifier id="34C786C0E5AE8A44F47E0A0C24A9554A" type="ISSN">0132-8069</mods:identifier>
</mods:mods>
<treatment id="03FAC740614FFFDDB4B6FED7C23EFC17" LSID="urn:lsid:plazi:treatment:03FAC740614FFFDDB4B6FED7C23EFC17" httpUri="http://treatment.plazi.org/id/03FAC740614FFFDDB4B6FED7C23EFC17" lastPageId="14" lastPageNumber="510" pageId="12" pageNumber="508">
<subSubSection id="C34925DD614FFFDFB4B6FED7C14AFE8E" pageId="12" pageNumber="508" type="nomenclature">
<paragraph id="8BEC7656614FFFDFB4B6FED7C14AFE8E" blockId="12.[208,768,303,385]" pageId="12" pageNumber="508">
<heading id="D0A4C13A614FFFDFB4B6FED7C14AFE8E" centered="true" fontSize="10" level="2" pageId="12" pageNumber="508" reason="8">
<taxonomicName id="4C530DD5614FFFDFB4B6FED7C1B4FEAC" authority="Shamshev" authorityName="Shamshev" box="[208,764,303,326]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="508" phylum="Arthropoda" rank="species" species="subtransbaicalica" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA44614FFFDFB4B6FED7C25BFEAC" box="[208,275,303,326]" italics="true" pageId="12" pageNumber="508">Empis</emphasis>
(
<emphasis id="B927AA44614FFFDFB544FED7C2F0FEAC" box="[290,440,303,326]" italics="true" pageId="12" pageNumber="508">Polyblepharis</emphasis>
)
<emphasis id="B927AA44614FFFDFB5A1FED7C1CEFEAC" box="[455,646,303,326]" italics="true" pageId="12" pageNumber="508">subtransbaicalica</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA44614FFFDFB5ABFEB5C14AFE8E" bold="true" box="[461,514,333,356]" pageId="12" pageNumber="508">
<taxonomicNameLabel id="A214173F614FFFDFB5ABFEB5C14AFE8E" box="[461,514,333,356]" pageId="12" pageNumber="508" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614FFFDFB5CFFE92C16FFE6B" box="[425,551,362,385]" pageId="12" pageNumber="508" type="description">
<paragraph id="8BEC7656614FFFDFB5CFFE92C16FFE6B" blockId="12.[208,768,303,385]" box="[425,551,362,385]" pageId="12" pageNumber="508">
<figureCitation id="13686AD3614FFFDFB5CFFE92C16BFE6B" box="[425,547,362,385]" captionStart="Figs 23–28" captionStartId="13.[190,228,1921,1940]" captionTargetBox="[208,1396,556,1859]" captionTargetId="figure-106@13.[200,1410,536,1874]" captionTargetPageId="13" captionText="Figs 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., male: 23 — habitus, holotype, lateral view; 24 — head and thorax, holotype, anterodorsal view; 25 — hypopygium, lateral view; 26 — hypandrium, ventral view; 27 — cerci, dorsal view; 28 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., самец: 23 — габитус, голотип, вид сбоку; 24 — голова и грудь, голотип, вид спереди-сверху; 25 — гипопигий, вид сбоку; 26 — гипандрий, вид сниЗу; 27 — церки, вид сверху; 28 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="12" pageNumber="508">Figs 23–28</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614FFFDFB485FE5CC1B6FB42" pageId="12" pageNumber="508" type="materials_examined">
<paragraph id="8BEC7656614FFFDFB485FE5CC1D5FDE0" blockId="12.[189,788,420,2076]" pageId="12" pageNumber="508">
<materialsCitation id="3B3B7C0B614FFFDFB485FE5CC1D5FDE0" collectionCode="TYPE, MATERIAL, ZISP, INS" country="Mongolia" county="Ara-Khangayskiy" latitude="47.466667" location="Nartshuk" longLatPrecision="1116" longitude="101.45" municipality="Tsetserleg" pageId="12" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Arhangay" typeStatus="holotype">
<collectionCode id="ED42EE93614FFFDFB485FE5CC2E1FE53" box="[227,425,420,441]" pageId="12" pageNumber="508">TYPE MATERIAL</collectionCode>
.
<typeStatus id="54E8C8F4614FFFDFB5DEFE5CC15CFE53" box="[440,532,420,441]" pageId="12" pageNumber="508" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D55BDDF614FFFDFB642FE5CC17EFE53" box="[548,566,420,441]" count="1" pageId="12" pageNumber="508" type="male">♂</specimenCount>
labelled [in Cyrillic, Russian]:
<collectingCountry id="F34436C6614FFFDFB579FE47C237FE3E" box="[287,383,447,468]" name="Mongolia" pageId="12" pageNumber="508">Mongolia</collectingCountry>
,
<collectingCounty id="628D0EDA614FFFDFB5EFFE47C17CFE3E" box="[393,564,447,468]" pageId="12" pageNumber="508">Ara-Khangayskiy</collectingCounty>
/ aymak [=
<collectingRegion id="4997B8B4614FFFDFB6C2FE47C05AFE3E" box="[676,786,447,468]" country="Mongolia" name="Arhangay" pageId="12" pageNumber="508">Arkhangay</collectingRegion>
aymag (province)],
<quantity id="4CABDBB3614FFFDFB51DFE22C2F0FE05" box="[379,440,474,495]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="508" unit="km" value="30.0">30 km</quantity>
E /
<collectingMunicipality id="6B88EC2C614FFFDFB5B8FE22C177FE05" box="[478,575,474,495]" pageId="12" pageNumber="508">Tsetserleg</collectingMunicipality>
[~
<geoCoordinate id="EE671091614FFFDFB63EFE22C1E2FE05" box="[600,682,474,495]" degrees="47" direction="north" minutes="28" orientation="latitude" pageId="12" pageNumber="508" precision="925" value="47.466667">47°28′N</geoCoordinate>
<geoCoordinate id="EE671091614FFFDFB6D7FE22C044FE05" box="[689,780,474,495]" degrees="101" direction="east" minutes="27" orientation="longitude" pageId="12" pageNumber="508" precision="925" value="101.45">101°27′E</geoCoordinate>
] /
<location id="8E8C208D614FFFDFB4ACFE0DC26AFDE0" LSID="urn:lsid:plazi:treatment:03FAC740614FFFDDB4B6FED7C23EFC17:8E8C208D614FFFDFB4ACFE0DC26AFDE0" box="[202,290,501,522]" country="Mongolia" county="Ara-Khangayskiy" latitude="47.466667" longLatPrecision="1116" longitude="101.45" municipality="Tsetserleg" name="Nartshuk" pageId="12" pageNumber="508" stateProvince="Arhangay">Nartshuk</location>
2.4.[1]975 (
<collectionCode id="ED42EE93614FFFDFB5FEFE0DC283FDE0" box="[408,459,501,522]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="12" pageNumber="508">ZISP</collectionCode>
,
<collectionCode id="ED42EE93614FFFDFB5B2FE0DC2B2FDE0" box="[468,506,501,522]" pageId="12" pageNumber="508">INS</collectionCode>
_DIP_0000625).
</materialsCitation>
</paragraph>
<paragraph id="8BEC7656614FFFDFB485FDF7C1B6FB42" blockId="12.[189,788,420,2076]" pageId="12" pageNumber="508">
<materialsCitation id="3B3B7C0B614FFFDFB485FDF7C1DCFDD5" collectorName="Bogdo-ula" country="Mongolia" location="Tsentralny" pageId="12" pageNumber="508" specimenCount="1" stateProvince="Tov" typeStatus="paratype">
<typeStatus id="54E8C8F4614FFFDFB485FDF7C20BFDCE" box="[227,323,527,548]" pageId="12" pageNumber="508" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F34436C6614FFFDFB52BFDF7C29AFDCE" box="[333,466,527,548]" name="Mongolia" pageId="12" pageNumber="508">MONGOLIA</collectingCountry>
,
<collectingRegion id="4997B8B4614FFFDFB5BAFDF7C102FDCE" box="[476,586,527,548]" country="Mongolia" name="Tov" pageId="12" pageNumber="508">Töv aymag</collectingRegion>
[=
<location id="8E8C208D614FFFDFB605FDF7C184FDCE" LSID="urn:lsid:plazi:treatment:03FAC740614FFFDDB4B6FED7C23EFC17:8E8C208D614FFFDFB605FDF7C184FDCE" box="[611,716,527,548]" country="Mongolia" name="Tsentralny" pageId="12" pageNumber="508" stateProvince="Tov">Tsentralny</location>
aymak (province)]: northern slope of
<collectorName id="26A61380614FFFDFB59CFDD2C117FDD5" box="[506,607,554,575]" pageId="12" pageNumber="508">Bogdo-ula</collectorName>
near
</materialsCitation>
<materialsCitation id="3B3B7C0B614FFFDFB6C7FDD2C144FDB0" collectingDate="1967-06-22" collectorName="Zaitzev" country="Mongolia" location="Ulan-Bator" pageId="12" pageNumber="508" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Ulaanbaatar" typeStatus="paratype">
<collectingRegion id="4997B8B4614FFFDFB6C7FDD2C047FDD5" box="[673,783,554,575]" country="Mongolia" name="Ulaanbaatar" pageId="12" pageNumber="508">Ulan-Bator</collectingRegion>
,
<date id="FFED5096614FFFDFB4DBFDBDC26EFDB0" box="[189,294,581,602]" pageId="12" pageNumber="508" value="1967-06-22">
<collectingDate id="EFA9A97E614FFFDFB4DBFDBDC26EFDB0" box="[189,294,581,602]" pageId="12" pageNumber="508" value="1967-06-22">22.vi.1967</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB557FDBDC231FDB0" box="[305,377,581,602]" pageId="12" pageNumber="508">Zaitzev</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB5EEFDBDC2F7FDB0" box="[392,447,581,602]" count="3" pageId="12" pageNumber="508" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D55BDDF614FFFDFB5AAFDBDC14BFDB0" box="[460,515,581,602]" count="2" pageId="12" pageNumber="508" type="female">2 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614FFFDFB67FFDBDC2EAFD9F" collectingDate="1967-06-29" collectorName="Zaitzev" country="Mongolia" location="Ulan-Bator" pageId="12" pageNumber="508" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Ulaanbaatar" typeStatus="paratype">
same locality,
<date id="FFED5096614FFFDFB6C3FDBDC046FDB0" box="[677,782,581,602]" pageId="12" pageNumber="508" value="1967-06-29">
<collectingDate id="EFA9A97E614FFFDFB6C3FDBDC046FDB0" box="[677,782,581,602]" pageId="12" pageNumber="508" value="1967-06-29">29.vi.1967</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB4DBFD98C24DFD9F" box="[189,261,608,629]" pageId="12" pageNumber="508">Zaitzev</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB570FD98C218FD9F" box="[278,336,608,629]" count="4" pageId="12" pageNumber="508" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="9D55BDDF614FFFDFB506FD98C2D1FD9F" box="[352,409,608,629]" count="2" pageId="12" pageNumber="508" type="female">2 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614FFFDFB5D4FD98C3BFFD7A" collectingDate="1967-06-29" collectorName="Kerzhner" country="Mongolia" location="Ulan-Bator" pageId="12" pageNumber="508" specimenCount="1" specimenCount-female="1" stateProvince="Ulaanbaatar" typeStatus="paratype">
same locality,
<date id="FFED5096614FFFDFB625FD98C1E3FD9F" box="[579,683,608,629]" pageId="12" pageNumber="508" value="1967-06-29">
<collectingDate id="EFA9A97E614FFFDFB625FD98C1E3FD9F" box="[579,683,608,629]" pageId="12" pageNumber="508" value="1967-06-29">29.vi.1967</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB6DFFD98C05AFD9F" box="[697,786,608,629]" pageId="12" pageNumber="508">Kerzhner</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB4A0FD83C3A6FD7A" box="[198,238,635,656]" count="1" pageId="12" pageNumber="508" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614FFFDFB561FD83C131FD41" collectingDate="1967-06-15" collectorName="Kerzhner" country="Mongolia" location="Zaysan" municipality="Bogdo-ula Mt." pageId="12" pageNumber="508" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
northern slope of
<collectingMunicipality id="6B88EC2C614FFFDFB5A6FD83C11DFD7A" box="[448,597,635,656]" pageId="12" pageNumber="508">Bogdo-ula Mt.</collectingMunicipality>
,
<location id="8E8C208D614FFFDFB605FD83C1E3FD7A" LSID="urn:lsid:plazi:treatment:03FAC740614FFFDDB4B6FED7C23EFC17:8E8C208D614FFFDFB605FD83C1E3FD7A" box="[611,683,635,656]" country="Mongolia" municipality="Bogdo-ula Mt." name="Zaysan" pageId="12" pageNumber="508">Zaysan</location>
, meadowsteppe slopes,
<date id="FFED5096614FFFDFB534FD6EC2F2FD41" box="[338,442,662,683]" pageId="12" pageNumber="508" value="1967-06-15">
<collectingDate id="EFA9A97E614FFFDFB534FD6EC2F2FD41" box="[338,442,662,683]" pageId="12" pageNumber="508" value="1967-06-15">15.vi.1967</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB5AFFD6EC16AFD41" box="[457,546,662,683]" pageId="12" pageNumber="508">Kerzhner</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB653FD6EC138FD41" box="[565,624,662,683]" count="3" pageId="12" pageNumber="508" type="male">3 ♂♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614FFFDFB6ECFD6EC146FD2F" collectingDate="1967-06-15" collectorName="Zaitzev" country="Mongolia" location="Zaysan" municipality="Bogdo-ula Mt." pageId="12" pageNumber="508" specimenCount="6" specimenCount-female="1" specimenCount-male="5" typeStatus="paratype">
same locality,
<date id="FFED5096614FFFDFB4DBFD48C26DFD2F" box="[189,293,688,709]" pageId="12" pageNumber="508" value="1967-06-15">
<collectingDate id="EFA9A97E614FFFDFB4DBFD48C26DFD2F" box="[189,293,688,709]" pageId="12" pageNumber="508" value="1967-06-15">15.vi.1967</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB553FD48C235FD2F" box="[309,381,688,709]" pageId="12" pageNumber="508">Zaitzev</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB5F6FD48C283FD2F" box="[400,459,688,709]" count="5" pageId="12" pageNumber="508" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D55BDDF614FFFDFB5BAFD48C14DFD2F" box="[476,517,688,709]" count="1" pageId="12" pageNumber="508" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614FFFDFB679FD48C17CFD0A" collectingDate="1970-06-24" collectingDateMax="1970-06-29" collectingDateMin="1970-06-24" collectorName="Nartshuk" country="Mongolia" location="Ulan-Bator" pageId="12" pageNumber="508" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Ulaanbaatar" typeStatus="paratype">
<collectingRegion id="4997B8B4614FFFDFB679FD48C1C3FD2F" box="[543,651,688,709]" country="Mongolia" name="Ulaanbaatar" pageId="12" pageNumber="508">Ulan-Bator</collectingRegion>
env., steppe,
<date id="FFED5096614FFFDFB4DBFD33C202FD0A" box="[189,330,715,736]" pageId="12" pageNumber="508" value="1970-06-24" valueMax="1970-06-29" valueMin="1970-06-24">
<collectingDate id="EFA9A97E614FFFDFB4DBFD33C202FD0A" box="[189,330,715,736]" pageId="12" pageNumber="508" value="1970-06-24" valueMax="1970-06-29" valueMin="1970-06-24">24–29.vi.1970</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB530FD33C2E6FD0A" box="[342,430,715,736]" pageId="12" pageNumber="508">Nartshuk</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB5DBFD33C2BEFD0A" box="[445,502,715,736]" count="2" pageId="12" pageNumber="508" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D55BDDF614FFFDFB663FD33C163FD0A" box="[517,555,715,736]" count="1" pageId="12" pageNumber="508" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614FFFDFB624FD33C29AFD11" collectingDate="1970-06-24" collectingDateMax="1970-06-29" collectingDateMin="1970-06-24" collectorName="Kerzhner" country="Mongolia" location="Ulan-Bator" pageId="12" pageNumber="508" specimenCount="2" specimenCount-female="2" stateProvince="Ulaanbaatar" typeStatus="paratype">
same locality,
<date id="FFED5096614FFFDFB6B6FD33C240FD11" pageId="12" pageNumber="508" value="1970-06-24" valueMax="1970-06-29" valueMin="1970-06-24">
<collectingDate id="EFA9A97E614FFFDFB6B6FD33C240FD11" pageId="12" pageNumber="508" value="1970-06-24" valueMax="1970-06-29" valueMin="1970-06-24">24–29. vi.1970</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB57CFD1EC23BFD11" box="[282,371,742,763]" pageId="12" pageNumber="508">Kerzhner</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB5ECFD1EC281FD11" box="[394,457,742,763]" count="2" pageId="12" pageNumber="508" type="female">2 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614FFFDFB580FD1EC2FAFCFC" collectingDate="1970-06-24" collectingDateMax="1970-06-29" collectingDateMin="1970-06-24" collectorName="Kandybina" country="Mongolia" location="Ulan-Bator" pageId="12" pageNumber="508" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Ulaanbaatar" typeStatus="paratype">
same locality,
<date id="FFED5096614FFFDFB6E7FD1EC047FD11" box="[641,783,742,763]" pageId="12" pageNumber="508" value="1970-06-24" valueMax="1970-06-29" valueMin="1970-06-24">
<collectingDate id="EFA9A97E614FFFDFB6E7FD1EC047FD11" box="[641,783,742,763]" pageId="12" pageNumber="508" value="1970-06-24" valueMax="1970-06-29" valueMin="1970-06-24">24–29.vi.1970</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB4DBFCF9C26EFCFC" box="[189,294,769,790]" pageId="12" pageNumber="508">Kandybina</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB55FFCF9C217FCFC" box="[313,351,769,790]" count="1" pageId="12" pageNumber="508" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D55BDDF614FFFDFB508FCF9C2E0FCFC" box="[366,424,769,790]" count="2" pageId="12" pageNumber="508" type="female">2 ♀♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="3B3B7C0B614FFFDFB5D9FCF9C218FCA1" collectingDate="1970-08-10" collectorName="Kerzhner" country="Mongolia" location="Ubsunurskiy" pageId="12" pageNumber="508" specimenCount="9" specimenCount-female="3" specimenCount-male="6" stateProvince="Uvs" typeStatus="paratype">
<collectingRegion id="4997B8B4614FFFDFB5D9FCF9C167FCFC" box="[447,559,769,790]" country="Mongolia" name="Uvs" pageId="12" pageNumber="508">Uvs aymag</collectingRegion>
[=
<location id="8E8C208D614FFFDFB62DFCF9C181FCFC" LSID="urn:lsid:plazi:treatment:03FAC740614FFFDDB4B6FED7C23EFC17:8E8C208D614FFFDFB62DFCF9C181FCFC" box="[587,713,769,790]" country="Mongolia" name="Ubsunurskiy" pageId="12" pageNumber="508" stateProvince="Uvs">Ubsunurskiy</location>
aymak (province)]:
<locationDeviation id="9A5E6F74614FFFDFB55BFCE4C168FCDB" box="[317,544,796,817]" location="Khyargas" pageId="12" pageNumber="508">
<quantity id="4CABDBB3614FFFDFB55BFCE4C23DFCDB" box="[317,373,796,817]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="508" unit="km" value="5.0">5 km</quantity>
N of Khyargas
</locationDeviation>
,
<date id="FFED5096614FFFDFB657FCE4C1EFFCDB" box="[561,679,796,817]" pageId="12" pageNumber="508" value="1970-08-10">
<collectingDate id="EFA9A97E614FFFDFB657FCE4C1EFFCDB" box="[561,679,796,817]" pageId="12" pageNumber="508" value="1970-08-10">10.viii.1970</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB6DFFCE4C05AFCDB" box="[697,786,796,817]" pageId="12" pageNumber="508">Kerzhner</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB4A2FCCEC3B6FCA1" box="[196,254,822,843]" count="6" pageId="12" pageNumber="508" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="9D55BDDF614FFFDFB568FCCEC20FFCA1" box="[270,327,822,843]" count="3" pageId="12" pageNumber="508" type="female">3 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614FFFDFB539FCCEC044FCA1" box="[351,780,822,843]" collectingDate="1970-08-10" collectorName="Nartshuk" country="Mongolia" location="Ubsunurskiy" pageId="12" pageNumber="508" specimenCount="1" specimenCount-female="1" stateProvince="Uvs" typeStatus="paratype">
same locality,
<date id="FFED5096614FFFDFB589FCCEC12DFCA1" box="[495,613,822,843]" pageId="12" pageNumber="508" value="1970-08-10">
<collectingDate id="EFA9A97E614FFFDFB589FCCEC12DFCA1" box="[495,613,822,843]" pageId="12" pageNumber="508" value="1970-08-10">10.viii.1970</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB615FCCEC183FCA1" box="[627,715,822,843]" pageId="12" pageNumber="508">Nartshuk</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB6B8FCCEC04BFCA1" box="[734,771,822,843]" count="1" pageId="12" pageNumber="508" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614FFFDFB4DBFCA9C1DBFC8C" box="[189,659,849,870]" collectingDate="1970-08-10" collectorName="Kozlov" country="Mongolia" location="Ubsunurskiy" pageId="12" pageNumber="508" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Uvs" typeStatus="paratype">
same locality,
<date id="FFED5096614FFFDFB528FCA9C28CFC8C" box="[334,452,849,870]" pageId="12" pageNumber="508" value="1970-08-10">
<collectingDate id="EFA9A97E614FFFDFB528FCA9C28CFC8C" box="[334,452,849,870]" pageId="12" pageNumber="508" value="1970-08-10">10.viii.1970</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB5B4FCA9C151FC8C" box="[466,537,849,870]" pageId="12" pageNumber="508">Kozlov</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB64AFCA9C11BFC8C" box="[556,595,849,870]" count="1" pageId="12" pageNumber="508" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D55BDDF614FFFDFB604FCA9C1C2FC8C" box="[610,650,849,870]" count="1" pageId="12" pageNumber="508" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614FFFDFB6C4FCA9C3BFFC76" collectingDate="1970-08-07" collectorName="Nartshuk" country="Mongolia" location="Shil Ridge" pageId="12" pageNumber="508" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
Togtokhyn-
<location id="8E8C208D614FFFDFB4DBFC94C267FC6B" LSID="urn:lsid:plazi:treatment:03FAC740614FFFDDB4B6FED7C23EFC17:8E8C208D614FFFDFB4DBFC94C267FC6B" box="[189,303,876,897]" country="Mongolia" name="Shil Ridge" pageId="12" pageNumber="508">Shil Ridge</location>
,
<locationDeviation id="9A5E6F74614FFFDFB527FC94C163FC6B" box="[321,555,876,897]" location="Ulangom" pageId="12" pageNumber="508">
<quantity id="4CABDBB3614FFFDFB527FC94C2CFFC6B" box="[321,391,876,897]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="508" unit="km" value="50.0">50 km</quantity>
ESE Ulangom
</locationDeviation>
,
<date id="FFED5096614FFFDFB65BFC94C1EFFC6B" box="[573,679,876,897]" pageId="12" pageNumber="508" value="1970-08-07">
<collectingDate id="EFA9A97E614FFFDFB65BFC94C1EFFC6B" box="[573,679,876,897]" pageId="12" pageNumber="508" value="1970-08-07">7.viii.1970</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB6DCFC94C05AFC6B" box="[698,786,876,897]" pageId="12" pageNumber="508">Nartshuk</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB4A0FC7FC3A5FC76" box="[198,237,903,924]" count="1" pageId="12" pageNumber="508" type="female">1 ♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="3B3B7C0B614FFFDFB562FC7FC1DEFC5D" collectingDate="1970-07-03" collectorName="Kandybina" country="Mongolia" location="Tevshrulekh" pageId="12" pageNumber="508" specimenCount="1" specimenCount-female="1" stateProvince="Arhangay" typeStatus="paratype">
<collectingRegion id="4997B8B4614FFFDFB562FC7FC227FC76" box="[260,367,903,924]" country="Mongolia" name="Arhangay" pageId="12" pageNumber="508">Arkhangay</collectingRegion>
aymag [=Ara-Khangayskiy aymak (province)]
<location id="8E8C208D614FFFDFB49AFC5AC23DFC5D" LSID="urn:lsid:plazi:treatment:03FAC740614FFFDDB4B6FED7C23EFC17:8E8C208D614FFFDFB49AFC5AC23DFC5D" box="[252,373,930,951]" country="Mongolia" name="Tevshrulekh" pageId="12" pageNumber="508" stateProvince="Arhangay">Tevshrulekh</location>
,
<date id="FFED5096614FFFDFB5E6FC5AC2ABFC5D" box="[384,483,930,951]" pageId="12" pageNumber="508" value="1970-07-03">
<collectingDate id="EFA9A97E614FFFDFB5E6FC5AC2ABFC5D" box="[384,483,930,951]" pageId="12" pageNumber="508" value="1970-07-03">3.vii.1970</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB588FC5AC11FFC5D" box="[494,599,930,951]" pageId="12" pageNumber="508">Kandybina</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB60FFC5AC1C5FC5D" box="[617,653,930,951]" count="1" pageId="12" pageNumber="508" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614FFFDFB6C5FC5AC14CFC3B" collectingDate="1970-07-24" collectorName="Kandybina" country="Mongolia" location="Tevshrulekh" pageId="12" pageNumber="508" specimenCount="1" specimenCount-female="1" stateProvince="Arhangay" typeStatus="paratype">
same locality,
<date id="FFED5096614FFFDFB484FC44C21AFC3B" box="[226,338,956,977]" pageId="12" pageNumber="508" value="1970-07-24">
<collectingDate id="EFA9A97E614FFFDFB484FC44C21AFC3B" box="[226,338,956,977]" pageId="12" pageNumber="508" value="1970-07-24">24.vii.1970</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB53AFC44C28DFC3B" box="[348,453,956,977]" pageId="12" pageNumber="508">Kandybina</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB5B0FC44C2B2FC3B" box="[470,506,956,977]" count="1" pageId="12" pageNumber="508" type="female">1 ♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="3B3B7C0B614FFFDFB669FC44C1E3FBED" collectingDate="1970-08-14" collectorName="Nartshuk" country="Mongolia" location="Khasagt-Khayrkhan Ridge" pageId="12" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Govi-Altay" typeStatus="paratype">
<collectingRegion id="4997B8B4614FFFDFB669FC44C13FFC3B" box="[527,631,956,977]" country="Mongolia" name="Govi-Altay" pageId="12" pageNumber="508">Govi-Altai</collectingRegion>
aymag [=Gobi-Altayskiy aymak (province)]:
<location id="8E8C208D614FFFDFB666FC2FC047FC06" LSID="urn:lsid:plazi:treatment:03FAC740614FFFDDB4B6FED7C23EFC17:8E8C208D614FFFDFB666FC2FC047FC06" box="[512,783,983,1004]" country="Mongolia" name="Khasagt-Khayrkhan Ridge" pageId="12" pageNumber="508" stateProvince="Govi-Altay">Khasagt-Khayrkhan Ridge</location>
,
<locationDeviation id="9A5E6F74614FFFDFB4DBFC0AC2C0FBED" box="[189,392,1010,1031]" location="Dzhargalan" pageId="12" pageNumber="508">
<quantity id="4CABDBB3614FFFDFB4DBFC0AC3B3FBED" box="[189,251,1010,1031]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="508" unit="km" value="15.0">15 km</quantity>
S Dzhargalan
</locationDeviation>
,
<date id="FFED5096614FFFDFB5F5FC0AC141FBED" box="[403,521,1010,1031]" pageId="12" pageNumber="508" value="1970-08-14">
<collectingDate id="EFA9A97E614FFFDFB5F5FC0AC141FBED" box="[403,521,1010,1031]" pageId="12" pageNumber="508" value="1970-08-14">14.viii.1970</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB672FC0AC124FBED" box="[532,620,1010,1031]" pageId="12" pageNumber="508">Nartshuk</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB61BFC0AC1E9FBED" box="[637,673,1010,1031]" count="1" pageId="12" pageNumber="508" type="male">1 ♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="3B3B7C0B614FFFDFB6D1FC0AC3B9FBD7" collectingDate="1913-07-08" collectorName="Tolstov" country="Russia" location="Verkhnie Sayany" municipality="Okinskiy Stan" pageId="12" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Buryatiya" typeStatus="paratype">
<collectingCountry id="F34436C6614FFFDFB6D1FC0AC058FBED" box="[695,784,1010,1031]" name="Russia" pageId="12" pageNumber="508">RUSSIA</collectingCountry>
.
<collectingRegion id="4997B8B4614FFFDFB4DBFBF5C25BFBC8" box="[189,275,1037,1058]" country="Russia" name="Buryatiya" pageId="12" pageNumber="508">Buryatia</collectingRegion>
:
<collectingMunicipality id="6B88EC2C614FFFDFB57AFBF5C2EFFBC8" box="[284,423,1037,1058]" pageId="12" pageNumber="508">Okinskiy Stan</collectingMunicipality>
,
<location id="8E8C208D614FFFDFB5D6FBF5C11DFBC8" LSID="urn:lsid:plazi:treatment:03FAC740614FFFDDB4B6FED7C23EFC17:8E8C208D614FFFDFB5D6FBF5C11DFBC8" box="[432,597,1037,1058]" country="Russia" municipality="Okinskiy Stan" name="Verkhnie Sayany" pageId="12" pageNumber="508" stateProvince="Buryatiya">Verkhnie Sayany</location>
,
<date id="FFED5096614FFFDFB638FBF5C189FBC8" box="[606,705,1037,1058]" pageId="12" pageNumber="508" value="1913-07-08">
<collectingDate id="EFA9A97E614FFFDFB638FBF5C189FBC8" box="[606,705,1037,1058]" pageId="12" pageNumber="508" value="1913-07-08">8.vii.1913</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB6ACFBF5C05AFBC8" box="[714,786,1037,1058]" pageId="12" pageNumber="508">Tolstov</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB4A0FBD0C3A0FBD7" box="[198,232,1064,1085]" count="1" pageId="12" pageNumber="508" type="male">1 ♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614FFFDFB49AFBD0C216FBBD" collectingDate="1971-06-24" collectionCode="V" collectorName="Richter" country="Russia" location="10 km SW Selendum" pageId="12" pageNumber="508" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="8E8C208D614FFFDFB49AFBD0C28FFBD7" LSID="urn:lsid:plazi:treatment:03FAC740614FFFDDB4B6FED7C23EFC17:8E8C208D614FFFDFB49AFBD0C28FFBD7" box="[252,455,1064,1085]" country="Russia" name="10 km SW Selendum" pageId="12" pageNumber="508">
<quantity id="4CABDBB3614FFFDFB49AFBD0C270FBD7" box="[252,312,1064,1085]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="508" unit="km" value="10.0">10 km</quantity>
SW Selendum
</location>
, pine forest,
<date id="FFED5096614FFFDFB627FBD0C1E2FBD7" box="[577,682,1064,1085]" pageId="12" pageNumber="508" value="1971-06-24">
<collectingDate id="EFA9A97E614FFFDFB627FBD0C1E2FBD7" box="[577,682,1064,1085]" pageId="12" pageNumber="508" value="1971-06-24">24.vi.1971</collectingDate>
</date>
,
<collectionCode id="ED42EE93614FFFDFB6D4FBD0C18BFBD7" box="[690,707,1064,1085]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="508" type="Museum">V</collectionCode>
.
<collectorName id="26A61380614FFFDFB6ADFBD0C05AFBD7" box="[715,786,1064,1085]" pageId="12" pageNumber="508">Richter</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB4A0FBBAC3A4FBBD" box="[198,236,1090,1111]" count="1" pageId="12" pageNumber="508" type="male">1 ♂</specimenCount>
, dissected)
</materialsCitation>
.
