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<mods:title id="D5307007EE28B1BBBCE016C2C698D0BF">A revision of Anadolua (Orthoptera: Tettigoniidae: Tettigoniinae) based on morphological and bioacoustic characters: Newly confirmed data suggesting a new synonym and two new species</mods:title>
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<emphasis id="1AD7A37DFFC3814435FDFB18FE80FB58" bold="true" box="[151,348,1230,1257]" italics="true" pageId="32" pageNumber="233">Anadolua davisi</emphasis>
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<paragraph id="281C7F6FFFC3814435FDFB24FCDFFABC" blockId="32.[151,771,1230,1293]" box="[151,771,1266,1293]" pageId="32" pageNumber="233">https://orthoptera.speciesfile.org/otus/804975/overview</paragraph>
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<emphasis id="1AD7A37DFFC3814435FDFAE1FEE1FAFE" box="[151,317,1335,1359]" italics="true" pageId="32" pageNumber="233">Anadolua davisi</emphasis>
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:
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<bibRefCitation id="4C32029EFFC381443420FAE1FE07FAFE" author="Karabag, T." box="[330,475,1335,1359]" pageId="32" pageNumber="233" pagination="139 - 143" refId="ref25717" refString="Karabag, T. (1952) Anadolua Ramme (Orthoptera, Tettigoniidae) genusunun revizyonu. Istanbul Universitesi Fen Fakultesi Mecmuasi, 17 (2), 139 - 143. [in Turkish]" type="journal article" year="1952">Karabağ 1952</bibRefCitation>
, p.143
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;
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, p.53
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;
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<bibRefCitation id="4C32029EFFC38144366BFAEFFCB2FAFE" author="Unal, M." box="[769,878,1335,1362]" pageId="32" pageNumber="233" pagination="1 - 66" refId="ref27801" refString="Unal, M. (2018) Tettigoniinae (Orthoptera: Tettigoniidae) from Turkey with key to genera and descriptions of six new species. Zootaxa, 4432 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4432.1.1" type="journal article" year="2018">Ünal 2018</bibRefCitation>
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:
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(
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) (
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)
</materialsCitation>
.
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: Male
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<paragraph id="281C7F6FFFC3814435ADFA3FFD28F996" blockId="32.[151,1437,1405,2008]" pageId="32" pageNumber="233">
<emphasis id="1AD7A37DFFC3814435ADFA3FFE85F9B2" box="[199,345,1513,1539]" italics="true" pageId="32" pageNumber="233">Distribution:</emphasis>
The
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is an Anatolian endemic species and is known only from
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, ÖdemiŞ Bozdağ Mountain (
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;
<bibRefCitation id="4C32029EFFC3814434ABF9DBFDBEF999" author="Karabag, T." box="[449,610,1549,1576]" pageId="32" pageNumber="233" refId="ref25753" refString="Karabag, T. (1958). Turkiye'nin Orthoptera faunasi. Sirketi Murettebiye Basimevi, Istanbul, 192 pp. [in Turkish]" type="book" year="1958">Karabağ 1958</bibRefCitation>
;
<bibRefCitation id="4C32029EFFC381443704F9D9FD34F996" author="Unal, M." box="[622,744,1549,1578]" pageId="32" pageNumber="233" pagination="1 - 66" refId="ref27801" refString="Unal, M. (2018) Tettigoniinae (Orthoptera: Tettigoniidae) from Turkey with key to genera and descriptions of six new species. Zootaxa, 4432 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4432.1.1" type="journal article" year="2018">Ünal 2018</bibRefCitation>
).
</paragraph>
<paragraph id="281C7F6FFFC3814435ADF9E7FB1FF922" blockId="32.[151,1437,1405,2008]" pageId="32" pageNumber="233">
<emphasis id="1AD7A37DFFC3814435ADF9E7FE74F9FA" box="[199,424,1585,1611]" italics="true" pageId="32" pageNumber="233">Material examined.</emphasis>
<materialsCitation id="98CB7532FFC3814434D8F9E7FB63F922" collectingDate="1950-08-16" collectionCode="NHMUK" collectorName="P. H. Davis &amp; Beulah Garner" country="Turkey" county="Tmolus" elevation="1450" location="Lydia" municipality="Bozdag" pageId="32" pageNumber="233" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="50B43FFFFFC3814434D8F9E7FDCBF9FA" box="[434,535,1585,1611]" name="Turkey" pageId="32" pageNumber="233">Anatolia</collectingCountry>
: OdemiŞ Prov. (
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)
<collectingMunicipality id="C878E515FFC381443644F9E7FC58F9FA" box="[814,900,1585,1611]" pageId="32" pageNumber="233">Bozdağ</collectingMunicipality>
(
<collectingCounty id="C17D07E3FFC3814436FDF9E7FC2CF9FA" box="[919,1008,1585,1611]" pageId="32" pageNumber="233">Tmolus</collectingCounty>
)
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.,
<date id="5C1D59AFFFC3814431DBF9E7FAE7F9FD" box="[1201,1339,1585,1612]" pageId="32" pageNumber="233" value="1950-08-16">
<collectingDate id="4C59A047FFC3814431DBF9E7FAE7F9FD" box="[1201,1339,1585,1612]" pageId="32" pageNumber="233" value="1950-08-16">16.viii.1950</collectingDate>
</date>
,
<specimenCount id="3EA5B4E6FFC381443026F9E7FA40F9FA" box="[1356,1436,1585,1611]" count="1" pageId="32" pageNumber="233" type="male">1 male</specimenCount>
(
<typeStatus id="F718C1CDFFC3814435F5F983FED5F9DE" box="[159,265,1621,1647]" pageId="32" pageNumber="233" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="EFA304ECFFC381443447F980FE53F9DE" authorityName="Karabag" authorityYear="1952" box="[301,399,1621,1648]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="1AD7A37DFFC381443447F980FE53F9DE" box="[301,399,1621,1648]" italics="true" pageId="32" pageNumber="233">A. davisi</emphasis>
</taxonomicName>
),
<specimenCount id="3EA5B4E6FFC3814434CCF983FDDBF9DE" box="[422,519,1621,1647]" count="1" pageId="32" pageNumber="233" type="female">1 female</specimenCount>
(
<typeStatus id="F718C1CDFFC38144377FF983FDA5F9DE" box="[533,633,1621,1647]" pageId="32" pageNumber="233" type="paratype">Paratype</typeStatus>
of
<taxonomicName id="EFA304ECFFC3814437F4F980FD23F9DE" authorityName="Karabag" authorityYear="1952" box="[670,767,1621,1648]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="1AD7A37DFFC3814437F4F980FD23F9DE" box="[670,767,1621,1648]" italics="true" pageId="32" pageNumber="233">A. davisi</emphasis>
</taxonomicName>
) (leg.:
<collectorName id="85561AB9FFC38144363AF980FC15F9DE" box="[848,969,1621,1648]" pageId="32" pageNumber="233">P.H. Davis</collectorName>
) (
<collectionCode id="4EB2E7AAFFC3814436B4F983FB97F9DE" box="[990,1099,1621,1647]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="32" pageNumber="233" type="Museum">NHMUK</collectionCode>
) (detailed and scaled photo of these two samples were send to us for this research by scientist
<collectorName id="85561AB9FFC38144360AF9AFFBD8F922" box="[864,1028,1657,1683]" pageId="32" pageNumber="233">Beulah Garner</collectorName>
from
<collectionCode id="4EB2E7AAFFC381443122F9AFFB6BF922" box="[1096,1207,1657,1683]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="32" pageNumber="233" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="281C7F6FFFC3814435ADF94BFA4FF866" blockId="32.[151,1437,1405,2008]" pageId="32" pageNumber="233">
<emphasis id="1AD7A37DFFC3814435ADF94BFE40F906" box="[199,412,1693,1720]" italics="true" pageId="32" pageNumber="233">
Remarks.
<taxonomicName id="EFA304ECFFC381443452F948FE40F906" authorityName="Karabag" authorityYear="1952" box="[312,412,1693,1720]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="davisi">A. davisi</taxonomicName>
</emphasis>
species was first described in
<bibRefCitation id="4C32029EFFC381443792F94BFC76F909" author="Karabag, T." box="[760,938,1693,1720]" pageId="32" pageNumber="233" pagination="139 - 143" refId="ref25717" refString="Karabag, T. (1952) Anadolua Ramme (Orthoptera, Tettigoniidae) genusunun revizyonu. Istanbul Universitesi Fen Fakultesi Mecmuasi, 17 (2), 139 - 143. [in Turkish]" type="journal article" year="1952">Karabağ (1952)</bibRefCitation>
and since then, it has not been found in the same locality by any other researcher. However, two other species belonging to the genus have been recorded. Although there are some records of
<taxonomicName id="EFA304ECFFC381443752F930FD61F94E" authorityName="Ramme" authorityYear="1939" box="[568,701,1765,1792]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFC381443752F930FD61F94E" box="[568,701,1765,1792]" italics="true" pageId="32" pageNumber="233">A. schwarzi</emphasis>
</taxonomicName>
and
<taxonomicName id="EFA304ECFFC381443792F930FC8FF94E" authorityName="Karabag" authorityYear="1958" box="[760,851,1765,1792]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="burri">
<emphasis id="1AD7A37DFFC381443792F930FC8FF94E" box="[760,851,1765,1792]" italics="true" pageId="32" pageNumber="233">A. burri</emphasis>
</taxonomicName>
from the
<typeStatus id="F718C1CDFFC3814436ADF930FC2BF8B1" box="[967,1015,1766,1792]" pageId="32" pageNumber="233">type</typeStatus>
locality of
<taxonomicName id="EFA304ECFFC381443115F930FB3FF94E" authorityName="Karabag" authorityYear="1952" box="[1151,1251,1765,1792]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="1AD7A37DFFC381443115F930FB3FF94E" box="[1151,1251,1765,1792]" italics="true" pageId="32" pageNumber="233">A. davisi</emphasis>
</taxonomicName>
in the literature (
<bibRefCitation id="4C32029EFFC3814435F5F8DFFEB4F892" author="Onder, F. &amp; Pehlivan, E. &amp; Karsavuran, Y. &amp; Serdar, T. &amp; Kismali, S." box="[159,360,1801,1827]" pageId="32" pageNumber="233" pagination="243 - 250" refId="ref26592" refString="Onder, F., Pehlivan, E., Karsavuran, Y., Serdar, T. &amp; Kismali, S., (1999) Catalogeu of collection of Tettigoniidae (Orthoptera) preserved in the Prof. Dr. Niyazi LODOS Museum, Izmir, Turkey. Turkish Journal of Entomology, 23 (4), 243 - 250." type="journal article" year="1999">
Onder
<emphasis id="1AD7A37DFFC381443584F8DCFEFAF892" box="[238,294,1801,1827]" italics="true" pageId="32" pageNumber="233">et al.</emphasis>
1999
</bibRefCitation>
;
<bibRefCitation id="4C32029EFFC38144341EF8DFFDA6F892" author="Tazegul, E. &amp; Onder, F." box="[372,634,1801,1830]" pageId="32" pageNumber="233" pagination="109 - 123" refId="ref27607" refString="Tazegul, E. &amp; Onder, F. (2012) Izmir ilinde bulunan Tettigoniidae (Orthoptera) familyasi turleri uzerinde sistematik araStirmalar. Turk Entomoloji Bulteni, 2 (2), 109 - 123." type="journal article" year="2012">Tazegül &amp; Önder 2012</bibRefCitation>
), there is no record of
<taxonomicName id="EFA304ECFFC38144361DF8DCFC06F892" authorityName="Karabag" authorityYear="1952" box="[887,986,1801,1828]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="1AD7A37DFFC38144361DF8DCFC06F892" box="[887,986,1801,1828]" italics="true" pageId="32" pageNumber="233">A. davisi</emphasis>
</taxonomicName>
. In addition,
<bibRefCitation id="4C32029EFFC38144311AF8DDFB25F892" author="Unal, M." box="[1136,1273,1801,1830]" pageId="32" pageNumber="233" pagination="1 - 66" refId="ref27801" refString="Unal, M. (2018) Tettigoniinae (Orthoptera: Tettigoniidae) from Turkey with key to genera and descriptions of six new species. Zootaxa, 4432 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4432.1.1" type="journal article" year="2018">Ünal (2018)</bibRefCitation>
stated that the specimens of
<taxonomicName id="EFA304ECFFC381443458F8F8FE57F8F6" authorityName="Karabag" authorityYear="1958" box="[306,395,1837,1864]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="burri">
<emphasis id="1AD7A37DFFC381443458F8F8FE57F8F6" box="[306,395,1837,1864]" italics="true" pageId="32" pageNumber="233">A. burri</emphasis>
</taxonomicName>
(
<specimenCount id="3EA5B4E6FFC3814434F0F8FBFDDEF8F6" box="[410,514,1837,1863]" count="33" pageId="32" pageNumber="233" type="male">33 males</specimenCount>
,
<specimenCount id="3EA5B4E6FFC381443764F8FBFD56F8F6" box="[526,650,1837,1863]" count="24" pageId="32" pageNumber="233" type="female">24 females</specimenCount>
) and
<taxonomicName id="EFA304ECFFC3814437A2F8F8FC97F8F6" authorityName="Ramme" authorityYear="1939" box="[712,843,1837,1864]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFC3814437A2F8F8FC97F8F6" box="[712,843,1837,1864]" italics="true" pageId="32" pageNumber="233">A. schwarzi</emphasis>
</taxonomicName>
(
<specimenCount id="3EA5B4E6FFC381443630F8FBFC1FF8F6" box="[858,963,1837,1863]" count="24" pageId="32" pageNumber="233" type="male">24 males</specimenCount>
,
<specimenCount id="3EA5B4E6FFC3814436A4F8FBFBE0F8F6" box="[974,1084,1837,1863]" count="9" pageId="32" pageNumber="233" type="female">9 females</specimenCount>
) collected by
<bibRefCitation id="4C32029EFFC3814431B0F8FBFF3EF8DA" author="Tazegul, E. &amp; Onder, F." pageId="32" pageNumber="233" pagination="109 - 123" refId="ref27607" refString="Tazegul, E. &amp; Onder, F. (2012) Izmir ilinde bulunan Tettigoniidae (Orthoptera) familyasi turleri uzerinde sistematik araStirmalar. Turk Entomoloji Bulteni, 2 (2), 109 - 123." type="journal article" year="2012">Tazegül &amp; Önder (2012)</bibRefCitation>
from ÖdemiŞ were only
<taxonomicName id="EFA304ECFFC38144376AF884FD59F8DA" authorityName="Ramme" authorityYear="1939" box="[512,645,1873,1900]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFC38144376AF884FD59F8DA" box="[512,645,1873,1900]" italics="true" pageId="32" pageNumber="233">A. schwarzi</emphasis>
</taxonomicName>
. In addition, no individuals of
<taxonomicName id="EFA304ECFFC38144368CF884FB95F8DA" authorityName="Karabag" authorityYear="1952" box="[998,1097,1873,1900]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="1AD7A37DFFC38144368CF884FB95F8DA" box="[998,1097,1873,1900]" italics="true" pageId="32" pageNumber="233">A. davisi</emphasis>
</taxonomicName>
were encountered in the field studies in ÖdemiŞ during this study (20212023). The male cercus character used in the description by Karabağ is an important distinctive character for the genus and it would be appropriate to continue the species status of the species until fresh materials are found and their songs are recorded and/or DNA-based up-to-date analyses are performed.
</paragraph>
<paragraph id="281C7F6FFFC2814535ADFF41FE1FFF00" blockId="33.[151,1437,151,285]" box="[199,451,151,177]" pageId="33" pageNumber="234">
<emphasis id="1AD7A37DFFC2814535ADFF41FE60FF00" box="[199,444,151,177]" italics="true" pageId="33" pageNumber="234">
<collectionCode id="4EB2E7AAFFC2814535ADFF41FED6FF00" box="[199,266,151,177]" pageId="33" pageNumber="234">IUCN</collectionCode>
status proposal
</emphasis>
.
</paragraph>
<paragraph id="281C7F6FFFC2814535ADFF6DFED6FEAC" blockId="33.[151,1437,151,285]" pageId="33" pageNumber="234">
This species has not been assessed for the
<collectionCode id="4EB2E7AAFFC2814537DEFF6DFD24FF64" box="[692,760,187,213]" pageId="33" pageNumber="234">IUCN</collectionCode>
Red List (2023-1). There is also not enough data about
<taxonomicName id="EFA304ECFFC2814530EEFF6DFF06FF48" authorityName="Karabag" authorityYear="1952" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="1AD7A37DFFC2814530EEFF6DFF06FF48" italics="true" pageId="33" pageNumber="234">A. davisi</emphasis>
</taxonomicName>
, and it is not possible to consider it on the basis of the extent of occurrence criteria of
<collectionCode id="4EB2E7AAFFC2814531BEFF09FAC4FF48" box="[1236,1304,223,249]" pageId="33" pageNumber="234">IUCN</collectionCode>
(DD =data deficient).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,112 @@
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<mods:title id="2878E6240E82F8B1A0C096905E1FF4B6">A revision of Anadolua (Orthoptera: Tettigoniidae: Tettigoniinae) based on morphological and bioacoustic characters: Newly confirmed data suggesting a new synonym and two new species</mods:title>
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Anadolua
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,
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. III. Mitteilungen aus dem Zoologischen Museum in
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, 24:41150.
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species:
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<emphasis id="1AD7A37DFFF78170344CF85DFE35F812" box="[294,489,1931,1955]" italics="true" pageId="20" pageNumber="221">Anadolua schwarzi</emphasis>
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View file

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Şirin, Sevgili and Taylan
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<emphasis id="1AD7A37DFFF8817F35FDFEF4FEF5FE8D" box="[151,297,290,316]" italics="true" pageId="27" pageNumber="228">Distribution:</emphasis>
The specimens belonging to the populations of the new species have been collected until now only from the borders of
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provinces (
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(deposited in alcohol,
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pass,
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,
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<elevation id="A38E985CFFF8817F34DDFE58FDD7FE18" box="[439,523,398,425]" metricMagnitude="3" metricUnit="m" metricValue="1.825" pageId="27" pageNumber="228" unit="m" value="1825.0">1825 m</elevation>
</quantity>
N.N.,
<date id="5C1D59AFFFF8817F373FFE58FD36FE19" box="[597,746,398,424]" pageId="27" pageNumber="228" value="2022-08-07">
<collectingDate id="4C59A047FFF8817F373FFE58FD36FE19" box="[597,746,398,424]" pageId="27" pageNumber="228" value="2022-08-07">07.VIII.2022</collectingDate>
</date>
,
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(including
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) and
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, leg.:
<collectorName id="85561AB9FFF8817F3071FE59FAAAFE19" box="[1307,1398,398,425]" pageId="27" pageNumber="228">D. Sirin</collectorName>
, H. Sevgili &amp;
<collectorName id="85561AB9FFF8817F3461FE64FE49FE7D" box="[267,405,434,460]" pageId="27" pageNumber="228">M.S. Taylan</collectorName>
-
<collectingRegion id="EA67B18DFFF8817F34C1FE64FDDBFE7D" box="[427,519,434,460]" country="Turkey" name="Antalya" pageId="27" pageNumber="228">Antalya</collectingRegion>
, GündoğmuŞ-Susambeli road,
<geoCoordinate id="4D9719A8FFF8817F360AFE65FC2AFE7D" box="[864,1014,434,461]" degrees="36.906202" direction="north" orientation="latitude" pageId="27" pageNumber="228" precision="1" value="36.9062">N 36.906202</geoCoordinate>
,
<geoCoordinate id="4D9719A8FFF8817F316BFE65FB48FE7C" box="[1025,1172,434,461]" degrees="32.076154" direction="east" orientation="longitude" pageId="27" pageNumber="228" precision="1" value="32.076153">E 32.076154</geoCoordinate>
,
<quantity id="EF5BD28AFFF8817F31F5FE64FB2BFE7C" box="[1183,1271,434,461]" metricMagnitude="3" metricUnit="m" metricValue="1.9369999999999998" pageId="27" pageNumber="228" unit="m" value="1937.0">
<elevation id="A38E985CFFF8817F31F5FE64FB2BFE7C" box="[1183,1271,434,461]" metricMagnitude="3" metricUnit="m" metricValue="1.9369999999999998" pageId="27" pageNumber="228" unit="m" value="1937.0">1937 m</elevation>
</quantity>
,
<date id="5C1D59AFFFF8817F3069FE64FA44FE7D" box="[1283,1432,434,460]" pageId="27" pageNumber="228" value="2022-08-06">
<collectingDate id="4C59A047FFF8817F3069FE64FA44FE7D" box="[1283,1432,434,460]" pageId="27" pageNumber="228" value="2022-08-06">06.VIII.2022</collectingDate>
</date>
,
<specimenCount id="3EA5B4E6FFF8817F35FDFE00FF39FE41" box="[151,229,470,496]" count="1" pageId="27" pageNumber="228" type="male">1 male</specimenCount>
, leg.:
<collectorName id="85561AB9FFF8817F3443FE01FE58FE41" box="[297,388,470,497]" pageId="27" pageNumber="228">D. Sirin</collectorName>
, H. Sevgili &amp;
<collectorName id="85561AB9FFF8817F374FFE00FD69FE41" box="[549,693,470,496]" pageId="27" pageNumber="228">M.S. Taylan.</collectorName>
</materialsCitation>
</paragraph>
<caption id="7CDC2FE7FFF8817F35FDFB2AFD5CFAEC" ID-DOI="http://doi.org/10.5281/zenodo.14022306" ID-Zenodo-Dep="14022306" httpUri="https://zenodo.org/record/14022306/files/figure.png" pageId="27" pageNumber="228" startId="27.[151,250,1276,1300]" targetBox="[164,1422,529,1237]" targetPageId="27" targetType="figure">
<paragraph id="281C7F6FFFF8817F35FDFB2AFD5CFAEC" blockId="27.[151,1436,1276,1373]" pageId="27" pageNumber="228">
<emphasis id="1AD7A37DFFF8817F35FDFB2AFEC3FAA5" bold="true" box="[151,287,1276,1300]" pageId="27" pageNumber="228">FIGURE 18.</emphasis>
Distribution map (A) and male calling song of
<taxonomicName id="EFA304ECFFF8817F3797FB2BFC73FAA4" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[765,943,1277,1301]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="bergeri" status="sp. nov.">
<emphasis id="1AD7A37DFFF8817F3797FB2BFC73FAA4" box="[765,943,1277,1301]" italics="true" pageId="27" pageNumber="228">Anadolua bergeri</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFF8817F36DFFB2BFBD9FAA4" bold="true" box="[949,1029,1277,1301]" pageId="27" pageNumber="228">
<taxonomicNameLabel id="01E41E06FFF8817F36DFFB2BFBD9FAA4" box="[949,1029,1277,1301]" pageId="27" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(B—two consecutive complete phrases, C—a complete phrase and D—two syllables, black arrow is the hypothetic opening hemisyllable movement and grey arrow is the hypothetic closing hemisyllable movement).