<materialsCitation id="3B3B7C0B614FFFDFB50CFBBAC2A6FB98" collectingDate="1966-06-27" collectorName="A. Pleshanov" country="Russia" county="Irkutskaya Prov." location="Chernorud" municipality="Baykal Lake" pageId="12" pageNumber="508" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCounty id="628D0EDA614FFFDFB50CFBBAC147FBBD" box="[362,527,1090,1111]" pageId="12" pageNumber="508">Irkutskaya Prov.</collectingCounty>
:
<collectingMunicipality id="6B88EC2C614FFFDFB67CFBBAC1D1FBBD" box="[538,665,1090,1111]" pageId="12" pageNumber="508">Baykal Lake</collectingMunicipality>
,
<location id="8E8C208D614FFFDFB6C3FBBAC046FBBD" LSID="urn:lsid:plazi:treatment:03FAC740614FFFDDB4B6FED7C23EFC17:8E8C208D614FFFDFB6C3FBBAC046FBBD" box="[677,782,1090,1111]" country="Russia" county="Irkutskaya Prov." municipality="Baykal Lake" name="Chernorud" pageId="12" pageNumber="508">Chernorud</location>
,
<date id="FFED5096614FFFDFB4DBFBA5C26DFB98" box="[189,293,1117,1138]" pageId="12" pageNumber="508" value="1966-06-27">
<collectingDate id="EFA9A97E614FFFDFB4DBFBA5C26DFB98" box="[189,293,1117,1138]" pageId="12" pageNumber="508" value="1966-06-27">27.vi.1966</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB549FBA5C2F9FB98" box="[303,433,1117,1138]" pageId="12" pageNumber="508">A. Pleshanov</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB5A7FBA5C2ACFB98" box="[449,484,1117,1138]" count="1" pageId="12" pageNumber="508" type="male">1 ♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="3B3B7C0B614FFFDFB59EFBA5C19AFB67" collectingDate="1986-06-21" collectorName="Bagachanova" country="Russia" location="Mikhaylovka" municipality="Yakutia" pageId="12" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Sakha" typeStatus="paratype">
<collectingMunicipality id="6B88EC2C614FFFDFB59EFBA5C10AFB98" box="[504,578,1117,1138]" pageId="12" pageNumber="508">Yakutia</collectingMunicipality>
(
<collectingRegion id="4997B8B4614FFFDFB629FBA5C1C7FB98" box="[591,655,1117,1138]" country="Russia" name="Sakha" pageId="12" pageNumber="508">Sakha</collectingRegion>
):
<location id="8E8C208D614FFFDFB6C7FBA5C39EFB67" LSID="urn:lsid:plazi:treatment:03FAC740614FFFDDB4B6FED7C23EFC17:8E8C208D614FFFDFB6C7FBA5C39EFB67" country="Russia" municipality="Yakutia" name="Mikhaylovka" pageId="12" pageNumber="508" stateProvince="Sakha">Mikhaylovka</location>
,
<locationDeviation id="9A5E6F74614FFFDFB484FB80C2C7FB67" box="[226,399,1144,1165]" location="Amga" pageId="12" pageNumber="508">
<quantity id="4CABDBB3614FFFDFB484FB80C26AFB67" box="[226,290,1144,1165]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="508" unit="km" value="60.0">60 km</quantity>
NE Amga
</locationDeviation>
,
<date id="FFED5096614FFFDFB5FDFB80C14CFB67" box="[411,516,1144,1165]" pageId="12" pageNumber="508" value="1986-06-21">
<collectingDate id="EFA9A97E614FFFDFB5FDFB80C14CFB67" box="[411,516,1144,1165]" pageId="12" pageNumber="508" value="1986-06-21">21.vi.1986</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB677FB80C1DAFB67" box="[529,658,1144,1165]" pageId="12" pageNumber="508">Bagachanova</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB6C2FB80C181FB67" box="[676,713,1144,1165]" count="1" pageId="12" pageNumber="508" type="male">1 ♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B3B7C0B614FFFDFB687FB80C1B3FB42" collectingDate="1985-06-04" collectionCode="ZISP" collectorName="Maksimova" country="Russia" location="Mikhaylovka" municipality="Yakutia" pageId="12" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Sakha" typeStatus="paratype">
same locality,
<date id="FFED5096614FFFDFB569FB6BC224FB42" box="[271,364,1171,1192]" pageId="12" pageNumber="508" value="1985-06-04">
<collectingDate id="EFA9A97E614FFFDFB569FB6BC224FB42" box="[271,364,1171,1192]" pageId="12" pageNumber="508" value="1985-06-04">4.vi.1985</collectingDate>
</date>
,
<collectorName id="26A61380614FFFDFB510FB6BC2AFFB42" box="[374,487,1171,1192]" pageId="12" pageNumber="508">Maksimova</collectorName>
(
<specimenCount id="9D55BDDF614FFFDFB591FB6BC151FB42" box="[503,537,1171,1192]" count="1" pageId="12" pageNumber="508" type="male">1 ♂</specimenCount>
, all
<typeStatus id="54E8C8F4614FFFDFB625FB6BC1D7FB42" box="[579,671,1171,1192]" pageId="12" pageNumber="508" type="paratype">paratypes</typeStatus>
in
<collectionCode id="ED42EE93614FFFDFB6D8FB6BC1BCFB42" box="[702,756,1171,1192]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="12" pageNumber="508">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614FFFDFB485FB56C289FA5E" pageId="12" pageNumber="508" type="diagnosis">
<paragraph id="8BEC7656614FFFDFB485FB56C289FA5E" blockId="12.[189,788,420,2076]" pageId="12" pageNumber="508">
DIAGNOSIS. Mid-sized species (body length
<quantity id="4CABDBB3614FFFDFB6C4FB56C04FFB29" box="[674,775,1198,1219]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="12" pageNumber="508" unit="mm" value="5.25" valueMax="5.5" valueMin="5.0">5–5.5 mm</quantity>
); male eyes dichoptic but frons very narrow at middle, narrower than anterior ocellus; postpedicel nearly 2.5X longer than basal width; labrum nearly 2.5X longer than eye height, palpus mostly yellow; mesoscutum with 4 brownish vittae, acrostichals biserial, dorsocentrals 1–2-serial, laterotergite with pale setae; legs extensively yellow, hind femur slender; abdomen mostly pale setose, tergites 2–5 faintly greyish pruinose dorsally; sternite 8 posteriorly and hypopygium yellow; female legs without pennate setae.
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614FFFDDB485FA42C2B4FC78" lastPageId="14" lastPageNumber="510" pageId="12" pageNumber="508" type="description">
<paragraph id="8BEC7656614FFFDFB485FA42C196F87C" blockId="12.[189,788,420,2076]" pageId="12" pageNumber="508">
DESCRIPTION. Body length
<quantity id="4CABDBB3614FFFDFB590FA42C12CFA25" box="[502,612,1466,1487]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.6" metricValueMin="5.1" pageId="12" pageNumber="508" unit="mm" value="5.35" valueMax="5.6" valueMin="5.1">5.1–5.6 mm</quantity>
, wing
<quantity id="4CABDBB3614FFFDFB6C6FA42C058FA25" box="[672,784,1466,1487]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" metricValueMax="6.5" metricValueMin="6.1" pageId="12" pageNumber="508" unit="mm" value="6.3" valueMax="6.5" valueMin="6.1">6.1–6.5 mm</quantity>
.
<emphasis id="B927AA44614FFFDFB4DBFA2CC3B9FA03" bold="true" box="[189,241,1492,1513]" pageId="12" pageNumber="508">Male</emphasis>
(
<figureCitation id="13686AD3614FFFDFB564FA2CC206FA03" box="[258,334,1492,1513]" captionStart="Figs 23–28" captionStartId="13.[190,228,1921,1940]" captionTargetBox="[208,1396,556,1859]" captionTargetId="figure-106@13.[200,1410,536,1874]" captionTargetPageId="13" captionText="Figs 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., male: 23 — habitus, holotype, lateral view; 24 — head and thorax, holotype, anterodorsal view; 25 — hypopygium, lateral view; 26 — hypandrium, ventral view; 27 — cerci, dorsal view; 28 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., самец: 23 — габитус, голотип, вид сбоку; 24 — голова и грудь, голотип, вид спереди-сверху; 25 — гипопигий, вид сбоку; 26 — гипандрий, вид сниЗу; 27 — церки, вид сверху; 28 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="12" pageNumber="508">Fig. 23</figureCitation>
). Head capsule regions mostly densely greyish pruinose; face on lower margin and entire clypeus shiny; head setation mostly black. Eyes dichoptic, ommatidia equally small. Frons slightly broadened just below ocellar triangle, very narrow before middle (narrower than anterior ocellus), broadened towards antennae on remaining portion; bare. Face broad, bare. Ocellar triangle with 2 long and several short setae. Occiput with almost regular row of moderately long setae on each side; postoculars moderately long on upper part and short laterally; postgena with pale hair-like setae. Antenna with scape and pedicel brown, postpedicel and stylus black; scape nearly 1.5X longer than subglobular pedicel, both with very short setulae; postpedicel nearly 2.5X as long as basal width, with straight margins; stylus slightly longer than postpedicel basal width. Proboscis with labrum brownish yellow, nearly 2.5X longer than eye height; palpus short, mostly yellow, brownish close to base; with scattered, dark setulae.
</paragraph>
<paragraph id="8BEC7656614FFFDFB485F864C6C1FD62" blockId="12.[189,788,420,2076]" lastBlockId="12.[824,1422,303,2076]" pageId="12" pageNumber="508">
Thorax black in ground-colour, mostly densely greyish pruinose and black setose; mesoscutum (
<figureCitation id="13686AD3614FFFDFB61FF84FC18AF826" box="[633,706,1975,1996]" captionStart="Figs 23–28" captionStartId="13.[190,228,1921,1940]" captionTargetBox="[208,1396,556,1859]" captionTargetId="figure-106@13.[200,1410,536,1874]" captionTargetPageId="13" captionText="Figs 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., male: 23 — habitus, holotype, lateral view; 24 — head and thorax, holotype, anterodorsal view; 25 — hypopygium, lateral view; 26 — hypandrium, ventral view; 27 — cerci, dorsal view; 28 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., самец: 23 — габитус, голотип, вид сбоку; 24 — голова и грудь, голотип, вид спереди-сверху; 25 — гипопигий, вид сбоку; 26 — гипандрий, вид сниЗу; 27 — церки, вид сверху; 28 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="12" pageNumber="508">Fig. 24</figureCitation>
) with 4 brown vittae (lateral vittae less distinct). Antepronotum with 4–5 black to yellow setae dorsally on each side. Postpronotal lobe with 1 long and 3–4 shorter setae (sometimes with additional 3–4 yellow to pale setulae). Proepisternum with tuft of several pale, fine setae on lower part. Prosternum bare. Mesonotal setae: acrostichals arranged in 2 close irregular rows, short, absent on prescutellar depression; dorsocentrals uniserial, mostly short, 2 long prescutellars; 1 short presutural intra-alar, 1 long presutural supra-alar, 3–4 notopleurals (usually 3 strong, longer and 1 short, finer), 1 postsutural supra-alar (usually with 1 short, fine seta anteriorly), 1 long and 1 minute postalars, usually 2 scutellars (sometimes 1–2 additional, lateral, short, fine seta(e) present); sometimes 1–3 brownish yellow setulae in anterior part of notopleuron (usually absent). Laterotergite with a few long, strong, yellow and additional short, fine, pale setae. Anterior and posterior spiracles yellow.
</paragraph>
<paragraph id="8BEC7656614FFFDFB738FD76C602FB87" blockId="12.[824,1422,303,2076]" pageId="12" pageNumber="508">Legs with coxae densely greyish pruinose, remaining podomeres subshiny; black setose; almost entirely yellow (including coxae), only tarsomere 1 at apex, most part of tarsomere 2 (except base) and entire tarsomere 3–5 brownish. Fore coxa with scattered setae anteriorly.All femora whitish pilose ventrally.Fore femur with rows of minute anteroventral and posteroventral setae. Fore tibia with 1–2 short, fine anterodorsal setae. Mid femur with row of mostly minute anteroventral setae (slightly longer and stronger near base) and more numerous, spinule-like setae posteroventrally (slightly longer in about basal half). Mid tibia with 2–3 anterodorsal and 3–4 posterodorsal, short setae (number, position and robustness variable). Hind femur slender; bearing complete row of rather short, numerous, anteroventral setae (longest setae nearly half as long as femur maximal width); covered with very short setae posteroventrally. Hind tibia simple, slender; with 3–4 anterodorsal and 3–4 posterodorsal, short setae; simple setulae ventrally; no seta in posteroapical comb.Tarsomeres of all legs slender; basitarsi with scattered, spine-like setae ventrally.</paragraph>
<paragraph id="8BEC7656614FFFDFB738FB8BC726FA90" blockId="12.[824,1422,303,2076]" pageId="12" pageNumber="508">
Wing membrane faintly brownish infuscate; entirely covered with microtrichia; veins yellowish brown, well-sclerotized. Veins R
<subScript id="17D77413614FFFDFB7B1FB4FC096FB29" attach="left" box="[983,990,1207,1219]" fontSize="5" pageId="12" pageNumber="508">5</subScript>
and M
<subScript id="17D77413614FFFDFB04EFB4FC767FB29" attach="left" box="[1064,1071,1207,1219]" fontSize="5" pageId="12" pageNumber="508">1</subScript>
almost parallel towards wing margin; radial fork acute; R
<subScript id="17D77413614FFFDFB056FB2AC77FFB34" attach="left" box="[1072,1079,1234,1246]" fontSize="5" pageId="12" pageNumber="508">5</subScript>
meeting costa before wing apex; CuA+CuP complete (slightly weakened beyond middle). Cell dm moderately large, with strongly elongate apex; apical portion of M
<subScript id="17D77413614FFFDFB7B9FADBC0AEFAC5" attach="left" box="[991,998,1315,1327]" fontSize="5" pageId="12" pageNumber="508">4</subScript>
nearly 3X longer than its middle portion. Pterostigma inconspicuous. Basal costal seta absent. Anal lobe well-developed; axillary incision nearly 90°. Squama yellow, pale fringed. Halter pale yellow.
</paragraph>
<paragraph id="8BEC7656614FFFDFB738FA78C64BF7F6" blockId="12.[824,1422,303,2076]" pageId="12" pageNumber="508">
Abdomen almost entirely brown in ground colour, only sternite 8 broadly yellow posteriorly (size of yellow space somewhat variable). Pruinescence: tergites
<quantity id="4CABDBB3614FFFDFB0BFFA4EC65EFA21" box="[1241,1302,1462,1483]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="12.7" metricValueMin="5.08" pageId="12" pageNumber="508" unit="in" value="3.5" valueMax="5.0" valueMin="2.0">2–5 in</quantity>
lateral view mostly densely greyish pruinose (faintly pruinose along posterior margin), in dorsal view mostly faintly greyish pruinose; tergites 6–7 with yellowish grey iridescent spot dorsally, greyish pruinose laterally (dorsal view); tergite 1 and sternites 2–7 densely greyish pruinose, sternite 8 faintly pruinose. Chaetotaxy: tergites 1–7 mostly covered with numerous, very short, fine, pale setulae; tergites 1–3 with moderately long, yellowish, posteromarginal setae laterally (sometimes less distinct on tergite 3); tergite 8 bearing black setulae; sternite 1 bare, sternites 2–6 with pale setulae only, sternite 7 with additional moderately long, brownish to pale posteromarginal setae, sternite 8 covered with black setae only (posteromarginals moderately long). Pregenital segments: segment 6 unmodified; tergite 7 broadly concave posteriorly; sternite 7 with somewhat produced posterior corners; segment 8 with separated tergite and sternite; tergite 8 somewhat flattened, represented by two subtriangular sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 large, scoop-shaped, somewhat constricted anteriorly, with 2 antero-lateral projections separated by depression (upper projection slightly smaller and with bifid apex; lower projection larger, subconical) and small- er projection on posterior margin of depression.
</paragraph>
<paragraph id="8BEC7656614EFFDDB485FED7C13BFE20" blockId="13.[189,786,303,457]" lastBlockId="14.[189,787,303,1021]" lastPageId="14" lastPageNumber="510" pageId="13" pageNumber="509">
Hypopygium (
<figureCitation id="13686AD3614EFFDEB514FED7C2ABFEAE" box="[370,483,303,324]" captionStart="Figs 23–28" captionStartId="13.[190,228,1921,1940]" captionTargetBox="[208,1396,556,1859]" captionTargetId="figure-106@13.[200,1410,536,1874]" captionTargetPageId="13" captionText="Figs 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., male: 23 — habitus, holotype, lateral view; 24 — head and thorax, holotype, anterodorsal view; 25 — hypopygium, lateral view; 26 — hypandrium, ventral view; 27 — cerci, dorsal view; 28 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., самец: 23 — габитус, голотип, вид сбоку; 24 — голова и грудь, голотип, вид спереди-сверху; 25 — гипопигий, вид сбоку; 26 — гипандрий, вид сниЗу; 27 — церки, вид сверху; 28 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="13" pageNumber="509">Figs 25–28</figureCitation>
) moderately large; almost entirely yellow, only apical semicircular part of phallus yellow; epandrial lamella with short, black setae more numerous along lower margin; cercus covered with black setulae. Epandrium entirely broadly divided (epandrial bridge absent); epandrial lamella (
<figureCitation id="13686AD3614EFFDEB572FE4CC215FE23" box="[276,349,436,457]" captionStart="Figs 23–28" captionStartId="13.[190,228,1921,1940]" captionTargetBox="[208,1396,556,1859]" captionTargetId="figure-106@13.[200,1410,536,1874]" captionTargetPageId="13" captionText="Figs 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., male: 23 — habitus, holotype, lateral view; 24 — head and thorax, holotype, anterodorsal view; 25 — hypopygium, lateral view; 26 — hypandrium, ventral view; 27 — cerci, dorsal view; 28 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., самец: 23 — габитус, голотип, вид сбоку; 24 — голова и грудь, голотип, вид спереди-сверху; 25 — гипопигий, вид сбоку; 26 — гипандрий, вид сниЗу; 27 — церки, вид сверху; 28 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="13" pageNumber="509">Fig. 25</figureCitation>
) rather subtriangular (lateral view), broadly rounded apically. Hypandrium (
<figureCitation id="13686AD3614EFFDEB00AFED7C7FBFEAE" box="[1132,1203,303,324]" captionStart="Figs 23–28" captionStartId="13.[190,228,1921,1940]" captionTargetBox="[208,1396,556,1859]" captionTargetId="figure-106@13.[200,1410,536,1874]" captionTargetPageId="13" captionText="Figs 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., male: 23 — habitus, holotype, lateral view; 24 — head and thorax, holotype, anterodorsal view; 25 — hypopygium, lateral view; 26 — hypandrium, ventral view; 27 — cerci, dorsal view; 28 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., самец: 23 — габитус, голотип, вид сбоку; 24 — голова и грудь, голотип, вид спереди-сверху; 25 — гипопигий, вид сбоку; 26 — гипандрий, вид сниЗу; 27 — церки, вид сверху; 28 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="13" pageNumber="509">Fig. 26</figureCitation>
) separated from epandrium; mostly membranous, undivided, narrowly sclerotized along margin, subtriangular in ventral view; bare; gonocoxal apodeme small. Cerci (
<figureCitation id="13686AD3614EFFDEB07EFE87C717FE7E" box="[1048,1119,383,404]" captionStart="Figs 23–28" captionStartId="13.[190,228,1921,1940]" captionTargetBox="[208,1396,556,1859]" captionTargetId="figure-106@13.[200,1410,536,1874]" captionTargetPageId="13" captionText="Figs 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., male: 23 — habitus, holotype, lateral view; 24 — head and thorax, holotype, anterodorsal view; 25 — hypopygium, lateral view; 26 — hypandrium, ventral view; 27 — cerci, dorsal view; 28 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., самец: 23 — габитус, голотип, вид сбоку; 24 — голова и грудь, голотип, вид спереди-сверху; 25 — гипопигий, вид сбоку; 26 — гипандрий, вид сниЗу; 27 — церки, вид сверху; 28 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="13" pageNumber="509">Fig. 27</figureCitation>
) separated with each other and from epandrium; cercus elongate ovate (lateral view), shorter than epandrial lamella, without additional lobe dorsally, serrate on about apical third of inner margin. Phallus (
<figureCitation id="13686AD3614DFFDDB6C0FED7C1A4FEAE" box="[678,748,303,324]" captionStart="Figs 23–28" captionStartId="13.[190,228,1921,1940]" captionTargetBox="[208,1396,556,1859]" captionTargetId="figure-106@13.[200,1410,536,1874]" captionTargetPageId="13" captionText="Figs 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., male: 23 — habitus, holotype, lateral view; 24 — head and thorax, holotype, anterodorsal view; 25 — hypopygium, lateral view; 26 — hypandrium, ventral view; 27 — cerci, dorsal view; 28 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm. Рис. 23–28. Empis (Polyblepharis) subtransbaicalica Shamshev, sp.n., самец: 23 — габитус, голотип, вид сбоку; 24 — голова и грудь, голотип, вид спереди-сверху; 25 — гипопигий, вид сбоку; 26 — гипандрий, вид сниЗу; 27 — церки, вид сверху; 28 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм." pageId="14" pageNumber="510">Fig. 28</figureCitation>
) almost entirely hidden; zigzag bent, strongly constricted beyond middle; basal portion thick, gently curved; apical bend portion semicircular, thickened at base, with its tip pointing to rear. Ejaculatory apodeme moderately large, extended far beyond basal curvature of phallus, with lateral wings.
</paragraph>
<caption id="DF2C26DE614EFFDEB4D8F879C622F7E6" pageId="13" pageNumber="509" startId="13.[190,228,1921,1940]" targetBox="[208,1396,556,1859]" targetPageId="13" targetType="figure">
<paragraph id="8BEC7656614EFFDEB4D8F879C6CEF82E" blockId="13.[190,1424,1921,2060]" pageId="13" pageNumber="509">
<emphasis id="B927AA44614EFFDEB4D8F879C26CF87E" bold="true" box="[190,292,1921,1940]" pageId="13" pageNumber="509">Figs 23–28.</emphasis>
<taxonomicName id="4C530DD5614EFFDEB54FF879C1AEF87E" authority="Shamshev" authorityName="Shamshev" box="[297,742,1921,1940]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="509" phylum="Arthropoda" rank="species" species="subtransbaicalica" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA44614EFFDEB54FF879C216F87E" box="[297,350,1921,1940]" italics="true" pageId="13" pageNumber="509">Empis</emphasis>
(
<emphasis id="B927AA44614EFFDEB50DF879C2ABF87E" box="[363,483,1921,1940]" italics="true" pageId="13" pageNumber="509">Polyblepharis</emphasis>
)
<emphasis id="B927AA44614EFFDEB589F879C1C0F87E" box="[495,648,1921,1940]" italics="true" pageId="13" pageNumber="509">subtransbaicalica</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA44614EFFDEB688F879C050F87E" bold="true" box="[750,792,1921,1940]" pageId="13" pageNumber="509">
<taxonomicNameLabel id="A214173F614EFFDEB688F879C050F87E" box="[750,792,1921,1940]" pageId="13" pageNumber="509" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, male: 23 — habitus, holotype, lateral view; 24 — head and thorax, holotype, anterodorsal view; 25 — hypopygium, lateral view; 26 — hypandrium, ventral view; 27 — cerci, dorsal view; 28 — phallus, lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 0.1 mm.
</paragraph>
<paragraph id="8BEC7656614EFFDEB4D8F831C622F7E6" blockId="13.[190,1424,1921,2060]" pageId="13" pageNumber="509">
<emphasis id="B927AA44614EFFDEB4D8F831C261F836" bold="true" box="[190,297,1993,2012]" pageId="13" pageNumber="509">Рис. 23–28.</emphasis>
<taxonomicName id="4C530DD5614EFFDEB557F831C1BDF836" authority="Shamshev" authorityName="Shamshev" box="[305,757,1993,2012]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="509" phylum="Arthropoda" rank="species" species="subtransbaicalica" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B927AA44614EFFDEB557F831C22EF836" box="[305,358,1993,2012]" italics="true" pageId="13" pageNumber="509">Empis</emphasis>
(
<emphasis id="B927AA44614EFFDEB513F831C2A5F836" box="[373,493,1993,2012]" italics="true" pageId="13" pageNumber="509">Polyblepharis</emphasis>
)
<emphasis id="B927AA44614EFFDEB59AF831C1DDF836" box="[508,661,1993,2012]" italics="true" pageId="13" pageNumber="509">subtransbaicalica</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B927AA44614EFFDEB766F831C062F836" bold="true" box="[768,810,1993,2012]" pageId="13" pageNumber="509">
<taxonomicNameLabel id="A214173F614EFFDEB766F831C062F836" box="[768,810,1993,2012]" pageId="13" pageNumber="509" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, самец: 23 — габитус, голотип, вид сбоку; 24 — голова и грудь, голотип, вид спереди- сверху; 25 — гипопигий, вид сбоку; 26 — гипандрий, вид сниЗу; 27 — церки, вид сверху; 28 — фаллус, вид сбоку. СокращениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Масштаб 0,1 мм.
</paragraph>
</caption>
<paragraph id="8BEC7656614DFFDDB485FE28C139FD51" blockId="14.[189,787,303,1021]" pageId="14" pageNumber="510">
<emphasis id="B927AA44614DFFDDB485FE28C27BFE0F" bold="true" box="[227,307,464,485]" pageId="14" pageNumber="510">Female.</emphasis>
Similar to male except as follows. Frons broad, parallel-sided, with scattered marginal setulae. Mid and hind femora mostly covered with simple setulae ventrally, hind femur only with 1–3 short, strong, anteroventral setae in about apical 1/3. Abdomen mostly densely greyish pruinose; mostly with pale setulae, yellow posteromarginal setae present only on tergites 1–2; tergite 7 faintly pruinose; tergite 8 yellow to brownish yellow, shiny; sternite 8 subshiny. Cercus brown, short, covered with dense, pale, erect setulae.
</paragraph>
<paragraph id="8BEC7656614DFFDDB485FD39C2B4FC78" blockId="14.[189,787,303,1021]" pageId="14" pageNumber="510">
DIFFERENTIAL DIAGNOSIS. The new species is similar to
<taxonomicName id="4C530DD5614DFFDDB493FD24C174FD1B" authority="Shamshev, 2006" authorityName="Shamshev" authorityYear="2006" box="[245,572,732,753]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="510" phylum="Arthropoda" rank="species" species="transbaicalica">
<emphasis id="B927AA44614DFFDDB493FD24C24CFD1B" box="[245,260,732,753]" italics="true" pageId="14" pageNumber="510">E</emphasis>
.
<emphasis id="B927AA44614DFFDDB569FD24C2D2FD1B" box="[271,410,732,753]" italics="true" pageId="14" pageNumber="510">transbaicalica</emphasis>
<bibRefCitation id="EFC20BA7614DFFDDB5F9FD24C174FD1B" author="Shamshev I. V." box="[415,572,732,753]" pageId="14" pageNumber="510" pagination="231 - 264" refId="ref15746" refString="Shamshev I. V. 2006. Revision of the genus Empis Linnaeus (Diptera: Empididae) from Russia and neighbouring lands. 3. Descriptions of thirteen new species of the subgenus Polyblepharis Bezzi // International Journal of Dipterological Research. Vol. 17. P. 231 - 264." type="journal article" year="2006">Shamshev, 2006</bibRefCitation>
</taxonomicName>
known from the Eastern Siberia [
<bibRefCitation id="EFC20BA7614DFFDDB552FD0FC292FCE6" author="Shamshev I. V." box="[308,474,759,780]" pageId="14" pageNumber="510" pagination="231 - 264" refId="ref15746" refString="Shamshev I. V. 2006. Revision of the genus Empis Linnaeus (Diptera: Empididae) from Russia and neighbouring lands. 3. Descriptions of thirteen new species of the subgenus Polyblepharis Bezzi // International Journal of Dipterological Research. Vol. 17. P. 231 - 264." type="journal article" year="2006">Shamshev, 2006</bibRefCitation>
].
<taxonomicName id="4C530DD5614DFFDDB58DFD0FC193FCE6" box="[491,731,759,780]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="510" phylum="Arthropoda" rank="species" species="subtransbaicalica" status="sp. nov.">
<emphasis id="B927AA44614DFFDDB58DFD0FC193FCE6" box="[491,731,759,780]" italics="true" pageId="14" pageNumber="510">Empis subtransbaicalica</emphasis>
</taxonomicName>
<emphasis id="B927AA44614DFFDDB684FD0FC05BFCE6" bold="true" box="[738,787,759,780]" pageId="14" pageNumber="510">
<taxonomicNameLabel id="A214173F614DFFDDB684FD0FC05BFCE6" box="[738,787,759,780]" pageId="14" pageNumber="510" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C530DD5614DFFDDB55AFCE9C2ADFCCC" authorityName="Shamshev" authorityYear="2006" box="[316,485,785,806]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="510" phylum="Arthropoda" rank="species" species="transbaicalica">
<emphasis id="B927AA44614DFFDDB55AFCE9C203FCCC" box="[316,331,785,806]" italics="true" pageId="14" pageNumber="510">E</emphasis>
.
<emphasis id="B927AA44614DFFDDB53CFCE9C2ADFCCC" box="[346,485,785,806]" italics="true" pageId="14" pageNumber="510">transbaicalica</emphasis>
</taxonomicName>
by pale laterotergal setae and pale setose abdomen. In addition, the male of the new species has pruinose abdominal tergites 2–5 (shiny in
<taxonomicName id="4C530DD5614DFFDDB61DFCBFC39CFC9D" authorityName="Shamshev" authorityYear="2006" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="510" phylum="Arthropoda" rank="species" species="transbaicalica">
<emphasis id="B927AA44614DFFDDB61DFCBFC1C2FCB6" box="[635,650,839,860]" italics="true" pageId="14" pageNumber="510">E</emphasis>
.
<emphasis id="B927AA44614DFFDDB6F0FCBFC39CFC9D" italics="true" pageId="14" pageNumber="510">transbaicalica</emphasis>
</taxonomicName>
) and spots of iridescent pruinosity on tergites 6 and 7 (only on tergite
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<taxonomicName id="4C530DD5614DFFDDB52FFC85C2A6FC78" authorityName="Shamshev" authorityYear="2006" box="[329,494,893,914]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="510" phylum="Arthropoda" rank="species" species="transbaicalica">
<emphasis id="B927AA44614DFFDDB52FFC85C210FC78" box="[329,344,893,914]" italics="true" pageId="14" pageNumber="510">E</emphasis>
.
<emphasis id="B927AA44614DFFDDB505FC85C2A6FC78" box="[355,494,893,914]" italics="true" pageId="14" pageNumber="510">transbaicalica</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C34925DD614DFFDDB485FC6FC2ABFC2D" pageId="14" pageNumber="510" type="etymology">
<paragraph id="8BEC7656614DFFDDB485FC6FC2ABFC2D" blockId="14.[189,787,303,1021]" pageId="14" pageNumber="510">
ETYMOLOGY. The epithet of the new species refers to its similarity to
<taxonomicName id="4C530DD5614DFFDDB55EFC4AC295FC2D" authorityName="Shamshev" authorityYear="2006" box="[312,477,946,967]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="510" phylum="Arthropoda" rank="species" species="transbaicalica">
<emphasis id="B927AA44614DFFDDB55EFC4AC20FFC2D" box="[312,327,946,967]" italics="true" pageId="14" pageNumber="510">E</emphasis>
.
<emphasis id="B927AA44614DFFDDB534FC4AC295FC2D" box="[338,477,946,967]" italics="true" pageId="14" pageNumber="510">transbaicalica</emphasis>
</taxonomicName>
.
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<paragraph id="8BEC7656614DFFDDB485FC35C23EFC17" blockId="14.[189,787,303,1021]" pageId="14" pageNumber="510">
DISTRIBUTION.
<collectingCountry id="F34436C6614DFFDDB5F1FC35C2BFFC08" box="[407,503,973,994]" name="Mongolia" pageId="14" pageNumber="510">Mongolia</collectingCountry>
,
<collectingCountry id="F34436C6614DFFDDB666FC35C108FC08" box="[512,576,973,994]" name="Russia" pageId="14" pageNumber="510">Russia</collectingCountry>
(
<collectingRegion id="4997B8B4614DFFDDB62BFC35C1EBFC08" box="[589,675,973,994]" country="Russia" name="Buryatiya" pageId="14" pageNumber="510">Buryatia</collectingRegion>
, Irkutskaya Province, Yakutia).