</paragraph>
</caption>
<paragraph id="281C7F6FFFF8817835ADFA66FA99FE60" blockId="27.[151,1437,1455,2022]" lastBlockId="28.[151,1437,151,465]" lastPageId="28" lastPageNumber="229" pageId="27" pageNumber="228">
<emphasis id="1AD7A37DFFF8817F35ADFA66FE8FFA78" box="[199,339,1456,1481]" italics="true" pageId="27" pageNumber="228">Description.</emphasis>
Head equal width with pronotum in both sexes. Fastigium mostly wider than scapus in both sexes. Maximum width of fastigium / Maximum width of scapus
<quantity id="EF5BD28AFFF8817F362DFA05FC0BFA5C" box="[839,983,1491,1518]" metricMagnitude="-2" metricUnit="m" metricValue="2.9844999999999997" metricValueMax="3.5559999999999996" metricValueMin="2.413" pageId="27" pageNumber="228" unit="in" value="1.1749999999999998" valueMax="1.4" valueMin="0.95">0.951.40 in</quantity>
male,
<quantity id="EF5BD28AFFF8817F3143FA05FB65FA5C" box="[1065,1209,1491,1517]" metricMagnitude="-2" metricUnit="m" metricValue="2.8575" metricValueMax="2.8701999999999996" metricValueMin="2.8448" pageId="27" pageNumber="228" unit="in" value="1.125" valueMax="1.13" valueMin="1.12">1.121.13 in</quantity>
female. Dark color between the eyes but not in the form of a band. The dorsal pronotum has a weakly hourglass shape and is weakly flattened in the prozona but distinctly flattened in the metazona, and has a deep transverse depression behind the middle, and its posterior edge is narrowly rounded. Maximum distance between lateral carinae / minimum distance between lateral carinae
<quantity id="EF5BD28AFFF8817F34F7F9B5FDF5F9CC" box="[413,553,1635,1661]" metricMagnitude="-2" metricUnit="m" metricValue="5.5245" metricValueMax="5.9182" metricValueMin="5.1308" pageId="27" pageNumber="228" unit="in" value="2.175" valueMax="2.33" valueMin="2.02">2.022.33 in</quantity>
male,
<quantity id="EF5BD28AFFF8817F3718F9B5FD22F9CC" box="[626,766,1635,1662]" metricMagnitude="-2" metricUnit="m" metricValue="6.7056" metricValueMax="7.0104" metricValueMin="6.4008" pageId="27" pageNumber="228" unit="in" value="2.6399999999999997" valueMax="2.76" valueMin="2.52">2.522.76 in</quantity>
female. The median carinae is absent, and the lateral carinae are prominent as round in the second part of the metazona. Hind femur long, its length 4.04.5 times of its maximum width in male, 4.0
<quantity id="EF5BD28AFFF8817F341DF97DFE6BF974" box="[375,439,1707,1733]" metricMagnitude="-1" metricUnit="m" metricValue="1.0413999999999999" pageId="27" pageNumber="228" unit="in" value="4.1">4.1 in</quantity>
female. Male tegmina macropterous, reaching the end of second abdominal tergite, the female tegmina is squamipterous with only the tip protruding. The speculum is in quadriform shape (generally trapezoidal), and lateral vein development is usually at the apical corner of the speculum. Costal vein very weak; subcostal vein reach to the apex, generally parallel to radial vein, and rarely, they may merge with a small vein near the apex. Medial vein sinuate at the apex and usually with a dichotomy at the ¾ from the base. The male stridulatory file bears fewer than 50 teeth and the space between the teeth is wider than the
<taxonomicName id="EFA304ECFFF8817F3142F889FB71F8C8" authorityName="Ramme" authorityYear="1939" box="[1064,1197,1887,1913]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFF8817F3142F889FB71F8C8" box="[1064,1197,1887,1913]" italics="true" pageId="27" pageNumber="228">A. schwarzi</emphasis>
</taxonomicName>
and
<taxonomicName id="EFA304ECFFF8817F318CF889FAE5F8C8" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[1254,1337,1887,1913]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="moli" status="sp. nov.">
<emphasis id="1AD7A37DFFF8817F318CF889FAE5F8C8" box="[1254,1337,1887,1913]" italics="true" pageId="27" pageNumber="228">A. moli</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFF8817F302BF889FA40F8C8" bold="true" box="[1345,1436,1887,1913]" pageId="27" pageNumber="228">
<taxonomicNameLabel id="01E41E06FFF8817F302BF889FA40F8C8" box="[1345,1436,1887,1913]" pageId="27" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<tableCitation id="65214AD4FFF8817F35F5F855FF29F82C" box="[159,245,1923,1949]" captionStart="TABLE 3" captionStartId="11.[152,244,495,521]" captionText="TABLE 3. Means and standard deviations for 16 morphometric characters measured from males representing 12 populations belonging to Anadolua genus." pageId="27" pageNumber="228">Table 3</tableCitation>
,
<figureCitation id="B09863EAFFF8817F3468F855FE5CF82C" box="[258,384,1923,1949]" captionStart-0="FIGURE 19" captionStart-1="FIGURE 20" captionStartId-0="28.[151,250,1210,1234]" captionStartId-1="29.[151,250,1782,1806]" captionTargetBox-0="[159,1428,494,1176]" captionTargetBox-1="[159,1428,187,1752]" captionTargetId-0="figure-196@28.[151,1436,486,1184]" captionTargetId-1="figure-19@29.[151,1436,179,1756]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionText-0="FIGURE 19. SEM images of stridulatory file: A. bergeri sp. nov. (A, Konya (Rezebeli) and B, Antalya (GündoğmuŞ)) and A. moli sp. nov. (C, Konya (Sultandag)). Number “1”—Stridulatory file on underside of left tegmen of males and Number “2”—stridulatory file of medial teeth at higher magnification." captionText-1="FIGURE 20. SEM images of stridulatory file: A. schwarzi (A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, Antalya (Tahtalıdağ); G, UŞak (Şaphane); H, Kütahya (Civanaga); I, Denizli (Çameli)). Number “1”—Stridulatory file on underside of left tegmen of males and Number “2”—stridulatory file of medial teeth at higher magnification." figureDoi-0="http://doi.org/10.5281/zenodo.14022308" figureDoi-1="http://doi.org/10.5281/zenodo.14022310" httpUri-0="https://zenodo.org/record/14022308/files/figure.png" httpUri-1="https://zenodo.org/record/14022310/files/figure.png" pageId="27" pageNumber="228">Fig. 1920</figureCitation>
). Cercus with internal tooth in male. Male cercus long (
<quantity id="EF5BD28AFFF8817F3698F855FBA3F82C" box="[1010,1151,1923,1949]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="27" pageNumber="228" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.22.4 mm</quantity>
), equal to half of anterior femur, with a strong tooth in the 1/2 parts (
<figureCitation id="B09863EAFFF8817F37EFF871FD26F870" box="[645,762,1959,1985]" captionStart="FIGURE 9" captionStartId="13.[151,250,1642,1666]" captionTargetBox="[317,1270,192,1603]" captionTargetId="figure-19@13.[304,1283,179,1616]" captionTargetPageId="13" captionText="FIGURE 9. Cerci of the males. AI, A. schwarzi: A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, Antalya (Tahtalıdağ); G, UŞak (Şaphane); H, Kütahya (Civanaga); I, Denizli (Çameli); JK, A. bergeri sp. nov.: J, Konya (Rezebeli); K, Antalya (GündoğmuŞ); L, A. moli sp. nov.: L, Konya (Sultandağ)." figureDoi="http://doi.org/10.5281/zenodo.14022288" httpUri="https://zenodo.org/record/14022288/files/figure.png" pageId="27" pageNumber="228">Fig. 9JK</figureCitation>
), last tergite with a deep excision which is narrowed in the first half and widened in the second (
<figureCitation id="B09863EAFFF8817F3753F81DFD70F857" box="[569,684,1995,2022]" captionStart="FIGURE 5" captionStartId="9.[151,250,1637,1661]" captionTargetBox="[211,1377,643,1598]" captionTargetId="figure-244@9.[198,1390,630,1610]" captionTargetPageId="9" captionText="FIGURE 5. Dorsal view of male last tergite. AI, A. schwarzi: A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, Antalya (Tahtalıdağ); G, UŞak (Şaphane); H, Kütahya (Civanaga); I, Denizli (Çameli); JK, A. bergeri sp. nov.: J, Konya (Rezebeli); K, Antalya (GündoğmuŞ); L, A. moli sp. nov.: L, Konya (Sultandağ)." figureDoi="http://doi.org/10.5281/zenodo.14022279" httpUri="https://zenodo.org/record/14022279/files/figure.png" pageId="27" pageNumber="228">Fig. 5JK</figureCitation>
), Male subgenital plate with a rounded obtusangular excision (
<figureCitation id="B09863EAFFF881783005F81DFF0AFF00" captionStart="FIGURE 7" captionStartId="10.[151,250,1868,1892]" captionTargetBox="[163,1424,914,1830]" captionTargetId="figure-80@10.[151,1436,902,1842]" captionTargetPageId="10" captionText="FIGURE 7. Subgenital plate of the males. AI, A. schwarzi: A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, Antalya (Tahtalıdağ); G, UŞak (Şaphane); H, Kütahya (Civanaga); I, Denizli (Çameli); JK, A. bergeri sp. nov.: J, Konya (Rezebeli); K, Antalya (GündoğmuŞ); L, A. moli sp. nov.: L, Konya (Sultandağ)." figureDoi="http://doi.org/10.5281/zenodo.14022283" httpUri="https://zenodo.org/record/14022283/files/figure.png" lastPageId="28" lastPageNumber="229" pageId="27" pageNumber="228">Fig. 7JK</figureCitation>
), female subgenital plate large, with a distinct wide angled excision, and a weak depression at the base of plate (
<figureCitation id="B09863EAFFFF817835F5FF6DFF31FF64" box="[159,237,187,213]" captionStart="FIGURE 8" captionStartId="12.[151,250,989,1013]" captionTargetBox="[162,1425,190,952]" captionTargetId="figure-23@12.[151,1436,179,963]" captionTargetPageId="12" captionText="FIGURE 8. Subgenital plate of the females. AH, A. schwarzi: A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, UŞak (Şaphane); G, Kütahya (Civanaga); H, Denizli (Çameli); I, A. bergeri sp. nov.: I, Konya (Rezebeli); J, A. moli sp. nov.: J, Konya (Sultandağ)." figureDoi="http://doi.org/10.5281/zenodo.14022285" httpUri="https://zenodo.org/record/14022285/files/figure.png" pageId="28" pageNumber="229">Fig. 8I</figureCitation>
). The basal arms of the titillator spineless, while the apical arms bear a small number of spines near the tip. The number of spines can differ not only between individuals but also between the two arms of an individual (
<figureCitation id="B09863EAFFFF81783005FF09FF38FEAC" captionStart="FIGURE 11" captionStartId="18.[151,250,1325,1349]" captionTargetBox="[156,1430,184,1294]" captionTargetId="figure-23@18.[151,1435,179,1299]" captionTargetPageId="18" captionText="FIGURE 11. Titillators of the males. AI, A. schwarzi: A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, Antalya (Tahtalıdağ); G, UŞak (Şaphane); H, Kütahya (Civanaga); I, Denizli (Çameli); JK, A. bergeri sp. nov.: J, Konya (Rezebeli); K, Antalya (GündoğmuŞ); L, A. moli sp. nov.: L, Konya (Sultandağ)." figureDoi="http://doi.org/10.5281/zenodo.14022292" httpUri="https://zenodo.org/record/14022292/files/figure.png" pageId="28" pageNumber="229">Fig. 11JK</figureCitation>
). The ovipositor is shorter than
<quantity id="EF5BD28AFFFF81783728FED4FD4CFEAC" box="[578,656,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="229" unit="mm" value="15.0">15 mm</quantity>
and weakly structured, straight or slightly curved upwards. Males and females mostly blackish brown, dark orange or dark yellowish dorsally, and brownish or yellowish ventrally in their general appearance. Male individuals generally have a pattern of brownish and/or blackish spots on their bodies on a grey background. Some individuals, especially females, with yellowish and/or dirty yellowish body colour may not have spot patterns. The tegmina from base to the apex is dark in colour up to nearly 1/2 part of it and the rest of it is yellowish or brownish (
<figureCitation id="B09863EAFFFF817834BBFE61FD99FE60" box="[465,581,439,465]" captionStart="FIGURE 4" captionStartId="8.[151,250,1457,1481]" captionTargetBox="[161,1426,189,1421]" captionTargetId="figure-23@8.[151,1436,179,1431]" captionTargetPageId="8" captionText="FIGURE 4. Tegmina of the males. AI, A. schwarzi: A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, Antalya (Tahtalıdağ); G, UŞak (Şaphane); H, Kütahya (Civanaga); I, Denizli (Çameli); JK, A. bergeri sp. nov.: J, Konya (Rezebeli); K, Antalya (GündoğmuŞ); L, A. moli sp. nov.: L, Konya (Sultandağ)." figureDoi="http://doi.org/10.5281/zenodo.14022277" httpUri="https://zenodo.org/record/14022277/files/figure.png" pageId="28" pageNumber="229">Fig. 4JK</figureCitation>
). The whole leg colour matches the general body colour and pattern.
</paragraph>
<caption id="7CDC2FE7FFFF817835FDFB6CFCD5FAAA" ID-DOI="http://doi.org/10.5281/zenodo.14022308" ID-Zenodo-Dep="14022308" httpUri="https://zenodo.org/record/14022308/files/figure.png" pageId="28" pageNumber="229" startId="28.[151,250,1210,1234]" targetBox="[159,1428,494,1176]" targetPageId="28" targetType="figure">
<paragraph id="281C7F6FFFFF817835FDFB6CFCD5FAAA" blockId="28.[151,1437,1210,1307]" pageId="28" pageNumber="229">
<emphasis id="1AD7A37DFFFF817835FDFB6CFEFEFB62" bold="true" box="[151,290,1210,1235]" pageId="28" pageNumber="229">FIGURE 19.</emphasis>
SEM images of stridulatory file:
<taxonomicName id="EFA304ECFFFF817837E9FB6DFD37FB62" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[643,747,1211,1235]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="bergeri" status="sp. nov.">
<emphasis id="1AD7A37DFFFF817837E9FB6DFD37FB62" box="[643,747,1211,1235]" italics="true" pageId="28" pageNumber="229">A. bergeri</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFF81783799FB6DFC9AFB62" bold="true" box="[755,838,1211,1235]" pageId="28" pageNumber="229">
<taxonomicNameLabel id="01E41E06FFFF81783799FB6DFC9AFB62" box="[755,838,1211,1235]" pageId="28" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A, Konya (Rezebeli) and B, Antalya (GündoğmuŞ)) and
<taxonomicName id="EFA304ECFFFF817835FDFB09FF3FFB46" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[151,227,1247,1271]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="moli" status="sp. nov.">
<emphasis id="1AD7A37DFFFF817835FDFB09FF3FFB46" box="[151,227,1247,1271]" italics="true" pageId="28" pageNumber="229">A. moli</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFF81783586FB09FEE3FB46" bold="true" box="[236,319,1247,1271]" pageId="28" pageNumber="229">
<taxonomicNameLabel id="01E41E06FFFF81783586FB09FEE3FB46" box="[236,319,1247,1271]" pageId="28" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(C, Konya (Sultandag)). Number “1”—Stridulatory file on underside of left tegmen of males and Number “2”—stridulatory file of medial teeth at higher magnification.
</paragraph>
</caption>
<paragraph id="281C7F6FFFFF817835ADFAB8FB5CF886" blockId="28.[151,1437,1389,1955]" pageId="28" pageNumber="229">
<emphasis id="1AD7A37DFFFF817835ADFAB8FD48FA36" box="[199,660,1389,1415]" italics="true" pageId="28" pageNumber="229">
Descriptive diagnosis.