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Thomson, 1859: 18
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Olivier, 1793
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(
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(
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<subSubSection id="C34918832F1BFFCCDDDFF9D8C5E60807" box="[245,730,1548,1571]" pageId="11" pageNumber="441" type="description">
<paragraph id="8BEC4B082F1BFFCCDDDFF9D8C5E60807" blockId="11.[211,765,1489,1571]" box="[245,730,1548,1571]" pageId="11" pageNumber="441">
<figureCitation id="1368578D2F1BFFCCDDDFF9D8C6510807" box="[245,365,1548,1571]" captionStart-0="Figs 9–12" captionStart-1="Figs 13–16" captionStartId-0="4.[190,228,1943,1962]" captionStartId-1="5.[188,226,1968,1987]" captionTargetBox-0="[280,1330,464,1857]" captionTargetBox-1="[264,1302,572,1891]" captionTargetId-0="figure-119@4.[276,1334,442,1873]" captionTargetId-1="figure-144@5.[258,1316,565,1922]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Figs 9–12. Enochrus, habitus in dorsal view. 9–10 — Enochrus simulans, 9 — male, North Korea, 10 — female, North Korea, 11–12 — E. umbratus stat.rev., 11 — lectotype of Philydrus umbratus, 12 — female, paralectotype of Philydrus umbratus, Japan, Chiuzenji. Scale bar: 1 mm. Рис. 9–12. Enochrus, габитус сверху. 9–10 — Enochrus simulans, 9 — самец, СевернаЯ КореЯ, 10 — самка, СевернаЯ КореЯ, 11–12 — E. umbratus stat.rev., 11 — лектотип Philydrus umbratus, 12 — самка, паралектотип Philydrus umbratus, ЯпониЯ, ТюЗенджи. МасштабнаЯ линейка: 1 мм." captionText-1="Figs 13–16. Enochrus elytral contour, dorsal view. 13-14 — Enochrus umbratus stat. rev., 13 — male, 14 — female. 15–16 — E. simulans, 15 — male, 16 — female. Scale bar: 1 mm. Рис. 13–16. Контур надкрылий Enochrus, вид сверху. 13–14 — Enochrus umbratus stat. rev., 13 — самец, 14 — самка. 15–16 — E. simulans, 15 — самец, 16 — самка. МасштабнаЯ линейка: 1 мм." figureDoi-0="http://doi.org/10.5281/zenodo.14722318" figureDoi-1="http://doi.org/10.5281/zenodo.14722320" httpUri-0="https://zenodo.org/record/14722318/files/figure.png" httpUri-1="https://zenodo.org/record/14722320/files/figure.png" pageId="11" pageNumber="441">Figs 11–14</figureCitation>
,
<figureCitation id="1368578D2F1BFFCCDC53F9D8C6810807" box="[377,445,1548,1571]" captionStart="Figs 17–24" captionStartId="6.[189,227,1861,1880]" captionTargetBox="[200,1405,365,1769]" captionTargetId="figure-6@6.[200,1410,360,1774]" captionTargetPageId="6" captionText="Figs 17–24. Enochrus, head, dorsal view. 17–18 — E. umbratus stat. rev., 17 — lectotype of Philydrus umbratus, 18 — female, paralectotype of Philydrus umbratus, Japan, Chiuzenji. 19-24 — Enochrus simulans, 19 — holotype of Philydrus simulans, 20 — male, Japan, Tochigi-ken, 21 — male, Russia, Primorsky Krai, Lazo, 22 — male, North Korea, Khoankhado, 23 — female, Japan, Shimane-ken, 24 — female, Russia, Primorsky Krai, Troitskoe. Scale bar: 0.5 mm. Рис. 17–24. Enochrus, голова, вид сверху. 17–18 — E. umbratus stat. rev., 17 — лектотип Philydrus umbratus, 18 — самка, паралектотип Philydrus umbratus, ЯпониЯ, ЧиуЗенджи. 19-24 — E. simulans, 19 — голотип Philydrus simulans, 20 — самец, ЯпониЯ, Тотиги-кен, 21 — самец, РоссиЯ, Приморский край, ЛаЗо, 22 — самец, СевернаЯ КореЯ, Хоанхадо, 23 — самка, ЯпониЯ, СиманЭ-кен, 24 — самка, РоссиЯ, Приморский край, Троицкое. МасштабнаЯ линейка: 0,5 мм." figureDoi="http://doi.org/10.5281/zenodo.14722322" httpUri="https://zenodo.org/record/14722322/files/figure.png" pageId="11" pageNumber="441">17–18</figureCitation>
,
<figureCitation id="1368578D2F1BFFCCDCE3F9D8C5320807" box="[457,526,1548,1571]" captionStart="Figs 25–33" captionStartId="7.[190,228,1861,1880]" captionTargetBox="[245,1366,306,1810]" captionTargetId="figure-10@7.[241,1367,302,1822]" captionTargetPageId="7" captionText="Figs 25–33. Enochrus, base of elytra. 25–26 — E. umbratus stat.rev., 25 — lectotype of Philydrus umbratus, 26 — female, paralectotype of Philydrus umbratus, Japan, Chiuzenji. 27–33 — E. simulans, 27 — holotype of Philydrus simulans, 28 — male, paralectotype of Philydrus umbratus, Japan, Hakodate, 29 — male, Japan, Higashikami, 30 — male, Japan, Fujioka-machi, 31 — male, Russia, Kunashir Isl., 32 — male, Russia, Primorsky Krai, Lazo, 33 — female, North Korea, Khoankhado. Рис. 25–33. Enochrus, основание надкрылий. 25–26 — E. umbratus stat.rev., 25 — лектотип Philydrus umbratus, 26 — самка, паралектотип Philydrus umbratus, ЯпониЯ, ЧиуЗенджи. 27–33 — E. simulans, 27 — голотип Philydrus simulans, 28 — самец, паралектотип Philydrus umbratus, ЯпониЯ, ХакодатЭ, 29 — самец, ЯпониЯ, Хигашиками, 30 — самец, ЯпониЯ, ФудЗиока-мати, 31 — самец, РоссиЯ, о-в Кунашир, 32 — самец, РоссиЯ, Приморский край, ЛаЗо, 33 — самка, СевернаЯ КореЯ, Хоанхадо." figureDoi="http://doi.org/10.5281/zenodo.14722324" httpUri="https://zenodo.org/record/14722324/files/figure.png" pageId="11" pageNumber="441">25–26</figureCitation>
,
<figureCitation id="1368578D2F1BFFCCDF33F9D8C5620807" box="[537,606,1548,1571]" captionStart="Figs 34–47" captionStartId="8.[190,227,1790,1809]" captionTargetBox="[203,1405,496,1726]" captionTargetId="figure-70@8.[200,1410,488,1732]" captionTargetPageId="8" captionText="Figs 34–47. Enochrus, scutellum, claw and protarsomeres of male. 34–43 — scutellum. 44–47 — claw and protarsomeres of male. 34–41, 45–47 — E. simulans, 42–44 — E. umbratus stat.rev., 34, 45 — holotype of Philydrus simulans, 35 — male, Japan, Higashikami. 36–38 — males, Japan, Fujioka-machi, 39 — female, Japan, Okinoshima, 40 — male, Russia, Primorsky Krai, Lazo, 41 — female, North Korea, Khoankhado. 42, 44 — lectotype of Philydrus umbratus, 43 — female, paralectotype of Philydrus umbratus, Japan, Chiuzenji. 46 — male, paralectotype of Philydrus umbratus, Japan, Hakodate, 47 — male, E. simulans, Russia, Kunashir Isl. Scale bar for figs 34–43: 0.3 mm. Scale bar for figs 44–47: 0.5 mm. Рис. 34–47. Enochrus, щиток, коготок и протарсомеры самца. 34–43 — щиток. 44–47 — коготок и протарсомеры самца. 34–41, 45–47 — E. simulans, 42–44 — E. umbratus stat.rev., 34, 45 — голотип Philydrus simulans, 35 — самец, ЯпониЯ, Хигашиками. 36–38 — самцы, ЯпониЯ, ФудЗиока-мати, 39 — самка, ЯпониЯ, Окиносима, 40 — самец, РоссиЯ, Приморский край, ЛаЗо, 41 — самка, СевернаЯ КореЯ, Хоанхадо. 42, 44 — лектотип Philydrus umbratus, 43 — самка, паралектотип Philydrus umbratus, ЯпониЯ, ЧиуЗенджи. 46 — самец, паралектотип Philydrus umbratus, ЯпониЯ, ХакодатЭ, 47 — самец, E. simulans, РоссиЯ, о-в Кунашир. МасштабнаЯ линейка длЯ рис. 34–43: 0,3 мм. МасштабнаЯ линейка длЯ рис. 44–47: 0,5 мм." figureDoi="http://doi.org/10.5281/zenodo.14722326" httpUri="https://zenodo.org/record/14722326/files/figure.png" pageId="11" pageNumber="441">42–44</figureCitation>
,
<figureCitation id="1368578D2F1BFFCCDF43F9D8C5BA0807" box="[617,646,1548,1571]" captionStart="Figs 48–51" captionStartId="9.[190,227,1939,1958]" captionTargetBox="[306,1285,971,1886]" captionTargetId="figure-427@9.[306,1304,971,1890]" captionTargetPageId="9" captionText="Figs 48–51. Enochrus, mesoventral elevation. 48 — E. umbratus stat. rev., male, paralectotype of Philydrus umbratus. 49–51 — E. simulans, 49 — male, Russia, Kunashir Isl., 50 — male, Russia, Primorsky Krai, Lazo, 51 — male (juvenile), Russia, Primorsky Krai, Lazo. Scale bar: 0.5 mm. Рис. 48–51. Enochrus, меЗовентральный отросток. 48 — E. umbratus stat. rev., самец, паралектотип Philydrus umbratus. 49–51 — E. simulans, 49 — самец, РоссиЯ, о-в Кунашир, 50 — самец, РоссиЯ, Приморский край, ЛаЗо, 51 — самец (недоокрашенный), РоссиЯ, Приморский край, ЛаЗо. МасштабнаЯ линейка: 0,5 мм." figureDoi="http://doi.org/10.5281/zenodo.14722328" httpUri="https://zenodo.org/record/14722328/files/figure.png" pageId="11" pageNumber="441">48</figureCitation>
,
<figureCitation id="1368578D2F1BFFCCDFBBF9D8C5EB0807" box="[657,727,1548,1571]" captionStart="Figs 52–56" captionStartId="10.[189,227,1395,1414]" captionTargetBox="[191,785,306,1321]" captionTargetId="figure-684@10.[188,786,302,1327]" captionTargetPageId="10" captionText="Figs 52–56. Enochrus, aedeagus. 52, 53 — E. umbratus stat.rev., 53 — paralectotype of Philydrus umbratus, 53 — apex of aedeagus, lectotype of Philydrus umbratus, 54–56 — E. simulans, 54 — holotype of Philydrus simulans, 55 — E. simulans, aedeagus, Russia, Kunashir Isl. 56 — apex of aedeagus, holotype of Philydrus simulans. Scale bar: 0.5 mm. Рис. 52–56. Enochrus, Эдеагус. 52, 53 — E. umbratus stat.rev., 53 — паралектотип Philydrus umbratus, 53 — вершина Эдеагуса, лектотип Philydrus umbratus, 54–56 — E. simulans, 54 — голотип Philydrus simulans, 55 — E. simulans, aedeagus, РоссиЯ, остров Кунашир. 56 — вершина Эдеагуса, голотип Philydrus simulans. МасштабнаЯ линейка: 0,5 мм." figureDoi="http://doi.org/10.5281/zenodo.14722330" httpUri="https://zenodo.org/record/14722330/files/figure.png" pageId="11" pageNumber="441">52, 53</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34918832F1BFFCCDDC9F99EC5490879" box="[227,629,1610,1629]" pageId="11" pageNumber="441" type="reference_group">
<paragraph id="8BEC4B082F1BFFCCDDC9F99EC5490879" blockId="11.[189,786,1610,2056]" box="[227,629,1610,1629]" pageId="11" pageNumber="441">
<treatmentCitationGroup id="AB436C262F1BFFCCDDC9F99EC5490879" box="[227,629,1610,1629]" pageId="11" pageNumber="441">
<treatmentCitation id="0AF26D192F1BFFCCDDC9F99EC51F0879" author="Sharp D." box="[227,547,1610,1629]" page="454" pageId="11" pageNumber="441" year="1884">
<taxonomicName id="4C53308B2F1BFFCCDDC9F99EC51F0879" authority="Sharp, 1884: 454" authorityName="Sharp" authorityPageNumber="454" authorityYear="1884" box="[227,547,1610,1629]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="B927971A2F1BFFCCDDC9F99EC6B00879" box="[227,396,1610,1629]" italics="true" pageId="11" pageNumber="441">Philydrus umbratus</emphasis>
<bibRefCitation id="EFC236F92F1BFFCCDCBBF99EC51F0879" author="Sharp D." box="[401,547,1610,1629]" pageId="11" pageNumber="441" pagination="439 - 464" refId="ref10112" refString="Sharp D. 1884. The water beetles of Japan // Transactions of the Entomological Society of London 1884. P. 439 - 464. https: // doi. org / 10.1111 / j. 1365 - 2311.1884. tb 01616. x" type="book chapter" year="1884">Sharp, 1884: 454</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(partim).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34918832F1BFFCCDDC9F9B7C49A083C" pageId="11" pageNumber="441" type="materials_examined">
<paragraph id="8BEC4B082F1BFFCCDDC9F9B7C49A083C" blockId="11.[189,786,1610,2056]" lastBlockId="11.[824,1422,1352,2067]" pageId="11" pageNumber="441">
MATERIAL. Type material.
<typeStatus id="54E8F5AA2F1BFFCCDF2BF9B7C55D085C" box="[513,609,1635,1656]" pageId="11" pageNumber="441" type="lectotype">Lectotype</typeStatus>
(present designation). Male,
<collectingCountry id="F3440B982F1BFFCCDC13F9AAC64E08B7" box="[313,370,1662,1683]" name="Japan" pageId="11" pageNumber="441">Japan</collectingCountry>
, “
<taxonomicName id="4C53308B2F1BFFCCDCA3F9AAC57608B7" authorityName="Sharp" authorityYear="1884" box="[393,586,1662,1683]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="B927971A2F1BFFCCDCA3F9AAC57608B7" box="[393,586,1662,1683]" italics="true" pageId="11" pageNumber="441">Philydrus umbratus</emphasis>
</taxonomicName>
, Types D.S., [Lake] Chiuzenji [Nikko,
<collectingRegion id="499785EA2F1BFFCCDC44F94CC5190889" box="[366,549,1688,1709]" country="Japan" name="Tochigi" pageId="11" pageNumber="441">Tochigi Prefecture</collectingRegion>
],
<collectingCountry id="F3440B982F1BFFCCDF18F94CC5570889" box="[562,619,1688,1709]" name="Japan" pageId="11" pageNumber="441">Japan</collectingCountry>
,
<date id="FFED6DC82F1BFFCCDF5EF94CC5EA0889" box="[628,726,1688,1709]" pageId="11" pageNumber="441" value="1881-08-24">24.8.1881</date>
, Lewis” [Sharp’s writing on the face of the card], “Type” [printed label], “
<collectingCountry id="F3440B982F1BFFCCDC21F91AC67A08C7" box="[267,326,1742,1763]" name="Japan" pageId="11" pageNumber="441">Japan</collectingCountry>
, G. Lewis” [printed label], “Sharp Coll. 1905- 313” [printed label], ”
<typeStatus id="54E8F5AA2F1BFFCCDCB9F93CC53E08D9" box="[403,514,1768,1789]" pageId="11" pageNumber="441" type="lectotype">Lectotypus</typeStatus>
,
<taxonomicName id="4C53308B2F1BFFCCDF27F93CC7D1093C" authority="Sharp, 1884" authorityName="Sharp" authorityYear="1884" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="B927971A2F1BFFCCDF27F93CC5F108D9" box="[525,717,1768,1789]" italics="true" pageId="11" pageNumber="441">Philydrus umbratus</emphasis>
<bibRefCitation id="EFC236F92F1BFFCCDFFEF93CC7D1093C" author="Sharp D." pageId="11" pageNumber="441" pagination="439 - 464" refId="ref10112" refString="Sharp D. 1884. The water beetles of Japan // Transactions of the Entomological Society of London 1884. P. 439 - 464. https: // doi. org / 10.1111 / j. 1365 - 2311.1884. tb 01616. x" type="book chapter" year="1884">Sharp, 1884</bibRefCitation>
</taxonomicName>
<specimenCount id="9D5580812F1BFFCCDDDEF8D7C63A093C" box="[244,262,1795,1816]" count="1" pageId="11" pageNumber="441" type="male">♂</specimenCount>
, des. Shödl, 1996” [red label Shödl 's writing],
<typeStatus id="54E8F5AA2F1BFFCCDFF9F8D7C7D40917" pageId="11" pageNumber="441" type="lectotype">Lectotype</typeStatus>
” [blue bordered round printed label]; “
<taxonomicName id="4C53308B2F1BFFCCDF7CF8CAC7CB0969" authority="Sharp" authorityName="Sharp" authorityYear="1884" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="B927971A2F1BFFCCDF7CF8CAC42E0917" box="[598,786,1822,1843]" italics="true" pageId="11" pageNumber="441">Philydrus umbratus</emphasis>
Sharp
</taxonomicName>
,
<typeStatus id="54E8F5AA2F1BFFCCDC2BF8ECC6670969" box="[257,347,1848,1869]" pageId="11" pageNumber="441" type="lectotype">lectotype</typeStatus>
, S. Ryndevich &amp; A. Prokin des., 2021” [white printed label].
<typeStatus id="54E8F5AA2F1BFFCCDC7CF887C6D9094C" box="[342,485,1875,1896]" pageId="11" pageNumber="441" type="paralectotype">Paralectotypes</typeStatus>
.
<collectingCountry id="F3440B982F1BFFCCDCDEF887C512094C" box="[500,558,1875,1896]" name="Japan" pageId="11" pageNumber="441">Japan</collectingCountry>
:
<specimenCount id="9D5580812F1BFFCCDF14F887C59D094C" box="[574,673,1875,1896]" count="2" pageId="11" pageNumber="441" type="female">2 females</specimenCount>
[originally mounted on one card with
<typeStatus id="54E8F5AA2F1BFFCCDCECF8BAC51F09A7" box="[454,547,1902,1923]" pageId="11" pageNumber="441" type="lectotype">lectotype</typeStatus>
], same data as for
<typeStatus id="54E8F5AA2F1BFFCCDFF1F8BAC7D509B9" pageId="11" pageNumber="441" type="lectotype">lectotype</typeStatus>
, but “
<typeStatus id="54E8F5AA2F1BFFCCDC0AF85CC68E09B9" box="[288,434,1928,1949]" pageId="11" pageNumber="441" type="paralectotype">Paralectotypus</typeStatus>
,
<taxonomicName id="4C53308B2F1BFFCCDC91F85CC5C809B9" authority="Sharp, 1884" authorityName="Sharp" authorityYear="1884" box="[443,756,1928,1949]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="B927971A2F1BFFCCDC91F85CC54609B9" box="[443,634,1928,1949]" italics="true" pageId="11" pageNumber="441">Philydrus umbratus</emphasis>
<bibRefCitation id="EFC236F92F1BFFCCDFAAF85CC5C809B9" author="Sharp D." box="[640,756,1928,1949]" pageId="11" pageNumber="441" pagination="439 - 464" refId="ref10112" refString="Sharp D. 1884. The water beetles of Japan // Transactions of the Entomological Society of London 1884. P. 439 - 464. https: // doi. org / 10.1111 / j. 1365 - 2311.1884. tb 01616. x" type="book chapter" year="1884">Sharp, 1884</bibRefCitation>
</taxonomicName>
<specimenCount id="9D5580812F1BFFCCDFD0F85CC43009B9" box="[762,780,1928,1949]" count="1" pageId="11" pageNumber="441" type="female">♀</specimenCount>
, des. Shödl, 1996” [2 red label Shödl 's writing], “
<typeStatus id="54E8F5AA2F1BFFCCDF84F877C7D409F7" pageId="11" pageNumber="441" type="paralectotype">Paralectotype</typeStatus>
” [blue bordered round printed label]; “
<taxonomicName id="4C53308B2F1BFFCCDF4DF86AC62509C9" authority="Sharp" authorityName="Sharp" authorityYear="1884" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="B927971A2F1BFFCCDF4DF86AC7E509C9" italics="true" pageId="11" pageNumber="441">Philydrus umbratus</emphasis>
Sharp
</taxonomicName>
,
<typeStatus id="54E8F5AA2F1BFFCCDC09F80CC69B09C9" box="[291,423,2008,2029]" pageId="11" pageNumber="441" type="paralectotype">paralectotype</typeStatus>
, S. Ryndevich &amp; A. Prokin des., 2021 [2 red printed labels]”; male and female [originally mounted on one card] “
<taxonomicName id="4C53308B2F1BFFCCDEEEFA9CC3CD0B79" authority=", Chiuzenji" authorityName="Chiuzenji" box="[964,1265,1352,1373]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="B927971A2F1BFFCCDEEEFA9CC3B80B79" box="[964,1156,1352,1373]" italics="true" pageId="11" pageNumber="441">Philydrus umbratus</emphasis>
, Chiuzenji
</taxonomicName>
, 24.8.81,
<collectingCountry id="F3440B982F1BFFCCD87AFA9CC2B50B79" box="[1360,1417,1352,1373]" name="Japan" pageId="11" pageNumber="441">Japan</collectingCountry>
, Lewis“[Sharp’s writing], “
<collectingCountry id="F3440B982F1BFFCCD915FAB7C3460B5C" box="[1087,1146,1379,1400]" name="Japan" pageId="11" pageNumber="441">Japan</collectingCountry>
, G. Lewis” [printed label], “Sharp Coll. 1905-313” [printed label], ”
<typeStatus id="54E8F5AA2F1BFFCCD9EDFAAAC2650BB7" box="[1223,1369,1406,1427]" pageId="11" pageNumber="441" type="paralectotype">Paralectotypus</typeStatus>
,
<taxonomicName id="4C53308B2F1BFFCCD84EFAAAC3700B89" authority="Sharp, 1884" authorityName="Sharp" authorityYear="1884" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="B927971A2F1BFFCCD84EFAAAC4E80B89" italics="true" pageId="11" pageNumber="441">Philydrus umbratus</emphasis>
<bibRefCitation id="EFC236F92F1BFFCCDEF3FA4CC3700B89" author="Sharp D." box="[985,1100,1432,1453]" pageId="11" pageNumber="441" pagination="439 - 464" refId="ref10112" refString="Sharp D. 1884. The water beetles of Japan // Transactions of the Entomological Society of London 1884. P. 439 - 464. https: // doi. org / 10.1111 / j. 1365 - 2311.1884. tb 01616. x" type="book chapter" year="1884">Sharp, 1884</bibRefCitation>
</taxonomicName>
<specimenCount id="9D5580812F1BFFCCD97BFA4CC35F0B89" box="[1105,1123,1432,1453]" count="1" pageId="11" pageNumber="441" type="male">♂</specimenCount>
(&amp;
<specimenCount id="9D5580812F1BFFCCD9A8FA4CC3A80B89" box="[1154,1172,1432,1453]" count="1" pageId="11" pageNumber="441" type="female">♀</specimenCount>
), des. Shödl, 1996” [2 red labels Shödl 's writing], “
<typeStatus id="54E8F5AA2F1BFFCCD901FA67C38E0BEC" box="[1067,1202,1459,1480]" pageId="11" pageNumber="441" type="paralectotype">Paralectotype</typeStatus>
” [blue bordered round printed label]; “
<taxonomicName id="4C53308B2F1BFFCCDEFDFA1AC3E70BC7" authority="Sharp" authorityName="Sharp" authorityYear="1884" box="[983,1243,1486,1507]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="B927971A2F1BFFCCDEFDFA1AC3A40BC7" box="[983,1176,1486,1507]" italics="true" pageId="11" pageNumber="441">Philydrus umbratus</emphasis>
Sharp
</taxonomicName>
,
<typeStatus id="54E8F5AA2F1BFFCCD9C3FA1AC2510BC7" box="[1257,1389,1486,1507]" pageId="11" pageNumber="441" type="paralectotype">paralectotype</typeStatus>
, S. Ryndevich &amp; A. Prokin des., 2021 [2 red printed labels] (all in
<collectionCode id="ED42D3CD2F1BFFCCDE7BF9D7C4A0083C" box="[849,924,1539,1560]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="11" pageNumber="441">NHML</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C34918832F1BFFCCDE74F9CAC2480907" pageId="11" pageNumber="441" type="discussion">
<paragraph id="8BEC4B082F1BFFCCDE74F9CAC35E084C" blockId="11.[824,1422,1352,2067]" pageId="11" pageNumber="441">
REMARKS. Type material has labels by Stefan Schödl (1957–2005), but the
<typeStatus id="54E8F5AA2F1BFFCCD926F9ECC3580869" box="[1036,1124,1592,1613]" pageId="11" pageNumber="441" type="lectotype">lectotype</typeStatus>
of the species had never been officially designated until now.
</paragraph>
<paragraph id="8BEC4B082F1BFFCCDE74F9BAC20208C9" blockId="11.[824,1422,1352,2067]" pageId="11" pageNumber="441">
<materialsCitation id="3B3B41552F1BFFCCDE74F9BAC20208C9" county="The" location="Yokohama" municipality="Nikko" pageId="11" pageNumber="441" specimenCount="2" stateProvince="Niigata" typeStatus="lectotype">
<collectingCounty id="628D33842F1BFFCCDE74F9BAC4BF08A7" box="[862,899,1646,1667]" pageId="11" pageNumber="441">The</collectingCounty>
species was described from
<collectingMunicipality id="6B88D1722F1BFFCCD985F9BAC3D108A7" box="[1199,1261,1646,1667]" pageId="11" pageNumber="441">Nikko</collectingMunicipality>
,
<collectingRegion id="499785EA2F1BFFCCD9D1F9BAC27808A7" box="[1275,1348,1646,1667]" country="Japan" name="Niigata" pageId="11" pageNumber="441">Niigata</collectingRegion>
,
<location id="8E8C1DD32F1BFFCCD878F9BAC45008B9" LSID="urn:lsid:plazi:treatment:03FAFA1E2F1BFFCBDDF9FA05C4830DFB:8E8C1DD32F1BFFCCD878F9BAC45008B9" county="The" municipality="Nikko" name="Yokohama" pageId="11" pageNumber="441" stateProvince="Niigata">Yokohama</location>
and
<location id="8E8C1DD32F1BFFCCDEB7F95CC4C708B9" LSID="urn:lsid:plazi:treatment:03FAFA1E2F1BFFCBDDF9FA05C4830DFB:8E8C1DD32F1BFFCCDEB7F95CC4C708B9" box="[925,1019,1672,1693]" county="The" municipality="Nikko" name="Hakodate" pageId="11" pageNumber="441" stateProvince="Niigata">Hakodate</location>
, but we haven't studied the type material from Yokohama.
<typeStatus id="54E8F5AA2F1BFFCCDECEF977C378089C" box="[996,1092,1699,1720]" pageId="11" pageNumber="441" type="lectotype">Lectotype</typeStatus>
is placed on one card (left) with
<specimenCount id="9D5580812F1BFFCCDE12F96AC4D008F7" box="[824,1004,1726,1747]" count="2" pageId="11" pageNumber="441" type="generic" typeStatus="paralectotype">two paralectotypes</specimenCount>
(females). Male genitalia of males mounted on a plastic plates and placed below the card on a pin.
</materialsCitation>
</paragraph>
<paragraph id="8BEC4B082F1BFFCCDE74F927C2480907" blockId="11.[824,1422,1352,2067]" pageId="11" pageNumber="441">
<typeStatus id="54E8F5AA2F1BFFCCDE74F927C4D6092C" box="[862,1002,1779,1800]" pageId="11" pageNumber="441" type="paralectotype">Paralectotypes</typeStatus>
of
<taxonomicName id="4C53308B2F1BFFCCD923F927C3FB092C" authorityName="Sharp" authorityYear="1884" box="[1033,1223,1779,1800]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="B927971A2F1BFFCCD923F927C3FB092C" box="[1033,1223,1779,1800]" italics="true" pageId="11" pageNumber="441">Philydrus umbratus</emphasis>
</taxonomicName>
) from Hakodate and
<collectingRegion id="499785EA2F1BFFCCDE12F8DAC4430907" box="[824,895,1806,1827]" country="Japan" name="Niigata" pageId="11" pageNumber="441">Niigata</collectingRegion>
(
<figureCitation id="1368578D2F1BFFCCDEA1F8DAC4EA0907" box="[907,982,1806,1827]" captionStart="Figs 25–33" captionStartId="7.[190,228,1861,1880]" captionTargetBox="[245,1366,306,1810]" captionTargetId="figure-10@7.[241,1367,302,1822]" captionTargetPageId="7" captionText="Figs 25–33. Enochrus, base of elytra. 25–26 — E. umbratus stat.rev., 25 — lectotype of Philydrus umbratus, 26 — female, paralectotype of Philydrus umbratus, Japan, Chiuzenji. 27–33 — E. simulans, 27 — holotype of Philydrus simulans, 28 — male, paralectotype of Philydrus umbratus, Japan, Hakodate, 29 — male, Japan, Higashikami, 30 — male, Japan, Fujioka-machi, 31 — male, Russia, Kunashir Isl., 32 — male, Russia, Primorsky Krai, Lazo, 33 — female, North Korea, Khoankhado. Рис. 25–33. Enochrus, основание надкрылий. 25–26 — E. umbratus stat.rev., 25 — лектотип Philydrus umbratus, 26 — самка, паралектотип Philydrus umbratus, ЯпониЯ, ЧиуЗенджи. 27–33 — E. simulans, 27 — голотип Philydrus simulans, 28 — самец, паралектотип Philydrus umbratus, ЯпониЯ, ХакодатЭ, 29 — самец, ЯпониЯ, Хигашиками, 30 — самец, ЯпониЯ, ФудЗиока-мати, 31 — самец, РоссиЯ, о-в Кунашир, 32 — самец, РоссиЯ, Приморский край, ЛаЗо, 33 — самка, СевернаЯ КореЯ, Хоанхадо." figureDoi="http://doi.org/10.5281/zenodo.14722324" httpUri="https://zenodo.org/record/14722324/files/figure.png" pageId="11" pageNumber="441">Figs 28</figureCitation>
,
<figureCitation id="1368578D2F1BFFCCDECAF8DAC3220907" box="[992,1054,1806,1827]" captionStart="Figs 34–47" captionStartId="8.[190,227,1790,1809]" captionTargetBox="[203,1405,496,1726]" captionTargetId="figure-70@8.[200,1410,488,1732]" captionTargetPageId="8" captionText="Figs 34–47. Enochrus, scutellum, claw and protarsomeres of male. 34–43 — scutellum. 44–47 — claw and protarsomeres of male. 34–41, 45–47 — E. simulans, 42–44 — E. umbratus stat.rev., 34, 45 — holotype of Philydrus simulans, 35 — male, Japan, Higashikami. 36–38 — males, Japan, Fujioka-machi, 39 — female, Japan, Okinoshima, 40 — male, Russia, Primorsky Krai, Lazo, 41 — female, North Korea, Khoankhado. 42, 44 — lectotype of Philydrus umbratus, 43 — female, paralectotype of Philydrus umbratus, Japan, Chiuzenji. 46 — male, paralectotype of Philydrus umbratus, Japan, Hakodate, 47 — male, E. simulans, Russia, Kunashir Isl. Scale bar for figs 34–43: 0.3 mm. Scale bar for figs 44–47: 0.5 mm. Рис. 34–47. Enochrus, щиток, коготок и протарсомеры самца. 34–43 — щиток. 44–47 — коготок и протарсомеры самца. 34–41, 45–47 — E. simulans, 42–44 — E. umbratus stat.rev., 34, 45 — голотип Philydrus simulans, 35 — самец, ЯпониЯ, Хигашиками. 36–38 — самцы, ЯпониЯ, ФудЗиока-мати, 39 — самка, ЯпониЯ, Окиносима, 40 — самец, РоссиЯ, Приморский край, ЛаЗо, 41 — самка, СевернаЯ КореЯ, Хоанхадо. 42, 44 — лектотип Philydrus umbratus, 43 — самка, паралектотип Philydrus umbratus, ЯпониЯ, ЧиуЗенджи. 46 — самец, паралектотип Philydrus umbratus, ЯпониЯ, ХакодатЭ, 47 — самец, E. simulans, РоссиЯ, о-в Кунашир. МасштабнаЯ линейка длЯ рис. 34–43: 0,3 мм. МасштабнаЯ линейка длЯ рис. 44–47: 0,5 мм." figureDoi="http://doi.org/10.5281/zenodo.14722326" httpUri="https://zenodo.org/record/14722326/files/figure.png" pageId="11" pageNumber="441">45–46</figureCitation>
) are belongs to
<taxonomicName id="4C53308B2F1BFFCCD99CF8DAC2510907" baseAuthorityName="Sharp" baseAuthorityYear="1873" box="[1206,1389,1806,1827]" class="Insecta" family="Hydrophilidae" genus="Enochrus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="B927971A2F1BFFCCD99CF8DAC2510907" box="[1206,1389,1806,1827]" italics="true" pageId="11" pageNumber="441">Enochrus simulans</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34918832F1BFFCBDE74F8FCC48F0CCB" lastPageId="12" lastPageNumber="442" pageId="11" pageNumber="441" type="description">
<paragraph id="8BEC4B082F1BFFCBDE74F8FCC5EB0C3D" blockId="11.[824,1422,1352,2067]" lastBlockId="12.[189,787,303,1364]" lastPageId="12" lastPageNumber="442" pageId="11" pageNumber="441">
REDESCRIPTION.
<emphasis id="B927971A2F1BFFCCD90FF8FCC3F50919" box="[1061,1225,1832,1853]" italics="true" pageId="11" pageNumber="441">Form and colour</emphasis>
. Body oval, slightly convex (
<figureCitation id="1368578D2F1BFFCCDEA1F897C4C7097C" box="[907,1019,1859,1880]" captionStart-0="Figs 9–12" captionStart-1="Figs 13–16" captionStartId-0="4.[190,228,1943,1962]" captionStartId-1="5.[188,226,1968,1987]" captionTargetBox-0="[280,1330,464,1857]" captionTargetBox-1="[264,1302,572,1891]" captionTargetId-0="figure-119@4.[276,1334,442,1873]" captionTargetId-1="figure-144@5.[258,1316,565,1922]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Figs 9–12. Enochrus, habitus in dorsal view. 9–10 — Enochrus simulans, 9 — male, North Korea, 10 — female, North Korea, 11–12 — E. umbratus stat.rev., 11 — lectotype of Philydrus umbratus, 12 — female, paralectotype of Philydrus umbratus, Japan, Chiuzenji. Scale bar: 1 mm. Рис. 9–12. Enochrus, габитус сверху. 9–10 — Enochrus simulans, 9 — самец, СевернаЯ КореЯ, 10 — самка, СевернаЯ КореЯ, 11–12 — E. umbratus stat.rev., 11 — лектотип Philydrus umbratus, 12 — самка, паралектотип Philydrus umbratus, ЯпониЯ, ТюЗенджи. МасштабнаЯ линейка: 1 мм." captionText-1="Figs 13–16. Enochrus elytral contour, dorsal view. 13-14 — Enochrus umbratus stat. rev., 13 — male, 14 — female. 15–16 — E. simulans, 15 — male, 16 — female. Scale bar: 1 mm. Рис. 13–16. Контур надкрылий Enochrus, вид сверху. 13–14 — Enochrus umbratus stat. rev., 13 — самец, 14 — самка. 15–16 — E. simulans, 15 — самец, 16 — самка. МасштабнаЯ линейка: 1 мм." figureDoi-0="http://doi.org/10.5281/zenodo.14722318" figureDoi-1="http://doi.org/10.5281/zenodo.14722320" httpUri-0="https://zenodo.org/record/14722318/files/figure.png" httpUri-1="https://zenodo.org/record/14722320/files/figure.png" pageId="11" pageNumber="441">Figs 11–14</figureCitation>
), length
<quantity id="4CABE6ED2F1BFFCCD978F897C3F5097C" box="[1106,1225,1859,1880]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="6.1" metricValueMin="5.4" pageId="11" pageNumber="441" unit="mm" value="5.75" valueMax="6.1" valueMin="5.4">5.4–6.1 mm</quantity>
, width
<quantity id="4CABE6ED2F1BFFCCD838F897C2B6097C" box="[1298,1418,1859,1880]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.3" metricValueMin="2.9" pageId="11" pageNumber="441" unit="mm" value="3.0999999999999996" valueMax="3.3" valueMin="2.9">2.9–3.3 mm</quantity>
. Maximum body width in basal part, in front of middle of elytra. Lateral margins of the elytra is smoothly rounded at the apices (
<figureCitation id="1368578D2F1BFFCCDE15F847C493098C" box="[831,943,1939,1960]" captionStart="Figs 13–16" captionStartId="5.[188,226,1968,1987]" captionTargetBox="[264,1302,572,1891]" captionTargetId="figure-144@5.[258,1316,565,1922]" captionTargetPageId="5" captionText="Figs 13–16. Enochrus elytral contour, dorsal view. 13-14 — Enochrus umbratus stat. rev., 13 — male, 14 — female. 15–16 — E. simulans, 15 — male, 16 — female. Scale bar: 1 mm. Рис. 13–16. Контур надкрылий Enochrus, вид сверху. 13–14 — Enochrus umbratus stat. rev., 13 — самец, 14 — самка. 15–16 — E. simulans, 15 — самец, 16 — самка. МасштабнаЯ линейка: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.14722320" httpUri="https://zenodo.org/record/14722320/files/figure.png" pageId="11" pageNumber="441">Figs 13–14</figureCitation>
). Dorsal side reddish-yellow to yellowish-brown. Basal part of the head and central triangular spot are black. Central dark spot reaches the labrum and occupies most part of head. Triangular spot of the female is larger than that of male. Labrum of male and female dark (
<figureCitation id="1368578D2F1BFFCCD9A7F82AC3C30637" box="[1165,1279,2046,2067]" captionStart="Figs 17–24" captionStartId="6.[189,227,1861,1880]" captionTargetBox="[200,1405,365,1769]" captionTargetId="figure-6@6.[200,1410,360,1774]" captionTargetPageId="6" captionText="Figs 17–24. Enochrus, head, dorsal view. 17–18 — E. umbratus stat. rev., 17 — lectotype of Philydrus umbratus, 18 — female, paralectotype of Philydrus umbratus, Japan, Chiuzenji. 19-24 — Enochrus simulans, 19 — holotype of Philydrus simulans, 20 — male, Japan, Tochigi-ken, 21 — male, Russia, Primorsky Krai, Lazo, 22 — male, North Korea, Khoankhado, 23 — female, Japan, Shimane-ken, 24 — female, Russia, Primorsky Krai, Troitskoe. Scale bar: 0.5 mm. Рис. 17–24. Enochrus, голова, вид сверху. 17–18 — E. umbratus stat. rev., 17 — лектотип Philydrus umbratus, 18 — самка, паралектотип Philydrus umbratus, ЯпониЯ, ЧиуЗенджи. 19-24 — E. simulans, 19 — голотип Philydrus simulans, 20 — самец, ЯпониЯ, Тотиги-кен, 21 — самец, РоссиЯ, Приморский край, ЛаЗо, 22 — самец, СевернаЯ КореЯ, Хоанхадо, 23 — самка, ЯпониЯ, СиманЭ-кен, 24 — самка, РоссиЯ, Приморский край, Троицкое. МасштабнаЯ линейка: 0,5 мм." figureDoi="http://doi.org/10.5281/zenodo.14722322" httpUri="https://zenodo.org/record/14722322/files/figure.png" pageId="11" pageNumber="441">Figs 17–18</figureCitation>
). Lateral parts of labrum of male slightly paler. Pronotum with four small and one large central dark spots (
<figureCitation id="1368578D2F1CFFCBDCD7FE9EC5510F7B" box="[509,621,330,351]" captionStart="Figs 9–12" captionStartId="4.[190,228,1943,1962]" captionTargetBox="[280,1330,464,1857]" captionTargetId="figure-119@4.[276,1334,442,1873]" captionTargetPageId="4" captionText="Figs 9–12. Enochrus, habitus in dorsal view. 9–10 — Enochrus simulans, 9 — male, North Korea, 10 — female, North Korea, 11–12 — E. umbratus stat.rev., 11 — lectotype of Philydrus umbratus, 12 — female, paralectotype of Philydrus umbratus, Japan, Chiuzenji. Scale bar: 1 mm. Рис. 9–12. Enochrus, габитус сверху. 9–10 — Enochrus simulans, 9 — самец, СевернаЯ КореЯ, 10 — самка, СевернаЯ КореЯ, 11–12 — E. umbratus stat.rev., 11 — лектотип Philydrus umbratus, 12 — самка, паралектотип Philydrus umbratus, ЯпониЯ, ТюЗенджи. МасштабнаЯ линейка: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.14722318" httpUri="https://zenodo.org/record/14722318/files/figure.png" pageId="12" pageNumber="442">Figs 11–12</figureCitation>
). Lateral parts of pronotum and elytra paler. Punctate rows not darkened (
<figureCitation id="1368578D2F1CFFCBDFCDFEB0C7C70FB0" captionStart="Figs 9–12" captionStartId="4.[190,228,1943,1962]" captionTargetBox="[280,1330,464,1857]" captionTargetId="figure-119@4.[276,1334,442,1873]" captionTargetPageId="4" captionText="Figs 9–12. Enochrus, habitus in dorsal view. 9–10 — Enochrus simulans, 9 — male, North Korea, 10 — female, North Korea, 11–12 — E. umbratus stat.rev., 11 — lectotype of Philydrus umbratus, 12 — female, paralectotype of Philydrus umbratus, Japan, Chiuzenji. Scale bar: 1 mm. Рис. 9–12. Enochrus, габитус сверху. 9–10 — Enochrus simulans, 9 — самец, СевернаЯ КореЯ, 10 — самка, СевернаЯ КореЯ, 11–12 — E. umbratus stat.rev., 11 — лектотип Philydrus umbratus, 12 — самка, паралектотип Philydrus umbratus, ЯпониЯ, ТюЗенджи. МасштабнаЯ линейка: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.14722318" httpUri="https://zenodo.org/record/14722318/files/figure.png" pageId="12" pageNumber="442">Figs 11–12</figureCitation>
). Maxillary palpomeres and antennae yellowish to reddish-yellow or brownish- yellow except for darker club. The last segment is darkened on apex. Ventral surface black to dark brown, posterior margins of abdominal ventrites paler, brown or reddish-brown. Legs reddish-yellow. Posterior margins of femora darker, tarsi brownish-yellow or reddish-yellow.
</paragraph>
<paragraph id="8BEC4B082F1CFFCBDDC9FDCBC6DB0D50" blockId="12.[189,787,303,1364]" pageId="12" pageNumber="442">
<emphasis id="B927971A2F1CFFCBDDC9FDCBC62B0C10" box="[227,279,543,564]" italics="true" pageId="12" pageNumber="442">Head</emphasis>
. Dorsal surface with dense evenly punctation with moderately coarse punctures being of equal size, without microsculpture. Groups of larger and coarser punctures are located around the inner edge of eyes and along anterior margin of head. Clypeus with dense, moderately coarse punctures, interstices without microsculpture. Anterior margin of clypeus with very narrow bead. Eyes big, hardly protruding, emarginated anteriorly, separated by ca. 4.2–4.3× width of one eye (
<figureCitation id="1368578D2F1CFFCBDDEEFD20C6090D2D" box="[196,309,756,777]" captionStart="Figs 17–24" captionStartId="6.[189,227,1861,1880]" captionTargetBox="[200,1405,365,1769]" captionTargetId="figure-6@6.[200,1410,360,1774]" captionTargetPageId="6" captionText="Figs 17–24. Enochrus, head, dorsal view. 17–18 — E. umbratus stat. rev., 17 — lectotype of Philydrus umbratus, 18 — female, paralectotype of Philydrus umbratus, Japan, Chiuzenji. 19-24 — Enochrus simulans, 19 — holotype of Philydrus simulans, 20 — male, Japan, Tochigi-ken, 21 — male, Russia, Primorsky Krai, Lazo, 22 — male, North Korea, Khoankhado, 23 — female, Japan, Shimane-ken, 24 — female, Russia, Primorsky Krai, Troitskoe. Scale bar: 0.5 mm. Рис. 17–24. Enochrus, голова, вид сверху. 17–18 — E. umbratus stat. rev., 17 — лектотип Philydrus umbratus, 18 — самка, паралектотип Philydrus umbratus, ЯпониЯ, ЧиуЗенджи. 19-24 — E. simulans, 19 — голотип Philydrus simulans, 20 — самец, ЯпониЯ, Тотиги-кен, 21 — самец, РоссиЯ, Приморский край, ЛаЗо, 22 — самец, СевернаЯ КореЯ, Хоанхадо, 23 — самка, ЯпониЯ, СиманЭ-кен, 24 — самка, РоссиЯ, Приморский край, Троицкое. МасштабнаЯ линейка: 0,5 мм." figureDoi="http://doi.org/10.5281/zenodo.14722322" httpUri="https://zenodo.org/record/14722322/files/figure.png" pageId="12" pageNumber="442">Figs 17–18</figureCitation>
). Mentum almost flat with setae in anterior part, 1.6 times as wide as long. Antennae with 9 antennomeres, scapus about 1× as long as antennomeres 2–5 combined, club non compact. Maxillary palpomere 4 symmetrical with, ca. 1.4× as long as, palpomere
<quantity id="4CABE6ED2F1CFFCBDC53FC8BC6A20D50" box="[377,414,863,884]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="442" unit="in" value="3.0">3 in</quantity>
length.
</paragraph>
<paragraph id="8BEC4B082F1CFFCBDDC9FCAEC3F50FC0" blockId="12.[189,787,303,1364]" lastBlockId="12.[824,1421,303,991]" pageId="12" pageNumber="442">
<emphasis id="B927971A2F1CFFCBDDC9FCAEC6140DAB" box="[227,296,890,911]" italics="true" pageId="12" pageNumber="442">Thorax</emphasis>
. Elytra and pronotum without microsculpture, have very rare large punctures (with rare trichobothria). Pronotum 3.2–3.4 times as long as elytra. Pronotum ca. 2.0–2.1× as wide as long, strongly narrowed anteriorly. Pronotal punctation similar to that on head. In lateral part of pronotum large punctures forming oval which interrupted from inside at the center level of pronotum. Margins of pronotum narrowly rimmed. Prosternum well developed, protruding anteromedially, with a transverse groove, carina, which can only be detectable posteriorly. Mesoventral elevation high acute and dentiform (
<figureCitation id="1368578D2F1CFFCBDF93FBBEC43B0A5B" box="[697,775,1130,1151]" captionStart="Figs 48–51" captionStartId="9.[190,227,1939,1958]" captionTargetBox="[306,1285,971,1886]" captionTargetId="figure-427@9.[306,1304,971,1890]" captionTargetPageId="9" captionText="Figs 48–51. Enochrus, mesoventral elevation. 48 — E. umbratus stat. rev., male, paralectotype of Philydrus umbratus. 49–51 — E. simulans, 49 — male, Russia, Kunashir Isl., 50 — male, Russia, Primorsky Krai, Lazo, 51 — male (juvenile), Russia, Primorsky Krai, Lazo. Scale bar: 0.5 mm. Рис. 48–51. Enochrus, меЗовентральный отросток. 48 — E. umbratus stat. rev., самец, паралектотип Philydrus umbratus. 49–51 — E. simulans, 49 — самец, РоссиЯ, о-в Кунашир, 50 — самец, РоссиЯ, Приморский край, ЛаЗо, 51 — самец (недоокрашенный), РоссиЯ, Приморский край, ЛаЗо. МасштабнаЯ линейка: 0,5 мм." figureDoi="http://doi.org/10.5281/zenodo.14722328" httpUri="https://zenodo.org/record/14722328/files/figure.png" pageId="12" pageNumber="442">Fig. 48</figureCitation>
). Metaventrite not abruply raised, with dense punctate pubescence median pentagonal area ca. 0.9–1 × as long as wide, center of pentagonal area glabrous. Scutellum expanded, ca. &lt;1 × as long as wide. Elytra with 9 punctate striae, barely noticeable punctate row at outer edge of elytra. Punctate striae near elytral base smoothed out and poorly distinguishable. Short scutellary stria in base of first elytral interval absent. Large punctures (with trichobothria) in elytra situated in the 3
<superScript id="7C26E6402F1CFFCBDF52FAEAC5B50B6E" attach="left" box="[632,649,1342,1354]" fontSize="5" pageId="12" pageNumber="442">rd,</superScript>
5
<superScript id="7C26E6402F1CFFCBDFB0FAEAC5990B6E" attach="none" box="[666,677,1342,1354]" fontSize="5" pageId="12" pageNumber="442">th</superScript>
, 7
<superScript id="7C26E6402F1CFFCBDF95FAEAC5F60B6E" attach="left" box="[703,714,1342,1354]" fontSize="5" pageId="12" pageNumber="442">th</superScript>
and 9
<superScript id="7C26E6402F1CFFCBDE2DFAEAC42E0B6E" attach="none" box="[775,786,1342,1354]" fontSize="5" pageId="12" pageNumber="442">th</superScript>
elytral intervals. Humeral bulge not distinct. Epipleura rather weakly oblique anteriorly, more so posteriorly. Femora with sparse and shallow punctures ventrally, pubescence part of metafemora occuped about 1/7 of femoral length. Tarsi with very short pale setae ventrally. Fifth tarsomere about as long as tarsomeres 3–4 combined. Claw not long, not wide spaced from fifth protarsomere of male (
<figureCitation id="1368578D2F1CFFCBD95FFE1BC3810FC0" box="[1141,1213,463,484]" captionStart="Figs 34–47" captionStartId="8.[190,227,1790,1809]" captionTargetBox="[203,1405,496,1726]" captionTargetId="figure-70@8.[200,1410,488,1732]" captionTargetPageId="8" captionText="Figs 34–47. Enochrus, scutellum, claw and protarsomeres of male. 34–43 — scutellum. 44–47 — claw and protarsomeres of male. 34–41, 45–47 — E. simulans, 42–44 — E. umbratus stat.rev., 34, 45 — holotype of Philydrus simulans, 35 — male, Japan, Higashikami. 36–38 — males, Japan, Fujioka-machi, 39 — female, Japan, Okinoshima, 40 — male, Russia, Primorsky Krai, Lazo, 41 — female, North Korea, Khoankhado. 42, 44 — lectotype of Philydrus umbratus, 43 — female, paralectotype of Philydrus umbratus, Japan, Chiuzenji. 46 — male, paralectotype of Philydrus umbratus, Japan, Hakodate, 47 — male, E. simulans, Russia, Kunashir Isl. Scale bar for figs 34–43: 0.3 mm. Scale bar for figs 44–47: 0.5 mm. Рис. 34–47. Enochrus, щиток, коготок и протарсомеры самца. 34–43 — щиток. 44–47 — коготок и протарсомеры самца. 34–41, 45–47 — E. simulans, 42–44 — E. umbratus stat.rev., 34, 45 — голотип Philydrus simulans, 35 — самец, ЯпониЯ, Хигашиками. 36–38 — самцы, ЯпониЯ, ФудЗиока-мати, 39 — самка, ЯпониЯ, Окиносима, 40 — самец, РоссиЯ, Приморский край, ЛаЗо, 41 — самка, СевернаЯ КореЯ, Хоанхадо. 42, 44 — лектотип Philydrus umbratus, 43 — самка, паралектотип Philydrus umbratus, ЯпониЯ, ЧиуЗенджи. 46 — самец, паралектотип Philydrus umbratus, ЯпониЯ, ХакодатЭ, 47 — самец, E. simulans, РоссиЯ, о-в Кунашир. МасштабнаЯ линейка длЯ рис. 34–43: 0,3 мм. МасштабнаЯ линейка длЯ рис. 44–47: 0,5 мм." figureDoi="http://doi.org/10.5281/zenodo.14722326" httpUri="https://zenodo.org/record/14722326/files/figure.png" pageId="12" pageNumber="442">Fig. 44</figureCitation>
).
</paragraph>
<paragraph id="8BEC4B082F1CFFCBDE74FE3EC2B70C10" blockId="12.[824,1421,303,991]" pageId="12" pageNumber="442">
<emphasis id="B927971A2F1CFFCBDE74FE3EC4850FDB" box="[862,953,490,511]" italics="true" pageId="12" pageNumber="442">Abdomen</emphasis>
. Abdomen with five exposed ventrites, dense uniformpubescence; first ventrite not carinate; ca. as long as second ventrite, fifth ventrite arcuate, not emarginate apically.
</paragraph>
<paragraph id="8BEC4B082F1CFFCBDE74FDEEC27E0C9D" blockId="12.[824,1421,303,991]" pageId="12" pageNumber="442">
<emphasis id="B927971A2F1CFFCBDE74FDEEC4DA0C6B" box="[862,998,570,591]" italics="true" pageId="12" pageNumber="442">Male genitalia</emphasis>
(
<figureCitation id="1368578D2F1CFFCBDED8FDEEC35C0C6B" box="[1010,1120,570,591]" captionStart="Figs 52–56" captionStartId="10.[189,227,1395,1414]" captionTargetBox="[191,785,306,1321]" captionTargetId="figure-684@10.[188,786,302,1327]" captionTargetPageId="10" captionText="Figs 52–56. Enochrus, aedeagus. 52, 53 — E. umbratus stat.rev., 53 — paralectotype of Philydrus umbratus, 53 — apex of aedeagus, lectotype of Philydrus umbratus, 54–56 — E. simulans, 54 — holotype of Philydrus simulans, 55 — E. simulans, aedeagus, Russia, Kunashir Isl. 56 — apex of aedeagus, holotype of Philydrus simulans. Scale bar: 0.5 mm. Рис. 52–56. Enochrus, Эдеагус. 52, 53 — E. umbratus stat.rev., 53 — паралектотип Philydrus umbratus, 53 — вершина Эдеагуса, лектотип Philydrus umbratus, 54–56 — E. simulans, 54 — голотип Philydrus simulans, 55 — E. simulans, aedeagus, РоссиЯ, остров Кунашир. 56 — вершина Эдеагуса, голотип Philydrus simulans. МасштабнаЯ линейка: 0,5 мм." figureDoi="http://doi.org/10.5281/zenodo.14722330" httpUri="https://zenodo.org/record/14722330/files/figure.png" pageId="12" pageNumber="442">Figs 52–53</figureCitation>
). Phallobase symmetrical, about 1.2 times as long as paramere. Parameres strong narrow in apical half; rounded at apex, noticeably joined at apex (
<figureCitation id="1368578D2F1CFFCBD833FDBBC2620CA0" box="[1305,1374,623,644]" captionStart="Figs 52–56" captionStartId="10.[189,227,1395,1414]" captionTargetBox="[191,785,306,1321]" captionTargetId="figure-684@10.[188,786,302,1327]" captionTargetPageId="10" captionText="Figs 52–56. Enochrus, aedeagus. 52, 53 — E. umbratus stat.rev., 53 — paralectotype of Philydrus umbratus, 53 — apex of aedeagus, lectotype of Philydrus umbratus, 54–56 — E. simulans, 54 — holotype of Philydrus simulans, 55 — E. simulans, aedeagus, Russia, Kunashir Isl. 56 — apex of aedeagus, holotype of Philydrus simulans. Scale bar: 0.5 mm. Рис. 52–56. Enochrus, Эдеагус. 52, 53 — E. umbratus stat.rev., 53 — паралектотип Philydrus umbratus, 53 — вершина Эдеагуса, лектотип Philydrus umbratus, 54–56 — E. simulans, 54 — голотип Philydrus simulans, 55 — E. simulans, aedeagus, РоссиЯ, остров Кунашир. 56 — вершина Эдеагуса, голотип Philydrus simulans. МасштабнаЯ линейка: 0,5 мм." figureDoi="http://doi.org/10.5281/zenodo.14722330" httpUri="https://zenodo.org/record/14722330/files/figure.png" pageId="12" pageNumber="442">Fig. 53</figureCitation>
). Penis with narrowly spaced ароdemes, apical part about 1.4 times as long as ароdemes, corona large, situated subapically.
</paragraph>
<paragraph id="8BEC4B082F1CFFCBDE74FD6BC48F0CCB" blockId="12.[824,1421,303,991]" pageId="12" pageNumber="442">
PUBLISHED RECORDS.
<collectingCountry id="F3440B982F1CFFCBD94CFD6BC3A10CF0" box="[1126,1181,703,724]" name="Japan" pageId="12" pageNumber="442">Japan</collectingCountry>
[
<bibRefCitation id="EFC236F92F1CFFCBD980FD6BC20F0CF0" author="Zaitsev F. A." box="[1194,1331,703,724]" pageId="12" pageNumber="442" pagination="283 - 420" refId="ref10325" refString="Zaitsev F. A. 1908. Catalogue des Coleopteres aquaticues des familles des Dryopidae, Georyssidae, Cyathoceridae, Heteroceridae et Hydrophilidae // Horae Societatis entomologicae Rossicae. Vol. 38. P. 283 - 420." type="journal article" year="1908">Zaitsev, 1908</bibRefCitation>
;
<bibRefCitation id="EFC236F92F1CFFCBD815FD6BC4560CCB" author="Hansen M." pageId="12" pageNumber="442" refId="ref9169" refString="Hansen M. 1999. World Catalogue of Insects. Vol. 2: Hydrophiloidea (s. str.) (Coleoptera). Stenstrup: Apollo Books. 416 pp." type="book" year="1999">Hansen, 1999</bibRefCitation>
,
<bibRefCitation id="EFC236F92F1CFFCBDE5EFD0EC4950CCB" author="Hansen M." box="[884,937,730,751]" pageId="12" pageNumber="442" pagination="44 - 68" refId="ref9202" refString="Hansen M. 2004. Family Hydrophilidae // I. Lobl, A. Smetana (eds). Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Histeroidea - Staphylinoidea. Stenstrup: Apollo Books. P. 44 - 68." type="book chapter" year="2004">2004</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="C34918832F1CFFCBDE74FD20C27A0D8D" pageId="12" pageNumber="442" type="distribution">
<paragraph id="8BEC4B082F1CFFCBDE74FD20C23C0D2D" blockId="12.[824,1421,303,991]" box="[862,1280,756,777]" pageId="12" pageNumber="442">
DISTRIBUTION.
<collectingCountry id="F3440B982F1CFFCBD938FD20C3700D2D" box="[1042,1100,756,777]" name="Japan" pageId="12" pageNumber="442">Japan</collectingCountry>
: Honshu (Fig. 60).
</paragraph>
<paragraph id="8BEC4B082F1CFFCBDE74FCDBC27A0D8D" blockId="12.[824,1421,303,991]" pageId="12" pageNumber="442">
NOTE. All records of
<taxonomicName id="4C53308B2F1CFFCBD969FCDBC3810D00" authorityName="Sharp" authorityYear="1884" box="[1091,1213,783,804]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="442" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="B927971A2F1CFFCBD969FCDBC3810D00" box="[1091,1213,783,804]" italics="true" pageId="12" pageNumber="442">E. umbratus</emphasis>
</taxonomicName>
from mainland Eurasia (Russian Far East,
<collectingCountry id="F3440B982F1CFFCBD90CFCFEC39A0D1B" box="[1062,1190,810,831]" name="South Korea" pageId="12" pageNumber="442">South Korea</collectingCountry>
), the Kuril Islands and most localities from Japanese islands (with the exception of Honshu) refer to
<taxonomicName id="4C53308B2F1CFFCBDEF7FC8BC3710D50" baseAuthorityName="Sharp" baseAuthorityYear="1873" box="[989,1101,863,884]" class="Insecta" family="Hydrophilidae" genus="Enochrus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="442" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="B927971A2F1CFFCBDEF7FC8BC3710D50" box="[989,1101,863,884]" italics="true" pageId="12" pageNumber="442">E. simulans</emphasis>
</taxonomicName>
[
<bibRefCitation id="EFC236F92F1CFFCBD973FC8BC3DA0D50" author="Kamiya K." box="[1113,1254,863,884]" pageId="12" pageNumber="442" pagination="113 - 139" refId="ref9366" refString="Kamiya K. 1940. Aquatic beetles of Manchoukuo. // Report of the Limnological Survey of Kwangtung and Manchoukuo Otzu. P. 113 - 139." type="book chapter" year="1940">Kamiya, 1940</bibRefCitation>
,
<bibRefCitation id="EFC236F92F1CFFCBD9DAFC8BC2B50D50" author="Lee S. H. &amp; Cho Y. B. &amp; Lee C. E." box="[1264,1417,863,884]" pageId="12" pageNumber="442" pagination="45 - 60" refId="ref9577" refString="Lee S. H., Cho Y. B., Lee C. E. 1992. The water beetles of Quelpart Island (Coleoptera) // Nature and Life. Vol. 22. No. 2. P. 45 - 60." type="book chapter" year="1992">
Lee
<emphasis id="B927971A2F1CFFCBD830FC8BC2790D50" box="[1306,1349,863,884]" italics="true" pageId="12" pageNumber="442">et al</emphasis>
., 1992
</bibRefCitation>
,
<bibRefCitation id="EFC236F92F1CFFCBDE12FCAEC4A70DAB" author="Lee S. H." box="[824,923,890,911]" pageId="12" pageNumber="442" pagination="14 - 21" refId="ref9547" refString="Lee S. H. 1994. Water beetles of Hyeong San river // Corentomon. Vol. 5. No. 1. P. 14 - 21." type="book chapter" year="1994">Lee, 1994</bibRefCitation>
, Kim
<emphasis id="B927971A2F1CFFCBDEFDFCAEC33F0DAB" box="[983,1027,890,911]" italics="true" pageId="12" pageNumber="442">et al</emphasis>
., 1994,
<bibRefCitation id="EFC236F92F1CFFCBD978FCAEC3E50DAB" author="Hansen M." box="[1106,1241,890,911]" pageId="12" pageNumber="442" refId="ref9169" refString="Hansen M. 1999. World Catalogue of Insects. Vol. 2: Hydrophiloidea (s. str.) (Coleoptera). Stenstrup: Apollo Books. 416 pp." type="book" year="1999">Hansen, 1999</bibRefCitation>
,
<bibRefCitation id="EFC236F92F1CFFCBD9CEFCAEC22A0DAB" author="Hansen M." box="[1252,1302,890,911]" pageId="12" pageNumber="442" pagination="44 - 68" refId="ref9202" refString="Hansen M. 2004. Family Hydrophilidae // I. Lobl, A. Smetana (eds). Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Histeroidea - Staphylinoidea. Stenstrup: Apollo Books. P. 44 - 68." type="book chapter" year="2004">2004</bibRefCitation>
,
<bibRefCitation id="EFC236F92F1CFFCBD80BFCAEC4DE0D8D" author="Kirejtshuk A. G. &amp; Shatrovskiy A. G." pageId="12" pageNumber="442" pagination="696 - 725" refId="ref9452" refString="Kirejtshuk A. G., Shatrovskiy A. G. 2001. [Family Hydrophilidae] // S. J. Tsalolikhin (ed.). Opredelitel' presnovodnykh bespozvonochnykh Rossii i sopredel'nykh territoriy. St. - Petersburg: Nauka. P. 300 - 326, 696 - 725 [In Russian]." type="book chapter" year="2001">Kirejtshuk, Shatrovskiy 2001</bibRefCitation>
,
<bibRefCitation id="EFC236F92F1CFFCBDEC6FC40C3B60D8D" author="Cho Y. &amp; Park S. W." box="[1004,1162,916,937]" pageId="12" pageNumber="442" pagination="83 - 157" refId="ref8875" refString="Cho Y. B, Park S. W. 2010. Coleoptera // M. K. Park (ed.). Checklist of Korean Insects. Seoul: Nature and Ecology. P. 83 - 157." type="book chapter" year="2010">Cho, Park, 2010</bibRefCitation>
,
<bibRefCitation id="EFC236F92F1CFFCBD9BEFC40C2000D8D" author="Ryndevich S. K." box="[1172,1340,916,937]" pageId="12" pageNumber="442" pagination="19 - 35" refId="ref9823" refString="Ryndevich S. K. 2014. [Hydrophiloid fauna (Coleoptera: Hydrophiloidea) of Palaearctic subtaiga zone] // Vestnik Bar- GU. Seriya &quot; Biologicheskiye nauki (obshchaya biologiya). Sel'skokhozyaystvennyye nauki. Agronomiya &quot;. Is. 2. P. 19 - 35 [in Russian]." type="journal article" year="2014">Ryndevich, 2014</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="C34918832F1CFFCBDE74FC7BC4830DFB" pageId="12" pageNumber="442" type="description">
<paragraph id="8BEC4B082F1CFFCBDE74FC7BC4830DFB" blockId="12.[824,1421,303,991]" pageId="12" pageNumber="442">
BIONOMICS. Water beetles.