<taxonomicName id="EFA304ECFFFF817834A7FABBFD48FA36" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[461,660,1389,1415]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="bergeri" status="sp. nov.">Anadolua bergeri</taxonomicName>
</emphasis>
<emphasis id="1AD7A37DFFFF817837F4FABBFD25FA36" bold="true" box="[670,761,1389,1415]" pageId="28" pageNumber="229">
<taxonomicNameLabel id="01E41E06FFFF817837F4FABBFD25FA36" box="[670,761,1389,1415]" pageId="28" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a member of the genus by the character combination as defined to characterize the genus. Among the specimens of this species, it shares similarities with other members of genus by its general morphological characters. The new species shows similarity to
<taxonomicName id="EFA304ECFFFF81783151FA63FB40FA7E" baseAuthorityName="Karabag" baseAuthorityYear="1952" box="[1083,1180,1461,1487]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="1AD7A37DFFFF81783151FA63FB40FA7E" box="[1083,1180,1461,1487]" italics="true" pageId="28" pageNumber="229">A. davisi</emphasis>
</taxonomicName>
with long cerci length, the cerci length of these two species being more than
<quantity id="EF5BD28AFFFF8178379FFA0FFC97FA42" box="[757,843,1497,1523]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="28" pageNumber="229" unit="mm" value="2.2">2.2 mm</quantity>
and equal or longer than half of anterior femur (
<figureCitation id="B09863EAFFFF81783005FA0FFF09F9A6" captionStart="FIGURE 9" captionStartId="13.[151,250,1642,1666]" captionTargetBox="[317,1270,192,1603]" captionTargetId="figure-19@13.[304,1283,179,1616]" captionTargetPageId="13" captionText="FIGURE 9. Cerci of the males. AI, A. schwarzi: A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, Antalya (Tahtalıdağ); G, UŞak (Şaphane); H, Kütahya (Civanaga); I, Denizli (Çameli); JK, A. bergeri sp. nov.: J, Konya (Rezebeli); K, Antalya (GündoğmuŞ); L, A. moli sp. nov.: L, Konya (Sultandağ)." figureDoi="http://doi.org/10.5281/zenodo.14022288" httpUri="https://zenodo.org/record/14022288/files/figure.png" pageId="28" pageNumber="229">Fig. 9JK</figureCitation>
, Appendix 1A). Also, the shape of the last tergite, with a deep excision, is similar in these two species. These characters are important for the differentiation within the genus allowing separation of
<taxonomicName id="EFA304ECFFFF8178311CF9F7FB3BF98A" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[1142,1255,1569,1595]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="bergeri" status="sp. nov.">
<emphasis id="1AD7A37DFFFF8178311CF9F7FB3BF98A" box="[1142,1255,1569,1595]" italics="true" pageId="28" pageNumber="229">A. bergeri</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFF8178319AF9F7FA97F98A" bold="true" box="[1264,1355,1569,1595]" pageId="28" pageNumber="229">
<taxonomicNameLabel id="01E41E06FFFF8178319AF9F7FA97F98A" box="[1264,1355,1569,1595]" pageId="28" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="EFA304ECFFFF817830EEF9F7FF06F9EE" baseAuthorityName="Karabag" baseAuthorityYear="1952" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="1AD7A37DFFFF817830EEF9F7FF06F9EE" italics="true" pageId="28" pageNumber="229">A. davisi</emphasis>
</taxonomicName>
from all the other species.
<taxonomicName id="EFA304ECFFFF8178376EF993FDAFF9EE" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[516,627,1605,1631]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="bergeri" status="sp. nov.">
<emphasis id="1AD7A37DFFFF8178376EF993FDAFF9EE" box="[516,627,1605,1631]" italics="true" pageId="28" pageNumber="229">A. bergeri</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFF81783713F993FD0EF9EE" bold="true" box="[633,722,1605,1631]" pageId="28" pageNumber="229">
<taxonomicNameLabel id="01E41E06FFFF81783713F993FD0EF9EE" box="[633,722,1605,1631]" pageId="28" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs prominently from others by its distinctive morphological and bioacustical characters. Male cercus length is between 2.2 and
<quantity id="EF5BD28AFFFF817836E8F9BFFC04F932" box="[898,984,1641,1667]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="28" pageNumber="229" unit="mm" value="2.4">2.4 mm</quantity>
(longer than
<taxonomicName id="EFA304ECFFFF81783106F9BFFB33F932" authorityName="Ramme" authorityYear="1939" box="[1132,1263,1641,1667]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFFF81783106F9BFFB33F932" box="[1132,1263,1641,1667]" italics="true" pageId="28" pageNumber="229">A. schwarzi</emphasis>
</taxonomicName>
and
<taxonomicName id="EFA304ECFFFF8178304EF9BFFAA8F932" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[1316,1396,1641,1667]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="moli" status="sp. nov.">
<emphasis id="1AD7A37DFFFF8178304EF9BFFAA8F932" box="[1316,1396,1641,1667]" italics="true" pageId="28" pageNumber="229">A. moli</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFF81783011F9BFFF14F916" bold="true" pageId="28" pageNumber="229">
<taxonomicNameLabel id="01E41E06FFFF81783011F9BFFF14F916" pageId="28" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and shorter than
<taxonomicName id="EFA304ECFFFF817834E4F95BFE2DF916" baseAuthorityName="Karabag" baseAuthorityYear="1952" box="[398,497,1677,1703]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="1AD7A37DFFFF817834E4F95BFE2DF916" box="[398,497,1677,1703]" italics="true" pageId="28" pageNumber="229">A. davisi</emphasis>
</taxonomicName>
). The length of hind femur generally shorter than
<taxonomicName id="EFA304ECFFFF81783146F95BFB6CF916" authorityName="Ramme" authorityYear="1939" box="[1068,1200,1677,1703]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFFF81783146F95BFB6CF916" box="[1068,1200,1677,1703]" italics="true" pageId="28" pageNumber="229">A. schwarzi</emphasis>
</taxonomicName>
and
<taxonomicName id="EFA304ECFFFF81783182F95BFAE6F916" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[1256,1338,1677,1703]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="moli" status="sp. nov.">
<emphasis id="1AD7A37DFFFF81783182F95BFAE6F916" box="[1256,1338,1677,1703]" italics="true" pageId="28" pageNumber="229">A. moli</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFF81783028F95BFA40F916" bold="true" box="[1346,1436,1677,1703]" pageId="28" pageNumber="229">
<taxonomicNameLabel id="01E41E06FFFF81783028F95BFA40F916" box="[1346,1436,1677,1703]" pageId="28" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<tableCitation id="65214AD4FFFF817835F5F967FF2AF97A" box="[159,246,1713,1739]" captionStart="TABLE 3" captionStartId="11.[152,244,495,521]" captionText="TABLE 3. Means and standard deviations for 16 morphometric characters measured from males representing 12 populations belonging to Anadolua genus." pageId="28" pageNumber="229">Table 3</tableCitation>
), and ratio hind femur/cercus ranges between 7.0 and 8.2 (
<quantity id="EF5BD28AFFFF817836F2F967FC07F97A" box="[920,987,1713,1740]" metricMagnitude="-1" metricUnit="m" metricValue="1.3208" pageId="28" pageNumber="229" unit="in" value="5.2">5.2 in</quantity>
<taxonomicName id="EFA304ECFFFF8178368EF967FB9BF97A" baseAuthorityName="Karabag" baseAuthorityYear="1952" box="[996,1095,1713,1739]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="1AD7A37DFFFF8178368EF967FB9BF97A" box="[996,1095,1713,1739]" italics="true" pageId="28" pageNumber="229">A. davisi</emphasis>
</taxonomicName>
, 9.212.0 in
<taxonomicName id="EFA304ECFFFF8178318AF967FAB9F97A" authorityName="Ramme" authorityYear="1939" box="[1248,1381,1713,1739]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFFF8178318AF967FAB9F97A" box="[1248,1381,1713,1739]" italics="true" pageId="28" pageNumber="229">A. schwarzi</emphasis>
</taxonomicName>
, and
<quantity id="EF5BD28AFFFF817835FDF903FEF8F95E" box="[151,292,1749,1776]" metricMagnitude="-1" metricUnit="m" metricValue="3.4162999999999997" metricValueMax="3.6576" metricValueMin="3.175" pageId="28" pageNumber="229" unit="in" value="13.45" valueMax="14.4" valueMin="12.5">12.514.4 in</quantity>
<taxonomicName id="EFA304ECFFFF81783441F903FEA0F95E" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[299,380,1749,1775]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="moli" status="sp. nov.">
<emphasis id="1AD7A37DFFFF81783441F903FEA0F95E" box="[299,380,1749,1775]" italics="true" pageId="28" pageNumber="229">A. moli</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFF817834E9F903FE00F95E" bold="true" box="[387,476,1749,1775]" pageId="28" pageNumber="229">
<taxonomicNameLabel id="01E41E06FFFF817834E9F903FE00F95E" box="[387,476,1749,1775]" pageId="28" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). While the phrase of
<taxonomicName id="EFA304ECFFFF817837BCF903FC9AF95E" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[726,838,1749,1775]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="bergeri" status="sp. nov.">
<emphasis id="1AD7A37DFFFF817837BCF903FC9AF95E" box="[726,838,1749,1775]" italics="true" pageId="28" pageNumber="229">A. bergeri</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFF81783624F903FC7BF95E" bold="true" box="[846,935,1749,1775]" pageId="28" pageNumber="229">
<taxonomicNameLabel id="01E41E06FFFF81783624F903FC7BF95E" box="[846,935,1749,1775]" pageId="28" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male calling song consists of single-syllable series, the phrases of other three species consist of double or triple syllables group
<emphasis id="1AD7A37DFFFF81783138F92CFB4FF8A2" box="[1106,1171,1786,1811]" italics="true" pageId="28" pageNumber="229">series</emphasis>
. Also, other diagnostic characters, namely morphological and acoustical, are presented in the identification keys.
</paragraph>
<paragraph id="281C7F6FFFFF817835ADF897FD51F812" blockId="28.[151,1437,1389,1955]" pageId="28" pageNumber="229">
<emphasis id="1AD7A37DFFFF817835ADF897FE97F8EA" box="[199,331,1857,1883]" italics="true" pageId="28" pageNumber="229">Coloration:</emphasis>
The colour variation of individuals of the species is very similar to that of
<taxonomicName id="EFA304ECFFFF817831B8F897FA86F8EA" authorityName="Ramme" authorityYear="1939" box="[1234,1370,1857,1883]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFFF817831B8F897FA86F8EA" box="[1234,1370,1857,1883]" italics="true" pageId="28" pageNumber="229">A. schwarzi</emphasis>
</taxonomicName>
. Two populations of this species are known, and it is seen that this species has dark yellow, grey, light brown and dark brown colour scale in general. (Appendix 7).
</paragraph>
<caption id="7CDC2FE7FFFE817935FDF920FE1FF8CB" ID-DOI="http://doi.org/10.5281/zenodo.14022310" ID-Zenodo-Dep="14022310" httpUri="https://zenodo.org/record/14022310/files/figure.png" pageId="29" pageNumber="230" startId="29.[151,250,1782,1806]" targetBox="[159,1428,187,1752]" targetPageId="29" targetType="figure">
<paragraph id="281C7F6FFFFE817935FDF920FE1FF8CB" blockId="29.[151,1437,1782,1914]" pageId="29" pageNumber="230">
<emphasis id="1AD7A37DFFFE817935FDF920FEFCF8BF" bold="true" box="[151,288,1782,1806]" pageId="29" pageNumber="230">FIGURE 20.</emphasis>
SEM images of stridulatory file:
<taxonomicName id="EFA304ECFFFE81793712F921FD32F8BF" authorityName="Ramme" authorityYear="1939" box="[632,750,1782,1806]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFFE81793712F921FD32F8BF" box="[632,750,1782,1806]" italics="true" pageId="29" pageNumber="230">A. schwarzi</emphasis>
</taxonomicName>
(A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, Antalya (Tahtalıdağ); G, UŞak (Şaphane); H, Kütahya (Civanaga); I, Denizli (Çameli)). Number “1”—Stridulatory file on underside of left tegmen of males and Number “2”—stridulatory file of medial teeth at higher magnification.