<typeStatus id="54E8F5AA2F1CFFCBD9A2FC7BC3840DE0" box="[1160,1208,943,964]" pageId="12" pageNumber="442">Type</typeStatus>
specimens were collected in lake.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="20C64D76FF81FD3E5E933319FB73B657" authority="Konow, 1899" authorityName="Konow" authorityYear="1899" box="[1003,1241,1835,1858]" class="Insecta" family="Argidae" genus="Cibdela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="D5B2EAE7FF81FD3E5E933319FB73B657" bold="true" box="[1003,1241,1835,1858]" pageId="3" pageNumber="469">
<emphasis id="D5B2EAE7FF81FD3E5E933319FB95B657" bold="true" box="[1003,1087,1835,1858]" italics="true" pageId="3" pageNumber="469">Cibdela</emphasis>
Konow, 1899
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AFDC657EFF81FD385E263357FD64B0DC" lastPageId="5" lastPageNumber="471" pageId="3" pageNumber="469" type="materials_examined">
<paragraph id="E77936F5FF81FD3E5E263357FAB9B66F" blockId="3.[824,1421,1893,2076]" box="[862,1299,1893,1914]" pageId="3" pageNumber="469">
<typeStatus id="387D8857FF81FD3E5E263357FC24B66F" box="[862,910,1893,1914]" pageId="3" pageNumber="469">Type</typeStatus>
species:
<taxonomicName id="20C64D76FF81FD3E5E9D3357FAA5B66F" authority="Klug, 1834" authorityName="Klug" authorityYear="1834" box="[997,1295,1893,1914]" class="Insecta" family="Argidae" genus="Hylotoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="janthina">
<emphasis id="D5B2EAE7FF81FD3E5E9D3357FB30B66F" box="[997,1178,1893,1914]" italics="true" pageId="3" pageNumber="469">Hylotoma janthina</emphasis>
Klug, 1834
</taxonomicName>
.
</paragraph>
<paragraph id="E77936F5FF81FD3E5E2633B2FB44B909" blockId="3.[824,1421,1893,2076]" pageId="3" pageNumber="469">
Body size 8.0–10.0 mm. Head not enlarged behind eyes in dorsal view. Antennae of males simple. Fore wings with 4 cells Rs; cell
<collectionCode id="81D7AE30FF81FD3E5EA83384FC4AB6DE" box="[976,992,1974,1995]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="469">R</collectionCode>
closed; vein Sc present; vein 2r–m present; vein B of fore wing joins vein
<collectionCode id="81D7AE30FF81FD3E592733E3FBC5B6F3" box="[1119,1135,2001,2022]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="469">R</collectionCode>
well before (longer than vein Sc) junction point of vein M; cell 2A closed. All tibiae without preapical spurs. Hind tibiae and spurs simple.
</paragraph>
<caption id="B3B9667DFF86FD395DC53324FB95B90C" pageId="4" pageNumber="470" startId="4.[189,227,1814,1833]" targetBox="[200,1409,368,1714]" targetPageId="4" targetType="figure">
<paragraph id="E77936F5FF86FD395DC53324FDFFB686" blockId="4.[189,1422,1814,2073]" pageId="4" pageNumber="470">
<emphasis id="D5B2EAE7FF86FD395DC53324FEB3B63C" bold="true" box="[189,281,1814,1833]" pageId="4" pageNumber="470">Figs 2–15.</emphasis>
<taxonomicName id="20C64D76FF86FD395C653324FECEB63C" box="[285,356,1814,1833]" class="Insecta" family="Argidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="family">Argidae</taxonomicName>
, details of morphology: 2, 10, 12 —
<taxonomicName id="20C64D76FF86FD395FDA3324FC38B63C" authority="(Linne, 1758)" baseAuthorityName="Linne" baseAuthorityYear="1758" box="[674,914,1814,1833]" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="ustulata">
<emphasis id="D5B2EAE7FF86FD395FDA3324FCBFB63C" box="[674,789,1814,1833]" italics="true" pageId="4" pageNumber="470">Arge ustulata</emphasis>
(Linné, 1758)
</taxonomicName>
; 3, 11 —
<taxonomicName id="20C64D76FF86FD395E9F3324FA82B63C" authority="Takeuchi, 1939" authorityName="Takeuchi" authorityYear="1939" box="[999,1320,1814,1833]" class="Insecta" family="Argidae" genus="Aproceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="leucopoda">
<emphasis id="D5B2EAE7FF86FD395E9F3324FB37B63C" box="[999,1181,1814,1833]" italics="true" pageId="4" pageNumber="470">Aproceros leucopoda</emphasis>
Takeuchi, 1939
</taxonomicName>
; 4, 8, 14 —
<taxonomicName id="20C64D76FF86FD395DC53302FE41B656" authority="Konow, 1902" authorityName="Konow" authorityYear="1902" box="[189,491,1840,1859]" class="Insecta" family="Argidae" genus="Kokujewia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="ectrapela">
<emphasis id="D5B2EAE7FF86FD395DC53302FEC5B656" box="[189,367,1840,1859]" italics="true" pageId="4" pageNumber="470">Kokujewia ectrapela</emphasis>
Konow, 1902
</taxonomicName>
; 5 —
<taxonomicName id="20C64D76FF86FD395F593302FC90B656" authority="(Klug, 1834)" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[545,826,1840,1859]" class="Insecta" family="Argidae" genus="Spinarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="D5B2EAE7FF86FD395F593302FD68B656" box="[545,706,1840,1859]" italics="true" pageId="4" pageNumber="470">Spinarge metallica</emphasis>
(Klug, 1834)
</taxonomicName>
; 6 —
<taxonomicName id="20C64D76FF86FD395E083302FB1FB656" authority="(Klug, 1814)" baseAuthorityName="Klug" baseAuthorityYear="1814" box="[880,1205,1840,1859]" class="Insecta" family="Argidae" genus="Aprosthema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="D5B2EAE7FF86FD395E083302FB94B656" box="[880,1086,1840,1859]" italics="true" pageId="4" pageNumber="470">Aprosthema melanurum</emphasis>
(Klug, 1814)
</taxonomicName>
; 7, 9 —
<taxonomicName id="20C64D76FF86FD39587A3302FE0AB64B" authority="Gussakovskij, 1935" authorityName="Gussakovskij" authorityYear="1935" class="Insecta" family="Argidae" genus="Pseudarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="rubicunda">
<emphasis id="D5B2EAE7FF86FD39587A3302FF5BB64B" italics="true" pageId="4" pageNumber="470">Pseudarge rubicunda</emphasis>
<bibRefCitation id="83574B04FF86FD395D8E3379FE0AB64B" author="Gussakovskij V. V." box="[246,416,1867,1886]" pageId="4" pageNumber="470" refId="ref8958" refString="Gussakovskij V. V. 1935. [Chalastogastra (pt. 1)] // Fauna SSSR. Nov. ser. Nasekomye Pereponchatokrylye. Vol. 2. No. 2. Moscow-Leningrad: AN SSSR Publ. 453 p. [In Russian, with French summary]" type="book" year="1935">Gussakovskij, 1935</bibRefCitation>
</taxonomicName>
; 13 —
<taxonomicName id="20C64D76FF86FD395CA43379FC80B64B" authority="(Gmelin, 1790)" baseAuthorityName="Gmelin" baseAuthorityYear="1790" box="[476,810,1867,1886]" class="Insecta" family="Argidae" genus="Sterictiphora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="geminata">
<emphasis id="D5B2EAE7FF86FD395CA43379FD0AB64B" box="[476,672,1867,1886]" italics="true" pageId="4" pageNumber="470">Sterictiphora geminata</emphasis>
(Gmelin, 1790)
</taxonomicName>
; 15 —
<taxonomicName id="20C64D76FF86FD395E1E3379FB21B64B" authority="Gussakovskij, 1935" authorityName="Gussakovskij" authorityYear="1935" box="[870,1163,1867,1886]" class="Insecta" genus="Asiarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="regeli">
<emphasis id="D5B2EAE7FF86FD395E1E3379FC77B64B" box="[870,989,1867,1886]" italics="true" pageId="4" pageNumber="470">Asiarge regeli</emphasis>
<bibRefCitation id="83574B04FF86FD395E9A3379FB21B64B" author="Gussakovskij V. V." box="[994,1163,1867,1886]" pageId="4" pageNumber="470" refId="ref8958" refString="Gussakovskij V. V. 1935. [Chalastogastra (pt. 1)] // Fauna SSSR. Nov. ser. Nasekomye Pereponchatokrylye. Vol. 2. No. 2. Moscow-Leningrad: AN SSSR Publ. 453 p. [In Russian, with French summary]" type="book" year="1935">Gussakovskij, 1935</bibRefCitation>
</taxonomicName>
. 2–4 — veins R and Rs of fore wing, dorsally; 5–6 — male antenna, dorsally; 7–8 — hind tibia, ventrally; 9–11 — cells 1A and 2A of fore wing, dorsally; 12–13 — vein R+M of fore wing, dorsally; 14–15 — head, laterally.
</paragraph>
<paragraph id="E77936F5FF86FD395DC533A9FB95B90C" blockId="4.[189,1422,1814,2073]" pageId="4" pageNumber="470">
<emphasis id="D5B2EAE7FF86FD395DC533A9FEB4B6BB" bold="true" box="[189,286,1947,1966]" pageId="4" pageNumber="470">Рис. 2–15.</emphasis>
<taxonomicName id="20C64D76FF86FD395C5D33A9FEC6B6BB" box="[293,364,1947,1966]" class="Insecta" family="Argidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="family">Argidae</taxonomicName>
, детали морфологии: 2, 10, 12 —
<taxonomicName id="20C64D76FF86FD395FCC33A9FC04B6BB" authority="(Linne, 1758)" baseAuthorityName="Linne" baseAuthorityYear="1758" box="[692,942,1947,1966]" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="ustulata">
<emphasis id="D5B2EAE7FF86FD395FCC33A9FC81B6BB" box="[692,811,1947,1966]" italics="true" pageId="4" pageNumber="470">Arge ustulata</emphasis>
(Linné, 1758)
</taxonomicName>
; 3, 11 —
<taxonomicName id="20C64D76FF86FD39597633A9FAF2B6BB" authority="Takeuchi, 1939" authorityName="Takeuchi" authorityYear="1939" box="[1038,1368,1947,1966]" class="Insecta" family="Argidae" genus="Aproceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="leucopoda">
<emphasis id="D5B2EAE7FF86FD39597633A9FB6DB6BB" box="[1038,1223,1947,1966]" italics="true" pageId="4" pageNumber="470">Aproceros leucopoda</emphasis>
Takeuchi, 1939
</taxonomicName>
; 4, 8, 14 —
<taxonomicName id="20C64D76FF86FD395D8C3384FD8BB6DC" authority="Konow, 1902" authorityName="Konow" authorityYear="1902" box="[244,545,1974,1993]" class="Insecta" family="Argidae" genus="Kokujewia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="ectrapela">
<emphasis id="D5B2EAE7FF86FD395D8C3384FE0CB6DC" box="[244,422,1974,1993]" italics="true" pageId="4" pageNumber="470">Kokujewia ectrapela</emphasis>
Konow, 1902
</taxonomicName>
; 5 —
<taxonomicName id="20C64D76FF86FD395F2E3384FCC7B6DC" authority="(Klug, 1834)" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[598,877,1974,1993]" class="Insecta" family="Argidae" genus="Spinarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="metallica">Spinarge metallica (Klug, 1834)</taxonomicName>
; 6 —
<taxonomicName id="20C64D76FF86FD395EDB3384FB4DB6DC" authority="(Klug, 1814)" baseAuthorityName="Klug" baseAuthorityYear="1814" box="[931,1255,1974,1993]" class="Insecta" family="Argidae" genus="Aprosthema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="D5B2EAE7FF86FD395EDB3384FBDAB6DC" box="[931,1136,1974,1993]" italics="true" pageId="4" pageNumber="470">Aprosthema melanurum</emphasis>
(Klug, 1814)
</taxonomicName>
; 7, 9 —
<taxonomicName id="20C64D76FF86FD39584B3384FE6CB6F6" authority="Gussakovskij, 1935" authorityName="Gussakovskij" authorityYear="1935" class="Insecta" family="Argidae" genus="Pseudarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="rubicunda">
<emphasis id="D5B2EAE7FF86FD39584B3384FEBFB6F6" italics="true" pageId="4" pageNumber="470">Pseudarge rubicunda</emphasis>
<bibRefCitation id="83574B04FF86FD395C6333E2FE6CB6F6" author="Gussakovskij V. V." box="[283,454,2000,2019]" pageId="4" pageNumber="470" refId="ref8958" refString="Gussakovskij V. V. 1935. [Chalastogastra (pt. 1)] // Fauna SSSR. Nov. ser. Nasekomye Pereponchatokrylye. Vol. 2. No. 2. Moscow-Leningrad: AN SSSR Publ. 453 p. [In Russian, with French summary]" type="book" year="1935">Gussakovskij, 1935</bibRefCitation>
</taxonomicName>
; 13 —
<taxonomicName id="20C64D76FF86FD395F7F33E2FCF0B6F6" authority="(Gmelin, 1790)" baseAuthorityName="Gmelin" baseAuthorityYear="1790" box="[519,858,2000,2019]" class="Insecta" family="Argidae" genus="Sterictiphora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="geminata">
<emphasis id="D5B2EAE7FF86FD395F7F33E2FD67B6F6" box="[519,717,2000,2019]" italics="true" pageId="4" pageNumber="470">Sterictiphora geminata</emphasis>
(Gmelin, 1790)
</taxonomicName>
; 15 —
<taxonomicName id="20C64D76FF86FD395EE333E2FB6FB6F6" authority="Gussakovskij, 1935" authorityName="Gussakovskij" authorityYear="1935" box="[923,1221,2000,2019]" class="Insecta" genus="Asiarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="470" phylum="Arthropoda" rank="species" species="regeli">
<emphasis id="D5B2EAE7FF86FD395EE333E2FBB9B6F6" box="[923,1043,2000,2019]" italics="true" pageId="4" pageNumber="470">Asiarge regeli</emphasis>
<bibRefCitation id="83574B04FF86FD39596233E2FB6FB6F6" author="Gussakovskij V. V." box="[1050,1221,2000,2019]" pageId="4" pageNumber="470" refId="ref8958" refString="Gussakovskij V. V. 1935. [Chalastogastra (pt. 1)] // Fauna SSSR. Nov. ser. Nasekomye Pereponchatokrylye. Vol. 2. No. 2. Moscow-Leningrad: AN SSSR Publ. 453 p. [In Russian, with French summary]" type="book" year="1935">Gussakovskij, 1935</bibRefCitation>
</taxonomicName>
. 2–4 — жилки R и Rs переднего крыла, дорсально; 5–6 — антенна самца, дорсально; 7–8 — Задние голени, вентрально; 9–11 — Ячейки 1А и 2А переднего крыла, дорсально; 12–13 — жилка R+M переднего крыла, дорсально; 14–15 — голова, латерально.
</paragraph>
</caption>
<paragraph id="E77936F5FF87FD385D9B351DFD64B0DC" blockId="5.[189,787,303,457]" pageId="5" pageNumber="471">
The genus is distributed in the Palaearctic (East), Oriental and Australasian regions.Number of species:World — 14; Palaearctic — 8.
<taxonomicName id="20C64D76FF87FD385D973556FE7BB06C" authority="(Klug, 1834)" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[239,465,356,377]" class="Insecta" family="Argidae" genus="Hylotoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="species" species="janthina">
<emphasis id="D5B2EAE7FF87FD385D973556FEFCB06C" box="[239,342,356,377]" italics="true" pageId="5" pageNumber="471">C. janthina</emphasis>
(Klug, 1834)
</taxonomicName>
has been introduced to
<collectingCountry id="9FD17665FF87FD385FDE3556FCB9B06C" box="[678,787,356,377]" name="Réunion" pageId="5" pageNumber="471">La Réunion</collectingCountry>
(
<collectingCountry id="9FD17665FF87FD385DBB354DFEAFB081" box="[195,261,383,404]" name="France" pageId="5" pageNumber="471">France</collectingCountry>
, Indian Ocean) in an attempt at biological control of an invasive species of
<taxonomicName id="20C64D76FF87FD385C0935A8FE01B0BA" box="[369,427,410,431]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="471" phylum="Tracheophyta" rank="genus">
<emphasis id="D5B2EAE7FF87FD385C0935A8FE01B0BA" box="[369,427,410,431]" italics="true" pageId="5" pageNumber="471">Rubus</emphasis>
</taxonomicName>
(e.g.,
<bibRefCitation id="83574B04FF87FD385C9D35A8FD16B0BA" author="Bourgeois T. &amp; Baret S. &amp; Chenon R. D." box="[485,700,410,431]" pageId="5" pageNumber="471" pagination="153 - 160" refId="ref8177" refString="Bourgeois T., Baret S., Chenon R. D. 2011. Biological Control of Rubus alceifolius (Rosaceae) in La Reunion Island (Indian Ocean): From Investigations on the Plant to the Release of the Biological Control Agent Cibdela janthina (Argidae) // XIII International Symposium on Biological Control of Weeds. P. 153 - 160." type="book chapter" year="2011">
Bourgeois
<emphasis id="D5B2EAE7FF87FD385F3135A8FDD8B0BA" box="[585,626,410,431]" italics="true" pageId="5" pageNumber="471">et al</emphasis>
. [2011]
</bibRefCitation>
;
<bibRefCitation id="83574B04FF87FD385FBE35A8FE9AB0DC" author="Dumont Y. &amp; Mathieu A. &amp; Quilici S." pageId="5" pageNumber="471" pagination="369 - 371" refId="ref8686" refString="Dumont Y., Mathieu A., Quilici S. 2013. Mathematical Modelling of the biocontrol of Rubus alceifolius, an invasive plant in Reunion Island // Finnish Society of Forest Science, Vantaa, Finland Finnish Forest Research Institute, Vantaa, Finland Department of Forest Sciences, University of Helsinki. P. 369 - 371." type="book chapter" year="2013">
Dumont
<emphasis id="D5B2EAE7FF87FD385DC53586FF4FB0DC" box="[189,229,436,457]" italics="true" pageId="5" pageNumber="471">et al</emphasis>
. [2013]
</bibRefCitation>
;
<bibRefCitation id="83574B04FF87FD385C403586FE54B0DC" author="Mathieu A. &amp; Dumont Y. &amp; Chiroleu F. &amp; Duyck P. - F. &amp; Flores O. &amp; Lebreton G. &amp; Reynaud B. &amp; Quilici S." box="[312,510,436,457]" pageId="5" pageNumber="471" pagination="461 - 471" refId="ref10549" refString="Mathieu A., Dumont Y., Chiroleu F., Duyck P. - F., Flores O., Lebreton G., Reynaud B., Quilici S. 2014. Predicting the altitudinal distribution of an introduced phytophagous insect against an invasive alien plant from laboratory controlled experiments: case of Cibdela janthina (Hymenoptera: Argidae) and Rubus alceifolius (Rosaceae) in La Reunion // BioControl. Vol. 59. No. 4. P. 461 - 471." type="book chapter" year="2014">
Mathieu
<emphasis id="D5B2EAE7FF87FD385CF33586FE18B0DC" box="[395,434,436,457]" italics="true" pageId="5" pageNumber="471">et al</emphasis>
. [2014]
</bibRefCitation>
;
<bibRefCitation id="83574B04FF87FD385F7E3586FD6EB0DC" author="Florens B. V. &amp; Bissessur P. &amp; Bunsy Y. &amp; Ramdonee D." box="[518,708,436,457]" pageId="5" pageNumber="471" pagination="271 - 274" refId="ref8798" refString="Florens B. V., Bissessur P., Bunsy Y., Ramdonee D. 2017. Cibdela janthina (Kiug 1834) (Hymenoptera: Tenthredinidae), Reunion's biocontrol agent of Rubus alceifolius Poir., recorded on Mauritius // African Entomology. Vol. 25. No. 1. P. 271 - 274." type="journal article" year="2017">
Florens
<emphasis id="D5B2EAE7FF87FD385F283586FDD2B0DC" box="[592,632,436,457]" italics="true" pageId="5" pageNumber="471">et al</emphasis>
. [2017]
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="6B087224DECFC981C6B944FF24BCA1F3">Illustrated key to the Palaearctic genera of the sawfly family Argidae (Hymenoptera: Symphyta), with taxonomic notes</mods:title>
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<mods:namePart id="388FA6DFFD67D0FD2D36DF1AE2813643">Basov, S. A.</mods:namePart>
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<mods:title id="62B74F8D41C29EDD60873D73AC0DDADE">Russian Entomological Journal</mods:title>
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<mods:date id="BE19B7134E3C3D3CBB3BAC1662193780">2024</mods:date>
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<treatment id="6F6F87E3FF81FD3E5C0F333FFAD3B23F" ID-DOI="http://doi.org/10.5281/zenodo.14722338" ID-Zenodo-Dep="14722338" LSID="urn:lsid:plazi:treatment:6F6F87E3FF81FD3E5C0F333FFAD3B23F" httpUri="http://treatment.plazi.org/id/6F6F87E3FF81FD3E5C0F333FFAD3B23F" lastPageNumber="469" pageId="3" pageNumber="469">
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<paragraph id="E77936F5FF81FD3E5C0F333FFDFDB631" blockId="3.[375,599,1805,1857]" box="[375,599,1805,1828]" pageId="3" pageNumber="469">
<heading id="BC318199FF81FD3E5C0F333FFDFDB631" box="[375,599,1805,1828]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="469" reason="8">
<taxonomicName id="20C64D76FF81FD3E5C0F333FFDFDB631" ID-CoL="32L7" authority="Schrank, 1802" authorityName="Schrank" authorityYear="1802" box="[375,599,1805,1828]" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="D5B2EAE7FF81FD3E5C0F333FFDFDB631" bold="true" box="[375,599,1805,1828]" pageId="3" pageNumber="469">
<emphasis id="D5B2EAE7FF81FD3E5C0F333FFE06B631" bold="true" box="[375,428,1805,1828]" italics="true" pageId="3" pageNumber="469">Arge</emphasis>
Schrank, 1802
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<paragraph id="E77936F5FF81FD3E5CF63318FDEBB654" blockId="3.[375,599,1805,1857]" box="[398,577,1834,1857]" pageId="3" pageNumber="469">
<figureCitation id="7FFD2A70FF81FD3E5CF63318FD94B654" box="[398,574,1834,1857]" captionStart-0="Fig" captionStart-1="Figs 2–15" captionStartId-0="2.[189,220,1838,1857]" captionStartId-1="4.[189,227,1814,1833]" captionTargetBox-0="[241,1387,474,1750]" captionTargetBox-1="[200,1409,368,1714]" captionTargetId-0="figure-86@2.[201,1411,444,1795]" captionTargetId-1="figure-4@4.[200,1410,360,1714]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="Fig. 1. Venation of wings of Arge ustulata (Linné, 1758). Cells (in red): R — radial; 1Rs, 2Rs, 3Rs, 4Rs — 1st, 2nd, 3rd, 4th subradial; Sc– subcostal; B — basal; 1M, 2M, 3M — 1st, 2nd, 3rd medial; 1Cu, 2Cu — 1st, 2nd cubital; 1A — apical anal; 2A+3A — basal anal. Veins (in black): C — costal; Sc– subcostal; R — radial; Rs — subradial; M — medial; B — basal; Cu — cubital; 1A, 2A, 3A, 4A — 1st, 2nd, 3rd, 4th anal; 1r–m, 2r–m, 3r–m — 1st, 2nd, 3rd radio–medial; 1m–cu, 2m–cu — 1st, 2nd medial– cubital; cu–a — cubital–anal. Рис. 1. Жилкование крыльев Arge ustulata (Linné, 1758). Ячейки (красным цветом): R — радиальнаЯ; 1Rs, 2Rs, 3Rs, 4Rs — 1–Я, 2–Я, 3–Я, 4–Я субрадиальные; Sc — субкостальнаЯ; B — баЗальнаЯ; 1M, 2M, 3M — 1–Я, 2–Я, 3–Я медиальные; 1Cu, 2Cu — 1–Я, 2–Я кубитальные; 1A — апикальнаЯ анальнаЯ; 2A+3A — баЗальнаЯ анальнаЯ. Жилки (черным цветом): C — костальнаЯ; Sc — субкостальнаЯ; R — радиальнаЯ; Rs — субрадиальнаЯ; M — медиальнаЯ; B — баЗальнаЯ; Cu — кубитальнаЯ; 1A, 2A, 3A, 4A — 1–Я, 2–Я, 3–Я, 4–Я анальные; 1r–m, 2r–m, 3r–m — 1–Я, 2–Я, 3–Я радиально–медиальные; 1m–cu, 2m–cu — 1–Я, 2–Я медиально–кубитальные; cu–a — кубитально–анальнаЯ." captionText-1="Figs 2–15. Argidae, details of morphology: 2, 10, 12 — Arge ustulata (Linné, 1758); 3, 11 — Aproceros leucopoda Takeuchi, 1939; 4, 8, 14 — Kokujewia ectrapela Konow, 1902; 5 — Spinarge metallica (Klug, 1834); 6 — Aprosthema melanurum (Klug, 1814); 7, 9 — Pseudarge rubicunda Gussakovskij, 1935; 13 — Sterictiphora geminata (Gmelin, 1790); 15 — Asiarge regeli Gussakovskij, 1935. 2–4 — veins R and Rs of fore wing, dorsally; 5–6 — male antenna, dorsally; 7–8 — hind tibia, ventrally; 9–11 — cells 1A and 2A of fore wing, dorsally; 12–13 — vein R+M of fore wing, dorsally; 14–15 — head, laterally. Рис. 2–15. Argidae, детали морфологии: 2, 10, 12 — Arge ustulata (Linné, 1758); 3, 11 — Aproceros leucopoda Takeuchi, 1939; 4, 8, 14 — Kokujewia ectrapela Konow, 1902; 5 — Spinarge metallica (Klug, 1834); 6 — Aprosthema melanurum (Klug, 1814); 7, 9 — Pseudarge rubicunda Gussakovskij, 1935; 13 — Sterictiphora geminata (Gmelin, 1790); 15 — Asiarge regeli Gussakovskij, 1935. 2–4 — жилки R и Rs переднего крыла, дорсально; 5–6 — антенна самца, дорсально; 7–8 — Задние голени, вентрально; 9–11 — Ячейки 1А и 2А переднего крыла, дорсально; 12–13 — жилка R+M переднего крыла, дорсально; 14–15 — голова, латерально." pageId="3" pageNumber="469">Figs 1–2, 10, 12</figureCitation>
.
</paragraph>
<paragraph id="E77936F5FF81FD3E5D9B3357FD3AB66F" blockId="3.[189,787,1893,2076]" box="[227,656,1893,1914]" pageId="3" pageNumber="469">
<typeStatus id="387D8857FF81FD3E5D9B3357FEB9B66F" box="[227,275,1893,1914]" pageId="3" pageNumber="469">Type</typeStatus>
species:
<taxonomicName id="20C64D76FF81FD3E5C123357FD26B66F" authority="Linne, 1767" authorityName="Linne" authorityYear="1767" box="[362,652,1893,1914]" class="Insecta" family="Tenthredinidae" genus="Tenthredo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="enodis">
<emphasis id="D5B2EAE7FF81FD3E5C123357FDA5B66F" box="[362,527,1893,1914]" italics="true" pageId="3" pageNumber="469">Tenthredo enodis</emphasis>
Linné, 1767
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.
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<paragraph id="E77936F5FF81FD3E5D9B33B2FD4FB909" blockId="3.[189,787,1893,2076]" pageId="3" pageNumber="469">Body size 6.5–12.0 mm. Right mandible without basal tooth. Antennae of males simple. Fore wings with 4 cells Rs; cell R closed; vein Sc present; vein 2r–m present; veins B and M join up at same point on vein R; cell 2A closed. Cell A of hind wing closed. Claws simple. Middle and hind tibiae with preapical spurs. The colour is varied, often with a metallic sheen.</paragraph>
<paragraph id="E77936F5FF81FD3E5E26351DFAD3B23F" blockId="3.[824,1422,303,810]" pageId="3" pageNumber="469">
The genus is widely distributed in the Palaearctic, Nearctic, Oriental, Neotropical,Afrotropical and Australasian regions. Number of species: World — 396; Palaearctic — 187. Since the publication of the world catalogue of sawflies [
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]. Another 11 species,
<taxonomicName id="20C64D76FF81FD3E5EDD3641FB28B39D" authority="Wei, 2010" authorityName="Wei" authorityYear="2010" box="[933,1154,627,648]" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="laticincta">
<emphasis id="D5B2EAE7FF81FD3E5EDD3641FBB3B39D" box="[933,1049,627,648]" italics="true" pageId="3" pageNumber="469">A. laticincta</emphasis>
<bibRefCitation id="83574B04FF81FD3E59663641FB28B39D" author="Wei M. &amp; Niu G." box="[1054,1154,627,648]" pageId="3" pageNumber="469" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei, 2010</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="20C64D76FF81FD3E59F43641FA23B39D" authority="Wei, 2010" authorityName="Wei" authorityYear="2010" box="[1164,1417,627,648]" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="renemalaisei">
<emphasis id="D5B2EAE7FF81FD3E59F43641FAB5B39D" box="[1164,1311,627,648]" italics="true" pageId="3" pageNumber="469">A. renemalaisei</emphasis>
<bibRefCitation id="83574B04FF81FD3E585D3641FA23B39D" author="Wei M. &amp; Niu G." box="[1317,1417,627,648]" pageId="3" pageNumber="469" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei, 2010</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="20C64D76FF81FD3E5E4036BCFC43B3B6" authority="Wei, 2010" authorityName="Wei" authorityYear="2010" box="[824,1001,654,675]" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="qini">
<emphasis id="D5B2EAE7FF81FD3E5E4036BCFCD0B3B6" box="[824,890,654,675]" italics="true" pageId="3" pageNumber="469">A. qini</emphasis>
<bibRefCitation id="83574B04FF81FD3E5EFB36BCFC43B3B6" author="Wei M. &amp; Niu G." box="[899,1001,654,675]" pageId="3" pageNumber="469" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei, 2010</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="20C64D76FF81FD3E5E8E36BCFB47B3B6" authority="Wei, 2010" authorityName="Wei" authorityYear="2010" box="[1014,1261,654,675]" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="nigroharpa">
<emphasis id="D5B2EAE7FF81FD3E5E8E36BCFBD7B3B6" box="[1014,1149,654,675]" italics="true" pageId="3" pageNumber="469">A. nigroharpa</emphasis>
<bibRefCitation id="83574B04FF81FD3E59FE36BCFB47B3B6" author="Wei M. &amp; Niu G." box="[1158,1261,654,675]" pageId="3" pageNumber="469" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei, 2010</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="20C64D76FF81FD3E598236BCFCC0B3AB" authority="Wei, 2010" authorityName="Wei" authorityYear="2010" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="shenae">
<emphasis id="D5B2EAE7FF81FD3E598236BCFAF2B3B6" box="[1274,1368,654,675]" italics="true" pageId="3" pageNumber="469">A. shenae</emphasis>
<bibRefCitation id="83574B04FF81FD3E581936BCFCC0B3AB" author="Wei M. &amp; Niu G." pageId="3" pageNumber="469" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei, 2010</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="20C64D76FF81FD3E5E0E369BFB89B3AB" authority="Wei, 2010" authorityName="Wei" authorityYear="2010" box="[886,1059,681,702]" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="xini">
<emphasis id="D5B2EAE7FF81FD3E5E0E369BFC1FB3AB" box="[886,949,681,702]" italics="true" pageId="3" pageNumber="469">A. xini</emphasis>
<bibRefCitation id="83574B04FF81FD3E5EC5369BFB89B3AB" author="Wei M. &amp; Niu G." box="[957,1059,681,702]" pageId="3" pageNumber="469" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei, 2010</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="20C64D76FF81FD3E5957369BFA9AB3AB" authority="Wei, 2010" authorityName="Wei" authorityYear="2010" box="[1071,1328,681,702]" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="longiflagella">
<emphasis id="D5B2EAE7FF81FD3E5957369BFB69B3AB" box="[1071,1219,681,702]" italics="true" pageId="3" pageNumber="469">A. longiflagella</emphasis>
<bibRefCitation id="83574B04FF81FD3E59B2369BFA9AB3AB" author="Wei M. &amp; Niu G." box="[1226,1328,681,702]" pageId="3" pageNumber="469" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei, 2010</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="20C64D76FF81FD3E5844369BFC35B3CC" authority="Wei, 2010" authorityName="Wei" authorityYear="2010" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="fanae">
<emphasis id="D5B2EAE7FF81FD3E5844369BFA26B3AB" box="[1340,1420,681,702]" italics="true" pageId="3" pageNumber="469">A. fanae</emphasis>
<bibRefCitation id="83574B04FF81FD3E5E4036F6FC35B3CC" author="Wei M. &amp; Niu G." box="[824,927,708,729]" pageId="3" pageNumber="469" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei, 2010</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="20C64D76FF81FD3E5ED536F6FB18B3CC" authority="Wei, 2010" authorityName="Wei" authorityYear="2010" box="[941,1202,708,729]" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="striatitergita">
<emphasis id="D5B2EAE7FF81FD3E5ED536F6FBEBB3CC" box="[941,1089,708,729]" italics="true" pageId="3" pageNumber="469">A. striatitergita</emphasis>
<bibRefCitation id="83574B04FF81FD3E593336F6FB18B3CC" author="Wei M. &amp; Niu G." box="[1099,1202,708,729]" pageId="3" pageNumber="469" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei, 2010</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="20C64D76FF81FD3E59B836F6FCC0B3E1" authority="Wei, 2010" authorityName="Wei" authorityYear="2010" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="rugosipleura">
<emphasis id="D5B2EAE7FF81FD3E59B836F6FAFDB3CC" box="[1216,1367,708,729]" italics="true" pageId="3" pageNumber="469">A. rugosipleura</emphasis>
<bibRefCitation id="83574B04FF81FD3E581936F6FCC0B3E1" author="Wei M. &amp; Niu G." pageId="3" pageNumber="469" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei, 2010</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="20C64D76FF81FD3E5EE736EDFB61B3E1" authority="Wei et Niu, 2010" authorityName="Wei et Niu" authorityYear="2010" box="[927,1227,735,756]" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="stenolutea">
<emphasis id="D5B2EAE7FF81FD3E5EE736EDFBB6B3E1" box="[927,1052,735,756]" italics="true" pageId="3" pageNumber="469">A. stenolutea</emphasis>
<bibRefCitation id="83574B04FF81FD3E595B36EDFB61B3E1" author="Wei M. &amp; Niu G." box="[1059,1227,735,756]" pageId="3" pageNumber="469" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei et Niu, 2010</bibRefCitation>
</taxonomicName>
, have been reported as new [
<bibRefCitation id="83574B04FF81FD3E5EF036C8FB8CB21A" author="Wu X. - Y. &amp; Xin X. &amp; Zhang X. - Z." box="[904,1062,762,783]" pageId="3" pageNumber="469" pagination="15 - 19" refId="ref12137" refString="Wu X. - Y., Xin X., Zhang X. - Z. 2010. [The Investigation and Taxonomical Research of the Sawflies Species from Gansu Province. III [recte: IV]. A list of the genera and species (Hymenoptera: Argidae).] // Journal of Gansu Forestry Science and Technology. Vol. 35. No. 1. P. 15 - 19 [in Chinese]." type="journal article" year="2010">
Wu
<emphasis id="D5B2EAE7FF81FD3E5ECC36C8FC4AB21A" box="[948,992,762,783]" italics="true" pageId="3" pageNumber="469">et al</emphasis>
., 2010
</bibRefCitation>
] but their descriptions have not been published to date, and they should be treated as nomina nuda.
</paragraph>
</subSubSection>
</treatment>
</document>

91
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<subSubSection id="AFDC657EFF81FD3E5C2030F8FDDCB5F4" box="[344,630,1226,1249]" pageId="3" pageNumber="469" type="nomenclature">
<paragraph id="E77936F5FF81FD3E5C2030F8FDDCB5F4" blockId="3.[344,630,1226,1279]" box="[344,630,1226,1249]" pageId="3" pageNumber="469">
<heading id="BC318199FF81FD3E5C2030F8FDDCB5F4" box="[344,630,1226,1249]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="469" reason="8">
<taxonomicName id="20C64D76FF81FD3E5C2030F8FDDCB5F4" authority="Konow, 1899" authorityName="Konow" authorityYear="1899" box="[344,630,1226,1249]" class="Insecta" family="Argidae" genus="Aprosthema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="D5B2EAE7FF81FD3E5C2030F8FDDCB5F4" bold="true" box="[344,630,1226,1249]" pageId="3" pageNumber="469">
<emphasis id="D5B2EAE7FF81FD3E5C2030F8FE77B5F4" bold="true" box="[344,477,1226,1249]" italics="true" pageId="3" pageNumber="469">Aprosthema</emphasis>
Konow, 1899
</emphasis>
</taxonomicName>
</heading>
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</subSubSection>
<subSubSection id="AFDC657EFF81FD3E5CD730DAFCB9B7FD" pageId="3" pageNumber="469" type="description">
<paragraph id="E77936F5FF81FD3E5CD730DAFD8AB5EA" blockId="3.[344,630,1226,1279]" box="[431,544,1256,1279]" pageId="3" pageNumber="469">
<figureCitation id="7FFD2A70FF81FD3E5CD730DAFE59B5EA" box="[431,499,1256,1279]" captionStart="Figs 2–15" captionStartId="4.[189,227,1814,1833]" captionTargetBox="[200,1409,368,1714]" captionTargetId="figure-4@4.[200,1410,360,1714]" captionTargetPageId="4" captionText="Figs 2–15. Argidae, details of morphology: 2, 10, 12 — Arge ustulata (Linné, 1758); 3, 11 — Aproceros leucopoda Takeuchi, 1939; 4, 8, 14 — Kokujewia ectrapela Konow, 1902; 5 — Spinarge metallica (Klug, 1834); 6 — Aprosthema melanurum (Klug, 1814); 7, 9 — Pseudarge rubicunda Gussakovskij, 1935; 13 — Sterictiphora geminata (Gmelin, 1790); 15 — Asiarge regeli Gussakovskij, 1935. 2–4 — veins R and Rs of fore wing, dorsally; 5–6 — male antenna, dorsally; 7–8 — hind tibia, ventrally; 9–11 — cells 1A and 2A of fore wing, dorsally; 12–13 — vein R+M of fore wing, dorsally; 14–15 — head, laterally. Рис. 2–15. Argidae, детали морфологии: 2, 10, 12 — Arge ustulata (Linné, 1758); 3, 11 — Aproceros leucopoda Takeuchi, 1939; 4, 8, 14 — Kokujewia ectrapela Konow, 1902; 5 — Spinarge metallica (Klug, 1834); 6 — Aprosthema melanurum (Klug, 1814); 7, 9 — Pseudarge rubicunda Gussakovskij, 1935; 13 — Sterictiphora geminata (Gmelin, 1790); 15 — Asiarge regeli Gussakovskij, 1935. 2–4 — жилки R и Rs переднего крыла, дорсально; 5–6 — антенна самца, дорсально; 7–8 — Задние голени, вентрально; 9–11 — Ячейки 1А и 2А переднего крыла, дорсально; 12–13 — жилка R+M переднего крыла, дорсально; 14–15 — голова, латерально." pageId="3" pageNumber="469">Figs 6</figureCitation>
,
<figureCitation id="7FFD2A70FF81FD3E5C8630DAFDB6B5EA" box="[510,540,1256,1279]" captionStart="Figs 16–24" captionStartId="5.[189,227,1843,1862]" captionTargetBox="[200,1409,783,1777]" captionTargetId="figure-260@5.[200,1410,783,1777]" captionTargetPageId="5" captionText="Figs 16–24. Argidae, details of morphology: 16, 24, 25 — Tanyphatnidea sinensis (W.F. Kirby, 1882); 17, 18 — Spinarge metallica (Klug, 1834); 19, 20, 21, 26 — Aproceros leucopoda Takeuchi, 1939; 22 — Sterictiphora geminata (Gmelin, 1790); 23 — Aprosthema melanurum (Klug, 1814); 16 — head and thorax, dorsally; 17–18 — medial tergites of abdomen, dorsally; 19 — labrum and right mandible, dorsally; 20 — claw, laterally; 21 — right mandible, dorsally; 22, 23, 26 — cell A of hind wing, dorsally; 24 — saw sheath, dorsally; 25 — saw sheath, laterally. Рис. 16–26. Argidae, детали морфологии: 16, 24, 25 — Tanyphatnidea sinensis (W.F. Kirby, 1882); 17, 18 — Spinarge metallica (Klug, 1834); 19, 20, 21, 26 — Aproceros leucopoda Takeuchi, 1939; 22 — Sterictiphora geminata (Gmelin, 1790); 23 — Aprosthema melanurum (Klug, 1814); 16 — голова и грудь, дорсально; 17–18 — медиальные тергиты брюшка, дорсально; 19 — нижнЯЯ губа и праваЯ мандибула, дорсально; 20 — коготок, латерально; 21 — праваЯ мандибула, дорсально; 22, 23, 26 — Ячейка A Заднего крыла, дорсально; 24 — ножны пилки, дорсально; 25 — ножны пилки, латерально." pageId="3" pageNumber="469">23</figureCitation>
.
</paragraph>
<paragraph id="E77936F5FF81FD3E5D9B3110FD6AB422" blockId="3.[189,788,1314,1768]" box="[227,704,1314,1335]" pageId="3" pageNumber="469">
<typeStatus id="387D8857FF81FD3E5D9B3110FEB9B422" box="[227,275,1314,1335]" pageId="3" pageNumber="469">Type</typeStatus>
species:
<taxonomicName id="20C64D76FF81FD3E5C123110FD16B422" authority="Fallen, 1808" authorityName="Fallen" authorityYear="1808" box="[362,700,1314,1335]" class="Insecta" family="Argidae" genus="Hylotoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="D5B2EAE7FF81FD3E5C123110FD91B422" box="[362,571,1314,1335]" italics="true" pageId="3" pageNumber="469">Hylotoma brevicornis</emphasis>
Fallén, 1808
</taxonomicName>
.
</paragraph>
<paragraph id="E77936F5FF81FD3E5D9B310FFDC9B705" blockId="3.[189,788,1314,1768]" pageId="3" pageNumber="469">Body size 4.5–8.5 mm. Labial palps with 4 segments, maxillary palps with 6 segments. Right mandible with basal tooth. Antennae of males bifurcate. Fore wings with 4 cells Rs; cell R open; vein Sc absent; vein 2r–m present; vein B of fore wing joins vein R well before (longer than vein Sc) junction point of vein M; cell 2A absent. Petiole of cell A of hind wing about 2.0 times its width. Claws simple.All tibiae without preapical spurs. The colour often yellow with black and without metallic sheen.</paragraph>
<paragraph id="E77936F5FF81FD3E5D9B3224FCB9B7FD" blockId="3.[189,788,1314,1768]" pageId="3" pageNumber="469">
The genus is distributed in the Palaearctic, Nearctic and Oriental regions. Number of species: World — 55; Palaearctic — 50. The life history of
<taxonomicName id="20C64D76FF81FD3E5CAA327EFCB8B774" authority="(Klug, 1814)" baseAuthorityName="Klug" baseAuthorityYear="1814" box="[466,786,1612,1633]" class="Insecta" family="Argidae" genus="Aprosthema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="tardum">
<emphasis id="D5B2EAE7FF81FD3E5CAA327EFD25B774" box="[466,655,1612,1633]" italics="true" pageId="3" pageNumber="469">Aprosthema tardum</emphasis>
(Klug, 1814)
</taxonomicName>
was investigated by
<bibRefCitation id="83574B04FF81FD3E5CFF3255FDE0B769" author="Liston A. D. &amp; Kan P. &amp; Kan-Van Limburg Stirum B." box="[391,586,1639,1660]" pageId="3" pageNumber="469" pagination="97 - 106" refId="ref10268" refString="Liston A. D., Kan P., Kan-Van Limburg Stirum B. 2018. The life - history of Aprosthema tardum (Klug, 1814) (Hymenoptera, Tenthredinoidea, Argidae) // Contributions to Entomology: Beitrage zur Entomologie. Vol. 68. No. 1. P. 97 - 106." type="journal article" year="2018">
Liston
<emphasis id="D5B2EAE7FF81FD3E5CB53255FE50B769" box="[461,506,1639,1660]" italics="true" pageId="3" pageNumber="469">et al</emphasis>
. [2018]
</bibRefCitation>
. The most complete key to the genus for the Palaearctic region was compiled by
<bibRefCitation id="83574B04FF81FD3E5DC532AFFE2FB7A7" author="Gussakovskij V. V." box="[189,389,1693,1714]" pageId="3" pageNumber="469" refId="ref8958" refString="Gussakovskij V. V. 1935. [Chalastogastra (pt. 1)] // Fauna SSSR. Nov. ser. Nasekomye Pereponchatokrylye. Vol. 2. No. 2. Moscow-Leningrad: AN SSSR Publ. 453 p. [In Russian, with French summary]" type="book" year="1935">Gussakovskij [1935]</bibRefCitation>
, but it is now considerably out of date and requires substantial revision and correction. Key to Chinese species see in
<bibRefCitation id="83574B04FF81FD3E5C3A32E1FE74B7FD" author="Wu R. &amp; Niu G. &amp; Wei M. C." box="[322,478,1747,1768]" pageId="3" pageNumber="469" pagination="225 - 229" refId="ref12088" refString="Wu R., Niu G., Wei M. C. 2021. A new species of Aprosthema Konow (Hymenoptera: Argidae) with a key to Chinese species // Entomotaxonomia. Vol. 43. No. 3. P. 225 - 229." type="book chapter" year="2021">
Wu
<emphasis id="D5B2EAE7FF81FD3E5C1032E1FE38B7FD" box="[360,402,1747,1768]" italics="true" pageId="3" pageNumber="469">et al</emphasis>
. [2021]
</bibRefCitation>
, European — in
<bibRefCitation id="83574B04FF81FD3E5F0432E1FCA5B7FD" author="Lacourt J." box="[636,783,1747,1768]" pageId="3" pageNumber="469" refId="ref9958" refString="Lacourt J. 2020. Sawflies of Europe: Hymenoptera of Europe 2. Verrieres-le-Buisson: N. A. P. editions. 876 p." type="book" year="2020">Lacourt [2020]</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
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<taxonomicName id="20C64D76FF81FD3E5C2635A6FDDBB0BE" authority="Malaise, 1931" authorityName="Malaise" authorityYear="1931" box="[350,625,404,427]" class="Insecta" family="Argidae" genus="Aproceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="genus">
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<emphasis id="D5B2EAE7FF81FD3E5C2635A6FE65B0BE" bold="true" box="[350,463,404,427]" italics="true" pageId="3" pageNumber="469">Aproceros</emphasis>
Malaise, 1931
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<paragraph id="E77936F5FF81FD3E5C0C3583FDF6B0DD" blockId="3.[350,625,404,456]" box="[372,604,433,456]" pageId="3" pageNumber="469">
<figureCitation id="7FFD2A70FF81FD3E5C0C3583FE74B0DD" box="[372,478,433,456]" captionStart="Figs 2–15" captionStartId="4.[189,227,1814,1833]" captionTargetBox="[200,1409,368,1714]" captionTargetId="figure-4@4.[200,1410,360,1714]" captionTargetPageId="4" captionText="Figs 2–15. Argidae, details of morphology: 2, 10, 12 — Arge ustulata (Linné, 1758); 3, 11 — Aproceros leucopoda Takeuchi, 1939; 4, 8, 14 — Kokujewia ectrapela Konow, 1902; 5 — Spinarge metallica (Klug, 1834); 6 — Aprosthema melanurum (Klug, 1814); 7, 9 — Pseudarge rubicunda Gussakovskij, 1935; 13 — Sterictiphora geminata (Gmelin, 1790); 15 — Asiarge regeli Gussakovskij, 1935. 2–4 — veins R and Rs of fore wing, dorsally; 5–6 — male antenna, dorsally; 7–8 — hind tibia, ventrally; 9–11 — cells 1A and 2A of fore wing, dorsally; 12–13 — vein R+M of fore wing, dorsally; 14–15 — head, laterally. Рис. 2–15. Argidae, детали морфологии: 2, 10, 12 — Arge ustulata (Linné, 1758); 3, 11 — Aproceros leucopoda Takeuchi, 1939; 4, 8, 14 — Kokujewia ectrapela Konow, 1902; 5 — Spinarge metallica (Klug, 1834); 6 — Aprosthema melanurum (Klug, 1814); 7, 9 — Pseudarge rubicunda Gussakovskij, 1935; 13 — Sterictiphora geminata (Gmelin, 1790); 15 — Asiarge regeli Gussakovskij, 1935. 2–4 — жилки R и Rs переднего крыла, дорсально; 5–6 — антенна самца, дорсально; 7–8 — Задние голени, вентрально; 9–11 — Ячейки 1А и 2А переднего крыла, дорсально; 12–13 — жилка R+M переднего крыла, дорсально; 14–15 — голова, латерально." pageId="3" pageNumber="469">Figs 3, 11</figureCitation>
,
<figureCitation id="7FFD2A70FF81FD3E5C923583FD85B0DD" box="[490,559,433,456]" captionStart="Figs 16–24" captionStartId="5.[189,227,1843,1862]" captionTargetBox="[200,1409,783,1777]" captionTargetId="figure-260@5.[200,1410,783,1777]" captionTargetPageId="5" captionText="Figs 16–24. Argidae, details of morphology: 16, 24, 25 — Tanyphatnidea sinensis (W.F. Kirby, 1882); 17, 18 — Spinarge metallica (Klug, 1834); 19, 20, 21, 26 — Aproceros leucopoda Takeuchi, 1939; 22 — Sterictiphora geminata (Gmelin, 1790); 23 — Aprosthema melanurum (Klug, 1814); 16 — head and thorax, dorsally; 17–18 — medial tergites of abdomen, dorsally; 19 — labrum and right mandible, dorsally; 20 — claw, laterally; 21 — right mandible, dorsally; 22, 23, 26 — cell A of hind wing, dorsally; 24 — saw sheath, dorsally; 25 — saw sheath, laterally. Рис. 16–26. Argidae, детали морфологии: 16, 24, 25 — Tanyphatnidea sinensis (W.F. Kirby, 1882); 17, 18 — Spinarge metallica (Klug, 1834); 19, 20, 21, 26 — Aproceros leucopoda Takeuchi, 1939; 22 — Sterictiphora geminata (Gmelin, 1790); 23 — Aprosthema melanurum (Klug, 1814); 16 — голова и грудь, дорсально; 17–18 — медиальные тергиты брюшка, дорсально; 19 — нижнЯЯ губа и праваЯ мандибула, дорсально; 20 — коготок, латерально; 21 — праваЯ мандибула, дорсально; 22, 23, 26 — Ячейка A Заднего крыла, дорсально; 24 — ножны пилки, дорсально; 25 — ножны пилки, латерально." pageId="3" pageNumber="469">19–21</figureCitation>
, 26.
</paragraph>
<paragraph id="E77936F5FF81FD3E5D9B35DEFD60B314" blockId="3.[189,787,492,1189]" box="[227,714,492,513]" pageId="3" pageNumber="469">
<typeStatus id="387D8857FF81FD3E5D9B35DEFEB9B314" box="[227,275,492,513]" pageId="3" pageNumber="469">Type</typeStatus>
species:
<taxonomicName id="20C64D76FF81FD3E5C1235DEFD6CB314" authority="Malaise, 1931" authorityName="Malaise" authorityYear="1931" box="[362,710,492,513]" class="Insecta" family="Argidae" genus="Aproceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="umbricola">
<emphasis id="D5B2EAE7FF81FD3E5C1235DEFD9CB314" box="[362,566,492,513]" italics="true" pageId="3" pageNumber="469">Aproceros umbricola</emphasis>
Malaise, 1931
</taxonomicName>
.
</paragraph>
<paragraph id="E77936F5FF81FD3E5D9B3635FD77B3AB" blockId="3.[189,787,492,1189]" pageId="3" pageNumber="469">Body size 4.5–7.0 mm. Labial palps with 4 segments, maxillary palps with 6 segments. Right mandible with basal tooth. Antennae of males bifurcate. Fore wings with 4 cells Rs; cell R open; vein Sc absent; vein 2r–m present; veins B and M join up at same point on vein R; cell 2A absent. Cell A of hind wing widely open. All tibiae without preapical spurs. The colour is usually black or brownish–red, without a metallic sheen.</paragraph>
<paragraph id="E77936F5FF81FD3E5D9B36F6FDEBB5B0" blockId="3.[189,787,492,1189]" pageId="3" pageNumber="469">
The genus is distributed in the Palaearctic (East and West) and Oriental regions. Number of species:World — 10; Palaearctic — 9. The larvae feed on the leaves of elms (Ulmus spp.).
<taxonomicName id="20C64D76FF81FD3E5FF636C8FE3EB23F" authority="Takeuchi, 1939" authorityName="Takeuchi" authorityYear="1939" class="Insecta" family="Argidae" genus="Aproceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="leucopoda">
<emphasis id="D5B2EAE7FF81FD3E5FF636C8FEAAB23F" italics="true" pageId="3" pageNumber="469">Aproceros leucopoda</emphasis>
Takeuchi, 1939
</taxonomicName>
is an invasive pest of elms (Ulmus spp.) that has spread widely over the last decade in central Europe (e.g.,
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<bibRefCitation id="83574B04FF81FD3E5DC5378AFEE3B2D8" author="Holling D." box="[189,329,952,973]" pageId="3" pageNumber="469" pagination="149 - 152" refId="ref9444" refString="Holling D. 2018. Aproceros leucopoda Takeuchi, 1939 - Erstbeobachtung der Zickzack - Ulmenblattwespe in der Schweiz (Hymenoptera: Argidae) // Entomo Helvetica. Vol. 11. S. 149 - 152." type="journal article" year="2018">Hölling [2018]</bibRefCitation>
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<bibRefCitation id="83574B04FF81FD3E5C2C378AFDBBB2D8" author="Burton L. &amp; Weiss J. &amp; Schneider N." box="[340,529,952,973]" pageId="3" pageNumber="469" pagination="185 - 188" refId="ref8243" refString="Burton L., Weiss J., Schneider N. 2019. Premiers signalements de cinq especes de symphytes (Hymenoptera, Symphyta) au Luxembourg: Aproceros leucopoda Takeuchi, 1939, Neurotoma saltuum (L., 1758), Craesus latipes (Villaret, 1832), Periclista pubescens (Zaddach, 1859) et Pristiphora geniculata (Hartig, 1840) // Bulletin de la Societe des Naturalistes Luxembourgeois. Vol. 121. P. 185 - 188." type="journal article" year="2019">
Burton
<emphasis id="D5B2EAE7FF81FD3E5CE3378AFE6EB2D8" box="[411,452,952,973]" italics="true" pageId="3" pageNumber="469">et al</emphasis>
. [2019]
</bibRefCitation>
;
<bibRefCitation id="83574B04FF81FD3E5F64378AFD49B2D8" author="Sinchuk A. &amp; Vaicekauskaite K. &amp; Sinchuk N." box="[540,739,952,973]" pageId="3" pageNumber="469" pagination="111 - 113" refId="ref11311" refString="Sinchuk A., Vaicekauskaite K., Sinchuk N. 2021. First record of Aproceros leucopoda Takeuchi, 1939 (Hymenoptera: Argidae) in Lithuania // Bulletin of the Lithuanian Entomological Society. Vol. 5. No. 33. P. 111 - 113." type="journal article" year="2021">
Sinchuk
<emphasis id="D5B2EAE7FF81FD3E5F16378AFD3DB2D8" box="[622,663,952,973]" italics="true" pageId="3" pageNumber="469">et al</emphasis>
. [2021]
</bibRefCitation>
) and
<collectingCountry id="9FD17665FF81FD3E5DC537E1FF53B2FD" box="[189,249,979,1000]" name="Russia" pageId="3" pageNumber="469">Russia</collectingCountry>
(e.g.,
<bibRefCitation id="83574B04FF81FD3E5C5537E1FE58B2FD" author="Artokhin K. S. &amp; Ignatova P. K. &amp; Terskov E. N." box="[301,498,979,1000]" pageId="3" pageNumber="469" pagination="199 - 202" refId="ref7586" refString="Artokhin K. S., Ignatova P. K., Terskov E. N. 2012. [New insects including invasive species for the fauna of Rostov Region (Russia)] // Caucasian Entomological Bulletin. Vol. 8. No. 2. P. 199 - 202 [in Russian]." type="book chapter" year="2012">
Artokhin
<emphasis id="D5B2EAE7FF81FD3E5CFB37E1FE03B2FD" box="[387,425,979,1000]" italics="true" pageId="3" pageNumber="469">et al</emphasis>
. [2012]
</bibRefCitation>
;
<bibRefCitation id="83574B04FF81FD3E5C8237E1FD35B2FD" author="Lengesova N. A." box="[506,671,979,1000]" pageId="3" pageNumber="469" pagination="136 - 139" refId="ref10097" refString="Lengesova N. A. 2012. [Elm sawfly (Aproceros leucopoda (Takeuchi, 1939) - new pest of elm for the territory of the Ulyanovsk Region] // Priroda Simbirskogo Povolzhyia. Vol. 13. P. 136 - 139 [in Russian]." type="book chapter" year="2012">Lengesova [2012]</bibRefCitation>
;
<bibRefCitation id="83574B04FF81FD3E5FDE37E1FEB2B516" author="Shchurov V. I. &amp; Gninenko Yu. I. &amp; Lengesova N. A. &amp; Gninenko M. Yu." pageId="3" pageNumber="469" pagination="37 - 38" refId="ref10843" refString="Shchurov V. I., Gninenko Yu. I., Lengesova N. A., Gninenko M. Yu. 2012. [Elm sawfly in the European part of Russia.] // Zashchita i karantin rastenii. No. 2. P. 37 - 38 [in Russian]." type="book chapter" year="2012">
Shchurov
<emphasis id="D5B2EAE7FF81FD3E5E7937E1FF64B516" italics="true" pageId="3" pageNumber="469">et al</emphasis>
. [2012]
</bibRefCitation>
;
<bibRefCitation id="83574B04FF81FD3E5C5937DCFE5BB516" author="Gninenko Yu. I. &amp; Gninenko M. Yu. &amp; Rakov A. G." box="[289,497,1006,1027]" pageId="3" pageNumber="469" pagination="40 - 41" refId="ref8908" refString="Gninenko Yu. I., Gninenko M. Yu., Rakov A. G. 2013. [New discoveries of the elm zigzag sawfly in Russia] // Zashchita i karantin rastenii. No. 3. P. 40 - 41 [in Russian]." type="book chapter" year="2013">
Gninenko
<emphasis id="D5B2EAE7FF81FD3E5CF837DCFE0CB516" box="[384,422,1006,1027]" italics="true" pageId="3" pageNumber="469">et al</emphasis>
. [2013]
</bibRefCitation>
; Lengesova, Mischenko [2013];
<bibRefCitation id="83574B04FF81FD3E5DC5303BFEE5B50B" author="Sorokin N. S." box="[189,335,1033,1054]" pageId="3" pageNumber="469" pagination="35 - 37" refId="ref11453" refString="Sorokin N. S. 2013. [Elm sawfly in the Rostov region] // Zashchita i karantin rastenii. No. 11. P. 35 - 37 [in Russian]." type="book chapter" year="2013">Sorokin [2013]</bibRefCitation>
;
<bibRefCitation id="83574B04FF81FD3E5C26303BFE54B50B" author="Blummer A. G." box="[350,510,1033,1054]" pageId="3" pageNumber="469" pagination="13 - 16" refId="ref8064" refString="Blummer A. G. 2015. [Elm sawfly-ziczag Aproceros leucopoda Takeuchi, 1939 (Hymenoptera, Argidae) - economically significant pest of elm from East Asia, expanding invasive area in the Europe- an part of Russia] // Karantin rasteniy. Nauka i praktika. Vol. 4. No. 14. P. 13 - 16 [in Russian]." type="journal article" year="2015">Blummer [2015]</bibRefCitation>
;
<bibRefCitation id="83574B04FF81FD3E5F75303BFCA5B50B" author="Martynov V. V. &amp; Nikulina T. V." box="[525,783,1033,1054]" pageId="3" pageNumber="469" pagination="25 - 34" refId="ref10494" refString="Martynov V. V., Nikulina T. V. 2017. Population surge of zigzag elm sawfly (Aproceros leucopoda (Takeuchi, 1939): Hymenoptera: Argidae) in Northern Sea of Azov region. // Rossiyskiy zhurnal biologicheskikh invaziy. Vol. 1. P. 25 - 34." type="journal article" year="2017">Martynov, Nikulina [2017]</bibRefCitation>
;
<bibRefCitation id="83574B04FF81FD3E5DC53016FE0EB52C" author="Kondratyeva A. M. &amp; Rzhevsky S. G. &amp; Aksenenko E. V. &amp; Kirpach D. A. &amp; Kornev I. I." box="[189,420,1060,1081]" pageId="3" pageNumber="469" pagination="63 - 68" refId="ref9885" refString="Kondratyeva A. M., Rzhevsky S. G., Aksenenko E. V., Kirpach D. A., Kornev I. I. 2023. Detection of a mass breeding site of the elm sawfly-zigzag Aproceros leucopoda Takeuchi, 1939 (Hymenoptera: Argidae) in Voronezh // Samara Journal of Science. Vol. 12. No. 3. P. 63 - 68." type="journal article" year="2023">
Kondratyeva
<emphasis id="D5B2EAE7FF81FD3E5C4E3016FEF1B52C" box="[310,347,1060,1081]" italics="true" pageId="3" pageNumber="469">et al</emphasis>
. [2023]
</bibRefCitation>
), and has also been accidently introduced to North America [
<bibRefCitation id="83574B04FF81FD3E5C1D300DFDBAB541" author="Martel V. &amp; Morin O. &amp; Monckton S. K. &amp; Eiseman C. S. &amp; Beliveau C. &amp; Cusson M. &amp; Blank S. M." box="[357,528,1087,1108]" pageId="3" pageNumber="469" pagination="1 - 18" refId="ref10421" refString="Martel V., Morin O., Monckton S. K., Eiseman C. S., Beliveau C., Cusson M., Blank S. M. 2021. Elm zigzag sawfly, Aproceros leucopoda (Hymenoptera: Argidae), recorded for the first time in North America through community science // The Canadian Entomologist. Vol. 154. P. 1 - 18." type="journal article" year="2021">
Martel
<emphasis id="D5B2EAE7FF81FD3E5CD1300DFE7AB541" box="[425,464,1087,1108]" italics="true" pageId="3" pageNumber="469">et al</emphasis>
., 2021
</bibRefCitation>
;
<bibRefCitation id="83574B04FF81FD3E5F61300DFD1EB541" author="Oten K. L. F. &amp; Day E. &amp; Dellinger T. &amp; Disque H. H. &amp; Barringer L. E. &amp; Cancelliere J. &amp; Somers L. &amp; Bertone M. A." box="[537,692,1087,1108]" pageId="3" pageNumber="469" pagination="1 - 7" refId="ref10719" refString="Oten K. L. F., Day E., Dellinger T., Disque H. H., Barringer L. E., Cancelliere J., Somers L., Bertone M. A. 2023. First records of elm zigzag sawfly (Hymenoptera: Argidae) in the United States // Journal of Integrated Pest Management. Vol. 14 (1). No. 12. P. 1 - 7." type="book chapter" year="2023">
Oten
<emphasis id="D5B2EAE7FF81FD3E5F33300DFDD8B541" box="[587,626,1087,1108]" italics="true" pageId="3" pageNumber="469">et al</emphasis>
., 2023
</bibRefCitation>
]. Males of
<taxonomicName id="20C64D76FF81FD3E5DC53068FD98B57A" authority="(Mocsary, 1909)" baseAuthorityName="Mocsary" baseAuthorityYear="1909" box="[189,562,1114,1135]" class="Insecta" family="Argidae" genus="Aproceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="pallidicornis">
<emphasis id="D5B2EAE7FF81FD3E5DC53068FE39B57A" box="[189,403,1114,1135]" italics="true" pageId="3" pageNumber="469">Aproceros pallidicornis</emphasis>
(Mocsáry, 1909)
</taxonomicName>
are further described in
<bibRefCitation id="83574B04FF81FD3E5DC53047FECAB59F" author="Choi J. - K. &amp; Lee S. B. &amp; Wei M. C. &amp; Lee J. W." box="[189,352,1141,1162]" pageId="3" pageNumber="469" pagination="323 - 327" refId="ref8377" refString="Choi J. - K., Lee S. B., Wei M. C., Lee J. W. 2015. Study on two genera, Yasumatsua Togashi and Aproceros Malaise (Hymenoptera: Argidae: Sterictiphorinae), from South Korea with description of a new species // Entomological Research. Vol. 45. P. 323 - 327." type="journal article" year="2015">
Choi
<emphasis id="D5B2EAE7FF81FD3E5D973047FEBFB59F" box="[239,277,1141,1162]" italics="true" pageId="3" pageNumber="469">et al</emphasis>
. [2015]
</bibRefCitation>
. The most complete identification key of
<taxonomicName id="20C64D76FF81FD3E5FAC3047FE83B5B0" pageId="3" pageNumber="469">
<emphasis id="D5B2EAE7FF81FD3E5FAC3047FF49B5B0" italics="true" pageId="3" pageNumber="469">Aproceros</emphasis>
species
</taxonomicName>
is presented in
<bibRefCitation id="83574B04FF81FD3E5CCA30A2FD97B5B0" author="Togashi I." box="[434,573,1168,1189]" pageId="3" pageNumber="469" pagination="59 - 61" refId="ref11525" refString="Togashi I. 2003. A new Aproceros Malaise (Hymenoptera, Symphyta, Argidae) from Japan // Biogeography. P. 59 - 61." type="book chapter" year="2003">Togashi [2003]</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

91
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<emphasis id="D5B2EAE7FF81FD3E5ECF3110FAA7B42C" bold="true" box="[951,1293,1314,1337]" pageId="3" pageNumber="469">
<emphasis id="D5B2EAE7FF81FD3E5ECF3110FBEDB42C" bold="true" box="[951,1095,1314,1337]" italics="true" pageId="3" pageNumber="469">Athermantus</emphasis>
W.F. Kirby, 1882
</emphasis>
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<paragraph id="E77936F5FF81FD3E5E26316EFAEFB464" blockId="3.[824,1422,1372,1798]" box="[862,1349,1372,1393]" pageId="3" pageNumber="469">
<typeStatus id="387D8857FF81FD3E5E26316EFC24B464" box="[862,910,1372,1393]" pageId="3" pageNumber="469">Type</typeStatus>
species:
<taxonomicName id="20C64D76FF81FD3E5E9D316EFAEBB464" authority="F. Smith, 1860" authorityName="F. Smith" authorityYear="1860" box="[997,1345,1372,1393]" class="Insecta" family="Argidae" genus="Hylotoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="469" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="D5B2EAE7FF81FD3E5E9D316EFB01B464" box="[997,1195,1372,1393]" italics="true" pageId="3" pageNumber="469">Hylotoma imperialis</emphasis>
F. Smith, 1860
</taxonomicName>
.