</paragraph>
</caption>
<paragraph id="281C7F6FFFFD817A35ADFF41FC05FD18" blockId="30.[151,1437,151,1185]" pageId="30" pageNumber="231">
<emphasis id="1AD7A37DFFFD817A35ADFF41FE71FF01" box="[199,429,151,177]" italics="true" pageId="30" pageNumber="231">Description of song:</emphasis>
A total number of 23 calling song recordings of
<specimenCount id="3EA5B4E6FFFD817A36ADFF41FBE1FF00" box="[967,1085,151,177]" count="4" pageId="30" pageNumber="231" type="male">four males</specimenCount>
have been examined. The male calling song consists of quite similar phrases in duration (
<figureCitation id="B09863EAFFFD817A3649FF6DFC56FF64" box="[803,906,187,213]" captionStart="FIGURE 18" captionStartId="27.[151,250,1276,1300]" captionTargetBox="[164,1422,529,1237]" captionTargetId="figure-118@27.[151,1436,518,1251]" captionTargetPageId="27" captionText="FIGURE 18. Distribution map (A) and male calling song of Anadolua bergeri sp. nov. (B—two consecutive complete phrases, C—a complete phrase and D—two syllables, black arrow is the hypothetic opening hemisyllable movement and grey arrow is the hypothetic closing hemisyllable movement)." figureDoi="http://doi.org/10.5281/zenodo.14022306" httpUri="https://zenodo.org/record/14022306/files/figure.png" pageId="30" pageNumber="231">Fig. 18B</figureCitation>
). The calling songs are recorded from males at 2830°C in room conditions (by D. Şirin and M.S. Taylan). The phrase structure is a distinct different song
<typeStatus id="F718C1CDFFFD817A3025FF09FAA3FF48" box="[1359,1407,223,249]" pageId="30" pageNumber="231">type</typeStatus>
in the genus
<taxonomicName id="EFA304ECFFFD817A346DFED5FEA8FEAC" authorityName="Ramme" authorityYear="1939" box="[263,372,259,285]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="1AD7A37DFFFD817A346DFED5FEA8FEAC" box="[263,372,259,285]" italics="true" pageId="30" pageNumber="231">Anadolua</emphasis>
</taxonomicName>
and contains continuous single-syllable series. Phrase duration varies from 0.48 to 0.86 s ((0.65 ± 0.11 s). Each phrase contains variable number (1220) single syllables series. The phrase usually begins with a low amplitude syllable, and then continues and lasts with a similar amplitude syllable series. Each syllable shows a standard structure, formulated as a short and rather indistinct soft opening hemisyllable + loud closing hemisyllable (
<figureCitation id="B09863EAFFFD817A35F5FE45FEF6FE1C" box="[159,298,403,429]" captionStart="FIGURE 18" captionStartId="27.[151,250,1276,1300]" captionTargetBox="[164,1422,529,1237]" captionTargetId="figure-118@27.[151,1436,518,1251]" captionTargetPageId="27" captionText="FIGURE 18. Distribution map (A) and male calling song of Anadolua bergeri sp. nov. (B—two consecutive complete phrases, C—a complete phrase and D—two syllables, black arrow is the hypothetic opening hemisyllable movement and grey arrow is the hypothetic closing hemisyllable movement)." figureDoi="http://doi.org/10.5281/zenodo.14022306" httpUri="https://zenodo.org/record/14022306/files/figure.png" pageId="30" pageNumber="231">Fig. 18CD</figureCitation>
). Oscillographic analyses show that the period durations of the syllables vary between 28.8 and 45.2 ms (38.6 ± 3.43). The durations opening and closing hemisyllables vary between 6.9 and 17.6 ms (9.8 ± 1.21) and between 19.1 and 34.6 ms (29.1 ± 2.54), respectively. Each closing hemisyllable begins with a low amplitude, reaches the highest level at approximately the midpoint, and lasts by decreasing in amplitude after this point. The spectral analyses show that Delta and Peak frequency of the
<emphasis id="1AD7A37DFFFD817A365EFDF5FB77FD8D" box="[820,1195,547,573]" italics="true" pageId="30" pageNumber="231">first low amplitude syllable group</emphasis>
varies between 23.55 and 39.67 kHz (29.97 ± 4.66) for Delta and between 20.25 and 33.00 kHz (25.58 ± 3.88) for Peak. Delta and Peak frequencies of the
<emphasis id="1AD7A37DFFFD817A3401FDBDFDB9FD35" box="[363,613,619,645]" italics="true" pageId="30" pageNumber="231">Phrase syllable group</emphasis>
(except first one) vary between 37.00 and 54.38 kHz (47.34 ± 5.52) for Delta and between 19.13 and 31.50 kHz (27.19 ± 3.53) (
<tableCitation id="65214AD4FFFD817A3664FD59FCB8FD18" box="[782,868,655,681]" captionStart="TABLE 7" captionStartId="18.[152,244,1469,1493]" captionText="TABLE 7. Spectral parameters belong to male calling songs of Anadolua shcwarzi, A. moli sp. nov. and A. bergeri sp. nov. populations." pageId="30" pageNumber="231">Table 7</tableCitation>
) for Peak.
</paragraph>
<paragraph id="281C7F6FFFFD817A35ADFD65FE51FD40" blockId="30.[151,1437,151,1185]" pageId="30" pageNumber="231">
<emphasis id="1AD7A37DFFFD817A35ADFD65FE68FD7C" box="[199,436,691,717]" italics="true" pageId="30" pageNumber="231">Measurements (mm).</emphasis>
<typeStatus id="F718C1CDFFFD817A34D6FD65FDF5FD7C" box="[444,553,691,717]" pageId="30" pageNumber="231" type="holotype">Holotype</typeStatus>
: Measurements (mm): Body length: 24.0, Pronotum: 6.58, Cercus: 2.44, Hind femur: 17.1 (
<tableCitation id="65214AD4FFFD817A3442FD01FEA3FD40" box="[296,383,727,753]" captionStart="TABLE 3" captionStartId="11.[152,244,495,521]" captionText="TABLE 3. Means and standard deviations for 16 morphometric characters measured from males representing 12 populations belonging to Anadolua genus." pageId="30" pageNumber="231">Table 3</tableCitation>
).
</paragraph>
<paragraph id="281C7F6FFFFD817A35ADFD2DFDA2FCEC" blockId="30.[151,1437,151,1185]" pageId="30" pageNumber="231">
<emphasis id="1AD7A37DFFFD817A35ADFD2DFE99FCA4" box="[199,325,763,789]" italics="true" pageId="30" pageNumber="231">Etymology.</emphasis>
The new species is named after Dr. Dirk Berger, who has provided great support to the first two authors with different techniques of bioacoustics analysis. He has also made a major contribution to the study of Turkish
<taxonomicName id="EFA304ECFFFD817A359EFC95FE54FCEC" authorityName="Fischer" authorityYear="1853" box="[244,392,835,861]" class="Insecta" family="Acrididae" genus="Stenobothrus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="1AD7A37DFFFD817A359EFC95FE54FCEC" box="[244,392,835,861]" italics="true" pageId="30" pageNumber="231">Stenobothrus</emphasis>
</taxonomicName>
in his doctoral thesis.
</paragraph>
<paragraph id="281C7F6FFFFD817A35ADFCB1FBA2FBA0" blockId="30.[151,1437,151,1185]" pageId="30" pageNumber="231">
<emphasis id="1AD7A37DFFFD817A35ADFCB1FEF5FC30" box="[199,297,871,897]" italics="true" pageId="30" pageNumber="231">Remarks</emphasis>
. The populations previously diagnosed and published (
<bibRefCitation id="4C32029EFFFD817A36FAFCBEFBCCFC30" author="Unal, M." box="[912,1040,871,899]" pageId="30" pageNumber="231" pagination="1 - 66" refId="ref27801" refString="Unal, M. (2018) Tettigoniinae (Orthoptera: Tettigoniidae) from Turkey with key to genera and descriptions of six new species. Zootaxa, 4432 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4432.1.1" type="journal article" year="2018">Ünal, 2018</bibRefCitation>
) as
<taxonomicName id="EFA304ECFFFD817A3151FCB1FB48FC30" authorityName="Karabag" authorityYear="1958" box="[1083,1172,871,897]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="burri">
<emphasis id="1AD7A37DFFFD817A3151FCB1FB48FC30" box="[1083,1172,871,897]" italics="true" pageId="30" pageNumber="231">A. burri</emphasis>
</taxonomicName>
from Rezebeli (
<collectingRegion id="EA67B18DFFFD817A302DFCB1FA49FC30" box="[1351,1429,871,897]" country="Turkey" name="Konya" pageId="30" pageNumber="231">Konya</collectingRegion>
) and GündoğmuŞ (
<collectingRegion id="EA67B18DFFFD817A3430FC5DFE6AFC14" box="[346,438,907,933]" country="Turkey" name="Antalya" pageId="30" pageNumber="231">Antalya</collectingRegion>
) have been studied in the present study on a large scale.According to the findings obtained, based on morphology and the male calling songs, these populations exhibit distinct differences from
<taxonomicName id="EFA304ECFFFD817A3045FC79FF2CFC5C" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="1AD7A37DFFFD817A3045FC79FA40FC78" box="[1327,1436,943,969]" italics="true" pageId="30" pageNumber="231">Anadolua</emphasis>
species.
</taxonomicName>
The identification of these populations as
<taxonomicName id="EFA304ECFFFD817A37AFFC05FCC1FC5C" authorityName="Karabag" authorityYear="1958" box="[709,797,979,1005]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="burri">
<emphasis id="1AD7A37DFFFD817A37AFFC05FCC1FC5C" box="[709,797,979,1005]" italics="true" pageId="30" pageNumber="231">A. burri</emphasis>
</taxonomicName>
by
<bibRefCitation id="4C32029EFFFD817A362FFC02FC11FC5C" author="Unal, M." box="[837,973,979,1007]" pageId="30" pageNumber="231" pagination="1 - 66" refId="ref27801" refString="Unal, M. (2018) Tettigoniinae (Orthoptera: Tettigoniidae) from Turkey with key to genera and descriptions of six new species. Zootaxa, 4432 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4432.1.1" type="journal article" year="2018">Ünal (2018)</bibRefCitation>
is an expected situation, because the only key that exists is that of
<bibRefCitation id="4C32029EFFFD817A34CDFC21FD84FBA0" author="Karabag, T." box="[423,600,1014,1041]" pageId="30" pageNumber="231" pagination="139 - 143" refId="ref25717" refString="Karabag, T. (1952) Anadolua Ramme (Orthoptera, Tettigoniidae) genusunun revizyonu. Istanbul Universitesi Fen Fakultesi Mecmuasi, 17 (2), 139 - 143. [in Turkish]" type="journal article" year="1952">Karabağ (1952)</bibRefCitation>
and the characters in that key are not distinctive.
</paragraph>
<paragraph id="281C7F6FFFFD817A35ADFBCDFAC8FB10" blockId="30.[151,1437,151,1185]" pageId="30" pageNumber="231">
<emphasis id="1AD7A37DFFFD817A35ADFBCDFE18FB84" box="[199,452,1051,1077]" italics="true" pageId="30" pageNumber="231">IUCN status proposal.</emphasis>
This species has not been assessed for the IUCN Red List (2023-1). The distribution of the
<taxonomicName id="EFA304ECFFFD817A35AAFBE9FEF3FBE8" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[192,303,1087,1113]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="bergeri">
<emphasis id="1AD7A37DFFFD817A35AAFBE9FEF3FBE8" box="[192,303,1087,1113]" italics="true" pageId="30" pageNumber="231">A. bergeri</emphasis>
</taxonomicName>
is restricted and its habitats are strongly under the threat of the anthropogenic effects such as camping areas, picnic areas, hiking routes, and overgrazing. The species should be considered as a
<emphasis id="1AD7A37DFFFD817A31CAFBB5FA40FBCC" box="[1184,1436,1123,1149]" italics="true" pageId="30" pageNumber="231">Critically Endangered</emphasis>
(B1ab (iii+v)) status based on the extent of occurrence criteria of IUCN (https://www.iucnredlist.org/).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,413 @@
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<mods:title id="B8CD761DB4CFB78EDB6437A0CD30E612">A revision of Anadolua (Orthoptera: Tettigoniidae: Tettigoniinae) based on morphological and bioacoustic characters: Newly confirmed data suggesting a new synonym and two new species</mods:title>
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<emphasis id="1AD7A37DFFFD817A35FDFB24FE90FABC" bold="true" box="[151,332,1266,1293]" italics="true" pageId="30" pageNumber="231">Anadolua moli</emphasis>
Şirin, Sevgili and Taylan
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,
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<emphasis id="1AD7A37DFFFD817A35FDFAEDFEF5FAE4" box="[151,297,1339,1365]" italics="true" pageId="30" pageNumber="231">Distribution:</emphasis>
Specimens belongs population of new species have been collected only from the
<collectingRegion id="EA67B18DFFFD817A318EFAEDFAEDFAE4" box="[1252,1329,1339,1365]" country="Turkey" name="Konya" pageId="30" pageNumber="231">Konya</collectingRegion>
, AkŞehir, Sultandağ Mountains (
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).
</paragraph>
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<typeStatus id="F718C1CDFFFD817A35ADFA55FEEDFA2C" box="[199,305,1411,1437]" pageId="30" pageNumber="231" type="holotype">Holotype</typeStatus>
:
</emphasis>
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(deposited in alcohol,
<collectionCode id="4EB2E7AAFFFD817A3712FA55FD38FA2C" box="[632,740,1411,1437]" pageId="30" pageNumber="231">NKUEM</collectionCode>
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<collectingCountry id="50B43FFFFFFD817A379CFA55FC8EFA2C" box="[758,850,1411,1437]" name="Turkey" pageId="30" pageNumber="231">Türkiye</collectingCountry>
,
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:
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,
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,
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N.N.,
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, leg.:
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, H. Sevgili &amp;
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Head equal width with pronotum in both sexes. Fastigium slightly wider than scapus in male and distinct wider than scapus in female. Maximum width of fastigium / Maximum width of scapus 1.0
<quantity id="EF5BD28AFFFD817A3026F9C5FA40F99C" box="[1356,1436,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="3.5559999999999996" pageId="30" pageNumber="231" unit="in" value="1.4">1.40 in</quantity>
male,
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female. A black band between the eyes present. Disk of pronotum with hourglass shape and surface weakly flattened in prozona but distinctly flattened in metazona, and also with a deep transverse depression behind the middle, and its posterior edge is broadly rounded. Maximum distance between lateral carinae / minimum distance between lateral carinae
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male,
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female. The median carinae is absent or slightly distinct, and the lateral carinae is prominent as round in the second part of the metazona. Hind femur long, its length 4.255.0 times of its maximum width in male
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female. Male tegmina micropterous form and barely reaches the end of second abdominal tergite; female tegmina is the squamipterous not touching and only slightly protruding beyond posterior margin of pronotum. The speculum is in quadriform shape (generally trapezoidal), and, usually, no lateral vein development occurs. Costal vein absent or very weak; subcostal vein is not reach to the apex, parallel to radial vein, and generally, they may merge with radial vein near the apex. Medial vein sinuate at the apex and usually has a dichotomy at the ¾ from the base (
<figureCitation id="B09863EAFFFD817A378CF849FCE2F808" box="[742,830,1951,1977]" captionStart="FIGURE 4" captionStartId="8.[151,250,1457,1481]" captionTargetBox="[161,1426,189,1421]" captionTargetId="figure-23@8.[151,1436,179,1431]" captionTargetPageId="8" captionText="FIGURE 4. Tegmina of the males. AI, A. schwarzi: A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, Antalya (Tahtalıdağ); G, UŞak (Şaphane); H, Kütahya (Civanaga); I, Denizli (Çameli); JK, A. bergeri sp. nov.: J, Konya (Rezebeli); K, Antalya (GündoğmuŞ); L, A. moli sp. nov.: L, Konya (Sultandağ)." figureDoi="http://doi.org/10.5281/zenodo.14022277" httpUri="https://zenodo.org/record/14022277/files/figure.png" pageId="30" pageNumber="231">Fig. 4L</figureCitation>
). Male cercus short (
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), shorter than half of anterior femur, with a strong tooth near the middle of it or just before (
<figureCitation id="B09863EAFFFD817A36D0F815FBCEF86C" box="[954,1042,1987,2013]" captionStart="FIGURE 9" captionStartId="13.[151,250,1642,1666]" captionTargetBox="[317,1270,192,1603]" captionTargetId="figure-19@13.[304,1283,179,1616]" captionTargetPageId="13" captionText="FIGURE 9. Cerci of the males. AI, A. schwarzi: A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, Antalya (Tahtalıdağ); G, UŞak (Şaphane); H, Kütahya (Civanaga); I, Denizli (Çameli); JK, A. bergeri sp. nov.: J, Konya (Rezebeli); K, Antalya (GündoğmuŞ); L, A. moli sp. nov.: L, Konya (Sultandağ)." figureDoi="http://doi.org/10.5281/zenodo.14022288" httpUri="https://zenodo.org/record/14022288/files/figure.png" pageId="30" pageNumber="231">Fig. 9L</figureCitation>
), with internal tooth; last tergite of male with a wide excision, which nearly divides the plate in two lobes (
<figureCitation id="B09863EAFFFC817B36C2FF41FC22FF00" box="[936,1022,150,177]" captionStart="FIGURE 5" captionStartId="9.[151,250,1637,1661]" captionTargetBox="[211,1377,643,1598]" captionTargetId="figure-244@9.[198,1390,630,1610]" captionTargetPageId="9" captionText="FIGURE 5. Dorsal view of male last tergite. AI, A. schwarzi: A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, Antalya (Tahtalıdağ); G, UŞak (Şaphane); H, Kütahya (Civanaga); I, Denizli (Çameli); JK, A. bergeri sp. nov.: J, Konya (Rezebeli); K, Antalya (GündoğmuŞ); L, A. moli sp. nov.: L, Konya (Sultandağ)." figureDoi="http://doi.org/10.5281/zenodo.14022279" httpUri="https://zenodo.org/record/14022279/files/figure.png" pageId="31" pageNumber="232">Fig. 5L</figureCitation>
). Subgenital plate with a deep deltoid excision (
<figureCitation id="B09863EAFFFC817B346EFF6DFE87FF64" box="[260,347,187,213]" captionStart="FIGURE 7" captionStartId="10.[151,250,1868,1892]" captionTargetBox="[163,1424,914,1830]" captionTargetId="figure-80@10.[151,1436,902,1842]" captionTargetPageId="10" captionText="FIGURE 7. Subgenital plate of the males. AI, A. schwarzi: A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, Antalya (Tahtalıdağ); G, UŞak (Şaphane); H, Kütahya (Civanaga); I, Denizli (Çameli); JK, A. bergeri sp. nov.: J, Konya (Rezebeli); K, Antalya (GündoğmuŞ); L, A. moli sp. nov.: L, Konya (Sultandağ)." figureDoi="http://doi.org/10.5281/zenodo.14022283" httpUri="https://zenodo.org/record/14022283/files/figure.png" pageId="31" pageNumber="232">Fig. 7L</figureCitation>
). Female subgenital plate large, with a distinct angled excision, and a diamond like depression at the base of plate (
<figureCitation id="B09863EAFFFC817B3433FF09FE7BFF48" box="[345,423,223,249]" captionStart="FIGURE 8" captionStartId="12.[151,250,989,1013]" captionTargetBox="[162,1425,190,952]" captionTargetId="figure-23@12.[151,1436,179,963]" captionTargetPageId="12" captionText="FIGURE 8. Subgenital plate of the females. AH, A. schwarzi: A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, UŞak (Şaphane); G, Kütahya (Civanaga); H, Denizli (Çameli); I, A. bergeri sp. nov.: I, Konya (Rezebeli); J, A. moli sp. nov.: J, Konya (Sultandağ)." figureDoi="http://doi.org/10.5281/zenodo.14022285" httpUri="https://zenodo.org/record/14022285/files/figure.png" pageId="31" pageNumber="232">Fig. 8J</figureCitation>
). Basal arms of titillator spineless, while the apical arms bear a small number of spines near the tip (
<figureCitation id="B09863EAFFFC817B3587FED5FE8CFEAC" box="[237,336,259,285]" captionStart="FIGURE 11" captionStartId="18.[151,250,1325,1349]" captionTargetBox="[156,1430,184,1294]" captionTargetId="figure-23@18.[151,1435,179,1299]" captionTargetPageId="18" captionText="FIGURE 11. Titillators of the males. AI, A. schwarzi: A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, Antalya (Tahtalıdağ); G, UŞak (Şaphane); H, Kütahya (Civanaga); I, Denizli (Çameli); JK, A. bergeri sp. nov.: J, Konya (Rezebeli); K, Antalya (GündoğmuŞ); L, A. moli sp. nov.: L, Konya (Sultandağ)." figureDoi="http://doi.org/10.5281/zenodo.14022292" httpUri="https://zenodo.org/record/14022292/files/figure.png" pageId="31" pageNumber="232">Fig. 11L</figureCitation>
). Ovipositor longer than
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, straight. Males mostly blackish brown or dark yellowish dorsally and brownish or yellowish ventrally in their general appearance. Although the colour of females is similar to that of males, they generally have lighter tones. Male and female individuals usually have a pattern of brownish and/or blackish spots on their bodies on a grey background. Some individuals with yellowish and/or dirty yellowish body colour may not have spot patterns. The tegmina from base to apex is dark in colour up to nearly 1/2 part of it and the rest of it is yellowish or brownish (Figure Tegmina). The entire leg colour matches the general body colour and pattern.
</paragraph>
<caption id="7CDC2FE7FFFC817B35FDFB28FD04FAEF" ID-DOI="http://doi.org/10.5281/zenodo.14022312" ID-Zenodo-Dep="14022312" httpUri="https://zenodo.org/record/14022312/files/figure.png" pageId="31" pageNumber="232" startId="31.[151,250,1278,1302]" targetBox="[161,1418,540,1239]" targetPageId="31" targetType="figure">
<paragraph id="281C7F6FFFFC817B35FDFB28FD04FAEF" blockId="31.[151,1436,1278,1374]" pageId="31" pageNumber="232">
<emphasis id="1AD7A37DFFFC817B35FDFB28FEFEFAA7" bold="true" box="[151,290,1278,1302]" pageId="31" pageNumber="232">FIGURE 21.</emphasis>
Distribution map (A) and male calling song of
<taxonomicName id="EFA304ECFFFC817B367EFB28FC77FAA7" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[788,939,1278,1302]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="moli" status="sp. nov.">
<emphasis id="1AD7A37DFFFC817B367EFB28FC77FAA7" box="[788,939,1278,1302]" italics="true" pageId="31" pageNumber="232">Anadolua moli</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFC817B36D9FB28FBDAFAA7" bold="true" box="[947,1030,1278,1302]" pageId="31" pageNumber="232">
<taxonomicNameLabel id="01E41E06FFFC817B36D9FB28FBDAFAA7" box="[947,1030,1278,1302]" pageId="31" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(B—two consecutive complete phrase, C—a group of syllables series and D—three syllables, black arrow is the hypothetic opening hemisyllable movement and grey arrow is the hypothetic closing hemisyllable movement).
</paragraph>
</caption>
<paragraph id="281C7F6FFFFC817B35ADFA67FC09F8E7" blockId="31.[151,1437,1456,2022]" pageId="31" pageNumber="232">
<emphasis id="1AD7A37DFFFC817B35ADFA67FDB1FA7B" box="[199,621,1456,1482]" italics="true" pageId="31" pageNumber="232">
Descriptive diagnosis.
<taxonomicName id="EFA304ECFFFC817B34ADFA66FDB1FA7B" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[455,621,1456,1482]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="moli" status="sp. nov.">Anadolua moli</taxonomicName>
</emphasis>
<emphasis id="1AD7A37DFFFC817B3719FA66FD10FA7B" bold="true" box="[627,716,1456,1482]" pageId="31" pageNumber="232">
<taxonomicNameLabel id="01E41E06FFFC817B3719FA66FD10FA7B" box="[627,716,1456,1482]" pageId="31" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a member of
<emphasis id="1AD7A37DFFFC817B36EAFA66FC28FA7B" box="[896,1012,1456,1482]" italics="true" pageId="31" pageNumber="232">
<taxonomicName id="EFA304ECFFFC817B36EAFA66FC2DFA7B" authorityName="Ramme" authorityYear="1939" box="[896,1009,1456,1482]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="genus">Anadolua</taxonomicName>
.