</paragraph>
<paragraph id="E77936F5FF81FD3E5E263145FAEDB771" blockId="3.[824,1422,1372,1798]" pageId="3" pageNumber="469">
Body size 13.0–16.0 mm. Head strongly enlarged behind eyes in dorsal view. Face roundly and strongly elevated, without middle ridge. Fore wings with 4 cells Rs; cell
<collectionCode id="81D7AE30FF81FD3E5848319FFAEAB4D7" box="[1328,1344,1453,1474]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="469">R</collectionCode>
closed; vein Sc present; vein 2r–m present; vein B of fore wing joins vein
<collectionCode id="81D7AE30FF81FD3E5E1F31D1FCDDB4ED" box="[871,887,1507,1528]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="469">R</collectionCode>
well before (longer than vein Sc) junction point of vein M. All tibiae without preapical spurs. Hind tibiae and apical spurs flattened. Saw sheath short and very broad, round at apex in dorsal view. Body black with strong metallic purplish tinge. For additional generic characters see
<bibRefCitation id="83574B04FF81FD3E59E6327DFAE9B771" author="Luo X. &amp; He H. &amp; Wei M. C." box="[1182,1347,1615,1636]" pageId="3" pageNumber="469" pagination="313 - 317" refId="ref10376" refString="Luo X., He H., Wei M. C. 2019. A new species of Athermantus Kirby (Hymenoptera: Argidae) from China // Entomotaxonomia. Vol. 41. No. 4. P. 313 - 317." type="book chapter" year="2019">
Luo
<emphasis id="D5B2EAE7FF81FD3E59B2327DFB5FB771" box="[1226,1269,1615,1636]" italics="true" pageId="3" pageNumber="469">et al</emphasis>
. [2019]
</bibRefCitation>
.
</paragraph>
<paragraph id="E77936F5FF81FD3E5E263258FAB9B613" blockId="3.[824,1422,1372,1798]" pageId="3" pageNumber="469">
This small genus is distributed in the Palaearctic (East) and Oriental regions. Number of species: World — 3; Palaearctic — 1. The body size of
<taxonomicName id="20C64D76FF81FD3E59383292FAA2B7A0" box="[1088,1288,1696,1717]" pageId="3" pageNumber="469">
<emphasis id="D5B2EAE7FF81FD3E59383292FB11B7A0" box="[1088,1211,1696,1717]" italics="true" pageId="3" pageNumber="469">Athermantus</emphasis>
species
</taxonomicName>
is the largest among the members of
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. Recently, a description of a new species from
<collectingCountry id="9FD17665FF81FD3E5E9332E4FB8EB7FE" box="[1003,1060,1750,1771]" name="China" pageId="3" pageNumber="469">China</collectingCountry>
was published with an identification key to the species of the genus [
<bibRefCitation id="83574B04FF81FD3E591132C3FAA3B613" author="Luo X. &amp; He H. &amp; Wei M. C." box="[1129,1289,1777,1798]" pageId="3" pageNumber="469" pagination="313 - 317" refId="ref10376" refString="Luo X., He H., Wei M. C. 2019. A new species of Athermantus Kirby (Hymenoptera: Argidae) from China // Entomotaxonomia. Vol. 41. No. 4. P. 313 - 317." type="book chapter" year="2019">
Luo
<emphasis id="D5B2EAE7FF81FD3E59E032C3FB68B613" box="[1176,1218,1777,1798]" italics="true" pageId="3" pageNumber="469">et al</emphasis>
., 2019
</bibRefCitation>
].
</paragraph>
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</treatment>
</document>

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<mods:title id="651CD108D3DF553FC5AE427CF799AAC8">Illustrated key to the Palaearctic genera of the sawfly family Argidae (Hymenoptera: Symphyta), with taxonomic notes</mods:title>
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<taxonomicName id="20C64D76FF84FD3B5EB0336CFB57B660" authority="Billberg, 1820" authorityName="Billberg" authorityYear="1820" box="[968,1277,1886,1909]" class="Insecta" family="Argidae" genus="Sterictiphora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="472" phylum="Arthropoda" rank="genus">
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<emphasis id="D5B2EAE7FF84FD3B5EB0336CFBF2B660" bold="true" box="[968,1112,1886,1909]" italics="true" pageId="6" pageNumber="472">Sterictiphora</emphasis>
Billberg, 1820
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<figureCitation id="7FFD2A70FF84FD3B5952334EFBC4B686" box="[1066,1134,1916,1939]" captionStart="Figs 2–15" captionStartId="4.[189,227,1814,1833]" captionTargetBox="[200,1409,368,1714]" captionTargetId="figure-4@4.[200,1410,360,1714]" captionTargetPageId="4" captionText="Figs 2–15. Argidae, details of morphology: 2, 10, 12 — Arge ustulata (Linné, 1758); 3, 11 — Aproceros leucopoda Takeuchi, 1939; 4, 8, 14 — Kokujewia ectrapela Konow, 1902; 5 — Spinarge metallica (Klug, 1834); 6 — Aprosthema melanurum (Klug, 1814); 7, 9 — Pseudarge rubicunda Gussakovskij, 1935; 13 — Sterictiphora geminata (Gmelin, 1790); 15 — Asiarge regeli Gussakovskij, 1935. 2–4 — veins R and Rs of fore wing, dorsally; 5–6 — male antenna, dorsally; 7–8 — hind tibia, ventrally; 9–11 — cells 1A and 2A of fore wing, dorsally; 12–13 — vein R+M of fore wing, dorsally; 14–15 — head, laterally. Рис. 2–15. Argidae, детали морфологии: 2, 10, 12 — Arge ustulata (Linné, 1758); 3, 11 — Aproceros leucopoda Takeuchi, 1939; 4, 8, 14 — Kokujewia ectrapela Konow, 1902; 5 — Spinarge metallica (Klug, 1834); 6 — Aprosthema melanurum (Klug, 1814); 7, 9 — Pseudarge rubicunda Gussakovskij, 1935; 13 — Sterictiphora geminata (Gmelin, 1790); 15 — Asiarge regeli Gussakovskij, 1935. 2–4 — жилки R и Rs переднего крыла, дорсально; 5–6 — антенна самца, дорсально; 7–8 — Задние голени, вентрально; 9–11 — Ячейки 1А и 2А переднего крыла, дорсально; 12–13 — жилка R+M переднего крыла, дорсально; 14–15 — голова, латерально." pageId="6" pageNumber="472">Figs 6</figureCitation>
,
<figureCitation id="7FFD2A70FF84FD3B5901334EFB3DB686" box="[1145,1175,1916,1939]" captionStart="Figs 16–24" captionStartId="5.[189,227,1843,1862]" captionTargetBox="[200,1409,783,1777]" captionTargetId="figure-260@5.[200,1410,783,1777]" captionTargetPageId="5" captionText="Figs 16–24. Argidae, details of morphology: 16, 24, 25 — Tanyphatnidea sinensis (W.F. Kirby, 1882); 17, 18 — Spinarge metallica (Klug, 1834); 19, 20, 21, 26 — Aproceros leucopoda Takeuchi, 1939; 22 — Sterictiphora geminata (Gmelin, 1790); 23 — Aprosthema melanurum (Klug, 1814); 16 — head and thorax, dorsally; 17–18 — medial tergites of abdomen, dorsally; 19 — labrum and right mandible, dorsally; 20 — claw, laterally; 21 — right mandible, dorsally; 22, 23, 26 — cell A of hind wing, dorsally; 24 — saw sheath, dorsally; 25 — saw sheath, laterally. Рис. 16–26. Argidae, детали морфологии: 16, 24, 25 — Tanyphatnidea sinensis (W.F. Kirby, 1882); 17, 18 — Spinarge metallica (Klug, 1834); 19, 20, 21, 26 — Aproceros leucopoda Takeuchi, 1939; 22 — Sterictiphora geminata (Gmelin, 1790); 23 — Aprosthema melanurum (Klug, 1814); 16 — голова и грудь, дорсально; 17–18 — медиальные тергиты брюшка, дорсально; 19 — нижнЯЯ губа и праваЯ мандибула, дорсально; 20 — коготок, латерально; 21 — праваЯ мандибула, дорсально; 22, 23, 26 — Ячейка A Заднего крыла, дорсально; 24 — ножны пилки, дорсально; 25 — ножны пилки, латерально." pageId="6" pageNumber="472">22</figureCitation>
.
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species:
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<emphasis id="D5B2EAE7FF84FD3B5E9D3384FB24B6DE" box="[997,1166,1974,1995]" italics="true" pageId="6" pageNumber="472">Hylotoma furcata</emphasis>
(Villers, 1789)
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. Body size 5.0–7.5 mm. Labial palps with 4 segments, maxillary palps with 6 segments. Right mandible with basal tooth. Antennae of males bifurcate. Fore wings with 4 cells Rs; cell
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open; vein
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absent; vein 2r–m present; veins B of fore wing joins vein
<collectionCode id="81D7AE30FF85FD3A5C583578FE9AB04A" box="[288,304,330,351]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="473">R</collectionCode>
well before (longer than vein Sc) junction point of vein M; cell 2A closed; petiole of cell A of hind wing shorter than its width.
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of hind wing closed.
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simple
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The genus is distributed in the Palaearctic, Nearctic, and Oriental regions. Number of species: World — 42; Palaearctic — 37. Two new species have been recently described from
<collectingCountry id="9FD17665FF85FD3A5DC535D8FE92B0EA" box="[189,312,490,511]" name="South Korea" pageId="7" pageNumber="473">South Korea</collectingCountry>
[
<bibRefCitation id="83574B04FF85FD3A5C3D35D8FE40B0EA" author="Wei M. C. &amp; Lee S. B. &amp; Choi J. - K. &amp; Le J. W." box="[325,490,490,511]" pageId="7" pageNumber="473" pagination="63 - 73" refId="ref12028" refString="Wei M. C., Lee S. B., Choi J. - K., Le J. W. 2017. Taxonomy of Sterictiphora Billberg (Hymenoptera: Argidae: Sterictiphorinae) in South Korea with descriptions of two new species // Entomological Research. Vol. 47. P. 63 - 73." type="book chapter" year="2017">
Wei
<emphasis id="D5B2EAE7FF85FD3A5C0E35D8FE08B0EA" box="[374,418,490,511]" italics="true" pageId="7" pageNumber="473">et al</emphasis>
., 2017
</bibRefCitation>
]. Rosa multiflora Thunb. (
<taxonomicName id="20C64D76FF85FD3A5F9535D8FF57B30C" class="Magnoliopsida" family="Rosaceae" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="473" phylum="Tracheophyta" rank="family">Rosaceae</taxonomicName>
) was recorded as host plant for
<taxonomicName id="20C64D76FF85FD3A5F4D3636FF47B321" authority="Takeuchi, 1939" authorityName="Takeuchi" authorityYear="1939" class="Insecta" family="Argidae" genus="Sterictiphora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="473" phylum="Arthropoda" rank="species" species="nipponica">
<emphasis id="D5B2EAE7FF85FD3A5F4D3636FD04B30C" box="[565,686,516,537]" italics="true" pageId="7" pageNumber="473">S. nipponica</emphasis>
Takeuchi, 1939
</taxonomicName>
[
<bibRefCitation id="83574B04FF85FD3A5D84362DFE6EB321" author="Hideho H. &amp; Shinichi I. &amp; Akihiko S." box="[252,452,543,564]" pageId="7" pageNumber="473" pagination="163 - 188" refId="ref9383" refString="Hideho H., Shinichi I., Akihiko S. 2021. Taxonomic Notes and New Distribution and Host Plant Records for Sawflies and Woodwasps (Hymenoptera, Symphyta) of Japan VI // Bulletin of the National Science Museum, Series A, Zoology. Vol. 47. No. 4. P. 163 - 188." type="journal article" year="2021">
Hideho
<emphasis id="D5B2EAE7FF85FD3A5C37362DFED7B321" box="[335,381,543,564]" italics="true" pageId="7" pageNumber="473">et al</emphasis>
., 2021
</bibRefCitation>
] and
<taxonomicName id="20C64D76FF85FD3A5C86362DFD60B321" authority="Medik." authorityName="Medik." box="[510,714,543,564]" class="Magnoliopsida" family="Rosaceae" genus="Amelanchier" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="473" phylum="Tracheophyta" rank="genus">Amelanchier Medik.</taxonomicName>
(
<taxonomicName id="20C64D76FF85FD3A5FA1362DFF43B35A" class="Magnoliopsida" family="Rosaceae" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="473" phylum="Tracheophyta" rank="family">Rosaceae</taxonomicName>
) for
<taxonomicName id="20C64D76FF85FD3A5C653608FDB0B35A" authority="(Norton, 1867)" baseAuthorityName="Norton" baseAuthorityYear="1867" box="[285,538,570,591]" class="Insecta" family="Argidae" genus="Sterictiphora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="473" phylum="Arthropoda" rank="species" species="sericea">
<emphasis id="D5B2EAE7FF85FD3A5C653608FED7B35A" box="[285,381,570,591]" italics="true" pageId="7" pageNumber="473">S. sericea</emphasis>
(Norton, 1867)
</taxonomicName>
[
<bibRefCitation id="83574B04FF85FD3A5F503608FCA3B35A" author="Eiseman C. S. &amp; Smith D. R." box="[552,777,570,591]" pageId="7" pageNumber="473" pagination="299 - 307" refId="ref8747" refString="Eiseman C. S., Smith D. R. 2020. New sawfly (Hymenoptera: Argidae, Tenthredinidae) host records from northeastern North America // Proceedings of the Entomological Society of Washington. Vol. 122. No. 2. P. 299 - 307." type="book chapter" year="2020">Eiseman, Smith, 2020</bibRefCitation>
].
<taxonomicName id="20C64D76FF85FD3A5DC53666FE0EB37C" authority="Wei, 2010" authorityName="Wei" authorityYear="2010" box="[189,420,596,617]" class="Insecta" family="Argidae" genus="Sterictiphora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="473" phylum="Arthropoda" rank="species" species="brevinerva">
<emphasis id="D5B2EAE7FF85FD3A5DC53666FE96B37C" box="[189,316,596,617]" italics="true" pageId="7" pageNumber="473">S. brevinerva</emphasis>
<bibRefCitation id="83574B04FF85FD3A5C3A3666FE0EB37C" author="Wei M. &amp; Niu G." box="[322,420,596,617]" pageId="7" pageNumber="473" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei, 2010</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="20C64D76FF85FD3A5CAB3666FCB8B37C" authority="Wei, 2010" authorityName="Wei" authorityYear="2010" box="[467,786,596,617]" class="Insecta" family="Argidae" genus="Sterictiphora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="473" phylum="Arthropoda" rank="species" species="nigrometathoracina">
<emphasis id="D5B2EAE7FF85FD3A5CAB3666FD00B37C" box="[467,682,596,617]" italics="true" pageId="7" pageNumber="473">S. nigrometathoracina</emphasis>
<bibRefCitation id="83574B04FF85FD3A5FC83666FCB8B37C" author="Wei M. &amp; Niu G." box="[688,786,596,617]" pageId="7" pageNumber="473" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei, 2010</bibRefCitation>
</taxonomicName>
have been reported as new taxa [
<bibRefCitation id="83574B04FF85FD3A5F73365DFD05B391" author="Wu X. - Y. &amp; Xin X. &amp; Zhang X. - Z." box="[523,687,623,644]" pageId="7" pageNumber="473" pagination="15 - 19" refId="ref12137" refString="Wu X. - Y., Xin X., Zhang X. - Z. 2010. [The Investigation and Taxonomical Research of the Sawflies Species from Gansu Province. III [recte: IV]. A list of the genera and species (Hymenoptera: Argidae).] // Journal of Gansu Forestry Science and Technology. Vol. 35. No. 1. P. 15 - 19 [in Chinese]." type="journal article" year="2010">
Wu
<emphasis id="D5B2EAE7FF85FD3A5F41365DFDCDB391" box="[569,615,623,644]" italics="true" pageId="7" pageNumber="473">et al</emphasis>
., 2010
</bibRefCitation>
] but their descriptions have not been published to date. These species should be treated as nomina nuda.
</paragraph>
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Konow, 1899
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species:
<taxonomicName id="20C64D76FF84FD3B5C123652FD24B360" authority="Konow, 1899" authorityName="Konow" authorityYear="1899" box="[362,654,608,629]" class="Insecta" family="Argidae" genus="Pampsilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="472" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="D5B2EAE7FF84FD3B5C123652FDAFB360" box="[362,517,608,629]" italics="true" pageId="6" pageNumber="472">Pampsilota afer</emphasis>
Konow, 1899
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.
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Body size 5.3–15.3 mm. Eyes not convex, inner margins of eyes nearly parallel. Width of head slightly less than width of thorax. Antennae of males simple. Third segment of antennae distinctly compressed and expanded. Fore wings with 4 cells Rs; cell
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closed; vein Sc present; vein 2r–m present; veins B and M join up at same point on vein
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; cell 2A closed; upper margin of cell 3Rs almost 2.0 times as large as lower. All tibiae without preapical spurs. Saw sheath obtuse in lateral view, width at the base view almost 2.0 times its length in dorsal.
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Representatives of the genus
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<emphasis id="D5B2EAE7FF84FD3B5C83375FFDC2B297" box="[507,616,877,898]" italics="true" pageId="6" pageNumber="472">Pampsilota</emphasis>
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have a very large morphological diversity of genital structures, which raises doubts about the monophyly of the genus.
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Liston
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. [2017]
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suggested that
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might reasonably be treated as merely a species group within
<taxonomicName id="20C64D76FF84FD3B5C9B37EBFDBBB2FB" authorityName="Schrank" authorityYear="1802" box="[483,529,985,1006]" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="472" phylum="Arthropoda" rank="genus">
<emphasis id="D5B2EAE7FF84FD3B5C9B37EBFDBBB2FB" box="[483,529,985,1006]" italics="true" pageId="6" pageNumber="472">Arge</emphasis>
</taxonomicName>
.
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The genus is distributed in the Palaearctic (East), Oriental and Afrotropical regions. Number of species: World — 8; Palaearctic — 2. Within the Palaearctic, members of
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<emphasis id="D5B2EAE7FF84FD3B5FDD3018FCB8B52A" box="[677,786,1066,1087]" italics="true" pageId="6" pageNumber="472">Pampsilota</emphasis>
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have been found only in northern
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. For a key to Afrotropical
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species
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, see
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Liston
<emphasis id="D5B2EAE7FF84FD3B5F413052FDC9B560" box="[569,611,1120,1141]" italics="true" pageId="6" pageNumber="472">et al</emphasis>
. [2017]
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<figureCitation id="7FFD2A70FF84FD3B5CCE32BBFDBFB7B5" box="[438,533,1673,1696]" captionStart="Figs 2–15" captionStartId="4.[189,227,1814,1833]" captionTargetBox="[200,1409,368,1714]" captionTargetId="figure-4@4.[200,1410,360,1714]" captionTargetPageId="4" captionText="Figs 2–15. Argidae, details of morphology: 2, 10, 12 — Arge ustulata (Linné, 1758); 3, 11 — Aproceros leucopoda Takeuchi, 1939; 4, 8, 14 — Kokujewia ectrapela Konow, 1902; 5 — Spinarge metallica (Klug, 1834); 6 — Aprosthema melanurum (Klug, 1814); 7, 9 — Pseudarge rubicunda Gussakovskij, 1935; 13 — Sterictiphora geminata (Gmelin, 1790); 15 — Asiarge regeli Gussakovskij, 1935. 2–4 — veins R and Rs of fore wing, dorsally; 5–6 — male antenna, dorsally; 7–8 — hind tibia, ventrally; 9–11 — cells 1A and 2A of fore wing, dorsally; 12–13 — vein R+M of fore wing, dorsally; 14–15 — head, laterally. Рис. 2–15. Argidae, детали морфологии: 2, 10, 12 — Arge ustulata (Linné, 1758); 3, 11 — Aproceros leucopoda Takeuchi, 1939; 4, 8, 14 — Kokujewia ectrapela Konow, 1902; 5 — Spinarge metallica (Klug, 1834); 6 — Aprosthema melanurum (Klug, 1814); 7, 9 — Pseudarge rubicunda Gussakovskij, 1935; 13 — Sterictiphora geminata (Gmelin, 1790); 15 — Asiarge regeli Gussakovskij, 1935. 2–4 — жилки R и Rs переднего крыла, дорсально; 5–6 — антенна самца, дорсально; 7–8 — Задние голени, вентрально; 9–11 — Ячейки 1А и 2А переднего крыла, дорсально; 12–13 — жилка R+M переднего крыла, дорсально; 14–15 — голова, латерально." pageId="6" pageNumber="472">Figs 7, 9</figureCitation>
.
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<emphasis id="D5B2EAE7FF84FD3B5C1232F6FD93B7CC" box="[362,569,1732,1753]" italics="true" pageId="6" pageNumber="472">Pseudarge rubicunda</emphasis>
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<paragraph id="E77936F5FF84FD3B5D9B32EDFCB9B683" blockId="6.[189,788,1732,2076]" pageId="6" pageNumber="472">Body size 8.0–11.5 mm. Supraclypeal area of face conspicuously, roundly protruding in lateral view. Antennae of males simple. Fore wings with 4 cells Rs; cell R closed; vein Sc present; vein 2r–m present; veins B and M join up at same point on vein R; cell 2A open; cell 3Rs shorter than in height. Cell A of hind wing closed. Middle and hind tibiae with preapical spurs. For additional generic characters, see Koch and Goergen [2008].</paragraph>
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The genus is distributed in the Palaearctic and Oriental regions. Number of species: World — 7; Palaearctic — 4. Recently, the males of two species have been described for the first time [
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] and a new species from the Crimean Peninsula described [
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<emphasis id="D5B2EAE7FF84FD3B5C6D30ABFE73B5A5" bold="true" box="[277,473,1177,1200]" italics="true" pageId="6" pageNumber="472">Pseudaprosthema</emphasis>
<bibRefCitation id="83574B04FF84FD3B5CA730ABFD10B5A5" author="Gussakovskij V. V." box="[479,698,1177,1200]" pageId="6" pageNumber="472" refId="ref8958" refString="Gussakovskij V. V. 1935. [Chalastogastra (pt. 1)] // Fauna SSSR. Nov. ser. Nasekomye Pereponchatokrylye. Vol. 2. No. 2. Moscow-Leningrad: AN SSSR Publ. 453 p. [In Russian, with French summary]" type="book" year="1935">Gussakovskij, 1935</bibRefCitation>
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species:
<taxonomicName id="20C64D76FF84FD3B5C0030E6FF5AB411" authority="Gussakovskij, 1935" authorityName="Gussakovskij" authorityYear="1935" class="Insecta" family="Argidae" genus="Pseudaprosthema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="472" phylum="Arthropoda" rank="species" species="barbara">
<emphasis id="D5B2EAE7FF84FD3B5C0030E6FDD4B5FC" box="[376,638,1236,1257]" italics="true" pageId="6" pageNumber="472">Pseudaprosthema barbara</emphasis>
<bibRefCitation id="83574B04FF84FD3B5FF330E6FF5AB411" author="Gussakovskij V. V." pageId="6" pageNumber="472" refId="ref8958" refString="Gussakovskij V. V. 1935. [Chalastogastra (pt. 1)] // Fauna SSSR. Nov. ser. Nasekomye Pereponchatokrylye. Vol. 2. No. 2. Moscow-Leningrad: AN SSSR Publ. 453 p. [In Russian, with French summary]" type="book" year="1935">Gussakovskij, 1935</bibRefCitation>
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.
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<paragraph id="E77936F5FF84FD3B5D9B3138FDD9B4D4" blockId="6.[189,787,1236,1607]" pageId="6" pageNumber="472">
Body size about 5.0 mm. Labial palps with 4 segments, maxillary palps with 6 segments. Right mandible with basal tooth. Antennae of males bifurcate. Fore wings with 4 cells Rs; cell
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open; vein 2r–m present; vein Sc present; vein B of fore wing joins vein
<collectionCode id="81D7AE30FF84FD3B5C253144FEC7B49E" box="[349,365,1398,1419]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="472">R</collectionCode>
well before (longer than vein Sc) junction point of vein M; pterostigma large; cell 2A open. Cell A of hind wing open. All tibiae without preapical spurs.
</paragraph>
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<materialsCitation id="57AE3CA8FF84FD3B5D9B31F4FE94B752" county="Palaearctic" location="The" municipality="World" pageId="6" pageNumber="472" specimenCount="1">
<location id="E219602EFF84FD3B5D9B31F4FEA2B4CE" LSID="urn:lsid:plazi:treatment:6F6F87E3FF84FD3B5C6D30ABFEE8B752:E219602EFF84FD3B5D9B31F4FEA2B4CE" box="[227,264,1478,1499]" county="Palaearctic" municipality="World" name="The" pageId="6" pageNumber="472">The</location>
genus is distributed in the
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.
<location id="E219602EFF84FD3B5DC531D3FEA6B4E3" LSID="urn:lsid:plazi:treatment:6F6F87E3FF84FD3B5C6D30ABFEE8B752:E219602EFF84FD3B5DC531D3FEA6B4E3" box="[189,268,1505,1526]" county="Palaearctic" municipality="World" name="Number" pageId="6" pageNumber="472">Number</location>
of species:
<collectingMunicipality id="071DAC8FFF84FD3B5C0231D3FE1FB4E3" box="[378,437,1505,1526]" pageId="6" pageNumber="472">World</collectingMunicipality>
and
<collectingCounty id="0E184E79FF84FD3B5C9931D3FDE0B4E3" box="[481,586,1505,1526]" pageId="6" pageNumber="472">Palaearctic</collectingCounty>
— 1.
<bibRefCitation id="83574B04FF84FD3B5FFB31D3FCB9B4E3" author="Lacourt J." box="[643,787,1505,1526]" pageId="6" pageNumber="472" refId="ref9958" refString="Lacourt J. 2020. Sawflies of Europe: Hymenoptera of Europe 2. Verrieres-le-Buisson: N. A. P. editions. 876 p." type="book" year="2020">Lacourt [2020]</bibRefCitation>
synonymized all three species with each other, but noted that this requires further verification with a re-examination of the
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material
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.
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149
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<heading id="BC318199FF84FD3B5E8E36D7FB65B3E9" box="[1014,1231,741,764]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="472" reason="8">
<taxonomicName id="20C64D76FF84FD3B5E8E36D7FB65B3E9" authority="Wei, 1998" authorityName="Wei" authorityYear="1998" box="[1014,1231,741,764]" class="Insecta" family="Argidae" genus="Spinarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="472" phylum="Arthropoda" rank="genus">
<emphasis id="D5B2EAE7FF84FD3B5E8E36D7FB65B3E9" bold="true" box="[1014,1231,741,764]" pageId="6" pageNumber="472">
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Wei, 1998
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<paragraph id="E77936F5FF84FD3B596E3730FB05B20C" blockId="6.[1014,1231,741,793]" box="[1046,1199,770,793]" pageId="6" pageNumber="472">
<figureCitation id="7FFD2A70FF84FD3B596E3730FBF0B20C" box="[1046,1114,770,793]" captionStart="Figs 2–15" captionStartId="4.[189,227,1814,1833]" captionTargetBox="[200,1409,368,1714]" captionTargetId="figure-4@4.[200,1410,360,1714]" captionTargetPageId="4" captionText="Figs 2–15. Argidae, details of morphology: 2, 10, 12 — Arge ustulata (Linné, 1758); 3, 11 — Aproceros leucopoda Takeuchi, 1939; 4, 8, 14 — Kokujewia ectrapela Konow, 1902; 5 — Spinarge metallica (Klug, 1834); 6 — Aprosthema melanurum (Klug, 1814); 7, 9 — Pseudarge rubicunda Gussakovskij, 1935; 13 — Sterictiphora geminata (Gmelin, 1790); 15 — Asiarge regeli Gussakovskij, 1935. 2–4 — veins R and Rs of fore wing, dorsally; 5–6 — male antenna, dorsally; 7–8 — hind tibia, ventrally; 9–11 — cells 1A and 2A of fore wing, dorsally; 12–13 — vein R+M of fore wing, dorsally; 14–15 — head, laterally. Рис. 2–15. Argidae, детали морфологии: 2, 10, 12 — Arge ustulata (Linné, 1758); 3, 11 — Aproceros leucopoda Takeuchi, 1939; 4, 8, 14 — Kokujewia ectrapela Konow, 1902; 5 — Spinarge metallica (Klug, 1834); 6 — Aprosthema melanurum (Klug, 1814); 7, 9 — Pseudarge rubicunda Gussakovskij, 1935; 13 — Sterictiphora geminata (Gmelin, 1790); 15 — Asiarge regeli Gussakovskij, 1935. 2–4 — жилки R и Rs переднего крыла, дорсально; 5–6 — антенна самца, дорсально; 7–8 — Задние голени, вентрально; 9–11 — Ячейки 1А и 2А переднего крыла, дорсально; 12–13 — жилка R+M переднего крыла, дорсально; 14–15 — голова, латерально." pageId="6" pageNumber="472">Figs 5</figureCitation>
,
<figureCitation id="7FFD2A70FF84FD3B591D3730FB01B20C" box="[1125,1195,770,793]" captionStart="Figs 16–24" captionStartId="5.[189,227,1843,1862]" captionTargetBox="[200,1409,783,1777]" captionTargetId="figure-260@5.[200,1410,783,1777]" captionTargetPageId="5" captionText="Figs 16–24. Argidae, details of morphology: 16, 24, 25 — Tanyphatnidea sinensis (W.F. Kirby, 1882); 17, 18 — Spinarge metallica (Klug, 1834); 19, 20, 21, 26 — Aproceros leucopoda Takeuchi, 1939; 22 — Sterictiphora geminata (Gmelin, 1790); 23 — Aprosthema melanurum (Klug, 1814); 16 — head and thorax, dorsally; 17–18 — medial tergites of abdomen, dorsally; 19 — labrum and right mandible, dorsally; 20 — claw, laterally; 21 — right mandible, dorsally; 22, 23, 26 — cell A of hind wing, dorsally; 24 — saw sheath, dorsally; 25 — saw sheath, laterally. Рис. 16–26. Argidae, детали морфологии: 16, 24, 25 — Tanyphatnidea sinensis (W.F. Kirby, 1882); 17, 18 — Spinarge metallica (Klug, 1834); 19, 20, 21, 26 — Aproceros leucopoda Takeuchi, 1939; 22 — Sterictiphora geminata (Gmelin, 1790); 23 — Aprosthema melanurum (Klug, 1814); 16 — голова и грудь, дорсально; 17–18 — медиальные тергиты брюшка, дорсально; 19 — нижнЯЯ губа и праваЯ мандибула, дорсально; 20 — коготок, латерально; 21 — праваЯ мандибула, дорсально; 22, 23, 26 — Ячейка A Заднего крыла, дорсально; 24 — ножны пилки, дорсально; 25 — ножны пилки, латерально." pageId="6" pageNumber="472">17–18</figureCitation>
.