</emphasis>
Among the specimens of this species, it shares similarities with
<taxonomicName id="EFA304ECFFFC817B34D2FA02FD3EFA5F" authority="Ramme, 1939" authorityName="Ramme" authorityYear="1939" box="[440,738,1492,1519]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFFC817B34D2FA02FDE7FA5F" box="[440,571,1492,1518]" italics="true" pageId="31" pageNumber="232">A. schwarzi</emphasis>
<bibRefCitation id="4C32029EFFFC817B3728FA03FD3EFA5F" author="Ramme, W." box="[578,738,1492,1519]" pageId="31" pageNumber="232" pagination="41 - 150" refId="ref26818" refString="Ramme, W. (1939) Beitraege zur Kenntnis der palaearktischen Orthopterenfauna (Tettig. u. Acrid.). III. Mitteilungen aus dem Zoologischen Museum Berlin, 24, 41 - 150." type="journal article" year="1939">Ramme, 1939</bibRefCitation>
</taxonomicName>
. The new species shows similarity to
<taxonomicName id="EFA304ECFFFC817B31E0FA02FAD1FA5F" authorityName="Ramme" authorityYear="1939" box="[1162,1293,1492,1518]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFFC817B31E0FA02FAD1FA5F" box="[1162,1293,1492,1518]" italics="true" pageId="31" pageNumber="232">A. schwarzi</emphasis>
</taxonomicName>
with several characters, longest femur, and teeth shape of stridulatory file in male and longest ovipositor (
<tableCitation id="65214AD4FFFC817B31F4FA2EFB2FF9A3" box="[1182,1267,1528,1554]" captionStart="TABLE 3" captionStartId="11.[152,244,495,521]" captionText="TABLE 3. Means and standard deviations for 16 morphometric characters measured from males representing 12 populations belonging to Anadolua genus." pageId="31" pageNumber="232">Table 3</tableCitation>
,
<figureCitation id="B09863EAFFFC817B3195FA2EFA87F9A3" box="[1279,1371,1528,1554]" captionStart="FIGURE 12" captionStartId="20.[151,250,1294,1318]" captionTargetBox="[162,1425,190,1257]" captionTargetId="figure-23@20.[151,1436,179,1268]" captionTargetPageId="20" captionText="FIGURE 12. Lateral view of Ovipositor. AH, A. schwarzi: A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, UŞak (Şaphane); G, Kütahya (Civanaga); H, Denizli (Çameli); I, A. bergeri sp. nov.: I, Konya (Rezebeli); J, A. moli sp. nov.: J, Konya (Sultandağ)." figureDoi="http://doi.org/10.5281/zenodo.14022294" httpUri="https://zenodo.org/record/14022294/files/figure.png" pageId="31" pageNumber="232">Figs. 12</figureCitation>
,
<figureCitation id="B09863EAFFFC817B300DFA2EFA4BF9A3" box="[1383,1431,1528,1554]" captionStart="FIGURE 19" captionStartId="28.[151,250,1210,1234]" captionTargetBox="[159,1428,494,1176]" captionTargetId="figure-196@28.[151,1436,486,1184]" captionTargetPageId="28" captionText="FIGURE 19. SEM images of stridulatory file: A. bergeri sp. nov. (A, Konya (Rezebeli) and B, Antalya (GündoğmuŞ)) and A. moli sp. nov. (C, Konya (Sultandag)). Number “1”—Stridulatory file on underside of left tegmen of males and Number “2”—stridulatory file of medial teeth at higher magnification." figureDoi="http://doi.org/10.5281/zenodo.14022308" httpUri="https://zenodo.org/record/14022308/files/figure.png" pageId="31" pageNumber="232">19C</figureCitation>
,
<figureCitation id="B09863EAFFFC817B35FDF9CAFF6BF987" box="[151,183,1564,1590]" captionStart="FIGURE 20" captionStartId="29.[151,250,1782,1806]" captionTargetBox="[159,1428,187,1752]" captionTargetId="figure-19@29.[151,1436,179,1756]" captionTargetPageId="29" captionText="FIGURE 20. SEM images of stridulatory file: A. schwarzi (A, Muğla (Sandras); B, Denizli (Babadağ); C, İzmir (Bozdağ); D, Antalya (Gömbe-I); E, Antalya (Gömbe-II); F, Antalya (Tahtalıdağ); G, UŞak (Şaphane); H, Kütahya (Civanaga); I, Denizli (Çameli)). Number “1”—Stridulatory file on underside of left tegmen of males and Number “2”—stridulatory file of medial teeth at higher magnification." figureDoi="http://doi.org/10.5281/zenodo.14022310" httpUri="https://zenodo.org/record/14022310/files/figure.png" pageId="31" pageNumber="232">20</figureCitation>
). Also, these characters can separate both
<taxonomicName id="EFA304ECFFFC817B37FEF9CAFD3AF987" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[660,742,1564,1590]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="moli" status="sp. nov.">
<emphasis id="1AD7A37DFFFC817B37FEF9CAFD3AF987" box="[660,742,1564,1590]" italics="true" pageId="31" pageNumber="232">A. moli</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFC817B3784F9CAFC94F987" bold="true" box="[750,840,1564,1590]" pageId="31" pageNumber="232">
<taxonomicNameLabel id="01E41E06FFFC817B3784F9CAFC94F987" box="[750,840,1564,1590]" pageId="31" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="EFA304ECFFFC817B36EBF9CAFBD9F987" authorityName="Ramme" authorityYear="1939" box="[897,1029,1564,1590]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFFC817B36EBF9CAFBD9F987" box="[897,1029,1564,1590]" italics="true" pageId="31" pageNumber="232">A. schwarzi</emphasis>
</taxonomicName>
from all other
<taxonomicName id="EFA304ECFFFC817B31DAF9CAFA44F987" box="[1200,1432,1564,1590]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="1AD7A37DFFFC817B31DAF9CAFAC1F987" box="[1200,1309,1564,1590]" italics="true" pageId="31" pageNumber="232">Anadolua</emphasis>
species.
<emphasis id="1AD7A37DFFFC817B30EEF9CAFA44F987" box="[1412,1432,1564,1590]" italics="true" pageId="31" pageNumber="232">A</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFC817B30F3F9CAFF15F9EB" italics="true" pageId="31" pageNumber="232">. moli</emphasis>
<emphasis id="1AD7A37DFFFC817B35B9F996FEF3F9EB" bold="true" box="[211,303,1600,1626]" pageId="31" pageNumber="232">
<taxonomicNameLabel id="01E41E06FFFC817B35B9F996FEF3F9EB" box="[211,303,1600,1626]" pageId="31" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs prominently from others by its distinctive morphological and bioacustical characters. Male cercus length in between 1.3 and
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and it is the shortest cercus length in genus (
<quantity id="EF5BD28AFFFC817B310EF9B2FB31F9CE" box="[1124,1261,1636,1663]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="31" pageNumber="232" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.61.9 mm</quantity>
in
<taxonomicName id="EFA304ECFFFC817B3078F9B2FA4AF9CF" authorityName="Ramme" authorityYear="1939" box="[1298,1430,1636,1662]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFFC817B3078F9B2FA4AF9CF" box="[1298,1430,1636,1662]" italics="true" pageId="31" pageNumber="232">A. schwarzi</emphasis>
</taxonomicName>
, 2.2.
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in
<taxonomicName id="EFA304ECFFFC817B3427F95EFE62F913" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[333,446,1672,1698]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="bergeri" status="sp. nov.">
<emphasis id="1AD7A37DFFFC817B3427F95EFE62F913" box="[333,446,1672,1698]" italics="true" pageId="31" pageNumber="232">A. bergeri</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFC817B34ADF95EFDFEF913" bold="true" box="[455,546,1672,1698]" pageId="31" pageNumber="232">
<taxonomicNameLabel id="01E41E06FFFC817B34ADF95EFDFEF913" box="[455,546,1672,1698]" pageId="31" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
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in
<taxonomicName id="EFA304ECFFFC817B37B5F95EFC9EF913" baseAuthorityName="Karabag" baseAuthorityYear="1952" box="[735,834,1672,1698]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="1AD7A37DFFFC817B37B5F95EFC9EF913" box="[735,834,1672,1698]" italics="true" pageId="31" pageNumber="232">A. davisi</emphasis>
</taxonomicName>
). The length of hind femur generally slightly shorter than
<taxonomicName id="EFA304ECFFFC817B35BAF97AFE89F977" authorityName="Ramme" authorityYear="1939" box="[208,341,1708,1734]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFFC817B35BAF97AFE89F977" box="[208,341,1708,1734]" italics="true" pageId="31" pageNumber="232">A. schwarzi</emphasis>
</taxonomicName>
(
<tableCitation id="65214AD4FFFC817B340FF97AFE61F977" box="[357,445,1708,1734]" captionStart="TABLE 3" captionStartId="11.[152,244,495,521]" captionText="TABLE 3. Means and standard deviations for 16 morphometric characters measured from males representing 12 populations belonging to Anadolua genus." pageId="31" pageNumber="232">Table 3</tableCitation>
), and ratio hind femur/cercus ranges between 12.5 and
<quantity id="EF5BD28AFFFC817B3151F97AFB7DF976" box="[1083,1185,1708,1735]" metricMagnitude="-2" metricUnit="m" metricValue="1.44" pageId="31" pageNumber="232" unit="mm" value="14.4">14.4 mm</quantity>
(
<quantity id="EF5BD28AFFFC817B31DBF97AFAD6F976" box="[1201,1290,1708,1735]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="31" pageNumber="232" unit="mm" value="5.2">5.2 mm</quantity>
in
<taxonomicName id="EFA304ECFFFC817B305BF97AFA49F977" baseAuthorityName="Karabag" baseAuthorityYear="1952" box="[1329,1429,1708,1734]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="1AD7A37DFFFC817B305BF97AFA49F977" box="[1329,1429,1708,1734]" italics="true" pageId="31" pageNumber="232">A. davisi</emphasis>
</taxonomicName>
, 9.212.0 mm in
<taxonomicName id="EFA304ECFFFC817B343AF906FE0FF95B" authorityName="Ramme" authorityYear="1939" box="[336,467,1744,1770]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFFC817B343AF906FE0FF95B" box="[336,467,1744,1770]" italics="true" pageId="31" pageNumber="232">A. schwarzi</emphasis>
</taxonomicName>
, and 7.0
<quantity id="EF5BD28AFFFC817B3728F906FD4BF95A" box="[578,663,1744,1771]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="31" pageNumber="232" unit="mm" value="8.2">8.2 mm</quantity>
in
<taxonomicName id="EFA304ECFFFC817B37D0F906FCF6F95B" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[698,810,1744,1770]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="bergeri" status="sp. nov.">
<emphasis id="1AD7A37DFFFC817B37D0F906FCF6F95B" box="[698,810,1744,1770]" italics="true" pageId="31" pageNumber="232">A. bergeri</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFC817B3658F906FC57F95B" bold="true" box="[818,907,1744,1770]" pageId="31" pageNumber="232">
<taxonomicNameLabel id="01E41E06FFFC817B3658F906FC57F95B" box="[818,907,1744,1770]" pageId="31" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). The phrase content of
<taxonomicName id="EFA304ECFFFC817B31F6F906FB31F95B" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[1180,1261,1744,1770]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="moli" status="sp. nov.">
<emphasis id="1AD7A37DFFFC817B31F6F906FB31F95B" box="[1180,1261,1744,1770]" italics="true" pageId="31" pageNumber="232">A. moli</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFC817B319EF906FA92F95B" bold="true" box="[1268,1358,1744,1770]" pageId="31" pageNumber="232">
<taxonomicNameLabel id="01E41E06FFFC817B319EF906FA92F95B" box="[1268,1358,1744,1770]" pageId="31" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="EFA304ECFFFC817B30EEF906FF27F8BF" authorityName="Ramme" authorityYear="1939" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFFC817B30EEF906FF27F8BF" italics="true" pageId="31" pageNumber="232">A. schwarzi</emphasis>
</taxonomicName>
are very similar differing from that of
<taxonomicName id="EFA304ECFFFC817B37C5F922FCC3F8BF" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[687,799,1780,1806]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="bergeri" status="sp. nov.">
<emphasis id="1AD7A37DFFFC817B37C5F922FCC3F8BF" box="[687,799,1780,1806]" italics="true" pageId="31" pageNumber="232">A. bergeri</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFC817B364CF922FC5CF8BF" bold="true" box="[806,896,1780,1806]" pageId="31" pageNumber="232">
<taxonomicNameLabel id="01E41E06FFFC817B364CF922FC5CF8BF" box="[806,896,1780,1806]" pageId="31" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, whereas, the phrase of
<taxonomicName id="EFA304ECFFFC817B31FAF922FB3DF8BF" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[1168,1249,1780,1806]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="moli" status="sp. nov.">
<emphasis id="1AD7A37DFFFC817B31FAF922FB3DF8BF" box="[1168,1249,1780,1806]" italics="true" pageId="31" pageNumber="232">A. moli</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFFC817B3182F922FA9EF8BF" bold="true" box="[1256,1346,1780,1806]" pageId="31" pageNumber="232">
<taxonomicNameLabel id="01E41E06FFFC817B3182F922FA9EF8BF" box="[1256,1346,1780,1806]" pageId="31" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with its double syllables group series differs from
<taxonomicName id="EFA304ECFFFC817B3719F8CEFD2BF883" authorityName="Ramme" authorityYear="1939" box="[627,759,1816,1842]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFFC817B3719F8CEFD2BF883" box="[627,759,1816,1842]" italics="true" pageId="31" pageNumber="232">A. schwarzi</emphasis>
</taxonomicName>
which has triple syllables group series. There are additional morphological and bioacoustical diagnostic characters in the keys as well.
</paragraph>
<paragraph id="281C7F6FFFFC817B35ADF8B6FEE7F857" blockId="31.[151,1437,1456,2022]" pageId="31" pageNumber="232">
<emphasis id="1AD7A37DFFFC817B35ADF8B6FE97F8CB" box="[199,331,1888,1914]" italics="true" pageId="31" pageNumber="232">Coloration:</emphasis>
The intra-population colour variation of this species, of which only one population is known, almost completely coincides with the colour scale of nine different populations of
<taxonomicName id="EFA304ECFFFC817B3137F852FB3FF82F" authorityName="Ramme" authorityYear="1939" box="[1117,1251,1924,1950]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFFC817B3137F852FB3FF82F" box="[1117,1251,1924,1950]" italics="true" pageId="31" pageNumber="232">A. schwarzi</emphasis>
</taxonomicName>
(Apppendix 8).
<emphasis id="1AD7A37DFFFC817B35FDF87FFE5DF873" box="[151,385,1960,1986]" italics="true" pageId="31" pageNumber="232">Measurements (mm).</emphasis>
<typeStatus id="F718C1CDFFFC817B34ECF87EFE2FF873" box="[390,499,1960,1986]" pageId="31" pageNumber="232" type="holotype">Holotype</typeStatus>
: Measurements (mm): Body length: 22.0, Pronotum: 6.32, Cercus: 1.28, Hind femur: 18.0 (
<tableCitation id="65214AD4FFFC817B35BDF81AFEF1F857" box="[215,301,1996,2022]" captionStart="TABLE 3" captionStartId="11.[152,244,495,521]" captionText="TABLE 3. Means and standard deviations for 16 morphometric characters measured from males representing 12 populations belonging to Anadolua genus." pageId="31" pageNumber="232">Table 3</tableCitation>
).
</paragraph>
<paragraph id="281C7F6FFFC3814435ADFF41FE8CFF64" blockId="32.[151,1437,151,1149]" pageId="32" pageNumber="233">
<emphasis id="1AD7A37DFFC3814435ADFF41FE99FF00" box="[199,325,151,177]" italics="true" pageId="32" pageNumber="233">Etymology.</emphasis>
Named after Turkish Orthopterist Abbas Mol, who has done outstanding works on the
<taxonomicName id="EFA304ECFFC381443049FF41FA40FF00" box="[1315,1436,151,177]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Orthoptera" pageId="32" pageNumber="233" phylum="Tracheophyta" rank="order">Orthoptera</taxonomicName>
fauna of
<collectingCountry id="50B43FFFFFC381443590FF6DFE90FF64" box="[250,332,187,213]" name="Turkey" pageId="32" pageNumber="233">Turkey</collectingCountry>
.
</paragraph>
<paragraph id="281C7F6FFFC3814435ADFF09FB55FE38" blockId="32.[151,1437,151,1149]" pageId="32" pageNumber="233">
<emphasis id="1AD7A37DFFC3814435ADFF09FEEDFF48" box="[199,305,223,249]" italics="true" pageId="32" pageNumber="233">Remarks.</emphasis>
The distribution of the population is close to
<taxonomicName id="EFA304ECFFC38144365AFF09FC79FF49" authorityName="Karabag" authorityYear="1952" box="[816,933,223,249]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="rammei">
<emphasis id="1AD7A37DFFC38144365AFF09FC79FF49" box="[816,933,223,249]" italics="true" pageId="32" pageNumber="233">A. rammei</emphasis>
</taxonomicName>
which was synonymized to
<taxonomicName id="EFA304ECFFC381443189FF09FAE0FF48" authorityName="Karabag" authorityYear="1958" box="[1251,1340,223,249]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="burri">
<emphasis id="1AD7A37DFFC381443189FF09FAE0FF48" box="[1251,1340,223,249]" italics="true" pageId="32" pageNumber="233">A. burri</emphasis>
</taxonomicName>
by
<bibRefCitation id="4C32029EFFC38144300CFF36FF39FEAC" author="Unal, M." pageId="32" pageNumber="233" pagination="1 - 66" refId="ref27801" refString="Unal, M. (2018) Tettigoniinae (Orthoptera: Tettigoniidae) from Turkey with key to genera and descriptions of six new species. Zootaxa, 4432 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4432.1.1" type="journal article" year="2018">Ünal (2018)</bibRefCitation>
.
<taxonomicName id="EFA304ECFFC381443598FED5FEB6FEAD" authorityName="Karabag" authorityYear="1952" box="[242,362,259,285]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="rammei">
<emphasis id="1AD7A37DFFC381443598FED5FEB6FEAD" box="[242,362,259,285]" italics="true" pageId="32" pageNumber="233">A. rammei</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="4C32029EFFC38144372EFED5FD24FEAC" author="Karabag, T." box="[580,760,258,285]" pageId="32" pageNumber="233" pagination="139 - 143" refId="ref25717" refString="Karabag, T. (1952) Anadolua Ramme (Orthoptera, Tettigoniidae) genusunun revizyonu. Istanbul Universitesi Fen Fakultesi Mecmuasi, 17 (2), 139 - 143. [in Turkish]" type="journal article" year="1952">Karabağ (1952)</bibRefCitation>
based on the
<typeStatus id="F718C1CDFFC3814436F7FED5FC21FEAC" box="[925,1021,259,285]" pageId="32" pageNumber="233" type="paratype">paratype</typeStatus>
of
<emphasis id="1AD7A37DFFC381443143FED5FB69FEAC" box="[1065,1205,259,285]" italics="true" pageId="32" pageNumber="233">
<taxonomicName id="EFA304ECFFC381443143FED5FB6DFEAC" authorityName="Ramme" authorityYear="1939" box="[1065,1201,259,285]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="schwarzi">A. schwarzi</taxonomicName>
,
</emphasis>
which was a single female. The female characters mentioned by
<bibRefCitation id="4C32029EFFC3814437E1FEF1FCE7FEF0" author="Karabag, T." box="[651,827,294,321]" pageId="32" pageNumber="233" pagination="139 - 143" refId="ref25717" refString="Karabag, T. (1952) Anadolua Ramme (Orthoptera, Tettigoniidae) genusunun revizyonu. Istanbul Universitesi Fen Fakultesi Mecmuasi, 17 (2), 139 - 143. [in Turkish]" type="journal article" year="1952">Karabağ (1952)</bibRefCitation>
appear to be variational when we have examined large populations of the genus in our study. The synonym process made by
<bibRefCitation id="4C32029EFFC3814436F5FE9AFBFAFED4" author="Unal, M." box="[927,1062,331,359]" pageId="32" pageNumber="233" pagination="1 - 66" refId="ref27801" refString="Unal, M. (2018) Tettigoniinae (Orthoptera: Tettigoniidae) from Turkey with key to genera and descriptions of six new species. Zootaxa, 4432 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4432.1.1" type="journal article" year="2018">Ünal (2018)</bibRefCitation>
seems correct.
<taxonomicName id="EFA304ECFFC3814431B9FE9DFAF8FED4" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[1235,1316,331,357]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="moli" status="sp. nov.">
<emphasis id="1AD7A37DFFC3814431B9FE9DFAF8FED4" box="[1235,1316,331,357]" italics="true" pageId="32" pageNumber="233">A. moli</emphasis>
</taxonomicName>
<emphasis id="1AD7A37DFFC381443040FE9DFA5FFED4" bold="true" box="[1322,1411,331,357]" pageId="32" pageNumber="233">
<taxonomicNameLabel id="01E41E06FFC381443040FE9DFA5FFED4" box="[1322,1411,331,357]" pageId="32" pageNumber="233" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is clearly separated from other species in the genus both morphologically and bioacustically.
</paragraph>
<paragraph id="281C7F6FFFC3814435ADFE45FAD4FDA8" blockId="32.[151,1437,151,1149]" pageId="32" pageNumber="233">
<emphasis id="1AD7A37DFFC3814435ADFE45FE61FE1C" box="[199,445,403,429]" italics="true" pageId="32" pageNumber="233">IUCN status proposal</emphasis>
. This species has not been assessed for the IUCN Red List (2023-1). The distribution of the
<taxonomicName id="EFA304ECFFC3814435AAFE61FECCFE60" authorityName="Şirin &amp; Taylan &amp; Sevgili &amp; Akyildiz &amp; Güngör" authorityYear="2024" box="[192,272,439,465]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="moli">
<emphasis id="1AD7A37DFFC3814435AAFE61FECCFE60" box="[192,272,439,465]" italics="true" pageId="32" pageNumber="233">A. moli</emphasis>
</taxonomicName>
is restricted and its habitats are strongly under the threat of anthropogenic effects such as camping areas, picnic areas, hiking routes, and overgrazing. The species should be considered of a
<emphasis id="1AD7A37DFFC381443139FE0DFA93FE44" box="[1107,1359,475,501]" italics="true" pageId="32" pageNumber="233">Critically Endangered</emphasis>
(B1ab (iii+v)) status on the basis of the extent of occurrence criteria of IUCN (https://www.iucnredlist.org/).
</paragraph>
<paragraph id="281C7F6FFFC3814435ADFDF5FDCCFBCC" blockId="32.[151,1437,151,1149]" pageId="32" pageNumber="233">
<emphasis id="1AD7A37DFFC3814435ADFDF5FE70FD8D" box="[199,428,547,573]" italics="true" pageId="32" pageNumber="233">Description of song:</emphasis>
Twenty calling song recordings of
<specimenCount id="3EA5B4E6FFC381443658FDF5FC6DFD8C" box="[818,945,547,573]" count="3" pageId="32" pageNumber="233" type="male">three males</specimenCount>
have been examined. The male calling song consists of quite variable
<emphasis id="1AD7A37DFFC3814434D3FD91FDCCFDD0" box="[441,528,583,609]" italics="true" pageId="32" pageNumber="233">phrases</emphasis>
in duration, just as in
<taxonomicName id="EFA304ECFFC38144367AFD91FC48FDD0" authorityName="Ramme" authorityYear="1939" box="[784,916,583,609]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFC38144367AFD91FC48FDD0" box="[784,916,583,609]" italics="true" pageId="32" pageNumber="233">A. schwarzi</emphasis>
</taxonomicName>
species (
<figureCitation id="B09863EAFFC381443697FD91FB5AFDD0" box="[1021,1158,583,609]" captionStart="FIGURE 21" captionStartId="31.[151,250,1278,1302]" captionTargetBox="[161,1418,540,1239]" captionTargetId="figure-196@31.[151,1436,522,1251]" captionTargetPageId="31" captionText="FIGURE 21. Distribution map (A) and male calling song of Anadolua moli sp. nov. (B—two consecutive complete phrase, C—a group of syllables series and D—three syllables, black arrow is the hypothetic opening hemisyllable movement and grey arrow is the hypothetic closing hemisyllable movement)." figureDoi="http://doi.org/10.5281/zenodo.14022312" httpUri="https://zenodo.org/record/14022312/files/figure.png" pageId="32" pageNumber="233">Fig. 21B, C</figureCitation>
). The phrase structure is similar to the
<taxonomicName id="EFA304ECFFC38144345FFDBDFE65FD34" authorityName="Ramme" authorityYear="1939" box="[309,441,619,645]" class="Insecta" family="Tettigoniidae" genus="Anadolua" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="1AD7A37DFFC38144345FFDBDFE65FD34" box="[309,441,619,645]" italics="true" pageId="32" pageNumber="233">A. schwarzi</emphasis>
</taxonomicName>
species song
<typeStatus id="F718C1CDFFC38144373CFDBDFD55FD34" box="[598,649,619,645]" pageId="32" pageNumber="233">type</typeStatus>
, with some differences, and consists of double syllables group series. Phrase duration varies between 3.12 s and 17.21 s (6.63 ± 3.14 s). Each phrase contains variable number (31167) double syllables group. The phrase usually starts with a low amplitude syllable group series ranging from 3 to 7 and continues and lasts with a similar amplitude syllable group series. In a double syllables group, syllable durations are quite similar, and the amplitude of the calling song varies and the first syllable with almost low amplitude according to second syllable. Oscillographic analyses show that the period durations of the double syllables group vary between 87.7 and 164.2 ms (104.1 ± 13.5). The durations of the first and second syllables in the double syllables group vary between 23.247.4 ms (28.5 ± 4.64) and 28.543.0 ms (32.5 ± 3.52), respectively. Each syllable shows a standard structure, formulated as a short and quite indistinct soft opening hemisyllable + loud closing hemisyllable (
<figureCitation id="B09863EAFFC3814435F5FC79FED5FC78" box="[159,265,943,969]" captionStart="FIGURE 21" captionStartId="31.[151,250,1278,1302]" captionTargetBox="[161,1418,540,1239]" captionTargetId="figure-196@31.[151,1436,522,1251]" captionTargetPageId="31" captionText="FIGURE 21. Distribution map (A) and male calling song of Anadolua moli sp. nov. (B—two consecutive complete phrase, C—a group of syllables series and D—three syllables, black arrow is the hypothetic opening hemisyllable movement and grey arrow is the hypothetic closing hemisyllable movement)." figureDoi="http://doi.org/10.5281/zenodo.14022312" httpUri="https://zenodo.org/record/14022312/files/figure.png" pageId="32" pageNumber="233">Fig. 21D</figureCitation>
). Each closing hemisyllable begins with a low amplitude, reaches the highest level at approximately the midpoint, and lasts by decreasing in amplitude after this point. The spectral analysis shows that Delta and Peak frequencies of the
<emphasis id="1AD7A37DFFC381443401FC21FD30FBA1" box="[363,748,1015,1041]" italics="true" pageId="32" pageNumber="233">first low amplitude syllable group</emphasis>
vary between 23.96 and 41.89 kHz (30.43 ± 4.73) for Delta and between 18.75 and 31.50 kHz (25.24 ± 3.98) for Peak. Delta and Peak frequencies of the
<emphasis id="1AD7A37DFFC3814431CDFBCDFA40FB85" box="[1191,1436,1051,1077]" italics="true" pageId="32" pageNumber="233">Phrase syllable group</emphasis>
(except first one) vary between 27.67 and 42.73 kHz (37.05 ± 4.25) for Delta and between 18.38 and 35.63 kHz (25.27 ± 4.13) for Peak (
<tableCitation id="65214AD4FFC3814434C1FBB5FDDDFBCC" box="[427,513,1123,1149]" captionStart="TABLE 7" captionStartId="18.[152,244,1469,1493]" captionText="TABLE 7. Spectral parameters belong to male calling songs of Anadolua shcwarzi, A. moli sp. nov. and A. bergeri sp. nov. populations." pageId="32" pageNumber="233">Table 7</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>