</paragraph>
<paragraph id="E77936F5FF84FD3B5E26370FFA80B247" blockId="6.[824,1422,829,1849]" box="[862,1322,829,850]" pageId="6" pageNumber="472">
<typeStatus id="387D8857FF84FD3B5E26370FFC24B247" box="[862,910,829,850]" pageId="6" pageNumber="472">Type</typeStatus>
species:
<taxonomicName id="20C64D76FF84FD3B5E9D370FFA8CB247" authority="Wei, 1998" authorityName="Wei" authorityYear="1998" box="[997,1318,829,850]" class="Insecta" family="Argidae" genus="Spinarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="472" phylum="Arthropoda" rank="species" species="sichuanensis">
<emphasis id="D5B2EAE7FF84FD3B5E9D370FFB16B247" box="[997,1212,829,850]" italics="true" pageId="6" pageNumber="472">Spinarge sichuanensis</emphasis>
Wei, 1998
</taxonomicName>
.
</paragraph>
<paragraph id="E77936F5FF84FD3B5E26376AFAF8B51A" blockId="6.[824,1422,829,1849]" pageId="6" pageNumber="472">Body size 7.2–11.9 mm. Antennae of males simple. Fore wings with 4 cells Rs; cell R closed; vein Sc present; vein 2r–m present; cell 2A closed. Cell A of hind wing closed. Hind tibiae with or without preapical spurs. Fifth abdominal tergite of male has a long medial process, in females represented by a scarcely noticeable triangular swelling and inconspicuous median groove on tergites 4–6. Hind tibia with or without preapical spur.</paragraph>
<paragraph id="E77936F5FF84FD3B5E263027FB86B417" blockId="6.[824,1422,829,1849]" pageId="6" pageNumber="472">
Representatives of this genus are very similar to the closely related genus
<taxonomicName id="20C64D76FF84FD3B5EA03002FBACB550" authorityName="Schrank" authorityYear="1802" box="[984,1030,1072,1093]" class="Insecta" family="Argidae" genus="Arge" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="472" phylum="Arthropoda" rank="genus">
<emphasis id="D5B2EAE7FF84FD3B5EA03002FBACB550" box="[984,1030,1072,1093]" italics="true" pageId="6" pageNumber="472">Arge</emphasis>
</taxonomicName>
. Furthermore, whereas the males have a pronounced process of the 7
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tergite, which makes it possible to reliably place them in
<taxonomicName id="20C64D76FF84FD3B59523054FB2AB56E" authorityName="Wei" authorityYear="1998" box="[1066,1152,1126,1147]" class="Insecta" family="Argidae" genus="Spinarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="472" phylum="Arthropoda" rank="genus">
<emphasis id="D5B2EAE7FF84FD3B59523054FB2AB56E" box="[1066,1152,1126,1147]" italics="true" pageId="6" pageNumber="472">Spinarge</emphasis>
</taxonomicName>
, in females the corresponding character is very weak and in some cases can be confused with folds and deformations of the abdomen caused by drying. In collections, females significantly predominate, while males are rare, which in our experience leads to a large number of erroneous identifications.
</paragraph>
<paragraph id="E77936F5FF84FD3B5E26313AFBAFB73E" blockId="6.[824,1422,829,1849]" pageId="6" pageNumber="472">
The genus is distributed in the Palaearctic and Oriental regions. Number of species: World — 13; Palaearctic — 10. Since the publication of the world catalogue of sawflies [
<bibRefCitation id="83574B04FF84FD3B5818310CFC64B47B" author="Taeger A. &amp; Blank S. M. &amp; Liston A. D." pageId="6" pageNumber="472" pagination="1 - 1064" refId="ref11488" refString="Taeger A., Blank S. M., Liston A. D. 2010. World Catalog of Symphyta (Hymenoptera) // Zootaxa. Vol. 2580. P. 1 - 1064." type="book chapter" year="2010">
Taeger
<emphasis id="D5B2EAE7FF84FD3B5E25316BFC23B47B" box="[861,905,1369,1390]" italics="true" pageId="6" pageNumber="472">et al</emphasis>
., 2010
</bibRefCitation>
] two new species have been described [
<bibRefCitation id="83574B04FF84FD3B5829316BFC3BB49C" author="Liu M. &amp; Li Z. &amp; Wei M." pageId="6" pageNumber="472" pagination="230 - 236" refId="ref10327" refString="Liu M., Li Z., Wei M. 2021. A new species of Spinarge Wei (Hymenoptera, Argidae) from China with a key to Chinese species // Entomotaxonomia. Vol. 43. No. 3. P. 230 - 236." type="book chapter" year="2021">
Liu
<emphasis id="D5B2EAE7FF84FD3B5804316BFCE1B49C" italics="true" pageId="6" pageNumber="472">et al</emphasis>
., 2021
</bibRefCitation>
;
<bibRefCitation id="83574B04FF84FD3B5EE33146FB96B49C" author="Vasilenko S. V." box="[923,1084,1396,1417]" pageId="6" pageNumber="472" pagination="1213 - 1217" refId="ref11602" refString="Vasilenko S. V. 2022. A New Species of the Sawfly Genus Spinarge Wei, 1998 (Hymenoptera, Argidae) from the Dzhungarian Ala Tau Mountains, Kazakhstan // Entomological Review. Vol. 102. No. 8. P. 1213 - 1217." type="journal article" year="2022">Vasilenko, 2022</bibRefCitation>
]. New data on the distribution and biology of members of the genus were also published [
<bibRefCitation id="83574B04FF84FD3B583331BDFC75B4AA" author="Shinohara A. &amp; Hara H." pageId="6" pageNumber="472" pagination="389 - 403" refId="ref10899" refString="Shinohara A., Hara H. 2012. Arge cerasus n. sp. (Hymenoptera, Argidae) Feeding on Cerasus spp. in Japan // Japanese journal of systematic entomology. Vol. 18. No. 2. P. 389 - 403." type="book chapter" year="2012">Shinohara, Hara, 2012</bibRefCitation>
;
<bibRefCitation id="83574B04FF84FD3B5E933198FB63B4AA" author="Shinohara A. &amp; Kiyoshi T. &amp; Kameda Y." box="[1003,1225,1450,1471]" pageId="6" pageNumber="472" pagination="81 - 85" refId="ref11154" refString="Shinohara A., Kiyoshi T., Kameda Y. 2016. New distribution records of two species of Arge and Spinarge (Hymenoptera, Argidae) from Kyushu, Japan // Bulletin of the National Museum of Nature and Science, Series A. Vol. 42. P. 81 - 85." type="journal article" year="2016">
Shinohara
<emphasis id="D5B2EAE7FF84FD3B592C3198FB2AB4AA" box="[1108,1152,1450,1471]" italics="true" pageId="6" pageNumber="472">et al</emphasis>
., 2016
</bibRefCitation>
].
<taxonomicName id="20C64D76FF84FD3B59A23198FC61B4CF" authority="Wei, 2010" authorityName="Wei" authorityYear="2010" class="Insecta" family="Argidae" genus="Spinarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="472" phylum="Arthropoda" rank="species" species="acutithecata">
<emphasis id="D5B2EAE7FF84FD3B59A23198FCCBB4CF" italics="true" pageId="6" pageNumber="472">Spinarge acutithecata</emphasis>
<bibRefCitation id="83574B04FF84FD3B5E1031F7FC61B4CF" author="Wei M. &amp; Niu G." box="[872,971,1477,1498]" pageId="6" pageNumber="472" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei, 2010</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="20C64D76FF84FD3B5E8331F7FAF3B4CF" authority="Wei, 2010" authorityName="Wei" authorityYear="2010" box="[1019,1369,1477,1498]" class="Insecta" family="Argidae" genus="Spinarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="472" phylum="Arthropoda" rank="species" species="xanthofemorata">
<emphasis id="D5B2EAE7FF84FD3B5E8331F7FB5AB4CF" box="[1019,1264,1477,1498]" italics="true" pageId="6" pageNumber="472">Spinarge xanthofemorata</emphasis>
<bibRefCitation id="83574B04FF84FD3B598E31F7FAF3B4CF" author="Wei M. &amp; Niu G." box="[1270,1369,1477,1498]" pageId="6" pageNumber="472" pagination="337 - 362" refId="ref11972" refString="Wei M., Niu G. 2010. [Hymenoptera: Argidae, Cimbicidae, Tenthredinidae] // H. Xu, X. Ye (eds.). [Insects of Fengyangshan National Nature Reserve]. Beijing: China Forestry Press. P. 337 - 362 [in Chinese]." type="book chapter" year="2010">Wei, 2010</bibRefCitation>
</taxonomicName>
have been reported as new taxa [
<bibRefCitation id="83574B04FF84FD3B594631D2FB73B4E0" author="Wu X. - Y. &amp; Xin X. &amp; Zhang X. - Z." box="[1086,1241,1504,1525]" pageId="6" pageNumber="472" pagination="15 - 19" refId="ref12137" refString="Wu X. - Y., Xin X., Zhang X. - Z. 2010. [The Investigation and Taxonomical Research of the Sawflies Species from Gansu Province. III [recte: IV]. A list of the genera and species (Hymenoptera: Argidae).] // Journal of Gansu Forestry Science and Technology. Vol. 35. No. 1. P. 15 - 19 [in Chinese]." type="journal article" year="2010">Wu et al., 2010</bibRefCitation>
], but their descriptions have not been published to date. They should be considered as nomina nuda.
</paragraph>
<paragraph id="E77936F5FF84FD3B5E263203FAB3B62C" blockId="6.[824,1422,829,1849]" pageId="6" pageNumber="472">
In the latest key to
<taxonomicName id="20C64D76FF84FD3B595B3203FBDAB753" box="[1059,1136,1585,1606]" class="Insecta" family="Argidae" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="472" phylum="Arthropoda" rank="family">Argidae</taxonomicName>
genera for Europe [
<bibRefCitation id="83574B04FF84FD3B58423203FCC7B774" author="Lacourt J." pageId="6" pageNumber="472" refId="ref9958" refString="Lacourt J. 2020. Sawflies of Europe: Hymenoptera of Europe 2. Verrieres-le-Buisson: N. A. P. editions. 876 p." type="book" year="2020">Lacourt, 2020</bibRefCitation>
], the thesis “hind tibiae without preapical spurs” leads to the genus
<taxonomicName id="20C64D76FF84FD3B5ECB3255FBA3B769" authorityName="Wei" authorityYear="1998" box="[947,1033,1639,1660]" class="Insecta" family="Argidae" genus="Spinarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="472" phylum="Arthropoda" rank="genus">
<emphasis id="D5B2EAE7FF84FD3B5ECB3255FBA3B769" box="[947,1033,1639,1660]" italics="true" pageId="6" pageNumber="472">Spinarge</emphasis>
</taxonomicName>
. This is erroneous, because, as noted by
<bibRefCitation id="83574B04FF84FD3B5E4032B0FBEEB782" author="Hara H. &amp; Shinohara A." box="[824,1092,1666,1687]" pageId="6" pageNumber="472" pagination="61 - 94" refId="ref9066" refString="Hara H., Shinohara A. 2006. The sawfly genus Spinarge (Hymenoptera, Argidae) // Bulletin of the National Science Museum, Series A, Zoology. Vol. 32. No. 2. P. 61 - 94." type="journal article" year="2006">Hara and Shinohara [2006]</bibRefCitation>
, only two species actually do not have them, while the rest, including the only one inhabiting Europe,
<taxonomicName id="20C64D76FF84FD3B5EF6328AFB7CB7D8" authority="(Klug, 1834)" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[910,1238,1720,1741]" class="Insecta" family="Argidae" genus="Spinarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="472" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="D5B2EAE7FF84FD3B5EF6328AFBE2B7D8" box="[910,1096,1720,1741]" italics="true" pageId="6" pageNumber="472">Spinarge metallica</emphasis>
(Klug, 1834)
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, have such spurs. This makes it impossible to correctly determine the genus according to that key. In all existing keys for
<collectingCountry id="9FD17665FF84FD3B598332DCFA91B616" box="[1275,1339,1774,1795]" name="Russia" pageId="6" pageNumber="472">Russia</collectingCountry>
[
<bibRefCitation id="83574B04FF84FD3B583032DCFC14B60B" author="Gussakovskij V. V." pageId="6" pageNumber="472" refId="ref8958" refString="Gussakovskij V. V. 1935. [Chalastogastra (pt. 1)] // Fauna SSSR. Nov. ser. Nasekomye Pereponchatokrylye. Vol. 2. No. 2. Moscow-Leningrad: AN SSSR Publ. 453 p. [In Russian, with French summary]" type="book" year="1935">Gussakovskij, 1935</bibRefCitation>
;
<bibRefCitation id="83574B04FF84FD3B5EB1333BFB3FB60B" author="Zhelochovtsev A. N." box="[969,1173,1801,1822]" pageId="6" pageNumber="472" pagination="7 - 234" refId="ref12365" refString="Zhelochovtsev A. N. 1988. Suborder Symphyta (Chalastogastra) // A. N. Zhelochovtsev, V. I. Tobias, M. A. Kozlov (eds.). Opredelitel' nasekomykh Evropeiskoi chasti SSSR. Leningrad: Nauka. Vol. 3. No. 6. P. 7 - 234 [in Russian]." type="journal article" year="1988">Zhelochovtsev, 1988</bibRefCitation>
], the species of the genus
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<emphasis id="D5B2EAE7FF84FD3B5E403316FC24B62C" box="[824,910,1828,1849]" italics="true" pageId="6" pageNumber="472">Spinarge</emphasis>
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are considered as part of the genus
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<emphasis id="D5B2EAE7FF84FD3B599E3316FABEB62C" box="[1254,1300,1828,1849]" italics="true" pageId="6" pageNumber="472">Arge</emphasis>
</taxonomicName>
.
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species:
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Body size 5.0–7.0 mm. Labial palps with 4 segments, maxillary palps with 6 segments. Right mandible with basal tooth. Antennae of males bifurcate. Fore wings with 4 cells Rs; cell
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open; vein Sc absent; vein 2r–m present; vein B of fore wing joins vein
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well before (longer than vein Sc) junction point of vein M; cell 2A closed. Cell A of the hind wing widely open. For additional generic characters, see
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.
</paragraph>
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The genus is distributed in the Palaearctic (East), Nearctic, Neotropical and Australasian regions. Number of species: World — 5; Palaearctic — 1. In the Palaearctic, the genus is known only from
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, following synonymization of
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with
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[
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Blank
<emphasis id="D5B2EAE7FF84FD3B58773699FA90B3D5" box="[1295,1338,683,704]" italics="true" pageId="6" pageNumber="472">et al</emphasis>
., 2009
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<emphasis id="D5B2EAE7FF87FD385C1835DFFE7FB311" bold="true" box="[352,469,493,516]" italics="true" pageId="5" pageNumber="471">Kokujewia</emphasis>
Konow, 1902
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<figureCitation id="7FFD2A70FF87FD385CDA3639FD80B337" box="[418,554,523,546]" captionStart="Figs 2–15" captionStartId="4.[189,227,1814,1833]" captionTargetBox="[200,1409,368,1714]" captionTargetId="figure-4@4.[200,1410,360,1714]" captionTargetPageId="4" captionText="Figs 2–15. Argidae, details of morphology: 2, 10, 12 — Arge ustulata (Linné, 1758); 3, 11 — Aproceros leucopoda Takeuchi, 1939; 4, 8, 14 — Kokujewia ectrapela Konow, 1902; 5 — Spinarge metallica (Klug, 1834); 6 — Aprosthema melanurum (Klug, 1814); 7, 9 — Pseudarge rubicunda Gussakovskij, 1935; 13 — Sterictiphora geminata (Gmelin, 1790); 15 — Asiarge regeli Gussakovskij, 1935. 2–4 — veins R and Rs of fore wing, dorsally; 5–6 — male antenna, dorsally; 7–8 — hind tibia, ventrally; 9–11 — cells 1A and 2A of fore wing, dorsally; 12–13 — vein R+M of fore wing, dorsally; 14–15 — head, laterally. Рис. 2–15. Argidae, детали морфологии: 2, 10, 12 — Arge ustulata (Linné, 1758); 3, 11 — Aproceros leucopoda Takeuchi, 1939; 4, 8, 14 — Kokujewia ectrapela Konow, 1902; 5 — Spinarge metallica (Klug, 1834); 6 — Aprosthema melanurum (Klug, 1814); 7, 9 — Pseudarge rubicunda Gussakovskij, 1935; 13 — Sterictiphora geminata (Gmelin, 1790); 15 — Asiarge regeli Gussakovskij, 1935. 2–4 — жилки R и Rs переднего крыла, дорсально; 5–6 — антенна самца, дорсально; 7–8 — Задние голени, вентрально; 9–11 — Ячейки 1А и 2А переднего крыла, дорсально; 12–13 — жилка R+M переднего крыла, дорсально; 14–15 — голова, латерально." pageId="5" pageNumber="471">Figs 4, 8, 14</figureCitation>
.
</paragraph>
<paragraph id="E77936F5FF87FD385D9B3676FD6AB34C" blockId="5.[189,787,580,708]" box="[227,704,580,601]" pageId="5" pageNumber="471">
<typeStatus id="387D8857FF87FD385D9B3676FEB9B34C" box="[227,275,580,601]" pageId="5" pageNumber="471">Type</typeStatus>
species:
<taxonomicName id="20C64D76FF87FD385C123676FD16B34C" authority="Konow, 1902" authorityName="Konow" authorityYear="1902" box="[362,700,580,601]" class="Insecta" family="Argidae" genus="Kokujewia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="species" species="ectrapela">
<emphasis id="D5B2EAE7FF87FD385C123676FD98B34C" box="[362,562,580,601]" italics="true" pageId="5" pageNumber="471">Kokujewia ectrapela</emphasis>
Konow, 1902
</taxonomicName>
.
</paragraph>
<paragraph id="E77936F5FF87FD385D9B366DFCB9B3D1" blockId="5.[189,787,580,708]" pageId="5" pageNumber="471">Body size 10.0–12.0 mm. Fore wings with 3 cells Rs; cell R closed; vein Sc present; vein 2r–m absent; veins B and M join up at same point on vein R; cell 2A cell 2A absent or open, vein A2+A3 barely distinguishable, straight, without clear apex. Cell</paragraph>
<paragraph id="E77936F5FF87FD385E40351DFACEB04A" blockId="5.[824,1422,303,511]" pageId="5" pageNumber="471">
A of hind wing closed. All tibiae without preapical spurs. For additional generic characters, see
<bibRefCitation id="83574B04FF87FD38590C3578FACAB04A" author="Blank S. M. &amp; Taeger A." box="[1140,1376,330,351]" pageId="5" pageNumber="471" pagination="505 - 515" refId="ref7956" refString="Blank S. M., Taeger A. 1998. Revision of the sawfly genera Asiarge Gussakovskij, 1935 and Kokujewia Konow, 1902 // Beitrage zu Entomologie. Vol. 48. No. 2. P. 505 - 515." type="journal article" year="1998">Blank and Taeger [1998]</bibRefCitation>
.
</paragraph>
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The genus is distributed in the Palaearctic (West) region. Number of species: World and Palaearctic — 3. A small genus, whose representatives are mainly distributed in West Asia. Biology and host specificity of
<taxonomicName id="20C64D76FF87FD3859083586FCC2B0F1" authority="Konow, 1902" authorityName="Konow" authorityYear="1902" class="Insecta" family="Argidae" genus="Kokujewia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="species" species="ectrapela">
<emphasis id="D5B2EAE7FF87FD3859083586FA90B0DC" box="[1136,1338,436,457]" italics="true" pageId="5" pageNumber="471">Kokujewia ectrapela</emphasis>
Konow, 1902
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was treated in
<bibRefCitation id="83574B04FF87FD38597C35FDFB13B0F1" author="Karimpour Y." box="[1028,1209,463,484]" pageId="5" pageNumber="471" pagination="1 - 10" refId="ref9526" refString="Karimpour Y. 2015. Biology, host specificity and feeding potential of the Dock's leaf defoliator sawfly, Kokujewia ectrapela Konow (Hymenoptera: Argidae), a biocontrol agent of Rumex spp. (Polygonaceae) // Journal of Entomological Society of Iran. Vol. 35. No. 1. P. 1 - 10." type="journal article" year="2015">Karimpour [2015]</bibRefCitation>
. For an identification key to the three known species, see
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.
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<emphasis id="D5B2EAE7FF87FD385E913611FBCDB32F" bold="true" box="[1001,1127,547,570]" italics="true" pageId="5" pageNumber="471">Ortasiceros</emphasis>
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species:
<taxonomicName id="20C64D76FF87FD385E9D366EFAAFB364" authority="Wei, 1997" authorityName="Wei" authorityYear="1997" box="[997,1285,604,625]" class="Insecta" family="Argidae" genus="Ortasiceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="species" species="zhengi">
<emphasis id="D5B2EAE7FF87FD385E9D366EFB31B364" box="[997,1179,604,625]" italics="true" pageId="5" pageNumber="471">Ortasiceros zhengi</emphasis>
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.
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Body size 4.2–6.1 mm. Right mandible simple, without basal tooth. Labial palps with 3 segments, maxillary palps with 5 segments. Antennae of males bifurcate. Fore wings with 4 cells Rs; cell
<collectionCode id="81D7AE30FF84FD3B5D9A351DFF58B051" box="[226,242,303,324]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="472">R</collectionCode>
open; vein Sc present; vein B of fore wing joins vein
<collectionCode id="81D7AE30FF84FD3B5FAE351DFD4CB051" box="[726,742,303,324]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="472">R</collectionCode>
well before (longer than vein Sc) junction point of vein M; cell 2A of fore wing obliterate. Cell A of hind wing widely open. Cenchri very large. Fore tibial spurs about two thirds the length of fore basitarsus; claw simple. For additional generic characters, see
<bibRefCitation id="83574B04FF84FD3B5FFA35A9FD40B0A5" author="Wei M." box="[642,746,411,432]" pageId="6" pageNumber="472" pagination="295 - 305" refId="ref11819" refString="Wei M. 1997. Taxonomical studies on Argidae (Hymenoptera) of China - A new genus and five new species of sterictiphorinae // Insect Science. Vol. 4. No. 4. P. 295 - 305." type="journal article" year="1997">Wei [1997]</bibRefCitation>
.
</paragraph>
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<emphasis id="D5B2EAE7FF87FD385DC53301FE88B653" bold="true" box="[189,290,1843,1862]" pageId="5" pageNumber="471">Figs 16–24.</emphasis>
<taxonomicName id="20C64D76FF87FD385C5E3301FEC7B653" box="[294,365,1843,1862]" class="Insecta" family="Argidae" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="family">Argidae</taxonomicName>
, details of morphology: 16, 24, 25 —
<taxonomicName id="20C64D76FF87FD385FD53301FBB3B653" authority="(W. F. Kirby, 1882)" baseAuthorityName="W. F. Kirby" baseAuthorityYear="1882" box="[685,1049,1843,1862]" class="Insecta" genus="Tanyphatnidea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="D5B2EAE7FF87FD385FD53301FCD9B653" box="[685,883,1843,1862]" italics="true" pageId="5" pageNumber="471">Tanyphatnidea sinensis</emphasis>
(W.F. Kirby, 1882)
</taxonomicName>
; 17, 18 —
<taxonomicName id="20C64D76FF87FD38590D3301FA23B653" authority="(Klug, 1834)" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[1141,1417,1843,1862]" class="Insecta" family="Argidae" genus="Spinarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="D5B2EAE7FF87FD38590D3301FABFB653" box="[1141,1301,1843,1862]" italics="true" pageId="5" pageNumber="471">Spinarge metallica</emphasis>
(Klug, 1834)
</taxonomicName>
; 19, 20, 21, 26 —
<taxonomicName id="20C64D76FF87FD385C24337CFD08B674" authority="Takeuchi, 1939" authorityName="Takeuchi" authorityYear="1939" box="[348,674,1870,1889]" class="Insecta" family="Argidae" genus="Aproceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="species" species="leucopoda">
<emphasis id="D5B2EAE7FF87FD385C24337CFDB9B674" box="[348,531,1870,1889]" italics="true" pageId="5" pageNumber="471">Aproceros leucopoda</emphasis>
Takeuchi, 1939
</taxonomicName>
; 22 —
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<emphasis id="D5B2EAE7FF87FD385F9E337CFC07B674" box="[742,941,1870,1889]" italics="true" pageId="5" pageNumber="471">Sterictiphora geminata</emphasis>
(Gmelin, 1790)
</taxonomicName>
; 23 —
<taxonomicName id="20C64D76FF87FD3859F8337CFF5BB66E" authority="(Klug, 1814)" baseAuthorityName="Klug" baseAuthorityYear="1814" class="Insecta" family="Argidae" genus="Aprosthema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="D5B2EAE7FF87FD3859F8337CFAE5B674" box="[1152,1359,1870,1889]" italics="true" pageId="5" pageNumber="471">Aprosthema melanurum</emphasis>
(Klug, 1814)
</taxonomicName>
; 16 — head and thorax, dorsally; 17–18 — medial tergites of abdomen, dorsally; 19 — labrum and right mandible, dorsally; 20 — claw, laterally; 21 — right mandible, dorsally; 22, 23, 26 — cell A of hind wing, dorsally; 24 — saw sheath, dorsally; 25 — saw sheath, laterally.
</paragraph>
<paragraph id="E77936F5FF87FD385DC533ACFD27B90E" blockId="5.[189,1422,1843,2075]" pageId="5" pageNumber="471">
<emphasis id="D5B2EAE7FF87FD385DC533ACFE8FB6A4" bold="true" box="[189,293,1950,1969]" pageId="5" pageNumber="471">Рис. 16–26.</emphasis>
<taxonomicName id="20C64D76FF87FD385C5133ACFEDAB6A4" box="[297,368,1950,1969]" class="Insecta" family="Argidae" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="family">Argidae</taxonomicName>
, детали морфологии: 16, 24, 25 —
<taxonomicName id="20C64D76FF87FD385FD133ACFBBCB6A4" authority="(W. F. Kirby, 1882)" baseAuthorityName="W. F. Kirby" baseAuthorityYear="1882" box="[681,1046,1950,1969]" class="Insecta" genus="Tanyphatnidea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="D5B2EAE7FF87FD385FD133ACFCDAB6A4" box="[681,880,1950,1969]" italics="true" pageId="5" pageNumber="471">Tanyphatnidea sinensis</emphasis>
(W.F. Kirby, 1882)
</taxonomicName>
; 17, 18 —
<taxonomicName id="20C64D76FF87FD38590C33ACFA23B6A4" authority="(Klug, 1834)" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[1140,1417,1950,1969]" class="Insecta" family="Argidae" genus="Spinarge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="D5B2EAE7FF87FD38590C33ACFABEB6A4" box="[1140,1300,1950,1969]" italics="true" pageId="5" pageNumber="471">Spinarge metallica</emphasis>
(Klug, 1834)
</taxonomicName>
; 19, 20, 21, 26 —
<taxonomicName id="20C64D76FF87FD385C24338AFD08B6DE" authority="Takeuchi, 1939" authorityName="Takeuchi" authorityYear="1939" box="[348,674,1976,1995]" class="Insecta" family="Argidae" genus="Aproceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="species" species="leucopoda">
<emphasis id="D5B2EAE7FF87FD385C24338AFDB9B6DE" box="[348,531,1976,1995]" italics="true" pageId="5" pageNumber="471">Aproceros leucopoda</emphasis>
Takeuchi, 1939
</taxonomicName>
; 22 —
<taxonomicName id="20C64D76FF87FD385F9E338AFB96B6DE" authority="(Gmelin, 1790)" baseAuthorityName="Gmelin" baseAuthorityYear="1790" box="[742,1084,1976,1995]" class="Insecta" family="Argidae" genus="Sterictiphora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="species" species="geminata">
<emphasis id="D5B2EAE7FF87FD385F9E338AFC07B6DE" box="[742,941,1976,1995]" italics="true" pageId="5" pageNumber="471">Sterictiphora geminata</emphasis>
(Gmelin, 1790)
</taxonomicName>
; 23 —
<taxonomicName id="20C64D76FF87FD3859F8338AFF5BB6F3" authority="(Klug, 1814)" baseAuthorityName="Klug" baseAuthorityYear="1814" class="Insecta" family="Argidae" genus="Aprosthema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="471" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="D5B2EAE7FF87FD3859F8338AFAE5B6DE" box="[1152,1359,1976,1995]" italics="true" pageId="5" pageNumber="471">Aprosthema melanurum</emphasis>
(Klug, 1814)
</taxonomicName>
; 16 — голова и грудь, дорсально; 17–18 — медиальные тергиты брюшка, дорсально; 19 — нижнЯЯ губа и праваЯ мандибула, дорсально; 20 — коготок, латерально; 21 — праваЯ мандибула, дорсально; 22, 23, 26 — Ячейка A Заднего крыла, дорсально; 24 — ножны пилки, дорсально; 25 — ножны пилки, латерально.
</paragraph>
</caption>
<paragraph id="E77936F5FF84FD3B5D9B3584FDB6B314" blockId="6.[189,788,303,513]" pageId="6" pageNumber="472">
The genus is distributed in the Palaearctic (East) region. Number of species: World and Palaearctic — 6. For a key to
<taxonomicName id="20C64D76FF84FD3B5DC535DEFED1B314" box="[189,379,492,513]" pageId="6" pageNumber="472">
<emphasis id="D5B2EAE7FF84FD3B5DC535DEFE87B314" box="[189,301,492,513]" italics="true" pageId="6" pageNumber="472">Ortasiceros</emphasis>
species
</taxonomicName>
, see
<bibRefCitation id="83574B04FF84FD3B5CD135DEFDB2B314" author="Wei M." box="[425,536,492,513]" pageId="6" pageNumber="472" pagination="295 - 305" refId="ref11819" refString="Wei M. 1997. Taxonomical studies on Argidae (Hymenoptera) of China - A new genus and five new species of sterictiphorinae // Insect Science. Vol. 4. No. 4. P. 295 - 305." type="journal article" year="1997">Wei [1997]</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>